Studies on the genus Leptodactylus (Amphibia:Leptodactylidae). 2. Diagnosis and distribution of the Leptodactylus of Costa Rica. Revta
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Heyer, W. Ronald. 1968. "<a href="https://repository.si.edu/handle/10088/2826">Studies on the genus Leptodactylus (Amphibia:Leptodactylidae). 2. Diagnosis and distribution of the Leptodactylus of Costa Rica. Revta</a>." <em>Revista De Biología Tropical</em>, 16 171–205,5.