Systematics of the Marmoratus Group of the Frog Genus Leptodactylus (Amphibia, Leptodactylidae)
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Heyer, W. Ronald. 1973. "<a href="https://repository.si.edu/handle/10088/1902">Systematics of the Marmoratus Group of the Frog Genus Leptodactylus (Amphibia, Leptodactylidae)</a>." <em>Contributions in Science</em>, (251) 1–50.