Moths: Lepidoptera 
V?tor O. Becker   -   Scott E. Miller 
THE FOLLOWING LIST summarizes identi- 
fications of the so-called Macrolepidoptera 
and pyraloid families from Guana Island. 
Methods are detailed in Becker and Miller 
(2002). Data and illustrations for Macrolepi- 
doptera are provided in Becker and Miller 
(2002). Data for Crambidae and Pyralidae will 
be provided in Becker and Miller (in prepara- 
tion). General, but outdated, background infor- 
mation on Crambidae and Pyralidae are pro- 
vided by Schaus (1940). Data for Pterophoridae 
are provided in Gielis (1992) and Landry and 
Gielis (1992). Author and date of description 
are given for each species name. Earlier dates 
were not always printed on publications; those 
in square brackets indicate that the year was 
determined from external sources not the pub- 
lication itself As in previous lists, authors' 
names are put in parentheses when their 
generic placement has been revised. Detailed 
acknowledgments are provided in Becker and 
Miller (2002), but, in addition, we are espe- 
cially grateful to C. Gielis, E.G. Munroe, M. 
Shaffer, and M. A. Solis for assistance with iden- 
tifications and to K. Darrow for assistance in all 
aspects of this project. Sampling on Guana Is- 
land   was   supported   by  The   Conservation 
Agency, through grants from the Falconwood 
Corporation. 
SPHINGIDAE 
SPHINGINAE 
Agrius cingulatus (Fabricius 1775). United States 
south to Argentina. 
Cocytius antaeus (Drury 1773). Southern United 
States to Argentina. 
Manduca sexta (Linnaeus 1763). Widespread in 
the New World. 
Manduca rustica (Fabricius 1775). Widespread in 
the New World. 
Manduca brontes (Drury 1773). Antilles north to 
Central Florida. 
MACROGLOSSINAE 
Pseudosphinx tetrio (Linnaeus 1771). (See plate 
37.)   United  States  through  the Antilles  to 
Argentina. 
Erinnyis alope (Drury 1773). Widespread in the 
New World. 
Erinnyis ello (Linnaeus 1758). Neotropical. 
Erinnyis   crameri   (Schaus   1898).   Southern 
United States through the Caribbean south to 
Brazil. 
303 
Erinnyis domingonis (Butler  1875).  Southern 
United States through the Caribbean south to 
Brazil. 
Erinnyis  obscura   (Fabricius   1775).   Southern 
United States through the Caribbean south to 
Brazil. 
Pachylia ficus (Linnaeus 1758). Widespread in 
the New World. 
Callionima fixlcifera (Gehlen 1943). Southern 
United States to Argentina. 
Perigonia lusca (Fabricius 1777). Southern Florida 
to Argentina. 
Enyo lugubris (Linnaeus 1771). Southern United 
States, Antilles, south to Argentina. 
Aellopos tantalus (Linnaeus 1758). New York 
south to Argentina. 
Eumorpha vitis (Linnaeus 1758). United States 
through the Caribbean to Argentina. 
Cauthetia      noctuiformis      (Walker      1856). 
Caribbean. 
Xylophanes chiron (Drury 1770). Mexico through 
the Antilles to Argentina. 
Xylophanes pluto  (Fabricius   1777).   Southern 
United States to Brazil. 
Xylophanes tersa (Linnaeus 1771). Ontario, Antilles 
south to Argentina. 
Hyles lineata (Fabricius 1775). Worldwide. 
NOTODONTIDAE 
Nystalea nyseus (Cramer 1775). Mexico through 
the Caribbean south to Brazil. 
NOCTUIDAE 
HELIOTHINAE 
Heliothis suhflexa (Guen?e 1852). North America, 
Antilles, south to Argentina. 
NOCTUINAE 
Anida infecta (Ochsenheimer 1816). Central 
United States south to Argentina. 
HADENINAE 
Leucania humidicola Guen?e 1852. Antilles to 
Brazil. 
Leucania dorso/is Walker 1856. Southern Florida 
through the Neotropics. 
AMPHIPYRINAE 
Neogalea   sunia   (Guen?e   1852).   Florida   to 
Argentina. 
Catabenoides terminellus (Grote 1883). Southern 
United States, Antilles. 
Catabenoides lazelli Becker & Miller 2002. Guana, 
Anegada, St. Croix. 
Spodoptera albulum (Walker 1857). United States, 
Antilles, south to Argentina. 
Spodoptera fi??giperda (J.E. Smith 1797). Wide- 
spread in the New World. 
Spodoptera latifascia (Walker 1856). U.S. gulf 
states, Antilles south to Costa Rica. 
Spodoptera pulchella (Herrich-Sch?ffer 1868). 
Florida, Greater Antilles. 
Spodoptera dolichos (Fabricius 1794). U.S. gulf 
states, Antilles south to Costa Rica. 
Magusa orbifera (Walker 1857). Widespread from 
Canada to Argentina. 
Condica albigera (Guen?e 1852). Mexico through 
the Antilles to Paraguay. 
Condica mohilis (Walker [1857]). Southern United 
States through Antilles to Argentina. 
Condica sutor (Guen?e 1852). Southern United 
States, Antilles to Argentina. 
Perigea gloria Becker & Miller 2002. Guana, 
T?rtola. 
Elaphria agrotina (Guen?e 1852). Florida, Antilles 
south to Argentina. 
Elaphria nucicolora (Guen?e 1852). Neotropical. 
Anateinoma affabilis M?schler 1890.  Puerto 
Rico, Virgin Islands. 
Micrathetis triplex (Walker 1857). Southern United 
States through South America. 
ACARISTINAE 
Caularis undulans Walker [1858]. Hispaniola, 
Jamaica, Puerto Rican Bank. 
BACISARINAE 
Bagisara repanda (Fabricius 1793). Southeast 
United States to Paraguay. 
ACONTIINAE 
Amyna axis (Guen?e 1852). Pantropical. 
Ponometia exigua (Fabricius 1793).  Southern 
United States, Antilles south to Brazil. 
304 TH B   CAST 
Cydosia  nobilite?a  (Cramer  1779).   Southern 
United States, Antilles south to Argentina. 
Tripudia  quadrifera  (Zeller   1874).   Southern 
United States, Antilles south to Brazil. 
Tripudia balteata Smith 1900. Southern United 
States, Antilles south to Brazil. 
Ommatochila mundula (Zeller 1872). Southern 
United States, Antilles south to Argentina. 
Cobubatha metaspilaris Walker 1863. Antilles. 
Eumicremma minima (Guen?e 1852). Southern 
United States, Antilles south to Argentina. 
Eublemma  rectum  (Guen?e  1852).   Southern 
United States, Antilles south to Argentina. 
Spragueia margana (Fabricius 1794). Southern 
United States, the Neotropics. 
Spragueia perstructana (Walker 1865). Southern 
United States, Antilles south to Costa Rica. 
Thioptera aurif?re  (Walker  [1858]).   Southern 
United States, Antilles south to Brazil. 
SARROTHRIPINAE 
Characoma nilotica (Rogenhofer 1882). Pantrop- 
ical. 
Collomena filifera (Walker 1857). Florida, Antilles 
south to Brazu. 
Motya abseuzalis Walker 1859. Florida, Antilles 
south to Brazil. 
EUTELIINAE 
Paectes  obrotunda   (Guen?e   1852).   Southern 
United States, Antilles south to Paraguay. 
PLUSIINAE 
Pseudoplusia includens (Walker [1858]). United 
States to northern Chile and Argentina. 
CATOCALINAE 
Ptichodis immunis (Guen?e 1852). Mexico through 
the Antilles to Brazil. 
Mods latipes (Guen?e 1852). Southern United 
States, Antilles south to Argentina. 
Mods antillesia Hampson 1913. Lesser Antilles, 
Bahamas. 
Mods   repanda   (Fabricius    1794).    Antilles, 
Guatemala. 
Ophisma tropicalis Guen?e 1852. Southern United 
States, Antilles south to Argentina. 
OPHIDERINAE 
Azeta versicolor (Fabricius 1794). Neotropical. 
Metallata absumens (Walker 1862). Southern 
United States, Antilles south to Brazil. 
Plusiodonta thomae (Guen?e 1852). Endemic to 
the Antilles. 
Syllectra erycata (Cramer 1780). Florida, Antilles 
south to Brazil. 
Litoprosopus puncticosta Hampson 1926. Haiti, 
Virgin Islands. 
Diphthera festiva (Fabricius 1775). Neotropical. 
Gonodonta bidens Geyer 1832. Florida, Antilles 
south to Argentina. 
Melipotis acontioides (Guen?e 1852). Florida, An- 
tilles south to Brazil. 
Melipotis fasdolaris (H?bner [1831]). Southern 
United States, Antilles south to Uruguay. 
Melipotis contorta (Guen?e 1852). Florida, An- 
tilles. 
Melipotis fam?lica  (Guen?e  1852).   Southern 
United States, Antilles south to Venezuela. 
Melipotis ochrodes (Guen?e 1852). Mexico through 
the Antilles to Brazil. 
Melipotis januaris (Guen?e 1852). Southern 
United States, Antilles, northern South America. 
Ascalapha odorata (Linnaeus 1758). Pan- 
tropical. 
Epidromia lienaris (H?bner 1823). Neotropical. 
Manbuta pyraliformis (Walker 1858).  Florida, 
Antilles. 
Ephyrodes cacata Guen?e 1852. Southern United 
States, Antilles south to Colombia. 
Concana mundissima Walker [1858]. Florida, An- 
tilles south to Brazil. 
Massala asema Hampson 1926. Antilles. 
Lesmone formularis (Geyer 1837). Neotropical. 
Lesmone hinna (Geyer 1837). Southern United 
States, Antilles south to Brazil. 
Eulepidotis addens (Walker 1858). Antilles. 
Eulepidotis modestula (Herrich-Sch?ffer 1869). 
Antilles. 
Toxonprucha diffundens (Walker 1858). Antilles 
south to Brazil. 
Kakopoda progenies (Guen?e 1852). Florida, 
Antilles south to Brazil. 
Parachabora abydas (Herrich-Sch?fFer [1869]). 
Mexico through the Antilles to Brazil. 
TH E   CAST 3?5 
Cecharismena abarusalis (Walker 1859). Florida, 
Antilles south to Brazil. 
Cecharismena cara M?schler 1890. Antilles. 
Glympis eubolialis (Walker [1866]). Antilles. 
HERMINIINAE 
Drepanopalpia lunifera (Butler 1878). Antilles. 
Lascar?a omeodalis (Guen?e 1854). Florida, An- 
tilles. 
Bleptina   hydrillalis   Guen?e   1854.   Southern 
United States, Central America, Antilles. 
Bleptina caradrinalis Guen?e 1854. Southern 
United States, Antilles south to Brazil. 
Bleptina menalcasalis Walker [1859]. Antilles 
south to Venezuela. 
Bleptina araealis (Hampson 1901). Antilles, 
Florida. 
HYPENINAE 
Hypena lividalis (H?bner 1790). Southern 
Palearctic, Pantropical. 
Hypena minualis (Guen?e 1854). Antilles south 
to Brazil. 
ARCTIIDAE 
ARCTIINAE 
Hypercompe simplex (Walker 1855). Puerto Rican 
Bank, Lesser Antilles. 
Calidota strigosa (Walker 1855). Antilles, Southern 
United States. 
Eupseudosoma involutum (Sepp [1855J). South- 
ern United States, Antilles south to Ar- 
gentina. 
Utetheisa omatrix (Linnaeus 1758). Southern 
United States, Antilles south to Argentina. 
Utetheisa pulchella (Linnaeus 1758). Africa, Asia, 
the Neotropics. 
PERICOPINAE 
Composta cr?dula (Fabricius 1775). Antilles, 
Puerto Rican Bank. 
CTENUCHINAE 
Empyreuma pugione (Linnaeus 1767). Restricted 
to the Puerto Rican Bank. 
Horama pretus (Cramer 1777). Antilles. 
Horama panthalon (Fabricius 1793). Southern 
United States, Antilles to southern Brazil. 
Cosmosoma achemon (Fabricius 1781), Antilles 
south to Brazil. 
Eunomia colombina (Fabricius 1793). Antilles. 
Nyridela chalciope  (H?bner  [1831]).  Antilles, 
Central America. 
LITHOSIINAE 
Afiida char?entisma Dyar 1913. Antilles. 
Progona pallida (M?schler 1890). Puerto Rico. 
Lomuna nigr?puncta (Hampson 1900). Puerto 
Rico. 
GEOMETRIDAE 
OENOCHROMINAE 
Almodes   terraria   Guen?e   [1858].   Southern 
United States, Antilles to Colombia. 
ENNOMINAE 
Pero rectisectaria (Herrich-Sch?ffer [1855]). Puerto 
Rican Bank, Lesser Antilles. 
Oxydia   vesulia   (Cramer   [1779]).    Southern 
United States, Antilles south to Argentina. 
Erastria decrepitaria (H?bner [1823]). Southern 
United States, Antilles south to Brazil. 
Sphacelodes fusilineatus  (Walker  i860).  An- 
tilles. 
Macar?a paleolata (Guen?e [1858]). Antilles. 
Patalene ephyrata (Guen?e [1858]). Antilles. 
Cyclomia   mopsaria   Guen?e   [1858].   Antilles 
south to Brazil. 
CEOMETRINAE 
Eueana   simplar?a   Herbulot   1986.    Guana, 
Guadaloupe. 
Phrudocentra centr?fugar?um (Herrich-Sch?ffer 
1870). Florida through the Greater Antilles to 
Puerto Rico. 
Chloropteryx paular?a (M?schler 1886). Florida, 
Antilles. 
Synchlora frondaria (Guen?e  [1858]).  United 
States, Antilles south to Argentina. 
Synchlora cupedinaria (Grote 1880).  Florida, 
Greater Antilles to Virgin Islands, Nevis. 
306 THB   CAST 
STERRHINAE 
Semaeopus malefidarius (M?schler 1890). Puerto 
Rican Bank. 
Leptostales noctuata (Guen?e [1858]). Antilles. 
Acratodes suavata (H?lst 1900). Southern United 
States, Antilles. 
Lobocleta nymphidiata  (Guen?e [1858]).  An- 
tilles. 
Scopula laresaria Schaus 1940. Puerto Rican 
Bank. 
Idaea sp., probably Idaeafemar?a (Schaus 1940). 
Antilles. 
"Idaea" monata (Forbes in Ramos [1947]). An- 
tilles. 
[daea minuta (Schaus 1901). Antilles, United 
States. 
Idaea eupitheciata (Guen?e [1858]). Antilles. 
Leptostales phorcana (Guen?e [1858]). Antilles. 
Leptostales oblinatana M?schler 1890. Southern 
United States, Antilles to South America. 
LARENTIINAE 
Obila praecurrar?a (M?schler 1890). Antilles. 
Pterocypha defensata Walker  1862.  Southern 
United States, Antilles. 
HYBLAEIDAE 
Hyblaea puera species complex. 
COSSIDAE 
Psychonoctua personalis Grote 1865. Antilles, 
Mexico. 
CRAMBIDAE 
CRAMBINAE 
Argyr?a diplomochalis Dyar 1913. Puerto Rico. 
Donacoscaptes micralis (Hampson 1919). An- 
tilles. 
Fissicrambusfissiradiellus (Walker 1863). Antilles, 
Panama, Colombia. 
Fissicrambus minuellus (Walker 1863). Florida, 
Antilles, Central America. 
Microcrambus atr?strigellus (Hampson   1919). 
Jamaica. 
ODONTIINAE 
Cliniodes nomadalis Dyar 1912. Central America, 
Antilles. 
Microthcons ophionalis (Walker 1859). Southern 
United States, Central America, Antilles. 
Mimoschinia rufofascialis (Stephens 1834). An- 
tilles. 
GLAPHYRIINAE 
Alatuncusia bergii (M?schler 1891). Southern 
Florida, Antilles, perhaps South America. 
Chalcoela iphitalis (Walker 1859). North America 
to Guatemala. 
Chalcoela pegasalis (Walker 1859). North America, 
Antilles. 
Contortipalpia santiagalis (Schaus 1920). An- 
tilles. 
Dichogama amabilis M?schler 1891. Southern 
Florida to Antilles. 
Dichogama innocua (Fabricius 1793). Antilles, 
Central America. 
Dichogama redtenbacher? Lederer 1863. Southern 
Florida, the Neotropics. 
Galphyria badierana (Fabricius 1794). Antilles, 
Central America. 
Schacontia new species Neotropical. 
EVERGESTINAE 
Evergestella evincalis (M?schler 1891). Antilles, 
Florida Keys. 
PYRAUSTINAE 
Achyra bifidalis (Fabricius 1794). Neotropical. 
Achyra rantalis (Guen?e 1854). New World. 
Arthromastix   lauralis   (Walker   1859).    Neo- 
tropical. 
Asciodes gordialis Guen?e 1854. Neotropical. 
Atomopteryx serpentifera (Hampson 1913). An- 
tilles. 
Bicilia iarchasalis (Walker 1859). Neotropical. 
Cryptobotys   zoilusalis   (Walker   1859).   Neo- 
tropical. 
Desmia ufeus (Cramer 1777). Neotropical. 
Diaphania costata (Fabricius 1794). Antilles. 
Diaphania elegans (M?schler 1890). Antilles, 
Central America. 
TH E   CAST 307 
Diaphania hyalinata (Linnaeus 1767). Neotrop- 
ical. 
Diaphantania species near D. ceresalis (Walker 
1859). Antilles. 
Ennomosia basalis (Hampson 1897). Antilles. 
Epicorsia cerata (Fabricius 1795).  Lesser An- 
tilles. 
Epipagis algarroholis (Schaus 1940). Lesser An- 
tilles. 
Ercta vittata (Fabricius 1794). Neotropical. 
Eulepte gastralis (Guen?e 1854). Antilles. 
Glyphodes sibillalis Walker 1859. Neotropical. 
Herpetogramma  hipunctalis  (Fabricius  1794). 
Neotropical. 
Herpetogramma phaeopteralis (Guen?e  1854). 
Neotropical. 
Loxomorpha cambogialis (Guen?e 1854). Neotrop- 
ical. 
Marasmia   cochrusalis   (Walker   1859).   Neo- 
tropical. 
Marasmia sp. Virgin Islands. 
Microthyris anormalis (Guen?e 1854). Neotrop- 
ical. 
Oenobotys vinotinctalis (Hampson 1895). Neo- 
tropical. 
Omiodes albicinctalis (Hampson  1904).  An- 
tilles. 
Omiodes indicata (Fabricius 1775). Pantropical. 
Palpita flegia (Cramer 1777). Neotropical. 
Palpita persmilis Munroe 1959. Neotropical. 
Palpita viettei Munroe 1959. Neotropical. 
Palpusia species near P. eurypalpalis (Hampson 
1912). 
Phaedropsis principaloides (M?schler 1890). An- 
tilles. 
Portentomorpha    xanthialis    (Guen?e    1854). 
Neotropical. 
Pseudopyrausta   minima   (Hedemann   1894). 
Lesser Antilles. 
Pyrausta insignitalis (Guen?e 1854). Lesser An- 
tilles. 
Pyrausta laresalis Schaus 1940.  Lesser An- 
tilles. 
Samea ecdesialis Guen?e 1854. Antilles. 
Sathria onophasalis (Walker 1859). Antilles. 
Spoladea   recurvalis   (Fabricius   1775).    Pan- 
tropical. 
Steniodes  mendica  (Hedemann   1894).   An- 
tilles. 
Synclera jarbusalis (Walker 1859). Antilles. 
Syngamia florella (Stoll 1781). Neotropical. 
Uresiphita   reversalis   (Guen?e   1854).   Neo- 
tropical. 
PYRALIDAE 
PYRALINAE 
Ocrasa nostralis (Walker 1854). Pantropical. 
CHRYSAUCINAE 
Bonchis munitalis (Lederer 1863). Neotropical. 
Murgisca subductella (M?schler 1890). Lesser 
Antilles. 
Streptopalpia   minusculalis   (M?schler   1890). 
Neotropical. 
CALLERIINAE 
Pogrima palmasalis Schaus 1940. Antilles. 
Thyridopyralis    gallaerandialis    Dyar     1901. 
Florida. 
EPIPASCHIINAE 
Phidotricha erigens Ragonot 1888. Antilles, Cen- 
tral America. 
Phidotricha insularella (Ragonot 1888). Antilles, 
Central America. 
Tineopaschia minuta Hampson 1916. Antilles. 
PHYCITINAE 
Anabasis ochrodesma (Zeller 1881). Southeast 
United States to northern South America. 
Ancylostomia stercorea (Zeller 1848). Neotropi- 
cal. 
Australephestiodes stictdla (Hampson 1901). An- 
tilles. 
Bema neuricella (Zeller 1848). Neotropical. 
Cactoblastis cactorum (Berg  1885).  Native to 
southern South America, now widespread be- 
cause of biological control introductions. 
Cassiana malacella (Dyar 1914). Mexico. 
Crocidomera fissuralis (Walker 1863). Hispan- 
iola, Puerto Rico. 
Ectomyelois   decolor   (Zeller   1881).   Antilles, 
Central America, South America. 
3 O? TH E   CAST 
Erdieva quantuldla (H?lst 1887). United States, 
Antilles. 
Etiella   zinckenella   (Treitschke   1832).   Cos- 
mopolitan. 
Eurythmia   hospitella   (Zeller   1875).   United 
States. 
Fundella argentina Dyar 1919. Neotropical. 
Hypargyria   definitella   (Zeller   1881).   Neo- 
tropical. 
Oryctometopia fossulatella Ragonot 1888. Texas, 
Antilles, Central America to Brazil. 
Ozamia lucidalis (Walker 1863). Neotropical. 
Sarasota fiirculella (Dyar 1919). Antilles. 
Unadilla errondla (Zeller 1881). Neotropical. 
Zamagiria laidion (Zeller 1881). Neotropical. 
PTEROPHORIDAE 
Adania   ambrosiae   (Murtfeldt   ? 
America, Antilles, Gal?pagos. 
Adania thomae (Zeller 1877). Antilles, South 
America. 
Exelastis cervinicolor (Barnes & McDunnough 
1913). Florida, Antilles, Gal?pagos. 
Exelastis pumilio (Zeller 1873). Neotropical. 
Lantanophaga   pusillidactyla   (Walker   1864). 
Pantropical. 
Megalorrhipida defectalis (Walker 1864). Pantrop- 
ical. 
There are literally hundreds of additional 
species of moths we have collected on Guana 
Island. They are generally small species infor- 
mally lumped together as "microlepidoptera." 
Sorting them out is taking decades. Meanwhile 
Davis (1986) clarified the taxonomic status of 
just one of them in family Tineidae: Acrolophus 
triatomellus. 
North 
THE   CAST 309 
ISLAND 
Fact and Theory in Nature 
James Lazell 
UNIVERSITY OF CALIFORNIA PRESS 
Berkeley    Los Angeles    London