NEW DIPTEEA OR TWO-WINGED FLIES IN THE UNITEDSTATES NATIONAL MUSEUM
By J. M. AldrichAssociate Curator, Division of Insects, United States Xational Museum
The following descriptions of two new genera and 26 new speciesof Diptera. with notes on little-known species and several tables ofspecies and genera, are the result of general work on the collectionin this order. Family MILICHIIDAEGenus PHOLEOMYIA HendeiPholeomyia Bilimek, Verh. Zool.-Bot. Ges. Wien, p. 903, 1867.
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Hen-del, Wien. Ent. Zeit., vol. 30, p. 40, 1911.
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Melander, Journ. N. Y., Ent.Soc, vol. 21, pp. 234-238, 1913.?Malloch, Proc. U. S. Nat. Mus., vol.46, pp. 130-134, 1913.Rhynchomilichia Hendel, Wien. Ent. Zeit., vol. 22, p. 250, 1903.PHOLEOMYIA EXPANSA. new speciea3Ia7e.?Dull black in color, the abdomen expanded and circular inoutline, silvery-white except on basal segment.Front brownish black, slightly converging toward the antennae,where it is about one-fourth the width of the head. Besides theorbital row of bristles, which extends slightly below the attach-ment of the antennae, there are two variable rows of small bristles orhairs beginning just below the lowest ocellus and converging so as tounite at the lunule. Antennae black, the third joint round; aristashort ; vibrissae about the middle of the face, close to the eyes, sepa-rated by tAvice the greatest diameter of the third antennal joint ; palpiblack ; proboscis not very long ; the labella folding back, as long asthe preceding segment. Thorax with well developed chaetotaxy:dorsocentrals 4 ; acrostichals 3 or 4 pairs behind the suture, the hind-most large; humeral 2; presutural 2; notopleural 2; supraalar 2;intraalar 2 or 3 ; scutellum with 2 pairs ; mesopleura with a clusterof about 8; sternopleura with 3. Halteres black including stem.Calypters brown with blackish rim and brownish fringe. AbdomenNo. 2555.?Proceedings U. S. National Museum, Vol. 66, Art. 18.9099? 25 1 1
2 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 66
circular, very thin and flat, the dorsal surface entirely silvery, butunder a high power showing a single row of delicate black hairsalong the hind margin of each segment. First abdominal segmentdull black, venter black, the sternites narrow, the membrane greatlydeveloped between them and the tergites, so as to cover almost thewhole. Wings sometimes milky, usually subhyaline, the veins black;the costa deeply and obliquely notched before the tip of the firstvein. Legs black.Length 4.5 mm.Described from 9 males taken by the writer on Mount Lowe, Cali-fornia, near the upper end of the electric car line, on July 3, 1917.Type-.?Male, Cat. No. 27242, U.S.N.M.Family CHLOROPIDAEGenus CHLOROPS MeigenChlorops Meigen, Illiger's Magazine, vol. 2, p. 27S, 1S03.CHLOROPS KUWANAE, new speciesMale and female.?General color light yellow, thorax with threebroad, opaque, black stripes, the inner much abbreviated behind, anda small black 'stripe above each wing. Front about half as wide ashead, dull yellow with only small and scattered hairs; frontal tri-angle shining, the sides convex, apex drawn out in a long pointwhich reaches the lunule; a roundish black spot covers all the tri-angle except the point and the basal angles, the latter j^ellow to theocelli; antennae of ordinary size, first joint yellow, second brown,third black, with a very blunt upper angle, arista white with yellowbasal joint. Face light yellow, white in male, bucca of same color,one-fifth the eye height; epistoma not much projecting. Palpi,proboscis and edge of mouth pale yellow. Occiput yellow, a largeblack spot extending downward from the vertex, widening below.Thorax shining except the dorsal stripes, with dark hairs and veryminute bristles; pleurae light yellow, with a small black spot belowthe anterior spiracle, an interrupted oblique brown one on the meso-pleura, the lower three-fifths of sternopleura light brown, sometimeswith blackish upper edge. Scutellum convex, with two pairs ofsmall bristles. Halteres pale yellow. Abdomen wholly pale yellowexcept four transverse black bands at the bases of the segments,which do not quite reach the lateral margin. Legs wholly yellowexcept the last tarsal joint of the middle and hind ones, and the lasttwo joints of the front ones, which are brown. Wings hyaline;crossveins separated by a distance equal to two-thirds of the lastsegment of the fifth vein ; third and fourth veins hardly divergent
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ART 18 DIPTERA OR TWO-WINGED FLIES ALDRICH 3the costal segment before the second vein neai"!}^ one and one-halftimes the following one.Length 2.6 mm.One male, six females, Nishigahara, Tokyo, Japan, from larvaeboring in rice stems; one vial of affected stems in alcohol, and twojDiiparia dry in vial. Two females will be returned to ProfessorKiiwana from the type lot.Type.?M?i\%, Cat No. 23927, U.S.N.M.Family TRYPETIDAEGenus ANASTREPHA SchinerAimstreplia Schinek, Novara Reise, 1868, p. 263.ANASTREPHA SCHAUSI, new speciesMale.?A reddish-yellow species with Aving pattern of 'pwrallelo..,but the month strikingly ornamented.Head yellow, third antennal joint blackened on apical third, palpiyelloAv ; the whole edge of mouth swollen and expanded, with a nar-row shining black line on the prominent part ; above this line on tliesides the color is contrasting white, but across the face the black lineis less sharply defined and above it the color is yellow like the restof the face.Thorax yellow, a white pruinose stripe on middle of mesonotum,wider behind, and a narrow whitish stripe above root of v\-ing.Pleura yellow, whitish along the suture above. JNIetanotum andhalteres yelloAv.Abdomen yellow, unmarked, the fifth segment darker and moreshining, and almost equal in length to the third and fourth com-bined. Legs yellow, the larger bristles brown.Wings with the clear area including the second basal cell con-tinued distall}^ and forward to the costa Avithout interruption; theinverted V-shaped hyaline area beyond this is not interrupted,though somewhat narrowed, at the third vein. Thus there are threeseparated areas of brown and yellow coloration, almost exactly as inLoeAv's figure of parallella.^Length, 10.6 mm.One male, Juan Vinas, Costa Hica, January 11, collected by Wil-liam Schaus and J. T. Barnes, and named in honor of the former,the distinguished lepidopterist.Typc?UalQ, Cat. No. 26837, U.S.N.M.ANASTREPHA BARNES!, new speciesFemale.?A yellow species with wing pattern like parallela, fourthvein more strongly curved forward than usual, and exceptionallylong ovipositor.
1 Mod. N. Amer. Dipt., vol. Z, 1S73, pi. 11, fig. 20.
4 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 66Head yellow, including proboscis, palpi, and antennae. Thoraxyellow, mesonotum without median stripe, but with one above rootof wing ending at suture, and one on mesopleura just below noto-pleural suture; the double inflated sclerite between halter and rootof wing is white on the mesial portion ; otherwise the whole thorax isyellow. Halteres yellow in one specimen, the knob infuscated in theother. Abdomen short and wide, wholly yellow; ovipositor as longas all the rest of the insect, the parts measuring by micrometer asfollows: Head, 18; thorax, 50; abdomen, 35 (total, 103) ; ovipositor,103 ; the ovipositor is darker brown than the body, covered with darkhair, the basal third tapers gradually while the remainder is cylin-drical. Legs yellow.Wings with yellow and brown pattern as in parallela^ but thefourth vein curves so far forward that the first posterior cell ismore nearly closed than usual (by micrometer 7 units wide at tip and12 units a little before).Length with ovipositor, 19 mm.; without, 9.5 mm.Two females, Cayuga, Guatemala (Schaus and Barnes).Ty/^e.?Female, Cat. No. 26838, U.S.N.M.Named in honor of J. T. Barnes, the companion of WilliamSchaus, and discoverer of this species.ANASTREPHA CORDATA, new speciesFemale.?A black-marked species with long ovipositor and strik-ing heavy blackish spot covering hind cross vein.Head yellow, the ocellar triangle, orbits at vertex and an in-definite occipital spot shining black: antennae yellow, palpi nar-rowly infuscated at tip. Thorax yellow, the dorsum with a pair ofinner black stripes abbreviated behind, and an outer pair inter-rupted at the suture and abbreviated in front; a transverse blackband just in front of scutellum; pleurae yellow, metanotum with aheart-shaped, shining black spot, notched with yellow in the middleabove. Halteres yellow.Abdomen yellow, the second to fifth segments with successivelynarrower basal black bands, that on the fifth interrupted. By mi-crometer the measurements are as follows : Head 16 ; thorax 34 ; ab-domen 30 (total 80) ; ovipositor 49. Thus the ovipositor is aboutfive-eighths as long as all the rest of the insect. It is yellow, morebrown apically, tapering on the basal half, and densely hairy.Wing with the pattern of uniform clear j^ellow color except theinverted V, of which one arm covers the posterior cross vein; thisV is all blackish in color, and the part covering the cross vein isexpanded and very striking. The hyaline stripe extending fromthe second basal cell to the costa is interrupted at the third vein
ART 18 DIPTEEA OR TWO-WINGED FLIES ALDRICH 5
while the tip of the inverted V is also cDimected on its basal sidealong the third vein with the yellow of the middle band. Thefourth vein curves forward at tip only a very little.Length with ovipositor 11.5 mm.; without, 6.75 mm.One female, Belize, British Honduras, collector unknown. Fromthe writer's collection, now a part of the United States NationalMuseum.Type.?F^m^lQ, Cat. No. 26839, U.S.N.M.ANASTREPHA OBSCURA, new speciesMale.?Head, thorax and abdomen yellow, a black spot in ocellartriangle and one just behind the root of the wing on the outer endof the postalar declivity. Mesonotum more reddish-yellow withpale yellow scutellum, and five more or less distinct pale longi-tudinal stripes, the outer including the humeri and notopleuralsuture. Upper edge of sternopleura and sides of mesonotum alsopale.Front with two orbitals turned back. Thoracic chaetotaxy as inserpentina Wiedemann. Abdomen unicolorous, the fifth segment notmuch longer than the fourth. Legs yellow, including the tarsi.Wings of very characteristic color, the usual undulating bandsobscured by a general brown infuscation, which leaves as subhyalineor distinctly lighter only a triangular spot on the costa beyond thefirst vein, the anal angle and two triangles on the hind margin,occupying a part of the second and third posterior cells. On exami-nation with a lens there is a rather distinct division in color betweenthe yellow-brown typical pattern and the plain brown obscuration:the outer border of the former crosses the discal cell diagonally be-yond the anterior cross vein, continuing straight on to the thirdvein and thence along it to the costa, receiving on the way a narrowstreak from behind which in its posterior part incloses the hindcross vein. The stigma and the base of the first basal cell andsome indefinite expansions from the latter are deeper brown as wellas the base of the third posterior cell. The fourth vein curves for-ward at tip as usual, and the first and third veins are setulose.Female.?The ovipositor (sixth apparent segment) is about twiceas long as the rest of the abdomen, cylindrical or slightly tapering,reddish in color.Length of male, 8.5 to 10 mm.; of female, over all, 13.5 to 14,5mm. ; of o'vipositor, 5 to 6.8 mm.Described from four males and four females, from Trinidad, WestIndies. Three were reared at Maraval, Trinidad, from larvae inLucuma multifloi'a, the tropical fruit called jacana, by W. Biithn.One pair are deposited in the British Museum.rype.?Female, Cat. No. 27246, U.S.N.M.
b PBOCEEDIXGS OF THE NATIONAL MUSEUM vol. CUANASTREPHA ORNATA, new speciesFemale. Head yellow, the third antennal joint twice the second,bordered with brown on front edge ; palpi also tinged with brown attip ; upper two frontals turned backward ; back of head with a black-ish spot on each side of the occiput.Thorax black with the following parts bright yellow: humeri; anarrow median stripe widening suddenly just before the prescutel-lars where it ends ; a lateral stripe above the root of the wing, extend-ing forward to the suture and inward a short distance on this; allthe scutellum except a basal border above ; a stripe on upper mesop-leura and a space below the root of the wing including most of theliypopleura and the side of the metanotum; propleura and regionabout front coxae ; and a stripe on the upper edge of the sternopleuraonce interrupted. There is also a reddish rather square spot on themiddle of the dorsum, divided by the median stripe ; and the sterno-pleurae are red along the median line.Abdomen blackish with wide hind borders of the segments yellow
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sixth segment (ovipositor ) much longer than preceding part ofabdomen, round, brown, haiiy. Legs entirely yellow.Wings hyaline with yellow and dark-brown pattern much like thatof serpentina, but very distinct. The base of the costa has a blackishstripe ending at the tip of the first vein. A second blackish stripebegins at the base of the main stem-vein, fills the first basal to be-yond the end of the second basal, then tapers off on the third veinand ends some distance before the anterior cross vein ; the space be-tween this and the costal stripe is largely filled with yellow. A thirdstripe begins narrowly at the outer hind corner of the second basal,follows the fifth vein (bulging behind it) more than halfway to hindcross vein, then becoming narrower runs straight in a diagonal direc-tion to the costa, including the anterior cross vein (which is itself de-cidedly oblique to correspond ) ; at the costa it widens again and runsto the apex of the wing, its widest part being where it includes thetip of the third vein. A fourth black stripe includes the anal cell,follows the anal vein to the margin, follows the margin to the pos-terior cross vein, then includes the latter and ends narrowly just infront of the fourth vein, without any trace of an arm turning backto the hind border. The basal half of the discal cell is faintly tingedwith yellow ; disregarding this we may consider that the hyaline por-tion is continuous fi^om the second basal to the costa. The obliqueposition of the anterior cross vein is a striking character. -Male.?Only the first and second abdominal segments have blackbasal border, the others are wholly yellow.Length of female, without ovijDOsitor, 6 mm ; with ovipositor 9 mm.Leno-th of male 6.5 mm.
ART 18 DTPTEEA OR TWO-WINGED FLIES ALDRICH 7Described from one female and two males, collected by F. X. Will-iams at Bancs, Oriente, Ecuador, altitude 6,000 feet. One of eachsex was captured on October 30, 1922 ; the other specimen was taken" On Luma Tree, " January 19, 1923. Received from the HawaiianSugar Planters' Experiment Station.Type.?Female, Cat. No. 27130, U.S.N.M.PHOBEMA, new genusWing like Anastrepha; ovipositor like Toxotinjpana; face withdistinct antennal grooves, separated by a broad, rounded carina,which becomes more prominent upward, projecting in a rounded knobbetween tlie antennae at their base, widely separating them. Thefront is wide and flat and protuberant, making approximately aright angle with the face ; there are two upper frentals turned back,the postverticals are present, the ocellars present but very small.Thoracic chaetotaxy: posterior dorcentral 1, far back; humeral 1;notopleural 2; presutural 1; supraalar 1; postalar 2; mesopleural 1near upper hind edge; scutellar 2 pairs.The relationship is with Anastrepha, from which the greatlyelongated ovipositor would not separate it; but the facial structureis very different.Type of the genus.?Phohema atrox, new species,PHOBEMA ATROX, new speciesFerimle.?General color brown tending toward yellow. Head darkyellow, tlie carina shining and translucent, the antennal grooves withwhitish pollen ; antennae dark yellow, reaching a little over halfwayto the epistoma ; third joint hardly twice the second, arista thin andbare. Palpi rather large and flat, yellow; probocis short, fleshy.Bucca (beloAv eye) hardly one-third eye height. Back of headsomewliat translucent.Thorax dark yellow, scutellum triangular, short, halteres withbrown knobs. Abdomen broAvn; first and second tergites unitedwithout suture, their sternites however distinct; sixth tergite veryshort, with a row of black hairs behind ; seventh abdominal segment(ovipositor) longer than the whole of the rest of the fly, round incross section, dark and curved upward at base, thence nearly straightiiud yellow, the extreme tip blackish.Wing large and long, hyaline with yellow pattern as in Ancistrepha,pseudoparallela as figured by Loew.- Fourth vein distinctly curvedforward just before reaching the margin as in the genus Anastrepha.Male.?Front not quite so protuberant; abdomen clavate, lateralborders of tergites 3?5 with large hairs slanting backward,
2 Mod. N. a. Diptera, vol. 3, pi. 11, fig. 24.
8 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 66Length of female "without ovipositor 12 mm; with ovipositor 28mm. Length of male 12 mm.Described from two females and one male, collected by F. X. Wil-liams at Banos, Oriente, Ecuador, January 19, 1923, " On LumaTree." Received from The Hawaiian Sugar Planters' ExperimentStation.Type.?Femsi]e, Cat. No. 27129, U.S.N.M.Family SAPROMYZIDAEGenus LONCHAEA FallenLoncliea Fallen, Ortalides, p. 25, 1820.?Becker, Berl. Eat. Zeitsch.,vol. 40, p. 322, 1895.?Melander, Psyche, vol. 20, p. 61, 1913.?Bezzi,Bull. Ent. Research, vol. 9, p. 250, 1918 ; vol. 11, p. 199, 1920.LONCHAEA HIRTITHORAX, new speciesMale.?Shining black. Wings and two basal joints of tarsi yellow.Front velvet black, clothed with numerous long hairs, which aremostly in four rows; width of the front above the antennae aboutone-seventh of the headwiclth; third antenal joint slightly elongated^about one-half longer than wide, hardly reaching epistoma. Para-facial with very slight gra}^ pruinosity, hardly visible except infavorable light. Palpi black, rather broad; epistoma at the sideswith numerous large hairs, manj^ of which are upturned; lunulebare. Thorax shining, covered with long, erect hair, among whichno distinct acrostichals or dorsocentrals are visible except close tothe scutellum where there appear to be two pairs of each. Scutellumshining black with two pairs of longer bristles and a marginal rowof hairs between them. Pleurae shining, the mesopleura with abun-dant long hairs largely curved forward and upward, those alongthe hind margin bristlelike. Upper edge of sternopleura with acluster of upturned large hairs. Halteres entirely black. Calyptersblack with fringe of same color. Abdomen wholly shining withrather abundant long hairs especially along the sides. Wings yellow,more infuscated apically and at the extreme base; the small cross-vein is opposite the tip of the first vein. Legs shining black exceptthe tarsi, of which the first joints are yellow, the remainder brownishor black.Female.?Front somewhat wider than in the male, with shorterhairs; the hairs of the epistoma, mesonotum and abdomen alsonoticeably shorter than in the male. Two distinct pairs of dorso-centrals with some hairlike ones anterior to them.Length. Male, 4 mm., female, 3.G mm.Described from 14 specimens reared at Forest Grove, Oregon, byL. P. Rockwood, from Lufinus polyjyhylhis.
ART 18 DIPTEEA OR TWO-WINGED FLIES ALDRICH 9Type.?M.2i\Q, Cat. No. 27243, U.S.N.M.The nearest related form is LoncJiaea aheiTans Malloch, whichis much less hairy, has a narrower front and the third antennaljoint almost circular.Mr. Rockwood writes that the larvae of this species are found inthe stems of the plant just above the ground, often in such numbersthat the plants are weakened and fall over by their own weight.Family MUSCIDAEGenus MESEMBRINA MeigenMesemirina Meigen, Syst. Beschr., vol. 5, p. 10, 1826.MESEMBRINA MAGNIFICA, new speciesFemale.?Black, the parafacials golden pollinose to the edge of themouth, sharply dividing behind from the shining black bucca;width of front 0.30 of the headwidth, much less than in 7neridiana^mystacea^ etc. Antennae slender, the arista yellow nearly to tip,plumose; palpi black. Thorax entirely black, the humeri and amedian dorsal stripe reaching the transverse suture are pale yellow,pollinose. The dorsum has only small black hair and a few bristleswhich are very delicate except those at the margins and behind.Posterior dorsocentrals 2; anterior 1, hairlike, just before the suture.Posterior acrostichals 1 ; anterior none ; humeral 3 ; posthumeral 1,very slender; prescutellar 1; notopleural 3 (the usual hind onedoubled on both sides) ; supraalar 4 or 5 (only one large) ; postalar2 (postalar declivity bare) ; sternopleural 2. Calypters deep orange.Abdomen black, first and second segments with black hair, exceptsome reddish at base of first. Third segment covered with goldenpile, its ground color tending toward reddish. Fourth segment withlonger and more erect golden pile, its ground color distinctly red.Legs black. Wings deep yellow at base, infuscated toward the tipand anal angle. Venation as in inystacea, except that the openingof the first posterior cell is before the extreme apex of the wing.Length 18 mm.Described from one female specimen in excellent condition, col-lected at Suifu, Szechuen, China, by D. C. Graham.Ty/j^.?Female, Cat. No. 27244, U.S.N.M.BALIOGLUTUM, new genu3Hypopleural bristles wanting; fourth vein curved forward, theapical cell at tip slightly more than half as wide as at its widestpart; third vein with a few distinct hairs below, none above; stemof venation not (as in Ghrysomyia, etc.) ciliated behind; facial9099?25- 2
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IDlate with a low but very sharp keel, beginning at the top, lowerpart of the plate strongly narrowed by the ridges, the vibrissaetwice the length of the second antennal joint above the mouth;facial ridges swollen below and convergent, covered with severalrows of short, spiny hairs, which extend upward in one or tworows almost to the level of the arista ; palpi rather short, flat ; pro-boscis very short, with fleshy labella, front of male nl)ove aboutone-seventh the head-width, ocellar bristles, verticals, and upperfrontals reduced to hairs, the lower frontals gradually larger, end-ing at insertion of antennae; parafrontals hairy down to samepoint. Lower part of head without bristles except about three pairsbelow vibrissae. Third antennal joint three to four times as longas second; arista with a few rays of moderate length above nearbase, and one or two below. Eyes bare.Thorax nearly bare of bristles above, of the dorsocentrals onlythe hindmost are distinct; scutellum without discal bristles; post-scutellum not developed; prostemum, pteropleura, and hypopleurabare ; sternopleura with a single bristle in the anterior upper cornerand a considerable row of mixed bristles and long hairs along theupper edge posteriorly. Abdomen entirely destitute of bristles;first sternite hairy, 2 to 5 broad and hairy ; genitalia small. Calyp-ters large, bare, the hind ones much larger and longer than theothers. Hind tibia without calcar.BAUOGLUTUM ILLINGWORTHI, new speciesMale.?Front 0.14 of the head-width at vertex, very graduallyv.idening below; parafrontals and parafacials golden poUinose, achangeable dark spot at the level of the antennal insertion; para-facials without hairs, shining; antennae reddish, the third joint in-fuscated at tip and on upper side ; palpi reddish-yellow ; bucca one-fourth the height of head. Mesonotum, scutellum, and abdomen dot-ted all over with minute darker spots on a dense gray poUinoseground, the mesonotum showing four rather distinct darker stripesin front, abdomen not at all tessellated, scutellum with shiningblack border. Pleurae subshining black. Chaetotaxy : Dorsocentrals0,1 (and a few hairs in the row) ; acrostichals 0,1 (an indistinctsmaller pair close to the prescutellars) ; humeral 3 ; posthumeral 1
;
presutural 1; supraalar 3; intraalar 1 (behind); postalar 2 or 3;scutellum with 4 marginal, 2 submarginal, no discal; prothoracica strong tuft. Calypters white. Wings glassy hyaline, veins brown.Legs black, with few bristles except the front femora, whichhave the usual two rows above and one on lower hind side. Hindtibia with short cilia on outer hind side.Female.?Front at vertex 0.24 of head-width ; no orbital nor cru-ciate bristles.
ART 18 DIPTERA OR TWO-WINGED FLIES?ALDRICH 11Length 8.4 to 10.4 mm.Three males, one female, Cairns, North Queensland, Australia,collected by A. P. Dodd and J. F. Illingworth. Two males arereturned to Dr. Illingworth, after whom I name the species.Ty'pe.?M2lQ, Cat. No. 26840, U.S.N.M.Family CALLIPHORIDAEGenus MESEMBRINELLAMesembrinclla Giglio-tos, Bull. Mus. Zool. ed Anat. Comp. R. Uciv.,Torino, vol. 7, No. 132, 1892, p. 4; Mem. R. Acad. Sci., Torino, ser. 2,vol. 45, 1895, p. 11.?Aldkich, Proc. U. S. Nat. Mus., vol. 62, art. 11,1922, p. 8.An interesting character of the genus not previously mentionedis the existence of a post-scutellum, the absence of which has beenconsidered a family character in Calliphoridae and Sarcophagidae.As here developed, however, it is much less bulging than in Dexidaeand Tachinidae.The discovery of four new species in collections submitted foridentification by Prof. A. L. Melander makes a new analytical tableof the genus desirable. For convenience it is put in the same formas the one previously published by me.ANALYTICAL TABLE OF THE GENUS MESEMBRINELLAA\ Stem- vein bare (subgenus Mesembrinella).a\ Two presutural bristles present.b\ Legs almost black, but middle and hind femora yellow on apical half.&. Wing with heavy subcostal black stripe not reaching the third vein,the posterior portion paler; 3 posterior acrostichals (BoliviaSurinam) brunnipes Surcouf.c^ Wings deep brown, the second fourth except behind yellow (Bolivia).pictipennis Aldrich.b'. Femora and tibiae yellow.c\ Apical cell very wide open, the included costal section more than halfas long as the preceding one (Costa Rica, Ecuador).umbrosa Aldrich.(;^ Apical cell less widely open, the included costal section less than halfthe preceding one.d\ Wing with only diffuse and not very strong infuscation (wide-spread neotropical) bicolor Pabricius.d\ Wing with heavy blackish subcostal stripe, beyond middle, beforethird vein (Brazil) batesi Aldrich.a'. Only one presutural present.b\ Fourth abdominal segment with a discal row of bristles.c". Femora, pleurae, and abdomen bluegreen or blackish ; 2 pairs acrosti-chals before suture.(f. Discal scutellar bristles small, almost in line with the much largerbasal lateral pair ; female with but one pair of proclinate orbitals,which are almost in the frontal row (Costa Rica).uniseta, new species.
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d^. Discal scutellar bristles but little smaller than the lateral basalpair, and forming with them a strong curve ; female with twopairs of orbitals, just outside the frontal row, which is here veryhairlike ( Peru ) cruciata Townsend.c". Femora, pleurae, and base of abdomen yellow.(f. One pair anterior acrostichals.e\ Posthumeral 1 (Panama) tibialis Aldrich.e^. Posthumeral 2 (South America) aeneiventris Wiedmann.d^. No anterior acrostichals.e^. Mesonotum viewed from behind shows three dark stripes, sepa-rating four pollinose ones (Brazil) purpurata Aldrich.e^. Viewed from beliind the pollen is not distinctly divided into 4stripes (Costa Rica) semiflava, new species.h\ Fourth abdominal segment without discals.c\ Facial ridges high and sharp, hairy to middle ; middle and hind tibiaenot at all infuscated ; sternopleurals 2, 1 facialis Aldrich.cl Facial ridges lower, not hairy except close to vibrissae.d'. With 1 or 2 pairs of anterior acrostichals.e^ Legs, pleurae, and base of abdomen largely yellow.f. Second to fourth abdominal segments with a posterior sharplydefined violet band ; third segment without marginal bristles(Brazil) cyaneicincta Surcouf.f. Second to fourth abdominal segments not banded with violet
;
third segment with row of marginals (Costa Rica).flavicrura, new species,
e*. Legs, thorax, and abdomen bluegreen or blackish ; fifth sterniteof male produced in two shining black styles (Costa Rica).spicata, new species.(Z^ Without anterior acrostichals.e^. Only one intra-alar (the posterior) ; abdominal segments 2-4with sharply defined posterior violet bands (Brazil).pauciseta Aldrich.e^ With 2 intra-alars ; abdomen not violet-banded.f. Second abdominal segment with only weak hairs along hindmargin (South America) randa Walker.f. Second abdominal segment with a distinct row of marginalbristles.g^. Middle and hind tibiae black, in male the middle ones elon-gated and with only minute bristles (South America).quadrilineata Fabricus.g*. Middle and hind tibiae not or hardly infuscated; male withthe usual bristles on middle tibiae (Brazil).dorsimacula Aldrich.A'. Stem-vein ciliated behind (subgenus Mesembolia Aldrich).a\ Greatest width of apical cell exceeding the length of the hind crossvein.b\ Apical cell moderately wide open, the included costal segment not morethan half the preceding one; no acrostichals immediately behind thesuture (Mexico to Paraguay) bellardiana Aldrich.h'. Apical cell very wide open, the included costal segment more than halfthe preceding (Brazil) fulvipes Aldrich.a: Greatest width of apical cell less than hind crossvein (Brazil).peregrina Aldrich.
ART 18 DIPTEEA OR TWO-WINGED FLIES ALDRICH 13MESEMBRINELLA SPICATA, new speciesMale.?Purplish-black in color, only the palpi, face, anteni:aeand lower part of front bright j'ellow. Front wide for a male, 0.115of the head width by micrometer (one specimen), the frontal rowscomposed of delicate hairs to the middle, below about 7 larger, thelowest just below antennal insertion ; ocellars large, proclinate, !a pairalmost as large behind the triangle; vertical only one pair. Thirdantennal joint more than three times the second ; facial ridges ratherhigh and sharp, hairy almost to the middle of the third antennaljoint; bucca one-fifth the eye height.Mesonotum not with distinct pollinose stripes. Chaetotaxy : doi-socentral 2, 3 ; acrostichal 2, 1 ; humeral 3 ; posthumeral 2 ; presutural1; notopleural 2; supraalar 3; intraalar 2: postalar 3; scutellar 2lateral, 1 large apical, 1 large discal; sternopleural 2. Pleurae con-colorous with mesonotum. Calypters transparent with black rimand conspicuous black fringe. Both thoracic spiracles large, dark.Abdomen purplish-black, with rather dense, erect, short hair, nobristles whatever. Genital segments large and conspicuous, shiningblack; inner forceps shining black, parallel and close together, nottapering, blunt at tip, the base behind united and swollen into asudden hump which is paler in color, divided into two arms back-ward toward the anus, and these arms bear a pair of black, converg-ing processes ending in tufts of black hair which touch each other onthe middle line. Outer forceps shining black, twisted, blunt. Fifthsternite narrow, shining black including its sides, with two erect,blunt, shining black processes in the place of the usual lobes.Legs blackish; middle tibia with flexor bristle; hind tibia withlong calcar just below middle.Wings subhyaline, small cross vein infuscated ; the opening of theapical cell at costa is about one-fifth the preceding costal segment.Length 8.3 mm.Described from one male. La Suiza de Turrialba, Costa Rica,Februaiy 22, 1923 (Pablo Schild). Through the kindness of ProlA. L. Melander we retain this striking unique for the UnitedState National Collection.Ty/^e.?Male, Cat. No. 26796, U.S.N.M.MESEMBRINELLA UXISETA, new speciesA blue-black species with face, antennae, palpi, and thoracic spi-racles yellow.Male.?Front rather wide, 0.13 the head width (the same in threespecimens measured by micrometer), black to antennae; no frontalsof any size above the middle ; ocellars long, proclinate, a post-ocellarpair also long; only one moderate pair of verticals. Third antennal
14 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 6Gjoint two and one-half times the second ; arista with long but sparsepiumosity; facial ridges flat, only a few hairs above vibrissae; buccayellow, about one-ninth the eye height.Mesonotum blue-black, with white pollen visible more from behind,not distinctly striped. Chaetotaxy: dorsocentral 2, 3; acrostichal2. 1; humeral 3; posthumeral 2; presutural 1; notopleural 2; supra-alar 3; intraalar 2; postalar 3; scutellum with 1 lateral; 1 apical, 1rather small discal; sternopleural 2, 1. Calypters rather dark, theanterior with black rim. Spiracles large, yellow.Abdomen blue-black, wdth a little Avhite pollen visible in certaindirections, not tessellated ; first segment with one large lateral margi-nal ; second with two laterals and sometimes a median marginal pair(21^ times in five males) ; third segment with strong marginal row of10; fourth segment with distinct discal row^ of 4 to G (usually notcontinuing down the sides), and an apical row of 6 to 8. Genitalsegments rather large, shining black, with hair but no bristles. Innerforceps black, flat and broad at base, but tapering to a slender, sharptip ; outer forceps black, narrowed at base, broader in middle, withsharp tip. Fifth sternite with ordinary deep incision in middle andtv?^o large, black, subshining, flat lobes.Legs black, middle tibia with flexor bristle, hind tibia with largecalcar below middle.Wings lightly infuscated, hind crossvein deeply so; fourth veinbeyond hind crossvein bowed a little backward so as to widen theapical cell, which includes at its tip a costal segment about one-seventh of the preceding.Female.?Front of equal width almost to antennae, 0.25 the headwidth (average of three, 0.24, 0.25, 0.27), the middle stripe red overhalfway up.Length 8 to 8.5 mm. in both sexes.Described from 6 males and 4 females, I>a Suiza de Turrialba,Costa Rica, February 22 to March 29, 1923, and September 5 andOctober 1, 1921. In Prof. L. Melander's collection.Paratypes.?M^ilQ and female. Cat. No. 26797, U.S.N.M.MESEMBRINELLA SEMIFLAVA, new speciesMale.?Front almost as wide as the narrow ocellar triangle, thenarrow parafrontals touching for some distance; frontal bristlesbeginning about the middle; ocellars long, proclinate, the post-ocellar pair about half as long ; one pair of smallish verticals. Thehead is yellow except upper two-thirds of back and upper third offront; antennae and palpi yellow; third antennal joint not muchmore than twice the second which is a litle longer than in somespecies; arista long, with long but sparse piumosity; facial ridges
AET18 DIPTEKA OR TWO-WINGED FLIES ALDRICH 15
rather fiat, only a few hairs above vibrissae; bucca very narrow,hardly one-eighth the eye height.Mesonotum, except humeri, metallic blue-green, overlaid withsome white pollen, especially anteriorly; the two outer of the usualdark stripes are rather distinct, the inner not so. Chaetotaxy:dorsocentrals 2, 3 ; acrostichals 0, 1 ; humeral 2 ; posthumeral 1
;
presutural 1; notopleural 2; supraalar 2 and a small behind; in-trallar 2; postalar 2 (the front one small); scutellura with onelateral, 1 apical, 1 discal; sternopleural 2, and a small hairlike onebelow the posterior. Pleurae and humeri yellow. Calypters lightlyinfuscated, the front one with black rim. Thoracic spiracles large,yellow.Abdomen pale yellow at base, blue and violet at tip. The firstsegment is yellow with a blackish narrow line at hind edge, extend-ing below; the second segment is yellow with a purplish hindborder, wider at middle, where it is nearly half the segment andvery narrow below; third segment yellow anteriorly at side andmore below, the rest blue, but the hind border purple; fourthsegment blue above and below with a distinct white poUinose spoteach side of the genitalia and hardly a trace of purple apically.The first segment has several lateral bristles, the second one, thethird a strong marginal row, the fourth a discal row of 8 and asmaller marginal row. Genital segments of moderate size, brownishor piceous, shining, with hairs but no bristles. Inner forceps yellow,slender, nearly straight, with sharp black tips ; outer forceps yellow,slender, strongly bowed in at tip. Fifth sternite small yellowish,cleft in middle.Legs yelloAY, middle and hind tibiae and tips of their femorablack; hind tarsi lighter than their tibiae; middle tibia withoutflexor bristle, hind tibia with long calcar a little below middle.Wings long and narrow, infuscated, more distinctly beyond tipof auxiliary vein, but with no definite pattern; apical cell openingon costa just before apex for a distance equal to one-seventh thepreceding costal segment.Female.?The front is narrowest at vertex, where it is 0.18 of thehead-width in both specimens ; cruciate bristles distinct, only 3 to 4lower frontals of any size; orbitals represented only feebly byhairs but the lower pair in one specimen a trifle stouter. Abdomenwith much less yellow, none above on the third segment, and a widerdark margin on the second.Length of male, 8.5 to 9 mm. ; of female, the same.Described from three males and two females, La Suiza de Tur-rialba, Costa Kica, February 23-28 and March 16, 1923 (PabloSchild). In Prof. P. L. Melander's collection.Paratypes.?M^ile and female, Cat. No. 26799, U.S.N.M.
16 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 66MESEMBRINELLA FLAVICRURA, new speciesBlue-black, the following parts yellow: Front except upper halfor less, antennae, face, bucca, palpi, humeri and lower prothoracicregion, thoracic spiracles, first abdominal segment in large part inthe male (less in the female), coxae, all the femora except the tips.Male.?Eyes almost contiguous, the front at narrowest onlj' aswide as the anterior ocellus ; the rows of frontal bristles begin belowthe narrow part; ocellars and post-ocellars of equal size, not large;verticals small. Facial ridges quite flat, only a few hairs abovevibrissae; bucca one-fifth the eye height.Thorax not with distinct pollinose stripes. Chetotaxy: dorsocentral 2, 3 ; acrostichal 2, 1 ; humeral 3 ; posthumeral 2 ; presutural 1
;
notopleural 2; supraalar 3; intraalar 2; postalar 3; scutellum with 2lateral, 1 apical, 1 discal. Calypters infuscated, especially the pos-terior, with dark rims. Spiracles large, yellow.Abdomen bluish-purple, the first segment yellow except a narrow-black posterior border above and below which widens suddenly atthe sides ; second segment with trace of yellow above anteriorly. Thefirst segment has one small lateral marginal, the second a large one,the third and fourth a marginal row, no discals. One male has asmall but unmistakable pair of median marginals on the secondsegment.Legs black except as indicated, middle tibia with flexor bristle,hind tibia with large calcar below middle.Wings lightly infuscated, paler toward base ; small cross vein de-cidedly infuscated; fourth vein and apical cell as in uniseta.Female.?Fmnt 0.23 of headwidth (average of three, 0.22, 0.24,0.24); two pairs of orbitals; only one vertical. First abdominalsegment varying in amount of yellow, sometimes with much less thanindicated for male.Length of males, 7 and 7.8 mm. ; of females, 8 to 8.5 mm.Described from two males and five females, La Suiza de Turrialba,Costa Rica, February 8 to July 26, 1923 (Pablo Schild). In Prof.A. L. Melander's collection.Paratypes.?lA?i\^ and female, Cat. No. 26800. U.S.N.M.Tribe CHRYSOMYIINITABLE OF GENERA OF THE WOELD
1. Hind calypters covered with hairs on upper side (Eastern Hemisphereexcept Chrysomyia desvoidyi Hough, noted below) 2Hind calypters bare except in the basal depression (Western Hemi-sphere) ^2. Vibrissae at least the length of the second antennal joint above oralmargin (type, marginalis Fabricius) Chrysomyia Robineau Desvoidy.Vibrissae at oral margin; male with broad front; small Australian species(type, varipes Macquart) Microcalliphora Townsend.
ART 18 DIPTEEA OR TWO-WINGED FLIES?ALDRICH 17
3. Palpi slender and very short ; vibrissae at least the length of the secondantennal joint above the oral margin and distinctly approximated
;
dorsocentrals only one or two pairs just before the scutellura (type,macellaria Fabricius) mesonotum striped Cochliomyia Townsend.Palpi normal, clavate 44. Vibrissae at oral margin, hardly approximated ; dorsocentrals 2, 4, small butdistinct (type, segmcntaria Fabricius) mesonotum not striped.Hemilucilia Brauer.Vibrissae at least the length of the second antennnl joint above the oralmargin 55. Without dorsocentrals except one or two pairs next to the scutellum
;
mesonotum striped (type, fulvlpes Macquart; Compsomijiops Townsend,same type) Paralucilia Brauer and Bergeustamm.Dorsocentrals 2, 4, small but distinct (tyi>e, semivirklis Van der Wulp)mesonotum not striped Chloroprocta Van der Wulp.Neopollenia Brauer, Neocalliphora Brauer and Bergenstamm, andParacalliphora Townsend, all from the oriental and Australian
' regions, whicli were placed in Chrysomyiini in the National Museumcollection by Townsend, have bare stem vein and I would refer themto the tribe Calliphonni^ with which head structure also agrees.Malloch in a recent paper ^ has gone still further in this direction,expressing the opinion that these three genera are at most only sub-genera of Callijyhora.Genus COCHLIOMYIA TownsendCochliomi/ia Townsend, Journ. Wash. Acad. Sci., vol. 5, 1915, p. 646.
?
Shannon, Insecutor Ins. Menst, vol. 11, 1923, p. 106.fCallitroga " Schiner MS " Bkauer, Denkschriften Kais. Akad., vol. 47,1883, p. 74.?Johnson, Bull. Amer. Mus. Nat. Hist., vol. 41, 1919, p. 439.Townsend considered that Schiner's manuscript name, being
"cited in synonymy," had no standing. Johnson asserted that" Brauer and Bergenstamm had a perfect right to adopt CallitrogaSchiner MS." They did not adopt it, but they merely mentioned itin an ambiguous way as a collection name of Schiner's, apparentlyconnecting it first and most clearly with Lucilia JioniinivoraxCoquerel. This is supposed to be a synonym of macellaria^ but maybe different, and other related species are gradually coming to light.Even admitting the validity of the name for Jiominivorax^ I doubtthe advisability of using it as if macella?^ were its type.The species described below may be separated from the commonand widespread North and South American macellaria by the fol-lowing characters:o\ Thorax evidently metallic blue or green, with four white pollinose stripes,the inner not continued on scutellum ; abdomen almost wholly shiningabove, or with thin and uniform pruinosity, the fourth segment with awhite pollinose spot on each side widely separated__macellaria Fabricius.
? Trans. New Zealand Inst., vol. 55, p. 640, 1924.9099?25 3
18 PROCEEDINGS OF THE NATIONAL MUSEUM vol.66
a'. Thorax black with only slight traces of metallic color, the four whitepoUinose stripes very distinct and the inner pair continued on the scutel-lum ; abdomen metallic green on the second and third segments with
"broad interrupted fascia of silvery pollen (on the third in some lightsbreaking up into four spots partly connected in pairs), fourth segment
"with a pair of silvery spots close together, elsewhere more coppery thanpreceding segments laniaria Wiedemann.COCHLIOMYIA LANIARIA (Wiedemann)Mnsca laniaria WiieajEMANN, Auss. Zweifl., vol. 2, 1830, p. 40G taniaria',corrected, p. 683).Co^mpsomyia laniaria Enrique Lynch A.. Anales Soc. Cient. Argentina,vol. 7, 1879, p. 256; vol. 10, 1880, p. 75 {taniaria, corrected, vol. 10, p. 249).Cited but not identified.Male.?Front as wide as ocellar triangle, quite black near vertex,gradually covered with white pollen below, with numerous smallwhite hairs which continue close to the eye as far down as the middleof the third antennal joint ; bucca two-fifths the eye height, translu-cent yellow and shining except before and behind; back of headblack to the proboscis; antennae, palpi, proboscis and facial struc-ture as in '??iaceUaria, except that the vibrissae are a little nearer theepistoma. Pleurae black. Hind ealypter brown on disk with whiterim, a few pale hairs in the concavity close to base. Postalar decliv-ity with tuft of long hairs on its center as in macellai'ia (above baseof front ealypter). Genitalia on same plan as in macellaria but theinner and outer forceps notably longer, and the former more slender
;
the penis at apex also more drawn out. Legs entirely black. Wingsas in macellaria.Female.?Front 0.31 of head width (average of three, 0.29, 0.30and 0.33) ; parafrontals shining black above almost to middle; backof head yellow below changing abruptly to black just below neck.Fifth abdominal segment conical, polished, metallic. Otherwise asin male.Length 5.5 to 7 mm., averaging distinctly smaller than macellaria.Redescribed from one male and three females, Key West, Florida,January 31, February 1 and C, 1869 (labels in handwriting; collectordoubtful, perhaps Burgess) ; one female evidently collected manyyears ago with only the label " Fla." In addition to this old mate-rial, the United States National Museum has lately received 65 speci-mens of both sexes in alcohol (now pinned) from Dr. Paul Bartsch,curator of moUusks, United States National Museum, which he col-lected at one time on San Salvador Island, Bahamas, in the summerof 1923, on dead moUusks.One Key West female bears the label " Chrysomyia certima AVlk.,"in Coquillett's handwriting. This would appear from Walker'sdescription to be a mistake, as certima is quite certainly a synonym
ART 18 DIPTERA OR TWO-WINGED FLIES?ALDRICH 19
of macellaria (not of Paralucilia comicma Fabricius, as suggestedin my Catalogue, 1905, p. 516).Wiedemann's allusion to this form by name occurs in a note fol-lowing his description of macellaria Fabricius. Discussing thevariations of the species, h6 says that those specimens having thewhite dorsal abdominal pollinose spots or interrupted cross-bands inthe collection of Count Hoffmansegg were labelled Blusca laniaria,adding that they are not otherwise materially different from macel-laria. Although in his description, based on the Fabrician type ortypes and additional material of his own, he had described thesefasciae as if i\\Qj were typical of maceUarla, his note seems to makeit clear that laniaria differs from macellaria in possessing them.Genus CHRYSOMYIA Robineau-DesvoidyChrysomyia Robineau-Desvoidy, Myiodaries, 1830, p. 444.
?
Townsend,Journ. Wash. Acad. Sci., vol. 5, 101.5, p. 646.CHRYSOMYIA DESVOIDYI HoughChrysomyia desvoidyi Hough, Kans. Univ. Quart., vol. 9, 1900, p. 203.This was described from Brazilian specimens. As far as knownit is the only American species with the hind calypter hairy, a char-acter fortunately mentioned by Hough, It must be admitted how-ever that the species is somewhat intermediate in this regard, since aconsiderable area of the lateral apical portion is bare.Eyes of male almost contiguous for a considerable distance, sepa-rated only by the width of the front ocellars. The female has thefront slightly narrowed just above the antennae, where it is aboutone-fourth as wide as the head (0.26 in each of tv/o measured bymicrometer). Halfway between the posterior ocelli and the eye, onthe vertex of the female, there is one distinct bristle curved back-ward and laterally. The ocellar bristles in the female are close tothe anterior ocellars and are directed straight to the side, oppositeto each other. In the male, however, the ocellars are parallel andproclinate. Thorax in both sexes with much less distinct stripesthan in the genera Paralucilia and Cochlioinyia, In all the speci-mens seen the body color is deep blue-green. The hind calypter isdark brown, with white rim only in two females. The second andthird abdominal segments have each a black band on the hind mar-gin. Additional characters are given in the original description.Six males and ten females ; Quebrada Secca, Venezuela ; Valera,Venezuela (Dr, C, Uribe) ; Cano Saddle, Canal Zone, Panama(Shannon) ; Erwin Island, Canal Zone, Panama (Shannon) ; Trini-dad Rio, Panama (Busck) ; Las Cascadas, Canal Zone, Panama(A, H, Jennings) ; San Carlos. Costa Rica (Schild and Burgdorf )
;
Cordoba, Mexico (Knab).
20 PROCEEDINGS OF THE NATIONAL, MUSEUM vol. 66.Genus MICROCALLIPHORA TewnsendMicrocalliphora Townsend, Proc. U. S. Nat. Mus., vol. 49, 1916, p. 618.The type species designated by Townsend was described as Luciliavaripes by Macquart,* It may be distinguished from the new specieshere described by the following characters. Both species areAustralian.
a' All the femora mainly black, tifeiae largely so; front in both sexes blackalmost to the lunule ; male with dense erect white hairs on upper side offront femur, which are shorter toward apes; front femora yellow on.anterior side varipes Mac-quart.a" Male only; femora and tibiae entirely yellow; front black from vertex onlyas far as the upper third, then abruptly changing to pure yellow ; front femurwith only the usual black bristles flavifrons, new species.MICROCALLIPHORA FLAVIFRONS, new speciesMale.?Front broad, narrowest just at the level of the lunule,where it is by micrometer 0.33 in one, 0.32 in the other, of theheadwidth (two males of varipes measure 0.33 and 0.35, and inthem the front is not narrowed below). Parafrontals shining greenat vertex and as far forward as the tip of the ocellar triangle, thenchanging suddenly to a pure light yellow, which color extends down-ward and covers the whole buccal region; frontal stripe a littledarker yellow, wider than either parafrontal, blackish around theocellar triangle; frontal bristles small, reaching as far as middle ofsecond antennal joint ; on the upper metallic part of the parafrontaleach side are two distinct orbitals, which are reclinate and divaricatCythe upper farther from the eye ; two large verticals ; the yellow partof the parafrontals bears small white hairs, which extend downon the parafacials as far as the middle of the third antennal jointbut are almost imperceptible. Antennae yellow, the third jointbroadly infuscated from the arista, five times as long as second joint.Arista rather short, the plumosity consisting of only a few long raysabove and about three more appressed below ; penultimate joint short.Vibrissae large and distinct, black, no black hairs above them, butthe ridges rather thick and well covered with small pale hairs.Bucca one-third the eye height. Palpi yellow, clavate, of averagesize. Proboscis small. Mesonotum shining green, without stripes,with a very delicate pale pruinosity; dorsocentrals 3 anterior, 4posterior; acrostichals, 1 just before suture and 1 prescutellar
;
sternopleurals 2; postalar declivity with several long bristly hairsin middle; lower lateral prong of scutellum (above base of calypters)bare; intraalar one large before the suture, two behind. Calyptersyellowish. Abdomen bright green, first segment black and a black
* Dipteres Exetiques, Suppl., vol. 4, 1851, p. 222,
ART 18 DIPTEKA OR TWO-WINGED FLIES ALDEICH 21posterior margin on second and third segments, that on formerwidened in middle. Genital segments rather small, shining green.Fifth sternite as usual with deep V-shaped cleft. Legs includingcoxae yellow, the tarsi only becoming brown near tip. Claws andpulvilli are small.Wings with evident brown tinge, less so posteriorly; third veinhairy almost to the crossvein. Third and fifth costal segments equal
;
fourth vein with short, rectangular bend near margin of wing, theapical crossvein deeply concave, ending almost in the apex.Length, 4 mm.Described from two males collected by Dr. J. F. Illingworth atGordonvale, North Queensland, Australia, in 1919. One is labeled
*? Ex carrion."r?//>e.?Male, Cat. No. 26841, U.S.N.M.Family SARCOPHAGIDAEGenus NOTOCHAETA AldrichXotochaeta Aldrich, Sarcophaga and Allies, 1916, p. 52.Front moderately narrow in male, not protruding at antennae;parafrontals and parafacials with only a few almost imperceptibleminute hairs; vibrissae at edge of mouth, not approximated; facialridges bare except close to the vibrissae; second antennal joint shortythird three times as long, almost reaching vibrissae ; arista with longplumosity to tip or nearly to it ; epistoma not produced, face a littlereceding; palpi and proboscis normal; back of head flattened.Thorax distinctly striped, with no acrostichals except a small pairbefore scutellum; dorsocentrals 2 anterior, 2 or 3 posterior; pre-sutural 1, notopleural 2, postalar declivity bare. Postscutellum notdeveloped; calypters bare. Abdomen without discals; no medianmarginals on first and second segments, third and fourth with strongerect rows.Wings as in Sarcophaga; first vein bare, third bristly nearly tocrossvein.The foregoing characters are taken from the type species, suh-polita Aldrich. KEY TO SPECIES OF NOTOCHAETA
1. "With two postsutiiral dorsocentrals 2With three postsutural dorsocentrals 42. Facial ridges somewhat prominent, with small hairs extending above thelevel of the middle of the third antennal joint plumigera Van der WulpFacial ridges more flattened, bare except close to vibrissae 33. With small but distinct prescutellars ; scutellum with indistinctly definedcontinuation of median dark thoracic stripe ; male with dense, suberecthairs on flexor surface of middle tibia subpolita Aldrich.
22 PROCEEDINGS OF THE NATIONAL MUSEUM vol.66With no prescutellars ; scutellum with greenish-black disk, bordered uni-formly with yellow pollen ; male with only appressed hair on flexorsurface of middle tibia townsendi, new species.4. Abdomen shining blue in color comata, new species.Abdomen black with golden pollinose pattern angusta, new species.NOTOCHAETA COMATA, new speciesMale.?Front 0.12 and 0.14 of the headwidtli in the two specimens.Parafrontais and parafacials golden pollinose ; frontal bristles about10, the uppermost 3 pairs reclinate, the lowest of all reaching to thefirst third of the second antennal joint, the rows diverging onlygradually; antennae black, third joint three times the second, aristalong plumose not quite to tip. Palpi black, of ordinary size; pro-boscis short. Back of head with black hairs, only a few pale aroundthe neck and below. Buca one-fifth the eye height.Thorax black with green reflections; mesonotum when viewedfrom behind with two white pollinose stripes just inside the dorso-central row^s, and another pair from humeri to suture; behind thesuture these begin again a little higher up and converge to follow thesides of the scutellum nearly to its apex. Presutural acrostichalssmall but distinct; sternopleurals 3, the intermediate smaller andalmost in line with the others.Abdomen subshining, blue-green, with very faint pollen exceptbelow ; bristles as in generic characters ; genital segments of moderatesize, without bristles, but just below the anus on the inner forcepswith a striking tuft of hairs. Fifth sternite with large, diverging,bare lobes, which turn up suddenly in a lobe directed forward andare truncate beyond this.
? Legs black, the femora slightly bluish ; middle tibia on inner hindside Avith suberect hair, hind tibia on inner flexor side with 2 to 3longer, fine hairs, on outer side with one bristle.Wings slightly smoky ; third vein curved so as to widen the apicalcell beyond its middle; third vein hairy almost to small cross vein.Length, 7.5 and 8 mm.Described from tw^o males collected at La Suiza, Costa Rica,April 20 and 24, 1923, by Pablo Schild ; they were sent to the Mu-seum for identification by Prof. A. L. Melander, and the type is re-turned to him.Paratype.?M^lQ, Cat. No. 26842, U.S.N.M.NOTOCHAETA TOWNSENDI, new speciesMale.?Front 0.18 of the head wddth in each of the two specimens
;
parafrontais and parafacials light golden pollinose, the latter witha few just distinguishable, minute black hairs in a single row;frontals about 14, the upper 3 reclinate, the remainder rather fine
ART 18 DIPTEKA OR TWO-WINGED FLIES ALDRICH 23and close together reaching down to middle of second antenna!joint, and only moderately diverging. Back of head with all blackhairs. Head otherwise as in comata.Thorax black, striped as in comata, but the pollinose parts aremore yellowish, the scutellum is wholly bordered (seen from be-hind) with this color, and it extends across the prescutellar regionexcept a brown space at middle. No prescutellar or other acros-tichals; the dorsocentral usually occurring behind the suture ismissing, leaving only two posterior. Sternopleurals 3. Calypterswhite, bare.Abdomen decidedly bluish in ground color, bases of the segmentsthinly white pollinose, the pollen denser at the sides and below anddiminishes gradually behind, leaving only the apical third anda median stripe on segments 2 to 4 entirely shining. No medianmarginals on segments 1 and 2, a strong erect row on 3 and 4.Genital segments rather large, black, with erect hair, which becomesalmost bristlelike on the apex of the first. The thick broad penis isthe most characteristic part. Fifth sternite with simple divergingsides, inconspicuous. Wings as in comata. Legs as in comata, butno suberect or villous hairs on the middle tibiae.Length, 7.5 and 8.5 mm.Described from two males collected by C. H. T, Townsend onHuascaray Ridge, Jaen Province, Peru, on September 21.Type.?M2lQ, Cat. No. 26843, U.S.N.M.XOTOCHAETA ANGUSTA, new speciesMale.?Blackish, the pollinose markings distinctly golden in colorexcept on the lovrer part of the pleurae and legs. Head some-what narrower in general outline than in comata, but the front atnarrowest slightly wider; parafacials a little wider than the narrowthird joint. Thorax with four golden pollinose stripes, the innerones nearly contiguous in front; the median brownish-black stripereaches beyond the middle of the scutellum, the two pollinosestripes on each side coalesce just in front of the scutellum; one pairof small, but distinct prescutellars ; sternopleurals 2. Abdomenwith a median shining blackish stripe which expands on the hindmargins of the second and third segments, extending around to theA'enter and more or less forward on the lat'?!!il dorsal portion. Nomedian marginals on the second segment, tlie tiiird with three pairs.Oenital segments black, yellow pollinose, the genitalia small, black-ish. Legs black, middle tibia with one bristle on outer front side.Wings subhyaline: third vein with about six hairs at base.Length, 7.2 mm.One male. Corazal. Canal Zone, Panama, June, 1911 (Busck).Type.?M?L\e, Cat. No. 26844, U.S.N.M.
24 PROCEEDINGS OF THE XATIOXAL MUSEUM vol. 66Genus SARCOPHAGA MeigenSarcopJiaga Meigen, Syst. Besclir., vol. 5, p. 14, 1826.In the following three species the puparia will be described andfigured in a forthcoming comprehensive work by Charles T. Greene.SARCOPHAGA PLACIDA, new speciesFig. b.31ale.?Front 0.25 of head width (the same in both specimens) ;parafacials and parafrontals golden yellow pollinose, the formerwith only a few hairs near the eye; frontals eight, the upper onelarge and' eclinate, the lowest at the level of the middle of the secondantennal joint, hardly diverging toward the eye, about as in com-munis. Antennae black, third joint rather slender, fully twice thesecond, arista plumose for nearly three-fourths its length; facialridges black below, the hairs ascending although sparsely to the mid-dle ; palpi black ; bucca one-third the eye height, golden pollinose onanterior half. Beard pale except two orbital rows above, reduced toone at lower curve of eye.Thorax gray, with three strongly marked black stripes, the innerreaching the tip of the scutellum, and a narrow, short stripe each sidealong the supraalars; pleura with a shining black stripe along thesuture above sternopleura. No anterior acrostichals; posteriordorsocentrals four, but only the posterior two of any size; sterno-pleurals three; scutellum with two lateral and a subdiscal pair ofbristles, and in addition to these a tuft of dense white or yellowhair on the vertical border near the base.Abdomen tessellated as usual, toward its tip becoming reddish inground color; first and second segments without median marginals;third with a large pair, fourth with a row of about eight. Genitalsegments wholly yellow, narrow and elongate; the first w^th only afew small hairs, the second with moderate black hair and a few smallbristles. Forceps black, long and narrow and closely touching eachother throughout their length, beyond the middle strongly taperingand becoming larger again near the apices, which are rounded andclavate; on the anterior side the profile is nearly straight. Accessoryplate small, yellow, its anterior end bearing a tuft of black short hair.Posterior clasper small, slender, yellow, bearing a long hair near itsbase; anterior clasper long and flat, yellow, fitting against the penisas if a part of it, the tip truncate. Penis short, mostly black, theterminal segment consisting mostly of three structures: {a) A flattransverse divided posterior plate; (&) two slender black filamentscoming up in the middle and recurved; {c) two lateral black rods,,tapering and curving forward, connected on the anterior side witlia transparent membrane which forms a half-cylinder or trough, clos-
ART 18 DIPTEEA OR TWO-WIXGED FLIES ALDRICH 25ing the anterior side of the organ. Fifth sternite delicate, yellow,retracted, in the form of a broad U, with rather dense hair on theinner side of the arms. The inflexed ends of the fourth tergite bearlong hair.
^crcophaqa subaenescsns rllJricfr Ma^icera arafor/Jle/nch
S^rcophs^a p.ladd^ .-H :'drich SdrcOOhaCa mOr053/I/drichI JFig. 1.
?
Male genitalia, a, Sarcophaga subaenescbns, new species ; 6, SarcophagaPLACIDA, NEW SPECIES; C, MASICERA ARATOR, NEW SPECIES; d, SARCOPHAGA MOKOSA,NEW SPECIES. Drawn by C. T. Greene.Legs black, femora stout, middle ones with oomb on posterior apicaledge below; middle tibia with a single bristle on outer front side;hind tibia without viliosity; claws and pulvilli large, the latter in-fuscated.
26 PROCEEDINGS OF THE NATIONAL MUSEUM vol.66Wings hyaline, with the usual venation; third vein with eight toten hairs at base, reaching halfway to cross vein ; third costal segmentslightly longer than fifth : no costal spine. Epaulet black, subepauletyellow.Female.?Front 0.31 of head width (average of two, 0,30 and0.32) ; the usual orbitals present, lower frontals as in male. Middletibiae Avith two or three bristles on outer front side. Tufts of whiteor yellow hair on sides of scutellum as in male. Genital segments alittle reddish, the organs much retracted.Length of male, 10 to 11 mm.; of female, 8 to 10 mm.Described from one male, Brownsville, Texas, collected by T. C.Barber in June, 1922; and from three lots, all from James Zetek:(a) One male and six females, reared at Ancon, Canal Zone, fromdead Murex; (&) one male, one female, reared at Fort Amador,Canal Zone, from Hylesia., species, which we assume was dead tobegin with; (c) four males and four females, reared at Ancon byone of Mr. Zetek's predecessors, but the data are now lost. Thisthird lot are paler in color of pollen, and the lateral scutellar tuftsare yellow instead instead of whitish, but the puparia show no dif-ferences. The type and allotype are from the lot a. Mr. Zetek'snumbers for the three lots are Z-2305, Z-2303, and Z-1834, re-spectively.ry;?e.?Male, Cat. No. 27097, U.S.N.M.SARCOPHAGA MOROSA, new speciesFig. d.J/aZe.?Front about 0.22 of head width, the head being damagedon the sides the measurement can not be taken exactly ; parafrontalsand parafacials pollinose with a distinct but not deep golden tinge,the former narrower than the middle stripe, the latter with a rowof hairs next the eye becoming bristly below and a few additionalhairs; two or three upper frontal pairs decreasingly reclinate, theuppermost not especially strong; lowest frontals strongly divergent;antennae black, third joint less than twice the second, moderatelywide, arista plumose more than to middle. Palpi black; buccaabout one-third eye height, with black hair except behind; back ofhead with mostly black hair.Thorax with the usual three black stripes and an outer pairshorter and weaker. Acrostichals, 0, 1; dorsocentrals, 4, 3, alllarge; sternopleural, 3; scutellum with two lateral, one apical, andone discal near tip. Postalar declivity with hairs in the middle.Abdomen tessellated as usual; first and second segments withoutmedian marginals, third with a pair, fourth with a marginal row.Genital segments rather large, wholly black, with erect, soft hair;forceps red on the attached basal part, the rest black, divergent, of
ART 18 DIPTERA OR TWO-WINGED FLIES ALDKICH 27uniform breadth to the tip, where the hind edge curves forward toform a sharp tooth at the middle of the apex; accessory plate red-dish, triangular, the apical side broad. Posterior clasper small andinconspicuous, anterior clasper long and low, rounded at tip. Penisthick and rather short, the distal segment globose, blackish; theapical posterior part is suddenly narrowed into a curved beak ex-tending forward, underneath which two blunt filaments emerge; theside pieces of the distal segment are in the form of truncated plates,diverging apically from each other. Fifth sternite yellow, retracted,apparently a plain V.Legs black ; middle femora with combs before and behind at tip
;
middle tibiae with two bristles on outer front side; hind tibiae withlong villosity on outer and inner flexor sides. Claws and pulvillilarge, the latter infuscated.Wings hyaline ; third vein at base with four to six hairs ; bend offourth vein with stump or heavy fold; third costal segment con-siderably longer than fifth ; no costal spine.Length 11 mm.Described from one male specimen, reared by F. Johansen from alarva taken near Ottawa, Canada; the fly emerged July 11, 1918.ry;>e.?Male, Cat. No. 27098, U.S.N.M.The species is close to pulla Aldrich, but differs in the genitalia,especially the form of the accessory plate, which is entirely different,and in the forceps.SARCOPHAGA SUBAENESCENS, new speciesFig. a.Male.?Front 0.21 of head width (one specimen) ; parafrontalsmuch narrower than median stripe, metallic above, thinly whitepollinose below; frontal bristles ten, the uppermost reclinate andsomewhat larger, the lowest diverging toward eye ; a single vertical
;
ocellars normal; parafacials with thin white pollen, quite narrowbelow, the usual row of hairs bristly below; antennae black, thirdjoint reddish basally, over twice the second, arista with somewhatshort plumosity extending only to middle. Palpi black; buccahardly one-third eye height, with black hair; back of head with onlya little pale hair about foramen and below.Thorax thinly glaucous pollinose, subshining, but when viewedfrom behind showing the usual three dark stripes well separated.Acrostichal 0, 1 ; dorsocentral 2, 3, sternopleural 3 ; scutellum Avith twolateral, one apical, and one discal ; postalar declivity with a few hairsin middle.Abdomen black with very thin tessellation, subshining and with aslight aeneous reflection; first segment without median marginals;
28 PEOCEEDINGS OF THE NATIONAL MUSEUM vol.66second with a small, depressed pair rather far apart in the describedspecimen; third with a large pair, fourth with a marginal row.Genital segments small, wholly black, with small black hair, the sec-ond segment with two small bristles turned up behind. Forcepsminute, black, diverging only at tip, in profile of rather even width,bearing a small tooth at the middle of the blunt tip. Accessory plateyellow, shining, rounded, not much produced apically. Posteriorclasper not visible in the specimen, anterior minute, depressed for-ward. Penis small, blackish, distinctly jointed, the distal segmentsomewhat globose, its main sclerite forming the back and sides hasa thick, rounded, transverse rim at apex, in front of which in themiddle is a small protuberance. Fifth sternite wholly retracted inthe specimen.Legs black, middle femur without comb, middle tibia with onebristle on outer front side, hind tibia without villosity; claws andpulvillia large, the latter infuscated.Wings hyaline ; third vein with a row of six or eight hairs ; fourthvein ending nearer apex than in most species (less than half the fifthcostal segment) ; third costal segment slightly shorter than fifth.Length 5 mm.Described from a single male, reared by Eay T. Webber from apuparium which he took out of a spider's web at Somerville, NewJersey, on June 23, 1922.Type.?MdlQ, Cat. No. 27099, U.S.N.M.The nearest relative is davidsoni, Coquillett, which has been rearedfrom spiders' eggs; subaenescens differs from this species in havingno anterior acrostichals, thinner pollen, a slight aeneous cast to thecolor of the abdomen, etc. It is highly probable that subaenescensattacks spiders' eggs. Family TACHINIDAEGenus ATACTA SchinerAtacta ScHiNEK, Novara Reise, 1868, p. 328. Type and sole species hrasil-iensis, new.Atactomima Townsend, Bulletin Amer. Mus. Nat. Hist., vol. 35, 1916, p. 15.Type and sole species crescentis, new, from Brazil.The characters of Atacta are in brief as follows : Head wider thanthorax, subhemispherical ; front in male strikingly narrow above(about twice the ocellar triangle), the eyes diverging at a wide angleto the level of the antennae, parafrontals someAvhat triangular inform, usually silvery, covered with dense hair, frontal stripe onlyabout as wide as ocellar triangle; only one vertical on each side.Female with wide front, the parafrontals slightly inflated, verybroad, with a long dark, translucent reflecting spot on each bearing
ART 18 DIPTERA OR TWO-WINGER FLIES ALDRICH 29
a row of five orbitals, the uppermost of which is reclinate. In bothsexes the ocellars are present, the frontals diverge below in a broadcurve almost to the eye margin at the level of the middle of thesecond antennal joint; the antennae are slender and small, the sec-ond joint equal to the third ; the face is very flat, the parafacials bareand wide, the facial ridges bare, the vibrissae distinctly above theedge of the mouth. Bucca in profile about one-fourth the eye height.Palpi and proboscis ordinary. The thoracic chaetotaxy is the sameas in Belvosia unifasciata Robineau-Desvoidy {Triachora of Town-send) and in thorax, abdomen, legs and wings there are no genericcharacters unlike the latter.Brauer and Bergenstamm include Brachycoma nigriceps Van derWulp in Atacta from a specimen * ; but as the third antennal jointis twice the second I doubt if they understood the genus as hereinrestricted.The species crescentis Townsend seems clearly congeneric from themale in the National Museum ; the characters given by Townsend forthe genus Atactovmna are all specific in my opinion.TABLE OF SPECIES
1. Fourth abdominal segment covered with dense golden pollen ; hair of me-dian portion of second and third abdominal segments recumbent ; secondabdominal segment almost always destitute of median marginals; malewith silvery parafrontals brasiliensis Schiner.Fourth abdominal segment with gray pollen ; hairs of median region ofsecond and third abdominal segments erect ; second abdominal segmentwith a pair of median marginal bristles 22. Antennae and palpi black, tip of latter yellow crassiceps, new species.Antennae with second joint yellow, palpi yellow 33. Male with bright silvery pollen on parafrontals, strongly contrasting withthe white or yellowish parafacials argentifrens, new species.Male with the strongly widened parafrontals more chalky white, almostconcolorous with parafacials crescentis Townsend.ATACTA BRASILIENSIS SchinerAtacta 'brasiliensis Schiner, Novara Reise, 1S6S, p. 328.
?
Brauer andBergenstamm, Zweifl. Kais. Mus., pt. 4, 18S9, p. 96, fig. 57 ; pt. 5, 1891,pp. 340, 365; pt. 6, 1893.?Townsend, Journ. N. Y. Ent. Soc, vol. 23,1915, p. 64.?C. S. Brimley, Ent. News, vol. 33, 1922, p. 21.Brachycoma laticeps Van deb Wulp, Biologia Cent.-Amer., Dipt., vol. 2.1890, p. 92.Atacta apicalis Coquillett, Revis. Tachin., 1897, p. 83.Originally described from a female taken in Brazil, and afterwardcollected by Townsend in Peru, the species ranges northward to thevicinity of Washington. Specimens from North America in theUnited States National Museum are as follows: One male, ChiriquiProvince, Canal Zone, reared from Remigia repanda Fabricius by
* Zweifl. Kais. Mus., pt. 5, 1891, p. 365.
30 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 6GZetek (No. 223(M) ; one female, Higuito, Costa Rica (Pablo Scliild)
;
one female, Puerto Barrios, Gnat., February 2-1, 1905 (C. C. Deam) :one male, Tifton, Georgia, September 8, 1896 (G. R. Pilate, type ofaplcalis) ; one female, Raleigh, North Carolina, July, 1906 (Brimley)
:
one female. Falls Church, Virginia, May 30 (Banks) ; and onefemale, Great Falls, Virginia, August 9, 1923 (Aldrich).ATACTA CRASSICEPS. new speciesMale.?front very narrow at the ocelli, only about twice the widthof the ocellar triangle, widening very rapidly to the lower end of thefrontal row. The parafrontals are covered with somewhat goldenpollen which in most lights gives a brownish or almost black reflec-tion; they are quite protuberant, a little inflated and rather denselycovered with erect black hairs. The frontal bristles begin a littlebelow the ocelli and the rows diverge rapidly below, ending close tothe eye at the level of the middle of the second antennal joint ; ocellarbristles distinct and a dense tuft of hair behind them to the vertex.Inner verticals are developed; face very flat with yellow pollenthrough which a darker ground color shows on each side ; parafacialsbare with yellow pollen; its least width nearly equal to the lengthof the third antennal joint. Antennae black, very slender, the aristashort, gradually tapering; its penultimate joint twice as long asbroad. Vibrissae considerably above the mouth (two-thirds thelength of the second antennal joint) with a group of half a dozeosmall hairs and bristles above them, the highest a little above the tipsof the antennae; palpi and proboscis of ordinary size; bucca one-third the eyeheight. Thorax black with conspicuous stripes of whitepollen, which leave between them a pair of abbreviated black stripesin front between the acrostichal and dorsocentral ; a pair of completeblack stripes beginning just mesad of the humerus and extending tothe scutellum; and a short median black stripe beginning at thescutellum and extending forward nearly to the suture. Chaetotaxy
:
acrostichals, anterior 3, posterior 3; dorsocentrals, anterior 2, pos-terior 4; intraalars, posterior 3, anterior 1; supraalar 3; postalar 2;humeral 3 and 4; sternopleural 4; scutellum with three equal pairsof the margin, the last of which might be called apical, and one pairdiscal. Abdomen black with gray reflecting pollen which on thefourth segment becomes more dense and yellowish-gray in color; thesecond segment with a single pair of marginals ; third segment withabout five pairs; fourth segment with five or six pairs in a single rowconsiderably before the apex. Genitalia small and concealed, of arather common type. Fifth sternite with a U-shaped incision, thelobes black and almost bare. Legs black, the middle tibia with threeor four bristles on the outer front side, the hind tibia distinctlyciliated on the outer side with one longer bristle below the middle.
ART 18 DIPTEKA OR TWO-WINGED FLIES ALDRICH 31Wings hyaline, third vein with four or five bristles at base; fourthvein rather suddenly bent, almost at a right angle from which itcurA'es a little outward reaching the costa considerably before the tipof the wing.Length 9 mm.Female.?The front has the two large dark, reflecting spots on theparafrontals as in hrasiliensis., but the color of the remaining pollenis gray rather than golden. This with the characters given in thetable will readily distinguish the two species.Length 7.5 mm.Described from three males and one female. The males are fromGreat Falls, Virginia, August 9, 1923 (Aldrich) : Tupelo, Missis-sippi, September 30, 1921 (H. W. Allen) ; Hope, Arkansas, August21, 1922 (received from C. W. Johnson and returned to him). Thesingle female is from Opelousas, Louisiana, April, 1897 (Pilate).r^/^oe.?Male, Cat. No. 26845, U.S.N.M., from Great Falls, Vir-ginia. ATACTA ARGENTIFRONS. new speciesMale.?This species is most nearly related to crassiceps, the singlemale specimen has the front much the same as in hrasiliensis, theparafrontals having very decided silvery color and being less in-flated than in crassiceps. The thoracic stripes are the same in allthe species which I have seen, but hrasiliensis is the only one withdeep golden pollen on the fourth abdominal segment and withoutmedian marginals on the second. The female of this species is un-known.Length 9.5 mm.Described from a single male collected in May by H. H. Smithat Corumba, Brazil.Type in the collection of the American Museum of National His-tory. ATACTA CRESCENTIS (Townsend)Atactomina crescenhs Townsend, Bull. Amer. Mus. Nat. Hist., vol. 35,1916, p. 15.Described from four males and a female in the American Museum.Locality, Chapada, Brazil. One paratype male is in the UnitedStates National Museum. The supposed generic characters are veryslight except that the parafrontals are quite rapidly widened below,the eye being rather more crescent-shaped than in the other species.ATACTA NIGRIPALPIS (Van der Wulp)Brachycoma nigripalpis Van der Wulp, Biologia Cent. Amer. Dipt.,vol. 2, p. 98.Atacta nigripalpis Brauee and Bergenstamm, Zweifl. Kais. Mus., pt. 5,1891, p. 365.
32 PEOCEEDINGS OF THE NATIONAL MUSEUM vol. 66Not seen by the writer and placed here on account of the statementof Brauer and Bergenstamm. who saw a type specimen. I doubtthe generic reference very much as none of the other species havesuch an elongated third antenna! joint.Genus MASICERA MacquartMasicera Macqiart, Ins. Dipt, du Nord de la France, 1S34, p. 285.
?
Coquil-LETT, Revision of the Tachinidae (Tech. Bull. No. 7, Division of Entom-ology), p. 113, 1897.MASICERA ARATOR, new speciesFig. c.Male.?Front 0.28 of head width (average of four, 0.28, 0.28, 0.26,0.29) ; parafrontals light golden pollinose; frontal bristles abouteight, the two uppermost rather large, reclinate, the lowest reachingthe level of the arista and strongly diverging toward the eyes; onepair of verticals: ocellars large; parafacials silvery from the lowestfrontals, at narrowest less than half the width of third antennaljoint; first two joints of antennae and usually the base of thirdred, the third broad and long, almost reaching the vibrissae, four orfive times the second; arista of moderate length, hardly thickenedbasally; face concolorous with parafacials, its ridges rather sharp,bare except close to vibrissae ; palpi yellow, ordinary, proboscis short,fleshy ; bucca over one-fourth of eye height.Thorax gray pollinose, with very indistinct darker stripes.Acrostichal 3, 3 ; dorsocentral 3, 4 ; humeral 3 ; posthumeral 2 ; presu-tural 2; notopleural 2; supraalar 3; intraalar 3; postalar 2; sterno-pleural 3; pteropleural 0; scutellum with 3 lateral, 1 apical, notupturned, 1 discal.Abdomen black with subsilvery basal bands of pollen on segmentstwo to four, which to the naked eye give the impression of beingequal to the alternating black bands; under the lens in some angleshowever the pollen covers most of the segments. First segment withone median marginal pair ; second segment with a discal and a margi-nal pair; third with a discal pair and a marginal row; fourth withtwo to eight discal and a marginal row. Genital segments ratherlarge, wholly black, with black hair and the second with a pair ofbristles directed backward. Inner forceps black, slender, long,deeply divided but not divergent, the tips blunt and slightly bentback. Outer forceps with very peculiar and characteristic shape,long and flat, shining black, beyond the middle suddenly wideningbackward in a thin, concave margin, the apex sharp and curved alittle forward so that the whole apical part suggests a plough share.Fifth sternite large and prominent, black, without special bristles orhairs.
ART 18 DIPTEEA OR TWO-WIXGED FLIES ALDRICH 33Legs black, claws and pulvilli iong, especially the front ones;front tibi'a with two bristles on outer hind side; middle tibia withone bristle on outer front side near middle; hind tibia on outerhind side subciliate, with one long about middle.Wings hyaline, fourth vein w^ith oblique and rounded bend, thedistance from its tip to extreme apex of wing barely equal to one-half the hind cross vein. Third vein with two to three hairs atbase.Female.?Front 0.30 of head width at vertex, wider 'anteriorly(one specimen) ; parafacials slightly wider and third antennal jointnarrower (hardly three times the second) ; abdomen with narrowersubsilvery bands, the shining black intervening portion wider;genital org'ans retracted, no indications of a piercing organ ; middletibia with two bristles on outer front side ; hind tibia with irregularbristles on outer hind side, not subciliate.Length of male, 8 to 8.5 mm; of female, 9 mm.Described from four males and one female. The type male andallotype female Avere collected at Linglestown, Pennsylvania, June15, 1913, and were received from the State Bureau of Plant Industry,Harrisburg, Pennsylvania, through the courtesy of A. B. Champlain.One male was bred " from a I'arge Tipula larva ' collected by JamesFletcher at Chelsea, Quebec (near Ottawa), on May 27, 1906; thepuparium was formed on June 3, and the fly emerged on June 27.The puparium will be figured by Charles T. Greene in a later paper.One m'ale was collected by H. C. Fall at Tyngsboro, Massachusetts,on July 26, 1916, and is deposited in the Boston Society of NaturalHistory. The remaining male was collected by E. C. Shannon atDead Run, Virginia (close to Washington), on June 22, 1913.Type^ allotype, and one paratype.?Both sexes. Cat. No. 27100,U.S.N.M. Family DEXIIDAEGenus DEXIA MeigenBexia Meigex, Systemat. Beschreib.. vol. 5, 1826. p. 33.
?
Baek, Die Tach-ininen, 1921, p. 160.DEXIA VENTRALIS, new speciesMale.?Front O.IS of headwidth (average of three, 0.16, 0.18, and0.19), the middle stripe reddish-brown, the parafrontrals, parafacials.posterior orbits and all below the eye light golden pollinose except abroad brown stripe from the eye downward and forward. Parafa-cials bare. Head from in front obviously higher than wide. Verti-cals small (only one pair), frontals 7 or 8 irregular, beginning belowthe ocelli and ending at base of antennae; vibrissae above mouth, a
34 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 66:few small hairs above them. Antennae small, yellow, separated by awell-developed carina which extends between them at base, althouglilow and narrow here: third joint slender, twice the second, not nearlyleaching the vibrissae: arista short with dense erect plumosity.Palpi smallish, yellow; proboscis small, the labella yellow. Biiccaone-half the eye height.Thorax with dense yellow pollen becoming more gray on pleurae,,with four narrow incomplete darker stripes. Chaetotaxy : acrostichal1, 1 (sometimes a second small pair before the prescutellars) ; dorso-central 4, 3 ; humeral 2 ; posthumeral 1 ; presutural 1 ; notopleural 2
;
supraalar 2 ; intraalar 2 ; postalar 2 ; scutellum with 2 lateral, a largedecussate apical pair and a small discal ; sternopleural 2 ; pteropleuralminute.Abdomen mostly yellow in ground color, a variable interruptedmedian black stripe, and narrow variable black hind margins on lastthree segments; the inflexed ends of the second and third tergitescome together below and are more or less tipped with black, thusforming a variable black median ventral stripe. First segment with-out median marginals; second with one pair discal and one marginal,other marginals toward the sides; third segment with one or twopairs of discals, and a stout marginal row of 8 ; fourth segment withirregular discal and apical rows. Genitalia small, yellow, the lobesof the fifth sternite black. The pollen of the abdomen is yellowishand confined to the bases of the last three segments, covering half ofthe fourth.Legs 3"ellow, tarsi however black : claws and pulvilli long. Middletibia with only one small bristle on outer front side ; hind tibia withtwo small on outer hind side.Wings somewhat brownish; costal spine distinct; fourth A'einsharply bent at a right angle, with a slight or distinct stump at thebend, ending not very far before the apex; third vein with only 2 to5 small hairs at base.Female.?Looks like a distinct species, but was reared with thesemales. The abdomen is but little or hardly at all yellow in groundcolor, the basal pollinose bands contrasting with the shining blackapical half on segments 2 to 4. The wing is broader, and the fourthvein curves more distinctly backward beyond the crossvein. Widthof front at narrowest (vertex) is 0.38 in one specimen and 0.45 in theother. There are only 5 to 6 frontals, the upper one turned back andoutward ; the usual two orbitals are large and proclinate. The usualtAvo pairs of verticals are present.Length of male, 8.6 to 10.3 mm. ; of female. 7.6 and 8.4 mm.Described from seven males and two females, reared from scara-baeid beetles at Suigen, Korea, by C. P. Clausen and J. L. King.r.v;?^.?Male, Cat. No. 27245, ij.S.N.M.
AUTlS DIPTEKA OR TWO-WINGED FLIES ALDRICH 35The species is strictly congeneric with Dexia rustica Fabricius ofEurope, type of the genns. It agrees very well with the descriptionof Dexia divergens Walker, described from Mount Ophir ^ ; but oneof our specimens was compared with Walker's type in the BritishMuseum by Dr. J. D. Tothill and showed differences apparentlyspecific. Genus EUTRIXOPSIS TownsendEutrixopsis Townsend, Insecutor In.scitiae Menst., vol. G, p. 166, 1918.This genus was based upon the single species javana, describedby Townsend on the same page ; only a single specimen was known,a male from JaA-a. The species has recently been found by Clausenand King, of the United States Bureau of Entomology, to be animportant parasite of the " Japanese Beetle," Popillia japonicaNewm., in Japan. Their studies of its biology and economic rela-tions will be published elsewhere.The genus differs from Eutrixa (inclusive of Eutrixoides Walton)chiefly in having mucli narrower parafacials and broader and flatterfacial ridges: in other Avorcls, the suture encloses a much broaderportion of the head below. The parafacial also bears close to theeye a more or less double row of hairs. Both genera are parasiticon melolonthine beetles in the adult stage.Since javana has been only briefly described hitherto, its eco-nomic importance requires that it be given a full description here.EUTRIXOPSIS JAVANA TownsendEutrixopsis javaiHi Townsend, Insecutor lascitiae Menst., vol. 6, p. 166,1918.Male.?A brown fly with mostly yellow abdomen and legs. Eyesalmost contiguous on the front, separated by less than the width ofthe anterior ocellus, ocellar triangle small, elevated ; ocellar and ver-tical bristles absent, no hairs on front above middle, a few smallbristles below stopping short of the lunule; small hairs begin onlower parafrontais and continue down the narrow parafacials in amostly double row to the lower end of the lunule. Antennae brown,very small, third joint twice the second, arista pale, about threetimes as long as the third joint, bare, its penultimate joint short.Facial ridges flat, converging below and at the closest point sepa-rated by hardly more than the width of the third antennal joint. Novibrissae, the ridges hairy in this region and bearing a few smallbut increasing bristles toward the mouth, which is some distancebelow. Palpi and proboscis ordinary, the former yellow. Back ofhead concave above, convex below. Bucca half the eye height, the
eProc. Linn. Soc. vol. 1, 1856, p. 21.
36 PROCEEDINGS OF THE NATIONAL MUSEUM vol.66large transverse impression ending vertically below the eye. Thoraxbrown with smooth, silky gray pollen and tAvo narrow brown stripesnear middle. Chaetotaxj'^ : Acrostichals, 1 next to scutellum ; dorso-centrals, 3 anterior, 4 posterior; intraalar 1, supraalar 1; postalaar2; humeral 1 or 2; prescutellar 1; notopleural 2; scutellum with 2lateral, 1 apical of same size, 1 discal; sternopleural 2. Calypterswhitish.Abdomen yellow in ground color, with thin gray pollen; seg-ments 1-3 with narrow posterior dark band and a dark middle lineof same width. First segment without marginal bristles; secondwith a small median pair; third with marginal row of about 12,not large. Genital segments yellow, small, hairy; inner forcepsunited into a small slender yellow process curving forward, blunt attip; outer forceps yellow, slender, as large and long as thecombined inner ones. Fifth sternite with broad yellow lobes, sepa-rated by a broad V-shaped incision.Legs yellow; tips of hind femora infuscated and all the tibiaewith faint dark reflections in certain lights. Claws long and pul-villi nearly equal to last tarsal joint. Wings subhyaline, third veinwith two or three hairs at base.Length, 6 to 7 mm.Female.?Front considerably wider than front ocellus; as theonly specimen is somewhat shriveled, the front may normally bealmost as wide as the ocellar triangle. Claws long for a female,but not so long as in the male ; pulvilli rounded, more than half aslong as last tarsal joint. 'Wliether any sort of piercing larvipositoris present is not clear; the organs have been unsuccessfully pulledapart. There may be a minute piercer, but in any event smallerthan the very distinct one of EutHxa exilis and not comparableVv ith the large one of jonesi.Length, 6 mm.Five males and six females, reared at the Japanese beetle labora-tory from adults of the so-called Japanese beetle, Popillia japonicaNewman ; the infested beetles were obtained in Japan.Type.?Male, Cat. No. 26846, U.S.N.M.o