NEW TWO-WINGED FLIES OF THE FAMILY CALLIPHO-RIDAE FROM CHINA
By J. M. AldrichAssociate Curator, Division of Insects, United States Naiional Museum
The present paper includes four new species and two new generaof muscoid flies from China; one of the species has recently beenintroduced into California.Genus CALLIPHORA Robineau-DesvoidyCalUphora Robineau-Desvoidy, Myoclaires, 1830, p. 433.
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Van deb Wulp;.Biologia, Dipt., vol. 2, 1896, p. 294.?Hough, Ent. News, vol 10, 1899, p.65; Zool. Bull., vol. 2, 1899, p. 285.
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Shannon, Insecutor Ins. Menst, vol.11, 1923, p. 107; Proc. Eut. Soc. Wash., vol. 28, 1926, p. 134.?Malloch,Trans. New Zealanfl Inst., vol. 55, 1924, p. 640; Proc. Linn. Soc. NewSouth Wales, vol. 52, 1927, p. 303.?Bezzi, Bull. Ent. Research, vol. 17,1927, p. 240.?LuNDBECK, Dipt. Danica, vol. 7, 1927, p. 149.CALLIPHORA GRAHAMl, new speciesOf the size and appearance of vomitoria Linnaeus, but having inboth sexes no inner presutural bristle (presutural intraalar of Hough,,hindmost sublateral of Shannon), beard wholly black, gray pollenof thorax and abdomen considerably more dense; the male also dif-fers in having the front about three times as wide (0.14 of head widthbelow front ocellus), and the genitalia greatly enlarged and of quiteunique form.The abdomen of the male is short and broad and the shining blackgenital segments (fig. 1) are very large, the first vertical, the secondextending forward more than halfway to the base of the abdomen;first visible sternite short, second longer than usual, third and fourthshort, curving downward, the fourth with more dense pile than pre-ceding. Fifth prominent at base and bearing a dense tuft of shortspines each side of middle line, not much separated ; lobes large andwidely divergent, the large median concavity bearing tv\"o long paral-lel lobes of soft structure, their tips turned outward, which are sepa-No. 2844.?Proceedings U. S. National Museum, Vol. 78, Art. I.740?30 I
PROCEEDINGS OF THE NATIONAL MUSEUM
rated from the more chitinized main iobes by a deep groove. Outerforceps shinino; black, giving off a pair of blunt processes at base,which project backward, the remainder long, extending forward andcurving downward, of uniform width to tip, which is rounded.Inner forceps rudimentary, narrow, less than half as long as theouter. In repose the closely approximated outer forceps evidentlylie in the median groove between the two soft inner lobes of the fifthsternite.Thoracic chaetotaxy : acrostichal 3, 3; dorsocentral 3, 3; humeral 4;posthumeral 3 (sublateral of Shannon 3) ; presutural 1 ; notopleural2; supraalar 3; intraalar 2 (none close behind suture); postalar 2;scutellum with four lateral (api-cal included), one discal ; sterno-pleural 2. 1.Wing subhyaline, as in vomi-toria; basicosta black, subcostalsclerite yellow, bare : third veinwith three or four small hairsaboA'e and below. Hind caiypterbrown with white rim.Length, 8-10 mm.Described from 62 specimens ofboth sexes: 34. including typeand allotype, were collected atSuifu. Szechuen Province, China,by Rev. D. C. Graham, afterwdiom the species is named; onemale from the same collector wascollected at the summit of therange of mountains, Washan,same Province ; and another maleat Yellow Dragon Gorge near Songpan, in the same Province, wastaken by the same collector at an altitude exceeding 12,000 feet.From E. Brunetti were received two males taken at Hang Kow,China, and two females from Hong Kong; while Prof. T. D. A.Cockerell sent a female which he took at Kongaus, Siberia, inAugust, 1923.The remaining 29 specimens were received from H. H. Keifer,department of agriculture, California, who sent them in December,1929, writing that he reared them all from a single batch of eggsthat he found on meat at Sacramento, Calif. Thus it appears thatthe species has lately established itself in the United States. TheCalifornia specimens are dated June 10, 1929. No other NorthAmerican specimens are known.
Figure 1.
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Calliphoea grahami^ newSPECIES. Male, side view of genitalSEGMENTS (DRAWN BV D. G. HALL)
AKT. 1 NEW TWO-WINGED FLIES EEOM CHINA ALDEICH 6Twenty of the Sacramento paratypes are returned to the CaliforniaDepartment of Agriculture.l\jpe.?Male, Cat. No. 26865, U.S.N.M.XANTHOTRYXUS, new genusNear Pollenia, from which it differs especially in having the para-facials bare, and the face with a broad keel.Hypopleural bristles present, postscutellum not developed; pro-pleura and prosternum bare ; e3'es bare ; vibrissae as high aboveoral margin as in PoJJenia rudls, but farther apart; facial ridgeswith only a few setules above vibrissae; palpi normal, proboscisshort, with ordinary labella ; antennal axis and vibrissa! axis aboutequal in length; frontal bristles reaching oidy to base of antennae;arista plumose. Thorax with dense deciduous curly pale hair ; hindcalypters bare, large. Abdomen robust, first sternite pilose, densehair of dorsum becoming bristlelike apically. Wing with apicalcrossvein, the first posterior cell ending distinctly before apex : thirdvein with a few very delicate pale hairs above and belovv' at base
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subcostal sclerite with a tuft of yellow hairs; stemvein bare.Genotype.?Xanthotryxus mongol, new species.XANTHOTRYXUS MONGOL, new speciesBlack with dense appressed golden deciduous hair on thorax,the abdomen with dense golden pollen covering dorsum and becom-ing gray underneath.Male.?Front narrow above, at narrowest hardly wider than dis-tance across hind ocelli, thence rapidly widening and the face wid-ening at same angle; parafrontals, parafacials, and upper part offace golden pollinose, lower face and cheek more gray ; frontal stripetriangular, wide anteriorly, brown. Ocellars normal, proclinate, nolarge verticals nor reclinate frontals. The single row of convergentfrontals begins some distance from the anterior ocellus and consistsof about 15 bristles. Antennae black, third joint rather slender,twice the second; arista pale except at base, of moderate length,plumose almost to tip. Palpi black. Beard yellow, hairs of cheekblack, cheek three-fourths of eye height.Thoracic chaetotaxy : acrostichal 2, 3; dorsocentral 3, 4; humeral3 or 4; posthumeral 1 (inner) ; presutural 1 (so far forward it mightbe called posthumeral); notopleural 2; supraalar 3; intraalar 2;postalar 2; sternopleural 1, 1; scutellum with 2 lateral, apical ofsame length. Postalar declivity with large tuft of yellow hairs;hind calypters brown, front ones yellow^, the long pile at junctionyellow.
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two verticals, oneMiddle tibia withCiaws
Abdomen with suberect dense black hair, third segment withmarginal row; fourth segment with irregular bristles and bristlyhairs. Genital segments black, rather large, the first with spot ofj^ellow pollen and numerous bristly hairs. Genitalia as figure (fig.2), the inner forceps slender, as long as outer, united except at ex-treme tip. Legs black; claws and pulvilli elongated, the latter yel-low ; middle tibia with one small bristle on outer front side ; base ofhind coxse bare behind. Wings brownish, especially along veins,yellowish near base; bend of fourth vein oblique, rounded, thencewith slight concavitj^ to costa.Female.?Front at vertex 0.29 of head widthsmall reclinate frontal, two proclinate orbitals.two bristles on outer front sideand puvilli small.Length, 11-13 mm.Described from seven males and fivefemales, collected in Szechuen Province,China, by Kev. D. C. Graham; eightwere taken at Chuan Chien and four onMount Omei.T^ype.?Male, Cat. No. 29168, U.S.N.M,XANTHOTEYXUS DRACO, new speciesMale.?Like niongol except that thepollen of the head is grayish-yellow,not golden; the face has a less distinctcarina, the antennae closer together atbase, and the genitalia are different.The inner forceps are distinctly sepa-rated on about the apical half, and the outer ones have expandedtips, about twice as wide as in mongol.Length, 9 mm.Described from one male collected at Yellow Dragon Gorge,Szechuen Province, China, by Rev, D. C. Graham, in altitude above12,000 feet.r^^e.?IMale, Cat. No. 29164, U.S.N.M.
FiGUEB 2. XANTHOTEYXUS MON-GOL, NEW SPECIES. Male, sideVIEW OF GEXITAL SEGMENTS(DRAWN BY D. G. HALL)
LISPOPAREA, new genusAllied to Pollenia, but has bare parafacials, vibrissae not so closetogether, facial carina low and rounded except just between bases ofantennae, and bend of fourth vein in form of a gentle curve. Thethorax has the same covering of deciduous yellow hair as in Pollenia.Differs from Xanthotryxus in the course of the fourth vein, and inhaving the facial carina less prominent. The sole species is paleyellow except mesonotum, and more slender than either of the
AET. 1 iq-EW TWO-WINGED FLIES FEOM CHINA A.LDE.IGH Ogenera mentioned, having a very diiferent appearance. Runs toAnthraco'niyia new genus in Mailoch's Key/ but that genus has noneof the short silky thoracic hair characteristic of Pollenia.Genotype.?Lisfoparea f.ava^ new species.LISPOPAREA FLAVA. new speciesMale.?Wholly yellow except the following parts : Mesonotum (ex-cept narrow sides), metanotum, and apical part of tarsi. Eyes al-most contiguous on front, separated by less than width of anteriorocellus, then diverging rapidly to their lower curve; frontal stripea broad triangle, disappearing at narrow part of front. No verti-cals, no reclinate frontals. Ocel-lars merely hairs; frontals begin-ning far forward, weak, about 10in number, barely reaching baseof antennae; third antennal jointmore than twice the second; aristashort, plumose to tip. Palpi rathersmall, proboscis slightly elongate;cheek two-thirds the eye height, fiquhb 3.?lispopabea flava, new sph-Eyes with distinctly larger facets cies. apical portion of wing (drawnBY C. T. Greene)on upper half or more whichgradually diminish downward. Head rather dull yellow but notdistinctly pollinose.Thoracic chaetotaxy: Acrostichal, 1, 1 (?); dorsocentral, 2, 3;humeral, 2; posthumeral, 1 ; presuturai, 1 (far forward as in Xantho-tryxus) ; notopleural, 2; supraalar, 2; intraalar, 2; postaiar, 2; ster-nopleural, 1, 1 ; scutellum with 2 lateral, one equally strong apical.Calypters light yellow. Abdomen wholly pale yellow, subshining,without distinct pollen. Genital segments the same. First segmentwith marginal bristles only at sides; second, third, and fourth withcomplete row, no discals. Tarsi gradually brown toward tip. Clawsand pulvilli not enlarged; front tibia with one bristle on outer hindside; middle tibia with one on outer front. Wings hyaline withyellow veins; venation as in Figure 3, the fourth vein convergingquite gently to third.Length, 6.6 mm.Described from one male collected on Mount Omei in SzechuenProvince, China, by Rev. D. G. Graham.rype.?Male, Cat. No. 42878, U.S.N.M.
1 Proc. Liim. Soc, New South Wales, Vol. 54, 1927, p. 303.
U. S. GOVERNMEWT ?RINTINS OfrlCE: 1930