NEW DIPTERA, OR TWO-WINGED FLIES, FROM AMER-ICA, ASIA, AND JAVA, WITH ADDITIONAL NOTES
By J. M. AldriciiAssociate Curator, Division of Injects, United States National Museu?n
This paper contains descriptions of 4 new genera, 13 new species,and 1 new variety of Diptera, together witli some miscellaneousnotes. It constitutes a report, which for various reasons it seemsdesirable to publish, on an accumulation of specimens collected frommany parts of the world. All the species are represented in theUnited States National Museum collections by type or paratypespecimens. Family CYRTIDAEGenus OCNAEA ErichsonEriosoma Macquakt (preoccupied), Dipteres exotiques, vol. 1, pt. 2, p. 288(Sep. p. 172), 1839.Ocnaea Erichson, Eutomographieu, vol. 1, p. 155, 1840.
?
Cole, Traus. Amer.Ent. Soc, vol. 45, p. 23, 1919.Exetasis Walker, lusecta Saundersiana, p. 203, 1S52.Two species were originally included in Ocnaea^ both new, micansfrom Mexico and longlcornis from Brazil; the second was figured.Coquillett ^ designated Ocnaea tnicans as the type species. Mac-quart designated Acrocera calida Wiedemann as type of his Erio-soma. Exetasis contained originally but one species, tumens, new,which is therefore the type ; Loew - placed the genus as a synonymof Ocnaea.The genus has very striking characters. The proboscis is veryshort; the densely hairy eyes are contiguous from the mouth almostto the vertex ; there is a short frontal triangle bearing the antennae
;
the anterior ocellus is absent; the antennae have a greatly elon-gated third antennal joint, curving downward, not bearing a styleor arista. The venation is complete and in general not unlike thatof Tdbanus^ except in a few variable details and in the regular oc-currence of a cross vein in the first posterior cell at or beyond the
iProc. U. S. Nat. Mus., vol. 37, p. 577, 1010.
" Wiener Ent. Monatsschr., a'oI. 1, p. 34, 1856.No. 2932.?Proceedings U. S. National Museum, Vol. 81, Art. 9.110571?32 1 1
2 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 81
tip of the discal. The third vein is not always forked near the tip.but this fork occurs in the genotype (mentioned in the note after thedescription of longicomis, which lacks it) ; it does not occur in thegenotype of Exetasis. There is a marked tendency toward the devel-opment of adventitious cross veins, which apparently have veryslight taxonomic value. Considerable variation exists in the apicalpart of the first posterior cell, which may be petiolate or closed inthe margin, or partially coalesced with the second posterior by thedisappearance of the last part of the fourth vein, or widely open.In drawing up a key largely from figures and descriptions, I hadmade considerable use of these differences, until I observed that mynew species differed in the two wings of the single specimen andwould run to what I had supposed were two groups of species. Thisdiscrepancy compelled me to view these differences with more caution.The nearest related genus is Apelleia Bellardi.^ The only speciesincluded was mttata, new, from Mexico; it has bare eyes, and thiswas put forward as the main difference from Ocnaea. Osten Sackendescribed in Ocnaea a species {grossa) with bare eyes, indicatingthat the character is not of generic value; but it seems so to me, andI therefore transfer grossa to Apelleia. I recognize the fact thatgreat caution should be exercised in proposing new genera in thefamily, where great plasticity seems to exist in characters whichwould elsewhere be of generic or even familj^ rank. The problemis to find constant characters, and this implies the examination ofconsiderable series in many species?a requirement which can hardlyas yet be met in any collection. It may be that Exetasis will ul-timately be restored to rank on the absence of the fork of the thirdvein; it would include tu7nens Walker (genotype), calida Wiede-mann, and longicomis Erichson.In preparing the following key, I have been able to examine onlyschwarzi^ gigas, falcifer, flavipes, and trivittata (all in type mate-rial) ; Cole's figures and those of Wiedemann, Erichson, Walker, andOsten Sacken have been of great assistance, and I have studied thedescriptions closely. Nevertheless it must be considered as a pre-liminary effort. Cole has given a key to the five species from NorthAmerica, which he had seen, in the reference cited above. I stillhave misgivings as to the distinctness of m}^ gigas and falcifer,which may be a single unusually variable species.In response to a request from me. Dr. G. Enderlein very kindlysent me notes on the types of three species in the Berlin Museum(micans, longicomis, and luguhris), together with excellent figuresof the venation in each case.
? Mem. Acad. Sci. Torino, vol. 21, p. 214, 1861 (or " Saggio di Ditt. Mess.," appendix,p. 17).
ART. 9 DIPTERA FEOM AMERICA, ASIA, AND JAVA ALDRICH 6KEY TO SPECIES OF OCNAEA
1. Third antennal joint clavate 2Third antennal joint of uniform width, slightly tapering 32. Legs brown, knees yellow (Mexico) micans ErichsonLegs yellow (Texas) loewi Cole3. Third antennal joint with scattered hairs on upper edge(Panama) trichocera Osten SackenThird antennal joint bare 44. Third vein not forked 5Third vein with widely divergent anterior fork near tip, aboutas in Tabanus 75. Mesonotum luteous, legs brown (Brazil) calida WiedemannMesonotum black on disk, legs yellow 66. Scutellum yellow, third antennal joint comparatively short, lessthan height of eye (Brazil) tumens WalkerScutellum black, third antennal joint very long, exceedingheight of eye (Brazil) longicornis Erichson7. Femora yellow 8Femora more or less infuscated 118. Mesonotum yellow (Texas) auripilosa JohnsonMesonotum black, or with distinct dark area 99. Second, third, and fourth abdominal segments reddish yellowwith a decreasing row of black spots in middle (Honduras).flavipes AldrichSecond, third, and fourth abdominal segments with basal trans-verse black bands 1010. First posterior cell petiolate by the union of the third andfourth veins before margin ; adventitious cross veins presentin marginal, and first and fourth posterior cells (Texas).helluo Osten SackenFirst posterior cell united with second near tip by the disap-I)earance of the fourth vein near base of second cell ; no ad-ventitious cross veins (the one dividing the first posterior isnot here regarded as adventitious) (Cuba) schwarzi Cole11. Mesonotum yellow with three broad black stripes confluentbehind, the outer much abbreviated in front ; scutellum darkyellow (Honduras) trivittata, new speciesMesonotum wholly black or blue except more or less of lateralmargins ; scutellum concolorous 1212. Abdomen metallic blue with narrow interrupted apical yellowcross bands on second, third, and fourth segments (Texas) ?coerulea ColeAbdomen brown, hind edges of same segments indistinctly paler(Brazil) lugubris GerstaeckerAbdomen without cross bands, except a broad one on secondsegment 1313. Blackish in color, with reddish third antennal joint (Ecuador).falcifer AldrichBrown in color, third antennal joint almost black (Ecuador)?gigas Aldrich
4 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 81OCNAEA TRIVITTATA, new species
Third vein forked at apex; on one side the fourth vein is incom-plete, on the other it joins the third just beyond the middle of its lastsection, making the first posterior cell petiolate. Coxae brownish,femora irregularly brownish, .not very dark ; tibiae and tarsi yel-low, apical half of last tarsal joint and the claws black; pulvilliyellow.Male.?Head rather large, eyes with dense yellow pile; palpi andproboscis very minute, not visible; antennae reddish brown, firstjoint black above, third slender, longer than height of head, andlonger than front tibia.Thorax and abdomen with dense erect yellow pile, venter bare;mesonotum yellow with three broad black stripes confluent behind,rounded in front, the outer much abbreviated anteriorly; pleuraebrown, both spiracles white; scutellum brownish yellow; calyptersroughened, margin brownish yellow with pale fringe.Abdomen shining blackish brown, first four segments with whit-ish hind margin of uniform width, that on first segment narrow,on the following ones conspicuous, not widening laterally except atextreme edge, where they include half the segment; they extendacross venter at the width which they assume at the edge. Fifthsegment with narrow pale hind border, sixth with only a lateralpale spot.Wings small ; the cross vein in the first posterior cell is more thanits length beyond the discal; no indications of other unusual crossveins.Length, 9.5 mm.Type.?MQl^^ U.S.N.M. No. 43480, collected by Dr. J. Bequaertat Sangrelaya, Honduras, March 13, 1924, and presented by him tothe National Museum.Remarks.?Described from one male, the type. This is the onlyspecies of Ocnaea with distinct thoracic stripes.
Family DOLICHOPODIDAECOLLINELLULA, new genusVery minute. Posterior half of mesonotum depressed. Antemiaeminute, third joint rounded, with af)ical bare arista; first antennaljoint bare; head concave behind; proboscis and palpi small. Ocel-lars large, proclinate, divergent, situated high up ; one outer vertical,and one frontal near eye. Face narrow in female, the eyes almosttouching near epistoma ; in male the ej^es are contiguous below theantennae to the mouth. Thorax with two rows of rather strongacrostichals ; two dorsocentrals on each side near scutellum, the row
AHT. 9 DIPTERA FROM AMERICA, ASIA, AND JAVA ALDRICH 5
continuing forward as coarse setules, not much larger than the acro-stichals; scuteUuni with a single pair of bristles, far apart. Nobristle on outer sioe of hind coxa. Male with globose and somewhatexserted hypopygium, the abdomen rather cylindrical. Wing ofmale (pi. 1, fig. 2) with distorted venation; in the female (pi. 1,fig. 3) the first vein is short, the second parallel with costa nearly totip, third vein ending just at tip, fourth diverging moderately, end-ing as far from tip of third as the second does, or very little farther
;
hind cross vein behind the center of the wing, a little shorter thanlast segment of fifth vein; basal cells and sixth vein absent.Related to Achalcus, but that has five pairs of acrostichals, twobefore the suture. In head structure the new genus is very muchlike Thrypticus^ but that has the venation quite different, the hindcross vein small and retracted, and the anterior part of the thoraxis peculiarly bulging and prominent above. Thrypticus also hasbetter developed dorsocentrals.Named in honor of J. E. Collin.Genotype.?CoUinellula magistri, new species.COLLINELLULA MAGISTRI, new speciesPlate 1, Figures 2-4Male.?Ver}'^ minute; dark blue-green; legs, antennae, and palpiblack. Venation as figured, which alone would make the speciesinstantly recognizable. The groups of long hairs give under lowpower the effect of slight clouds in the wing. Front tarsi compli-cated in structure; the first joint rather thick and short, widenedapically; the second shorter and paler, forming an irregular collar;third somewhat like the second but excised below, the excision partlyclosed by a transverse plate, and the upper side of the segment witha striking spine at apex; fourth segment very short and tapering;fifth segment as long as the preceding three, very narrow at base,rapidly widening into a triangular flat shape, with dense fine hairand the usual claws and pulvilli. The tarsus is described from aspecimen mounted in balsam, its small size making the details prac-tically impossible to see otherwise. Abdomen deep green, the globosegenitalia shown in posterior view and spread out (pi. 1, fig. 4), wherethe rest of the structures are shown in the side view, considerablypressed down. The two median ventral organs anterior to the hypo-pygium are especially noteworthy. Middle femora with a longbristle at base beloAv,Length, 1.2 mm.Female.?Like the male, but the wings and tarsi are normal, andthe eyes do not come entirely together below the antennae, exceptjust at the mouth.Length, 1.1 mm.
6 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 81
Paratypes.?M.2i\Q and female, U.S.N.M. No. 43658.i?6marZjs.?Described from 11 males and 9 females, all collectedat Taughannock Falls, near Ithaca, N. Y., on August 19, 1928, byJ. E. Collin, of Newmarket, England ; and 1 female collected by meat Washington, D. C, on July 1, 1920. In the specific name I meanto celebrate the ability of Mr. Collin as a coUector.^^Four males and three females of the Ithaca material are retainedin the National Museum, a gift from Mr. Collin; the rest of theseries, including the type, is returned to him.A single female in the National Museum, collected by me at LaFayette, Ind., August 24, 1916, seems to belong to another species ofthis genus. Genus LEPTOCERA OlivierLeptocera Oliviee, Mem. Soc. Agr. Dept. Seine, vol. 16, p. 16, 1813.?CoQuiLLmT.Proc. U. S. Nat. Mus., vol. 37, p. 559, 1910.LEPTOCERA (LIMOSINA) OPACA, new speciesHead, thorax, and abdomen black. Front opaque, two-thirds thewidth of head, with two pairs of convergent bristles back of vertex(one outside the other) ; a small divergent pair just behind ocelli;outer verticals reclinate, inner large and convergent but not decus-sate; ocellars proclinate and divergent; two divergent orbitals eachside; three pairs convergent bordering middle region, lowest justabove suture. Antennae black, decidedly divergent, separated by thetriangular lunule ; arista long and slender, pubescent. Face shiningblack, concave, epistoma protruding, narrow, prelabrum visible butnot prominent ; a large bristle laterally below end of sirture at edgeof mouth, directed forward. Cheek at narrowest hardly one-halfthe eye height, bare except a few hairs below. Mouth large. Meso-notum opaque, nearly circular, with 14 regular rows of hairs ; suturenot distinguishable. Thoracic bristles as follows: Dorsocentral, 1;prescutellar, 1 small; humeral, 1; notopleural, 1; presutural, 1;postalar, 1; mesopleural, 0; sternopleural, 1; scutellum flat, moder-ately long, bare, with two pairs of marginals. Halteres brownishyellow. Abdomen opaque black. Legs black. Coxae and trochan-ters yellow, the former more brownish; tarsi brownish, middle andhind often yellowish. Mid tibia of female with one smallish ventralbristle on middle and large apical on same side (both lacking inmale), also two widely spaced bristles in both sexes on anterior dor-sal surface and two beyond middle and nearly side by side on pos-terodorsal. Wings with slight infuscation, moderately rounded;costa without striking bristles, the segment between tips of first andsecond veins equal to the following; cross veins strikingly approx-
?? Dr. G. Enderlein, of Berlin, Germany, also distinguished himself as a collector bycapturing both sexes of the same fly on the same occasion, as I learned from a letterreceived after the galley proofs of the present paper had been corrected.
ART. 9 DIPTERA FROM AMERICA, ASIA, AND JAVA ALDRICH 7imated, their distance apart on fourth vein generally less than halfthe length of hind cross vein ; third vein almost imperceptibly curvedforward toward tip, ending only a little before tip of wing; costaextending distinctly beyond it, fourth and fifth evanescent fromslightly beyond hind cross vein.Length, 1.5-1.6 mm,Type.?M.^\Q, U. S. N. M. No. 42847.Remarks.?Described from 16 mounted and 14 alcoholic specimens,collected February 2, 1930, in a dahlia cellar at Fort Collins, Colo-rado, and sent to the Museum by Sam C. McCampbell, Deputy StateEntomologist.The species is exceedingly like heteroneura Haliday of Europe,which I have not seen. Duda's full description of heteroneura *seems to leave few differences to note. It has the second abdominalsegment in the male much elongated, but ours has it hardly any longerthan the third or fourth. In the European species the face is dirtyyellow; in ours it is black. These are the chief differences I see.The wing of opaca agrees with Duda's figure.This species would go in the subgenus Scotophilella Duda.^ Thissubgenus, however, is a synonym of Lhnosina Macquart,? as alreadypointed out by Richards.'^Family OTITIDAE(Ortalidae of authors)The generic name Ortalis being preoccupied in birds, and in factin common use there, the family should be based upon the still oldergenus Otites Latreille^ (type Musca formosa Panzer). This genusbelongs to the subfamily Ortalinae of authors, hence the change toOtitinae does not affect the other subfamilies,DYSCRASIS, new genus
Belongs in subfamily Pterocallinae, but is widely different fromany known genus. The presence of five pairs of well-developeddorsocentrals, and of the same number of equally large acrostichals,one pair of each being before the suture, separates the genus fromall known to me. Numerous venational peculiarities, which areimpressively shown in Plate 1, Figure 1, also emphasize the distinct-ness of this form, I was at a loss to place it in any subfamily, andreferred a specimen to Professor Hendel, whose extensive publica-
*Abh. K. K. Ges. Wlen, toI. 10, no. 1, p. 188, 1918.?Abh. zool.-bot. Ges., vol. 10, p. 34, 1918.
? Histoire naturelle des insectes, Diptferes, vol. 2, p. 271, 1835.
' Proc. Zool. Soc. London, 1930, p. 291.
' Histoire naturelle, g6n6rale et particuli&re des crustac^s et des insectes, vol. 14, p. 383,1805.
8 PEOCEEDINGS OF THE NATIONAL. MUSEUM vol. 81
tions in the group are well known; he places it as a peculiar formof the above subfamily.Head broad and short (flattened from before) ; frons wider thanone eye, with parallel sides, flat; face receding below antennae,slightly protruding at epistoma, without antennal grooves ; antennaewith both basal joints very short, third oval, shorter than face, witharista microscopically pubescent. Proboscis short, palpi of ordinary
size, flat. Cheek a little more than half the eye height. Postverticalbristles divergent ; other head bristles are inner and outer verticals
;
two fronto-orbitals, reclinate, the anterior halfway between innervertical and lunule; and the usual ocellars, which are of good size.Ocellar triangle small.Thorax narrower anteriorly than the head, almost bare of smallhair, with the following bristles: Acrostichal, 1,4; dorsocentral, 1,4;prothoracic, ; humeral, 1 ; notopleural, 2 ; supraalar, 2 ; intraalar, ;postalar, 2 (the outer might be taken for a low supraalar, and theinner for a high one) ; mesopleural, 2 small on hind edge; sterno-pleural, 1 ; scutellars, 2 pairs. Scutellum swollen, polished.Abdomen broader than thorax, sixth joint in female (base of ovi-positor) broad, flat, shining.Legs not elongated, without noticeable bristles, except a doubleposterodorsal and a single longer posteroventral row on anteriorfemora, and a row of small ones on middle of anterior side of midfemora.Wings (pi. 1, fig. 1) with striking pattern somewhat resemblingthat of the trypetid Ceratitis capitata Wiedemann. Costa withouta break ; costal cell very wide, the auxiliary vein sinuous, ending farbefore tip of first vein; first vein long, hairy above on apical half;second and third veins converging beyond small cross vein, thendiverging; anal cell with a long acute extension reaching two-thirdsof the way to the wing margin, and narrowed at its base.Genotype.?Dyscrasis hendeli,) new species.
DYSCKASIS HENDELI, new speciesMale.?Frons gray pollinose, its lower half or less shining brown
;
ocellar triangle also pollinose. Face except immediately below theantennae white in ground color with very thin white pollen; cheeksand lower back of head the same but not quite so white. Thoraxwith two pairs of large polished black spots, above the notopleuralarea and wing, one before and the other behind the suture; thesespots show a little velvety in some lights on their outer edge. Scu-tellum swollen and polished black. Kemainder of dorsal regionwith thin gray pollen, except a small, opaque, black semicircle justbefore the scutellum and traces of dark spots at the bases of the
ART. 9 DIPTERA FROM AMERICA, ASIA, AND JAVA AliDRICH 9
bristles. Upper half of pleura and notopleural region with whitepollen forming a sharply limited band, which passes around themetanotum below the small infrascutellum ; lower half or a littleless of the pleura brown. Abdomen shining black, the dorsum ofthe second segment, however, white pollinose, and a band of whitepollen on hind edge of third and fourth segments, widest on middleof third segment. Legs brownish yellow, the femora nearly blackand tarsi yellow, except near tips, where they become brownish.Wings as shown in figure and as described under the genus ; the colorpattern is brown, but yellow in a considerable area around the an-terior cross vein.Length, 3.5-3.7 mm.Female.?Like the male, but the fifth segment is white pollinosedorsally for about two-thirds of its length from the base.Length, 4.2 to 5 mm.ry^e.?Male, U.S.N.M. No. 43575.Remarks.?Described from 17 specimens of both sexes: Eightspecimens, including type and allotype, are from Dallas, Tex., col-lected by F. C. Bishopp, the dates being March 29, 1907 ; March 17and 20, 1908; and May 30, 1908; one male, Uvalde, Tex., in trap,November 16, 1915 (Bishopp No. 5672) ; one male and three females,Matamoras, Mexico, in traps (T. R. Stephens) ; one male, Mercedes,Tex., in trap, August 26, 1931 (W. R. Heard) ; one male, San Benito,Tex., on office window. May 26, 1930 (L. F, Greer) ; one female,Allen, Tex., August 14, 1931 (F. O. Swan) ; one female, Texas, noother data. Family RHOPAEOMERIDAE
This family has been revised by Professor Lindner in DeutscheEntomologische Zeitschrift, 1930, pp. 122-137.
Genus KROBERIA Lindner
Kroieria Lindnee, Deutsche Ent. Zeitschr., 1930, p. 127.The only species known is fuliginosa Lindner from southern Brazil,described in the same place. The arista is bare, the scutellum notprolonged into a blunt, shining, knoblike tip, but rounded, flatabove; front flat, not concave, covered with hairs except on theocellar triangle, which extends narrowly forward to the lunule;one or two small orbitals present; anterior edge of front somewhatoverhanging the antennae in a transverse rim. Auxiliary vein welldeveloped; fourth vein ending just before tip of wing, rather closeto third.
10 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 81KROBERIA FLORIDENSIS, new speciesDark brown, legs yellowish brown, basal and apical third of tibiaeand tarsi from tip of first joint blackish. Differs from fuliginosain having the scutellum opaque brown with pale pollinose border ; thehind femora only moderately thickened, with only a few bristlesabove ; and the hind tibiae only slightly flattened.Male.?Front black in ground color, with thin pale pollen, ocellartriangle more densely pollinose ; a silvery pollinose spot between eyeand third antennal joint; face dark yellow, the protuberance some-times blackened, the epistoma below it transversely wrinkled ; cheekbrown, wrinkled, nearly as high as the eye, with pale hair. Mesono-tum black, with five distinct pale pollinose stripes ending atscutellum; mesopleurae with pale pollen, covered with small darkdots where the hairs arise, but these do not extend upon the portionbelow the spiracle. Chaetotaxy: Acrostichal, 1 (prescutellar)
;
dorsocentral, 1 or 2 (far back); humeral, 1; notopleural, 2; pre-sutural, 1; supraalar, 1; postalar, 2; intraalar, 1 (before the innerpostalar) ; scutellar, 2 pairs (a third in one male and an odd one ina female) ; mesopleural, 2 or 3 ; sternopleural, 1. Most of thesebristles are quite small. Abdomen blackish, dorsum opaque, secondand following segments with interrupted band of pale pollen onposterior half or more, widening toward sides to include whole lengthof segment, most distinct from behind. Legs as described; hindfemur with anterodorsal row of bristles from base nearly to tip, andtwo short rows near tip, none of striking size. Wings ratheruniformly pale brownish. Knob of halteres very pale yellow.Length, 6-6.6 mm.Female.?Abdomen with pale pollinose stripe on each side, centralregion opaque dark brown; sixth and seventh segments shiningblack, the latter becoming dark yellow toward tip.Length, 7 to 8 mm.Type.?KslQ, U.S.N.M. No. 43761.ReTYiarks.?Described from two males and four females, rearedfrom rotten wood of Sabal palmetto in Putnam County, Fla., byMark Dodd, February 21, 1931; and one female reared by D. J.Nicholson from a larva found in deca3'^ed wood of Sahal minor^ 6inches under water, l^/^ miles east of Fort Christmas, Fla., emergedMarch 25, 1931.The discovery of this species in northern Florida, only about 70miles from the Georgia line, extends the known range of the familyin a remarkable way. Lindner recognizes about 13 species in 6genera. All these are distinctly tropical, and nearly all rangesouthward from the northern limits of South America to Paraguay
ART. 9 DIPTEEA FROM AMERICA, ASIA, AND JAVA ALDRICH 11
and northern Argentina. None have been reported from the WestIndies, nor from the United States.All the references to occurrence from Panama north are the fol-lowing :Rhinotora diversa Giglio-Tos was described from Tuxpango,Mexico.Rhopalouiera wanthops Williston was described from Yucatan;the collector being Gaumer, the species was probably taken nearMerida.Rhopalotnera femorata Fabricius, described from South America,is reported by Lindner from Guatemala and Mexico.WiUistoniella pleuropunctata Wiedemann, described from SouthAmerica, is reported from Playa Vicente, Mexico, by Giglio-Tos;from the Volcano Colima, Mexico, by Lindner; and I have identi-fied it from Corozal, Canal Zone, where Mr. Zetek bred it fromtrunk of coconut palm.Genus SCATOPHAGA Meigen
Scatophaga Meigen, Illiger's Mag. fiir Insekt., vol. 2, p. 277, 1803.SCATOPHAGA GIGANTEA, new speciesA very large species, the male with dense long fulvous hair onabdomen, legs, and pleurae ; hair on mosonotum black.Male.?Front prominent, angular at antennae; eye oblique; cheekhigh, especially behind, about equal to height of eye; back of headvery bulging; palpi a little shorter than proboscis. Front broad,with a narrow black orbit covered with yellow pollen ; frontal stripealmost half the width of the head, deep red, the middle portionblackish and having a bluish reflection; parafacial and anteriorhalf of cheek red with bluish reflections ; antennae black, third jointhardly twice the second and reaching five-sixths of the way to prin-cipal vibrissae; arista pubescent; palpi reddish yellow, with longblack hairs below from near base to tip ; proboscis black, its princi-pal segment shorter than height of head. Back of head with blackhair on upper half or about to lower edge of eye, the rest coveredwith the same long fulvous pile as the pleurae and femora. Innervertical bristles long, oeellars slightly shorter, the row of frontals
still shorter, all slender. About a dozen slender bristles in a rowfrom epistoma to and a little above the main vibrissa.Thorax black with dense fulvous pollen above and on the scutel-lum and pleurae ; mesonotum with erect long delicate black hair anda few bristles; the most distinct bristles are the posterior notopleu-ral; three supraalar; on intraalar, far back; two postalar; and
12 PROCEEDINGS OF THE NATIONAL MUSEUM vol.81
the posterior pair of dorsocentral, far apart. Scutellum with threepairs. Sternopleural with one posterior. The long fulvous pilecovers the mesopleura, propleura, and sternopleura, leaving thepteropleura and all behind it bare. Calypters with fulvous rimand marginal hairs, which are very long on the fold.Abdomen black in ground color, with very striking long densefulvous hair, which seems even more brightly fulvous than that ofthe sides of the body and the femora. This dense covering extendsover the sides of the abdomen and even somewhat underneath, sothat no bare area below is visible.Legs reddish yellow, the coxae and basal three-fourths or more offemora black. Coxae with dense long fulvous pile to the outer hindside of the middle ones, this area and all the outer side of the hindones bare; femora and tibiae with the same pile as the pleurae, butit is mainly black close to the apices of the femora, and considerablymixed with black on the extensor side of all the tibiae. Tarsi withcomparatively short black hair above, about as long as the depth ofthe segment. Pulvilli brown, claws black. The middle tibia hastwo spines on the anterodorsal side, three or four on the postero-dorsal, two on the posteroventral, and one on the anteroventral, notincluding the apicals. The hind tibiae have three spines on theanterodorsal and two on the posterodorsal. None of the femora havespines.The wings are distinctly yellow, more intense near base.Length, 12-15 mm.FeTnale.?The characteristic red color of front, parafacials, andcheek as in male ; the pile, which is so striking in the male, is reducedto insignificant size, though present. The mesonotum is deep ful-vous, with the following chaetotaxy : Acrostichal, 4 delicate rowsof hairs ; dorsocentral, 2, 3 ; humeral, 2 ; presutural, 1 ; notopleural, 2
;
supraalar, 2; intraalar, 1 far back; postalar, 2; sternopleural, 0, 1;scutellum, with 2 pairs marginal. Abdomen depressed, with densegray pollen, seventh segment wholly shining, except hind edge,which is notched in the middle above. Legs reddish yellow, only thefront femora a little black above. No bristles on front femora;middles ones with two on front side beyond middle and two neartogether near tip on hind side; hind femora with irregular row ofabout six small on anterodorsal side. Front tibiae with two bristleson front side; middle tibiae with two anterodorsal, two postero-dorsal, and one anteroventral; hind tibiae with three anterodorsaland posterodorsal.Length, 9 mm.Type.?M^\Q, U.S.N.M. No. 43G92, from Yu Long Gong, Tibet,August 1, 1923. Allotype from same lot.
ART. DIPTEEA FROM AMERICA, ASIA, AND JAVA ALDRICH 13
Remark's.?Described from 63 males and 22 females, all collectedby Dr. D. C. Graham, in the high conntry of western China alongthe Tibet border and in the edge of Tibet. Ten males and onefemale were collected August 1, 1923, at Yu Long Gong, Tibet,near Tatsienlu, altitude 12,000 feet; 13 males and 3 females at thesame place, August 12, 1930; 12 males 9 miles south of Tatsienlu,June 26 and 27, 1923, altitude 8,500 to 13,000 feet ; 12 males and 1female at Yu Long Si, July 26-28, 1930, altitude 14,000 to 15,900feet; 9 males and 1 female in Yu Long Si Gorge, 13,000 to 15,000feet, no date ; and 7 males and 16 females near Wa Hu Pass, Tibet,August 6 and 7, 1930, altitude 14,000 to 16,000 feet.In chaetotaxy the species resembles Scatophaga vulpiiia Coquil-lett, from Point Barrow, Alaska. It is the largest species of thegenus as far as I know. The lot first cited was labeled, '* Fond ofcowdung." SCATOPHAGA GIGANTEA OBSCURA, new varietyA series of males differ from typical gigantea in being smaller, thepollen of the dorsum dull brown, pile of pleurae and abdomenshorter and darker, that of the femora and tibiae almost whollyblack, and shorter than in the typical form. The appearance is sodifferent that it is hard to believe the relationship so close as itappears on closer study. The head structure is the same, with thecharacteristic deep red color of front and face, but the pale pile ofthe back of the cheek is much less conspicuous. I am not able torecognize a corresponding series of females, as these males approachthe typical females in their appearance, except for having in generala darker color of the thoracic pollen.ry^?e.?Male, IT.S.N.M. No. 43693, from Yu Long Gong, Tibet.Length, 8 to 9 mm.Remarks.?Described from 14 males collected with typicalgigantea as above cited. Ten are from Yu Long Gong, three fromWa Hu Pass, and one from Yu Long Si.SCATOPHAGA AMPLIPENNIS Portschinsky
Scatophaffa ampliijcnnis Portschinsky, Horae Soc. Ent. Ross., vol. 21, p. 199,1887.Portschinsky's description is so brief that it seems worth whileto give a fuller one, as a long series is available from DoctorGraham's collecting-.A slender blackish species with inconspicuous pile, but with verylong and broad wings, which are uniformly dark brown in color.It agrees with Scatophaga scyhalaria Linnaeus in having the attach-
14 PROCEEDINGS OF THE NATIONAL MUSEUM vol.81
ment of the abdomen distinctly elevated above the hind coxae, sothat the sclerites above the latter (metathoracic epimerit of Hendel)are completely united on the middle line, not separated by a mem-branous portion as in other species. It differs from scyhalarixi in.having the arista pubescent, not pilose, in much darker color, andin other respects.Male.?Bliick in ground color, including femora; tibiae reddishyellow, tarsi brownish yellow; palpi yellow with brownish apices;frontal stripe very dark red with a glaucous reflection. Head darkbrown above, the frontal stripe narrowed upward, the parafrontalportions wide, elevated in middle, with dense long slender bristles,of which about half a dozen of the upper series turn out over theeye, the lower ones being mesially directed. Parafacials reddish onupper half along facial ridges, elsewhere with brown pollen. Vibris-sal region with dense row of bristles. Antennae wholly black, thirdjoint less than twice the second, arista black, densely pubescent.Cheek two-thirds as high as eye, which has a somewhat diagonalposition, the cheek with upturned slender hairs below, its posteriorhalf with long and mostly yellow hairs. Palpi infuscated and withmany black hairs beyond the middle, just before the middle the hairsare yellow. Thorax with dark pollen and a median glaucous stripereaching the scutellum, on each side of this a partly double blackish,subshining stripe ; scutellum glaucous on middle, connecting with thethoracic stripe. Chaetotaxy: Acrostichal, only fine, erect hairs allthe way; dorsocentral, 2 hairlike anteriorly, about 3 bristles pos-teriorly, the hindmost gradually larger; notopleural, 2; presutural,1; supraalar, 2; postalar, 2; intraalar, 0; sternopleural, 0, 1; scutel-lum, with 2 lateral pairs and an equally long apical pair close to-gether. Pleurae with rather dense but slender long brownish hair,Calypters brown, smallish, with fringe of dark yellow hair. Ptero-pleural and all behind the sternopleura bare. Halteres with brownloiob.Abdomen narrow, not much deflexed, shining brownish black, cov-ered above and on sides with dense, erect, but rather short yellowpile, which is more or less mixed with black close to tip; this pileseems to cover the under side also.Legs with erect but not very long pile, which varies on the femorafrom black to brown and a considerable admixture of yellow; fronttibiae with long black pile on outer side, middle and hind ones withthe same standing out all round; front tarsi with the three inter-mediate joints short, of equal length; pulvilli blackish, claws black;middle tibia with two anterodorsal spines beyond middle, twoposterodorsal, one flexor far down toward tip ; hind tibia with threeanterodorsal, three posterodorsal (one near base).
ART. 9 DIPTERA FEOM AMERICA, ASIA, AND JAVA ALDRICH 15Wings strikingly long and wide, and of uniform dark browncolor, only the anterior part near base gradually a little yellowish.In a male measuring 9.5 mm, the wings measured 12.5 long and 5wide at middle. First posterior cell not narrowed at tip, hindcross vein bisinuate.Length, 7.5-10 mm.Fetnale.?Wing not so strikingly enlarged; in a specimen 8 mmlong the wing measured 8 mm long and 2.5 mm wide; wing color alittle paler than in male ; abdomen depressed as in most species andmuch wider across middle than in the male. Front tibia with twohairlike extensor bristles ; middle tibia with two anterodorsal spines
;
three posterodorsal ; one flexor; hind tibia with three anterodorsal;three posterodorsal.Length, 6.4-8 mm.Remarks.?Redescribed from 35 males and 13 females, collected byDr. David C. Graham in the high altitudes along the Tibet-Chinaboundary. Four males were collected in Yellow Dragon Gorge nearSong-Pan, altitude 12,000 to 14,000 feet, in the summer of 1924; allthe others were collected at Yu Long Si, near Tatsienlu, July 26-28,1930. altitude 14,000 to 15,900 feet.
Genus SARCOPHAGA Meigen
Sarcophaffa Meigen, Systeinatische Besclireibung, vol. 5, p. 14, 1826.SARCOPHAGA (BLAESOXIPHA) VALANGAE. new speciesFigure 1
Male.?First vein bare, posterior dorsocentrals generally three,but a small or even a normal additional one may occur as the secondbehind the suture ; without villous hairs on inner side of hind tibiagenital segments dull black or dark brown.Front 0.18 of head width, at narrowest, above middle (four meas-ured 0.18, 0.18, 0.19, 0.18) ; ocellars normal, proclinate ; outer verticalnot differentiated; two upper frontals reclinate but the second pairpartly convergent; eight other frontals in row, which extends tonear tip of second antennal joint and diverges toward eye; para-frontals narrow, brownish above, subsilvery below; frontal stripebrown; parafacials subsilvery, a changeable dark spot in the pollenat lowest frontals, the parafacial hairs rather numerous and bristlybelow; facial ridges broadly curved outward in middle of the face,converging markedly below, with small irregular setules extendinghalfway up ; middle of face blackish ; cheek two-fifths of eye height,with black hair except on hind edge ; back of head with black hair,only a little pale about neck and below. Palpi black, proboscis or-dinary. Antennae black; third joint one and one-half times the110571?32 2
16 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. SI
second; arista with rather short plumosity for less than half itslength.Thorax gray, with three black stripes and a partial one above thewing extending a little before the suture, the median stripe extend-ing nearly to tip of scutelluni ; at least three pairs of large and some-what irregular anterior and four pairs posterior acrostichals ; sterno-pleural 3; scutellum with three pairs lateral, one discal, the apicalpair quite large, subdepressed, decussate. Hind calypter distinctlybrownish.Abdomen gray with medium black stripe becoming slender onfourth segment, and one less distinct lateral black stripe each side,changeable on third segment and ending at middle of fourth ; a pairof median marginals on second segment, marginal row on thirdand fourth. Genital segmentsbrownish, small; genitalia as inFigure 1. Forceps flattened be-hind, brown basally; outer for-ceps (accessory plate of Parker)rather large for a Sarcophaga.brown basally ; penis verj^ small.In the iigure the anterior andposterior claspers change places,crossing each other at base.Fifth sternite deeply cleft, with-out striking characters.Legs black, front tibia withone bristle on hind side; middletibia with one on outer front.Wings subhyaline, costal segment before first vein equal to the onebeyond second ; third vein with bristles extending more than half-way to anterior cross vein.Length, 7 mm.Female.?Much grayer, the black thoracic stripes narrowerand the median one hardly visible on scutellum. Front near vertex0.29 of head width (four measured 0.29, 0.3, 0.28, 0.28) ; parafacialswith fewer setules: abdomen more tessellated, median stripe muchless distinct, the lateral hardly noticeable. Second segment withoutmedian marginals, at most with depressed bristly hairs. Terminalventral sclerite yellowish, a little elongated, tapering, slightly curveddownward. Posterior dorsocentrals uniformly three.Length, 6 mm.Ty^e.?Male, U.S.N.M. No. 43689.
Figure 1.
?
Saroophaga {Blacsoxipha) va-laiigac, new species. Side view of malegenitalia with forceps from behind.(Drn^vn by David G. Hall)
AET. 9 DIPTERA FEOM AMERICA, ASIA, AND JAVA AL.DRICH 17Remarks.?Described from five males and 15 females, reared fromthe locust Valanga nigricornis Burmeister, at Gedangan, centralJava, by F. Verbeek, December, 1929, and sent to the National Mu-seum by Dr. S. Leefmans, of Buitenzorg, to whom a pair of para-types are returned.The species is very similar to those that in Europe have been re-garded as belonging to the genus Blaesoxipha. The male is readilydistinguishable from all European forms known to me, in having theforceps much broader toward the base and the anterior claspers witha lobe on the front near the middle.
Genus LESKIOMIMA Brauer and Bergenstamm
Leskiomima Braueb and Bergenstamm. Zweifliigler des kaiserl. Museums zuWien, pt. 5, p. 372, 1891 (Deukschr. kaiserl. Akad. Wiss., vol. 58).LESKIOMIMA JAYNESI, new speciesA slender yellowish fly, differing from the type species tenera inhaving only one to three coarse hairs at middle of first vein insteadof a full series on all its length becoming more densely placed neartip. Palpi of ordinary size ; proboscis (last joint) long and slender(one and one-third times the head height), curved downward, withsmall labella. Eyes bare. Face slightly receding, epistoma notprominent. Ocellars present, proclinate and divergent ; a single pairof large verticals ; two upper frontals reclinate, the uppermost, how-ever, very small, the lowest at middle of second antennal joint ; para-facial about as wide as third antennal joint. Cheek two-fifths theoye height. Front in both sexes wide, about 0.35 of head width, andwith proclinate orbitals in both; pollen of head mostly white, butnot silvery. Antennae red, third joint blackened on apical two-thirds; second joint about half the third. Arista black, stronglypubescent. Mesonotum mostly black in ground color, with j^ellowishpollen, forming indistinct stripes. Acrostichal, 2, 1 (none just be-fore suture) ; dorsocentral, 2, 3; humeral, 2; posthumeral, 1; presu-tural, 1; notopleural, 2; supraalar, 2 (hind small) ; intraalar, 2; post-alar, 2 ; sternopleural. 2, 1 ; scutellum, with 1 lateral and 1 long,depressed, parallel apical. Pleurae yellow in ground color. Abdo-men yellow, shining, with distinct narrow basal band of pale pollenon segments 2 to 4; no median marginals on first segment, a pair onsecond, marginal row on third and fourth ; no discals even on fourth.Legs yellow, tarsi black. Wings with yellowish tinge, not at allelongated; fourth vein with rounded oblique bend ending not veryfar before tip. Costal segment before tip of first vein, considerablyshorter than that between tips of second and third; hind cross veinstraight and suberect.
18 PEOCEEDINQS OF THE NATIONAL MUSEUM vol. 81Male with small claws and pulvilli; genitalia yellow; a smallblackish spot at hind edge of third and fourth abdominal segments.Length of male, 5.5 mm ; of female, 5.6-7 mm.Ty^e.?Male, U.S.N.M. No. 43062.Remarks.?Described from 3 males and 10 females, all but one ofwhich were reared from the sugarcane borer, Diatraea saccharalisFabricius, at Tucuman, Argentina. Eleven were reared by H. A.Jaynes and one by H. E. Box. The other specimen was collectedby H. H. Smith at " Piedras B.," Brazil, in April ; it belongs to thecollection of the American Museum of Natural History, to which itis returned. SCHISTOCHILUS, new genuaRuns to Atractochaeta Brauer and Bergenstamm in Stein's 1924
" Key to the Central European Tachinidae," ^ but differs in numer-ous characters. Eyes bare, head in profile almost triangular, thelength at vibrissae being less than one-half of that at antennae.Front long, almost horizontal, in male a little more than one-thirdthe head width; face much receding, longer than front; third an-tennal joint elongated, four times the second, its upper edge straight,lower rounded forward at the apex, the sharp tip in line with upperedge. Parafacial bare, as wide as third antennal joint; facial ridgesbristly on lower third. Clypeus deeply depressed, forming a singlegroove for the reception of the antennae, which are out of sight inprofile when depressed; the groove continues to the mouth throughepistoma. Arista bare, shorter than the third antennal joint, verythick and blunt, the terminal joint shorter than the preceding one.Proboscis short and small, palpi rather small. Cheek two-fifthsof eye height. Outer vertical small, inner large, not decussate.Ocellars of ordinary size, proclinate and diverging. Frontals ina single row, one upper reclinate, the lowest at about the middle ofthe second antennal joint. One proclinate orbital in male. Proster-num with a row of black setules on each side, propleura bare. Scu-tellum with two pairs of laterals, the apicals small, depressed, notdecussate; postscutellum well developed, but joined rather closelyto the scutellum so that the transverse groove below it is muchdeeper than the one above. Abdomen of ordinary length, widerbeyond the middle than the thorax, no discal bristles. First veinof wing bare, third with a single bristle at base. First basal cellnarrow, the anterior cross vein short ; apical cell widening uniformlyto bend, which is rectangular and slightly rounded ; apical cross veinerect, joining third vein before the tip, the petiole more than one-third as long as the cross vein and ending far before wing tip.Squamae bare, rounded, not very large.Genotype.?Schistochilus aristatum., new species.^"
?Arch, fiir Naturg., vol. 90A, p. 19, 1924."The genus is of neuter gender, derived from the Greek oheiloa (lip).
ART. 9 DIPTERA FEOM AMERICA, ASIA, AND JAVA ALDRICH 19SCHISTOCHILUS ARISTATUM, new speciesColor black, mostly gray-pollinose.Male.?Frontal stripe brownish toward arista and the same colorextending vaguely down along the edge of the luniile. Pollen ofparafrontals and parafacials and posterior orbit dull gray. Palpiand basal joints of antennae dark yellow. Thorax black, with graypollen, leaving two slender submedian stripes interrupted at thesuture, an indistinct wider interrupted outer stripe. ChaetotaxyAcrostichal, 2, 4; dorsocentral, 3, 4 (the posterior may be only 3)humeral, 2; notopleural, 2; posthumeral, 1 (inner); presutural, 1supraalar, 2; intraalar, apparently 0; postalar, 2; sternopleural, 1, 1pteropleural small. Scutellum black at base and margin. Abdomenshining black, with basal gray-pollinose bands on second to fourthsegments, narrowed in middle on the first and covering about one-third of the segment except on the fourth where in certain lightsthe pollen extends almost to the tip ; first and second segments with-out median marginals; third with a depressed pair; fourth with arow at apex of varying sizes. Genitalia small, of ordinary structure,the outer forceps almost as long as the inner. Wing quite distinctlyinfuscated, the color following the veins broadly, leaving a sub-hyaline spot in the middle of the apical cell and the margin irregu-larly of the same color. Base of wing distinctly paler. Squamawhite. Costal spine small but distinct. The costal segment beforetip of second vein about one and one-third times as long as the onebeyond it. Hind cross vein concave outward, joining fourth veinbarely beyond middle between small cross vein and bend. Legsblack; claws and pulvilli not at all elongated; middle tibia with asingle bristle on anterodorsal side and one on ventral ; hind tibia witha single sparse row of bristles of varying size on anterodorsal side.Length, 5.8 mm.Ty^pe.?Male, U. S. N. M. No. 43690.Remarhs.?Described from three males, reared from Diatraeastriatalis Snellen, at Pasoeroean, Java, April, 1931, by Dr. P. C.Hart, received from Dr. S. Leefmans, to whom one paratype isreturned.I am unable to find a genus more nearly related to this thanAtractochaeta., from which it differs markedly in having the deepfacial groove for the antennae, the face more receding, apical crossvein more erect, and first vein bare and third with only a singlesetule, as well as in minor characters.
Genus ZENILLIA Robineau-Desvoidy
Zenillia Robineau-Dbsvoidy, Essai sur les Myodaires, p. 152, 1830.
?
Aldbichand Webbee, Proc. U. S. Nat. Mus., vol. 63, art. 17, p. 7, 1924.
20 PROCEEDINGS OF THE NATIONAL. MUSEUM vol. 81ZENILLIA PALPALIS, new species
Figure 2Black with ochraceous pollen on the head, thorax, and abdomen.Female.?Head 0.27 to 0.31 of the head width ; the frontal stripevelvet brown, as wide as one parafrontal; parafrontal, parafacial,and posterior orbit with almost golden yellow pollen, that of the faceand cheek gray, with a slight tinge of yellow ; front somewhat promi-nent at the insertion of the antennae, forming a slightly obliqueangle with the face, the frontal profile and the facial of about thesame length. Parafacial at middle as wide as third antennal joint,facial ridges with only three or four hairs above the vibrissae, whichare far apart at the oral margin. Cheek one-fifth the eye height.Antennae black, tip of second joint reddish, third joint long andslender, fully four times the second, rounded at tip, reaching almostto vibrissae. Arista rather long and slender, slightly thickened onbasal third. Palpi (fig. 2) considerablyswollen, with a very distinct and strikingpocketlike depression or hole on the outerside just before the tip, which is the same in
_ all the specimens. Inner verticals large,FiGDKB 2.
?
Zendlldu palpalis, ^ , .new species. Palpus of outer much smaller; ocellars of good size;female, outer side. (Drawn (^^.^ upper froiitals recliuate. the remainingby C. T. Greene) . ^^ ? ' i , ,in a Single row diverging below towardorbit, reaching to the base of third joint; the usual two proclinateorbitals; parafrontal with only inconspicuous hairs besides thebristles. Mesonotum covered with yellowish pollen, which formsprincipally three stripes on the sides and middle, leaving a ratherheavy black stripe on each side, which is divided anteriorly,the inner and narrower portion extending to the neck, Scutellumwith yellow pollen. Thoracic bristles rather stout. Chaetotaxyas follows: Acrostichal, 4, 3; dorsocentral, 3, 4; humeral, 4; noto-pleural, 2; posthumeral, 3; presutural, 2 (the inner large) ; supraalar,3; intraalar, 3; postalar, 3 (the middle one large, the others small)
;
sternopleural, 2, 1; pteropleural small. Scutellum with 3 pairslateral, 1 discal, the apicals slender and upturned, rather long.Propleura bare, prosternum setose, postalar declivity bare, no infra-squamal setules. Scutellum densely covered with coarse erect rathershort hairs. Abdomen largely shining black, the yellow pollenforming basal bands on the second and third segments, which arewide at the side and prolonged posteriorly in a blunt point in themiddle, leaving considerably more than half the surface shining;fourth segment with yellow pollen except an oval median spot above,which covers the apical two-thirds, but does not extend on the sides.
ART. 9 DIPTERA FEOM AMERICA, ASIA, AND JAVA AL.DRICH 21
First and second abdominal segments each bearing one pair ofmedian marginals ; second and third with one pair of smallish discals,which are quite far forward on the yellow pollinose portion; thirdwith a marginal row, the middle pair very large and stout; fourthsegment with irregular bristles extending forward nearly to thebase, no distinct row on posterior margin. The terminal ventral seg-ment when exposed appears as a cylindrical tubular structure, notvery long, bearing at its apex a minute, shining black prolongation,grooved above, bearing microscopic hairs. Legs black, middle tibiawith one bristle on the outer front side and one flexor; hind tibiawith an inconspicuous row of small bristles and two larger onesrather far apart almost dividing the length into thirds. Wings alittle brownish ; third vein with two to four bristles at base ; fourthvein with a round oblique curve, the apical cross vein more slender,straight for two-thirds of its length and almost parallel with themargin, then gradually curved toward the tip, joining the costaquite close to the tip a little distance from the third ; hind cross veinsuberect, joining fourth at two-thirds of the distance from the smallcross vein to the bend. Hind calypter light brownish with palemargin ; the front one white and subtransparent.Length, 8-9 mm.Male.?Front at level of anterior ocellus 0.27 of head width; noorbitals, third antennal joint a little broader than in female, of thesame length; palpi distinctly flattened and a little widened at tip,but lacking the pore which is so striking in the female; eyes withsame pilosity as in female. Genitalia small, black, the inner forcepsstraight, slender and close together, the outer slender and almost aslong; penis black, slender, with a delicate white flap on front attip. Claws and pulvilli moderately long.Paratypes.?F^m^lQ, U.S.N.M. No. 43691.Remarks.?Described from five females and one male, reared atWanaina, Northwest District, British Guiana, March, 1931, by J.G. Myers, from Castnia Ucoides Boisduval. The specimens were re-ceived from the Imperial Institute of Entomology, to which the typefemale, allotype male, and two paratype females are returned.The peculiar depression, or perhaps a sensory pit, in the palpusdoes not occur in any other tachinid known to me. It occurs onlyin the female. The species is distinguished by rather stout bristlesthroughout. I have compared the type series carefully with Zenillialihatrix Panzer (det. Bezzi) and find it agrees well except in havinga single bristle on the anteroventral side of the middle tibia. Nofine hairlike bristles extending up facial ridges, and in the visiblestructures at the tip of the abdomen in the female; in libatnx theterminal organs are concealed in our specimens by closure of thefourth abdominal segment in a longitudinal slit.
22 PROCEEDINGS OF THE NATIONAL MUSEUM vol.81TROPHOPS, new genusSomewhat allied to Pexomyia {ruhrifrons Perris, the genotype),but differing in having the vibrissae distinctly above oral margin, nocostal spine nor infrasquamal setules.Hypopleural bristles and postscutellum present; eyes and aristabare ; palpi and proboscis of ordinary form ; parafrontals and para-facials broad, especially the latter, which are bare and about halfas wide as clypeal depression; facial ridges bare, low, vibrissae farapart and much above the epistoma, only a little below the lowestcurve of the eye; cheeks broad and bulging, almost half the eyeheight. Third antennal joint nearly three times the second, ratherslender and tapering, not quite reaching vibrissae. Length of headat antennae slightly greater than at vibrissae ; frontal profile a littlegreater than facial ; clypeus flat, moderately broad.Scutellum with three lateral pairs of bristles, the apical pair al-most equally large, decussate or not, depressed. No discal abdominalbristles on second and third segments.First vein bare, third with one or two bristles at base, fourthwith bend slightly rounded, thence a little concave, ending distinctly,yet not very far before apex of wing. Hind cross vein concaveand a little oblique, joining fourth vein at three-fifths of the distancefrom anterior cross vein to bend.Genotype.?Trophops clauseni, new species.TROPHOPS CLAUSENI, new speciesBlack with yellowish pollen on head and thorax. Abdomen shin-ing black with broad bands of gray pollen on the second, third, andfourth segments.Male.?Front 0.31 of the head width. Head with somewhat silverypollen except on front and upper orbits where it is distinctly yellow.Frontal stripe wider than one parafacial. Verticals one pair; ocel-lars of normal size and proclinate; two upper orbitals reclinate,strong, the others rather weak, the lowest at middle of secondantennal joint. Antennae black, tip of second joint and basal thirdof third joint reddish; third joint decidedly slender and tapering,a little more than twice the second, not reaching to vibrissae. Cheekcovered with fine dark hair. Palpi brown. Thorax with a pair ofvery slender stripes outside the acrostichals, much enlarged behindthe suture; an outer stripe is broadly divided into two spots, thehindmost elongated. Pleurae black with thin gray pollen. Chae-totaxy : Acrostichal, 3, 3 ; dorsocentral, 3, 4 ; humeral, 2 ; posthumeral,2; notopleural, 2 ; presutural, 2 (the inner rather large) ; supraalar, 3;intraalar, 3 ; postalar, 2 ; sternopleural, 2, 1 ; pteropleural small. No
ART. 9 DIPTERA FEOM AMERICA, ASIA, AND JAVA ALDEICH 23infrasquamal setules. Propleura and prosternum bare. Abdomenmostly shining black, basal two-thirds of second segment, one-half ofthird segment, and two-thirds of fourth segment with gray pollen. Adelicate median black line on these segments. No marginals onfirst segment, those of second depressed and inconspicuous, third witha median pair and two lateral, fourth with a marginal row and asingle subdiscal pair far back. Genitalia black. Legs black (onlyhind ones present) ; hind tibiae with rather dense fringe of uniformbristles on anterodorsal side. Hind pulvilli and claws elongated.Wing considerably brownish except along the veins. Costal seg-ment beyond the first vein only slightly longer than the one beforeit. Calypters white; margin narrow, brownish yellow.Length, 7.4 mm.Female.?Front 0.37 of head width (the same in two) ; usual twopairs of orbital bristles, below the lowest frontals a distinct pollinosedark band extends from the eye to the suture (faintly indicated inmale). Hindmost sclerite of venter broadly rounded. Middle tibiawith two bristles on anterodorsal side. Claws and pulvilli small.Length, 5.7-7.7 mm.Type.?Male, IT. S. N. M. No. 43695.Remai'ks.?Described from one male and three females (one of thelatter considerably broken), reared from Popillia japonica Newman;male and two females at Toyona, Japan, July 9, 1930, by T. R. Gard-ner; the other female, which is broken, at Takarozawa, Japan, byC. P. Clausen, July, 1928. The species is named in honor of C. P.Clausen, who has made an extensive study of the Japanese beetleparasites in Japan and adjacent regions.
Genus EXORISTOIDES Coquillett
Exoristoides Coquillett, Eevisiou of the Tachiuidae of America north of Mex-ico, p. 90, 1897.?Walton, Proc. Ent. Soc. Washington, vol. 17, p. 98, 1915.?AxDEiCH and Webber. Proc. U. S. Nat. Mus., vol. 63, art. 17, p. 10, 1924.
?
CuKEAN, Can. Ent., vol. 58, p. 85, 1926.Exoristopsis Townsend. Proc. U. S. Nat. Mus., vol. 49, p. 426, 1915.The genotype of Exoristoides, designated by Coquillett in 1897, isjohnsoni Coquillett; that of Exoristopsis, designated by Townsendin 1915, is setifera Townsend.The genus has the general characters of ExoHsta of authors{Zenillia sens. lat. of Aldrich and Webber, 1924), with the additionof a large pteropleural bristle and almost invariably some setules onthe first vein. Curran suggests that Lypha is a near relative, whichis true. Townsend placed one of our species {slossonae) in Lydina{Polidea of authors), in the National Museum collection some yearsago, and this also expressed a true relation. Lypha may be distin-
24 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 81guished by the frontal bristles, which extend remarkably far downon the parafacials ; and both genera differ from Exoristoides in hav-ing the eyes smaller and cheek wider, as well as in lacking any hairson the first vein. In the single known specimen of Exoristopsisboth pteropleurals are broken off, but the large scars are present.Three species were originally included, johnso7ii, slossonae, andharringtoni, of which the last has been removed as type species ofthe genus Homalactia Townsend.KEY TO SPECIES OF EXORISTOIDESMALES
1. Third antennal joint very wide, two-thirds as wide as long, withobliquely truncate apex (North Carolina to Louisiana andCalifornia) johnsoni CoquillettThird antennal joint less than half as wide as long 22. First vein with few setules, rarely none ; fourth abdominal seg-ment wholly black (New Hampshire to Alabama) slossonae CoquillettFirst vein with complete series of setules from near humeralcross vein (Trinidad, West Indies) uricM, new speciesFEMALES
1. First vein setulose from near humeral cross vein to tip 2First vein with only a few setules, at most on basal half 32. Third antennal joint concave on front edge, widened at tip.urichi, new speciesThird antennal joint straight, not widened apically (Peru).setifera Townsend3. Sternopleurals two ; fourth abdominal segment red jolinsoni CoquillettSternopleurals three ; fourth abdominal segment wholly black.slossonae CoquillettEXORISTOIDES JOHNSONI Coquillett
Exoristoides johnsoni Coqihixett, Revision of the Tachinidae of America northof Mexico, p. 91, 1897.
?
Walton, Proc. Ent. Soe. Washington, vol. 17, p. 97,1915.?Bbimlet, Ent. News, vol. 33, p. 22, 1922.The material in the National Museum at present referred to thisspecies is the following: Holotype, female, Hertford County, N. C.(collection Coquillett) ; one female, Raleigh, N. C. (Brimley) ; onefemale, Palo Alto, Calif. (W. F. Derby coll., through the Aldrich col-lection) ; one female, Opelousas, La. (Pilate) ; one female, Lindsey,Calif. (McGregor) ; one dwarf male, reared at Capa, S. Dak., fromGryllus abhreviatus Serville, by Prof. H. C. Severin, emerged Sep-tember 12, 1919; two males and one female, reared at Sacramento,Calif., from Gryllus assim'dis Fabricius, by C. C. Wilson, emergedMarch 27, 1930; and one female, reared at Winters, Calif., by thesame entomologist from the same host, emerged April 10, 1931.Thus there are three different rearing records from Gryllus^ andthese are the only ones yet known.
AET. 9 DIPTERA FROM AMERICA, ASIA, AND JAVA ALDRICH 25EXORISTOIDES SLOSSONAE CoqniUett
Ex&ristoides slossonae Coqitillett, Revision of the Tachinidae of America northof Mexico, p. 91, 1807.?Johnson, Catalogue of the insects of New Jersey,p. 671, 1899, and p. 779, 1910.?Banks, Ent. News, vol. 23, p. 110, 1912.?Walton, Proc. Ent. Soc. Washington, vol. 17, p. 97, 1915.
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Britton, Check-list of the insects of Connecticut, p. 192, 1920.?Bbimley, Ent. News, vol.33, p. 22, 1922.?Johnson, List of the Dipteva of New England, p. 199,1925.
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West, A list of the insects of New York (Cornell Univ. Agr. Exp.Sta. Mem. 101), p. 815, 1928.
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Axlen, Ann. Ent. Soc. America, vol. 22, p.687, 1929.?CuRr.AN, Bull. Amer. Mus. Nat. Hist., vol. 61, p. 106, 1930.Exorisia spinipennis Coquillett, Revision of the Tachinidae of America northof Mexico, p. 95, 1897.
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Johnson, Ent. News, vol. 15, p. 157, 1904; Cata-logue of insects of New Jersey, p. 780, 1910.
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Smith, Psyche, vol. 24, p.58, 1917 (syn.).?Gibson. Ann. Rep. Ent. Soc. Ontario, p. 119, 1918.?AxDBiCH and Webber, Proc. U. S. Nat. Mus. vol. 63, art. 17, p. 10, 1924.The material in the National Museum referred to slossonae is asfollows : Four types of slossonae^ both sexes, from Franconia, N. H.(Mrs. Slosson), Eastport, Me. (coll. C. V. Riley), and Westville, N.J. (Johnson) ; the female type of spinipennis Coquillett, from Tif-ton, Ga. (Pilate) ; a female from Franconia, N. H. (Townsend)
;
a male from Chevy Chase Lake, Md. (Townsend) ; a female fromPotomac Run, Va. (McAtee) ; two males from College Park, Md.(Walton) ; a male from La Fayette, Ind. (Aldrich) ; and a malefrom Birmingham, Ala., reared by H. L. Weatherbee from EpiJachnacorrupta Mulsant, the Mexican bean beetle. The last emerged onAugust 3, 1922, and this is the only rearing record.
EXORISTOIDES SETIFERA Townsend
Exoristopsis setifera Townsend, Proc. U. S. Nat. Mus., vol. 49, p. 426, 1915.The only specimen in the National Museum is the type, a femalefrom Peru.The characters given for the genus Exoristopsis by Townsend seemmostlj^ specific, especially when considered in relation to the otherspecies than johnsoni, which I here include. Perhaps the most impor-tant is the small size of the ocellars, which is shared by urichi, newspecies. No mention is made of the pteropleurals, which are broken
off, but the scars are distinct.
EXORISTOIDES URICHI. new speciesBlack with silvery-gray pollen. Fourth abdominal segment redor reddish in ground color with yellow pollen.Male.?Front 0.27 of head width, pollen of the head silvery gray,the posterior orbit and upper part of front slightly yellow. Ocellarshairlike, proclinate, two upper frontals reclinate, six others, thelowest at the level of the base of third antennal joint. Antennae
26 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 81
mostly black; tip of second and base of third antennal joints reddish,this color extending to the middle of the third on inner side ; thirdantennal joint three times the second, rather wide, at its middlealmost twice as wide as the parafacial, somewhat prominent at baseand tip on the front side, leaving a distinct concavity between, theapex truncate, reaching nearly to vibrissae. Arista with penultimatejoint a little elongate, barely twice as long as thick, the apical jointrather short, thickened nearly to the middle. Eyes with long pile.Cheek a little more than one-fourth the eye height. Palpi slender,yellow apically, dark brown on basal half; outer verticals distinct,three-fourths as long as the inner. Mesonotum with rather thinsilvery-gray pollen, leaving four longitudinal shining black stripes,the inner narrow and reaching halfway from the suture to the scutel-lum; an elongate median spot before the scutellum. Acrostichal3, 3; dorsocentral, 3, 3; sternopleural, 2, 1. Scutellum, with 3 lateralbristles, a small erect nondecussate hairlike apical pair. Propleuraand prosternum bare. Second and third segments of abdomen ofsame color as thorax, in some lights showing pollen to the apices ofthe segments ; fourth segment with more yellowish pollen, the groundcolor rather dark red ; first segment with no median marginals, secondwith one pair, third with a median pair and two or three at themargin; second and third segments with a single pair of discalsmuch smaller than marginals; fourth segment with a median rowof six large bristles and a marginal row of the same number con-siderably smaller. Genitalia red, small, very similar to those of theother species ; the inner forceps blackish, being united into a beaklikeprocess curved upward and acute at tip; outer forceps slender,curved like the inner with a minute tooth at apex. Legs black, allthe tarsi rather short and decreasing in width. All the claws andpulvilli very small. Front tibiae with two bristles near middle onposteroventral side; middle tibiae with four bristles on anterodorsalside, the two middle ones quite long, and with one ventral; hindtibiae with an uneven sparse row on anterodorsal side. Wings hya-line, the first vein hairy to tip from humeral cross vein; third veinwith coarse hairs nearly to cross vein; fourth vein with roundedalmost rectangular bend, then concave, reaching costa only a littlebefore apex. The first posterior cell open. Hind cross veinstraight, somewhat oblique, joining fourth vein considerably beyondmiddle of the distance between anterior cross vein and bend. Calyp-ters white, the hind ones subtransparent ; infrasquamal setules presentbut delicate.Length, 6. 5 mm.Female.?Front at vertex 0.27 of the head width. The pollen onupper two-thirds much more distinctly yellow than in the male.Third antennal joint more than twice the second, distinctly wider
AET. 9 DIPTERA FROM AMEEICA, ASIA, AND JAVA ALDEICH 27than the parafacial, its tip widened about as in the male but thebasal part not protruding noticeably in the vicinity of the arista.Palpi as in the male. Fourth abdominal segment distinctly red inground color, shining in some lights over most of its surface. Gen-italia simple. Front tarsi a little widened from the second joint,the claws and pulvilli very small.Length, 6.2 mm.Type.?In the British Museum of Natural Histor3\Remarhs.?Described from one male and one female, reared inTrinidad, West Indies, by F. W. Urich, from pupae of Calyodesethlius Cramer. The specimens were received from Sir Guy A. K.Marshall, director of the Imperial Institute of Entomology, inLondon, to whom they are returned for ultimate deposit in theBritish Museum.Genus ACHAETONEURA Brauer and Bergenstamm
Achaetoneura Brauer and Bergenstamm, Zweifliigler des kaiserl. Museums zuWien, pt. 5, p. 334, 1891 (Denkschr. kaiserl. Akad. Wiss., vol. 58).
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Webbeb,Proc. U. S. Nat. Mus., vol. 78, art. 10, p. 1, 1930.ACHAETONEURA NIGRIPALPIS, new 8P?cies
Entirely black, including antennae, palpi, and scutellum.Female.?Front at vertex 0.3, 0.31 (in the two specimens) of thehead height. Head with silvery pollen tinged with light yellow onthe front; third antennal joint two and one-half times the second.Arista slender, bare, third joint very slightly thickened basally.Parafacial a little wider than third antennal joint. Facial ridgeswith rather small bristles extending to middle of third antennaljoint. Cheek one-fourth eye height. Thorax gray pollinose withfour black stripes, the inner pair abbreviated behind and the outerinterrupted at suture. Scutellum subshining black at base. Acro-stichal, 3, 3; dorsocentral, 3, 4; scutellum, with 3 lateral pairs ofbristles, the apicals rather strong, decussate, depressed; sterno-pleural, 2, 2, the anterior of the hindmost somewhat hairlike; pro-jjleura bare. First abdominal segment black, the second and thirdblack on apical two-fifths, their bases with smooth gray pollen whichis sharply defined behind; fourth segment wholly pollinose with aslight yellow tinge. No discals on second and third segments, a pairof marginals on first and second, marginal row of six on the third
;
fourth with a row of bristles in the middle, the marginals small andinconspicuous. Venter with a somewhat reddish tinge. Legs blackanterior tibiae with two posteroventral bristles; middle tibiae withtwo large anterodorsal, two small posterodorsal and one ventral;hind tibiae with a complete row of small bristles on posterodorsalside, one in middle slightly larger. Anterior tarsi with third and
28 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 81 : akt. 9fourth joints slightly flattened; claws and pulvilli small. Wingssubhyaline with the usual venation for the genus; third vein withthree setules at base. Calypters white.Length, 7.6-8.5 mm.Paratype.?Female, U. S. N. M. No. 43694.Remarks.?Described from two females reared by F. W. Urich,in Trinidad, from pupae of Calfodes ethlius Cramer. The speci-mens were received from Sir Guy A. K. Marshall, of the ImperialInstitute of Entomology, and the type is returned to him for ulti-mate deposit in the British Museum.Compared with the females of the genotype A. frenchii Williston,the head appears a little rounder, the eye longer vertically, the thirdantennal joint shorter and the bristles of the facial ridges do notextend so high. In chaetotaxy and other generic characters it agreesvery well. o
U. S. NATIONAL MUSEUM PROCEEDINGS. VOL. 81 . ART. 9 PL 1
DYSCRASIS AND COLLI NELLULA. NEW GENERA
1, Dyscrasis hendeli, new species: Wing. (Enlarged 16 dianietens.) 2-4, Cntlinellula magistri, new
.species: 2, Wing of male; '-i, wing of female; 4, abdomen of male, side view, flattened, with genitaliatwisted to show dorsal view. ( Enlarged CO diameters.) \ 11 photographed from micro.scopic slide.s.