Diversity in the Weapons of Sexual Selection: Horn Evolution in the Beetle Genus Onthophagus (Coleoptera: Scarabaeidae)
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Emlen, Douglas J., Marangelo, Jennifer, Ball, Bernard, and Cunningham, Clifford W. 2005. "<a href="https://repository.si.edu/handle/10088/3859">Diversity in the Weapons of Sexual Selection: Horn Evolution in the Beetle Genus Onthophagus (Coleoptera: Scarabaeidae)</a>." <em>Evolution</em>, (5) 1060–1084.