SMITHSONIAN INSTITUTION
Contributions from the United States National Herbarium
Volume 55: 1-584
Checklist of the Plants of the Guiana Shield
(Venezuela: Amazonas, Bolivar, Delta Amacuro;
Guyana, Surinam, French Guiana)
Department of Botany
National Museum of Natural History
Washington, DC
2007

SMITHSONIAN INSTITUTION
Contributions from the United States National Herbarium
Volume 55: 1-584
Checklist of the Plants of the Guiana Shield
(Venezuela: Amazonas, Bolivar, Delta Amacuro;
Guyana, Surinam, French Guiana)
by
V Funk,T. Hollowell, R Berry, C. Kelloff, and S. N. Alexander
•Varies
Department of Botany
National Museum of Natural History
Washington, DC
2007
ABSTRACT
Funk, v., T. HoUowell, P. Berry, C. Kelloff, and S. N. Alexander. Checklist of the Plants of the Guiana
Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contributions
from the United States National Herbarium, volume 55, 584 pages. This volume contains a checklist of the
plants of the Guiana Shield Area (northeastern South America) produced using information gathered from historical and recent
collections as well as many recent expeditions. The Guiana Shield, as treated here, includes the countries of Guyana and
Surinam, French Guiana (a part of France) and the Venezuelan states ofAmazonas, Bolivar, and Delta Amacuro. The checklist
contains 257 vascular plant families with 13,367 species. According to the list 6,791 of those species are from Amazonas, 6,669
from Bolivar, 1,651 from Delta Amacuro, 7,1 12 from Guyana, 4,984 from Surinam, and 5,406 are from French Guiana. Of these
species, 293 (2.2%) are estimated to be introduced and naturalized and even fewer are invasive. The small number of naturalized
species is a good indication of the large amount^of intact vegetation from this area of South America. The largest family is the
Leguminoseae with 146 genera containing 1032 species; 41 families are represented by only one species. Over 250 scientists
from around the world participated in this project by identifying specimens and checking species lists. This volume should be of
use to students, taxonomists, ecologists, and conservation biologists, as well as to interested amateurs.
KEY WORDS (not in title): South America, biodiversity, endemic species, invasive species
DATE OF PUBLICATION: 31 July 2007
Cover Design by Alice R. Tangerini: front, illustration by Alice R. Tangerini: new species ofPaloue, habit,
leaf base detail, ovary detail, anther and stipules (K.M. Redden 4027); back, illustration by Alice R.
Tangerini: Nautilocalyx pemphidius L. Skog; habit, flower bud, open flower, corolla face, longitudinal
section of flower, stamens, pistil, and nectary, fruit, seeds (Bell 324)
Contributions from the United States National Herbarium (ISSN 0097- 1618) Department ofBotany, National
Museum of Natural History, MRC-166, Smithsonian Institution, Washington, DC, 20013-7012, USA.
POSTMASTER: Send address changes to Contributions from the U.S. National Herbarium, Department
of Botany, National Museum of Natural History, MRC-166, RO. Box 37012, Smithsonian Institution,
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The paper used in this publication meets the minimum requirements of the American National Standard
for Permanence of Paper for Printed Library Materials Z39.48—1984.
The periodical, Contributions from the United States National Herbarium, was first published in 1890 by
The United States Department ofAgriculture. From 1 July 1902 forward it was published as a Bulletin of
the United States National Museum. The series was discontinued after volume 38 (1974), and has been
revived with volume 39 (2000) as a venue for publishing longer taxonomic papers, checklists, floras, and
monographs, produced by the staff and associates at the U.S. National Herbarium. It is externally peer
reviewed, and published at irregular intervals. Subscription and other correspondence should be addressed
to CUSNH, Department of Botany, National Museum of Natural History, MRC-166, Smithsonian
Institution, Washington, DC, 20013-7012, USA e-mail: CUSNH@si.edu. The present issue is available
for free while supplies last and PDF files of the most recent issues (vol. 49-present) of the Contr. U.S. Natl.
Herb, are available at: http://www.nmnh.si.edu/botany/pubs/CUSNH/ If you have any questions about
manuscript submission or comments on previous volumes, please contact the editor, Paul M. Peterson
(peterson@si.edu).
CONTENTS
FOREWORD by Peter H. Raven 5
INTRODUCTION by V. A. Funk & T. Hollowell 7
Geology of the Guiana Shield 8
Climate 9
Diversity 9
C onservation 10
Discussion 10
Using the Checklist 13
Nomenclatorial considerations 14
Acknowledgements 14
References 14
CHECKLIST 17
Psilotophyta 17
Lycophyta 17
Pterophyta 21
Coniferophyta 56
Cycadophyta 56
Gnetophyta 57
Anthophyta 57
Monocotyledonea 58
Dicotyledonea 183
LITERATURE CITATION FOR ILLUSTRATIONS 562
APPENDIX: Individuals who identified plants 563
INDEX TO GENERA 565
INDEX TO FAMILIES 584
0.10 mm
Henriettea multiflora Naudin (Melastomataceae)
Checklist of the Plants of the Guiana Shield
FOREWORD
At a time when human activities have placed enormous stress on the natural world, there is still
much that we do not know about the Earth's biodiversity; the great majority of species have not yet been
described, and very few of those that have been named are known in any detail. Checklists provide us
with a first approximation of what we know, and can provide a profile of the biodiversity in relatively
poorly-known parts of the globe. For many groups, they may represent the most complete information
available. Since basic taxonomic information provides essential information for studies in systematic
and evolutionary biology, we continually seek to improve and update it. Collecting specimens and writing
checklists, flora treatments, and monographs are all essential elements in our quest to understand the
natural world and how it evolved.
In northern South America, one of the richest regions of the world in terms of biodiversity, our
knowledge has grown with the recent publication of the Checklist ofthe Plants ofPeru, Checklist ofthe
Plants ofEcuador, and the nearly-completed Checklist of the Plants ofBolivia. This current volume,
which covers all of the plants known from the Guiana Shield region in northeastern South America, is
another outstanding contribution to this global effort. All perrtinent information about this region will
be brought together in a comprehensive New World Checklist of Vascular Plants and Bryophytes
(http://mobot.mobot.org/W3T/Search/nwc.html). All of these projects contribute to our overall mission
to understand, preserve, and protect the wealth of biodiversity that is such an outstanding feature of this
incredible part of the world.
Checklists have many uses: they are aids in the identification and correct naming of species, essential
resources for biodiversity estimates and biogeographic studies, and the starting points for more detailed
studies ofan area's biota. Reviewed by many specialists, they often represent the most advanced state of
knowledge available in their field. They also increase national and regional pride and provide important
public outreach and fundamental information to use in addressing the biodiversity crisis. This checklist
of the plants of the Guiana Shield, combined with the recently published Checklist of the Terrestrial
Vertebrates ofthe Guiana Shield, increases our knowledge ofthis fascinating region and acts as a starting
point from which additional research may be developed. It gives insight into the species richness,
endemicity, and floral and faunal affinities of this region. Such lists can also provide the baseline
information for analyzing species turnover rates, migration, invasion events, and many other biological
phenomena.
In the face of the growing biodiversity crisis, we celebrate the collaborative nature of a project such
as this. It is an excellent example of effective interaction, involving over 250 people from around the
world who contributed in diverse ways to its completion. These contributors variously collected plants
from the area, provided identifications of and information about those plants, or reviewed drafts of the
checklist, and together made possible its successful completion.
It is with great pleasure that I welcome this important addition to our knowledge of the biodiversity
of Earth, and at the same time congratulate the many individuals and institutions who made it possible.
Peter H. Raven
President, Missouri Botanical Garden
George Engelmann Professor of Botany, Washington University in St. Louis
Checklist of the Plants of the Guiana Shield
70°W 65°W 60°W
_L
55°W
1 ^
50°W
10°N-
5°N-
Atlantic Ocean
O-N-
Guiana Shield
, Shield Outliers
-10°N
•5°N
70°W 65°W 60°W 55°W sew
Figure 1 : The Guiana Shield
Checklist of the Plants of the Guiana Shield
Checklist of the Plants of the Guiana Shield
(VENEZUELA: Amazonas, Bolivar, Delta Amacuro;
GUYANA, SURINAM, FRENCH GUIANA)
by
V. Funk^ T. HollowelP, P. Berry^ C. Kelloff , and S. N. Alexander'
INTRODUCTION
V. A. Funk and T. Hollowell
The Guiana Shield region has long held a
fascination for tropical biologists because of its
unique geography that includes table-top
mountains (tepuis), tropical savannas, and broad
expanses of rainforests. These areas are home to
many endemic taxa, as well as to unique
ecosystems such as the Greenheart
{Chlorocardium) forests of Guyana and the
montane savannas of Brazil, Venezuela, and
Guyana. Some of the largest expanses of
untouched Neotropical forests occur in these areas.
The Checklist of the Plants of the Guiana
Shield covers all vascular plants known to occur
in the Guiana Shield region of northeastern South
America. It is the product of the combined efforts
of two research initiatives, the Smithsonian
Institution's Biological Diversity of the Guiana
Shield Program (BDG) and the Missouri Botanical
Garden's Flora ofthe Venezuelan Guayana project
(FVG). The database created for the Checklist of
the Plants of the Guianas (Boggan et al., 1992,
1997) has been extensively updated and refmed,
and most importantly the nomenclature has been
coordinated with a database extracted from the
Flora of the Venezuelan Guayana (Steyermark,
Berry, and Hoist, 1995—2005).
This plant checklist is a companion to the
recently published Checklist of the Terrestrial
Vertebrates of the Guiana Shield (Hollowell and
Reynolds, eds. 2005). Together they represent a
new research and conservation resource which
highlights three critical facets oftaxonomic work:
research, collections, and expeditions. These
volumes should be of particular use to students,
taxonomists, ecologists, and conservation
biologists, as well as to interested amateurs. Both
of these checklists were produced using
information gathered from historical and recent
collections as well as many recent expeditions to
northeastern South America. The expertise of
scientists from many organizations around the
globe has been drawn together and made available
to the scientific and conservation communities in
both hard copy and on the web (Hollowell et al.,
2000). Research projects such as this would not
be possible without considerable international
collaboration from over 100 scientists who provide
determinations for the collections (see Appendix).
Checklists have many uses. They are aids in
the identification of organisms, resources for
biodiversity estimates and biogeographic studies.
' Department of Botany and Biological Diversity of the Guiana Shield Program (BDG), National Museum of Natural History,
MRC-166, P.O.Box 37012, Smithsonian Institution, Washington, DC 20013-7012 U.S.A.
-BDG and Office of Information Technology, National Museum of Natural History, PO.Box 37012, Smithsonian Institution,
Washington, DC 20013-7012 U.S.A.
^ University of Michigan Herbarium & Dept. of Ecology and Evolutionary Biology, 3600 Varsity Drive, Ann Arbor, Michigan
48108-2287 U.S.A.
8 Checklist of the Plants of the Guiana Shield
nomenclators, and essential starting points for more
detailed studies of an area's biota. The Guiana
Shield is a geologic formation, and as such it
defines a section of northeastern South America
that is ideally a natural unit rather than a political
one (Kelloffand Funk 2004). This gives these two
checklists (plants and terrestrial vertebrates) an
environmental focus that is a first step toward
thinking in terms of ecosystems, evolution, and
systematics rather than simple lists of organisms.
For instance, checklists can tell us how many
organisms ofa certain kind are in a particular area,
thus giving us an idea of the species richness.
Properly utilized, the data can serve as an indicator
of endemicity within an area and provide
information on the number of introduced species,
and they can be used in a comparative manner
between and among areas. Since this is a natural
area, they can be used to formulate questions about
adaptive radiation, dispersal, and evolutionary
history. In fact, data from these checklists are
already being used to examine endemicity levels,
species turnover rates, and the location of areas
most in need of additional study. The checklists
are particularly useful for these types of
investigations because they encompass several
groups of organisms (plants, amphibians, reptiles,
birds, and mammals) facilitating comparisons
among the different groups. Conservation efforts
are improved because the checklists provide
standardized species nomenclature that will be
used in both governmental and academic
undertakings, including impact studies, reserve
planning, ecological research, production of floral
and faunal studies, and biogeographical analyses,
allowing data sets from different sources to be
compared with greater accuracy and confidence.
Both checklists exemplify the collaborative
nature of studies in biological diversity science,
being reflections of the efforts of so many
specialists essential to their production. They are
truly international efforts, for while the volumes
were organized by the BDG staff, the authors and
experts who supplied identifications are based all
over the world and in many types of organizations.
The result affirms the common interests of all of
the people, institutions, and nations involved in
studying, understanding, and conserving the
irreplaceable natural heritage of our world.
GEOLOGY OF THE GUIANA SHIELD
The Guiana Shield (Figure 1) is a distinct
ancient geological formation that today is roughly
bounded by the Atlantic Ocean to the east, the
Orinoco River to the north and west, the Rio Negro
(a major tributary of the Amazon River) to the
southwest, and the Amazon River to the south
(Gibbs and Barron 1993). The Shield includes the
mountain systems that form the watershed
boundary between the Amazon and Orinoco rivers.
On the Shield's western side, the Orinoco River
and Rio Negro are connected by the Rio Casiquiare.
The base of the Guiana Shield is composed of
crystalline rocks of Proterozoic origin; these are
mainly granites and gneisses formed between 3.6
and 0.8 billion years ago (Mendoza 1977; Schubert
and Huber 1990). Large areas of the Shield were
overlain with sediments from 1 .6 to 1 billion years
ago (Huber 1995a), and remnants of these
sandstones comprise the Roraima group of
formations that are scattered across the central
portion of the Shield. Erosion has decreased the
size of these formations, and the remaining
sediments extend west from the Potaro escarpment
of the Pakaraima Mountains in central Guyana
through parts of Venezuela and Colombia
(Arbelaez and Callejas 1999) and south into
northern Brazil (Leechman 1913; Gibbs and Barron
1993). Within this area, erosion has resulted in
numerous vertical-walled, frequently flat-topped
mountains called "tepuis," among them Chimanta-
tepui (2,550 m), Cerro Duida (2,358 m) and Auyan-
tepui (2,450 m). Pico de Neblina (3,014 m) is the
Shield's westernmost tepui and highest point,
located on the southernmost segment ofthe border
between Venezuela and Brazil. Mount Roraima
(2,810 m) is located at the juncture of Guyana,
Venezuela, and Brazil and includes the highest
point within Guyana. The easternmost peaks or
the shield are located in Guyana and reach
approximately 2,000 m elevation, including Mt.
Ayanganna (2,080 m) and Mt. Wokomung (2,000
m). Most Venezuelan tepuis are between 2,000
and 2,400 m and the highest point is Pico de
Neblina in Venezuela/Brazil (over 3,000 m).
Several ofthese mesa-like formations are virtually
inaccessible by foot and are so unusual that a
fictional scientific expedition account referred to
one as "The Lost World" (Doyle 1912), a term
sometimes applied to all tepuis. Notable waterfalls
Checklist of the Plants of the Guiana Shield
of the region include Angel Falls (979 m) on
Auyan-tepui in Venezuela and Kaieteur Falls (226
m) on the Potaro River in Guyana's Pakaraima
Mountains.
Granitic dome mountains occur in the southern
part ofthe three Guianas (Guyana, Surinam, French
Guiana), where they are known as "inselbergs," as
well as in Venezuela where they are known as
"lajas" and extend in a broad arc all the way from
northeastern Bolivar state west and south along the
Orinoco and then further south as far as Brazil.
Deposits of low-nutrient white sands occur inland
of the coastal plain, in belts across the region and
in isolated pockets. Large areas of savanna are
found in the region, particularly the complex of
savannas that includes the Rupununi Savanna in
southwestern Guyana, the Gran Sabana in eastern
Bolivar, Venezuela, and the savannas of northern
Roraima, Brazil. In some parts ofthese areas there
is limited forest growth because sands overlay a
clay hardpan that is resistant to penetration by tree
roots and because they flood during the heavy rainy
season. Tertiary and Quaternary sediments
separate the southern edge of the Guiana Shield
from the Amazon River and the eastern edge of
the Shield from the Atlantic Ocean.
The boundaries of the Guiana Shield are
difficult to define precisely. In the south between
mountains on the southern boundary of the Shield
and the Amazon and Negro rivers there is a broad
band of outwash materials resulting from erosion.
In the Venezuelan state of DeltaAmacuro much of
the area is covered by thick sediments deposited
primarily by the Orinoco River. In the west, there
are remnants of the Shield at least as far as
Colombia. In the east and south the hills give way
in some areas to lowland tropical rainforest more
similar to that found in the Amazon basin.
However, some remains ofthe Guiana Shield occur
in the southern section, and some portion of the
sediments are derived from outwash from the
highlands of the Shield. For more detailed
discussions of the geology of the area, readers
should refer to Gibbs and Barron ( 1 993) and Huber
(1995a).
CLIMATE
As a whole, the Guiana Shield region has a
tropical climate characterized by a relatively high
mean annual temperature exceeding 25° C at sea
level, an annual monthly maximum temperature
range of less than 5° C, and an average daily
temperature range ofapproximately 6° C. Because
of the Guiana Shield's location just north of the
equator, its climate varies primarily according to
elevation and effects of the trade winds that
combine to affect rainfall patterns. The trade winds
blow consistently from the east and northeast, off
of the Atlantic Ocean onto northeastern South
America. Due to orographic effects, the
easternmost escarpments of the mountains of the
Guiana Shield are generally localities of increased
precipitation where these moisture-laden winds
meet the slopes (Clarke et al. 2001). Seasonal
oscillations ofthe Inter-tropical Convergence Zone
(ITCZ) also bring variations in rainfall as the
locations of low pressure zones near the equator
change (Snow 1976). Varying primarily by
latitude, one or two rainy seasons result from shifts
in the ITCZ. The heaviest rains usually occur
between May andAugust, whereas the rainy season
running from December to January is shorter and
less intense, with rains that do not penetrate as far
inland. In areas where there is only one rainy
season, the driest part of the year is January to
March; in areas where there are two rainy seasons
the driest months are March and October.
However, even during most dry seasons, frequent
storms provide adequate moisture to allow
evergreen tropical moist forests to persist in most
low elevation parts of the region.
DIVERSITY
The varied topology of the Guiana Shield
includes sandstone tepuis, granite inselbergs, white
sands, seasonally flooded tropical savannas,
lowlands with numerous rivers, isolated mountain
ranges, and coastal swamps, each supporting a
characteristic vegetation (Huber et al. 1 995; Huber
1995b). This variety accounts for a great deal of
the high diversity and endemicity of the Shield's
biota. The Shield has a flora and fauna with
numerous endemic species, ca. 40% according to
a recent analysis (Kelloff and Funk 2004). Some
tepui endemic species occur as low as 300 m
elevation, with increasing numbers by 1,500 to
1,800 m, and fully developed communities
occurring by 2,000 m (Funk and Berry, 2005).
10 Checklist of the Plants of the Guiana Shield
Most parts of Brazil north of the Amazon River
and much ofeastern Colombia are poorly explored,
and they are not included in this publication (see
below).
CONSERVATION
With the exceptions of the few populated
localities such as Puerto Ayacucho, Ciudad
Guayana, Ciudad Bolivar, and the agricultural
coastal areas, the environment ofthe Guiana Shield
has benefited from limited access and low
population densities, although this sarae isolation
has hindered biodiversity research. Estimates vary,
but approximately 70% (Funk & Richardson 2003)
of the vegetation is still relatively undisturbed by
human activities. Recently, however, the pace of
disturbance has greatly increased; current threats
to the environment include large-scale logging by
Asian and local companies, large- and small-scale
gold and diamond mining, oil prospecting, bauxite
mining, massive hydroelectric projects, wildlife
trade, and population-related pressures such as
burning, grazing, agriculture, and the expansion
of Amerindian villages. Taken together, these
impacts have begun to take their toll, with vast
areas previously left undisturbed are now
vulnerable to increasing disturbance.
The status of conservation efforts varies by
country. Throughout the Guiana Shield, many
areas that are designated as protected or otherwise
restricted are often only "paper" parks because of
a lack of infrastructure and funds to protect them.
Over the last four decades, Venezuela has
established seven national parks, 29 natural
monuments, and two biosphere reserves covering
about 142,280 km^, more than 30% of its share of
the Guiana Shield (Funk and Berry 2005). In
Guyana, the progress of conservation efforts has
been slower, with the only substantial protected
area being Kaieteur National Park, its 627 km^
comprising about 0.3% of the country's area
(Kelloff 2003), with additional reserves under
consideration. Guyana's 3,710 km^ Iwokrama
Forest (Clarke et al. 2001) is dedicated to
sustainable use. Surinam's protected areas system
includes one national park and a network of II
reserves, totaling almost 20,000 km^ over 12% of
its total area. This includes the recently created
16,000 km^ Central Suriname Nature Reserve, an
UNESCO World Heritage Site that joined and
expanded three existing reserves (see http://
www.stinasu.com). French Guiana has no
officially designated protected areas, but 18
proposed sites total 6,710 km^, about 7.5%) of its
area (Lindeman and Mori 1989).
The Guiana Shield encompasses part or all of
five countries (excluding the small part in
Colombia) with five different governments, five
official languages, and many more indigenous
languages. Cooperation is sometimes hampered
by border disputes and illegal cross-border
transportation of gold, wildlife, and more recently,
illegal drugs. The implementation of conservation
practices is further complicated by many issues
concerning the indigenous peoples of the region.
All of these challenges will have to be overcome
on the way to designing and maintaining a viable
reserve system for the Guiana Shield.
DISCUSSION
This checklist includes 257 vascular plant
families containing 2,072 accepted genera. We
have recorded over 14,732 accepted subgeneric
taxa, 1,358 of those including a subspecific or
varietal epithet and 13,367 distinct species.
According to our list 6,791 of those species are
from Amazonas, 6,669 from Bolivar, 1 ,65 1 from
Delta Amacuro, 7,112 from Guyana, 4,984 from
Surinam, and 5,406 from French Guiana. Ofthese
species, 293 (2.2%) are estimated to be introduced
and naturalized. In the three Guianas there are
8,933 species and there are 9,814 species in the
Venezuelan Guayana. With 5,482 species found
in both areas there is 41% overlap between the two
areas. Within the checklist area 3,451 species are
listed only for the Guianas and 4,332 are found
only in the Venezuelan Guayana. See Table I for
a summary of the numbers of taxa in each family
and Table 2 for the families with the largest number
of species.
The top 1 5 families (in number of species)
have a total of7,094 species, which represents over
half of the species in the Guiana Shield area.
According to our list 154 families have 25 species
or less, and 34 families have 26-50 species.
Consequently, 73% of families have no more than
50 species and 41 families (16%) are represented
by only one species.
Checklist of the Plants of the Guiana Shield 11
Table 1 . Number of species in each plant family.
PSILOPHYTA(l) GNETOPHYTA (6) Acanthaceae 115
Psilotaceae 1 Gnetaceae 6 Aizoaceae
Amaranthaceae
1
30
LYCOPHYTA(107) ANTHOPHYTA Anacardiaceae 23
Isoetaceae 5 MONOCOTYLEDONEA (3301) Anisophylleaceae 1
Lycopodiaceae 27 Agavaceae 2 Annonaceae 178
Selaginellaceae 75 Alismataceae 13 Apiaceae 4
Araceae 226 Apocynaceae 211
PTEROPHYTA (754) Arecaceae 111 Aquifoliaceae 76
Adiantaceae 71 Bromeliaceae 327 Araliaceae 64
Aspleniaceae 36 Burmanniaceae 29 Aristolochiaceae 32
Blechnaceae 23 Cannaceae 2 Asclepiadaceae 132
Cyatheaceae 46 Commelinaceae 26 Balanophoraceae 2
Davalliaceae 1 Costaceae 16 Balsaminaceae 1
Dennstaedtiaceae 63 Cyclanthaceae 33 Basellaceae 2
Dicksoniaceae 1 Cyperaceae 413 Bataceae 1
Dryopteridaceae 26 Dioscoreaceae 54 Begoniaceae 14
Gleicheniaceae 18 Eriocaulaceae 94 Bignoniaceae 175
Grammitidaceae 62 Haemodoraceae 3 Bixaceae 4
Hymenophyllaceae 89 Heliconiaceae 19 Bombacaceae 55
Hymenophyllopsidaceae 8 Hydrocharitaceae 2 Bonnetiaceae 28
Lomariopsidaceae 79 Iridaceae 13 Boraginaceae 62
Lygodiaceae 4 Lemnaceae 8 Brassicaceae 8
Marattiaceae 7 Liliaceae 22 Bromeliaceae 1
Marsileaceae 3 Limnocharitaceae 3 Brunelliaceae 2
Metaxyaceae 1 Marantaceae 85 Burseraceae 81
Oleandraceae 8 Mayacaceae 3 Buxaceae 1
Ophioglossaceae 3 Musaceae 2 Cabombaceae 4
Osmundaceae 2 Najadaceae 3 Cactaceae 24
Parkeriaceae 4 Orchidaceae 1020 Campanulaceae 10
Plagiogyriaceae 1 Poaceae 526 Canellaceae 1
Polypodiaceae 59 Pontederiaceae 9 Capparaceae 33
Pteridaceae 16 Rapateaceae 83 Caprifoliaceae 3
Salviniaceae 5 Ruppiaceae 1 Caricaceae 3
Schizaeaceae 21 Smilacaceae 24 Caryocaraceae 10
Tectariaceae 21 Strelitziaceae 1 Caryophyllaceae 3
Thelypteridaceae 44 Taccaceae 1 Casuarinaceae 1
Vittariaceae 16 Thurniaceae 3 Cecropiaceae 42
Woodsiaceae 16 Triuridaceae 9 Celastraceae 22
Typhaceae 1 Ceratophyllaceae 1
CONIFEROPHYTA(IO) Velloziaceae 2 Chenopodiaceae 1
Pinaceae 1 Xyridaceae 100 Chloranthaceae 3
Podocarpaceae 9 Zingiberaceae 11 Chrysobalanaceae 186
Zosteraceae 1 Clethraceae 1
CYCADOPHYTA (5) Clusiaceae 198
Cycadaceae 1 DICOTYLEDONEA (9 1 83) Combretaceae 48
Zamiaceae 4 Incertae sedis 7 Compositae 303
12 Checklist of the Plants of the Guiana Shield
Table 1 . Number of species in each plant family, (contd.)
Connaraceae 30 Lythraceae 35 Proteaceae 20
Convolvulaceae 107 Magnoliaceae 2 Quiinaceae 32
Crassulaceae 2 Malpighiaceae 205 Rafflesiaceae 2
Cucurbitaceae 56 Malvaceae 85 Ranunculaceae 2
Cunoniaceae 13 Marcgraviaceae 17 Rhabdodendraceae 1
Cuscutaceae 5 Melastomataceae 534 Rhamnaceae 26
Cyrillaceae 2 Meliaceae 49 Rhizophoraceae 13
Dichapetalaceae 10 Mendonciaceae 14 Rosaceae 10
Dilleniaceae 38 Menispermaceae 48 Rubiaceae 742
Droseraceae 14 Menyanthaceae 1 Rutaceae 81
Ebenaceae 25 Molluginaceae 3 Sabiaceae 11
Elaeocarpaceae 50 Monimiaceae 4 Santalaceae 1
Eremolepidaceae 3 Monotaceae 1 Sapindaceae 158
Ericaceae 73 Moraceae 83 Sapotaceae 148
Erythroxylaceae 37 Moringaceae 1 Sarraceniaceae 10
Euphorbiaceae 317 Muntingiaceae 1 Scrophulariaceae 52
Euphroniaceae 3 Myoporaceae 1 Simaroubaceae 12
Flacourtiaceae 64 Myricaceae 1 Siparunaceae 10
Gentianaceae 92 Myristicaceae 33 Solanaceae 116
Gesneriaceae 94 Myrsinaceae 70 Sphenocleaceae 1
Haloragaceae 2 Myrtaceae 270 Sterculiaceae 67
Hemandiaceae 6 Nelumbonaceae 1 Styracaceae 22
Hippocrateaceae 52 Nyctaginaceae 50 Surianaceae 1
Hugoniaceae 8 Nymphaeaceae 15 Symplocaceae 13
Humiriaceae 30 Ochnaceae 165 Tepuianthaceae 4
Hydrophyllaceae 3 Olacaceae 35 Ternstroemiaceae 31
Icacinaceae 22 Oleaceae 4 Tetrameristaceae 1
Ixonanthaceae 10 Onagraceae 21 Theaceae 1
Juglandaceae 1 Opiliaceae 2 Theophrastaceae 4
Juncaceae 1 Oxalidaceae 12 Thymelaeaceae 15
Krameriaceae 3 Papaveraceae 1 Tiliaceae 37
Lacistemataceae 3 Passifloraceae 107 Trigoniaceae 12
Lamiaceae 38 Pedaliaceae 2 Tropaeolaceae 1
Lauraceae 214 Peridiscaceae 2 Tumeraceae 31
Lecythidaceae 67 Phytolaccaceae 12 Ulmaceae 5
Leguminosae Picramniaceae 9 Urticaceae 15
- Caesalpinioideae 294 Piperaceae 172 Valerianaceae 1
- Faboideae 500 Plantaginaceae 2 Verbenaceae 90
- Mimosoideae 238 Plumbaginaceae 2 Violaceae 39
Lentibulariaceae 55 Podostemaceae 67 Viscaceae 60
Lissocarpaceae 3 Polygalaceae 76 Vitaceae 16
Loasaceae 2 Polygonaceae 53 Vochysiaceae 84
Loganiaceae 43 Portulacaceae 15 Winteraceae 1
Loranthaceae 57 Primulaceae 1 Zygophyllaceae 5
Checklist of the Plants of the Guiana Shield 13
Table 2: Families with largest numbers of species.
Rank Family No. of Genera No. of Species
1 Leguminoseae 146 1032
- Mimosoideae 31 238
- Caesalpinioideae 41 294
- Faboideae 74 500
2 Orchidaceae 171 1020
3 Ferns* 95 754
4 Rubiaceae 97 742
5 Melastomataceae 54 534
6 Poaceae 116 526
7 Cyperaceae 37 413
8 Bromeliaceae 24 327
9 Euphorbiaceae* * 60 317
10 Compositae 104 303
11 Myrtaceae 18 270
12 Araceae 24 226
13 Lauraceae 18 214
14 Apocynaceae*** 38 211
15 Malpighiaceae 25 205
* Includes many families
** Includes the small families Putrangiivaceae, Picrodendraceae, and Phyllanthaceae
which do not belong in a monophyletic Euphorbiaceae
*** Does not include the Asclepidaceae (18 genera and 132 species)
The largest family on the Shield is the
Leguminoseae (= Fabaceae) with 1032 species.
The phylogeny of this family is under revision but
it is well enough understood to know that this
family has a rich and interesting history in the
Guiana Shield area. We know from the molecular
data that there are a number of genera that have
their primary radiation in the area and that there
are many endemics. Very little is known about the
phylogeny of the Guiana Shield members of the
other large family, the Orchidaceae.
In a broad scope of botanical understanding,
the combination of such checklists looks to the
evolutionary and ecological importance ofphysical
conditions beyond those of political units and
boundaries. This process is simply a step in the
direction of looking at a flora at the level of the
distinct geological unit of the Guiana Shield. In
the future there will be additional field trips and
further progress on the Flora of the Guianas
(Nationaal Herbarium Nederland 2006) which will
provide additions and corrections to this list. Also,
future developments will hopefully include the
ability to analyze plant community composition
on finer levels of the landscape, and to produce
customized checklists for analysis and
conservation with the aid of Geographic
Information System (GIS) technologies drawing
upon improved databases incorporating
georeferenced historical collections from many
herbaria.
USING THE CHECKLIST
For each family, accepted names are listed
followed by the known distributions.
Abbreviations used are Amazonas (AM), Bolivar
(BO), and Delta Amacuro (DA); and for the three
Guianas: Guyana (GU), Surinam (SU), and French
Guiana (FG).
Comments following the accepted name or
synonym are enclosed in brackets and include
aspects such as: exotic species, distributional
expectations or details, and in the case of
misidentifications an indication that the taxon is
not found in the flora area. Some taxa are listed
14 Checklist of the Plants of the Guiana Shield
with comments naming Mt. Roraima, Serrania de
la Neblina or other locaHties without distribution
codes. Specimens from these areas may lack a
country designation or may be incorrect or
unknown, especially in historical collections.
Alternatively, a species may be known from
neighboring Brazil and if it is thought that it is
likely to occur in the checklist area it is included
as part of the flora.
NOMENCLATURAL CONSIDERATIONS
Family names generally follow the
classification used by Cronquist ( 1 98 1 ) but the list
has been modified to reflect some recent changes
based on molecular and morphological data and
presented on the website ofPeter Stevens (Stevens,
2001). An occasional dilemma in combining plant
checklists is the necessity to choose among
classifications used by different specialists. When
decisions had to be made we selected the most
recent comprehensive work for the group in
question. However, our decision does not imply
an endorsement of one or rejection of another and
it is hoped that an adequate reference system has
been provided to allow the use of most systems
and to facilitate work towards future clarification.
Authors ofplant names are standardized according
to Brummitt and Powell (1992).
In some cases, a species has been reported
from the flora area either because ofa broad species
circumscription that has since been restricted (or
as an infraspecific taxon that has since been raised
to species status), or due to misidentifications or
other errors. Such names are listed in the
Synonyms and Misidentification section as "not
in the flora area."
In some ofthe cases source data for one region
lists subspecific taxa while data for the other region
lists only the specific level. In those cases, both
the specific and subspecific distributions are given
rather than eliminating the partial information for
the subspecific taxa.
When the list has been reviewed by a specialist
that persons name is listed under the taxon name.
All other Family lists were compiled by T.
HoUowell.
ACKNOWLEDGMENTS
Special thanks go to botanists at the Missouri
Botanical Garden (MO), particularly Kay
Yatskievych, James Miller, Ronald Liesner, Robert
Magill, and many others who assisted in the Flora
of the Venezuelan Guayana project and shared the
electronic manuscript with the BDG program.
Also, we thank several colleagues including Scott
Mori (NY) and Jean-Jacques de Granville (CAY)
who provided data in the development of the first
checklist of the three Guianas. In addition, our
work would not have been possible without the
dedication of botanical specialists from many
institutions who contributed to checklists,
taxonomic treatments, did countless identifications,
and reviewed drafts of the combined lists. Each
family treatment lists the name ofthe collaborating
specialists for that family and the Appendix lists
all of those who helped with identifications. We
owe a great debt to the many plant collectors and
their field crews who over more than two centuries
have provided specimens for the museum
collections essential to an understanding of the
flora of the Guiana Shield (Ek, 1990, 1991;
HoUowell et al. 2000, 2003, 2005). Equally
indispensable to studies of biodiversity are the
many institutions that have financed those
expeditions and those that house the collections,
and their staffmembers that maintain and organize
those collections. In addition to the home
institutions we must also thank the various funding
agencies that have made field work possible, chief
among these is the National Geographic Society
who funded quite a few expeditions for botanists
and zoologists and USAID in Guyana. In Guyana
the collecting program was hosted by the Centre
for the Study of Biological Diversity, University
ofGuyana, and we appreciate their help. Particular
thanks for assistance are given to Marilyn Hansel,
Christian Feuillet, Bob DeFilipps, Susan Grose,
Karen Redden, Matt Sewell, H. David Clarke, Julie
Barcelona, Greg McKee, Sylvia Stone, and Mark
Strong (all from US). Finally, we must thank Peter
Jorgensen (MO) who provided a careful review of
the manuscript.
This publication was prepared under the
auspices of the Biological Diversity of the Guiana
Shield Program. This is number 125 in the
Smithsonian's Biological Diversity of the Guiana
Shield publication series.
Checklist of the Plants of the Guiana Shield 15
REFERENCES
Arbelaez, M. v., and R. Callejas. 1999. Florula
de la Meseta de Arensica de la comunidad de
Monochoa (Region de Araracuara, Medio
Caquetd. Tropenbos - Colombia, Bogota.
Boggan, J., V. A. Funk, C. L. Kelloff, M. Hoff, G.
Cremers, & C. Feuillet. 1992. Checklist of
the Plants of the Guianas. Biological
Diversity of the Guianas Program,
Smithsonian Institution, Washington, DC. 381
pp.
Boggan, J., V. Funk, C. L. Kelloff, M. Hoff, G
Cremers, & C. Feuillet. 1997. Checklist of
the Plants of the Guianas, 2nd Edition.
Biological Diversity of the Guianas Program,
Smithsonian Institution, Washington, DC. 238
pp.
Brummitt, R. K., and C. E. Powell, eds. 1992.
Authors of Plant Names. Royal Botanic
Gardens, Kew.
Clarke, H. D., V. A. Funk, and T. Hollowell. 2001
.
Using checklists and collections data to
investigate plant diversity. I: A comparative
checklist of the plant diversity of the
Iwokrama Forest, Guyana. Sida Botanical
Miscellany 21: 1-86.
Cronquist, A. 1981. An Integrated System of
Classification ofFlowering Plants . Columbia
University Press, New York.
Doyle, A. C. 1912. TheLost World. PuffmBooks,
London
Ek, R. C. 1 990. Index ofGuyana Plant Collectors.
Flora of the Guianas, A.R.A. Gorts-van Rijn
ed.. Supplementary Fascicle 1. Koeltz
Scinetific Books, Koenigstein, Germany.
Ek, R. C. 1991. Index of Suriname Plant
Collectors. Flora of the Guianas, A.R.A.
Gorts-van Rijn ed.. Supplementary Fascicle
2. Koeltz Scinetific Books, Koenigstein,
Germany.
Funk, V. A. and P E. Berry. 2005. Chapter 4.3
The Guiana Shield. In Plant Conservation: A
natural history approach. University of
Chicago Press, pages 76-79.
Funk, V. A. and K. Richardson. 2002. Biological
Specimen Data in Biodiversity Studies: Use
it or lose it. Systematic Biology 5 1 : 303-3 1 6.
Funk, V. A., K. S. Richardson, and S. Ferrier. 2005.
Survey-gap analysis in expeditionary research,
where do we go from here? BiologicalJournal
oftheLinnean Society 85: 549-567.
Gibbs,A.K. and C.N. Barron. 1993. The Geology
of the Guiana Shield. Oxford University
Press, New York.
Hollowell, T., V. Funk, and C. Kelloff 2000
(onwards). Biological Diversity ofthe Guiana
Shield, www.mnh.si.edu/biodiversity/bdg
Hollowell, V. A. Funk, C. L. Kelloff & G
Gharbarran. 2000. Smithsonian Plant
Collections, Guyana: 1986-1987, John J.
Pipoly III. Smithsonian Institution,
Washington, D.C.
Hollowell, T., L. J. Gillespie, V. A. Funk, and C.
L. Kelloff 2003. Smithsonian Plant
Collections, Guyana: 1989-1991, Lynn J.
Gillespie. Contributionsfrom the UnitedStates
National Herbarium AA'. 1-104.
Hollowell, T., T. McDowell, V. A. Funk, C. L.
Kelloff, and D. Gopaul. 2005. Smithsonian
Plant Collections, Guyana: 1990-1991, Tim
McDowell. Contributions from the United
States National Herbarium 50: 1-150.
Hollowell, T. and R. P. Reynolds, eds. 2005.
Checklist of the Terrestrial Vertebrates of the
Guiana Shield. Bulletin of the Biological
Society of Washington, no. 12.
Huber, O. 1995a. Geography and Physical
Features. Pp 1-61 in: PE. Berry, B.K. Hoist,
and K. Yatskievych (eds.). Flora of the
Venezuelan Guayana. Volume 1 : Introduction.
J.A. Steyermark, Paul E. Berry, and B.K. Hoist
(general eds.). Missouri Botanical Garden, St.
Louis.
Huber, O. 1995b. Vegetation. Pp 97-160 in: PE.
Berry, B.K. Hoist, and K. Yatskievych (eds.),
Flora ofthe Venezuelan Guayana. Volume 1
:
Introduction. J.A. Steyermark, PE. Berry, and
B.K. Hoist (general eds.) Missouri Botanical
Garden, St. Louis.
Huber, O., G Gharbarran, and VA. Funk. 1995.
Preliminary vegetation map of Guyana.
Biological diversity of the Guianas program,
Smithsonian Institution, Washington, D.C.
Kelloff, C. L. 2003. The use of biodiversity data
in developing Kaieteur National Park, Guyana
for ecotourism and conservation.
Contributions to the Study of Biological
Diversity 1: 1-44. University of Guyana,
Georgetown.
Kelloff, C. L., and VA. Funk. 2004.
16 Checklist of the Plants of the Guiana Shield
Phytogeography ofthe Kaieteur Falls, Potaro
Plateau, Guyana: floral distributions and
affinities. Journal ofBiogeography 31: 501-
513.
Leechman, A. 1913. The British Guiana
Handbook. "The Argosy" Co., Ltd.;
Georgetown, British Guiana and Dulau & Co.,
London.
Lindeman, J. C, and S. A. Mori. 1989. The
Guianas. In: D.G Campbell and H.D.
Hammond (eds.), Floristic Inventory of
Tropical Countries. New York Botanical
Garden, New York. pp. 375-391. ,
Mendoza, V. 1977. Petrogenesis de rocas
volcanicas Precambrics del noroeste del
Escudo de Guayana Venezuela. Mem. V
Congreso Geol. Venezolano, Caracas, Venez.
Tomo II, Pp. 555-589.
Nationaal Herbarium Nederland. 2006. www.bio.
uu.nl/~herba/Guyana/Flora_of_the_Guianas
Schubert, C, and O. Huber. 1990. The Gran
Sabana: Panorama of a Region. Caracas:
Lagoven Booklets.
Snow, J. W. 1976. Climates of northern South
America. In: W. Schwerdtfeger (ed.), Climates
of Central and South America. Elsevier
Scientific Publishing Company, Amsterdam, pp
295-403.
Stevens, P. F. 2001 (onwards). Angiosperm
Phylogeny Website. www.mobot.org/MOBOT/
research/APweb/.
Steyermark, J. A., P. E. Berry, and B. K. Hoist, eds.
1995-2005. Flora ofthe Venezuelan Guayana,
9 vols. Missouri Botanical Garden, St. Louis.
Guapira eggersiana (Heimerl) Lundell (Nyctaginaceae)
Checklist of the Plants of the Guiana Shield 17
CHECKLIST OF THE PLANTS OF THE GUIANA SHIELD
PTERIDOPHYTES & ALLIES
G. Cremers, R.J. Hickey, D.B. Lellinger, J.T. Mickel, R.C. Moran, B. 011gaard, A.R. Smith
PSILOTOPHYTA
PSILOTACEAE
Psilotum nudum (L.) R Beauv. AM GU SU FG
Synonyms and Misidentifications
Psilotum triquetrum Sw. =Psilotum nudum
References
Kramer, K.U. 1990. Psilotaceae. In K.U. Kramer & RS. Green, Families and Genera of Vascular Plants. 1. Pteridophytes and
Gymnosperms. New York: Springer-Verlag.
Smith, A.R. 1995. Psilotaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 250-251.
LYCOPHYTA
General references
Berry, RE., B.K. Hoist, & K. Yatskievych. 1995. Flora of the Venezuelan Guayana, vol. 2. Pteridophytes, Spermatophytes
Acanthaceae - Araceae. Missouri Botanical Garden, St. Louis, & Timber Press, Portland, Oregon.
Cremers, G. 1997. Lycophyta. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide
to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs of the
New York Botanical Garden 76, pt. 1 : 56-65.
Cremers, G, C. Feuillet, J.J. de Granville, & M. Hoff. 1988. Liste des especes de Phanerogames et de Pteridophytes de la
Guyane fran9aise d'apres I'Herbier du Centre ORSTOM de Cayenne. ORSTOM, Cayenne.
Cremers, G. & M. Hoff. 1990. Inventaire taxonomique des plantes de la Guyane fran9aise. 1. Les Pteridophytes. Mus. Nat. Hist.
Nat., Inventaires de Faune et de Flore 54: 1-133.
Kramer, K.U. 1978. The Pteridophytes of Suriname. An enumeration with keys of the ferns and fern-allies. Uitgaven
"Natuurwetenschappelijke Studiekring voor Suriname en de Nederlandse Antillen" Utrecht no. 93.
Kramer, K.U. & RS. Green (ed.). 1990. The Families and Genera of Vascular Plants. I. Pteridophytes and Gymnosperms. New
York: Springer-Verlag.
Posthumous, O. 1928. The Ferns of Surinam and of French and British Guiana. In A. Pulle, Flora of Suriname supplement: 1-
196.
Tryon, R.M. & A.F. 1981. Ferns and Allied Plants. New York: Springer-Verlag.
ISOETACEAE
Isoetes killipii C.V Morton [Mt. Roraima] BO
Isoetes ovata N. Pfeiff.
Isoetes schinzii H.P. Fuchs [nomem nudum]
Isoetes triangula U. Weber AM BO
Isoetes vermiculata Hickey AM
Synonyms and Misidentifications
Isoetes ovatum auct. non N. Pfeiff: Vareschi (pro parte) =Isoetes killipii
References
Fuchs-Eckert, H.P. von. 1982. Zur heutigen Kenntnis von Vorkommen und Verbreitung der Sudamerikanischen Isoetes-Arten.
Proc. Kon. Ned. Akad. Wetensch. (ser. C) 85: 205-260.
Hickey, R.J. & A.R. Smith. 1995. Isoetaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana
2: 189-190.
Lycopodiaceae
Huperzia aqualupiana (Spring) Rothm. BO GU
Huperzia beitelii B. 011g. [Mt. Roraima] AM BO
GU
GU FG
GU FG
GU
18 Checklist of the Plants of the Guiana Shield
Huperzia capillaris (Sodiro) Holub
Huperzia dichotoma (Jacq.) Trevis.
Huperziafuniformis (Cham, ex Spring) Trevis.
Huperzia hippuridea (H. Christ) Holub
Huperzia huberi B. 011g.
Huperzia intermedia (Spring) Trevis. [Mt. Roraima]
Huperzia killipii (Herter) B. 011g.
Huperzia linifolia (L.) Trevis.
Huperzia linifolia var. jenmanii (Underw. & F.E. Lloyd) B. 011g. & P.G
Windisch
Huperzia linifolia van linifolia
Huperzia myrsinites (Lam.) Trevis.
Huperzia recurvifolia Rolleri
Huperzia reflexa (Lam.) Trevis. [Mt. Roraima]
Huperzia robusta (Klotzsch) Holub
Huperzia taxifolia (Sw.) Trevis.
Huperzia tenuicaulis (Underw. & F.E. Lloyd) B. 011g.
Huperzia wilsonii (Underw. & F.E. Lloyd) B. 011g.
Lycopodiella alopecuroides (L.) Cranfill
Lycopodiella alopecuroides var. integerrima (Spring) B. 011g. & RG.
Windisch
Lycopodiella camporum B. 011g. & RG. Windisch
Lycopodiella caroliniana (L.) Pic. Serm.
Lycopodiella caroliniana var. meridionalis (Underw. & F.E. Lloyd)
B. 011g. & RG. Windisch
Lycopodiella caroliniana var. paradoxa (Mart.) B. 011g. & P.G. Windisch
Lycopodiella cernua (L.) Pic. Serm.
Lycopodiella cernua var. cernua
Lycopodiella cernua var. curvata (Sw.) Kartesz & Gandhi
Lycopodiella contexta (Mart.) Holub
Lycopodiella iuliformis (Underw. & F.E. Lloyd) B. 011g.
Lycopodiella iuliformis var. iuliformis
Lycopodiella iuliformis var. tatei (A.C. Sm.) B. 011g. [Mt. Roraima]
Lycopodiella riofrioi (Sodiro) B. 011g.
Lycopodiella steyermarkii B. 011g.
Lycopodium clavatum L.
Lycopodium clavatum subsp. clavatum
Lycopodium clavatum subsp. contiguum (Klotzsch) B. 011g.
Lycopodium Jussiaei Poir.
Synonyms and Misidentifications
Diphasiumjussiaei (Desv. ex Poir.) Rothm. =Lycopodium jussiaei
Huperziajenmanii (Underw. & F.E. Lloyd) Holub =Huperzia linifolia var. jenmanii
Huperzia passerinoides (Kunth) Trevis. =Huperzia taxifolia
Huperzia phelpsii (Vareschi) Holub =Huperzia robusta
Huperzia underwoodiana (Maxon) Holub =Huperzia capillaris
Lepidotis cernua (L.) P. Beauv. =Lycopodiella cernua
Lepidotis contexta (Mart.) Rothm. =Lycopodiella contexta
Lycopodiella duidae (A.C. Sm.) B. 011g. =Lycopodiella iuliformis var. tatei
Lycopodium alopecuroides L. =Lycopodiella alopecuroides
Lycopodium anomalum Hook. & Grev. (Selaginellaceae) =Selaginella porelloides
Lycopodium aqualupianum Spring =Huperzia aqualupiana
Lycopodium aristatum Willd. =Lycopodium clavatum
Lycopodium barbatum H. Christ =Huperzia dichotoma
Lycopodium capillaceum (Spring) Hieron. =Lycopodiella cernua
Lycopodium capillare Sodiro =Huperzia capillaris
Lycopodium carolinianum L. =Lycopodiella caroliniana
Lycopodium carolinianum var. meridionale (Underw. & F.E. Lloyd) Nessel =Lycopodiella caroliniana var. meridionalis
Lycopodium carolinianum var. paradoxum (Mart.) H. Christ =Lycopodiella caroliniana var. paradoxa
Lycopodium cernuum L. =Lycopodiella cernua
Lycopodium cernuum var. capillaceum Spring =Lycopodiella cernua
Lycopodium chamaepeuce Herter =Huperzia dichotoma
Lycopodium contextum Mart. =Lycopodiella contexta
Lycopodium contiguum Klotzsch =Lycopodium clavatum subsp. contiguum
Lycopodium convolutum Am. (Selaginellaceae) =Selaginella convoluta
BO
BO DA GU SU FG
BO GU
AM BO
AM BO
BO
AM BO FG
AM BO DA GU su FG
AM BO DA GU SU FG
BO GU FG
AM BO GU
AM BO GU
AM BO
AM BO
AM BO GU
GU
su FG
AM BO su
AM BO GU
GU
su
su
AM BO GU su FG
AM BO GU su FG
AM BO
BO
GU su FG
AM BO GU su FG
GU su FG
GU?
AM BO
AM BO GU su
AM BO GU su
AM BO
AM BO
AM BO
AM BO GU
AM BO
BO GU
AM BO
Checklist of the Plants of the Guiana Shield 19
Lycopodium dichotomum Jacq. =Huperzia dichotoma
Lycopodium duidae A.C. Sm. =Lycopodiella iuliformis var. tatei
Lycopodium funiforme Cham, ex Spring =Huperzia funiformis
Lycopodium gramineum Spring =Huperzia dichotoma
Lycopodium guatemalense Maxon =Huperzia capillaris
Lycopodium hippuhdeum H. Christ =Huperzia hippuridea
Lycopodium intermedium Spring =Huperzia intermedia
Lycopodium iuliforme Underw. & F.E. Lloyd =Lycopodiella iuliformis
Lycopodiumjenmanii Underw. & F.E. Lloyd =Huperzia linifolia var.jenmanii
Lycopodium killipii (Herter) C.V. Morton =Huperzia killipii
Lycopodium linifolium L. =Huperzia linifolia
Lycopodium marginatum Humb. & Bonpl. ex Willd. (Selaginellaceae) =Selaginella marginata
Lycopodium meridionale Underw. & F.E. Lloyd =Lycopodiella caroliniana var. meridionalis
Lycopodium myrsinites Lam. =Huperzia myrsinites
Lycopodium nudum L. (Psilotaceae) =Psilotum nudum
Lycopodium paradoxum Mart. =Lycopodiella caroliniana var. paradoxa
Lycopodium parkeri Hook. & Grev. (Selaginellaceae) =Selaginella parkeri
Lycopodium passerinoides Kunth =Huperzia taxifolia
Lycopodium penniforme Lam. (Selaginellaceae) =Selaginella radiata
Lycopodium pensum Lellinger & Mickel =Lycopodiella riofrioi
Lycopodium phelpsii Vareschi =Huperzia robusta
Lycopodium plumosum sensu Aubl., non L. (Selaginellaceae) =Selaginella epirrhizos
Lycopodium porelloides Lam. (Selaginellaceae) =Selaginella porelloides
Lycopodium radiatum Aubl. (Selaginellaceae) =Selaginella radiata
Lycopodium reflexum Lam. =Huperzia reflexa
Lycopodium reflexum Lam. var. caesium Vareschi [nom. inval.] =Huperzia reflexa
Lycopodium riofrioi Sodiro =Lycopodiella riofrioi
Lycopodium robustum Klotzsch =Huperzia robusta
Lycopodium roraimense Underw. & F.E. Lloyd =Huperzia myrsinites
Lycopodium skutchii Maxon =Huperzia myrsinites
Lycopodium sprucei Baker =Lycopodiella contexta
Lycopodium tatei A.C. Sm. =Lycopodiella iuliformis var. tatei
Lycopodium taxifolium Sw. =Huperzia taxifolia
Lycopodium underwoodianum Maxon =Huperzia capillaris
Lycopodium wilsonii Underw. & F.E. Lloyd =Huperzia wilsonii
Palhinhaea capillacea (Spring) Holub =Lycopodiella cernua
Palhinhaea cernua (L.) Vase. & Franco =Lycopodiella cernua
Phlegmariurus taxifolius (Sw.) A. Love & D. Love -Huperzia taxifolia
Pseudolycopodiella caroliniana (L.) Holub =Lycopodiella caroliniana
Pseudolycopodiella iuliformis (Underw. & F.E. Lloyd) Holub =Lycopodiella iuliformis
Pseudolycopodiella meridionalis (Underw. & F.E. Lloyd) Holub =Lycopodiella caroliniana var. meridionalis
Pseudolycopodiella paradoxa (Mart.) Holub =Lycopodiella caroliniana var paradoxa
Urostachys commutatus Herter =Huperzia intermedia
Urostachys hippurideus (H. Christ) Nessel =Huperzia hippuridea
Urostachys intermedius (Trevis.) Herter =Huperzia intermedia
Urostachys killipii Herter =Huperzia killipii
Urostachys linifolius (L.) Herter =Huperzia linifolia
Urostachys myrsinites (Lam.) Herter =Huperzia myrsinites
Urostachys reflexus (Lam.) Herter =Huperzia reflexa
Urostachys roraimensis (Underw. & F.E. Lloyd) Nessel =Huperzia myrsinites
Urostachys wilsonii (Underw. & F.E. Lloyd) Herter =Huperzia wilsonii
References
011gaard, B. 1987. A revised classification of the Lycopodiaceae s.l. Opera Bot. 92: 153-178.
011gaard, B. 1995. Lycopodiaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 190-206.
011gaard, B. & RG. Windisch. 1987. Sinopse das Licopodiaceas do Brasil. Bradea 5(1): 1-43.
Selaginellaceae
Selaginella albolineata A.R. Sm. AM
Selaginella amazonica Spring AM GU
Selaginella anaclasta Alston ex Crabbe & Jermy AM
Selaginella applanata A. Braun AM
Selaginella arrecta A.R. Sm. AM BO
20 Checklist of the Plants of the Guiana Shield
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Selagine,
Selagine
Selagine,
Selagine
Selagine
Selagine
Selagine
Selagine
Selagine
Selagine
Selagine
Selagine
Selagine
Selagine
Selagine,
Selagine
Selagine
Selagine,
Selagine,
Selagine,
Selagine,
Selagine.
Selagine.
Selagine.
Selagine.
Selagine.
Selagine
Selagine
Selagine
Selagine
Selagine
Selagine
Selagine,
Selagine,
Selagine.
Selagine
Selagine
Selagine
Selagine,
Selagine,
Selagine,
Selagine.
la arthritica Alston
la aspenila Spring
la beitelii A.R. Sm.
la brevifolia Baker
la breweriana A.R. Sm.
la breynii Spring
la cabrerensis Hieron.
la calceolata Jermy & J.M. Rankin
la cardiophylla Valdespino
la cladorrhizans A. Braun
la coarctata Spring
la conduplicata Spring
la convoluta (Am.) Spring
la cyclophylla A.R. Sm.
la dendricola Jenman
la densifolia Spruce ex Hook.
la diffusa (C. Presl) Spring
la epirrhizos Spring
lafalcata (R Beauv.) Spring
laflabellata (L.) Spring
laflagellata Spring
lafragilis A. Braun
la hemicardia Valdespino
la hirsuta Alston ex Crabbe & Jermy
la humboldtiana A. Braun
la imbricans A.R. Sm.
la karowtipuensis Valdespino, sp. nov. ined.
la kochii Hieron.
la kunzeana A. Braun
la lechleri Hieron.
a marahiiacae A.R. Sm.
a marginata (Humb. & Bonpl. ex Willd.) Spring
la mazariiniense Jenman
la microdonta A.C. Sm.
la minima Spring
la muscosa Spring
la neblinae A.R. Sm.
la novae-hollandiae (Sw.) Spring
la pallescens (C. Presl) Spring
la palmiformis Alston ex Crabbe & Jermy
la parkeri (Hook. & Grev.) Spring
la plana (Desv. ex Poir.) Hieron. [introduced]
la porelloides (Lam.) Spring
la porphyrospora A. Braun
la potaroensis Jenman
la producta Baker
la pruskiana Valdespino
la pubens A.R. Sm.
la radiata (Aubl.) Spring
la revoluta Baker
la rhodostachya Baker
la roraimensis Baker
la sandwithii Alston
la scalariformis A.C. Sm.
la schultesii Alston ex Crabbe & Jermy
la seemannii Baker
la smithiorum Valdespino
la sobolifera A.R. Sm.
la sp. A
la substipitata Spring
la sulcata (Desv. ex Poir.) Spring ex Mart.
la terezoana Bautista
la thysanophylla A.R. Sm.
la tuberculata Spruce ex Baker
BO
BO
BO
BO
BO
AM
BO
AM
AM BO
AM
BO
AM
AM
AM
AM
AM
AM
AM BO
AM BO
AM
AM
AM
AM BO
BO
BO
AM BO
AM BO
AM
AM
AM BO
BO
AM BO
AM BO
AM
BO
AM
BO
AM BO
BO
BO
BO
DA
DA
DA
GU
GU
GU
GU
GU
SU
FG?
FG
FG
GU
GU SU FG
GU FG?
GU SU FG
SU? FG?
SU FG
GU SU FG
GU?
SU
FG
GU SU FG
FG
FG
FG?
SU
GU SU FG
GU
GU FG
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU
GU
GU SU FG
GU
GU SU FG
GU
GU SU FG
GU
GU SU FG
AM
AM BO
AM BO GU
AM BO GU
AM
AM BO
Checklist of the Plants of the Guiana Shield 21
Selaginella tyleri A.C. Sm.
Selaginella umbrosa Lem. ex Hieron.
Selaginella valdepilosa Baker
Selaginella vernicosa Baker
Selaginella versatilis A.R. Sm.
Selaginella wurdackii Alston
Synonyms and Misidentifications
Selaginella affinis A. Braun =Selaginella epirrhizos
Selaginella brachyclada Baker =Selaginella rhodostachya
Selaginella brevispicata Hieron. ex Bautista =Selaginella rhodostachya
Selaginella calcarata A. Braun =Selaginella conduplicata
Selaginella caudorhiza Baker =Selaginella sulcata
Selaginella cayennensis Baker =Selaginella producta
Selaginella diminiitifolia Jenman =Selaginella rhodostachya
Selaginella duidae A.C. Sm. ^Selaginella scalariformis
Selaginella guianensis Spring =Selaginellafalcata
Selaginellajenmanii Baker =Selaginella porelloides
Selaginella karsteniana A. Braun =Selaginella siibstipitata
Selaginella lemairei Hieron. =Selaginella umbrosa
Selaginella lucidinervia Spring =Selaginella parkeri
Selaginella macroclada Baker =Selaginella parkeri
Selaginella parkeri Spring var. pedata (Klotzsch) Baker =Selaginella parkeri
Selaginella pedata Klotzsch =Selaginella parkeri
Selaginella penniformis (Lam.) Hieron. =SelagineUa radiata
Selaginella platyphylla Baker =Selaginella porelloides
Selaginella platyphylla var. laxa Jenman =Selaginella porelloides
Selaginella poeppigiana var. guyanensis Spring =Selaginella epirrhizos
Selaginella puberula Klotzsch ^Selaginella coarctata
Selaginella rigidula Baker =Selaginella substipitata
Selaginella scintillata Alston =Selaginella roraimensis
Selaginella stellata Spring =Selaginella conduplicata
Selaginella valdepilosa subsp. tricholoma Jermy & J.M. Rankin =Selaginella rhodostachya
Selaginella vernicosa var. oligoclada Baker =Selaginella vernicosa
References
Alston, A.H.G., A.C. Jermy, & J.M. Rankin. 1981. The genus Selaginella in tropical South America. Bull. Brit. Mus. (Nat. Hist.)
Bot. 9: 233-330.
Smith, A.R. 1995. Selaginellaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 296-314.
PTEROPHYTA
General references
Berry, RE., B.K. Hoist, & K. Yatskievych. 1995. Flora of the Venezuelan Guayana, vol. 2. Pteridophytes, Spermatophytes
Acanthaceae - Araceae. Missouri Botanical Garden, St. Louis, & Timber Press, Portland, Oregon.
Cremers, G 1997. Pterophyta. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1 : 65-162.
Cremers, G, C. Feuillet, J.J. de Granville, & M. Hoff. 1988. Liste des especes de Phanerogames et de Pteridophytes de la
Guyane fran9aise d'apres I'Herbier du Centre ORSTOM de Cayenne. ORSTOM, Cayenne.
Cremers, G & M. Hoff. 1990. Inventaire taxonomique des plantes de la Guyane fran^aise. 1. Les Pteridophytes. Mus. Nat. Hist.
Nat., Inventaires de Faune et de Flore 54: 1-133.
Jenman, GS. 1909. The ferns and fern allies of the British West Indies and Guiana. Trinidad: Bull. Misc. Int. Bot. Dept.
Kramer, K.U. 1978. The Pteridophytes of Suriname. An enumeration with keys of the ferns and fern-allies. Uitgaven
"Natuurwetenschappelijke Studiekring voor Suriname en de Nederlandse Antillen" Utrecht no. 93.
Kramer, K.U. & P.S. Green, eds. 1990. The Families and Genera of Vascular Plants. 1. Pteridophytes and Gymnosperms. New
York: Springer-Verlag.
Lellinger, D.B. 1989. The ferns and fern-allies of Costa Rica, Panama, and the Choco. Part 1: Psilotaceae through Dicksoniaceae.
Pteridologia2A: 1-364.
Mickel, J.T., and J.M. Beitel. 1988. Pteridophytc flora of Oaxaca, Mexico. Memoirs of the New York Botanical Garden 46: 1-
568.
Pichi Sermolli, R.E.G 1977. Tentamen Pteridophytorum genera in taxonomicum ordinem redigendi. Webbia 31: 313-512.
Proctor, GR. 1977. Pteridophytes. Vol. 2 in Flora of the Lesser Antilles, edited by R. A. Howard. Cambridge, Massachusetts:
Harvard University.
Proctor, GR. 1985. Ferns of Jamaica. London: British Museum (Natural History).
22 Checklist of the Plants of the Guiana Shield
Proctor, G.R. 1989. Ferns of Puerto Rico and the Virgin Islands. Memoirs of the New York Botanical Garden 53: 1-389.
Posthumous, O. 1928. The Ferns of Surinam and of French and British Guiana. In A. Pulle, Flora of Suriname supplement: 1-
196.
Smith, A.R. 1981. Pteridophytes. Vol. 2 in Flora of Chiapas, edited by D. E. Breedlove. San Francisco: California Academy of
Sciences.
Tryon, R.M. 1972. Endemic areas and geographic speciation in tropical American ferns. Biotropica 4: 121-131.
Tryon, R.M., and R.G. Stolze. 1989. Pteridophyta of Peru. Part I. 1. Ophioglossaceae— 12. Cyatheaceae. Fieldiana, Bot., n.s. 20:
1-145.
Tryon, R.M., and A.F. Tryon. 1982. Ferns and Allied Plants with Special Reference to Tropical America. New York: Springer-
Verlag.
Vareschi, V. 1968. Helechos. In T. Lasser, Flora de Venezuela 1: 1-1033.
Adiantaceae
Adiantopsis monticola (Gardner) T. Moore
Adiantopsis radiata (L.) Fee
Adiantum adiantoides (J. Sm.) C. Chr.
Adiantum amazonicum A.R. Sm.
Adiantum argutum Splitg.
Adiantum asperum (Poir.) Desv.
Adiantum cajennense Willd. ex Klotzsch
Adiantum concinnum Humb. & Bonpl. ex Willd.
Adiantum cordatum Maxon [specimens ident. as sp. nov. by G. Cremers]
Adiantum decoratum Maxon & Weath.
Adiantum deflectens Mart.
Adiantumfructuosum Poepp. ex Spreng.
Adiantum fuliginosum Fee
Adiantum glaucescens Klotzsch
Adiantum glaziovii Baker
Adiantum humile Kunze
Adiantum latifolium Lam.
Adiantum leprieurii Hook.
Adiantum lucidum (Cav.) Sw.
Adiantum hmulatum Burm. f.
Adiantum macrophyllum Sw.
Adiantum nudum A.R. Sm.
Adiantum obliquum Willd.
Adiantum olivaceum Baker
Adiantum oyapokense Jenman
Adiantum paraense Hieron.
Adiantum petiolatum Desv.
Adiantum phyUitidis J. Sm.
Adiantum pulverulentum L.
Adiantum serratodentatum Humb. & Bonpl. ex Willd.
Adiantum terminatum Kunze ex Miq. [possibly conspecific with Adiantum
humile}
Adiantum tetraphyllum Humb. & Bonpl. ex Willd.
Adiantum tomentosum Klotzsch
Adiantum tripteris K.U. Kramer
Adiantum villosum L.
Cassebeera pinnata Kaulf.
Cheilanthes bonahensis (Willd.) Proctor
Cheilanthes concolor (Langsd. & Fisch.) R.M. Tryon & A.F. Tryon
Cheilanthes eriophora (Fee) Mett.
Doryopteris collina (Raddi) J. Sm.
Doryopteris conformis K.U. Kramer & R.M. Tryon
Doryopteris cyclophylla A.R. Sm.
Doryopteris davidsei A.R. Sm.
Doryopteris lomariacea Kunze ex Klotzsch
Doryopteris pedata (L.) Fee
Doryopteris pedata var. multipartita (Fee) R.M. Tryon
Doryopteris pedata var. palmata (Willd.) Hicken
Doryopteris rediviva Fee
Doryopteris sagittifolia (Raddi) J. Sm.
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
AM BO
BO
AM BO
AM BO
BO
BO
BO
BO
AM BO
BO
AM
AM BO
BO
AM BO
AM BO
AM BO
AM
AM BO
AM BO
BO
BO
AM BO
AM
BO
AM
BO
BO
BO
BO
AM BO
DA
DA
DA
DA
DA
DA
DA
GU
GU SU FG
GU SU FG
GU SU
GU SU FG
GU SU FG
FG
FG
FG
GU SU FG
GU SU FG
GU SU FG
FG
SU FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU
FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU SU
SU
FG
GU SU
GU SU
GU
GU SU
GU SU
GU SU
FG?
FG
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
AM BO
AM BO
AM BO
AM
AM
AM BO
AM BO
AM
AM BO
BO
BO
AM
AM BO
GU
GU
GU
GU
GU
GU?
GU SU FG
GU SU
SU
FG
GU SU FG
GU
GU SU FG
Checklist of the Plants of the Guiana Shield 23
Doryopteris varians (Raddi) J. Sm.
Ehosonis flexuosus (Kunth) Copel.
Eriosorusflexuosus \2ir. flexuosus
Eriosorus hispidulus (Kunze) Vareschi
Eriosorus hispidulus var. hispidulus
Eriosorus paucifolius (A.C. Sm.) Vareschi [Mt. Roraima]
Hemionitis palmata L.
Hemionitis rufa (L.) Sw. BO
Notholaena solitaria R.M. Tryon
Pityrogramma calomelanos (L.) Link AM BO DA
Pityrogramma calomelanos var. aureoflava (Hook.) Weath. ex L.H. Bailey
Pityrogramma calomelanos var. calomelanos DA
Pityrogramma ebenea (L.) Proctor
Pityrogramma ebenea var. ebenea [Mt. Roraima]
Pityrogramma trifoliata (L.) R.M. Tryon
Pterozonium brevifrons (A.C. Sm.) Lellinger GU
Pterozonium cyclophyllum (Baker) Diels GU
Pterozonium cyclosorum A.C. Sm.
Pterozonium elaphoglossoides (Baker) Lellinger GU
Pterozonium lineare Lellinger
Pterozonium maguirei Lellinger
Pterozonium paraphysatum (A.C. Sm.)Le\\'mger GU SU
Pterozonium reniforme (Mart.) Fee GU
Pterozonium retroflexum Mickel
Pterozonium scopulinum Lellinger GU
Pterozonium spectabile Maxon & A.C. Sm.
Pterozonium steyermarkii Vareschi
Pterozonium tatei A.C. Sm.
Pterozonium terrestre Lellinger
Synonyms and Misidentifications
Adiantum dolosum Kunze (considered valid in FVG) =Adiantum asperum
Adiantum guianense Aubl. (Dennstaedtiaceae) =Lindsaea guianensis
Adiantum hirtum Klotzsch =Adiantum terminatum
Adiantum lancea L. (Dennstaedtiaceae) =Lindsaea lancea
Adiantum lunulatum Houtt. (Dryopteridaceae) =Didymochlaena truncatula
Adiantum philippense L. =Adiantum lunulatum
Adiantum propinquum Fee =Adiantum pulverulentum
Adiantum radiatum L. =Adiantopsis radiata
Adiantum sagittatum Aubl. (Dennstaedtiaceae) =Lindsaea sagittata
Adiantum strictum Sw. (Dennstaedtiaceae) =Lindsaea stricta
Cheilanthes glandulosa Sw. (Dennstaedtiaceae) =Paesia glandulosa
Cheilanthes radiata (L.) J. Sm. =Adiantopsis radiata
Cheilanthes viscosa H. Karst. [nomen nudum] (Dennstaedtiaceae) =Hypolepis viscosa
Doryopteris concolor (Langsd. & Fisch.) Kuhn =Cheilanthes concolor
Doryopteris palmata (Willd.) J. Sm. =Doryopteris pedata var. palmata
Eriosorus ottonis (Kunze ex Klotzsch) Vareschi =Eriosorus hispidulus var. hispidulus
Eriosorus paucifolius var. neblinae A.F. Tryon =Eriosorus paucifolius
Eriosorus paucifolius var. steyermarkii A.¥. Tryon =Eriosorus paucifolius
Eriosorus schomburgkianus (Kunze ex Klotzsch) Copel =Eriosorus hispidulus var. hispidulus
Gymnogramma cyclophylla Baker =Pterozonium cyclophyllum
Gymnogramma diplazioides Desv. (Thelypteridaceae) =Thelypteris linkiana
Gymnogramma elaphoglossoides Baker =Pterozonium elaphoglossoides
Gymnogrammaflexuosa Desv. =Eriosorusflexuosus
Gymnogramma hispidula (Kunze) Klotzsch =Eriosorus hispidulus
Gymnogramma ottonis Kunze ex Klotzsch =Eriosorus hispidulus var. hispidulus
Gymnogramma reniformis Mart. =Pterozonium reniforme
Gymnogramma rufa (L.) Desv. =Hemionitis rufa
Gymnogramma schomburgkiana Kunze ex Klotzsch =Eriosorus hispidulus var. hispidulus
Gymnopteris rufa (L.) Bernh. ex Underw. =Hemionitis rufa
Gymnopteris semipinnatifida Fee (Lomariopsidaceae) =Bolbitis semipinnatifida
Hemionitis cajenensis Desv. (Vittariaceae) =Antrophyum cajenense
Hemionitis lineata Sw. (Vittariaceae) =Antrophyum lineatum
Jamesonia hispidula Kunze =Eriosorus hispidulus
Notholaena aurea (Poir.) Desv. =Cheilanthes bonariensis
24 Checklist of the Plants of the Guiana Shield
Notholaena eriophora Fee =Cheilanthes eriophora
Notholaena steyermarkii Vareschi =Cheilanthes eriophora
Pellaea lomariacea (Kunze ex Klotzsch) Hook. =Doryopteris lomariacea
Pellaea pinnata (Kaulf.) Prantl (Pteridaceae) =Cassebeera pinnata
Pityrogramma tartarea (Cav.) Maxon =Pityrogramma ebenea
Psilogramme paucifolia A.C. Sm. =Eriosorus paucifolius
Pterozonium reniforme var. ulei Brause =Pterozonium reniforme
Pterozonium schneei Vareschi =Pterozonium cyclophyllum
Pterozonium spathulatum Vareschi =Pterozonium cyclophyllum
Syngramma brevifrons A.C. Sm. =Pterozonium brevifrons
Syngramma elaphoglossoides (Baker) Diels ^Pterozonium elaphoglossoides
Syngramma paraphysata A.C. Sm. =Pterozomum paraphysatum
Syngramma versicolor Vareschi =Pterozonium brevifrons
Syngrammatopsis elaphoglossoides (Baker) Alston =Pterozonium elaphoglossoides
Syngrammatopsis paraphysata (A.C. Sm.) Alston =Pterozonium paraphysatum
Trismeria trifoliata (L.) Diels =Pityrogramma trifoliata
References
Lellinger, D.B. 1967. Pterozonium. In B. Maguire & collaborators, Botany of the Guayana Highland part 7. Mem. New York
Bot. Card. 17(1): 1-439.
Lellinger, D.B. 1972. Polypodiaceae subfam. Adiantoideae. In B. Maguire & collaborators, Botany of the Guayana Highland
part 9. Mem. New York Bot. Gard. 23: 1-832.
Smith, A.R. & D.B. Lellinger. 1995. Pteridaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana
2: 250-286.
Tryon, A.F. 1970. A monograph of the fern genus Eriosorus. Contrib. Gray Herb. 200: 54-174.
Tryon, R.M. 1942. A revision of the genus Doryopteris. Contrib. Gray Herb. 143: 1-80.
ASPLENIACEAE
Asplenium abscissum Willd.
Asplenium angustum Sw.
Asplenium auriculatum Sw.
Asplenium auritum Sw.
Asplenium chimantae A.R. Sm.
Asplenium claussenii Hieron.
Asplenium cowanii A.R. Sm.
Asplenium cristatum Lam.
Asplenium cuneatum Lam.
Asplenium cuspidatum Lam.
Asplenium delitescens (Maxon) L.D. Gomez
Asplenium dissectum Sw.
Asplenium feci Kunze ex Fee
Asplenium flabellulatum Kunze
Aspleniumformosum Willd.
Asplenium hallii Hook.
Asplenium harpeodes Kunze
Asplenium hastatum Klotzsch ex Kunze
Asplenium hostmannii Hieron.
Aspleniumjuglandifolium Lam.
Asplenium laetum Sw.
Asplenium macilentum Kunze ex Klotzsch
Asplenium pearcei Baker
Asplenium pedicularifolium A. St.-Hil.
Asplenium praemorsum Sw.
Asplenium pteropus Kaulf.
Asplenium pumilum Sw.
Asplenium radicans L.
Asplenium radicans var. cirrhatum (Rich, ex Willd.) Rosenst.
Asplenium radicans var. radicans
Asplenium repandulum Kunze
Asplenium riparium Liebm.
Asplenium rutaceum (Willd.) Mett.
Asplenium salicifolium L.
Asplenium serra Langsd. & Fisch.
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
AM BO
BO
AM BO
AM BO
AM BO
AM
AM
AM BO
DA
DA
GU
GU
GU
GU
GU?
GU
GU
GU
GU
GU
GU
su
su
su
su
su
FG
FG
FG
FG
FG
FG
FG
FG
DA
GU SU FG
GU SU FG
GU SU
su
FG
GU su FG
GU su FG
GU
GU su
GU FG
GU FG
GU
FG
FG
GU su FG
GU su FG
GU su
Checklist of the Plants of the Guiana Shield 25
Asplenium serratum L. AM BO DA GU SU FG
Asplenium sessilifoliiim Desv. FG?
Asplenium zamiifolium Willd. AM GU SU FG
Synonyms and Misidentifications
Asplenium alloeoptewn Kunze ex Klotzsch =Asplenium radicans
Asplenium amazonicum H. Christ =Asplenium pearcei
Asplenium anisophyllum auct. non Kunze =Asplenium feei
Asplenium arboreum Willd. (Woodsiaceae) ^Diplazium cristatum
Asplenium asplenioides (A.C. Sm.) Pittier et al. (Hymenophyllopsidaceae) =Hymenophyllopsis asplenioides
Asplenium auritum var. auriculatum (Hook, f.) C.V. Morton & Lellinger ^Asplenium auritum
Asplenium auritum var. bipinnatifidum Kunze =Asplenium auritum
Asplenium auritum var. obtusum Kunze ex Mett. =Asplenium macilentum
Asplenium blechnoides Lag. ex Sw. (Blechnaceae) =Blechnum polypodioides
Asplenium centripetale Baker (Woodsiaceae) =Diplazium centripetale
Asplenium cicutarium Sw. =Asplenium cristatum
Asplenium cirrhatum Rich, ex Willd. =Asplenium radicans van cirrhatum
Asplenium cyrtopteron Kunze =Asplenium radicans
Asplenium dejectum (Baker) Pittier (Hymenophyllopsidaceae) =Hymenophyllopsis dejecta
Asplenium dimidiatum auct. non Sw. =Asplenium zamiifolium
Asplenium fragrans Sw. =Asplenium cuspidatum
Asplenium grandifolium Sw. (Woodsiaceae) =Diplazium grandifolium
Asplenium haplophyllum Domin =Asplenium pearcei
Asplenium integerrimum Spreng. =Asplenium juglandifolium
Asplenium kappleriamim Kunze =Asplenium juglandifolium
Asplenium lechleri Mett. (Woodsiaceae) =Diplazium lechleri
Asplenium marginatum L. (Woodsiaceae) =Hemidictyum marginatum
Asplenium marinum Hook. f. =Asplenium auritum
Asplenium perkinsii Jenman =Asplenium rutaceum
Asplenium plantaginifolium L. (Woodsiaceae) =Diplazium plantaginifolium
Asplenium pseudoerectum Hieron. =Asplenium pteropus
Asplenium sanguinolentum Kunze ex Mett. =Asplenium feei
Asplenium schomburgkianum Klotzsch =Asplenium serratum
Asplenium semicordatum Raddi =Asplenium salicifolium
Asplenium spruceanum Hieron. =Asplenium hallii
Asplenium sulcatum auct. non Lam. =Asplenium auritum
Asplenium wilsonii Baker (Woodsiaceae) =Diplazium wilsonii
References
(Included in Pteridaceae in Flora of the Venezuelan Guayana)
Morton, C.V. & D.B. Lellinger. 1966. The Polypodiaceae subfam. Asplenioideae in Venezuela. Mem. New York Bot. Gard. 15:
1-49.
Smith, A.R. 1995. Aspleniaceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 12-22.
Blechnaceae
Blechnum asplenioides Sw. BO GU SU FG
Blechnum atropurpureum A.R. Sm. AM BO
Blechnum auratum (Fee) R.M. Tryon & Stolze AM
Blechnum binervatum (Poir.) C.V. Morton & Lellinger AM
Blechnum brasiliense Desv. BO GU
Blechnum x caudatum Cav. BO
Blechnum cordatum (Desv.) Hieron. AM BO
Blechnum divergens (Kunze) Mett. AM
Blechnum fraxineum Willd.
Blechnum gracile Kaulf
Blechnum lanceola Sw.
Blechnum lehmannii Hieron. AM BO
Blechnum Iherminieri (Bory) C. Chr.
Blechnum Iherminieri subsp. Iherminieri
Blechnum lineatum (Sw.) C. Chr.
Blechnum Occidentale L. AM BO
Blechnum polypodioides Raddi BO
Blechnum proliferum Rosenst. [Possibly misident. B. violaceum in the
Guianas]
GU SU FG
GU? FG
GU SU
SU
FG
GU
GU
GU
GU
GU SU FG
GU SU FG
GU?
26 Checklist of the Plants of the Guiana Shield
AM BO GU
AM BO
BO
DA GU
GU
su FG
AM GU
AM GU su FG
AM BO GU su FG
Blechnum schomburgkii (Klotzsch) C. Chr.
Blechnum serrulatum Rich.
Blechnum stipitellatum (Sodiro) C. Chr.
Blechnum violaceum (Fee) C. Chr.
Salpichlaena hookeriana (Kuntze) Alston
Salpichlaena volubilis (Kaulf.) J. Sm.
Synonyms and Misidentifications
Blechnum x antillanum Proctor =Blechnum x caudatum
Blechnum arborescens (Klotzsch & H. Karst.) Hieron =Blechnum cordatum
Blechnum blechnoides (Lag. ex Sw.) C. Chr. =Blechnum polypodioides
Blechnum buchtienii Rosenst. =Blechnum auratum
Blechnum ensiforme (Liebm.) C. Chr. =Blechnum binervatum
Blechnum exaltatum (Fee) Duek =Blechnum divergens
Blechnum indicum auct. non Burm. f. =Blechnum serrulatum
Blechnum maxonii (Broadh.) C. Chr. =Blechnum lehmannii
Blechnum ornifolium (C. Presl) Ettingsh. =Blechnum cordatum
Blechnum unilaterale Sw. [nom. superfl.] =Blechnum polypodioides
Blechnum volubile Kaulf. =Salpichlaena volubilis
Blechnum volubile van lomarioidea Baker =Salpichlaena hookeriana
Lomaria arborescens Klotzsch & H. Karst. =Blechnum cordatum
Lomaria aurata Fee =Blechnum auratum
Lomaria cordata Desv. =Blechnum cordatum
Lomaria divergens Kunze =Blechnum divergens
Lomaria ensiformis Liebm. =Blechnum binervatum
Lomaria exaltata Fee =Blechnum divergens
Lomaria ornifolia C. Presl =Blechnum cordatum
Lomaria schomburgkii Klotzsch =Blechnum schomburgkii
Lomaria stipitellata Sodiro =Blechnum stipitellatum
Lomaria violacea Fee =Blechnum violaceum
Salpichlaena lomarioidea (Baker) A.R. Sm. =Salpichlaena hookeriana
Spicanta hookeriana Kuntze =Salpichlaena hookeriana
Stenochlaenafendleri (D.C. Eaton) Underw. (Lomariopsidaceae) =Lomariopsis fendleri
Stenochlaenajapurensis (Mart.) L. Underw. (Lomariopsidaceae) =Lomariopsis japurensis
Stenochlaena marginata auct. non (Schrad.) C. Chr. (Lomariopsidaceae) =Lomariopsis japurensis
Struthiopteris maxonii Broadh. =Blechnum lehmannii
References
Smith, A.R. 1995. Blechnaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 23-29.
Cyatheaceae
Alsophila cuspidata (Kunze) D.S. Conant
Alsophila imrayana (Hook.) D.S. Conant
Alsophila imrayana (Hook.) D.S. Conant var. basilaris (H. Christ) Lellinger
Alsophila miquelii Kunze
Cnemidaria cruciata (Desv.) Stolze
Cnemidaria horrida (L.) C. Presl
Cnemidaria roraimensis (Domin) R.M. Tryon
Cnemidaria spectabilis (Kunze) R.M. Tryon
Cnemidaria spectabilis var. spectabilis
Cyathea akawaiorum P.J. Edwards
Cyathea andina (H. Karst.) Domin
Cyathea bipinnatifida (Baker) Domin
Cyathea caracasana (Klotzsch) Domin [Mt. Roraima]
Cyathea cyatheoides (Desv.) K.U. Kramer
Cyathea cyclodium (Tryon) Lellinger
Cyathea delgadii Stemb.
Cyathea demissa (C.V. Morton) A.R. Sm. ex Lellinger
Cyathea dissimilis (C.V. Morton) Stolze
Cyathea divergens Kunze
Cyathea divergens var. divergens
Cyatheafulva (M. Martens & Galeotti) Fee
Cyathea gibbosa (Klotzsch) Domin
Cyathea infesta (Kunze) Domin
Cyathea intramarginalis (P.G. Windisch) Lellinger
FG
AM BO
GU
SU
SU
SU?
FG
FG
FG
AM
AM BO GU
BO DA GU su FG
BO DA GU
GU
su FG
AM BO GU su FG
AM BO
BO
GU
GU?
AM BO DA GU su FG
AM
AM
BO
BO
BO
GU
GU
AM BO GU su
AM BO GU su
AM BO
GU
FG
BO
BO
BO
AM BO
AM
AM
AM BO
AM BO
BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
GU su FG
GU
GU su FG
GU su FG
GU
GU
Checklist of the Plants of the Guiana Shield 27
Cyathea kalbreyeri (Baker) Domin
Cyathea lasiosora (Kuhn) Domin FG
Cyathea lechleh Mett.
Cyathea liesneri A.R. Sm.
Cyathea macrocarpa (C. Presl) Domin
Cyathea macrosora (Baker) Domin
Cyathea maa-osora var. macrosora
Cyathea macrosora var. vaupensis (P.G. Windisch) A.R. Sm.
Cyathea marginalis (Klotzsch) Domin
Cyathea microdonta (Desv.) Domin DA
Cyathea multiflora Sm.
Cyathea nanna (Barrington) Lellinger
Cyathea neblinae A.R. Sm.
Cyathea platylepisiWook.) X:>om\n GU
Cyathea praeceps A.R. Sm.
Cyathea pubens Domin GU
Cyathea pimgens {W\M.) Domin DA GU SU FG
Cyathea schlimii (Kuhn) Domin
Cyathea simplex R.M. Tryon AM GU?
Cyathea sipapoensis (R.M. Tryon) Lellinger BO
Cyathea steyermarkii R.M. Tryon AM
Cyathea surinamensis (Miq.) Domin BO DA GU SU FG
Cyathea thysanolepis (Barrington) A.R. Sm. BO
Cyathea traillii (Baker) Dom.in AM GU
Cyathea ulei (H. Christ) Domin AM BO
Cyathea villosa Humb. & Bonpl. ex Willd. BO GU
Cyathea williamsii (Maxon) Domin AM
Synonyms and Misidentifications
Alsophila bipinnatifida Baker =Cyathea bipinnatifida
Alsophila blechnoides Hook. (Metaxyaceae) =Metaxya rostrata
Alsophila caracasana Klotzsch =Cyathea caracasana
Alsophila demissa C.V. Morton =Cyathea demissa
Alsophilafulva M. Martens & Galeotti =Cyatheafulva
Alsophila gibbosa Klotzsch =Cyathea gibbosa
Alsophila gleasonii Maxon =Cyathea pungens
Alsophila kalbreyeri Baker =Cyathea kalbreyeri
Alsophila lasiosora Mett. ex Kuhn =Cyathea lasiosora
Alsophila lechleri Mett. =Cyathea ulei
Alsophila macrosora Baker =Cyathea macrosora
Alsophila marginalis Klotzsch =Cyathea marginalis
Alsophila microdonta (Desv.) Desv. -Cyathea microdonta
Alsophila nigra Mart. =Cyafhea lasiosora
Alsophila oblonga Klotzsch =Cyathea pungens
Alsophila phalerata var. infesta Kunze =Cyathea infesta
Alsophila podophylla Baker =Cyathea kalbreyeri
Alsophila procera (Willd.) Desv. =Cyathea pungens
Alsophila pubescens Baker =Cyathea bipinnatifida
Alsophila pungens (Willd.) C. Presl =Cyathea pungens
Alsophila schlimii Kuhn =Cyathea schlimii
Alsophila subaculeata Splitg. =Cyathea pungens
Alsophila ulei H. Christ =Cyathea ulei
Alsophila vestita C. Presl =Cyathea spp.
Alsophila villosa (Humb. & Bonpl.) Desv. =Cyathea villosa
Alsophila williamsii Maxon =Cyathea williamsii
Cyathea arborea aucl. non (L.) Sm. =Alsophila imrayana
Cyathea aurea Klotzsch -Cyathea fulva
Cyathea boconensis H. Karst. =Cyathea caracasana
Cyathea calva H. Karst. —Cyathea divergens var. divergens
Cyathea caracasana var. meridensis (H. Karst.) R.M. Tryon =Cyathea caracasana
Cyathea cuspidata Kunze =Alsophila cuspidata
Cyathea horrida (L.) Sm. -Cnemidaria horrida
Cyathea imrayana Hook. =Alsophila imrayana
Cyathea macrosora var. reginae (P.G. Windisch) A.R. Sm. =Cyathea traillii
Cyathea meridensis H. Karst. =Cyathea caracasana
28 Checklist of the Plants of the Guiana Shield
Cyathea oblonga (Klotzsch) Domin =Cyathea piingens
Cyathea oyapoka Jenman =Alsophila cuspidata
Cyathea petiohilata H. Karst. =Cyathea divergens var. divergens
Cyathea procera (Willd.) Domin =Cyathea pungens
Cyathea purpurea C.V. Morton =Cyathea lechleri
Cyathea roraimemis (Domin) Domin =Cnemidaria roraimensis
Cyathea spectabilis (Kunze) Domin =Cnemidaria spectabilis
Cyathea squamipes H. Karst. =Cyathea caracasana
Cyathea subtropica Domin =Cyathea ulei
Cyathea surukunensis V. Marcano =Cyathea kalbreyeri
Cyathea vestita Mart. =Cyathea delgadii
Hemitelia andina H. Karst. =Cyathea andina
Hemitelia boryana Mett. ex Kuhn =Cyathea andina
Hemitelia cyatheoides Desv. =Cyathea cyatheoides
Hemitelia dissimilis C.V. Morton =Cyathea dissimilis
Hemitelia escuquensis H. Karst. =Cyathea andina
Hemitelia hirsuta (Desv.) Weath. =Cyathea surinamensis
Hemitelia horrida (L.) R. Br. =Cnemidaria horrida
Hemitelia hostmannii Hook. =Cyathea cyatheoides
Hemitelia macrocarpa C. Presl =Cyathea macrocarpa
Hemitelia macrosora (Baker) Jenman =Cyathea macrosora
Hemitelia minima C.V. Morton =Cyathea platylepis
Hemitelia multiflora (Desv.) Spreng. =Cyathea cyatheoides
Hemitelia multiflora (Desv.) Spreng. var. sprucei Baker =Cyathea cyatheoides
Hemitelia parkeri Hook. =Cyathea surinamensis
Hemitelia platylepis Hook. =Cyathea platylepis
Hemitelia roraimensis Domin =Cnemidaria roraimensis
Hemitelia spectabilis Kunze =Cnemidaria spectabilis
Hemitelia stigmosa Desv. =Cyathea cyatheoides
Hemitelia surinamensis Miq. =Cyathea surinamensis
Nephelea imrayana (Hook.) R.M. Tryon =Alsophila imrayana
Sphaeropteris cyatheoides (Desv.) RG. Windisch =Cyathea cyatheoides
Sphaeropteris dissimilis (C.V. Morton) RG. Windisch =Cyathea dissimilis
Sphaeropteris hirsuta (Desv.) R.M. Tryon =Cyathea surinamensis
Sphaeropteris intramarginalis RG Windisch =Cyathea intramarginalis
Sphaeropteris macrocarpa (C. Presl) R.M. Tryon =Cyathea macrocarpa
Sphaeropteris macrosora (Baker) RG. Windisch =Cyathea macrosora
Sphaeropteris macrosora var. reginae RG. Windisch =Cyathea traillii
Sphaeropteris macrosora var. vaupensis RG. Windisch =Cyathea macrosora var. vaupensis
Sphaeropteris marginalis (Klotzsch) R.M. Tryon =Cyathea marginalis
Sphaeropteris sipapoensis R.M. Tryon =Cyathea sipapoensis
Trichipteris cyclodium R.M. Tryon =Cyathea cyclodium
Trichipteris demissa (C.V. Morton) R.M. Tryon =Cyathea demissa
Trichipteris demissa var. thysanolepis Barrington =Cyathea thysanolepis
Trichipteris kalbreyeri (Baker) R.M. Tryon =Cyathea kalbreyeri
Trichipteris lechleri (Mett.) R.M. Tryon =Cyathea ulei
Trichipteris maguirei R.M. Tryon =Cyathea williamsii
Trichipteris microdonta (Desv.) R.M. Tryon =Cyathea microdonta
Trichipteris nigra (Mart.) R.M. Tryon =Cyathea lasiosora
Trichipteris procera (Willd.) R.M. Tryon =Cyathea pungens
Trichipteris pubescens (Baker) R.M. Tryon =Cyathea bipinnatifida
Trichipteris schlimii (Kuhn) Barrington =Cyathea schlimii
Trichipteris villosa (Humb. & Bonpl. ex Willd.) R.M. Tryon =Cyathea villosa
Trichipteris williamsii (Maxon) R.M. Tryon =Cyathea williamsii
References
Conant, D.C. 1983. A revision of the genus Alsophila in the Americas. Joum. Am. Arb. 64: 333-382.
Smith, A.R. 1995. Cyatheaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 30-43.
Stolze, R.G. 1974. A taxonomic revision of the genus Cnemidaria. Fieldiana Bot. 37: 1-98.
Tryon, R.M. 1970. The classification of the Cyatheaceae. Contrib. Gray Herb. 200: 3-53.
Tryon, R.M. 1976. A revision of the genus Cyathea. Contrib. Gray Herb. 206: 19-101.
Windisch, RG. 1977. Synopsis of the genus Sphaeropteris, with a revision of the neotropical exindusiate species. Bot. Jahrb.
Syst. 98(2): 176-198.
GU su FG
GU
GU
GU
GU su FG
GU
GU FG
GU su FG
GU
GU
GU?
Checklist of the Plants of the Guiana Shield 29
Davalliaceae
Rumohra adiantiformis (G. Forst.) Ching BO GU
Synonyms and Misidentifications
Davallia domingensis Spreng. (Dennstaedtiaceae) =Saccoloma domingense
Davallia inaequalis Kunze (Dennstaedtiaceae) =Saccoloma inaequale
Davallia meifolia Kunth (Dennstaedtiaceae) =Lindsaea meifolia
Davallia parkeri Hook. (Dennstaedtiaceae) =Lindsaea parkeri
References
Cremers, G, K.U. Kramer, R.C. Moran, & A.R. Smith. 1994. Dryopteridaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas
(series B) 6: 3-65.
Dennstaedtiaceae
Blotiella lindeniana (Hook.) R.M. Tryon AM
Dennstaedtia bipinnata (Cav.) Maxon
Dennstaedtia dissecta (Sw.) T. Moore AM
Dennstaedtia globidifera (Poir.) Hieron.
Histioptehs incisa (Thunb.) J. Sm. AM
Hypolepis guianensis Klotzsch [Mt. Roraima]
Hypolepis hostilis (Kunze) C. Presl
Hypolepis nigrescens Hook. [Mt. Roraima] AM
Hypolepis parallelogramma (Kunze) C. Presl
Hypolepis repens (L.) C. Presl
Hypolepis ^p. 1 [Mt. Roraima]
Hypolepis sp. 2 [Mt. Roraima]
Hypolepis sp. 3 [Mt. Roraima]
Hypolepis sp. A
Hypolepis viscosa Mett. [Cheilanthes]
Lindsaea arcuata Kunze
Lindsaea bolivarensis V. Marcano
Lindsaea coarctata K.U. Kramer
Lindsaea culthformis K.U. Kramer
Lindsaea cyclophylla K.U. Kramer
Lindsaea divahcata Klotzsch
Lindsaea diibia Spreng.
Lindsaeafalciformis Hook. [Mt. Roraima] [taxon of uncertain status]
Lindsaea guianensis (Aubl.) Dryand.
Lindsaea guianensis subsp. guianensis
Lindsaea hemiptera K.U. Kramer
Lindsaea javitensis Humb. & Bonpl. ex Willd. AM BO GU
Lindsaea klotzschiana Moritz ex Ettingsh.
Lindsaea lancea (L.) Bedd.
Lindsaea lancea var. elatior (Kunze) K.U. Kramer
Lindsaea lancea \ar.falcata (Dryand.) Rosenst.
Lindsaea lancea var. lancea
Lindsaea lancea var. leprieurii (Hook.) K.U. Kramer
Lindsaea lancea var. remota (Kunze) K.U. Kramer
Lindsaea lancea var. submontana Boudrie & Cremers
Lindsaea macrophylla Kaulf.
Lindsaea mazarunensis Jenman [species of uncertain status, probably a hybrid] GU
Lindsaea meifolia (Kunth) Mett. ex Kuhn AM
Lindsaea mesarum K.U. Kramer AM BO
Lindsaea pallida K\otzsch BO
Lindsaea parkeri {Hook.) Kuhn AM BO
Lindsaea parkeri subsp. parkeri BO
Lindsaea parkeri subsp. steyermarkiana K.\J. Kramer AM BO
Lindsaea pendula Klotzsch AM BO
Lindsaea pleioptera K.V.Kramer AM BO
Lindsaea portoricensis Desv. AM BO
Lindsaea pumila Klotzsch [dubious species]
Lindsaea quadrangularis Raddi
AM
AM BO
AM
BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM
AM BO
AM BO
AM BO
AM BO
AM BO
AM
AM BO
GU
GU
GU
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU SU FG
GU su FG
GU su FG
GU su FG
GU su FG
GU su FG
FG
FG
GU SU FG
GU SU FG
GU SU FG
GU SU
GU
GU SU FG
GU
GU SU FG
AM BO
BO
AM BO
AM BO
AM
AM BO
AM BO
AM BO
AM BO
AM BO
AM
BO
AM BO
AM BO
GU SU FG
GU SU FG
GU SU FG
GU
GU
GU
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU
GU
GU
SU
GU SU FG
GU FG
GU FG
GU SU FG
30 Checklist of the Plants of the Guiana Shield
Lindsaea quadrangularis subsp. antillensis K.U. Kramer
Lindsaea reniformis Dryand.
Lindsaea rigidhiscula Lindm.
Lindsaea sagittata (Aubl.) Dryand. DA
Lindsaea schomburgkii Klotzsch
Lindsaea semihmata (C. Chr.) C. Chr.
Lindsaea sphenomeridopsis K.U. Kramer
Lindsaea stenomeris K.U. Kramer
Lindsaea striata (Sw.) Dryand.
Lindsaea striata \a.r. jamesoniiformis K.U. Kramer
Lindsaea striata var. parvula (Fee) K.U. Kramer
Lindsaea striata var. striata
Lindsaea surinamensis Posth.
Lindsaea taeniata K.U. Kramer
Lindsaea tenuis Klotzsch
Lindsaea tetraptera K.U. Kramer
Lindsaea ulei Hieron.
Lonchitis hirsuta L. SU FG
Miarolepia speluncae (L.) T. Moore [introduced, fide Lellinger] FG
Odontosoria fiimarioides (Sw.) J. Sm. FG?
Ormoloma imrayanum (Hook.) Maxon [cited from FG in error] BO GU
Paesia aaalivis (Kunze) Kuhn BO
Paesia glandulosa (Sw.) Kuhn AM
Pteridium araahnoideum (Kaulf.) Maxon AM BO
Pteridium aaudatum (L.) Maxon AM
Sacaoloma domingense (Spreng.) C. Chr. BO
Saacoloma elegans Kaulf.
Saaaoloma elegans subsp. ahartaaeum Nair ex Cremers & K.U. Kramer
Saaaoloma inaequale (Kunze) Mett. AM BO
Synonyms and Misidentifications
Allosorus aaalivis Kunze =Paesia aaalivis
Dennstaedtia adiantoides (Willd.) T. Moore =Dennstaedtia disseata
Dennstaedtia obtusifolia (Willd.) T. Moore =Dennstaedtia disseata
Ithyaaulon inaequale (Kunze) Copel. =Saaaoloma inaequale
Lindsaea aquatiaa Jenman =Lindsaea striata var. parvula
Lindsaea aoriifolia Lindm. =Lindsaea sahomburgkii
Lindsaea arenata Klotzsch =Lindsaea portoriaensis
Lindsaea elegans Fee [nom. illeg.] =Lindsaea klotzsahiana
Lindsaeafalaata Dryand. =Lindsaea lanaea var.falaata
Lindsaeafalaata var. elatior Kunze =Lindsaea lanaea var. elatior
Lindsaeafalaata var. lanaea (L.) Jenman =Lindsaea lanaea var. lanaea
Lindsaeafalaata var. leprieurii (Hook.) Jenman =Lindsaea lanaea var. leprieurii
Lindsaeafalaata var. subrotundifolia Jenman [name of uncertain application]
Lindsaea feei C. Chr. =Lindsaea klotzsahiana
Lindsaea filiformis Hook. =Lindsaea tenuis
Lindsaea gracilis Klotzsch =Lindsaea striata var. parvula
Lindsaea guianensis var. venosa Jenman [name of uncertain application]
Lindsaea horizontalis Hook. =Lindsaea arauata
Lindsaea krameriana V. Marcano =Lindsaea mesarum
Lindsaea lanaea var. longifolia K.U. Kramer =Lindsaea lanaea var. elatior
Lindsaea lanaea var. semilunata C. Chr. =Lindsaea semilunata
Lindsaea leprieurii Hook. =Lindsaea lanaea var. leprieurii
Lindsaea nervosa Lindm. [nom. illeg.] =Lindsaea rigidiusaula
Lindsaea nitidissima Rich, ex Willd. =Lindsaea lanaea
Lindsaea parvula Fee =Lindsaea striata var. parvula
Lindsaea pumila var. remota Kunze =Lindsaea lanaea var. remota
Lindsaea pusilla Splitg. =Lindsaea lanaea var. remota
Lindsaea raddiana Klotzsch =Lindsaea striata
Lindsaea rigesaens Willd. =Lindsaea striata
Lindsaea rufesaens Kunze ^Lindsaea portoriaensis
Lindsaea sahomburgkii f aoriifolia (Lindm.) K.U. Kramer =Lindsaea sahomburgkii
Lindsaea sahomburgkii f. densa K.U. Kramer =Lindsaea sahomburgkii
Lindsaea trapeziformis Dryand. =Lindsaea lanaea
Lindsaea trapeziformis var. erythromera Kunze =Lindsaea divariaata
Checklist of the Plants of the Guiana Shield 31
Lonchitis lindeniana Hook. =Blotiella lindeniana
Lonchitis pedata L. (Pteridaceae) =Pteris podophylla
Odontoloma parkeri (Hook.) C. Presl =Lindsaea parkeri
Orthiopteris domingensis (Spreng.) Copel. =Saccoloma domingense
Orthiopteris inaequalis (Kunze) Copel. =Saccoloma inaequale
Paesia viscosa A. St.-Hil. =Paesia glandulosa
Pteridium aquilinum var. arachnoideum (Kaulf.) Brade =Pteridium arachnoideum
Pteridium aquilinum subsp. caudatum (L.) Bonap. =Pteridium caudatum
Saccoloma imrayana Hook. =Ormoloma imrayanum
Saccoloma imrayanum Hook. =Ormoloma imrayanum
Schizoloma macrophyllum (Kaulf.) C. Presl =Lindsaea macrophylla
Schizoloma reniforme (Dryand.) Diels =Lindsaea reniformis
Schizoloma sagittatum (Aubl.) Diels =Lindsaea sagittata
Stenoloma gratissima Fee =Lindsaea meifolia
References
Cremers, G. & K.U. Kramer. 1991. Dennstaedtiaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series B) 4: 20-81.
Kramer, K.U. 1957. A revision of the genus Lindsaea in the New World. Acta Bot. Need. 6: 97-290.
Smith, A.R. & K.U. Kramer. 1995. Dennstaedtiaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan
Guayana 2: 46-71.
DiCKSONIACEAE
Culcita coniifolia (Hook.) Maxon AM BO GU
Synonyms and Misidentifications
Dicksonia adiantoides Humb. & Bonpl. ex Willd. (Dennstaedtiaceae) =Dennstaedtia bipinnata
Dicksonia bipinnata Cav. (Dennstaedtiaceae) =Dennstaedtia bipinnata
Dicksonia coniifolia Hook. =Culcita coniifolia
Dicksonia dissecta Sw. (Dennstaedtiaceae) =Dennstaedtia dissecta
Dicksonia obtusifolia Willd. (Dennstaedtiaceae) =Dennstaedtia dissecta
References
Maxon, W.R. 1922. The genus Culcita. J. Wash. Acad. Sci. 12: 454-460.
Smith, A.R. 1995. Dicksoniaceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 72-73.
Dryopteridaceae
Arachniodes denticulata (Sw.) Ching [Mt. Roraima]
Arachniodes leucostegioides (C. Chr.) Ching [Mt. Roraima]
Arachniodes macrostegia (Hook.) R.M. Tryon & D.S. Conant [Mt. Roraima]
Arachniodes ochropteroides (Baker) Lellinger
Cyclodium akawaiorum A.R. Sm. [Mt. Roraima]
Cyclodium calophyllum (C.V. Morton) A.R. Sm.
Cyclodium guianense (Klotzsch) van der Werff ex L.D. Gomez
Cyclodium inerme (Fee) A.R. Sm.
Cyclodium meniscioides (Willd.) C. Presl
Cyclodium meniscioides var. meniscioides
Cyclodium meniscioides var. rigidissimum (C. Chr.) A.R. Sm.
Cyclodium rheophilum A.R. Sm.
Cyclodium varians (Fee) A.R. Sm.
Didymochlaena truncatula (Sw.) J. Sm.
Olfersia cervina (L.) Kunze
Polybotrya caudata Kunze
Polybotrya cyathifolia Fee
Polybotrya fractiserialis (Baker) Sm.
Polybotrya glandulosa Mett. ex Kuhn
Polybotrya lechleriana Mett. [Mt. Roraima]
Polybotrya osmundacea Humb. & Bonpl. ex Willd.
Polybotrya sessilisora R.C. Moran [Bra/il/Guyana border area]
Polybotrya sorbifolia Mett. ex Kuhn [expected in the Guianas]
Polystichum platylepis Fee GU?
Polystichum platyphyllum (Willd.) C. Presl AM
Polystichum sp. A BO
Stigmatopteris longicaudata (Liebm.) C. Chr. AM FG
AM BO GU
AM GU
AM BO GU?
AM BO
BO
BO
GU
GU
GU?
SU
AM BO DA GU SU FG
AM BO GU SU FG
AM BO? DA GU SU FG
AM BO
BO?
BO
DA GU
GU
GU
SU FG
FG
AM BO DA GU SU FG
AM BO GU SU FG
AM BO DA GU SU FG
FG
GU SU FG
AM
GU
AM
BO
GU
GU?
SU FG
AM BO
32 Checklist of the Plants of the Guiana Shield
Stigmatopteris rotundata (Humb. & Bonpl. ex Willd.) C. Chr. BO GU SU FG
Synonyms and Misidentifications
Aspidium articulatum Sw. (Oleandraceae) =Oleandra articulata
Aspidium biolleyi H. Christ (Thelypteridaceae) =Thelyptens biolleyi
Aspidium biserratum Sw. (Oleandraceae) =Nephrolepis biserrata
Aspidium capense Willd. (Davalliaceae) =Rumohra adiantiformis
Aspidium coarctatum Kunze (Thelypteridaceae) =Thelypteris opposita
Aspidium confertum Kaulf. =Cyclodium meniscioides var. meniscioides
Aspidium crenulans Fee (Tectariaceae) =Megalastrum crenulans
Aspidium dicksonioides Fee (Tectariaceae) =Triplophyllum dicksonioides
Aspidium extensum Blume (Thelypteridaceae) =Thelypteris opulenta
Aspidium funestum Kunze (Tectariaceae) =Triplophyllum funestum
Aspidium gongylodes Schkuhr (Thelypteridaceae) =Thelypteris interrupta
Aspidium guianense Klotzsch =Cyclodium guianense
Aspidium hispidulum Decne. (Thelypteridaceae) =Thelypteris hispidula
Aspidium lizarzaburui (Sodiro) C. Chr. (Tectariaceae) =Tectaria lizarzaburui
Aspidium martinicense Spreng. (Tectariaceae) =Tectaria incisa
Aspidium meniscioides Willd. =Cyclodium meniscioides
Aspidium nodosum Willd. (Oleandraceae) =Oleandra articulata
Aspidium opulentum Kaulf. (Thelypteridaceae) =Thelypteris opulenta
Aspidium pectinatum Willd. (Oleandraceae) =Nephrolepis pectinata
Aspidium pendulum Raddi (Oleandraceae) =Nephrolepis cordifolia
Aspidium pendulum Splitg., non Raddi 1819 (Oleandraceae) =Oleandra pilosa
Aspidium plantagineum (Jacq.) Griseb. (Tectariaceae) =Tectaria plantaginea
Aspidium platyphyllum Willd. =Polystichum platyphyllum
Aspidium plumieri C. Presl (Tectariaceae) =Tectaria trifoliata
Aspidium rostratum Kunth (Metaxyaceae) =Metaxya rostrata
Aspidium rotundatum Humb. & Bonpl. ex Willd. =Stigmatoptehs rotundata
Aspidium rutaceum Willd. (Aspleniaceae) =Asplenium rutaceum
Aspidium schomburgkii Klotzsch (Tectariaceae) =Ctenitis falciculata
Aspidium sprengelii Kaulf. (Thelypteridaceae) =Thelypteris balbisii
Aspidium truncatulum Sw. =Didymochlaena truncatula
Cyclodium confertum (Kaulf.) C. Presl =Cyclodium meniscioides var. meniscioides
Cyclodium rigidissimum C. Chr. =Cyclodium meniscioides var. rigidissimum
Dryopteris amplissima (C. Presl) Kuntze (Tectariaceae) =Lastreopsis amplissima
Dryopteris amplissima var. subeffusa C. Chr. (Tectariaceae) =Lastreopsis amplissima
Dryopteris angustifolia (Willd.) Urb. (Thelypteridaceae) =Thelypteris angustifolia
Dryopteris arborea Brause (Thelypteridaceae) =Thelypteris arborea
Dryopteris balbisii (Spreng.) Urb. (Thelypteridaceae) =Thelypteris balbisii
Dryopteris brachypoda (Baker) C. Chr. (Thelypteridaceae) =Thelypteris brachypoda
Dryopteris calophylla C.V. Morton =Cyclodium calophyllum
Dryopteris carrii auct. non (Baker) C. Chr. =Stigmatopteris rotundata
Dryopteris chrysodioides (Fee) Maxon & C.V. Morton (Thelypteridaceae) =Thelypteris chrysodioides
Dryopteris chrysodioides var. goyazensis Maxon & C.V. Morton (Thelypteridaceae) =Thelypteris chrysodioides var.
goyazensis
Dryopteris crenulans (Fee) C. Chr. (Tectariaceae) =Megalastrum crenulans
Dryopteris decussata (L.) Urb. (Thelypteridaceae) =Thelypteris decussata
Dryopteris dejecta (Jenman) C. Chr. (Thelypteridaceae) =Thelypteris glandulosa
Dryopteris demerarana (Baker) C. Chr. (Thelypteridaceae) =Thelypteris demerarana
Dryopteris densiloba C. Chr. [nom. illeg.] (Thelypteridaceae) =Thelypteris gardneriana
Dryopteris dentata auct. non (Forssk.) C. Chr. (Thelypteridaceae) =Thelypteris hispidula
Dryopteris denticulata (Sw.) Kuntze =Arachniodes denticulata
Dryopteris denticulata var. aperta C. Chr. =Arachniodes denticulata
Dryopteris effusa (Sw.) Urb. (Tectariaceae) =Lastreopsis effusa
Dryopteris engelii Hieron. (Thelypteridaceae) =Thelypteris rudis
Dryopteris extensa (Blume) Kuntze (Thelypteridaceae) =Thelypteris opulenta
Dryopterisfunesta (Kunze) C. Chr. (Tectariaceae) =Triplophyllum funestum
Dryopteris gongylodes (Schkuhr) Kuntze (Thelypteridaceae) =Thelypteris interrupta
Dryopteris gonophora Weath. (Thelypteridaceae) =Thelypteris gonophora
Dryopteris guianensis (Klotzsch) Posth. =Cyclodium guianense
Dryopteris hostmannii Klotzsch (Thelypteridaceae) =Thelypteris arborescens
Dryopteris kanukuana Weath. (Thelypteridaceae) =Thelypteris pennata
Dryopteris killipii C. Chr. & Maxon (Tectariaceae) =Lastreopsis killipii
Dryopteris latiuscula Maxon =Thelypteris abrupta
Checklist of the Plants of the Guiana Shield 33
Dryopteris leprieurii (Hook.) Kuntze (Thelypteridaceae) =Thelypteris leprieurii
Dryopteris leucostegioides C. Chr. =Arachniodes leucostegioides
Dryopteris lindigii C. Chr. (Thelypteridaceae) =Thelypteris deflexa
Dryopteris linkiana (C. Presl) Maxon (Thelypteridaceae) =Thelypteris linkiana
Dryopteris longicaudata (Liebm.) Maxon =Stigmatopteris longicaudata
Dryopteris macrophylla (Kunze) C. Chr. (Thelypteridaceae) ^Thelypteris macrophylla
Dryopteris macrostegia (Hook.) Kuntze =Arachniodes macrostegia
Dryopteris tnartinii C. Chr. (Thelypteridaceae) [not in flora area, type apparently erroneously cited from FG] =Thelypteris
connexa
Dryopteris megalodus (Schkuhr) Urb. (Thelypteridaceae) =Thelypteris pennata
Dryopteris meniscioides (Willd.) Kuntze =Cyclodium meniscioides
Dryopteris meniscioides var. conferta (Kaulf.) C.V. Morton =Cyclodium meniscioides var. meniscioides
Dryopteris mercurii A. Braun ex Hieron. (Thelypteridaceae) =Thelypteris balbisii
Dryopteris nephrodioides var. bioUeyi (H. Christ) C. Chr. (Thelypteridaceae) =Thelypteris biolleyi
Dryopteris nesiotica Maxon & C.V. Morton (Thelypteridaceae) =Thelypteris nesiotica
Dryopteris ochropteroides (Baker) C. Chr. =Arachniodes ochropteroides
Dryopteris oligocarpa (Humb. & Bonpl. ex Willd.) Kuntze (Thelypteridaceae) =Thelypteris oligocarpa
Dryopteris opposita (Vahl) Urb. (Thelypteridaceae) =Thelypteris opposita
Dryopteris poiteana (Bory) Urb. (Thelypteridaceae) =Thelypteris poiteana
Dryopteris protensa var. dicksonioides (Fee) C. Chr. (Tectariaceae) =Triplophyllum dicksonioides
Dryopteris protensa \av.funesta (Kunze) C. Chr. (Tectariaceae) =TriplophyUumfimestum
Dryopteris pteroidea (Klotzsch) C. Chr. (Thelypteridaceae) =Thelypteris pteroidea
Dryopteris pyramidata (Fee) Maxon (Thelypteridaceae) =Thelypteris abrupta
Dryopteris quadrangularis (Fee) Alston (Thelypteridaceae) =Thelypteris hispidula
Dryopteris refulgens (Klotzsch ex Mett.) C. Chr. (Tectariaceae) =Ctenitis refulgens
Dryopteris rolandii C. Chr. (Thelypteridaceae) =Thelypteris x rolandii
Dryopteris roraimensis Brause (Thelypteridaceae) =Thelypteris arborea
Dryopteris rotundata (Humb. & Bonpl. ex Willd.) C. Chr. =Stigmatopteris rotundata
Dryopteris rudis (Kunze) C. Chr. (Thelypteridaceae) =Thelypteris rudis
Dryopteris sancti-gabrieli (Hook.) Kuntze =Cyclodium guianense
Dryopteris serrata (Cav.) C. Chr. (Thelypteridaceae) =Thelypteris serrata
Dryopteris sorbifolia auct. non (Jacq.) Hieron. (Thelypteridaceae) =Thelypteris arborescens
Dryopteris subcuneata (Baker) Kuntze (Thelypteridaceae) =Thelypteris abrupta
Dryopteris subincisa (Willd.) Urb. (Tectariaceae) =Megalastrum subincisum
Dryopteris subobliquata (Hook.) Kuntze =Cyclodiiim inerme
Dryopteris tetragona (Sw.) Urb. (Thelypteridaceae) =Thelypteris tetragona
Dryopteris tristis (Kunze) Kuntze (Thelypteridaceae) =Thelypteris tristis
Dryopteris varians (Fee) Kuntze =Cyclodium varians
Nephrodium antioqidamtm Baker (Tectariaceae) =Tectaria antioquioana
Nephrodium aureoviridum Jenman [dubious species; =Thelypteris tetragona, Thelypteridaceae?]
Nephrodium brachypodum Baker (Thelypteridaceae) =Thelypteris brachypoda
Nephrodium connexum Kuhn ex Baker (Thelypteridaceae) [not in flora area, type erroneously cited from FG] =Thelypteris
connexa
Nephrodium deflexum C. Presl (Thelypteridaceae) =Thelypteris deflexa
Nephrodium dejectum Jenman (Thelypteridaceae) =Thelypteris glandidosa
Nephrodium gardnerianum Baker (Thelypteridaceae) =Thelypteris gardneriana
Nephrodium leprieurii Hook. (Thelypteridaceae) =Thelypteris leprieurii
Nephrodium lizarzaburui Sodiro (Tectariaceae) =Tectaria lizarzaburui
Nephrodium macrostegium Hook. =Arachniodes macrostegia
Nephrodium ochropteroides Baker =Arachniodes ochropteroides
Nephrodium quadrangulare Fee (Thelypteridaceae) ^^Thelypteris hispidula
Nephrodium subcuneatum Baker (Thelypteridaceae) =Thelypteris abrupta
Nephrodium subfuscum Baker (Thelypteridaceae) =Thelypteris germaniana
Nephrodium varians Fee =Cyclodium varians
Polybotrya aristeguietae Brade =Polybotrya osmundacea
Polybotrya cerxnna (L.) Kaulf. =Olfersia cervina
Polybotrya pubens auct. non Mart. =Polyhotrya caudata
Polybotrya salicifolia Lellinger ^Polybotrya sorbifolia
Polybotrya suhelliptica Lellinger =Polybotrya glandulosa
Polybotrya vareschii Brade ^Polybotrya osmundacea
Polybotrya villosula H. Christ =Polybotrya caudata
Polystichopsis ochropteroides (Baker) C.V. Morton =Arachniodes ochr-opteroides
Polystichum adiantiforme (G. Forst.) J. Sm. (Davalliaceae) =Rumohr-a adiantiformis
Polystichum amplissimum C. Presl (Tectariaceae) =Lastreopsis amplissima
Polystichum capense (Willd.) J. Sm. (Davalliaceae) =Rumohr-a adiantiformis
34 Checklist of the Plants of the Guiana Shield
Polystichum inerme Fee =Cyclodmm inerme
Stigmatopteris guianensis (Klotzsch) C. Chr. =Cyclodium guianense
Stigmatopteris meniscioides (Willd.) K.U. Kramer =Cyclodium meniscioides
Stigmatopteris sancti-gabrieli (Hook.) C. Chr. =Cyclodium guianense
Stigmatopteris varians (Fee) Alston =Cyclodiiim varians
References
Cremers, G, K.U. Kramer, R.C. Moran, & A.R. Smith. 1994. Dryopteridaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas
(series B) 6: 3-65.
Moran, R.C. 1986. The Neotropical fern genus Olfersia. Amer. Fern J. 76: 161-178.
Moran, R.C. 1987. Monograph of the Neotropical fern genus Polybotrya (Dryopteridaceae). III. Nat. Hist. Surv. Bull. 34(1): 1-
138.
Moran, R.C. 1991 . Monograph of the Neotropical fern genus Stigmatopteris (Dryopteridaceae). Ann. Missouri Bot. Gard. 78(4):
857-914.
Smith, A.R. 1986. Revision of the Neotropical fern genus Cyclodium. Am. Fern J. 76(2): 56-98.
Smith, A.R., J.T. Mickel, & R.C. Moran. 1995. Dryopteridaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the
Venezuelan Guayana 2: 73-128.
Gleicheniaceae
Dicranopteris flexiiosa (Schrad.) Underw. AM BO GU SU FG
Dicranopteris nervosa (Kaulf.) Maxon
Dicranopteris pectinata(Wi\\d.) Underw. AM BO GU SU FG
Dicranopteris schombitrgkiana (J.W. Sturm) C.V. Morton [Mt. Roraima]
Diplopterygiiim bancroftii (Hook.) A.R. Sm.
Sticherus bifidus {W\M.) Ching AM BO GU SU FG
Sticherus compactus (H. Christ) Nakai
Sticheriis hypoleiicus (Sodiro) Copel.
Sticherus longipinnatus (Hook.) Ching GU SU
Sticherus melanoblastus (Alston) 0sterg. & B. 011g.
Sticherus nervatus J. Gonzales
Sticherus pallescens (Mett.) Vareschi
Sticherus penniger (Mart.) Copel.
Sticherus remotus {¥^2i\x\f.) C\ixys\Qx AM BO GU SU FG
Sticherus revolutus (Kunth) Ching [Mt. Roraima]
Sticherus sp. A
Sticherus tepuiensis A.R. Sm.
Sticherus tomentosus (Cav. ex Sw.) A.R. Sm.
Synonyms and Misidentifications
Dicranopteris bifida (Willd.) Maxon =Sticherus bifidus
Dicranopteris nitidula (Rosenst.) Suess. ex Losch =Sticherus nitidulus
Dicranopteris remota (Kaulf.) Maxon ^Sticherus remotus
Gleichenia bancroftii Hook. =Diplopterygium bancroftii
Gleichenia bifida (Willd.) Spreng. =Sticherus bifidus
Gleichenia hypoleuca Sodiro =Sticherus hypoleucus
Gleichenia linearis auct. non (Burm.) C.B. Clarke =Dicranopteris flexuosa
Gleichenia longipinnata Hook. =Sticherus longipinnatus
Gleichenia melanoblasta Alston
Gleichenia nitidula Rosenst. =Sticherus nitidulus
Gleichenia pallescens Mett. =Sticherus pallescens
Gleichenia pectinata (Willd.) C. Presl ^Dicranopteris pectinata
Gleichenia pennigera (Mart.) T. Moore =Sticherus penniger
Gleichenia remota (Kaulf) Spreng. =Sticherus remotus
Gleichenia revoluta Kunth =Sticherus revolutus
Gleichenia tomentosa (Cav. ex Sw.) Spreng. =Sticherus tomentosus
Hicriopteris bancroftii (Hook.) Ching =Diplopterygium bancroftii
Mertensia bifida Willd. ^Sticherus bifidus
Mertensiaflexuosa Schrad. =Dicranopteris flexuosa
Mertensia pectinata Willd. =Dicranopteris pectinata
Mertensia pennigera Mart. ^Sticherus penniger
Mertensia pubescens Humb. & Bonpl. ex Willd. =Sticherus bifidus
Mertensia remota Kaulf. =Sticherus remotus
Mertensia schombitrgkiana J.W. Sturm =Dicranopteris schomburgkiana
Mertensia tomentosa Cav. ex Sw. ^Sticherus tomentosus
GU
AM BO
AM BO
GU
GU
GU
GU
AM
BO
AM BO
BO
GU
AM BO GU
AM BO GU
Checklist of the Plants of the Guiana Shield 35
Mertensia velata Kunze =Sticherus tomentosus
Sticherus nitidiilus (Rosenst.) Copel. [not in flora area]
Sticherus piibescens (Humb. & Bonpl. ex Willd.) Nakai =Sticherus bifidus
Sticherus velatus (Kunze) Copel. =Sticherus tomentosus
References
Smith, A.R. 1995. Gleicheniaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 128-135.
Grammitidaceae
Ceradenia arthrothrix L.E. Bishop & A.R. Sm.
Ceradenia capillaris (Desv.) L.E. Bishop [Mt. Roraima] AM BO GU?
Ceradenia discolor (Hook.) L.E. Bishop [Mt. Roraima] AM BO GU
Ceradeniafragillima (Copel.) L.E. Bishop [Mt. Roraima]
Ceradeniajungermannioides (Klotzsch) L.E. Bishop
Ceradenia kalbreyeri (Baker) L.E. Bishop AM BO GU SU EG
Ceradenia kookenamae (Jenman) L.E. Bishop [Mt. Roraima] AM BO GU
Ceradenia microcystis L.E. Bishop & A.R. Sm.
Ceradenia pruinosa (Maxon) L.E. Bishop
Ceradenia spixiana (Mart, ex Mett.) L.E. Bishop
Cochlidium attenuatum A.C. Sm.
Cochlidium connellii (Baker ex C.H. Wright) A.C. Sm.
Cochlidium furcatum (Hook. & Grev.) C. Chr.
Cochlidium linearifolium (Desv.) Maxon ex C. Chr.
Cochlidium rostratum (Hook.) Maxon ex C. Chr.
Cochlidium seminudum (Willd.) Maxon
Cochlidium serrulatum (Sw.) L.E. Bishop
Cochlidium tepuiense (A.C. Sm.) L.E. Bishop
Cochlidium wurdackii L.E. Bishop
Enterosora campbellii Baker
Enterosora campbellii subsp. campbellii
Enterosora parietina (Klotzsch) L.E. Bishop
Enterosora trifurcata (L.) L.E. Bishop
Grammitis anfractuosa (Kunze ex Klotzsch) Proctor [=Melpomene
anfractuosa]
Grammitis apiculata (Kunze ex Klotzsch) F. Seym.
Grammitis asplenifolia (L.) Proctor [=Terpsichore asplenifolia]
Grammitis blanchetii (C. Chr.) A.R. Sm. [=Micropolypodium nanum]
Grammitis bryophila (Maxon) F. Seym.
Grammitis caucana (Hieron.) C.V. Morton
Grammitis cultrata (Bory ex Willd.) Proctor [=Terpsichore cultrata]
Grammitis firma (J. Sm.) C.V. Morton [=Melpomenefirma]
Grammitis flabelliformis (Poir.) C.V. Morton [Mt. Roraima] [=Melpomene
flabelliformis]
Grammitis kegeliana (Kunze) Lellinger [^Terpsichore kegeliana] SU FG
Grammitis leptopoda (C.H. Wright) Copel. [Mt. Roraima] BO GU
Grammitis liesneri A.R. Sm.
Grammitis limula (H. Christ) L.D. Gomez
Grammitis major (Copel.) C.V. Morton
Grammitis melanosticta (Kunze) F. Seym. [=Melpomene melanosticta] AM BO GU SU
Grammitis melanotrichia (Baker) Lellinger [Mt. Roraima] [=Lellingeria
melanotrichia]
Grammitis mollissima (Fee) Proctor [=Terpsichore mollissima] AM BO GU SU FG
Grammitis moniliformis (Lag. ex Sw.) Proctor [=Melpomene moniliformis] AM BO GU
Grammitis myosuroides (Sw.) Sw. [Mt. Roraima] [=Lellingeria myosuroides]
Grammitis paramicola L.E. Bishop [Mt. Roraima]
Grammitis peritimundi L.E. Bishop & A.R. Sm. [Mt. Roraima]
Grammitis phlegmaria (J. Sm.) Proctor
Grammitis phlegmaria var. phlegmaria
Grammitis pilosissima (M. Martens & Galeotti) C.V. Morton [=Melpomene SU
pilosissima]
Grammitis plicata A.R. Sm. AM BO
Grammitis randallii (Maxon) Proctor GU
Grammitis shaferi (Maxon) Lellinger [=Lellingeria shaferi] SU
AM
BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
AM BO
AM
BO
AM BO
AM
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM
AM
AM
BO
AM BO
BO
AM BO
AM BO
AM BO
GU
GU
GU SU FG
GU SU FG
GU?
GU? FG?
GU SU FG
GU
GU FG?
GU FG?
GU SU
GU
GU
GU SU FG
GU
GU
GU
GU?
AM BO
BO
BO
AM BO
BO
AM BO
AM BO
BO
AM BO
AM BO
AM BO
AM BO
AM BO
GU
GU SU FG
GU SU FG
GU SU FG
GU
GU FG
36 Checklist of the Plants of the Guiana Shield
Grammitis sinuosa A.R. Sm. [=Lellingeria sinuosa] GU SU
Grammitis sp. A
Grammitis staheliana (Posth.) Lellinger [=Terpsichore staheliana] SU FG
Grammitis subsessilis (Baker) C.V. Morton [=LeUingeria subsessilis]
Grammitis suspensa (L.) Proctor [=Lellingeria suspensa]
Grammitis taenifolia (Jenman) Proctor [=MicropoIypodium taenifolium]
Grammitis taxifolia (L.) Proctor [=Terpsichore taxifolia]
Grammitis tegetiformis L.E. Bishop
Grammitis tenuicula (Fee) Proctor
Grammitis trichomanoides (Sw.) Ching {=Micropolypodium trichomanoides]
Grammitis truncicola (Klotzsch) C.V. Morton
Grammitis xanthotrichia (Klotzsch) Duek & Lellinger [Mt. Roraima]
[=Terpsichore xanthotrichia']
Grammitis xiphopteroides (Liebm.) A.R. Sm.
Zygophlebia sectifrons (Kunze ex Mett.) L.E. Bishop
Synonyms and Misidentifications
Ctenopteris anfractuosa (Kunze ex Klotzsch) Copel. =Grammitis anfractuosa
Ctenopteris apiculata (Kunze ex Klotzsch) Copel. =Grammitis apiculata
Ctenopteris asplenifolia (L.) Copel. =Grammitis asplenifolia
Ctenopteris capillaris (Desv.) Copel. =Ceradenia capillaris
Ctenopteris cultrata (Bory ex Willd.) Copel. =Grammitis cultrata
Ctenopterisfirma J. Sm. =Grammitis firma
Ctenopterisfragillima Copel. =Ceradeniafragillima
Ctenopterisjubiformis (Kaulf.) J. Sm. =Grammitis suspensa
Ctenopteris kaieteura (Jenman) Copel. =Grammitis tenuicula
Ctenopteris kalbreyeri (Baker) Copel. =Ceradenia kalbreyeri
Ctenopteris kegeliana (Kunze) K.U. Kramer =Grammitis kegeliana
Ctenopteris kookenamae (Jenman) Copel. =Ceradenia kookenamae
Ctenopteris major Copel. =Grammitis major
Ctenopteris melanosticta (Kunze) Copel. =Grammitis melanosticta
Ctenopteris melanotrichia (Baker) Copel. =Grammitis melanotrichia
Ctenopteris mollissima (Fee) Copel. ^Grammitis mollissima
Ctenopteris moniliformis (Lag. ex Sw.) J. Sm. =Grammitis moniliformis
Ctenopteris phlegmaria (J. Sm.) Copel. =Grammitis phlegmaria
Ctenopteris pilosissima (M. Martens & Galeotti) Copel. =Grammitis pilosissima
Ctenopteris pruinosa (Maxon) Copel. =Ceradenia pruinosa
Ctenopteris rigens (Maxon) Copel. =Grammitis xiphopteroides
Ctenopteris rigescens (Bory ex Willd.) J. Sm. =Grammitis flabelliformis
Ctenopteris staheliana (Posth.) K.U. Kramer =Grammitis staheliana
Ctenopteris subsessilis (Baker) Copel. =Grammitis subsessilis
Ctenopteris suspensa (L.) Copel. =Grammitis suspensa
Ctenopteris taxifolia (L.) Copel. =Grammitis taxifolia
Ctenopteris tenuicula (Fee) Copel. =Grammitis tenuicula
Ctenopteris trifurcata (L.) Pic. Serm. =Enterosora trifurcata
Ctenopteris xanthotrichia (Klotzsch) Copel. ^Grammitis xanthotrichia
Enterosora Integra Lellinger & C.V. Morton =Enterosora campbellii subsp. campbellii
Glyphotaenium campbellii (Baker) Copel. =Enterosora campbellii subsp. campbellii
Glyphotaenium integrum (Lellinger & C.V. Morton) Lellinger =Enterosora campbellii subsp. campbellii
Glyphotaenium trifurcatum (L.) Lellinger =Enterosora trifurcata
Grammitis campbellii (Baker) Proctor =Enterosora campbellii
Grammitis capillaris (Desv.) Proctor =Ceradenia capillaris
Grammitis dicranophylla (C. Chr.) Vareschi =Cochlidium furcatum
Grammitis discolor (Hook.) C.V. Morton =Ceradenia discolor
Grammitis flexuosa Kunth (Adiantaceae) =Eriosorus flexuosus
Grammitis fragillima (Copel.) C.V. Morton =Ceradeniafragillima
Grammitis furcata Hook. & Grev. =Cochlidium furcatum
Grammitis haplophlebica (A.C. Sm.) Vareschi =Grammitis bryophila
Grammitis Jamesonii (Hook.) C.V. Morton =Grammitis myosuroides
Grammitisjubiformis (Kaulf.) Proctor =Grammitis suspensa
Grammitis jungermannioides (Klotzsch) Ching =Ceradeniajungermannioides
Grammitis kaieteura (Jenman) C.V. Morton =Grammitis tenuicula
Grammitis kalbreyeri (Baker) C.V. Morton =Ceradenia kalbreyeri
Grammitis kirkbridei Mickel =Ceradenia pruinosa
Grammitis leptodon (C.H. Wright) Vareschi [incorrect spelling, combination in Grammitis previously made] =Grammitis
Checklist of the Plants of the Guiana Shield 37
leptopoda
Grammitis linearifolia (Desv.) Steud. =Cochlidium lineahfolium
Grammitis linkiana C. Presl (Thelypteridaceae) =Thelypteris linkiana
Grammitis nana Fee =Grammitis blanchetii
Grammitis parietina (Klotzsch) Fee =Enterosora parietina
Grammitis pruinosa (Maxon) C.V. Morton =Ceradenia pruinosa
Grammitis revohita Spreng. ex Willd. (Polypodiaceae) =Pleopeltis astrolepis
Grammitis rigens (Maxon) Proctor =Grammitis xiphopteroides
Grammitis rigescens (Bory ex Willd.) Lellinger =Grammitis flabelliformis
Grammitis sectifrons (Kunze ex Mett.) F. Seym. =Zygophlebia sectifrons
Grammitis serndata (Sw.) Sw. =Cochlidium serridatiim
Grammitis sprucei (Hook.) J. Sm. =Enterosora parietina
Grammitis tatei Copel. =Cochlidium attenuatiim
Grammitis tepuiensis (A.C. Sm.) Vareschi =Cochlidiiim tepuiense
Grammitis transiens (Lindm.) F. Seym. =Ceradenia spixiana
Grammitis trifurcata (L.) Copel. =Enterosora trifurcata
Grammitis yarumalensis (Hieron.) Proctor =Enterosora parietina
Lellingeria apiculata (Kunze ex Klotzsch) A.R. Sm. & R.C. Moran =Grammitis apicidata
Lellingeria limula (H. Christ) A.R. Sm. & R.C. Moran =Grammitis limula
Lellingeria major (Copel.) A.R. Sm. & R.C. Moran =Grammitis major
Lellingeria melanotrichia (Baker) A.R. Sm. & R.C. Moran =Grammitis melanotrichia
Lellingeria myosuroides (Sw.) A.R. Sm. & R.C. Moran =Grammitis myosuroides
Lellingeria shaferi (Maxon) A.R. Sm. & R.C. Moran =Grammitis shaferi
Lellingeria sinuosa (A.R. Sm.) A.R. Sm. & R.C. Moran =Grammitis sinuosa
Lellingeria subsessilis (Baker) A.R. Sm. & R.C. Moran =Grammitis subsessilis
Lellingeria suspensa (L.) A.R. Sm. & R.C. Moran =Grammitis suspense
Lellingeria tenicida A.R. Sm. & R.C. Moran =Grammitis tenuicula
Melpomene anfractuosa (Kunze ex Klotzsch) A.R. Sm. & R.C. Moran =Grammitis anfractuosa
Melpomenefirma (J. Sm.) A.R. Sm. & R.C. Moran =Grammitis firma
Melpomeneflabelliformis (Poir.) A.R. Sm. & R.C. Moran =Grammitis flabelliformis
Melpomene melanosticta (Kunze) A.R. Sm. & R.C. Moran =Grammitis melanosticta
Melpomene moniliformis (Lag. ex Sw.) A.R. Sm. & R.C. Moran =Grammitis moniliformis
Melpomene pilosissima (M. Martens & Galeotti) A.R. Sm. & R.C. Moran ^Grammitis pilosissima
Melpomene xiphopteroides (Liebm.) A.R. Sm. & R.C. Moran =Grammitis xiphopteroides
Micropolypodium caiicamim (Hieron.) A.R. Sm. =Grammitis caiicana
Micropolypodiiim liesneri (A.R. Sm.) A.R. Sm. =Grammitis liesneri
Micropolypodium namim (Fee) A.R. Sm. =Grammitis blanchetii
Micropolypodium plicatum (A.R. Sm.) A.R. Sm. =Grammitis plicata
Micropolypodium taenifolium (Jenman) A.R. Sm. =Grammitis taenifolia
Micropolypodium trichomanoides (Sw.) A.R. Sm. ^Grammitis trichomanoides
Micropolypodium truncicola (Klotzsch) A.R. Sm. =Grammitis truncicola
Monogramma linearifolia Desv. =Cochlidium linearifolium
Nanopteris caucana (Hieron.) Vareschi [nom. inval.] =Grammitis caucana
Nanopteris limula (H. Christ) Pic. Serm. =Grammitis limula
Nanopteris nana (Fee) Vareschi [nom. inval.] =Grammitis blanchetii
Nanopteris taenifolia (Jenman) Vareschi [nom. inval.] =Grammitis taenifolia
Nanopteris truncicola (Klotzsch) Vareschi =Grammitis truncicola
Terpsichore asplenifolia (L.) A.R. Sm. =Grammitis asplenifolia
Terpsichore cultrata (Bory ex Willd.) A.R. Sm. =Grammitis cultrata
Terpsichore kegeliana (Kunze) A.R. Sm. =Grammitis kegeliana
Terpsichore mollissima (Fee) A.R. Sm. =Grammitis mollissima
Terpsichore staheliana (Posth.) A.R. Sm. =Grammitis staheliana
Terpsichore taxifolia (L.) A.R. Sm. =Grammitis taxifolia
Terpsichore xanthotrichia (Klotzsch) A.R. Sm. =Grammitis xanthotrichia
Xiphopteris auyantepuiensis Vareschi =Cochlidium serrulatum
Xiphopteris duale (Maxon) Vareschi =Cochlidium serrulatum
Xiphopteris jamesonii Hook. =Grammitis myosuroides
Xiphopteris limula (H. Christ) Pic. Serm. ^Grammitis limula
Xiphopteris myosuroides (Sw.) Kaulf. =Grammitis myosuroides
Xiphopteris nana (Fee) Copel. =Grammitis blanchetii
Xiphopteris rigescens (Bory ex Willd.) Alston =Grammitis flabelliformis
Xiphopteris serrulata (Sw.) Kaulf. =Cochlidium serrulatum
Xiphopteris shaferi (Maxon) K.U. Kramer =Grammitis shaferi
Xiphopteris strictissima (Hook.) Vareschi =Grammitis myosuroides
Xiphopteris taenifolia (Jenman) Copel. =Grammitis taenifolia
AM BO
BO
AM BO
BO
AM BO
AM
AM BO
AM BO
AM BO
AM BO
38 Checklist of the Plants of the Guiana Shield
Xiphopteris trichomanoides (Sw.) Copel. =Grammitis trichomanoides
Xiphopteris truncicola (Klotzsch) Copel. =Grammitis truncicola
References
Bishop, L.E. 1977. The American species of Grammitis sect. Grammitis. Amer. Fern J. 67: 101-106.
Bishop, L.E. 1978. Revision of the genus Cochlidium. Amer. Fern J. 68: 76-94.
Bishop, L.E. 1988. Ceradenia, a new genus of Grammitidaceae. Amer. Fern J. 78: 1-5.
Bishop, L.E. & A.R. Smith. 1992. Revision of the fern genus Enterosora (Grammitidaceae) in the New World. Syst. Bot. 17:
345-362.
Bishop, L.E. & A.R. Smith. 1995. Grammitidaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan
Guayana2: 135-158.
Copeland, E.B. 1952. The American species of Xiphopteris. Amer. Fern J. 42: 41-52, 93-1 10.
Smith, A.R. 1992. A review of the fern genus Micropolypodium (Grammitidaceae). Novon 2(4): 419-425.
Smith, A.R. 1993. Terpsichore, a new genus of Grammitidaceae (Pteridophyta). Novon 3(4): 478-489.
Smith, A.R. & R.C. Moran. 1992. Melpomene, a new genus of Grammitidaceae (Pteridophyta). Novon 2(4): 426-432.
Smith, A.R., R.C. Moran, & L.E. Bishop. 1994. Lellingeria, a new genus of Grammitidaceae. Amer. Fern J. 81: 76-88.
Hymenophyllaceae
Hymenophyllum angustum Bosch
Hymenophyllum antillense (Jenman) Jenman [Mt. Roraima]
Hymenophyllum apiculatum Mett. ex Kuhn
Hymenophyllum asplenioides (Sw.) Sw.
Hymenophyllum axilIare Sw.
Hymenophyllum brevifrons Kunze
Hymenophyllum decurrens (Jacq.) Sw.
Hymenophyllum dependens C.V. Morton
Hymenophyllum elegans Spreng.
Hymenophyllum fendlerianum J.W. Sturm
Hymenophyllumfucoides (Sw.) Sw.
Hymenophyllum funckii Bosch
Hymenophyllum hirsutum (L.) Sw. BO DA
Hymenophyllumjamesonii Hook. AM
Hymenophyllum kaieteurum Jenman
Hymenophyllum karstenianum J.W. Sturm
Hymenophyllum lanatum Fee
Hymenophyllum lehmannii Hieron.
Hymenophyllum lindenii Hook.
Hymenophyllum microcarpum Desv. [Mt. Roraima]
Hymenophyllum nanostellatum Lellinger
Hymenophyllum plumieri Hook. & Grev.
Hymenophyllum polyanthos (Sw.) Sw.
Hymenophyllum roraimense C.V. Morton [Mt. Roraima]
Hymenophyllum sericeum (Sw.) Sw.
Hymenophyllum sp. A
Hymenophyllum sp. B
Hymenophyllum trichomanoides Bosch
Hymenophyllum trichophyllum Kunth
Hymenophyllum undulatum (Sw.) Sw.
Hymenophyllum valvatum Hook. & Grev. [Mt. Roraima]
Trichomanes accedens C. Presl
Trichomanes angustifrons (Fee) Wess. Boer
Trichomanes ankersii C. Parker ex Hook. & Grev.
Trichomanes anomalum Maxon & C.V. Morton
Trichomanes arbuscula Desv.
Trichomanes bancroftii Hook. & Grev.
Trichomanes bicorne Hook.
Trichomanes botryoides Kaulf.
Trichomanes caliginum Lellinger
Trichomanes capillaceum L.
Trichomanes cellulosum Klotzsch
Trichomanes crinitum Sw.
Trichomanes crispum L.
Trichomanes cristatum Kaulf.
AM BO
BO
AM
AM BO
BO
AM
AM BO
BO
BO
BO
AM BO
AM BO
AM
BO
AM BO
AM BO
BO
AM BO
AM BO
AM
AM
AM BO
AM BO
AM BO
AM BO
GU SU
GU
FG
GU SU FG
GU
GU
GU SU
GU SU
FG?
GU SU FG
GU
SU
GU
GU
GU SU FG
GU?
GU
GU
GU
GU SU FG
GU SU FG
GU SU FG
GU SU?
GU SU FG
GU
GU
GU SU FG
GU SU FG
GU SU?
GU
GU SU FG
GU SU? FG
Checklist of the Plants of the Guiana Shield 39
Trichomanes dactylites Sodiro [Mt. Roraima] AM
Trichomanes diaphanum Kunth AM
Trichomanes diversifrons (Bory) Mett. ex Sadeb. AM
Trichomanes egleri P.G. Windisch AM
Trichomanes ekmanii Wess. Boer [Guianas records due to misidentification AM
fide Lellinger]
Trichomanes elegans Rich. AM
Trichomanes fimbriatum Backh. ex T. Moore [Guianas records due to AM
misidentification fide Lellinger]
Trichomanes guidoi RG Windisch AM
Trichomanes hostmannianum (Klotzsch) Kunze AM
Trichomanes hiimboldtii (Bosch) Lellinger AM
Trichomanes hymenophylloides Bosch [Mt. Roraima] AM
Trichomanes jenmanii Lellinger
Trichomanes kapplerianum J.W. Sturm
Trichomanes krausii Hook. & Grev.
Trichomanes longifolium Desv.
Trichomanes lucens Sw. [Mt. Roraima] AM
Trichomanes macilentum Bosch AM
Trichomanes martiusii C. Presl AM
Trichomanes membranaceum L.
Trichomanes mougeotii Bosch
Trichomanes osmundoides DC. ex Poir.
Trichomanes ovale (E. Foum.) Wess. Boer AM
Trichomanes pedicellatum Desv.
Trichomanes pellucens Kunze AM
Trichomanes pinnatinervium Jenman
Trichomanes pinnatum Hedw. AM
Trichomanes plumosum Kunze
Trichomanes polypodioides L. AM
Trichomanes punctatum Poir. AM
Trichomanes punctatum subsp. labiatum (Jenman) Wess. Boer AM
Trichomanes pyxidiferum L.
Trichomanes radicans Sw. AM
Trichomanes radicans var. radicans AM
Trichomanes resinosum R.C. Moran
Trichomanes rigidum Sw. AM
Trichomanes robustum E. Foum. AM
Trichomanes roraimense Jenman AM
Trichomanes rupestre (Raddi) Bosch
Trichomanes spruceanum Hook.
Trichomanes sprucei Baker AM
Trichomanes steyermarkii P.G. Windisch & A.R. Sm. [Mt. Roraima] AM
Trichomanes tanaicum Hook, ex J.W. Sturm AM
Trichomanes trollii Bergdolt AM
Trichomanes tuerckheimii H. Christ AM
Trichomanes vandenboschii P.G. Windisch AM
Trichomanes vittaria DC. ex Poir. AM
Synonyms and Misidentifications
Davalliopsis elegans (Rich.) Copel. =Trichomanes elegans
Didymoglossum krausii (Hook. & Grev.) C. Presl -Trichomanes krausii
Didymoglossum membranaceum (L.) Vareschi =Trichomanes membranaceum
Didymoglossum ovale E. Foum. =Trichomanes ovale
Didymoglossum palmarum Vareschi =Trichomanes kapplerianum
Didymoglossum punctatum (Poir.) Desv. =Trichomanes punctatum
Feea boryi Bosch, nom. superfi. =Trichomanes diversifrons
Feea botryoides (Kaulf.) Bosch =Trichomanes botryoides
Feea humboldtii Bosch ^Trichomanes humboldtii
Feea nana Bory =Trichomanes botryoides
Feea trollii (Bergdolt) Vareschi =Trichomanes trollii
Homoeotes heterophylla (Humb. & Bonpl. ex Willd.) C. Presl =Trichomanes humboldtii
Hymenophyllum amoenum (Klotzsch) J.W. Sturm =Hymenophyllum fendlerianum
Hymenophyllum ciliatum (Sw.) Sw. =Hymenophyllum hirsutum
BO GU SU FG
BO GU SU FG
BO GU SU
BO GU SU FG
BO GU? SU?
BO
BO GU SU FG
BO GU
BO GU?
GU
BO DA GU SU FG
BO GU
GU
SU FG
BO GU?
BO GU
BO GU SU
BO DA GU SU FG
SU FG
GU
SU
FG
BO GU SU FG
BO GU SU FG
GU SU FG
BO GU
GU
SU FG
GU SU FG
BO DA GU SU FG
BO DA GU SU FG
FG
BO GU SU FG
BO GU SU FG
BO GU
BO GU SU FG
BO FG
BO GU SU?
FG
BO GU
BO GU SU
BO GU
BO GU
BO GU SU FG
BO GU SU FG
BO GU SU
GU SU FG
40 Checklist of the Plants of the Guiana Shield
Hymenophyllum ciliatum var. ornifolium Kunze =Hymenophylhim hirsutiim
Hymenophyllum dejectiim Baker (Hymenophyllopsidaceae) =Hymenophyllopsis dejecta
Hymenophyllum lineare var. antillense Jenman ^Hymenophyllum antillense
Hymenophyllum myriocarpum Hook, [records due to misidentification fide Lellinger]
Hymenophyllum protrusum Hook, [records due to misidentification fide Lellinger]
Hymenophyllum schomburgkii C. Presl ex J.W. Sturm ^Hymenophyllum decurrens
Hymenophyllum siliquosum H. Christ =Hymenophyllum apiculatum
Hymenophyllum surinamense Bosch, nom. superfl. =Hymenophyllum hirsutum
Hymenophyllum trapezoidale Liebm. [records due to misidentification fide Lellinger]
Hymenophyllum undulatum var. fendlerianum (J.W. Sturm) Stolze =Hymenophyllum fendlerianum
Hymenostachys diversifrons Bory =Trichomanes diversifwns
Hymenostachys elegans (Rudge) K. Presl ^Trichomanes diversifrons
Hymenostachys osmundoides (DC. ex Poir.) C. Presl =Trichomanes osmundoides
Lacostea ankersii (C. Parker ex Hook. & Grev.) Prantl =Trichomanes ankersii
Lacostea tanaica (Hook, ex J.W. Sturm) Prantl =Trichomanes tanaicum
Lecanium membranaceum (L.) C. Presl =Trichomanes membranaceum
Mecodium apiculatum (Mett. ex Kuhn) Vareschi ^Hymenophyllum apiculatum
Mecodium asplenioides (Sw.) Copel. =Hymenophyllum asplenioides
Mecodium axillare (Sw.) Copel. =Hymenophyllum axillare
Mecodiumfendlerianum (J.W. Stumi) Copel. =Hymenophyllum fendlerianum
Mecodium microcarpum (Desv.) Copel. =Hymenophyllum microcarpum
Mecodium polyanthos (Sw.) Copel. =Hymenophyllum polyanthos
Mecodium trichomanoides (Bosch) Pic. Serm. =Hymenophyllum trichomanoides
Mecodium undulatum (Sw.) Copel. =Hymenophyllum undulatum
Microgonium kapplerianum (J.W. Sturm) Pic. Serm. =Trichomanes kapplerianum
Neuromanes abruptum Bosch, nom. superfl. =Trichomanes hostmannianum
Neuromanes kaulfussii Bosch, nom. superfl. =Trichomanes pinnatum
Neuromanes schomburgkianum (J.W. Sturm) Bosch =Trichomanes pinnatum
Neuromanes vittaria (DC. ex Poir.) Bosch =Trichomanes vittaria
Neurophyllum hostmannianum Klotzsch =Trichomanes hostmannianum
Neurophyllum pennatum (Kaulf.) K. Presl =Trichomanes pinnatum
Neurophyllum pinnatum (Hedw.) K. Presl =Trichomanes pinnatum
Neurophyllum rhizophyllum (Cav.) Fee =Trichomanes pinnatum
Neurophyllum vittaria (DC. ex Poir.) Prantl =Trichomanes vittaria
Odontomanes hostmannianum (Klotzsch) C. Presl =Trichomanes hostmannianum
Ptilophyllum accedens (C. Presl) Prantl =Trichomanes accedens
Ptilophyllum bancroftii (Hook. & Grev.) Prantl =Trichomanes bancroftii
Ptilophyllum bicorne (Hook.) Prantl =Trichomanes bicorne
Ptilophyllum botryoides (Kaulf) Prantl =Trichomanes botryoides
Ptilophyllum elegans (Rudge) Prantl =Trichomanes diversifrons
Ptilophyllum hostmannianum (Klotzsch) Prantl =Trichomanes hostmannianum
Ptilophyllum pinnatum (Kaulf) Prantl =Trichomanes pinnatum
Ptilophyllum schomburgkianum (J.W. Sturm) Prantl =Trichomanes pinnatum
Ptilophyllum vittaria (DC. ex Poir.) Prantl ^^Trichomanes vittaria
Ragatelus eriophorus C. Presl =Trichomanes crispum
Selenodesmium rigidum (Sw.) Copel. =Trichomanes rigidum
Sphaerocionium antillense (Jenman) Copel. =Hymenophyllum antillense
Sphaerocionium crispum var. amoenum Klotzsch =Hymenophyllum fendlerianum
Sphaerocionium dependens (C.V. Morton) Vareschi =Hymenophyllum dependens
Sphaerocionium elegans (Spreng.) Copel. =Hymenophyllum elegans
Sphaerocionium hirsutum (L.) C. Presl =Hymenophyllum hirsutum
Sphaerocionium karstenianum (J.W. Sturm) Vareschi =Hymenophyllum karstenianum
Sphaerocionium lindenii (Hook.) Vareschi =Hymenophyllum lindenii
Sphaerocionium microcarpum (Desv.) Copel. =Hymenophyllum microcarpum
Sphaerocionium plumieri (Hook. & Grev.) C. Presl =Hymenophyllum plumieri
Sphaerocionium roraimense (C.V. Morton) Vareschi =Hymenophyllum roraimense
Sphaerocionium trichophyllum (Kunth) Copel. =Hymenophyllum trichophyllum
Sphaerocionium valvatum (Hook. & Grev.) Copel. =Hymenophyllum valvatum
Trichomanes abruptum (Bosch) H. Christ =Trichomanes hostmannianum
Trichomanes ankersii var. tanaicum (Hook, ex J.W. Sturm) Sadeb. ^Trichomanes tanaicum
Trichomanes asplenioides Sw. =Hymenophyllum asplenioides
Trichomanes carolianum Vareschi =Trichomanes tanaicum
Trichomanes ciliatum Sw. ^Hymenophyllum hirsutum
Trichomanes commutatum J.W. Sturm =Trichomanes pedicellatum
Checklist of the Plants of the Guiana Shield 41
Trichomanes ciirrami Weath. [Guianas records due to misidentifications of 7] caliginuni] =Trichomanes fimbriatum
Trichomanes digitatum Sodiro =Trichomanes dactylites
Trichomanes elegans Rudge, non Rich. [nom. illeg.] =Trichomanes diversifrons
Trichomanes floribundiim var. vittaria (DC. ex Poir.) Splitg. =Trichomanes vittaria
Trichomanes fruticiilosiim Jenman =Trichomanes punctatiim subsp. labiatum
Trichomanesfucoides Sw. =^Hymenophylhim fucoides
Trichomanes furcatum Bosch, non Kaulf. ex Spreng. =Trichomanes pedicellatum
Trichomanes gemmatum J. Sm. ex Hook. & Baker, nom. superfl. =Trichomanes cellulosum
Trichomanes guianense J.W. Sturm =Trichomanes ankers ii
Trichomanes heterophyllum Humb. & Bonpl. ex Willd. ^Trichomanes humboldtii
Trichomanes hirsutum L. =Hymenophyllum hirsutum
Trichomanes hookeri auct. non C. Presl =Trichomanes kapplerianum
Trichomanes hiiberi H. Christ =Trichomanes hostmannianum
Trichomanes kunzeaniim Hook. =Trichomanes radicans
Trichomanes labiatum Jenman =Trichomanes punctatum subsp. labiatum
Trichomanes lambertianum Hook. =Trichomanes lucens
Trichomanes leprieurii Hook., nom. superfl. =Trichomanes elegans
Trichomanes leptophyllum Bosch [nom. illeg.] =Trichomanes hymenophylloides
Trichomanes myrioneuron Lindm. =Trichomanes punctatum subsp. labiatum
Trichomanes nanum (Bory) Hook. =Trichomanes botryoides
Trichomanes pennatum Kaulf. =Trichomanes pinnatum
Trichomanes pilosum Raddi =Trichomanes martiusii
Trichomanes pinnatum var. kaulfussii Domin =Trichomanes pinnatum
Trichomanes pinnatum var. vittaria (DC. ex Poir.) Baker =Trichomanes vittaria
Trichomanes polyanthos Sw. ='Hymenophyllum polyanthos
Trichomanes polyphlebius V. Marcano =Trichomanes arbuscula
Trichomanes prieurii Kunze =Trichomanes elegans
Trichomanes rhizophyllum Cav. =Trichomanes pinnatum
Trichomanes schomburgkianum J.W. Sturm =Trichomanes pinnatum
Trichomanes schomburgkii Bosch =Trichomanes crispum
Trichomanes sellowianum C. Presl =Trichomanes cristatum
Trichomanes sphenoides auct. non Kunze =Trichomanes punctatum
Trichomanes spicisorum Desv. =Trichomanes osmundoides
Trichomanes subsessile Splitg. =Trichomanes pedicellatum
Trichomanes superbum Bosch =Trichomanes fimbriatum
Trichomanes torotumanii Vareschi =Trichomanes tuerckheimii
Trichomanes trigonum auct. non Desv. [records due to misidentification fide Lellinger] =Trichomanes fimbriatum
Trichomanes trigonum vm. fimbriatum (Backh. ex T. Moore) Proctor =Trichomanes fimbriatum
Trichomanes undulatum Sw. =Hymenophyllum undulatum
Trichomanes undulatum Bosch =Trichomanes vandenboschii
Trichomanes windischianum Lellinger =Trichomanes sprucei
Vandenboschia capillacea (L.) Copel. =Trichomanes capillaceum
References
Lellinger, D.B. 1984. Hymenophyllaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 12. Mem. New
York Bot. Card. 38: 1-84.
Lellinger, D.B. 1991. Notes on neotropical Hymenophyllaceae. Amer. Fern J. 81: 42-46.
Lellinger, D.B. 1994. Trichomanes polypodioides and its allies. Amer. Fern J. 84(1): 1-4.
Lellinger, D.B. 1994. Hymenophyllaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series B) 3: 1-66.
Smith, A.R. 1995. Hymenophyllaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 159-
185.
Hymenophyllopsidaceae
Hymenophyllopsis asplenioides A.C. Sm. [Mt. Roraima]
Hymenophyllopsis ctenitoides Lellinger [Mt. Roraima]
Hymenophyllopsis dejecta (Baker) K.l. Goebel [Mt. Roraima]
Hymenophyllopsis hymenophylloides L.D. Gomez
Hymenophyllopsis incognita Lellinger
Hymenophyllopsis steyermarkii Vareschi
Hymenophyllopsis superba Lellinger
Hymenophyllopsis universitatis Vareschi BO
AM BO GU
AM BO GU'
AM BO GU
AM BO
BO
AM BO
AM
42 Checklist of the Plants of the Guiana Shield
References
Lellinger, D.B. 1984. Hymenophyllopsidaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 12. Mem.
New York Bot. Card. 38: 1-84.
Lellinger, D.B. 1991. Hymenophyllopsidaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series B) 4: 82-86.
Lellinger, D.B. 1995. Hymenophyllopsidaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana
2: 186-188.
LOMARIOPSIDACEAE
Bolbitis lindigii (Mett.) Ching
Bolbitis nicotianifolia (Sw.) Alston AM
Bolbitis portoricensis (Spreng.) Hennipman
Bolbitis semipinnatifida (Fee) Alston AM
Bolbitis serratifolia (Mert. ex Kaulf ) Schott
Elaphoglossum acrocarpum (Mart.) T. Moore AM
Elaphoglossum andicola (Fee) T. Moore AM
Elaphoglossum antioquianum Hieron. AM
Elaphoglossum apodum (Kaulf.) Schott ex J. Sm.
Elaphoglossum auricomum (Kunze) T. Moore
Elaphoglossum brachyneuron (Fee) J. Sm. AM
Elaphoglossum chnitum (L.) H. Christ
Elaphoglossum crispatum Mickel AM
Elaphoglossum crispatum var. beitelii Mickel AM
Elaphoglossum crispatum var. crispatum AM
Elaphoglossum cuspidatum (Willd.) T. Moore AM
Elaphoglossum decoratum (Kunze) T. Moore AM
Elaphoglossum discolor (Kuhn) C. Chr. AM
Elaphoglossum dolichopus Mickel AM
Elaphoglossum engelii (H. Karst.) H. Christ
Elaphoglossum erinaceum (Fee) T. Moore AM
Elaphoglossum eximium (Mett.) H. Christ
Elaphoglossum flaccidum (Fee) T. Moore AM
Elaphoglossum floccosum Mickel
Elaphoglossum foldatsii Vareschi AM
Elaphoglossum glabellum J. Sm. AM
Elaphoglossum hayesii (Mett. ex Kuhn) Maxon
Elaphoglossum herminieri (Bory & Fee ex Fee) T. Moore
Elaphoglossum hoffmannii (Mett. ex Kuhn) H. Christ AM
Elaphoglossum horridulum (Kaulf.) J. Sm. AM
Elaphoglossum incubus Mickel AM
Elaphoglossum laminarioides (Bory ex Fee) T. Moore AM
Elaphoglossum langsdorffii (Hook. & Grev.) T. Moore AM
Elaphoglossum latifolium (Sw.) J. Sm. [see also Elaphoglossum sporadolepis]
Elaphoglossum leptophlebium (Baker) C. Chr.
Elaphoglossum lingua (Raddi) Brack.
Elaphoglossum lloense (Hook.) T. Moore
Elaphoglossum longicaudatum Mickel AM
Elaphoglossum luridum (Fee) H. Christ AM
Elaphoglossum macrophyllum (Mett. ex Kuhn) H. Christ AM
Elaphoglossum maguirei Mickel AM
Elaphoglossum meridense (Klotzsch) T. Moore
Elaphoglossum minutum (Pohl ex Fee) T. Moore [Mt. Roraima]
Elaphoglossum moritzianum (Klotzsch) T. Moore
Elaphoglossum nigrescens (Hook.) T. Moore ex Diels AM
Elaphoglossum obovatum Mickel AM
Elaphoglossum paleaceum (Hook. & Grev.) Sledge
Elaphoglossum papillosum (Baker) H. Christ AM
Elaphoglossum parvulum Mickel AM
Elaphoglossum peltatum (Sw.) Urb. AM
Elaphoglossum piloselloides (C. Presl) T. Moore
Elaphoglossum plumosum (Fee) T. Moore AM
Elaphoglossum polyblepharum Mickel AM
Elaphoglossum praetermissum Mickel AM
Elaphoglossum productum Rosenst. [Mt. Roraima] AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
DA
DA
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU?
GU
GU
GU
GU
GU
GU
GU?
SU
SU
SU
SU
SU
SU
FG
FG
FG
FG
GU
GU SU FG
GU SU
SU
FG
GU SU FG
GU
GU SU FG
GU SU FG
GU
GU FG
SU FG?
FG
GU SU FG
GU FG
FG
FG
FG?
su FG
su FG
FG
su? FG?
su FG
GU SU FG
FG?
GU SU FG
GU SU FG
Checklist of the Plants of the Guiana Shield 43
Elaphoglossiim pteropodiim (Sodiro) C. Chr.
Elaphoglossum pteropus C.Chr. AM BO GU
Elaphoglossum pusillum (Mett. ex Kuhn) C. Chr. BO
Elaphoglossum raywaense (Jenman) Alston AM BO GU
Elaphoglossum revolutum (Liebm.) T. Moore
Elaphoglossum scandens (Bory ex Fee) T. Moore
Elaphoglossum sporadolepis (Kunze ex Kuhn) T. Moore ex C. Chr. AM BO GU
[=Elaphoglossum latifoliuml]
Elaphoglossum squarwsum (Klotzsch) T. Moore AM
Elaphoglossum steyermarkii Mickel BO
Elaphoglossum strictum (Raddi) T. Moore GU SU FG
Elaphoglossum styriacum Mickel AM BO FG?
Elaphoglossum succubus Mickel BO
Elaphoglossum tantalinum Mickel AM
Elaphoglossum tectum (Humb. & Bonpl. ex Willd.) T. Moore AM
Elaphoglossum tenuifolium (Liebm.) T. Moore FG
Elaphoglossum tovarense (Mett. ex Kuhn) T. Moore ex C. Chr. [Mt. Roraima] AM BO
Elaphoglossum vanderwerffii Mickel AM
Elaphoglossum vareschianum Mickel BO
Elaphoglossum wurdackii Vareschi [Mt. Roraima]
' AM BO
Elaphoglossum xiphoides Mickel BO
Elaphoglossum zosteriformis Mickel BO
Lomagramma guianensis (Aubl.) Ching AM BO
Lomariopsis erythrodes (Kunze) Fee
Lomariopsis fendleri D.C. Eaton BO
Lomariopsis japurensis (Mart.) J. Sm. AM BO DA
Lomariopsis prieuriana Fee
Synonyms and Misidentifications
Bolbitis aliena (Sw.) auct. non Alston =Bolbitis semipinnatifida
Bolbitis cladorrhizans (Spreng.) Ching =Bolbitis portoricensis
Bolbitis guianensis (Aubl.) K.U. Kramer ex Vareschi =Lomagramma guianensis
Bolbitis maguirei Maxon & C.V. Morton =Bolbitis serratifolia
Bolbitis scopulina Maxon & C.V. Morton =Bolbitis semipinnatifida
Elaphoglossum agostinii Vareschi ^Elaphoglossum sporadolepis
Elaphoglossum attenuatum (Kunze ex Kuhn) H. Christ =Elaphoglossum hoffmannii
Elaphoglossum cordigerum H. Christ =Elaphoglossum eximium
Elaphoglossum fraseri (Mett. ex Kuhn) H. Christ =Elaphoglossum pusillum
Elaphoglossum gayanum (Fee) T. Moore =Elaphoglossum minutum
Elaphoglossum leptophyllum (Fee) T. Moore =Elaphoglossum minutum
Elaphoglossum lineare var. klotzschii (Moritz) Hieron. =Elaphoglossum eximium
Elaphoglossum longifolium (C. Presl) J. Sm. =Elaphoglossum latifolium
Elaphoglossum reichenbachii (Moritz ex Kuhn) H. Christ ^Elaphoglossum eximium
Elaphoglossum rigidum (Aubl.) Urb. =Elaphoglossum flaccidum
Elaphoglossum schomburgkii (Fee) T. Moore =Elaphoglossum luridum
Elaphoglossum spathulatum auct. non (Bory) T. Moore =Elaphoglossum piloselloides
Elaphoglossum squamosum (Sw.) J. Sm. =Elaphoglossum paleaceum
Elaphoglossum stenopteris (Klotzsch) T. Moore ^Elaphoglossum brachyneuron
Elaphoglossum williamsii Vareschi ^Elaphoglossum nigrescens
Hymenodium crinitum (L.) Fee =Elaphoglossum crinitum
Leptochilus alienus auct. non (Sw.) C. Chr. =Bolbitis semipinnatifida
Leptochilus guianensis (Aubl.) C. Chr. =Lomagramma guianensis
Leptochilus nicotianifolius (Sw.) C. Chr. =Bolbitis nicotianifolia
Leptochilus serratifolius (Mert. ex Kaulf.) C. Chr. =Bolbitis serratifolia
Peltapteris peltata (Sw.) C.V. Morton =Elaphoglossum obovatum
Rhipidopteris peltata (Sw.) Schott =Elaphoglossum peltatum
References
Hennipman, E. 1977. A monograph of the fern genus Bolbitis. Leiden Bot. Ser. 2: 1-331.
Smith, A.R., J.T. Mickel, & R.C. Moran. 1995. Dryopterldaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the
Venezuelan Guayana 2: 73-128.
Lygodiaceae
Lygodium flexuosum (L.) Sw. GU
44 Checklist of the Plants of the Guiana Shield
GU
BO GU su FG
BO DA GU su FG
Lygodium microphyllum (Cav.) R. Br. [introduced from Old World tropics]
Lygodium venustum Sw.
Lygodium volubile Sw.
Synonyms and Misidentifications
Lygodium mexicanum C. Presl =Lygodium venustum
Lygodium micans J.W. Sturm =Lygodium volubile
Lygodium polymorphum auct. non (Cav.) Kunth =Lygodium venustum
References
Duek, J.J. 1978. A taxonomic revision of Lygodium in America. Feddes Repert. 89: 41 1-423.
Lellinger, D.B. 1969. Schizaeaceae. In B. Maguire & collaborators, Botany of the Guayana Highland - Part VIII. Mem. New
York Bot. Gard. 18(2): 2-11.
Smith, A.R. 1995. Schizaeaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 288-296.
Marattiaceae
Danaea elliptica Sm.
Danaea leprieurii Kunze
Danaea moritziana C. Presl
Danaea nodosa (L.) Sm.
Danaea simplicifolia Rudge
Danaea trifoliata Rchb. ex Kunze
Marattia laevis Sm.
AM
AM
AM
AM
BO
BO
BO
BO
BO
GU? SU FG
GU SU FG
FG
SU FG
GU SU FG
GU su FG
Synonyms and Misidentifications
Danaea elliptica subsp. simplicifolia (Rudge) Rolleri =Danaea simplicifolia
Danaea nigrescens Jenman =Danaea nodosa
Marattia kaulfussii J. Sm. =Marattia laevis
References
Smith, A.R. 1995. Marattiaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 206-209.
Marsileaceae
Marsilea deflexa A. Braun
Marsilea minuta L.
Marsilea polycarpa Hook. & Grev. BO DA
GU
GU?
GU SU FG
References
Johnson, D.M. 1986. Systematics of the New World species of Marsilea. Syst. Bot. Monogr. 11: 1-87.
Reed, C.F. 1965. Index Marsileata et Salviniata. Suppl. Bol. Soc. Brot. II, 39: 259-302.
Smith, A.R. 1995. Marsileaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 209-211,
Metaxyaceae
Metaxya rostrata (Kunth) C. Presl AM BO GU SU FG
Synonyms and Misidentifications
Amphidesmium blechnoides (Hook.) J. Sm. =Metaxya rostrata
Amphidesmium rostratum (Kunth) J. Sm. =Metaxya rostrata
References
Smith, A.R. 1995. Metaxyaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 211-212.
Nephrolepis biserrata (Sw.) Schott
Nephrolepis cordifolia (L.) C. Presl
Nephrolepis multiflora (Roxb.) Jarrett ex C.V Morton
Nephrolepis pectinata (Willd.) Schott
Nephrolepis rivularis (Vahl) Mett. ex Krug
Oleandra articulata (Sw.) C. Presl
Oleandra duidae A.C. Sm.
Oleandra pilosa Hook.
AM BO
AM BO
BO
AM BO
AM BO
AM BO
AM BO
DA
DA
GU SU FG
GU su
SU
FG
GU
GU su FG
GU su FG
GU SU FG
Checklist of the Plants of the Guiana Shield 45
Synonyms and Misidentifications
Nephrolepis pendula (Raddi) J. Sm. =Nephrolepis cordifolia
Oleandra nodosa (Willd.) C. Presl =Oleandra articulata
References
Cremers, G. & K.U. Kramer. 1994. Nephrolepidaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series B) 6: 66-72.
Cremers, G. & K.U. Kramer. 1994. Oleandraceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series B) 6: 73-76.
Smith, A.R. 1995. Davalliaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 43-46.
Smith, A.R., J.T. Mickel, & R.C. Moran. 1995. Dryopteridaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the
Venezuelan Guayana 2: 73-128.
BO DA GU SU
BO GU SU FG
GU SU FG?
Ophioglossaceae
Cheiroglossa palmata (L.) C. Presl AM
Ophioglossiim nudicaule L. f.
Ophioglossum reticidatum L.
Synonyms and Misidentifications
Ophioglossum eUipticum Hook. & Grev. =Ophioglossum nudicaule
Ophioglossum macrorhizum Kunze =Ophioglossum nudicaule
Ophioglossum palmatum L. =Cheiroglossa palmata
References
Clausen, R.T. 1938. A monograph of the Ophioglossaceae. Mem. Torrey Bot. Club 19(2): 1-177.
Smith, A.R. 1995. Ophioglossaceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 212-214.
OSMUNDACEAE
Osmunda cinnamomea L. BO
Osmunda regalis L. BO
Synonyms and Misidentifications
Osmunda cervina L. (Dryopteridaceae) =Olfersia cervina
Osmunda humilis Cav. (Schizaeaceae) =Anemia oblongifolia
Osmunda oblongifolia Cav. (Schizaeaceae) =Anemia oblongifolia
Osmunda peltata Sw. (Lomariopsidaceae) =Elaphoglossum obovatum
Parkeriaceae
Ceratoptehs deltoidea Benedict [=Ceratopteris pteridoides?]
Ceratopteris pteridoides (Hook.) Hieron. BO DA
Ceratopteris richardii Brongn.
Ceratopteris thalictroides (L.) Brongn. BO DA
Synonyms and Misidentifications
Ceratopteris calomelanos (L.) Link (Pteridaceae) =Pityrogramma calomelanos
Ceratopteris lockhartii (Hook. & Grev.) Kunze =Ceratopteris pteridoides or Ceratopteris thalictroides
Parkeria pteridoides Hook. =Ceratopteris pteridoides
References
Lloyd, R.M. 1974. Systematics of the genus Ceratopteris Brongn. II. Taxonomy. Brittonia 26: 139-160.
Smith, A.R. 1995. Parkeriaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 215-217.
GU SU
GU SU FG
GU SU FG
GU SU FG
Plagiogyriaceae
Plagiogyria semicordata (C. Presl) H. Christ AM
Synonyms and Misidentifications
Lomaridium semicordatum C. Presl =Plagiogyria semicordata
POLYPODIACEAE
Campyloneurum amphostenon (Kunze ex Klotzsch) Fee [Mt. Roraima] AM BO
46 Checklist of the Plants of the Guiana Shield
Campyloneurum angustifolium (Sw.) Fee
Campyloneunim coarctatum (Kunze) Fee
Campyloneurum fuscosqiiamatum Lellinger
Campyloneurum latum T. Moore
Campyloneurum occultum (H. Christ) L.D. Gomez
Campyloneurum phyllitidis (L.) C. Presl
Campyloneurum repens (Aubl.) C. Presl
Campyloneurum wurdackii B. Leon
Dicranoglossum desvauxii (Klotzsch) Proctor
Dicranoglossumfurcatum (L.) J. Sm.
Microgramma baldwinii Brade
Microgrammafuscopunctata (Hook.) Vareschi
Microgramma geminata (Schrad.) R.M. Tryon & A.F. Tryon
Microgramma lycopodioides (L.) Copel.
Microgramma megalophylla (Desv.) de la Sota
Microgramma persicariifolia (Schrad.) C. Presl
Microgramma piloselloides (L.) Copel.
Microgramma reptans (Cav.) A.R. Sm.
Microgramma tecta (Kaulf.) Alston
Microgramma thurnii (Baker) R.M. Tryon & Stolze
Neurodium lanceolatum (L.) Fee
Niphidium crassifolium (L.) Lellinger
Pecluma camptophyllaria (Fee) M.G. Price
Pecluma camptophyllaria van lachnifera (Hieron.) Lellinger
Pecluma consimilis (Mett.) M.G Price
Pecluma consimilis var. consimilis
Pecluma dispersa (A.M. Evans) M.G. Price
Pecluma emybasis (C. Chr.) M.G. Price [Mt. Roraima]
Pecluma hygrometrica (Splitg.) M.G. Price
Pecluma pectinata (L.) M.G. Price
Pecluma plumula (Humb. & Bonpl. ex Willd.) M.G. Price
Pecluma ptilodon (Kunze) M.G. Price
Pecluma ptilodon var. caespitosa (Jenman) Mickel & Beitel
Pecluma ptilodon var. pilosa (A.M. Evans) Stolze
Pecluma ptilodon var. ptilodon
Phlebodium aureum (L.) J. Sm.
Phlebodium decumanum (Willd.) J. Sm.
Phlebodium pseudoaureum (Cav.) Lellinger
Pleopeltis astrolepis (Liebm.) E. Foum.
Pleopeltis macrocarpa (Bory ex Willd.) Kaulf.
Pleopeltis percussa (Cav.) Hook. & Grev.
Pleopeltis repanda A.R. Sm.
Polypodium adnatum Kunze ex Klotzsch
Polypodium attenuatum Humb. & Bonpl. ex Willd.
Polypodium aturense Maury
Polypodium bombycimim Maxon
Polypodium caceresii Sodiro
Polypodium dissimile L.
Polypodium dulce Poir.
Polypodium flagellare H. Christ
Polypodium fraxinifolium Jacq.
Polypodium funckii Mett.
Polypodium lasiopus Klotzsch
Polypodium levigatum Cav.
Polypodium loriceum L.
Polypodium panorense C. Chr.
Polypodium polypodioides (L.) Watt
Polypodium polypodioides var. burchellii (Baker) Weath.
Polypodium polypodioides var. polypodioides
Polypodium sessilifolium Desv.
Polypodium thyssanolepis A. Braun ex Klotzsch
Polypodium triseriale Sw.
Polypodium wagneri Mett.
Terpsichore bradeana Labiak
AM BO
BO
GU SU FG
GU SU FG
GU SU FG
FG
AM BO DA GU SU FG
AM
AM
BO
BO
DA GU SU FG
AM BO GU SU
SU
FG
FG
AM BO GU
AM BO GU
GU
SU FG
AM BO DA GU SU FG
AM GU SU?
AM BO DA GU
GU
SU FG
FG
AM BO DA GU SU FG
AM GU SU FG
AM BO GU SU FG
FG?
AM BO
BO
BO
BO
GU
GU
GU
SU FG
AM BO
AM BO GU SU
AM BO GU SU FG
AM BO GU SU FG
AM BO GU SU
SU
FG
FG
AM BO GU
GU
SU FG
BO DA GU SU FG
AM BO DA GU SU FG
AM BO GU SU FG
BO GU SU FG
AM BO
AM BO
BO
GU SU FG
AM BO GU SU FG
AM BO GU SU FG
AM BO
AM BO GU
AM BO GU SU FG
GU SU FG
AM BO GU SU
SU
FG
FG
AM BO GU
AM BO
BO SU FG
AM
BO
GU
GU
GU
SU FG
AM BO GU SU FG
AM
AM
BO
BO
BO
GU
GU
GU
SU FG
FG?
AM
AM
BO
BO
BO
DA GU SU FG
Checklist of the Plants of the Guiana Shield 47
Terpsichore steyermarkii Lobiak BO
Synonyms and Misidentifications
Campyloneurum costatum (Kunze) C. Presl =Campyloneurum phyllitidis
Campyloneiiriim sphenodes (Kunze) Fee [not in flora area]
Drynaria acuminata Fee =Microgramma thurnii
Eschatogramme desvauxii (Klotzsch) C. Chr. =Dicranoglossum desvauxii
Eschatogranmiefurcata (L.) C. Chr. =Dicranoglossum fwcatiim
Eschatogramme fiircata auct. non (L.) C. Chr. =Dicranoglossum desvauxii
Goniophlebium adnatum (Kunze ex Klotzsch) T. Moore =Polypodium adnatum
Goniophlebium attenuatum (Humb. & Bonpl. ex Willd.) C. Presl =Polypodium attenuatum
Goniophlebiumfraxinifolium (Jacq.) Moore =Polypodium fraxinifolium
Goniophlebium glaucophyllum (Kunze ex Klotzsch) Fee ^Polypodium levigatum
Goniophlebium lasiopus (Klotzsch) T. Moore =Polypodium lasiopus
Goniophlebium loriceum (L.) J. Sm. ex Hook. =Polypodium loriceum
Goniophlebium triseriale (Sw.) Pic. Serm. =Polypodium triseriale
Marginaria bombycina (Maxon) Pic. Serm. =Polypodium bombycinum
Marginaria polypodioides (L.) Tidestr. =Polypodium polypodioides
Marginaria thyssanolepis (A. Braun ex Klotzsch) Farw. =Polypodium thyssanolepis
Microgramma acuminata (Fee) Lellinger =Microgramma thurnii
Microgramma ciliata (Willd.) Alston =Microgramma reptans
Microgramma percussa (Cav.) de la Sota =Pleopeltis percussa
Paltonium lanceolatum (L.) C. Presl =Neurodium lanceolatum
Pessopteris crassifolia (L.) Underw. & Maxon =Niphidium crassifolium
Phymatodes prominula Maxon =Microgramma lycopodioides
Pleopeltisfuscopunctata (Hook.) R.M. Tryon & A.F. Tryon =Microgrammafuscopunctata
Pleopeltis lanceolata (L.) C. Presl ^Pleopeltis macrocarpa
Pleopeltis polypodioides (L.) E.G. Andrews & Windham =Polypodium polypodioides
Pleopeltis revoluta (Spreng. ex Willd.) A.R. Sm. =Pleopeltis astrolepis
Pleopeltis thyssanolepis (A. Braun ex Klotzsch) E.G. Andrews & Windham =Polypodium thyssanolepis
Polypodium abruptum Desv. (Thelypteridaceae) ^Thelypteris abrupta
Polypodium adiantiforme G Forst. (Davalliaceae) =Rumohra adiantiformis
Polypodium adiantoides Aubl., non Burm. (Dryopteridaceae) =Polybotrya caudata
Polypodium amphostenon Kunze ex Klotzsch =Campyloneurum amphostenon
Polypodium anfractuosum Kunze ex Klotzsch (Grammitidaceae) =Grammitis anfractuosa
Polypodium angustifolium Sw. =Campyloneurum angustifolium
Polypodium angustifolium var. amphostenon (Kunze ex Klotzsch) Baker =Campyloneurum amphostenon
Polypodium apiculatum Kunze ex Klotzsch (Grammitidaceae) =Grammitis apiculata
Polypodium areolatum Humb. & Bonpl. ex Willd. =Phlebodium pseudoaureum
Polypodium aromaticum Maxon (Grammitidaceae) =Grammitis firma
Polypodium asplenifolium L. (Grammitidaceae) =Grammitis asplenifolia
Polypodium astrolepis Liebm. =Pleopeltis astrolepis
Polypodium aureum L. =Phlebodium aureum
Polypodium aureum var. areolatum (Humb. & Bonpl. ex Willd.) Baker =Phlebodium pseudoaureum
Polypodium balbisii Spreng. (Thelypteridaceae) =Thelypteris balbisii
Polypodium binervatum Poir. (Blechnaceae) =Blechnum binervatum
Polypodium blanchetii C. Chr. (Grammitidaceae) =Grammitis blanchetii
Polypodium bolivari de la Sota =Polypodium bombycinum
Polypodium brasiliense Poir. =Polypodium triseriale
Polypodium bryophilum Maxon (Grammitidaceae) =Grammitis bryophila
Polypodium camptophyllarium Fee =Pecluma camptophyllaria
Polypodium camptophyllarium var. lachniferum (Hieron.) A.M. Evans =Pecluma camptophyllaria var. lachnifera
Polypodium capillare Desv. (Grammitidaceae) =Ceradenia capillaris
Polypodium caucanum Hieron. (Grammitidaceae) =Grammitis caucana
Polypodium chimantense Vareschi (Grammitidaceae) =Ceradenia discolor
Polypodium ciliatum Willd. =Microgramma reptans
Polypodium coarctatum Kunze =Campyloneurum coarctatum
Polypodium confusum J. Sm. (Grammitidaceae) =Grammitis suspensa
Polypodium connellii Baker ex C.H. Wright (Grammitidaceae) =Cochlidium connellii
Polypodium consimile Mett. =Pecluma consimilis
Polypodium crassifolium L. =Niphidium crassifolium
Polypodium crispatum (J. Sm.) C.V. Morton (Grammitidaceae) [not in flora area] =Enterosora ecostata (Sodiro) L.E.Bishop
Polypodium cultratum auct. non Willd. (Grammitidaceae) =Grammitis mollissima
Polypodium decumanum Willd. =Phlebodium decumanum
48 Checklist of the Plants of the Guiana Shield
Polypodium decussatum L. (Thelypteridaceae) =Thelypteris decussata
Polypodium demeraranum Baker (Thelypteridaceae) =Thelypteris demerarana
Polypodium denticidatum Sw. (Dryopteridaceae) =Arachniodes denticulata
Polypodium dicranophyllum C. Chr. (Grammitidaceae) =Cochlidium furcatum
Polypodium discolor Hook. (Grammitidaceae) =Ceradenia discolor
Polypodium dispersum A.M. Evans =Pecluma dispersa
Polypodium dissectum Sw. (Dennstaedtiaceae) =Dennstaedtia dissecta
Polypodium divergens Willd. (Tectariaceae) =Lastreopsis effusa subsp. divergens
Polypodium duale Maxon (Grammitidaceae) =Cochlidium serrulatum
Polypodium effusum Sw. (Tectariaceae) =Lastreopsis effusa
Polypodium eurybasis C. Chr. =Pecluma eurybasis
Polypodium exiguum Fee (Grammitidaceae) =Grammitis blanchetii
Polypodiumfirmum Klotzsch (Grammitidaceae) =Grammitis firma
Polypodium flabelliforme Poir. (Grammitidaceae) =Grammitis flabelliformis
Polypodium furcatum (Hook. & Grev.) Mett. (Grammitidaceae) =Cochlidium furcatum
Polypodium fuscopunctatum Hook. =Microgrammafuscopunctata
Polypodium glaucophyllum Kunze ex Klotzsch =Polypodium levigatum
Polypodium guianense Aubl. (Lomariopsidaceae) =Lomagramma guianensis
Polypodium haplophlebicum A.C. Sm. (Grammitidaceae) =Grammitis bryophila
Polypodium hirsutum Desv. (Cyatheaceae) =Cyathea surinamensis
Polypodium horridum L. (Cyatheaceae) =Cnemidaria horrida
Polypodium hostmannii Klotzsch (Thelypteridaceae) =Thelypteris arborescens
Polypodium hygrometricum Splitg. =Pecluma hygrometrica
Polypodium incanum var. burchelli Baker =Polypodium polypodioides var. burchellii
Polypodium induens Maxon (Grammitidaceae) =Grammitis anfractuosa
Polypodium jubiforme Kaulf. (Grammitidaceae) =Grammitis suspensa
Polypodium jungermannioides Klotzsch (Grammitidaceae) =Cerademajungermannioides
Polypodium kaieteurum Jenman (Grammitidaceae) =Grammitis tenuicula
Polypodium kalbreyeri Baker (Grammitidaceae) =Ceradenia kalbreyeri
Polypodium kegelianum Kunze (Grammitidaceae) =Grammitis kegeliana
Polypodium kookenamae Jenman (Grammitidaceae) =Ceradenia kookenamae
Polypodium lachniferum Hieron. =Pecluma camptophyllaria var. lachnifera
Polypodium lanceolatum auct. non L. =Pleopeltis astrolepis
Polypodium leptopodon C.H. Wright (Grammitidaceae) =Grammitis leptopoda
Polypodium I 'herminieri Fee (Grammitidaceae) =Grammitis taxifolia
Polypodium limulum H. Christ (Grammitidaceae) =Grammitis limula
Polypodium longicaudatum Liebm. (Dryopteridaceae) =Stigmatopteris longicaudata
Polypodium lycopodioides L. =Microgramma lycopodioides
Polypodium macrocarpum Bory ex Willd. =Pleopeltis macrocarpa
Polypodium megalodus Schkuhr (Thelypteridaceae) =Thelypteris pennata
Polypodium megalophyllum Desv. =Microgramma megalophylla
Polypodium melanostictum Kunze (Grammitidaceae) =Grammitis melanosticta
Polypodium melanotrichium Baker (Grammitidaceae) =Grammitis melanotrichia
Polypodium microdonton Desv. (Cyatheaceae) =Cyathea microdonta
Polypodium mollissimum Fee (Grammitidaceae) =Grammitis mollissima
Polypodium moniliforme Lag. ex Sw. (Grammitidaceae) =Grammitis moniliformis
Polypodium myosuroides Sw. (Grammitidaceae) =Grammitis myosuroides
Polypodium nanum Fee (Grammitidaceae) =Grammitis blanchetii
Polypodium nematorhizon D.C. Eaton =Microgrammafuscopunctata
Polypodium neriifolium Schkuhr =Polypodium triseriale
Polypodium nervosum Klotzsch (Thelypteridaceae) =Thelypteris rudis
Polypodium oligocarpum Humb. & Bonpl. ex Willd. (Thelypteridaceae) =Thelypteris oligocarpa
Polypodium oppositum Vahl (Thelypteridaceae) =Thelypteris opposita
Polypodium parietinum Klotzsch (Grammitidaceae) =Enterosora parietina
Polypodium pectinatum L. =Pecluma pectinata
Polypodium pennatum Poir. (Thelypteridaceae) =Thelypteris pennata
Polypodium percussum Cav. =Pleopeltis percussa
Polypodium persicariifolium Schrad. =Microgramma persicariifolia
Polypodium phlegmaria J. Sm. (Grammitidaceae) =Grammitis phlegmaria
Polypodium phyllitidis L. =Campyloneurum phyllitidis
Polypodium plantagineum Jacq. (Tectariaceae) =Tectaria plantaginea
Polypodium plumula Humb. & Bonpl. ex Willd. =Pecluma plumula
Checklist of the Plants of the Guiana Shield 49
Polypodium plumulum Humb. & Bonpl. ex Willd. =Pecluma plumula
Polypodium pressiim Brause (Grammitidaceae) =Ceradema kalbreyeri
Polypodium procerum Willd. (Cyatheaceae) =Cyathea pungens
Polypodium priiinatiim Baker (Grammitidaceae) =Ceradenia pruinosa
Polypodium pruinosum Maxon (Grammitidaceae) =Ceradenia pruinosa
Polypodium pseudoaureum Cav. =Phlebodium pseudoaureum
Polypodium pseudofraternum A.C. Sm. ^Polypodium sessilifolium
Polypodium pteroideum Klotzsch (Thelypteridaceae) =Thelypteris pteroidea
Polypodium pteropus Hook. (Grammitidaceae) =Grammitis subsessilis
Polypodium ptilodon Kunze =Pecluma ptilodon
Polypodium ptilodon war. pilosum A.M. Evans =Pecluma ptilodon var. pilosa
Polypodium pungens Willd. (Cyatheaceae) =Cyathea pungens
Polypodium repens Aubl. =Campyloneurum repens
Polypodium rigens Maxon (Grammitidaceae) =Grammitis xiphopteroides
Polypodium rigescens Bory ex Willd. (Grammitidaceae) =Grammitis flabelliformis
Polypodium rivulare Vahl (Oleandraceae) =Nephrolepis rivularis
Polypodium roraimense Brause (Grammitidaceae) =Ceradenia kalbreyeri
Polypodium roraimense Baker (Thelypteridaceae) =Thelypteris roraimensis
Polypodium rostratum Willd. (Metaxyaceae) =Metaxya rostrata
Polypodium rude Kunze (Thelypteridaceae) =Thelypteris rudis
Polypodium sancti-gabrieli Hook. (Dryopteridaceae) =Cyclodium guianense
Polypodium sectifrons Kunze ex Mett. (Grammitidaceae) =Zygophlebia sectifrons
Polypodium semicordatum Sw. (Tectariaceae) =Cyclopeltis semicordata
Polypodium serrulatum (Sw.) Mett. (Grammitidaceae) =Cochlidium serrulatum
Polypodium serrulatum var. strictissimum Hook. (Grammitidaceae) ^Grammitis myosuroides
Polypodium sororium Humb. & Bonpl. ex Willd. =Polypodium dulce
Polypodium sphenodes Kunze [not in flora area] =Campyloneurum sphenodes (Kunze) Fee
Polypodium spixianum Mart, ex Mett. (Grammitidaceae) =Ceradenia spixiana
Polypodium sprucei Hook. (Grammitidaceae) =Enterosora parietina
Polypodium stahelianum Posth. (Grammitidaceae) =Grammitis staheliana
Polypodium strictissimum (Hook.) Hieron. (Grammitidaceae) =Grammitis myosuroides
Polypodium subincisum Willd. (Tectariaceae) =Megalastrum subincisum
Polypodium subobliquatum Hook. (Dryopteridaceae) =Cyclodium inerme
Polypodium subsessile Baker (Grammitidaceae) =Grammitis subsessilis
Polypodium subtetragonum Link (Thelypteridaceae) =Thelypteris tetragona
Polypodium surinamensis Jacq. =Microgramma lycopodioides
Polypodium suspensum L. (Grammitidaceae) =Grammitis suspensa
Polypodium taenifolium Jenman (Grammitidaceae) =Grammitis taenifolia
Polypodium taxifolium L. (Grammitidaceae) =Grammitis taxifolia
Polypodium tectum Kaulf. =Microgramma tecta
Polypodium tenuiculum Fee (Grammitidaceae) =Grammitis tenuicula
Polypodium tenuiculum var. acrosorum Hieron. (Grammitidaceae) =Grammitis major
Polypodium tepuiense A.C. Sm. (Grammitidaceae) =Cochlidium tepuiense
Polypodium tetragonum Sw. (Thelypteridaceae) =Thelypteris tetragona
Polypodium thurnii Baker =Microgramma thurnii
Polypodium tottum Thunb. (Thelypteridaceae) =Thelypteris interrupta
Polypodium transiens Lindm. (Grammitidaceae) =Ceradenia spixiana
Polypodium trichomanoides Sw. (Grammitidaceae) =Grammitis trichomanoides
Polypodium trifoliatum L. (Tectariaceae) =Tectaria trifoliata
Polypodium trifurcatum L. (Grammitidaceae) =Enterosora trifurcata
Polypodium triste Kunze (Thelypteridaceae) =Thelypteris tristis
Polypodium truncicola Klotzsch (Grammitidaceae) =Grammitis truncicola
Polypodium xanthotrichium Klotzsch (Grammitidaceae) =Grammitis xanthotrichia
Polypodium xiphopteroides Liebm. (Grammitidaceae) =Grammitis xiphopteroides
Polypodium yarumalense Hieron. (Grammitidaceae) =Enterosora parietina
Taenitis desvauxii Klotzsch =Dicranoglossum desvauxii
References
Lellinger, D.B. 1972. A revision of the fern genus Niphidium. Amer. Fern J. 62: 101-120.
Lellinger, D.B. 1988. Some new species of Campyloneurum and a provisional key to the genus. Amer. Fern J. 78(1): 14-35.
Price, M.G 1983. Pecluma, a new tropical American fern genus. Amer. Fern J. 73(3): 109-116.
Smith, A. R. 1995. Polypodiaceae. In J. A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 219-249.
50 Checklist of the Plants of the Guiana Shield
Pteridaceae
Acrostichwn aiireiim L. DA
Acrostichum danaeifoliiim Langsd. & Fisch. [expected in DA]
Pteris altissima Poir. AM BO
Pteris biaurita L.
Pteris deflexa Link
Pteris denticulata Sw.
Pteris pearcei Baker AM BO
Pteris plumula Desv.
Pteris podophylla Sw. AM
Pteris polita Link [see also Pteris propinqua]
Pteris propinqua i. Agardh [=Pteris polital] AM BO
Pteris pungens W\M. AM DA
Pteris quadriaiirita Retz. [neotropical material =P. plumula, fide D. Lellinger] BO
Pteris splendens Kaulf.
Pteris tripartita Sw. [naturalized] DA
Pteris vittata L. [naturalized]
Synonyms and Misidentifications
Acrostichum acrocarpiim Mart. (Lomariopsidaceae) =Elaphoglossum acrocarpum
Acrostichum alatum Fee (Lomariopsidaceae) =Elaphoglossum pferopus
Acrostichum andicola Fee (Lomariopsidaceae) =Elaphoglossum andicola
Acrostichum auricomum Kunze (Lomariopsidaceae) =Elaphoglossum auricomum
Acrostichum bonariense Willd. (Adiantaceae) =Cheilanthes bonariensis
Acrostichum brachyneuron Fee (Lomariopsidaceae) =Elaphoglossum brachyneuron
Acrostichum calomelanos L. (Adiantaceae) =Pityrogramma calomelanos
Acrostichum chrysolepis Fee (Lomariopsidaceae) =Elaphoglossum plumosum
Acrostichum citrifolium L. (Vittariaceae) =Anetium citrifolium
Acrostichum cladorrhizans Spreng. (Lomariopsidaceae) =Bolbitis portoricensis
Acrostichum compactum Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum sporadolepis
Acrostichum corpulentum Kunze ex Kuhn (Lomariopsidaceae) =Elaphoglossum sporadolepis
Acrostichum crinitum L. (Lomariopsidaceae) =Elaphoglossum crinitum
Acrostichum cuspidatum Willd. (Lomariopsidaceae) =Elaphoglossum cuspidatum
Acrostichum decoratum Kunze (Lomariopsidaceae) =Elaphoglossum decoratum
Acrostichum discolor Kuhn (Lomariopsidaceae) ^Elaphoglossum discolor
Acrostichum ebeneum L. (Adiantaceae) =Pityrogramma ebenea
Acrostichum elegans Vahl (Schizaeaceae) =Schizaea elegans
Acrostichum erinaceum Fee (Lomariopsidaceae) =Elaphoglossum erinaceum
Acrostichum eximium Mett. (Lomariopsidaceae) =Elaphoglossum eximium
Acrostichum flaccidum Fee (Lomariopsidaceae) =Elaphoglossum flaccidum
Acrostichum fraseri Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum pusillum
Acrostichum gayanum (Fee) T. Moore (Lomariopsidaceae) =Elaphoglossum minutum
Acrostichum hayesii Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum hayesii
Acrostichum herminieri Bory ex Fee (Lomariopsidaceae) =Elaphoglossum herminieri
Acrostichum hoffmannii Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum hoffmannii
Acrostichum horridulum Kaulf. (Lomariopsidaceae) =Elaphoglossum horridulum
Acrostichum japurense Mart. (Lomariopsidaceae) =Lomariopsis japurensis
Acrostichum laminarioides Fee (Lomariopsidaceae) =Elaphoglossum laminarioides
Acrostichum langsdorffii Hook. & Grev. (Lomariopsidaceae) =Elaphoglossum langsdorffii
Acrostichum leptophlebium Baker (Lomariopsidaceae) =Elaphoglossum leptophlebium
Acrostichum leptophyllum Fee (Lomariopsidaceae) =Elaphoglossum minutum
Acrostichum luridum Fee (Lomariopsidaceae) =Elaphoglossum luridum
Acrostichum macrophyllum Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum macrophyllum
Acrostichum meridense Klotzsch (Lomariopsidaceae) =Elaphoglossum meridense
Acrostichum minutum Pohl ex Fee (Lomariopsidaceae) =Elaphoglossum minutum
Acrostichum nicotianifolium Sw. (Lomariopsidaceae) =Bolbitis nicotianifolia
Acrostichum nigrescens Hook. (Lomariopsidaceae) =Elaphoglossum nigrescens
Acrostichum nodosum L. (Marattiaceae) '=Danaea nodosa
Acrostichum paleaceum Hook. & Grev. (Lomariopsidaceae) =Elaphoglossum paleaceum
Acrostichum papillosum Baker (Lomariopsidaceae) =Elaphoglossum papillosum
Acrostichum piloselloides C. Presl (Lomariopsidaceae) =Elaphoglossum piloselloides
Acrostichum plumosum Fee (Lomariopsidaceae) =Elaphoglossum plumosum
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU
GU
FG
SU FG
SU FG
GU SU FG
FG
GU
SU FG
SU
Checklist of the Plants of the Guiana Shield 51
Acrostichum polypodioides L. (Polypodiaceae) =Polypodium polypodioides
Acrostichum portoricense Spreng. (Lomariopsidaceae) =Bolbitis portoricensis
Acrostichum pusillum Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum pusillum
Acrostichum raywaense Jenman (Lomariopsidaceae) =Elaphoglossum raywaense
Acrostichum reichenbachii Moritz ex Kuhn (Lomariopsidaceae) =Elaphoglossum eximium
Acrostichum reptans Cav. (Polypodiaceae) =Microgramma reptans
Acrostichum rufum (L.) L. (Adiantaceae) =Hemionitis rufa
Acrostichum schomburgkii Fee (Lomariopsidaceae) =Elaphoglossum luridum
Acrostichum serratifolium Mert. ex Kaulf. (Lomariopsidaceae) =Bolbitis serratifolia
Acrostichum serrulatum Sw. (Grammitidaceae) =Cochlidium serrulatum
Acrostichum sporadolepis Kunze ex Kuhn (Lomariopsidaceae) =Elaphoglossum sporadolepis
Acrostichum squamosum Sw. (Lomariopsidaceae) =Elaphoglossum paleaceum
Acrostichum squarrosum Klotzsch (Lomariopsidaceae) =Elaphoglossum squarrosum
Acrostichum stenopteris Klotzsch (Lomariopsidaceae) =Elaphoglossum brachyneuron
Acrostichum tartareum Cav. (Adiantaceae) =Pityrogramma ebenea
Acrostichum tectum Humb. & Bonpl. ex Willd. (Lomariopsidaceae) ^Elaphoglossum tectum
Acrostichum thalictroides L. (Parkeriaceae) =Ceratopteris thalictroides
Acrostichum tovarense Mett. ex Kuhn (Lomariopsidaceae) =Elaphoglossum tovarense
Acrostichum trifoliatum L. (Adiantaceae) =Pityrogramma trifoliata
Litobrochia podophylla (Sw.) C. Presl =Pteris podophylla
Litobrochia tripartita (Sw.) C. Presl =Pteris tripartita
Neurogramme elaphoglossoides (Baker) H. Christ (Adiantaceae) =Pterozonium elaphoglossoides
Ormopteris pinnata (Kaulf.) Lellinger (Adiantaceae) =Cassebeera pinnata
Pteris angustifolia Sw. (Vittariaceae) =Ananthacorus angustifolius
Pteris arachnoidea Kaulf. (Dennstaedtiaceae) =Pteridium arachnoideum
Pteris aurea Poir. (Adiantaceae) =Cheilanthes bonariensis
Pteris caudata L. (Dennstaedtiaceae) =Pteridium caudatum
Pteris concolor Langsd. & Fisch. (Adiantaceae) =Cheilanthes concolor
Pteris incisa Thunb. (Dennstaedtiaceae) =Histiopteris incisa
Pteris interrupta Willd. (Thelypteridaceae) =Thelypteris interrupta
Pteris kunzeana J. Agardh =Pteris altissima
Pteris lindeniana (Hook.) H. Christ (Dennstaedtiaceae) =Blotiella lindeniana
Pteris lineata L. (Vittariaceae) ^Vittaria lineata
Pteris lucida Cav. (Adiantaceae) =Adiantum lucidum
Pteris palmata Willd. (Adiantaceae) =Doryopteris pedata var. palmata
Pteris pedata L. (Adiantaceae) ^^Doryopteris pedata
Pteris petiolulata R.M. Tryon =Pteris pearcei
Pteris sagittifolia Raddi (Adiantaceae) =Doryopteris sagittifolia
References
Smith, A.R. & D.B. Lellinger. 1995. Pteridaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana
2: 250-286.
Salviniaceae
Azolla caroliniana Willd. AM DA
Azolla mexicana C. Presl
Azolla microphylla Kaulf.
Salvinia auriculata Aubl. BO DA
Salvinia radula Baker
Synonyms and Misidentifications
Salvinia laevigata Humb. & Bonpl. ex Willd. (Hydrocharitaceae) =Limnobium laevigatum
Salvinia rotundifolia Willd. =Salvinia auriculata
References
Reed, C.F. 1965a. Distribution of Salvinia and Azolla in South America and Africa in connection with studies for control by
insects. Phytologia 12(3): 121-130.
Reed, C.F. 1965b. Index Marsileata et Salviniata. Suppl. Bol. Soc. Brot. II, 39: 259-302.
Smith, A.R. 1995. Azollaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 22-23.
Smith, A.R. 1995. Salviniaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 287-288.
Svenson, H.K. 1944. The New World species of Azolla. Amer. Fern J. 34: 69-84.
GU SU FG
GU
GU
GU SU FG
GU SU FG
52 Checklist of the Plants of the Guiana Shield
SCHIZAEACEAE
Actinostachys penmila (Sw.) Hook.
Actinostachys subtrijiiga (Mart.) C. Presl
Anemia ayacuchensis Mickel
Anemia buniifolia (Gardner) T. Moore
Anemia collina Raddi
Anemiaferruginea Kunth
Anemiaferruginea var. ahenobarba (H. Christ) Mickel
Anemiaferrugineavar. ferriiginea
Anemiaflexuosa (Sav.) Sw.
Anemia hirsuta (L.) Sw.
Anemia hirta (L.) Sw.
Anemia mandioccana Hook.
Anemia millefolia (Gardner) C. Presl
Anemia oblongifolia (Cav.) Sw.
Anemia pastinacaria Moritz ex Prantl
Anemia phyllitidis (L.) Sw.
Anemia porrecta Mickel
Schizaea elegans (Vahl) Sw.
Schizaeafluminensis Miers ex J.W. Sturm
Schizaea incurvata Schkuhr
Schizaea poeppigiana J.W. Sturm
Schizaea spnicei Hook.
Schizaea sthcta Lellinger
Synonyms and Misidentifications
Actinostachys penicillata (Humb. & Bonpl. ex Willd.) Maxon =Actinostachys pennula
Anemia humilis (Cav.) Sw. =Anemia oblongifolia
Anemia villosa Humb. & Bonpl. ex Willd. [not in flora area] ^Anemiaflexuosa
Coptophyllum buniifolium Gardner =Anemia buniifolia
Coptophyllum millefolium Gardner =Anemia millefolia
Lophidium elegans (Vahl) C. Presl =Schizaea elegans
Lophidium poeppigianum (J.W. Sturm) Underw. =Schizaea poeppigiana
Schizaeaflabellum Mart. =Schizaea elegans
Schizaea pennula Sw. =Actinostachys pennula
Schizaea subtrijuga Mart. =Actinostachys subtrijuga
References
Lellinger, D.B. 1969. Schizaeaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 8. Mem. New York
Bot. Gard. 18(2): 1-290.
Smith, A.R. 1995. Schizaeaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 288-296.
Tectariaceae
AM BO GU su FG
AM GU su FG
AM
AM BO
GU
su
AM BO GU su
su
AM BO GU su
AM BO
BO
GU
GU
GU
GU
GU
su
su
FG
AM BO GU su
BO GU
GU
su
su
FG
AM
AM BO GU su FG
AM BO GU su FG
AM BO GU su FG
AM BO GU
AM
AM BO GU
Ctenitis falciculata (Raddi) Ching
Ctenitis paranaensis (C. Chr.) Lellinger
Ctenitis refulgens (Klotzsch ex Mett.) C. Chr. ex Vareschi
Cyclopeltis semicordata (Sw.) J. Sm.
Lastreopsis amplissima (C. Presl) Tindale
Lastreopsis effusa (Sw.) Tindale
Lastreopsis effusa subsp. divergens (Willd.) Tindale
Lastreopsis killipii (C. Chr. & Maxon) Tindale
Megalastrum crenulans (Fee) A.R. Sm. & R.C. Moran
Megalastrum subincisum (Willd.) A.R. Sm. & R.C. Moran
Tectaria antioquioana (Baker) C. Chr.
Tectaria heracleifolia (Willd.) Underw.
Tectaria incisa Cav.
Tectaria lizarzaburui (Sodiro) C. Chr.
Tectaria plantaginea (Jacq.) Maxon
Tectaria plantaginea var. macrocarpa (Fee) C.V Morton
Tectaria plantaginea var. plantaginea
Tectaria transiens (C.V. Morton) A.R. Sm.
Tectaria trifoliata (L.) Cav.
AM BO
BO
AM BO
BO
BO
AM
AM BO
AM
BO
AM BO
AM
AM BO
AM BO
AM BO
DA
DA
GU
GU
GU SU FG
GU SU FG
GU
GU SU FG
GU su FG
GU?
BO DA
GU SU
GU su FG
GU SU FG
GU su FG
GU FG
GU su FG
Checklist of the Plants of the Guiana Shield 53
GU su
GU?
GU su FG
BO GU su FG
BO GU su FG
Tectaria trin itens is Maxon
Triplophyllum acutilobum Holttum
Triplophyllum angustifolium Holttum
Triplophyllum dicksonioides (Fee) Holttum
Triplophyllum funestum (Kunze) Holttum
Synonyms and Misidentifications
Bathmium macrocarpon Fee =Tectaria plantaginea var. macrocarpa
Bathmium sinuatum Fee =Tectaria plantaginea var. macrocarpa
Ctenitis crenulans (Fee) Ching =Megalastrum crenulans
Ctenitis effusa (Sw.) Copel. =Lastreopsis effusa
Ctenitisfunesta (Kunze) Vareschi =Triplophyllum funestum
Ctenitis protensa (Afzel. ex Sw.) Ching var. funesta (Kunze) Proctor =Triplophyllum funestum
Ctenitis subincisa (Willd.) Ching =Megalastrum subincisum
Lastreopsis effusa subsp. effusa [not in flora area]
Phegopteris refulgens Klotzsch ex Mett. =Ctenitis refulgens
Tectaria martinicensis (Spreng.) Copel. =Tectaria incisa
Triplophyllum crassifolium Holttum =Triplophyllum funestum
Triplophyllum crassifolium var. caudatum Holttum =Triplophyllumfunestum
Triplophyllum funestum var. hirsutum Holttum =Triplophyllum funestum
References
(Included in Dryopteridaceae in Flora of the Venezuelan Guayana and Flora of the Guianas)
Cremers, G, K.U. Kramer, R.C. Moran, & A.R. Smith. 1994. Dryopteridaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas
(series B) 6: 3-65.
Holttum, R.E. 1986. Studies in the fern genera allied to Tectaria Cav. V. Triplophyllum, a new genus ofAfrica and America. Kew
Bull. 41:237-260.
Smith, A.R., J.T. Mickel, & R.C. Moran. 1995. Dryopteridaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the
Venezuelan Guayana 2: 73-128.
Tindale, M.D. 1965. A monograph of the genus Lastreopsis Ching. Contr. New South Wales Nat. Herb. 3: 249-339.
Thelypteridaceae
Thelypteris abrupta (Desv.) Proctor
Thelypteris angustifolia (Willd.) Proctor
Thelypteris arborea (Brause) A.R. Sm. [Mt. Roraima]
Thelypteris arborescens (Humb. & Bonpl. ex Willd.) C.V Morton
Thelypteris balbisii (Spreng.) Ching
Thelypteris binervata A.R. Sm.
Thelypteris biolleyi (H. Christ) Proctor
Thelypteris brachypoda (Baker) C.V. Morton [Mt. Roraima]
Thelypteris chrysodioides (Fee) C.V. Morton
Thelypteris chrysodioides var. goyazensis (Maxon & C.V. Morton) C.V.
Morton
Thelypteris decussata (L.) Proctor
Thelypteris decussata var. decussata
Thelypteris deflexa (C. Presl) R.M. Tryon
Thelypteris demerarana (Baker) C.F. Reed [Mt. Roraima]
Thelypteris funckii (Mett.) Alston [Mt. Roraima]
Thelypteris gardneriana (Baker) C.F. Reed
Thelypteris gemmulifera (Hieron.) A.R. Sm.
Thelypteris germaniana (Fee) Proctor
Thelypteris glandulosa (Desv.) Proctor
Thelypteris glandulosa var. glandulosa
Thelypteris gonophora (Weath.) A.R. Sm.
Thelypteris hispidula (Decne.) C.F. Reed
Thelypteris hispidula var. hispidula
Thelypteris holodictya K.U. Kramer
Thelypteris interrupta (Willd.) K. Iwats.
Thelypteris juruensis (C. Chr.) R.M. Tryon & D.S. Conant
Thelypteris kunthii (Desv.) C.V. Morton
Thelypteris leprieurii (Hook.) R.M. Tryon
Thelypteris leprieurii var. leprieurii
Thelypteris linkiana (C. Presl) R.M. Tryon [Mt. Roraima]
Thelypteris macrophylla (Kunze) C.V. Morton
BO DA
DA
GU SU FG
AM BO GU
AM BO
BO
GU
GU
SU FG
AM
AM
BO
BO
BO
GU
GU
BO GU su FG
BO GU su FG
AM
BO GU?
GU
AM
FG
FG
GU su FG
GU su FG
GU
AM BO DA GU su FG
AM BO DA GU su
su
FG
FG
BO GU
GU
su FG
FG
AM BO GU su FG
AM BO
BO
GU
GU
su FG
AM GU su FG
54 Checklist of the Plants of the Guiana Shield
Thelypteris nephrodioides (Klotzsch) Proctor
Thelypteris nesiotica (Maxon & C.V. Morton) C.V. Morton
Thelypteris oligocarpa (Humb. & Bonpl. ex Willd.) Ching
Thelypteris opposita (Vahl) Ching
Thelypteris opulenta (Kaulf.) Fosberg [naturalized]
Thelypteris pennata (Poir.) C.V. Morton
Thelypteris peradenia A.R. Sm. [Mt. Roraima]
Thelypteris pilosissima C.V. Morton [Mt. Roraima]
Thelypteris poiteana (Bory) Proctor
Thelypteris pteroidea (Klotzsch) R.M. Tryon [Mt. Roraima]
Thelypteris x rolandii (C. Chr.) R.M. Tryon
Thelypteris roraimensis (Baker) C.F. Reed [Mt. Roraima]
Thelypteris riidis (Kunze) Proctor
Thelypteris salzmannii (Fee) C.V. Morton
Thelypteris schombiirgkii A.R. Sm.
Thelypteris serrata (Cav.) Alston
Thelypteris tetragona (Sw.) Small
Thelypteris tristis (Kunze) R.M. Tryon
GU FG
BO GU
BO
BO
AM BO GU SU FG?
BO DA GU SU FG
BO
BO
GU SU FG
AM BO
AM
BO GU
AM BO GU
GU
GU
DA GU SU FG
DA GU SU FG
AM BO DA GU SU FG
Synonyms and Misidentifications
Amauropelta deflexa (C. Presl) A. Love & D. Love =Thelypteris deflexa
Amauropelta diplazioides (Desv.) Pic. Serm. =Thelypteris linkiana
Amauropelta oligocarpa (Humb. & Bonpl. ex Willd.) Pic. Serm. =Thelypteris oligocarpa
Amauropelta opposita (Vahl) Pic. Serm. =Thelypteris opposita
Amauropelta rudis (Kunze) Pic. Serm. =Thelypteris rudis
Amphineuron opulentum (Kaulf.) Holttum =Thelypteris opulenta
Christella hispidula (Decne.) Holttum =Thelypteris hispidula
Cyclosorus gongylodes (Schkuhr) Link =Thelypteris interrupta
Glaphyropteris decussata (L.) Fee =Thelypteris decussata
Goniopteris biolleyi (H. Christ) Pic. Serm. =Thelypteris biolleyi
Goniopteris juriiensis (C. Chr.) Brade =Thelypterisjuruensis
Goniopteris megalodus (Schkuhr) C. Presl =Thelypteris pennata
Goniopteris pennata (Poir.) Pic. Serm. =Thelypteris pennata
Goniopteris poiteana (Bory) Ching =Thelypteris poiteana
Goniopteris pyramidata Fee =Thelypteris abrupta
Goniopteris subtetragona (Link) Vareschi =Thelypteris tetragona
Goniopteris tetragona (Sw.) C. Presl =Thelypteris tetragona
Goniopteris tristis (Kunze) Brade =Thelypteris tristis
Lastrea adiantiformis (G Forst.) Vareschi (Davalliaceae) =Rumohra adiantiformis
Lastrea brachypoda (Baker) Vareschi =Thelypteris brachypoda
Lastrea coarctata (Kunze) T. Moore =Thelypteris opposita
Lastrea demerarana (Baker) Vareschi =Thelypteris demerarana
Lastrea denticulata (Sw.) T. Moore (Dryopteridaceae) =Arachniodes denticulata
Lastrea engelii (Hieron.) Vareschi [nomen nudum] ^Thelypteris rudis
Lastrea lindigii (C. Chr.) Vareschi =Thelypteris deflexa
Lastrea linkiana (C. Presl) Copel. =Thelypteris linkiana
Lastrea nervosa (Klotzsch) Vareschi [comb, illeg.] ^Thelypteris rudis
Lastrea oligocarpa (Humb. & Bonpl. ex Willd.) T. Moore ^Thelypteris oligocarpa
Lastrea opposita (Vahl) Gilli =Thelypteris opposita
Lastrea pilosissima (C.V. Morton) Vareschi ^Thelypteris pilosissima
Lastrea poiteana Bory =Thelypteris poiteana
Lastrea roraimensis (Baker) Brade =Thelypteris roraimensis
Meniscium angustifolium Willd. =Thelypteris angustifolia
Meniscium arborescens Humb. & Bonpl. ex Willd. ^Thelypteris arborescens
Meniscium chyrsodioides Fee =Thelypteris chrysodioides
Meniscium cristatum Desr. (Woodsiaceae) =Diplazium cristatum
Meniscium guyanense Fee =Thelypteris macrophylla
Meniscium macrophyllum Kunze =Thelypteris macrophylla
Meniscium serratum Cav. =Thelypteris serrata
Steiropteris gardneriana (Baker) Pic. Serm. =Thelypteris gardneriana
Steiropteris glandulosa (Desv.) Pic. Serm. =Thelypteris glandulosa
Steiropteris leprieurii (Hook.) Pic. Serm. ^Thelypteris leprieurii
Thelypteris coarctata (Kunze) R.M. Tryon =Thelypteris opposita
Thelypteris connexa (Kuhn ex Baker) C.F. Reed [not in flora area, type erroneously cited from FG]
Thelypteris extensa (Blume) C.V. Morton ^Thelypteris opulenta
Checklist of the Plants of the Guiana Shield 55
Thelypteris gongylodes (Schkuhr) Small =Thelypteris interrupta
Thelypteris guyanensis (Fee) C.V. Morton =Thelypteris macwphylla
Thelypteris hostmannii (Klotzsch) C.V. Morton ^Thelypteris arborescens
Thelypteris lindigii (C. Chr.) Alston =Thelypteris deflexa
Thelypteris megalodus (Schkuhr) Proctor =Thelypteris pennata
Thelypteris mercurii (A. Braun ex Hieron.) C.F. Reed =Thelypteris balbisii
Thelypteris nervosa (Klotzsch) R.M. Tryon =Thelypteris rudis
Thelypteris quadrangularis (Fee) Schelpe =Thelypteris hispidula
Thelypteris sprengelii (Kaulf.) Proctor =Thelypteris balbisii
Thelypteris totta (Thunb.) Schelpe =Thelypteris interrupta
References
Smith, A.R. 1994. Thelypteridaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series B) 6: 77-1 15.
Smith, A.R. 1995. Thelypteridaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 315-326.
VlTTARIACEAE
AM BO GU SU FG
AM DA GU SU FG
AM BO DA GU SU FG
AM BO GU
GU
GU
SU
SU
SU
FG
FG
FG
AM BO
AM
GU
AM BO GU SU FG
AM BO GU SU
AM BO GU SU
AM BO DA GU
GU
SU FG
AM BO
BO
DA GU SU FG
FG
Ananthacorus angustifolius (Sw.) Underw. & Maxon
Anetium citrifolium (L.) Splitg.
Antrophyum cajenense (Desv.) Spreng.
Antrophyum guayanense Hieron.
Antrophyum jenmanii Benedict
Antrophyum lanceolatum (L.) Kaulf.
Antrophyum lineatum (Sw.) Kaulf.
Antrophyum ophioglossoides Lellinger
Hecistopteris kaieteurensis Kelloff & G.S. McKee
Hecistopteris pumila (Spreng.) J. Sm.
Vittaria gardneriana Fee
Vittaria graminifolia Kaulf.
Vittaria lineata (L.) Sm.
Vittaria moritziana Mett.
Vittaria remota Fee
Vittaria stipitata Kunze
Synonyms and Misidentifications
Gymnogramme pumila Spreng. =Hecistopteris pumila
Polytaenium brasilianum (Desv.) Benedict [records due to misidentification fide Lellinger]
Polytaenium cajenense (Desv.) Benedict =Antrophyum cajenense
Polytaenium guayanense (Hieron.) Alston =Antrophyum guayanense
Polytaenium lineatum (Sw.) J. Sm. =Antrophyum lineatum
Pteridanetium citrifolium (L.) Copel. =Anetium citrifolium
Vittaria angustifolia (Sw.) Baker =Ananthacorus angustifolius
Vittaria costata Kunze =Ananthacorus angustifolius
Vittaria filifolia Fee = Vittaria graminifolia
References
Lellinger, D.B. 1972. Polypodiaceae subfam. Vittarioideae. In B. Maguire & colaborators. Botany of the Guayana Highland part
9. Mem. New York Bot. Gard. 23: 1-832.
Smith, A.R. 1995. Vittariaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 327-334.
WOODSIACEAE
Diplaz
Diplaz
Diplaz
Diplaz
Diplaz
Diplaz
Diplaz
Diplaz
Diplaz
Diplaz
Diplaz
Diplaz,
Diplaz,
um celtidifolium Kunze
um centripetale (Baker) Maxon [Mt. Roraima]
um cristatum (Desr.) Alston
um expansum Willd.
um gracilescens (Mett.) C. Chr.
um grandifolium (Sw.) Sw.
um hians Kunze ex Klotzsch
um lechleri (Mett.) T. Moore
um plantaginifolium (L.) Urb.
um radicans Desv.
um riedelianum Kunze
um roraimense Cremers & K.U. Kramer [Mt. Roraima]
um striatum (L.) C. Pre si
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
SU
SU
SU
SU
SU
FG
FG
FG
FG
FG
FG
FG
56 Checklist of the Plants of the Guiana Shield
Diplazium tabaloseme Hieron. AM
Diplazhim wilsonii (Baker) Diels BO
Hemidictyiim marginatum (L.) C. Presl AM SU FG
Synonyms and Misidentifications
Diplazium arboreum (Willd.) C. Presl ^Diplazium cristatum
Diplazium calliptehs Fee =Diplazium celtidifolium
Diplazium delitescem Maxon (Aspleniaceae) =Asplenium delitescem
Diplazium shepherdii (Spreng.) Link =Diplazium cristatum
References
(Included in Dryopteridaceae in Flora of the Venezuelan Guayana and in Flora of the Guyanas)
Cremers, G, K.U. Kramer, R.C. Moran, & A.R. Smith. 1994. Dryopteridaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas
(series B) 6: 3-65.
Smith, A.R., J.T. Mickel, & R.C. Moran. 1995. Dryopteridaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the
Venezuelan Guayana 2: 73-128.
GYMNOSPERMS
D.J. de Laubenfels, D.W. Stevenson
CONIFEROPHYTA
PiNACEAE
Pinus caribaea Morelet [cultivated, naturalized] GU SU FG
References
Stevenson, D. & T. Zanoni. 1991. Pinaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 9: 19-22.
PODOCARPACEAE
Podocarpus acuminatus de Laub.
Podocarpus aracensis de Laub. & Silba
Podocarpus brasiliensis de Laub. [expected in GU]
Podocarpus buchholzii de Laub. [expected in GU]
Podocarpus celatus de Laub.
Podocarpus magnifolius J. Buchholz & N.E. Gray [expected in GU]
Podocarpus roraimae Pilg. [Mt. Roraima]
Podocarpus steyermarkii J. Buchholz & N.E. Gray
Podocarpus tepuiensis J. Buchholz & N.E. Gray
References
Laubenfels, D.J. de. 1982. Podocarpaceae. In Flora de Venezuela 11(2): 7-41.
Laubenfels, D.J. de. 1985. Ataxonomic revision of the genus Podocarpus. Blumea 30: 251-278.
Laubenfels, D.J. de. 2004. Podocarpaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 8: 297-300.
Stevenson, D. 1991. Podocarpaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 9: 23-30.
CYCADOPHYTA
Cycadaceae
Cycas spp. [species of this Old World family are widely cultivated]
References
Stevenson, D.W. 1991. Cycadaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 9: 3-7.
Zamiaceae
Zamia amazonum D.W. Stev. AM
Zamia lecointei Ducke [expected in the Guianas] AM BO
Zamia muricata Willd. [expected in the Guianas] BO
Zamia ulei Dammer [expected in the Guianas]
Synonyms and Misidentifications
Zamia jirijirmensis R.E. Schult. =Zamia lecointei
Zamia obidensis Ducke =Zamia lecointei
Zamia ulei subsp. lecointei (Ducke) Ducke =Zamia lecointei
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO GU?
AM BO GU
AM BO GU
GNETOPHYTA
Gnetaceae
T. Zanoni BO GU
AM BO GU
AM BO GU su FG
AM BO GU FG
AM GU
AM BO GU su FG
Checklist of the Plants of the Guiana Shield 57
References
Jones, D.L. 1993. Cycads of the World. Washington, DC: Smithsonian Institution Press.
Sabato, S. 1990. West Indian and South American Cycads. Mem. New York Bot. Card. 57: 173-185.
Stevenson, D. 1991. Zamiaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 9: 7-11.
Gnetum camporum (Markgr.) D.W. Stev.
.
Gnetum Jeyboldii Tul.
Gnetum nodiflorum Brongn.
Gnetum paniculatum Spruce ex Benth.
Gnetum schwackeanum Taub. ex Markgr.
Gnetum urens (Aubl.) Blume
Synonyms and Misidentifications
Gnetum amazonicum Tul. =Gnetum nodiflorum
Gnetum cruzianum Gleason =Gnetum nodiflorum
Gnetum leyboldii var. woodsonianum Markgr. =Gnetiim urens
Gnetum melinonii Benoist =Gnetum urens
Gnetum microstachyum Benth. ex Tul. =Gnetum paniculatum
Gnetum oblongifolium Huber =Gnetum nodiflorum
Gnetum paraense Huber =Gnetum leyboldii
Gnetum thoa Brongn. [nom. illeg.] =Gnetum urens
Gnetum urens var. camporum Markgr. =Gnetum camporum
Thoa urens Aubl. =Gnetum urens
References
Markgraf, F. 1965. New discoveries of Gnetum in tropical America. Ann. Missouri Bot. Gard. 52: 379-386.
Stevenson, D.W. 1997. Gnetaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 162
Stevenson, D.W. 1999. Gnetaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan Guayana
5: 573-576.
Stevenson, D. & T. Zanoni. 1991. Gnetaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 9: 12-18.
Incertae sedis
Aubletella macoucou (Aubl.) Pierre [excluded from Sapotaceae. FG
Chrysophyllum macoucou Aubl.?]
Macahanea guianensis Aubl. [excluded from Hippocrateaceae. Salacia FG
guianensis (Aubl.) Lemee?]
Managa guianensis Aubl. [excluded from Hippocrateaceae] FG
Sarcaulus melinonianum Pierre [name of unknown origin, placed by Lemee in FG
Myrsinaceae]
Senapea guianensis Aubl. [excluded from Passifloraceae] FG
Tampoa guianensis Aubl. [excluded from Hippocrateaceae] FG
Vouapa simira Aubl. [excluded from Leguminosae-Caesalp. Macrolobium FG
simira (Aubl.) J.F. Gmelin?]
ANTHOPHYTA
General references
Cremers, G. 1985. Especes nouvelles de Guyane fran^aise: richesse de I'herbier de Cayenne (CAY). Proc. Kon. Ned. Akad.
Wet. ser.C, 88(1): 15-37.
Cremers, G, C. Feuillet, J.J. de Granville, & M. Hoff. 1988. Liste des especes de Phanerogames et de Pteridophytes de Guyane
fran9aise d'apres I'Herbier du Centre ORSTOM de Cayenne. ORSTOM, Cayenne.
Cremers, G, J.J. de Granville, & M. Hoff. 1994. Les plantes "endemiques" de Guyane fran^aise. Mus. Nat. Hist. Nat., Coll.
Patr. Nat. (Ser. Patr. Genetique) 18: 1-50.
Cremers, G & M. Hoff. 1994. Inventaire taxonomique des plantes de la Guyane fran9aise. IV - Les Monocotyledones. Mus.
Nat. Hist. Nat., Coll. Patr. Nat. (Ser. Patr. Genetique) 17: 1-252.
DeFilipps, R.A. 1992. Ornamental Garden plants of the Guianas: An Historical Perspective of Selected Garden Plants from
Guyana, Surinam and French Guiana. 363 pp. Washington, D.C.: Smithsonian Institution.
Fanshawe, D.B. 1952. The vegetation of British Guiana: A preliminary review. Imperial Forestry Institute paper no. 29: 1-96.
58 Checklist of the Plants of the Guiana Shield
Funk, V.A. & C.L. Kelloff. 1991. A Preliminary Checklist of the Flowering Plants and Ferns of Kaieteur Falls National Park
Guyana. Unpublished manuscript. Biological Diversity of the Guianas program, Smithsonian Institution, Washington, DC.
Gleason, H.A. 1929. Studies on the flora of northern South America - XIII. The Tate collection from Mount Roraima and
vicinity. Bull. Torr. Bot. Club 56(8): 391-408.
Graham, E.H. 1934. Flora of the Kartabo Region, British Guiana. Reprinted from Ann. Cam. Mus. 22(1) (1933).
Howard, R.A. 1983. The plates of Aublet's Histoire des Plantes de la Guiane Fran9oise. J. Arn. Arb. 64: 255-292.
Humboldt, A. von, A. Bonpland, and K.S. Kunth. 1815-1825. Nova Genera et Species Plantarum. 7 volumes. Paris: Sumtibus
Librariae Graeco-Latino-Germanicae
Jansen-Jacobs, M.J. & B.J.H. ter Welle. 1989. Botanical explorations in Guyana, 1985. Annex: identifications. Institute of
Systematic Botany, Utrecht.
Jansen-Jacobs, M.J. & B.J.H. ter Welle. 1990. Botanical explorations in the northern part of the western Kanuku Mountains
(Guyana). Annex: identifications. Institute of Systematic Botany, Utrecht.
Jansen-Jacobs, M.J. & B.J.H. ter Welle. 1992. Botanical exploration in the Wai-Wai area of southern Guyana. Annex:
identifications. Herbarium Division, Department of Plant Ecology and Evolutionary Biology, University of Utrecht.
Judd, W.S., C.S. Campbell, E.A. Kellogg, RF. Stevens, and M.J. Donoghue. 2002. Plant Systematics: A Phylogenetic Approach.
Sinauer Associates, Inc., Sunderland, MA-
Lemee, A. 1953-1956. Flore de la Guyane Fran9aise. Lechevalier, Paris.
Lindeman, J.C. 1953. The Vegetation of the Coastal Region of Suriname. Drukkerij en Uitg.-Mij. v/h Kemink, Utrecht.
Maas, P.J.M. & L.Y.Th. Westra. 1990. Botanical exploration in the northern part of the Rupununi Savanna and in the Mabura Hill
area (Guyana). Appendix: identifications. Institute of Systematic Botany, Utrecht.
Maguire, B. & collaborators. 1948. Plant explorations in Guiana in 1944, chiefly to the Tafelberg and the Kaieteur Plateau. Bull.
Torrey Bot. Club 75: 56-115, 189-230, 286-323, 374-438, 523-580, 633-671.
Mennega, E.A., W.C.M. Tammens-de Rooij, & M.J. Jansen-Jacobs. 1988. Check-List ofWoody Plants ofGuyana. The Tropenbos
Foundation, Ede, The Netherlands.
Mori, S.A. & collaborators. 1987. The Lecythidaceae of a lowland neotropical forest: La Fumee Mountain, French Guiana.
Mem. New York Bot. Gard. 44: 1-190.
Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular
plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2). 776 pp.
Omawale. 1973. Guyana's Edible Plants. University of Guyana.
Plotkin, M.J., B.M. Boom, & M. Allison. 1991. The ethnobotany of Aublet's Histoire des Plantes de la Guiane Fran9oise
(1775). Monogr. Syst. Bot. 35: 1-108.
Polak, A.M. 1992. Major Timber Trees of Guyana, a Field Guide. The Tropenbos Foundation, Wageningen.
Pulle, A. 1906. An Enumeration of the Vascular Plants Known from Surinam, Together with their Distribution and Synonymy.
E.J. Brill, Ltd., London.
Roosmalen, M.G.M. van. 1985. Fruits of the Guianan Flora. Institute of Systematic Botany, Utrecht University, Utrecht;
Silvicultural Department of Wageningen Agricultural University, Wageningen.
Smith, N., S.A. Mori, A. Henderson, D.W. Stevenson, & S.V. Heald. Flowering Plants of the Neotropics. Princeton University
Press, Princeton. 616 pp.
Steege, H. ter, M.J. Jansen-Jacobs, & M. Polak. 1993. Checklist ofplants collected around Mabura Hill, Guyana. The Tropenbos
Foundation, Wageningen and Department of Plant Ecology and Evolutionary Biology, Utrecht University.
Steyermark, J.A., RE. Berry, & B.K. Hoist. 1995-2005. Flora of the Venezuelan Guayana. Missouri Botanical Garden, St.
Louis, & Timber Press, Portland, Oregon.
MONOCOTYLEDONEA
Agavaceae
RE. Berry
Furcraea cabuya Trel. BO
Furcraeafoetida (L.) Haw. GU SU FG
Synonyms and Misidentifications
Cordyline micrantha Baker (Bromeliaceae) =Brocchinia micrantha
Furcraea gigantea Vent. =Furcraeafoetida
References
Berry, RE. 1995. Agavaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 374-375.
Bogler, D.J. and B.B. Simpson. 1995. A chloroplast DNA study of the Agavaceae. Syst. Bot. 20 (2): 191-205.
Bos, J.J. 1984. Dracaena in West Africa. 126 pp. Wageningen, The Netherlands: Agricultural University Wageningen.
Boterenbrood, M.J.A. 1943. Amaryllidaceae. In: Pulle, A., Flora of Suriname 1(1): 443-454.
Chahinian, B.J. 1986. The Sansevieria trifasciata varieties. 109 pp. Reseda, California: Trans Terra Publishing.
Fosberg, F.R. 1985. Cordyline fruticosa (L.) Chevalier (Agavaceae). Baileya 22 (4): 180-181.
Gentry, H.S. 1982. Agaves of Continental North America. 670 pp. Tucson, Arizona: University ofArizona Press.
Grenand, P., Moretti, C. and H. Jacquemin. 1987. Pharmacopees Traditionnelles en Guyane: Creoles, Palikur, Wayapi. 569 pp.
GU
GU
GU su FG
GU su FG?
GU
GU su FG?
GU su?
su
GU su
GU
GU su
GU
GU su FG
GU su FG
GU su FG
GU su FG
GU su FG?
Checklist of the Plants of the Guiana Shield 59
Paris: ORSTOM.
Howard, R.A. 1979. Agavaceae. Flora of the Lesser Antilles 3: 485-504.
Morgenstem, K.D. 1979. Sansevierias. 112 pp. Kempten, Germany: Illertaler Offsetdruck.
Ostendorf, F.W. 1962. Nuttige Planten en Sierplanten in Suriname. 325 pp. Landbouwproefstation in Suriname Bulletin No. 79.
SECAB. 1989. Agave americana. Especies Vegetales Promisorias 1: 9-37.
Alismataceae
R.R. Haynes, L.B. Holm-Nielsen
Echinodorus berteroi (Spreng.) Fassett
Echinodorus bolivianus (Rusby) Holm-Niels. BO
Echinodorus grisebachii Small BO
Echinodorus horizontalis Rataj DA
Echinodorus macrophyllus (Kunth) Micheli
Echinodorus macrophyllus subsp. macrophyllus
Echinodorus macrophyllus subsp. scaber (Rataj) Haynes & Holm-Niels.
Echinodorus paniculatus Micheli BO
Echinodorus reticulatus Haynes & Holm-Niels.
Echinodorus subalatus (Mart.) Griseb.
Echinodorus subalatus subsp. andrieuxii (Hook. & Am.) Haynes & Holm-
Niels.
Echinodorus subalatus subsp. subalatus
Echinodorus tenellus (Mart.) Buchenau AM
Sagittaria guayanensis Kunth BO
Sagittaria guayanensis subsp. guayanensis BO
Sagittaria lancifolia L.
Sagittaria lancifolia subsp. lancifolia
Sagittaria rhombifolia Cham. BO
Sagittaria sprucei Micheli DA
Synonyms and Misidentifications
Alisma boliviana Rusby =Echinodorus bolivianus
Alisma tenellum Mart. =Echinodorus tenellus
Alisma tenellum f. latifolia Seub. =Echinodorus bolivianus
Echinodorus andrieuxii (Hook. & Am.) Small ^Echinodorus subalatus subsp. andrieuxii
Echinodorus gracilis Rataj =Echinodorus grisebachii
Echinodorus grandiflorus (Cham. & Schltdl.) Micheli [not in flora area]
Echinodorus guianensis (Kunth) Griseb. =Sagittaria guayanensis
Echinodorus intermedius Griseb. {=E. grisebachii, fide Lindeman]
Echinodorus latifolius (Seub.) Rataj =Echinodorus bolivianus
Echinodorus scaber Rataj =Echinodorus macrophyllus subsp. scaber
Lophotocarpus guayanensis (Kunth) J.G. Sm. =Sagittaria guayanensis
Sagittaria pugioniformis L. =Sagittaria lancifolia
References
Bogin, C. 1955. Revision of the genus Sagittaria. Mem. New York Bot. Gard. 9: 179-233.
Fassett, N.C. 1955. Echinodoms in the American tropics. Rhodora 57: 133-212.
Haynes, R.R. & L.B. Holm-Nielsen. 1986. Notes on Echinodorus. Brittonia 38: 325-332.
Haynes, R.R. & L.B. Holm-Nielsen. 1994. Alismataceae. Flora Neotropica Monograph 64: 1-112.
Haynes, R.R. & L.B. Holm-Nielsen. 1995. Alismataceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan
Guayana2: 377-383.
Jonker, F.P 1943. Alismataceae. In A. Pulle, Flora of Suriname 1(1): 472-482.
Jonker, F.P. 1968. Alismataceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 378-379.
Rataj, K. 1972. Revision of the genus Sagittaria. Part II. Annot. Zool. Bot. Bratislava 78: 1-61.
Rataj, K. 1975. Revizion (sic) of the genus Echinodoms Rich. Studie CSAV (Praha) 2: 1-156.
Araceae
G.S. Bunting, T. Croat, D. Nicolson
Alocasia macrorrhiza (L.) G. Don [cultivated, escaped] SU
Anaphyllopsis americana (Engl.) A. Hay SU FG
Anaphyllopsis pinnata A . Hay AM
Anthurium acrobates Sodiro AM
Anthurium apaporanum R.E. Schult. AM
Anthurium atropurpureum R.E. Schult. & Maguire AM?
60 Checklist of the Plants of the Guiana Shield
Anthurium bakeri Hook. f.
Anthurium bonplandii G.S. Bunting
Anthurium bonplandii subsp. bonplandii
Anthurium bonplandii subsp. guayanum (G.S. Bunting) Croat
Anthurium bonplandii subsp. rionegrense G.S. Bunting
Anthurium cataniapoense Croat
Anthurium clavigerum Poepp.
Anthurium corocoroense GS. Bunting
Anthurium cowanii Croat
Anthurium cremersii Croat, sp. nov. ined.
Anthurium digitatum (Jacq.) G Don
Anthurium eminens Schott
Anthurium expansum Gleason
Anthurium friedrichsthalii Schott
Anthurium glaucophyllum Gleason
Anthurium gracile (Rudge) Schott
Anthurium guaiquinimae GS. Bunting
Anthurium guanchezii GS. Bunting
Anthurium guayanum G.S. Bunting
Anthurium hookeri Kunth
Anthurium iramirezae GS. Bunting
Anthuriumjenmanii Engl.
Anthurium kunthii Poepp.
Anthurium lanjouwii A.M.E. Jonker & Jonker
Anthurium maguirei A.D. Hawkes
Anthurium martianum K. Koch & Kolb
Anthurium moonenii Croat & E.GGon^.
Anthurium nymphaeifolium K. Koch & Bouche
Anthurium obtusum (Engl.) Grayum
Anthurium pentaphyllum (Aubl.) G Don
Anthurium ptarianum Steyerm.
Anthurium roraimense N.E. Br.
Anthurium rubrinervium (Link) G Don
Anthurium scandens (Aubl.) Engl.
Anthurium sinuatum Benth. ex Schott
Anthurium subscriptum GS. Bunting AM
Anthurium tatei G.S. Bunting AM
Anthurium thrinax Madison GU FG
Anthurium triphyllum Brongn. ex Schott AM
Anthurium vinillense G.S. Bunting AM
Anthurium wurdackii G.S. Bunting AM
Anthurium xanthoneurum G.S. Bunting AM
Anthurium yutajense G.S. Bunting AM
Caladium bicolor (Aiton) Vent. AM BO
Caladium humboldtii Schott [cultivated in Guyana] AM
Caladium macrotites Schott AM
Caladium picturatum K.Koch & Bouche [=Caladium bicolor?] AM BO DA
Caladium schomburgkii Schott BO
Colocasia esculenta (L.) Schott [cultivated, naturalized] BO DA
Cyrtosperma spruceanum (Schott) Engl.
Dieffenbachia duidae (Steyerm.) GS. Bunting AM BO
Dieffenbachia elegans A.M.E. Jonker & Jonker
Dieffenbachia humilis Poepp.
Dieffenbachiajenmanii Veitch ex Kegel
Dieffenbachia paludicola N.E. Br. ex Gleason AM
Dieffenbachia parvifolia Engl. AM
Dieffenbachia seguine (Jacq.) Schott AM BO DA
Dieffenbachia seguine van lingulata (Mart, ex Schott) Engl.
Dieffenbachia seguine var. seguine
Dieffenbachia seguine var. venteniana (Schott) Engl.
Dieffenbachia seguine var. viridis Engl.
Dieffenbachia sp. nov. 1
Dieffenbachia sp. nov. 2 GU
Dracontium asperum K. Koch [expected in GU] BO SU FG
AM GU
AM BO GU SU FG
AM BO GU FG
GU SU FG
AM
AM SU
AM BO GU SU FG
AM
GU SU
FG
BO DA FG
AM GU SU FG
AM BO GU
GU?
GU
SU FG
AM BO
BO
DA GU SU FG
AM
BO
BO GU SU FG
AM
BO GU SU FG
AM GU?
GU
GU
GU
SU
SU
SU
FG
FG
AM BO DA GU SU FG
GU SU FG
AM BO GU
AM BO GU
GU SU FG
AM BO DA GU SU FG
GU SU FG
GU SU FG
GU
GU FG
GU SU FG
GU SU
SU
FG
GU
GU SU FG
GU SU FG
GU
GU SU FG
SU FG
GU SU
SU
FG
GU SU
SU
FG
FG
SU FG
AM BO GU
GU su
su FG
AM BO GU su FG
AM BO GU su
AM BO
GU?
BO GU
GU
su FG
AM BO GU FG
AM
AM
BO
BO
FG
AM GU? FG
AM BO GU su FG
GU su FG
BO GU
AM BO DA GU su FG
AM BO GU su FG
AM BO DA GU su FG
AM BO GU su? FG?
AM
AM BO
AM BO DA GU su FG
AM BO GU su FG
AM BO DA GU su FG
AM? BO DA? GU su FG
AM BO DA GU su FG
AM BO
Checklist of the Plants of the Guiana Shield 61
Dracontium dubium Kunth
Dracontiumfoecundum Hook. f.
Dracontium polyphyllum L.
Heteropsis flexuosa (Kunth) G.S. Bunting
Heteropsis flexuosa wax. flexuosa
Heteropsis flexuosa var. maguirei G.S. Bunting
Heteropsis longispathacea Engl.
Heteropsis melinonii (Engl.) A.M.E. Jonker & Jonker
Heteropsis oblongata (Kunth) Croat, comb. nov. ined.
Heteropsis spruceana Schott
Heteropsis spruceana var. robusta GS. Bunting
Heteropsis spruceana var. spruceana
Heteropsis steyermarkii G.S. Bunting
Heteropsis tenuispadix G.S. Bunting
Homalomena picturata (Linden & Andre) Regel [cultivated only?]
Jasarum steyermarkii G.S. Bunting
Monstera adansonii Schott
Monstera adansonii var. klotzschiana (Schott) Madison
Monstera adansonii var. laniata (Schott) Madison
Monstera dubia (Kunth) Engl. & K. Krause
Monstera gracilis Engl.
Monstera lechleriana Schott
Monstera obliqua Miq.
Monstera spruceana (Schott) Engl.
Montrichardia arborescens (L.) Schott
Montrichardia linifera (Arruda) Schott
Philodendron acutatum Schott
Philodendron anaadu G.S. Bunting
Philodendron asplundii Croat FG
Philodendron atabapoense G.S. Bunting AM
Philodendron auyantepuiense G.S. Bunting BO GU
Philodendron barrosoanum G.S. Bunting AM
Philodendron billietiae Croat
Philodendron bipennifolium Schott [expected in GU]
Philodendron brevispathum Schott
Philodendron brevispathum subsp. holmquistii (G.S. Bunting) G.S. Bunting
Philodendron callosum K. Krause
Philodendron callosum subsp. callosum
Philodendron callosum suhsp. ptarianum (Steyerm.) G.S. Bunting
Philodendron camposportoanum G.M. Barroso
Philodendron canaimae G.S. Bunting BO
Philodendron cataniapoense GS. Bunting AM
Philodendron chimantae GS. Bunting BO
Philodendron conforme G.S. Bunting AM
Philodendron cremersii Croat & Grayum
Philodendron deflexum Poepp. ex Schott AM BO
Philodendron delascioi G.S. Bunting
Philodendron dioscoreoides Gleason
Philodendron duckei Croat & Grayum
Philodendron dunstervilleorum GS. Bunting BO
Philodendron dyscarpium R.E. Schult. AM
Philodendron dyscarpium var. dyscarpium AM
Philodendron dyscarpium var. ventuarianum G.S. Bunting AM
Philodendron ecordatum Schott
Philodendron englerianum Steyerm. AM BO
Philodendron englerianum subsp. duidae (Steyerm.) GS. Bunting AM
Philodendron englerianum subsp. englerianum BO
Philodendron exile G.S. Bunting AM
Philodendron fendleri K. Krause
Philodendron fragrantissimum (Hook.) G Don AM BO
Philodendron goeldii G.M. Barroso AM
Philodendron grandifolium (Jacq.) Schott
Philodendron guaiquinimae GS. Bunting BO
Philodendron guianense Croat & Grayum GU SU FG
GU FG
AM BO SU FG
AM BO GU SU FG
AM BO
AM BO GU SU FG
BO GU SU FG
AM BO GU
GU SU FG
SU? FG
DA GU SU FG
DA
GU SU
GU SU FG
GU SU FG
GU SU
SU FG
DA
GU SU FG
su FG
DA GU su FG
AM FG?
AM BO SU FG?
AM
FG
AM BO DA GU SU FG
AM
A A/fAM
BO GU SU
SU
FG
AM BO
BO
DA GU SU FG
AM
AM BO DA GU SU FG
AM BO DA
62 Checklist of the Plants of the Guiana Shield
Philodendron guttiferum Kunth
Philodendron hederaceim (Jacq.) Schott [expected in GU]
Philodendron holstii G.S. Bunting
Philodendron hoiilletianum Engl.
Philodendron hylaeae G.S. Bunting
Philodendron inaequilaterum Liebm.
Philodendron inaequilaterum subsp. anthoblastum GS. Bunting
Philodendron insigne Schott
Philodendronjonkerianum Croat
Philodendron krauseanum Steyerm. [possibly not distinct from P. platypodum
Gleason]
Philodendron lemae G.S. Bunting
Philodendron liesneri G.S. Bunting
Philodendron linnaei Kunth
Philodendron linnaei var. linnaei
Philodendron linnaei var. rionegrense G.S. Bunting AM
Philodendron macropodum K. Krause AM BO FG
Philodendron maguirei GS. Bunting AM
Philodendron marahuacae G.S. Bunting AM
Philodendron maroae GS. Bunting AM
Philodendron mawarinumae G.S. Bunting AM
Philodendron melinonii Brongn. ex Regel AM
Philodendron multinerve GS. Bunting AM
Philodendron muricatum Willd. ex Schott AM
Philodendron nebulense GS. Bunting AM
Philodendron ornatum Schott AM
Philodendron panduriforme (Kunth) Kunth AM
Philodendron panduriforme var. panduriforme AM
Philodendron panduriforme var. reichenbachianum (Schott) Croat AM
Philodendron pedatum (Hook.) Kunth AM
Philodendron peperomioides GS. Bunting
Philodendron peraiense G.S. Bunting
Philodendron phlebodes G.S. Bunting AM
Philodendron phlebodes var. kermesinum GS. Bunting AM
Philodendron phlebodes var. phlebodes AM
Philodendron pimichinense G.S. Bunting AM
Philodendron placidum Schott
Philodendron platypodum Gleason
Philodendron quercifolium Engl.
Philodendron remifolium R.E. Schult. AM
Philodendron remifolium subsp. sabulosum (G.S. Bunting) G.S. Bunting AM
Philodendron roraimae K. Krause AM
Philodendron roraimae subsp. aracamuniense G.S. Bunting AM
Philodendron roraimae subsp. roraimae AM
Philodendron rudgeanum Schott
Philodendron samayense G.S. Bunting
Philodendron scandens K. Koch & Sello AM
Philodendron scitulum GS. Bunting AM
Philodendron simsii Sweet
Philodendron simulans G.S. Bunting AM
Philodendron solimoesense A.C. Sm. AM
Philodendron sphalerum Schott
Philodendron splitgerberi Schott
Philodendron spruceanum G.S. Bunting AM
Philodendron squamiferum Poepp. SU FG
Philodendron steyermarkii GS. Bunting AM BO
Philodendron subsessile Gleason
Philodendron surinamense (Miq.) Engl. AM BO GU SU FG
Philodendron tatei K. Krause AM
Philodendron tatei subsp. melanochlorum (G.S. Bunting) G.S. Bunting AM
Philodendron tatei subsp. tatei AM
Philodendron triplum GS. Bunting BO
Philodendron venezuelense GS. Bunting AM FG
Philodendron venustum G.S. Bunting AM
SU FG
BO DA GU SU FG
BO DA GU SU
BO DA GU
GU
GU
SU FG
BO DA GU SU FG
GU SU FG
GU SU FG
GU
BO GU FG
BO
DA GU SU FG
BO
BO DA GU
GU
SU FG
FG
BO GU SU FG
GU SU
SU
FG
FG
GU
BO GU
BO
BO DA GU su FG
BO
AM
AM
GU
GU
su FG?
FG
AM BO GU su FG
AM GU su
su
FG
FG
AM
A A/f
BO
AM
AM
AM BO DA GU
BO GU su FG?
AM GU su FG
Checklist of the Plants of the Guiana Shield 63
Philodendrort vinaceiim G.S. Bunting BO
Philodendron werkhoveniae Croat SU
Philodendron wittianum Engl. GU FG
Philodendron wurdackii G.S. Bunting
Philodendron yavitense G.S. Bunting
Philodendron yutajense G.S. Bunting
Pistia stratiotes L.
Rhodospatha bolivarana G.S. Bunting
Rhodospatha brachypoda G.S. Bunting
Rhodospatha cardonae G.S. Bunting
Rhodospatha latifolia Poepp.
Rhodospatha oblongata Poepp.
Rhodospatha venosa Gleason
Schismatoglottis americana A.M.E. Jonker & Jonker
Schismatoglottis bolivarana GS. Bunting & Steyerm.
Schismatoglottis spruceana (Schott) G.S. Bunting
Spathiphyllum bariense G.S. Bunting
Spathiphyllum cannifolium (Dryand. ex Sims) Schott
Spathiphyllum ciispidatum Schott
Spathiphyllum humboldtii Schott
Spathiphyllumjejunum G.S. Bunting AM
Spathiphyllum maguirei G.S. Bunting [=Spathiphyllum cuspidatum fide Croat] GU
Spathiphyllum mawarinumae G.S. Bunting AM
Spathiphyllum monachinoi GS. Bunting AM
Spathiphyllum monachinoi var. monachinoi
Spathiphyllum monachinoi var. perangustum G.S. Bunting AM
Spathiphyllum neblinae G.S. Bunting AM
Spathiphyllum schomburgkii Schott
Stenospermation ammiticum G.S. Bunting AM
Stenospermation amomifolium (Poepp.) Schott AM
Stenospermation maguirei A.M.E. Jonker & Jonker
Stenospermation multiovulatum (Engl.) N.E. Br.
Stenospermation nebulense G.S. Bunting AM
Stenospermation spruceamim Schott AM
Stenospermation ulei K. Krause AM
Syngonium hastifolium Engl.
Syngonium podophyllum Schott AM
Syngonium podophyllum var. vellozianum (Schott) Croat AM
Typhonium blumei Nicolson & Sivad. [naturalized]
Urospatha angustiloba Engl. AM
Urospatha sagittifolia (Rudge) Schott AM
Urospatha sagittifolia var. sagittifolia AM
Urospathella wurdackii (G.S. Bunting) G.S. Bunting AM
Xanthosoma acutum E.G. Gong.
Xanthosoma akkermansii (G.S. Bunting) Croat AM
Xanthosoma belophyllum (Willd.) Schott [cultivated?]
Xanthosoma bolivaranum G.S. Bunting
Xanthosoma caracu K. Koch & Bouche [cultivated?]
Xanthosoma caulotuberculatum GS. Bunting
Xanthosoma conspurcatum Schott AM
Xanthosoma contractum G.S. Bunting
Xanthosoma cordatum N.E. Br.
Xanthosoma exiguum GS. Bunting AM
Xanthosoma granvillei Croat & S.A. Thomps.
Xanthosoma helleborifolium (Jacq.) Schott AM
Xanthosoma mafaffoides G.S. Bunting AM
Xanthosoma maroae GS. Bunting AM
Xanthosoma orinocense G.S. Bunting AM
Xanthosoma sagittifolium (L.) Schott [widely cultivated]
Xanthosoma striatipes (K. Koch & Bouche) Madison AM
Xanthosoma striolatum Mart, ex Schott
Xanthosoma trilobum G.S. Bunting AM
Xanthosoma undipes (K. Koch & CD. Bouche) K. Koch AM
Xanthosoma violaceum Schott [widely cultivated]
BO
BO
BO GU
BO GU
GU SU
BO
BO
DA GU
GU
GU
su
su
su
FG
FG
BO DA GU su FG
BO DA GU su
su
FG
BO DA GU su FG
BO DA GU su FG
FG
GU su FG
BO
su?
BO
GU su FG
BO
GU
GU
FG
FG
BO GU su
DA FG
BO GU
GU?
FG
FG
DA GU SU FG
FG
64 Checklist of the Plants of the Guiana Shield
Synonyms and Misidentifications
Alocasia undipes K. Koch & CD. Bouche =Xanthosoma undipes
Anthurium amoenum Kunth [not in flora area]
Anthurium andersonii Schott [not in flora area]
Anthurium angelorum G.S. Bunting =Anthurium bakeri
Anthurium bonplandii subsp. cuatrecasii Croat =Anthurium bonplandii subsp. bonplandii
Anthurium cordatum G. Don [not in flora area]
Anthurium crassinervium (Jacq.) Schott [not in flora area]
Anthurium duidae Steyerm. =Anthurium ptarianum
Anthurium englerianum G.S. Bunting =Anthurium jenmanii
Anthurium galeottii (Hort.) K. Koch [not in flora area]
Anthurium huegelii Schott =Anthurium hookeri
Anthurium idroboanum R.E. Schult. =Anthurium ptarianum
Anthurium igarapeum G.S. Bunting =Anthurium acrobates
Anthurium krauseanum Steyerm. =Anthurium ptarianum
Anthurium liesneri Croat =Anthurium tatei
Anthurium martini Schott =Anthurium sinuatum
Anthurium monsteroides Steyerm. =Anthurium clavigerum
Anthurium neblinae G.S. Bunting =Anthurium triphyllum
Anthurium nigrescens Engl, [not in flora area]
Anthurium obovatum Gleason =Anthiirium clavigerum
Anthurium panduratum Mart, ex Schott =Anthurium clavigerum
Anthurium pentaphyHum var. digitatum (Jacq.) Madison =Anthurium digitatum
Anthurium polyrrhizum K. Koch & Augustin =Anthurium rubrinervium
Anthurium potarense Gleason =Anthurium expansum
Anthurium salvinii Hemsl. [not in flora area]
Anthurium scandens var. trinerve (Miq.) Matuda =Anthurium obtusum
Anthurium scolopendrinum (Ham.) Kunth =Anthurium gracile
Anthurium solitarium Schott [not in flora area]
Anthurium stahelii A.D. Hawkes =Philodendron deflexum
Anthurium steyermarkii GS. Bunting =Anthurium ptarianum
Anthurium surinamense Miq. =Philodendron surinamense
Anthurium trinerve Miq. [nom. illeg.] =Anthurium obtusum
Anthurium trinervium var. obtusum Engl. =Anthurium obtusum
Arum arborescens L. =Montrichardia arborescens
Arum bicolor Alton =Caladium bicolor
Arum esculentum L. =Colocasia esculenta
Arum grandifolium Jacq. =Philodendron grandifolium
Arum hederaceum Jacq. =Philodendron hederaceum
Arum helleborifolium Jacq. =Xanthosoma helleborifolium
Arum liniferum Arruda =Montrichardia linifera
Arum sagittaefolium L. =Xanthosoma sagittifolium
Arum sagittaefolium Rodschied =Urospatha sagittifolia
Arum seguine Jacq. =Dieffenbachia seguine
Arum xanthorrhizon Jacq. =Xanthosoma sagittifolium
Caladium aculeatum G. Mey. =Montrichardia arborescens
Caladium akkermansii GS. Bunting =Xanthosoma akkermansii
Caladium angustifolium Engl. =Xanthosoma striatipes
Caladium angustilobum Engl. =Caladium macrotites
Caladium argyrites Lem. =Caladium humboldtii
Caladium aturense GS. Bunting =Caladium picturatum
Caladium belleyenei hort. Chantin ex Lem. =Caladium picturatum
Caladium fragrantissimum Hook. =Philodendronfragrantissimum
Caladium maculatum Lodd. =Dieffenbachia seguine
Caladium pedatum Hook. =Philodendron pedatum
Caladium striatipes (Kunth & Bouche) Schott =Xanthosoma striatipes
Colocasia antiquorum Schott =Colocasia esculenta
Cyrtosperma americanum Engl. =Anaphyllopsis americana
Cyrtosperma wurdackii GS. Bunting =Urospathella wurdackii
Dieffenbachia bolivarana GS. Bunting =Dieffenbachia duidae
Dieffenbachia liesneri Croat =Dieffenbachia duidae
Dieffenbachia longipistila Croat =Dieffenbachia paludicola
Dieffenbachia maculata (Lodd.) G Don =Dieffenbachia seguine
Dieffenbachia picta Schott =Dieffenbachia seguine
Checklist of the Plants of the Guiana Shield 65
Dieffenbachia spruceana Schott =Dieffenbachia humilis
Dracontium bissectum Veil. (Cyclanthaceae) ^Thoracocarpus bissectus
Dracontium changuango G.S. Bunting =Dracontium dubium
Dracontium laciniatum Veil. =Philodendron pedatum
Dracontium pentaphyllum Aubl. =Anthurium pentaphyllum
Dracontium pertusum L. =Monstera adansonii
Dracontium scandens Aubl. =Anthurium scandens
Dracontium spruceanum (Schott) G.H. Zhu =Cyrtosperma spruceanum
Elopium surinamense (Miq.) Schott =Philodendron surinamense
Heteropsisjenmanii Oliv. =Heteropsis flexuosa
Heteropsis salicifolia Kunth [not in flora area]
Maguirea spathicarpoides A.D. Hawkes =Dieffenbachia paludicola
Monstera acreana K. Krause =Monstera dubia
Monstera amomifolia Poepp. =Stenospermation amomifolium
Monstera cannifolia Schott =Philodendron rudgeanum
Monstera egregia Schott =Monstera lechleriana
Monstera epipremnoides Engl, [not in flora area]
Monstera expilata Schott =Monstera obliqua
Monstera falcifolia Engl. =Monstera obliqua
Monsterafendleri Engl. =Monstera obliqua
Monstera henri-pittieri G.S. Bunting =Monstera lechleriana
Monstera irritans N.W. Simmonds =Monstera dubia
Monstera klotzschiana Schott =Monstera adansonii var. klotzschiana
Monstera maxima Engl. & K. BCrause ^Monstera lechleriana
Monstera milleriana Schott =Monstera adansonii
Monstera pertusa (L.) de Vriese =Monstera adansonii
Monstera sagotiana Engl. =Monstera obliqua
Monstera steyermarkii G.S. Bunting =Monstera spruceana
Monstera xanthospatha Madison [not in flora area]
Montrichardia arborescens var. aculeata (G. Mey.) Engl. =Montrichardia arborescens
Philodendron amplectens A.C. Sm. =Philodendron muricatum
Philodendron asperatum (K. Koch) K. Koch =Philodendron ornatum
Philodendron brevispathum var. wurdackii Croat =Philodendron brevispathum subsp. holmquistii
Philodendron broadwayi N.E. Br. [not in flora area] =Philodendron venosum (Willd.) Croat
Philodendron calophyllum Brongn. ex Linden & Andre =Philodendron insigne
Philodendron cannifolium (Schott) Engl. =Philodendron rudgeanum
Philodendron cuspidatum K. Koch & Bouche =Philodendron scandens
Philodendron cyclops A.D. Hawkes =Philodendron acutatum
Philodendron decurrens K. Krause =Philodendron linnaei
Philodendron demerarae Gleason =Philodendron fragrantissimum
Philodendron deviatum Schott =Philodendron hederaceum
Philodendron duidae Steyerm. "^^Philodendron englerianum subsp. duidae
Philodendron grandipes K. Krause [not in flora area]
Philodendron guttiferum var. rudgeanum (Schott) A.M.E. Jonker & Jonker =Philodendron rudgeanum
Philodendron haematinum R.E. Schult. ^Philodendron insigne
Philodendron hederaceum auct. non (Jacq.) Schott =Philodendron scandens
Philodendron hoffmannii Schott =Philodendron hederaceum
Philodendron holmquistii G.S. Bunting =Philodendron brevispathum subsp. holmquistii
Philodendron jacquinii Schott =Philodendron hederaceum
Philodendron jenmanii K. Krause =Philodendron muricatum
Philodendron laciniatum (Veil.) Engl. =Philodendron pedatum
Philodendron latilobum Schott =Philodendron panduriforme
Philodendron levelii G.S. Bunting =Philodendron barrosoanum
Philodendron longipetiolatum Engl. =Philodendron sphalerum
Philodendron megalophyllum Schott =Philodendron deflexum
Philodendron melanochlorum G.S. Bunting =Philodendron tatei subsp. melanochlorum
Philodendron micans Klotzsch ex K. Koch ^'Philodendron scandens
Philodendron milleri Croat =Philodendron barrosoanum
Philodendron myrmecophilum Engl. =Philodendron deflexum
Philodendron nobile W. Bull. =Philodendron linnaei
Philodendron oxycardium Schott =Philodendron scandens
Philodendron polypodioides A.M.E. Jonker & Jonker =Philodendron pedatum
Philodendron prieurianum Schott =Philodendron scandens
Philodendron ptarianum Steyerm. =Philodendron callosum subsp. ptarianum
66 Checklist of the Plants of the Guiana Shield
Philodendron reichenbachiamim Schott =PhiIodendron pandiiriforme var. reichenbachianum
Philodendron riedelianum Schott =Philodendron siihnamerjse
Philodendron rubens Schott ^Philodendron ornatum
Philodendron sabulosum G.S. Bunting =Philodendron remifolium subsp. sabulosum
Philodendron scabrum K. Krause =Philodendron muricatum
Philodendron smaragdinum G.S. Bunting =Philodendron muricatum
Philodendron striatipes Kunth & Bouche =Xanthosoma striatipes
Philodendron tessmannii K. Krause ^Philodendron guttiferum
Philodendron tobagense Engl. =Philodendron ornatum
Philodendron tripartitum (Jacq.) Schott [not in flora area]
Philodendron wayombense A.M.E. Jonker & Jonker =Philodendron bipennifolium
Philonotion spruceanum Schott =Schismatoglottis spruceana
Philonotion williamsii Steyerm. =Schismatoglottis spruceana
Pothos cannaefolia Dryand. =Spathiphyllum cannifolium
Pothos cannifolius Rudge =Philodendron rudgeamim
Pothos digitata Jacq. =Anthurium digitatum
Pothos flexuosus Kunth =Heteropsis flexuosa
Pothos gracilis Rudge =Anthurium gracile
Pothos panduraeformis Kunth =Philodendron panduriforme
Pothos sagittaefolia Rudge =Urospatha sagittifolia
Pothos scolopendrinus Ham. =Anthurium gracile
Rhodospatha blanda var. melinonii Engl. =Heteropsis melinonii
Rhodospatha melinonii (Engl.) Engl. & K. Krause =Heteropsis melinonii
Rhodospatha pachysperma Madison =Rhodospatha venosa
Rhodospatha spruceana (Schott) J.F. Macbr. =Stenospermation spruceanum
Spathicarpa duidae Steyerm. =Dieffenbachia duidae
Spathiphyllum candolleanum Schott =Spathiphyllum humboldtii
Spathiphyllum caudatum Poepp. =Urospatha sagittifolia
Spathiphyllum floribundum (Linden & Andre) N.E. Br. [not in flora area]
Spathiphyllum glaziovii Engl. "^Spathiphyllum humboldtii
Spathiphyllum huberi Engl. =Spathiphyllum humboldtii
Spathiphyllum lanceifolium (Jacq.) Schott [not in flora area]
Spathiphyllum liesneri Croat =Spathiphyllum humboldtii
Spathiphyllum sipapoanum G.S. Bunting =Spathiphyllum jejunum
Stenospermation ammiticum subsp. neblinae G.S. Bunting =Stenospermation ammiticum
Stenospermation spruceanum var. multiovulatum Engl. =Stenospermation multiovulatum
Stenospermation steyermarkii G.S. Bunting =Stenospermation multiovulatum
Stenospermation verecundum R.E. Schult. =Schismatoglottis spruceana
Syngonium connatum Gleason ^^Philodendron hylaeae
Syngonium ternatum Gleason =Syngonium podophyllum
Syngonium vellozianum Schott =Syngonium podophyllum var. vellozianum
Thaumatophyllum spruceanum Schott =Philodendron goeldii
Tornelia laniata Schott =Monstera adansonii var. laniata
Tornelia spruceana Schott =Monstera spruceana
Typhonium trilobatum (L.) Schott [misidentification of T. blumei]
Urospatha caudata (Poepp.) Schott =Urospatha sagittifolia
Urospatha decipiens Schott =Urospatha sagittifolia
Urospatha dubia Schott ^Urospatha sagittifolia
Urospatha hostmannii Schott =Urospatha sagittifolia
Urospatha sagittifolia var. spruceana (Schott) Engl. =Urospatha sagittifolia
Urospatha savannarum Steyerm. =Urospatha sagittifolia
Urospatha spruceana Schott =Urospatha sagittifolia
Urospatha wurdackii (G.S. Bunting) A. Hay =Urospathella wurdackii
Xanthosoma hoffmannii var. wendlandii (Schott) Engl, [not in flora area]
Xanthosomajacquinii Schott =Xanthosoma undipes p.p. ; X. sagittifolium p.p.
Zantedeschia asperata K. Koch =Philodendron ornatum
References
Bunting, G.S. 1960. A revision of Spathiphyllum. Mem. New York Bot. Gard. 10(3): 1-54.
Bunting, G.S. 1995. Araceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 600-679.
Croat, T.B. 1981. A revision of Syngonium. Ann. Missouri Bot. Gard. 68: 565-651.
Croat, T.B. 1991. A revision of Anthurium section Pachyneurium. Ann. Missouri Bot. Gard. 78(3): 539-855.
Croat, T.B. 1995 (1994). Taxonomic status of Neotropical Araceae. Aroideana 17: 33-60.
Croat, T.B. 1995. Two new species of Araceae for the Guianas. Novon 5(1): 24-29.
Checklist of the Plants of the Guiana Shield 67
Croat, T.B. 1997. Araceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide to
the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs of the
New York Botanical Garden 76, pt. 1: 167-190.
Croat, T.B. & M.H. Grayum. 1994. Philodendron subgenus Pteromischum from Saiil, French Guiana. Novon 4(3): 211-219.
Jonker-Verhoef, A.M.E. & F.R Jonker. 1953. Araceae. In A. Pulle, Flora of Suriname 1(2): 1-80.
Jonker-Verhoef, A.M.E. & F.P. Jonker. 1968. Araceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname
1(2): 380-412.
Madison, M. 1977. A revision of Monstera. Contrib. Gray Herb. 207: 3-100.
Madison, M. 1981. Notes on Caladium and its allies. Selbyana 5(3-4): 342-377.
Zhu, G, J.O. Boos, and T.B. Croat. 1998. Identity and typification of Draconitum dubium (Araceae). Novon 8(1): 101-103.
Arecaceae
M. Balick, J. -J. de Granville, A. Henderson
Acrocomia aciileata (Jacq.) Lodd. ex Mart.
Asterogyne guianensis Granv. & A.J. Hend.
Astrocaryum acaule Mart.
Astrocaryum aculeatum G. Mey.
Astrocaryum chambira Burret
Astrocaryum gynacanthum Mart.
Astrocaryumjauari Mart.
Astrocaryum murumuru Mart.
Astrocaryum paramaca Mart.
Astrocaryum sciophilum (Miq.) Pulle
Astrocaryum vulgare Mart.
Attalea butyracea (Mutis ex L. f ) Wess. Boer
Attalea camopiensis (Glassman) Zona
Attalea degranvillei (Glassman) Zona
Attalea guianensis (Glassman) Zona
Attalea luetzelburgii (Burret) Wess. Boer
Attalea maripa (Aubl.) Mart.
Attalea maripensis (Glassman) Zona
Attalea microcarpa Mart.
Attalea racemosa Spruce
Attalea speciosa Mart.
Bactris acanthocarpa Mart.
Bactris acanthocarpa van acanthocarpa
Bactris acanthocarpa var. intermedia A.J. Hend.
Bactris acanthocarpoides Barb. Rodr.
Bactris aubletiana Trail
Bactris balanophora Spruce
Bactris bidentula Spruce
Bactris brongniartii Mart.
Bactris campestris Poepp. ex Mart.
Bactris constanciae Barb. Rodr.
Bactris corossilla H. Karst.
Bactris cuspidata Mart.
Bactris elegans Barb. Rodr.
Bactris gasipaes Kunth [cultivated, escaped]
Bactris gastoniana Barb. Rodr.
Bactris guineensis (L.) H.E. Moore
Bactris hirta Mart.
Bactris hirta var. hirta
Bactris hirta var. Jenmanii A.J. Hend.
Bactris hirta vox. pulchra (Trail) A.J. Hend.
Bactris major Jacq.
Bactris major var. infesta (Mart.) Drude
Bactris major var. major
Bactris major var. megalocarpa (Trail) A.J. Hend.
Bactris maraja Mart, [see also Bactris brongniartii]
Bactris maraja var. juruensis (Trail) A.J. Hend.
Bactris x moorei Wess. Boer
Bactris oligocarpa Barb. Rodr. & Trail ex Barb. Rodr.
Bactris oligoclada Burret
AM BO GU SU FG
FG
AM BO GU
AM GU SU
AM
AM BO DA GU SU FG
AM BO GU SU FG
AM FG
GU SU FG
GU SU FG
GU SU FG
AM BO FG
FG
FG
FG
AM
AM BO DA GU SU FG
FG
AM GU SU FG
AM
GU SU
AM BO GU SU FG
AM BO GU
GU
SU FG
FG
GU? SU
SU
FG
FG
AM GU
AM
AM BO DA GU SU FG
AM GU SU FG
GU SU FG
AM BO GU?
SU FG
GU SU FG
AM BO
SU
FG
FG
AM BO
GU SU FG
GU SU FG
GU
GU
AM BO DA GU SU
SU
FG
GU SU FG
GU
AM BO
BO
BO
DA GU
GU
SU
SU
FG
FG
FG
68 Checklist of the Plants of the Guiana Shield
Bactris pliniana Granv. & A.J. Hend.
Bactris ptariana Steyerm.
Bactris rhaphidacantha Wess. Boer
Bactris setulosa H. Karst.
Bactris simplicifrons Mart.
Bactris sp. nov. 1 [=Bactris vesiculifera Trail, nomen nudum]
Bactris sp. nov. 2
Bactris sp. nov. 3
Bactris tomentosa Mart.
Bactris tomentosa var. tomentosa
Bactris trailiana Barb. Rodr.
Bactris turbinocarpa Barb. Rodr.
Cocos nucifera L. [cultivated, naturalized]
Copernicia tectonim (Kunth) Mart.
Desmonciis mitis Mart.
Desmonciis cf. mitis Mart.
Desmoncus orthacanthos Mart.
Desmoncus phoenicocarpus Barb. Rodr.
Desmoncus polyacanthos Mart.
Desmoncus polyacanthos var. polyacanthos
Desmoncus cf. pycnacanthos Mart.
Desmoncus cf. riparius Spruce [Desmoncus riparius = Desmoncus
polyacanthos var. polyacanthos]
Dictyocaryumfuscum (H. Karst.) H. Wendl.
Dictyocaryum ptarianum (Steyerm.) H.E. Moore & Steyerm.
Elaeis guineensis Jacq. [cultivated, naturalized]
Elaeis oleifera (Kunth) Cortes
Euterpe catinga Wallace
Euterpe catinga var. catinga
Euterpe catinga var. roraimae (Dammer) A.J. Hend. & Galeano
Euterpe longebracteata Barb. Rodr.
Euterpe oleracea Mart.
Euterpe precatoria Mart.
Euterpe precatoria var. precatoria
Geonoma appuniana Spruce
Geonoma aspidiifolia Spruce
Geonoma baculifera (Poit.) Kunth
Geonoma densa Linden & H. Wendl. ex H. Wendl. [Mt. Roraima]
Geonoma deversa (Poit.) Kunth
Geonoma interrupta (Ruiz & Pav.) Mart.
Geonoma interrupta var. euspatha (Burret) A.J. Hend.
Geonoma interrupta var. interrupta [not in flora area fide A. Henderson]
Geonoma leptospadix Trail
Geonoma macrostachys Mart.
Geonoma macrostachys var. acaulis (Mart.) Skov
Geonoma macrostachys var. poiteauana (Kunth) A.J. Hend.
Geonoma maxima (Poit.) Kunth
Geonoma maxima var. ambigua (Spruce) A.J. Hend.
Geonoma maxima var. chelidonura (Spruce) A.J. Hend.
Geonoma maxima var. maxima
Geonoma oldemanii Granv.
Geonoma oligoclona Trail
Geonoma sp. nov. 1
Geonoma sp. nov. 2
Geonoma stricta (Poit.) Kunth
Geonoma stricta var. piscicauda (Dammer) A.J. Hend.
Geonoma stricta var. stricta
Geonoma stricta var. trailii (Burrett) A.J. Hend.
Geonoma umbraculiformis Wess. Boer
Hyospathe elegans Mart.
Iriartea deltoidea Ruiz & Pav.
Iriartella setigera (Mart.) H. Wendl.
Leopoldinia major Wallace
Leopoldinia piassaba Wallace
GU su FG
BO GU
GU su FG
BO
AM BO GU
GU
su
su
su
FG
FG
FG
FG
FG
FG
AM BO
BO
DA GU su FG
AM
FG
AM BO DA GU su FG
AM GU su FG
AM BO DA GU su FG
GU su
su
FG
FG
GU
AM BO GU
GU su FG
GU su FG
AM BO GU
AM
AM BO GU
AM BO GU
BO DA GU su FG
AM BO DA GU su FG
GU su FG
AM BO GU
AM GU FG
AM BO
BO
GU
GU?
su FG
AM BO DA GU su FG
BO GU su FG
BO GU su
su
FG
BO GU su FG
AM BO GU su FG
AM BO
AM BO GU su FG
AM BO DA GU su FG
BO DA GU su FG
AM
AM BO DA GU su FG
FG
AM
FG
FG
AM GU su
su
FG
FG
AM GU su FG
FG
GU su FG
AM BO GU su FG
AM BO
AM BO GU
AM
AM
AM BO
AM
AM
BO
AM
AM BO
AM
AM BO
AM BO
AM
AM BO
GU
GU
GU SU FG
GU su FG
GU
GU SU
su
FG
GU
GU su FG
GU su FG
GU? su? FG
GU su FG
GU su FG
GU
GU
GU su FG
GU
su FG
Checklist of the Plants of the Guiana Shield 69
Leopoldinia piilchra Mart.
Lepidocaiyum temie Mart.
Lepidocaiyiim tenue var. gracile (Mart.) A.J. Hend.
Manicmia saccifera Gaertn. M. DA
Mauritia carana Wallace
Mauritia flexiiosa L. f. DA
Mauritiella acideata (Kunth) Burret
Mauritiella armata (Mart.) Burret
Maximbignya dahlgreniana (Bondar) Glassman [=AttaIea dahlgreniana]
Nypa fruticans Wurmb. [Asian native, invasive along tidal rivers]
Oenocarpus bacaba Mart. DA
Oenocarpus balickii F. Kahn
Oenocarpus bataua Mart. DA
Oenocarpus bataua var. bataua
Oenocarpus bataua var. oligocarpa (Griseb. & H. Wendl.) A.J. Hend.
Phoenix recUnata Jacq. [cultivated, naturalized]
Prestoea tenuiramosa (Dammer) H.E. Moore AM BO
Roystonea oleracea (Jacq.) O.F. Cook [also widely cultivated, naturalized] BO DA
Sabal mauritiiformis (H. Karst.) Griseb. & H. Wendl. BO
Socratea exorrhiza (Mart.) H. Wendl. AM BO DA
Syagrus cocoides Mart.
Syagrus inajai (Spruce) Becc.
Syagrus orinocensis (Spruce) Burret AM BO
Syagrus sancona H. Karst. BO
Syagrus stratincola Wess. Boer GU SU FG
Synonyms and Misidentifications
Acrocomia lasiospatha Mart. =Acrocomia aculeata
Acrocomia sclerocarpa Mart. =Acrocomia aculeata
Acrocomia wallaceana (Drude) Becc. =Acrocomia aculeata
Asterogyne spicata (H.E. Moore) Wess. Boer [not in flora area]
Astrocaryum caudescens Barb. Rodr. =Astrocaryum aculeatum
Astrocaryum farinosum Barb. Rodr. =Astrocaryum sciophihim
Astrocaryum guianense Splitg. ex Mart. =Astrocaryum vulgare
Astrocaiyum gymnopus Burret =Astrocaryum gy-nacanthum
Astrocaryum humile Wallace =Bactris acanthocarpa
Astrocaryum macrocalyx Burret =Astrocaryum munimuru
Astrocaryum minus Trail =Astrocaryum gynacanthum
Astrocaryum munbaca Mart. =Astrocaryum gynacanthum
Astrocaryum murumurii var. macrocalyx (Burret) A.J. Hend. [not in flora area]
Astrocaryum plicatum Drude =Astrocaryum sciophilum
Astrocaryum rodriguesii Trail =Astrocaryum paramaca
Astrocaryum segregatum Drude ^Astrocaryum vulgare
Astrocaryum tucuma Mart. =Astrocaryum aculeatum
Astrocaryum tucumoides Drude =Astrocaryum vulgare
Attalea attaleoides (Barb. Rodr.) Wess. Boer [non Scheelea attaleoides Karst.] =Scheelea sp.
Attalea cryptanthera Wess. Boer =Attalea maripa
Attalea dahlgreniana (Bondar) Wess. Boer =Maximbignya dahlgreniana
Attalea excelsa Mart, ex Spreng. [not in flora area] =AttaIea phalerata
Attaleaferruginea Burret =Attalea racemosa
Attalea funifera Mart, [not in flora area]
Attalea humboldtiana Spruce =Attalea butyracea
Attalea macrocarpa (H. Karst.) Wess. Boer =Attalea butyracea
Attalea macrolepis (Burret) Wess. Boer =Attalea butyracea
Attalea macropetala (Burret) Wess. Boer =Attalea maripa
Attalea monosperma Barb. Rodr. [not in flora area] =Attalea spectabilis
Attalea polysticha (Burret) Wess. Boer =Attalea microcarpa
Attalea pycnocarpa Wess. Boer =Attalea butyracea
Attalea regia (Mart.) Wess. Boer =Attalea maripa
Attalea sagotii (Trail ex Thum) Wess. Boer =Attalea microcarpa
Attalea spectabilis Mart, [not in flora area]
Bactris armata Barb. Rodr. [not in flora area] =Bactris maraja var. maraja
Bactris capinensis Huber =Bactris tomentosa var tomentosa
Bactris carolensis Spruce =Bactris simplicifrons
70 Checklist of the Plants of the Guiana Shield
Bactris chaetochlamys Burret =Bactris maraja
Bactris chaetospatha Mart. =Bactris maraja
Bactris concinna Mart, [not in flora area]
Bactris cruegeriana Griseb. & H. Wendl. ex Griseb. =Bactris major var. major
Bactris cuspidata var. tenuis (Wallace) Drude =Bactris simplicifrons
Bactris duidae Steyerm. =Bactris corossilla
Bactris exscapa (Barb. Rodr.) Barb. Rodr. =Bactris acanthocarpa var. acanthocarpa
Bactrisfloccosa Spruce =Bactris cuspidata
Bactris gaviona (Trail) Trail ex Drude =Bactris major var. infesta
Bactris geonomoides Drude ^Bactris hirta var hirta
Bactris gymnospatha Burret =Bactris maraja
Bactris humilis (Wallace) Burret =Bactris acanthocarpa var. acanthocarpa
Bactris integrifolia Wallace ^Bactris hirta var hirta
Bactris kamarupa Steyerm. =Bactris maraja
Bactris leptocarpa Trail ex Thum =Bactris campestris
Bactris longicuspis Burret =Bactris maraja^
Bactris hietzelburgii Burret =Bactris simplicifrons
Bactris megalocarpa Trail ex Thum =Bactris major var. megalocarpa
Bactris mitis Mart. =Bactris cuspidata
Bactris monticola Barb. Rodr. =Bactris maraja
Bactris pallidispina Mart. =Bactris brongniartii
Bactris pectinata Mart. =Bactris hirta var hirta
Bactris penicillata Barb. Rodr. =Bactris maraja var juruensis
Bactris rivularis Barb. Rodr. =Bactris brongniartii
Bactris sororpanae Steyerm. =Bactris simplicifrons
Bactris sphaerocarpa Trail [not in flora area] =Bactris tomentosa var. sphaerocarpa
Bactris tenuis Wallace =Bactris simplicifrons
Bactris tomentosa var. sphaerocarpa (Trail) A.J. Hend. [not in flora area]
Bactris trichospatha Trail =Bactris maraja
Bactris turbinata Spruce =Bactris hirta
Bactris ulei Burret =Bactris hirta var. pulchra
Bactris venezuelensis Steyerm. =Bactris corossilla
Chamaedorea gracilis Willd. [name of uncertain application]
Cocos aculeatus Jacq. =Acrocomia aculeata
Cocos butyracea Mutis ex L. f. ^Attalea butyracea
Cocos guineensis L. =Bactris guineensis
Cocos orinocensis Spruce =Syagrus orinocensis
Copernicia cerifera (Arruda) Mart, [cultivated only] =Copernicia prunifera (Miller) Moore
Corypha tectorum Kunth =Copernicia tectorum
Dahlgrenia ptariana Steyerm. =Dictyocaryum ptarianum
Desmoncus apureanus L.H. Bailey =Desmoncus orthacanthos
Desmoncus duidensis Steyerm. =Desmoncus mitis
Desmoncus guianensis Trail [apparently a nomen nudum]
Desmoncus horridus Splitg. ex Mart. =Desmoncus orthacanthos
Desmoncus kaieteurensis L.H. Bailey =Desmoncus phoenicocarpus
Desmoncus macroacanthos Mart, [specimens probably = B. phoenicocarpus according to Henderson] =Desmoncus
polyacanthos var. polyacanthos
Desmoncus maguirei L.H. Bailey =Desmoncus polyacanthos var. polyacanthos
Desmoncus multijugus Steyerm. =Desmoncus orthacanthos
Desmoncus oxyacanthos Mart. =Desmoncus polyacanthos
Desmoncus palustris Trail =Desmoncus orthacanthos
Desmoncus parvulus L.H. Bailey =Desmoncus phoenicocarpus
Desmoncus pumilus Trail =Desmoncus mitis
Desmoncus riparius Spruce ^Desmoncus polyacanthos var. polyacanthos
Desmoncus velezii L.H. Bailey & H.E. Moore =Desmoncus orthacanthos
Euterpe aurantiaca H.E. Moore =Euterpe catinga
Euterpe erubescens H.E. Moore =Euterpe catinga var. roraimae
Euterpe montis-duida Burret =Euterpe catinga
Euterpe ptariana Steyerm. =Euterpe catinga var. roraimae
Euterpe roraimae Dammer =Euterpe catinga var. roraimae
Euterpe stenophylla Trail =Euterpe precatoria var. precatoria
Euterpe subruminata Burret =Euterpe precatoria
Euterpe tenuiramosa Dammer =Prestoea tenuiramosa
Geonoma acaulis Mart. =Geonoma macrostachys var. acaulis
Geonoma acaulis (Poit.) Burret =Geonoma macrostachys var. poiteauana
Checklist of the Plants of the Guiana Shield 71
Geonoma ambigua Spruce =Geonoma maxima var. ambigua
Geonoma arundinacea Mart, [not in flora area]
Geonoma barthia Engel =Geonoma densa
Geonoma bartlettii Dammer ex Burret =Geonoma deversa
Geonoma chaimostachys Burret =Geonoma macrostachys var. poiteaitana
Geonoma chelidoneura Spruce =Geonoma maxima var. chelidomira
Geonoma elegans Mart, [not in flora area]
Geonoma euspatha Burret =Geonoma internipta var. euspatha
Geonoma fusca Wess. Boer =Geonoma aspidiifolia
Geonomajuniana Dammer =Geonoma maxima
Geonoma kariiaiana Steyerm. ^Geonoma interrnpta var. euspatha
Geonoma macrospatha Spruce =Geonoma bacidifera
Geonoma maguirei L.H. Bailey =Geonoma striata var. striata
Geonoma major Burret ='Geonoma deversa
Geonoma miarospatha Spruce =Geonoma deversa
Geonoma midtiflora Mart. =Geonoma maxima var. maxima
Geonoma negrensis Spruce =Geonoma maxima var. maxima
Geonoma panicidigera Mart. =Geonoma deversa
Geonoma pinnatifrons Willd. [not in flora area fide A. Henderson] =Geonoma interrupta var. interrupta
Geonoma piscicaiida Dammer ^Geonoma striata var. pisaicauda
Geonoma platyaaida Drude & Trail ex Drude [not in flora area] =Geonoma rubescens H. Wendl. ex Drude
Geonoma poiteaiiana Kunth =Geonoma maarostaahys var. poiteauana
Geonoma pyanostaahys Mart. =Geonoma striata var. striata
Geonoma robusta Burret =Geonoma maxima var ambigua
Geonoma roraimae Dammer =Geonoma appuniana p.p. ; Geonoma baaulifera p.p.
Geonoma saramaaaana L.H. Bailey =Geonoma leptospadix
Geonoma sahomburgkiana Spruce =Geonoma maxima var. ambigua
Geonoma spruaeana Trail =Geonoma maxima
Geonoma tamandua Trail [not in flora area] =Geonoma maarostaahys var. macrostachys
Geonoma triglochin Burret [not in flora area] [confused with G. umbraauliformis fide Granville]
Guilielma speciosa Mart. =Bactris gasipaes
Gynestum baculiferum Poit. =Geonoma baaulifera
Gynestum deversum Poit. ^Geonoma deversa
Gynestum maximum Poit. =Geonoma maxima
Hyospathe maculata Steyerm. =Hyospathe elegans
Iriartea exorrhiza Mart. =Soaratea exorrhiza
Iriarteafusca (H. Karst.) Drude [not in flora area] =Diatyoaaryumfusaum
Iriartea setigera Mart. =Iriartella setigera
Iriartea ventriaosa Mart. =Iriartea deltoidea
Jessenia bataua (Mart.) Burret =Oenoaarpus bataua
Jessenia bataua subsp. bataua =Oenoaarpus bataua var. bataua
Jessenia bataua subsp. oligoaarpa (Griseb. & H. Wendl.) Balick =Oenoaarpus bataua var. oligoaarpa
Jessenia oligoaarpa Griseb. & H. Wendl. =Oenoaarpus bataua var. oligoaarpa
Lepidoaaryum casiquiarense (Spruce) Drude =Lepidoaaryum tenue
Lepidocaryum graaile Mart. =Lepidoaaryum tenue var. gracile
Lepidoaaryum guainiensis (Spruce) Drude =Lepidocaryum tenue var. graaile
Maniaaria atriaha Burret =Mamaaria saaaifera
Maniaaria plukenetii Griseb. & H. Wendl. =Maniaaria saaaifera
Markleya dahlgreniana Bondar =Maximbignya dahlgreniana
Mauritia aaideata Kunth =Mauritiella aaideata
Mauritia aasiquiarensis Spruce =Lepidocaryum tenue
Mauritiaflexuosa var. venezuelana Steyerm. =Mauritiaflexuosa
Mauritia guainiensis Spruce =Lepidocaryum tenue
Mauritia martiana Spruce =Attalea maripa
Mauritia minor Burret =Mauritiaflexuosa
Mauritia subinermis Spruce =Mauritiella armata
Mauritiella martiana (Spruce) Burret =Mauritiella armata
Maximiliana elegans H. Karst. =Attalea maripa
Maximiliana maaropetala Burret =Attalea maripa
Maximiliana maripa (Aubl.) Drude =Attalea maripa
Maximiliana martiana H. Karst. =Attalea maripa
Maximiliana regia Mart. =Attalea maripa
Oenoaarpus baaaba var. grandis (Burret) Wess. Boer =Oenoaarpus baaaba
Oenocarpus baaaba var. parvus Wess. Boer =Oenoaarpus baliakii
Oenoaarpus grandis Burret =Oenocarpus baaaba
72 Checklist of the Plants of the Guiana Shield
Oenocarpus oligocarpa (Griseb. & H. Wendl.) Wess. Boer =Oenocarpus bataua var. oligocarpa
Orbignya barbosiana Burret =Attalea speciosa
Orbignya luetzelbiirgii Burret =Attalea hietzelbiirgii
Orbignya microcarpa (Mart.) Burret =Attalea microcarpa
Orbignya phalerata Mart. =Attalea speciosa
Orbignya polysticha Burret =Attalea microcarpa
Orbignya sagotii Trail ex Thum =Attalea microcarpa
Orbignya speciosa (Mart.) Barb. Rodr. =Attalea speciosa
Orbignya spectabilis (Mart.) Burret [not in flora area] =Attalea spectabilis
Palma maripa Aubl. =Attalea maripa
Parascheelea luetzelburgii (Burret) Dugand =Attalea luetzelbiirgii
Prestoea steyermarkii H.E. Moore =Prestoea tenuiramosa
Raphia taedigera (Mart.) Mart, [not in flora area]
Roystoneajenmanii (Wright) Burret [cultivated] =Roystonea regia (Kunth) Cook
Roystonea venezuelana L.H. Bailey =Roystonea oleracea
Scheelea butyracea (Mutis ex L. f.) H. Kar§t. ex H. Wendl. =Attalea butyracea
Scheelea camopiensis Glassman =Attalea camopiensis
Scheelea degranvillei Glassman =Attalea degranvillei
Scheelea guianensis Glassman =Attalea guianensis
Scheelea humboldtiana (Spruce) Burret =Attalea butyracea
Scheelea macrocarpa H. Karst. =Attalea butyracea
Scheelea macrolepis Burret =Attalea butyracea
Scheelea maripensis Glassman =Attalea maripensis
Scheelea passargei Burret =Attalea butyracea
Socratea albolineata Steyerm. =Socratea exorrhiza
Socratea gracilis Burret ^Socratea exorrhiza
Socratea microchlamys Burret =Socratea exorrhiza
Trithrinax mauritiaeformis H. Karst. =Sabal mauritiiformis
Yuyba dakamana L.H. Bailey =Bactris simplicifrons
Yuyba essequiboensis L.H. Bailey =Bactris simplicifrons
Yuyba gleasonii L.H. Bailey =Bactris simplicifrons
Yuyba maguirei L.H. Bailey =Bactris simplicifrons
Yuyba stahelii L.H. Bailey ^Bactris simplicifrons
References
Balick, M.J. 1986. Systematics and economic botany of the Oenocarpus-Jessenia (Palmae) complex. Advances in Economic
Botany 3: 1-140. The New York Botanical Garden, New York.
Dahlgren, B.E. 1936. Index ofAmerican Palms. Field Museum of Natural History, Botanical Series, 14: 1-436.
Dransfield, J., D. Johnson, & H. Synge. 1988. The Palms of the New World, A Conservation Census. lUCN: Kew, Richmond,
Surrey.
Glassman, S.F. 1972. A revision of B.E. Dahlgren's Index of American Palms. Phanerogamarum Monographiae Tomus VI: 1-
294.
Glassman, S.F. 1977. Preliminary taxonomic studies in the palm genus Attalea H.B.K. Fieldiana Bot. 38: 31-61.
Glassman, S.F. 1977. Preliminary studies in the palm genus Orbignya Mart. Phytologia 36: 89-1 15.
Glassman, S.F. 1978. Preliminary taxonomic studies in the palm genus Maximiliana Mart. Phytologia 38: 161-172.
Glassman, S.F. 1987. Revisions of the palm genus Syagrus Mart, and other selected genera in the Cocos alliance. 111. Biol.
Monogr. 56: 1-230.
Granville, J. -J. de. 1975. Un nouveau palmier en Guyane Fran^aise: Geonoma oldemanii. Adansonia, ser. 2 14(4): 553-559.
Granville, J. -J. de. 1986. The Palms of the Guianas. Flora of the Guianas Newsletter 3: 36-39.
Granville, J.-J. de. 1989. La distribucion de las palmas en Guyana Francesa. Acta Amaz. 19: 115-138.
Granville, J.-J. de. 1990. Les Palmiers de la Guyane Fran^aise. Bois et Forets des Tropiques 220: 43-54.
Granville, J.-J. de. 1992. The genus Bactris: taxonomy and distribution. Flora of the Guianas Newsletter 9: 6-11.
Granville, J.-J. de. 1997. Arecaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 190-215.
Granville, J.-J. de. 2002. Les Palmiers de Guyane fran9aise. Acta Botanica Gallica, Bulletin de la Societe Botanique de France
149 (3): 275-228.
Granville, J.-J. de & A. Henderson. 1 988. A new species ofAsterogyne (Palmae) from French Guiana. Studies on the Flora of the
Guianas 28. Brittonia 40(1): 76-80.
Granville, J.-J. de & A. Henderson. 1994. Anew species of Bactris (Palmae) from the Amazon region. Studies on the Flora of the
Guianas 78. Brittonia 46(2): 147-150.
Henderson, A. 1990. Arecaceae part I. Introduction and Iriarteinae. Flora Neotropica Monogr. 53: 1-100.
Henderson, A. 1994. The genus Attalea. Palm Journal 114: 49-53.
Henderson, A. 1995. The Palms of the Amazon. Oxford University Press.
Henderson, A. 1997. Arecaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 32-122.
AM BO GU FG
BO DA GU su FG
BO DA GU
GU
su
su
FG
FG
AM
AM BO DA GU
GU
su FG
AM BO su
AM
AM GU
AM
AM FG
Checklist of the Plants of the Guiana Shield 73
Henderson, A, G Galeano, & R. Bemal. 1995. Field Guide to the Palms of the Americas. Princeton University Press.
Kahn, F. & J. -J. de Granville. 1992. Palms in Forest Ecosystems ofAmazonia. Ecological Studies 95: 1-226. Springer-Verlag.
Kahn, F. & B. Millan. 1992. Astrocaryum (Palmae) in Amazonia. A preliminary treatment. Bull. Inst. fr. etudes andines 21(2):
459-531.
Lindeman, J.C. 1984. Palmae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 651-652.
Wessels Boer, J.G. 1965. Palmae. In J. Lanjouw, Flora of Suriname 5(1): 1-172.
Wessels Boer, J.G. 1968. The Geonomoid Palms. N.V. Noord-Hollandsche Uitgevers Maatschappij, Amsterdam.
Wessels Boer, J.G. 1972. Palmae. In B. Maguire & collaborators. Botany of the Guayana Highland part 9. Mem. New York Bot.
Gard.23: 1-832.
Bromeliaceae
E.J. Gouda, J. Grant, B.K. Hoist, H. Luther, M. Spencer
Aechmea angustifolia Poepp. & Endl.
Aechmea aquilega (Salisb.) Griseb.
Aechmea aquilega var. aquilega
Aechmea beeriana L.B. Sm. & M.A. Spencer [=Streptocalyx poeppigii]
Aechmea brassicoides Baker
Aechmea brevicoUis L.B. Sm.
Aechmea bromeliifolia (Rudge) Baker
Aechmea campanulata L.B. Sm.
Aechmea castelnavii Baker [expected in FG, fide Gouda]
Aechmea chantinii (Carriere) Baker
Aechmea contracta (Mart, ex Schult. f.) Baker
Aechmea corymbosa (Mart, ex Schult. f.) Mez
Aechmea egleriana L.B. Sm.
Aechmea egleriana var. egleriana
Aechmea egleriana var. major L.B. Sm. [expected in FG, fide Gouda] AM
Aechmea lanjouwii (L.B. Sm.) L.B. Sm.
Aechmea lingulata (L.) Baker
Aechmea lingulata var. lingulata
Aechmea longifolia (Rudge) L.B. Sm. & M.A. Spencer [=Streptocalyx
longifolius]
Aechmea melinonii Hook.
Aechmea mertensii (G. Mey.) Schult. & Schult. f.
Aechmea nudicaulis (L.) Griseb.
Aechmea nudicaulis var. cuspidata Baker
Aechmea nudicaulis var. nudicaulis
Aechmea pallida L.B. Sm.
Aechmea penduliflora Andre
Aechmea poitaei (Baker) L.B. Sm. & M.A. Spencer [=Streptocalyx poitaei]
Aechmea politii L.B. Sm.
Aechmea rubiginosa Mez [expected in FG]
Aechmea setigera Mart, ex Schult. f.
Aechmea tillandsioides (Mart, ex Schult. f.) Baker
Aechmea tocantina Baker
Ananas ananassoides (Baker) L.B. Sm.
Ananas comosus (L.) Merr. [cultivated, naturalized]
Ananas lucidus Mill.
Ananas nanus (L.B. Sm.) L.B. Sm. [records misident., fide Spencer]
Ananas parguazens is L.A. Camargo & L.B. Sm.
Araeococcus flagellifolius Harms
Araeococcus goeldianus L.B. Sm.
Araeococcus micranthus Brongn.
Ayensua uaipanensis (Maguire) L.B. Sm.
Billbergia brachysiphon L.B. Sm. [expected in FG, fide Gouda]
Billbergia macrolepis L.B. Sm. AM BO GU
Billbergia pyramidalis (Sims) Linden FG
Billbergia rosea hort. ex Beer SU? FG
Billbergia violacea Beer GU SU FG
Brewcaria brocchinioides (L.B. Sm.) B. Hoist AM
Brewcaria duidensis L.B. Sm., Steyerm. & H. Rob. AM
Brewcaria hechtioides (L.B. Sm.) B. Hoist AM
Brewcaria hohenbergioides (L.B. Sm.) B. Hoist AM
SU
DA GU SU FG
DA GU su FG
BO GU SU FG
GU su FG
AM BO DA GU su FG
BO DA
DA
GU
GU
su FG
BO DA GU
GU
su FG
AM BO GU
FG
AM GU FG?
AM BO
AM BO su FG
AM BO DA GU su
AM GU su FG
GU su FG
AM BO DA GU su FG
AM BO
su?
FG
FG?
AM BO GU su FG
AM BO su FG
AM BO
BO
DA GU su FG
74 Checklist of the Plants of the Guiana Shield
Brewcaria marahuacae L.B. Sm., Steyerm. & H. Rob.
Brewcaria reflexa (L.B. Sm.) B. Hoist
Brocchinia acuminata L.B. Sm. [expected in GU]
Brocchinia cowanii L.B. Sm.
Brocchinia delicatula L.B. Sm. [Sierra de la Neblina]
Brocchinia gilmartiniae G.S. Varad.
Brocchinia hechtioides Mez
Brocchinia hitchcockii L.B. Sm.
Brocchinia maguirei L.B. Sm.
Brocchinia melanacra L.B. Sm. [expected in GU]
Brocchinia micrantha (Baker) Mez
Brocchinia paniculata Schult. f.
Brocchinia prismatica L.B. Sm.
Brocchinia reducta Baker
Brocchinia steyermarkii L.B. Sm.
Brocchinia tatei L.B. Sm.
Brocchinia vestita L.B. Sm.
Brocchinia wurdackiana B. Hoist
Bromelia agavifolia Brongn. ex Houllett
Bromelia alta L.B. Sm.
Bromelia chrysantha Jacq. [expected in GU, fide Gouda]
Bromeliafosteriana L.B. Sm.
Bromelia goeldiana L.B. Sm.
Bromelia granvillei L.B. Sm. & Gouda
Bromelia humilis Jacq.
Bromelia morreniana (Regel) Mez [expected in FG, fide Gouda]
Bromelia oleveiriae L.B. Sm. [this species not expected in FG, fide Gouda]
Bromelia pinguin L. [probably cultivated, fide Grant]
Bromelia plumieri (E. Morren) L.B. Sm.
Bromelia rondoniana L.B. Sm. [expected in GU, fide Gouda]
Bromelia serra Griseb.
Bromelia tubulosa L.B. Sm.
Catopsis berteroniana (Schult. & Schult. f.) Mez
Catopsis sessiliflora (Ruiz & Pav.) Mez
Connellia augiistae (M.R. Schomb.) N.E. Br. [Mt. Roraima]
Connellia caricifolia L.B. Sm. [Mt. Roraima]
Connellia nutans L.B. Sm. [expected in GU]
Connellia quelchii N.E. Br. [Mt. Roraima]
Connellia varadarajanii L.B. Sm. & Steyerm.
Disteganthus basi-lateralis Lem.
Disteganthus lateralis (L.B. Sm.) Gouda
Guzmania altsonii L.B. Sm.
Guzmania brasiliensis Ule [possibly expected in GU]
Guzmania calothyrsa Mez
Guzmania lingulata (L.) Mez
Guzmania melinonis Regel
Guzmania monostachia (L.) Rusby ex Mez
Guzmania monostachia var. monostachia
Guzmania nubicola L.B. Sm. [Sierra de la Neblina]
Guzmania patula Mez & Werckle
Guzmania plumieri (Griseb.) Mez
Guzmania retusa L.B. Sm.
Guzmania roezlii (E. Morren) Mez
Guzmania sphaeroidea (Andre) Andre ex Mez
Guzmania squarrosa (Mez & Sodiro) L.B. Sm. & Pittendr.
Guzmania steyermarkii L.B. Sm. [expected in GU]
Guzmania terrestris L.B. Sm. & Steyerm. AM
Hohenbergia stellata Schult. f. [expected in GU, fide Gouda] BO DA
Lindmania arachnoidea (L.B. Sm., Steyerm. & H. Rob.) L.B. Sm. AM
Lindmania argentea L.B. Sm. BO
Lindmania atrorosea (L.B. Sm., Steyerm. &. H. Rob.) L.B. Sm. BO
Lindmania aurea L.B. Sm., Steyerm. & H. Rob. BO
Lindmania brachyphylla L.B. Sm. BO GU
Lindmania candelabriformis B. Hoist AM
AM
AM
AM BO GU?
AM
AM
BO
AM BO GU
AM
AM
AM BO
BO GU
AM
AM
BO
BO
GU
GU
AM BO GU
AM
AM
BO
GU SU
SU
FG
AM BO GU
GU
SU?
SU
FG
FG?
BO DA GU SU FG
FG
AM BO FG?
BO DA GU SU FG
BO DA GU SU FG
BO GU
BO GU?
BO
BO GU
BO
SU
FG
FG
AM BO GU SU FG
AM
GU
AM BO DA GU SU FG
AM GU SU FG
AM DA GU?
AM DA
AM
AM BO
GU? SU? FG
BO GU
BO DA GU
AM
AM
BO
BO
BO
GU
GU
Checklist of the Plants of the Guiana Shield 75
Lindmania cylindrostachya L.B. Sm.
Lindmania dendritica (L.B. Sm.) L.B. Sm. [Sierra de la Neblina]
Lindmania graciUima (L.B. Sm.) L.B. Sm. [expected in GU]
Lindmania guianensis (Beer) Mez
Lindmania guianensis var. vestita (L.B. Sm.) L.B. Sm. [recognized as distinct
in FVG, but appropriate level not ventured]
Lindmania holstii Steyerm. & L.B. Sm.
Lindmania huberi L.B. Sm., Steyerm. & H. Rob.
Lindmania imitans L.B. Sm., Steyerm. & H. Rob.
Lindmania lateralis (L.B. Sm. & Read) L.B. Sm. & H. Rob. [Sierra de la
Neblina]
Lindmania longipes (L.B. Sm.) L.B. Sm.
Lindmania maguirei (L.B. Sm.) L.B. Sm. [Sierra de la Neblina]
Lindmania marahuacae (L.B. Sm., Steyerm. & H. Rob.) L.B. Sm.
Lindmania minor L.B. Sm.
Lindmania navioides L.B. Sm.
Lindmania nubigena (L.B. Sm.) L.B. Sm. [Sierra de la Neblina]
Lindmania oliva-estevae L.B. Sm. & Steyerm. ex B. Hoist
Lindmania phelpsiae L.B. Sm.
Lindmania piresii L.B. Sm., Steyerm. & H. Rob. [Serra Araca, Brazil]
Lindmania riparia L.B. Sm., Steyerm. & H. Rob.
Lindmania saxicola L.B. Sm., Steyerm. & H. Rob.
Lindmania serrulata L.B. Sm.
Lindmania smithiana (Steyerm. & Luteyn) L.B. Sm.
Lindmania sp. A [Sierra de la Neblina]
Lindmania sp. B [Sierra de la Neblina]
Lindmania sp. C
Lindmania stenophylla L.B. Sm.
Lindmania steyermarkii L.B. Sm.
Lindmania subsimplex L.B. Sm.
Lindmania thyrsoidea L.B. Sm.
Lindmania tillandsioides L.B. Sm.
Lindmania wurdackii L.B. Sm.
Mezobromelia capituligera (Griseb.) J.R. Grant {=Vriesea capituligera]
[possibly expected in GU]
Mezobromelia pleiosticha (Griseb.) Utley & H. Luther [=Vriesea pleiosticha]
Navia abysmophila L.B. Sm. [Sierra de la Neblina]
Navia affinis L.B. Sm. [Sierra de la Neblina]
Navia albiflora L.B. Sm., Steyerm. & H. Rob.
Navia aliciae L.B. Sm., Steyerm. & H. Rob.
Navia aloifolia L.B. Sm. [Sierra de la Neblina]
Navia angustifolia (Baker) Mez
Navia arida L.B. Sm. & Steyerm.
Navia aurea L.B. Sm.
Navia barbellata L.B. Sm.
Navia berryana L.B. Sm., Steyerm. & H. Rob.
Navia brachyphylla L.B. Sm.
Navia breweri L.B. Sm. & Steyerm.
Navia cardonae L.B. Sm.
Navia caricifolia L.B. Sm.
Navia carnevalii L.B. Sm. & Steyerm.
Navia cataractarum Sandwith
Navia caurensis L.B. Sm.
Navia colorata L.B. Sm.
Navia connata L.B. Sm. & Steyerm.
Navia crassicaulis L.B. Sm., Steyerm. & H. Rob. [Sierra de la Neblina]
Navia cretacea L.B. Sm.
Navia crispa L.B. Sm.
Navia cucullata L.B. Sm.
Navia culcitaria L.B. Sm., Steyerm. & H. Rob.
Navia duidae L.B. Sm. [expected in Guyana]
Navia duidae var. duidae
Navia duidae var. glabrior L.B. Sm.
Navia emergens L.B. Sm., Steyerm. & H. Rob.
AM BO
AM
BO
AM BO
BO
BO
BO
GU
GU
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA GU su FG
GU
GU
GU
GU?
GU
GU?
76 Checklist of the Plants of the Guiana Shield
Naviafilifera L.B. Sm., Steyerm. & H. Rob. [Sierra de la Neblina]
Navia geaster L.B. Sm., Steyerm. & H. Rob.
Navia glandiilifera B. Hoist
Navia glaiica L.B. Sm.
Navia gleasonii L.B. Sm.
Navia huberiana L.B. Sm., Steyerm. & H. Rob.
Navia igneosicola L.B. Sm., Steyerm. & H. Rob.
Navia immersa L.B. Sm.
Navia incrassata L.B. Sm. & Steyerm.
Navia intermedia L.B. Sm. & Steyerm.
Navia involucrata L.B. Sm.
Naviajauana L.B. Sm., Steyerm. & H. Rob.
Navia lactea L.B. Sm., Steyerm. & H. Rob.
Navia lanigera L.B. Sm.
Navia lasiantha L.B. Sm. & Steyerm.
Navia latifolia L.B. Sm.
Navia lepidota L.B. Sm. [Sierra de la Neblina]
Navia liesneri L.B. Sm., Steyerm. & H. Rob. [Sierra de la Neblina]
Navia lindmanioides L.B. Sm.
Navia linearis L.B. Sm., Steyerm. & H. Rob.
Navia luzuloides L.B. Sm., Steyerm. & H. Rob.
Navia maguirei L.B. Sm.
Navia mima L.B. Sm.
Navia mosaica B. Hoist
Navia naviculars L.B. Sm. & Steyerm.
Navia nubicola L.B. Sm.
Navia ocellata L.B. Sm.
Navia octopoides L.B. Sm.
Navia ovoidea L.B. Sm., Steyerm. & H. Rob.
Navia paruana B. Hoist
Navia pai-vida L.B. Sm.
Navia pai-vula var. expansa L.B. Sm. [Sierra de la Neblina]
Navia parviila var. parviila [Sierra de la Neblina]
Navia patria L.B. Sm. & Steyerm.
Navia paiiciflora L.B. Sm.
Navia phelpsiae L.B. Sm.
Navia piresii L.B. Sm., Steyerm. & H. Rob.
Navia polyglomerata L.B. Sm., Steyerm. & H. Rob. [Sierra de la Neblina]
Navia pulvinata L.B. Sm.
Navia pungens L.B. Sm.
Navia robinsonii L.B. Sm.
Navia rupestris (Gleason) Sandwith
Navia sandwithii L.B. Sm.
Navia saxicola L.B. Sm.
Navia scirpiflora L.B. Sm., Steyerm. & H. Rob.
Navia scopulorum L.B. Sm.
Navia semiserrata L.B. Sm.
Navia serridata L.B. Sm.
Navia sp. A
Navia sp. B [Sierra de la Neblina]
Navia sp. C
Navia splendens L.B. Sm.
Navia stenodonta L.B. Sm.
Navia steyermarkii L.B. Sm.
Navia subpetiolata L.B. Sm.
Navia tentacidata B. Hoist
Navia terramarae L.B. Sm. & Steyerm.
Navia trichodonta L.B. Sm.
Navia umbratilis L.B. Sm.
Navia viridis L.B. Sm.
Navia wiirdackii L.B. Sm.
Navia xyridiflora L.B. Sm.
Neoregelia leviana L.B. Sm.
Neoregelia myrmecophila (Ule ex G. Karst. & Schenck) L.B. Sm.
Neoregelia stolonifera L.B. Sm.
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
GU
BO
BO
BO
BO
BO
BO
BO
SU
BO
BO
BO
GU
GU
BO
BO
BO
BO
GU
Checklist of the Plants of the Guiana Shield 77
Pitcairnia agavifolia L.B. Sm.
Pitcairnia armata Maury
Pitcairnia brittoniana Mez
Pitcairnia bulbosa L.B. Sm.
Pitcairnia cana B. Hoist
Pitcairnia caricifolia Mart, ex Schult. f. [=Pepinia caricifolia]
Pitcairnia ctenophylla L.B. Sm. [=Pepinia ctenophylla] [expected in GU]
Pitcairnia echinata Hook.
Pitcairnia echinata Hook. var. echinata
Pitcairnia epiphytica L.B. Sm.
Pitcairnia filispina L.B. Sm.
Pitcairnia geyskesii L.B. Sm. [=Pepinia geyskesii]
Pitcairnia graniticola B. Hoist
Pitcairnia heterophylla (Lindl.) Beer [expected in GU]
Pitcairnia incarnata Baker [=Pepinia incarnata]
Pitcairniajuncoides L.B. Sm.
Pitcairnia kunhardtiana L.B. Sm.
Pitcairnia leopoldii (W. Till & S. Till) B. Hoist
Pitcairnia leprieurii Baker
Pitcairnia maguirei L.B. Sm.
Pitcairnia maidifolia (C. Morren) Decne. ex Planch. & Linden
Pitcairnia nematophora L.B. Sm. & Read
Pitcairnia niida Baker [=Pepinia nuda]
Pitcairnia orchidifolia Mez
Pitcairnia patentiflora L.B. Sm. [=Pepinia patentiflora]
Pitcairnia patentiflora var. armata L.B. Sm.
Pitcairnia patentiflora var. patentiflora
Pitcairnia phelpsiae (L.B. Sm.) B. Hoist & L.B. Sm.
Pitcairnia pruinosa Kunth
Pitcairnia pusilla Mez
Pitcairnia rubiginosa (Brongn. ex E. Morren) Baker [=Pepinia rubiginosa]
Pitcairnia rubignosa (Baker) Mez
Pitcairnia rubignosa var. amazonica (Baker) L.B. Sm.
Pitcairnia sastrei L.B. Sm. & Read
Pitcairnia sp. A
Pitcairnia sp. B [Sierra de la Neblina]
Pitcairnia sp. C
Pitcairnia sprucei Baker [=Pepinia sprucei]
Pitcairnia uaupensis Baker
Puyafloccosa (Linden) E. Morren ex Mez
Puyafloccosa \ar.floccosa
Puya grafii Rauh
Racinaeajenmanii (Baker) M.A. Spencer & L.B. Sm.
Racinaea spiculosa (Griseb.) M.A. Spencer & L.B. Sm.
Racinaea spiculosa var. micrantha (Baker) M.A. Spencer & L.B. Sm.
Racinaea spiculosa var. spiculosa
Racinaea spiculosa var. stenoglossa (L.B. Sm.) M.A. Spencer & L.B. Sm.
Racinaea tetrantha (Ruiz & Pav.) M.A. Spencer & L.B. Sm.
Racinaea tetrantha var. aurantiaca (Griseb.) M.A. Spencer & L.B. Sm.
Racinaea tetrantha var. caribaea (L.B. Sm.) M.A. Spencer & L.B. Sm.
Steyerbromelia deflexa L.B. Sm. & H. Rob.
Steyerbromelia diffusa L.B. Sm., Steyerm. & H. Rob.
Steyerbromelia discolor L.B. Sm. & H. Rob.
Steyerbromelia plowmanii (L.B. Sm., Steyerm. & H. Rob.) H. Rob. & D.C.
Taylor [Sierra de la Neblina]
Steyerbromelia ramosa (L.B. Sm.) B. Hoist
Steyerbromelia thomasii (L.B. Sm., Steyerm. & H. Rob.) B. Hoist [Sierra de
la Neblina]
Tillandsia adpressiflora Mez
Tillandsia anceps Lodd.
Tillandsia balbisiana Schult. f. [possibly expected in GU]
Tillandsia brachycaulos Schltdl.
Tillandsia bulbosa Hook.
Tillandsia clavigera Mez
Tillandsia compacta Griseb. [possibly expected in GU]
AM
AM BO
AM BO GU
AM
AM
AM BO DA
BO
GU SU FG
AM
AM
AM
AM
GU SU FG
BO
AM BO
FG
AM
AM
AM
GU FG
AM
AM BO GU SU
AM
BO
GU SU
AM FG
AM
AM FG
AM
AM BO
FG
AM FG
AM
AM
GU SU? FG
AM
AM
AM
AM GU FG
AM
BO
BO
AM
AM BO GU
AM BO GU SU FG
AM BO GU
AM BO GU SU FG
AM BO GU
AM BO
BO
GU
AM BO GU
AM
AM
AM
AM
AM BO
AM
AM BO SU FG
AM BO DA
BO DA
BO
GU SU FG
AM BO DA GU SU FG
AM
AM BO
78 Checklist of the Plants of the Guiana Shield
Tillandsia compacta var. compacta
Tillandsia complanata Benth.
Tillandsia confinis L.B. Sm.
Tillandsia elongata Kunth
Tillandsia elongata var. subimbricata (Baker) L.B. Sm. [expected in GU]
Tillandsiafasciculata Sw.
Tillandsia fendleri Griseb.
Tillandsiafendleri var. fendleri
Tillandsiafendleri var. reducta (L.B. Sm.) L.B. Sm.
Tillandsiaflexuosa Sw.
Tillandsia gardneri Lindl.
Tillandsia geminiflora Brongn.
Tillandsiajuncea (Ruiz & Pav.) Poir.
Tillandsia kegeliana Mez
Tillandsia monadelpha (E. Morren) Baker
Tillandsia paraensis Mez
Tillandsia polystachia (L.) L.
Tillandsia pruinosa Sw.
Tillandsia pyramidata Andre
Tillandsia pyramidata var. pyramidata
Tillandsia recurvata (L.) L.
Tillandsia schiedeana Steud.
Tillandsia striata Sol. ex Ker Gawl.
Tillandsia striata var. striata
Tillandsia tenuifolia L.
Tillandsia tenuifolia var. tenuifolia
Tillandsia turneri Baker
Tillandsia turneri var. orientalis L.B. Sm.
Tillandsia usneoides (L.) L.
Vriesea bibeatriais Morillo
Vriesea duidae (L.B. Sm.) Gouda
Vrieseafibrosa L.B. Sm.
Vriesea glutinosa Lindl.
Vriesea heliaonioides (Kunth) Hook, ex Walp.
Vriesea inaurva (Griseb.) Read [=Tillandsia inaurva]
Vrieseajohnstonii (Mez) L.B. Sm. &. Pittendr. [=Vriesea hygrometrical']
Vrieseajonghei (K. Koch) E. Morren
Vriesea maguirei L.B. Sm. [Sierra de la Neblina]
Vriesea melgueiroi I. Ramirez & Camevali
Vriesea mitoura L.B. Sm. [Sierra de la Neblina]
Vriesea platynema Gaudich.
Vriesea platynema var. platynema
Vriesea proaera (Mart, ex Schult. f.) Wittm.
Vriesea proaera var. proaera
Vriesea rubra (Ruiz & Pav.) Beer
Vriesea sanguinolenta Cogn. & Marchand
Vriesea simplex (Veil.) Beer
Vriesea soaialis L.B. Sm.
Vriesea soderstromii L.B. Sm.
Vriesea splendens (Brongn.) Lem.
Vriesea splendens var. formosa Suringar ex Witte
Vriesea splendens var. splendens
Vriesea sulaata L.B. Sm.
Vriesea wurdaakii L.B. Sm.
Werauhia gigantea (Mart, ex Schult. f.) J.R. Grant [non Vriesea gigantea
Gaudich, 1846] [=Vriesea amazoniaa]
Werauhia gladioliflora (H. Wendl.) J.R. Grant [=Vriesea gladioliflora]
Werauhia hygrometrica (Andre) J.R. Grant {=Vriesea hygrometriaa]
Werauhia ringens (Griseb.) J.R. Grant [=Vriesea ringens]
Werauhia viridiflora (Regel) J.R. Grant [=Vriesea viridiflora]
Synonyms and Misidentifications
Aeahmea basi-lateralis (Lem.) L.B.Sm. =Disteganthus basi-lateralis
Aeahmeafernandae (E. Morren) Baker =Aeahmea rubiginosa
Aeahmea humilis Mez =Aeahmea mertensii
AM BO
AM BO GU
AM BO
AM BO DA
AM BO DA
BO DA GU SU FG
AM BO GU
AM BO GU
AM BO GU
AM BO
BO
BO
DA
DA
GU SU
SU
FG
BO GU SU FG
BO DA GU SU FG
AM BO
BO DA
GU SU FG
AM BO
BO
BO
SU
AM BO
AM
BO GU SU
BO
BO GU SU FG
BO
AM BO GU
AM BO GU
BO DA GU SU FG
AM BO
AM BO GU
AM
BO
AM GU SU FG
AM BO GU
GU
FG
AM
AM
AM
AM BO DA GU
AM BO DA
DA GU SU FG
DA GU SU
AM BO
BO
BO
DA GU
AM
GU
BO GU
GU
SU FG
BO GU SU FG
AM
AM
DA GU SU FG
GU SU FG
BO GU
GU
BO GU
Checklist of the Plants of the Guiana Shield 79
Aechmea kienastii E. Morren ex Mez =Aechmea tillandsioides
Aechmea lateralis L.B. Sm. =Disteganthus lateralis
Aechmea magdalenae auct. non (Andre) Andre ex Baker sensu L.B. Sm. =Aechmea rubiginosa
Aechmea megalantha Harms =Aechmea melinonii
Aechmea mucroniflora Hook. =Aechmea mertensii
Aechmea oligosphaera Baker =Hohenbergia stellata
Aechmea prancei L.B. Sm. [not in flora area, fide Gouda]
Aechmea spicata Mart, ex Schult. f. =Aechmea mertensii
Aechmea spriicei Mez =Aechmea castelnavii
Aechmea tillandsioides var. kienastii (E. Morren ex Mez) L.B. Sm. =Aechmea tillandsioides
Aechmea wiillschlaegeliana Mez =Aechmea mertensii
Ananas ananassoides var. nanus L.B. Sm. =Ananas nanus
Ananas sativus Schult. f. =Ananas comosus
Anoplophytum guianensis Beer =Lindmania guianensis
Billbergia chantinii Carriere =Aechmea chantinii
Billbergia contracta Mart, ex Schult. f. =Aechmea contracta
Billbergia corymbosa Mart, ex Schult. f. ^Aechmea corymbosa
Billbergia tillandsioides Mart, ex Schult. f. =Aechmea tillandsioides
Bonparteajiincea Ruiz & Pav. =TillandsiaJuncea
Brocchinia bernardii L.B. Sm. =Brocchinia melanacra
Brocchinia cordylinoides Baker [nom. illeg.] =Brocchinia micrantha
Brocchinia cryptantha L.B. Sm. =Brocchinia hechtioides
Brocchinia oliva-estevae Steyerm. & L.B. Sm. =Brocchinia tatei
Brocchinia secunda L.B. Sm. =Brocchinia tatei
Bromelia aquilega Salisb. =Aechmea aquilega
Bromelia comosa L. =Ananas comosus
Bromelia karatas L. =Bromelia plumieri
Bromelia lingulata L. =Aechmea lingulata
Bromelia longifolia Rudge =Aechmea longifolia
Bromelia lutea G. Mey. =Aechmea nudicaulis var. cuspidata
Bromelia mertensii G. Mey. =Aechmea mertensii
Bromelia nudicaulis L. =Aechmea nudicaulis
Caraguata hygrometrica Andre =Werauhia hygrometrica
Caraguata lingulata (L.) Lindl. =Guzmania lingulata
Caraguata sphaeroidea Andre =Guzmania sphaeroidea
Caraguata splendens Planch. =Guzmania lingulata
Catopsisfendleri Baker =Racinaea tetrantha var. caribaea
Catopsis inconspicua Baker [not in flora area] =Catopsis nitida (Hook.) Griseb.
Chevalierafernandae (E. Morren) L.B. Sm. & W.J. Kress =Aechmea rubiginosa
Chevaliera pallida (L.B. Sm.) L.B. Sm. & W.J. Kress =Aechmea pallida
Chevaliera rubiginosa (Mez) L.B. Sm. & W.J. Kress =Aechmea rubiginosa
Connellia smithiana Steyerm. & Luteyn =Lindmama smithiana
Cottendorfia arachnoidea L.B. Sm., Steyerm. & H. Rob. =Lindmania arachnoidea
Cottendorfia argentea (L.B. Sm.) L.B. Sm. =Lindmama argentea
Cottendorfia atrorosea L.B. Sm., Steyerm. & H. Rob. =Lindmania atrorosea
Cottendorfia brachyphylla (L.B. Sm.) L.B. Sm. =Lindmania brachyphylla
Cottendorfia brachyphylla var. angustior L.B. Sm., Steyerm. & H. Rob. =Lindmania brachyphylla
Cottendorfia cylindrostachya (L.B. Sm.) L.B. Sm. =Lindmania cylindrostachya
Cottendorfia dendritica L.B. Sm. =Lindmania dendritica
Cottendorfia ? dyckioides L.B. Sm. =Lindmania guianensis
Cottendorfia geniculata (L.B. Sm.) L.B. Sm. =Lindmania guianensis
Cottendorfia ? geniculata var. minor L.B. Sm., Steyerm. & H. Rob. =Lindmania guianensis
Cottendorfia gracillima L.B. Sm. =Lindmania gracillima
Cottendorfia guianensis (Beer) Klotzsch ex Baker =Lindmania guianensis
Cottendorfia ? guianensis var. vestita L.B. Sm. =Lindmania guianensis
Cottendorfia lateralis L.B. Sm. & Read =Lindmania lateralis
Cottendorfia longipes L.B. Sm. =Lindmania longipes
Cottendorfia maguirei L.B. Sm. =Lindmania maguirei
Cottendorfia marahuacae L.B. Sm., Steyerm. & H. Rob. =Lindmania marahuacae
Cottendorfia minor (L.B. Sm.) L.B. Sm. =Lindmania minor
Cottendorfia navioides (L.B. Sm.) L.B. Sm. =Lindmania navioides
Cottendorfia nubigena L.B. Sm. =Lindmania nubigena
Cottendorfia paludosa (L.B. Sm.) L.B. Sm. =Lindmania guianensis
Cottendorfia phelpsiae (L.B. Sm.) L.B. Sm. =Lindmania phelpsiae
80 Checklist of the Plants of the Guiana Shield
Cottendorfia savannemis L.B. Sm. =Lindmania nubigena
Cottendorfia serrulata (L.B. Sm.) L.B. Sm. =Lindmania serrulata
Cottendorfia serrulata van reducta (L.B. Sm.) L.B. Sm. =Lindmania serrulata
Cottendorfia stenophylla (L.B. Sm.) L.B. Sm. =Lindmama stenophylla
Cottendorfia steyermarkii (L.B. Sm.) L.B. Sm. =Lindmania steyermarkii
Cottendorfia subsimplex (L.B. Sm.) L.B. Sm. =Lindmania subsimplex
Cottendorfia thyrsoidea (L.B. Sm.) L.B. Sm. =Lindmania thyrsoidea
Cottendorfia tillandsioides (L.B. Sm.) L.B. Sm. =Lindmania tillandsioides
Cottendorfia wurdackii (L.B. Sm.) L.B. Sm. =Lindmania wurdackii
Disteganthus bromeliifolius L.B. Sm. [nomen nudum] =Disteganthus lateralis
Disteganthus calatheoides (L.B. Sm.) L.B. Sm. & Read =Disteganthus basi-lateralis
Dyckia augustae (M.R. Schomb.) Baker =Connellia augustae
Encholirium augustae M.R. Schomb. =Connellia augustae
Gravisia aquilega (Salisb.) Mez =Aechmea aquilega
Gravisia lanjouwii (L.B. Sm.) L.B. Sm. =Aechmea lanjouwii
Guzmania angustifolia (Baker) Wittm. [not in flora area]
Guzmania capituligera (Griseb.) Mez =Meiobromelia capituligera
Guzmania cryptantha L.B. Sm. =Guzmania squarrosa
Guzmania erythrolepis Brongn. ex Planch, [probably misident. of G. melinonis, fide E. Gouda]
Guzmania geniculata L.B. Sm. =Guzmania sphaeroidea
Guzmania lingulata var. minor (Mez) L.B. Sm. & Pittendr. =Guzmania lingulata
Guzmania lingulata var. splendens (Planch.) Mez =Guzmania lingulata
Guzmania minor Mez =Guzmania lingulata
Guzmania pleiosticha auct. =Guzmania altsonii
Guzmania pleiosticha (Griseb.) Mez =Mezobromelia pleiosticha
Guzmania plumieri auct. non (Griseb.) Mez sensu L.B. Sm. in Lasser =Guzmania steyermarkii
Guzmania splitgerberi Mez =Mezobromelia pleiosticha
Guzmania venamensis L.B. Sm. =Guzmania sphaeroidea
Hepetis amazonica (Baker) Mez =Pifcairnia rubignosa var. amazonica
Hepetis brittoniana (Mez) Mez =Pitcairnia brittoniana
Hepetis caricifolia (Mart, ex Schult. f.) Mez =Pitcairnia caricifolia
Hepetis kegeliana (Schltdl.) Mez =Pitcairnia caricifolia
Hepetis pauciflora (Baker) Mez =Pitcairnia caricifolia
Hepetis pruinosa (Kunth) Mez =Pitcairnia pruinosa
Hepetis rubiginosa (Baker) Mez =Pitcairnia rubiginosa
Hepetis sprucei (Baker) Mez =Pitcairnia sprucei
Hepetis uaupensis (Baker) Mez =Pitcairnia uaupensis
Karatas plumieri E. Morren =Bromelia plumieri
Lamprococcus brevicollis (L.B. Sm.) L.B. Sm. & W.J. Kress =Aechmea brevicollis
Lamprococcus campanulatus (L.B. Sm.) L.B. Sm. & W.J. Kress =Aechmea campanulata
Lindmania brachyphylla var. angustior (L.B. Sm., Steyerm. & H. Rob.) R.W. Read =Lindmania brachyphylla
Lindmania dyckioides (L.B. Sm.) L.B. Sm. =Lindmania guianensis
Lindmania geniculata L.B. Sm. =Lindmania guianensis
Lindmania geniculata var. minor (L.B. Sm., Steyerm. & H. Rob.) L.B. Sm. =Lindmania guianensis
Lindmania paludosa L.B. Sm. =Lindmania guianensis
Lindmania savannensis (L.B. Sm.) L.B. Sm. =Lindmania nubigena
Lindmania serrulata var. reducta L.B. Sm. =Lindmania serrulata
Lindmania sessilis L.B. Sm., Steyerm. & H. Rob. =Lindmania serrulata
Lindmania terramarae L.B. Sm., Steyerm. & H. Rob. =Lindmania marahuacae
Macrochordion bromeliifolia (Rudge) Beer =Aechmea bromeliifolia
Navia brocchinioides L.B. Sm. =Brewcaria brocchinioides
Navia delascionis L.B. Sm., Steyerm. & H. Rob. =Navia berryana
Navia diffusa L.B. Sm. non Steyerbromelia diffusa L.B. Sm., Steyerm. & H. Rob. 1986 [1987]. =Steyerbromelia plowmanii
Navia fluviatilis L.B. Sm. =Navia rupestris
Navia gracilis L.B. Sm. ^Brewcaria reflexa
Navia grafii Steyerm. & B. Hoist =Navia phelpsiae
Navia hechtioides L.B. Sm. =Brewcaria hechtioides
Navia hohenbergioides L.B. Sm. =Brewcaria hohenbergioides
Navia iosothrix L.B. Sm. & Steyerm. =Navia nubicola
Navia lopezii L.B. Sm. (Xyridaceae) =Aratitiyopea lopezii
Navia pedemontana L.B. Sm., Steyerm. & H. Rob. =Navia nubicola
Navia ? platyphylla L.B. Sm. & Steyerm. =Steyerbromelia ramosa
Navia plowmanii L.B. Sm., Steyerm. & H. Rob. =Steyerbromelia plowmanii
Navia ramosa L.B. Sm. =Steyerbromelia ramosa
Navia reflexa L.B. Sm. =Brewcaria reflexa
Checklist of the Plants of the Guiana Shield
Navia thomasii L.B. Sm., Steyerm. & H. Rob. =Steyerbromelia thomasii
Nidularium myrmecophilum Ule ex G. Karst. & Schenck =Neoregelia myrmecophila
Pepinia agavifolia (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia agavifolia
Pepinia armata (Maury) G.S. Varad. & Gilmartin =Pitcairnia armata
Pepinia breweri (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia armata
Pepinia bulbosa (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia bulbosa
Pepinia caricifolia (Mart, ex Schult. f.) G.S. Varad. & Gilmartin =Pitcaimia caricifolia
Pepinia ctenophylla (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia ctenophylla
Pepinia epiphytica (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcaimia epiphytica
Pepiniafilispina (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairniafilispina
Pepinia geyskesii (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia geyskesii
Pepinia incarnata (Baker) G.S. Varad. & Gilmartin =Pitcairnia incarnata
Pepiniajuncoides (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairniajuncoides
Pepinia kunhardtiana (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia kunhardtiana
Pepinia leopoldii W. Till & S. Till =Pitcaimia leopoldii
Pepinia maguirei (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia maguirei
Pepinia nuda (Baker) G.S. Varad. & Gilmartin =Pitcairnia nuda
Pepinia patentiflora (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia patentiflora
Pepinia pruinosa (Kunth) G.S. Varad. & Gilmartin =Pitcairnia pruinosa
Pepinia rubiginosa (Brongn.) G.S. Varad. & Gilmartin =Pitcairnia rubiginosa
Pepinia sprucei (Baker) G.S. Varad. & Gilmartin =Pitcaimia sprucei
Pepinia uaupensis (Baker) G.S. Varad. & Gilmartin =Pitcairnia uaupensis
Pepinia wurdackii (L.B. Sm.) G.S. Varad. & Gilmartin =Pitcairnia bulbosa
Phytarrhiza monadelpha E. Morren =Tillandsia monadelpha
Pitcairnia amazonica Baker =Pitcairnia rubignosa van amazonica
Pitcairnia breweri L.B. Sm. =Pitcairnia armata
Pitcairnia cinerea L.B. Sm. =Pitcairnia armata
Pitcairnia kegeliana Schltdl. =Pitcairnia caricifolia
Pitcairnia nuda auct. non Baker: sensu L.B. Sm. & R.J. Downs =Pitcairnia patentiflora var. patentiflora
Pitcairnia pauciflora Baker =Pitcairnia caricifolia
Pitcairnia viridiflora Regel =Werauhia viridiflora
Pitcairnia wurdackii L.B. Sm. =Pitcairnia bulbosa
Platyaechmea chantinii (Carriere) L.B. Sm. & W.J. Kress =Aechmea chantinii
Platyaechmea contracta (Mart, ex Schult. f.) L.B. Sm. & W.J. Kress ^Aechmea contracta
Platyaechmea tillandsioides (Mart, ex Schult. f.) L.B. Sm. & W.J. Kress =Aechmea tillandsioides
Pogospermum berteronianum (Schult. f.) Brongn. =Catopsis berteroniana
Pothuava nudicaulis (L.) Regel =Aechmea nudicaulis
Pothuava nudicaulis var. cuspidata (Baker) L.B. Sm. & W.J. Kress =Aechmea nudicaulis var. cuspidata
Pourretiafloccosa Linden =Puyafloccosa
Puya augustae (M.R. Schomb.) Mez =Connellia augustae
Puya guianensis Klotzsch ex M.R. Schomb. [nomen nudum] =Puyafloccosa var. floccosa
Puya guyanensis Baker =Puyafloccosa var floccosa
Puya heterophylla Lindl. =Pitcairnia heterophylla
Puya maidifolia E. Morren =Pitcairnia maidifolia
Puya phelpsiae L.B. Sm. =Pitcairnia phelpsiae
Puya quelchii (N.E. Br.) L.B. Sm. =Connellia quelchii
Puya roraimae Mez =Connellia quelchii
Quesnelia lamarckii Baker (Musaceae) [cultivated only] =Musa coccinea Andrews
Schlumbergeria roezli E. Morren =Guzmania roezlii
Steyerbromelia neblinae B. Hoist =Steyerbromelia plowmanii
Streptocalyx angustifolius Mez =Aechmea longifolia
Streptocalyx longifolius (Rudge) Baker =Aechmea longifolia
Streptocalyx poeppigii Beer =Aechmea beeriana
Streptocalyx poitaei Baker =Aechmea poitaei
Thecophyllum splitgerberi (Mez) Pittendr. =Mezobromelia pleiosticha
Thecophyllum squarrosum Mez & Sodiro =Guzmania squarrosa
Tillandsia abysmophila L.B. Sm. & Steyerm. =Tillandsia confinis
Tillandsia aloifolia Hook. =Tillandsiaflexuosa
Tillandsia amazonica Baker =Werauhia gigantea
Tillandsia appuniana Baker = Vriesea splendens var. splendens
Tillandsia archeri L.B. Sm. ^Tillandsia turneri
Tillandsia aurantiaca Griseb. =Racinaea tetrantha var. aurantiaca
Tillandsia berteroniana Schult. f. =Catopsis berteroniana
Tillandsia bromeliaefolia Rudge =Aechmea bromeliifolia
Tillandsia capituligera Griseb. =Mezobromelia capituligera
82 Checklist of the Plants of the Guiana Shield
Tillandsia caracasana Baker =TiUandsia compacta var. compacta
Tillandsia caribaea L.B. Sm. =Racinaea tetrantha var. caribaea
Tillandsia chinchiicuana Harms =Racinaea spiculosa var. micrantha
Tillandsia compressa Bertero [not in flora area]
Tillandsia conspersa Miq. =Tillandsia stricta
Tillandsia crinita Willd. ex Beer =Tillandsia usneoides
Tillandsia dactylifera E. Morren ex Baker =Vriesea incurva
Tillandsia deppeana var. clavigera (Mez) L.B. Sm. =Tillandsia clavigera
Tillandsia deppeana var. latifolia (Griseb.) L.B. Sm. =Tillandsia fendleri war. fendleri
Tillandsia deppeana var. reducta (L.B. Sm.) L.B. Sm. =Tillandsia fendleri var. reducta
Tillandsia duidae L.B. Sm. =Vnesea diiidae
Tillandsia excelsa var. latifolia Griseb. =Tillandsia fendleri var. fendleri
Tillandsiafendleri (Baker) Mez =Racinaea tetrantha var. caribaea
Tillandsiaferruginascens Mez [nom. inval.] =Racinaea tetrantha var. caribaea
Tillandsiaflexuosa var. vivipara Andre =Tillandsiaflexuosa
Tillandsia fiilgens (Klotzsch ex Beer) Mez =Racinaea tetrantha var. aurantiaca
Tillandsia gigantea Mart, ex Schult. f. =Weraiihia gigantea
Tillandsia graminifolia Baker =Tillandsia monadelpha
Tillandsia gy^mnophylla Baker =Gitzmania monostachia var. monostachia
Tillandsia heliconioides Kunth ^Vriesea heliconioides
Tillandsia heliconioides auct. non Kunth: sensu Griseb. =Guzmania monostachia var. monostachia
Tillandsia incurva Griseb. =Vriesea incurva
TillandsiaJenmanii Baker =Racinaeajenmanii
Tillandsia lineatifolia Mez =Tillandsia anceps
Tillandsia lingulata L. =Guzmania lingulata
Tillandsia micrantha Baker =Racinaea spiculosa var. micrantha
Tillandsia multifolia Mez =Tillandsia turneri var. orientalis
Tillandsia pachychlamys Mez = Werauhia gladioliflora
Tillandsia panicidata fifendleri (Griseb.) Mez =Tillandsia fendleri
Tillandsia parkeri Baker =Racinaea spiculosa var. spiculosa
Tillandsia parviflora auct. non Ruiz & Pav. 1802: sensu Griseb. =Racinaea tetrantha var. caribaea
Tillandsia patens Willd. ex Schult. f. =Tillandsiaflexuosa
Tillandsia pleiosticha Griseb. =Mezobromelia pleiosticha
Tillandsia procera Mart, ex Schult. f. =Vriesea procera
Tillandsia pulchella Hook. =Tillandsia tenuifolia
Tillandsia pulchella var. surinamensis Mez =Tillandsia tenuifolia
Tillandsia pumila Griseb. =Tillandsia bulbosa
Tillandsia punctidata Cham. & Schltdl. [not in flora area]
Tillandsia rhodocincta Baker =Tillandsia turneri var. orientalis
Tillandsia rhododactyla Mez =Vriesea rubra
Tillandsia rigidula Baker =Guzmania roezlii
Tillandsia rubra Ruiz & Pav. =Vriesea rubra
Tillandsia rubra var. fendleri (Griseb.) Mez =Tillandsiafendleri
Tillandsia rubra var. reducta L.B. Sm. =Tillandsia fendleri var. reducta
Tillandsia sessiliflora Ruiz & Pav. =Catopsis sessiliflora
Tillandsia simplex Veil. =Vriesea simplex
Tillandsia spiculosa var. micrantha (Baker) L.B. Sm. =Racinaea spiculosa var. micrantha
Tillandsia spiculosa var. spiculosa =Racinaea spiculosa var spiculosa
Tillandsia spiculosa var. stenoglossa (L.B. Sm.) Gouda =Racinaea spiculosa var. stenoglossa
Tillandsia splendens Brongn. =Vriesea splendens
Tillandsia stenoglossa L.B. Sm. =Racinaea spiculosa var stenoglossa
Tillandsia subimbricata Baker =Tillandsia elongata var subimbricata
Tillandsia surinamensis Miq. ex Mez =Tillandsia tenuifolia
Tillandsia tenuifolia var. surinamensis (Mez) L.B. Sm. =Tillandsia tenuifolia
Tillandsia tetrantha var. aurantiaca (Griseb.) L.B. Sm. =Racinaea tetrantha var. aurantiaca
Tillandsia tetrantha var. caribaea (L.B. Sm.) Gouda =Racinaea tetrantha var. caribaea
Tillandsia trichoides Kunth ^Tillandsia usneoides
Tillandsia usneoides var. fliformis (Andre) Mez =Tillandsia usneoides
Tillandsia vestita Schltdl. & Cham. =Tillandsia schiedeana
Tillandsia viscidula Britton =Vriesea procera
Tillandsia xiphostachys Griseb. =Tillandsia anceps
Tussaciafulgens Klotzsch ex Beer =Racinaea tetrantha var. aurantiaca
Vriesea amazonica (Baker) Mez =Werauhia gigantea
Vriesea capituligera (Griseb.) L.B. Sm. & Pittendr. =Mezobromelia capituligera
Vriesea gigantea (Mart, ex Schult. f.) Mez =Werauhia gigantea
Checklist of the Plants of the Guiana Shield 83
Vhesea gladioliflora (H. Wendl.) Antoine =Werauhia gladioliflora
Vriesea heliconioides var. polysticha L.B. Sm. =Vriesea heliconioides
Vriesea heterandra (Andre) L.B. Sm. [misident. of possibly non-Guianan material]
Vriesea hygrometrica (Andre) L.B. Sm. & Pittendr. =Werauhia hygrometrica
Vriesea pachychlamys Mez = Werauhia gladioliflora
Vriesea pleiosticha (Griseb.) Gouda =Mezobromelia pleiosticha
Vriesea ringens (Griseb.) Mez =Weranhia ringens
Vriesea splitgerberi (Mez) L.B. Sm. & Pittendr. =Mezobromelia pleiosticha
Vriesea viridiflora (Regel) Wittm. ex Mez =Werauhia viridiflora
Wittmackia lingulata (L.) Mez =Aechmea Ungulata
Wittmackia odora Mez =Aechmea lingulata
References
Grant, J.R. 1995. The resurrection ofAlcantarea and Werauhia, a new genus. Tropische und subtropische Pflanzenwelt 91 : 1-59.
Gouda, E.J. 1987. Bromeliaceae, subfamily Tillandsioideae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 3: 1-112.
Gouda, E. 1997. Bromeliaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 215-232.
Hoist, B.K. 1994. Checklist of Venezuelan Bromeliaceae with notes on species distribution by state and levels of endemism.
Selbyana 15(1): 132-149.
Hoist, B.K. 1997. Bromeliaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 548-676.
Kramer, K.U. & A.R.A. Gorts-van Rijn. 1968. Bromeliaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of
Suriname 1(2): 413-414.
Smith, L.B. 1957. Bromeliaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 94-148.
Smith, L.B. 1967. Bromeliaceae of the Guayana Highland. Mem. New York Bot. Gard. 14(3): 15-68.
Smith, L.B. 1971. Bromeliaceae. Flora of Venezuela 12(1): 1-361
Smith, L.B. 1986. Revision of the Guayana Highland Bromeliaceae. Ann. Missouri Bot. Gard. 73(4): 689-721.
Smith, L.B. & R.J. Downs. 1974. Pitcaimioideae. Flora Neotropica Monogr. 14, part 1: 1-658.
Smith, L.B. & R.J. Downs. 1977. Tillandsioideae. Flora Neotropica Monogr. 14, part 2: 663-1492.
Smith, L.B. & R.J. Downs. 1979. Bromelioideae. Flora Neotropica Monogr. 14, part 3: 1493-2142.
BURMANNIACEAE
P.J.M. Maas, H. Maas
Apteria aphylla (Nutt.) Bamhart ex Small
Burmannia bicolor Mart.
Burmannia capitata (Walter ex J.F. Gmel.) Mart.
Burmannia compacta Maas & H. Maas
Burmannia dasyantha Mart.
Burmanniaflava Mart.
Burmannia foliosa Gleason
Burmannia kalbreyeri Oliv.
Burmannia polygaloides Schltr.
Burmannia sanariapoana Steyerm.
Burmannia tenella Benth.
Campylosiphon purpurascens Benth.
Cymbocarpa saccata Sandwith
Dictyostega orobanchoides (Hook.) Miers
Dictyostega orobanchoides subsp. orobanchoides
Dictyostega orobanchoides suhsp. parviflora (Benth.) Snelders & Maas
Gymnosiphon brachycephalus Snelders & Maas
Gymnosiphon breviforus Gleason
Gymnosiphon capitatus (Benth.) Urb.
Gymnosiphon cymosus (Benth.) Benth. & Hook. f.
Gymnosiphon divaricatus (Benth.) Benth. & Hook. f.
Gymnosiphonfimbriatus (Benth.) Urb.
Gymnosiphon guianensis Gleason
Gymnosiphon minutus Snelders & Maas
Gymnosiphon recurvatus Snelders & Maas
Gymnosiphon suaveolens (H. Karst.) Urb.
Gymnosiphon tenellus (Benth.) Urb.
Hexapterella gentianoides Urb.
Hexapterella steyermarkii Maas & H. Maas
Miersiella umbellata (Miers) Urb.
Thismia saiilensis H. Maas & Maas
AM BO
AM BO
AM BO
AM
AM
BO
AM BO
BO
AM
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
AM BO
AM BO
AM BO
BO
AM
AM BO
AM BO
DA
GU su EG
GU su EG
GU su EG
GU su EG
GU
GU SU
GU
GU SU
GU SU
GU SU
GU su
GU su
GU
su
GU su
GU
GU su
GU
GU
GU su
GU
EG
EG
EG
EG
FG
EG?
FG
EG
84 Checklist of the Plants of the Guiana Shield
Synonyms and Misidentifications
Apteria aphylla var. hymenanthera (Miq.) Jonker =Apteria aphylla
Apteria gracillima Sagot [nomen nudum] =Apteria aphylla
Apteria hymenanthera Miq. =Apteria aphylla
Apteria orobanchoides Hook. =Dictyostega orobanchoides
Benitzia suaveolens H. Karst. =Gymnosiphon suaveolens
Burmannia brachyphylla Willd. ex Schult. & Schult. f. =Burmannia bicolor
Burmannia brachystachya Miq. =Burmannia bicolor
Burmannia bracteosa Gleason =Burmannia capitata
Burmannia carrenoi Steyerm. ^Burmannia kalbreyeri
Burmannia foliosa subsp. marahuacensis Maguire & Steyerm. =Burmanniafoliosa
Dictyostega schomburgkii Miers =Dictyostega orobanchoides subsp. orobanchoides
Dictyostega schomburgkii var. parviflora Benth. =Dictyostega orobanchoides subsp. parviflora
Dictyostega umbellata Miers =Miersiella umbellata
Gymnosiphon altsoni Gleason =Hexapterella gentianoides
Gymnosiphon arcuatus Urb. =Gymnosiphon divaricatus
Gymnosiphon cornutus (Benth.) Benth. & lH[ook. f. =Gymnosiphon divaricatus
Miersiella aristata Sandwith =Miersiella umbellata
Ptychomeria cymosa Benth. =Gymnosiphon cymosus
Ptychomeria divaricata Benth. =Gymnosiphon divaricatus
Ptychomeriafimbriata Benth. =Gymnosiphonfimbriatus
Ptychomeria tenella Benth. ^Gymnosiphon tenellus
Vogelia capitata Walter ex J.F. Gmel. =Burmannia capitata
References
Jonker, F.P. 1938. Burmanniaceae. In A. Pulle, Flora of Suriname 1(1): 176-187.
Jonker, F.P. 1968. Burmanniaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 321-325.
Maas, P.J.M., H. Maas-van de Kamer, J. van Benthem, H.C.M. Snelders, & T. Riibsamen. 1986. Burmanniaceae. Flora Neotropica
Monogr. 42: 1-189.
Maas, P.J.M. & H. Maas-van de Kamer. 1989. Burmanniaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 6: 1-43.
Maas, P.J.M. & H. Maas-van de Kamer. 1987. A new Thismia (Burmanniaceae) from French Guiana. Brittonia 39(3): 376-378.
Maas, P.J.M. & H. Maas-van de Kamer. 1997. Bromeliaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora ofthe Venezuelan
Guayana 3: 678-688.
Maas-van de Kamer, H. & P. Maas. 1997. Burmanniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, &
J.D. Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and
Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1: 232-237.
Cannaceae
P.J.M. Maas, B.A. Stein
Canna glauca L.
Canna indica L. [naturalized?]
Synonyms and Misidentifications
Canna angustifolia L. =Canna glauca
Canna coccinea Mill. =Canna indica
Cannaformosa Bouche =Canna indica
Canna heliconiifolia Bouche =Canna indica
Canna lambertii Lindl. =Canna indica
Canna pallida Roscoe =Canna indica
Canna surinamensis Bouche =Canna indica
References
Maas, RJ.M. 1979. Cannaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 412-415.
Maas, RJ.M. 1985. Cannaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 1: 69-73.
Maas, P. & H. Maas-van de Kamer. 1997. Cannaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D.
Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and
Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1: 237.
Miller, J.S. 1998. Cannaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 130-131.
GU SU FG
DA GU SU FG
Checklist of the Plants of the Guiana Shield 85
COMMELINACEAE
RE. Berry, R. Faden
BO
Aneilema brasiliense C.B. Clarke
Aneilema wnbrosum (Vahl) Kunth
Aneilema umbrosum subsp. ovato-oblongum (R Beauv.) J.K. Morton
Buforrestia candolleana C.B. Clarke
Callisia filiformis (M. Martens & Galeotti) D.R. Hunt
Callisia repens L.
Commelina benghalensis L. [naturalized]
Commelina caroliniana Walter
Commelina diffusa Burm. f.
Commelina erecta L.
Commelina obliqua Vahl
Commelina platyphylla Klotzsch ex C.B. Clarke
Commelina rufipes Seub.
Commelina rufipes var. glabrata (D.R. Hunt) Faden & D.R. Hunt
Commelina rufipes var. rufipes
Commelina schomburgkiana Klotzsch
Dichorisandra diederichsanae Steyerm.
Dichorisandra hexandra (Aubl.) Standi.
Dichorisandra sp. nov. [Saiil, Central French Guiana]
Dichorisandra villosula Mart, ex Schult. & Schult. f.
Floscopa peruviana Hassk. ex C.B. Clarke
Gibasis geniculata (Jacq.) Rohweder
Murdannia nudiflora (L.) Brenan
Murdannia schomburgkiana (Kunth) Briickn.
Tinantia umbellata (Vahl) Urb.
Tradescantia zanonia (L.) Sw.
Tradescantia zebrina hort. ex Bosse
Tripogandra glandulosa (Seub.) Rohweder
Tripogandra serrulata (Vahl) Handlos
Synonyms and Misidentifications
Aneilema geniculata (Jacq.) Woodson =Gibasis geniculata
Aneilema ovato-oblongum R Beauv. =Aneilema umbrosum subsp. ovato-oblongum
Athyrocarpus persicariifolius (DC.) Hemsl. =Commelina rufipes var. glabrata
Athyrocarpus rufipes (Seub.) Standi. =Commelina rufipes var. rufipes
Campelia zanonia (L.) Kunth =Tradescantia zanonia
Commelina cayennensis Rich. ^Commelina diffusa
Commelina elegans Kunth =Commelina erecta
Commelina hexandra Aubl. =Dichorisandra hexandra
Commelina longicaulis Jacq. =Commelina diffusa
Commelina nudiflora L. ^Murdannia nudiflora
Commelina nudiflora auct. non L.: sensu Burm. f. =Commelina diffusa
Commelina persicariaefolia auct. non Delile =Commelina rufipes var. glabrata
Commelina serrulata Vahl =Tripogandra serrulata
Commelina umbrosa Vahl =Aneilema umbrosum
Commelina virginica L. [not in flora area]
Commelina zanonia L. =Tradescantia zanonia
Commelinopsis glabrata D.R. Hunt =Commelina rufipes var. glabrata
Commelinopsis persicariifolia (DC.) Pichon =Commelina rufipes var. glabrata
Commelinopsis persicariifolium auct. non (Delile) Pichon ^Commelina rufipes var. glabrata
Dichorisandra aubletiana Schult. & Schult. f. =Dichorisandra hexandra
Dichorisandra incurva Roem. & Schult. [not in flora area]
Dichorisandra leucophthalmos Hook, [not in flora area]
Leptorhoeo filiformis (M. Martens & Galeotti) C.B. Clarke =Callisia filiformis
Phaeosphaerion persicariifolium auct. non (Delile) C.B. Clarke =Commelina rufipes var. glabrata
Rhoeo discolor (L'Her.) Hance ex Walp. [cultivated only] =Tradescantia spathacea Sw.
Tinantia erecta (Jacq.) Schltdl. [not in flora area]
Tradescantia elongata G. Mey. [not in flora area]
AM GU SU FG
AM
BO
BO
GU
GU
SU
SU
FG
FG
FG
FG
AM BO DA GU SU FG
BO GU
GU
SU FG
FG
AM BO DA GU SU FG
AM BO DA GU SU FG
BO GU
GU
GU
SU FG
AM BO DA GU SU FG
FG
AM
AM SU FG
BO GU SU FG
AM BO
BO
DA GU
GU
GU
SU FG?
AM BO GU
GU
AM FG
BO DA GU SU FG
86 Checklist of the Plants of the Guiana Shield
TradescantiaflUformis M. Martens & Galeotti =Callisia filiformis
Tradescantia geniculata Jacq. =Gibasis genicidata
Tradescantia glandulosa Seub. =Tripogandra glandulosa
Tradescantia umbellata Vahl =Tinantia umbellata
References
Aristeguieta, L. 1965. Notas sobre la familia Commelinaceae en Venezuela. Bol. Acad. Cienc. Fisicas, Matematicas y Naturales
25(69): 94-142.
Berry, RE. 1998. Commelinaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 353-365.
Oracle, C. 1997. Commelinaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 237-243.
Uittien, H. 1938. Commelinaceae. In A. Pulle, Flora of Suriname 1(1): 197-212.
COSTACEAE
H. Maas, P.J M. Maas
Costiis amazonicus (Loes.) J.F. Macbr.
Costus arabicus L.
Costus claviger Benoist
Costus comosus (Jacq.) Roscoe
Costus comosus var. bakeri (K. Schum.) Maas
Costus congestiflorus Rich, ex Gagnep.
Costus curcumoides Maas
Costus erythrothyrsus Loes.
Costus guanaiensis Rusby
Costus guanaiensis var. macrostrobilus (K. Schum.) Maas
Costus lanceolatus Petersen
Costus lanceolatus subsp. pulchriflorus (Ducke) Maas
Costus lasius Loes.
Costus lima K. Schum.
Costus longebracteolatus Maas
Costus scaber Ruiz & Pav.
Costus spiralis (Jacq.) Roscoe
Costus spiralis var. spiralis
Costus spiralis var. villosus Maas
Costus villosissimus Jacq.
Dimerocostus strobilaceus Kuntze
Dimerocostus strobilaceus subsp. gutierrezii (Kuntze) Maas
Dimerocostus strobilaceus subsp. strobilaceus
GU
BO DA GU SU FG
GU SU FG
GU
GU
DA GU SU FG
FG
GU SU FG
BO GU SU FG
BO GU
GU
GU
SU
SU
FG
FG
FG
BO DA GU SU FG
BO GU SU FG
BO GU SU FG
GU SU FG
BO GU FG
DA GU SU
DA GU SU
AM
Synonyms and Misidentifications
Costus bracteatus Gleason =Costus claviger
Costus ciliatus Miq. =Costus scaber
Costus congestiflorus var. glabrior K. Schum. =Costus congestiflorus
Costus cylindricus auct. non Jacq. =Costus scaber
Costus cylindricus var. ciliatus (Miq.) Petersen ^Costus scaber
Costus discolor Roscoe =Costus arabicus
Costus guianicus Loes. ^Costus claviger
Costus macrostrobilus K. Schum. =Costus guanaiensis var. macrostrobilus
Costus niveus G. Mey. =Costus arabicus
Costus pulchriflorus Ducke =Costus lanceolatus subsp. pulchriflorus
Costus ramosus Woodson =Costus arabicus
Costus scaberulus Rich, ex Gagnep. =Costus scaber
Dimerocostus gutierrezii Kuntze =Dimerocostus strobilaceus subsp. gutierrezii
References
Maas, H. & P.J.M. Maas. 1997. Costaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4:
424-430.
Maas, P.J.M. 1972. Costoideae (Zingiberaceae). Flora Neotropica Monogr. 8: 1-140.
Maas, RJ.M. 1977. Costoideae (additions). Flora Neotropica Monogr. 18: 162-218.
Maas, RJ.M. 1979. Zingiberaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 416-441.
Maas, P.J.M. 1985. Zingiberaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 1: 29-67.
AM BO
BO
AM
BO
AM BO
AM BO
AM BO
GU su FG
GU su
GU su FG
GU
GU
su FG
GU su
BO
BO
BO
Checklist of the Plants of the Guiana Shield 87
Maas, P. & H. Maas-van de Kamer. 1997. Costaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D.
Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and
Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1: 243-244.
Cyclanthaceae
R. Eriksson, G Harling
Asplundia brachyphylla Harling
Asplundiafanshawei (Maguire) Harling
Asplundiaflavovaginata Harling AM BO
Asplundia glandulosa (Gleason) Harling
Asplundia gleasonii Harling
Asplundia guianensis Harling
Asplundia heteranthera Harling
Asplundia luetzelburgii Harling
Asplundia maguirei Harling
Asplundia mohtziana (Klotzsch) Harling DA
Asplundia neblinae Harling
Asplundia nilssonii Harling
Asplundia vaupesiana Harling
,
Asplundia venezuelensis Harling
Asplundia xiphophylla Harling
Carludovica palmata Ruiz & Pav. PQ?
Cyclanthus bipartitus Poit. AM
Dicranopygium angustissimum (Sandwith) Harling
Dicranopygium bolivarense Harling
Dicranopygium imeriense Harling
Dicranopygium nanum (Gleason) Harling
Dicranopygium omichlophilum R.E. Schult. ex Harling
Dicranopygium pygmaeum (Gleason) Harling
Dicranopygium pygmaeum subsp. fimbrlatum (Maguire) Harling
Dicranopygium pygmaeum subsp. pygmaeum
Dicranopygium robustum Harling
Dicranopygium yacu-sisa Harling
Evodianthus funifer (Poit.) Lindm.
Evodianthus funifer sub^p. funifer
Evodianthus funifer subsp. trailianus (Drude) Harling
Ludovia lancifolia Brongn.
Sphaeradenia amazonica Harling
Sphaeradenia duidae Harling
Sphaeradenia lemaensis Harling
Stelestylis stylaris (Gleason) Harling
Stelestylis surinamensis Harling
Thoracocarpus bissectus (Veil.) Harling AM BO DA
Synonyms and Misidentifications
Carludovica acuminata (Ruiz & Pav.) Harling [not in flora area]
Carludovica angustissima Sandwith =Dicranopygium angustissimum
Carludovica bracteosa Gleason =Thoracocarpus bissectus
Carludovica coronata Gleason =Evodianthus funifer subsp. funifer
Carludovica disticha Neum. «Poit.?» =Ludovia lancifolia
Carludovicafanshawei Maguire =Asplundiafanshawei
Carludovicafimbriata Maguire =Dicranopygium pygmaeum subsp. fimbriatum
Carludovica glandulosa Gleason =Asplundia glandulosa
Carludovica kegeliana Lem. =Thoracocarpus bissectus
Carludovica latifrons Drude [not in flora area] =Asplundia latifrons (Drude) Harling
Carludovica moritziana Klotzsch =Asplundia moritziana
Carludovica nana Gleason ^Dicranopygium nanum
Carludovica pygmaea Gleason ^Dicranopygium pygmaeum
Carludovica sarmentosa Sagot ex Drude =Thoracocarpus bissectus
Carludovica stylaris Gleason =Stelestylis stylaris
Cyclanthus bipartitus var. gracilis Drude =Cyclanthus bipartitus
Cyclanthus cristatus Klotzsch =Cyclanthus bipartitus
Cyclanthus plumieri Poit. =Cyclanthus bipartitus
AM
AM BO
AM
AM
AM BO
AM
AM BO
BO
AM BO
DA GU SU FG
GU
GU
GU SU FG
SU FG
GU SU FG
DA GU su FG
DA GU su FG
GU su FG
GU su FG
GU
SU FG?
GU su FG
Checklist of the Plants of the Guiana Shield
Ludovia crenifolia Drude =Ludovia lancifolia
Ludovia fiinifera Poit. =Evodianthus funifer
Sphaeradenia maguirei Harling =Sphaeradenia duidae
Sphaeradenia rubescens Harling =Sphaeradenia duidae
References
Harling, G. 1958. Monograph of the Cyclanthaceae. Acta Horti Berg. 18: 1-428.
Harling, G. 1972. Cyclanthaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 9. Mem. New York Bot.
Gard. 23: 1-832.
Harling, G. & R. Eriksson. 1998. Cyclanthaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana
4:471-486.
Mori, S.A. 1997. Cyclanthaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1 : 244-249.
Cyperaceae
K. Camelbeke, P. Goetghebeur, M.S. Gonzalez-Elizondo, D.M. Keams, R. Krai, A.A. Reznicek,
D.A. Simpson, M.T. Strong, W. W. Thomas, GC. Tucker
Abildgaardia ovata (Burm. f.) Krai
Ascolepis brasiliensis (Kunth) Benth. ex C.B. Clarke
Becquerelia cymosa Brongn.
Becquerelia cymosa subsp. cymosa
Becquerelia cymosa subsp. merkeliana (Nees) T. Koyama
Becquerelia tuberculata (Bock.) H. Pfeiff.
Bisboeckelera irrigua (Nees) Kuntze
Bisboeckelera longifolia (Rudge) Kuntze
Bisboeckelera microcephala (Bock.) T. Koyama
Bulbostylis aturensis (Maury) C.B. Clarke
Bulbostylis barbata (Rottb.) C.B. Clarke
Bulbostylis capillaris (L.) Kunth ex C.B. Clarke
Bulbostylis capillaris var. capillaris
Bulbostylis capillaris var. major Uittien
Bulbostylis circinata (Nees) C.B. Clarke
Bulbostylis conifera (Kunth) C.B. Clarke
Bulbostylis fasciculata Uittien
Bulbostylis fluviatilis Krai & Davidse
Bulbostylis hispidula (Vahl) R. Haines
Bulbostylis junciformis (Kunth) C.B. Clarke
Bulbostylisjuncoides (Vahl) Kiik. ex Osten
Bulbostylis lanata (Kunth) C.B. Clarke [nom. cons]
Bulbostylis leucostachya (Kunth) C.B. Clarke
Bulbostylis medusae Camelbeke, Goetgh. & O. Huber, sp. nov. ined. [B.
sp "A" in FVG]
Bulbostylis paradoxa (Spreng.) Lindm.
Bulbostylis schomburgkiana (Steud.) M.T. Strong [type erroneously reported
from Guyana]
Bulbostylis stenocarpa Kuk.
Bulbostylis surinamensis Uittien
Bulbostylis svensoniana Steyerm.
Bulbostylis tenuifolia (Rudge) J.F. Macbr.
Bulbostylis truncata (Nees) M.T. Strong [expected in FG]
Bulbostylis vestita (Kunth) C.B. Clarke
Bulbostylis vestita var. glabrescens Bock.
Bulbostylis vestita var. vestita
Calyptrocarya bicolor (H. Pfeiff.) T. Koyama
Calyptrocarya glomerulata (Brongn.) Urb.
Calyptrocarya luzuliformis T. Koyama
Calyptrocarya monocephala Hochst. ex Steud.
Calyptrocarya poeppigiana Kunth [records may be referable to another
species]
Carex bonplandii Kunth GU?
Carexjamesonii Boott BO
Carex marahuacana Reznicek AM
BO GU
BO GU SU
AM BO DA GU su FG
AM BO DA GU SU FG
AM DA GU su FG
AM BO GU su
AM GU
GU su FG
GU su FG
AM
su
AM BO DA GU su FG
GU su
su
FG
GU su FG
BO GU su FG
GU su FG
AM
AM BO GU su FG
AM BO DA GU su FG
AM BO GU FG
AM
AM
AM
BO
BO
GU su FG
AM BO GU su FG
GU SU FG
GU SU FG?
BO DA
AM BO GU SU FG
AM BO GU su
AM BO su
su
su
FG
FG
AM BO GU su FG
AM BO DA GU su FG
AM BO DA GU su
AM BO GU su FG
AM BO GU su FG
Checklist of the Plants of the Guiana Shield 89^
Carex neblinensis Reznicek AM
Carex polystachya Sw. ex Wahlenb.
Carex porrecta Reznicek & Camelbeke
Carex roraimensis Steyerm.
Carex tamana Steyerm.
Cephalocarpus confertiis Gilly
Cephalocarpus dracaemila Nees
Cephalocarpus lineariifolius Gilly
Cephalocarpus obovoideus T. Koyama
Cephalocarpus rigidus Gilly ex Gleason & Killip
Cladium costatum Steyerm. [Mt. Roraima]
Cladium mariscus Pohl
Cyperus aggregatus (Willd.) Endl.
Cyperus altsonii Kiik.
Cyperus amabilis Vahl
Cyperus articulatus L.
Cyperus articulatus var. articulatus
Cyperus articulatus var. nodosus (Willd.) Kiik.
Cyperus bipartitus Torr.
Cyperus camphoratus Liebm.
Cyperus capillifolius A. Rich.
Cyperus chlorocephalus (C.B. Clarke) Kiik.
Cyperus comosus Poir.
Cyperus compressus L.
Cyperus cornelii-ostenii Kiik.
Cyperus croceus Vahl
Cyperus cuspidatus Kunth
Cyperus difformis L.
Cyperus digitatus Roxb.
Cyperus distans L. f.
Cyperus drummondii Torr. & Hook.
Cyperus elegans L.
Cyperus eragrostis Lam.
Cyperus esculentus L.
Cyperus esculentus var. leptostachyus Bock.
Cyperus felipponei Kiik.
Cyperus filifolius Willd. ex Kunth
Cyperus filiformis Sw.
Cyperusflavescens L.
Cyperus flavicomus Michx.
Cyperusfugax Liebm.
Cyperus gardneri Nees
Cyperus gayi (C.B. Clarke) Kuk.
Cyperus giganteus Vahl
Cyperus grandisimplex C.B. Clarke
Cyperus guianensis (C.B. Clarke) Kiik.
Cyperus haspan L. [subsp. haspan?]
Cyperus haspan subsp. Juncoides (Lam.) Kiik.
Cyperus haspan var. riparius (Nees) Kiik.
Cyperus hayesii (C.B. Clarke) Standi.
Cyperus hermaphroditus (Jacq.) Standi.
Cyperus hoppiifolius Uittien
Cyperus imbricatus Retz.
Cyperus imbricatus var. elongatus (Bock.) Kuk.
Cyperus imbricatus Retz. var. imbricatus
Cyperus involucratus Rottb. [escaped from cultivation]
Cyperus iria L. [introduced]
Cyperus kappleri Hochst. ex Steud.
Cyperus laxus Lam.
Cyperus ligularis L.
Cyperus luzulae (L.) Rottb. ex Retz.
Cyperus megapotamicus Kunth
Cyperus meridionalis Barros
Cyperus meyenianus Kunth
BO GU
BO
BO GU
BO
AM BO
BO
AM
AM
AM BO GU su
AM BO GU
su?
AM BO DA GU
GU
su FG
AM BO DA GU su
AM BO DA GU? su FG
AM BO
BO
DA GU
GU?
su FG
FG
AM BO GU
GU?
GU FG
GU su FG
DA GU su FG
BO DA GU su
AM BO GU su FG
AM BO GU
GU
su FG
BO DA GU su
AM DA GU
GU
GU
su
su
su
su
FG
AM
AM
AM BO DA
GU
AM
BO
BO
GU
GU
AM BO DA
GU FG
BO DA GU SU FG
BO
GU SU
AM BO DA GU SU FG
GU SU FG
GU FG
FG
BO GU FG?
AM BO SU FG
AM BO DA GU
GU
AM BO DA GU
GU
SU
BO GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU
SU
FG
AM BO
BO DA
GU
90 Checklist of the Plants of the Guiana Shield
Cyperus miliifolius Poepp. & Kunth
Cyperus mutisii (Kunth) Griseb.
Cyperus odoratus L.
Cyperus oxylepis Nees ex Steud.
Cyperus polystachyos Rottb.
Cyperus procerus Rottb.
Cyperus prolixus Kunth
Cyperus pseudodistans Uittien
Cyperus pumilus L.
Cyperus reflexus Vahl
Cyperus rofundus L.
Cyperus schomburgkianus Nees
Cyperus simplex Kunth
Cyperus sphacelatus Rottb.
Cyperus subumbellatus Kiik.
Cyperus surinamensis Rottb. [var. surinamensis7]
Cyperus surinamensis var. lutescens Bock.
"
Cyperus swartzii (A. Dietr.) Bock. & Kiik.
Cyperus tenuis Sw.
Cyperus tenuispica Steud. [sparingly naturalized Asian species]
Cyperus thyrsiflorus Jungh.
Cyperus trailii C.B. Clarke
Cyperus unicolor Bock.
Cyperus unioloides R. Br.
Cyperus virens Michx.
Didymiandrum stellatum (Bock.) Gilly
Diplacrum africanum (Benth.) C.B. Clarke
Diplacrum capitatum (Willd.) Bock.
Diplacrum guianense (Nees) T. Koyama
Diplasia karatifolia Rich.
Egleria fluctuans L.T. Eiten
Eleocharis acutangula (Roxb.) Schult.
Eleocharis alveolatoides S. Gonzalez & Reznicek
Eleocharis amazonica C.B. Clarke
Eleocharis atrospiculata S. Gonzalez & Reznicek
Eleocharis ayacuchensis S. Gonzalez & Reznicek
Eleocharis capillacea Kunth
Eleocharis debilis Kunth [f. anomala S. Gonzalez & Reznicek, found only in
Amazonas Territory in the flora area]
Eleocharis eglerioides S. Gonzalez & Reznicek AM BO
Eleocharis elegans (Kunth) Roem. & Schult. DA
Eleocharis filiculmis Kunth
Eleocharis flavescens (Poir.) Urb.
Eleocharis geniculata (L.) Roem. & Schult. BO DA
Eleocharis cf. glauca Bock.
Eleocharis interstincta (Vahl) Roem. & Schult.
Eleocharis jelskiana Bock.
Eleocharis liesneri S. Gonzalez & Reznicek
Eleocharis maculosa (Vahl) Roem. & Schult.
Eleocharis minima Kunth [var. minima?] AM BO DA
Eleocharis minima var. bicolor (Chapm.) Svenson
Eleocharis minima var. minima BO DA
Eleocharis cf. minima 'A'
Eleocharis cf. minima 'B'
Eleocharis cf. minima 'C
Eleocharis mitrata (Griseb.) C.B. Clarke
Eleocharis montana (Kunth) Roem. & Schult.
Eleocharis mutata (L.) Roem. & Schult. DA
Eleocharis nana Kunth
Eleocharis nigrescens (Nees) Steud.
Eleocharis aff nigrescens 'A'
Eleocharis aff. nigrescens 'B'
Eleocharis pachystyla (C. Wright) C.B. Clarke
Eleocharis plicarhachis (Griseb.) Svenson [expected in Venezuelan Guayana]
AM BO GU
GU
su FG
FG
AM BO DA GU
GU
su FG
DA GU
GU
su FG
AM BO DA GU su
su
FG^
FG
BO
BO DA GU
GU
su FG
AM BO GU su FG
AM BO DA GU su
su
FG
AM BO DA GU su
su
FG
FG
AM
BO
BO
BO
GU
AM
BO
BO
GU su
su
BO su FG
AM BO GU
su
AM BO GU su FG
AM BO GU su FG
AM BO GU su FG
AM
AM BO
BO
GU su
AM BO
AM
AM
BO
BO GU
AM BO
BO
BO
AM BO
AM BO
BO
BO
AM BO
AM
BO
BO
BO
BO
BO
BO
BO
AM BO
BO
GU SU FG
GU su FG
GU su FG
GU su FG
GU
GU su FG
GU su FG
GU
GU FG'
GU
GU SU
SU
FG
GU SU FG
GU su
GU
GU su FG
GU su FG
Checklist of the Plants of the Guiana Shield 91
Eleocharis retwflexa (Poir.) Urb.
Eleocharis sellowiana Kunth
Eleocharis sp. A
Eleocharis sp. B [Eleocharis retroflexa complex]
Eleocharis sp. C
Eleocharis sqiiamigera Svenson
Eleocharis sqiiamigera subsp. straminea S. Gonzalez & Reznicek
Eleocharis steyermarkii S. Gonzalez & Reznicek
Eleocharis subfoliata C.B. Clarke
Eleocharis venezuelensis S. Gonzalez & Reznicek
Everardia angusta N.E. Br.
Everardia debilis T. Koyama & Maguire
Everardia diffusa T. Koyama & Maguire
Everardia disticha T. Koyama & Maguire
Everardia flexifolia (Gilly) T. Koyama & Maguire
Everardia lanata T. Koyama & Maguire
Everardia longifolia Gilly ex Gleason & Killip
Everardia maguireana T. Koyama [Sierra de la Neblina]
Everardia montana Ridl. ex Thum
Everardia montana subsp. duidae (Gilly) T. Koyama & Maguire
Everardia montana subsp. glaucifolia (Gilly) T. Koyama & Maguire
Everardia montana subsp. guaiquinimae (Schnee) T. Koyama & Maguire
Everardia montana subsp. montana
Everardia montana subsp. ptariensis (Gilly) T. Koyama & Maguire
Everardia montana subsp. velutina T. Koyama
Everardia recurvigluma T. Koyama & Maguire [Mt. Roraima]
Everardia surinamensis Gilly
Everardia vareschii Maguire
Exochogyne amazonica C.B. Clarke
Exochogyne megalorrhyncha Tutin
Fimbristylis aestivalis (Retz.) Vahl
Fimbristylis annua (All.) Roem. & Schult.
Fimbristylis aspera Bock.
Fimbristylis autumnalis (L.) Roem. & Schult.
Fimbristylis complanata (Retz.) Link
Fimbristylis cymosa R. Br.
Fimbristylis decipiens Krai
Fimbristylis dichotoma (L.) Vahl
Fimbristylis dipsacea (Rottb.) C.B. Clarke
Fimbristylis ferruginea (L.) Vahl
Fimbristylis limosa Poepp. & Kunth
Fimbristylis littoralis Gaudich.
Fimbristylis quinquangularis (Vahl) Kunth
Fimbristylis spadicea (L.) Vahl
Fimbristylis squarrosa Vahl
Fimbristylis vahlii (Lam.) Link
Fuirena robusta Kunth
Fuirena simplex Vahl
Fuirena umbellata Rottb.
Hypolytrum amplum Poepp. & Kunth
Hypolytrum jenmanii C.B. Clarke
Hypolytrum jenmanii subsp. jenmanii
Hypolytrum jenmanii subsp. serratifolium (Uittien) T. Koyama
Hypolytrum laxum Kunth
Hypolytrum leptocalamum M. A Ives & W.W. Thomas
Hypolytrum longifolium (Rich.) Nees
Hypolytrum longifolium subsp. longifolium
Hypolytrum longifolium subsp. nicaraguense (Liebm.) T. Koyama
Hypolytrum longifolium subsp. rubescens (Huber ex C.B. Clarke) T. Koyama AM
Hypolytrum longifolium subsp. sylvaticum (Poepp. & Kunth) T. Koyama
Hypolytrum nudum C.B. Clarke
Hypolytrum pallidiceps S.S. Hooper & T. Koyama
Hypolytrum pulchrum (Rudge) H. Pfeiff.
Hypolytrum schraderianum Nees
AM BO GU su FG
BO GU FG
BO
AM
BO
AM BO
AM BO
BO
AM BO GU su
AM BO
AM BO GU
AM GU
AM BO
BO GU
AM
BO
BO
AM
AM BO GU su
AM BO
AM BO GU
AM
AM BO
BO
GU
AM
BO GU?
AM BO
BO
GU su
AM BO GU
GU
su FG
AM BO
AM BO GU
GU
su FG
DA GU? su FG
AM BO DA GU su FG
BO DA GU su FG
FG
AM BO
BO
DA GU su FG
GU su FG
GU
AM BO DA GU su FG
DA GU su FG
DA GU su FG
BO
AM BO DA GU su FG
GU su FG
FG
AM BO DA GU su FG
AM BO GU su FG
BO DA GU su FG
BO DA GU su FG
GU su FG
AM BO
AM BO GU
AM BO DA GU su FG
AM BO DA
DA
GU su FG
DA GU
AM BO DA GU su FG
BO GU
GU
AM BO GU su FG
AM BO GU FG
92 Checklist of the Plants of the Guiana Shield
Hypolytrum sphaerostachyum Bock.
Hypolytmm spongiosum T. Koyama
Hypolytrum stemonifolium T. Koyama
Hypolytrum striatum Poepp. & Kunth
Koyamaea neblinensis W.W. Thomas & Davidse
Kyllinga brevifolia Rottb.
Kyllinga nemoralis (J.R. Forst. & G. Forst.) Dandy ex Hutch.
Kyllinga odorata Vahl
Kyllinga pumila Michx.
Kyllinga pungens Link
Kyllinga vaginata Lam.
Lagenocarpus celiae T. Koyama & Maguire
Lagenocarpus eriopodus T. Koyama & Maguire
Lagenocarpus glomerulatus Gilly
Lagenocarpus guianensis Lindl. & Nees ex Nees
Lagenocarpus guianensis subsp. guianensis
Lagenocarpus guianensis subsp. hypochaeius T. Koyama
Lagenocarpus pendulus T. Koyama
Lagenocarpus rigidus (Kunth) Nees
Lagenocarpus rigidus subsp. rigidus
Lagenocarpus rigidus subsp. tremulus (Nees) T. Koyama & Maguire
Lagenocarpus sabanensis Gilly
Lagenocarpus venezuelensis Davidse
Lagenocarpus verticillatus (Spreng.) T. Koyama & Maguire
Lipocarpha humboldtiana Nees
Lipocarpha mexicana Liebm.
Lipocarpha micrantha (Vahl) G.C. Tucker
Lipocarpha salzmanniana Steud.
Lipocarpha schomburgkii (Friedl.) G.C. Tucker
Machaerina ayangannensis M.T. Strong
Mapania assimilis T. Koyama
Mapania assimilis subsp. guianensis D.A. Simpson
Mapania aturensis D.A. Simpson
Mapania cuatrecasasii T. Koyama
Mapania effusa (C.B. Clarke) T. Koyama
Mapania imeriensis (R. Gross) T. Koyama
Mapania insignis Sandwith
Mapania macrophylla (Bock.) H. Pfeiff.
Mapania maguireana T. Koyama & Steyerm.
Mapania micrococca (T. Koyama) D.A. Simpson
Mapania neblina D.A. Simpson
Mapania paradoxa J. Raynal
Mapania pedunculata D.A. Simpson
Mapania pycnocephala (Benth.) Benth.
Mapania pycnocephala subsp. fluviatiIis (Sandwith) T. Koyama
Mapania pycnocephala subsp. pycnocephala
Mapania pycnostachya (Benth.) T. Koyama
Mapania rionegrensis D.A Simpson
Mapania steyermarkii T. Koyama
Mapania surinamensis Uittien
Mapania sylvatica Aubl.
Mapania sylvatica subsp. sylvatica
Mapania tepuiana (Steyerm.) T. Koyama
Mapania theobromina D.A. Simpson
Oreobolus goeppingeri Suess.
Oxycaryum cubense (Poepp. & Kunth) Palla
Pleurostachys guianensis Uittien
Pleurostachys puberula Bock.
Remirea maritima Aubl.
Rhynchocladium steyermarkii (T. Koyama) T. Koyama
Rhynchospora agostiniana T. Koyama
Rhynchospora albescens (Miq.) Kiik.
Rhynchospora albida (Nees) Bock.
Rhynchospora albomarginata Kiik.
AM GU
GU
AM
AM
BO GU SU
SU
FG
FG?
AM BO DA GU SU FG
AM BO DA GU SU FG
GU SU FG
GU SU FG
AM GU
AM
AM BO GU
AM BO DA GU SU FG
AM BO DA GU SU FG
AM
AM
AM BO GU SU FG
AM BO GU
AM BO GU SU FG
AM BO SU FG
AM
BO GU SU
AM BO GU SU
AM BO
AM BO GU
GU
SU FG
AM GU
GU
FG
FG
AM
GU
BO GU FG
AM GU
GU
SU
BO GU SU FG
AM
BO
GU
GU
AM
FG
AM
AM BO GU SU FG
AM BO GU FG
AM
AM
AM BO
BO GU
GU SU
BO GU SU FG
BO GU SU FG
BO GU FG?
AM
AM
BO DA GU SU
SU
FG
AM BO GU
SU
FG
FG
AM BO GU
AM
AM GU SU FG
AM GU
BO GU FG
Checklist of the Plants of the Guiana Shield 93
Rhynchospora amazonica Poepp. & Kunth
Rhynchospora amazonica subsp. amazonica
Rhynchospora amazonica subsp. guianensis (Kuk.) T. Koyama
Rhynchospora angustipaniculata M.T. Strong
Rhynchospora arenicola Uittien
Rhynchospora armerioides J. Presl & C. Presl
Rhynchospora ayangannensis M.T. Strong
Rhynchospora bakhiiisensis M.T. Strong
Rhynchospora barbata (Vahl) Kunth
Rhynchospora biflora Bock.
Rhynchospora bolivarana Steyerm.
Rhynchospora brachychaeta C. Wright
Rhynchospora brasiliensis Bock.
Rhynchospora brevirostris Griseb.
Rhynchospora brownii Roem. & Schult.
Rhynchospora brownii subsp. americana Guaglianone
Rhynchospora cajennensis Bock.
Rhynchospora Candida (Nees) Bock.
Rhynchospora capillifolia W.W. Thomas
Rhynchospora capitata (Kunth) Roem. & Schult.
Rhynchospora caracasana (Kunth) Bock.
Rhynchospora cariciformis Nees
Rhynchospora cephalotes (L.) Vahl
Rhynchospora chimantensis W.W. Thomas
Rhynchospora ciliata (G. Mey) Kiik.
Rhynchospora comata (Link) Roem. & Schult.
Rhynchospora contracta (Nees) J. Raynal
Rhynchospora cordatachenia M.T. Strong
Rhynchospora corymbosa (L.) Britton
Rhynchospora curvula Griseb.
Rhynchospora dentinux C.B. Clarke
Rhynchospora divaricata (Ham.) M.T. Strong
Rhynchospora donselaarii M.T. Strong
Rhynchospora duckei R. Gross
Rhynchospora diddae Steyerm.
Rhynchospora eburnea Krai & W.W. Thomas
Rhynchospora elegantula Maury
Rhynchospora emaciata (Nees) Bock.
Rhynchospora exaltata Kunth
Rhynchospora exaltata var. microcephala Kiik.
Rhynchospora eximia (Nees) Bock.
Rhynchosporafallax Uittien
Rhynchospora fascicidaris (Michx.) Vahl
Rhynchosporafascicularis suhsp. fascicularis
Rhynchosporafdiformis Vahl
Rhynchospora gigantea Link
Rhynchospora globosa (Kunth) Roem. & Schult.
Rhynchospora harperi Small
Rhynchospora hassleri C.B. Clarke
Rhynchospora hieronymii Bock.
Rhynchospora hieronymii subsp. hieronymii
Rhynchospora hirsuta (Vahl) Vahl
Rhynchospora holoschoenoides (Rich.) Herter
Rhynchospora imeriensis (Kuk.) W.W. Thomas
Rhynchospora immensa Kiik.
Rhynchosporajunciformis (Kunth) Bock.
Rhynchospora lechleri Steud. ex Bock.
Rhynchospora lociiples C.B. Clarke
Rhynchospora longibracteata Bock.
Rhynchospora macrochaeta Steud. ex Bock.
Rhynchospora maguireana T Koyama
Rhynchospora mariscidus Nees
Rhynchospora mexicana (Liebm.) Steud.
Rhynchospora montana (Uittien) H. Pfeiff.
AM BO
AM BO
AM BO
AM BO
AM
AM BO
AM BO
AM BO
AM BO
AM BO
BO
BO
AM BO
AM BO
BO
AM
AM BO
AM BO
AM BO
BO
AM BO
AM BO
BO
AM BO
BO
AM BO
BO
AM
BO
AM BO
AM BO
AM BO
AM BO
BO
BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM
AM BO
AM BO
BO
AM BO
AM BO
AM
AM BO
AM
DA
GU SU FG
GU SU FG
GU FG?
GU
GU SU FG
GU SU
GU
SU
GU SU FG
GU
GU
GU
GU SU FG
GU FG
GU SU FG?
GU
GU SU
GU SU FG
GU SU FG
GU SU FG
FG
FG
GU SU FG
GU SU FG
FG
GU SU
SU
FG
GU SU
GU
GU
GU SU FG
GU
GU
SU
SU
GU SU FG
GU SU FG
GU SU FG
GU SU
GU SU FG
GU SU FG
GU SU FG
GU
GU SU FG
GU
GU
GU SU
SU
94 Checklist of the Plants of the Guiana Shield
Rhynchospora nervosa (Vahl) Bock.
Rhynchospora nitens (Vahl) A. Gray
Rhynchospora pallida (Nees) Steud.
Rhynchospora papulosa W.W. Thomas
Rhynchospora pilosa (Kunth) Bock.
Rhynchospora planifolia (T. Koyama) W.W. Thomas
Rhynchospora polyphylla Vahl
Rhynchospora pubera (Vahl) Bock.
Rhynchospora pubera subsp. /jcrrvw/a W.W. Thomas
Rhynchospora pubera swbs^. pubera
Rhynchospora pubisquama M.T. Strong
Rhynchospora radicans (Schltdl. & Cham.) H. Pfeiff.
Rhynchospora radicans subsp. microcephala (Bertero ex Spreng.) W.W.
Thomas
Rhynchospora reptans (Rich.) Bock.
Rhynchospora rigidifolia (Gilly) T. Koyama
Rhynchospora riparia (Nees) Bock.
Rhynchospora robusta (Kunth) Bock.
Rhynchospora roraimae Kiik. [Mt. Roraima]
Rhynchospora rugosa (Vahl) Gale
Rhynchospora ruiziana Bock.
Rhynchospora rupicola M.T. Strong
Rhynchospora sanariapensis Steyerm.
Rhynchospora saxisavannicola M.T. Strong
Rhynchospora schomburgkiana (Bock.) T. Koyama
Rhynchospora sp. A
Rhynchospora sp. B
Rhynchospora sp. C
Rhynchospora sp. D
Rhynchospora sp. E
Rhynchospora splendens Lindm.
Rhynchospora spruceana C.B. Clarke
Rhynchospora steyermarkii T. Koyama [expected in GU]
Rhynchospora subdicephala T. Koyama
Rhynchospora sublanata Lindeman & Donsel.
Rhynchospora subplumosa C.B. Clarke
Rhynchospora tenella (Nees) Bock.
Rhynchospora tenerrima Nees ex Spreng.
Rhynchospora tenuis Link
Rhynchospora tenuis subsp. austrobrasiliensis T. Koyama
Rhynchospora tenuis subsp. tenuis
Rhynchospora tepuiana Steyerm.
Rhynchospora trichochaeta C.B. Clarke
Rhynchospora trichophora Nees
Rhynchospora triflora Vahl
Rhynchospora trispicata (Nees) Schrad. ex Steud.
Rhynchospora tuerckheimii C.B. Clarke ex Kiik.
Rhynchospora uniflora Bock.
Rhynchospora unisetosa T. Koyama
Rhynchospora velutina (Kunth) Bock.
Rhynchospora watsonii (Britton) Davidse
Schoenoplectus maritimus (L.) Lye
Schoenoplectus robustus (Pursh) M.T. Strong
Scleria acanthocarpa Bock.
Scleria amazonica Camelbeke, M. Strong & Goetgh.
Scleria anceps Liebm.
Scleria bracteata Cav.
Scleria camaratensis Core
Scleria cerradicola T. Koyama
Scleria cyperina Willd. ex Kunth
Scleria cyperinoides C.B. Clarke
Scleria distans Poir.
Scleria eggersiana Bock.
Scleriaflagellum-nigrorum P.J. Bergius
AM BO GU
GU
su FG
AM
GU
GU
AM
AM BO GU
AM BO GU su FG
AM BO GU su
AM BO GU su
su
FG
GU su FG
GU su FG
GU su FG
AM BO
AM BO GU su FG
AM BO
AM BO GU
AM BO
BO
GU
GU
GU
su
su
FG
AM BO GU
su FG
AM GU
AM
AM BO GU
AM
BO
AM
GU
AM BO
BO
GU su FG?
AM
,
GU su
FG
AM BO GU su
AM BO GU su FG
BO GU su FG
AM BO GU
GU
su FG
AM BO GU
AM BO GU
AM BO GU su?
AM BO
BO GU su FG
AM BO
BO
BO
GU
GU
su FG
AM BO GU su FG
GU su
su
FG
FG?
DA GU su? FG?
FG
AM
GU su
AM BO
BO
GU
GU
su FG
AM BO GU su FG
AM BO
AM BO GU su FG
BO GU su
AM BO DA GU su FG
GU su FG
BO GU
GU
su
su
FG
AM BO
BO
DA GU
GU
su FG
FG?
FG
AM BO DA GU su FG
AM BO GU su
su
FG
FG
AM BO DA GU su FG
AM BO DA GU su FG
BO DA GU su FG
AM BO DA GU su
su
FG
AM
AM BO
AM BO
AM BO
Checklist of the Plants of the Guiana Shield 95
Scleria hirtella Sw. [records may be referable to Scleria distans Poir.]
Scleria interrupta Rich.
Scleria lacusths C. Wright
Scleria latifolia Sw.
Scleria lindleyana C.B. Clarke
Scleria lithosperma (L.) Sw.
Scleria longigluma Kiik.
Scleria macrogyne C.B. Clarke
Scleria macrophylla J. Presl & C. Presl
Scleria martii (Nees) Steud.
Scleria melaleuca Rchb. ex Schltdl. & Cham.
Scleria microcarpa Nees ex Kunth
Scleria mitis P.J. Bergius
Scleria muehlenbergii Steud.
Scleria neogranatensis C.B. Clarke
Scleria parallela C.B. Clarke
Scleria pinetorum Britton SU
Scleria plusiophylla Steud. GU
Scleria purdiei C.B. Clarke BO
Scleria ramosa C.B. Clarke SU
Scleria robusta Camelbeke & Goetgh.
Scleria scabra Willd.
Scleria secans (L.) Urb. DA
Scleria splitgerberiana Henrard ex Uittien
Scleria staheliana Uittien
Scleria stipitata Uittien
Scleria stipularis Nees
Scleria tenacissima Steud.
Scleria tepuiensis Core
Scleria triquetra M.T. Strong
Scleria cf. vaginata Steud.
Scleria vaginata Steud.
Scleria verticillata Muhl. ex Willd.
Scleria violacea Pilg.
Trilepis kanukuensis Gilly
Uncinia hamata (Sw.) Urb.
Websteria confervoides (Poir.) S.S. Hooper
Synonyms and Misidentifications
Abildgaardia monostachyos (L.) Vahl =Abildgaardia ovata
Ascolepis venezuelensis Schnee =Lipocarpha humboldtiana
Becquerelia bicolor H. Pfeiff. =Calyptrocarya bicolor
Becquerelia divaricata (Nees ex Bock.) Palla =Becquerelia cymosa subsp. merkeliana
Becquerelia glomerulata Brongn. =Calyptrocarya glomerulata
Becquerelia merkeliana Nees =Becquerelia cymosa subsp. merkeliana
Bisboeckelera angustifolia (Bock.) Kuntze =Bisboeckelera longifolia
Bolboschoenus maritimus (L.) Palla =Schoenoplectus maritimus
Bolboschoenus robustus (Pursh) Sojak ^Schoenoplectus robustus
Bulbostylis arenaria (Nees) Lindm. =Bulbostylis juncoides
Bulbostylis argentina Palla ^^Bulbostylis juncoides
Bulbostylis capillaris var. glabra Uittien =Bulbostylis truncata
Bulbostylis capillaris var. tenuifolia (Rudge) C.B. Clarke =Bulbostylis tenuifolia
Bulbostylis fendleri C.B. Clarke =Bulbostylis juncoides
Bulbostylis hirta (Thunb.) Svenson [misapplied to Bulbostylis hispidula]
Bulbostylis hirtella (Schrad. ex Schult.) Nees ex Urb. =Bulbostylis juncoides
Bulbostylis juncoides var. lorentzii (Bock.) Kuk. =Bulbostylis juncoides
Bulbostylis langsdorffiana (Kunth) C.B. Clarke =Bulbostylis juncoides
Bulbostylis papulosa Kiik. =Bulbostylis junciformis
Bulbostylis spadicea (Kunth) Kuk. =Bulbostylis paradoxa
Calyptrocaryafragifera Nees ex Kunth [nomen nudum] =Calyptrocarya glomerulata
Calyptrocarya intermedia C.B. Clarke =Calyptrocarya glomerulata
Calyptrostylis ceylanica (Nees) Nees =Rhynchospora triflora
Calyptrostylis longirostris Nees =Rhynchospora amazonica subsp. guianensis
Carex hamata Sw. = Uncinia hamata
Carex hermaphrodita Jacq. =Cyperus hermaphroditus
AM BO
AM BO
AM
BO
AM BO
AM
AM BO
AM BO
GU SU
GU SU FG
GU SU
SU FG
GU SU
GU SU FG
GU
su
FG
FG
GU su FG
FG
GU su FG
GU?
GU su? FG
96 Checklist of the Plants of the Guiana Shield
Carex ovata Burm. f. =Abildgaardia ovata
Cephalocarpus maguirei Gilly =Cephalocarpus rigidus
Cephalocarpus steyermarkii Gilly =Cephalocarpus rigidus
Cephaloschoenus ceylanicus Nees =Rhynchospora triflora
Chaetospora capitata Kunth =Rhynchospora capitata
Chaetospora globosa Kunth =Rhynchospora globosa
Chaetospora pterocarpa Kunth =Rhynchospora barbata
Cladium steyermarkii T. Koyama =Rhynchocladium steyermarkii
Cryptangium stellatum Bock. =Didymiandrum stellatum
Cyperus albomarginatus (Nees) Steud. =Cyperus flavicomus
Cyperus amabilis var. macrostachyus (Bock.) Kuk. =Cyperus amabilis
Cyperus amabilis var. oligostachyus (Kunth) Kiik. =Cyperus amabilis
Cyperus aurantiacus Kunth =Cyperus amabilis
Cyperus aureus Kunth =Cyperus amabilis
Cyperus aureus var. aurantiacus (Kunth) Bock. =Cyperus amabilis
Cyperus aureus var. oligostachyus (Kunth) Bock. ^Cyperus amabilis
Cyperus aureus x macrostachyus Bock. =Cyperus amabilis
Cyperus brevifolius (Rottb.) Endl. ex Hassk. =Kyllinga brevifolia
Cyperus cayennensis (Lam.) Britton ^Cyperus aggregatus
Cyperus chalaranthus J. Presl & C. Presl =Cyperus laxus
Cyperus corymbosus Rottb. =Cyperus articulatus
Cyperus densicaespitosus Mattf. & Kuk. =Kyllinga pumila
Cyperus diffusus Vahl =Cyperus laxus
Cyperus dissitiflorus Nees ex Torr. =Cyperus thyrsiflorus
Cyperus eggersii Bock. =Cyperus odoratus
Cyperus elegans auct. non L. =Cyperus laxus
Cyperusferax Rich. =Cyperus odoratus
Cyperus flavus (Vahl) Nees =Cyperus aggregatus
Cyperusfratemus Kunth =Cyperus reflexus
Cyperus globulosus Aubl. =Cyperus luzulae
Cyperus globulosus auct., non Aubl. =Cyperus croceus
Cyperus herndoniae G.C. Tucker =Cyperus camphoratus
Cyperus hirtus Thunb. =Bulbostylis hispidula
Cyperus huarmensis (Kunth) M.C. Johnst. =Cyperus aggregatus
Cyperus kyllingia Endl. =Kyllinga nemoralis
Cyperus macrocephalus Liebm. =Cyperus odoratus
Cyperus monostachyos L. =Abildgaardia ovata
Cyperus nodosus Willd. =Cyperus articulatus var. nodosus
Cyperus obtusatus (J. Presl & C. Presl) Mattf. & Kiik. =Kyllinga pungens
Cyperus oligostachyus Kunth =Cyperus amabilis
Cyperus pedunculatus (R. Br.) Kern =Remirea maritima
Cyperus planifolius Rich. [Not in flora area]
Cyperus rivularis Kunth =Cyperus bipartitus
Cyperus ruderalis Uittien =Cyperus trailii
Cyperus sesquiflorus (Torr.) Mattf. & Kiik. =Kyllinga odorata
Cyperus tolucensis Kunth =Cyperus laxus
Cyperus uncinatus Poir. [not in flora area]
Dichromena albida (Nees) Steud. =Rhynchospora albida
Dichromena brevicaulis Suess. =Rhynchospora duckei
Dichromena Candida (Nees) Ridl. =Rhynchospora Candida
Dichromena caracasana Kunth =Rhynchospora caracasana
Dichromena cariciformis (Nees) J.F. Macbr. =Rhynchospora cariciformis
Dichromena ciliata Vahl [nom. illeg., non Dichromena ciliata Pers. (1805)] =Rhynchospora ciliata
Dichromena ciliata var. cinnamomea Kiik. =Rhynchospora ciliata
Dichromena exaltata (Kunth) J.F. Macbr. =Rhynchospora exaltata
Dichromena filiformis (Vahl) Kunth =Rhynchosporafdiformis
Dichromenajunciformis Kunth =Rhynchosporajunciformis
Dichromena monostachya (Bock.) C.B. Clarke =Rhynchospora albescens
Dichromena montana Uittien =Rhynchospora montana
Dichromena nervosa Vahl =Rhynchospora nervosa
Dichromena persooniana Nees =Rhynchospora nervosa
Dichromena pilosa Kunth =Rhynchospora pilosa
Dichromena pubera Vahl =Rhynchospora pubera
Dichromena radicans Schltdl. & Cham. ^^Rhynchospora radicans
Checklist of the Plants of the Guiana Shield 91_
Dichromena repens Vahl =Rhynchospora reptans
Dichromena rigidifolia Gilly =Rhynchospora rigidifoHa
Dichromena robusta Kunth =Rhynchospora robusta
Dichromena squarrosa Willd. ex Link (Rapateaceae) =Duckea squarrosa
Dichromena tenella (Nees) Steud. =Rhynchospora tenella
Dichromena triflora (Vahl) Macbride =Rhynchospora triflora
Dichromena velutina Kunth =Rhynchospora vehitina
Didymiandrum flexifolium Gilly =Everardia flexifolia
Didymiandrum guaiquinimae Schnee =Didymiandriim stellatum
Diplacrum laevissimum (Kiik.) Uittien =Diplacrum guianense
Diplacrum longifolium (Griseb.) C.B. Clarke ex T. Durand & Schinz =Diplacrum capitatum
Diplacrum notopterum (Nees ex Steud.) C.B. Clarke =Diplacrum guianense
Diplasia pycnostachya Benth. =Mapania pycnostachya
Echinoschoenus beyrichii Nees =Rhynchospora exaltata
Eleocharis atropurpurea (Retz.) J. Presl & C. Presl [not in flora area]
Eleocharis bicolor Chapm. =EIeocharis minima var. bicolor
Eleocharis caribaea (Rottb.) S.F. Blake =Eleocharis geniculata
Eleocharis confervoides (Poir.) G. Tucker =Websteria confervoides
Eleocharis elata Bock. =Eleocharis plicarhachis
Eleocharis fistidosa (Poir.) Schult. [nom. superfl.] =Eleocharis acutangula
Eleocharis fistulosa (Poir.) Link [invalidly published] =Eleocharis acutangula
Eleocharis flaccida (Rchb. ex Spreng.) Urb. =Eleocharis flavescens
Eleocharisflaccida auct. non (Rchb. ex Spreng.) Urb. sensu Svenson =Eleocharis flavescens
Eleocharis homonyma Steud. =Eleocharis sellowiana
Eleocharis nodulosa (Roth) Schult. =Eleocharis montana
Eleocharis sagotii C.B. Clarke ^Eleocharis jelskiana
Eleocha?-is sagotii var. glochidiata C.B. Clarke =Eleocharis plicarhachis
Eleocharis sellowiana var. homonyma (Steud.) H. Pfeiff. =Eleocharis sellowiana
Eleocharis sulcata Nees [nomen nudum] =Eleocharis fdiculmis
Eleocharis sulcata Nees ex Bock. [nom. superfl.] =Eleocharis filiculmis
Ephippiorhynchium triflorum (Vahl) Nees =Rhynchospora triflora
Ephippiorhynchium trispicatum Nees =Rhynchospora trispicata
Everardia duidae Gilly =Everardia montana subsp. duidae
Everardia erecto-laxa T. Koyama & Maguire =Everardia angusta
Everardia glaucifolia Gilly =Everardia montana subsp. glaucifolia
Everardia gracilis Gilly =Everardia angusta
Everardia guaiquinimae Schnee =Everardia montana subsp. guaiquinimae
Everardia hitchcockii Lasser & Maguire =Everardia montana subsp. montana
Everardia montana var. pilosa Gilly =Everardia montana subsp. montana
Everardia ptariensis Gilly =Everardia montana subsp. ptariensis
Everardia revoluta Gilly ^Everardia montana subsp. montana
Everardia steyermarkii Gilly =Everardia angusta
Exochogyne steyermarkii Gilly =Exochogyne amazonica
Fimbristylis annua var. diphylla (Retz.) Kiik. =Fimbristylis dichotoma
Fimbristylis apus (A. Gray) S. Watson =Fimbristylis vahlii
Fimbristylis autumnalis var. complanata (Retz.) Barros =Fimbristylis complanata
Fimbristylis baldwiniana (Schult.) Torr. "^^Fimbristylis annua
Fimbristylis bufonia (Kunth) Alain =Bulbostylis tenuifolia
Fimbristylis capillaris var. pilosa Britton =Bulbostylis juncoides
Fimbristylis chirigota (C. Wright) Alain =Bulbostylis hispidula
Fimbristylis comata Nees =Fimbristylis squarrosa
Fimbristylis congesta Torr. =Fimbristylis vahlii
Fimbristylis cymosa subsp. spathacea (Roth) T. Koyama ^Fimbristylis cymosa
Fimbristylis dichotoma f. annua (All.) Owhi =Fimbristylis annua
Fimbristylis diphylla (Retz.) Vahl =Fimbristylis dichotoma
Fimbristylis divaricata Ham. =Rhynchospora divaricata
Fimbristylis frankii Steud. =Fimbristylis autumnalis
Fimbristylis glomerata (Retz.) Urb. =Fimbristylis cymosa
Fimbrist}'lis hirta (Kunth) Roem. & Schult. =Fimbristylis squarrosa
Fimbristylis hirta (Thunb.) Roem. & Schult. ex Kuk. [name has been misapplied to Bulbostylis hispidula] =Fimbristylis
squarrosa
Fimbristylis hispidula (Vahl) Kunth =Bulbostylis hispidula
Fimbristylis juncoides (Vahl) Alain =Bulbostylis juncoides
Fimbristylis laxa Vahl =Fimbristylis annua
98 Checklist of the Plants of the Guiana Shield
Fimbristylis miliacea (L.) Vahl [Guianas collections referable to F. littoralis] =Fimbristylis littoralis
Fimbristylis monostachyos (L.) Hassk. =Abildgaardia ovata
Fimbristylis obtusifolia (Vahl) Kunth ^Fimbristylis cymosa
Fimbristylis ovata (Burm. f.) J. Kern =Abildgaardia ovata
Fimbristylis papulosa (Kiik.) Alain =Bulbostylis jiinciformis
Fimbristylis polymorpha Bock. =Fimbrist)^lis dichotoma
Fimbristylis savannariim Alain =Biilbostylis juncoides
Fimbristylis schombiirgkiana Steud. =Bulbostylis schomburgkiana
Fimbristylis spadicea f. domingensis (Pers.) Kiik. =Fimbristylis spadicea
Fimbristylis spathacea Roth =Fimbristylis cymosa
Fimbristylis speciosa Rohde ex Spreng. =Fimbristylis spadicea
Fimbristylis verrucosa C. Presl =Fimbristylis annua
Fimbristylis vestita (Kunth) Hemsl. =Bulbostylis vestita
Fuirena paniculata L. f. =Fuirena umbellata
Fuirena tereticulmis C. Presl =Fuirena umbellata
Fuirena verticillata Spreng. =Lagenocarpus verticillatus
Haloschoenus albidus Nees =Rhynchospora albida
Haloschoenus caracasanus (Kunth) Nees =Rhynchospora caracasana
Haloschoenus contractus Nees =Rhynchospora contracta
Haloschoenus emaciatus Nees =Rhynchospora emaciata
Haloschoenus riparius Nees =Rhynchospora riparia
Haloschoenus tenellus Nees =Rhynchospora tenella
Haplostylis mexicana Liebm. =Rhynchospora mexicana
Hemicarpha micrantha (Vahl) Pax =Lipocarpha micrantha
Hemicarpha schomburgkii Friedl. =Lipocarpha schomburgkii
Hoppia irrigua Nees =Bisboeckelera irrigua
Hypolytrumfuscum Nees =Hypolytrum longifolium
Hypolytrum imeriense R. Gross =Mapania imeriensis
Hypolytrum irriguum Nees ^Hypolytrum longifolium
Hypolytrum macrophyllum Bock. =Mapania macrophylla
Hypolytrum micranthum Bock. =Hypolytrum strictum
Hypolytrum nicaraguense Liebman =Hypolytrum longifolium subsp. nicaraguense
Hypolytrum punctatum Nees =Hypolytrum longifolium subsp. sylvaticum
Hypolytrum pycnocephalum Benth. =Mapania pycnocephala
Hypolytrum rubescens Huber ex C.B. Clarke =Hypolytrum longifolium subsp. rubescens
Hypolytrum sandwithii Gilly =Mapania imeriensis
Hypolytrum scapigerum Nees ^Hypolytrum strictum
Hypolytrum sylvaticum Poepp. & Kunth =Hypolytrum longifolium subsp. sylvaticum
Hypolytrum tepuianum Steyerm. =Mapania tepuiana
Hypoporum micrococcum Liebm. =Scleria verticillata
Iria monostachya (L.) Kuntze =Abildgaardia ovata
Isolepis albescens Miq. =Rhynchospora albescens
Isolepis bufonia Kunth =Bulbostylis tenuifolia
Isolepis capillaris (L.) Roem. & Schult. =Bulbostylis capillaris
Isolepis conifera Kunth =Bulbostylis conifera
Isolepis dipsacea (Rottb.) Roem. & Schult. =Fimbristylis dipsacea
Isolepis hirta Kunth =Fimbristylis squarrosa
Isolepis hirtella Schrad. ex Schult. =Bulbostylis juncoides
Isolepis humboldtii Roem. & Schult. =Lipocarpha micrantha
Isolepisjunciformis Kunth =Bulbostylis junciformis
Isolepis lanata Kunth =Bulbostylis lanata
Isolepis langsdorffiana Kunth =Bulbostylis juncoides
Isolepis leucostachya Kunth =Bulbostylis leucostachya
Isolepis miliacea (L.) C. Presl =Fimbristylis littoralis
Isolepis paradoxa (Spreng.) Kunth =Bulbostylis paradoxa
Isolepis radiciflora Steud. =Bulbostylis capillaris
Isolepis vahlii (Lam.) Kunth =Fimbristylis vahlii
Isolepis vestita Kunth =Bulbostylis vestita
Isolepis willdenowii Roem. & Schult. =Fimbristylis complanata
Kyllinga cayennensis Lam. =Cyperus aggregatus
Kyllinga filiformis Sw. =Cyperus swartzii
Kyllinga monocephala Rottb., nom. illeg. =Kyllinga nemoralis
Kyllinga peruviana Lam. =Kyllinga vaginata
Kyllinga rigida Baldwin =Kyllinga vaginata
Checklist of the Plants of the Guiana Shield 99
Kyllinga sesquiflora Torr. =Kyllwga odorata
KyUinga squarrosa Baldwin =Cypenis aggregatiis
Lagenocarpus amazonicus Davidse =Lagenocarpus venezuelensis
Lagenocarpus amazonicus (C.B. Clarke) H. Pfeiff. =Exochogyne amazonica
Lagenocarpus kunthii (Miq.) Uittien =Lagenocarpus verticillatus
Lagenocarpus tremuhis Nees =Lagenocarpus rigidus subsp. tremulus
Lagenocarpus weigelti (Spreng.) Uittien =Lagenocarpus verticillatus
Leptoschoenus prolifer Nees =Rhynchospora schomburgkiana
Lipocarpha maculata auct. non (Michx.) Torrey 1836: sensu neotropical =Lipocarpha mexicana
Lipocarpha sellowiana Kunth =Lipocarpha humboldtiana
Mapania fluviatilis Sandwith =Mapania pycnocephala subsp. fluviatilis
Mapaniopsis effusa C.B. Clarke =Mapama effusa
Mapaniopsis micrococca T. Koyama =Mapania micrococca
Mariscus aggregatus Willd. =Cyperus aggregatus
Mariscus brizaeus (Vahl) C.B. Clarke =Cyperus planifolius
Mariscus cayennensis (Lam.) Urb. =Cyperus aggregatus
Mariscus chlorocephalus C.B. Clarke =Cyperus chlorocephalus
Mariscus cornelii-ostenii (Kiik.) T. Koyama =Cyperus cornelii-ostenii
Mariscus dissitiflorus (Torr.) C.B. Clarke =Cyperus thyrsiflorus
Mariscus elatus Kunth =Cyperus aggregatus
Mariscus flavus Vahl =Cyperus aggregatus
Mariscus flavus var. mayor Nees =Cyperus aggregatus
Mariscus gayi C.B. Clarke =Cyperus gayi
Mariscus guianensis C.B. Clarke =Cyperus guianensis
Mariscus laevis Kunth =Cyperus aggregatus
Mariscus ligularis (L.) Urb. =Cyperus ligularis
Mariscus mutisii Kunth =Cyperus mutisii
Mariscus pedunculatus (R. Br.) T. Koyama =Remirea maritima
Mariscus swartzii A. Dietr. =Cyperus swartzii
Mariscus tovarii Kunth =Cyperus hermaphroditus
Mariscus umbellatus (Rottb.) Vahl =Cyperus subumbellatus
Micropapyrus viviparoides Suess. =Rhynchospora schomburgkiana
Mitrospora cephalophora Nees =Rhynchospora exaltata
Mitrospora pallida Nees =Rhynchospora pallida
Oncostylis arenaria Nees =Bulbostylis juncoides
Oncostylis hispida Liebm. =Bulbostylis vestita
Oncostylis junciformis var. humboldtiana (Spreng.) Nees =Bulbostylis junciformis
Oncostylis paradoxa (Spreng.) Nees =Bulbostylis paradoxa
Oncostylis tenuifolia (Rudge) Nees =Bulbostylis tenuifolia
Oncostylis truncata Nees =Bulbostylis truncata
Oncostylis vestita (Kunth) Nees =Bulbostylis vestita
Oreobolus obtusangulus var. rubrovaginatus Kiik. =Oreobolus goeppingeri
Oreobolus obtusangulus subsp. rubrovaginatus (Kiik.) T. Koyama =Oreobolus goeppingeri
Platylepis brasiliensis Kunth ^Ascolepis brasiliensis
Pogonostylis squarrosus (Vahl) Bertol. =Fimbristylis squarrosa
Psilocarya Candida Nees =Rhynchospora Candida
Psilocarya robusta (Kunth) Nees =Rhynchospora robusta
Psilocarya rufa Nees =Rhynchospora velutina
Psilocarya velutina (Kunth) Nees =Rhynchospora velutina
Pteroscleria guianensis Nees =Diplacrum guianense
Pteroscleria longifolia Griseb. =Diplacrum capitatum
Pycreus albomarginatus Mart. & Schrad. ex Nees =Cyperus flavicomus
Pycreus capillifolius (A. Rich.) C.B. Clarke =Cyperus capillifolius
Pycreusflavescens (L.) Rchb. =Cyperus flavescens
Pycreus megapotamicus (Kunth) Nees ^Cyperus megapotamicus
Pycreus polystachyos (Rottb.) R Beauv. =Cyperus polystachyos
Pycreus pumilus (L.) Nees =Cyperus pumilus
Pycreus unioloides (R. Br.) Urb. =Cyperus unioloides
Rhynchospora amazonica var. guianensis Kuk. =Rhynchospora amazonica subsp. guianensis
Rhynchospora aristata var. immensa (Kiik.) Kuk. =Rhynchospora immensa
Rhynchospora arundinacea Bock. =Rhynchospora exaltata
Rhynchospora aurea Vahl ^Rhynchospora corymbosa
Rhynchospora baileyi Britton ex L.H. Bailey ^Rhynchospora albomarginata
Rhynchospora beyrichii (Nees) Steud. =Rhynchospora exaltata
100 Checklist of the Plants of the Guiana Shield
Rhynchospora borinquensis Britton =Rhynchospora marisculus
Rhynchospora buchii Urb. =Rhynchospora mexicana
Rhynchospora canaliculata Bock. =Rhynchospora riparia
Rhynchospora Candida var. pilosa Schnee =Rhynchospora Candida
Rhynchospora cariciformis var. trichophora (Nees) Kiik. =Rhynchospora trichophora
Rhynchospora cephalophora (Nees) Steud. =Rhynchospora exaltata
Rhynchospora cephalotes var. angustifolia Uittien =Rhynchospora pubisquama
Rhynchospora cephalotes var. imeriensis Kiik. =Rhynchospora imeriensis
Rhynchospora ceylanica (Nees) Kunth ^Rhynchospora triflora
Rhynchospora clarkei Rose =Rhynchospora brevirostris
Rhynchospora cyperoides (Sw.) Mart. =Rhynchospora holoschoenoides
Rhynchospora denticulata Huber =Rhynchospora trichochaeta
Rhynchospora distans Vahl =Rhynchospora fascicularis
Rhynchospora elegans Kiik. ^Rhynchospora eximia
Rhynchospora exaltata var. ovalis Kiik. =Rhynchospora exaltata
Rhynchosporafascicularis var. distans (Michx.) Chapm. [not in flora area] misidentifications =R. fascicularis subsp.
fascicularis^
Rhynchospora filifolia A. Gray [not in flora area. Identifications =R. harperi]
Rhynchosporaflexuosa var. tenerrima (Bock.) Kuk. =Rhynchospora tepuiana
Rhynchosporaflexuosa subsp. tepuiana (Steyerm.) T. Koyama ^^Rhynchospora tepuiana
Rhynchospora glauca Vahl =Rhynchospora rugosa
Rhynchospora graminea Uittien =Rhynchospora spruceana
Rhynchospora guianensis Lindeman & Donsel. =Rhynchospora trispicata
Rhynchospora heterocaulis C.B. Clarke =Rhynchospora barbata p.p. & R. holoschoenoides p.p.
Rhynchospora hispidula (Vahl) Bock. [nom. illeg.] ^^Rhynchospora divaricata
Rhynchospora igarapicola T. Koyama =Rhynchospora cariciformis
Rhynchosporajunciformis var. monocarpa Kiik. =Rhynchosporajunciformis
Rhynchospora karuaiana Steyerm. ^Rhynchospora triflora
Rhynchospora longispicata Bock. =Rhynchosporafdiformis
Rhynchospora macrochaeta var. ruiziana (Bock.) Kiik. =Rhynchospora ruiziana
Rhynchospora mexicana var. trichochaeta (C.B. Clarke) Kiik. =Rhynchospora trichochaeta
Rhynchospora micrantha Vahl [nom. illeg.] =Rhynchospora contracta
Rhynchospora minutiflora (Rich, ex Spreng.) Adams =Rhynchospora contracta
Rhynchospora monostachya Bock. =Rhynchospora albescens
Rhynchospora nervosa var. ciliata (G. Mey.) Kuk. ^Rhynchospora ciliata
Rhynchospora nervosa subsp. ciliata (G. Mey.) T. Koyama =Rhynchospora ciliata
Rhynchospora nervosa var. cinnamomea (Kuk.) Kiik. =Rhynchospora ciliata
Rhynchospora oxycephala C. Wright =Rhynchospora eximia
Rhynchospora patuligluma Lindm. =Rhynchospora pallida
Rhynchospora perrigida Bock. =Bulbostylis paradoxa
Rhynchospora persooniana (Nees) Griseb. =Rhynchospora nervosa
Rhynchospora pilosa subsp. arenicola (Uittien) T. Koyama =Rhynchospora arenicola
Rhynchospora podosperma C. Wright ^Rhynchosporafdiformis
Rhynchospora pringlei C.B. Clarke =Rhynchospora brevirostris
Rhynchospora prolifera (Nees) C.B. Clarke =Rhynchospora schomburgkiana
Rhynchospora psilocaryoides Griseb. =Rhynchospora eximia
Rhynchospora ptaritepuiana Steyerm. =Rhynchospora lechleri
Rhynchospora pterocarpa (Kunth) Roem. & Schult. =Rhynchospora barbata
Rhynchospora pungens Liebm. =Rhynchospora rugosa
Rhynchospora repens (Vahl) Bock. =Rhynchospora reptans
Rhynchospora rigidifolia subsp. capillacea T. Koyama ^Rhynchospora capillifolia
Rhynchospora rigidifolia suhsp. planifolia T. Koyama =Rhynchospora planifolia
Rhynchospora rufa (Nees) Bock. =Rhynchospora velutina
Rhynchospora setacea (Bergius) Bock. =Rhynchospora fallax
Rhynchospora sororopana Steyerm. =Rhynchospora biflora
Rhynchospora stellata (Lam.) Griseb. [not in flora area] =Rhynchospora colorata (L.) H. Pfeiffer
Rhynchospora stricta var. demerarensis (C.B. Clarke) Kiik. =Rhynchospora cajennensis
Rhynchospora tamana Steyerm. =Rhynchospora macrochaeta
Rhynchospora tenella f. haplostylis Uittien =Rhynchospora tenella
Rhynchospora tenella var. haplostylis (Uittien) Kiik. =Rhynchospora tenella
Rhynchospora tenerrima Bock. =Rhynchospora tepuiana
Rhynchospora tenerrima war. fallax (Uittien) Kuk. =Rhynchospora fallax
Rhynchospora tenerrima suhsp. fallax (Uittien) T. Koyama =Rhynchosporafallax
Rhynchospora tenuis subsp. riparia (Nees) T. Koyama =Rhynchospora riparia
Checklist of the Plants of the Guiana Shield 1 1
Rhynchospora tenuiseta C. Wright =Rhynchospora marisculus
Rhynchospora triceps Steud. =Rhynchospora exaltata
Rhynchospora trichochaeta var. venezuelensis Steyerm. =Rhynchospora subplumosa
Rhynchospora trichodes C.B. Clarke ^Rhynchospora divaricata
Rhynchospora venezolana Kiik. =Rhynchospora hirsuta
Rhynchospora viridUutea C.B. Clarke =Rhynchospora hassleri
Schoenus barbatus Vahl =Rhynchospora barbata
Schoenus ciliatus G. Mey. [nom. illeg., non Dichromena ciliata Pers. (1805)] =Rhynchospora ciliata
Schoenus comatus Link =Rhynchospora comata
Schoenus cyperoides Sw. =Rhynchospora holoschoenoides
Schoenusflavus Link (Rapateaceae) =Duckeaflava
Schoenus hirsutus Vahl =Rhynchospora hirsuta
Schoenus hispidulus Vahl =Rhynchospora divaricata
Schoenus holoschoenoides Rich. =Rhynchospora holoschoenoides
Schoenusjuncoides Vahl =Bulbostylis juncoides
Schoenus paradoxus Spreng. =Bulbostylis paradoxa
Schoenus rugosus Vahl ^Rhynchospora rugosa
Schoenus secans L. =Scleria secans
Schoenus setaceus RJ. Bergius =Rhynchospora fallax
Scirpidium nigrescens Nees =Eleocharis nigrescens
Scirpidium sulcatum Nees [nom. superfl.] =Eleocharis fdiculmis
Scirpus acutangulus Roxb. =Eleocharis acutangula
Scirpus aestivalis Retz. =Fimbristylis aestivalis
Scirpus annuus All. =Fimbristylis annua
Scirpus apus A. Gray =Fimbristylis vahlii
Scirpus aturensis Maury =Bulbostylis aturensis
Scirpus autumnalis L. =Fimbristylis autumnalis
Scirpus baldwinianus Schult. =Fimbristylis annua
Scirpus brachyphyllus Link =Bulbostylis capillaris
Scirpus bufonis (Kunth) Poir. =Bulbostylis tenuifolia
Scirpus capillaris L. =Bulbostylis capillaris
Scirpus caribaeus Rottb. =Eleocharis geniculata
Scirpus cephalotes L. =Rhynchospora cephalotes
Scirpus chirigota C. Wright =Bulbostylis hispidula
Scirpus complanatus Retz. =Fimbristylis complanata
Scirpus confervoides Poir. =Websteria confervoides
Scirpus corymbosus L. =Rhynchospora corymbosa
Scirpus cubensis Poepp. & Kunth =Oxycaryum cubense
Scirpus dichotomus L. =Fimbristylis dichotoma
Scirpus diphyllus Retz. =Fimbristylis dichotoma
Scirpus dipsaceus Rottb. =Fimbristylis dipsacea
Scirpus domingensis Pers. =Fimbristylis spadicea
Scirpus elegans Kunth =Eleocharis elegans
Scirpus fistulosus Poir. =Eleocharis acutangula
Scirpus flavescens Poir. =Eleocharis flavescens
Scirpus geniculatus L. =Eleocharis geniculata
Scirpus glomeratus Retz. =Fimbristylis cymosa
Scirpus hirtus Griseb. =Bulbostylis vestita
Scirpus hispidulus Vahl =Bulbostylis hispidula
Scirpus humboldtiana Spreng. =BulbostylisJunciformis
Scirpus interstinctus Vahl =Eleocharis interstincta
Scirpus lithospermus L. =Scleria lithosperma
Scirpus longifolius Rich. =Hypolytrum longifolium
Scirpus lorentzii Bock. =Bulbostylis juncoides
Scirpus luzulae L. =Cyperus luzulae
Scirpus maculosus Vahl =Eleocharis maculosa
Scirpus maritimus L. =Schoenoplectus maritimus
Scirpus maritimus var. robustus (Pursh) Kuk. =Schoenoplectus robustus
Scirpus maritimus fi macrostachyus Michx. =Schoenoplectus robustus
Scirpus micranthus Vahl ^Lipocarpha micrantha
Scirpus miliaceus L. =Fimbristylis littoralis
Scirpus mitratus Griseb. =Eleocharis mitrata
Scirpus montanus Kunth ^Eleocharis montana
Scirpus mucronulatus Michx. =Fimbristylis autumnalis
102 Checklist of the Plants of the Guiana Shield
Scirpus mutatus L. =Eleocharis mutata
Scirpus nodulosus Roth =Eleochahs montana
Scirpus obtusifolhis Vahl =Fimbristylis cymosa
Scirpus pachystylus C. Wright =Eleocharis pachystyla
Scirpus plicarhachis Griseb. =Eleocharis plicarhachis
Scirpus pulcher Rudge =Hypolytrum pulchrum
Scirpus quinquangularis Vahl =Fimbristylis quinquangularis
Scirpus retroflexus Poir. =Eleocharis retroflexa
Scirpus robustus Pursh =Schoenoplectus robustus
Scirpus schomburgkianus Bock. =Rhynchospora schomburgkiana
Scirpus spadiceus L. =Fimbristylis spadicea
Scirpus submersus C. Wright =Websteria confervoides
Scirpus sulcatus Roth =Eleocharis Jiliculmis
Scirpus tenuifolius Rudge =Bulbostylis tenuifolia
Scirpus triflorus (Vahl) Poir. =Rhynchospora triflora
Scirpus umbellatus (Rottb.) Kuntze =Fuirena umbellata
Scirpus vahlii Lam. =Fimbristylis vahlii
Scirpus vestitus (Kunth) Rchb. ex Bock. =Bulbostylis vestita
Scleria arundirtacea Kunth =Scleria latifolia
Scleria bracteata f. supragynacea H. Pfeiff. =Scleria bracteata
Scleria capitata Willd. =Diplacrum capitatum
Scleriaflagellum Sw. =Scleria secans
Scleria foliosa C. Wright, nom. illeg. ^Scleria microcarpa
Scleria grandis Core non Labill =Scleria robusta
Scleria hirtella auct. non Sw. =Scleria distans
Scleria latifolia Nees =Scleria latifolia
Scleria latifolia var. arundinacea (Kunth) H. Pfeiff. ^Scleria latifolia
Scleria liebmannii Steud. =Scleria verticillata
Scleria micrococca (Liebm.) Steud. =Scleria verticillata
Scleria minutiflora Rich, ex Spreng. [nom. inval.] =Rhynchospora contracta
Scleria nutans Willd. ex Kunth =Scleria distans
Scleria paludosa Poepp. & Kunth =Scleria macrophylla
Scleria pterota C. Presl [nomen nudum] =Scleria melaleuca
Scleria pterota var. melaleuca (Rchb. ex Schltdl. & Cham.) Uittien =Scleria melaleuca
Scleria pterota var. melanocarpa Nees [nomen nudum] =Scleria melaleuca
Scleria pyramidalis Hochst. ex Steud. [=Scleria stipularisl] =Scleria martii
Scleria reflexa Kunth =Scleria secans
Scleria reticularis Michx. [Not in flora area. Restricted to E. US]] "^Scleria muehlenbergii
Scleria rigida Kunth =Lagenocarpus rigidus
Scleria setacea Poir. =Scleria muehlenbergii
Scleria tenella Kunth =Scleria verticillata
Scleria tuberculata Bock. ^Becquerelia tuberculata
Scleria weigeltiana Schrad. ex Bock. =Scleria secans
Spermodon eximius Nees =Rhynchospora eximia
Spermodon filiformis (Vahl) Nees =Rhynchospora filiformis
Stenophyllus capillaris (L.) Britton =Bulbostylis capillaris
Stenophyllus paradoxus (Spreng.) Standi. =Bulbostylis paradoxa
Stenophyllus truncata (Nees) H. Pfeiff. =Bulbostylis truncata
Stenophyllus vestitus (Kunth) Britton =Bulbostylis vestita
Toruliniumferax (Rich.) Urb. =Cyperus odoratus
Torulinium odoratum (L.) S.S. Hooper =Cyperus odoratus
Trichelostylis complanata (Retz.) Nees =Fimbristylis complanata
Trichelostylis miliacea (L.) Nees =Fimbristylis littoralis
Trichelostylis mucronulata (Michx.) Torn =Fimbristylis autumnalis
Trichelostylis quinquangularis (Vahl) Nees =Fimbristylis quinquangularis
Trichelostylis stricta Nees =Bulbostylis schomburgkiana
Trichochaeta tenuis Steud. =Rhynchospora trichochaeta
References
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Tucker, G. C. 1987. The genera of Cyperaceae in the southeastern United States. Journal of the Arnold Arboretum 68: 361-445.
Tucker, G. C. 1994. A revision of the Mexican species of Cyperus (Cyperaceae). Systematic Botany Monographs 43. 212 pp.
Uittien, H. 1934. Cyperaceae. In A. Pulle, Flora of Suriname 1(1): 72-149.
Uittien, H. 1943. Cyperaceae (additions and corrections). In A. Pulle (ed.), Flora of Suriname 1(1): 492-505.
Dioscorea abysmophila Maguire & Steyerm.
Dioscorea alata L. [introduced] AM BO DA GU SU FG
Dioscorea altissima Lam.
Dioscorea amazonum Mart, ex Griseb. AM BO DA GU SU FG
Dioscorea aspera Humb. & Bonpl. ex Willd.
Dioscorea atrescens R. Knuth
Dioscorea bolivarensis Steyerm.
Dioscorea bulbifera L. [introduced] SU FG
Dioscorea cayenensis Lam. [introduced?] AM GU SU FG
Dioscorea cordifolia Laness. FG
Dioscorea crotalariifolia Uline AM GU
Dioscorea cuspidata Humb. & Bonpl. ex Willd. AM
Dioscorea decorticans C. Presl BO
Dioscorea discolor Kunth BO
Dioscorea dodecaneura Veil. GU SU FG
Dioscorea effusa Griseb. AM
Dioscorea esculenta (Lour.) Prain [cultivated only?] FG
Dioscorea georgensis R. Knuth [=Dioscorea amazonum?] GU
Dioscorea guianensis R. Knuth GU
Dioscorea hastata Veil. [=Dioscorea hassleranal] FG
Dioscorea hitchcockii R. Knuth GU
Dioscorea holmioidea Maury AM
Dioscorea laxiflora Mart, ex Griseb. AM
Dioscorea macrostachya Benth. GU?
Dioscorea manicorensis Clayton, sp. nov. ined. SU FG
Dioscorea megacarpa Gleason GU SU FG
Dioscorea melastomatifolia Uline ex Prain & Burkill [^Dioscorea truncaial] FG
Dioscorea microura R. Knuth GU
Dioscorea neblinensis Maguire & Steyerm. AM
Dioscorea nitida R. Knuth AM
Dioscorea oblonga Gleason GU SU
Dioscorea pachycarpa Kunth GU?
0. Tellez
AM
AM BO DA GU SU
BO
AM BO
BO
104 Checklist of the Plants of the Guiana Shield
Dioscorea panamensis R. Knuth
Dioscorea piloshisciila Bertero ex Spreng.
Dioscorea piperifolia Humb. & Bonpl. ex Willd.
Dioscorea pittieri R. Knuth
Dioscorea polygonoides Humb. & Bonpl. ex Willd.
Dioscorea pomeroonefis is R. Knuth
Dioscorea pofarens is R. Knuth
Dioscorea pubescens Poir. [insufficiently known species]
Dioscorea qiiinquefoliolata R. Knuth
Dioscorea riparia Kunth & R.H. Schomb. {=Dioscorea samyclea7]
Dioscorea sagittata Poir. [insufficiently known species]
Dioscorea samydea Griseb.
Dioscorea scabra Humb. & Bonpl. ex Willd.
Dioscorea sororopana Steyerm.
Dioscorea sp. A
Dioscorea syringifolia Kunth & R.H. Schomb.
Dioscorea trachyandra Griseb.
Dioscorea trachycarpa Kunth [insufficiently known species]
Dioscorea trichanthera Gleason
Dioscorea trifida L. f
Dioscorea trifoliata Kunth
Dioscorea truncata Miq. [=Dioscorea frifida?]
BO
AM BO GU
BO GU
AM
AM BO DA GU
GU
GU
GU
GU
GU
BO
BO
AM
GU
GU
GU
AM BO GU
AM GU
su
su
su
su
AM
SU
su
FG
FG
FG
FG
FG
FG
Synonyms and Misidentifications
Dioscorea affmis Kunth =Dioscorea trifida
Dioscorea amazomim var. consanguinea (Kunth) Uline =Dioscorea amazonum
Dioscorea bulbifera Griseb., non L. =Dioscorea pilosiuscula
Dioscorea ciispidata Klotzsch ex Kunth =Dioscorea amazonum
Dioscorea elegantiila Kunth =Dioscorea amazonum
Dioscorea holtii R. Knuth =Dioscorea amazonum
Dioscorea kegeliana Griseb. =Dioscorea polygonoides
Dioscorea lanosa Gleason =Dioscorea pilosiuscula
Dioscorea lindmanii Uline ex R. Knuth =Dioscorea pilosiuscula
Dioscorea megalobotrya Kunth & R.H. Schomb. ex Kunth =Dioscorea amazonum
Dioscorea pichinchensis R. Knuth =Dioscorea cuspidata
Dioscorea pimichinensis R. Knuth ex Sandwith =Dioscorea cuspidata
Dioscorea schomburgkiana Kunth & R.H. Schomb. =Dioscorea pilosiuscula
Dioscorea surinamensis G. Mey. =Dioscorea amazonum
Dioscorea truncata Prain, non Miq. =Dioscorea megacarpa
Dioscorea truncata R.H. Schomb., non Miq. =Dioscorea megacarpa
Helmia consanguinea Kunth =Dioscorea amazonum
References
Gleason, H.A. 1925. Studies on the flora of northern South America. V. Miscellaneous new or noteworthy species. Bull. Torrey
Bot. Club 52(5): 181-196.
Knuth, R. 1924. Dioscoreaceae. In A. Engler, Das Pflanzenreich IV(43): 1-387.
Mori, S.A. 1997. Dioscoreaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 264-266.
Tellez, O. 1998. Dioscoreaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 686-696.
Eriocaulaceae
M. Hakki, N. Hensold
Eriocaulon cinereum R. Br.
Eriocaulon dimorphopetalum Moldenke
Eriocaulon guyanense Kom.
Eriocaulon heterodoxum Moldenke
Eriocaulon humboldtii Kunth
Eriocaulonjauense Moldenke
Eriocaulon melanocephalum Kunth
Eriocaulon spongiola Hensold
Eriocaulon steyermarkii Moldenke
Eriocaulon stramineum Kom. [possibly ^Eriocaulon guyanense]
AM BO
BO
BO
AM BO
AM BO
AM BO
AM BO
DA
GU
GU
GU SU
GU SU
GU su
FG
FG
FG?
Checklist of the Plants of the Guiana Shield 05
Eriocaulon tenuifolium Klotzsch ex Korn.
Leiothrix celiae Moldenke
Leiothrixflavescens (Bong.) Ruhland
Leiothrixflavescens var. alpina Moldenke
Leiothrixflavescens var. flavescens
Leiothrix mucronata var. glabra Moldenke
Paepalanthus apacarensis Moldenke
Paepalanthus aristatus Moldenke
Paepalanthus auyantepidensis Moldenke
Paepalanthus bifidus (Schrad.) Kunth
Paepalanthus cardonae Moldenke
Paepalanthus chimantensis Hensold
Paepalanthus convexus Gleason
Paepalanthus cristatus Moldenke
Paepalanthus cumbricola Moldenke
Paepalanthus dichotomus Klotzsch ex Korn.
Paepalanthus dichotomus var. dichotomus
Paepalanthus dichotomus var. pumilus Moldenke
Paepalanthus fasciculatus (Rottb.) Kunth
Paepalanthus fasciculoides Hensold
^
Paepalanthusformosus Moldenke
Paepalanthus fraternus N.E. Br.
Paepalanthusfulgidus Moldenke
Paepalanthus gleasonii Moldenke
Paepalanthus holstii Steyerm.
Paepalanthus kunhardtii Moldenke
Paepalanthus lamarckii Kunth
Paepalanthus major (Moldenke) Hensold
Paepalanthus oyapockensis Herzog
Paepalanthus parvicephalus (Moldenke) Hensold
Paepalanthus parvicephalus var. parvicephalus
Paepalanthus parvicephalus var. wurdackii (Moldenke) Hensold
Paepalanthus pauper Moldenke
Paepalanthus perpusillus Kunth
Paepalanthus phelpsiae Moldenke
Paepalanthus polytrichoides Kunth
Paepalanthus roraimensis Moldenke
Paepalanthus schomburgkii Klotzsch ex Korn.
Paepalanthus scopulorum Moldenke
Paepalanthus sessiliflorus Korn.
Paepalanthus sessiliflorus var. venezuelensis Moldenke
Paepalanthus squamuliferus Moldenke
Paepalanthus stegolepoides Moldenke
Paepalanthus subcaulescens N.E. Br. [expected in BO]
Paepalanthus subtilis Miq.
Paepalanthus sulcatus Hensold
Paepalanthus tortilis (Bong.) Korn.
Paepalanthus turbinatus (Gleason) Hensold
Paepalanthus venustus Moldenke
Paepalanthus yapacanensis (Moldenke) Hensold
Paepalanthus yapacanensis var. hirsutus (Moldenke) Hensold
Paepalanthus yapacanensis var. yapacanensis
Philodice hoffmannseggii Mart.
Rondonanthus acopanensis (Moldenke) Hensold & Giul.
Rondonanthus acopanensis var. acopanensis
Rondonanthus acopanensis var. obtusifolius (Moldenke) Hensold & Giul.
Rondonanthus capillaceus (Klotzsch ex Korn.) Hensold & Giul.
Rondonanthus caulescens (Moldenke) Hensold & Giul.
Rondonanthus duidae (Gleason) Hensold & Giul.
Rondonanthus flabelliformis (Moldenke) Hensold & Giul.
Rondonanthus roraimae (Oliv.) Herzog
Syngonanthus acephalus Hensold
Syngonanthus albopulvinatus (Moldenke) Moldenke
Syngonanthus amapensis Moldenke [expected in the Guianas]
AM GU
AM
AM BO
BO
GU
AM BO
BO
BO
GU
AM
BO
AM BO
BO
BO
GU su
AM
BO
BO
GU?
BO GU
BO GU
BO
AM BO
BO
GU su
AM BO
BO GU
BO
AM
BO
GU
AM BO
AM BO GU su
AM
AM BO GU su
AM
AM
AM
GU
AM
BO
AM BO GU su
BO GU
BO GU
BO
BO GU
BO GU
BO
BO
BO? GU
AM BO GU su
AM
AM BO GU su
AM
BO
AM
AM
AM
AM BO
BO
BO
BO
GU su
AM BO
BO
GU
AM BO
BO
BO GU
AM
AM BO
AM
FG
FG
FG
FG
GU SU FG
GU su FG
GU
GU SU FG
GU
GU SU
GU?
GU su
GU SU
GU
GU SU
GU SU
106 Checklist of the Plants of the Guiana Shield
Syngonanthus amazonicus Moldenke
Syngonanthus anomalus (Kom.) Ruhland
Syngonanthus biformis (N.E. Br.) Gleason
Syngonanthus bisumbellatus (Steud.) Ruhland
Syngonanthus caulescens (Poir.) Ruhland
Syngonanthus cowanii Moldenke
Syngonanthus duidae Moldenke
Syngonanthus duidae var. duidae
Syngonanthus duidae var. humilis Hensold
Syngonanthus fenestratus Hensold
Syngonanthus gracilis (Bong.) Ruhland [possibly =Syngonanthus davidsei AM BO FG
Huft]
Syngonanthus heteropeploides Herzog
Syngonanthus humboldtii (Kunth) Ruhland
Syngonanthus humboldtii var. humboldtii
Syngonanthus humboldtii var. pai-vus (Moldenke) Hensold
Syngonanthusjenmanii (Gleason) Giul. & Hensold
Syngonanthus kegelianus (Kom.) Ruhland
Syngonanthus lepheurii (Kom.) Ruhland FG
Syngonanthus longipes Gleason
Syngonanthus macrocephalus (Moldenke) Hensold
Syngonanthus minutus (Moldenke) Hensold
Syngonanthus nitens (Bong.) Ruhland
Syngonanthus oblongus (Kom.) Ruhland
Syngonanthus oblongus var. aequinoctialis Ruhland
Syngonanthus ottohuberi Hensold
Syngonanthus pakaraimensis Moldenke BO GU
Syngonanthus pakaraimensis var. pakaraimensis BO GU
Syngonanthus pakaraimensis var. rivularis (Moldenke) Hensold
Syngonanthus reflexus Gleason
Syngonanthus setifolius Hensold
Syngonanthus simplex (Miq.) Ruhland AM BO GU SU
Syngonanthus surinamensis Moldenke SU
Syngonanthus tenuis (Kunth) Ruhland
Syngonanthus tenuis var. bulbifer (Huber) Hensold
Syngonanthus tenuis var. tenuis
Syngonanthus tiricensis Moldenke
Syngonanthus trichophyllus Moldenke AM BO GU
Syngonanthus umbellatus (Lam.) Ruhland AM BO GU SU FG
Syngonanthus williamsii (Moldenke) Hensold
Syngonanthus xeranthemoides (Bong.) Ruhland AM BO GU
TomnafluviatilisAubl AM BO DA GU SU FG
Synonyms and Misidentifications
Carptotepala insolita Moldenke =Syngonanthus Jenmanii
Carptotepalajenmanii (Gleason) Moldenke ^Syngonanthus jenmanii
Comanthera kegeliana (Kom.) Moldenke =Syngonanthus kegelianus
Comanthera linderi L.B. Sm. =Syngonanthus kegelianus
Eriocaulon ? aequinoctiale Ruhland =Eriocaulon humboldtii
Eriocaulon aquaticum Sagot ex Kom =Eriocaulon melanocephalum
Eriocaulon atabapense Moldenke =Eriocaulon tenuifolium
Eriocaulon bifidum Schrad. =Paepalanthus bifidus
Eriocaulon bisumbellatum Steud. =Syngonanthus bisumbellatus
Eriocaulon bonplandianum Steud. =Syngonanthus humboldtii
Eriocaulon brevifolium Klotzsch ex Kom. =Eriocaulon tenuifolium
Eriocaulon caulescens Poir. =Syngonanthus caulescens
Eriocaulonfasciculatum Rottb. =Paepalanthus fasciculatus
Eriocaulonfasciculatum Lam. =Paepalanthus lamarckii
Eriocaulonflavescens Bong. =Leiothrixflavescens
Eriocaulon gracile Bong. =Syngonanthus gracilis
Eriocaulon klotzschii Moldenke =Eriocaulon tenuifolium
Eriocaulon nitens Bong. =Syngonanthus nitens
Eriocaulon sieboldianum Siebold & Zucc. ex Steud. =Eriocaulon cinereum
Eriocaulon tenue Kunth =Syngonanthus tenuis
AM BO
AM BO
AM BO
BO
AM
A A/f
BO
AM
AM BO
AM BO
BO
BO
AM BO
AM BO
AM BO
BO
BO
AM BO
AM
BO
AM BO
AM BO
AM BO
AM
BO
AM
AM
AM BO
AM BO
AM BO
BO
AM
Checklist of the Plants of the Guiana Shield 107
Eriocaulon tortile Bong. =Paepalanthus tortilis
Eriocaidon umbellatum Kunth =Syngonanthus humboldtii
Eriocaulon umbellatum Lam. =Syngonanthus umbellatus
Eriocaulon xeranthemoides Bong. =Syngonanthus xeranthemoides
Lasiolepis aquatica Bock. =Eriocaulon melanocephalum
Leiothrix echinulata Moldenke =Syngonanthus reflexus
Leiothrixflavescens var. chimantensis Moldenke =Leiothrixflavescens
Leiothrix marahuacensis Moldenke =Syngonanthus duidae
Leiothrix steyermarkii Moldenke =Syngonanthus pakaraimensis
Leiothrix turbinata Gleason =Paepalanthus turbinatus
Leiothrix umbratilis Moldenke =Leiothrixflavescens
Leiothrix umbratilis var. brevipes Moldenke =Leiothrixflavescens var. alpina
Paepalanthus albopulvinatus Moldenke =Syngonanthus albopulvinatus
Paepalanthus anomalus Kom. =Syngonanthus anomalus
Paepalanthus apacarensis var. humilis Moldenke ^Paepalanthus dichotomus
Paepalanthus biformis N.E. Br. =Syngonanthus biformis
Paepalanthus brunneus Moldenke ^Paepalanthus oyapockensis
Paepalanthus bulbifer Huber =Syngonanthus tenuis var. bulbifer
Paepalanthus capillaceus Klotzsch ex Kom. =Rondonanthus capillaceus
Paepalanthus capillaceus f. proliferus (Gleason) Moldenke =Rondonanthus capillaceus
Paepalanthus capillaceus var. proliferus Gleason =Rondonanthus capillaceus
Paepalanthus caulescens (Poir.) Kunth =Syngonanthus caulescens
Paepalanthus congestus (Kunth) Kunth =Paepalanthus fasciculatus
Paepalanthus convexus var. major Moldenke ^Paepalanthus major
Paepalanthus convexus var. parvicephalus Moldenke =Paepalanthus parvicephalus
Paepalanthus convexus var. strigosus Moldenke =Paepalanthus gleasonii
Paepalanthus duidae Gleason =Rondonanthus duidae
Paepalanthus duidae var. parvifolius Moldenke =Rondonanthus duidae
Paepalanthus eriophyllus Kom. =Syngonanthus simplex
Paepalanthus fertilis Kom. =Syngonanthus humboldtii
Paepalanthus fdipes Moldenke =Syngonanthus simplex
Paepalanthus flavescens (Bong.) Kom. =Leiothrixflavescens
Paepalanthus fraternus var. chimantensis Moldenke =Paepalanthus stegolepoides
Paepalanthus fraternus var. marahuacensis Moldenke =Leiothrixflavescens
Paepalanthus fraternus var. spathulatus Moldenke =Paepalanthus squamuliferus
Paepalanthusfulgidus var. zuloagensis Moldenke =Paepalanthus major
Paepalanthus gracilis (Bong.) Kom. =Syngonanthus gracilis
Paepalanthus griseus Moldenke ^Paepalanthus oyapockensis
Paepalanthus guyanensis Klotzsch ex Kom. =Paepalanthus dichotomus
Paepalanthus heteropeplus Kom. =Syngonanthus caulescens
Paepalanthus humboldtii Kunth =Syngonanthus humboldtii
Paepalanthusjauensis Moldenke =Rondonanthus duidae
Paepalanthusjauensis var. caulescens Moldenke =Rondonanthus caulescens
Paepalanthus jenmanii Gleason =Syngonanthusjenmanii
Paepalanthus kegelianus Kom. ^Syngonanthus kegelianus
Paepalanthus killipii Moldenke =Paepalanthus tortilis
Paepalanthus leucocyaneus Tutin =Paepalanthus oyapockensis
Paepalanthus lilliputianus Moldenke =Paepalanthus sessiliflorus var. venezuelensis
Paepalanthus macrocaulon var. venamensis Moldenke =Paepalanthus schomburgkii
Paepalanthus maguirei Moldenke =Paepalanthus tortilis
Paepalanthus meseticola Moldenke & Steyerm. =Leiothrixflavescens
Paepalanthus minutus Moldenke =Syngonanthus minutus
Paepalanthus moldenkeanus R. Schult. =Paepalanthus formosus
Paepalanthus nitens (Bong.) Kunth =Syngonanthus nitens
Paepalanthus oblongus Korn. =Syngonanthus oblongus
Paepalanthus pauperrimus Herzog =Paepalanthus subtilis
Paepalanthus pendulus Moldenke =Paepalanthus schomburgkii
Paepalanthus perplexans Moldenke =Paepalanthus schomburgkii
Paepalanthus perplexans var. steyermarkii Moldenke =Paepalanthus fulgidus
Paepalanthus perplexans var. wurdackii Moldenke =Paepalanthus parvicephalus var. wurdackii
Paepalanthus robustus Gleason =Paepalanthus gleasonii
Paepalanthus roraimae Oliv. =Rondonanthus roraimae
Paepalanthus salticola Herzog =Paepalanthus tortilis
Paepalanthus savannarum (Moldenke) Moldenke =Paepalanthus dichotomus
Paepalanthus savannarum var. glabrescens (Moldenke) Moldenke =Paepalanthus dichotomus
108 Checklist of the Plants of the Guiana Shield
Paepalanthus saxicola var. conicus Moldenke =Syngonanthus trichophyllns
Paepalanthiis scopulorum var. aiiyantepuiensis Moldenke =Paepalanthus scopulorum
Paepalanthus simplex Miq. =Syngonanthus simplex
Paepalanthus stegolepoides var. acutalis Moldenke =Paepalanthus sulcatus
Paepalanthus steudelianus Kom. =Syngonanthus bisumbellatus
Paepalanthus steyermarkii Moldenke =Paepalanthus dichotomus
Paepalanthus subulatus Klotzsch =Paepalanthus subtilis
Paepalanthus tafelbergensis Moldenke =Paepalanthus oyapockensis
Paepalanthus tatei Moldenke =Paepalanthus tortilis
Paepalanthus tenuis (Kunth) Kunth =Syngonanthus tenuis
Paepalanthus umbellatus (Lam.) Kunth =Syngonanthus umbellatus
Paepalanthus ? venetifolius Moldenke & Steyerm. =Syngonanthus pakaraimensis var. pakaraimensis
Paepalanthus venustoides Moldenke =Paepalanthus venustus
Paepalanthus vigiensis Moldenke =Paepalanthus polytrichoides
Paepalanthus villipes Moldenke =Paepalanthus polytrichoides
Paepalanthus viscosus Moldenke ^Paepalanthus polytrichoides
Paepalanthus williamsii Moldenke =Syngonanthus williamsii
Rondonanthus micropetalus Moldenke =Paepalanthus roraimensis
Syngonanthus acopanensis Moldenke ^Rondonanthus acopanensis
Syngonanthus akurimensis Moldenke =Syngonanthus kegelianus
Syngonanthus allenii var. parvus Moldenke ^Syngonanthus humboldtii var. parvus
Syngonanthus baldwinii Moldenke ^Syngonanthus anomalus
Syngonanthus brevifolius Gleason =Syngonanthus anomalus
Syngonanthus bulbifer (Huber) Ruhland =Syngonanthus tenuis var. bulbifer
Syngonanthus caulescens var. hirsutus Moldenke =Syngonanthus caulescens
Syngonanthus cowanii var. involucratus Moldenke =Syngonanthus cowanii
Syngonanthus cowanii var. longipedunculatus Moldenke =Syngonanthus cowanii
Syngonanthus cowanii var. simplex Moldenke ^Syngonanthus cowanii
Syngonanthus cowanii var. tabulatus Moldenke =Syngonanthus cowanii
Syngonanthus densifolius var. venezuelensis Moldenke =Syngonanthus ottohuberi
Syngonanthus densus var. pumilus Moldenke ^Syngonanthus biformis
Syngonanthus drouetii L.B. Sm. =Syngonanthus tenuis var. bulbifer
Syngonanthus drouetii var. parviceps Moldenke =Syngonanthus tenuis
Syngonanthus duidae var. longifolius Moldenke =Syngonanthus pakaraimensis
Syngonanthus eriophyllus (Kom.) Ruhland =Syngonanthus simplex
Syngonanthus fertilis (Kom.) Ruhland =Syngonanthus humboldtii
Syngonanthus fertilis var. glandulosus (Gleason) Moldenke =Syngonanthus humboldtii
Syngonanthus fertilis var. orinocensis (Moldenke) Moldenke =Syngonanthus humboldtii
Syngonanthus flavipes Moldenke =Syngonanthus nitens
Syngonanthus glandulosus Gleason =Syngonanthus caulescens
Syngonanthus glandulosus var. epapillosus Moldenke =Syngonanthus caulescens
Syngonanthus glandulosus f. epapillosus (Moldenke) Moldenke =Syngonanthus caulescens
Syngonanthus guianensis Moldenke =Paepalanthus polytrichoides
Syngonanthus heteropeplus (Kom.) Ruhland =Syngonanthus caulescens
Syngonanthus huberi Ruhland ^Syngonanthus anomalus
Syngonanthus humboldtii var. elongatus Moldenke =Syngonanthus humboldtii
Syngonanthus humboldtii var. glabrescens Moldenke =Syngonanthus humboldtii
Syngonanthus humboldtii var. glandulosus Gleason =Syngonanthus humboldtii
Syngonanthus humboldtii var. macrocephalus Moldenke =Syngonanthus macrocephalus
Syngonanthus humboldtii var. orinocensis Moldenke =Syngonanthus humboldtii
Syngonanthus humboldtii var. simplex Moldenke =Syngonanthus umbellatus
Syngonanthus lanatus var. alpinus Moldenke ^Syngonanthus pakaraimensis
Syngonanthus macrocaulon Ruhland =Syngonanthus anomalus
Syngonanthus obtusifolius Moldenke =Rondonanthus acopanensis var. obtusifolius
Syngonanthus paraensis Ruhland =Syngonanthus anomalus
Syngonanthus phelpsiae Moldenke =Syngonanthus duidae
Syngonanthus phelpsiae var. elongatus Moldenke ^Syngonanthus duidae
Syngonanthus phelpsiae var. pilosus Moldenke =Rondonanthus duidae
Syngonanthus phelpsiae var. viridis Moldenke =Rondonanthus duidae
Syngonanthus reflexus var. longifolius Moldenke =Syngonanthus reflexus
Syngonanthus rivularis Moldenke =Syngonanthus pakaraimensis var. rivularis
Syngonanthus savannarum Moldenke =Paepalanthus dichotomus
Syngonanthus savannarum var. glabrescens Moldenke =Paepalanthus dichotomus
Syngonanthus similis var. venezuelensis Moldenke =Syngonanthus ottohuberi
Syngonanthus simplex var. appendiculifera Ruhland ^Syngonanthus biformis
Checklist of the Plants of the Guiana Shield 109
Syngonanthus ? tenuis var. minor Moldenke =Syngonanthus tenuis
Syngonanthus tricostatus Gleason =Syngonanthus xeranthemoides
Syngonanthus umbellatus f. proliferens Moldenke =Syngonanthus umbellatus
Syngonanthus vareschii Moldenke =Syngonanthus simplex
Syngonanthus vaupesanus Moldenke =Syngonanthus tenuis var. bulbifer
Syngonanthus venezuelensis Moldenke =Paepalanthus dichotomus
Syngonanthus xeranthemoides var. alpinus Moldenke =Syngonanthus xeranthemoides
Syngonanthus xeranthemoides var. angustifolius Moldenke ^Syngonanthus xeranthemoides
Syngonanthus xeranthemoides f. brevifolius Moldenke =Syngonanthus xeranthemoides
Syngonanthus xeranthemoides var. tricostatus (Gleason) Moldenke =Syngonanthus xeranthemoides
Syngonanthus yapacanensis Moldenke =Paepalanthus yapacanensis
Syngonanthus yapacanensis var. hirsutus Moldenke =Paepalanthus yapacanensis var. hirsutus
Wurdackia flabelliformis Moldenke ^Rondonanthus flabelliformis
References
Giulietti, A.M., M.C.E. Amaral, & V. Bittrich. 1995. Phylogenetic analysis of inter- and infrageneric relationships of Leiothrix
Ruhland (Eriocaulaceae). Kew Bull. 50(1): 55-71.
Giulietti, A.M. & N. Hensold. 1991. Nomenclatural changes and range extension in Leiothrix flavescens (Bong.) Ruhl.
(Eriocaulaceae). Novon 1(1): 45-49.
Giulietti, A.M. & N. Hensold. 1991. Synonymization of the genera Comanthera and Carptotepala with Syngonanthus
(Eriocaulaceae). Ann. Missouri Bot. Gard. 78 (2): 460-464.
Hensold, N. 1991. Revisionary studies in the Eriocaulaceae of Venezuela. Ann. Missouri Bot. Gard. 78(2): 424-440.
Hensold, N. 1999. Eriocaulaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
5: 1-58.
Hensold, N. & A.M. Giulietti. 1 99 1 . Revision and redefinition ofthe genus Rondonanthus Herzog (Eriocaulaceae). Ann. Missouri
Bot. Gard. 78(2): 441-459.
Lindeman, J.C. & A.R.A. Gorts-van Rijn. 1968. Eriocaulaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of
Suriname 1(2): 330-339.
Moldenke, H.N. 1971. A Fifth Summary of the Verbenaceae, Avicenniaceae, Stilbaceae, Dicrostylidaceae, Symphoremaceae,
Nyctanthaceae, and Eriocaulaceae of the World. H.N. Moldenke, Wayne, NJ.
Mori, S.A. 1997. Eriocaulaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 266.
Uittien, H. & A.N.J. Heyn. 1938. Eriocaulaceae. In A. Pulle, Flora of Suriname 1(1): 213-224.
Haemodoraceae
P.J.M. Maas, Hiltje Maas
AM BO DA GU SU FG
AM BO GU SU
AM BO DA GU SU FG
AM BO DA GU SU FG
Pyrrorhiza neblinae Maguire & Wurdack
Schiekia orinocensis (Kunth) Meisn.
Schiekia orinocensis subsp. orinocensis
Schiekia orinocensis subsp. silvestris Maas & Stoel
Xiphidium caeruleum Aubl.
Synonyms and Misidentifications
Schiekiaflavescens Maury =Schiekia orinocensis subsp. orinocensis
Schiekia orinocensis subsp. savannarum Maguire & Wurdack =Schiekia orinocensis subsp. orinocensis
Troschelia orinocensis (Kunth) Klotzsch & M.R. Schomb. [nomen nudum] =Schiekia orinocensis subsp. orinocensis
Wachendorfla orinocensis Kunth =Schiekia orinocensis subsp. orinocensis
Xiphidium floribundum Sw. =Xiphidium caeruleum
Xiphidiumfockeanum Miq. =Xiphidium caeruleum
Xiphidium giganteum Lindl. =Xiphidium caeruleum
References
Maas-van de Kamer, H. & P. Maas. 1997. Haemodoraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, &
J.D. Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and
Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1: 270.
Lanjouw, J. 1938. Haemodoraceae. In A. Pulle, Flora of Suriname 1(1): 255-257.
Maas, P.J.M. & H. Maas-van de Kamer. 1993. Haemodoraceae. Flora Neotropica Monogr. 61: 1-44.
Maas, P.J.M. & H. Maas-van de Kamer. 1994. Haemodoraceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 15:
103-112.
Maas, P.J.M. & H. Maas. 1999. Haemodoraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 5: 576-581.
Villarroel, F. 1971. Haemodoraceae. In T. Lasser, Flora de Venezuela 3(1): 207-222.
AM BO GU su FG
AM BO GU su FG
AM
FG?
BO DA GU su FG
AM BO GU su
su
FG
FG
BO GU su FG
BO GU
BO GU su FG
AM su
AM BO DA GU su FG
AM
BO GU su FG
DA GU
GU
GU
su
su
FG
BO
AM BO DA GU su FG
AM
BO DA GU su FG
BO DA GU
GU
su
su
FG
110 Checklist of the Plants of the Guiana Shield
Heliconiaceae
L.L. Andersson, W.J. Kress, P.J.M. Maas
Heliconia acuminata Rich.
Heliconia acuminata subsp. acuminata
Heliconia acuminata subsp. occidentalis L. Andersson
Heliconia aemygdiana Burle-Marx [CAY specimen not seen]
Heliconia bihai (L.) L.
Heliconia chartacea Lane ex Barreiros
Heliconia dasyantha K. Koch & Bouche
Heliconia densiflora B. Verl.
Heliconia densiflora subsp. angustifolia L. Andersson
Heliconia densiflora subsp. densiflora
Heliconia episcopalis Veil.
Heliconia hirsuta L. f.
Heliconiajuliani Barreiros
Heliconia lourteigiae Emygdio & E. Santos
Heliconia marginata (Griggs) Pittier
Heliconia x nickeriensis Maas & de Rooij
Heliconia pendula Wawra
Heliconia platystachys Baker
Heliconia psittacorum L. f.
Heliconia revoluta (Griggs) Standi. [Sierra de la Neblina]
Heliconia richardiana Miq.
Heliconia spathocircinata Aristeg. [orthographic variant: H. spathocircinada]
Heliconia stricta Huber
Synonyms and Misidentifications
Bihai marginata Griggs =Heliconia marginata
Bihai revoluta Griggs ^Heliconia revoluta
Bihai silvestris Gleason =Heliconia psittacorum
Heliconia aurea G. Rodr. =Heliconia bihai
Heliconia ballia Rich. =Heliconia psittacorum
Heliconia brasiliensis var. concolor Petersen =Heliconia psittacorum
Heliconia cannoidea Rich. =Heliconia psittacorum
Heliconia caribaea auct. non Lam. 1783 [1785], nee Aristeg. 1961 =Heliconia bihai
Heliconia costanensis Aristeg. =Heliconia hirsuta
Heliconia glauca Poit. ex B. Verl. =Heliconia richardiana
Heliconia hirsuta var. cannoidea (Rich.) Baker =Heliconia psittacorum
Heliconia humilis Aubl., auct. non Musa humilis =Heliconia bihai
Heliconia humilis (Aubl.) Jacq. =Heliconia psittacorum
Heliconia marginata f. lutea Aristeg. =Heliconia marginata
Heliconia psittacorum var. rhizomatosa Aristeg. =Heliconia psittacorum
Heliconia psittacorum var. schomburgkiana (Klotzsch) Baker =Heliconia psittacorum
Heliconia psittacorum var. spathacea Eichler ex Petersen =Heliconia psittacorum
Heliconia schaeferiana G. Rodr. =Heliconia bihai
Heliconia schomburgkiana Klotzsch =Heliconia psittacorum
Heliconia silvestris (Gleason) L.B. Sm. =Heliconia psittacorum
References
Anderson, P., S.A. Mori, & C. Gracie. 1997. Heliconiaceae. In: Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, M. Hoff, &
J.D. Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and
Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1 : 270-273.
Andersson, L.L. 1999. Heliconiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 5: 583-591.
Berry, F. & W.J. Kress. 1991. Heliconia: An Identification Guide. Washington, DC: Smithsonian Institution Press.
Maas, P.J.M. 1985. Musaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 1: 1-28.
Maas, RJ.M. & M.J.M. de Rooij. 1979. Musaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 393-411.
Hydrocharitaceae
R.R. Haynes, L.B. Holm-Nielsen
Elodea granatensis Bonpl. AM BO GU SU FG
Limnobium laevigatum (Humb. & Bonpl. ex Willd.) Heine DA GU SU FG
Checklist of the Plants of the Guiana Shield 1 1
1
Synonyms and MisidentificationsYNONYMS AND I
Apalanthe granatensis (Bonpl.) Planch. =Elodea granatensis
Elodea guianensis Rich. =Elodea granatensis
Elodea orinocensis Rich. =Elodea granatensis
Hydromystria laevigata (Willd.) Hanzeker =Limnobium laevigatum
Hydromystria laevigata (Humb. & Bonpl. ex Willd.) Diaz-Mir. & Philcox =Limnobium laevigatum
Limnobium spongia subsp. laevigatum (Humb. & Bonpl. ex Willd.) Lowden =Limnobium laevigatum
Limnobium stoloniferum (G. Mey.) Griseb. =Limnobium laevigatum
Philotria granatensis (Bonpl.) Vict. =Elodea granatensis
Philotria orinocensis (Rich.) Vict. =Elodea granatensis
Udora granatensis (Bonpl.) Spreng. =Elodea granatensis
References:
Cook, CD. 1985. A revision of the genus Apalanthe. Aquatic Bot. 21: 157-164.
Cook, CD. & K. Urmi-Konig, 1983. A revision of the genus Limnobium including Hydromystria. Aquatic Bot. 17: 1-27.
Haynes, R.R. & L.B. Holm-Nielsen. 1999. Hydrocharitaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 5: 641-644.
Jonker, F.R 1943. Hydrocharitaceae. In A. Pulle, Flora of Suriname 1(1): 467-471.
Lowden, R.M. 1992. Floral variation and taxonomy of Limnobium L.C Richard. Rhodora 94: 111-134.
Phillips, R.C & E.G. Meilez. 1988. Seagrasses. Smiths. Contrib. Marine Sci. 34: 1-104.
Iridaceae
R Goldblatt, J.E. Henrich
Alophia drummondii (Graham) R.C. Foster GU
Cipura gigas Cells, Goldblatt & Betancur AM
Cipura paludosa Auhl AM BO GU SU FG
Cipura rupicola Goldblatt & Henrich AM
Cypella linearis (Kunth) Baker AM BO
Eleutherinebulbosa{M\\\.)\]rh. GU SU FG
Neomarica caerulea (Ker Gawl.) Sprague [escaped from cultivation]
Sisyrinchium tinctorium Kunth AM
Sisyrinchium vaginatum Spreng. [expected in FG] AM BO GU SU
Trimezia chimantensis Steyerm. BO
Trimeziafosteriana Steyerm. BO GU
Trimezia guianensis Ravenna GU
Trimezia martinicensis (Jacq.) Herb. BO GU? FG
Synonyms and Misidentifications
Iris martinicencis Jacq. =Trimezia martinicensis
Ixia americana Aubl. =Eleutherine bulbosa
Ixia xiphidium Loefl. (Haemodoraceae) =Xiphidium caeruleum
Larentia linearis (Kunth) Klatt =Cypella linearis
Moraea linearis Kunth =Cypella linearis
Sisyrinchium alatum Hook. =Sisyrinchium vaginatum
Sisyrinchium americanum (Aubl.) Lemee =Eleutherine bulbosa
Sisyrinchium marchio (Veil.) Steud. =Sisyrinchium vaginatum
Souza marchio Veil. =Sisyrinchium vaginatum
References:
Boterenbrood, M.J.A. 1943. Iridaceae. In A. Pulle, Flora of Suriname 1(1): 455-460.
Goldblatt, R & James E. Henrich. 1999. Iridaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 5: 658-664.
Goldblatt, P. &T.M. Howard. 1992. Notes on Alophia and anew species, A. veracruzana, from Vera Cruz, Mexico. Ann. Missouri
Bot. Card. 79(4): 901-905.
Goldblatt, P. & N. Snow. 1991. Systematics and chromosome cytology of Eleutherine Herbert. Ann. Missouri Bot. Gard. 78(4):
942-949.
Mori, S.A. 1997. Iridaceae. In Mori, S.A., G Cremers, C Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide
to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs of the
New York Botanical Garden 76, pt. 1 : 273-274.
JUNCACEAE
CA. Vargas
Juncus densiflorus Kunth AM BO
112 Checklist of the Plants of the Guiana Shield
Synonyms and Misidentifications
Jimcus cyperinus Willd. ex Spreng. =Juncus densiflorus
Juncus densiflorus van cyperinus (Willd.) Buchenau =Juncus densiflorus
References
Vargas, C.A. 1999. Juncaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
5: 672-674.
Lemnaceae
L.J. Davenport, R.A. DeFilippst
DA
DA
GU
GU
SU
SU
SU
SU
FG
SU FG
SU
DA GU SU FG
Lemna aequinoctialis Welw.
Lemna valdiviana Phil.
Spirodela intermedia W. Koch
Spirodela punctata (G. Mey.) C.H. Thomps.
Wolffia brasiliensis Wedd.
^
BO
Wolffiella lingulata (Hegelm.) Hegelm.
Wolffiella neotropica Landolt
Wolffiella welwitschii (Hegelm.) Monod BO
Synonyms and Misidentifications
Lemna gibba L. [not in flora area]
Lemna minor L. [not in flora area]
Lemna paucicostata Hegelm. =Lemna aequinoctialis
Lemna perpusilla Torr. [not in flora area]
Lemna trinervis sensu Small, non (D.F. Austin) Small =Lemna aequinoctialis
Spirodela biperforata W. Koch =Spirodela intermedia
Spirodela polyrhiza (L.) Schleid. [not in flora area]
Wolffia welwitschii Hegelm. =Wolffiella welwitschii
Wolffiopsis welwitschii (Hegelm.) Hartog & Plas =Wolffiella welwitschii
References
Daubs, E.H. 1965. A monograph of Lemnaceae. 111. Biol. Monogr. 34: 1-118.
Davenport, L.J. 1999. Lemnaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan Guayana
5:779-781.
Hartog, C. den. & F. van der Plas. 1970. A synopsis of the Lemnaceae. Blumea 18: 355-368.
Landoh, E. 1986-1987. The family of Lemnaceae - a monographic study. Veroff Geobot. Inst. E.T.H. Stift. Rubel, Zurich 71:1-
566,95: 1-638.
LiLIACEAE
J.K. Boggan, R.W. Cruden, A.W. Meerow
Alstroemeria amazonica Ducke
Alstroemeria sp.
Bomarea edulis (Tussac) Herb.
Crinum erubescens L. f ex Sol.
Curculigo scorzonerifolia (Lam.) Baker
Eccremis coarctata (Ruiz & Pav.) Baker
Echeandia bolivarensis Cruden
Hippeastrum barbatum Herbert
Hippeastrum elegans (Spreng.) H.E. Moore
Hippeastrum puniceum (Lam.) Kuntze
Hippeastrum stylosum Herbert [distribution described as "Guiana"]
Hippeastrum vanleestenii (Traub) H.E. Moore [=Hippeastrum puniceum?]
Hymenocallis caribaea (L., emend. Ker Gawl.) Herb, [cultivated or escaped
from cultivation]
Hymenocallis littoralis (Jacq.) Salisb. [cultivated or escaped from cultivation]
Hymenocallis tubiflora Salisb.
Hypoxis decumbens L. [most, if not all, records represent Curculigo
scorzonerifolia]
Isidrogalvia duidae (Steyerm.) Cruden
Isidrogalvia schomburgkiana (Oliv.) Cruden
Nietneria corymbosa Klotzsch & M.R. Schomb. ex B.D. Jacks. [Mt. Roraima] AM
Nietneria paniculata Steyerm.
BO
FG
AM BO GU SU
AM DA GU SU FG
AM BO GU SU FG
AM BO
BO
SU
AM BO GU SU
AM GU SU
SU
FG
GU SU FG
GU? SU FG
BO DA GU SU FG
GU? SU? FG?
AM
AM BO GU
BO GU
BO GU
Checklist of the Plants of the Guiana Shield 113
Zephyranthes citrina Baker [type attributed to Guyana, possibly erroneously] GU?
Zephyranthes tubispatha (Ker Gawl.) Herb. SU
Synonyms and Misidentifications
Alstroemeria edulis Tussac =Bomarea edulis
Amaryllis belladonna L. [name incorrectly applied to Hippeastrum puniceum]
Amaryllis belladonna var. barbata (Herb.) Traub & Moldenke =Hippeastrum barbatiim
Amaryllis eguestris Alton =Hippeastrum puniceum
Amaryllis elegans Spreng. =Hippeastrum elegans
Amaryllis longiflora Steud. =Hippeastrum elegans
Amaryllis punicea Lam. =Hippeastrum puniceum
Amaryllis solandriflora Lindl. ^^Hippeastrum elegans
Amaryllis stylosa (Herb.) Sweet ^Hippeastrum stylosum
Amaryllis vanleestenii Traub =Hippeastrum vanleestenii
Anthericum coarctatum Ruiz & Pav. =Eccremis coarctata
Bomareafuscata Klotzsch ex M.R. Schomb. [nomen nudum]
Bomarea guianensis Kraenzl. =Bomarea edulis
Crinum commelyni Jacq. =Crinum erubescens
Crinum guianense M. Roem. ? =Crinum erubescens
Crinum lancei Herbert ex Sweet ? =Crinum erubescens
Crinum lindleyanum Schult. f. ex Seub. =Crinum erubescens
Dianella dubia Kunth =Eccremis coarctata
Echeandia spp. [recorded from Guianas in error, but possibly expected]
Hippeastrum equestre (Alton) Herb. =Hippeastrum puniceum
Hippeastrum puniceum (Lam.) Urb. =Hippeastrum puniceum
Hippeastrum puniceum (Lam.) Voss =Hippeastrum puniceum
Hippeastrum solandriflorum (Lindl.) Herb. =Hippeastrum elegans
Hymenocallis guianensis (Ker Gawl.) Herb. =Hymenocallis tubiflora
Hymenocallis guianensis var. undulata (Kunth) Herb. =Hymenocallis tubiflora
Hymenocallis moritziana Kunth =Hymenocallis tubiflora
Hymenocallis obtusata (Griseb.) Walp. =Hymenocallis caribaea
Hymenocallis petiolata (Willd. ex Schult.) M. Roem. =Hymenocallis tubiflora
Hymenocallis undulata (Kunth) Herb. =Hymenocallis tubiflora
Hypoxis scorzonerifolia Lam. =Curculigo scorzonerifolia
Isidrogalvia guianensis Klotzsch [nomen nudum] =Isidrogalvia schomburgkiana
Nietneria corymbosa Klotzsch & M.R. Schomb. [nomen nudum] =Nietneria corymbosa
Pancratium guianense Ker Gawl. =Hymenocallis tubiflora
Pancratium petiolatum Willd. ex Schult. =Hymenocallis tubiflora
Pancratium tubiflorum (Salisb.) Schult. =Hymenocallis tubiflora
Pancratium undulatum Kunth =Hymenocallis tubiflora
Tofieldia duidae Steyerm. =Isidrogalvia duidae
Tofieldia guianensis (Klotzsch) H. Schulz [nomen nudum] =Isidrogalvia schomburgkiana
Tofieldia rupestris Gleason (Bromeliaceae) =Navia rupestris
Tofieldia schomburgkiana Oliv. =Isidrogalvia schomburgkiana
References
Baker, J.G 1888. Handbook of the Amaryllideae. London: George Bell & Sons.
Bloomfield, F. 1985. Miracle Plants: Aloe vera. London: Century Publishing.
Boterenbrood, M.J.A. 1943. Amaryllidaceae. In A. PuUe, Flora of Suriname 1(1): 443-454.
Cruden, R.W. 1991. A revision of Isidrogalvia: recognition for Ruiz & Pavon's genus. Syst. Bot. 16: 270-282.
Howard, R.A. 1979. Liliaceae. Flora of the Lesser Antilles 3: 454-460.
Killip, E.P. 1 936. Bomarea, a genus of showy Andean plants. Nat. Hort. Mag. 1 5(2): 115-1 28.
Maas, P.J.M. 1968. Amaryllidaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 376.
Martyn, E.B. 1936. Amaryllidaceae of British Guiana. Herbertia 3: 71.
Meerow, A.W. & R.W. Cruden. 2001. Liliaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 6: 1-15.
Mori. S.A. 1997. Liliaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. Guide to the
vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs of the New
York Botanical Garden 76, pt. 1
.
Obermeyer, A.A. 1962. A revision of the South African species of Anthericum, Chlorophytum and Trachyandra. Bothalia 7 (4):
669-767.
Sealy, J.R. 1954. Review of the genus Hymenocallis. Kew Bull. 1954: 201-240.
Traub, H.P. 1958. The Amaryllis Manual. The Macmillan Company, New York.
Traub, H.P. & H.N. Moldenke. 1949. Amaryllidaceae: Tribe Amarylleae. American Plant Life Society.
Uphof, J.C. Th. 1942. A review of the species of Crinum. Herbertia 9: 63-84.
114 Checklist of the Plants of the Guiana Shield
LiMNOCHARITACEAE
R.R. Haynes, L.B. Holm-Nielsen
Hydrocleys nymphoides (Humb. & Bonpl. ex Willd.) Buchenau BO DA GU SU FG?
Limnocharisflava (L.) Buchenau [record not verified, but expected FG?
in the Guianas]
Limnocharis laforestii Duchass. ex Griseb. BO
Synonyms and Misidentifications
Limnocharis flava var. minor Micheli ^Limnocharis laforestii
Stratiotes nymphoides Humb. & Bonpl. ex Willd. =Hydrocleys nymphoides
References
Haynes, R.R. & L.B. Holm-Nielsen. 2001. Limnocharitaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 6: 16-18.
Haynes, R.R. & L.B. Holm-Nielsen. 1992. Limnocharitaceae. Flora Neotropica Monogr. 56: 1-34.
Jonker, F.R 1943. Butomaceae. In A. Pulle, Flora of Suriname 1(1): 483-485.
Marantaceae
L.L. Andersson, Mats Hagberg, H. Kennedy
Calathea acuminata Steyerm.
Calathea altissima (Poepp. & Endl.) Kom.
Calathea cannoides (Nicolson, Steyerm. & Sivad.) H. Kenn.
Calathea casupito (Jacq.) Schult.
Calathea cataractarum K. Schum.
Calathea comosa (L. f ) Lindl.
Calathea cyclophora Baker
Calathea cylindrica (Roscoe) K. Schum.
Calathea densa (K. Koch) Regel
Calathea dilabens L. Andersson & H. Kenn.
Calathea elliptica (Roscoe) K. Schum.
Calathea erecta L. Andersson & H. Kenn.
Calathea fragilis Gleason
Calathea grandis Petersen
Calathea granvillei L. Andersson & H. Kenn.
Calathea inocephala (Kuntze) H. Kenn. & Nicolson
Calathea lasseriana Steyerm.
Calathea latifolia (Willd. ex Link) Klotzsch
Calathea legrelleana (Linden) Regel
Calathea liesneri H. Kenn.
Calathea lutea (Aubl.) Schult.
Calathea maasioriim H. Kenn.
Calathea mansonis Kom.
Calathea micans (Mathieu) Korn.
Calathea microcephala (Poepp. & Endl.) Kom.
Calathea mishuyacu J.F. Macbr.
Calathea neblinensis H. Kenn.
Calathea ovata (Nees & Mart.) Lindl.
Calathea panamensis Rowlee ex Standi.
Calathea polytricha Baker
Calathea propinqua (Poepp. & Endl.) Kom.
Calathea splendida (Verschaff. ex Lem.) Regel
Calathea squarrosa L. Andersson & H. Kenn.
Calathea variegata Linden ex Korn.
Calathea villosa Lindl.
Calathea zingiberina Kom.
Ctenanthe compressa (A. Dietr.) Eichler
Hylaeanthe hexantha (Poepp. & Endl.) A.M.E. Jonker & Jonker
Hylaeanthe polystachya (Pulle) A.M.E. Jonker & Jonker
Hylaeanthe unilateralis (Poepp. & Endl.) A.M.E. Jonker & Jonker
Ischnosiphon arouma (Aubl.) Kom.
Ischnosiphon bambusaceus (Poepp. & Endl.) Kom.
Ischnosiphon cannoideus L. Andersson
AM
BO
AM BO
AM
AM BO
AM
BO
AM
AM BO
BO
AM BO
AM BO
AM BO
AM
AM BO
BO
BO
AM
AM
AM BO
BO
AM BO
AM BO
AM BO
AM BO
AM BO
DA
DA
DA
DA
DA
DA
DA
GU
GU
GU
SU FG
SU FG?
GU
SU
FG
FG
GU SU FG
FG
GU SU
GU SU FG
GU SU FG
GU SU FG
GU SU FG
SU FG
GU SU FG
GU SU FG
FG
SU? FG?
GU
GU SU FG
FG
SU FG
GU SU
GU SU FG
GU SU FG
SU FG
SU FG?
GU SU FG
GU SU FG
SU? FG?
AM
Checklist of the Plants of the Guiana Shield 115
Ischnosiphon centricifolius L. Andersson
Ischnosiphon enigmaticus L. Andersson
Ischnosiphon foJiosus Gleason
Ischnosiphon gracilis (Rudge) Kom.
Ischnosiphon gracilis subsp. gracilis
Ischnosiphon gracilis subsp. sandwithii L. Andersson
Ischnosiphon hirsutus Petersen
Ischnosiphon leucophaeus (Poepp. & Endl.) Kom.
Ischnosiphon leucophaeus subsp. leucophaeus
Ischnosiphon longiflorus K. Schum.
Ischnosiphon longiflorus subsp. angustifolius L. Andersson
Ischnosiphon martianus Eichler ex Petersen
Ischnosiphon obliquus (Rudge) Kom.
Ischnosiphon petiolatus (Rudge) L. Andersson
Ischnosiphon polyphyllus (Poepp. & Endl.) Kom.
Ischnosiphon puberulus Loes.
Ischnosiphon surumuensis Loes.
Ischnosiphon ursinus L. Andersson
Koernickanthe orbiculata (Kom.) L. Andersson
Maranta amplifolia K. Schum.
Maranta arundinacea L.
Marantafriedrichsthaliana Kom.
Maranta gibba Sm.
Maranta humHis Aubl.
Maranta linearis L. Andersson
Maranta protracta Miq.
Maranta ruiziana Kom.
Maranta rupicola L. Andersson
Monophyllanthe oligophylla K. Schum.
Monotagma contractum Huber
Monotagmajuruanum Loes.
Monotagma laxum (Poepp. & Endl.) K. Schum.
Monotagma ovatum Hagberg
Monotagma plurispicatum (Kom.) K. Schum.
Monotagma rhodanthum Maguire & Wurdack
Monotagma secundum (Petersen) K. Schum.
Monotagma sp. nov. ined.
Monotagma spicatum (Aubl.) J.F. Macbr.
Monotagma tomentosum K. Schum. ex Loes.
Monotagma ulei K. Schum. ex Loes.
Monotagma vaginatum Hagberg
Monotagma yapacanensis Steyerm. & G.S. Bunting
Myrosma cannifolia L. f.
Stromanthejacquinii (Roem. & Schult.) H. Kenn. & Nicolson
Stromanthe tonckat (Aubl.) Eichler
Thalia geniculata L.
Synonyms and Misidentifications
Calathea albicans Brongn. ex K. Schum. =Calathea micans
Calathea allouia (Aubl.) Lindl. [not in flora area] =Calathea legrelleana
Calathea allouia var. violacea auct. non Calathea violacea Lindl. 1825 =Calathea latifolia
Calathea altissima auct. non (Poepp. & Endl.) Kom. 1862 =Calathea inocephala
Calathea barbillana Cufod. =Calathea inocephala
Calathea discolor G. Mey. ^Calathea lutea
Calathea duidae Steyerm. =Calathea fragilis
Calathea hirsuta Standi. =Calathea villosa
Calathea klugii J.F. Macbr. =Calathea micans
Calathea laxa Poepp. & Endl. =Monotagma laxum
Calathea leucophaea Poepp. & Endl. =Ischnosiphon leucophaeus
Calathea magnifica C.V. Morton & Skutch =Calathea lutea
Calathea marantina (Willd.) K. Koch =Calathea casupito
Calathea micans var. amabilis (Linden) Petersen =Calathea micans
Calathea pardina Planch. & Linden =Calathea villosa
Calathea polyphylla Poepp. & Endl. ^Ischnosiphon polyphyllus
GU SU FG
BO GU
GU
FG
AM BO GU SU FG
AM BO GU
GU
SU FG
BO GU
AM BO GU? SU FG
AM BO GU? SU FG
AM GU SU
AM GU SU
SU FG
AM BO DA GU SU FG
GU SU FG
AM
AM BO DA GU SU FG
AM
AM SU
SU
FG
BO GU SU FG
AM BO GU SU FG
FG
BO DA GU SU
AM BO GU? SU FG
AM BO
BO GU SU
AM GU SU FG
BO GU SU
SU
SU
SU
FG
FG
FG
FG
AM BO
BO
GU
GU
SU FG
AM BO GU? SU FG
AM
AM BO GU SU
FG
AM BO DA GU SU FG
AM BO
FG
AM FG
AM
AM BO
BO
GU SU FG
BO GU SU FG
BO DA GU SU FG
116 Checklist of the Plants of the Guiana Shield
Calathea strobilifera (Miq.) Kom. (Zingiberaceae) =Renealmia cernua
Calathea trinilensis Britton =Calathea casupito
Calathea villosa \ar. pardina (Planch. & Linden) Kom. =Calathea villosa
Calathea vittata hort. ex Kom. =Calathea elliptica
Ischnosiphon gracilis var. scabra Petersen =Ischnosiphon pubenilus
Ischnosiphon lasseriana Steyemi. ^Ischnosiphon gracilis
Ischnosiphon phirispicatiis Kom. =Monotagma pliirispicatum
Ischnosiphon puberulus var. scaber (Petersen) L. Andersson =Ischnosiphon puberulus
Ischnosiphon puberuhis var. verruculosus (J.F. Macbr.) L. Andersson =Ischnosiphon puberulus
Ischnosiphon secundus Petersen =Monotagma secundum
Ischnosiphon surinamensis (Miq.) Kom. =Ischnosiphon leucophaeus
Ischnosiphon verruculosous J.F. Macbr. =Ischnosiphon puberulus
Ischnosiphon violaceus Pulle =Ischnosiphon petiolatus
Maranta allouia Aubl. =Calathea legrelleana
Maranta amabilis Linden =Calathea micans
Maranta arouma Aubl. =Ischnosiphon arouma
Maranta cachibou Jacq. =Calathea lutea
Maranta casupito Jacq. =Calathea casupito
Maranta compressa A. Dietr. =Ctenanthe compressa
Maranta divaricata K. Schum. 1902 auct. non Roscoe 1828, A.M.E. Jonker & Jonkerl957 =Maranta gibba
Maranta divaricata Roscoe [not in flora area]
Maranta gracilis Rudge =Ischnosiphon gracilis
Marantajacquinii Roem. & Schult. =Stromanthejacquinii
Maranta lutea Aubl. =Calathea lutea
Maranta lutea Jacq. =Stromanthejacquinii
Maranta micans Mathieu =Calathea micans
Maranta obliqua Rudge =Ischnosiphon obliquus
Maranta orbiculata (Kom.) K. Schum. =Koernickanthe orbiculata
Maranta spicata Aubl. =Monotagma spicatum
Maranta strobilifera Miq. (Zingiberaceae) =Renealmia cernua
Maranta tonckat Auhl. =Stromanthe tonckat
Monotagma duidae Steyerm. =Monotagma tomentosum
Monotagma exannulatum K. Schum. ^Monotagma laxum
Monotagma guianense (Kom.) K. Schum. =Monotagma plurispicatum
Monotagma parkeri (Roscoe) K. Schum. =Monotagma spicatum
Monotagma surinamense Pulle =Monotagma secundum
Myrosma polystachya Pulle =Hylaeanthe polystachya
Myrosma unilateralis (Poepp. & Endl.) K. Schum. =Hylaeanthe unilateralis
Phrynium altissimum Poepp. & Endl. =Calathea altissima
Phrynium densum K. Koch =Calathea densa
Phrynium ellipticum Roscoe =Calathea elliptica
Phrynium micans (Mathieu) Klotzsch =Calathea micans
Phrynium ovatum Nees & Mart. ^Calathea ovata
Phrynium pusillum K. Koch =Calathea micans
Phrynium strobiliferum (Miq.) K. Koch (Zingiberaceae) =Renealmia cernua
Phyllodes inocelphalum Kuntze =Calathea inocephala
Phyllodes platyphyllum Kuntze =Calathea latifolia
Saranthe urceolata Petersen =Maranta humilis
Stromanthe lutea Eichler =Stromanthejacquinii
Thalia latifolia (Willd. ex Link) Link ex Schult. =Calathea latifolia
Thalia trichocalyx Gagnep. =Thalia geniculata
Thalia unilateralis Poepp. & Endl. =Hylaeanthe unilateralis
Thymocarpus cannoides Nicolson, Steyerm. & Sivad. =Calathea cannoides
References
Andersson, L. 1977. The genus Ischnosiphon. Opera Bot. 43: 1-113.
Andersson, L. 1986. Revision of Maranta subgen. Maranta. Nord. J. Bot. 6(6): 729-756.
Andersson, L. 1997. Marantaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 176-284.
Andersson, L. & H. Kennedy. 1986. Four new species of Calathea from French Guiana. Nordic J. Bot. 6(4): 447-455.
Andersson, L., H. Kennedy & M. Hagberg. Marantaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of
the Venezuelan Guayana 6: 219-248.
Checklist of the Plants of the Guiana Shield 117
Hagberg, M. 1990. Two new species of Monotagma from the Venezuelan Guayana. Ann. Missouri Bot. Gard. 77(2): 421-424.
Jonker-Verhoef, A.M.E. & P.P. Jonker. 1957. Marantaceae. In A. Pulle & J. Lanjouw, Plora of Suriname 1(2): 149-208.
Kennedy, H. 1995. Calathea maasiorum (Marantaceae), a new species from Prench Guiana and Surinam. Brittonia 47(2): 156-
159.
A. Lourteigt
AM BO DA GU SU FG
AM BO GU SU FG
AM BO GU SU FG
Mayacafluviatilis Aubl.
Mayaca longipes Mart, ex Seub.
Mayaca sellowiana Kunth
Synonyms and Misidentifications
Mayaca aubletii Michx. =Mayaca fluviatilis
References
Lanjouw, J. 1938. Mayacaceae. In A. Pulle, Flora of Suriname 1(1): 249-251.
Lourteig, A. 1952. Mayacaceae. Notul. Syst. 14: 234-248.
Lourteig, A. 1971. Mayacaceae. In T. Lasser, Flora de Venezuela 3(1): 197-203.
Lourteig, A. 2001. Mayacaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
6: 260-262.
Mori, S.A. 1997. Mayacaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 284.
MUSACEAE
K. Yatskievych
Musa X paradisiaca L. [widely cultivated]
Musa spp. [cultivated only] GU SU FG
Synonyms and Misidentifications
Musa bihai L. (Heliconiaceae) =Helicoma bihai
Musa humilis Auhl. (HeUconiaceae) =Heliconia psittacorum
Musa sapientum L. =Musa ^paradisiaca
References
Maas, P.J.M. 1985. Musaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (Series A) 1: 1-28.
Maas, P.J.M. & M.J.M. de Rooij. 1979. Musaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 393-411.
Mori, S.A. 1997. Musaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide
to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs of the
New York Botanical Garden 76, pt. 1 : 284-286.
Yatskievych, K. 2001 . Musaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan Guayana
6: 731-732.
Najadaceae
R.R. Haynes, L.B. Holm-Nielsen
Najas guadalupensis (Spreng.) Magnus FG?
Najas microcarpa K. Schum. [expected in GU, SU; Mt Roraima] BO FG
Najas wrightiana A. Braun [as circumscribed by Lowden, not in Guianas] BO FG?
Synonyms and Misidentifications
Najas conferta (A. Braun) A. Braun [not in flora area]
Najas marina L. [widespread, perhaps expected]
Najas microdon A. Braun ^Najas guadalupensis
References
Haynes, R.R. & L.B. Holm-Nielsen. 2003. Najadaceae. In J. A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of
the Venezuelan Guayana 7: 100-101.
Lowden, R.M. 1986. Taxonomy of the genus Najas in the Neotropics. Aquatic Bot. 24: 147-184.
118 Checklist of the Plants of the Guiana Shield
Orchidaceae
G. Camevali, E. Christenson, E. Foldats, I.M. Ramirez-Morillo, G.A. Romero-Gonzalez,
C.A. Vargas, M. Werkhoven
Acacallis cyanea Lindl.
Acacallisfimbriata (Rchb. f.) Schltr.
Acineta alticola C. Schweinf.
Acineta superba (Kunth) Rchb. f.
Ada keiliana (Rchb. f. ex Lindl.) N.H. Williams
Aganisia pulchella Lindl.
Aracamunia liesneri Camevali & I. Ramirez
Aspasia variegata Lindl.
Aspidogyne confusa (C. Schweinf.) Garay
Aspidogynefoliosa (Poepp. & Endl.) Garay
Aspidogyne longicornu (Cogn.) Garay
Aspidogyne multifoliata (C. Schweinf.) Garay
Aspidogyne pumila (Cogn.) Garay
Aspidogyne robusta (C. Schweinf.) Garay
Aspidogyne steyermarkii Camevali & Foldats
Barbosella orbicularis Luer
Baskervilla colombiana Garay
Batemannia armillata Rchb. f.
Batemannia colleyi Lindl.
Batemannia lepida Rchb. f.
Beloglottis costaricensis (Rchb. f.) Schltr.
Bifrenaria longicornis Lindl.
Bifrenaria steyermarkii (Foldats) Garay & Dunst.
Bifrenaria venezuelana C. Schweinf.
Bletia altilamellata Garay
Bletia purpurea (Lam.) DC.
Bletia stenophylla Schltr.
Bollea hemixantha Rchb. f.
Bollea violacea (Lindl.) Rchb. f.
Brachionidium brevicaudatum Rolfe
Brachionidium juliani Camevali & L Ramirez
Brachionidium longicaudatum Ames & C. Schweinf.
Brachionidium neblinense Camevali & I. Ramirez
Brachionidium parvum Cogn.
Brachystele guayanensis (Lindl.) Schltr.
Braemia vittata (Lindl.) Jenny
Brassavola cucullata (L.) R. Br.
Brassavola gardneri Cogn.
Brassavola martiana Lindl.
Brassavola nodosa (L.) Lindl.
Brassia angusta Lindl.
Brassia bidens Lindl.
Brassia caudata (L.) Lindl.
Brassia chloroleuca Barb. Rodr.
Brassiaforgetiana hort. ex C. Schweinf.
Brassia huebneri Schltr.
Brassia lanceana Lindl.
Brassia lawrenceana Lindl.
Brassia macrostachya Lindl.
Brassia neglecta Rchb. f.
Brassia pumila Lindl.
Brassia villosa Lindl.
Brassia wageneri Rchb. f.
Bulbophyllum bracteolatum Lindl.
Bulbophyllum dunstervillei Garay
Bulbophyllum exaltatum Lindl.
Bulbophyllum geraense Rchb. f
Bulbophyllum meridense Rchb. f.
Bulbophyllum meristorhachis Garay & Dunst.
Bulbophyllum oerstedii (Rchb. f.) Hemsl.
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
AM BO
AM
AM BO
BO
AM BO
AM BO
AM BO
AM? BO
AM
AM
A \A
BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
GU su
GU?
su
GU
GU su FG
GU su FG
FG
GU su FG
GU su
GU
GU su
GU
SU
GU SU
SU
su
FG
GU su FG
GU
GU
su? FG
GU FG?
GU su FG
GU
GU
GU su FG
GU su FG
GU su FG
su? FG
GU su FG
GU FG
GU
GU
GU su FG
GU FG
FG
GU su FG
GU su FG
GU su
GU su FG
GU FG
GU
GU
GU su FG
GU
GU
SU FG
Checklist of the Plants of the Guiana Shield 119
Bulbophyllum pachyrachis (A. Rich.) Griseb. FG
Bulbophyllum quadrisetum Lindl. SU FG
Bulbophyllum roraimense Rolfe BO GU FG
Bulbophyllum setigerum Lindl. AM GU
Caluera suhnamemis Dodson & Determann SU
Caluera vulpina Dodson & Determann FG
Campylocerrtrumfasciola (Lindl.) Cogn. AM BO DA GU SU FG
Campylocentriim hondurense Ames AM GU
Campylocentrum huebneri Mansf. AM
Campylocentrum lansbergii (Rchb. f.) Schltr. BO DA
Campylocentrum micranthum (Lindl.) Rolfe BO DA GU SU FG
Campylocentrum nataliae Camevali & I. Ramirez BO
Campylocentrum pachyrrhizum {Rchh.f.)Ro\fQ BO GU SU FG
Campylocentrum panamense Ames AM BO
Campylocentrum poeppigii (Rchb. f.) Rolfe AM BO GU
Catasetum barbatum (Lindl) Lindl. AM BO DA GU SU FG
Catasetum barbatum var. spinosum Rolfe FG?
Catasetum bergoldianum Foldats AM
Catasetum bicallosum Cogn. AM GU?
Catasetum bicolor Klotzsch AM BO GU?
Catasetum callosum Lindl. AM GU SU? FG
Catasetum callosum var. carunculatum Mansf. GU SU? FG
Catasetum collare Cogn. AM BO
Catasetum costatum Rchb. f. AM
Catasetum cristatum Lindl. AM BO GU SU FG
Catasetum deltoideum (Lindl.) MvXol GU SU FG?
Catasetum discolor (Lindl.) Lindl. AM BO GU SU FG
Catasetum x dunstervillei G.A. Romero & Camevali [natxiral hybrid of AM BO GU?
C. pileatum and C. discolor]
Catasetum ferox Kraenzl. AM GU?
Catasetum gnomus Linden & Rchb. f. GU?
Catasetum gomezii G.A. Romero & Camevali AM
Catasetum x guianense Jenny & G.A. Romero GU SU? FG
Catasetum longifolium Lindl. AM BO GU SU FG
Catasetum luridum (Link) Lindl. GU
Catasetum macrocarpum Rich, ex Kunth AM BO GU SU FG
Catasetum maroaense G.A. Romero & C. Gomez AM
Catasetum merchae GA. Romero AM
Catasetum microglossum Rolfe AM
Catasetum ochraceum Lindl. GU?
Catasetum parguazense G.A. Romero & Camevali BO
Catasetum pileatum Rchb. L AM BO GU?
Catasetum planiceps Lindl. AM BO GU SU FG?
Catasetum poriferum R.H. Schomb. ex Lindl. GU
Catasetum rivularium Barb. Rodr. AM
Catasetum X roseo-album (Hook..) Lindl. [nsitnxaWxyhndoi C discolor and AM BO GU SU FG?
C longifolium]
Catasetum saccatum Lindl.
Catasetum X tapiriceps Rchb. f [natural hybrid of C. macrocarpum and C.
pileatum]
Catasetum thompsonii Dodson
Catasetum x wendlingeri Foldats [natural hybrid of C. pileatum and C.
planiceps]
Catasetum yavitaense G.A. Romero & C Gomez
Cattleyajenmanii Rolfe
Cattleya lawrenceana Rchb. f
Cattleya violacea (Kunth) Rolfe
Caularthron bicornutum (Hook.) Raf.
Caularthron bilamellatum (Rchb. f.) R.E. Schult.
Caularthron bilamellatum f indivisa (Bradf ex Griseb.) Camevali & I.
Ramirez
Chaubardia surinamensis Rchb. f.
Chaubardiella tigrina (Garay & Dunst.) Garay
Cheiradenia cuspidata Lindl.
GU
AM BO GU?
GU
FG'
AM BO GU
AM
BO GU
AM BO GU
AM BO DA GU SU FG
AM BO
BO
GU
GU?
SU
BO GU SU FG
BO GU SU FG
BO GU SU FG
120 Checklist of the Plants of the Guiana Shield
Chelyorchis ampliata (Lindl.) Dressier & N.H. Williams
Cleistes grandiflora (Aubl.) Schltr.
Cleistes huberi Camevali & I. Ramirez
Cleistes lepida (Rchb. f.) Schltr.
Cleistes pahidosa Rchb. f.
Cleistes parviflora Lindl.
Cleistes rosea Lindl.
Cleistes striata (C. Schweinf.) Garay & Dunst.
Cleistes tenuis (Rchb. f. ex Griseb.) Schltr.
Cleistes triflora (C. Schweinf.) Camevali & I. Ramirez [Mt. Roraima]
Cleistes unifoliata (C. Schweinf.) Camevali & I. Ramirez
Clowesia warczewiczii (Lindl. & Paxton) Dodson
Cochleanthes flabelliformis (Sw.) R.E. Schult. & Garay
Cochleanthes guianensis Lafontaine, G Gerlach & Senghas
Comparettiafalcata Poepp. & Endl.
Coryanthes albertinae H. Karst.
Coryanthes cataniapoensis GA. Romero & Camevali
Coryanthes feildingii Lindl.
Coryanthes gomezii GA. Romero & G Gerlach
Coryanthes macrantha (Hook.) Hook.
Coryanthes macidata Hook.
Coryanthes pegiae GA. Romero
Coryanthes punctata Beer
Coryanthes rutkisii Foldats
Coryanthes speciosa (Hook.) Hook.
Corymborkis sp.
Cranichis diphylla Sw.
Cranichis sp.
Cryptarrhena guatemalensis Schltr.
Cryptarrhena kegelii Rchb. f.
Cryptocentrum dunstervilleorum Camevali & GA. Romero
Cryptocentrum peruvianum (Cogn.) C. Schweinf.
Cryptocentrum peruvianum subsp. peruvianum
Cryptocentrum sp. nov. {Cryptocentrum peruvianum auct., Cryptocentrum
gracillimum auct.]
Cyclopogon cranichoides (Griseb.) Schltr.
Cyclopogon elatus (Sw.) Schltr.
Cyclopogon olivaceus (Rolfe) Schltr.
Cycnoches chlorochilon Klotzsch
Cycnoches haagii Barb. Rodr.
Cycnoches loddigesii Lindl.
Cycnoches lusiae GA. Romero & Garay
Cycnoches thurstonorum Dodson
Cyrtopodium andersonii (Lamb, ex Andrews) R. Br.
Cyrtopodium cristatum Lindl.
Cyrtopodium fowliei L.C. Menezes
Cyrtopodium graniticum GA. Romero & Camevali
Cyrtopodium intermedium Brade
Cyrtopodium paniculatum (Ruiz & Pav.) Garay
Cyrtopodium parviflorum Lindl.
Degranvillea dermaptera Determann
Diadenium barkeri (Lindl.) Benth. & Hook. f.
Dichaea ancoraelabia C. Schweinf.
Dichaea angustifolia H.G Jones
Dichaea brevicaulis Cogn.
Dichaea biyophila Rchb. f.
Dichaea camaridioides Schltr. [Mt. Roraima]
Dichaea cornuta S. Moore
Dichaea histrio Rchb. f.
Dichaea hookeri Garay & H.R. Sweet
Dichaea hystricina Rchb. f.
Dichaea kegelii Rchb. f.
Dichaea latifolia Lindl.
Dichaea latifolia var. longa (Schltr.) Folsom
BO DA
GU FG
BO
BO GU
su FG
AM BO GU su FG
AM BO GU su FG
AM BO
AM BO
BO
BO
GU su
su
FG
AM BO
BO
GU? su?
GU su FG
AM BO FG
AM BO GU su FG
AM
BO GU
AM
AM BO GU su FG
BO GU su FG
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
AM
AM
BO
BO
BO
AM BO
BO
BO
AM
AM BO
AM BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
GU
GU SU FG
GU su?
su
su FG
GU su FG
GU su FG
GU
su
GU su FG
GU su? FG
GU su FG
GU su FG
GU su FG
GU su FG
GU su
su
GU su FG
GU su FG
FG
GU su FG
GU
FG?
FG?
GU?
su
GU? su?
GU SU FG
GU
GU su FG
GU?
GU
AM BO DA
AM BO GU FG
AM BO DA GU
GU SU FG
AM BO DA
AM BO DA GU SU FG
AM
AM GU SU FG
AM BO GU SU FG
AM BO GU
GU SU
AM BO DA GU SU FG
BO DA GU SU FG
BO
BO
BO
AM BO
AM
BO
AM BO GU
FG
GU SU FG
GU SU FG
AM BO
AM BO
AM BO
AM BO
Checklist of the Plants of the Guiana Shield 121_
Dichaea morris ii Fawc. & Rendle
Dichaea ochracea Lindl.
Dichaea panamensis Lindl.
Dichaea pendula (Aubl.) Cogn.
Dichaea picta Rchb. f.
Dichaea rendlei Gleason
Dichaea robiista Schltr.
Dichaea splitgerberi Rchb. f.
Dichaea tachirensis G.A. Romero & Camevali
Dichaea trinitensis Gleason
Dichaea trulla Rchb. f.
Dichaea venezuelensis Camevali & I. Ramirez
Dichaea weigeltii Rchb. f.
Dimerandra elegans (H. Focke) Siegerist
Dimerandra emarginata (G. Mey.) Hoehne
Dipteranthus planifolius (Rchb. f.) Garay
Discyphus scopiilariae (Rchb. f.) Schltr.
Dryadella lueriana Camevali & G.A. Romero
Duckeella alticola C. Schweinf. [expected in GU]
Duckeella pauciflora Garay
Dunstervillea mirabilis Garay
Elleanthus arpophyllostachys (Rchb. f.) Rchb. f.
Elleanthiis brasiliensis (Lindl.) Rchb. f.
Elleanthus caravata (Aubl.) Rchb. f. AM BO
Elleanthus cephalotus Garay & H.R. Sweet
Elleanthus columnaris (Lindl.) Rchb. f.
Elleanthus confusus Garay
Elleanthus crinipes Rchb. f. FG
Elleanthusfurfuraceus (Lindl.) Rchb. f.
Elleanthus gracilis (Rchb. f.) Rchb. f.
Elleanthus graminifolius {^?i.rh.Kodx.)L0]indin\ GU SU FG
Elleanthus linifolius C. Presl
Elleanthus malpighiiflorus Camevali & G.A. Romero [expected in AM;
Serra da Neblina]
Elleanthus norae Garay & Dunst.
Elleanthus sphaerocephalus Schltr.
Elleanthus trilobatus Ames & C. Schweinf.
Elleanthus wageneri (Rchb. f.) Rchb. f.
Eloyella panamensis (Dressier) Dodson
Eltroplectris calcarata (Sw.) Garay & H.R. Sweet
Encyclia auyantepuiensis Camevali & L Ramirez
Encyclia chloroleuca (Hook.) Neumann
Encyclia cochleata (L.) Lemee
Encyclia conchaechila (Barb. Rodr.) Porto & Brade
Encyclia cordigera (Kunth) Dressier
Encyclia dichroma (Lindl.) Schltr.
Encyclia diurna (Jacq.) Schltr.
Encyclia granitica (Bateman ex Lindl.) Schltr.
Encyclia guianensis Camevali & G.A. Romero
Encyclia ionosma (Lindl.) Schltr.
Encyclia ivonae Camevali & G.A. Romero
Encyclia leucantha Schltr.
Encyclia longifolia Barb. Rodr.
Encyclia odoratissima (Lindl.) Schltr.
Encyclia pachyantha (Lindl.) Hoehne
Encyclia picta (Lindl.) Hoehne
Encyclia pilosa (C. Schweinf.) Camevali & I. Ramirez
Encyclia remotiflora (C. Schweinf) Camevali & I. Ramirez AM
Encyclia selligera (Bateman ex Lindl.) Schltr. [not in flora area, fide E. SU?
Christenson, specimens =E. granitica]
Encyclia tarumana Schltr. FG
Epidendrum agathosmicum Rchb. f
Epidendrum alsum Ridl. ex Thum
Epidendrum althausenii A.D. Hawkes AM DA GU SU FG
GU SU
SU
AM BO
AM BO
BO
BO
BO
BO
GU
GU
SU
AM BO DA GU SU FG
GU SU? FG
AM GU
BO GU
SU
FG
BO GU SU FG
BO DA GU SU FG
BO GU SU
GU SU? FG
BO GU
AM BO
GU
SU?
FG
DA GU
GU
SU? FG
AM GU
AM
BO GU
122 Checklist of the Plants of the Guiana Shield
AM
AM
Epidendrum amazonicoriifolium Hagsater AM
Epidendrum anceps Jacq.
Epidendrum apaganum Mansf.
Epidendrum apuahuense Mansf.
Epidendrum attenuatum Lindl.
Epidendrum bakrense Hagsater & Cremers
Epidendrum calanthum Rchb. f. & Warsz.
Epidendrum carpophorum Barb. Rodr.
Epidendrum caurense Camevali & G.A. Romero
Epidendrum chimantense Hagsater & Camevali
Epidendrum churunense Garay & Dunst.
Epidendrum ciliare L.
Epidendrum ciliare var. squamatum Schnee
Epidendrum commelinispathum Camevali & I. Ramirez [expected in AM;
Serra da Neblina]
Epidendrum compressum Griseb. AM
Epidendrum congestoides Ames & C. Schweinf.
Epidendrum cooperianum Bateman
Epidendrum coronatum Ruiz & Pav. AM
Epidendrum cristatum Ruiz & Pav.
Epidendrum densiflorum Hook.
Epidendrum dichaeoides Camevali & G.A. Romero AM
Epidendrum difforme Jacq.
Epidendrum diffusum Sw.
Epidendrum diffusum var. depauperatum Cogn. [=Epidendrum compressuml]
Epidendrum dipus Lindl.
Epidendrum durum Lindl. AM
Epidendrum ecostatum Pabst ex Angley
Epidendrumflexuosum G Mey. AM
Epidendrum fraudulentum Rchb. f.
Epidendrum geminiflorum Kunth
Epidendrum gracillimum Lindl.
Epidendrum gramineum Lindl.
Epidendrum gratissimum (Rchb. f.) Hagsater
Epidendrum hombersleyi Summerh. AM
Epidendrum ibaguense Kunth AM
Epidendrum ibaguense Kunth x secundum Jacq.
Epidendrum imitans Schltr.
Epidendrum imthurnii Ridl. ex Thum AM
Epidendrum isomerum Schltr.
Epidendrum klotzscheanum Rchb. f. AM
Epidendrum latilabre Lindl.
Epidendrum leucochilum Klotzsch
Epidendrum longicolle Lindl. AM
Epidendrum macrocarpum Rich. AM
Epidendrum magnicallosum C. Schweinf. AM
Epidendrum micronocturnum Camevali & GA. Romero
Epidendrum microphyllum Lindl. AM
Epidendrum miserrimum Rchb. f. AM
Epidendrum montigenum Ridl. ex Thum
Epidendrum myrmecophorum Barb. Rodr. AM
Epidendrum nocturnum Jacq. AM
Epidendrum norae Camevali & G.A. Romero AM
Epidendrum nuriense Camevali & Hagsater
Epidendrum nutans Sw.
Epidendrum oldemanii Christenson
Epidendrum orchidiflorum Salzm. AM
Epidendrum pachyphyton Garay AM
Epidendrum paniculatum Ruiz & Pav. [=Epidendrum densiflorum?]
Epidendrum paruimense G.A. Romero & Camevali [expected in BO]
Epidendrum patens Sw.
Epidendrum prostratum (Lindl.) Cogn.
Epidendrum pseudoramosum Schltr.
GU
GU SU
su
BO
FG
FG
BO GU?
BO GU SU FG
BO
BO
BO GU
BO GU SU FG
BO
BO DA
DA
GU
GU
SU FG
BO SU
BO GU SU
BO GU SU FG
GU SU FG
GU SU
GU SU
FG
BO GU
GU FG
BO DA GU
GU?
GU
SU
SU
SU?
SU?
FG
BO GU SU? FG
BO
GU FG
BO GU
GU
FG
GU SU? FG
BO
BO DA GU
BO GU SU FG
BO
BO DA GU SU FG
BO GU SU
BO GU
BO
BO DA GU SU FG
BO GU
BO
SU
FG
BO GU SU FG
BO GU
GU SU FG
GU
GU?
BO
BO
GU
GU
Checklist of the Plants of the Guiana Shield 123
Epidendrum purpurascens H. Focke
Epidendrum ramosum Jacq.
Epidendrum repens Cogn.
Epidendrum revertianum (Stehle) Hagsater
Epidendrum rigidum Jacq.
Epidendrum rostratum Garay & Dunst.
Epidendrum saxatile Lindl.
Epidendrum schlechterianum Ames
Epidendrum schlimii Rchb. f.
Epidendrum sculptum Rchb. f.
Epidendrum sculptum van linearifolium Rchb. f.
Epidendrum sculptum var. sculptum
Epidendrum secundum Jacq.
Epidendrum sertorum Garay & Dunst.
Epidendrum smaragdinum Lindl.
Epidendrum sp. A
Epidendrum splendens Schltr.
Epidendrum stalkyi Camevali & GA. Romero
Epidendrum stamfordianum Bateman
Epidendrum stiliferum Dressier
Epidendrum strobiliferum Rchb. f.
Epidendrum strobiloides Garay & Dunst.
Epidendrum tenuifolium Lindl.
Epidendrum tridens Poepp. & Endl.
Epidendrum tumuc-humaciense (Veyret) Camevali & G.A. Romero
Epidendrum ulei Schltr.
Epidendrum unguiculatum (C. Schweinf.) Garay & Dunst.
Epidendrum urbanianum Cogn.
Epidendrum urichianum Camevali, Foldats & I. Ramirez
Epidendrum vincentinum Lindl.
Epidendrum violascens Ridl. ex Thum
Epidendrum viviparum Lindl.
Epidendrum xanthium Lindl.
Epistephium amplexicaule (Ruiz & Pav.) Poepp. & Endl.
Epistephium duckei Huber
Epistephium elatum Kunth
Epistephium ellipticum R.O. Williams & Summerh.
Epistephium hernandii Garay
Epistephium lucidum Cogn.
Epistephium parviflorum Lindl.
Epistephium sclerophyllum Lindl.
Epistephium subrepens Hoehne
Eriopsis biloba Lindl.
Eriopsis rutidobulbon Hook.
Eriopsis sceptrum Rchb. f. & Warsz.
Erycina glossomystax (Rchb. f.) N.H. Williams & M.W. Chase
Erycina pumilio (Rchb. f.) N.H. Williams & M.W. Chase
Eiycina pusilla (L.) N.H. Williams & M.W. Chase
Erythrodes arietina (Rchb. f. & Warm.) Ames
Erythrodes hirtella (Sw.) Fawc. & Rendle
Erythrodes major (Presl) Ames
Erythrodes paleacea (Schltr.) Ames
Erythrodes pumila (Cogn.) Pabst
Eulophia alta (L.) Fawc. & Rendle
Eurystyles cotyledon Wawra
Galeandra badia Garay & G.A. Romero
Galeandra baueri Lindl.
Galeandra baueri var. baueri
Galeandra baueri var. lutea S. Watson
Galeandra beyrichii Rchb. f.
Galeandra carnevaliana GA. Romero & Warford
Galeandra devoniana R.H. Schomb. ex Lindl.
Galeandra dives Rchb. f.
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
AM BO
AM BO
AM BO
AM
AM BO
AM
AM BO
AM BO
BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
BO
BO
BO
DA
DA
DA
DA
DA
DA
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
SU
SU
SU
SU
SU
SU
GU SU
GU
FG
FG
FG
FG
FG
FG
SU FG
SU?
GU SU FG
GU SU FG
GU
GU SU
GU SU FG
GU
SU FG
GU
GU
GU
GU FG
GU?
GU
GU SU FG
GU
GU SU
SU?
FG
GU
GU SU FG
GU SU FG
GU SU FG
SU FG
GU SU FG
GU SU? FG
GU SU
GU
SU
GU SU FG
GU SU FG
FG
FG
FG
GU SU FG
AM GU
GU SU FG
124 Checklist of the Plants of the Guiana Shield
Galeandra diiidemis Garay & G.A. Romero AM
Galeandra lagoensis Rchb. f. & Warm. SU
Galeandra macroplectra G.A. Romero & Warford AM
Galeandra magnicohimna G.A. Romero & Warford AM
Galeandra minax Rchb. f. AM
Galeandra stangeana Rchb. f. AM
Galeandra stylomisantha (Veil.) Hoehne
Galeottia burkei (Rchb. f.) Dressier & Christenson
Galeottiajorisiana (Rolfe) Schltr.
Gomphichis adnata (Ridl. ex Thum) Schltr. [=Gomphichis viscosal]
Gomphichis costaricensis (Schltr.) Ames, Hubb. & C. Schweinf.
Gomphichis viscosa (Rchb. f.) Schltr.
Gongora atropurpurea Hook.
Gongora histrionica Rchb. f.
Gongora maculata Lindl.
Gongora nigrita Lindl.
Gongora pleiochroma Rchb. f.
Govenia utricidata (Sw.) Lindl.
Guanchezia maguirei (C. Schweinf.) G.A. Romero & Camevali
Habenaria abortiens Lindl.
Habenaria alata Hook.
Habenaria alterosula Snuv. & Westra
Habenaria amambayensis Schltr. [expected in BO and AM]
Habenaria armata Rchb. f.
Habenaria ayangannensis Renz [expected in BO]
Habenaria caldensis Kraenzl.
Habenaria distans Griseb.
Habenaria dusenii Schltr.
Habenaria entomantha (La Llave & Lex.) Lindl.
Habenaria ernestii Schltr.
Habenariafloribunda Lindl.
Habenaria heptadactyla Rchb. f
Habenaria huberi Carnevali & Morillo
Habenaria lehmanniana Kraenzl.
Habenaria leprieiiri Rchb. f.
Habenaria longicaiida Hook.
Habenaria longicauda subsp. ecalcarata Snuv. & Westra
Habenaria longicauda subsp. longicauda
Habenaria macilenta (Lindl.) Rchb. f.
Habenaria mesodactyla Griseb. BO
Habenaria monorrhiza (Sw.) Rchb. f. [expected in the Guianas] BO
Habenaria montis-wilhelminae Renz SU
Habenaria nilssonii Foldats BO
Habenaria obtusa Lindl. BO
Habenaria odontopetala Rchb. f.
Habenaria parvidens Lindl.
Habenaria parviflora Lindl. BO
Habenaria platydactyla Kraenzl.
Habenaria pratensis {L'\nd\.)KQ\\bA. BO
Habenaria pratensis \ax. parviflora Cogn. BO
Habenaria pygmaea C. Schweinf. & R.E. Schult. AM
Habenaria quadrata Lindl. AM BO
Habenaria quinqueseta (Michx.) A. Eaton BO
Habenaria repens Nutt. BO DA
Habenaria rodeiensis Barb. Rodr.
Habenaria rodriquezii Cogn.
Habenaria roraimensis Rolfe [Mt. Roraima] AM BO
Habenaria santens is Barb. Rodr.
Habenaria schomburgkii Lindl. ex Benth. BO
Habenaria setacea Lindl. BO
Habenaria setacea var. setacea BO
Habenaria seticauda Lindl. ex Benth.
Habenaria sp. A AM
Habenaria sp. B AM
AM BO GU SU FG
AM BO GU SU?
AM BO
BO
BO
GU
GU
GU
AM BO DA GU SU FG
GU? SU FG?
GU FG?
GU SU?
AM BO DA GU SU
SU?
FG
AM
SU?
FG?
GU SU FG
GU FG
AM BO GU
GU
BO GU SU FG?
BO
AM BO SU FG
GU SU FG?
AM BO
BO
GU
GU
FG
AM BO GU SU
AM
BO GU FG
AM BO GU SU FG
AM BO DA GU SU FG
GU SU FG
GU SU
GU SU FG
GU SU FG?
GU
SU?
GU SU? FG?
GU SU FG
GU? SU FG
GU? SU FG
GU SU FG?
GU SU FG
GU FG
FG
GU
SU?
GU SU
GU FG
GU FG
GU FG
Checklist of the Plants of the Guiana Shield 125
Habenaria spnicei Cogn.
Habenaha trifida Kunth
Helonoma americana (C. Schweinf. & Garay) Garay
Helonoma bifida (Ridl. ex Thum) Garay
Helonoma chiropterae (Szlach.) Camevali & G.A. Romero
Heterotaxis proboscidea (Rchb. f.) F. Barros
Heterotaxis superfiua (Rchb. f.) F. Barros
Heterotaxis villosa (Barb. Rodr.) F. Barros
Heterotaxis violaceopimctata (Rchb. f.) F. Barros
Hexisea bidentata Lindl.
Hexisea imbricata (Lindl.) Rchb. f.
Houlletia odoratissima Linden ex LindL
Hoidletia roraimensis Rolfe
Himtleya lucida (Rolfe) Rolfe
Huntleya meleagris Lindl.
Hundeya sessilifiora Bateman ex Lindl.
Hylaeorchis petiolaris (Schltr.) Camevali & G.A. Romero
lonopsis satyrioides (Sw.) Rchb. f.
lonopsis utricularioides (Sw.) Lindl.
Isochihis linearis (Jacq.) R. Br.
Jacquiniella globosa (Jacq.) Schltr.
Jacqiiiniella leucomelana (Rchb. f ) Schltr.
Jacquiniella steyermarkii Camevali & Dressier
Jacquiniella teretifolia (Sw.) Britton & R Wilson
Kefersteinia lafontainei Senghas & G. Gerlach
Kegeliella houtteana (Rchb. f.) L.O. Williams
Kegeliella orientalis G. Gerlach
Koellensteinia carraoensis Garay & Dunst.
Koellensteinia graminea (Lindl.) Rchb. f.
Koellensteinia hyacinthoides Schltr.
Koellensteinia kellneriana Rchb. f.
Koellensteinia lilijae Foldats
Koellensteinia roraimae Schltr.
Koellensteinia tricolor (Lindl.) Rchb. f
Leochihts labiatus (Sw.) Kuntze
Lepanthes aithalos Camevali & I. Ramirez
Lepanthes cercion Luer & Escobar
Lepanthes duidensis Ames & C. Schweinf.
Lepanthes dussii Urb.
Lepanthes exilis C. Schweinf
Lepanthes helicocephala Rchb. f.
Lepanthes marahuacensis Camevali & I. Ramirez
Lepanthes pariaensis Foldats
Lepanthes pectinata Luer
Lepanthes ruscifolia Rchb. f.
Lepanthes iinitrinervis Camevali & 1. Ramirez
Lepanthopsis fioripecten (Rchb. f.) Ames
Lepanthopsis obliquipetala (Ames & C. Schweinf.) Luer
Lepanthopsis pulchella Garay & Dunst.
Lepanthopsis vinacea C. Schweinf
Leucohyle mutica (Lindl.) Schltr.
Leucohyle subulata (Sw.) Schltr.
Ligeophila amazonica Garay
Ligeophila clavigera (Rchb. f ) Garay
Ligeophilajuruensis (Hoehne) Garay
Ligeophila peterana (Cogn.) Garay
Ligeophila rosea (Lindl.) Garay
Ligeophila stigmatoptera (Rchb. f ) Garay
Liparis jamaicensis Lindl. ex Griseb.
Liparis nervosa (Thunb.) Lindl. subsp. A
Liparis nervosa subsp. nervosa
Liparis verticillata G.A. Romero & Garay
Lockhartia acuta (Lindl.) Rchb. f DA
Lockhartia biserra (Rich.) Christenson & Garay GU
BO GU SU FG
AM BO GU SU FG
AM
AM
BO
BO
GU
GU
AM
AM
AM BO GU SU FG
AM BO GU SU FG
AM BO DA GU SU FG
AM BO GU SU? FG
AM BO GU SU
AM BO GU SU?
AM BO
BO
GU
GU
GU
GU
AM BO GU
BO DA GU SU FG
AM BO DA GU SU FG
BO GU SU FG
AM BO GU
GU
SU FG
AM BO GU
AM BO GU SU FG
GU? SU
SU
FG
FG
BO
BO GU SU FG
AM BO DA GU SU FG
AM SU? FG?
GU SU FG
AM
BO
AM BO
BO
BO
BO
GU
AM BO
AM BO
BO
AM BO GU SU FG
AM
AM BO
AM BO
BO
SU FG
AM
AM
BO
BO
BO
BO
GU FG
A\/f
BO DA GU
GU
SU
AM
GU SU FG
AM BO SU
SU
SU?
FG
AM
AM
BO
BO
BO
GU
GU
SU FG
AM BO
BO
DA GU? SU FG
AM BO
BO
BO
BO
BO
BO
AM
AM BO
AM BO
BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
BO
AM BO
AM BO
AM BO
BO
AM BO
AM BO
AM BO
AM
BO
AM BO
BO
AM BO
AM BO
BO
AM
AM
AM
BO
AM BO
AM BO
AM BO
BO
GU su FG
GU
GU
su FG
GU su
su
FG
GU su FG
GU su
GU su FG
su FG
GU su
GU
su FG
GU su
su
FG
GU su FG
su FG
126 Checklist of the Plants of the Guiana Shield
Lockhartia imbricata (Lam.) Hoehne DA
Lockhartia latilabhs C. Schweinf.
Lockhartia parthenocomos Rchb. f.
Lueddemannia pescatorei (Lindl.) Linden & Rchb. f.
Lycastefulvescens Hook.
Lycaste macrophylla (Poepp. & Endl.) Lindl.
Lyroglossa grisebachii (Cogn.) Schltr.
Macradenia brassavolae Rchb. f.
Macradenia loxoglottis H. Focke & Rchb. f. ex Rchb. f.
Macradenia lutescens R. Br.
Macradenia rubescens Barb. Rodr. DA
Macroclinium mirabile (C. Schweinf.) Dodson
Macroclinium wullschlaegelianum (H. Focke) Dodson
Malaxis excavata (Lindl.) Kuntze
Malaxis maguirei C. Schweinf. AM BO
Malaxis umbelUfera Sw.
Masdevallia cuprea Lindl.
Masdevallia guayanensis Lindl. BO
Masdevallia infracta Lindl.
Masdevallia manarana Camevali & I. Ramirez AM
Masdevallia minuta Lindl.
Masdevallia norae Luer
Masdevallia picturata Rchb. f. GU
Masdevallia sprucei Rchb. f
Masdevallia vargasii C. Schweinf. GU
Masdevallia wendlandiana Rchb. f.
Maxillaria acutifolia Lindl.
Maxillaria alba (Hook.) Lindl.
Maxillaria albiflora Ames & C. Schweinf.
Maxillaria alpestris Lindl.
Maxillaria alticola C. Schweinf.
Maxillaria aurea (Poepp. & Endl.) L.O. Williams
Maxillaria auyantepuiensis Foldats
Maxillaria bolivarensis C. Schweinf. DA
Maxillaria brachybulbon Schltr.
Maxillaria caespitifica Rchb. f
Maxillaria camaridii Rchb. f. DA
Maxillaria Candida Lodd. ex Lindl.
Maxillaria chlorantha Lindl.
Maxillaria christobalensis Rchb. f.
Maxillaria colemanii Camevali & W. Fritz [expected in GU]
Maxillaria conferta (Griseb.) C. Schweinf ex Leon DA
Maxillaria connellii Rolfe
Maxillaria crassifolia (Lindl.) Rchb. f.
Maxillaria cryptobulbon Camevali & J.T. Atwood
Maxillaria desvauxiana Rchb. f.
Maxillaria discolor (Lodd. ex Lindl.) Rchb. f DA
Maxillaria x dunstervillei Camevali & I. Ramirez [natural hybrid ofM
meridensis Lindl. and M. quelchii Rolfe]
Maxillaria eburnea Lindl. GU?
Maxillaria equitans (Schltr.) Garay
Maxillariafoldatsiana Camevali & 1. Ramirez
Maxillaria grobyoides Garay & Dunst.
Maxillaria guadalupensis Cogn.
Maxillaria guareimensis Rchb. f.
Maxillaria imbricata Barb. Rodr.
Maxillariajenischiana (Rchb. f.) C. Schweinf.
Maxillaria kegelii Rchb. f. AM
Maxillaria lasallei Foldats BO
Maxillaria leucaimata Barb. Rodr.
Maxillaria liparophylla Summerh. GU
Maxillaria longipetiolata Ames & C. Schweinf AM BO SU
Maxillaria loretoensis C. Schweinf. AM BO
Maxillaria luteoalba Lindl. AM BO
GU FG
GU SU FG
GU
GU
GU
GU su
GU
GU SU FG
GU su FG
GU su FG
GU
GU
GU su FG
GU
GU
GU FG
GU su FG
GU su FG
GU
GU
su
GU SU
GU su
GU su FG
GU
FG
Checklist of the Plants of the Guiana Shield 127
Maxillaha mapihemis (Kraenzl.) L.O. Williams AM BO GU SU
Maxillaria marmoUana Dodson AM BO
Maxillaria melina Lindl. GU
Maxillaria meridensis Lindl. AM BO DA GU
Maxillaria monacensis Kraenzl. GU?
Maxillaria nasuta Rchb. f. AM BO GU
Maxillaria notylioglossa Rchb. f.
Maxillaria miriensis Camevali & I. Ramirez
Maxillaria ochroleuca Lodd. ex Lindl.
Maxillaria ochroleuca var. longipes Sander
Maxillaria pannieri Foldats
Maxillaria parkeri Hook.
Maxillaria parviflora (Poepp. & Endl.) Garay
Maxillaria pauciflora Barb. Rodr.
Maxillaria ponerantha Rchb. f.
Maxillaria porrecta Lindl.
Maxillaria prolifera Ruiz & Pav.
Maxillaria pterocarpa Barb. Rodr.
Maxillaria pumila Hook.
Maxillaria quelchii Rolfe AM BO DA
Maxillaria ramosa Ruiz & Pav.
Maxillaria reichenheimiana Endres & Rchb. f.
Maxillaria rigida Barb. Rodr.
Maxillaria nifescens Lindl. AM BO DA
Maxillaria santanae Camevali & I. Ramirez
Maxillaria schlechteri Foldats
Maxillaria scorpioidea Kraenzl.
Maxillaria setigera Lindl.
Maxillaria splendens Poepp. & Endl.
Maxillaria squamata Barb. Rodr.
Maxillaria stenophylla Rchb. f.
Maxillaria tenuis C. Schweinf.
Maxillaria uncata Lindl. AM BO DA
Maxillaria venezuelana (C. Schweinf.) Camevali ined
Maxillaria xylobiiflora Schltr.
Mesadenella angustisegmenta Garay
Mesadenella cuspidata (Lindl.) Garay
Miltoniopsis santanaei Garay & Dunst. BO
Mormodes atropurpurea Lindl.
Mormodes buccinator Lindl. SU EG
Mormodes carnevaliana Salazar & G.A. Romero AM BO DA
Mormodes cucumerina Pabst AM
Mormodes igneum Lindl. & Paxton FG
Mormodes orinocoensis Salazar & G.A. Romero AM BO
Mormodes vernixioidea Pabst
Mormodes vernixioidea subsp. autanensis G.A. Romero & Salazar
Mormodes vernixium Rchb. f.
Myoxanthus octomeriae (Schltr.) Luer
Myoxanthus parahybunensis (Barb. Rodr.) Luer
Myoxanthus parvilabius (C. Schweinf.) Luer
Myoxanthus reymondii (H. Karst.) Luer
Myoxanthus simplicicaulis (C. Schweinf.) Luer
Myoxanthus speciosus (Luer) Luer
Myoxanthus trachychlamys (Schltr.) Luer
Nidema ottonis (Rchb. f.) Britton & Millsp.
Notylia angustifolia Cogn.
Notylia aromatica Barker ex Lindl.
Notylia bungerothii Rchb. f.
Notylia durandiana Cogn.
Notylia fragrans H. Focke
Notylia laxa Rchb. f
Notylia longispicata Hoehne & Schltr.
Notylia micrantha Lindl.
Notylia microchila Cogn.
AM BO
BO
AM BO
AM BO
AM BO
AM
AM BO
BO
BO
AM BO
AM
AM BO
BO
BO
AM BO
AM BO
AM BO
BO
AM
AM BO
BO
GU
GU SU FG
GU SU FG
GU
GU SU FG
GU SU
SU
FG
GU
GU
GU SU
GU SU FG
GU SU FG
GU
GU SU FG
GU FG
GU
GU SU?
GU SU FG
GU SU FG
GU SU FG
GU
GU SU
GU SU
GU?
AM
A Ayr
GU? SU
AM
GU?
GU
GU SU
AM GU SU
BO
AM BO GU
AM BO
AM BO GU SU
AM BO GU
AM BO FG
AM BO GU
GU
SU
SU
FG
AM
AM
BO
GU
GU
SU
SU
FG
AM BO GU
128 Checklist of the Plants of the Guiana Shield
Notylia pentachne Rchh. f. BO
Notylia peruviana (Sch\tr.)C.Schwemf. AM BO FG
Notylia platyglossa Schltr. AM BO
Notylia rhombilabia C. Schweinf. AM BO GU
Notylia sagittifera (Kunth) Link, Klotzsch & Otto GU SU FG
Notylia yauaperyemis Barb. Rodr. AM BO FG
Octomeria amazonica Pabst FG
Octomeria anomala Garay & Dunst. AM BO
Octomeria apiculata (Lindl.) Garay & H.R. Sweet BO FG?
Octomeria connellii Rolfe BO GU
Octomeria cordilabia C. Schweinf. AM BO GU
Octomeria dentifera C. Schweinf. BO GU
Octomeria erosilabia C. Schweinf. AM BO GU
Octomeria exigua C. Schweinf. AM BO GU SU FG
Octomeria filifolia C. Schweinf. AM BO GU
Octomeriaflaviflora C. Schweinf. AM BO GU
Octomeria gemmula Camevali & I. Ramirez AM
Octomeria graminifolia R. Br. FG
Octomeria heleneana Camevali & Delascio AM
Octomeria integrilabia C. Schweinf. AM BO GU
Octomeria lancipetala C. Schweinf. AM BO GU
Octomeria minor C. Schweinf. AM BO GU SU
Octomeria monticola C. Schweinf. BO GU
Octomeria nana C. Schweinf. AM BO
Octomeria oxychela Barb. Rodr. GU
Octomeria parvifolia Rolfe AM BO GU
Octomeria parvula C. SchwQinf. BO GU
Octomeria petulans Rchb. f. FG
Octomeria rhizomatosa C. Schweinf. AM BO
Octomeria romerorum Camevali & I. Ramirez AM
Octomeria sarthouae Luer FG
Oc/owerw 5c/>po/Wea (Poepp. &Endl.) Rchb. f. AM BO GU SU FG
Octomeria sp. A AM
Octomeria spathulata Rchb. f. AM BO
Octomeria splendida Garay & Dunst. BO
Octomeria steyermarkii Garay & Dunst. BO
Octomeria surinamensis H. Focke AM BO GU SU FG
Octomeria tapiricataractae GA. Romero & C.A. Luer AM
Octomeria taracuana Schltr. AM BO
Octomeria yauaperyensis Barh. Rodr. AM GU
Odontoglossum naevium Lindl. BO GU
Oeceoclades maculata (Lindl.) Lindl AM BO GU SU FG
Oliveriana egregia Rchb. f. GU
Oncidium baueri Lindl. BO DA GU SU FG
Oncidium boothianum Rchb. f. AM
Oncidium citrinum Lindl. BO
Oncidium dactyliferum Garay & Dunst. AM BO
Oncidium heteranthum Poepp. & Endl. SU
Oncidium luridum Lindl. GU FG
Oncidium nigratum Lindl. BO GU
Oncidium orthostates Ridl. ex Thum AM BO GU
Oncidium pirarense Rchb. f. GU
Oncidium sphacelatum Lindl. BO
Oncidium tenellum F. Gerard FG
Oncidium warmingii Rchb. f.
Ondidium pardalis Rchb. f.
Ophidion pleurothallopsis (Kiaenzl.) Luer BO GU
Orleanesia amazonica Barb. Rodr. AM BO GU SU FG
Orleanesia maculata Garay
Orleanesia yauaperyensis (Schltr.) Barb. Rodr.
Ornithocephalus bicornis Lindl. GU SU FG
Ornithocephalus bonplandii Rchb. f. SU
Ornithocephalus brachyceras G.A. Romero & Camevali BO
Ornithocephalus ciliatus Lindl. GU SU FG
AM BO
BO
AM BO
AM
AM
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
FG?
GU SU FG
GU su FG
SU FG
BO
BO
BO
BO
BO
BO
BO
Checklist of the Plants of the Guiana Shield 129
Ornithocephalus dasyrhizus Kraenzl.
Ornithocephalus falcatus H. Focke BO
Ornithocephalus gJadiatus Hook.
Ornithocephalus kruegeri Rchb. f.
Otoglossum arminii (Rchb. f.) Garay & Dunst.
Otoglossum globidiferum (Kunth) N.H. Williams & M.W. Chase
Otoglossum scansor (Rchb. f.) Camevali & 1. Ramirez
Otostylis brachystalix (Rchb. f.) Schltr.
Otostylis lepida (Linden & Rchb. f.) Schltr.
Palmorchis guianensis (Schltr.) C. Schweinf. & Correll
Palmorchis kuhlmannii (Schltr.) L.O. Williams
Palmorchis lobulata (Mansf.) C. Schweinf. & Correll
Palmorchis pabstii Veyret
Palmorchis prospectorum Veyret
Palmorchis puber (Cogn.) Garay AM
Palmorchis pubescens Barb. Rodr.
Palmorchis trilobulata L.O. Williams
Paphiniacristata{\.md\.)l.\ndX. AM DA
Paphinia dunstervillei Dodson & G.A. Romero
Paphinia lindeniana Rchb. f.
Pelexia callifera (C. Schweinf.) Garay
Pelexia delicatida Szlach. & Veyret
Pelexia goninensis (Pulle) Schltr.
Pelexia laxa (Poepp. & Endl.) Lindl.
Peristeria aspersa Rolfe
Peristeria cerina Lindl.
Peristeria pendula Hook.
Peristeria rossiana Rchb. f
Peristeria selligera Rchb. f.
Phragmipedium caudatum (Lindl.) Rolfe
Phragmipedium klotzschianum (Rchb. f.) Rolfe
Phragmipedium lindleyanum (R.H. Schomb. ex Lindl.) Rolfe
Pinelianthe alticola (Garay & Dunst.) S. Rauschert
Platystelejohnstonii (Ames) Garay
Platystele ovalifolia (H. Focke) Garay & Dunst. BO GU SU FG
Platystele oxyglossa (Schltr.) Garay
Platystele stenostachya (Rchb. f.) Garay
Platythelys maculata (Hook.) Garay
Plectrophora cultrifolia (Barb. Rodr.) Cogn.
Plectrophora iridifolia H. Focke
Pleurothallis aondae Camevali & G.A. Romero
Pleurothallis arachnopsis H.G. Jones
Pleurothallis archidiaconi Ames
Pleurothallis arenicola (Camevali & I. Ramirez) Camevali & I. Ramirez
Pleurothallis aristata Hook.
Pleurothallis aspasicensis Rchb. f.
Pleurothallis barboselloides Schltr.
Pleurothallis barbulata L\nd\. AM BO DA
Pleurothallis barthelemyi Luer
Pleurothallis biglandulosa Schltr.
Pleurothallis brevipes H. Focke AM
Pleurothallis cabellensis Rchb. f
Pleurothallis callifera C. Schweinf. BO
Pleurothallis ciliolata Schltr.
Pleurothallis coriacardia Rchb. f AM BO
Pleurothallis corniculata Lindl. BO DA
Pleurothallis curvifructa H.G. Jones [possibly =P. archidiaconi]
Pleurothallis deborana Camevali & L Ramirez BO
Pleurothallis determannii Luer
Pleurothallis discoidea Lindl.
Pleurothallis discophylla Luer & Camevali AM
Pleurothallis elvirana Camevali & I. Ramirez AM
Pleurothallis ephemera Lindl.
Pleurothallis erebatensis Camevali & G.A. Romero BO
BO
BO
BO
BO
BO
GU SU FG
GU?
GU SU
SU
FG
FG
SU FG
GU
SU
FG
GU SU FG
GU?
SU FG
FG
GU SU FG
GU SU?
GU SU FG
GU SU FG
GU? SU
GU
FG
GU
GU SU FG
BO
BO
BO
BO
BO
BO
SU
SU FG
FG
GU SU FG
GU
GU SU FG
GU
GU SU FG
GU
SU
GU SU FG
FG
GU FG
GU SU FG
FG
GU SU FG
GU SU
GU
GU SU
SU FG
GU SU FG
130 Checklist of the Plants of the Guiana Shield
Pleurothallis erinacea Rchb. f.
PleiiwthalUs exigiia Cogn.
Pleurothallisfloribimda Poepp. & Endl.
Pleurothallisfockei Lindl.
Pleurothallis funerea (Barb. Rodr.) Cogn.
Pleurothallis galeafa Lindl.
Pleurothallis garciae Luer
Pleurothallis geographica Luer
Pleurothallis glandulosa Ames
Pleurothallis granitica Luer & G.A. Romero
Pleurothallis grobyi Bateman ex Lindl.
Pleurothallis hexandra Garay & Dunst.
Pleurothallis hilariana Camevali & G.A. Romero
Pleurothallis hitchcockii Ames
Pleurothallis holstii Camevali & I. Ramirez
Pleurothallis humilis C. Schweinf.
Pleurothallis hymenantha Lindl.
Pleurothallis imraei Lindl.
Pleurothallis ionantha Rchb. f.
Pleurothallis kerrii Braga
Pleurothallis lanceana Lodd.
Pleurothallis lancipetala (H. Karst.) Schltr.
Pleurothallis lappiformis A.H. Heller & L.O. Williams
Pleurothallis longisepala Barb. Rodr.
Pleurothallis loranthophylla Rchb. f.
Pleurothallis maguirei Luer
Pleurothallis mentosa Cogn.
Pleurothallis minima C. Schweinf.
Pleurothallis miqueliana (H. Focke) Lindl.
Pleurothallis monophylla (Hook.) Fawc. & Rendle
Pleurothallis morilloi Camevali & I. Ramirez
Pleurothallis moritzii Rchb. f. [Mt. Roraima]
Pleurothallis nanifolia Foldats
Pleurothallis obovata (Lindl.) Lindl.
Pleurothallis omissa Luer
Pleurothallis omoglossa Luer
Pleurothallis parvifolia Lindl.
Pleurothallis pemonum Camevali & L Ramirez
Pleurothallis phoenicoptera Camevali & G.A. Romero
Pleurothallis picta Lindl.
Pleurothallis polygonoides Griseb.
Pleurothallis prolifera Herb, ex Lindl.
Pleurothallis pruinosa Lindl.
Pleurothallis pubescens Lindl.
Pleurothallis revoluta (Ruiz & Pav.) Garay
Pleurothallis rhombipetala Rolfe
Pleurothallis ruscifolia (Jacq.) R. Br.
Pleurothallis samacensis Ames
Pleurothallis sandaliorum G.A. Romero & Camevali
Pleurothallis sarcosepala Camevali & L Ramirez
Pleurothallis sclerophylla Lindl.
Pleurothallis secunda Poepp. & Endl.
Pleurothallis semperflorens Lindl.
Pleurothallis seriata Lindl.
Pleurothallis sp. A
Pleurothallis spiculifera Lindl.
Pleurothallis steinbuchiae Camevali & G.A. Romero
Pleurothallis stenocardium Schltr.
Pleurothallis suspensa Luer
Pleurothallis tepuiensis Camevali & I. Ramirez
Pleurothallis testifolia (Sw.) Lindl.
Pleurothallis tridentata Klotzsch
Pleurothallis trinitensis (Griseb.) Camevali & G.A. Romero
Pleurothallis uniflora Lindl.
BO GU SU
GU
AM BO
AM BO GU su
BO DA GU FG
AM BO
BO GU SU FG
BO
BO DA GU su FG
AM BO
AM BO GU su FG
BO GU su
BO
AM BO
BO
GU
AM BO
GU
AM BO
BO
GU su
AM
AM BO GU su FG
AM
AM
BO
BO
BO
AM BO
AM BO
AM BO
BO
AM BO
AM BO
BO
AM BO
AM BO
AM BO
BO
AM BO
AM BO
BO
AM BO
BO
AM
BO
AM BO
BO
BO
AM
BO
BO
BO
AM? BO
BO
AM BO
DA
DA
GU
GU
GU
GU
GU
GU
GU
GU
GU
SU
SU
FG?
FG
GU SU FG
GU SU FG
GU
GU SU FG
su FG
su
GU
GU su FG
GU SU? FG
FG
GU su FG
GU su FG
FG
GU
GU SU FG
GU
su?
GU
GU SU FG
GU su FG
BO
AM BO
BO
AM BO
AM
AM BO
BO
AM BO
AM BO
BO
AM BO
BO
AM BO
BO
AM BO
AM
AM BO
AM
AM BO
AM BO
BO
BO
AM BO
AM BO
AM BO
Checklist of the Plants of the Guiana Shield 131
PlewothaUis velaticaiilis Rchb. f. AM
Pleiirothallis wilsonii Lindl.
PlewothaUis yauaperyensis Barb. Rodr.
Pleiirothallis zephyrina Rchb. f.
Polycycnis siirinamensis C. Schweinf.
Polyotidium huebneri (Mansf.) Garay
Polystachya amazonica Schltr. DA
Polystachya cavanayensis Garay & Dunst.
Polystachya concreta (Jacq.) Garay & H.R. Sweet DA
Polystachya d'agremondiana Kraenzl. [dubious species]
Polystachya foliosa (Lindl.) Rchb. f. DA
Ponthieva diptera Linden & Rchb. f.
Ponthieva ovatilabia C. Schweinf.
Prescottia auyantepuiensis Camevali & GA. Romero
Prescottia carnosa C. Schweinf.
Prescottia oligantha (Sw.) Lindl.
Prescottia stachyodes (Sw.) Lindl.
Prescottia tepuyensis Camevali & C.A. Vargas
Prosthechea aemula (Lindl.) W.E. Higgins DA
Prosthechea calamaria (Lindl.) W.E. Higgins
Prosthechea crassilabia (Poepp. & Endl.) Camevali & L Ramirez
Prosthechea fragrans (Sw.) W.E Higgins [in Guianas records may
=Prosthechea aemula]
Prosthecheajauana (Camevali & I. Ramirez) W.E. Higgins
Prosthechea pygmaea (Hook.) W.E. Higgins
Prosthechea tigrina (Linden ex Lindl.) W.E. Higgins
Prosthechea vespa (Veil.) W.E. Higgins DA
Psilochilus dusenianus Kraenzl. ex Garay & Dunst.
Psilochilus macrophyllus (Lindl.) Ames
Psilochilus physurifolius (Rchb. f.) Lojtnant
Psychopsis kramerianum (Rchb. f.) H.G. Jones
Psychopsis papilio (Lindl.) H.G. Jones BO
Psychopsis versteegianum (Pulle) Liickel & Braem
Pterichis galeata Lindl.
Quekettia microscopica Lindl.
Quekettia papulosa Garay
Quekettia vermeuleniana Determann
Reichenbachanthus reflexus (Lindl.) Brade AM BO DA
Restrepiopsis tubulosa (Lindl.) Luer
Rodriguezia batemanii Poepp. & Endl.
Rodriguezia Candida (Bateman ex Lindl.) Christenson
Rodriguezia cinnabarina H.G. Jones
Rodrigueziaflavida Garay & Dunst.
Rodriguezia lanceolata Ruiz & Pav. AM BO DA
Rodriguezia leeana Rchb. f.
Rodriguezia luteola N.E. Br.
Rudolfiella aurantiaca (Lindl.) Hoehne AM BO DA
Rudolfiella bicornaria (Rchb. f ) Hoehne
Sacoila hassleri (Cogn.) Garay
Sacoila lanceolata (Aubl.) Garay BO
Sarcoglottis acaulis (Sm.) Schltr. BO
Sarcoglottis amazonica Pabst
Sarcoglottis aphylla (Ridl. ex Thum) Schltr. BO
Sarcoglottis degranvillei Szlach. & Veyret
Sarcoglottis grandiflora (Lindl.) Klotzsch
Sarcoglottis metallica (Rolfe) Schltr. AM BO
Sarcoglottis rupestris (Lindl.) Barb. Rodr.
Sarcoglottis sagittata (Rchb. f. & Warm.) SchUr.
Sarcoglottis simplex (Griseb.) Schltr.
Sarcoglottis stergiosii Camevali & 1. Ramirez
Scaphosepalum breve (Rchb. f ) Rolfe
Scaphyglottis bifida (Rchb. f ) C. Schweinf
Scaphyglottis boliviensis (Rolfe) B.R. Adams
Scaphyglottis dunstervillei (Garay) Foldats
BO
BO
BO
AM BO
AM
BO
AM BO
AM BO
BO
SU
GU su FG
GU SU FG
GU su
su
FG
GU su FG
GU
GU
GU
GU
GU su FG
GU su FG
GU su FG
GU
GU su FG
GU su FG
GU
GU su FG
GU
GU
GU
su
su FG
su
GU su
su
FG
GU su
GU su FG
GU su FG
GU
GU su
GU su
su
FG
GU
GU su
su
su
FG
GU su FG
GU su FG
su FG
GU
FG
GU su FG?
GU
su
FG
GU
GU
GU
su FG
su
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
GU su FG
su FG
GU su
su
FG
GU
GU su FG
GU su FG
FG
GU
GU? su? FG'
GU su?
GU su FG
GU
GU
GU
GU su FG
GU su FG
GU
GU su? FG
su FG
132 Checklist of the Plants of the Guiana Shield
Scaphyglottisfelskyi{Kchh.i.)S>cmx. BO GU SU
Scaphyglottisfusiformis (Griseb.) R.E. Schult. AM BO DA GU SU FG
^ca;7/7>;g/om"5 gramw//b/za (Ruiz & Pav.) Poepp. & Endl. AM BO DA GU SU FG
Scaphyglottis grandiflora Ames & C. Schweinf. AM BO GU FG
Scaphyglottis leucantha Rchb. f. BO
Scaphyglottis lindeniana (A. Rich. & Galeotti) L.O. Williams GU SU
Scaphyglottis michelangeliorum Camevali & Steyerm. AM
Scaphyglottis modesta (Rchb. f.) Schltr. AM BO
Scaphyglottis ochroleuca Schltr.
Scaphyglottis prolifera Cogn. BO
Scaphyglottis propinqua C. Schweinf.
Scaphyglottis sessilis (Rchb. f.) Foldats AM BO
Scaphyglottis sickii Pabst AM BO DA
Scaphyglottis stellata Lodd. ex Lindl. AM BO
Scelochilus ecalcaratus Determann
Scelochilus ottonis Klotzsch AM
Scelochilus paraguaensis Garay &. Dunst.' BO
Schlimia alpina Rchb. f. & Warsz.
Schomburgkia crispa Lindl. [records in Guianas may =S. marginata]
Schomburgkia gloriosa Rchb. f.
Schomburgkia heidii Camevali AM BO
Schomburgkia marginata Lindl. BO
Schomburgkia rosea Linden ex Lindl.
Schomburgkia undulata Lindl. BO
Scuticaria hadwenii hort. ex Planch.
Scuticaria hadwenii hort. var. dodgsoni (B.S. Wms.) Stein
Scuticaria steelei (Hook.) Lindl.
Selenipedium palmifolium (Lindl.) Rchb. f.
Selenipedium steyermarkii Foldats
Sievekingiajenmanii Rchb. f.
Sigmatostalix amazonica Schltr.
Sigmatostalix huebneri Mansf.
Sobralia Candida (Poepp. & Endl.) Rchb. f.
Sobralia crocea (Poepp. & Endl.) Rchb. f. SU?
Sobralia dichotoma Ruiz & Pav.
Sobraliafimbriata Poepp. & Endl.
Sobraliafragrans Lindl.
Sobralia granitica G.A. Romero & Camevali
Sobralia infundibuligera Garay & Dunst. AM DA
Sobralia liliastrum Lindl.
Sobralia macrophylla Rchb. f.
Sobralia oliva-estevae Camevali & L Ramirez
Sobralia sessilis Lindl. AM DA
Sobralia speciosa C. Schweinf.
Sobralia stenophylla Lindl.
Sobralia suaveolens Rchb. f.
Sobralia valida Rolfe AM DA
Sobralia violacea Linden ex Lindl.
Solenidium lunatum (Lindl.) Kraenzl.
Spiranthes tenuis Lindl.
Spiranthes torta (Thunb.) Garay & H.R. Sweet
Stanhopea Candida Barb. Rodr.
Stanhopea grandiflora (Lodd.) Lindl. AM DA
Stanhopea wardii Lodd. ex Lindl.
Stelis alata Lindl.
Stelis aprica Lindl.
Stelis argentata Lindl. DA
Stelis aviceps Lindl.
Stelis bangii Rolfe
Stelis cucullata Ames
Stelis cuprea Lindl. [dubious record] FG
Stelis effusa Schltr.
Stelis fendleri Lindl.
Stelisfraterna Lindl AM GU
SU FG
GU su FG
GU
GU su FG
GU su FG
GU
GU su FG
GU su FG
GU su FG
GU
su?
GU
GU su FG
FG?
GU?
GU su FG
GU
GU su
GU su FG
GU
GU
Checklist of the Plants of the Guiana Shield 133
Stelis garayi (Dunst.) Carnevali & I. Ramirez AM BO
Stelis grandiflora Lindl. GU
Stelis guianensis Rolfe BO GU FG
Stelis intermedia Poepp. & Endl. BO GU SU
Stelis latisepala C. Schweinf. BO
Stelis maxima Lindl. AM
Stelis minimiflora Schltr. BO
Stelis muscifera Lindl. GU
Stelis obovata C. Schweinf.
Stelis ophioglossoides (Jacq.) Jacq. BO GU SU? FG
Stelis papaquerensis Rchb. f. SU? FG
Stelis perparva C. Schweinf. GU
Stelis pusilla Kunth
Stelis pygmaea Cogn. BO FG
Stelis santiagoensis Mansf AM BO DA FG
Stelis schomburgkii Fawc. & Rendle AM BO GU
Stelis tolimensis Schltr.
Stelis tridentata Lindl.
Stelis tristyla Lindl. GU SU
Stelis zonata Rchb. f. [=Stelis muscifera?] BO GU
Stellilabium alticolum Dodson & R. Escobar
Stenia pallida Lindl. BO DA GU
Stictophyllorchis pygmaea (Cogn.) Carnevali & Dodson
Teuscheria dodsonii Dressier SU?
Teuscheria wageneri (Rchb. f.) Garay
Trichocentrum carthagenense (Jacq.) M.W. Chase & N.H. Williams
Trichocentrum cebolleta (Jacq.) M.W. Chase & N.H. Williams AM BO DA
Trichocentrumfuscum Lindl.
Trichocentrum lanceanum (Lindl.) M.W. Chase & N.H. Williams AM BO DA
Trichocentrum nanum (Lindl.) M.W. Chase & N.H. Williams
Trichocentrum purpureum Lindl. ex Rchb. f.
Trichocentrum recurvum Lindl.
Trichopilia aenigma Garay
Trichopilia oicophylax Rchb. f.
Trichopilia wageneri Rchb. f.
Trichosalpinx cedralensis (Ames) Luer
Trichosalpinx ciliaris (Lindl.) Luer
Trichosalpinx dura (Lindl.) Luer
Trichosalpinx egleri (Pabst) Luer BO DA
Trichosalpinx intricata (Lindl.) Luer
Trichosalpinx memor (Rchb. f ) Luer
Trichosalpinx orbicularis (Lindl.) Luer
Trichosalpinx oxychilos Carnevali & G.A. Romero
Trichosalpinx patula Luer
Trichosalpinx pumila (Luer) Luer
Trichosalpinx pusilla (Kunth) Luer
Trichosalpinx roraimensis (Rolfe) Luer AM BO GU
Trichosalpinx steyermarkii Luer
Trigonidium acuminatum Bateman ex Lindl. AM BO DA
Trigonidium egertonianum Bateman ex Lindl.
Trigonidium macranthum Barb. Rodr.
Trigonidium obtusum Lindl.
Triphora amazonica Schltr.
Triphora surinamensis (Lindl. ex Benth.) Britton & P. Wilson
Trisetella triglochin (Rchb. f ) Luer
Trizeuxis falcata Lindl. BO DA
Uleiorchis liesneri Carnevali & 1. Ramirez
Uleiorchis ulaei (Cogn.) Handro
Vanilla acuta Rolfe
Vanilla appendiculata Rolfe
Vanilla barrereana Veyret & Szlach.
Vanilla bicolor Lindl.
Vanilla chamissonis Klotzsch
Vanilla cristato-callosa Hoehne
AM BO
BO
AM BO
AM
BO
AM BO
BO
AM
BO
AM BO
BO
BO
AM BO
AM BO
BO
AM BO
AM BO
BO
BO
BO
AM
BO
AM BO
AM BO
AM
AM
AM BO
GU SU FG
SU FG
GU SU FG
GU FG
GU
GU SU
GU SU
GU SU
SU
FG
GU SU FG
GU SU
GU SU FG
GU SU FG
GU
GU
GU SU FG
GU SU? FG
GU SU FG
SU FG
GU
GU FG
GU SU FG
GU
FG
GU SU FG
FG
GU
134 Checklist of the Plants of the Guiana Shield
Vanillafimbrlata Rolfe
Vanilla gardneri Rolfe
Vanilla grandiflora Lindl.
Vanilla grandifolia Lindl.
Vanilla guianensis Splitg. [name of uncertain application; =V. mexicanal
V. grandiflora?]
Vanilla hostmanni Rolfe
Vanilla latisegmenta Ames & C. Schweinf
Vanilla leprieurii Porteres
Vanilla marowynensis Pulle
Vanilla mexicana Mill.
Vanilla odorata C. Presl
Vanilla ovata Rolfe
Vanilla palmarum (Salzm. ex Lindl.) Lindl.
Vanilla penicillata Garay & Dunst.
Vanilla planifolia Andrews
Vanilla porteresiana Veyret & Szlach.
Vanilla surinamensis Rchb. f.
Vanilla wrightii Rchb. f
Vargasiella venezuelana C. Schweinf.
Wullschlaegelia calcarata Benth.
Xerorchis amazonica Schltr.
Xerorchis trichorhiza (Kraenzl.) Garay
Xylobium chapadense (Barb. Rodr.) Cogn.
Xylobium colleyi (Bateman ex Lindl.) Rolfe
Xylobiumfoveatum (Lindl.) G. Nicholson
Xylobium pallidiflorum (Hook.) G. Nicholson
Xylobium palmifolium (Sw.) Benth. ex Fawc.
Xylobium variegatum (Ruiz & Pav.) Garay & Dunst.
Zygosepalum angustilabium (C. Schweinf.) Garay
Zygosepalum kegelii (Rchb. f ) Rchb. f
Zygosepalum labiosum (Rich.) Garay
Zygosepalum lindeniae (Rolfe) Garay & Dunst.
Zygosepalum tatei (Ames & C. Schweinf) Garay & Dunst.
Zygostates sp. nov.
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
AM BO
BO
BO
BO
BO
AM BO
AM
AM BO
DA
DA
GU
GU SU
su
SU
FG:
GU
su
FG
GU su FG
FG
GU? FG
GU su FG
GU
GU su
su
FG
GU su FG
GU su
su
FG
GU FG
GU
GU su FG
GU su
su
FG
GU
su FG
GU
su
GU su FG
GU FG
GU
SU
Synonyms and Misidentifications
Aciantherafockei (Lindl.) Pridgeon & M.W. Chase =Pleurothallis fockei
Acianthera garciae (Luer) Pridgeon & M.W. Chase =Pleurothallis garciae
Acianthera lanceana Pridgeon & M.W. Chase '=Pleurothallis lanceana
Acianthera miqueliana (H. Focke) Pridgeon & M.W. Chase =Pleurothallis miqueliana
Acianthera omissa (Luer) Pridgeon & M.W. Chase =Pleurothallis omissa
Acianthera prolifera (Herb, ex Lindl.) Pridgeon & M.W. Chase =Pleurothallis prolifera
Acianthera wilsonii (Lindl.) Pridgeon & M.W. Chase =Pleurothallis wilsonii
Acianthera yauaperyensis (Barb. Rodr.) Pridgeon & M.W. Chase =Pleurothallis yauaperyensis
Acoidiumfuscum Lindl. [nomen nudum] =Trichocentrum fuscum
Adeneleutherophora graminifolia Barb. Rodr. =Elleanthus graminifolius
Adipe longicornis (Lindl.) M. Wolfe =Bifrenaria longicornis
Aeranthus pachyrrhizus Rchb. f. =Campylocentrum pachyrrhizum
Aganisia alba Ridl. ex Thum =Otostylis lepida
Aganisia brachypoda Schltr. =Aganisia pulchella
Aganisia cyanea (Lindl.) Rchb. f. =Acacallis cyanea
Aganisiafimbriata Rchb. f. =Acacallis fimbriata
Aganisia lepida Linden & Rchb. f. =Otostylis lepida
Aganisia venusta (Ridl. ex Oliv.) Rolfe =Otostylis lepida
Anacheilium calamarium (Lindl.) Pabst, Moutinho & Pinto =Prosthechea calamaria
Anathallis barbulata (Lindl.) Pridgeon & M.W. Chase =Pleurothallis barbulata
Anathallis brevipes (H. Focke) Pridgeon & M.W. Chase =Pleurothallis brevipes
Anathallis maguirei (Luer) Pridgeon & M.W. Chase =Pleurothallis maguirei
Anathallis sclerophylla (Lindl.) Pridgeon & M.W. Chase =Pleurothallis sclerophylla
Angraecumfasciola Lindl. =Campylocentrum fasciola
Angraecum lansbergii Rchb. f. =Campylocentrum lansbergii
Angraecum maculatum Lindl. =Oeceoclades maculata
Angraecum micranthum Lindl. =Campylocentrum micranthum
Checklist of the Plants of the Guiana Shield 135
Angraecum poeppigii Rchb. f. =Campylocentrum poeppigii
Anthereon mentosus (Cogn.) Pridgeon & M.W. Chase =Pleurothallis mentosa
Apatostelis alata (Lindl.) Garay =Stelis alata
Apatostelis garayi Dunst. =Stelis garayi
Apatostelis minimiflora (Schltr.) Garay =Stelis minimiflora
Aspegrenia scirpoidea Poepp. & Endl. =Octomeria scirpoidea
Aulizia ciliaris (L.) Salisb. =Epidendrum ciliare
Barbosella kegelii (Rchb. f.) Schltr. =Pleurothallis hexandra
Baskervilla venezuelana Garay & Dunst. [not in flora area]
Beadlea bifida (Ridl.) Garay & Dunst. =Helonoma chiropterae
Beadlea bifida (Ridl. ex Oliv.) Garay & Dunst. =Helonoma bifida
Beadlea cranichoides (Griseb.) Small =Cyclopogon cranichoides
Beadlea elata (Sw.) Small ex Britton =Cyclopogon elatus
Beloglottis americana (C. Schweinf. & Garay) Szlach. =Helonoma americana
Beloglottis bifida (Ridl. ex Oliv.) Szlach. =Helonoma bifida
Beloglottis chiropterae Szlach. =Helonoma chiropterae
Bifrenaria aurantiaca Lindl. '=Rudolfiella aurantiaca
Bifrenaria bicornaria Rchb. f. =Rudolfiella bicornaria
Bifrenaria maguirei C. Schweinf. =Guanchezia maguirei
Bifrenaria minuta Garay ^Hylaeorchis petiolaris
Bifrenaria rudolfii (Hoehne) Camevali & G.A. Romero =Hylaeorchis petiolaris
Bifrenaria sabiilosa Barb. Rodr. =Bifrenaria longicornis
Bifrenaria wageneri Rchb. f. =Teuscheria wageneri
Bletia verecunda R. Br. =Bletia purpurea
Bolbophyllaria oerstedii Rchb. f. =Bulbophyllum oerstedii
Bollea cardonae Schnee =Bollea hemixantha
Bollea guyanensis Klotzsch [nomen nudum]
Bonatea macilenta (Lindl.) Rchb. f. =Habenaria macilenta
Bonatea paucifiora Lindl. =Habenaria trifida
Bonatea pratensis Lindl. =Habenaria pratensis
Brassavola angustata Lindl. =Brassavola martiana
Brassavola surinamensis H. Focke =Brassavola martiana
Brassia attenuata H.G Jones =Brassia chloroleuca
Brassia cochleata Knowles & Westc. =Brassia lawrenceana
Brassia verrucosa auct., non Lindl. =Brassia spp.
Bulbophyllum correae Pabst =Bulbophyllum setigerum
Bulbophyllum pachyrrhachis auct. non (A. Rich.) Griseb. 1864, sensu Dunst. & Garay =Bulbophyllum oerstedii
Burlingtonia Candida Bateman ex Lindl. =Rodriguezia Candida
Camaridium album (Hook.) Hoehne =Maxillaria alba
Camaridium amazonicum Schltr. =Maxillaria camaridii
Camaridium equitans Schltr. =Maxillaria equitans
Camaridium iguapensis (Hoehne & Schltr.) Hoehne =Maxillaria imbricata
Camaridium imbricatum (Barb. Rodr.) Hoehne =Maxillaria imbricata
Camaridium ochroleucum Lindl. =Maxillaria camaridii
Camaridium pendulum Barb. Rodr. =Maxillaria rigida
Camaridium squamatum (Barb. Rodr.) Hoehne =Maxillaria squamata
Camaridium uncatum (Lindl.) Hoehne =Maxillaria uncata
Camaridium vandiforme Schltr. =Maxillaria equitans
Campylocentrum burchellii Cogn. [not in flora area]
Campylocentrum micranthum auct. non (Lindl.) Rolfe 1903: sensu Foldats in Lasser ^^Campylocentrum panamense p.p., C.
lansbergii p.p.
Campylocentrum steyermarkii Foldats ^Campylocentrum hondurense
Campylocentrum uroplectron Pabst =Campylocentrum huebneri
Catasetum apertum Rolfe =Catasetum ^tapiriceps
Catasetum bicallosum auct. non Cogn. 1894: sensu Dunst. & Garay =Catasetum parguazense
Catasetum bungerothii N.E. Br. =Catasetum pileatum
Catasetum chloranthum Cogn. =Catasetum planiceps
Catasetum comosum Cogn. =Catasetum barbatum
Catasetum discolor var. roseo-album (Hook.) Mansf =Catasetum x roseo-album
Catasetumfuliginosum Lindl. ? =Catasetum callosum var carunculatum
Catasetum gongoroides Kraenzl. =Catasetum bicolor
Catasetum recurvatum Link ^Catasetum planiceps
Catasetum scurra Rchb. f =Clowesia warczewiczii
Catasetum splendens Cogn. =Catasetum x tapiriceps
136 Checklist of the Plants of the Guiana Shield
Catasetum stenoglossum Pabst =Catasetum bicallosum
Catasetum warczewiczii Lindl. & Paxton =Clowesia warczewiczii
Cattleya liliastrum (Lindl.) Beer =Sobralia liliastnim
Cattleya piimila R.H. Schomb. =Cattleya lawrenceana
Cattleya schomburgkii Lodd. ex Lindl. =Cattleya violacea
Cattleya superba R.H. Schomb. ex Lindl. =Cattleya violacea
Cattleya violacea (Linden ex Lindl.) Beer =Sobralia violacea
Caularthron indivisum (Bradf. ex Griseb.) Garay & Dunst =Caularthron bilamellatum f. indivisa
Centrogenium calcaratum (Sw.) Schltr. =Eltroplectris calcarata
Centrogenium setaceum (Lindl.) Schltr. =Eltroplectris calcarata
Centroglossa peruviana Cogn. =C}yptocentriim peruvianum
Ceratochilus grandiflorus Lodd. =Stanhopea grandiflora
Chaenanthe barkeri Lindl. =Diadeniiim barkeri
Chamelophyton kegelii (Rchb. f.) Garay =Pleurothallis hexandra
Chaubardia tigrina Garay & Dunst. =Chaubardiella tigrina
Cheiradenia imthurnii Cogn. =Cheiradenia^cuspidata
Cladobium violaceum Lindl. =Scaphyglottis graminifolia
Cleistes moritzii (Rchb. f.) Garay & Dunst. =Cleistes lepida
Cochleanthes amazonica (Rchb. f.) Schult. & Garay [not in flora area]
Cohniella cebolleta (Jacq.) Christenson =Trichocentrum cebolleta
Coryanthes speciosa var. punctata (Beer) G. Gerlach =Coryanthes punctata
Cranichis alfredi Schltr. =Cranichis diphylla
Cranichis guatemalensis Schltr. =Cranichis diphylla
Cranichis monophylla Lindl. =Cranichis diphylla
Cranichis nigrescens Schltr. =Cranichis diphylla
Cranichis oligantha Sw. =Prescottia oligantha
Cranichis ovatilabia Schltr. ^Cranichis diphylla
Cranichis stachyodes Sw. =Prescottia stachyodes
Cranichis stictophylla Schltr. =Cranichis diphylla
Cryptarrhena acrensis Schltr. =Cryptarrhena guatemalensis
Cryptarrhena ghillanyi Pabst =Cryptarrhena guatemalensis
Cryptarrhena lunata auct., non R. Br. =Cryptarrhena kegelii
Cryptocentrum gracillimum auct. non Ames & C. Schweinf. 1925: sensu Dunst. & Garay ^Cryptocentrum dunstervilleorum
Cryptocentrum hoppii Schltr. =Cryptocentrum peruvianum subsp. peruvianum
Cryptophoranthus pleurothallopsis Kraenzl. =Ophidion pleurothallopsis
Cyathoglottis Candida Poepp. & Endl. =Sobralia Candida
Cyclopogon bifidus (Ridl. ex Oliv.) Schltr. =Helonoma bifida
Cyclopogon stictophyllus Schltr. =Cyclopogon olivaceus
Cycnoches cucullata Lindl. =Cycnoches loddigesii
Cycnoches pescatorei Lindl. =Lueddemannia pescatorei
Cycnoches ventricosum var. chlorochilon (Klotzsch) P. Allen =Cycnoches chlorochilon
Cycnoches versicolor Rchb. f. =Cycnoches haagii
Cymbidium andersonii Lamb, ex Andrews ^Cyrtopodium andersonii
Cymbidium cordigerum Kunth =Encyclia cordigera
Cymbidium diurnum (Jacq.) Sw. =Encyclia diurna
Cymbidium ochroleucum (Lindl.) Lindl. =Maxillaria camaridii
Cymbidium violaceum Kunth =Cattleya violacea
Cypripedium klotzschianum Rchb. f. =Phragmipedium klotzschianum
Cypripedium lindleyanum R.H. Schomb. ex Lindl. =Phragmipedium lindleyanum
Cypripedium palmifolium Lindl. =Selenipedium palmifolium
Cypripedium schomburgkianum Klotzsch ex M.R. Schomb. =Phragmipedium klotzschianum
Cyrtopera longifolia Rchb. f. =Eulophia alta
Cyrtopodium broadwayi Ames =Cyrtopodium parviflorum
Cyrtopodium eburnea Barb. Rodr. =Koellensteinia tricolor
Cyrtopodium flavescens Cogn. =Cyrtopodium andersonii
Cyrtopodium glutiniferum Raddi [dubious in Guianas]
Cyrtopodium punctatum (L.) Lindl. [dubious in Guianas]
Dendrobium album Hook. =Maxillaria alba
Dendrobium cebolleta Jacq. =Trichocentrum cebolleta
Diacrium amazonicum Schltr. =Caularthron bicornutum
Diacrium bicornutum (Hook.) Benth. =Caularthron bicornutum
Diacrium bilamellatum (Rchb. f.) Hemsl. ^Caularthron bilamellatum
Diacrium venezuelanum Schltr. =Caularthron bilamellatum
Dichaea ancoraelabia C. Schweinf. [not in flora area. Peruvian taxon]
Checklist of the Plants of the Guiana Shield 137
Dichaea brachypoda auct. non Rchb. f 1866: sensu Foldats in Lasser =Dichaea camaridioides
Dichaea brachypoda Rchb. f. [Costa Rican taxon, name widely misapplied]
Dichaea echinocarpa (Sw.) Lindl. [nom. illeg.] =Dichaea pendula
Dichaea gramiuoides (Sw.) Lindl. [name (mis?)applied to Dichaea rendlei in Guianas]
Dichaea histrio auct. non Rchb. f. 1859: sensu Dunst. & Garay =Dichaea tachirensis
Dichaea kegelii auct. non Rchb. f. 1877: sensu Dunst. and Garay =Dichaea splitgerberi
Dichaea longa Schltr. =Dichaea latifolia van longa
Dichaea morrisii auct. non Fawc. & Rendle 1910: sensu Dunst. and Garay =Dichaea kegelii, p.p., Dichaea robiista, p.p.
Dichaea muricata auct. non (Sw.) Lindl. 1833: sensu Dunst. and Garay =Dichaea latifolia var. longa
Dichaea muricata auct. non (Sw.) Lindl. 1833: sensu J. Folsom =Dichaea robiista
Dichaea muricata (Sw.) Lindl. [rejected name]
Dichaea pendula var. swartzii C. Schweinf. [nom. illeg.] =Dichaea pendula
Dichaea rendlei var. rendlei auct. non Gleason 1927: sensu Foldats in Lasser =Dichaea trinitensis
Dichaea rendlei var. trinitensis auct. non (Gleason) Foldats 1959: sensu Dunst. and G =Dichaea hookeri
Dichaea rendlei var. trinitensis (Gleason) Foldats =Dichaea trinitensis
Dichaea swartzii (C. Schweinf.) Garay & H.R. Sweet =Dichaea pendula
Diciypta bicolor Paxton ex Planch. =Maxillaria discolor
Dicrypta discolor Lodd. ex Lindl. =Maxillaria discolor
Dicrypta longifolia Barb. Rodr. =Heterotaxis superflua
Dicrypta villosa Barb. Rodr. =Heterotaxis villosa
Didactyle exaltata (Lindl.) Lindl. =Bulbophyllum exaltatum
Didactyle meridensis (Rchb. f.) Lindl. =Bulbophyllum meridense
Dikylikostigma preusii Kraenzl. =Discyphus scopulariae
Dimerandra isthmii Schltr. =Dimerandra elegans
Dimerandra stenopetala auct., non (Hook.) Schltr. [name widely misapplied] =Dimerandra spp.
Diothonea gratissima Rchb. f. =Epidendrum gratissimum
Diothonea imbricata Lindl. =Hexisea imbricata
Dipteranthus peruvianus Schltr. =Notylia peruviana
Duboisia reymondii H. Karst. =Myoxanthus reymondii
Duckeella adolphii auct. non Porto et Brade 1940: sensu Foldats in Lasser =Duckeella pauciflora
Echinella aspasicensis (Rchb. f.) Pridgeon & M.W. Chase =Pleurothallis aspasicensis
Echinella lappiformis (A.H. Heller & L.O. Williams) Pridgeon & M. W. Chase =Pleurothallis lappiformis
Echinosepala aspasicensis (Rchb. f.) Pridgeon & M.W. Chase =Pleurothallis aspasicensis
Echinosepala lappiformis (A.H. Heller & L.O. Williams) Pridgeon & M. W. Chase =Pleiirothallis lappiformis
Elleanthus attenuatus J.R. Johnst. =Elleanthus arpophyllostachys
Elleanthus kermesinus auct. non (Lindl.) Rchb. f. 1862: sensu Foldats in Lasser =Elleanthus confusus
Elleanthus linifolius auct., non C. Presl =Elleanthus graminifolius
Elleanthus puber Cogn. =Palmorchis puber
Elleanthus pusillus Schltr. =Elleanthus graminifolius
Elleanthus strobilifer (Poepp. & Endl.) Rchb. f. [not in flora area]
Elleanthus virgatus auct. non (Rchb. f.) C. Schweinf. 1938: sensu Dunst. & Garay =Elleanthus confusus
Encyclia abbreviata auct., non (Schltr.) Dressier =Prosthechea calamaria
Encyclia acuta auct. non Schltr. 1825: sensu Camevali & I. Ramirez =Encyclia chloroleuca
Encyclia aemula (Lindl.) Camevali & 1. Ramirez =Prosthechea aemula
Encyclia amicta auct. non (Linden & Rchb.) Schltr. 1919: sensu Camevali & 1. Ra =Encyclia pilosa p.p., Encyclia
conchaechila p.p.
Encyclia atropurpurea Schltr. =Encyclia cordigera
Encyclia calamaria (Lindl.) Pabst =Prosthechea calamaria
Encyclia chloroleuca (Hook.) Schltr. =Encyclia chloroleuca
Encyclia ciliaris (L.) Lemee =Epidendrum ciliare
Encyclia crassilabia (Poepp. & Endl.) Dressier =Prosthechea crassilabia
Encycliafragrans subsp. aemula (Lindl.) Dressier & G.E. Pollard =Prosthechea aemula
EncycliaJauana Camevali & I. Ramirez =Prosthecheajauana
Encyclia latipetala (C. Schweinf) Pabst ^Encyclia pachyantha
Encyclia maravalensis Withner ^Encyclia chloroleuca
Encyclia oncidioides auct. non (Lindl.) Schltr. 1833: sensu Camevali & 1. Ramirez =Encyclia granitica
Encyclia oncidioides (Lindl.) Schltr. [not in flora area]
Encyclia palacea (Rchb. f ) Lemee [not in flora area] =Nidema boothii
Encyclia pamplonense (Rchb. f ) Carncvali =Prosthechea tigrina
Encyclia picta auct. non (Lindl.) Hoehne 1952: sensu Camevali & 1. Ramirez =Encyclia guianensis
Encyclia pygmaea (Hook.) Dressier =Prosthechea pygmaea
Encyclia recurvata auct. non Schltr. 1919: sensu Dunst. & Garay =Encyclia ivonae
Encyclia recurvata Camevali & I. Ramirez =Encyclia ivonae
Encyclia tigrina (Linden ex Lindl.) Camevali & 1. Ramirez =Prosthechea tigrina
Encyclia tumuc-humaciense (Veyret) Hagsater =Epidendrum tumuc-humaciense
138 Checklist of the Plants of the Guiana Shield
Encyclia vespa (Veil.) Dressier =Prosthechea vespa
Encyclia wageneri (Klotzsch, Otto & Dietr.) Schltr. =Encyclia diurna
Epidendropsis vincentina (Lindl.) Garay & Dunst. =Epidendrum vincentinum
Epidendropsis violascens (Ridl. ex Thum) Garay & Dunst. =Epidendrum violascens
Epidendrum aemulum Lindl. =Prosthechea aemula
Epidendrum ahum Ridl. [nom. superfl.] ^Epidendrum alsum
Epidendrum amazonicum Schltr. =Epidendrum coronatum
Epidendrum amictum auct. non Linden & Rchb. f. 1855: sensu Dunst. & Garay =Encyclia leucantha
Epidendrum atropurpureum auct. non Willd. 1805: sensu Dunst. & Garay =Encyclia cordigera
Epidendrum bathyschistum Schltr. ^Epidendrum cristatum
Epidendrum bicornutum Hook. =Caularthron bicornutum
Epidendrum bifidum Aubl. [not in flora area]
Epidendrum bilamellatum Rchb. f. =Caularthron bilamellatum
Epidendrum boothii (Lindl.) L.O. Williams [not in flora area] =Nidema boothii
Epidendrum caespitosum Barb. Rodr. =Prosthechea pygmaea
Epidendrum calamarium Lindl. =Prosthechea calamaria
Epidendrum carnosum Lindl. =Epidendrum durum
Epidendrum carthagenense Jacq. =Trichocentrum carthagenense
Epidendrum caudatum L. =Brassia caudata
Epidendrum chloranthum Lindl. =Encyclia chloroleuca
Epidendrum chloroleucum Hook. =Encyclia chloroleuca
Epidendrum clavatum Lindl. =Epidendrum purpurascens
Epidendrum cochleatum L. =Encyclia cochleata
Epidendrum conchaechilum Barb. Rodr. =Encyclia conchaechila
Epidendrum concretum Jacq. =Polystachya concreta
Epidendrum cordigerum (Kunth) Foldats =Encyclia cordigera
Epidendrum coriifolium auct. non Lindl. 1851: sensu Dunst. & Garay =Epidendrum amazonicoriifolium
Epidendrum coriifolium Lindl. [all material probably referable to E. imitans in flora area]
Epidendrum crassifolium Lindl. =Epidendrum secundum
Epidendrum crassilabium Poepp. «fe Endl. =Prosthechea crassilabia
Epidendrum cucullatum L. =Brassavola cucullata
Epidendrum decipiens auct., non Lindl. =^Epidendrum ibaguense
Epidendrum dendrobioides auct. non Thunb. 1818: sensu Dunst. & Garay =Epidendrum ulei
Epidendrum dendrobioides Thunb. [not in flora area]
Epidendrum dichotomum C. Presl =Epidendrum secundum
Epidendrum diurnum (Jacq.) Rchb. f. =Encyclia diurna
Epidendrum diurnum auct. non (Jacq.) Rchb. f. 1866 [1867]: sensu Foldats in Lasser =Encyclia spp.
Epidendrum diurnum (Jacq.) Cogn. [nom. illeg.] =Encyclia diurna
Epidendrum duckei Porto & Brade =Epidendrum hombersleyi
Epidendrum durum auct. non Lindl. 1841: sensu Foldats in Lasser =Epidendrum alsum p.p.; Epidendrum ulei p.p.
Epidendrum echinocarpum Sw. [nom. superfl.] =Dichaea pendula
Epidendrum ellipticum Graham ex Hook. =Epidendrum secundum
Epidendrum elongatum Jacq. =Epidendrum secundum
Epidendrum ernstii Schltr. =Epidendrum klotzscheanum
Epidendrumfastigiatum auct. non Lindl. 1853: sensu Dunst. & Garay =Epidendrum unguiculatum
Epidendrum flabelliformis Sw. =Cochleanthes flabelliformis
Epidendrum floribundum Kunth =Epidendrum densiflorum
Epidendrum florijugum Barb. Rodr. =Epidendrum sculptum
Epidendrum fragrans auct. non Sw. 1788: sensu Foldats in Lasser =Prosthechea aemula
Epidendrum fragrans Sw. [Guianas records =Prosthechea aemula'] =Prosthecheafragrans
Epidendrum fragrans var. aemulum (Lindl.) Barb. Rodr. =Prosthechea aemula
Epidendrum frigidum var. stenophyton auct. non (Schltr.) C. Schweinf. 1944: sensu Foldats =Epidendrum imthurnii
Epidendrum fulgens Brongn. ? =Epidendrum ibaguense
Epidendrum fuscatum Sm. =Epidendrum anceps
Epidendrum globosum Jacq. =Jacquiniella globosa
Epidendrum graniticum Bateman ex Lindl. =Encyclia granitica
Epidendrum huebneri auct. non Schltr. 1925: sensu Dunst. & Garay =Epidendrum orchidiflorum
Epidendrum huebneri Schltr. =Epidendrum myrmecophorum
Epidendrum ibaguense var. confluens (Lindl.) C. Schweinf. =Epidendrum calanthum
Epidendrum ibaguense var. schomburgkii (Lindl.) C. Schweinf. =Epidendrum macrocarpum
Epidendrum imatophyllum Lindl. =Epidendrum flexuosum
Epidendrum imbricatum Lam. =Lockhartia imbricata
Epidendrum imitans auct. non Schltr. 1921: sensu G.A. Romero & Camevali =Epidendrum amazonicoriifolium
Epidendrum incisum Veil. =Epidendrum macrocarpum
Checklist of the Plants of the Guiana Shield 139
Epidendrum inconstans Ames =Epidendrum secundum
Epidendntm indivisum Bradf. ex Griseb. =Caularthron bilamellatum f. indivisa
Epidendrum ionosmum Lindl. =Encyclia ionosma
Epidendrum labiatum Sw. =Leochilus labiatus
Epidendrum labiosum Rich. =Zygosepalum labiosum
Epidendrum latifolium (Lindl.) Garay & H.R. Sweet =Epidendrum carpophorum
Epidendrum latipetalum C. Schweinf. =Encyclia pachyantha
Epidendrum laxum Poepp. & Endl. =Epidendrum compressum
Epidendrum lechleri auct. non Rchb. f. 1876: sensu Foldats in Lasser =Epidendrum chimantense
Epidendrum leucanthum (Schltr.) Schnee =Encyclia leucantha
Epidendrum leucanthum (Schltr.) C. Schweinf. =Encyclia leucantha
Epidendrum lindenii Lindl. =Epidendrum secundum
Epidendrum lineare Jacq. =Isochilus linearis
Epidendrum maculatum Aubl. =Oncidium luridum
Epidendrum miersii Lindl. ^Epidendrum saxatile
Epidendrum moyobambae Kraenzl. =Epidendrum coronatum
Epidendrum musciferum Lindl. =Epidendrum anceps
Epidendrum nocturnum var. latifolium Lindl. =Epidendrum carpophorum
Epidendrum nocturnum var. minus Cogn. =Epidendrum longicolle
Epidendrum nocturnum var. tumuc-humacience Veyret =Epidendrum tumuc-humaciense
Epidendrum odoratissimum Lindl. =Encyclia odoratissima
Epidendrum oncidioides auct. non Lindl. 1833: sensu Dunst. & Garay =Encyclia auyantepuiensis
Epidendrum oncidioides Lindl. =Encyclia oncidioides
Epidendrum ophioglossoides Jacq. =Stelis ophioglossoides
Epidendrum ottonis Rchb. f. =Nidema ottonis
Epidendrum pachyanthum Lindl. =Encyclia pachyantha
Epidendrum palmarum Salzm. ex Lindl. =Vanilla palmarum
Epidendrum pamplonense Rchb. f. =Prosthechea tigrina
Epidendrum paniculatum auct. non Ruiz & Pav. 1798: sensu Foldats in Lasser =Epidendrum unguiculatum p.p., E.
densiflorum p.p.
Epidendrum paniculatum var. unguiculatum C. Schweinf. =Epidendrum unguiculatum
Epidendrum pictum Lindl. —Encyclia picta
Epidendrum pilosum C. Schweinf. =Encyclia pilosa
Epidendrum pusillum L. =Erycina pusilla
Epidendrum pygmaeum Hook. =Prosthechea pygmaea
Epidendrum pygmaeum Hook. [nom. superfl.] =Prosthechea pygmaea
Epidendrum raniferum auct., non Lindl. =Epidendrum cristatum
Epidendrum rectopedunculatum C. Schweinf. ^Epidendrum myrmecophorum
Epidendrum rectopedunculatum C. Schweinf. f. denticulatum C. Schweinf. ^Epidendrum orchidiflorum
Epidendrum remotiflorum C. Schweinf. =Encyclia remotiflora
Epidendrum ruscifolium Jacq. =Pleurothallis ruscifolia
Epidendrum satyrioides Sw. =Ionopsis satyrioides
Epidendrum schlechterianum auct. non Ames 1924: sensu Foldats in Lasser =Epidendrum congestoides
Epidendrum schomburgkii Lindl. ^Epidendrum macrocarpum
Epidendrum schomburgkii var. confluens Lindl. =Epidendrum calanthum
Epidendrum secundum auct. non Jacq. 1760: sensu Dunst. & Garay =Epidendrum anceps
Epidendrum selligerum auct. non Bateman ex Lindl. 1838: sensu Dunst. & Garay =Encyclia granitica
Epidendrum serricardium Schltr. =Epidendrum vincentinum
Epidendrum spilotum Garay & Dunst. =Epidendrum myrmecophorum
Epidendrum spruceanum Lindl. =Epidendrum nocturnum
Epidendrum stenopetalum Hook, [name misapplied] =Dimerandra stenopetala (Hook.) Schltr
Epidendrum teretifolium Sw. ^Jacquiniella teretifolia
Epidendrum tigrinum Linden ex Lindl. =Prosthechea tigrina
Epidendrum trichorhizum Kraenzl. =Xerorchis trichorhiza
Epidendrum trigoniflorum Sw. =Stelis ophioglossoides
Epidendrum uniflorum Barb. Rodr. =Epidendrum althausenii
Epidendrum utricularioides Sw. =Ionopsis utricularioides
Epidendrum variegatum Hook. =Prosthechea vespa
Epidendrum vespa Veil. =Prosthechea vespa
Epidendrum wageneri Klotzsch, Olio & Dietr. =Encyclia diurna
Epidendrum weddellii Lindl. =Epidendrum saxatile
Epidendrum xanthinum Lindl. ^^^Epidendrum secundum
Epidendrum yatapuense Barb. Rodr. =Epidendrum compressum
Epistephium cruegeri Rchb. f ex Griseb. [nomen nudum] =Epistephium subrepens
140 Checklist of the Plants of the Guiana Shield
Epistephium petiolatum Huber =Epistephium parviflorum
Eriopsis colombiana Schltr. =Eriopsis rutidobulbon
Eriopsis grandibiilbosa Ames «& C. Schweinf. =Eriopsis biloba
Eriopsis schomburgkii (Rchb. f.) Rchb. f. =Eriopsis biloba
Eriopsis sprucei Rchb. f. =Eriopsis sceptrum
Erythrodes confusa C. Schweinf. =Aspidogyne confusa
Erythrodes petersiana (Cogn.) Ames =Ligeophila peterana
Erythrodes plantaginea (L.) Fawc. & Rendle [dubious in Guianas]
Erythrodes pumilus (Cogn.) Pabst =Aspidogyne pumila
Erythrodes robusta C. Schweinf. =Aspidogyne robusta
Erythrodes santensis (Kraenzl.) C. Schweinf. =Ligeophilajuruensis
Erythrodes steyermarkii (Camevali & Foldats) Camevali & Dodson =Aspidogyne steyermarkii
Erythrodes stigmatoptera (Rchb. f ) Pabst =Ligeophila stigmatoptera
Eulophidium maculatum (Lindl.) Pfitzer =Oeceoclades maculata
Evelyna arpophyllostachys Rchb. f. =Elleanthus arpophyllostachys
Evelyna columnaris Lindl. =Elleanthus columnaris
Evelyna gracilis Rchb. f. =Elleanthus gracilis
Evelyna wageneri Rchb. f. =EUeanthus wageneri
Fernandezia acuta Lindl. =Lockhartia acuta
Fernandezia graminifolia Ruiz & Pav. =Scaphyglottis graminifolia
Fractiunguis reflexa Schltr. =Reichenbachanthus reflexus
Galeandrajuncea Lindl. =Galeandra stylomisantha
Galeandra lacustris Barb. Rodr. =Galeandra minax
Galeandra montana Barb. Rodr. [dubious in Guianas] =Galeandra lagoensis Reichb.f. & Warm.
Garayella hexandra (Garay & Dunst.) Brieger [nom. inval.] =Pleurothallis hexandra
Gongora macrantha Hook. =Coryanthes macrantha
Gongora quinquenervis Ruiz & Pav. [not in flora area]
Goodyera guayanensis Lindl. =Brachystele guayanensis
Habenaria amalfitana F. Lehm. & Kraenzl. =Habenaria parviflora
Habenaria amazonica Schltr. =Habenaria platydactyla
Habenaria arecunarum Schltr. =Habenaria parviflora
Habenaria caldensis auct. non Kraenzl.: sensu Dunst. & Garay =Habenaria ernestii
Habenaria caldensis auct. non Kraenzl. 1892: sensu Foldats in Lasser =Habenaria lehmanniana
Habenaria culmiformis Schltr. =Habenaria leprieuri
Habenaria demerarensis Rchb. f. ex M.R. Schomb. =Habenaria parviflora
Habenaria enthomantha auct. non (La Llave & Lex) Lindl. 1835: sensu Foldats in Lasser =Habenaria armata
Habenaria ernestii auct. non Schltr. 1914: sensu Foldats in Lasser =Habenaria caldensis
Habenariafastor auct., non Warm. =Habenaria longicauda
Habenaria georgii Schltr. =Habenaria pratensis var. parviflora
Habenaria goyazensis Cogn. [as circumscribed by Renz, not in flora area]
Habenaria guanchezii Camevali & I. Ramirez =Habenaria pygmaea
Habenaria leaoana Schltr. =Habenaria sprucei
Habenaria leprieuri auct. non Rchb. f. 1846: sensu Foldats in Lasser =Habenaria heptadactyla
Habenaria leprieuri var. heptadactyla (Rchb. f ) R.E. Schult. =Habenaria heptadactyla
Habenaria macroceratitis Willd. =Habenaria quinqueseta
Habenaria maculosa Lindl. =Habenaria monorrhiza
Habenaria moritzii Ridl. =Habenaria armata
Habenaria muelleriana Cogn. =Habenaria macilenta
Habenaria pauciflora (Lindl.) Rchb. f. =Habenaria trifida
Habenaria petalodes auct. non Lindl. 1835: sensu Dunst. & Garay =Habenariafloribunda
Habenaria rodeiensis auct. non Barb. Rodr.: sensu Dunst. & Garay =Habenaria ernestii
Habenaria sartor auct. non Lindl. 1843: sensu Foldats in Lasser =Habenaria longicauda
Habenaria savannensis Renz =Habenaria parviflora
Habenaria speciosa Poepp. & Endl. =Habenaria monorrhiza
Habenaria staminodiata Schltr. =Habenaria macilenta
Habenaria viridiaurea Lindl. ex Kraenzl. =Habenaria heptadactyla
Hexadesmia bifida Rchb. f =Scaphyglottis bifida
Hexadesmia boliviensis Rolfe =Scaphyglottis boliviensis
Hexadesmia dunstervillei Garay =Scaphyglottis dunstervillei
Hexadesmiafusiformis Griseb. =Scaphyglottis fusiformis
Hexadesmia sessilis Rchb. f =Scaphyglottis sessilis
Hexisea bidentata var. imbricata (Lindl.) C. Schweinf. =Hexisea imbricata
Hexisea reflexa (Lindl.) Rchb. f. ex Griseb. =Reichenbachanthus reflexus
Hoehneella santos-nevesii Ruschi =Chaubardia surinamensis
Checklist of the Plants of the Guiana Shield 141
Houlletia vittata Lindl. =Braemia vittata
Huebneria yauaperyensis Schltr. =Orleanesia yauaperyensis
Huebneria yauaperyemis auct. non Schltr. 1925: sensu Hoehne =Epidendrum sculptum
Humboldtia aspasicensis (Rchb. f.) Kuntze =Pleurothallis aspasicensis
Humboldtia biflora (H. Focke) Kuntze =Trichosalpinx orbicularis
Humboldtia ciliata (Knowles & Westc.) Kuntze =Pleurothallis lanceana
Humboldtia dura (Lindl.) Kuntze =Trichosalpinx dura
Humboldtiafockei (Lindl.) Kuntze =Pleurothallis fockei
Humboldtia foliata (Griseb.) Kuntze =Trichosalpinx dura
Humboldtia grobyi (Lindl.) Kuntze =Pleurothallis grobyi
Humboldtia memor (Rchb. f.) Kuntze =Trichosalpinx memor
Humboldtia miqueliana (H. Focke) Kuntze =Pleurothallis miqueliana
Humboldtia orbicularis (Lindl.) Kuntze =Trichosalpinx orbicularis
Humboldtia pruinosa (Lindl.) Kuntze =Pleurothallis pruinosa
Humboldtia revoluta Ruiz & Pav. =Pleurothallis revoluta
Humboldtia zephyrina (Rchb. f.) Kuntze =Pleurothallis zephyrina
Huntleya albido-fulva Lem. =Huntleya meleagris
Huntleya violacea Lindl. =Bollea violacea
Hybochilus huebneri Mansf. =Polyotidium huebneri
lonopsis paniculata Lindl. =Ionopsis utricularioides
lonopsis pygmaea Cogn. =Stictophyllorchis pygmaea
lonopsis teres Lindl. =Ionopsis satyrioides
Isochilus elegans H. Focke =Dimerandra elegans
Kefersteinia pusilla auct., non (C. Schweinf.) C. Schweinf. =Kefersteinia lafontainei
Kegelia houtteana Rchb. f. =Kegeliella houtteana
Koellensteinia alba (Ridl. ex Thurn) Schltr. =Otostylis lepida
Koellensteinia eburnea (Barb. Rodr.) Schltr. =Koellensteinia tricolor
Laelia crispa (Lindl.) Lemee =Schomburgkia crispa
Laelia marginata (Lindl.) L.O. Williams =Schomburgkia marginata
Laelia undulata (Lindl.) L.O. Williams =Schomburgkia undulata
Lanium microphyllum (Lindl.) Benth. =Epidendrum microphyllum
Lanium subulatum Rolfe =Epidendrum stiliferum
Leaoa reedii (Rchb. f.) Garay =Scaphyglottis sessilis
Lepanthesfunerea Barb. Rodr. =Pleurothallis funerea
Lepanthes orbiculata Lindl. ex Rchb. f. =Trichosalpinx orbicularis
Lepanthes yauaperyensis Barb. Rodr. =Pleurothallis mentosa
Leptorchis kappleri (Rchb. f.) Kuntze =Liparis nervosa
Limodorum altum L. =Eulophia alta
Limodorum diurnum Jacq. =Encyclia diurna
Limodorum grandiflorum Aubl. =Cleistes grandiflora
Limodorum lanceolatum Aubl. =Sacoila lanceolata
Limodorum pendulum Aubl. =Dichaea pendula
Lindleyella aurantiaca (Lindl.) Schltr. ^Rudolfiella aurantiaca
Liparis elata Lindl. =Liparis nervosa subsp. nervosa
Liparis elliptica (Rchb. f.) Rchb. f. [non Wight] =Liparisjamaicensis
Liparis kappleri (Rchb. f.) Rchb. f. =Liparis nervosa
Liparis nervosa f. kappleri (Rchb. f.) Christenson & Camevali =Liparis nervosa
Liparis vexillifera auct. non (La Llave & Lex.) Cogn.: sensu Dunst. & Garay "^Liparis jamaicensis
Lockhartia elegans Hook. =Lockhartia imbricata
Lockhartia lasseri Schnee ^Lockhartia acuta
Lockhartia micrantha Rchb. f. [not in flora area]
Lockhartia pallida Rchb. f. =Lockhartia acuta
Lockhartia weigeltii Rchb. & Rchb. f. ex Rchb. f. ^Lockhartia imbricata
Lophiaris carthagenensis (Jacq.) Braem =Trichocentrum carthagenense
Lophiaris fragrans Raf. =Trichocentrum lanceanum
Lophiaris lanceana (Lindl.) Braem =Trichocentrum lanceanum
Lophiaris nana (Lindl.) Braem =Trichocentrum nanum
Lycaste lindeniana (Rchb. f.) G. Nicholson =Paphinia lindeniana
Lycomormium minus Kraenzl. =Peristeria cerina
Macradenia modesta (Rolfe) Rolfe =Macradenia brassavolae
Macradenia mutica Lindl. =Leucohyle mutica
Macradenia surinamensis Rchb. f. & Wullschl. -^Macradenia lutescens
Macradenia triandra Lindl. =Macradenia lutescens
Macroclinium norae (Garay) Dodson =Macroclinium mirabile
142 Checklist of the Plants of the Guiana Shield
Malaxis excavata auct. non (Lindl.) Kuntze 1891: sensu Foldats in Lasser =Malaxis maguirei
Malaxis umbelliflora Sw. [not in flora area]
Manniella americana C. Schweinf. & Garay =Helonoma amehcana
Marsupiaha equitans Schltr. =MaxiUaria equitans
Marsupiaria mattogrossensis (Brade) Hoehne =Maxillaha equitans
Masdevallia brevis Rchb. f. =Scaphosepalum breve
Masdevallia cayennensis Rchb. f. =Masdevallia ciiprea
Masdevallia guttulata auct. non Rchb. f. 1877: sensu Dunst. & Garay =Masdevallia norae
Masdevallia huebneri Schltr. =Trisetella triglochin
Masdevallia surinamensis H. Focke =Masdevallia mimita
Masdevallia triahstella auct. non Rchb. f. 1876: sensu Foldats in Lasser =Trisetella triglochin
Masdevallia triglochin Rchb. f. =Trisetella triglochin
Maxillaria amazonica Schltr. =Maxillaria kegelii
Maxillaria amazonica auct. non Schltr. 1925: sensu Dunst. & Garay =Maxillaria porrecta
Maxillaria anatamorum auct. non Rchb. f.: sensu Foldats in Lasser =Maxillaria albiflora
Maxillaria arachnitis auct non Rchb. f.: sensu Foldats in Lasser =Maxillaria marmoliana
Maxillaria attenuata Ames & C. Schweinf. =Maxillaria giiadalupensis
Maxillaria auyantepuiensis auct. non Foldats sensu: Foldats in Lasser =Maxillaria longipetiolata
Maxillaria auyantepuiensis subsp. epiphytica Camevali & 1. Ramirez =Maxillaria longipetiolata
Maxillaria brevipedunculata Ames & C. Schweinf. =Maxillaria nasuta
Maxillaria brunnea auct. non Linden & Rchb. f. 1854: sensu Dunst. & Garay =Maxillaria cryptobulbon
Maxillaria brunnea Linden & Rchb. f. ^Maxillaria porrecta
Maxillaria buchtienii Schltr. =Maxillaria splendens
Maxillaria colleyi Bateman ex Lindl. =Xylobium colleyi
Maxillaria condensata C. Schweinf. =Maxillaria mapiriensis
Maxillaria coriacea Barb. Rodr. =Maxillaria desvauxiana
Maxillaria cozieriana H.G. Jones [possibly distinct in GU] =Maxillaria camaridii
Maxillaria cristata Lindl. =Paphinia cristata
Maxillariafoveata Lindl. =Xylobiumfoveatum
Maxillaria graminea Lindl. =Koellensteinia graminea
Maxillaria grandiflora auct. non (Kunth) Lindl. 1832: sensu Foldats in Lasser =MaxiUaria eburnea
Maxillaria iguapensis Hoehne & Schltr. =Maxillaria imbricata
Maxillaria lactea Schltr. ^Maxillaria alba
Maxillaria lactea Schltr., non Schltr. 1923 =Maxillaria kegelii
Maxillaria lactiflora Pabst =Maxillaria kegelii
Maxillaria longifolia (Barb. Rodr.) Cogn. =Heterotaxis superflua
Maxillaria macrophylla Poepp. & Endl. =Lycaste macrophylla
Maxillaria melina auct. non Lindl.: sensu Foldats in Lasser =Maxillaria albiflora
Maxillaria nana Hook. =Maxillaria uncata
Maxillaria nasuta auct. non Rchb. f. 1866: sensu Dunst. & Garay =Heterotaxis proboscidea
Maxillaria pallidiflora Hook. =Xylobium pallidiflorum
Maxillaria pendens Pabst =Maxillaria rigida
Maxillaria perparva Garay & Dunst. =Hylaeorchis petiolaris
Maxillaria petiolaris Schltr. =Hylaeorchis petiolaris
Maxillaria petiolaris A. Rich, [nomen nudum] =Maxillaria desvauxiana
Maxillaria proboscidea Rchb. f. =Heterotaxis proboscidea
Maxillaria purpurea auct. non (Spreng.) Ames & Correll 1943: sensu Foldats in Lasser =Maxillaria conferta
Maxillaria ringens auct. non Rchb. f. 1863: sensu Dunst. & Garay =Maxillaria porrecta
Maxillaria riopalenquensis Dodson =Maxillaria longipetiolata
Maxillaria rudolfii Hoehne [nomen nudum] =Hylaeorchis petiolaris
Maxillaria rufescens auct. non Lindl. 1835 [1836]: sensu Foldats in Lasser =Maxillaria acutifolia
Maxillaria rugosa Schltr. =Maxillaria schlechteri
Maxillaria simulans Ames & C. Schweinf. ^Maxillaria conferta
Maxillaria spilotantha auct. non Rchb. f. 1854: sensu Foldats in Lasser =Maxillaria guareimensis
Maxillaria steelei Hook. =Scuticaria steelei
Maxillaria superflua Rchb. f. ='Heterotaxis superflua
Maxillaria surinamensis H. Focke ex Rchb. f. =Maxillaria conferta
Maxillaria tafallae (Rchb. f.) C. Schweinf. =Maxillaria ramosa
Maxillaria taracuana Schltr. =Maxillaria kegelii
Maxillaria tarumaensis Hoehne =Heterotaxis superflua
Maxillaria trinitatis Ames ex Broadway [nom. superfl.] =Maxillaria porrecta
Maxillaria trinitatis auct. non Ames 1923: Dunst. & Garay =Maxillaria cryptobulbon
Maxillaria trinitatis Ames =Maxillaria porrecta
Maxillaria uncata auct. non Lindl. 1837: sensu Foldats in Lasser =Maxillaria squamata
Checklist of the Plants of the Guiana Shield 143
Maxlllario vandifonnis (Schltr.) C. Schweinf. [possibly distinct species] =Maxillaria equitans
Maxillaria variabilis auct., non Bateman ex Lindl. =MaxiUaria caespitifica
Maxillaria verrucifera C. Schweinf. =Maxillaria desvanxiana
Maxillaria vestita (Sw.) Lemee =MaxiUaria parviflora
Maxillaria villosa (Barb. Rodr.) Cogn. =Heterotaxis villosa
Maxillaria violaceopimctata Rchb. f. =Heterotaxis violaceopiinctata
Medadeniitm labiosiim (Rich.) Cogn. =Zygosepalurn labiosum
Medadenium lindeniae (Rolfe) Cogn. =Zygosepaliim lindeniae
Menadeniiim kegelii (Rchb. f.) Cogn. =Zygosepalum kegelii
Menadenium labiosum (Rich.) Cogn. =Zygosepalum labiosum
Mendoncella burkei (Rchb. f.) Garay =Galeottia burkei
Mendoncellajorisiana (Rolfe) A.D. Hawkes =Galeottiajorisiana
Mesadenella esmeraldae (Lindl. & Rchb. f.) Pabst & Garay =Mesadenella cuspidata
Monachanthus discolor Lindl. =Catasetum discolor
Monachanthus roseo-albus Hook. =Catasetum xroseo-album
Myanthus barbatus Lindl. =Catasetum barbatum
Myanthus deltoideus Lindl. =Catasetum deltoideum
Myoxanthus aspasicensis (Rchb. f ) Luer =Pleurothallis aspasicensis
Myoxanthus aspasicensis subsp. arenicola Camevali & I. Ramirez =Pleurothallis arenicola
Myoxanthus lappiformis (A.H. Heller & L.O. Williams) Luer =Pleurothallis lappiformis
Nanodes discolor Lindl. =Epidendrum schlechterianum
Neobartlettia guianensis Schltr. =Palmorchis guianensis
Neolehmannia apagana (Mansf.) Garay & Dunst. =Epidendrum apaganum
Neolehmannia barbeyana auct. non (Kraenzl.) Garay & Dunst. 1976: sensu Dunst. & Garay =Epidendrum althausenii
Neolehmannia pabstii Braga =Epidendrum dichaeoides
Neottia acaulis Sm. =Sarcoglottis acaidis
Neottia calcarata Sw. =Eltroplectris calcarata
Nidema boothii (Lindl.) Schltr. [not in flora area]
Notylia amesii L.B. Sm. & S.K. Harris =Notylia microchila
Notylia amesii L.B. Sm. & S.K. Harris =Notylia microchila
Notylia bicolor Lindl. [not in flora area] =Macroclinium bicolor (Lindl.) Dodson
Notylia mirabilis C. Schweinf. =Macroclinium mirabile
Notylia nana Cogn. =Notylia angustifolia
Notylia norae Garay =Macroclinium mirabile
Notylia trullifera Brade =Notylia longispicata
Notylia venezuelana Schltr. =Notylia fragrans
Notylia wullschlaegeliana H. Focke =Macroclinium wullschlaegelianum
Octomeria brevifolia Cogn. =Octomeria scirpoidea
Octomeria complanata C. Schweinf. =Octomeria yauaperyensis
Octomeria deltoglossa Garay ^'Octomeria exigua
Octomeria exigua var. elata C. Schweinf. =Octomeria exigua
Octomeria grandiflora Lindl. [not in flora area]
Octomeria kestrochila Garay & Dunst. =Octomeria exigua
Octomeria pusilla Lindl. [not in flora area]
Odontoglossum arminii Rchb. f. =Otoglossum arminii
Odontoglossum brevifolium auct. non. Lindl. 1844: sensu Foldats in LassQr =Otoglossum arminii
Oncidium altissimum auct., non (Jacq.) Sw. =Oncidium baueri
Oncidium ampllatum Lindl. =Chelyorchis ampliata
Oncidium bernouillanum Kraenzl. =Chelyorchis ampliata
Oncidium carthagenense (Jacq.) Sw. =Trichocentrum carthagenense
Oncidium cebo 11eta (Jacq.) Sw. ^Trichocentrum cebolleta
Oncidium convolvulaceum Lindl. =Otoglossum globuliferum
Oncidium emarginatum G. Mey. =Dimerandra emarginata
Oncidium globuliferum auct. non Kunth 1815 [1816]: sensu Dunst. & Garay =Otoglossum scansor
Oncidium globuliferum Kunth =Otoglossum globuliferum
Oncidium glossomystax Rchb. f ^Eiycina glossomystax
Oncidium guianense (Aubl.) Garay [not in flora area]
Oncidium kappleri Rchb. f ex Lindl. =Oncidium baueri
Oncidium lanceanum Lindl. =Trichocentrum lanceanum
Oncidium lunatum Lindl. =Solenidium lunatum
Oncidium maculatum Urb. =Oncidium luridum
Oncidium nanum Lindl. =Trichocentrum nanum
Oncidium papilio Lindl. =Psychopsis papilio
Oncidium pusillum (L.) Rchb. f =Eiycina pusilla
144 Checklist of the Plants of the Guiana Shield
Oncidium sancti-pauli auct. non Kaenzl. 1922: sensu Foldats in Lasser =Otoglossum globuliferum
Oncidium scansor Rchb. f. =Otoglossum scansor
Oncidium uaipanense Schnee =Oncidium nigratum
Oncidium ultrajectinum Pulle =Trichocentrum cebolleta
Oncidium vagans C. Schweinf. =Oncidium warmingii
Oncidium versteegianum Pulle =Psychopsis versteegianum
Onychium flavescens Blume =Polystachya concreta
Ophrys guianensis Aubl. [not in flora area] =Oncidium guianense (Aubl.) Garay
Ophrys nervosa Thunb. =Liparis nervosa subsp. nervosa
Ophrys nervosa Thunb. =Liparis nervosa
Ophrys peruviana Aubl. =Spiranthes torta
Orchis monorrhiza Sw. =Habenaria monorrhiza
Orchis quinqueseta Michx. =Habenaria quinqueseta
Orchis stylomisantha Veil. =Galeandra stylomisantha
Ornithidium aureum Poepp. & Endl. =Maxillaria aurea
Ornithidium chloroleucum Barb. Rodr. =MaxiUaria conferta
Ornithidium confertum Griseb. =Maxillaria conferta
Ornithidium mapiriense Kraenzl. =Maxillaria mapiriensis
Ornithidium serrulatum Lindl. [non Maxillaria serrulata Ames & Correll 1943] =Maxillaria alticola
Ornithidium squamatum (Barb. Rodr.) Barb. Rodr. =Maxillaria squamata
Ornithidium vestitum Rchb. f. =Maxillaria conferta
Ornithocephalus avicula Rchb. f. =Ornithocephalus kruegeri
Ornithocephalus bicornis auct. non Lindl. 1846: sensu Garay & Dunst. =Ornithocephalus brachyceras
Ornithocephalus planifolius Rchb. f. =Dipteranthus planifolius
Otostylis alba (Ridl. ex Thum) Summerh. =Otostylis lepida
Otostylis paludosa (Cogn.) Schltr. =Otostylis brachystalix
Otostylis venusta (Ridl. ex Oliv.) Schltr. =Otostylis lepida
Paphiopedilum klotzschianum Pfitzer =Phragmipedium klotzschianum
Paphiopedilum lindleyanum Pfitzer =Phragmipedium lindleyanum
Pelexia aphylla Ridl. ex Thum =Sarcoglottis aphylla
Pelexia aphylla Ridl. ex Oliv. [nom. superfl.] =Sarcoglottis aphylla
Pelexia cranichoides Griseb. =Cyclopogon cranichoides
Pelexia maculata Rolfe =Pelexia laxa
Pelexia setacea Lindl. =Eltroplectris calcarata
Peristeria guttata Knowles & Westc. =Peristeria cerina
Phragmipedium kaieteurum (N.E. Br.) Garay =Phragmipedium lindleyanum
Phymatidium panamense Dressier =Eloyella panamensis
Physinga prostrata Lindl. =Epidendrum prostratum
Physosiphon obliquipetalus Ames & C. Schweinf. =Lepanthopsis ohliquipetala
Physurus arietinus Rchb. f. & Warm. =Erythrodes arietina
Physurusjuruensis Hoehne =Ligeophilajuruensis
Physurus paleaceus Schltr. =Erythrodes paleacea
Physurus pumilus Cogn. =Aspidogyne pumila
Physurus santensis Kraenzl. =Ligeophilajuruensis
Physurus stigmatopterus Rchb. f. =Ligeophila stigmatoptera
Pilumna wageneri Rchb. f. =Trichopilia wageneri
Pinelia alticola Garay & Dunst. =Pinelianthe alticola
Platystele lancilabris auct. non (Rchb. f.) Schltr. 1923: sensu Garay & Dunst. =Platystele oxyglossa
Plectrophora cultrifolia auct. non (Barb. Rodr.) Cogn. 1904: sensu Garay & Dunst. =Plectrophora iridifolia
Pleurothallis acuminata H. Focke =Pleurothallis semperflorens
Pleurothallis acutissima Lindl. =Pleurothallis spiculifera
Pleurothallis acutissima auct. non Lindl. 1856: sensu Foldats in Lasser =Pleurothallis trinitensis
Pleurothallis amygdalina Rolfe ex Schltr. [nomen nudum] =Trichosalpinx dura
Pleurothallis anomala Hoehne =Trichosalpinx dura
Pleurothallis apiculata Lindl. =Octomeria apiculata
Pleurothallis barbata H. Focke =Pleurothallis barbulata
Pleurothallis barberiana Rchb. f. =Pleurothallis aristata
Pleurothallis biflora H. Focke =Trichosalpinx orbicularis
Pleurothallis blaisdellii S. Watson [not in flora area] =Trichosalpinx blaisdellii (S. Watson) Luer
Pleurothallis brevis Schltr. =Trichosalpinx memor
Pleurothallis breviscapa C. Schweinf. ^Pleurothallis funerea
Pleurothallis broadwayi KmQ?, =Trichosalpinx dura
Pleurothallis broadwayi subsp. anomala (Hoehne) Garay =Trichosalpinx dura
Pleurothallis cedralensis Ames =Trichosalpinx cedralensis
Checklist of the Plants of the Guiana Shield 145
Pleurothallis ciliahs auct. non (Lindl.) L.O. Williams 1842: sensu Foldats in Lasser =Trichosalpinx memor
Pleurothallis ciliaris (Lindl.) L.O. Williams =Trichosalpinx ciliaris
Pleurothallis ciliata Knowles & Westc. =Pleurothallis lanceana
Pleurothallis ciliata var. elongata C. Schweinf. =Pleurothallis lanceana
Pleurothallis ciliolata auct. non Schltr. 1926: sensu Foldats in Lasser =Pleurothallis funerea
Pleurothallis coffeicola auct. non Schltr. 1929: sensu Foldats in Lasser =Pleurothallis discophylla p.p., P. sandaliorum p.p.
Pleurothallis connata Luer =Lepanthopsis obliquipetala
Pleurothallis consimilis Ames =Pleurothallis yauaperyensis
Pleurothallis consimilis auct. non Ames 1922: sensu Garay & Dunst. ^Pleurothallis garciae
Pleurothallis corazonica F. Lehm. & Kraenzl. =Trichosalpinx dura
Pleurothallis delascioi Camevali & I. Ramirez =Pleurothallis kerrii
Pleurothallis diffusiflora C. Schweinf. =Pleurothallis seriata
Pleurothallis dura Lindl. =Trichosalpinx dura
Pleurothallis egleri Pabst =Trichosalpinx egleri
Pleurothallis elegans auct. non (Kunth) Lindl. 1842: sensu Foldats in Lasser =Pleurothallis moritzii
Pleurothallis exasperata auct. non Lindl. 1859: sensu Foldats in Lasser =Myoxanthus speciosus
Pleurothallis fimbriata Lindl. =Pleurothallis miqueliana
Pleurothallis floripecten Rchb. f. =Lepanthopsis floripecten
Pleurothallis foliata Griseb. =Trichosalpinx dura
Pleurothallis gnomonifera Ames =Trichosalpinx memor
Pleurothallis guadalupensis Cogn. =Trichosalpinx dura
Pleurothallis huebneri Schltr. =Pleurothallis lanceana
Pleurothallis intricata Lindl. =Trichosalpinx intricata
Pleurothallis johnstonii Ames =Platystelejohnstonii
Pleurothallis kegelii Rchb. f. =Pleurothallis lanceana
Pleurothallis lancifera Schltr. =Trichosalpinx orbicularis
Pleurothallis lancilabris var. oxyglossa (Schltr.) C. Schweinf. =Platystele oxyglossa
Pleurothallis lansbergii auct. non Regel 1856: sensu Dunst. & Garay =Pleurothallis coriacardia
Pleurothallis lepanthopsis Schltr. =Trichosalpinx dura
Pleurothallis leptopetala Cogn. =Pleurothallis discoidea
Pleurothallis linguifera auct. non Lindl. 1859: sensu Foldats in Lasser =Pleurothallis coriacardia
Pleurothallis longirostris H. Focke =Pleurothallis ephemera
Pleurothallis marginalis Rchb. f. =Pleurothallis grobyi
Pleurothallis memor Rchb. f. =Trichosalpinx memor
Pleurothallis minutipetala Ames & C. Schweinf =Trichosalpinx memor
Pleurothallis monocardia auct. non Rchb. f. 1855: sensu Garay & Dunst. ^Pleurothallis archidiaconi
Pleurothallis myrmecophila Hoehne =Pleurothallisfockei
Pleurothallis orbicularis (Lindl.) Lindl. =Trichosalpinx orbicularis
Pleurothallis ovalifolia (H. Focke) Rchb. f =Platystele ovalifolia
Pleurothallis oxyglossa Schltr. =Platystele oxyglossa
Pleurothallis papulosa auct. non. Lindl. 1836: sensu Dunst. & Garay =Pleurothallis granitica
Pleurothallis parvilabia C. Schweinf =Myoxanthus parvilabius
Pleurothallis peduncularis Lindl. =Myoxanthus parahybunensis
Pleurothallis penduliflora auct. non Kraenzl. 1905: sensu Garay & Dunst. =Pleurothallis suspensa
Pleurothallis pertenuis C. Schweinf. =Pleurothallis glandulosa
Pleurothallis plumosa Lindl. =Pleurothallis lanceana
Pleurothallis pluriracemosa Garay =Pleurothallis floribunda
Pleurothallis pulchella auct. non (Kunth) Lindl. 1825: sensu Luer =Pleurothallis tepuiensis
Pleurothallis pumila Luer =Trichosalpinx pumila
Pleurothallis revoluta auct. non (Ruiz & Pav.) Garay 1962: sensu Foldats in Lasser ^Pleurothallis suspensa
Pleurothallis reymondii (H. Karst.) Rchb. f =Myoxanthus reymondii
Pleurothallis ringens C. Schweinf ^Pleurothallis hitchcockii
Pleurothallis roraimensis Rolfe =Trichosalpinx roraimensis
Pleurothallis scandens var. simplicicaulis C. Schweinf =Myoxanthus simplicicaulis
Pleurothallis sertularioides var. trinitensis Griseb. =Pleurothallis trinitensis
Pleurothallis setigera Lindl. =Pleurothallis zephyrina
Pleurothallis simplicicaulis (C. Schweinf.) Luer =Myoxanthus simplicicaulis
Pleurothallis speciosus Luer =Myoxanthus speciosus
Pleurothallis spiculifera auct. non Lindl. 1859: sensu Dunst. & Garay ^Pleurothallis trinitensis
Pleurothallis stenopetala Lodd. ex Lindl. =Pleurothallis sclerophylla
Pleurothallis striata H. Focke =Pleurothallis uniflora
Pleurothallis tepuiensis auct. non Camevali & 1. Ramirez 1993: sensu G.A. Romero & Came =Pleurothallis moritzii
Pleurothallis trachyclamys Schltr. =Myoxanthus trachychlamys
Pleurothallis trachytheca F. Lehm. & Kraenzl. =Trichosalpinx orbicularis
146 Checklist of the Plants of the Guiana Shield
Pleurothallis tricarinata H. Focke =Pleurothallisfockei
Pleurothallis tubulosa Lindl. =Restrepiopsis tubulosa
Pleurothallis uncinata auct. non Fawc. 1895: sensu Foldats in Lasser =Pleurothallis arenicola
Pleurothallis uncinata auct. non Fawc. 1895, sensu Foldats p.p., Dunst. & Garay p.p. ^Pleurothallis aspasicensis
Pleurothallis uniflora auct non Lindl. 1836: sensu Dunst. & Garay =Pleurothallis corniculata
Pleurothallis vaginata Schltr. =Pleurothallis imraei
Pleurothallis velaticaulis auct. non Rchb. f. 1849: sensu Foldats in Lasser =Pleurothallis floribunda
Pleurothallis velatipes Rchb. f. =Pleurothallis velaticaulis
Pleurothallis villosa Knowles & Westc. =Trichosalpinx egleri
Pleurothallis viridula Lindl. =Restrepiopsis tubulosa
Pleurothallis vitatta auct. non Lindl. 1838: sensu Foldats in Lasser =Pleurothallis sarcosepala
Pleurothallis vittariifolia Schltr. =Pleurothallis glandulosa
Pleurothallis wendlandiana auct. non Rchb. f. 1888: sensu Dunst. & Garay =Pleurothallis ruscifolia
Pleurothallis williamsii Ames =Trichosalpinx dura
Pogonia grandiflora Rchb. f. =Cleistes grandiflora
Pogonia lepida Rchb. f. =Cleistes lepida
Pogonia macrophylla Lindl. =Psilochilus°macrophyllus
Pogonia moritzii Rchb. f. =Cleistes lepida
Pogonia paludosa (Rchb. f.) Rchb. f. =Cleistes paludosa
Pogonia parviflora (Lindl.) Rchb. f. =Cleistes parviflora
Pogonia physuraefolia Rchb. f. ='Psilochilus macrophyllus
Pogonia physurifolia Rchb. f. =Psilochilus physurifolius
Pogonia rosea (Lindl.) Rchb. f. =Cleistes rosea
Pogonia stricta C. Schweinf. =Cleistes stricta
Pogonia surinamensis Lindl. =Triphora surinamensis
Pogonia tenuis Rchb. f. =Cleistes tenuis
Pogonia triflora C. Schweinf. =Cleistes triflora
Pogonia unifoliata C. Schweinf. =Cleistes unifoliata
Polycycnis vittata (Lindl.) Rchb. f. =Braemia vittata
Polystachya estrellensis Rchb. f. [not in flora area]
Polystachyaflavescens (Blume) J.J. Sm. =Polystachya concreta
Polystachya geraensis Barb. Rodr. [not in flora area]
Polystachya luteola (Sw.) Hook. =Polystachya concreta
Polystachya minuta Britton [not in flora area]
Polystachya nana Poepp. & Endl. [not in flora area]
Polystachya stenophylla Schltr. ^Polystachya amazonica
Ponera amethystina Rchb. f. =Scaphyglottis stellata
Ponerafelskyi Rchb. f. =Scaphyglottis felskyi
Ponera leucantha (Rchb. f.) Rchb. f. =Scaphyglottis leucantha
Prescottia orchioides auct. non Lindl. 1845: sensu Foldats in Lasser =Prescottia auyantepuiensis
Promenaea graminea (Lindl.) Lindl. =Koellensteinia graminea
Prosthechea pamplonensis (Rchb. f.) W.E. Higgins =Prosthechea tigrina
Pseuderiopsis schomburgkii Rchb. f. =Eriopsis biloba
Pseudomaxillaria chloroleuca (Barb. Rodr.) Hoehne =Maxillaria conferta
Psychopsis picta Raf. =Psychopsis papilio
Psygmorchis glossomystax (Rchb. f.) Dodson & Dressier =Erycina glossomystax
Psygmorchis pusilla (L.) Dodson & Dressier =Erycina pusilla
Quekettia pygmaea (Cogn.) Garay & R.E. Schult. =Stictophyllorchis pygmaea
Reichenbachanthus modestus Barb. Rodr. =Reichenbachanthus reflexus
Restrepia kegelii Rchb. f. [non Pleurothallis kegelii Rchb. f. 1877] =Pleurothallis hexandra
Restrepiella viridula (Lindl.) Garay & Dunst. =Restrepiopsis tubulosa
Restrepiopsis viridula (Lindl.) Luer =Restrepiopsis tubulosa
Rhynchopera punctata H. Karst. =Pleurothallis loranthophylla
Rodriguezia secunda Kunth =Rodriguezia lanceolata
Sarcoglottis acaulis auct. non (Sm.) Schltr. 1919: sensu Foldats in Lasser =Sarcoglottis metallica
Sarcoglottis grandiflora auct. non (Lindl.) Klotzsch 1842: sensu Dunst. & Garay =Sarcoglottis acaulis
Sarcoglottis maasorum Pabst =Sarcoglottis metallica
Sarcoglottis picta auct. non Klotzsch 1842: sensu Garay & Dunst. =Sarcoglottis metallica
Satyrium elatum Sw. =Cyclopogon elatus
Satyrium orchioides Sw. =Sacoila lanceolata
Scaphosepalum ochthodes Pfitzer [not in flora area]
Scaphosepalum verrucosum auct. non (Rchb. f.) Pfitzer 1888: sensu Dunst. & Garay =Scaphosepalum breve
Scaphyglottis amethystina (Rchb. f.) Schltr. =Scaphyglottis stellata
Scaphyglottis cuneata Schltr. =Scaphyglottis prolifera
Checklist of the Plants of the Guiana Shield 147
Scaphyglottis huebneri Schltr. =Scaphyglottis boliviensis
Scaphyglottis propinqita auct. non C. Schweinf. 1955; Foldats in Lasser =Scaphyglottis sickii
Scaphyglottis reedii (Rchb. f.) Ames =Scaphyglottis sessilis
Scaphyglottis reflexa Lindl. =Reichenbachanthus reflexus
Scaphyglottis tafallae Rchb. f. =Maxillaria ramosa
Scaphyglottis violacea (Lindl.) Lindl. =Scaphyglottis graminifolia
Scaphyglottis wercklei var. major C. Schweinf. =Scaphyglottis leucantha
Schlechterella aurantiaca (Lindl.) Hoehne =Rudolfiella aurantiaca
Schomburgkia crispa auct. non Lindl. 1838: sensu Foldats in Lasser =Schomburgkia marginata
Schomburgkia rosea auct. non Linden ex Lindl. 1845: sensu Foldats in Lasser =Schomburgkia heidii
Schomburgkia weberbaueriana auct. non Kraenzl. 1906: sensu Garay & Dunst. ^Schomburgkia marginata
Selenipedium kaieteiirum N.E. Br. =Phragmipedium lindleyanum
Selenipedium lindleyanum Rchb. f. =Phragmipedium lindleyanum
Selenipedium schomburgkianum Rchb. f. =Phragmipedium klotzschianum
Serapias caravata Aubl. =Elleanthus caravata
Serrastylis modesta Rolfe =Macradenia brassavolae
Sievekingia dunstervilleorum Foldats =SievekingiaJenmanii
Sobralia biflora auct. non Ruiz & Pav. 1798: sensu Schweinf. =Sobralia valida
Sobralia cattleya Rchb. f. =Sobralia dichotoma
Sobralia elisabethae R.H. Schomb. =Sobralia liliastrum
Sobralia liliastrum var. rosea Lindl. =Sobralia infundibuligera
Sobralia sclerophylla auct. non Lindley =Sobralia stenophylla
Sobralia sessilis auct. non Lindl. 1841 : sensu Foldats in Lasser =Sobralia valida
Sobralia viclacea auct. non Linden & Rchb. f. 1846: sensu Garay & Dunst. =Sobralia sessilis
Sobralia yauaperyensis Barb. Rodr. =Sobralia sessilis
Sobralia yauaperyensis auct. non Barb. Rodr. 1881: sensu Foldats in Lasser =Sobralia violacea
Spathiger rigidus (Jacq.) Small =Epidendrum rigidum
Specklinia aristata (Hook.) Pridgeon & M.W. Chase =Pleurothallis aristata
Specklinia grobyi (Bateman ex Lindl.) Pridgeon & M.W. Chase =Pleurothallis grobyi
Specklinia miqueliana H. Focke =Pleurothallis miqueliana
Specklinia orbicularis Lindl. =Trichosalpinx orbicularis
Specklinia parvifolia (Lindl.) Pridgeon & M.W. Chase =Pleurothallis parvifolia
Specklinia picta (Lindl.) Pridgeon & M.W. Chase =Pleurothallis picta
Specklinia samacensis (Ames) Pridgeon & M.W. Chase =Pleurothallis samacensis
Specklinia seriata (Lindl.) Pridgeon & M.W. Chase =Pleurothallis seriata
Specklinia spiculifera (Lindl.) Pridgeon & M.W. Chase =Pleurothallis spiculifera
Specklinia vittariifolia (Schltr.) Pridgeon & M.W. Chase =Pleurothallis glandulosa
Spiranthes acaulis (Sm.) Cogn. =Sarcoglottis acaulis
Spiranthes aphylla (Ridl. ex Thum) Cogn. =Sarcoglottis aphylla
Spiranthes bifida Ridl. ex Oliv. [nom. superfl.] =Helonoma bifida
Spiranthes bifida Ridl. ex Thum =Helonoma bifida
Spiranthes callifera C. Schweinf. =Pelexia callifera
Spiranthes cranichoides (Griseb.) Cogn. =Cyclopogon cranichoides
Spiranthes elata (Sw.) Rich. =Cyclopogon elatus
Spiranthes goninens is (Pulle) C. Schweinf =Pelexia goninensis
Spiranthes grisebachii Cogn. =Lyroglossa grisebachii
Spiranthes guayanensis (Lindl.) Cogn. =Brachystele guayanensis
Spiranthes lanceolata (Aubl.) Leon =Sacoila lanceolata
Spiranthes laxa (Poepp. & Endl.) C. Schweinf =Pelexia laxa
Spiranthes maculata (Rolfe) C. Schweinf =Pelexia laxa
Spiranthes metallica Rolfe =Sarcoglottis metallica
Spiranthes orchioides (Sw.) A. Rich. =Sacoila lanceolata
Spiranthes quinqueloba (Poir.) Urb. =Spiranthes torta
Spiranthes scopulariae Rchb. f. =Discyphus scopulariae
Spiranthes simplex Griseb. =Sarcoglottis simplex
Stanhopea aurea Lodd. ex Lindl. =Stanhopea wardii
Stanhopea eburnea Lindl. =Stanhopea grandifora
Stanhopea randii Rolfe =Stanhopea Candida
Stanhopea venusta Lindl. =Stanhopea wardii
Stelis aprica auct. non Lindl. 1836: sensu Dunst. & Garay =Stelis schomburgkii
Stelis braccata auct. non Rchb. f & Warsz. 1 854: sensu Foldats in Lasser =Stelis minimiflora
Stelis cupuligera auct. non Rchb. f 1854: sensu Dunst. & Garay =Stelis aviceps
Stelis drosophila Barb. Rodr. =Stelis intermedia
Stelis flavida H. Focke =Pleurothallis pruinosa
148 Checklist of the Plants of the Guiana Shield
Stelis floribunda (Poepp. & Endl.) Pridgeon & M.W. Chase =Pleurothallisfloribunda
Stelis foliosa Lindl. =Polystachyafoliosa
Stelis galeata (Lindl.) Pridgeon & M.W. Chase =Pleurothallis galeata
Stelis grossilabris auct. non Rchb. f. 1881: sensu Foldats in Lasser =Stelis zonata
Stelis imraei (Lindl.) Pridgeon & M.W. Chase =Pleurothallis imraei
Stelis maderoi auct. non Schltr. 1920: sensu G.A. Romero & Camevali =Stelis obovata
Stelis moritzii (Rchb. f.) Pridgeon & M.W. Chase =Pleurothallis moritzii
Stelis obliquipetala (Ames & C. Schweinf.) L.O. Williams =Lepanthopsis obliquipetala
Stelis ovalifolia H. Focke =Platystele ovalifolia
Stelis papaquerensis auct. non Rchb. f. 1849: sensu Foldats in Lasser =Stelis fraterna
Stelis tenuilabris Lindl. [possibly distinct species] =Stelis alata
Stelis trichorrachis auct. non Rchb. f. 1855: sensu Dunst. & Garay =Stelis pygmaea
Stelis trigoniflora (Sw.) Garay =Stelis ophioglossoides
Stelis tristyla auct. non Lindl. 1838: sensu Foldats in Lasser =Stelis bangii
Stelis velaticaulis (Rchb. f.) Pridgeon & M.W. Chase =Pleurothallis velaticaulis
Stelis vulgaris Pridgeon & M.W. Chase =Pleurothallis floribunda
Stenia tigrina (Garay & Dunst.) Foldats =Chaubardiella tigrina
Stenocoryne longicornis (Lindl.) Lindl. =Bifrenaria longicornis
Stenoptera adnata Ridl. ex Oliv. [nom. superfl.] =Gomphichis adnata
Stenoptera adnata Ridl. ex Thum =Gomphichis adnata
Stenoptera costaricensis Schltr. =Gomphichis costaricensis
Stenoptera roehlii Schnee =Eurystyles cotyledon
Stenorrhynchos goninensis Pulle =Pelexia goninensis
Stenorrhynchos lanceolatum (Aubl.) Rich, ex Spreng. =Sacoila lanceolata
Stenorrhynchos laxus Poepp. & Endl. =Pelexia laxa
Stictophyllum pygmaeum (Cogn.) Dodson & M.W. Chase =Stictophyllorchis pygmaea
Sturmia elliptica Rchb. f. =Liparis jamaicensis
Sturmia kappleri Rchb. f. =Liparis nervosa
Tetragamestus modestus Rchb. f. =Scaphyglottis modesta
Teuscheria venezuelana Garay =Teuscheria wageneri
Triaristella triglochin (Rchb. f.) Luer =Trisetella triglochin
Triaristellina triglochin (Rchb. f.) Rauschert ='Trisetella triglochin
Trichocentrum cornucopiae Linden & Rchb. f. =Trichocentrum fuscum
Trichocentrum hartii Rolfe =Trichocentrumfuscum
Trichopilia albida auct. non H. Wendl. 1851: sensu Dunst. & Garay =Trichopilia aenigma
Trichopiliafragrans auct. non (Lindl.) Rchb. f. 1858: sensu Foldats in Lasser =Trichopilia aenigma
Trichopilia mutica (Lindl.) Rchb. f. & Wullschl. ex Rchb. f. =Leucohyle mutica
Trichosalpinx blaisdellii auct., non (S. Watson) Luer =Trichosalpinx egleri
Trichosalpinxfoliata (Griseb.) Luer =Trichosalpinx dura
Trichosalpinx greenwoodiana Soto Arenas =Trichosalpinx memor
Trichosalpinx minutipetala (Ames & Schweinf.) Luer =Trichosalpinx memor
Trichosalpinx nageliana Soto Arenas =Trichosalpinx memor
Trichosalpinx obliquipetala (Ames & C. Schweinf.) Luer =Lepanthopsis obliquipetala
Trichosalpinx patula Luer [nom. inval.] ^Trichosalpinx patula
Trichosalpinx zephyrina (Rchb. f.) Luer =Pleurothallis zephyrina
Trigonidium tenue Lodd. ex Lindl. =Trigonidium acuminatum
Trisetella huebneri (Schltr.) Luer =Trisetella triglochin
Trizeuxis pygmaea (Cogn.) Schltr. =Stictophyllorchis pygmaea
Uleiorchis cogniauxiana f. maior Hoehne =Uleiorchis ulaei
Vanilla aromatica Sw. =Vanilla mexicana
Vanilla ensifolia Rolfe =Vanilla odorata
Vanilla inodora Schiede = Vanilla mexicana
Vanilla pompona auct., non Schiede =Vanilla grandiflora
Veyretia aphylla (Ridl. ex Thum) Szlach. =Sarcoglottis aphylla
Veyretia simplex (Griseb.) Szlach. =Sarcoglottis simplex
Wullschlaegelia aphylla (Sw.) Rchb. f. [not in flora area]
Wullschlaegelia ulaei Cogn. ^Uleiorchis ulaei
Xylobium steyermarkii Foldats =Bifrenaria steyermarkii
Zygopetalum brachystalix Rchb. f. =Otostylis brachystalix
Zygopetalum burkei Rchb. f. =Galeottia burkei
Zygopetalum jorisianum Rolfe =Galeottiajorisiana
Zygopetalum kegelii Rchb. f. ='Zygosepalum kegelii
Zygopetalum lindeniae Rolfe =Zygosepalum lindeniae
Zygopetalum paludosum Cogn. =Otostylis brachystalix
Checklist of the Plants of the Guiana Shield 149
Zygopetalum rostratum Hook. =Zygosepalum labiosum
Zygopetalum tatei Ames & C. Schweinf. =Zygosepalum tatei
Zygopetalum tricolor Lindl. =Koellensteinia tricolor
Zygopetalum venustum Ridl. ex Oliv. =Otostylis lepida
Zygosepalum tatei var. angustilabium C. Schweinf. =Zygosepalum angustilabium
References
Ames, O., and C. Schweinfurth. 1931. Orchidaceae. In Botanical Results of the Tyler-Duida expedition, by A. Gleason. Bull.
Torrey Bot. Club 68: 345-353.
Anonymous. 1887. Catasetum bungerothii. Gard. Chron. ser. 3, 1: 139-140.
Camevali, G, I.M. Ramirez-Morillo, G.A. Romero-Gonzalez, C.A. Vargas, E. Foldats. 2003. Orchidaceae. In J.A. Steyermark,
RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 7: 200-619.
Christenson, E.A. 1997. Orchidaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds.
1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and Monocotyledons.
Memoirs of the New York Botanical Garden 76, pt. 1 : 286-342.
Cremers, G, M. Hoff, D. Barthelemy, E.A. Christenson, Ly. & L. Leneveu, & Ch. &. CI. Pawilowski. 1992. Inventaire taxonomique
des plantes de la Guyane fran9aise II - les Orchidacees. Mus. Nat. Hist. Nat., Coll. Patr. Nat. (Ser. Patr. Genetique) 7: 1-144.
Dunsterville, G.C.K., and L.A. Garay. 1959-1976. Venezuelan Orchids Illustrated. 6 volumes. London: Andre Deutsch.
Dunsterville, G.C.K., and L.A. Garay. 1979. Venezuelan Orchids: An Illustrated Field Guide. 3 volumes. Cambridge: Harvard
University Press.
Dunsterville, G.C.K., and E. Dunsterville 1982. Auyan-tepui: Reminiscences of an orchid search. In Orchid Botany: Reviews
and Perspectives, edited by J. Arditti, 19-38. Ithaca: Cornell University Press.
Foldats, E. 1 969-1 970. Orchidaceae. In Flora de Venezuela, edited by T. Lasser, volume 1 5, parts 1-5. Caracas: Instituto Botanico.
Gleason, H.A. 1931. Botanical results of the Tyler-Duida expedition. Bull. Torrey Bot. Club 58: 277-506.
Im Thum, E.F. 1884a. Cattleya lawrenceana. Gard. Chron. n.s. 25: 168.
Im Thum, E.F. 1884b. New plants from Guiana. Timehri 3: 156-158.
Im Thum, E.F. 1885a. The first ascent of Roraima. Timehri 4: 1-48.
Im Thum, E.F. 1885b. Roraima. Timehri 4: 255-267.
Im Thum, E.F. 1886. Notes on the plants observed during the Roraima expedition of 1884. Timehri 5: 147-223.
Jirak, I.L. 1968. Angel Falls Expedition 1968. Mimeo.
Kromer, E. 191 1-1912. An orchid collector's travels through British Guiana to Brazil. Orchid World 2: 55-57, 88-90, 107-1 10,
136-139, 175-177.
Linden, L., A. Cogniaux, and G. Grignan. 1894. Les Orchidees Exotiques. Gand: Eug. Vander Haeghen.
Reichenbach, H.G 1885. Cattleya lawrenceana. Gard. Chron. n.s. 23: 338.
Renz, J. 1992. The genus Habenaria in the Guianas. Candollea 47: 483-512.
Rolfe, R.A. 1889. Catasetum bungerothii var. aureum. Gard. Chron. ser. 3, 6: 466.
Rolfe, R.A. 1891. Peristeria aspersa. Lindenia 6: 57-58, t. 267.
Romero-Gonzalez, G.A., and G. Camevali Fernandez-Concha. 2000. Orchids of Venezuela: An Illustrated Field Guide. Second
edition. Caracas, Venezuela: Armitano Editores.
Sander, F. 1888. Reichenbachia. Volume 1. London: H. Sotheran.
Schomburgk, R.H. 1931. Travels in Guiana and on the Orinoco. Translated by W.E. Roth. Georgetown, Guyana: Argosy.
Schweinfurth, C. 1967. Orchidaceae of the Guayana Highland. Mem. New York Bot. Gard. 14(3): 69-214.
Seidel, E. 1890. Voyage aux montagnes de Roraima. Orchidophile 10: 5-7, 76-77.
Szlachetko, D.L. & Y. Veyret. 1994. Two new species of Spiranthinae (Orchidaceae) from French Guyana. Fragm. Flor. Geobot.
39(2): 471-474.
Ule, E. 1914. Die vegetation des Roraima. Bot. Jahrb. Syst. 52(Beibl. 115): 42-53.
Veitch, J.H. 1906. Hortus Veitchii. London: Gilbert & Rivington.
Werkhoven, M.C.M. 1986. Orchids of Suriname. Uitgeversmaatschappij: Paramaribo, Suriname.
POACEAE
G. Davidse, E.J. Judziewicz, F.O. Zuloaga
Acroceras excavatum (Henrard) Zuloaga & Morrone BO
Acroceras macrum Stapf [naturalized]
Acroceras zizanioides (Kunth) Dandy [naturalized]
Aegopogon cenchroides Humb. & Bonpl. ex Willd. [Mt. Roraima]
Andropogon angustatus (J. Presl) Steud.
Andropogon bicornis L.
Andropogon bicornis var. bicornis
Andropogon bicornis var. burchellii Hack.
Andropogon carinatus Nees
Andropogon diuturnus Sohns
Andropogon fastigiatus Sw.
Andropogon gayanus Kunth [cultivated, naturalized] SU
GU SU FG?
AM BO
BO
DA GU
GU
SU
SU
FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM
AM
AM BO
AM BO GU
150 Checklist of the Plants of the Guiana Shield
Andropogon glomeratus (Walter) E. Britton, Steams & Poggenb.
Andropogon hypogynus Hack.
Andropogon indetonsus Sohns
Andropogon insolitus Sohns
Andropogon leucostachyus Kunth [expected in DA]
Andropogon longiramosus Sohns
Andropogon selloanus (Hack.) Hack, [expected in DA]
Andropogon vetus Sohns
Andropogon virgatus Desv. [expected in DA]
Anthephora hermaphrodita (L.) Kuntze
Arberella aff. bahiensis Soderstr. & Zuloaga
Arberellaflaccida (Doll) Soderstr. & C.E. Calderon
Arberella venezuelae Judz. & Davidse
Aristida adscensionis L.
Aristida capillacea Lam.
Aristida gibbosa (Nees) Kunth
Aristida longifolia Trin.
Aristida moritzii Henrard
Aristida recurvata Kunth
Aristida riparia Trin.
Aristida setifolia Kunth
Aristida ternipes Cav.
Aristida torta (Nees) Kunth
Arthrostylidium pubescens Rupr.
Arthrostylidium scandens McClure
Arthrostylidium schomburgkii (Benn.) Munro
Arthrostylidium sp. A
Arthrostylidium sp. B
Arthrostylidium sp. C
Arthrostylidium sp. D
Arthrostylidium venezuelae (Steud.) McClure
Arundinella hispida (Humb. & Bonpl. ex Willd.) Kuntze
Arundo donax L. [cuhivated and naturalized]
Atractantha amazonica Judz. & L.G. Clark
Aulonemia chimantaensis Judz. & Davidse
Aulonemia deflexa (N.E. Br.) McClure
Aulonemiajauaensis Judz. & Davidse
Aulonemia nitida Judz.
Aulonemia sp. A
Aulonemia sp. B
Aulonemia sp. C
Aulonemia aff. subpectinata (Kuntze) McClure
Axonopus anceps (Mez) Hitchc.
Axonopus arundinaceus G.A. Black
Axonopus aureus P. Beauv.
Axonopus canescens (Nees ex Trin.) Pilg.
Axonopus capillaris (Lam.) Chase
Axonopus casiquiarensis Davidse
Axonopus casiquiarensis var. A
Axonopus caulescens (Mez) Henrard
Axonopus chimantensis Davidse
Axonopus chrysoblepharis (Lag.) Chase [expected in BO]
Axonopus chrysostachyus (Schrad.) Pilg.
Axonopus compressus (Sw.) P. Beauv.
Axonopus cuatrecasasii G.A. Black
Axonopus eminens (Nees) G.A. Black
Axonopus equitans Hitchc. & Chase
Axonopus excavatus (Nees ex Trin.) Henrard
Axonopus fissifolius (Raddi) Kuhlm.
Axonopus flabelliformis Swallen
Axonopus gracilis G.A. Black
Axonopus iridifolius (Poepp.) G.A. Black
Axonopus leptostachyus (Flugge) Hitchc.
Axonopus longispicus (Doll) Kuhlm.
GU?
AM BO
BO
BO
AM BO GU SU FG
AM
AM BO GU su FG
AM BO
AM BO
BO
GU SU FG
FG
AM
su FG
AM su FG
AM BO GU?
AM BO
BO
GU
GU
su FG
AM BO
BO
GU su FG
AM BO DA GU su
AM BO DA GU su FG
BO GU su
BO
AM BO
BO
GU su FG
AM BO GU su
AM
AM
AM
BO
AM
AM BO GU
AM BO GU su
su
FG
FG?
AM
BO
BO
BO
GU
GU
AM
AM
AM
AM BO
AM BO DA GU su FG
AM
AM BO GU su FG
AM BO GU su FG
AM BO GU su FG
AM
AM
BO
BO
GU
su
FG
AM BO DA GU su FG
AM
BO GU su
AM GU su FG
AM BO FG
AM BO GU su FG
AM BO GU su FG
AM
AM BO GU
AM BO GU su FG
AM BO GU su FG
Checklist of the Plants of the Guiana Shield 1 5
1
Axonopus magallanesiae Giraldo-Canas BO
Axonopus oiapocensis G.A. Black
Axonopus passourae G.A. Black
Axonopus pennellii G.A. Black
Axonopus pubivaginatus Henrard
Axonopus purpusii (Mez) Chase
Axonopus ramosus Swallen [expected in GU and adjacent Brazil]
Axonopus schultesii G.A. Black
Axonopus senescens (Doll) Henrard
Axonopus sp. A
Axonopus sp. B
Axonopus sp. C
Axonopus steyermarkii Swallen
Axonopus suffultiformis GA. Black
Axonopus suffultiformis var. A
Axonopus surinamensis (Hochst. ex Steud.) Henrard
Axonopus triglochinoides (Mez) Dedecca
Axonopus villosus Swallen
Axonopus yutajensis GA. Black
Bothriochloa bladhii (Retz.) S.T. Blake [naturalized]
Bouteloua americana (L.) Scribn. [introduced? no recent collections]
Brachiaria arrecta (Hack, ex T. Durand & Schinz) Stent [culitvated,
naturalized?]
Brachiaria decumbens Stapf
Brachiariafasciculata (Sw.) Parodi
Brachiaria humidicola (Rendle) Schweick.
Brachiaria mollis (Sw.) Parodi
Brachiaria mutica (Forssk.) Stapf [naturalized]
Brachiaria plantaginea (Link) Hitchc.
Briza minor L.
Cenchrus brownii Roem. & Schult.
Cenchrus echinatus L.
Chloris barbata Sw.
Chloris elata Desv.
Chloris radiata (L.) Sw.
Chusquea linearis N.E. Br.
Chusquea sp. A
Coelorachis aurita (Steud.) A. Camus
Coix lacryma-jobi L. [cultivated, naturalized]
Cortaderia roraimensis (N.E. Br.) Pilg.
Cryptochloa capillata (Trin.) Soderstr.
Ctenium cirrosum (Nees) Kunth
Cymbopogon citratus (DC.) Stapf [cultivated, naturalized]
Cynodon dactylon (L.) Pers.
Dactyloctenium aegyptium (L.) Willd.
Dichanthelium aequivaginatum (Swallen) Zuloaga
Dichanthelium davidsei (Zuloaga & Morrone) Zuloaga
Dichanthelium hebotes (Trin.) Zuloaga
Dichanthelium pycnoclados (Tutin) Davidse
Dichanthelium sciurotoides (Zuloaga & Morrone) Davidse
Dichanthelium telmatum (Swallen) Zuloaga [expected in GU]
Dichanthium annulatum (Forssk.) Stapf [cultivated, naturalized]
Dichanthium aristatum (Poir.) C.E. Hubb. [cultivated, naturalized]
Digitaria ciliaris (Retz.) Koeler
Digitaria decumbens Stent [cultivated, naturalized]
Digitaria fragilis (Steud.) Luces
Digitariafuscescens (J. Presl) Henrard [naturalized]
Digitaria horizontalis Willd.
Digitaria insularis (L.) Fedde
Digitaria lanuginosa (Nees) Henrard
Digitaria laxa (Rchb. ex A. Spreng.) Parodi
Digitaria longiflora (Retz.) Pers. [naturalized?]
Digitaria nervalis Henrard
Digitaria nuda Schumach. [naturalized?]
FG
FG
BO
GU su FG
AM BO GU su FG
AM
AM
su FG
FG
AM BO
AM
BO
AM
AM BO GU
AM
BO GU su FG
AM
AM
AM
BO
GU
GU
su
BO GU su FG
AM GU su
BO GU su FG
BO GU su FG
BO GU FG
BO DA
DA
GU su FG
FG
BO
BO GU su FG
AM BO GU su FG
BO DA GU su FG
BO GU su
BO DA GU su FG
AM BO GU
AM
AM BO GU su FG
BO DA GU su FG
AM BO GU
su
FG
GU su FG
BO DA GU su FG
BO GU su FG
AM BO
BO
BO
GU su FG
BO GU su
BO GU FG
BO
GU
GU
AM DA GU
GU
GU
su
su
FG
AM BO DA GU su FG
AM BO DA GU su FG
BO GU
GU
su FG
FG
GU su FG
GU su
AM
AM BO GU su FG
GU su
su
FG
BO GU
BO su FG
BO GU su FG
BO DA GU su FG
BO DA GU su FG
DA GU su FG
FG
BO
BO GU su FG
BO DA GU su FG
BO GU su FG
BO DA GU su FG
BO GU su FG
BO
BO DA GU su FG
BO DA
BO
BO GU FG
BO DA GU su FG
BO DA GU
BO GU su FG
BO su FG
BO su FG
BO GU
GU
FG
BO GU
152 Checklist of the Plants of the Guiana Shield
Digitaria setigera Roth ex Roem. & Schult. [naturalized]
Digitaria swazilandemis Stent [cultivated, naturalized]
Digitaria tenuis (Nees) Henrard
Digitaria veneziielae Henrard
Digitaria violascens Link [cultivated, naturalized]
Echinochloa colona (L.) Link
Echinochloa crus-pavonis (Kunth) Schult.
Echinochloa polystachya (Kunth) Hitchc.
Echinochloa pyramidalis (Lam.) Hitchc. & Chase [cultivated, naturalized]
Echinolaena gracilis Swallen
Echinolaena inflexa (Poir.) Chase
Eleusine indica (L.) Gaertn. [naturalized]
Elionurus muticus (Spreng.) Kuntze
Enteropogon mollis (Nees) Clayton
Eragrostis acutiflora (Kunth) Nees
Eragrostis airoides Nees [expected in GU]
Eragrostis amabilis (L.) Wight & Am. ex Nees [naturalized]
Eragrostis atrovirens (Desf.) Trin. ex Steud.
Eragrostis ciliaris (L.) R. Br. [naturalized]
Eragrostis ciliaris var. ciliaris
Eragrostis gangetica (Roxb.) Steud.
Eragrostis guianensis Hitchc.
Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb.
Eragrostis japonica (Thunb.) Trin. [naturalized]
Eragrostis maypurensis (Kunth) Steud.
Eragrostis pectinacea (Michx.) Nees
Eragrostis pectinacea var. pectinacea
Eragrostis pilosa (L.) P. Beauv. [naturalized]
Eragrostis plurinodis Swallen ex Luces
Eragrostis polytricha Nees
Eragrostis refracta (Muhl. ex Elliott) Scribn. SU
Eragrostis riobrancensis Judz. & P.M. Peterson GU
Eragrostis scaligera Salzm. ex Steud. FG
Eragrostis secundiflora J. Presl [expected in SU]
Eragrostis tephrosanthos Schult.
Eragrostis imioloides (Retz.) Nees ex Steud. [naturalized]
Eragrostis viscosa (Retz.) Trin.
Eriochloa distachya Kunth
Eriochloa polystachya Kunth
Eriochloa procera (Retz.) C.E. Hubb. [naturalized]
Eriochloa punctata (L.) Desv. ex Ham.
Eriochrysis cayennensis P. Beauv.
Eriochrysis aff. holcoides (Nees) Kuhlm.
Froesiochloa boutelouoides G.A. Black
Gouinia latifolia (Griseb.) Vasey
Guadua angustifolia Kunth
Guadua ciliata Londoiio & Davidse
Guaduafascicularis Doll
Guadua aff. glomerata Munro
Guadua latifolia (Bonpl.) Kunth
Guadua macrostachya Rupr. [expected in SU]
Guadua paniculata Munro [expected in GU]
Guadua superba Huber
Guadua venezuelae Munro
Guadua weberbaueri Pilg.
Gymnopogon fastigiatus Nees
Gymnopogon foliosus (Willd.) Nees
Gymnopogon spicatus (Spreng.) Kuntze
Gynerium sagittatum (Aubl.) P. Beauv.
Hackelochloa granularis (L.) Kuntze [naturalized]
Hemarthria altissima (Poir.) Stapf& C.E. Hubb. [naturalized]
Heteropogon contortus (L.) P. Beauv. ex Roem. & Schult.
Heteropogon melanocarpus (Elliott) Elliott ex Benth. [expected in GU]
Homolepis aturensis (Kunth) Chase [naturalized?]
GU
GU SU FG
BO GU su FG
BO
AM BO
GU su FG
GU su
AM DA GU su FG
AM BO GU su FG
AM BO
BO
FG
AM GU
AM
BO
BO DA GU FG
AM BO
BO DA
GU
GU
su
su
FG
FG
AM GU
AM BO
AM BO
BO
GU
GU
su FG
AM BO DA GU su FG
AM BO
BO
GU
GU
GU
su FG
FG
FG
AM BO GU su FG
BO GU FG?
AM BO GU SU FG
BO DA GU SU FG
BO DA GU SU FG
BO GU SU
AM BO GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU
GU SU FG
Checklist of the Plants of the Guiana Shield 153
Homolepis glutinosa (Sw.) Zuloaga & Soderstr.
Homolepis isocalycia (G. Mey.) Chase
Hymenachne amplexicaulis (Rudge) Nees
Hymenachne donacifolia (Raddi) Chase
Hyparrhenia bracteata (Willd.) Stapf
Hyparrhenia nifa (Nees) Stapf [cultivated, naturalized]
Ichnanthus breviscrobs Doll
Ichnartthus calvescens (Nees) Doll
Ichnanthus dasycoleus Tutin
Ichnanthus ephemeroblepharis GA. Black & Froes ex GA. Black & Pires AM BO
Ichnanthus hojfmannseggii (Roem. & Schult.) Doll FG
Ichnanthus lancifolius Mez
Ichnanthus lancifolius var. weberbaueri (Mez) Stieber
Ichnanthus leiocarpus (Spreng.) Kunth
Ichnanthus nemoralis (Schrad. ex Schult.) Hitchc. & Chase
Ichnanthus nemorosus (Sw.) Doll [Sierra de la Neblina]
Ichnanthus pallens (Sw.) Munro ex Benth.
Ichnanthus panicoides R Beauv.
Ichnanthus procurrens (Nees exTrin.) Swallen
Ichnanthus ruprechtii Doll
Ichnanthus tenuis (J. Presl) Hitchc. & Chase
Imperata brasiliensis Trin.
Imperata contracta (Kunth) Hitchc.
Isachne polygonoides (Lam.) Doll
Isachne rigens (Sw.) Trin.
Ischaemum arenosum Sohns
Ischaemum guianense Kunth ex Hack.
Ischaemum indicum (Houtt.) Merr. [cultivated, naturalized]
Ischaemum latifolium (Spreng.) Kunth
Ischaemum rugosum Salisb. [naturalized]
Ischaemum timorense Kunth [cultivated, naturalized]
Ixophorus unisetus (J. Presl) Schltdl. [cultivated but expected in GU]
Lasiacis anomala Hitchc.
Lasiacis ligulata Hitchc. & Chase
Lasiacis nigra Davidse
Lasiacis procerrima (Hack.) Hitchc.
Lasiacis ruscifolia (Kunth) Hitchc.
Lasiacis scabrior Hitchc.
Lasiacis sloanei (Griseb.) Hitchc.
Lasiacis sorghoidea (Desv. ex Ham.) Hitchc. & Chase
Leersia hexandra Sw.
Leersia ligularis Trin. [expected in GU]
Leptochloafdiformis (Lam.) P. Beauv.
Leptochloa scabra Nees
Leptochloa virgata (L.) P. Beauv.
Leptocoryphium lanatum (Kunth) Nees
Lithachne pauciflora (Sw.) P. Beauv. [expected in GU]
Luziola bahiensis (Steud.) Hitchc.
Luziola doelliana Prodoehl
Luziola subintegra Swallen
Melinis minutiflora P. Beauv. [cultivated, naturalized]
Merostachys maguireorum McClure
Merostachys aff. retrorsa McClure
Mesosetum cayennense Steud.
Mesosetum chaseae Luces
Mesosetum chlorostachyum (Doll) Chase
Mesosetum filifolium Hubb.
Mesosetum loliiforme (Hochst. ex Steud.) Chase
Mesosetum rottboellioides (Kunth) Hitchc.
Microchloa indica (L. f.) P. Beauv. [introduced species expected in GU]
Myriocladus cardonae Swallen
Myhocladus chwunensis Swallen
Myriocladus distantiflorus Swallen [other species of this genus expected as
well]
AM
AM
GU SU FG
BO DA GU SU FG
AM
AM BO DA GU SU FG
AM BO
BO
DA GU SU FG
BO GU SU FG
AM BO GU SU FG
AM BO GU SU FG
BO GU SU FG
AM BO GU SU FG
AM BO
BO
GU
AM BO GU
GU
SU
SU
FG
AM
AM GU SU FG
GU SU FG
AM BO DA GU SU FG
AM BO
BO
DA GU
GU
SU FG
AM BO GU
AM GU
AM
BO
AM BO GU SU FG
BO DA GU SU FG
BO
BO GU
AM DA GU SU FG
AM BO DA GU SU FG
AM BO GU SU FG
BO SU FG
BO GU SU FG
BO
AM DA GU SU FG
AM BO
BO
GU SU
AM BO GU
BO GU SU FG
BO GU
AM
AM BO
AM BO GU SU FG
AM BO
BO
BO
BO
GU
GU
SU FG
154 Checklist of the Plants of the Guiana Shield
Myriocladus exsertus Swallen
Myriocladus grandifolius Swallen
Myriocladus invohttus Judz. & Davidse
Myriocladus longiramosus Swallen
Myriocladus neblinaensis Swallen
Myriocladus paludicolus Swallen [Sierra de la Neblina]
Myriocladus simplex Swallen
Myriocladus sp. A
Myriocladus steyermarkii Swallen
Myriocladus virgatus Swallen
Neurolepis angusta Swallen
Neurolepis diversiglumis Soderstr. [Sierra de la Neblina]
Neurolepis glomerata Swallen
Neurolepis pittieri McClure
Neurolepis sp. A
Olyra caudata Trin.
Olyra ciliatifolia Raddi
Olyra ecaudata Doll
Olyra latifolia L.
Olyra longifolia Kunth
Olyra obliquifolia Steud.
Olyra standleyi Hitchc.
Olyra wurdackii Swallen
Oplismenus burmannii (Retz.) R Beauv.
Oplismenus hirtellus (L.) R Beauv.
Orthoclada laxa (Rich.) R Beauv.
Oryza grandiglumis (Doll) Prodoehl
Oryza latifolia Desv.
Oryza rufipogon Griff, [naturalized]
Oryza sativa L. [cultivated, occasionally escaping]
Otachyrium grandiflorum Send. & Soderstr.
Otachyrium succisum (Swallen) Send. & Soderstr.
Otachyrium versicolor (Doll) Henrard
Panicum altum Hitchc. & Chase
Panicum arctum Swallen
Panicum bergii Arechav.
Panicum caricoides Nees
Panicum cayennense Lam.
Panicum cervicatum Chase [expected in GU]
Panicum chnoodes Trin.
Panicum cordovense E. Foum.
Panicum cyanescens Nees ex Trin.
Panicum deciduum Swallen
Panicum dichotomiflorum Michx.
Panicum discrepans Doll
Panicum elephantipes Nees ex Trin.
Panicum eligulatum N.E. Br.
Panicumfontanale Swallen
Panicum fonticolum Swallen
Panicum grande Hitchc. & Chase
Panicum granuliferum Kunth
Panicum haenkeanum J. Presl
Panicum hirsutum Sw.
Panicum hirtum Lam.
Panicum hispidifolium Swallen
Panicum hylaeicum Mez
Panicum ichunense Swallen
Panicumjauanum Davidse
Panicum ligulare Nees ex Trin.
Panicum maximum Jacq. [naturalized]
Panicum mertensii Roth
Panicum micranthum Kunth
Panicum miliaceum L. [cultivated; naturalized?]
Panicum millegrana Poir.
BO
AM
AM
AlVf
BO
/\iVl
AM
AM BO
BO
AM BO
BO GU
AM
AM
BO
BO
BO
BO GU su FG
BO GU FG
AM BO GU su FG
AM BO DA GU su FG
AM BO DA GU su
su
FG
FG
AM
AM
BO
AM BO DA GU su FG
AM BO DA GU su FG
FG
AM BO DA GU su FG
BO DA GU su FG
AM BO DA
AM
GU su FG
AM BO GU
AM DA GU su FG
BO GU
GU
su FG
AM BO GU su FG
AM DA GU su FG
AM BO
AM BO GU
BO GU su FG
AM BO GU su FG
AM
DA GU su FG
AM BO GU su FG
AM BO DA GU su FG
AM BO
BO
GU
AM
BO DA GU su FG
AM BO
BO
DA
GU
GU
FG
AM BO
BO
GU su FG
AM
BO
BO
BO
GU su FG
AM BO DA GU su FG
AM DA GU su FG
AM BO GU
GU
su FG
FG
BO GU su FG
BO
AM
AM BO
BO
AM BO
AM
AM BO
AM BO
AM BO
AM
AM BO
BO
AM BO
AM
BO
AM BO
AM BO
BO
AM BO
AM
BO
AM
AM
BO
GU su FG
GU su FG
GU
FG
GU su FG
GU su FG
GU su FG
GU FG
GU su FG
GU su FG
GU su FG
GU su FG
GU su FG
GU
GU su FG
GU su FG
GU su
su
FG
GU su FG
Checklist of the Plants of the Guiana Shield 155
Panicum nervosum Lam.
Panicum olyroides Kunth
Panicum orinocanum Luces
Panicum pandum Swallen
Panicum pantrichum Hack, [expected in GU]
Panicum parvifolium Lam. DA
Panicum petrense Swallen
Panicum pilosum Sw. DA
Panicum polycomum Trin.
Panicum polygonatum Schrad.
Panicum pulchellum Raddi
Panicum pyrularium Hitchc. & Chase
Panicum rivale Swallen
Panicum rudgei Roem. & Schult.
Panicum scabridum Doll DA
Panicum schwackeanum Mez
Panicum sellowii Nees
Panicum sipapoense Swallen
Panicum stenodes Griseb.
Panicum steyermarkii Swallen
Panicum stoloniferum Poir. DA
Panicum tepuianum Davidse & Zuloaga
Panicum trichanthum Nees
Panicum trichocladum Hack, ex K. Schum. [naturalized]
Panicum trichoides Sw.
Panicum tricholaenoides Steud.
Panicum yavitaense Swallen
Pappophorum mucronulatum Nees
Pappophorum pappiferum (Lam.) Kuntze [naturalized]
Paratheria prostrata Griseb.
Pariana campestris Aubl.
Pariana radiciflora Sagot ex Doll
Parodiolyra lateralis (J. Presl ex Nees) Soderstr. & Zuloaga
Parodiolyra luetzelburgii (Pilg.) Soderstr. & Zuloaga
Parodiolyra micrantha (Kunth) Davidse & Zuloaga
Paspalidium geminatum (Forssk.) Stapf
Paspalum albidulum Henrard
Paspalum altsonii Chase
Paspalum ammodes Trin.
Paspalum apiculatum Doll
Paspalum arenarium Schrad.
Paspalum arundinaceum Poir.
Paspalum aspidiotes Trin.
Paspalum atabapense Davidse & Zuloaga
Paspalum bifidifolium Soderstr.
Paspalum boscianum Fliigge
Paspalum carinatum Humb. & Bonpl. ex Fliigge
Paspalum chaffanjonii Maury
Paspalum cinerascens (Doll) A.G. Burm. & M.N.C. Bastos
Paspalum clavuliferum C. Wright
Paspalum conjugatum P.J. Bergius
Paspalum conspersum Schrad.
Paspalum convexum Humb. & Bonpl. ex Flugge
Paspalum corcovadense Raddi
Paspalum coryphaeum Trin.
Paspalum decumbens Sw.
Paspalum delicatum Swallen
Paspalum densum Poir.
Paspalum dilatatum Poir. [cultivated, naturalized]
Paspalum distichum L.
Paspalum exaltatum J. Presl
Paspalum fasciculatum Willd. ex Flugge [cultivated; naturalized]
Paspalumfimbriatum Kunth [introduced, no recent collections]
Paspalum gardnerianum Nees
GU
AM GU FG
GU? SU FG
AM GU SU FG
AM BO GU su
AM BO GU su FG
AM BO GU su FG
BO GU su
su
FG
AM BO
BO
BO
GU
GU
su
BO GU su FG
FG
AM BO GU
AM
GU
GU su FG
AM BO GU su
BO
su FG
BO GU
AM BO DA GU su FG
AM GU su FG
AM BO GU su FG
BO GU
AM BO GU su FG
AM BO DA GU su FG
AM FG
AM BO GU
GU
GU
GU
su
su
su
FG
FG
AM DA
GU?
su FG
AM BO DA GU su FG
GU su FG
GU su FG
156 Checklist of the Plants of the Guiana Shield
Paspalum goeldii Swallen
Paspahim hyalinum Nees ex Trin. AM
Paspalum intermedium Munro ex Morong & Britton
Paspalum lacustre Swallen [records erroneous, but perhaps expected]
Paspalum lanciflorum Trin. AM BO GU SU
Paspalum laxum Lam. FG
Paspalum lineare Trin.
Paspalum loefgrenii Ekman
Paspalum maculosum Trin.
Paspalum maritimum Trin.
Paspalum melanospermum Desv. ex Poir.
Paspalum millegrana Schrad.
Paspalum morichalense Davidse, Zuloaga & Filg. [specimens previously
identified as Paspalum lacustre']
Paspalum multicaule Poir.
Paspalum multinervium A.G. Burm.
Paspalum notatum Flugge
Paspalum nudatum Luces
Paspalum nutans Lam.
Paspalum oligostachyum Salzm. ex Steud.
Paspalum orbiculatum Poir.
Paspalum paniculatum L.
Paspalum parviflorum Rhode ex Fliigge
Paspalum pectinatum Nees ex Trin.
Paspalum petilum Chase
Paspalum pictum Ekman
Paspalum pilosum Lam.
Paspalum plicatulum Michx.
Paspalum pulchellum Kunth
Paspalum pumilum Nees
Paspalum repens P.J. Bergius
Paspalum riparium Nees
Paspalum saccharoides Nees ex Trin.
Paspalum scrobiculatum L. [cultivated; naturalized?]
Paspalum serpentinum Hochst. ex Steud.
Paspalum stellatum Humb. & Bonpl. ex Flugge [expected in GU] BO
Paspalum subciliatum Chase BO GU FG
Paspalum subfalcatum (Doll) Tutin AM
Paspalum tillettii Davidse & Zuloaga AM
Paspalum trinii Swallen
Paspalum vaginatum Sw.
Paspalum virgatum L. AM BO
Paspalum wrightii Hitchc. & Chase
Pennisetum ciliare (L.) Link [cultivated; naturalized?]
Pennisetum polystachion (L.) Schult.
Pennisetum purpureum Schumach. [cultivated, naturalized]
Pharus lappulaceus Aubl.
Pharus latifolius L.
Pharus mezii Prodoehl
Pharus parvifolius Nash
Pharus parvifolius subsp. elongatus Judz.
Pharus parvifolius subsp. parvifolius
Pharus virescens Doll AM BO
Phragmites australis (Cav.) Trin. ex Steud. BO
Piresia goeldii Swallen
Piresia macrophylla Soderstr.
Piresia sympodica (Doll) Swallen BO
Pseudechinolaena polystachya (Kunth) Stapf
Raddia guianensis (Brongn.) Hitchc.
Raddiella esenbeckii (Steud.) C.E. Calderon & Soderstr. AM BO
Raddiella aff kaieteurana Soderstr. BO
Raddiella aff. potaroensis Soderstr. BO
Rehia nervata (Swallen) Fijten
Reimarochloa aberrans (Doll) Chase
GU
GU
AM BO GU SU
GU su FG
AM BO DA GU su FG
AM BO DA GU su FG
AM BO su
AM BO GU su FG
FG
GU su FG
BO GU su
BO DA GU su
su
FG
FG
AM BO DA GU su FG
BO DA GU su FG
AM BO GU su FG
AM BO GU su FG
AM BO GU
AM GU
AM BO GU su FG
AM BO GU su FG
AM BO DA GU su FG
GU su FG
GU su FG
GU su FG
BO
su FG
GU su FG
BO
BO
BO
GU
DA GU SU FG
DA GU
GU
GU
SU FG
GU su FG
GU FG
GU SU FG
DA GU su FG
DA GU
GU
su FG
DA GU su FG
GU su FG
GU su FG
su FG
FG
GU su FG
GU
su FG
GU su FG
GU su
GU
GU su FG
FG
Checklist of the Plants of the Guiana Shield 157
Reimarochloa acuta (Flugge) Hitchc.
Rhipidocladum aff. maxonii (Hitchc.) McClure
Rhipidocladum aff. racemiflorum (Steud.) McClure
Rhipidocladum sibilans Davidse, Judz. & L.G. Clark [Mt. Roraima]
Rhipidocladum sp. A
Rhipidocladum sp. B
Rhynchelytrum repens (Willd.) C.E. Hubb. [naturalized]
Rhytachne gonzalezii Davidse
Rhytachne guianensis (Hitchc.) Clayton
Rhytachne rottboellioides Desv. ex Ham.
Rhytachne subgibbosa (Winkl. ex Hack.) Clayton
Rottboellia cochinchinensis (Lour.) Clayton [naturalized]
Saccharum asperum (Nees) Steud. [expected in GU]
Saccharum trinii (Hack.) Renvoize
Sacciolepis angustissima (Hochst. ex Steud.) Kuhkn.
Sacciolepis indica (L.) Chase [naturalized?]
Sacciolepis myuros (Lam.) Chase
Sacciolepis otachyrioides Judz.
Sacciolepis striata (L.) Nash
Sacciolepis vilvoides (Trin.) Chase
Schizachyrium brevifolium (Sw.) Nees ex Btise
Schizachyrium condensatiim (Kunth) Nees
Schizachyrium maclaudii (Jacq.-Fel.) S.T. Blake
Schizachyrium sanguineum (Retz.) Alston
Schizachyrium tenerum Nees
Setaria barbata (Lam.) Kunth [naturalized]
Setaria italica (L.) R Beauv. [naturalized?]
Setaria macrostachya Kunth
Setaria magna Griseb.
Setaria palmifolia (J. Konig) Stapf [cultivated, native to tropical S. Asia]
Setaria parviflora (Poir.) Kerguelen
Setaria setosa (Sw.) R Beauv. [naturalized?]
Setaria sulcata Raddi
Setaria tenacissima Schrad. ex Schult.
Setaria tenax (Rich.) Desv.
Setaria vulpiseta (Lam.) Roem. & Schult.
Sorghastrum setosum (Griseb.) Hitchc.
Sorghastrum stipoides (Kunth) Nash
Sorghum bicolor (L.) Moench [cultivated, sparingly naturalized]
Sorghum halepense (L.) Pers. [cultivated, naturalized]
Spartina alterniflora Loisel.
Spheneria kegelii (Miill. Berol.) Pilg.
Sporobolus ciliatus J. Presl
Sporobolus cubensis Hitchc.
Sporobolus jacquemontii Kunth
Sporobolus pyramidatus (Lam.) Hitchc.
Sporobolus temomairemensis Judz. & P.M. Peterson
Sporobolus tenuissimus (Mart, ex Schrank) Kuntze
Sporobolus virginicus (L.) Kunth
Steinchisma decipiens (Nees ex Trin.) W.V. Br.
Steinchisma laxa (Sw.) Zuloaga
Steinchisma stenophylla (Hack.) Zuloaga & Morrone
Steirachne barbata (Trin.) Renvoize
Stenotaphrum secundatum (Walter) Kuntze
Steyermarkochloa angustifolia (Spreng.) Judz.
Streptochaeta spicata Schrad. ex Nees [expected in GU]
Streptogyna americana C.E. Hubb.
Streptostachys asperifolia Desv.
Thrasya achlysophila Soderstr.
Thrasya axillaris (Swallen) A.G. Burm. ex Judz. [expected in GU]
Thrasya granitica A.G. Burm.
Thrasya hitchcockii Chase FG
Thrasya paspaloides Kunth AM BO
Thrasya petrosa {Ix'm.) ChdiSQ AM BO GU SU
GU SU FG
GU
AM BO
BO
GU? SU
AM BO
BO
AM BO DA GU
SU
AM BO
BO
GU SU
AM BO
BO
SU FG
BO GU SU
AM BO GU
GU
SU FG
AM BO GU SU FG
AM GU
DA GU SU FG
DA GU SU FG
AM BO GU SU? FG
AM BO GU SU FG
GU SU FG
AM BO GU SU FG
AM BO
BO
GU
GU
SU
SU
FG
FG
FG
AM BO DA GU SU FG
FG
BO GU SU FG
BO
AM BO GU SU FG
BO GU SU FG
AM BO GU FG
GU SU FG
AM GU
GU
SU FG
FG
DA GU
GU
SU
SU
FG
GU SU FG
AM BO GU SU FG
AM BO GU SU
SU
SU
FG
FG
BO GU SU FG
GU SU FG
BO
AM BO GU SU FG
AM
BO GU
SU
GU SU FG
AM
AM BO DA GU SU FG
BO GU
GU
SU FG
AM BO SU
SU
AM
AM BO
AM BO
BO
AM BO
158 Checklist of the Plants of the Guiana Shield
Thrasya robusta Hitchc. & Chase BO
Thrasya scandem (Tutin) Soderstr. ex A.G. Burm.
Thrasya striata A.G. Burm.
Thrasya trinitensis Mez
Trachypogon spicatus (L. f.) Kuntze DA
Tridens flaccidus (Doll) Parodi
Tripogon spicatus (Nees) Ekman [expected in GU]
Tripsacum andersonii J.R. Gray [introduced]
Tripsacum australe H.C. Cutler & E.S. Anderson AM
Tripsacum dactyhides (L.) L.
Urochloa reptans (L.) Stapf [naturalized]
Vetiveria zizanioides (L.) Nash [cultivated; naturalized?]
Zea luxurians (Durieu & Asch.) Bird [cultivated or escaped from cuhivation]
Zeugites sp. A AM BO
Synonyms and Misidentifications
Acicarpa sacchariflora Raddi =Digitaria insularis
Aegopogon cenchroides Kunth =Aegopogon cenchroides
Agrosticula muralis Raddi =Sporobolus tenuissimus
Agrostis brasiliensis Spreng. =Reimarochloa acuta
Agrostis compressa (Sw.) Poir. =Axonopus compressus
Agrostis littoralis Lam. =Sporobolus virginicus
Agrostis minutiflora (Trin.) Desf. ex Steud. =Sporobolus tenuissimus
Agrostis radiata L. =Chloris radiata
Agrostis villosa Rchb. ex A. Spreng. ^Sporobolus ciliatus
Aira laxa Rich. =Orthoclada laxa
Anastrophus capillaris (Lam.) Nash =Axonopus capillaris
Anastrophus platycaulis (Poir.) Nash ex Small =Axonopus compressus
Anatherum holcoides Nees =Eriochrysis holcoides
Anatherum zizanioides (L.) Hitchc. & Chase =Vetiveria zizanioides
Andropogon adustus Trin. =Elionurus muticiis
Andropogon angustifolius Kunth =Trachypogon spicatus
Andropogon barbatus L. =Chloris elata
Andropogon barbatus L., 1771 =Chloris barbata
Andropogon bracteatus Humb. & Bonpl. ex Willd. =Hyparrhenia bracteata
Andropogon brevifolius Sw. =Schizachyrium brevifolium
Andropogon canescens (Nees) Kunth =Trachypogon spicatus
Andropogon condensatus Kunth =Schizachyrium condensatum
Andropogon contortus L. =Heteropogon contortus
Andropogon crassus Sohns =Andropogon vetus
Andropogon crucianus Renvoize =Andropogon insolitus
Andropogon dactyloides Steud. =Trachypogon spicatus
Andropogon dissolutus (Nees) Steud. =Trachypogon spicatus
Andropogon hirtiflorus (Nees) Kunth =Schizachyrium sanguineum
Andropogon hispidus Humb. & Bonp. ex Willd. =Arundinella hispida
Andropogon insularis L. =Digitaria insularis
Andropogon latifolius Spreng. =Ischaemum latifolium
Andropogon leucostachyus subsp. selloanus Hack. =Andropogon selloanus
Andropogon ligularis (Nees) Kunth =Trachypogon spicatus
Andropogon macroglossus (Trin.) Steud. =Trachypogon spicatus
Andropogon microstachyum Desv. =Schizachyrium condensatum
Andropogon mollis (Nees) Kunth =Trachypogon spicatus
Andropogon montufarii Kunth =Trachypogon spicatus
Andropogon nodosus (Willemet) Nash =Dichanthium aristatum
Andropogon perdignus Sohns =Andropogon vetus
Andropogon plumosus Humb. & Bonpl. ex Willd. =Trachypogon spicatus
Andropogon polydactylon L. [nom. superfl.] =Chloris elata
Andropogon riedelii Trin. =Schizachyrium sanguineum
Andropogon secundus (J. Presl) Kunth =Trachypogon spicatus
Andropogon segetum (Trin.) Steud. =Ischaemum rugosum
Andropogon semiberbis (Nees) Kunth =Schizachyrium sanguineum
Andropogon setosus Griseb. =Sorghastrum setosum
Andropogon sorghum (L.) Brot. =Sorghum bicolor
Andropogon spicatus Steud. =Trachypogon spicatus
su FG
GU su FG
GU
GU su FG
GU
GU su FG
GU su FG
GU su FG
GU FG
GU su FG
FG
Checklist of the Plants of the Guiana Shield 159
Andropogon squarrosus sensu Pulle non L. f. =Vetiveria zizanioides
Andropogon stipoides (J. Presl) Kunth =Trachypogon spicatus
Andropogon tener (Nees) Kunth =Schizachyrium tenerum
Andropogon trichospirus Hack. =Trachypogon spicatus
Andropogon truncatus (Nees) Steud. =Trachypogon spicatus
Andropogon vestitus Steud. =Trachypogon spicatus
Anthaenantia lanata (Kunth) Benth. =Leptocoryphium lanatum
Apluda zeugites sensu Aubl. non L. =Ichnanthus pallens
Aristida adscensionis var. bromoides (Kunth) Henrard =Aristida adscensionis
Aristida adscensionis subsp. bromoides (Kunth) Henrard =Aristida adscensionis
Aristida adscensionis var. coarctata (Kunth) Kuntze =Aristida adscensionis
Aristida americana L. =Bouteloua americana
Aristida arenaria Trin. ^'Aristida setifolia
Aristida bromoides Kunth =Aristida adscensionis
Aristida coarctata Kunth =Aristida adscensionis
Aristida dispersa var. humilis (Kunth) Trin. & Rupr. =Aristida adscensionis
Aristida doelliana Henrard =Aristida setifolia
Aristida elegans Rudge =Aristida capillacea
Aristida humilis Kunth =Aristida adscensionis
Aristida implexa var. aequa Trin. & Rupr. =Aristida riparia
Aristida marginalis Ekman =Aristida gibbosa
Aristida neesiana Trin. & Rupr. =Aristida recurvata
Aristida orizabensis E. Foum. =Aristida gibbosa
Aristida pittieri Henrard =Aristida moritzii
Aristida planifolia Swallen =Aristida riparia
Aristida riedeliana Trin. & Rupr. ^Aristida recurvata
Aristida scabra (Kunth) Kunth =Aristida ternipes
Aristida setifolia var. arenaria (Trin.) Trin. & Rupr. =Aristida setifolia
Aristida setifolia var. grandiflora Trin. & Rupr. =Aristida setifolia
Aristida setifolia var. intermedia Trin. & Rupr. =Aristida setifolia
Aristida tarapotana Mez =Aristida setifolia
Aristida tenuis (Kunth) Kunth ^Aristida ternipes
Aristida tincta Trin. & Rupr. =Aristida torta
Arthrostylidium cacuminis McClure =Arthrostylidium scandens
Arthrostylidium capillifolium Griseb. [West Indian species, not in flora area] =Arthrostylidium farctum
Arthrostylidium farctum (Aubl.) Soderstr. & Lourteig [West Indian species, not in flora area]
Arthrostylidium racemiflorum Steud. =Rhipidocladum racemiflorum
Arthrostylidium steyermarkii McClure =Aulonemia deflexa
Arthrostylidium subpectinatum Kuntze =Aulonemia subpectinata
Arundafarcta Aubl. [West Indian species, not in flora area] =Arthrostylidium farctum
Arundinaria deflexa N.E. Br. =Aulonemia deflexa
Arundinaria schomburgkii Benn. =Arthrostylidium schomburgkii
Arundinella confinis (Schult.) Hitchc. & Chase =Arundinella hispida
Arundo altissima Benth. =Phragmites australis
Arundo australis Cav. =Phragmites australis
Arundo phragmites L. =Phragmites australis
Arundo roraimensis N.E. Br. =Cortaderia roraimensis
Aulonemia steyermarkii (McClure) McClure =Aulonemia deflexa
Axonopus "perpusillus Swallen" [invalid name, error for Axonopus purpurellus Swallen]
Axonopus affinis sensu Lindeman & Gorts non Chase =Axonopus purpusii
Axonopus affinis Chase =Axonopus fissifolius
Axonopus anomalus Swallen =Axonopus purpusii
Axonopus appendiculatus (J. Presl) Hitchc. & Chase =Axonopus excavatus
Axonopus ater Chase =Axonopus fissifolius
Axonopus attenuatus non (J. Presl) Hitchc, sensu Hitchc. =Axonopus flabelliformis
Axonopus aturensis Luces =Axonopus anceps
Axonopus aureus war. pilosus (Doll) Henrard =Axonopus chrysoblepharis
Axonopus canescens war. psilachne (Doll) G.A. Black =Axonopus canescens
Axonopus caracarahyensis G.A. Black & Froes =Axonopus anceps
Axonopus carinato-vaginatus (Mez) H. Scholz =Axonopus aureus
Axonopus carinato-vaginatus var. sprucei (G.A. Black) H. Scholz =Axonopus canescens
Axonopus caulescens sensu Swallen non (Mez) Henrard =Axonopus ramosus
Axonopus chrysites (Steud.) Kuhlm. =Axonopus aureus
Axonopus chrysodactylus (Trin.) Kuhlm. =Axonopus canescens
160 Checklist of the Plants of the Guiana Shield
Axonopus compressus var. affinis (Chase) M.R. Hend. =Axonopus fissifolius
Axonopus compressus var. australis G.A. Black =Axonopus compressus
Axonopus compressus var. macropodius (Steud.) G.A. Black =Axonopus compressus
Axonopus erectus Swallen =Axonopus anceps
Axonopus exasperatus (Nees ex Steud.) G.A. Black =Axonopus aureus
Axonopus extenuatus (Nees) Kuhlm. =Axonopus capillaris
Axonopus fissifolius sensu Amshoff& Henrard non (Raddi) Kuhlm. =Axonopus purpusii
Axonopus fissifolius var. coronatus G.A. Black =Axonopus fissifolius
Axonopus flabelliformis var. camporum G.A. Black =Axonopus flabelliformis
Axonopus flabelliformis var. decipiens G.A. Black =Axonopus flabelliformis
Axonopus flexilis (Mez) Henrard =Axonopus purpusii
Axonopus fockei (Mez) Henrard =Axonopus surinamensis
Axonopus fockei sensu Amshoff& Henrard non (Mez) Henrard, =Axonopus equitans
Axonopus gentilis Henrard =Axonopus eminens
Axonopus hirsutus G.A. Black =Axonopus fissifolius
Axonopus hitchcockii G.A. Black =Axonopus longispicus
Axonopus immersus (Nees) Kuhlm. =Axonopus chrysoblepharis
Axonopus kaietukensis Swallen =Axonopus flabelliformis
Axonopus laxus Luces =Axonopus capillaris
Axonopus leptostachyus non (Fliigge) Hitchc. sensu Hitchc. =Axonopus longispicus
Axonopus macrostachyus Hitchc. & Chase =Axonopus leptostachyus
Axonopus maguirei GA. Black [not in flora area]
Axonopus minutus Luces =Axonopus aureus
Axonopus multipes Swallen =Axonopus compressus
Axonopus paranaensis Parodi =Axonopus leptostachyus
Axonopus paucisetosus G.A. Black =Axonopus canescens
Axonopus piccae Giraldo-Caiias =Axonopus iridifolius
Axonopus pruinosus Henrard =Axonopus anceps
Axonopus pulcher (Nees) Kuhlm. =Axonopus canescens
Axonopus purpurellus Swallen =Axonopus flabelliformis
Axonopus rivularis G.A. Black =Axonopus longispicus
Axonopus scoparius (Fliigge) Hitchc. sensu Hitchc. =Axonopus pubivaginatus
Axonopus sprucei GA. Black =Axonopus canescens
Axonopus sprucei var. glabratus (Doll) GA. Black =Axonopus canescens
Axonopus stragulus Chase =Axonopus fissifolius
Axonopus stragulus sensu Amshoff & Henrard non Chase ^Axonopus ramosus
Axonopus tamayonis Luces =Axonopus flabelliformis
Bambusa guadua Bonpl. =Guadua angustifolia
Bambusa latifolia Bonpl. =Guadua latifolia
Bambusa macrostachya (Rupr.) McClure ^Guadua macrostachya
Bambusa surinamensis Rupr. [cultivated] =Bambusa vulgaris
Bambusa venezuelae (Munro) McClure =Guadua venezuelae
Bambusa vulgaris Schrad. ex J.C. Wendl. [cultivated]
Bothriochloa ischaemum sensu Hitchc. non (L.) Keng =Bothriochloa bladhii
Bothriochloa pertusa sensu Hitchc. non (L.) Willd. =Bothriochloa bladhii
Brachiaria purpurascens (Raddi) Henrard =Brachiaria mollis
Bromus spicatus Nees =Tripogon spicatus
Bulbulus nervatus Swallen =Rehia nervata
Cabrera chrysoblepharis Lag. =Axonopus chrysoblepharis
Capriola dactylon (L.) Kuntze =Cynodon dactylon
Caryochloa bahiensis Steud. =Luziola bahiensis
Cenchrus ciliaris L. =Pennisetum ciliare
Cenchrus dactylolepis Steud. =Cenchrus brownii
Cenchrus granulans L. =Hackelochloa granulans
Cenchrus inflexus Poir. =Echinolaena inflexa
Cenchrus marginalis Rudge ^Echinolaena inflexa
Cenchrus parviflorus Poir. ^^Setaria parviflora
Cenchrus setosus Sw. =Pennisetum polystachion
Cenchrus viridis Spreng. =Cenchrus brownii
Chaetaria adscensionis (L.) P. Beauv. =Aristida adscensionis
Chaetaria bromoides (Kunth) Roem. & Schult. =Aristida adscensionis
Chaetaria capillacea (Lam.) P. Beauv. =Aristida capillacea
Chaetaria coarctaia (Kunth) Roem. & Schult. =Aristida adscensionis
Chaetaria gibbosa Nees =Aristida gibbosa
Checklist of the Plants of the Guiana Shield 161
Chaetaha humilis (Kunth) Roem. & Schult. =Ahstida adscensionis
Chaetaria recurvata (Kunth) Roem. & Schult. =Aristida recurvata
Chaetaha setifolia (Kunth) Nees ^Aristida setifolia
Chaetaria torta Nees =Aristida torta
Chaetochloa geniculata (Lam.) Millsp. & Chase =Setaria parviflora
Chaetochloa poiretiana (Schult.) Hitchc. =Setaria poiretiana
Chaetochloa tenax (Rich.) Hitchc. =Setaria tenax
Chloris barbata (L.) Nash =Chloris elata
Chloris dandyana CD. Adams [nom. illeg.] =Chloris elata
Chlorisfoliosa Willd. =Gymnopogon foliosus
Chloris inflata Link =Chloris barbata
Chloris mollis (Nees) Swallen =Enteropogon mollis
Chloris polydactyla (L.) Sw. [nom. illeg.] =Chloris elata
Chusquea venezuelae Steud. =Arthrostylidium venezuelae
Coelorachis loricata (Trin.) Nash =Rhytachne rottboellioides
Cryptochloa concinna sensu Swallen non (Hook, f.) Swallen =Piresia goeldii
Cymbopogon bracteatus (Willd.) Hitchc. =Hyparrhenia bracteata
Cynosurus aegyptius L. =Dactyloctenium aegyptium
Cynosurus domingensis Jacq. =Leptochloa virgata
Cynosurus indicus L. =Eleusine indica
Cynosurus virgatus L. =Leptochloa virgata
Diandrochloa diplachnoides (Steud.) A.N. Henry =Eragrostisjaponica
Diandrochloa glomerata (Walter) Burkart ^^Eragrostisjaponica
Diandrochloajaponica (Thunb.) A.N. Henry ^Eragrostisjaponica
Diectomis fastigiata (Sw.) R Beauv. =Andropogon fastigiatus
Digitaria adscendens (Kunth) Henrard =Digitaria ciliaris
Digitaria argillacea sensu Swallen non (Hitchc. & Chase) Femald =Digitariafragilis
Digitaria aurea (P. Beauv.) Spreng. =Axonopus aureus
Digitaria glabriculmis Swallen =Digitaria venezuelae
Digitaria insularis Mez ex Ekman =Digitaria insularis
Digitaria malacophylla (Hitchc.) Henrard =Digitaria lanuginosa
Digitaria microbachne (J. Presl) Henrard ^Digitaria setigera
Digitaria platicaulis (Poir.) Desv. =Axonopus compressus
Digitaria sacchariflora (Raddi) Henrard =Digitaria insularis
Digitaria sanguinalis auct. non (L.) Scop. 1771 : sensu Luces =Digitaria ciliaris
Digitaria uniflora Salzm. ex Steud. =Axonopus compressus
Elionurus adustus (Trin.) Ekman ^Elionurus muticus
Elionurus planifolius Renvoize =Elionurus muticus
Eragrostis acuminata Doll =Eragrostis maypurensis
Eragrostis acutiflora var. humilior J. Presl =Eragrostis acutiflora
Eragrostis amabilis var. plumosa (Retz.) E.G. Camus & A. Camus =Eragrostis amabilis
Eragrostis amoena J. Presl =Eragrostis maypurensis
Eragrostis aturensis (Kunth) Trin. ex Steud. =Eragrostisjaponica
Eragrostis bahiensis sensu Lemee non Schrad. ex Schult. =Eragrostis scaligera
Eragrostis barbata Trin. =Steirachne barbata
Eragrostis capillaris sensu Lemee non (L.) Nees =Eragrostis pilosa
Eragrostis chariis (Schult.) Hitchc. =Eragrostis atrovirens
Eragrostis ciliaris var. patens Chapm. ex Beal =Eragrostis amabilis
Eragrostis conferta (Elliott) Trin. =Eragrostisjaponica
Eragrostis diplachnoides Steud. ^Eragrostis japonica
Eragrostis elegantula (Kunth) Nees ex Steud. ^Eragrostis gangetica
Eragrostis filiformis Link =Eragrostis pilosa
Eragrostis flamignii De Wild. =Eragrostis gangetica
Eragrostisfloridana Hitchc. =Eragrostis polytricha
Eragrostis fragilis Swallen =Eragrostis polytricha
Eragrostis glomerata (Walter) L.H. Dewey =Eragrostisjaponica
Eragrostis hapalantha Trin. =Eragrostis japonica
Eragrostis interrupta var. diplachnoides (Steud.) Sid^^i ^Eragrostis japonica
Eragrostis interrupta var. laxiflora Doll ^Eragrostis japonica
Eragrostis interrupta var. parvijlora Doll =Eragrostis japonica
Eragrostis lindeniana Steud. =Eragrostis maypurensis
Eragrostis linkii (Kunth) Steud. =Eragrostis pilosa
Eragrostis maypurensis var. densiuscula Pilg. =Eragrostis maypurensis
Eragrostis maypurensis var. meratiana (Kunth) Pilg. =Eragrostis maypurensis
162 Checklist of the Plants of the Guiana Shield
Eragrostis meratiana (Kunth) Steud. =Eragrostis maypurensis
Eragrostis mexicana auct., non (Lag.) Link =Eragrostis pilosa
Eragrostis namaquensis var. diplachnoides (Steud.) Clayton =Eragrostisjaponica
Eragrostis pallida Vasey =Eragrostisjaponica
Eragrostis panamensis J. Presl =Eragrostis maypurensis
Eragrostis plumosa (Retz.) Link =Eragrostis amabilis
Eragrostis poaeoides sensu PuUe non P. Beauv. =Eragrostis tephrosanthos
Eragrostis polytricha var. glabrior Doll =Eragrostis polytricha
Eragrostis polytricha var. hirsutior Doll =Eragrostis polytricha
Eragrostis purpusa Jedwabn. =Eragrostis polytricha
Eragrostis reptans (Michx.) Nees =Eragrostis hypnoides
Eragrostis reptans var. contractu Doll =Eragrostis hypnoides
Eragrostis reptans var. laxior Doll =Eragrostis hypnoides
Eragrostis reptans var. pygmaea Doll =Eragrostis hypnoides
Eragrostis rufescens var. laurentiana Benoist =Eragrostis scaligera
Eragrostis stenoclada J. Presl =Eragrostis gangetica
Eragrostis tenella (L.) P. Beauv. ex Roem. & Schult. =Eragrostis amabilis
Eragrostis tenella \av. japonica (Thunb.) Roem. & Schult. =Eragrostis japonica
Eragrostis tenella war. plumosa (Retz.) Stapf =Eragrostis amabilis
Eragrostis tenella var. viscosa (Retz.) Siapf =Eragrostis viscosa
Eragrostis trichocolea Hack. & Arechav. =Eragrostis polytricha
Eragrostis trichocolea \ar. floridana (Hitchc.) Witherspoon -Eragrostis polytricha
Eragrostis vahlii (Roem. & Schult.) Nees =Eragrostis maypurensis
Eragrostis vahlii var. polyantha Doll =Eragrostis maypurensis
Eragrostis viscosa var. pilosissima (Hochst. ex A. Rich.) Hochst. =Eragrostis viscosa
Eragrostis warmingii Hack. =Eragrostis viscosa
Erianthus asper Nees =Saccharum asperum
Erianthus saccharoides var. fi trinii Hack. =Saccharum trinii
Erianthus trinii (Hack.) Hack. =Saccharum trinii
Eriochloa kunthii G. Mey. =Eriochloa punctata
Eriochloa ramosa (Retz.) Kuntze =Eriochloa procera
Eriochloa subglabra (Nash) Hitchc. =Eriochloa polystachya
Erosion hypnoides (Lam.) Lunell =Eragrostis hypnoides
Euchlaena luxurians Durieu & Asch. =Zea luxurians
Festucafiliformis Lam. =Leptochloa filiformis
Guadua dioica Steud. =Guadua macrostachya
Guadua tagoara sensu Pulle non (Nees) Kunth =Guadua latifolia
Gymnopogon mollis Nees =Enteropogon mollis
Gynerium saccharoides Humb. =Gynerium sagittatum
Helopus cognatus Steud. =Eriochloa punctata
Helopus kunthii (G. Mey.) Trin. ex Steud. =Eriochloa punctata
Heteropogon secundus J. Presl =Trachypogon spicatus
Heteropogon stipoides J. Presl =Trachypogon spicatus
Heteropogon truncatus Nees =Trachypogon spicatus
Holcus bicolor L. =Sorghum bicolor
Holcus sorghum L. ^Sorghum bicolor
Holcus sorghum var. sudanensis (Piper) Hitchc. =Sorghum halepense
Holcus striatus L. =Sacciolepis striata
Homalocenchrus hexandrus (Sw.) Kuntze =Leersia hexandra
Hymenachne auriculata (Spreng.) Chase =Hymenachne donacifolia
Hypogynium virgatum (Desv.) Dandy =Andropogon virgatus
Ichnanthus acuminatus Swallen =Ichnanthus calvescens
Ichnanthus amplus Swallen =Ichnanthus calvescens
Ichnanthus angustifolius Swallen =Ichnanthus tenuis
Ichnanthus angustus Swallen =Ichnanthus ephemeroblepharis
Ichnanthus areolatus K.E. Rogers =Ichnanthus pallens
Ichnanthus axillaris (Nees) Hitchc. & Chase =Ichnanthus pallens
Ichnanthus candicans sensu Lemee non (Nees) Doll =Ichnanthus tenuis
Ichnanthus candicans var. virescens Doll =Ichnanthus tenuis
Ichnanthus chaseae Swallen =Ichnanthus calvescens
Ichnanthus confertus K.E. Rogers =Ichnanthus pallens
Ichnanthus duidensis Swallen =Ichnanthus lancifolius var. weberbaueri
Ichnanthus hispidus Swallen =Ichnanthus calvescens
Ichnanthus hitchcockii K.E. Rogers =Ichnanthus pallens
Checklist of the Plants of the Guiana Shield 163
Ichnanthus ichnodes (Griseb.) Hitchc. & Chase =Ichnanthus calvescens
Ichnanthus longifolius Swallen =Ichnanthus ephemewblepharis
Ichnanthus neblinaensis Swallen =Ichnanthus ephemeroblepharis
Ichnanthus nubilis Chase =Ichnanthus nemorosus
Ichnanthus pallidus Swallen =Ichnanthus calvescens
Ichnanthus petiolatus var. lanceolatus Doll =Ichnanthus nemoralis
Ichnanthus polythyrsus (Nees ex Steud.) K.E. Rogers =Ichnanthus tenuis
Ichnanthus riedelii sensu Hitchc. non (Trin.) Doll =Ichnanthus nemoralis
Ichnanthus sabulosus K.E. Rogers =Ichnanthus tenuis
Ichnanthus serratus Swallen =Ichnanthus ephemeroblepharis
Ichnanthus sucrensis K.E. Rogers =Ichnanthus tenuis
Ichnanthus tamayonis Chase =Ichnanthus tenuis
Ichnanthus tectus Swallen =Ichnanthus ephemeroblepharis
Ichnanthus tenuifolius K.E. Rogers =Ichnanthus nemorosus
Ichnanthus venezuelanus Mez ^Ichnanthus tenuis
Ichnanthus vestitus Swallen =Ichnanthus calvescens
Ichnanthus weberbaueri Mez =Ichnanthus lancifolius var. weberbaueri
Imperata arundinacea var. americana L. Andersson =Imperata brasiliensis
Imperata caudata (G. Mey.) Trin. =Imperata contracta
Imperataflexuosa Swallen =Imperata contracta
Isachne arundinacea auct. non (Sw.) Griseb. 1864: sensu Luces =Isachne rigens
Isachne ligulata Swallen =Isachne rigens
Ischaemopogon latifolius (Spreng.) Griseb. =Ischaemum latifolium
Ischaemum ciliare Retz. =Ischaemum indicum
Ischaemum guianense var. schomburgkii Hack. =Ischaemum guianense
Ischaemum latifolium subsp. hirtivaginum Hitchc. =Ischaemum latifolium
Ischaemum latifolium var. minus E. Foum. =Ischaemum latifolium
Ischaemum rugosum var. segetum (Trin.) Hack. =Ischaemum rugosum
Ischaemum segetum Trin. =Ischaemum rugosum
Lasiacis acuminata Swallen =Lasiacis sorghoidea
Lasiacis compacta (Sw. ex Wikstr.) Hitchc. =Lasiacis ruscifolia
Lasiacis divaricata sensu Amshoff& Henrard non (L.) Hitchc. =Lasiacis ligulata
Lasiacis glabra Swallen =Lasiacis ruscifolia
Lasiacis guaraniticum (Speg.) Parodi =Lasiacis sorghoidea
Lasiacis liebmanniana (E. Foum.) Hitchc. =Lasiacis ruscifolia
Lasiacis patentiflora Hitchc. & Chase =Lasiacis sorghoidea
Lasiacis sorghoidea var. patentiflora (Hitchc. & Chase) Davidse =Lasiacis sorghoidea
Lasiacis swartziana (Hitchc.) Hitchc. =Lasiacis sorghoidea
Leersia distichophylla Balansa & Poitr. =Leersia ligularis
Leersia grandiflora (Doll) Prodoehl =Leersia ligularis
Leersia ligularis var. glabriflora Pyrah =Leersia ligularis
Leersia ligularis var. grandiflora (Doll) Pyrah =Leersia ligularis
Leptochloa domingensis (Jacq.) Trin. ^Leptochloa virgata
Leptochloa mutica Steud. =Leptochloa virgata
Leptochloa virgata var. mutica (Steud.) Doll =Leptochloa virgata
Leptostachys domingensis G. Mey. =Leptochloa virgata
Leptostachys filiformis G. Mey. =Leptochloafiliformis
Leptostachys virgata G Mey. =Leptochloa virgata
Luziola pittieri Luces =Luziola doelliana
Luziola spruceana auct. non Benth. ex Doll 1871: sensu Hitchc. =Luziola subintegra
Lycurus muticus Spreng. =Elionurus muticus
Manisuris aurita (Steud.) Kuntze =Coelorachis aurita
Manisuris granularis (L.) Naezen =Hackelochloa granularis
Manisuris guianensis Hitchc. =Rhytachne guianensis
Manisuris loricata (Trin.) Kuntze =Rhytachne rottboellioides
Megastachya acutiflora (Kunth) Roem. & Schult. =Eragrostis acutiflora
Megastachya breviflora E. Foum. =Eragrostis hypnoides
Megastachya corymbifera E. Foum. =Eragrostis hypnoides
Megastachya gouinii E. Foum. =Eragrostis hypnoides
Megastachya hypnoides (Lam.) P. Beau v. =Eragrostis hypnoides
Megastachya maypurensis (Kunth) Roem. & Schult. =Eragrostis maypurensis
Melinis repens (Willd.) Zizka =Rhynchelytrum repens
Meoschium rugosum (Salisb.) Nees =Ischaemum rugosum
Mesosetum cardonum Luces =Mesosetum chaseae
164 Checklist of the Plants of the Guiana Shield
Mesosetum elongatum Mez =Mesosetum loliiforme
Mesosetum tenuifolium Swallen =Mesosetum loliiforme
Milium compressum Sw. =Axonopus compressus
Milium punctatiim L. =Eriochloa punctata
Mnesithea aurita (Steud.) de Koning & Sosef =Coelorachis aurita
Mnesithea subgibbosa (Winkl. ex Hack.) de Koning & Sosef =Rhytachne subgibbosa
Myriocladus affinis Swallen =Myriocladus longiramosus
Myriocladus confertus Swallen =Myriocladus distantiflorus
Myriocladus gracilis Swallen =Myriocladus steyermarkii
Myriocladus maguirei Swallen =Myriocladus virgatus
Myriocladus paraquensis Swallen =Myriocladus grandifolius
Myriocladus paruensis Swallen =Myriocladus longiramosus
Myriocladus purpureus Swallen =Myriocladus distantiflorus
Myriocladus variabilis Swallen =Myriocladus distantiflorus
Myriocladus wurdackii Swallen =Myriocladus distantiflorus
Neeragrostis hypnoides (Lam.) Bush =Eragrostis hypnoides
Neurolepis densiflora Swallen =Neurolepi^glomerata
Neurolepis nigra Swallen =Neurolepis glomerata
Olyra cordifolia Kunth =Olyra latifolia
Olyra dimidiata Hochst. ex Steud. =Olyra caudata
Olyrafasciculata auct. non Trin. 1834 =Olyra standleyi
Olyrafloribunda var. microphylla Doll =Raddia guianensis
Olyra kegelii Mez =Olyra longifolia
Olyra lateralis (J. Presl ex Nees) Chase =Parodiolyra lateralis
Olyra lateralis sensu Amshoff& Henrard non (J. Presl ex Nees) Chase =Parodiolyra luetzelburgii
Olyra latifolia var. pubescens (Raddi) Doll =Olyra latifolia
Olyra latifolia var. vestita Henrard =Olyra latifolia
Olyra luetzelburgii Pilg. =Parodiolyra luetzelburgii
Olyra micrantha Kunth =Parodiolyra micrantha
Olyra nana Doll [nom. superfl.] =Raddiella esenbeckii
Olyra ovata Desv. =Ichnanthus panicoides
Olyra pauciflora Sw. =Lithachne pauciflora
Olyra pubescens Raddi =Olyra latifolia
Olyra sarmentosa Doll =Parodiolyra lateralis
Olyra scabra Nees =Olyra latifolia
Olyra speciosa Mez =Olyra caudata
Olyra surinamensis Hochst. ex Steud. =Olyra latifolia
Olyra surinamensis auct. non Hochst. ex Steud. =Olyra longifolia
Olyra sympodica Doll =Piresia sympodica
Oplismenus crus-pavonis Kunth =Echinochloa crus-pavonis
Oplismenus polystachyus Kunth =Echinochloa polystachya
Oplismenus pseudo-undulatifolius (Roem. «fe Schult.) Kunth =Panicum stoloniferum
Oplismenus tenuis J. Presl =Ichnanthus tenuis
Oplismenus velutinus (G. Mey.) Schult. =Oplismenus hirtellus
Orthoclada rariflora (Lam.) P. Beauv. =Orthoclada laxa
Orthoclada rariflora var. lanceolata Doll =Orthoclada laxa
Orthoclada rariflora var. sesquiflora Doll =Orthoclada laxa
Oryza alta Swallen =Oryza latifolia
Oryza glumaepatula Hochst. ex Steud. =Oryza rufipogon
Oryza monandra var. grandiflora Doll =Leersia ligularis
Oryza perennis Moench [nom. confus.] =Oryza rufipogon
Panicum aequivaginatum Swallen =Dichanthelium aequivaginatum
Panicum agglutinans Kunth =Lasiacis sorghoidea
Panicum albociliatum Swallen =Dichanthelium pycnoclados
Panicum altissimum G. Mey. =Panicum mertensii
Panicum amplexicaule Rudge =Hymenachne amplexicaulis
Panicum amplifolium Steud. =Setaria vulpiseta
Panicum angulosum Swallen =Panicum rivale
Panicum angustissimum Hochst. ex Steud. =Sacciolepis angustissima
Panicum appressifolium Swallen =Dichanthelium aequivaginatum
Panicum appressum (Lam.) Doll =Paspalidium geminatum
Panicum arrectum Hack, ex T. Durand & Schinz =Brachiaria arrecta
Panicum asperifolium (Desv.) Hitchc. =Streptostachys asperifolia
Panicum aturense Kunth =Homolepis aturensis
Checklist of the Plants of the Guiana Shield 165
Panicum auyanense Swallen =Panicum eligulatum
Panicum axillare Nees =Ichnanthus pollens
Panicum axipilium Steud. =Panicum hirtum
Panicum blackii Swallen =Panicum micranthum
Panicum breviscrobs (Doll) Pilg. =Ichnanthus breviscrobs
Panicum burmannii Retz. =Oplismenus burmannii
Panicum calvescens Nees ^Ichnanthus calvescens
Panicum caricoides sensu Lemee non Nees =Pamcum stenodes
Panicum caricoides var. pilosum Doll =Panicum caricoides
Panicum caroniense Luces =Steinchisma laxa
Panicum caudatum Lam. =Setaria setosa
Panicum cayennense var. curtatum Doll =Panicum cayennense
Panicum cayennense var. divaricatum Doll =Panicum rudgei
Panicum cayennense var. patulum Doll =Panicum cayennense
Panicum chlorostachyum Doll =Mesosetum chlorostachyum
Panicum chloroticum Nees =Panicum dichotomiflorum
Panicum chrysites Steud. =Axonopus aureus
Panicum chrysoblephare (Lag.) Steud. =Axonopus chrysoblepharis
Panicum chrysodactylon Trin. =Axonopus canescens
Panicum churunense Swallen =Panicum chnoodes
Panicum ciliare Retz. =Digitaria ciliaris
Panicum coenosum Doll ^Panicum pilosum
Panicum colonum L. =Echinochloa colona
Panicum commelinaefolium Rudge =Panicum nervosum
Panicum compactum Sw. ex Wikstr. =Lasiacis ruscifolia
Panicum compositum auct., non L. =Oplismenus hirtellus
Panicum cordovense sensu Amshoff& Henrard non E. Foum. =Dichanthelium pycnoclados
Panicum cowanii Swallen =Panicum chnoodes
Panicum crus-galli var. aristatum G. Mey. =Echinochloa crus-pavonis
Panicum curvifolium Swallen ^Panicum chnoodes
Panicum cyanescens var. lamarckianum Doll ^Panicum cyanescens
Panicum dactylon L. =Cynodon dactylon
Panicum davidsei Zuloaga & Morrone =Dichanthelium davidsei
Panicum decipiens Nees ex Trin. =Steinchisma decipiens
Panicum densifolium Swallen =Panicum micranthum
Panicum distichum var. lancifolium Griseb. =Panicum pilosum
Panicum distichum var. lancifolium Griseb. ex Hitchc. =Panicum pilosum
Panicum distichum var. luxurians G. Mey. =Panicum pilosum
Panicum divaricatum var. lanatum Schltdl. & Cham. =Lasiacis sorghoidea
Panicum divaricatum var. latifolium Schltdl. & Cham. =Lasiacis sorghoidea
Panicum donacifolium Raddi =Hymenachne donacifolia
Panicum echinolaena Nees =Echinolaena inflexa
Panicum elatior Kunth =Panicum mertensii
Panicum errabundum Hitchc. =Panicum schwackeanum
Panicum esenbeckii Nees ex Steud. =Raddiella esenbeckii
Panicum exasperatum Nees ex Steud. =Axonopus aureus
Panicum excavatum Henrard =Acroceras excavatum
Panicum fasciculatum Sw. =Brachiariafasciculata
Panicum fistulosum Hochst. ex Steud. =Panicum elephantipes
Panicum frondescens G. Mey. =Panicum stoloniferum
Panicumfunckianum Steud. =Panicum olyroides
Panicum fusciflorum Steud. =Leptocoryphium lanatum
Panicumfuscum Sw. =Brachiariafasciculata
Panicum geminatum Forssk. =Paspalidium geminatum
Panicum geniculatum Willd. —Setaria parviflora
Panicum glutinosum Lam. =Lasiacis sorghoidea
Panicum glutinosum Sw. =Homolepis glutinosa
Panicum gracilissimum Swallen =Panicum polycomum
Panicum graniticum Swallen =Panicum pandum
Panicum guaraniticum Speg. =Lasiacis sorghoidea
Panicum guianense Hitchc. =Panicum hylaeicum
Panicum hebotes Trin. =Dichanthelium hebotes
Panicum hirtellum L. —Oplismenus hirtellus
Panicum hoffmannseggii Roem. & Schult. =Ichnanthus hoffmannseggii
1 66 Checklist of the Plants of the Guiana Shield
Panicum horizontale (Willd.) G. Mey. =Digitaria horizontalis
Panicum humidicolum Rendle =Brachiaria humidicola
Panicum imberbe Poir. =Setaria parviflora
Panicum immersum (Nees) Trin. =Axonopus chrysoblepharis
Panicum insulare (L.) G. Mey. =Digitaria insularis
Panicum inversum Swallen =Panicum chnoodes
Panicum isocalycium G Mey. =Homolepis isocalycia
Panicum kaietukense Tutin =Panicum parvifolium
Panicum kappleri Steud. =Panicum micranthum
Panicum kavanayense Swallen =Panicum chnoodes
Panicum lanatum Sw. =Lasiacis sorghoidea
Panicum lanatum var. sorghoideum (Desv. ex Ham.) Griseb. =Lasiacis sorghoidea
Panicum laterale C. Presl. ex Nees =Parodiolyra lateralis
Panicum latifolium Ham. =Lasiacis sloanei
Panicum laxum Sw. =Steinchisma laxa
Panicum leprieurii Steud. =Panicum stoloniferum
Panicum leptachyrium Doll =Paratheria prbstrata
Panicum liebmannianum E. Foum. =Lasiacis ruscifolia
Panicum loliaceum auct., non Lam. =Oplismenus hirtellus
Panicum loliiforme Hochst. ex Steud. =Mesosetum loliiforme
Panicum longepaniculatum Benoist =Setaria poiretiana
Panicum luticola Hitchc. =Steinchisma laxa
Panicum macrostachyum (Kunth) Doll =Setaria vulpiseta
Panicum magnum Hitchc. =Ichnanthus breviscrobs
Panicum maguirei Swallen =Panicum chnoodes
Panicum manacalense Swallen =Panicum scabridum
Panicum manacalensis Swallen =Panicum scabridum
Panicum mauryi Swallen =Panicum micranthum
Panicum megiston Schult. =Panicum mertensii
Panicum micranthum var. hirtum Maury =Panicum micranthum
Panicum milleflorum Hitchc. & Chase -Panicum pilosum
Panicum mirandum Luces =Dichanthelium hebotes
Panicum molle Sw. =Brachiaria mollis
Panicum monostachyum Kunth =Panicum pilosum
Panicum multiflorum sensu Pulle non Poir. =Panicum grande
Panicum musei Steud. =Echinochloa colona
Panicum muticum Forssk. =Brachiaria mutica
Panicum myosurus Rich. =Sacciolepis myuros
Panicum myrianthum Mez =Panicum grande
Panicum myuros Lam. =Sacciolepis myuros
Panicum nemorale Schrad. ex Schult. =Ichnanthus nemoralis
Panicum nemorosum Sw. =Ichnanthus nemorosus
Panicum numidianum sensu Pulle non Lam. =Brachiaria mutica
Panicum obovatum Doll =Panicum polycomum
Panicum pallens Sw. =Ichnanthus pallens
Panicum pedunculare Willd. ex Steud. =Panicum cayennense
Panicum perpusillum Benoist =Panicum polycomum
Panicum petrosum Trin. =Thrasya petrosa
Panicum pilcomayense Hack. ^Panicum bergii
Panicum pilisparsum G Mey. =Panicum pilosum
Panicum pilosissimum Roth =Panicum rudgei
Panicum pilosum var. lancifolium (Griseb.) R. Pohl =Panicum pilosum
Panicum plantagineum Link =Brachiaria plantaginea
Panicum platycaulon (Poir.) Kuntze =Axonopus compressus
Panicum poiretianum Schult. =Setaria poiretiana
Panicum politii Swallen =Panicum granuliferum
Panicum polygonoides Lam. =Isachne polygonoides
Panicum polystachion L. =Pennisetum polystachion
Panicum praegnans Steud. =Lasiacis sorghoidea
Panicum prieurii Mez =Panicum scabridum
Panicum procerrimum Hack. =Lasiacis procerrima
Panicum procurrens Nees ex Trin. =Ichnanthus procurrens
Panicum proximum Steud. =Panicum mertensii
Panicum pseudo-undulatifolium Roem. & Schult. ^Panicum stoloniferum
Checklist of the Plants of the Guiana Shield 167
Panicum purpurascens Raddi =Brachiaria mutica
Panicum pycnoclados Tutin =Dichanthelium pycnoclados
Panicum quetameense Mez ^Panicum pyrularium
Panicum racemiferum Steud. ? =Setaria poiretiana
Panicum ramuliflorum Hochst. ex Steud. =Steinchisma laxa
Panicum rariflorum Lam. =Orthoclada laxa
Panicum recalvum sensu Pulle non (Nees) Kunth =Digitaria laxa
Panicum reptans L. =Urochloa reptans
Panicum retroflexum Steud. ? =Setaria sp.
Panicum rigens Sw. =Isachne rigens
Panicum rottboellioides Kunth =Mesosetum rottboellioides
Panicum rugulosum var. subvelutinum Doll =Panicum millegrana
Panicum ruscifolium Kunth =Lasiacis ruscifolia
Panicum sanguinale sensu Pulle non L. =Digitaria ciliaris
Panicum savannarum Soderstr. =Panicum cyanescens
Panicum savannarum Schltdl. ex G.A. Black =Axonopus chrysoblepharis
Panicum sciurotis auct., non Trin. =Dichanthelium sciurotoides
Panicum sciurotoides Zuloaga & Morrone =Dichanthelium sciurotoides
Panicum scoparium Rudge =Panicum rudgei
Panicum sellowii var. subvelutinum (Doll) Henrard =Panicum millegrana
Panicum setarioides Steud. =Echinochloa pyramidalis
Panicum siccaneum Trin. =Panicum polycomum
Panicum sloanei Griseb. =Lasiacis sloanei
Panicum sorghoideum Desv. ex Ham. =Lasiacis sorghoidea
Panicum spectabile Nees ex Trin. =Echinochloa polystachya
Panicum spissifolium Swallen =Panicum granuliferum
Panicum stenodoides F.T. Hubb. =Panicum caricoides
Panicum stenophyllum Hack. =Steinchisma stenophylla
Panicum subcordatum Swallen =Panicum granuliferum
Panicum subfalcatum Doll =Paspalum subfalcatum
Panicum subinclusum Swallen =Panicum granuliferum
Panicum sulcatum Aubl. =Setaria palmifolia
Panicum supernum Swallen =Panicum micranthum
Panicum surinamense Hochst. ex Steud. =Axonopus surinamensis
Panicum swartzianum Hitchc. =Lasiacis sorghoidea
Panicum tamayonis Luces =Panicum polycomum
Panicum tatei Swallen =Panicum chnoodes
Panicum telmatum Swallen =Dichanthelium telmatum
Panicum tenax Rich. =Setaria tenax
Panicum tenuiculmum G. Mey. =Steinchisma laxa
Panicum tenuissimum Mart, ex Schrank =Sporobolus tenuissimus
Panicum tiricaense Swallen =Dichanthelium pycnoclados
Panicum tiricaoides Swallen =Dichanthelium pycnoclados
Panicum tropidoblephare Tutin =Panicum chnoodes
Panicum vannum Swallen =Panicum chnoodes
Panicum velutinum G. Mey. =Oplismenus hirtellus
Panicum versicolor Doll =Otachyrium versicolor
Panicum vigoratum Swallen =Panicum altum
Panicum vilvoides Trin. =Sacciolepis vilvoides
Panicum vinaceum Swallen =Panicum cervicatum
Panicum vulpisetum Lam. =Setaria vulpiseta
Panicum wurdackii Swallen =Panicum eligulatum
Panicum zizanioides Kunth =Acroceras zizanioides
Pariana angustifolia Spreng. =Steyermarkochloa angustifolia
Pariana debilis Tutin =Pariana radiciflora
Pariana gleasonii Hitchc. =Pariana radiciflora
Pariana inaequalis Miq. =Pariana campestris
Pariana longiflora Tutin =Pariana radiciflora
Pariana lunata Nees =Pariana campestris
Pariana maynensis Huber =Pariana radiciflora
Pariana mollis Nees =Pariana campestris
Pariana obtusa Swallen =Pariana radiciflora
Pariana pallida Swallen =Pariana radiciflora
Pariana pulcherrima Tutin =Pariana radiciflora
168 Checklist of the Plants of the Guiana Shield
Pariana trichosticha Tutin =Pariana radiciflora
Pariana violascens Swallen =Pahana radiciflora
Pariana vulgaris Tutin =Pariana radiciflora
Pariana zingiberina Rich, ex Doll =Pariana radiciflora
Paspaliim abrahamii Chase =Paspalum maritimum
Paspahim abstrusum Trin. =Paspaliim hyalinum
Paspaliim anceps Mez =Axonopiis anceps
Paspalum appendiculatum J. Presl =Axonopus excavatus
Paspaliim arenicoliim Miill. Hal. =Paspaliim piilchellum
Paspalum argyrocondylon Steud. =Paspalum serpentiniim
Paspalum aureolatum Swallen ^Paspalum lancifloriim
Paspalum aiireum (P. Beauv.) Kunth =Axonopus aureus
Paspalum axillare Swallen =Thrasya axillaris
Paspalum bistipulatum Hochst. ex Steud. =Paspalum repens
Paspalum caespitosiim sensu Pulle non Fliigge -Paspalum maritimum
Paspalum caespitosum Steud. =Paspalum melanospermum
Paspalum canum Sohns =Paspalum ammodes
Paspalum capillare Lam. =Axonopus capillaris
Paspalum carinato-vaginatum Mez =Axonopus aureus
Paspalum carinatum var. kappleri (Steud.) Doll ^Paspalum carinatum
Paspalum caulescens Mez =Axonopus caulescens
Paspalum chrysites (Steud.) Doll =Axonopus aureus
Paspalum chrysoblephare (Lag.) Doll =Axonopus chrysoblepharis
Paspalum chrysodactylon (Trin.) Doll =Axonopus canescens
Paspalum chrysodactylon var. glabratum Doll =Axonopus canescens
Paspalum chrysodactylon var. psilachne Doll =Axonopus canescens
Paspalum chrysodactylon var. villosum Doll =Axonopus canescens
Paspalum ciliatum Lam. =Paspalum conjugatum
Paspalum compressum (Sw.) Raspail -Axonopus compressus
Paspahim compressum var. arenarium Bertoni =Axonopus fissifolius
Paspalum conjugatum var. pubescens Doll =Paspalum conjugatum
Paspalum conjugatum var. subcordatum Griseb. =Axonopus compressus
Paspalum contractum Pilg. =Paspalum lanciflorum
Paspalum densiflorum sensu Pulle non Doll =Paspalum maritimum
Paspalum densum var. ciliatum Doll =Paspalum densum
Paspalum depressum Steud. =Axonopus compressus
Paspalum dispar var. marahuacense H. Rodr. =Paspalum petilum
Paspalum echinotrichum Mez =Paspalum lanciflorum
Paspalum elatiim Rich, ex Doll =Paspalum ariindinaceum
Paspalum eminens Nees =Axonopus eminens
Paspalum erectifolium Swallen =Paspalum aspidiotes
Paspalum excavatum Nees ex Trin. =Axonopus excavatus
Paspalum extenuatum Nees =Axonopus capillaris
Paspalum familiare Steud. =Paspaliim coryphaeum
Paspalum filostachyum A. Rich, ex Steud. =Axonopus compressus
Paspalum fissifolium Raddi =Axonopus fissifolius
Paspalum flexile Mez ^Axonopus purpusii
Paspalum fockei Mez ^Axonopus surinamensis
Paspalum foveolatum Steud. =Paspalum melanospermum
Paspalum froesianum Swallen =Paspaliim loefgrenii
Paspalum furcatum sensu Pulle non Fliigge =Axonopus surinamensis
Paspalum furcatum vav.flssum Doll =Axonopus compressus
Paspalum furcatum war. parviflorum Doll =Axonopus compressus
Paspalum fuscescens J. Presl =Digitariafuscescens
Paspalum geniculatum Steud. =Paspalum orbiculatum
Paspalum gnaphalioideum Miill. Hal. =Axonopus chrysostachyus
Paspalum gossypinum Mez =Paspalum hyalinum
Paspalum gracile Rudge =Paspalum repens
Paspalum guadaloupense Steud. =Axonopus compressus
Paspalum hirtigluma Steud. =Paspalum maritimum
Paspalum hiimile Steud. ^Paspalum melanospermum
Paspalum immersum Nees =Axonopus chrysoblepharis
Paspalum immersum var. pilosum Doll =Axonopus chrysoblepharis
Paspalum indiitum Luces =Paspalum coryphaeum
Checklist of the Plants of the Guiana Shield 169
Paspalum iridifoliiim Poepp. =Axonopus iridifolius
Paspalum kappleri Hochst. ex Steud. =PaspaJum carinatum
Paspalum kegelii Miill. Hal. =Spheneria kegelii
Paspalum lacustre sensu Judz., non Swallen =Paspalum morichalense
Paspalum lanatum Kunth =Leptocoryphium lanatum
Paspalum laticulmum Spreng. =Axonopus compressus
Paspalum leptostachyum Fliigge =Axonopus leptostachyus
Paspalum longebrachiatum Steud. =Paspalum arundinaceum
Paspalum longispica Doll =Axonopus longispicus
Paspalum longispicum Doll =Axonopus longispicus
Paspalum longissimum Hochst. ex Steud. =Paspalum conjugatum
Paspalum lucidulum Swallen =Paspalum altsonii
Paspalum macropodium Steud. =Axonopus compressus
Paspalum maculosum var. rotundiflorum Doll =Paspalum serpentinum
Paspalum meyerianum Spreng. =Paspalum exaltatum
Paspalum minutum Trin. =Axonopus capillaris
Paspalum montanum Henrard =Paspalum oligostachyum
Paspalum orbiculatum var. lanuginosum Henrard =Paspalum orbiculatum
Paspalum orbiculatum var. potaroense Chase ^Paspalum orbiculatum
Paspalum pectinatum sensu Hitchc. non Nees ex Trin. =Paspalum lanciflorum
Paspalum pedunculatum Poir. =Paspalum decumbens
Paspalum piligerum Swallen =Paspalum lanciflorum
Paspalum platycaulon Poir. =Axonopus compressus
Paspalum platycaulon var. gracilius Doll =Axonopus purpusii
Paspalum platycaulon var. parviflorum Doll =Axonopus purpusii
Paspalum plicatulum var. microspermum Doll =Paspalum melanospermum
Paspalum polychaetum Mez =Paspalum hyalinum
Paspalum pulchrum Nees =Axonopus canescens
Paspalum pulchrum var. angustifolium Doll =Axonopus aureus
Paspalum purpusii Mez =Axonopus purpusii
Paspalum pusillum Vent, ex Fliigge =Paspalum orbiculatum
Paspalum raunkiaerii Mez =Axonopus compressus
Paspalum scandens Tutin =Thrasya scandens
Paspalum scoparium var. parviflorum Doll =Axonopus anceps
Paspalum senescens Doll =Axonopus senescens
Paspalum setifolium Doll =Spheneria kegelii
Paspalum setiglume Chase =Paspalum aspidiotes
Paspalum triglochinoides Mez =Axonopus triglochinoides
Paspalum tristachyon Lam. =Axonopus compressus
Paspalum vaginiflorum Steud. =Paspalum decumbens
Paspalum wagenerianum Schltdl. =Paspalum stellatum
Paspalum wullschlaegelii Mez =Paspalum maritimum
Pennisetum dasistachyum Desv. =Pennisetum polystachion
Pennisetum richardii Kunth =Pennisetum polystachion
Pennisetum setosum (Sw.) Rich. =Pennisetum polystachion
Pharus glaber Kunth =Pharus lappulaceus
Pharus glaber sensu Lemee non Kunth =Pharus latifolius
Pharus scaber var. pictus Doll =Pharus latifolius
Phragmites communis Trin. =Phragmites australis
Piptatherum confine Schult. =Arundinella hispida
Poa acutiflora Kunth =Eragrostis acutiflora
Poa amabilis L. =Eragrostis amabilis
Poa atrovirens Desf. =Eragrostis atrovirens
Poa aturensis Kunth =Eragrostis japonica
Poa chariis Schult. =Eragrostis atrovirens
Poa ciliaris L. =Eragrostis ciliaris
Poa conferta Elliott =Eragrostis Japonica
Poa elegantula Kunth =Eragrostis gangetica
Poa gangetica Roxb. =Eragrostis gangetica
Poa glomerata Walter =Eragrostis japonica
Poa hypnoides Lam. =Eragrostis hypnoides
Poajaponica Thunb. =Eragrostisjaponica
Poa linkii Kunth =Eragrostis pilosa
Poa maypurensis Kunth =Eragrostis maypurensis
170 Checklist of the Plants of the Guiana Shield
Poa meratiana Kunth =Eragrostis maypurensis
Poa pectinacea Michx. =Eragrostis pectinacea
Poapilosa L. =Eragrostis pilosa
Poa plumosa Retz. =Eragwstis amabilis
Poa polytricha (Nees) Kunth =Eragrostis polytricha
Poa racemosa Vahl =Eragrostis maypurensis
Poa tenella L. =Eragrostis amabilis
Poa unioloides Retz. =Eragrostis unioloides
Poa vahlii Roem. & Schult. ^Eragrostis maypurensis
Poa viscosa Retz. =Eragrostis viscosa
Poa viscosa var. pilosissima Hochst. ex A. Rich. =Eragrostis viscosa
Polypogon spicatus Spreng. =Gymnopogon spicatus
Raddia capillata (Trin.) Hitchc. =Cryptochloa capillata
Raddia malmeana sensu Hitchc. non (Ekman) Hitchc. =Raddiella kaieteurana
Raddia nana (Doll) Chase =Raddiella esenbeckii
Raddia sympodica (Doll) Hitchc. =Piresia sympodica
Raddiella maipuriensis Soderstr. =Raddiellti potaroensis
Raddiella malmeana sensu Hitchc. non (Ekman) Swallen =Raddiella kaieteurana
Raddiella nana (Doll) Swallen =Raddiella esenbeckii
Raddiella truncata Swallen =Parodiolyra lateralis
Reimaria aberrans Doll =Reimarochloa aberrans
Reimaria acuta Fliigge =Reimarochloa acuta
Reimaria laxa Rchb. ex A. Spreng. =Digitaria laxa
Reimarochloa brasiliensis (Spreng.) Hitchc. =Reimarochloa acuta
Rhynchelytrum roseum (Nees) Stapf& C.E. Hubb. =Rhynchelytrum repens
Roshevitzia glomerata (Walter) Tzvelev =Eragrostisjaponica
Roshevitziajaponica (Thunb.) Tzvelev =Eragrostisjaponica
Rottboellia aurita Steud. =Coelorachis aurita
Rottboellia compressa L. f. =Hemarthria altissima
Rottboellia exaltata L. f. =Rottboellia cochinchinensis
Rottboellia loricata Trin. =Rhytachne rottboellioides
Rottboellia loricata subsp. glaberrima Hack. =Rhytachne subgibbosa
Rottboellia loricata subsp. subgibbosa Winkl. ex Hack. =Rhytachne subgibbosa
Rottboellia sanguinea Retz. =Schizachyrium sanguineum
Saccharum caudatum G. Mey. =Imperata contracta
Saccharum cayennense (R Beauv.) Benth. =Eriochrysis cayennensis
Saccharum contractum Kunth =Imperata contracta
Saccharum giganteum Trin. =Saccharum trinii
Saccharum repens Willd. =Rhynchelytrum repens
Saccharum sagittatum Aubl. =Gynerium sagittatum
Sacciolepis pungens Swallen =Sacciolepis angustissima
Schizachyrium hirtiflorum Nees =Schizachyrium sanguineum
Schizachyrium microstachyum (Desv. ex Ham.) Roseng., B.R. Arrill. & Izag. =Schizachyrium condensatum
Schizachyrium riedelii (Trin.) A. Camus =Schizachyrium sanguineum
Schizachyrium riedelii var. multirameum (Hack.) Henrard =Schizachyrium sanguineum
Schizachyrium semiberbe Nees =Schizachyrium sanguineum
Setaria geniculata (Willd.) R Beauv. [nom. illeg.] =Setaria parviflora
Setaria gracilis Kunth =Setaria parviflora
Setaria poiretiana (Schult.) Kunth = Setaria sulcata Raddi
Setaria sulcata (Aubl.) A. Camus =Setaria palmifolia
Sorghastrum parviflorum (Desv.) Hitchc. &. Chase =Sorghastrum setosum
Sorghum parviflorum Desv. =Sorghastrum setosum
Sorghum vulgare Pers. =Sorghum bicolor
Spartina brasiliensis Raddi =Spartina alterniflora
Spheneria setifolia (Doll) Kuhlm. =Spheneria kegelii
Sporobolus aeneus sensu Hitchc. non (Trin.) Kunth =Sporobolus cubensis
Sporobolus eflusus Desv. =Sporobolus tenuissimus
Sporobolus indicus auct., non (L.) R. Br. =Sporobolusjacquemontii
Sporobolus indicus war. pyramidalis (R Beauv.) Veldkamp =Sporobolusjacquemontii
Sporobolus littoralis (Lam.) Kunth =Sporobolus virginicus
Sporobolus minutiflorus (Trin.) Link =Sporobolus tenuissimus
Sporobolus muralis (Raddi) Hitchc. & Chase =Sporobolus tenuissimus
Sporobolus poiretii auct., non (Roem. & Schult.) Hitchc. =Sporobolus jacquemontii
Sporobolus pyramidalis R Beauv. =Sporobolus jacquemontii
Sporobolus pyramidalis vaY. jacquemontii (Kunth) Jovet & Guedes =Sporobolusjacquemontii
Checklist of the Plants of the Guiana Shield 171
Sporobolus villosus (A. Spreng.) Kunth =Sporobolus ciliatus
Steirachne diandra sensu Swallen non Eicman =Steirachne barbata
Stenotaphrum glabrum var. americanum (Schrank) Doll =Stenotaphrum secundatum
Steyermarkochloa unifolia Davidse & R.P. Ellis =Steyermarkochloa angustifolia
Stipa spicata L. f. =Trachypogon spicatus
Strephium guianense Brongn. =Raddia guianemis
Streptachne scabra Kunth =Aristida ternipes
Streptachne tenuis Kunth =Anstida ternipes
Streptogyna crinita auct., non P. Beauv. =Streptogyna americana
Streptostachys hirsuta P. Beauv. =Streptostachys asperifolia
Streptostachys vaginaeflorum Steud. =Streptostachys asperifolia
Syntherisma chinensis (Nees) Hitchc. =Digitaria violascens
Syntherisma cuyabensis (Trin.) Hitchc. =Digitaria lanuginosa
Syntherisma digitata sensu Hitchc. non (Sw.) Hitchc. =Digitaria horizontalis
Syntherisma longiflora (Retz.) Skeels =Digitaria longiflora
Syntherisma malacophylla Hitchc. =Digitaria lanuginosa
Syntherisma sanguinalis sensu Hitchc. non (L.) Dulac =Digitaria ciliaris
Thrasya campylostachya sensu Amshoff& Henrard non (Hack.) Chase =Thrasya granitica
Thrasya cinerascens (Doll) Judz. =Paspalum cinerascens
Thrasya guianensis Swallen =Thrasya scandens
Thrasya reticulata Swallen =Paspalum multinervium
Thrasya setosa Swallen =Thrasya paspaloides
Trachypogon angustifolius E. Foum. ex Hemsl. =Trachypogon spicatus
Trachypogon angustifolius Nees ex Hack. =Trachypogon spicatus
Trachypogon canescens Nees =Trachypogon spicatus
Trachypogon capensis Trin. =Trachypogon spicatus
Trachypogon capensis subvar mollis (Nees) Roberty =Trachypogon spicatus
Trachypogon dactyloides (Steud.) E. Foum. =Trachypogon spicatus
Trachypogon densus Swallen =Trachypogon spicatus
Trachypogon dissoluta Nees =Trachypogon spicatus
Trachypogon durus Stapf =Trachypogon spicatus
Trachypogon filifolius (Hack.) Hitchc. =Trachypogon spicatus
Trachypogon glaucescens Pilg. =Trachypogon spicatus
Trachypogon gouinii E. Foum. =Trachypogon spicatus
Trachypogon gracilis Andersson =Trachypogon spicatus
Trachypogon gracilis var. ciliatus Andersson =Trachypogon spicatus
Trachypogon gracilis var. hirtus Andersson ^Trachypogon spicatus
Trachypogon involutus Pilg. =Trachypogon spicatus
Trachypogon karwinskyi (Hack.) Nash =Trachypogon spicatus
Trachypogon ligularis Nees =Trachypogon spicatus
Trachypogon macroglossus Trin. =Trachypogon spicatus
Trachypogon mayaensis Wipff & S.D. Jones =Trachypogon spicatus
Trachypogon micans Andersson =Trachypogon spicatus
Trachypogon mollis Nees ^Trachypogon spicatus
Trachypogon montufarii (Kunth) Nees ^Trachypogon spicatus
Trachypogon montufarii var. bolivianus (Pilg.) Pilg. =Trachypogon spicatus
Trachypogon montufarii var. grandiflorus Andersson =Trachypogon spicatus
Trachypogon montufarii var. mollis (Nees) Andersson =Trachypogon spicatus
Trachypogon montufarii var. pauciflorus Andersson =Trachypogon spicatus
Trachypogon montufarii var. pilosus E. Foum. =Trachypogon spicatus
Trachypogon muelleri E. Foum. =Trachypogon spicatus
Trachypogon palmeri Nash =Trachypogon spicatus
Trachypogon parviflorus Swallen =Trachypogon spicatus
Trachypogon plumosus (Humb. & Bonpl. ex Willd.) Nees =Trachypogon spicatus
Trachypogon plumosus var. montufarii (Kunth) Hack. "^Trachypogon spicatus
Trachypogon plumosus var. secundus (J. PresI) Beetle =Trachypogon spicatus
Trachypogon plumosus suhvar secundus (J. Presl) Hack, ex Henrard =Trachypogon spicatus
Trachypogon polymorphus Hack. =Trachypogon spicatus
Trachypogon polymorphus var. bolivianus Pilg. ^Trachypogon spicatus
Trachypogon polymorphus var. canescens (Nees) Hack. ^Trachypogon spicatus
Trachypogon polymorphus subvar capensis (Trin.) Hack. ^Trachypogon spicatus
Trachypogon polymorphus subvar dactyloides (Steud.) Hack. ^Trachypogon spicatus
Trachypogon polymorphus var. dissolutus (Nees) Hack. =Trachypogon spicatus
Trachypogon polymorphus var. filifolius Hack. ^Trachypogon spicatus
Trachypogon polymorphus var. gouinii (E. Foum.) Hack. =Trachypogon spicatus
172 Checklist of the Plants of the Guiana Shield
Trachypogon polymorphus subvar gracilis (Andersson) Hack. =Trachypogon spicatus
Trachypogon polymorphus var. karwinskyi Hack. =Trachypogon spicatus
Trachypogon polymorphus var. ligularis (Nees) Hack. =Trachypogon spicatus
Trachypogon polymorphus var. macroglossus (Trin.) Hack. =Trachypogon spicatus
Trachypogon polymorphus subvar mollis (Nees) Hack. =Trachypogon spicatus
Trachypogon polymorphus var. montufarii (Kunth) Hack. =Trachypogon spicatus
Trachypogon polymorphus war. plumosus (Willd.) Hack. =Trachypogon spicatus
Trachypogon polymorphus subvar secundus (J. Presl) Hack. =Trachypogon spicatus
Trachypogon polymorphus var. thollonii Franch. =Trachypogon spicatus
Trachypogon polymorphus var. truncatus (Nees) Hack. =Trachypogon spicatus
Trachypogon polymorphus var. vestitus (Andersson) Hack. =Trachypogon spicatus
Trachypogon presla Andersson =Trachypogon spicatus
Trachypogon preslii f. secundus (J. Presl) Andersson =Trachypogon spicatus
Trachypogon ramosus Swallen ^Trachypogon spicatus
Trachypogon renvoizei Cat. Guerra ^^Trachypogon spicatus
Trachypogon rigidifolius Swallen =Trachypogon spicatus
Trachypogon rufus Nees =Hyparrhenia rufa^
Trachypogon thollonii (Franch.) Stapf =Trachypogon spicatus
Trachypogon truncatus (Nees) Andersson =Trachypogon spicatus
Trachypogon vestitus Andersson =Trachypogon spicatus
Trichachne insularis (L.) Nees =Digitaria insularis
Trichachne sacchariflora (Raddi) Nees =Digitaria insularis
Trichachne tenuis Nees =Digitaria tenuis
Tricholaena rosea Nees =Rhynchelytrum repens
Tricuspis latifolia Griseb. =Gouinia latifolia
Triodiaflaccida (Doll) Hitchc. =Tridens flaccidus
Tripsacum hermaphroditum L. =Anthephora hermaphrodita
Tripsacum latifolium sensu Hitchc. (1922) non Hitchc. (1906) =Tripsacum australe
Tripsacum laxum sensu Ostendorf non Nash =Tripsacum andersonii
Uralepis flaccida Doll =Tridens flaccidus
Urochloa arrecta (Hack, ex T. Durand & Schwinze) Morrone & Zuloaga =Brachiaria arrecta
Urochloa decumbens (Stapf) R.D. Webster =Brachiaria decumbens
Urochloafasciculata (Sw.) R.D. Webster =Brachiariafasciculata
Urochloafusca (Sw.) B.F. Hansen & Wunderlin =Brachiaria fasciculata
Urochloa humidicola (Rendle) Morrone & Zuloaga =Brachiaria humidicola
Urochloa maxima (Jacq.) R.D. Webster =Panicum maximum
Urochloa mollis (Sw.) Morrone & Zuloaga =Brachiaria mollis
Urochloa mutica (Forssk.) T.Q. Nguyen =Brachiaria mutica
Urochloa plantaginea (Link) R.D. Webster =Brachiaria plantaginea
Valota insularis (L.) Chase =Digitaria insularis
Valota laxa (A. Spreng.) Hitchc. & Chase =Digitaria laxa
Vilfajacquemontii (Kunth) Trin. =Sporobolus jacquemontii
Vilfa minutiflora Trin. =Sporobolus tenuissimus
Vilfa pyramidalis (P. Beauv.) Trin. ex Steud. ^Sporobolusjacquemontii
Vilfa villifera Steud. =Sporobolus ciliatus
References
(for a more extensive bibliography, see Judziewicz 1990)
Amshoff, G.J.H. & J.T. Henrard. 1943. Gramineae. In A. Pulle, Flora of Suriname 1(1): 273-442.
Cremers, G. & M. Hoff. 1993. Inventaire taxonomique des plantes de la Guyane fran9aise III - les Cyperaceae et les Poaceae.
Mus. Nat. Hist. Nat., Coll. Patr. Nat. (Ser. Patr. Genetique) 11: 1-213.
Davidse, G, F.O. Zuloaga, & T.S. Filrueiras. 1995. Paspalum morichalense (Poaceae: Paniceae), a new aquatic species from
South America. Novon 5(3): 234-237.
Davidse, G E.J. Judziewicz, & F.O. Zuloaga. 2004. Poaceae. In In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 8: 1-297.
Hitchcock, A.S. 1922. Grasses of British Guiana. Contr. U.S. Natl. Herb. 22: 443-514.
Judziewicz, E. 1990. Poaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 8: 1-725.
Judziewicz, E. 1997. Poaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1 : 342-360.
Lindeman, J.C. & A.R.A. Gorts-van Rijn. 1968. Gramineae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of
Suriname 1(2): 343-375.
Swallen, J. 1948. Gramineae. In B. Maguire et al.. Plant explorations in Guiana in 1944, chiefly to the Tafelberg and the Kaieteur
Plateau - 1. Bull. Torrey Bot. Club 75: 81-91.
Checklist of the Plants of the Guiana Shield 173
BO DA GU su FG
AM BO DA GU su FG
AM BO DA GU su FG
AM BO
DA
GU
GU
su FG
AM BO
BO
BO
DA GU
GU
GU
su FG
PONTEDERIACEAE
C.N. Horn
Eichhornia azurea (Sw.) Kunth
Eichhornia crassipes (Mart.) Solms
Eichhornia diversifolia (Vahl) Urb.
Eichhornia heterosperma Alexander
Eichhornia paniculata (Spreng.) Solms [probably naturalized]
Heteranthera multiflora (Griseb.) C.N. Horn
Pontederia rotundifolia L.
Pontederia subovata (Seub.) Lowden
Pontederia triflora (Endl. ex Seub.) G Agostini, D. Velasquez & Velasquez
Synonyms and Misidentifications
Eichhornia polysperma [dubious name of uncertain origin]
Eichhornia venezuelensis Velasquez =Eichhornia heterosperma
Heteranthera diversifolia Vahl "^Eichhornia diversifolia
Heterantheraformosa Miq. =Eichhornia crassipes
Heteranthera grandiflora Klotzsch [nomen nudum] =Eichhornia diversifolia
Heteranthera reniformis var. multiflora Griseb. =Heteranthera multiflora
Heteranthera zosterifolia Mart, [not in flora area, misident. for Eichhornia diversifolia]
Pontederia azurea Sw. =Eichhornia azurea
Pontederia cordata L. [not in flora area]
Pontederia cordifolia Mart, ex Schult. & Schult. f. =Pontederia rotundifolia
Pontederia crassipes Mart. =Eichhornia crassipes
Pontederia eriantha Miq. =Pontederia rotundifolia
Pontederia schomburgkiana Klotzsch [nomen nudum] =Pontederia subovata
Reussia triflora Endl. ex Seub. =Pontederia triflora
References
Castellanos, A. 1958. Las Pontederiaceae de Brasil. Arq. Jard. Bot. Rio de Janeiro 16: 149-216.
Horn, C.N. 1994. Pontederiaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 15: 91-102.
Horn, C.N. 2004. Pontederiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
8:371-376.
Jonker-Verhoef, A.M.E. 1953. Pontederiaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 81-90.
Lowden, R.M. 1973. A revision of the genus Pontederia. Rhodora 75: 426-487.
Rapateaceae
RE. Berry
Amphiphyllum rigidum Gleason
Cephalostemon affinis Kom.
Cephalostemon gracilis (Poepp. & Endl.) R.H. Schomb.
Cephalostemon microglochin Sandwith
Duckea cyperaceoidea (Ducke) Maguire
Duckeaflava (Link) Maguire
Duckeajunciformis Maguire
Duckea squarrosa (Willd.) Maguire
Epidryos guayanensis Maguire
Epidryos sp. A
Guacamaya superba Maguire
Kunhardtia radiata Maguire & Steyerm.
Kunhardtia rhodantha Maguire
Marahuacaea schomburgkii (Maguire) Maguire
Monotrema aemulans Kom.
Monotrema affine Maguire
Monotrema bracteatum Maguire
Monotrema bracteatum subsp. bracteatum
Monotrema bracteatum subsp. major Maguire
Monotrema xyridoides Gleason
Phelpsiella ptericaulis Maguire
Potarophytum riparium Sandwith
Rapatea angustifolia Spruce ex Kom.
Rapatea aracamuniana Steyerm.
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
SU
SU?
BO
BO GU
BO
BO
BO
GU
GU
174 Checklist of the Plants of the Guiana Shield
Rapatea chimantensis Steyerm.
Rapatea circasiana Garcia-Barr. & L.E. Mora
Rapateafanshawei Maguire
Rapateafanshawei yar.fanshawei
Rapateafanshawei var. minor Maguire
Rapatea linearis Gleason
Rapatea longipes Spruce ex Kom.
Rapatea membranacea Maguire
Rapatea paludosa Aubl.
Rapatea paludosa var. paludosa
Rapatea paludosa var. sessiliflora Maguire
Rapatea satilensis B.M. Boom
Rapatea scabra Maguire
Rapatea spruceana Kom.
Rapatea steyermarkii Maguire
Rapatea ulei Pilg.
Rapatea xiphoides Sandwith *
Rapatea yapacana Maguire
Saxofridericia aculeata Kom.
Saxofridericia compressa Maguire
Saxofridericia duidae Maguire
Saxofridericia grandis Maguire
Saxofridericia inermis Ducke
Saxofridericia petiolata Maguire
Saxofridericia regalis R.H. Schomb.
Saxofridericia sp. A
Saxofridericia spongiosa Maguire
Schoenocephalium cucullatum Maguire
Schoenocephalium teretifolium Maguire
Spathanthus bicolor Ducke
Spathanthus unilateralis (Rudge) Desv.
Spathanthus unilateralis var. abruptus Maguire
Spathanthus unilateralis war. jenmanii (N.E. Br.) Maguire
Spathanthus unilateralis var. unilateralis
Stegolepis albiflora Steyerm.
Stegolepis angustata Gleason
Stegolepis breweri Maguire
Stegolepis cardonae Maguire
Stegolepis celiac Maguire
Stegolepis choripetala Maguire
Stegolepisferruginea Baker f.
Stegolepis gleasoniana Steyerm.
Stegolepis grandis Maguire
Stegolepis guianensis Klotzsch ex Kom.
Stegolepis hitchcockii Maguire
Stegolepis hitchcockii subsp. hitchcockii
Stegolepis hitchcockii subsp. morichensis Maguire
Stegolepis huberi Steyerm.
Stegolepis humilis Steyerm.
Stegolepisjauaensis Maguire
Stegolepis ligulata Maguire
Stegolepis linearis Gleason
Stegolepis maguireana Steyerm.
Stegolepis membranacea Maguire
Stegolepis microcephala Maguire
Stegolepis minor Steyerm.
Stegolepis neblinensis Maguire [Sierra de la Neblina]
Stegolepis parvipetala Steyerm.
Stegolepis pauciflora Gleason
Stegolepis perligulata Maguire
Stegolepis ptaritepuiensis Steyerm.
Stegolepis pulchella Maguire
Stegolepis pungens Gleason
Stegolepis sp. nov. [Mt. Ayanganna]
BO
AM
BO GU
BO GU
GU
GU
AM
GU
AM BO DA GU SU FG
AM BO DA GU SU
SU
SU
FG
FG
AM
AM
BO GU
GU
GU
FG
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
GU SU FG
GU SU
GU
GU?
GU
GU
GU
GU
GU
GU
SU
SU
FG
FG
FG
FG
GU
GU
Checklist of the Plants of the Guiana Shield 175
Stegolepis squarrosa Maguire BO
Stegolepis steyermarkii Maguire BO GU
Stegolepis terramarensis Steyerm. AM
Stegolepis vivipara Maguire BO
Stegolepis wurdackii Maguire AM BO
Stegolepis wurdackii subsp. chimantensis Maguire BO
Stegolepis wurdackii subsp. wurdackii [Sierra de la Neblina] AM
Windsorina guianensis Gleason GU
Synonyms and Misidentifications
Amphiphyllum schomburgkii Maguire =Marahuacaea schomburgkii
Cephalostemon cyperaceoides Ducke =Duckea cyperaceoidea
Cephalostemon flavus (Link) Steyerm. ^Duckeaflava
Cephalostemonjunciformis (Maguire) Steyerm. =Duckeajunciformis
Cephalostemon squarrosus (Willd.) Kom. =Duckea squarrosa
Mnasium paludosum (Aubl.) Willd. =Rapatea pahidosa
Mnasium unilaterale Rudge =Spathanthus unilateralis
Mnasium unilateralis Rudge =Spathanthus unilateralis
Rapatea longipes van modesta (Maguire) Maguire =Rapatea longipes
Rapatea modesta Maguire =Rapatea longipes
Rapatea schultesiana Garcia-Barr. & L.E. Mora =Rapatea pahidosa
Rapatea unilateralis (Rudge) Schult. & Schult. f. =Spathanthus unilateralis
Rapatea viscosa Gleason =Saxofridericia regalis
Saxofridericia australis Gleason =Saxofridericia aculeata
Saxofridericia duidae subsp. marahuacensis Maguire =Saxofridericia duidae
Saxofridericia pandanoides Linden & Andre =Saxofridericia aculeata
Saxofridericia subcordata Kom. =Saxofridericia aculeata
Schoenocephalium coriaceum Maguire =Schoenocephalium cucullatum
Schoenocephalium sipapoanum Maguire =Kunhardtia rhodantha
SpathanthusJenmaniiN.E. Br. =Spathanthus unilateralis var. jenmanii
Stegolepis aemulans (Kom.) Benth. & Hook. =Monotrema aemulans
Stegolepis grandis subsp. Jauensis Maguire =Stegolepis grandis
Stegolepis grandis subsp. phelpsiae Maguire =Stegolepis grandis
Stegolepis parvipetala subsp. chimantensis Maguire =Stegolepis parvipetala
References
Berry, RE. 2004. Rapateaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
8:413-472.
Boom, B.M. 1994. A new species of Rapatea section Paludosa from French Guiana. Brittonia 46(4): 314-316.
Boom, B.M. 1997. Rapateaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1 : 360-362.
Lindeman, J.C. & A.R.A. Gorts-van Rijn. 1968. Rapateaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of
Suriname 1(2): 326-328.
Maguire, B. 1958. Rapateaceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part 3. Mem.
New York Bot. Gard. 10(1): 1-156.
Maguire, B. 1965. Rapateaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 6. Mem. New York Bot.
Gard. 12(3): 1-285.
Maguire, B. 1982. Rapateaceae. In Z. Luces de Febres, Flora de Venezuela 6(2): 85-203.
Uittien, H. 1938. Rapateaceae. In A. Pulle, Flora of Suriname 1(1): 191-196.
RUPPIACEAE
R. Haynes, L.B. Holm-Nielsen
Ruppia maritima L. GU SU FG
References
Haynes, R.R., L.B. Holm-Nielsen & D.H. Les. 1998. Ruppiaceae. Families and Genera of Vascular Plants 4: 445—448.
Smilacaceae
RE. Berry, R.A. DePhilippst, J.F. Gaskin
Smilax cinnamommea Desf ex DC. FG
Smitax cordato-ovata Rich. FG
176 Checklist of the Plants of the Guiana Shield
AM
AM
AM
AM
BO
BO
BO
GU SU FG
FG
GU
GU FG?
GU su FG
GU SU FG
su? FG
GU su FG?
GU?
GU?
FG
FG
FG
GU SU FG
FG
GU su FG
GU su FG
GU su FG
GU
Smilax cumanensis Humb. & Bonpl. ex Willd. AM BO DA
Smilax cuspidata Poir.
Smilax immersa A.C. Sm.
Smilax krukovii A.C. Sm.
Smilax kunthii Killip & C.V. Morton
Smilax lappacea Humb. & Bonpl. ex Willd.
Smilax latipes Gleason
Smilax longifolia Rich.
Smilax maypurensis Humb. &. Bonpl. ex Willd.
Smilax megalophylla Duhamel
Smilax pittieriana Steyerm.
Smilax poeppigii Kunth
Smilax pseudosyphilitica Kunth
Smilax quinquenervia Veil.
Smilax riedeliana A. DC.
Smilax santaremensis DC.
Smilax satilensis J.D. Mitch. i,
Smilax schomburgkiana Kunth AM BO DA
Smilax staminea Griseb. AM BO
Smilax syphilitica Humb. & Bonpl. ex Willd. AM BO DA
Smilax talbotiana A. DC.
Smilax triplinervia Humb. & Bonpl. ex Willd. AM
Synonyms and Misidentifications
Smilax auraimensis Steyerm. =Smilax pittieriana
Smilax chimantensis Steyerm. & Maguire =Smilax pittieriana
Smilax duidae Steyerm. =Smilax syphilitica
Smilaxflavescens Ham. =Smilax longifolia
Smilaxfloribunda Ham. ? =Smilax cuspidata
Smilax globifera G Mey. =Smilax cumanensis
Smilax hostmanniana Kunth =Smilax cumanensis
Smilaxjauaensis Steyerm. & Maguire =Smilax staminea
Smilax lasseriana Steyerm. =Smilax staminea
Smilax papyracea Duhamel =Smilax longifolia
Smilax pirarensis Kunth & M.R. Schomb. =Smilax cumanensis
Smilax procera Griseb. =Smilax quinquenervia
Smilax staminea Griseb. f. obtusata Steyerm. =Smilax staminea
Smilax surinamensis Miq. =Smilax cumanensis
References
Andreata, R.H.R 1980. Smilax Linnaeus (Smilacaceae): Ensaio para uma revisao das especies brasileiras. Arq. Jard. Bot. Rio de
Janeiro 24: 1 79-30 l.Lachman-White, D.A., Adams, CD. and U.O. Trotz. 1987. A Guide to Medicinal Plants of Coastal
Guyana: CSC Technical Publication Series No. 225. 350 pp. London: Commonwealth Science Council. (Smilax, pp. 256-
257
Gaskin, J.F. & RE. Berry. 2005. Smilacaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan
Guayana 9: 184-193.
Lemee, A. 1955. Smilax. Flore de la Guyane Fran^aise 1: 321-324. Paris: Paul Lechevalier.
Mitchell, J.D. 1994. Smilax saulensis, a new species from French Guiana. Brittonia 46(3): 225-227.
Mitchell, J.D. 1997. Smilacaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 362-365.
Ostendorf, F.W 1962. Nuttige Planten en Sierplanten in Suriname. Landbouwproefstation in Suriname Bulletin No. 79. 325 pp.
(Smilax, pp. 216-217).
Roosmalen, M.G.M. van. 1985. Fruits of the Guianan Flora. 483 pp. Utrecht: Institute of Systematic Botany, Utrecht University;
Wageningen: Silvicultural Department, Wageningen Agricultural University. (Smilax, pp. 254-256)
Sipman, H. 1979. Liliaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 442-456.
Strelitziaceae
L.L. Andersson, P.J.M. Maas
Phenakospermum guyannense (Rich.) Endl. ex Miq. AM
Synonyms and Misidentifications
Ravenala guyannensis (Rich.) Petersen =Phenakospermum guyannense
Urania guyannensis Rich. =Phenakospermum guyannense
BO GU SU FG
Checklist of the Plants of the Guiana Shield 177
References
Andersson, L.L. 2005. Strelitziaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 9: 281-283.
Maas, P.J.M. 1985. Musaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 1: 1-28.
Maas, P.J.M. & M.J.M. de Rooij. 1979. Musaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 393-411.
Maas, P. & H. Maas-van de Kamer. 1997. Strelitziaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D.
Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and
Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1 : 365-367.
Taccaceae
P.E. Berry, Z.S. Rogers
Tacca parkeri Seem. AM GU SU?
Synonyms and Misidentifications
Tacca parkeri var. lanceolata H. Limpr. =Tacca parkeri
Tacca sprucei Benth. =Tacca parkeri
Tacca ulei H. Limpr. =Tacca parkeri
References
Berry, P.E. & Z.S. Rogers. 2005. Taccaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan
Guayana 9: 295-296.
Drenth, E. 1972. A revision of the family Taccaceae. Blumea 20: 367-406.
Thurniaceae
RE. Berry, R. Riina, M.T. Strong
ThurniaJenmanii Hook. f. GU
Thurnia polycephala Schnee AM
Thurnia sphaerocephala (Rudge) Hook. f. AM BO GU SU FG
Thurnia sphaerocephala subsp. 1 GU
Synonyms and Misidentifications
Mnasium sphaerocephala Rudge =Thurnia sphaerocephala
References
Mori, S.A. 1997. Thurniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997.
Guide to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs
of the New York Botanical Garden 76, pt. 1: 367.
Schnee, L. 1943. El genero Thurnia (con descripcion de una nueva especie). Bol. Soc. Venez. Cienc. Nat. 8(55):_241-243.
Riina, R. & P.E. Berry. 2005. Thurniaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 9: 330-332.
Uittien, H. 1938. Thurniaceae. In A. PuUe, Flora of Suriname 1(1): 188-190.
Triuridaceae
P.J.M. Maas, H. Maas
Peltophyllum luteum Gardner
Sciaphila albescens Benth.
Sciaphila picta Miers
Sciaphila polygyna Maas
Sciaphila purpurea Benth.
Sciaphila rubra Maas
Soridium spruceanum Miers
Triuris hexophthalma Maas
Triuris hyalina Miers
Synonyms and Misidentifications
Sciaphila brevipes S.F. Blake =Soridium spruceanum
Sciaphila guianensis Sandwith =Sciaphila albescens
Sciaphila spruceana (Miers) Engl. =Soridium spruceanum
Sciaphila spruceana (Miers) Benth. & Hook. f. =Soridium spruceanum
Soridium guianense (Sandwith) Giesen =Sciaphila albescens
AM
AM
AM
AM
AM
BO
BO
GU
GU SU FG
GU?
FG
GU
SU FG
GU
GU SU
178 Checklist of the Plants of the Guiana Shield
References
Jonker, F.P. 1943. Triuridaceae. In A. Pulle, Flora of Suriname 1(1): 461-466.
Maas, P.J.M. & T. Rubsamen. 1986. Triuridaceae. Flora Neotropica Monogr. 40: 1-55.
Maas, P.J.M. & H. Maas-van de Kamer. 1989. Triuridaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 5: 1-16.
Maas, P.J.M. & H. Maas. 2005. Triuridaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan
Guayana9: 368-371.
Maas-van de Kamer, H. & P. Maas. 1997. Triuridaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D.
Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and
Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1:367-371.
Typhaceae
L.J. Raechal, M.T. Strong
Typha domingensis Pers.
Typha angiistifolia L. [not in flora area]
BO DA GU SU FG
Synonyms and Misidentifications
References
Lanjouw, J. 1938. Typhaceae. In A. Pulle, Flora of Suriname 1(1): 252-254.
Raechal, L.J. 2005. Typhaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
9: 385.
Velloziaceae
RE. Berry, J.S. Miller
Barbacenia celiae Maguire
Vellozia tubiflora (A. Rich.) Kunth
AM
AM BO GU
Synonyms and Misidentifications
Barbacenia alexandrinae R.H. Schomb. ex Hook. =Vellozia tubiflora
Barbacenia uaipanensis Maguire (Bromeliaceae) =Ayensua uaipanensis
Pleiirostima celiae (Maguire) Menezes =Barbacenia celiae
Radia tubiflora A. Rich. =Vellozia tubiflora
Vellozia alexandrinae (R.H. Schomb. ex Hook.) Goethart & Henrard =Vellozia tubiflora
Vellozia duidae Steyerm. =Vellozia tubiflora
Vellozia uaipanensis (Maguire) L.B. Sm. (Bromeliaceae) =Ayensua uaipanensis
References
Maguire, B. 1969. Velloziaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part VIII. Mem. New York
Bot. Gard. 18(2): 1-290.
Miller, J.S. & RE. Berry. 2005. Velloziaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan
Guayana 9: 404-406.
Smith, L.B. & E.S. Ayensu. 1976. A revision ofAmerican Velloziaceae. Smiths. Contrib. Bot. 30: 1-172.
Abolboda acaulis Maguire
Abolboda acaulis var. acaulis
Abolboda acaulis var. scaposa Krai
Abolboda acicularis Idrobo & L.B. Sm.
Abolboda acicularis var. acicularis
Abolboda acicularis var. granulans Maguire
Abolboda americana (Aubl.) Lanj.
Abolboda bella Maguire
Abolboda ciliata Maguire & Wurdack
Abolboda dunstervillei Maguire ex Krai
Abolboda ebracteata Maguire & Wurdack
Abolboda ebracteata var. brevifolia Maguire
Abolboda ebracteata var. ebracteata
Abolboda x glomerata Maguire [sterile hybrid]
Abolboda grandis Griseb.
Abolboda grandis var. grandis
Xyridaceae
L. Kral, M.T. Strong
AM BO
AM BO
AM BO
AM BO
AM BO
AM
AM BO
AM
AM
AM
AM
AM
AM
AM
AM BO
AM BO
GU
GU
GU
GU SU FG
GU
GU
SU
SU
FG
FG
Checklist of the Plants of the Guiana Shield 179
Abolboda grandis van rigida Malme
Abolboda killipii Lasser
Abolboda linearifolia Maguire
Abolboda macrostachya Spruce ex Malme
Abolboda macrostachya var. macrostachya
Abolboda macrostachya var. robustior Steyerm.
Abolboda neblinae Maguire
Abolboda paniculata Maguire
Abolboda poarchon Seub.
Abolboda pulchella Bonpl.
Abolboda scabrida Krai
Abolboda sprucei Malme
Abolboda uniflora Maguire
Achlyphila disticha Maguire & Wurdack
Aratitiyopea lopezii (L.B. Sm.) Steyerm. & RE. Berry
Aratitiyopea lopezii var. lopezii
Orectanthe ptaritepuiana (Steyerm.) Maguire
Orectanthe sceptrum (Oliv.) Maguire
Orectanthe sceptrum subsp. sceptrum
Xyris albescens Steyerm. [Mt. Roraima]
Xyris anceps Lam.
Xyris aquatica Idrobo & L.B. Sm.
Xyris aracamunae Krai
Xyris arachnoidea Maguire & L.B. Sm.
Xyris atriceps Malme
Xyris atriceps subsp. atriceps
Xyris atriceps subsp. chimantensis Maguire & L.B. Sm.
Xyris atriceps subsp. marahuacae Krai & L.B. Sm.
Xyris atriceps subsp. neblinensis Maguire & L.B. Sm.
Xyris bicephala Gleason
Xyris bicostata Maguire & L.B. Sm.
Xyris bicostata var. bicostata
Xyris bicostata var. tillettii (L.B. Sm.) Krai
Xyris byssacea Krai
Xyris carinata Maguire & L.B. Sm.
Xyris chimantae Krai & L.B. Sm.
Xyris concinna N.E. Br. [Mt. Roraima]
Xyris connosepala Lanj. & Lindeman
Xyris consolida Krai & L.B. Sm.
Xyris contracta Maguire & L.B. Sm.
Xyris cryptantha Maguire & L.B. Sm.
Xyris culmenicola Steyerm.
Xyris cyperoides Gleason
Xyris decussata Gleason [Mt. Roraima]
Xyris delicatula Maguire & L.B. Sm.
Xyris esmeraldae Steyerm.
Xyris fallax Malme
Xyris frequens Maguire & L.B. Sm.
Xyrisfrondosa Maguire & L.B. Sm.
Xyris fuliginea Krai & L.B. Sm.
Xyris globosa L.A. Nilsson
Xyris gongylospica Krai
Xyris graniticola Krai
Xyris guianensis Steud.
Xyris huberi Krai & L.B. Sm.
Xyris hymenachne Mart. [Mt. Roraima]
Xyris involucrata Nees
Xyris juncifolia Maguire & L.B. Sm.
Xyrisjupicai Rich.
Xyris kukenaniana Krai [expected in GU]
Xyris lacerata Pohl ex Seub. [expected in GU]
Xyris lanulobractea Steyerm.
Xyris laxifolia Mart.
Xyris laxifolia var. laxifolia
AM BO GU SU
AM BO GU SU
AM
AM BO GU
AM BO GU
AM BO GU
AM
AM BO
SU?
AM BO GU SU
AM
AM
AM
AM
AM
AM
BO GU
AM BO GU
GU
AM BO GU
GU? SU?
FG
FG
AM
AM
AM
AM BO
AM
BO
AM
AM
BO GU SU
AM
AM
AM
AM BO
AM BO
BO
BO
BO
GU
SU
AM
AM
AM
BO GU SU
BO GU
AM BO
AM
AM BO GU SU FG
AM
BO
AM
AM BO
BO
AM
AM BO GU SU
AM
AM BO GU SU FG
AM BO GU SU
AM BO
AM BO
BO
DA GU SU FG
AM BO
BO GU
AM BO DA GU SU FG
AM BO DA GU SU FG
180 Checklist of the Plants of the Guiana Shield
Xyris liesneri Krai
Xyris lithophila Krai & L.B. Sm.
Xyris lomatophylla Mart.
Xyris lugubris Malme
Xyris malmeana L.B. Sm.
Xyris melanovaginata Krai & L.B. Sm.
Xyris mima L.B. Sm. & Downs
Xyris neblinae Maguire & L.B. Sm.
Xyris oblata Krai & L.B. Sm.
Xyris oxylepis Idrobo & L.B. Sm.
Xyris paraensis Poepp. ex Kunth
Xyris paraens is van longiceps (Malme) L.B. Sm. & Downs
Xyris paraensis var. paraensis
Xyris pratensis Maguire & L.B. Sm.
Xyris prolificans Krai
Xyris ptariana Steyerm.
Xyris riparia Maguire & L.B. Sm.
Xyris roraimae Malme [Mt. Roraima]
Xyris rubrolimbata Heimerl
Xyris savanensis Miq.
Xyris schneeana L.B. Sm. & Steyerm.
Xyris setigera Oliv. ex Thum
Xyris seubertii L.A. Nilsson [Mt. Roraima]
Xyris spathacea Lanj.
Xyris spruceana Malme
Xyris stenophylloides Malme
Xyris stenostachya Steyerm.
Xyris subglabrata Malme
Xyris subuniflora Malme
Xyris sulcatifolia Krai
Xyris surinamensis Spreng.
Xyris tatei Malme
Xyris teinosperma Idrobo & L.B. Sm.
Xyris tenella Kunth
Xyris tenella var. leprieurii Malme
Xyris tenella var. tenella
Xyris thysanolepis Maguire & L.B. Sm.
Xyris thysanolepis var. sipapoa (Maguire & L.B. Sm.) Krai & L.B. Sm.
[expected in GU]
Xyris thysanolepis var. thysanolepis [expected in GU]
Xyris toronoana Krai
Xyris uleana Malme
Xyris uleana var. angustifolia Lanj.
Xyris uleana var. uleana
Xyris valdeapiculata Krai
Xyris witsenioides Oliv. ex Thum [Mt. Roraima]
Xyris wurdackii Maguire & L.B. Sm.
Xyris wurdackii subsp. wurdackii
Xyris xiphophylla Maguire & L.B. Sm.
AM
BO
AM
AM
AM BO
BO
BO
GU su FG
AM
AM/\iVl
AM
AM BO GU su FG
AM BO GU su FG
BO GU su FG
AM
AM
AM BO
BO
BO GU
AM
AM BO
BO
BO
BO
GU
GU
GU
su FG
AM GU su FG
AM
AM BO
AM BO
AM BO
AM BO GU su
AM BO
AM BO GU su FG
AM
AM
AM BO GU su FG
FG
AM BO GU su FG
AM BO
AM BO
AM BO
BO
BO GU su FG
AM BO GU su
AM BO GU su
AM
AM BO GU
AM
AM
AM
Synonyms and Misidentifications
Abolboda americana var. imberbis (Kunth) Maguire =Abolboda americana
Abolboda brasiliensis Kunth =Abolboda pulchella
Abolboda chapadensis vax. pauciflora Hoehne =Abolboda pulchella
Abolboda excelsa Malme =Abolboda macrostachya var robustior
Abolboda gracilis Ruber =Abolboda pulchella
Abolboda grandis var. guayanensis Maguire =Abolboda grandis var. rigida
Abolboda grandis var. minor Spruce ex Malme =Abolboda grandis
Abolboda imberbis Kunth =Abolboda americana
Abolboda macrostachya var. angustior Maguire "^Abolboda macrostachya
Abolboda minima Maguire =Abolboda killipii
Abolboda pervaginata Malme =Abolboda grandis var. rigida
Abolboda poeppigii Kunth =Abolboda americana
Abolboda psammophila Maguire =Abolboda killipii
Checklist of the Plants of the Guiana Shield 181
Abolboda ptaritepuiana Steyerm. =Orectanthe ptaritepuiana
Abolboda rigida Gleason =Abolboda macrostachya var. robustior
Abolboda rigida (Malme) Steyerm. =Abolboda grandis var. rigida
Abolboda sceptrum Oliv. =Orectanthe sceptrum
Abolboda schultesii Idrobo & L.B. Sm. =Abolboda sprucei
Orectanthe sceptrum subsp. occidentalis Maguire =Orectanthe sceptrum
Xyris acuminata Miq. ex Steud. =Xyrisjupicai
Xyris ambigua Beyr. ex Kunth =Xyrisjupicai
Xyris americana Aubl. =Abolboda americana
Xyris anceps Pers., [nom. illeg.] =Xyrisjupicai
Xyris applanata Idrobo & L.B. Sm. =Xyris spruceana
Xyris arenicola Miq. =Xyrisjupicai
Xyris arescens Kunth =Xyris hymenachne
Xyris asterocephala Seub. =Xyris involucrata
Xyris calcarata Heimerl =Xyris seubertii
Xyris capillaris Steyerm. =Xyris subuniflora
Xyris caroliniana var. major (Mart.) Idrobo & L.B. Sm. =Xyris laxifolia var. laxifolia
Xyris coerulea Lam. =Abolboda americana
Xyris communis Kunth =Xyris jupicai
Xyris connectens Malme =Xyris uleana var. angustifolia
Xyris dolichosperma Lanj. =Xyris fallax
Xyris duidensis Malme =Xyris uleana var. uleana
Xyris epicarae Krai & L.B. Sm. =Xyris cyperoides
Xyris eriophylla Rchb. [nomen nudum] ^Xyris surinamensis
Xyris eriophylla Rchb. ex Kunth =Xyris surinamensis
Xyris erythema Maguire & L.B. Sm. =Xyris fallax
Xyris exserta Idrobo & L.B. Sm. =Xyris spathacea
Xyrisfdiscapa Malme =Xyris guianensis
Xyrisfoveolata Krai & L.B. Sm. =Xyris huberi
Xyris fusiform is Maguire & L.B. Sm. =Xyris oxylepis
Xyris garcia-barrigae Idrobo & L.B. Sm. =Xyris subglabrata
Xyris gardneri Malme =Xyris guianensis
Xyris glabrata sensu Griseb. et auct. =Xyris malmeana
Xyris guianensis Klotzsch [nomen nudum] =Xyris jupicai
Xyris gymnoptera Griseb. =Xyris jupicai
Xyrisjauana L.B. Sm. & Steyerm. =Xyris thysanolepis var. thysanolepis
Xyrisjupicae Michx. =Xyris jupicai
Xyrisjupicae var. brachylepis Malme =Xyris jupicai
Xyris jupicae var. major (Mart.) L.B. Sm. & Downs =Xyris laxifolia var. laxifolia
Xyris laxifolia var. major Mart. =Xyris laxifolia var. laxifolia
Xyris laxifolia var. minor Mart. =Xyrisjupicai
Xyris leptostachya Malme =Xyris paraensis var. longiceps
Xyris longiceps Malme =Xyris paraensis var. longiceps
Xyris macrocephala auct. non Vahl =Xyris laxifolia
Xyris ? macrocephala Vahl [nom. confus.] =Xyris laxifolia var. laxifolia
Xyris macrocephala var. major (Mart.) L.A. Nilsson =Xyris laxifolia var. laxifolia
Xyris macrocephala f. minor (Mart.) M. Kuhlm. & E. Kuhn =Xyris jupicai
Xyris rugulosa Maguire & L.B. Sm. =Xyris esmeraldae
Xyris scabridula Steyerm., [nom. illeg.] =Xyris sulcatifolia
Xyris setigera var. elongata Steyerm. =Xyris setigera
Xyris sipapoa Maguire & L.B. Sm. =Xyris thysanolepis var. sipapoa
Xyris steyermarkii Maguire & L.B. Sm. =Xyris tenella var. tenella
Xyris submontana Gleason =Xyns hymenachne
Xyris surinamensis Miq. =Xyris jupicai
Xyris tillettii L.B. Sm. =Xyris bicostata var tilleftii
Xyris trisperma Krai & L.B. Sm. =Xyris mima
Xyris vaginata Spreng. =Abolboda pulchella
Xyris vaupesana Idrobo & L.B. Sm. =Xyris uleana var. uleana
Xyris yapobodensis Idrobo & L.B. Sm. =Xyris spathacea
Xyris yutajens is Maguire & L.B. Sm. ='Xyris tenella var. tenella
References
Krai, R. 1988. The genus Xyris in Venezuela and contiguous northern South America. Ann. Missouri Bot. Gard. 75: 522-722.
Krai, R. 1992. A treatment of American Xyridaceae exclusive of Xyris. Ann. Missouri Bot. Gard. 79: 819-885.
182 Checklist of the Plants of the Guiana Shield
Kj-al, R. 1994. Xyridaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 15: 3-90.
Krai, R. 2005. Xyridaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 9:
526-574.
Lanjouw, J. 1938. Xyridaceae. In A. Pulle, Flora of Suriname 1(1): 225-248.
Maguire, B. 1958. Xyridaceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part III. Mem.
New York Bot. Gard. 10(1): 1-156.
Maguire, B. & L.B. Smith. 1964. Xyridaceae. In B. Maguire, J.J. Wurdack, & collaborators. Botany of the Guayana Highland
part V. Mem. New York Bot. Gard. 10(5): 1-278.
Mennega, A.M.W., J.C. Lindeman, & A.R.A. Gorts-van Rijn. 1968. Xyridaceae (additions and corrections). In A. Pulle & J.
Lanjouw, Flora of Suriname 1(2): 340-342.
Mori, S.A. 1997. Xyridaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D. Mitchell, eds. 1997. Guide
to the vascular plants of central French Guiana: Part 1 . Pteridophytes, Gymnosperms, and Monocotyledons. Memoirs of the
New York Botanical Garden 76, pt. 1 : 371.
ZiNGIBERACEAE
H. Maas, P.J.M. Maas
Hedychium cownarium J. Koenig [cultivated, naturalized]
Renealmia alpinia (Rottb.) Maas
Renealmia aromatica (Aubl.) Griseb.
Renealmia cernua (Sw. ex Roem. & Schult.) J.F. Macbr.
Renealmiafloribunda K. Schum.
Renealmia guianensis Maas
Renealmia microcalyx Maas & H. Maas
Renealmia monosperma Miq.
Renealmia orinocensis Rusby
Renealmia thyrsoidea (Ruiz & Pav.) Poepp.
Renealmia thyrsoidea subsp. chrysantha Maas
Renealmia thyrsoidea subsp. thyrsoidea
Zingiber spp. [several species cultivated, possibly escaping]
GU SU FG
AM BO DA GU su FG
AM BO DA GU
SU
FG?
AM BO GU su FG
BO DA GU su FG
AM
BO DA GU su FG
BO DA GU su FG
AM GU
GU
su
su
AM
Synonyms and Misidentifications
Alpinia aromatica Aubl. =Renealmia aromatica
Alpinia latifolia Willd. ex Link (Marantaceae) =Calathea latifolia
Alpinia spiralis Jacq. (Costaceae) =Costus spiralis var. spiralis
Alpinia tubulata Ker Gawl. =Renealmia alpinia
Amomum alpinia Rottb. =Renealmia alpinia
Amomum thyrsoideum Ruiz & Pav. =Renealmia thyrsoidea
Renealmia exaltata L. f =Renealmia alpinia
Renealmia gracilis Petersen =Renealmia guianensis
Renealmia humilis Petersen =Renealmia monosperma
Renealmia lutea J.R. Johnst. =Renealmia aromatica
Renealmia monostachia L. (Bromeliaceae) =Guzmania monostachia
Renealmia occidentalis (Sw.) Sweet =Renealmia aromatica
Renealmia occidentalis var. macrocarpa Loes. =Renealmia aromatica
Renealmia occidentalis var. occidentalis ^Renealmia aromatica
Renealmia occidentalis yox.fi cylindrica Petersen =Renealmia aromatica
Renealmia pedicellaris Gleason =Renealmiafloribunda
Renealmia pendula C.F. Gaertn. (Bromeliaceae) =Catopsis berteroniana
Renealmia polystachia L. (Bromeliaceae) =Tillandsia polystachia
Renealmia raja Petersen =Renealmia alpinia
Renealmia recurvata \Qx.fi (Bromeliaceae) =Tillandsia recurvata
Renealmia usneoides (Bromeliaceae) =Tillandsia usneoides
References
Maas, P.J.M. 1977. Renealmia. Flora Neotropica Monogr. 18: 1-161.
Maas, RJ.M. 1979. Zingiberaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 416-441.
Maas, P.J.M. 1985. Zingiberaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A).
Maas, P. & H. Maas-van de Kamer. 1997. Zingiberaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, M. Hoff, & J.D.
Mitchell, eds. 1997. Guide to the vascular plants of central French Guiana: Part 1. Pteridophytes, Gymnosperms, and
Monocotyledons. Memoirs of the New York Botanical Garden 76, pt. 1: 371-374.
Maas-van de Kamer, H. & P. Maas. 2005. Zingiberaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of
the Venezuelan Guayana 9: 576-580.
Checklist of the Plants of the Guiana Shield 183
ZOSTERACEAE
Zostera marina L. [record by Lemee extremely dubious] FG?
References
Phillips, R.C. & E.G. Menez. 1988. Seagrasses. Smithsonian Contrib. Marine Sci. 34: 1-104.
DICOTYLEDONEA
ACANTHACEAE
D. Wasshausen
Anisacanthus secundus Leonard
Aphelandra aurantiaca (Schweidw.) Lindl.
Aphelandra aurantiaca var. aurantiaca
Aphelandrafasciculata Wassh.
Aphelandra pilosa Leonard
Aphelandra pulcherrima (Jacq.) Kunth
Aphelandra scabra (Vahl) Sm.
Asystasia gangetica (L.) T. Anderson [cultivated, persists as an escape]
Barleria cristata L.
Barleria lupulina Lindl. [introduced, possibly naturalized]
Blechum pyramidatum (Lam.) Urb.
Bravaisia integerrima (Spreng.) Standi.
Dicliptera ciliaris J. Juss.
Dicliptera granvillei Wassh.
Dicliptera knappii Wassh.
Dicliptera sexangularis (L.) J. Juss. [expected in GU]
Elytraria imbricata (Vahl) Pers.
Eranthemum pulchellum Andr. [cultivated, naturalized]
Gynocraterium guianense Bremek.
Hemigraphis alternata (Burm. f.) T. Anderson [cultivated, persists as an
escape]
Herpetacanthus rotundatus (Lindau) Bremek.
Hygrophila costata Nees
Hygrophila erecta (Burm. f.) Hochr. [introduced]
Hygrophila sandwithii Bremek.
Hygrophila surinamensis Bremek. [expected in GU]
Justicia betonica L. [introduced]
Justicia breteleri Wassh.
Justicia calycina (Nees) V.A.W. Graham
Justicia carthaginensis Jacq.
Justicia cataractae Leonard
Justicia cayennensis (Nees) Lindau
Justicia cayennensis var. cayennensis
Justicia cayennensis var. parvifolia Bremek.
Justicia coma ta (L.) Lam.
Justicia coppenamensis Wassh. [nom. nov. for Beloperone modesta Bremek.]
Justicia delascioi Wassh. AM
Justicia guianensis (N.E. Br.) Wassh.
Justicia huberi Wassh.
Justicia hylaea Leonard AM
Justicia kunhardtii Leonard
Justicia laevilinguis (Nees) Lindau [expected in GU] BO DA SU FG
Justicia maguirei Wassh.
Justicia mcdowellii Wassh.
Justicia monachinoi Wassh.
Justicia moritziana Wassh.
Justicia nuriana Wassh.
Justicia oldemanii Wassh. FG
Justicia panarensis Wassh.
Justicia parguazensis Wassh.
BO GU SU FG
GU SU FG
GU SU FG
AM
AM BO
BO DA GU SU
AM BO GU SU
AM DA
DA
DA
GU
FG
AM BO
BO
DA GU SU FG
GU? SU FG
FG
GU
SU FG
BO
GU
SU
SU
SU FG
BO GU
GU
SU
SU
FG
AM BO
GU
SU
AM
AM BO DA GU SU FG
BO GU SU FG
AM
GU SU FG
GU SU
SU
FG
AM BO DA GU SU
SU
FG
BO GU
BO
BO
BO
GU
GU
BO GU
BO DA
BO
BO
BO
AM
AM
AM BO GU SU FG
AM BO su FG
GU SU FG
FG
AM BO DA GU
AM BO DA GU su FG
FG
BO
BO
BO
BO
BO
BO
BO
SU FG
SU
SU FG
SU
SU FG
SU
FG?
184 Checklist of the Plants of the Guiana Shield
Justicia pahmensis Wassh.
Justicia paruana Wassh.
Justicia pectoralis Jacq.
Justicia polystachya Lam. [expected in GU]
Justicia potarensis (Bremek.) Wassh.
Justicia prevostiae Wassh.
Justicia schomburgkiana (Nees) V.A.W. Graham
Justicia secunda Vahl
Justicia sprucei V.A.W. Graham
Justicia uvida Wassh. BO
Justicia wilhelminensis Wassh. SU
Justicia wurdackii Leonard
Lepidagathis alopecuroidea (Vahl) R. Br. ex Griseb. AM DA GU
Lepidagathis gracilis (Bremek.) Wassh. [expected in GU]
Lepidagathis medicaginea (Bremek.) Wassh. [expected in GU]
Lepidagathis nickeriensis Wassh. [expected in GU]
Lepidagathis surinamensis (Bremek.) Was^. GU
Megaskepasma erythrochlamys Lindau
Nelsonia campestris R. Br.
Odontonema album V.M. Baum
Odontonema bracteolatum (Jacq.) Kuntze
Odontonema liesneri Wassh.
Odontonema mazaninensis Wassh. GU
Odontonema schomburgkianum (Nees) Kuntze GU
Pachystachys coccinea (Aubl.) Nees
Pachystachys spicata (Ruiz & Pav.) Wassh. AM
Polylychnis fulgens Bremek.
Polylychnis radicans (Nees) Wassh.
Pseuderanthemum maguirei Leonard AM
Pulchranthus congestus (Lindau) V.M. Baum, Reveal & Nowicke
Pulchranthus surinamensis (Bremek.) V.M. Baum, Reveal & Nowicke
Pulchranthus variegatus (Aubl.) V.M. Baum, Reveal & Nowicke AM
Ruellia bolivarensis Wassh. BO
Ruellia delascioi Wassh. AM
Ruellia geminiflora Kunth AM BO
Ruellia geminiflora van angustifolia (Nees) Griseb. AM BO
Ruellia geminiflora var. geminiflora
Ruellia inflata Rich.
Ruellia liesneri Wassh. AM
Ruellia longifolia Rich. SU FG
Ruellia macrophylla Vahl BO
Ruellia malaca Leonard AM BO
Ruellia menthoides (Nees) Hiem AM
Ruellia microcalyx (Nees) Lindau
Ruellia prostrata Poir.
Ruellia rubra Aubl. [includes several varieties without combinations in
Ruellia]
Ruellia saiilensis Wassh.
Ruellia sprucei Lindau AM
Ruellia steyermarkii Wassh. BO
Ruellia tuberosa L. [naturalized] BO
Ruellia violacea Aubl.
Ruellia wurdackii Wassh. AM BO
Sanchezia lutea Leonard GU
Sanchezia pennellii Leonard DA
Staurogynefockeana Bremek. SU
Staurogyne miqueliana Kuntze SU
Staurogyne repens (Nees) Kuntze GU
Staurogyne spraguei Wassh. AM BO GU
Staurogyne stahelii Bremek. SU
Staurogyne stolonifera (Nees) Kuntze SU
Staurogyne trinitensis Leonard BO DA GU SU FG
Staurogyne wullschlaegeliana Bremek. SU
Teliostachya cataractae Nees [=Lepidagathis sp.] GU FG
GU FG
GU FG
SU FG
GU
SU
FG
FG
SU FG
GU SU FG
GU SU FG
GU SU FG
SU FG
GU SU
GU
GU SU FG
FG
GU SU FG
FG
SU
SU
GU
GU SU
SU
SU
GU SU
GU SU
GU SU
Checklist of the Plants of the Guiana Shield 185
Teliostachya coppenamensis Bremek. [=Lepidagathis sp.]
Teliostachya lanceolata Nees [=Lepidagathis sp.]
Teliostachya mazaninensis Bremek. [=Lepidagathis sp.]
Thimbergia alata Bojer ex Sims [cultivated, naturalized]
Thimbergia erecta (Benth.) T. Anderson [cultivated, naturalized]
Thimbergiafragrans Roxb. [cultivated, naturalized]
Thimbergia grandiflora (Roxb. ex Rottl.) Roxb. [cultivated, naturalized]
Trichanthera gigantea (Bonpl.) Nees
Trichanthera gigantea var. guianensis Gleason
Synonyms and Misidentifications
Adhatoda carthaginensis (Jacq.) Nees =Justicia carthaginensis
Amphiscopia lancifolia var. minor Nees =Justicia moritziana
Anisacanthiis boliviensis (Nees) Wassh. [not in flora area; records referable to A. secimdiis]
Aphelandra deppeana Schltdl. & Cham. =Aphelandra scabra
Aphelandra inserta Leonard =Aphelandra pulcherrima
Aphelandra paraensis Lindau =Aphelandra aurantiaca
Aphelandra pectinata Willd. ex Nees =Aphelandra scabra
Aphelandra tetragona (Vahl) Nees [not in flora area]
Arrhostoxylum coccineum Nees [not in flora area]
Arrhostoxylum rubrum (Aubl.) Nees =Ruellia rubra
Asystasia coromandeliana Nees =Asystasia gangetica
Asystasia nemoriim Nees =Asystasia gangetica
Barleria pyramidata Lam. =Blechum pyramidatum
Beloperone calycina Nees =Justicia calycina
Beloperone modesta Bremek. =Justicia coppenamensis
Beloperone schombiirgkiana Nees =Justicia schombiirgkiana
Blechum bronmei J. Juss. =Blechum pyramidatum
Blechum brownei f. puberulum Leonard =Blechum pyramidatum
Bravaisiafloribunda DC. =Bravaisia integerrima
Chaetochlamys ciliata Lindau =Justicia sprucei
Chaetochlamys wurdackii Leonard =Justicia parguazensis
Dactylostegium assurgens (L.) Bremek. =Dicliptera sexangularis
Dianthera acuminatissima (Miq.) Benoist =Justicia calycina
Dianthera calycina (Nees) Benth. ex B.D. Jacks. =Justicia calycina
Dianthera comata L. =Justicia comata
Dianthera guianensis N.E. Br. =Justicia guianensis
Dianthera pectoralis (Jacq.) J.F. Gmel., nom. illeg. =Justicia pectoralis
Dicliptera assurgens (L.) J. Juss. =Dicliptera sexangularis
Dicliptera vahliana Nees =Dicliptera sexangularis
Dipteracanthus angustifolius (Nees) Bremek. =Ruellia geminiflora var. angustifolia
Dipteracanthus canescens Nees =Ruellia geminiflora var. angustifolia
Dipteracanthus geminiflorus (Kunth) Nees =Ruellia geminiflora
Dipteracanthus geminiflorus var. angustifolius Nees =Ruellia geminiflora var. angustifolia
Dipteracanthus menthoides Nees =Ruellia menthoides
Dipteracanthus paniculatus Nees ? [not in flora area] =Ruellia sp.
Dipteracanthus surinamensis Miq. =Ruellia microcalyx
Dipteracanthus vindex Nees =Ruellia geminiflora var. angustifolia
Dipteracanthus violaceus (Nees) Lemee =Ruellia violacea
Drejera boliviensis Nees =Anisacanthus secundus p.p.
Duvernoia potarensis Bremek. =Justicia potarensis
Duvernoia potarensis var. tetrapora Bremek. =Justicia potarensis
Dyspemptemorion cayennense (Nees) Bremek. =Justicia cayennensis
Ebermaiera repens Nees =Staurogyne repens
Ebermaiera spp. =Staurogyne spp.
Eranthemum nervosum (Vahl) R. Br. =Eranthemum pulchellum
Fittonia sp. [GU record =Justicia sp. nov., fide Wasshausen]
Hygrophila guianensis Nees ex Benth. =Hygrophila cosfata
Jacobinia venezuelica Lindau =Justicia guianensis
Juruasia rotundata Lindau =Herpetacanthus rotundatus
Justicia "divergens (Nees) Lindau" [invalid name, records based on misidentifications]
Justicia acuminatissima (Miq.) Bremek. =Justicia calycina
Justicia bracteolata Jacq. =Odontonema bracteolatum
Justicia caripensis Kunth =Justicia secunda
186 Checklist of the Plants of the Guiana Shield
Justicia coccinea Aubl. =Pachystachys coccinea
Justicia gangetica L. =Asystasia gangetica
Justicia glabribracteata (Lindau) V.A.W. Graham [records apparently in error]
Justicia imbricata Vahl =Elytraria imbricata
Justicia martiana (Nees) Lindau =Justicia comata
Justicia mohcandiana (Nees) Lemee =Justicia secunda
Justicia obtusifolia (Nees) Lindau =Justicia laevilinguis
Justicia pectoralis var. latifolia Bremek. =Justicia pectoralis
Justicia plectica Leonard =Justicia schomburgkiana
Justicia pulcherrima Jacq. =Aphelandra pulcherrima
Justicia scabra Vahl =Aphelandra scabra
Justicia spicata Ruiz & Pav. =Pachystachys spicata
Justicia variegata Aubl. =Pulchranthus variegatus
Leptostachya martiana var. hispida Nees =Justicia comata
Meyenia erecta Benth. =Thunbergia erecta
Nelsonia canescens (Lam.) Spreng. =Nelsonia campestris
Odontonema surinamense Bremek. =Pulchranthus surinamensis
Odontonema variegatum (Aubl.) Kuntze =Pulchranthus variegatus
Odontonema venezuelense Leonard =Odontonema bracteolatum
Onychacanthus arboreus H. Karst. =Bravaisia integerrima
Pachystachys riedeliana Nees =Pachystachys spicata
Polylychnis essequibensis Bremek. =Polylychnis radicans
Psadacocalymma spp. =Justicia spp.
Rhacodiscus acuminatissimus (Miq.) Lindau =Justicia calycina
Rhacodiscus calycinus (Nees) Bremek. =Justicia calycina
Rhacodiscus intermedius (Nees) Lindau =Justicia calycina
Rhacodiscus moricandianus (Nees) Bremek. =Justicia secunda
Rhacodiscus secundus (Vahl) Bremek. =Justicia secunda
Rhytiglossa acuminatissima Miq. =Justicia calycina
Rhytiglossa laevilinguis Nees =Justicia laevilinguis
Rhytiglossa moricandiana Nees =Justicia secunda
Rhytiglossa secunda (Vahl) Nees ^Justicia secunda
Rhytiglossa secunda var. intermedia Nees =Justicia calycina
Ruellia alopecuroidea Vahl =Lepidagathis alopecuroidea
Ruellia blechum L. -Blechum pyramidatum
Ruellia cordifolia Lindau =Ruellia inflata
Ruellia gigantea Bonpl. =Trichanthera gigantea
Ruellia pipericaulis Rizzini =Ruellia inflata
Ruellia radicans (Nees) Lindau =Polylychnis radicans
Ruellia tubiflora Kunth [not in flora area, based on misidentifications]
Ruellia vindex (Nees) Lindau =Ruellia geminiflora var. angustifolia
Sericographis caripensis (Kunth) Nees =Justicia secunda
Staurogyne leptocaulis Leonard =Staurogyne spraguei
Staurogyne linearifolia Bremek. =Staurogyne stolonifera
Staurogyne versteegii Bremek. =Staurogyne trinitensis
Stemonacanthus macrophyllus (Vahl) Nees =Ruellia macrophylla
Teliostachya alopecuroidea (Vahl) Nees =Lepidagathis alopecuroidea
Teliostachya glandulosa Bremek. =Lepidagathis nickeriensis
Teliostachya gracilis Bremek. =Lepidagathis gracilis
Teliostachya medicaginea Bremek. =Lepidagathis medicaginea
Teliostachya surinamensis Bremek. =Lepidagathis surinamensis
Thyrsacanthus bracteolatus (Jacq.) Nees =Odontonema bracteolatum
Thyrsacanthus variegatus (Aubl.) Nees =Pulchranthus variegatus
Ulleria angustifolia (Nees) Bremek. =Ruellia geminiflora var. angustifolia
References
(see also Mendonciaceae)
Baum, V.M. & J.L. Reveal. 1983. Pulchranthus, a new genus from northern South America. Syst. Bot. 8(2): 221-220.
Bremekamp, C.E.B. 1938. Acanthaceae. In A. Pulle, Flora of Suriname 4(2): 166-252.
Bremekamp, C.E.B. 1938. Notes on the Acanthaceae of Surinam. Rec. Trav. Bot. Neerl. 35: 130-171.
Bremekamp, C.E.B. 1938. Acanthaceae. In A. Pulle, Flora of Suriname 4(2): 166-252.
Bremekamp, C.E.B. 1938. Notes on the Acanthaceae of Surinam. Rec. Trav. Bot. Neerl. 35: 130-171.
Graham, V.A.W. 1988. Delimitation and infra-generic classification of Justicia. Kew Bull. 43(4): 551-624.
Leonard, E.C. 1961. Acanthaceae. In B. Maguire, J.J. Wurdack, & collaborators. Botany of the Guayana Highland part IV. Mem.
New York Bot. Gard. 10(4): 47-58.
Checklist of the Plants of the Guiana Shield 187
Wasshausen, D.C. 1975. The genus Aphelandra. Smiths. Contrib. Bot. 18: 1-157.
Wasshausen, D.C. 1990. Justicia maguirei, a new species of Acanthaceae from Guyana. Mem. New York Bot. Gard. 64: 235-
238.
Wasshausen, D.C. 1991. New species of Dicliptera from the Guianas. Brittonia 43(1): 1-6.
Wasshausen, D.C. 1992a. New species and new combinations of Justicia (Acanthaceae) from the Venezuelan Guayana. Novon
2: 62-80.
Wasshausen, D.C. 1992b. Nomenclatural changes in the Acanthaceae and two new species of Odontonema from Venezuela and
Guyana. Novon 2: 149-152.
Wasshausen, D.C. 1995. Acanthaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 335-
374.
Wasshausen, D.C. 1995. A new species, new combinations, and a new name in Acanthaceae from northern South America.
Brittonia 47(4): 422-425.
Wasshausen, D.C. 2002. Acanthaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 31-39.
AlZOACEAE
R.A. DeFilippst, J.A. Steyermarkf
Sesiivium portidacastrum (L.) L. DA GU SU FG
Synonyms and Misidentipications
Sesuvium acutifolhim Miq. =Sesuvium portidacastrum
REFERENCEs(see also Molluginaceae)
Burger, W. 1983. Aizoaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 199-212.
Eyma, RJ. 1934. Aizoaceae. In A. Pulle, Flora of Suriname 1(1): 158-161.
Nevling, L.I. Jr. 1961. Aizoaceae. InR.E. Woodson Jr., R.W. Schery, & collaborators. Flora ofPanama. Ann. Missouri Bot. Gard.
48(1): 80-85.
Amaranthaceae
R.A. DeFilippst, M.H. Nee
Achyranthes indica (L.) Mill, [naturalized]
Alternanthera brasUiana (L.) Kuntze
Alternanthera halimifolia (Lam.) Standi. {^Alternanthera tenella?]
Alternanthera littoralis P. Beauv.
Alternanthera littoralis var. maritima (Mart.) Pedersen
Alternanthera paronychioides A. St.-Hil. [specimens misidentified
Alternanthera tenella in the Guianas]
Alternanthera philoxeroides (Mart.) Griseb.
Alternanthera pulchella KunXh
Alternanthera sessilis (L.) R. Br. ex DC.
Alternanthera tenella Colla
Amaranthus australis (A. Gray) J.D. Sauer [introduced?]
Amaranthus blitum L. [introduced?]
Amaranthus caudatus L. [cultivated or escaped from cuUivation]
Amaranthus dubius Mart, ex Thell.
Amaranthus hybridus L.
Amaranthus spinosus L. [introduced?]
Amaranthus tricolor L. [cultivated only?]
Amaranthus viridis L. [introduced?]
Blutaparon vermiculare (L.) Mears
Celosia argentea L. [cultivated, naturalized]
Celosia virgata Jacq.
Chamissoa altissima (Jacq.) Kunth
Chamissoa altissima var. altissima
Chamissoa altissima var. swansonii Sohmer
Cyathula achyranthoides (Kunth) Moq. [introduced?]
Cyathula prostrata (L.) Blume [introduced?]
Froelichia humboldtiana (Schult.) Seub.
Gomphrena globosa L. [widely cultivated and escaping]
Iresine diffusa Humb. & Bonpl. ex Willd.
Pfaffia glabrata Mart.
BO DA
AM
BO
GU SU FG
GU SU FG
GU SU FG
FG?
FG?
DA GU SU FG
BO
BO GU SU FG
GU SU FG
DA GU SU FG
BO DA GU SU
SU
FG
FG
AM BO DA GU SU
SU
FG
FG
BO GU SU
SU
FG
FG
BO DA GU SU FG
DA GU SU FG
AM BO
DA
GU SU FG
BO DA GU
GU
SU
SU
SU
AM BO DA GU SU FG
BO DA GU SU FG
BO
AM BO DA GU SU FG
AM BO DA GU SU
FG?
i: Checklist of the Plants of the Guiana Shield
Pfaffia glabrata var. rostrata Stiitzer
Pfaffia glomerata (Sprang.) Pedersen
Pfaffia grandiflora (Hook.) R.E. Fr.
Pfaffia iresinoides (Kunth) Sprang.
AM
FG?
GU SU FG
DA GU
Synonyms and Misidentifications
Achyranthes altissima Jacq. =Chamissoa altissima var. altissima
Achyranthes aspera var. indica L. =Achyranthes indica
Achyranthes brasiliana (L.) Standi. =Alternanthera brasiliana
Achyranthes corymbosa L. (Caryophyllaceae) =Polycarpaea corymbosa
Achyranthes halimifolia Lam. ^^Alternanthera halimifolia
Achyranthes philoxeroides (Mart.) Standi. = Alternanthera philoxeroides (Mart.) Griseb.
Achyranthes prostrata L. =Cyathula prostrata
Achyranthes sessilis (L.) Desf. ex Steud. =Alternanthera sessilis
Acnida australis A. Gray =Amaranthus australis
Acnida cuspidata Bertero ex Spreng. =Amaranthus australis
Alternanthera crucis Bouldingh =Alternanthera halimifolia
Alternanthera dentata (Moench) Scheyg. =Alternanthera brasiliana
Alternantheraficoidea (L.) Sm. =Alternanthera paronychioides
Alternantheraficoidea (L.) R. Br. ex Roem. & Schult. [name often applied to A. tenella specimens] =Alternanthera
paronychioides
Alternantheraflavogrisea (Urb.) Urb. ^Alternanthera halimifolia
Alternantheraflavogrisea subsp. diffusa (Mart.) Mears ex Foum. =Alternanthera tenella
Alternanthera iresinoides Kunth =Pfaffia iresinoides
Alternanthera maritima (Mart.) A. St.-Hil. ^Alternanthera littoralis var. maritima
Alternanthera ramosissima R.E. Fr. =Alternanthera brasiliana
Amaranthus gracilis Desf. =Amaranthus viridis
Amaranthus lividus L. =Amaranthus blitum
Amaranthus paniculatus L. =Amaranthus hybridus
Bucholzia philoxeroides Mart. =Alternanthera philoxeroides
Caraxeron vermicularis (L.) Raf. =Blutaparon vermiculare
Celosia cristata L. =Celosia argentea
Celosia paniculata L. =Iresine diffusa
Desmochaeta achyranthoides Kunth =Cyathula achyranthoides
Froelichia lanata (Kunth) Moq. =Froelichia humboldtiana
Gomphrena brasiliana L. =Alternanthera brasiliana
Gomphrena humboldtiana Schult. =Froelichia humboldtiana
Gomphrena iresinoides (Kunth) Moq. =Pfaffia iresinoides
Gomphrena lanata Kunth =Froelichia humboldtiana
Gomphrena sessilis L. =Alternanthera sessilis
Gomphrena stenophylla Spreng. =Pfaffia glomerata
Gomphrena vermicularis L. =Blutaparon vermiculare
Hebanthe hookeriana Hemsl. =Pfaffia grandiflora
Illecebrum diffusum Schult. =Alternanthera pulchella
Iresine celosia L. =Iresine diffusa
Iresine celosioides L. =Iresine diffusa
Iresine glomerata Spreng. =Pfaffia glomerata
Iresine grandiflora Hook, (non R.E. Fr.) =Pfaffia grandiflora
Iresine polymorpha Mart. =Iresine diffusa
Iresine vermicularis (L.) Moq. ^Blutaparon vermiculare
Pfaffia glauca (Mart.) Spreng. ^Pfaffia glomerata
Pfaffia hookeriana (Hemsl.) Greenm. =Pfaffia grandiflora
Pfaffia stenophylla (Spreng.) Stuchlik =Pfaffia glomerata
Philoxerus vermicularis (L.) R. Br. =Blutaparon vermiculare
Serturnera glauca Mart. =Pfaffia glomerata
Telanthera maritima Moq. =Alternanthera littoralis var. maritima
References
Burger, W. 1983. Amaranthaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 142-180.
DeFilipps, R.A. and S.L. Maina. 2002. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 40-42.
Duke, J.A. 1961. Amaranthaceae. In R.E. Woodson, R.W. Schery, & collaborators, Flora of Panama 4(4): 348-392.
Kellogg, E.A. 1988. Amaranthaceae. In R.A. Howard, Flora of the Lesser Antilles.
Checklist of the Plants of the Guiana Shield 189
Mears, J.A. 1977. The nomenclature and type collections of the widespread taxa of Altemanthera. Proc. Acad. Nat. Sci. Phila.
129: 1-21.
Morros, M.E., B. Trujillo, & M. Ponce. 1990. Descripcion del genero Amaranthus L. con 3 nuevos registros para Venezuela y
consiguiente clave para las especies. Emstia 58-59-60: 45-51.
Nee, M.H. 1995. Amaranthaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 384-399.
Pedersen, T.M. 1967. Studies in South American Amaranthaceae. Darwiniana 14(2-3): 430-462.
Pedersen, T.M. 1976. Estudios sobre Amaranthaceae Sudamericanas, II. Darwiniana 20(1-2): 269-303.
Scheygrond, A. 1932. Amarantaceae. In A. Pulle, Flora of Suriname 1(1): 25-44.
Sohmer, S.H. 1977. Revision of Chamissoa. Bull. Torr. Bot. Club 104: 111-126.
Anacardiaceae
J.D. Mitchell
Anacardium amapaense J.D. Mitch. FG
Anacardiumfruticosum J.D. Mitch. & S.A. Mori
Anacardium giganteum W. Hancock ex Engl.
Anacardium occidentale L.
Anacardium spruceanum Benth. ex Engl.
Astroniumfraxinifolium Schott
Astronium graveolens Jacq.
Astronium lecointei Ducke
Astronium ulei Mattick
Campnosperma gummiferum (Benth.) Marchand
Cyrtocarpa velutinifolia (R.S. Cowan) J.D. Mitch. & Daly
Loxopterygium sagotii Hook, f
Mangifera indica L. [widely cultivated]
Spondias dulcis Parkinson
Spondias mombin L.
Spondias purpurea L.
Tapirira bethanniana J.D. Mitch.
Tapirira guianensis Aubl.
Tapirira obtusa (Benth.) J.D. Mitch.
Thyrsodium guianense Sagot ex Marchand
Thyrsodium herrerense Encamacion
Thyrsodium puberulum J.D. Mitch. & Daly
Thyrsodium spruceanum Benth.
Synonyms and Misidentifications
Anacardium microcarpum Ducke =Anacardium occidentale
Comocladia tapaculo Kunth =Tapirira guianensis
Cyrtospermum gummiferum Benth. =Campnosperma gummiferum
Drepanospermum gummiferum (Benth.) Benth. =Campnosperma gummiferum
Garuga gigantea Engl. =Thyrsodium spruceanum
Garuga guianensis (Sagot) Engl. =Thyrsodium guianense
Garuga schomburgkiana (Benth.) Engl. =Thyrsodium spruceanum
Garuga spruceana (Benth.) Engl. ^Thyrsodium spruceanum
Loxopterygium gutierrezii F.A. Barkley =Cyrtocarpa velutinifolia
Mauria multiflora Mart, ex Benth. =Tapirira guianensis
Mauria obtusa Benth. =Tapirira obtusa
Spondias cytherea Sonn. =Spondias dulcis
Spondias lutea L. =Spondias mombin
Tapirirafanshawei Sandwith ^Tapirira guianensis
Tapirira guianensis van cuneata Engl. =Tapirira guianensis
Tapirira guianensis van elliptica Engl. ^'Tapirira guianensis
Tapirira marchandii Engl. [nom. illeg.] =Tapirira obtusa
Tapirira myriantha Triana & Planch. =Tapirira guianensis
Tapirira pao-pombo van major Marchand =Tapirira obtusa
Tapirira peckoltiana Engl. =Tapirira obtusa
Tapirira velutinifolia (R.S. Cowan) Marc.-Berti "^Cyrtocarpa velutinifolia
Thyrsodium dasytrichum Sandwith =Thyrsodium spruceanum
Thyrsodium giganteum (Engl.) Engl. =Thyrsodium spruceanum
Thyrsodium paraense Huber =Thyrsodium spruceanum
Thyrsodium schomburgkianum Benth. =Thyrsodium spruceanum
GU
AM BO GU SU
AM BO GU SU FG
AM BO
BO
DA
GU SU
SU
SU
FG
AM BO GU SU FG
AM
BO GU
BO DA GU SU FG
AM BO DA GU SU FG
BO DA GU FG
AM BO DA GU
GU
SU FG
FG
FG
AM BO DA GU SU FG
AM BO DA GU SU FG
GU? SU FG
BO
SU FG
AM BO DA GU SU FG
190 Checklist of the Plants of the Guiana Shield
References
Barkley, F.A. 1968. Anacardiaceae: Rhoideae: Astronium. Phytologia 16: 107-152.
Jansen-Jacobs, M.J. 1976. Anacardiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2):
441-444.
Mitchell, J.D. 1992. Additions to Anacardium. Anacardium amapaense, a new species from French Guiana and eastern Amazonian
Brazil. Brittonia 44(3): 331-338.
Mitchell, J.D. 1993. Tapirira obtusa comb. nov. (Anacardiaceae). Novon 3(1): 66.
Mitchell, J.D. 1995. Anacardiaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 399-412.
Mitchell, J.D. 1997. Anacardiaceae. In A.R.A. Gorts-van Rijn & M.J. Jansen-Jacobs, Flora of the Guianas (series A) 129: 1-79.
Mitchell, J.D. 2002. Anacardiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
43-53.
Mitchell, J.D. & D.C. Daly. 1991. Cyrtocarpa Kunth (Anacardiaceae) in South America. Ann. Missouri Bot. Gard. 78: 184-189.
Mitchell, J.D. & D.C. Daly. 1993. A revision of Thyrsodium. Brittonia 45(2): 115-129.
Mitchell, J.D. & S. Mori. 1987. The cashew and its relatives (Anacardium: Anacardiaceae). Mem. New York Bot. Gard. 42: 1-
76.
Nannenga, E.T. 1934. Anacardiaceae. In A.^Pulle, Flora of Suriname 2(1): 132-145.
Anisophylleaceae
Anisophyllea guianensis Sandwith GU
References
A historical and taxonomic synopsis of Rhizophoraceae and Anisophylleaceae. Ann.Juncosa, A.M. & RB. Tomlinson. 1988.
Missouri Bot. Gard. 75: 1278-1295.
Sandwith, N.Y 1952. Contributions to the flora of tropical America LV. Discovery ofAnisophyllea in America. Kew Bull. 1952:
303-306.
Annonaceae
RE. Berry, D.M. Johnson, P.J.M. Maas, N.A. Murray, H. Rainer, J.A. Steyermarkf
Anaxagorea acuminata (Dunal) A. DC.
Anaxagorea angiistifolia Timmerman
Anaxagorea brachycarpa R.E. Fr.
Anaxagorea brevipedicellata Timmerman
Anaxagorea brevipes Benth.
Anaxagorea dolichocarpa Sprague & Sandwith
Anaxagorea gigantophylla R.E. Fr.
Anaxagorea inundata RE. Berry & R.B. Mill.
Anaxagorea petiolata R.E. Fr.
Anaxagorea phaeocarpa Mart.
Anaxagorea prinoides (Dunal) A. DC.
Anaxagorea rheophytica Maas & Westra
Anaxagorea riifa Timmerman
Annona ambotay Aubl.
Annona atabapensis Kunth
Annona densicoma Mart.
Annona echinata Dunal
Annonafoetida Mart.
Annona gigantophylla (R.E. Fr.) R.E. Fr.
Annona glabra L.
Annona haematantha Miq.
Annona hypoglaiica Mart.
Annonajahnii Saff.
Annona montana Macfad.
Annona muricata L.
Annona paludosa Aubl.
Annona prevostiae H. Rainer
Annona purpurea M09. & Sesse ex Dunal
Annona sericea Dunal
Annona symphyocarpa Sandwith
Annona temiiflora Mart.
Annona tenuipes R.E. Fr.
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
GU
BO GU
BO DA GU
BO
BO
GU
GU
SU
su
SU
su
su
FG
FG
FG
FG
FG
FG
GU su FG
GU su
su
FG
FG
AM
DA GU su FG
GU su FG
AM
BO
GU su FG
AM BO DA GU su FG
BO DA GU su FG
GU su FG
FG
BO
AM BO GU su FG
AM BO GU FG
FG
GU
su FG
su FG
FG
su FG
Checklist of the Plants of the Guiana Shield 191
Armona tnmciflora R.E. Fr. AM BO FG
Amiona iilei R.E. Fr. GU
Bocageopsis canescem (Spruce ex Benth.) R.E. Fr. AM
Bocageopsis miiUiflora (Mart.) R.E. Fr. AM BO GU
Cardiopetahim siihnamense R.E. Fr.
Cremastosperma brevipes (DC. ex Dunal) R.E. Fr.
Cymbopetalum brasilieme (Veil.) Benth. ex Baill. AM BO DA GU
Diclinanona calycina (Diels) R.E. Fr. AM
Duguetia aberrans Maas AM
DiigiietJa amplexifolia R.E. Fr. SU FG
Duguetia argentea (R.E. Fr.) R.E. Fr. AM
Duguetia cadavehca Ruber
Duguetia calycina Benoist
Duguetia cauliflora R.E. Fr.
Duguetia chrysea Maas
Duguetia decurrens R.E. Fr.
Duguetia dimorphopetala R.E. Fr.
Duguetia eximia Diels
Duguetia flagellaris Huber
Duguetia granvilleana Maas
Duguetia guianensis R.E. Fr.
Duguetia inconspicua Sagot
Duguetia latifolia R.E. Fr.
Duguetia lepidota (Miq.) Pulle
Duguetia lucida Urb.
Duguetia macrocalyx R.E. Fr.
Duguetia macwphylla R.E. Fr.
Duguetia megalophylla R.E. Fr.
Duguetia neglecta Sandwith
Duguetia oligocarpa Maas & van Dam
Duguetia paraensis R.E. Fr.
Duguetia pauciflora Rusby
Duguetia pycnastera Sandwith
Duguetia quitarensis Benth.
Duguetia hberensis Aristeg. ex Maas & Boon
Duguetia rigida R.E. Fr.
Duguetia honegrensis van Zuilen & Maas
Duguetia riparia Huber SU FG
Duguetia sancticawli Maas AM
Duguetia schuhii Jans.-Jac. SU
Duguetia surinamensis R.E. Fr. GU SU FG
Duguetia tenuis R.E. Fr. AM GU
Duguetia trunciflora Maas & A.H. Gentry AM FG
Duguetia uniflora (Dunal) Mart. AM
Duguetia vallicola J.F. Macbr. AM
Duguetia venezuelana R.E. Fr. AM
Duguetia yeshidan Sandwith BO DA GU SU FG
Ephedranthus guianensis R.E. Fr. AM GU SU FG
Fusaealongifolia{kuh\.)S?iii. AM BO GU SU FG
Guatteria atabapensis Aristeg. ex D.M. Johnson & N.A. Murray AM
Guatteria atra Sandwith GU
Guatteria axilliflora (DC.) R.E. Fr. FG
Guatteria brachypoda R.E. Fr.
Guatteria cardoniana R.E. Fr. [cf. Guatteria recurvisepala]
Guatteria citriodora Ducke
Guatteria clusiifolia D.M. Johnson & N.A. Murray
Guatteria conspicua R.E. Fr.
Guatteria discolor R.E. Fr.
Guatteria duckeana var. subcordata R.E. Fr.
Guatteria dura R.E. Fr.
Guatteria elongata Benth. [Brazilian species? "Rio Negro"]
Guatteria flexilis R.E. Fr.
Guatteria foliosa Benth.
Guatteria ? gracilipes R.E. Fr. SU
GU SU FG
AM GU SU FG
AM
AM
BO GU
GU
GU
su
su FG
AM
GU su
FG
GU su FG
AM GU
AM BO su
AM BO DA GU
GU
AM
BO DA GU
GU
GU
su
FG
FG
BO DA GU
AM BO DA GU su FG
AM
BO
GU
AM BO GU
AM
GU
AM BO GU
GU SU FG
GU
GU SU FG
FG
AM
AM
GU?
BO DA GU
AM BO FG
192 Checklist of the Plants of the Guiana Shield
Guatteria guianensis (Aubl.) R.E. Fr. FG
Guatteria insculpta R.E. Fr. AM
Guatteria ? imignis R.E. Fr. SU
Guatteria inundata Mart. AM
Guatteria latipetala R.E. Fr. AM
Guatteria liesneri D.M. Johnson & N.A. Murray AM BO
Guatteria longicuspis R.E. Fr. AM
Guatteria maguirei R.E. Fr. AM
Guatteria maypurensis Kunth AM BO
Guatteria megalophylla Diels
Guatteria monticola R.E. Fr.
Guatteria oblonga R.E. Fr.
Guatteria obovata R.E. Fr. [Brazilian species? "Rio Negro, Lagoon Pedrero"]
Guatteria ouregou (Aubl.) Dunal
Guatteria ovalifolia R.E. Fr. AM BO DA
Guatteria paludosa R.E. Fr.
Guatteria procera R.E. ¥r.
^
Guatteria ? pteropus Benth.
Guatteria punctata (Aubl.) R.A. Howard
Guatteria recurvisepala R.E. Fr. [cf. Guatteria cardoniana]
Guatteria richardii R.E. Fr.
Guatteria riparia R.E. Fr. AM
Guatteria rubrinervis R.E. Fr. BO DA
Guatteria scandens Ducke
Guatteria schomburgkiana Mart. AM BO
Guatteria stenopetala R.E. Fr. AM
Guatteria subsessilis Mart. AM
Guatteria ? trichostemon R.E. Fr.
Guatteria ? umbonata R.E. Fr.
Guatteria wachenheimi Benoist
Guatteria williamsii R.E. Fr. AM
Guatteriopsis blepharophylla (Mart.) R.E. Fr. BO
Heteropetalum brasiliense Benth. AM
Malmea guianensis R.E. Fr.
Malmea surinamensis Chatrou
Mosannona discolor (R.E. Fr.) Chatrou
Oxandra asbeckii (Pulle) R.E. Fr. AM
Oxandra espintana (Spruce ex Benth.) Baill. AM BO
Oxandra euneura Diels AM
Oxandra guianensis R.E. Fr. GU
Oxandra? krukoffii R.E. ¥r. SU
Oxandra leucodermis (Spruce ex Benth.) Warm. AM
Oxandra surinamensis Jans.-Jac.
Oxandra xylopioides Diels AM
Pseudephedranthus fragrans (R.E. Fr.) Aristeg. AM
Pseudoxandra coriacea R.E. Fr. AM
Pseudoxandra cuspidata Maas
Pseudoxandra guianensis (R.E. Fr.) R.E. Fr.
Pseudoxandra polyphleba (Diels) R.E. Fr. AM
Rollinia cuspidata Mart. AM
Rollinia elliptica R.E. Fr.
Rollinia exsucca (DC. ex Dunal) A. DC. AM
Rollinia mucosa (Jacq.) Baill.
Tetrameranthus duckei R.E. Fr. AM
Trigynaea caudata (R.E. Fr.) R.E. Fr.
Trigynaea duckei (R.E. Fr.) R.E. Fr.
Unonopsis glaucopetala R.E. Fr. AM
Unonopsis grandis (Benth.) R.E. Fr. AM
Unonopsis guatterioides (A. DC.) R.E. Fr.
Unonopsis perrottetii (A. DC.) R.E. Fr.
Unonopsis rufescens (Baill.) R.E. Fr.
Unonopsis stipitata Diels AM
Unonopsis velutina Maas AM
Xylopia amazonica R.E. Fr. AM
Xylopia aromatica (Lam.) Mart. AM
GU
GU
GU
SU? FG
GU?
FG
GU
GU
GU SU FG
SU FG
GU SU FG
GU FG
FG
GU
GU SU FG
GU SU FG
GU
SU
GU
GU SU FG
SU FG
GU
SU
GU
GU SU FG
SU
GU SU FG
FG
GU FG
BO GU
GU
SU
FG
FG
FG
BO DA GU SU FG
BO GU
GU
GU
SU
SU
FG
BO DA GU SU FG
BO GU
GU
SU FG
FG
GU SU FG
GU SU FG
BO
BO SU
BO DA GU SU FG
Checklist of the Plants of the Guiana Shield 193
Xylopia benthamii R.E. Fr.
Xylopia calophylla R.E. Fr.
Xylopia cayennensis Maas
Xylopia crinita R.E. Fr.
Xylopia discreta (L. f.) Sprague & Hutch.
Xylopia emarginata Mart.
Xylopia excellens R.E. Fr.
Xylopiafrutescens Aubl.
Xylopia ligustrifolia Humb. & Bonpl. ex Dunal
Xylopia nervosa (R.E. Fr.) Maas
Xylopia nitida Dunal
Xylopia parviflora Spruce
Xylopia plowmanii RE. Berry & D.M. Johnson
Xylopia pulcherrima Sandwith
Xylopia sericea A. St.-Hil.
Xylopia spruceana Benth. ex Spruce
Xylopia surinamensis R.E. Fr.
Xylopia venezuelana R.E. Fr.
AM BO
AM BO
AM
AM BO
AM
AM
AM BO
AM
BO
AM
AM
AM BO
AM
DA
DA
GU su FG
GU
GU su FG
FG
GU su FG
FG
GU su FG
FG
GU su FG
FG
GU su FG
GU su FG
GU SU FG
AM BO
Synonyms and Misidentifications
Aberemoa argentea R.E. Fr. =Duguetia argentea
Aberemoa guianensis Aubl. =Guatteria guianensis
Aberemoa longifolia (Aubl.) Baill. =Fusaea longifolia
Anaxagorea mutica R.E. Fr. =Anaxagorea dolichocarpa
Annona axilliflora DC. =Guatteria axilliflora
Annona exsucca DC. ex Dunal =Rollinia exsucca
Annona guaricensis Pittier =Annonajahnii
Annona hostmannii Steud. =Guatteria schomburgkiana
Annonajenmanii Saff. =Annona sericea
Annona lepidota Miq. =Duguetia lepidota
Annona longifolia Aubl. =Fusaea longifolia
Annona manirote Kunth =Annona purpurea
Annona mucosa Jacq. =Rollima mucosa
Annona palustris L. =Annona glabra
Annona punctata Aubl. =Guatteria punctata
Annona rhombipetala Ruiz & Pav. ex G. Don =Fusaea longifolia
Annona uniflora Dunal =Duguetia uniflora
Bocagea asbeckii Pulle =Oxandra asbeckii
Bocagea espintana Spruce ex Benth. =Oxandra espintana
Bocagea leucodermis Spruce ex Benth. =Oxandra leucodermis
Bocagea multiflora Mart. =Bocageopsis multiflora
Cananga ouregou Aubl. =Guatteria ouregou
Cremastosperma guianense R.E. Fr. =Pseudoxandra guianensis
Cremastosperma poiteaui (Diels) R.E. Fr. =Cremastosperma brevipes
Cremastosperma polyphlebum (Diels) R.E. Fr. =Pseudoxandra polyphleba
Cymbopetalum venosum Gleason (Lecythidaceae) =Lecythis corrugata subsp. corrugata
Diclinanona calycina var. macrophylla R.E. Fr. =Diclinanona calycina
Duguetia adiscandra Jans.-Jac. =Duguetia cadaverica
Duguetia cuspidata R.E. Fr. =Duguetia calycina
Duguetia elegans R.E. Fr. =Duguetia calycina
Duguetia friesii Jans.-Jac. =Duguetia cadaverica
Duguetia insculpta R.E. Fr. =Duguetia riparia
Duguetia longifolia (Aubl.) Baill. =Fusaea longifolia
Duguetia obovata R.E. Fr. =Duguetia yeshidan
Duguetia rostrata Rusby =Duguetia pauciflora
Duguetia sandwithii R.E. Fr. =Duguetia yeshidan
Ephedranthus fragrans R.E. Fr. =Pseudephedranthus fragrans
Froesiodendron surinamense (R.E. Fr.) R.E. Fr. =Cardiopetalum surinamense
Fusaea decurrens R.E. Fr. =Fusaea longifolia
Guatteria bernardii R.E. Fr. ^Guatteria schomburgkiana
Guatteria blepharophylla Mart. =Guatteriopsis blepharophylla
Guatteria calva R.E. Fr. =Guatteria maypurensis
Guatteria chrysopetala (Steud.) Miq. =Guatteria punctata
Guatteriaferruginea A. St.-Hil. [not in flora area]
Guatteriaflavovirens R.E. Fr. =Guatteria schomburgkiana
194 Checklist of the Plants of the Guiana Shield
Guatteria heteropetala Benth. =Heteropetaliim brasiliense
Guatteria maypwensis var. attenuata R.E. Fr. =Guatteria maypurensis
Guatteria maypurensis var. pulchra R.E. Fr. =Guatteria maypurensis
Guatteria microsperma R.E. Fr. =Guatteria wachenheimi
Guatteria ouregou Griseb., non Dunal [not in flora area] =Guatteria inuncta
Guatteria ouregou Mart., non Dunal ^Guatteria pteropus
Guatteria podocarpa DC. ex Dunal =Guatteria ouregou
Guatteria sagotiana R.E. Fr. =Guatteria punctata
Guatteria sandwithii R.E. Fr. =Guatteria schomburgkiana
Guatteria sessilis R.E. Fr. =Guatteria schomburgkiana
Guatteria spruceana R.E. Fr. =Guatteria schomburgkiana
Guatteria velezii R.E. Fr. =Guatteria maypurensis
Guatteria wessels-boerii Jans.-Jac. =Guatteria punctata
Heteropetahim spruceanum R.E. Fr. =Heferopetahim brasiliense
Heteropetahmi spruceanum var. longipetahim R.E. Fr. =Heteropetalum brasiliense
Hornschuchia caudata R.E. Fr. =Trigynaea caudata
Hornschuchia duckei R.E. Fr. =Trig}maea<duckei
Hornschuchia guianensis R.E. Fr. =Trig}maea duckei
Malmea discolor R.E. Fr. =Mosannona discolor
Oxandra nervosa R.E. Fr. -Xylopia nervosa
Pseudoxandra lucida R.E. Fr. =Pseudoxandra polyphleba
Raimondia tenuiflora (Mart.) R.E. Fr. =Annona tenuiflora
Rollinia broadwayi var. cuneata R.E. Fr. =Rollinia exsucca
Rollinia cardiantha Diels =Rollinia cuspidata
Rollinia multiflora Splitg. ^Rollinia exsucca
Rollinia puberula A. DC. =Rollinia exsucca
Rollinia pulchrinervia A. DC. =Rollinia mucosa
Rollinia sieberi A. DC. =Rollinia mucosa
Rollinia surinamensis R.E. Fr. =Rollinia exsucca
Trigynaea anastomosans Rusby =Cymbopetalum brasiliense
Trigynaea canescens Spruce ex Benth. =Bocageopsis canescens
Trigynaea grandis Benth. =Unonopsis grandis
Trigynaea guianensis (R.E. Fr.) R.E. Fr. =Trigynaea duckei
Unona crassipetala Dunal =Guatteria ouregou
Unona discreta L. f. =Xylopia discreta
Unonopsis polyphleba Diels =Pseudoxandra polyphleba
Uva brasiliensis (Veil.) Kuntze =Cymbopetalum brasiliense
Uvaria aromatica Lam. =Xylopia aromatica
Uvaria brasiliensis Veil. =Cymbopetalum brasiliense
Uvaria guatterioides A. DC. =Unonopsis guatterioides
Uvaria spectabills DC. =Fusaea longifolia
Xylopia acuminata Dunal =Anaxagorea acuminata
Xylopia calycina Diels =Diclinanona calycina
Xylopia cinerea Sandwith =Xylopia nitida
Xylopia gigantophylla R.E. Fr. =Annona gigantophylla
Xylopia grandiflora A. St.-Hil. ^Xylopia aromatica
Xylopia longifolia (Sagot) R.E. Fr. =Xylopia cayennensis
Xylopia longifolia A. DC. =Xylopia aromatica
Xylopia neglecta (Kuntze) R.E. Fr. =Xylopia parviflora
Xylopia nitida var. longifolia Sagot =Xylopia cayennensis
References
Chatrou, L.W. & R He. 1999. Studies in Annonaceae XXXIII. A revision of Fusaea. (Baill.) Saff. Brittonia 51(2): 181-203.
Fries, R.E. 1930-1939. Revision derArteneiniger Annonaceen-Gattungen. Acta Horti Berg. 10(1): 1-128(1930), 10(2): 129-341
(1931), 12(1): 1-220 (1934), 12(2): 221-288 (1937), 12(3): 289-577 (1939).
Fries, R.E. 1940. Annonaceae. In A. Pulle, Flora of Suriname 2(2): 341-383.
Jansen-Jacobs, M.J. 1976. Annonaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 658-
687.
Johnson, D.M. & N.A. Murray. 1995. Synopsis of the tribe Bocageeae (Annonaceae), with revisions of Cardiopetalum,
Froesiodendron, Trigynaea, Bocagea, and Hornschuchia. Brittonia 47(3): 248-319.
Maas, RJ.M. 1992. Rollinia. Flora Neotropica Monogr. 57: 1-192.
Maas, RJ.M. 1996. Studies in Annonaceae XXX. Monograph of Duguetia: Preliminary notes. Bot. Jahrb. 118(2): 187-227.
Maas, P.J.M., J. Koek-Noorman, & L.Y.Th. Westra. 1993. Studies in Annonaceae XVIIl. New species from the Neotropics and
miscellaneous notes. Bot. Jahrb. 115(1): 77-95.
Checklist of the Plants of the Guiana Shield 195
Maas, P.J.M. & H Maas-van de Kamer. 2002. Annonaceae.In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M.
Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New
York Bot. Gard. 76 (2): 53 - 67.
Maas, P.J.M., E.A. Mennega, & L.Y.Th. Westra. 1994. Studies in Annonaceae XXI. Index to species and infraspecific taxa of
neotropical Annonaceae. CandoUea 49(2): 389-481.
Maas, P.J.M. & L.Y.Th. Westra. 1984-1985. Studies in Annonaceae II. A monograph of the genus Anaxagorea A. St. Hil. Part 1.
Bot. Jahrb. Syst. 105: 73-134. Part 2. Bot. Jahrb. Syst. 105: 145-204.
Steyermark, J.A., P.J.M. Maas, RE. Berry, D.M. Johnson, N.A. Murray, & H. Rainer. 1995. Annonaceae. In J.A. Steyermark,
RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 413-469.
Zuilen, CM. van & P.J.M. Maas. 1994. Studies in Annonaceae XXII. A taxonomic revision of Duguetia A. St. -Hil., sections
Alcmene and Xylopipetalum. Bot. Jahrb. 116(2): 221-242.
Apiaceae
RE. Berry, J.C. Lindeman
BO
AM BO DA GU
GU?
SU FG
AM DA GU SU FG
Eiyngiiim ebracteatiim Lam.
Enngiwn foetidum L.
Hydrocotyle wtiindifolia Roxb. [introduced, no recent records]
Hydrocotyle umbellata L.
References
Berry, RE. 1995. Apiaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 470-471.
Mathias, M.E. 1936. The genus Hydrocotyle in northern South America. Brittonia 2: 201-237.
Mori, S.A. 2002. Apiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 67-68.
Stafleu, RA. 1951. Umbelliferae. In A. Pulle, Flora of Suriname 3(2): 252-256.
Apocynaceae
L. Allorge, M.E. Endress, B.F. Hansen, A.J.M. Leeuwenberg, G.N. Morillo, J.L. Zarucchi
Allamanda cathartica L.
Allamanda doniana Miill. Arg.
Allamanda oenotherifolia Pohl
Allamanda schottii Pohl
Allamanda setulosa Miq.
Allamanda thevetiifolia Miill. Arg.
Ambelania acida Aubl.
Aspidosperma album (Vahl) Benoist ex Pichon
Aspidosperma araracanga Marc.-Ferr.
Aspidosperma carapanauba Pichon
Aspidosperma cnientum Woodson
Aspidosperma cuspa (Kunth) S.F. Blake ex Pittier
Aspidosperma decussatum Woodson
Aspidosperma discolor A. DC.
Aspidosperma excelsum Benth.
Aspidosperma glaucum Monach.
Aspidosperma helstonei Donsel.
Aspidosperma leiicocymosum Kuhlm.
Aspidosperma macrophyllum Miill. Arg.
Aspidosperma macrophyllum subsp. macrophyllum
Aspidosperma macrophyllum subsp. morii L. Allorge
Aspidosperma marcgraviamim Woodson
Aspidosperma neblinae Monach.
Aspidosperma oblongum A. DC.
Aspidosperma pachypterum Miill. Arg.
Aspidosperma pichonianum Woodson
Aspidosperma sandwithianum Markgr.
Aspidosperma schultesii Woodson
Aspidosperma spruceanum Benth. ex Miill. Arg.
Aspidosperma steyermarkii Woodson
Aspidosperma ulei Markgr.
Aspidosperma vargasii A. DC.
Aspidosperma verruculosum Miill. Arg.
AM
AM
AM
BO
BO
BO
BO
AM
AM BO
AM BO
BO
BO
AM BO
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
DA GU
GU
DA
DA
DA
DA
SU
SU
FG
FG
GU SU FG
GU SU FG
FG
GU SU
SU
FG
GU SU FG
SU FG
GU FG
GU
FG
GU SU FG
GU SU FG
GU SU FG
FG
GU SU FG
GU SU
GU SU
196 Checklist of the Plants of the Guiana Shield
Catharanthus roseus (L.) G. Don [widely cultivated, escaping]
Condylocarpon amazonicum (Markgr.) Ducke
Condylocarpon guyanense Desf.
Condylocarpon intermedium Mull. Arg.
Condylocarpon intermedium var. intermedium
Condylocarpon myrtifolium (Miq.) Miill. Arg.
Condylocarpon pubiflorum Miill. Arg.
Couma catingae Ducke
Couma guianensis Aubl.
Couma macrocarpa Barb. Rodr.
Couma rigida Mull. Arg.
Couma utilis (Mart.) Miill. Arg.
Forsteronia acouci (Aubl.) A. DC.
Forsteronia adenobasis Miill. Arg.
Forsteronia affinis Miill. Arg.
Forsteronia diospyrifolia Mull. Arg.
Forsteronia duckei Markgr.
Forsteronia gracilis (Benth.) Miill. Arg.
Forsteronia guyanensis Miill. Arg.
Forsteronia laurifolia (Benth.) A. DC.
Forsteronia lucida Markgr.
Forsteronia obtusiloba Mull. Arg.
Forsteronia paraensis Hansen
Forsteronia schomburgkii A. DC.
Forsteronia umbellata (Aubl.) Woodson
Galactophora calycina (Huber ex Ducke) Woodson
Galactophora colellana Morillo
Galactophora crassifolia (Miill. Arg.) Woodson
Galactophora pulchella Woodson
Galactophora pumila Monach.
Galactophora schomburgkiana Woodson
Galactophora schomburgkiana var. megaphylla Monach.
Galactophora schomburgkiana var. schomburgkiana
Geissospermum argenteum Woodson
Geissospermum laeve (Thunb.) Miers
Geissospermum reticulatum A.H. Gentry AM
Geissospermum sericeum Benth. & Hook. f. ex Miers
Himatanthus articulatus (Vahl) Woodson AM BO DA
Himatanthus attenuatus (Benth.) Woodson
Himatanthus attenuatus var. attenuatus
Himatanthus attenuatus var. obtusifolius (Miill. Arg.) Plumel
Himatanthus bracteatus (A. DC.) Woodson
Himatanthus bracteatus var. bracteatus
Himatanthus bracteatus var. revolutus (Huber) Plumel
Himatanthus drasticus (Mart.) Plumel
Himatanthus fallax (Miill. Arg.) Plumel
Himatanthus obovatus (Mull. Arg.) Woodson
Himatanthus phagedaenicus (Mart.) Woodson AM
Himatanthus semilunatus Markgr. AM
Himatanthus speciosus (Miill. Arg.) Plumel
Himatanthus stenophyllus Plumel
Himatanthus sucuuba (Spruce ex Miill. Arg.) Woodson BO
Hylaea arborescens (Monach.) J.F. Morales AM
Kopsia arborea Blume
Lacmellea aculeata (Ducke) Monach.
Lacmellea arborescens (Miill. Arg.) Markgr.
Lacmelleafloribunda (Poepp.) Benth. & Hook. f.
Lacmellea guyanensis (Miill. Arg.) Monach.
Lacmellea microcarpa (Miill. Arg.) Markgr. AM
Lacmellea pygmaeaMon?ic\\
.
AM
Lacmellea pygmaea var. latifolia Monach. AM
Lacmellea pygmaea var. pygmaea AM
Lacmellea ramosissima (Miill. Arg.) Markgr. BO
Lacmellea utilis (Am.) Markgr. AM GU FG?
Laxoplumeria baehniana Monach. FG
AM BO GU SU FG
AM
GU
SU FG?
FG
DA GU SU FG
DA GU SU FG
DA GU SU FG
AM FG
AM GU
GU SU FG
AM BO GU
AM BO GU
AM BO
AM BO GU SU FG
BO GU FG
AM
BO GU
GU
DA GU SU
BO DA GU SU FG
AM
AM
BO GU
GU
GU
SU
AM FG
AM
AM
AM
AM
AM
AM BO GU
AM
AM BO GU
AM
AM
AM
AM BO
AM BO
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU
SU
GU SU FG
FG
GU SU FG
GU?
SU FG
FG
Checklist of the Plants of the Guiana Shield 197
Macoubea guianemis Aubl.
Macoubea sprucei (Miill. Arg.) Markgr.
Macropharynx spectabilis (Stadelm.) Woodson
Macropharynx strigillosa Woodson
Malouetia calva Markgr. AM
Malouetia duckei Markgr. [expected on or near Sierra de la Neblina]
Malouetiaflavescem (Willd. ex Roem. & Schult.) Mull. Arg. AM
Malouetia glandulifera Miers AM
Malouetia gracilis (Benth.) A. DC.
Malouetia grandiflora Woodson AM
Malouetia molongo M.E. Endress AM
Malouetia naias M.E. Endress AM
Malouetia parvifolia Woodson AM
Malouetia pubescens Markgr.
Malouetia pubescens var. glabra M.E. Endress
Malouetia tamaquarina (Aubl.) A. DC. AM
Malouetia virescens Spruce ex Miill. Arg. AM
Mandevilla anceps Woodson AM
Mandevilla annulariifolia Woodson AM
Mandevilla aracamunensis Morillo AM
Mandevilla benthamii (A. DC.) K. Schum.
Mandevilla caurensis Markgr. AM
Mandevilla duidae (Woodson) Woodson AM
Mandevilla filifolia Monach. AM
Mandevilla hirsuta (Rich.) K. Schum. AM
Mandevilla holstii Morillo AM
Mandevilla huberi Morillo AM
Mandevillajavitensis (Kunth) K. Schum. AM
Mandevilla lancifolia Woodson AM
Mandevilla lasiocarpa (A. DC.) Malme
Mandevilla leptophylla (A. DC.) K. Schum.
Mandevilla obtusifolia Monach. AM
Mandevilla pachyphylla Woodson
Mandevilla rugellosa (Rich.) L. Allorge
Mandevilla rugosa (Benth.) Woodson
Mandevilla scaberula N.E. Br.
Mandevilla scabra (Hoffmanns, ex Roem. & Schult.) K. Schum. AM
Mandevilla spruceana (Miill. Arg.) K. Schum. AM
Mandevilla steyermarkii Woodson AM
Mandevilla subcarnosa (Benth.) Woodson AM
Mandevilla subcarnosa var. angustata Steyerm.
Mandevilla subcarnosa var. subcarnosa AM
Mandevilla subspicata (Vahl) Markgr.
Mandevilla surinamensis (Pulle) Woodson AM
Mandevilla symphitocarpa (G. Mey.) Woodson
Mandevilla tenuifolia (Mik.) Woodson
Mandevilla trianae Woodson
Mandevilla turgida Woodson AM
Mandevilla vanheurckii (Miill. Arg.) Markgr.
Mesechites acutisepala Monach. AM
Mesechites trifida (Jacq.) Miill. Arg. AM
Microplumeria anomala (Mull. Arg.) Markgr. AM
Molongum laxum (Benth.) Pichon AM
Molongum lucidum (Kunth) Zarucchi AM
Mucoa pantchenkoana (Markgr.) Zarucchi
Neocouma parviflora (Markgr.) Zarucchi AM
Neocouma ternstroemiacea (Mull. Arg.) Pierre AM
Nerium oleander L. [cultivated, perhaps escaping]
Odontadeniafunigera Woodson AM
Odontadenia geminata (Hoffmanns, ex Roem. & Schult.) Miill. Arg. AM
Odontadenia glauca Woodson AM
Odontadenia macrantha (Roem. & Schult.) Markgr. AM
Odontadenia nitida (Vahl) Mull. Arg. AM
Odontadenia perrottetii (A. DC.) Woodson
Odontadenia puncticulosa (Rich.) Pulle AM
BO
BO
GU
GU
SU
su
FG
BO GU
DA GU
GU
SU EG
FG?
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
BO
BO DA
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
BO
BO DA
BO
DA
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
su
SU
su
su
su
su
GU
GU SU
SU
FG
SU FG
SU FG
FG
FG
FG
FG
FG
FG
FG
GU SU FG
GU?
GU SU FG
GU SU FG
GU FG
GU SU FG
198 Checklist of the Plants of the Guiana Shield
Odontadenia sandwithiana Woodson
Odontadenia verrucosa (Willd. ex Roem. & Schult.) K. Schum. ex Markgr.
Pacouria guianensis Aubl.
Parahancorniafasciculata (Poir.) Benoist ex Pichon
Parahancornia negroensis Monach. AM
Parahancornia oblonga (Benth. ex Miill. Arg.) Monach. AM
Parahancornia surrogata Zarucchi AM
Plumeria inodora Jacq. AM
Plumeria rubra L. [cultivated & escaped]
Prestonia acutifolia (Benth. ex Miill. Arg.) K. Schum. AM
Prestonia annularis (L. f.) G. Don
Prestonia cayennensis (A. DC.) Pichon
Prestonia coalita (Veil.) Woodson
Prestonia exserta (A. DC.) Standi.
Prestonia Jinitima Woodson AM
Prestonia guianensis Gleason AM
Prestonia ipomaeifolia A. DC. •
Prestonia latifolia Benth.
Prestonia lindleyana Woodson ex Gleason & A.C. Sm. AM
Prestonia megagros (Veil.) Woodson AM
Prestonia perplexa Woodson
Prestonia quinquangularis (Jacq.) Spreng.
Prestonia surinamensis Miill. Arg.
Prestonia tomentosa R. Br. AM
Prestonia vaupesiana Woodson AM
Rauvolfia ligustrina Willd. ex Roem. & Schult.
Rauvolfia pachyphylla Markgr.
Rauvolfia paraensis Ducke
Rauvolfia polyphylla Benth. AM
Rauvolfia tetraphylla L.
Rhabdadenia biflora (Jacq.) Mull. Arg.
Rhabdadenia macrostoma (Benth.) Miill. Arg.
Rhabdadenia pohlii Miill. Arg.
Rhigospira quadrangularis (Miill. Arg.) Miers AM
Salpinctes kalmiifoliiis Woodson AM
Secondatia densiflora A. DC. AM
Spongiosperma cataractarum Zarucchi
Spongiosperma macrophyllum (Miill. Arg.) Zarucchi AM
Spongiosperma oleifolium (Monach.) Zarucchi AM
Spongiosperma riparium (Monach.) Zarucchi AM
Stemmadenia grandiflora (Jacq.) Miers AM
Tabernaemontana albiflora (Miq.) Pulle [=Bonafousia albiflora]
Tabernaemontana angulata Mart, ex Miill. Arg. [=Bonafousia angulata]
Tabernaemontana attenuata (Miers) Urb. [=Anartia meyeri] AM
Tabernaemontana cerea (Woodson) Leeuwenb. [=Anartia cerea]
Tabernaemontana cymosa Jacq. [=Peschiera cymosa]
Tabernaemontana disticha A. DC. [=Bonafousia disticha] AM
Tabernaemontana divaricata (L.) R. Br. ex Roem. & Schult. [cultivated or
escaped from cultivation]
Tabernaemontanaflavicans Willd. ex Roem. & Schult. [=Anartia flavicans] AM
GU
AM GU SU
AM GU SU FG
DA GU SU FG
Tabernaemontana heterophylla Vahl [=Stenosolen heterophyllus]
Tabernaemontana lagenaria Leeuwenb.
Tabernaemontana lorifera (Miers) Leeuwenb. [=Bonafousia lorifera]
Tabernaemontana macrocalyx Miill. Arg. [=Bonafousia macrocalyx,
Bonafousia muelleriana]
Tabernaemontana maxima Markgr.
Tabernaemontana palustris Markgr.
Tabernaemontana pandacaqui Lam. [Asian species, cultivated only?]
Tabernaemontana rupicola Benth. [=Bonafousia rupicola]
Tabernaemontana sananho Ruiz & Pav. [=Bonafousia sananho]
Tabernaemontana siphilitica (L. f.) Leeuwenb. [=Bonafousia siphilitica]
Tabernaemontana undulata Vahl [=Bonafousia undulata]
Thevetia ahouai A. DC.
Thevetia peruviana (Pers.) K. Schum.
AM
AM
BO
BO
BO
BO
DA
GU
GU SU FG
GU SU
GU SU FG
GU FG
GU FG
GU
FG
GU
BO GU?
BO GU?
GU?
GU SU
GU SU FG
BO DA GU SU?
BO GU SU
BO GU
SU FG
FG
BO DA GU SU FG
DA GU SU FG
BO DA GU SU FG
BO
BO GU SU FG
BO
BO GU SU FG
GU SU FG
GU FG
BO DA GU SU FG
BO DA GU SU
BO DA GU
GU SU FG
GU
BO GU? SU? FG?
BO DA GU
GU
SU
SU
SU
FG
FG
BO GU SU FG
AM
AM
FG
AM GU SU FG
AM BO GU SU FG
AM BO DA GU SU FG
AM BO
BO
DA GU SU FG
AM BO DA GU SU FG
Checklist of the Plants of the Guiana Shield 199
Synonyms and Misidentifications
Ahouai nitida (Kunth) Pichon =Thevetia ahouai
Ambelania cucumerina Miers =Ambelania acida
Ambelania cimeata Miill. Arg. =Molongum lucidum
Ambelania lanceolata Markgr. =Spongiosperma oleifolium
Ambelania laxa (Benth.) Miill. Arg. =Molongum laxum
Ambelania lopezii Woodson =Spongiosperma macrophyllum
Ambelania lucida (Kunth) Markgr. =Molongum lucidum
Ambelania macrophylla Miill. Arg. =Spongiosperma macrophyllum
Ambelania oleifolia Monach. =Spongiosperma oleifolium
Ambelania oleifolia var. riparia Monach. ^Spongiosperma riparium
Ambelania pantchenkoana Markgr. =^Mucoa pantchenkoana
Ambelania parviflora Markgr. =Neocouma parviflora
Ambelania quadrangularis Miill. Arg. '=Rhigospira quadrangularis
Ambelania sagotii Miill. Arg. =Ambelania acida
Ambelania tenuiflora Miill. Arg. =Ambelania acida
Ambelania tenuiflora var. tenuiramea Miill. Arg. =Ambelama acida
Ambelania ternstroemiacea (Miill. Arg.) Monach. =Neocouma ternstroemiacea
Amblyanthera crassifolia Miill. Arg. =Galactophora crassifolia
Amblyanthera spruceana Miill. Arg. =Mandevilla spruceana
Anacampta albescens (Rusby) Markgr. =Tabernaemontana undulata
Anacampta disticha (A. DC.) Markgr. =Tabernaemontana disticha
Anacampta echinata (Aubl.) Markgr. [see Tabernaemontana echinata, Peschiera echinata]
Anacampta macrocalyx (Miill. Arg.) Markgr. =Tabernaemontana macrocalyx
Anacampta maxima (Markgr.) Markgr. =Tabernaemontana maxima
Anacampta muelleriana (Mart, ex Miill. Arg.) Markgr. =Tabernaemontana macrocalyx
Anacampta palustris (Markgr.) Markgr. =Tabernaemontana palustris
Anacampta rigida (Miers) Markgr. [not in flora area] =Tabernaemontana muricata Link ex Roem. & Schult.
Anacampta rupicola (Benth.) Markgr. ^Tabernaemontana rupicola
Anartia attenuata (Miers) Markgr. =Tabernaemontana attenuata
Anartia cerea (Woodson) L. Allorge ^Tabernaemontana cerea
Anartiaflavicans (Willd. ex Roem. & Schult.) Miers =Tabernaemontanaflavicans
Anartia meyeri (G. Mey. ex G. Don) Miers =Tabernaemontana attenuata
Anartia oblongifolia (A. DC.) Markgr. =Tabernaemontanaflavicans
Anartia olivacea (Miill. Arg.) Markgr. =Tabernaemontanaflavicans
Anechites amazonica Markgr. =Condylocarpon amazonicum
Anechites nerium Urb. [not in flora area]
Angadenia geminata (Hoffmanns, ex Roem. & Schult.) Miers ^Odontadenia geminata
Angadenia nitida (Vahl) Miers =Odontadenia nitida
Anisolobus cognatus (Stadelm.) Miill. Arg. =Odontadenia verrucosa
Anisolobus puncticulosus (Rich.) Miq. =Odontadenia puncticulosa
Apocynum acouci Aubl. =Forsteronia acouci
Apocynum cumanense Willd. ex Schult. (Asclepiadaceae) =Matelea cumanensis
Apocynum paniculatum Lam. =Forsteronia acouci
Apocynum umbellatum Aubl. =Forsteronia umbellata
Aspidosperma anomalum Miill. Arg. =Microplumeria anomala
Aspidosperma decipiens Miill. Arg. =Aspidosperma cuspa
Aspidosperma decussatum suhs^.jauaensis Steyerm. & Maguire =Aspidosperma decussatum
Aspidosperma desmanthum Benth. ex Mull. Arg. ^Aspidosperma album
Aspidosperma exalatum Monach. =Aspidosperma pachypterum
Aspidosperma lucentivenium S.F. Blake =Aspidosperma cuspa
Aspidosperma megalocarpon Miill. Arg. [not in flora area]
Aspidosperma nitidum Benth. ex Miill. Arg. [not in flora area]
Aspidosperma quadriovulatum Pittier =Rauvolfia pachyphylla
Aspidosperma rhombeosignatum Markgr. =Aspidosperma pachypterum
Aspidosperma verruculosum var. laeve Monach. ^Aspidosperma verruculosum
Aspidosperma woodsonianum Markgr. =Aspidosperma spruceanum
Bonafousia albiflora (Miq.) Boiteau & L. Allorge ^Tabernaemontana albiflora
Bonafousia angulata (Mart, ex Miill. Arg.) Boiteau & L. Allorge =Tabernaemontana angulata
Bonafousia attenuata Miers =Tabernaemontana attenuata
Bonafousia disticha (A. DC.) Boiteau & L. Allorge =Tabernaemontana disticha
Bonafousia lorifera (Miers) Boiteau & L. Allorge =Tabernaemontana lorifera
Bonafousia macrocalyx (Miill. Arg.) Boiteau & L. Allorge =Tabernaemontana macrocalyx
Bonafousia maxima (Markgr.) Boiteau & L. Allorge =Tabernaemontana maxima
200 Checklist of the Plants of the Guiana Shield
Bonafousia morettii L. Allorge =Tabernaemontana macrocalyx
Bonafousia miiellehana (Mart, ex Miill. Arg.) Boiteau & L. Allorge =Tabernaemontana macrocalyx
Bonafousia obliqua Miers =Tabernaemontana iindidala
Bonafousia oblongifolia (A. DC.) Miers =Tabernaemontanaflavicans
Bonafousia palustris (Markgr.) Boiteau & L. Allorge =Tabernaemontana palustris
Bonafousia rimuJosa (Woodson ex R.E. Schult.) Boiteau & L. Allorge =Tabernaemontana palustris
Bonafousia rupicola (Benth.) Miers =Tabernaemontana rupicola
Bonafousia rupicola van versicolor (Woodson) L. Allorge =Tabernaemontana rupicola
Bonafousia sananho (Ruiz & Pav.) Markgr. =Tabernaemontana sananho
Bonafousia siphilitica (L. f.) L. Allorge =Tabernaemontana siphilitica
Bonafousia speciosa (Poir.) Boiteau =Tabernaemontana siphilitica
Bonafousia tetrastachya (Kunth) Markgr. =Tabernaemontana siphilitica
Bonafousia undulata (Vahl) A. DC. =Tabernaemontana undulata
Cameraria tamaquarina Aubl. =Malouetia tamaquarina
Cerbera ahouai L. =Thevetia ahouai
Cerbera nitida Kunth =Thevetia ahouai
Cerbera peruviana Pers. =Thevetia peruvihna
Codonemma macrocalyx (Miill. Arg.) Miers =Tabernaemontana macrocalyx
CoUophora utilis Mart. =Couma utilis
Condylocarpon breviarticulatum Miill. Arg. =Condylocarpon intermedium var. intermedium
Condylocarpon ciliatum Miill. Arg. =Condylocarpon pubiflorum
Condylocarpon hirtellum Ducke =Condylocarpon pubiflorum
Condylocarpon intermedium var. brevifolium Miill. Arg. =Condylocarpon intermedium var. intermedium
Condylocarpon longii Standi. & L.O. Williams =Condylocarpon intermedium var. intermedium
Condylocarpon obtusiusculum Miill. Arg. =Condylocarpon intermedium var. intermedium
Condylocarpon occidentale Markgr. =Condylocarpon amazonicum
Condylocarpon rauwolfae Miill. Arg. [not in flora area] ^Condylocarpon isthmicum (Veil.) A. DC.
Condylocarpon reticulatum Ducke =Condylocarpon amazonicum
Conoria cuspa Kunth =Aspidosperma cuspa
Couma capiron Pittier =Couma macrocarpa
Couma caurensis Pittier =Couma macrocarpa
Coumafasciculata (Poir.) Benoist =Parahancorniafasciculata
Couma multinervis Monach. =Couma utilis
Couma oblonga Benth. ex Miill. Arg. =Parahancornia oblonga
Couma sapida Pittier =Couma macrocarpa
Cylindrosperma anomalum (Miill. Arg.) Ducke =Microplumeria anomala
Dipladenia calycina Ruber ex Ducke =Galactophora calycina
Dipladenia surinamensis Pulle =Mandevilla surinamensis
Dipladenia upatae Woodson =Mandevilla surinamensis
Echites angustifolia Benth. =Mandevilla benthamii
Echites benthamii A. DC. =Mandevilla benthamii
Echites biflora Jacq. =Rhabdadenia biflora
Echites brachystachya Benth. =Mandevilla scabra
Echites cognata Stadelm. =Odontadenia verrucosa
Echites elegans Benth. =Odontadenia geminata
Echites geminata Hoffmanns, ex Roem. & Schuh. =Odontadenia geminata
Echites grandiflora G. Mey. =Odontadenia macrantha
Echites hirsuta Rich. =Mandevilla hirsuta
Echites hoffmannseggiana Steud. =Odontadenia macrantha
Echites javitensis Kunth =Mandevilla javitensis
Echites laurentiae-disca Rusby =Prestonia acutifolia
Echites leptophylla A. DC. =Mandevilla leptophylla
Echites macrantha Roem. & Schult. =Odontadenia macrantha
Echites macrostoma Benth. =Rhabdadenia macrostoma
Echites megagros Veil. =Prestonia megagros
Echites nitida Vahl =Odontadenia nitida
Echites puncticulosa Rich. =Odontadenia puncticulosa
Echites quinquangularis Jacq. =Prestonia quinquangularis
Echites rugellosa Rich. =Mandevilla rugellosa
Echites scabra Hoffmanns, ex Roem. & Schult. =Mandevilla scabra
Echites siphilitica L. f. =Tabernaemontana siphilitica
Echites spectabilis Stadelm. =Macropharynx spectabilis
Echites subcarnosa Benth. =Mandevilla subcarnosa
Echites subspicata Vahl =Mandevilla subspicata
Checklist of the Plants of the Guiana Shield 201
Echites symphytocarpa G. Mey. =MandeviUa symphitocarpa
Echites trifida Jacq. =Mesechites trifida
Echites tubulosa Benth. =Mesechites trifida
Echites verrucosa Roem. & Schult. =Odontadenia verrucosa
En'atamia coronaria (Jacq.) Stapf ^Tabernaemontana divaricata
En'atatnia cumingiana (A. DC.) Markgr. =Tabernaemontana pandacaqui
Exothostemonjavitense (Kunth) G. Don =MandeviUajavitensis
Forsteronia benthamiana Miill. Arg. =Forsteronia acouci
Forsteronia coytnbifera (Miers) Sandwith =Forsteronia acouci
Forsteronia mediocris Woodson ^Forsteronia affinis
Forsteronia schoniburgkii (Benth.) Miill. Arg. =Forsteronia acouci
Forsteronia viridescens S.F. Blake =Forsteronia acouci
Haemadictyon acutifolium Benth. ex Miill. Arg. =Prestonia acutifolia
Haemadictyon cayennense A. DC. =Prestonia cayennensis
Haemadictyon exsertum A. DC. =Prestoma exserta
Haemadictyon marginatum Benth. =Prestonia cayennensis
Hancornia amapa Huber =Parahancorniafasciculata
Hancornia laxa (Benth.) A. DC. =Molongum laxum
Himatanthus rigida Willd. ex Roem. & Schult. =Himatanthus articulatus
Hortsmania myrtifolia Miq. =Condylocarpon myrtifolium
Lacmellea htcida (Kunth) Miers =Molongum lucidum
Lacmellea ramosissima (Miill. Arg.) Monach. =LacmeUea ramosissima
Lacmellea u tilis (Am.) Monach. =Lacmellea utilis
Landolphia guianensis (Aubl.) Pulle =Pacouria guianensis
Lochnera rosea (L.) Rchb. =Catharanthus roseus
Macaglia alba Vahl =Aspidosperma album
Macaglia excelsa (Benth.) Kuntze =Aspidosperma excelsum
Macaglia oblonga (A. DC.) Kuntze =Aspidosperma oblongum
Macaglia verruculosa (Miill. Arg.) Kuntze =Aspidosperma verruculosum
Macoubeafasciculata (Poir.) Lemee =Parahancorniafasciculata
Macoubea guianensis var. reticulata (A. DC.) L. Allorge =Macoubea guianensis
Macoubea paucifolia (Spruce ex Miill. Arg.) Markgr. ex LI. Williams =Macoubea guianensis
Macoubea reticulata (A. DC.) Markgr. =Macoubea guianensis
Macoubea sinuosa (Miers) Markgr. =Macoubea guianensis
Macoubea sprucei var. paucifolia (Spruce ex Mull. Arg.) Monach. =Macoubea guianensis
Malouetiafurfuracea Spruce ex Miill. Arg. =Malouetia tamaquarina
Malouetia guianensis (Aubl.) Miers =Malouetia tamaquarina
Malouetia lactifua Miers =Parahancornia fasciculata
Malouetia mexiae Woodson =Malouetiaflavescens
Malouetia obtusiloba A. DC. =Malouetia tamaquarina
Malouetia odorata DC. =Malouetia tamaquarina
Malouetia retroflexa Miill. Arg. [not in flora area] =Malouetia cubana A. DC.
Malouetia schomburgkii Miill. Arg. =Malouetiaflavescens
Malouetia tetrastachya (Kunth) Miers =Tabernaemontana siphilitica
Mandevilla angustissima Markgr. =Mandevilla lancifolia
Mandevilla dielsiana Markgr. =Mandevilla subcarnosa var. subcarnosa
Mandevilla glabra N.E. Br. =Mandevilla vanheurckii
Mandevilla lancifolia var. calva Monach. =Mandevilla lancifolia
Mandevilla linearis N.E. Br. =Mandevilla leptophylla
Merizadenia amplifolia Miers =Macoubea guianensis
Merizadenia sananho (Ruiz & Pav.) Miers =Tabernaemontana sananho
Mesechites syphilitica (L. f.) Lemee =Tabernaemontana siphilitica
Microplumeria anomala var. parvifolia Monach. =Microplumeria anomala
Microplumeria sprucei Baill. =Microplumeria anomala
Mitozus leptophyllus (A. DC.) Miers =Mandevilla leptophylla
Molongum cuneatum (Miill. Arg.) Pichon =Molongum lucidum
Molongum macrophyllum (Mull. Arg.) Pichon =Spongiosperma macrophyllum
Odontadenia "schomburgkiana (Nees) Kr." [invalid name, error for Odontonema schomburgkiana (Acanthaceae)?]
Odontadenia amazonica (Stadelm.) Malme =Odontadenia verrucosa
Odontadenia cognata (Stadelm.) Woodson ^Odontadenia verrucosa
Odontadenia cururu (Mart.) K. Schum. =Odontadenia puncticulosa
Odontadenia grandifora (G. Mey.) Kuntze =Odontadenia macrantha
Odontadenia hoffmannseggiana (Steud.) Woodson =Odontadenia macrantha
Odontadenia kochii Pilg. =Odontadenia puncticulosa
202 Checklist of the Plants of the Guiana Shield
Odontadenia neglecta Woodson =Odontadenia verrucosa
Odontadenia silvestris (A. DC.) Miill. Arg. ^'Odontadenia macrantha
Odontadenia speciosa Benth. =Odontadenia macrantha
Odontadenia spoliata Malme =Odontadenia puncticulosa
Odontadenia surinamensis (Pulle) Woodson =Odontadenia verrucosa
Orelia grandiflora Aubl. =Allamanda cathartica
Parahancornia amapa (Huber) Ducke =Parahancorniafasciculata
Parahancornia tabernaemontana Woodson =Macoubea guianensis
Peschiera cymosa (Jacq.) Dugand =Tabernaemontana cymosa
Peschiera echinata (Aubl.) A. DC. [name misapplied to Tabernaemontana lagenaria]
Peschiera heterophylla (Vahl) Miers =Tabernaemontana heterophylla
Peschiera psychotriifolia (Kunth) Miers =Tabernaemontana cymosa
Peschiera umbrosa (Kunth) Miers =Tabernaemontana cymosa
Plumeria alba var. inodora (Jacq.) G. Don ^Plumeria inodora
Plumeria articulata Vahl =Himatanthus articulatus
Plumeria attenuata Benth. =Himatanthus attenuatus
Plumeria attenuata var. obtusifolia Miill. Arg. =Himatanthus attenuatus var. obtusifolius
Plumeria bracteata A. DC. =Himatanthus bracteatus
Plumeria mollis Kunth =Plumeria rubra
Plumeria phagedaenica Mart. =Himatanthus phagedaenicus
Plumeria sucuuba Spruce ex Miill. Arg. =Himatanthus sucuuba
Plumeriopsis ahouai (L.) Rusby & Woodson =Thevetia ahouai
Prestonia arborescens Monach. =Hylaea arborescens
Prestonia marginata (Benth.) Woodson =Prestonia cayennensis
Quadricasaea inaequilateralis Woodson =Tabernaemontana macrocalyx
Rauvolfia hirsuta Jacq. =Rauvolfia tetraphylla
Rauvolfia lamarckii DC. =Rauvolfia tetraphylla
Rauvolfia parvifolia Bertero ex Spreng. =Rauvolfia ligustrina
Rauvolfia ternifolia Kunth =Rauvolfia ligustrina
Rhigospira paucifolia (Spruce ex Miill. Arg.) Miers =Macoubea guianensis
Rhigospira sinuosa Miers =Macoubea guianensis
Rhigospira sprucei (Miill. Arg.) Miers ^Macoubea sprucei
Rhigospira ternstroemiacea (Miill. Arg.) Miers =Neocouma ternstroemiacea
Rhigospira venulosa Miers =Spongiosperma macrophyllum
Rhipidia amazonica (Markgr.) Markgr. =Condylocarpon amazonicum
Salpinctes duidae Woodson =Mandevilla duidae
Stemmadenia cerea Woodson =Tabernaemontana cerea
Stenosolen heterophyllus (Vahl) Markgr. =Tabernaemontana heterophylla
Stenosolen holothuria Markgr. ^Tabernaemontana heterophylla
Taberna albiflora (Miq.) Markgr. =Tabernaemontana albiflora
Taberna cymosa (Jacq.) Miers =Tabernaemontana cymosa
Taberna disticha Miers ^Tabernaemontana disticha
Tabernaemontana albescens Rusby =Tabernaemontana undulata
Tabernaemontana aubletii Pulle, nom. illeg. =Macoubea guianensis
Tabernaemontana citrifolia Miill. Arg. [not in flora area]
Tabernaemontana coronaria (Jacq.) Willd. =Tabernaemontana divaricata
Tabernaemontana echinata Veil., non Aubl. [not in flora area] =Tabernaemontana hystrix Steud.
Tabernaemontanafasciculata Poir. =Parahancorniafasciculata
Tabernaemontanaflavescens Willd. ex Roem. & Schult. =Malouetiaflavescens
Tabernaemontana gracilis Benth. =Malouetia gracilis
Tabernaemontana grandiflora Jacq. =Stemmadenia grandiflora
Tabernaemontana holothuria (Markgr.) Leeuwenb. ^Tabernaemontana heterophylla
Tabernaemontana inaequilateralis (Woodson) Pichon =Tabernaemontana macrocalyx
Tabernaemontana laurifolia L. [not in flora area]
Tabernaemontana laxa Benth. =Molongum laxum
Tabernaemontana longifolia Benth. ^Tabernaemontana siphilitica
Tabernaemontana lucida Kunth =Molongum lucidum
Tabernaemontana macrophylla Miill. Arg. [not in flora area] =Tabernaemontana muricata Link ex Roem. & Schult.
Tabernaemontana meyeri G. Don =Tabernaemontana attenuata
Tabernaemontana muelleriana Mart, ex Miill. Arg. =Tabernaemontana macrocalyx
Tabernaemontana obliqua (Miers) Leeuwenb. ^Tabernaemontana undulata
Tabernaemontana oblongifolia A. DC. =Tabernaemontanaflavicans
Tabernaemontana olivacea Miill. Arg. =Tabernaemontanaflavicans
Tabernaemontana paucifolia Spruce ex Miill. Arg. =Macoubea guianensis
Tabernaemontana psychotriifolia Kunth ^Tabernaemontana cymosa
Checklist of the Plants of the Guiana Shield 203
Tabernaemontana repanda E. Mey. =Tabernaemontana siphilitica
Tabernaemontana rimulosa Woodson ex R.E. Schult. =Tabernaemontana palustris
Tabernaemontana sessilifolia Klotzsch =Tabernaemontana r-upicola
Tabernaemontana speciosa Poir. =Tabernaemontana siphilitica
Tabernaemontana sprucei Mull. Arg. =Macoubea sprucei
Tabernaemontana siibmoUis Mart, ex Mull. Arg. [not in flora area] =Tabernaemontana coriacea Link ex Roem. & Schult.
Tabernaemontana tenuis Monach. ^Tabernaemontana palustris
Tabernaemontana ternstroemiacea Miill. Arg. =Neocouma ternstroemiacea
Tabernaemontana tetrastachya Kunth =Tabernaemontana siphilitica
Tabernaemontana umbrosa Kunth =Tabernaemontana cymosa
Tabernaemontana utilis Am. =Lacmellea utilis
Tabernaemontana versicolor Woodson =Tabernaemontana rupicola
Thenardia laurifolia Benth. =Forsteronia laurifolia
Thenardia umbellata (Aubl.) Spreng. =Forsteroma umbellata
Thyrsanthus acouci (Aubl.) Miers =Forsteronia acouci
Thyrsanthus adenobasis (Miill. Arg.) Miers =Forsteronia adenobasis
Thyrsanthus ajfinis (Miill. Arg.) Miers =Forsteronia ajfinis
Thyrsanthus benthamianus (Miill. Arg.) Miers =Forsteronia acouci
Thyrsanthus diospyrifolius (Miill. Arg.) Miers =Forsteronia diospyrifolia
Thyrsanthus fasciculatus (Poir.) Miers =Parahancorniafasciculata
Thyrsanthus gracilis Benth. =Forsteronia gracilis
Thyrsanthus guyanensis (Miill. Arg.) Miers =Forsteronia guyanensis
Thyrsanthus laurifolius (Benth.) Miers =Forsteronia laurifolia
Thyrsanthus schomburgkii Benth. =Forsteronia acouci
Vinca rosea L. =Catharanthus roseus
Willughbeja acida (Aubl.) J.F. Gmel. =Ambelania acida
Willughbeja guianensis (Aubl.) J.F. Gmel. =Pacouria guianensis
Willughbeja scandans Willd. =Pacouria guianensis
Zschokkea aculeata Ducke =Lacmellea aculeata
Zschokkea microcarpa Miill. Arg. =Lacmellea microcarpa
Zschokkea ramosissima Miill. Arg. =Lacmellea ramosissima
Zschokkea utilis (Am.) Hemsl. =Lacmellea utilis
References
Allorge, L. 1985. Monographie des Apocynacees - Tabemaemontanoidees Americaines. Mem. Mus. Nat. Hist. Naturelle (n.s.)
Serie B, tome 30.
Allorge, L. 1985. Distribution, relations et interet biogeographique des Tabemaemontanoideae (Apocynaceae) dans la region
neotropicale. C.R. Soc. Biogeogr. 61(4): 137-155.
Allorge, L. 2002. Apocynaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
69-84.
Allorge, L. & C. Poupat. 1991. Position systematique et revision du genre Aspidosperma (Apocynaceae) pour les trois Guyanes.
Le point sur leur etude chimique. Bull. Soc. bot. Fr., Lettres bot. (4/5): 267-301.
Allorge, L. & C. Sastre. 1990. Distribution geographique dans les Guyanes, d'Apocynaceae et d'Ochnaceae. Essai de mise en
evidence de centres de speciation. C.R. Soc. Biogeogr. 67(2): 77-94.
Leeuwenberg, A.J.M. 1988. Series of revisions of Apocynaceae. XXI. Notes on Tabemaemontaneae. Agric. Univ. Wageningen
Papers 87-5: 1-32.
Leeuwenberg, A.J.M. 1994. A Revision of Tabemaemontana. Two. The New World Species and Stemmadenia. Royal Botanic
Gardens, Kew, Richmond, Surrey, England.
Markgraf, F. 1932. Apocynaceae. In A. Pulle, Flora of Suriname 4(1): 1-65.
Markgraf, F. 1937. Apocynaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 443-467.
Monachino, J.V. 1943 (1944). A revision of Couma and Parahancomia. Lloydia 6(4): 229-247.
Monachino, J. 1961. Apocynaceae. In B. Maguire, J.J. Wurdack, & collaborators. Botany of the Guayana Highland part IV.
Mem. New York Bot. Gard. 10(4): 59-65.
Plumel, M.-M. 1991. Le genre Himatanthus, revision taxonomique. Bradea 5 suppl.: 1-118.
Woodson, R.E., Jr. 1933-1936. Studies in the Apocynaceae. IV. The American genera of Echitoideae. Ann. Missouri Bot. Gard.
20: 605-790, 22: 153-306, 23: 169-439.
Zamcchi, J.L., G.N. Morillo, M.E. Endress, B.F. Hansen & A.J.M. Leeuwenberg. 1995. Apocynaceae. In J.A. Steyermark, P.E.
Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 2: 471-571.
Aquifoliaceae
P.E. Berry, P.-A. Loizeau, J.A. Steyermarkt
Ilex abscondita Steyerm. AM
Ilex acutidenticulata Steyerm. AM
204 Checklist of the Plants of the Guiana Shield
Ilex altiplana Steyerm.
Ilex amazonensis Edwin
Ilex apicidens N.E. Br.
Ilex aracamuniana Steyerm.
Ilex archeri Edwin
Ilex atabapoensis T.R. Dudley
Ilex attenuata Steyerm. [near Pico Neblina, Brazil]
Ilex brevipedicellata Steyerm.
Ilex cardonae Steyerm. [near Sierra Tapirapeco, Brazil]
Ilex casiquiarensis Loes.
Ilex ciliolata Steyerm.
Ilex costata Edwin
Ilex cowanii Wurdack
Ilex cidmenicola Steyerm.
Ilex daphnogenea Reissek
Ilex davidsei Steyerm.
Ilex diospyroides Reissek «'
Ilex divaricata Mart, ex Reissek
Ilex duidae Gleason
Ilexfanshawei Edwin
Ilex glabella Steyerm.
Ilex gleasoniana Steyerm.
Ilex gransabanensis Steyerm.
Ilex guaiquinimae Steyerm.
Ilex guianensis (Aubl.) Kuntze
Ilex holstii Steyerm.
Ilex huachamacariana Edwin
Ilex ignicola Steyerm.
Ilex inundata Poepp. ex Reissek
Ilexjauaensis Steyerm.
Ilexjenmanii Loes.
Ilex karuaiana Steyerm.
Ilex lasseri Edwin
Ilex laureola Triana
Ilex liesneri Steyerm.
Ilex longipilosa Steyerm.
Ilex macarenensis Cuatrec.
Ilex magnifructa Edwin
Ilex maguirei Wurdack
Ilex marahuacae Steyerm.
Ilex marginata Edwin
Ilex martiniana D. Don
Ilex neblinensis Edwin
Ilex oliveriana Loes.
Ilex ovalifolia G. Mey.
Ilex paruensis Steyerm.
Ilex parvifructa Edwin
Ilex paujiensis Steyerm.
Ilex polita Steyerm.
Ilex psammophila Reissek
Ilex ptariana Steyerm.
Ilex retusa Klotzsch ex Reissek
Ilex savannarum Wurdack
Ilex savannarum var. morichei Edwin
Ilex savannarum var. savannarum
Ilex sessilifructa Edwin
Ilex sipapoana Edwin
Ilex soderstromii Edwin
Ilex soderstromii var. ovata Edwin
Ilex soderstromii var. soderstromii
Ilex solida Edwin
Ilex spruceana Reissek
Ilex steyermarkii Edwin
Ilex subrotundifolia Steyerm.
AM
AM
AM
AM
BO
BO
AM
AM
AM
AM
AM
AM
BO
AM
AM
AM
AM BO
AM BO
AM
AM
AM
AM BO
BO
AM BO
AM
AM BO
AM BO
BO
BO
BO
AM BO
AM BO
AM
BO
AM
AM
AM BO
AM
BO
BO
AM
AM BO
BO
BO
AM BO
BO
AM BO
AM
AM
AM
AM BO
AM BO
BO
BO
BO
GU
GU?
GU
GU
GU
GU
FG
FG
FG
FG
DA GU SU
GU
GU
GU
SU
SU
FG
FG?
DA
DA
GU
GU
GU
GU
SU
SU
FG
FG
GU
GU
GU
GU
BO
BO
BO GU
BO GU
BO GU
BO GU
BO GU
BO
BO
BO GU
BO GU
BO
Checklist of the Plants of the Guiana Shield 205
Ilex sulcata Edwin [?later homonym of Ilex sulcata Wall., nomen nudum]
Ilex summa Steyerm.
Ilex tateana Steyerm. AM
Ilex tepuiana Steyerm. ex Edwin
Ilex theezam Mart.
Ilex theezans var. riedelii Loes.
Ilex thyrsiflora Klotzsch ex Reissek
Ilex thyrsiflora var. schomburgkii (Klotzsch ex Reissek) Loes.
Ilex tiricae Edwin
Ilex uaramae Edwin
Ilex umbellata Klotzsch ex Reissek
Ilex vacciniifoUa Klotzsch ex Reissek
Ilex venezuelensis Steyerm. AM
Ilex vismiifolia Reissek GU
Ilex wurdackiana Steyerm. BO
Ilexyutajensis^uxddick. AM
Synonyms and Misidentifications
Ilex acuminata Willd. =Ilex guianensis
Ilex bolivarensis Edwin =Ilex vacciniifoUa Klotzsch ex Reissek
Ilex brevipetiolata Steyerm. & Wurdack =Ilex subrotundifolia
Ilex celastroides Klotzsch ex Garcke =Ilex guianensis
Ilex congesta Reissek (Myrsinaceae) =Cybianthus lineatus
Ilex cumanensis Turcz. =Ilex guianensis
Ilex humirioides Reissek =Ilex umbellata
Ilex lanceolata Klotzsch [nomen nudum] =Ilex martiniana
Ilex laureola var. neglecta Loes. =Ilex laureola
Ilex laurina Klotzsch [nomen nudum] =Ilex daphnogenea
Ilex lucumifolia Reissek ex Maxim. (Sapotaceae) =Pouteria lucumifolia
Ilex lymanii Edwin =Ilex sipapoana
Ilex macoucoua Pers. =Ilex guianensis
Ilex magnifructa var. minor Steyerm. =Ilex magnifructa
Ilex neblinensis var. wurdackii Edwin =Ilex neblinensis
Ilex retusa Klotzsch [in R.M. Schomb., nom. inval] =Ilex retusa Klotzsch ex Reissek
Ilex retusa f. brunnescens Loes. =Ilex retusa
Ilex retusa f. genuina Loes. =Ilex retusa
Ilex retusa f. glabra Steyerm. =Ilex retusa
Ilex retusa f. glaucescens Steyerm. =Ilex retusa
Ilex retusa f. major Steyerm. =11ex retusa
Ilex riparia Reissek =Ilex inundata
Ilex schomburgkii Klotzsch ex Reissek =Ilex thyrsiflora var. schomburgkii
Ilex spathulata Steyerm., non Koidz 1925 =Ilex atabapoensis
Ilex spruceana var. genuina Loes. =Ilex spruceana
Ilex spruceana var. guainiensis Loes. =Ilex spruceana
Ilex stenophylla Steyerm. =Ilex retusa
Ilex sulcata var. pygmaea Edwin =Ilex sulcata
Ilex vacciniifoUa Klotzsch =Ilex vacciniifoUa Klotzsch ex Reissek
Macoucoua guianensis Aubl. =Ilex guianensis
References
Amshoff, G.J.H. 1951. Aquifoliaceae. In A. Pulle, Flora of Suriname 3(2): 159-165.
Edwin, G. 1965. Aquifoliaceae. In B. Maguire & collaborators, Botany of the Guayana Highland Part VI. Mem. New York Bot.
Gard. 12(3): 1-285.
Steyermark, J.A. & P.E. Berry. 1995. Aquifoliaceae. In J. A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan
Guayana 2: 571-599.
Wurdack, J.J. 1961. Aquifoliaceae. In B. Maguire, J.J. Wurdack, & collaborators. Botany of the Guayana Highland part IV.
Mem. New York Bot. Gard. 10(4): 1-6.
Araliaceae
D.G. Frodin, J.C. Lindeman
Dendropanax arboreus {L.)DQcnQ. 8c?\^x\q\\. AM BO DA GU
Dendropanax neblinae Maguire, Steyerm. & Frodin AM
Oreopanax capitatus (Jacq.) Decne. & Planch. AM BO DA GU FG
206 Checklist of the Plants of the Guiana Shield
Oreopanax capitatiis var. capitatiis AM BO DA GU FG
Schefflera acaropimctata Frodin AM
Schefflera argophyUa Frodin AM
Schefflera asymmetrica Frodin AM
Schefflera auyantepidensis (Steyerm.) Maguire, Steyerm. & Frodin BO
Schefflera ayangannensis Maguire, Steyerm. & Frodin GU
Schefflera baculosa Frodin AM
Schefflera brachypodia Frodin BO
Schefflera chimantensis (Steyerm. & Maguire) Maguire, Steyenn. & Frodin BO GU
Schefflera chimantensis subsp. chimantensis BO
Schefflera chimantensis var. iliiensis Frodin BO
Schefflera chimantensis subsp. miiltiramosa (Steyerm.) Frodin BO
Schefflera chimantensis var. nigosifolia (Maguire, Steyerm. & Frodin) Frodin BO GU
[Mt. Roraima]
Schefflera chimantensis subsp. nigosifolia (Maguire, Steyerm. & Frodin) BO GU
Frodin
Schefflera claiisa Frodin • BO
Schefflera clavigera Frodin BO
Schefflera clusietonim Frodin AM
Schefflera concolor Frodin AM
Schefflera contracta Frodin BO
Schefflera coriacea (Marchal) Harms BO GU
Schefflera cracens Frodin BO
Schefflera crassilimba Frodin AM
Schefflera decaphylla{SQQm.) W?irms GU SU FG
Schefflera disparifolia Frodin BO
Schefflera dissidens Frodin [Mt. Roraima] BO? GU
Schefflera diiidae Steyerm. [expected in GU] AM BO
Schefflera eximia Frodin [Sierra de la Neblina] AM
Schefflera gracillima Steyerm. & Maguire BO
Schefflera giianayensis Maguire, Steyerm. & Frodin AM
Schefflera hitchcockii (Lasser & Maguire) Maguire, Steyerm. & Frodin AM
Schefflera huachamacarii Maguire, Steyerm. & Frodin AM
Schefflera hiiberi Frodin AM
Schefflerajapurensis (Mart. & Zucc. ex Marchal) Harms AM BO
Schefflera jaiiaensis (Maguire, Steyerm. & Frodin) Frodin BO
Schefflera longistyla Frodin AM
Schefflera maguireanorum Frodin AM
Schefflera marahiiacensis (Maguire, Steyerm. & Frodin) Frodin AM GU?
Schefflera meiiirophylla Frodin AM
Schefflera monosperma Maguire, Steyerm. & Frodin [Mt. Roraima] BO GU
Schefflera montana (Gleason) Maguire, Steyerm. & Frodin AM BO
Schefflera morototoni (Aubl.) Maguire, Steyerm. & Frodin AM BO DA GU SU FG
Schefflera myrioneura Frodin AM
Schefflera neblinae (Maguire, Steyerm. & Frodin) Frodin [Sierra de la AM
Neblina]
Schefflera nigrescens Frodin AM
Schefflera pallens Maguire, Steyerm. & Frodin AM
Schefflera paruana (Maguire, Steyerm. & Frodin) Frodin AM
Schefflera pauciradiata Maguire, Steyerm. & Frodin AM
Schefflera pedicelligera Maguire, Steyerm. & Frodin AM
Schefflera pimichinensis Maguire, Steyerm. & Frodin AM
Schefflera phirispicata Maguire, Steyerm. & Frodin [expected in AM]
Schefflera psilophylla (Harms) Maguire, Steyerm. & Frodin [Mt. Roraima] BO
Schefflera quinquecarinata Steyerm. BO GU
Schefflera quinqiiestylorum Steyerm. BO GU
Schefflera sessiliflora Splithof-Heerschop ex Frodin AM BO GU
Schefflera simplex Steyerm. & B. Hoist AM
Schefflera sipapoensis Maguire, Steyerm. & Frodin AM
Schefflera spruceana (Seem.) Maguire, Steyerm. & Frodin AM
Schefflera steyermarkii Frodin BO
Schefflera siiaveolens Frodin AM
Schefflera tamatamaensis Maguire, Steyerm. & Frodin AM
Schefflera tremuloidea Maguire, Steyerm. & Frodin AM
Checklist of the Plants of the Guiana Shield 207
Schefflera iilocephala Frodin AM
Schefflera umbellata (N.E. Br.) R. Vig. [Mt. Roraima] BO GU
Schefflera umbraculifera Frodin AM
Schefflera varisiana Frodin [expected in AM]
Schefflera yutajensis Steyerm. & B. Hoist AM
Sciadodendron excelsimi Griseb. BO GU
Synonyms and Misidentifications
Aralia arborea L. =Dendropanax arboreus
Aralia capitata Jacq. =Oreopanax capitatus
Crepinella gracilis Marchal ex Thum =Schefflera monosperma
Didymopanax auyantepuiensis Steyerm. =Schefflera aiiyantepuiensis
Didymopanax chimantensis Steyerm. & Maguire =Schefflera chimantensis
Didymopanax hitchcockii Lasser & Maguire =Schefflera hitchcockii
Didymopanax montaniim Gleason =Schefflera montana
Didymopanax morototoni (Aubl.) Decne. & Planch. =Schefflera morototoni
Didymopanax multiramosus Steyerm. =Schefflera chimantensis subsp. multiramosa
Didymopanax psilophyllus Harms =Schefflera psilophylla
Didymopanax reticulatus Gleason =Schefflera sessiliflora
Didymopanax rugosum N.E. Br. =Schefflera chimantensis subsp. rugosifolia
Didymopanax spruceanus Seem. =Schefflera spruceana
Didymopanax spruceanus var. cuyabensis Hoehne =Schefflera spruceana
Gilibertia arborea (L.) Marchal =Dendropanax arboreus
Oreopanax capitatus var. multiflorus (DC.) Marchal [Guianan material agrees with var. capitatus fide Lindeman]
Panax morototoni Aubl. =Schefflera morototoni
Panax splendens Kunth =Schefflera morototoni
Panax undulata Aubl. =Schefflera morototoni
Schefflera arborea (L.) M. Gomez =Dendropanax arboreus
Schefflera coriacea subsp. neblinae Maguire, Steyerm. & Frodin =Schefflera neblinae
Schefflera duidae subsp. quinquestylorum (Steyerm.) Maguire, Steyerm. & Frodin =Schefflera quinquestylorum
Schefflera globulifera Maguire, Steyerm. & Frodin ^Schefflera ulocephala
Schefflera guanayensis subsp. /jan/ew^w Maguire, Steyerm. & Frodin =Schefflera paruana
Schefflera guanayensis subsp. sipapoensis Maguire, Steyerm. & Frodin =Schefflera crassilimba
Schefflera paraensis Huber ex Ducke =Schefflera decaphylla
Schefflera reticulata (Gleason) Maguire, Steyerm. & Frodin, non Philipson =Schefflera sessiliflora
Schefflera reticulata suhsp.jauaensis Maguire, Steyerm. & Frodin =Schefflerajauaensis
Schefflera reticulata subsp. yutajensis Maguire, Steyerm. & Frodin =Schefflera maguireanorum
Schefflera rugosifolia Maguire, Steyerm. & Frodin =Schefflera chimantensis subsp. rugosifolia
Schefflera splendens (Kunth) Frodin ex Lindeman =Schefflera morototoni
Schefflera tremuloides var. marahuacensis Maguire, Steyerm. & Frodin =Schefflera marahuacensis
Schefflera umbellata subsp. duidana Maguire, Steyerm. & Frodin =Schefflera acaropunctata
Sciadophyllum coriaceum Marchal =Schefflera coriacea
Sciadophyllum japurense Mart. & Zucc. ex Marchal =Schefflerajapurensis
Sciadophyllum umbellatum N.E. Br. =Schefflera umbellata
References
Frodin, D.G 1993. Studies in Schefflera, VI. New species and subordinate taxa in the Venezuelan Guayana and immediately
adjacent areas. Novon 3: 367-403.
Frodin, D.G. 1997. Araliaceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 1-31.
Maguire, B., J.A. Steyermark, & D.G. Frodin. 1 984. Araliaceae. In B. Maguire &. collaborators. Botany of the Guayana Highland
Part XII. Mem. New York Bot. Gard. 38: 1-84.
Mori, S.A. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide
to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 84-87.
Muller, F.M. 1934. Araliaceae. In A. Pulle, Flora of Suriname 3(1): 160-163.
Aristolochiaceae
K.A. Barringer, C. Feuillet, F.A. Gonzalez G, O. Poncy
Aristolochia acutifolia Duch. AM BO
Aristolochia amara (Aubl.) Poncy [taxon based on sterile material, of FG
uncertain application]
Aristolochia anguicida Jacq. AM BO
Aristolochia bukuti Poncy SU FG
Aristolochia chamissonis Duch. AM
208 Checklist of the Plants of the Guiana Shield
Aristolochia consimilis Mast.
Aristolochia cornuta Mast.
Aristolochia cremersii Poncy
Aristolochia daemoninoxia Mast.
Aristolochia didyma S. Moore
Aristolochia disticha Mast.
Aristolochiaflava Poncy
Aristolochia guianensis Poncy
Aristolochia guntheri O.C. Schmidt
Aristolochia hians Willd.
Aristolochia iquitensis O.C. Schmidt
Aristolochia kamikuensis Feuillt, ined.
Aristolochia leprieurii Duch.
Aristolochia maxima Jacq.
Aristolochia melgiieiroi Barringer & Guanchez
Aristolochia mossii S. Moore
Aristolochia nummularifolia Kunth
Aristolochia pannosoides Hoehne
Aristolochia paramaribensis Duch.
Aristolochia ringens Vahl
Aristolochia riigosa Lam.
Aristolochia silvatica Barb. Rodr.
Aristolochia sprucei Mast.
Aristolochia stahelii O.C. Schmidt
Aristolochia surinamensis Willd.
Aristolochia trilobata L.
Aristolochia weddellii Duch.
Aristolochia weddellii var. rondoniana Hoehne
AM BO
AM BO
AM BO
AM BO
BO
AM BO
BO
GU
GU
GU
GU
GU
GU
AM
DA
DA
GU
GU
GU
SU
SU
FG
FG
FG
FG
FG
FG
FG?
FG
FG
FG
SU FG
FG
SU FG
GU SU
GU SU FG
GU SU FG
GU SU FG
Synonyms and Misidentifications
Aristolochia amesiana R.E. Schult. =Aristolochia silvatica
Aristolochia barbata Jacq. =Aristolochia rugosa
Aristolochia dictyantha var. schomburgkii (Klotzsch) Duch. =Aristolochia rugosa
Aristolochia dubia Hoehne =Aristolochia mossii
Aristolochia gabrielis Briq. =Aristolochia leprieurii
Aristolochia glaucescens Kunth [not in flora area]
Aristolochia longecaudata Mast. =Aristolochia weddellii var. rondoniana
Aristolochia macrophylla Duch. =Aristolochia iquitensis
Aristolochia macrota Duch. =Aristolochia surinamensis
Aristolochia platyloba Garcke =Aristolochia surinamensis
References
Barringer, K.A., & G.A. Gonzalez G. 1997. Aristolochiaceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora ofthe Venezuelan
Guayana3: 122-129.
Feuillet, C. & O. Poncy. 1998. Aristolochiaceae. In A.R.A. Gorts-van Rijn & M.J. Jansen-Jacobs, Flora of the Guianas (series A)
20: 1-34.
Feuillet, C. & O. Poncy. 2002. Aristolochiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, &
J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot.
Gard. 76 (2): 87-89.
Hoehne, F.C. 1942. Aristolochiaceas. In F.C. Hoehne, Flora Brasilica 14(2): 1-141.
Poncy, O. 1988. Quatre especes nouvelles d'Aristolochia de Guyane fran9aise, et remarques nomenclaturales concemant une
espece d'Aublet. Bull. Mus. Hist. Nat. Paris, ser. 4, 10 (sect. B), Adansonia 4: 337-344.
ASCLEPIADACEAE
G.N. Morillo
Asclepias curassavica L.
Asclepiasfruticosa L.
Barjonia erecta (Veil.) K. Schum.
Blepharodon glaucescens (Decne.) Fontella
Blepharodon grandiflorus Benth.
Blepharodon grandiflorus subsp. crassifolius (Schltr.) Morillo
Blepharodon julianii Morillo [expected in GU]
Blepharodon maigualidae Morillo
AM
AM
BO
BO
DA GU
GU
SU
SU
SU
BO
BO
GU
GU
FG
Checklist of the Plants of the Guiana Shield 209
Blepharodon mucronatus (Schltdl.) Decne.
Blepharodon nitidus (Veil.) J.F. Macbr.
Blepharodon tillettii Morillo
Blepharodon idei Schltr.
Calotropis gigantea (L.) Dryand. [cultivated, naturalized]
Cryptostegia madagascariensis Bojer ex Decne. [escaped from cultivation?]
Cynanchum blandum (Decne.) Sundell
Cynanchum buntingii Morillo
Cynanchum chimantense Morillo
Cynanchum goertsianum Morillo [published as C. gortsianum]
Cynanchum guanchezii Morillo
Cynanchum hirtellum (Oliv.) V.M. Badillo
Cynanchum huberi Morillo
Cynanchum mirifolium (Gleason & Moldenke) R.W. Holm
Cynanchum montevidense Spreng.
Cynanchum neei Morillo
Cynanchum paraquense Morillo
Cynanchum parviflorum Sw.
Cynanchum prevostiae Morillo
Cynanchum sobradoi Morillo
Cynanchum stannardii Morillo
Cynanchum strictum R.W. Holm
Ditassa acerifolia Lasser & Maguire
Ditassa angustifolia Decne.
Ditassa auyantepuiensis (Steyerm.) Morillo
Ditassa ayangannensis Morillo
Ditassa bolivarensis (R.W. Holm) Morillo [expected in GU]
Ditassa carnevalii (Morillo) Morillo
Ditassa ciliata (Moldenke) Morillo
Ditassa colellae (Morillo) Morillo
Ditassa consanguinea Decne.
Ditassa duidae Gleason
Ditassa foldatsii Morillo
Ditassafranciscoi (Morillo) S. Liede
Ditassa gillespieae Morillo
Ditassajuliani (Morillo) Morillo
Ditassa liesneri Morillo
Ditassa lindemanii Morillo
Ditassa multinervia (Morillo) Morillo
Ditassa obovata (R.W. Holm) Morillo [expected in GU]
Ditassa olivaestevae Morillo
Ditassa ottohuberi Morillo
Ditassa pauciflora Decne.
Ditassa roraimensis Schltr.
Ditassa sipapoana Morillo
Ditassa surinamensis Morillo
Ditassa tatei Gleason & Moldenke
Ditassa taxifolia Decne.
Ditassa venamensis Morillo
Ditassa verticillata Morillo
Fischeria stellata (Veil.) E. Foum.
Gonolobus aristolochioides Kunth
Gonolobus germanianus Morillo [expected in GU]
Gonolobus glaber Decne. [specimens not seen, status uncertain]
Gonolobus grandiflorus R. Br. [specimens not seen, status uncertain]
Gonolobus tobagensis Urb.
Macroditassa Carolina Morillo [Mt. Roraima]
Macroditassa laurifolia (Decne.) Fontella
Marsdenia altissima (Jacq.) Dugand
Marsdenia gillespieae Morillo
Marsdenia guanchezii Morillo
Marsdenia macrophylla (Humb. & Bonpl. ex Schult.) E. Foum.
Marsdenia rubrofusca Benth. ex E. Fourn.
Marsdenia sprucei Rothe
AM BO
AM BO GU
GU
su FG
AM BO GU
BO GU su
AM BO GU
GU su FG
AM
BO
su FG
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
GU
GU
GU
GU
GU
GU
GU
GU
GU
FG
GU
GU
GU SU FG
GU
SU
SU
SU?
GU?
GU
BO GU
GU
BO GU
GU
SU
AM BO
AM BO GU SU
AM BO DA GU SU
AM
FG
FG
210 Checklist of the Plants of the Guiana Shield
Marsdenia undiilata (Jacq.) Dugand BO
Marsdenia weddellii (E. Foum.) Malme SU
Marsdenia xerohylica Dugand BO
Matelea amazonica Morillo AM
Matelea badilhi MoriWo BO GU FG
Matelea bolivarensis Morillo BO GU
Matelea cayennensis Morillo FG
Matelea coriacea Morillo [expected in GU] BO
Matelea cremersii Morillo FG
Matelea cumanensis (Willd. ex Schult.) W.D. Stevens BO
Matelea delascioi Morillo DA GU
Matelea denticiilata (Vahl) Fontella & E.A. Schwarz BO DA GU SU FG
Mateleafournieri Morillo SU
Mateleafucata Woodson BO
Matelea funkiana Morillo GU
Matelea glaziovii (E. Foum.) Morillo SU
Matelea gracieae Morillo • FG
Matelea grenandii Morillo FG
Matelea herbacea Woodson GU
Matelea hildegardiana Morillo BO
Matelea hirsuta (Vahl) Woodson BO GU
Matelea hoffmanii Morillo GU
Matelea holstii Morillo & Camevali BO
Mateleajansen-jacobsiae Morillo SU
Mateleajenmanii (Morillo) Morillo GU
Matelea liesneri Morillo AM
Matelea loiirteigiae Morillo AM
Matelea mahtima (Jacq.) Woodson AM
Matelea neblinae Morillo AM
Matelea oldemanii Morillo
Matelea palustris Aubl.
Matelea planiflora (Jacq.) Dugand
Matelea riparia Morillo
Matelea sastrei Morillo
Matelea squiresii (Rusby) Morillo
Matelea stenopetala Sandwith AM
Matelea stergiosii Morillo
Matelea suareziae Morillo AM
Matelea surinamensis Morillo SU'
Matelea yanomamica Morillo AM
Metalepis albiflora Urb.
Metalepis prevostiae Morillo
Nephradenia linearis Benth. ex E. Foum. AM
Oxypetalum capitatiim Mart.
Sarcostemma clausiim (Jacq.) Schult. AM
Stenomeria decalepis Turcz.
Tassadia aristata (Benth. ex E. Foum.) Fontella [expected in the Guianas] AM
Tassadia berteriana (Spreng.) W.D. Stevens
Tassadia castellanosii Fontella AM
Tassadia decaisneana Miq. GU SU
Tassadia guianensis Decne. GU SU FG
Tassadia ivonae Morillo AM BO
Tassadia kamaensis (Morillo) Morillo BO
Tassadia leptobotrys Decne. SU
Tassadia marahuacensis Morillo AM
Tassadia martiana Decne. [expected in the Guianas]
Tassadia medinae (Morillo) Morillo
Tassadia obovata Decne. AM BO DA GU SU FG
Tassadia ovalifolia (E. Foum.) Fontella [expected in the Guianas]
Tassadia propinqua Decne. [expected in FG]
Tassadia trailiana (Benth.) Fontella
Synonyms and Misidentifications
Asclepias altissima Jacq. =Marsdenia altissima
Asclepias clausa Jacq. =Sarcostemma clausum
GU SU FG
BO GU
FG
GU SU FG
BO GU
GU
SU FG
BO DA GU SU
BO GU FG
BO
FG
FG
BO GU
BO GU SU
BO DA GU
GU
SU FG
BO
BO GU SU
DA GU
GU
SU
Checklist of the Plants of the Guiana Shield 211
Asclepias gigantea L. =Calotropis gigantea
Asclepias macrophylla Humb. & Bonpl. ex Schult. =Marsdenia macrophylla
Asclepias planiflora Jacq. =Matelea planiflora
Asclepias undulata Jacq. =Marsdenia undulata
Astephamis mucronata Schltdl. =Blepharodon miicronatus
Blepharodon auyantepuiense Steyerm. =Blepharodon ulei
Blepharodon bifidus Schltr. =Blepharodon grandifloriis subsp. crassifolius
Blepharodon ciliatum Moldenke =Ditassa ciliata
Blepharodon crassifolius Schltr. ^Blepharodon grandiflorus subsp. crassifolius
Blepharodon hitchcockii Gleason =Blepharodon ulei
Blepharodon steudelianum (Miq.) Pulle =Blepharodon nitidus
Blepharodon steyermarkii R.W. Holm =Blepharodon nitidus
Blepharodon venezuelense Moldenke =Blepharodon nitidus
Blepharodon viscidum R.W. Holm =Nephradenia linearis
Cynanchum acerifolium (Lasser & Maguire) Morillo =Ditassa acerifolia
Cynanchum altissimum (Jacq.) Jacq. =Marsdenia altissima
Cynanchum angustifolium (Decne.) Morillo =Ditassa angustifolia
Cynanchum apocynellum (Gleason & Moldenke) Spellman =Tassadia obovata
Cynanchum auyantepuiense Steyerm. =Ditassa auyantepuiensis
Cynanchum bolivarense R.W. Holm =Ditassa bolivarensis
Cynanchum carnevalii Morillo =Ditassa carnevalii
Cynanchum clausum (Jacq.) Jacq. =Sarcostemma clausum
Cynanchum colellae Morillo =Ditassa colellae
Cynanchum denticulatum Vahl =Matelea denticulata
Cynanchum duidae R.W. Holm ^Ditassa ciliata
Cynanchum fimbriatum Kunth =Matelea cumanensis
Cynanchum franciscoi Morillo =Ditassafranciscoi
Cynanchum guianense Spreng. =Matelea denticulata
Cynanchum hirsutum Vahl =Matelea hirsuta
Cynanchum holmianum Steyerm. =Ditassa duidae
Cynanchumjenmanii Morillo =Mateleajenmanii
Cynanchum julianii Morillo =Ditassa juliani
Cynanchum kamaense Morillo =Tassadia kamaensis
Cynanchum maritimum Jacq. =Matelea maritima
Cynanchum medinae Morillo =Tassadia medinae
Cynanchum multinei'vium Morillo =Ditassa multinervia
Cynanchum nitidum Veil. =Blepharodon nitidus
Cynanchum obovatum R.W. Holm =Ditassa obovata
Cynanchum planiflorum Jacq. =Matelea planiflora
Cynanchum recurvum (Rusby) Spellman =Tassadia obovata
Cynanchum revolutum R.W. Holm =Cynanchum hirtellum
Cynanchum roraimense (Schlltr.) Morillo =Ditassa roraimensis
Cynanchum stellatum Veil. =Fischeria stellata
Cynanchum stenolobum (Decne.) Morillo =Tassadia berteriana
Cynanchum tatei (Gleason & Moldenke) R.W. Holm =Ditassa tatei
Cynanchum taxifolium (Decne.) R.W. Holm =Ditassa taxifolia
Cynanchum tepuiense Morillo =Ditassa ciliata
Cynanchum undulatum (Jacq.) Jacq. =Marsdenia undulata
Cynanchum viridiflorum G. Mey. ^Matelea denticulata
Ditassa aristata Benth. ex E. Foum. =Tassadia aristata
Ditassa blepharodontoides Schltr. =Blepharodon glaucescens
Ditassa glaucescens Decne. ^Blepharodon glaucescens
Esmeraldia stricta E. Foum. ^'Cynanchum strictum
Fischeria calycina Decne. -Fischeria stellata
Fischeria subaequalis S.F. Blake =Fischeria stellata
Funastrum clausum (Jacq.) Schltr. =Sarcostemma clausum
Glaziostelma ovalifolium E. Foum. =Tassadia ovalifolia
Gonolobus adenophorus Suess. =Matelea planiflora
Gonolobus ligustrinus Decne. [not in flora area]
Gonolobus maritimus (Jacq.) R. Br. =Matelea maritima
Gonolobus riparius Kunth [not in flora area]
Gonolobus squiresii Rusby =Matelea squiresii
Gonolobus surinamensis Jonker =Matelea lourteigiae
Gonolobus viridiflorus (G. Mey.) Schult. =Matelea denticulata
212 Checklist of the Plants of the Guiana Shield
Ibatia albiflora H. Karst. =MaTeIea cumanensis
Ibatia maritima (Jacq.) Decne. ^Matelea maritima
Macroscepis hirsuta (Vahl) Schltr. =Matelea hirsiita
Macroscepis trianae Decne. ex Triana =Matelea hirsuta
Madarosperma trailiana Benth. =Tassadia trailiana
Marsdenia amazonica Morillo =Marsdenia spriicei
Marsdenia caurensis Morillo =Marsdenia altissima
Marsdenia maculata Hook. =Marsdenia macrophyUa
Marsdenia margaritaria R.C. Foster =Marsdenia undulata
Marsdenia moUissima E. Foum. =Marsdenia altissima
Matelea albiflora (H. Karst.) Dugand =Matelea cumanensis
Mateleafimbriata (Kunth) Dugand =Matelea cumanensis
Matelea latifolia Aub\. =Matelea palustris
Matelea mediocris Woodson =Matelea squiresii
Matelea trianae (Decne. ex Triana) Spellman =Matelea hirsuta
Matelea viridiflora (G. Mey.) Woodson =Matelea denticulata
Meresaldia stricta (E. Foum.) Bullock =Cxnanchum strictum
Metastelma berterianum (Spreng.) Decne. =Tassadia berteriana
Metastelma brasiliense K. Schum. [not in flora area]
Metastelma campanulatum Decne. =Cynanchum parviflorum
Metastelma decaisneana Benth. & Hook. =Tassadia decaisneana
Metastelma mirifolium Gleason & Moldenke =Cynanchum mirifolium
Metastelma paraquense (Morillo) Liede =Cynanchum paraquense
Metastelma paniflorum (Sw.) R. Br. ex Schult. =Cynanchum parviflorum
Metastelma stenolobum Decne. =Tassadia berteriana
Metastelma steudelianum Miq. =Blepharodon nitidus
Metastelma strictum (E. Foum.) Spruce ex Knuth =Cynanchum strictum
Metastelma strictum Gleason & Moldenke =Cynanchum strictum
Oxypetalum berterianum Spreng. =Tassadia berteriana
Roulinia blanda Decne. =Cynanchum blandum
Roulinia voglii Suess. ^Cynanchum montevidense
Rouliniella guianensis (Decne.) Jonker =Cynanchum blandum
Sarcostemma cumanense Kunth =Sarcostemma clausum
Sarcostemma pubescens Kunth =Sarcostemma clausum
Stephanotella killipii Woodson =Marsdenia rubrofusca
Tassadia angustifolia Malme =Tassadia propinqua
Tassadia apocynella Gleason & Moldenke =Tassadia obovata
Tassadia recurva Rusby =Tassadia obovata
Telminostelma corymbosum (Decne.) Fontella & Schwarz =Cynanchum blandum
Vincetoxicum hirtellum Oliv. =Cynanchum hirtellum
References
Fontella Pereira, J. 1977. Revisao taxonomica do genero Tassadia Decaisne. Arq. Jard. Bot. Rio de Janeiro 21: 235-392.
Jonker, F.R 1940. Asclepiadaceae. In A. Pulle, Flora of Suriname 4(2): 326-352.
Morillo, G. 1986. Asclepiadaceae II. Especies nuevas o interesantes en Venezuela y Guyana. Emstia 37: 1-13.
Morillo, G. 1988. Asclepiadaceae III, one new species and four new records. Proc. Kon. Ned. Akad. Wet. ser. C, 91(2): 161-164.
Morillo, G. 1989. Contribucion al conocimiento de las Asclepiadaceae suramericanas, principalmente de las Guayanas. Emstia
51:2-15.
Morillo, G. 1989. Veinte Asclepiadaceae sudamericanas nuevas para la ciencia y una nueva combinacion. An. Jard. Bot. Madrid
47(2): 349-359.
Morillo, G. 1991. Once Asclepiadaceae suramericanas nuevas para la ciencia. Emstia 1(3): 109-120.
Morillo, G. 1993. Asclepiadaceae nuevas o interesantes de Venezuela y Guyana. Emstia 3(2): 61-70.Morillo, G 1994. Seis
Asclepiadaceae Sudamericanas nuevas para la ciencia. Emstia 4(1-2): 3-10.
Morillo, GN. 1997. In J.A. Steyermark, RE. Berry, «& B.K. Hoist, Flora of the Venezuelan Guayana 3: 129-177.
Morillo, G. 1988. Asclepiadaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
89-94.
Balanophoraceae
B. Hansen, J.S. Miller
Helosis cayennensis (Sw.) Spreng.
Helosis cayennensis var. cayennensis
Langsdorffia hypogaea Mart.
AM BO DA GU su FG
AM BO DA GU su EG
AM BO GU
Checklist of the Plants of the Guiana Shield 213
Synonyms and Misidentifications
Caldasia brasiliensis (Schott & Endl.) Kuntze =HeJosis cayennensis
Caldasia cayennensis (Sw.) Mutis ex Steud. =Helosis cayennensis
Cynomoriiim cayennense Sw. =Helosis cayennensis
Helosis brasiliensis Schott & Endl. =Helosis cayennensis
Helosis guyanensis Rich. [nom. illeg. superfl.] =Helosis cayennensis
Langsdorffia nibiginosa Wedd. =Langsdorffia hypogaea
References
Gracie, C.A. 2002. Balanophoraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76(2): 116.
Hansen, B. 1980. Balanophoraceae. Flora Neotropica Monogr. 23: 1-80.
Hansen, B. 1993. Balanophoraceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 14: 40-43.
Lanjouw, J. 1932. Balanophoraceae. In A. Pulle, Flora of Suriname 1(1): 45-46.
Miller, J.S. 1997. Balanophoraceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 393-395.
Balsaminaceae
Impatiens balsamina L. [probably escaped from cultivation] GU
Basellaceae
R.A. DeFilippst, J.A. Steyermarkt
Anredera vesicaria (Lam.) C.F. Gaertn. [probably escaped from cultivation in AM
FG]
Basella alba L. [cultivated & naturalized]
BO
GU SU
FG
FG
Synonyms and Misidentifications
Anredera leptostachys (Moq.) Steenis =Anredera vesicaria
Basella rubra L. =Basella alba
Boussingaidtia leptostachya Moq. =Anredera vesicaria
Boiissingaultia leptostachys Moq. =Anredera vesicaria
References
Sperling, C.R. 1987. Systematics of the Basellaceae. Ph.D. Dissertation, Harvard University. U.M.I. Dissertation Information
Service, Ann Arbor, Michigan.
Steyermark, J.A. 1997. Basellaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 395-397.
Bataceae
Batis maritima L. GU SU FG:
References
Jonker, F.P. 1953. Batidaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 91-93.
Begoniaceae
J.C. Lindeman, J.A. Steyermarkt, D. Wasshausen
Begonia fischeri Schrank
Begonia fiichsioides Hook.
Begonia glabra Aubl.
Begonia giiaduensis Kunth
Begonia heloisana Brade
Begonia hirsuta Aubl.
Begonia humHis Dryand.
Begoniajenmanii Tutin
Begonia meridensis A. DC.
Begonia nubicola L.B. Sm. & B.G. Schub. [Sierra de la Ncblina]
Begonia prieurii A. DC.
Begonia semiovata Liebm.
Begonia steyermarkii L.B. Sm. & B.G. Schub.
Begonia ulmifolia Willd.
BO DA GU
AM
AM GU SU FG
AM BO
GU SU FG
FG
AM BO
BO
GU
GU
SU FG
AM
BO GU SU FG
BO DA GU SU FG
BO GU
BO GU SU FG
214 Checklist of the Plants of the Guiana Shield
Synonyms and Misidentifications
Begonia diffusa L.B. Sm. & B.G. Schub. [not in flora area]
Begoniafilipes Benth. [not in flora area]
Begonia guyanensis A. DC. =Begonia semiovata
Begonia hirtella Link [not in flora area]
Begonia kaietukensis Tutin =Begoma fischeri
Begonia kvistii L.B. Sm. [nomen nudum] =Begoniajenmanii
Begonia roraimensis Tutin =Begonia fischeri
Begonia scandens Sw. =Begonia glabra
Begonia tovarensis Klotzsch =Begonia fischeri
References
Heald, S.V. 2002. Begoniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
117-118.
Lindeman, J.C. 1984. Begoniaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 576-584.
Smith, L.B., D.C. Wasshausen, J. Golding, & C.E. Karegeannes. 1986. Begoniaceae (illustrated key and annotated species list).
Smithsonian Contrib. Bot. 60: 1-584.
Smith, L and D. Wasshausen. 1989. Begoniaceae. Flora de Venezuela 4(1): 5-68
Steyermark, J.A. 1997. Begoniaceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora ofthe Venezuelan Guayana 3: 397-403.
Adenocalymna apurense (Kunth) Sandwith
Adenocalymna impressum (Rusby) Sandwith
Adenocalymna inundatum Mart, ex DC.
Adenocalymna inundatum van surinamense Bureau & K. Schum
Adenocalymna prancei A.H. Gentry
Adenocalymna purpurascens Rusby
Adenocalymna saulense A.H. Gentry
Adenocalymna subincanum Huber
Amphilophium aschersonii Ule
Amphilophium paniculatum (L.) Kunth
Amphilophium paniculatum var. imatacense A.H. Gentry
Amphilophium paniculatum var. molle (Schltdl. & Cham.) Standi
Amphilophium paniculatum var. paniculatum
Anemopaegma alatum A.H. Gentry
Anemopaegma cf brevipes S. Moore
Anemopaegma chamberlaynii (Sims) Bureau & K. Schum.
Anemopaegma chrysoleucum (Kunth) Sandwith
Anemopaegmafioridum Mart, ex DC.
Anemopaegmafoetidum Bureau & K. Schum.
Anemopaegma granvillei A.H. Gentry
Anemopaegma ionanthum A.H. Gentry
Anemopaegmajucundum Bureau & K. Schum.
Anemopaegma karstenii Bureau & K. Schum.
Anemopaegma longidens DC.
Anemopaegma maguirei Sandwith
Anemopaegma oligoneuron (Sprague & Sandwith) A.H. Gentry
Anemopaegma paraense Bureau & K. Schum.
Anemopaegma parkeri Sprague
Anemopaegma patelliforme A.H. Gentry
Anemopaegma robustum Bureau & K. Schum.
Anemopaegma salicifolium (Kunth) Sandwith
Arrabidaea arthrerion (DC.) Bureau ex K. Schum.
Arrabidaea bilabiata (Sprague) Sandwith
Arrabidaea brachypoda (DC.) Bureau
Arrabidaea candicans (Rich.) DC.
Arrabidaea carichanensis (Kunth) Bureau & K. Schum.
Arrabidaea chica (Bonpl.) B. Verl.
Arrabidaea cinerea Bureau ex K. Schum.
Arrabidaea cinerea var. cinerea
Arrabidaea cinerea var. longipila Sandwith
BiGNONIACEAE
L.H. Gentryt, S.O. Grose
BO
AM BO
AM BO
um. AM BO
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
DA
DA
DA
DA
DA
DA
DA
GU?
GU
GU
GU
GU
GU
GU
GU
GU?
GU
GU
GU
GU
su
su
su
su
su
su?
su
su
FG
FG
FG
FG
FG
FG
FG
FG
SU
GU
GU SU FG
GU SU
FG
GU FG
GU
GU
FG
GU? SU
GU SU FG
GU? SU FG
GU su FG
FG
FG
FG
SU? FG
FG
FG
Checklist of the Plants of the Guiana Shield 215
Arrabidaea cinnamomea (DC.) Sandwith
Arrabidaea conjugata (Veil.) Mart.
Arrabidaea corallina (Jacq.) Sandwith
Arrabidaea egensis Bureau & K. Schum.
Arrabidaeafanshawei Sandwith
Arrabidaea fJorida DC.
Arrabidaea grosourdyana (Baill.) Sandwith
Arrabidaea inaequalis (DC. ex Splitg.) K. Schum.
Arrabidaeajapurensis (DC.) Bureau & K. Schum.
Arrabidaea lachnaea (Bureau ex Bureau & K. Schum.) Sandwith
Arrabidaea mollis (Vahl) Bureau ex K. Schum.
Arrabidaea mollissima (Kunth) Bureau & K. Schum.
Arrabidaea nigrescens Sandwith
Arrabidaea oligantha Bureau & K. Schum.
Arrabidaea oxycarpa Urb.
Arrabidaea patellifera (Schltdl.) Sandwith
Arrabidaea prancei A.H. Gentry [Sierra de laNeblina]
Arrabidaea pubescens (L.) A.H. Gentry
Arrabidaea revillae A.H. Gentry
Arrabidaea selloi (Spreng.) Sandwith
Arrabidaea spicata Bureau & K. Schum. '
Arrabidaea trailii Sprague
Arrabidaea triplinervia (Mart, ex DC.) Baill. ex Bureau
Arrabidaea tuberculata DC.
Callichlamys latifolia (Rich.) K. Schum.
Ceratophytum tetragonolobum (Jacq.) Sprague & Sandwith
Clytostoma binatum (Thunb.) Sandwith
Clytostoma sciiiripabulum Bureau & K. Schum.
Crescentia amazonica Ducke
Crescentia cujete L. [cultivated only?]
Cuspidaria subincana A.H. Gentry
Cybistax antisyphilitica (Mart.) Mart, ex DC.
Cydista aequinoctialis (L.) Miers
Cydista lilacina A.H. Gentry
Digomphia ceratophora A.H. Gentry
Digomphia densicoma (Mart, ex DC.) Pilg.
Digomphia laurifolia Benth.
Distictella arenaria A.H. Gentry
Distictella cremersii A.H. Gentry
Distictella cuneifolia (DC.) Sandwith
Distictella dasytricha Sandwith
Distictella elongata (Vahl) Urb.
Distictella laevis (Sandwith) A.H. Gentry
Distictella magnoliifolia (Kunth) Sandwith
Distictella monophylla Sandwith
Distictella obovata Sandwith
Distictella parkeri (DC.) Sprague & Sandwith
Distictella pauciflora A.H. Gentry
Distictella porphyrotricha Sandwith
Distictis granulosa Bureau & K. Schum.
Distictis pulverulenta (Sandwith) A.H. Gentry
Godmania aesculifolia (Kunth) Standi.
Haplolophium rodriguesii AM. Gentry
Jacaranda copaia (Aubl.) D. Don
Jacaranda copaia subsp. copaia
Jacaranda copaia subsp. spectabilis (Mart, ex DC.) A.H. Gentry
Jacaranda duckei Vattimo
Jacaranda obtusifolia Bonpl.
Jacaranda obtusifolia subsp. obtusifolia
Jacaranda obtusifolia subsp. rhombifolia (G. Mey.) A.H. Gentry
Jacaranda orinocensis Sandwith
Lundia corymbifera (Vahl) Sandwith
Lundia densiflora DC.
Lundia erionema DC.
AM BO
GU
su FG
FG
AM
BO GU
GU
su? FG
BO DA GU su FG
AM BO GU su FG
BO GU FG
AM BO DA GU su FG
AM BO DA GU su FG
AM BO
BO DA GU su FG
BO
AM BO GU su FG
GU su FG
AM BO
AM BO GU su FG
AM GU
AM BO
BO
GU
GU
su
su
FG
BO GU su FG
FG
AM BO GU su FG
AM BO DA GU su FG
BO GU su
AM BO DA GU su FG
AM BO DA GU
AM BO
AM BO DA GU su FG
BO GU su
su
AM BO DA GU su FG
AM BO DA GU su FG
AM
AM BO GU
AM BO GU
AM BO
su FG
FG
AM
AM GU su FG
AM
AM BO DA GU su FG
AM
BO GU
GU
FG?
BO
BO GU
AM BO GU su? FG
AM BO GU su FG
AM BO GU
FG
AM BO DA GU su FG
AM BO DA GU su FG
AM BO DA GU su
su
FG
FG
AM BO DA GU su FG
AM BO
AM BO DA GU su FG
AM BO
BO GU FG
AM BO GU su
AM GU su FG
216 Checklist of the Plants of the Guiana Shield
Lundia puberula Pittier
Macfadyena uncata (T.F. Andrews) Sprague & Sandwith
Macfadyena unguis-cati (L.) A.H. Gentry
Manaosella cordifolia (DC.) A.H. Gentry
Mamoa alliacea (Lam.) A.H. Gentry
Mansoa hymenaea (DC.) A.H. Gentry [cultivated, escaping?]
Mamoa kerere (Aubl.) A.H. Gentry
Mansoa kerere var. incarnata (Aubl.) A.H. Gentry
Mansoa kerere var. kerere
Mansoa onohualcoides A.H. Gentry
Mansoa sp. nov. aff. standleyi
Mansoa standleyi (Steyerm.) A.H. Gentry
Mansoa verrucifera (Schltdl.) A.H. Gentry
Martinella iquitosensis A. Samp.
Martinella obovata (Kunth) Bureau & K. Schum.
Melloa quadrivalvis (Jacq.) A.H. Gentry
Memora alba (Aubl.) Miers
Memora bracteosa (DC.) Bureau ex K. Schum.
Memora cladotricha Sandwith
Memoraflavida (DC.) Bureau & K. Schum.
Memoraflaviflora (Miq.) Pulle
Memora heterophylla (Kraenzl.) Sandwith
Memora moringifolia (DC.) Sandwith
Memora patula Miers
Memora pseudopatula A.H. Gentry
Memora racemosa A.H. Gentry
Memora schomburgkii (DC.) Miers
Memora tanaeciicarpa A.H. Gentry
Mussatia hyacinthina (Standi.) Sandwith
Mussatia prieiirei (DC.) Bureau ex K. Schum.
Parabignonia steyermarkii Sandwith
Paragonia pyramidata (Rich.) Bureau
Phryganocydia corymbosa (Vent.) Bureau ex K. Schum.
Pithecocteniiim criicigerum (L.) A.H. Gentry
Pleonotoma albiflora (Salzm. ex DC.) A.H. Gentry
Pleonotoma clematis (Kunth) Miers
Pleonotoma dendrotricha Sandwith
Pleonotoma echitidea Sprague & Sandwith
Pleonotoma exserta A.H. Gentry
Pleonotomajasminifolia (Kunth) Miers
Pleonotoma moringifolia (DC.) Miers [unverified name] FG
Pleonotoma variabilis (Jacq.) Miers AM
Potamoganos microcalyx (G. Mey.) Sandwith AM
Pseudocalymma sagotii (Bureau & K. Schum.) Sandwith [unverified name]
Pyrostegia dichotoma Miers ex K. Schum. AM
Roentgenia sordida (Bureau & K. Schum.) Sprague & Sandwith
Schlegeliafuscata A.W. Gentry
Schlegelia macrophylla Ducke
Schlegelia paraensis Ducke
Schlegelia parviflora (Oerst.) Monach.
Schlegelia roseiflora Ducke
Schlegelia scandens (Briq. & Spruce) Sandwith AM
Schlegelia spruceana K. Schum. AM
Schlegelia violacea (Aubl.) Griseb. AM
Sparattosperma leucanthiim (Veil.) K. Schum.
Stizophyllum inaequilaterum Bureau & K. Schum. AM
Stizophyllum riparium (Kunth) Sandwith AM
Tabebuia aurea (Silva Manso) Benth. & Hook, f ex S. Moore
Tabebuia barbata (E. Mey.) Sandwith AM
Tabebuia capitata (Bureau & K. Schum.) Sandwith AM
Tabebuia chrysantha (Jacq.) G. Nicholson [cultivated?] AM
Tabebuia donnell-smithii Rose BO
Tabebuia fluviatilis {Auh\.) DC. DA GU SU FG
Tabebuia guayacan (Seem.) Hemsl. BO
BO
AM BO DA GU SU FG
AM BO DA GU SU FG
AM
GU SU
SU
FG
FG
AM BO DA GU SU FG
AM FG
AM BO DA GU SU FG
AM SU
FG
BO DA GU SU FG
AM BO DA GU
AM
AM
BO
GU
GU
SU
SU
FG
FG
AM BO GU SU FG
AM BO
GU SU FG
AM BO GU SU FG
AM GU
GU SU FG
BO DA
AM BO
GU SU FG
AM BO GU SU FG
AM BO GU FG
AM BO DA GU
AM BO GU SU FG
BO DA GU SU FG
GU SU FG
AM BO DA GU FG
AM BO GU SU FG
AM DA GU SU FG
AM BO GU SU FG
AM
GU SU
FG
AM BO
AM BO
BO
GU
SU
SU
FG
BO DA GU SU
BO DA GU SU FG
FG
FG
GU SU
SU
FG
FG?
FG
BO GU
BO DA GU SU FG
BO
BO GU SU FG
BO GU SU
SU
FG
BO
BO
BO
DA GU
GU
SU FG
AM BO su FG
AM BO DA GU su FG
AM BO DA GU su FG
BO DA GU su FG
AM BO DA
AM GU FG'
AM BO GU
AM BO
BO
GU
AM BO
AM BO
AM BO GU su FG
AM
BO DA GU su? FG
BO GU
AM BO GU
AM BO DA GU
GU
su FG
FG
AM FG
AM GU FG
FG
Checklist of the Plants of the Guiana Shield 217
Tabebiiia impetiginosa (Mart, ex DC.) Standi.
Tabebuia insignis (Miq.) Sandwith
Tabebiiia insignis var. insignis
Tabebuia insignis var. monophylla Sandwith
Tabebuia insignis var. pacimonensis Sandwith
Tabebuia obscura (Bureau & K. Schum.) Sandwith
Tabebuia ochracea (Cham.) Standi.
Tabebuia ochracea subsp. heterotricha (DC.) A.H. Gentry
Tabebuia ochracea subsp. neochrysantha (A.H. Gentry) A.H. Gentry
Tabebuia orinocensis (Sandwith) A.H. Gentry
Tabebuia pilosa A.U. Gentry
Tabebuia rosea (Bertol.) DC. [cultivated]
Tabebuia serratifoUa (Vahl) G Nicholson
Tabebuia sp. A
Tabebuia stenocalyx Sprague & Stapf
Tabebuia subtilis Sprague & Sandwith
Tabebuia uleana (Kraenzl.) A.H. Gentry
Tanaecium jaroba Sw.
Tanaecium nocturnum (Barb. Rodr.) Bureau & K. Schum.
Tecoma stans (L.) Juss. ex Kunth [cuhivated?]
Tynanthus polyanthus (Bureau) Sandwith
Tynanthus pubescens A.H. Gentry
Tynanthus sastrei A.Yi. Gentry
Xylophragma seemannianum (Kuntze) Sandwith AM BO DA GU SU
Synonyms and Misidentifications
Adenocalymna bilabiatum (Sprague) Sandwith =Arrabidaea bilabiata
Adenocalymna heterophyllum Kraenzl. =Memora heterophylla
Adenocalymna sagotii Bureau & K. Schum. =Pseudocalymma sagotii
Adenocalymna symmetricum Rusby =Mansoa kerere
Amphilophium molle Schltdl. & Cham. =Amphilophium paniculatum var. molle
Anemopaegma carrerense Arm. =Anemopaegma karstenii
Anomoctenium granulosum (Bureau & K. Schum.) Sandwith =Distictis granulosa
Arrabidaea obliqua (Kunth) Bureau =Arrabidaea corallina
Arrabidaea sordida Bureau & K. Schum. =Roentgenia sordida
Bignonia aequinoctialis L. =Cydista aequinoctialis
Bignonia aesculifolia Kunth =Godmania aesculifolia
Bignonia alba Aubl. =Memora alba
Bignonia albiflora Salzm. ex DC. =Pleonotoma albiflora
Bignonia alliacea Lam. =Mansoa alliacea
Bignonia apurensis Kunth =Adenocalymna apurense
Bignonia aquatilis E. Mey. =Tabebuia fluviatilis
Bignonia aurea Silva Manso =Tabebuia aurea
Bignonia barbata E. Mey. =Tabebuia barbata
Bignonia binata Thunb. =Clytostoma binatum
Bignonia brachypoda DC. =Arrabidaea brachypoda
Bignonia candicans Rich. =Arrabidaea candicans
Bignonia carichanensis Kunth =Arrabidaea carichanensis
Bignonia chamberlaynii Sims =Anemopaegma chamberlaynii
Bignonia chica Bonpl. =Arrabidaea chica
Bignonia chrysantha Jacq. =Tabebuia chrysantha
Bignonia chrysoleuca Kunth =Anemopaegma chrysoleucum
Bignonia cinnamomea DC. =Arrabidaea cinnamomea
Bignonia clematis Kunth -Pleonotoma clematis
Bignonia copaia Aubl. -Jacaranda copaia
Bignonia corallina Jacq. =Arrabidaea corallina
Bignonia cordifolia DC. =Manaosella cordifolia
Bignonia corymbifera Vahl =Lundia corymbifera
Bignonia crucigera L. =Pithecoctenium crucigerum
Bignonia digitata E. Mey. =Tabebuiafluviatilis
Bignonia echinata Aubl. =^Pithecoctenium crucigerum
Bignonia echinatum Jacq. =Pithecoctenium crucigerum
Bignonia elongata Vahl =DisticteUa elongata
Bignonia fdicifolia Anderson =Jacaranda obtusifolia subsp. rhombifolia
218 Checklist of the Plants of the Guiana Shield
Bignoniafluviatilis Aubl. =Tabebuiafluviatilis
Bignonia heterotricha DC. =Tabebuia ochracea subsp. heterotricha
Bignonia impressa Rusby =Adenocalymna impressum
Bignonia inaequalis DC. ex Splitg. =Arrabidaea inaequalis
Bignonia incarnata Aubl. =Mansoa kerere var. incarnata
Bignoniajasminifolia Kunth =Pleonotomajasminifolia
Bignonia kerere Aubl. =Mansoa kerere
Bignonia latifolia Rich. =Callichlamys latifolia
Bignonia leucantha Veil. ^Sparatlosperma leucanthum
Bignonia magnoliifolia Kunth =Distictella magnoliifolia
Bignonia martinii DC. =Martinella obovata
Bignonia microcalyx G. Mey. =Potamoganos microcalyx
Bignonia mollis Vahl =Arrabidaea mollis
Bignonia mollissima Kunth =Arrabidaea mollissima
Bignonia moringaefolia DC. =Pleonotoma moringifolia
Bignonia paniculata L. =Amphilophium paniculatum
Bignonia patellifera Schltdl. =Arrabidaea patellifera
Bignonia picta Kunth =Cydista aequinoctialis
Bignonia popidifolia DC. =MeUoa quadrivalvis
Bignonia prieurei DC. =Mussatia prieurei
Bignonia pubescens L. =Arrabidaea pubescens
Bignonia pyramidata Rich. =Paragonia pyramidata
Bignonia quadrivalvis Jacq. =Melloa quadrivalvis
Bignonia riparia Kunth =Stizophyllum riparium
Bignonia salicifolia Kunth =Anemopaegma salicifolium
Bignonia selloi Spreng. =Arrabidaea selloi
Bignonia serratifolia Vahl =Tabebuia serratifolia
Bignonia tetragonoloba Jacq. =Ceratophytum tetragonolobum
Bignonia tiliaefolia Kunth =Pithecoctenium crucigerum
Bignonia tinctoria Arruda ? ^Arrabidaea chica
Bignonia uncata Andrews =Macfadyena uncata
Bignonia unguis-cati L. =Macfadyena unguis-cati
Bignonia variabilis Jacq. =Pleonotoma variabilis
Bignonia verrucifera Schltdl. =Mansoa verrucifera
Bignonia verrucosa Kunth ^Arrabidaea carichanensis
Callichlamys riparia Miq. =Callichlamys latifolia
Ceratophytum tobagense (Urb.) Sprague & Sandwith =Ceratophytum tetragonolobum
Cremastus lachnaeus Bureau ex Bureau & K. Schum. =Arrabidaea lachnaea
Cybistax donnell-smithii (Rose) Seib. =Tabebuia donnell-smithii
Cybistax millsii Miranda =Tabebuia donnell-smithii
Distictella monophylla var. laevis Sandwith =Distictella laevis
Distictella pulverulenta Sandwith =Distictis pulverulenta
Distictella racemosa (Bureau «fe K. Schum.) Urb. =Distictella magnoliifolia
Distictis robinsonii J. Johnst. =Anemopaegma karstenii
Handroanthus durus (Bureau ex K. Schum.) Mattos =Tabebma insignis var. insignis
Jacaranda densicoma Mart, ex A. DC. =Digomphia densicoma
Jacaranda filicifolia D. Don =Jacaranda obtusifolia subsp. rhombifolia
Jacaranda filicifolia var. puberula K. Schum. =Jacaranda obtusifolia subsp. rhombifolia
Jacaranda obtusifolia var. rhombifolia (G. Mey.) Sandwith =Jacaranda obtusifolia subsp. rhombifolia
Jacaranda paraensis (Huber) Vattimo =Jacaranda copaia subsp. spectabilis
Jacaranda rhombifolia G Mey. =Jacaranda obtusifolia subsp. rhombifolia
Jacaranda spectabilis Mart, ex DC. ^Jacaranda copaia subsp. spectabilis
Jacaranda superba Pittier =Jacaranda copaia subsp. spectabilis
Lundia stricta Bureau & K. Schum. =Lundia densiflora
Martinella insculpta Sprague & Sandwith ^Martinella obovata
Martinella manaosiana A. Samp. =Martinella iquitosensis
Melloa populifolia (DC.) Britton =Melloa quadrivalvis
Memora bilabiata Sprague =Arrabidaea bilabiata
Memora ovata (Bureau & K. Schum.) Sprague & Sandwith =Memora schomburgkii
Pachyptera alliacea (Lam.) A.H. Gentry =Mansoa alliacea
Pachyptera kerere (Aubl.) Sandwith =Mansoa kerere
Pachyptera kerere var. incarnata (Aubl.) A.H. Gentry =Mansoa kerere var. incarnata
Pachyptera standleyi (Steyerm.) A.H. Gentry =Mansoa standleyi
Paramansoa grosourdyana Baill. =Arrabidaea grosourdyana
Petastoma kerere (Aubl.) Schnee =Mansoa kerere
Checklist of the Plants of the Guiana Shield 219
Petastoma patelliferum (Schltdl.) Miers =Arrabidaea patellifera
Pithecoctenium echinatum (Jacq.) Baill. =Pithecoctenium crucigerum
Pithecoctenium gramilosum (Bureau & K. Schum.) Sprague & Sandwith =Distictis granulosa
Pithecoctenium uleanum Kraenzl. =Distictis granulosa
Pseudocalymma alliaceum (Lam.) Sandwith =Mansoa alliacea
Pseudocalymma alliaceum var. macrocalyx Sandwith =Mansoa standleyi
Pseudocalymma standleyi Steyerm. =Mansoa standleyi
Pseudopaegma oligoneuron Sprague & Sandwith =Anemopaegma oligoneuron
Roentgenia tuberculata Sprague & Sandwith [name of uncertain application, apparently a nomen nudum]
Saldanhaea seemanniana Kuntze =Xylophragma seemannianum
Schizopsis polyantha Bureau =Tynanthus polyanthus
Schlegelia lilacina Miq. =Schlegelia violacea
Schlegelia parasitica (Sw.) Miers ex Griseb. [not in flora area]
Spathodea ? bracteosa DC. =Memora bracteosa
Spathodea corymbosa Vent. =Phryganocydia corymbosa
Spathodea flaviflora Miq. =Memora flaviflora
Spathodea obovata Kunth =Martinella obovata
Spathodea orinocensis Kunth =Phryganocydia corymbosa
Spathodea schomburgkii DC. =Memora schomburgkii
Tabebuia aquatilis (E. Mey.) Sprague & Sandwith =Tabebuia fluviatilis
Tabebuia caraiba (Mart.) Bureau =Tabebuia aurea
Tabebuia chrysotricha (Mart, ex DC.) Standi, [not in flora area]
Tabebuia dura (Bureau & K. Schum.) Sprague & Sandwith =Tabebuia insignis var. insignis
Tabebuiafloccosa (Bureau & K. Schum.) Sprague & Sandwith =Xylophragma seemannianum
Tabebuia glomerata Urb. =Tabebuia capitata
Tabebuia hypolepra Sprague & Sandwith =Tabebuia capitata
Tabebuia insignis var. orinocensis Sandwith =Tabebuia orinocensis
Tabebuia ipe var. integra (Sprague) Sandwith =Tabebuia impetiginosa
Tabebuiajapurensis DC. =Arrabidaeajapurensis
Tabebuia latifolia DC. =Martinella obovata
Tabebuia longipes Baker =Tabebuia insignis var. monophylla
Tabebuia millsii (Miranda) A.H. Gentry =Tabebuia donnell-smithii
Tabebuia moritziana (Kraenzl.) Schnee =Arrabidaea grosourdyana
Tabebuia neochrysantha A.H. Gentry =Tabebuia ochracea subsp. neochrysantha
Tabebuia neurophylla Miq. =Arrabidaea patellifera
Tabebuia obscura var. schultesiana (Sandwith) Sandwith =Tabebuia obscura
Tabebuia roraimae Oliv. =Tabebuia insignis var. insignis
Tabebuia rufescens J. Johnst. =Tabebuia chrysantha
Tabebuia rufinervis Hoffmanns, ex DC. =Callichlamys latifolia
Tabebuia subtilis var. schultesiana Sandwith =Tabebuia obscura
Tanaecium praelongum Miers =Tanaecium jaroba
Tecoma capitata Bureau & K. Schum. =Tabebuia capitata
Tecoma conspicua DC. =Tabebuia serratifolia
Tecoma dura Bureau ex K. Schum. =Tabebuia insignis var. insignis
Tecomafluviatilis (Aubl.) Miq. =Tabebuiafluviatilis
Tecoma guayacan Seem. =Tabebuia guayacan
Tecoma heterotricha DC. -Tabebuia ochracea subsp. heterotricha
Tecoma impetiginosa Mart, ex DC. =Tabebuia impetiginosa
Tecoma insignis Miq. =Tabebuia insignis
Tecoma meyeriana DC. =Tabebuiafluviatilis
Tecoma moritziana Kraenzl. =Arrabidaea grosourdyana
Tecoma obscura Bureau & K. Schum. =Tabebuia obscura
Tecoma ochracea Cham. =Tabebuia ochracea
Tecoma patrisiana DC. =^Tabebuia serratifolia
Tecoma roraimae (Oliv.) K. Schum. =Tabebuia insignis var insignis
Tecoma serratifolia (Vahl) G. Don =Tabebuia serratifolia
Tecoma uleana Kraenzl. =Tabebuia uleana
Tetrastichella inaequalis (DC. ex Splitg.) Pichon =Arrabidaea inaequalis
Tynanthus hyacinthinus Standi. =Mussatia hyacinthina
Tynanthus myrianthus Bureau & K. Schum. ^Tynanthus polyanthus
Zeyheria surinamensis Miq. =Paragonia pyramidata
References
Gentry, A.H. 1978. Bignoniaceae. In B. Maguire & collaborators, 1978. Botany of the Guayana Highland part 10. Mem. New
York Bot. Gard. 29: 245-283.
220 Checklist of the Plants of the Guiana Shield
Gentry, A.H. 1980. Bignoniaceae - Part I (Crescentieae and Tourretieae). Flora Neotropica Monogr. 25(1): 1-130.
Gentry, A.H. 1982. Bignoniaceae. In Z. Luces de Febres, Flora de Venezuela 8: 1-462.
Gentry, A.H. 1992. Bignoniaceae - Part 11 (Tribe Tecomeae). Flora Neotropica Monogr. 25(11): 1-335.
Gentry, A.H. 1997. Bignoniaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 403-491.
Lohmann, L.G, J.L. Brown, & S.A. Mori. 2002. Bignoniaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V
Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons.
Mem. New York Bot. Gard. 76 (2): 118-139.
Sandwith, N.Y. 1938. Bignoniaceae. In A. Pulle, Flora of Suriname 4(2): 1-86.
BlXACEAE
B.K. Hoist, J.A. Steyermarkt
GU su FG
AM BO DA GU su
AM BO DA GU su FG
AM BO GU su FG?
Bixa orellana L.
Bixa urucurana Willd. {=Bixa orellana?]
Cochlospermum orinocense (Kunth) Steud.
Cochlospermum vitifolium (Willd.) Spreng.
Synonyms and Misidentifications
Bixa orellana var. urucurana (Willd.) Kuntze ex Pilg. =Bixa urucurana
Cochlospermum insigne A. St.-Hil. [not in flora area] =Cochlospermum regium (Schr.) Pilger
Cochlospermum parkeri Planch. =Cochlospermum orinocense
Cochlospermum parviifolium Planch. =Cochlospermum orinocense
Cochlospermum triphyllum (S.F. Blake) Pittier =Cochlospermum vitifolium
Cochlospermum wentii Pulle =Cochlospermum orinocense
Maximilianea triphylla S.F. Blake =Cochlospermum vitifolium
Orellana americana var. urucurana Kuntze =Bixa urucurana
References
Heald, S.V. 2002. Bixaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 139.
Poppendieck, H.H. 1981. Cochlospermaceae. Flora Neotropica Monogr. 27: 1-33.
Steyermark, J.A., & B.K. Hoist. 1997. Bixaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana
3: 492-495.
BOMBACACEAE
W.S. Alverson, L.J. Dorr, J.A. Steyermarkt
Bombacopsis nervosa (Uittien) A. Robyns
Catostemma altsonii Sandwith
Catostemma commune Sandwith
Catostemma durifolius W.S. Alverson
Catostemma ebracteolatum Steyerm.
Catostemmafragrans Benth.
Catostemma hirsutulum Steyerm.
Catostemma marahuacense Steyerm.
Catostemma sancarlosianum Steyerm.
Cavanillesia sp. A
Ceiba pentandra (L.) Gaertn.
Eriotheca crassa (Uittien) A. Robyns
Eriotheca globosa (Aubl.) A. Robyns
Eriotheca longitubulosa A. Robyns
Eriotheca surinamensis (Uittien) A. Robyns
Huberodendron swietenioides (Gleason) Ducke
Matisia amplifolia Pittier
Matisia ochrocalyx K. Schum.
Ochroma pyramidale (Cav. ex Lam.) Urb.
Pachira amazonica (A. Robyns) W.S. Alverson
Pachira aquatica Aubl.
Pachira aracamuniana (Steyerm.) W.S. Alverson
Pachira coriacea (Mart.) W.S. Alverson
Pachira cowanii (A. Robyns) W.S. Alverson
Pachira dolichocalyx A. Robyns
Pachirafaroensis (Ducke) W.S. Alverson
Pachira flaviflora (Pulle) Fern. Alonso
AM BO
BO
AM BO
AM BO
AM
AM
AM
AM BO
BO
BO
AM BO
BO
AM BO
AM BO
AM
AM
AM
DA
GU
GU
GU
GU
SU
su
FG
FG
DA GU SU FG
su FG
DA GU SU FG
FG
GU su FG
FG
GU su FG
GU FG
AM
DA GU SU FG
GU FG
GU SU FG
Checklist of the Plants of the Guiana Shield 221
Pachirafuscolepidota (Steyerm.) W.S. Alverson AM
Pachira gracilis (A. Robyns) W.S. Alverson AM BO
Pachira gracilis subsp. bolivarensis (Steyerm.) W.S. Alverson BO
Pachira gracilis subsp. gracilis AM
Pachira humilis Spruce ex Decne. AM
Pachira insignis (Sw.) Sw. ex Savigny GU SU FG
Pachira liesneri (Steyerm.) W.S. Alverson AM
Pachira macrocalyx (Ducke) Fern. Alonso FG?
Pachira mawarinumae (Steyerm.) W.S. Alverson AM
Pachira minor (Sims) Hemsl. [Sierra de la Neblina] AM BO DA GU SU
Pachira nitida Kunth AM
Pachira obovata (A. Robyns) W.S. Alverson [Sierra de la Neblina] AM
Pachira orinocensis (A. Robyns) W.S. Alverson AM BO
Pachira paracusis (Ducke) W.S. Alverson AM
Pachira pseudofaroensis (A. Robyns) W.S. Alverson AM
Pachira quinata (Jacq.) W.S. Alverson AM BO GU
Pachira robynsii (Steyerm. & W.D. Stevens) W.S. Alverson [Sierra de la AM
Neblina]
Pachira rupicola (A. Robyns) W.S. Alverson BO
Pachira sordida (R.E. Schult.) W.S. Alverson
^
AM
Pachira tepuiensis (Steyerm.) W.S. Alverson BO
Pachira trinitensis Urb. BO
Pachira yapacanae Steyerm. ex W.S. Alverson AM
Pseudobombax croizatii A. Robyns AM BO
Quararibea amazonica Ulbr. FG
Quararibea duckei Huber SU FG
Quararibea guianensis Aubl. GU SU FG
Quararibea lasiocalyx (K. Schum.) Vischer FG
Quararibea spatulata Ducke FG
Scleronema guianense Sandwith GU
Scleronema micranthum (Ducke) Ducke [Sierra de la Neblina] AM
Synonyms and Misidentifications
Bombacopsis amazonica A. Robyns =Pachira amazonica
Bombacopsis coriacea A. Robyns =Pachira robynsii
Bombacopsis cowanii A. Robyns =Pachira cowanii
Bombacopsis fendleri (Seem.) Pittier =Pachira quinata
Bombacopsis jaris Pittier [nomen nudum] =Pachira quinata
Bombacopsis jenmanii (Oliv.) Lasser =Pachira minor
Bombacopsis macrocalyx (Ducke) A. Robyns =Pachira macrocalyx
Bombacopsis mucronulata Pittier =Pachira trinitensis
Bombacopsis obovata A. Robyns =Pachira obovata
Bombacopsis orinocensis A. Robyns =Pachira orinocensis
Bombacopsis pachiroides auct. non Pittier: sensu LI. Williams =Pachira quinata
Bombacopsis pachiroides auct. non Pittier: LI. Williams =Pachira rupicola
Bombacopsis pachiroides Pittier =Pachira trinitensis
Bombacopsis paraensis (Ducke) A. Robyns =Pachira paraensis
Bombacopsis pseudamazonica A. Robyns =Pachira amazonica
Bombacopsis quinata (Jacq.) Dugand =Pachira quinata
Bombacopsis rupicola A. Robyns =Pachira rupicola
Bombacopsis sepium Pittier =Pachira quinata
Bombacopsis trinitensis (Urb.) A. Robyns =Pachira trinitensis
Bombacopsis trinitensis var. mucronulata (Pittier) A. Robyns =Pachira trinitensis
Bombacopsis wurdackii A. Robyns =Pachira amazonica
Bombax aquaticum (Aubl.) K. Schum. =Pachira aquatica
Bombax coriaceum Mart. =Pachira coriacea
Bombax crassum Uittien =Eriotheca crassa
Bombaxfaroense Ducke =Pachirafaroensis
Bombaxflaviflorum Pulle =Pachira flaviflora
Bombax globosum Aubl. =Eriotheca globosa
Bombax humile (Spruce ex Decne.) Benth. ex K. Schum. =Pachira humilis
Bombaxjenmanii Oliv. =Pachira minor
Bombax macrocarpum (Schltdl. & Cham.) K. Schum. =Pachira aquatica
Bombax minus (Sims) Ducke =Pachira minor
222 Checklist of the Plants of the Guiana Shield
Bombax nervosum Uittien =Bombacopsis nervosa
Bombax orinocense Kunth (Bixaceae) =Cochlospermum orinocense
Bombax paraense Ducke =Pachira paraensis
Bombax pentandrum L. =Ceiba pentandra
Bombax poissonianum K. Schum. =Pachira nitida
Bombax pyramidale Cav. ex Lam. =Ochroma pyramidale
Bombax quinatum Jacq. =Pachira quinata
Bombax rigidifolium Ducke =Pachira aquatica
Bombax sordidum R.E. Schult. =Pachira sordida
Bombax spectabile Ulbr. =Pachira insignis
Bombax surinamense Uittien =Eriotheca surinamensis
Bombax vitifolium Willd. (Bixaceae) =Cochlospermum vitifolium
Carolinea macrocarpa Schltdl. & Cham. =Pachira aquatica
Carolinea minor Sims =Pachira minor
Catostemma clarkii Steyerm. =Catostemma sancarlosianum
Catostemma macrosperma (Sagot) Lemee =Catostemmafragrans
Catostemma micranthum Ducke =Scleronema micranthum
Catostemma pubistylum Steyerm. =Catostemma sancarlosianum
Ceiba occidentalis (Spreng.) Burkill =Ceiba pentandra
Matisia apaporiensis Cuatrec. =Matisia ochrocalyx
Matisia lasiocalyx K. Schum. =Quararibea lasiocalyx
Matisia muricata (Cuatrec.) Cuatrec. =Matisia ochrocalyx
Ochroma lagopus Sw. =Ochroma pyramidale
Pachira cardonae Cuatrec. =Pachira minor
Pachira fendleri Seem. ^Pachira quinata
Pachirajenmanii (Oliv.) A. Terrace. =Pachira minor
Pachira longiflora Hook. =Pachira aquatica
Pachira macrocarpa (Schhdl. & Cham.) Walp. =Pachira aquatica
Pachira obtusa Spruce ex K. Schum. =Pachira nitida
Pachira pustulifera Pittier =Pachira aquatica
Pachira villosula Pittier =Pachira aquatica
Pochota amazonica (A. Robyns) Steyerm. 8c W.D. Stevens =Pachira amazonica
Pochota aracamuniana Steyerm. =Pachira aracamuniana
Pochota coriacea (Mart.) Steyerm. & W.D. Stevens =Pachira coriacea
Pochota coriacea subsp. orinocensis (A. Robyns) Steyerm. & W.D. Stevens =Pachira coriacea
Pochota cowanii (A. Robyns) Steyerm. & W.D. Stevens =Pachira cowanii
Pochota ewelii Steyerm. =Pachira amazonica
Pochota flaviflora (Pulle) Steyerm. & W.D. Stevens =Pachira flaviflora
Pochotafuscolepidota Steyerm. =Pachirafuscolepidota
Pochota gracilis (A. Robyns) Steyerm. & W.D. Stevens =Pachira gracilis
Pochota gracilis subsp. bolivarensis Steyerm. =Pachira gracilis subsp. bolivarensis
Pochota humilis (Spruce ex Decne.) Steyerm. & W.D. Stevens =Pachira humilis
Pochota liesneri Steyerm. =Pachira liesneri
Pochota maguirei (A. Robyns) Steyerm. & W.D. Stevens =Pachira gracilis subsp. bolivarensis
Pochota mawarinumae Steyerm. =Pachira mawarinumae
Pochota minor (Sims) Steyerm. & W.D. Stevens =Pachira minor
Pochota nitida (Kunth) Steyerm. & W.D. Stevens =Pachira nitida
Pochota obovata (A. Robyns) Steyerm. & W.D. Stevens =Pachira obovata
Pochota orinocensis (A. Robyns) Steyerm. & W.D. Stevens =Pachira orinocensis
Pochota pseudamazonica (A. Robyns) Steyerm. & W.D. Stevens =Pachira amazonica
Pochota pseudofaroensis (A. Robyns) Steyerm. & W.D. Stevens =Pachira pseudofaroensis
Pochota quinata (Jacq.) W.D. Stevens =Pachira quinata
Pochota redmondii Steyerm. =Pachira obovata
Pochota robynsii Steyerm. & W.D. Stevens =Pachira robynsii
Pochota sordida (R.E. Schult.) Steyerm. & W.D. Stevens =Pachira sordida
Pochota tepuiensis Steyerm. =Pachira tepuiensis
Pochota trinitensis (Urb.) Steyerm. & W.D. Stevens =Pachira trinitensis
Pochota vulgaris Ram. Goyena =Pachira quinata
Pochota wurdackii (A. Robyns) Steyerm. & W.D. Stevens ^Pachira amazonica
Quararibea apaporiensis (Cuatrec.) A. Robyns =Matisia ochrocalyx
Quararibea muricata Cuatrec. =Matisia ochrocalyx
Quararibea ochrocalyx (K. Schum.) Vischer =Matisia ochrocalyx
Quararibea turbinata Poir. [specimens apparently referable to Quararibea duckei]
Rhodognaphalopsis coriacea (Mart.) A. Robyns =Pachira coriacea
Rhodognaphalopsis coriacea subsp. orinocensis A. Robyns =Pachira coriacea
Checklist of the Plants of the Guiana Shield 223
Rhodognaphalopsis coriacea var. sordida (R.E. Schult.) A. Robyns =Pachira sordida
Rhodognaphalopsis discolor A. Robyns =Pachira sordida
Rhodognaphalopsis faroensis (Ducke) A. Robyns =Pachirafaroensis
Rhodognaphalopsis flaviflora vox.flaviflora =Pachira flaviflora
Rhodognaphalopsis flaviflora var. glabra A. Robyns =Pachira flaviflora
Rhodognaphalopsis gracilis A. Robyns =Pachira gracilis
Rhodognaphalopsis humilis (Spruce ex Decne.) A. Robyns =Pachira humilis
Rhodognaphalopsis maguirei A. Robyns =Pachira gracilis subsp. bolivarensis
Rhodognaphalopsis minor (Sims) A. Robyns =Pachira minor
Rhodognaphalopsis minor var. amblyophylla A. Robyns =Pachira minor
Rhodognaphalopsis minor var. minor =Pachira minor
Rhodognaphalopsis nitida (Kunth) A. Robyns =Pachira nitida
Rhodognaphalopsis pseudofaroensis A. Robyns =Pachira pseudofaroensis
Scleronema neblinense Steyerm. =Scleronema micranthum
Sophia Carolina L. =Pachira aquatica
References
Alverson, W.S. 1994. New species and combinations of Catostemma and Pachira (Bombacaceae) from the Venezuelan Guayana.
Novon4(l):3-8.
Alverson, W.S. & J.A. Steyermark. 1997. Bombaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan
Guayana 3: 496-527.
Alverson, W.S. & S.A. Mori. 2002. Bombaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hofif, &
J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot.
Gard. 76(2): 139-145.
Robyns, A. 1963. Essai de monographic du genre Bombax s.l. Bull. Jard. Bot. Etat 33: 1-311.
Robyns, A. 1967. Bombacaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 7. Mem. New York Bot.
Gard. 17(1): 1-439.
Steyermark, J.A. 1987. Bombacaceae. In Flora of the Venezuelan Guayana - III. Ann. Missouri Bot. Gard. 74(3): 636-646.
Steyermark, J.A. & W.D. Stevens. 1988. Notes on Rhodognaphalopsis and Bombacopsis in the Guayanas. Ann. Missouri Bot.
Gard. 75: 396-398.
Uittien, H. 1932. Bombacaceae. In A. PuUe, Flora of Suriname 3(1): 26-33.
BONNETIACEAE
Archytaea angustifolia Maguire
Archytaea triflora Mart.
Bonnetia ahogadoi (Steyerm.) A.L. Weitzman & P.F. Stevens
Bonnetia bolivarensis Steyerm.
Bonnetia celiac Maguire
Bonnetia chimantensis Steyerm.
Bonnetia cordifolia Maguire
Bonnetia crassa Gleason
Bonnetia euryanthera Steyerm.
Bonnetiafasciculata A.L. Weitzman & P.F. Stevens
Bonnetia huberiana Steyerm.
Bonnetiajauaensis Maguire
Bonnetia kathleenae Lasser
Bonnetia lanceifolia Kobuski
Bonnetia liesneri Steyerm.
Bonnetia maguireorum Steyerm.
Bonnetia multinervia (Maguire) Steyerm. [expected in GU?]
Bonnetia neblinae Maguire
Bonnetia paniculata Spruce ex Benth.
Bonnetia ptariensis Steyerm.
Bonnetia roraimae 01 i v.
Bonnetia roseiflora Maguire
Bonnetia nibicunda (Sastre) A.L. Weitzman & P.F. Stevens
Bonnetia sessilis Benth.
Bonnetia steyermarkii Kobuski
Bonnetia tepuiensis Kobuski & Steyerm.
Bonnetia tristyla Gleason
Bonnetia wurdackii Maguire
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
GU
GU
GU
GU
GU
GU
Acopanea ahogadoi Steyerm.
Synonyms and Misidentifications
~-Bonnetia ahogadoi
224 Checklist of the Plants of the Guiana Shield
Archytaea multiflora Benth. =Archytaea triflora
Bonnetia dinizii Huber =Bonnetia paniculata
Bonnetia duidae Kobuski & Steyerm. (Clusiaceae) =Neotatea duidae
Bonnetia guaiquinimae Steyerm. =Bonnetia lanceifolia
Bonnetia longifolia Gleason (Clusiaceae) =Neotatea longifolia
Bonnetia martiana Maguire =Bonnetia sessilis
Bonnetia martiana var. venadoensis Maguire =Bonnetia sessilis
Bonnetia neblinensis Steyerm. (Clusiaceae) =Neotatea neblinae
Bonnetia parviflora Spruce ex Benth. =Bonnetia paniculata
Bonnetia phelpsii Gleason =Bonnetia sessilis
Bonnetia tepuiensis subsp. minor Steyerm. =Bonnetia tepuiensis
Bonnetia toronoensis Steyerm. ^'Bonnetia chimantensis
Bonnetia tristyla subsp. nervosa Steyerm. =Bonnetia tristyla
Kieseria sessilis (Benth.) Kuntze =Bonnetia sessilis
Neblinaria celiae Maguire =Bonnetia maguireorum
Neblinaria rubicunda Sastre =Bonnetia rubicunda
Neogleasonia duidae (Kobuski & Steyerm.) Maguire (Clusiaceae) =Neotatea duidae
Neogleasonia multinervia Maguire =Bonnetia multinervia
Neogleasonia wurdackii Maguire =Bonnetiafasciculata
References
Kobuski, C.E. 1948. Studies in the Theaceae, XVII. A review of the genus Bonnetia. J. Am. Arb. 29: 393-413.
Maguire, B. 1972. Bonnetiaceae. In B. Maguire &. collaborators, Botany of the Guayana Highland part 9. Mem. New York Bot.
Gard. 23: 1-832.
BORAGINACEAE
C. Feuillet, J. Gaviria, R. Gomez, J.S. Miller, G. Rodriguez
Bourreria cumanensis (Loefl.) O.E. Schulz
Cordia alliodora (Ruiz & Pav.) Oken
Cordia bicolor DC. AM
Cordia bullata (L.) Roem. & Schult.
Cordia bullata subsp. humilis (Jacq.) Gaviria
Cordia cabanayensis Gaviria
Cordia collococca Sandmark ex L.
Cordia cremersii Feuillet
Cordia curassavica (Jacq.) Roem. & Schult. AM
Cordia dentata Poir.
Cordia exaltata Lam.
Cordiafallax I.M. Johnst.
Cordiafanchoniae Feuillet FG
Cordiafulva I.M. Johnst. SU FG
Cordia goeldiana Huber FG
Corfi^/a gra«J//7ora (Desv.) Kunth AM BO DA GU
Cordia koemarae J.S. Mill. SU
Cordia laevifrons I.M. Johnst. SU FG
Cordia lomatoloba IM. ]ohnst. BO GU SU FG
Cordia marioniae Feuillet GU
Cordia multispicata Cham. GU
Cordia naidophila I.M. Johnst. AM FG
Cordia nervosa Lam. GU SU FG
Cordia nodosa Lsim. AM BO DA GU SU FG
Cordia panamens is Riley BO DA
Cordia panicularis Rudge GU SU FG
Cordia polycephala{L2im.)\M.io\m^i. BO GU SU FG
Cordia polystachya Kunth AM
Cordia roraimae I.M. Johnst. BO
Cordia sagotiilM.iolmsi. BO GU SU FG
Cordia scabrifolia DC. AM GU
Cordia schomburgkii DC. BO GU SU FG
Cordia sericicalyx DC. BO DA GU SU
Cordia sipapoi Gaviria AM
Cordia sp. A AM
Cordia sprucei Mez AM FG
BO
BO DA GU SU FG
BO GU SU FG
BO DA GU
BO DA GU
BO
BO DA GU
FG
BO DA GU SU FG
BO DA
BO DA GU SU FG
BO DA GU
AM
BO
BO DA GU su
su
FG
FG
BO GU FG
BO
BO
GU
BO
GU su FG?
AM BO GU su FG
AM BO
BO
BO
DA GU su
su
FG
AM BO
BO
BO
DA GU
GU
BO GU su
AM BO
BO
DA GU
GU
su FG
BO DA
DA
GU
GU
su FG
BO GU
GU
su FG
FG
GU su FG
GU su FG
AM BO GU su FG
BO GU FG
Checklist of the Plants of the Guiana Shield 225
Cordia stenostachya Killip ex Gaviria
Cordia steyermarkii Gaviria
Cordia tetrandra Aubl.
Cordia tomentosa Lam. ex Roem. & Schult.
Cordia toqueve Aubl. [expected in SU]
Cordia umbellifera Killip ex G Agostini
Cordia williamsii G. Agostini ex Gaviria
Cordia wurdackiana Feuillet
Heliotropium angiospermum Murray
Heliotropium curassavicum L.
Heliotropium filiforme Lehm.
Heliotropium indicum L. [cultivated, persists as an escape]
Heliotropium lagoense (Warm.) Giirke
Heliotropium polyphyllum Lehm.
Heliotropium procumbens Mill.
Heliotropium purdiei I.M. Johnst.
Heliotropium ternatum Vahl
Lepidocordia punctata Ducke
Tournefortia bicolor Sw.
Tournefortia candidula (Miers) LM. Johnst.
Tournefortia cuspidata Kunth
Tournefortia hirsutissima L.
Tournefortia maculata Jacq.
Tournefortia melanochaeta DC.
Tournefortia paniculata Cham.
Tournefortia paniculata var. spigeliiflora (DC.) I.M. Johnst.
Tournefortia ulei Vaupel
Tournefortia volubilis L.
Synonyms and Misidentifications
Bourreria viridis Rusby =Cordia sericicalyx
Cerdana alliodora Ruiz & Pav. =Cordia alliodora
Cordia alba auct. non DC. =Cordia dentata
Cordia alliodora (Ruiz & Pav.) Cham, ex DC. =Cordia alliodora (Ruiz & Pav.) Oken
Cordia aubletii DC. =Cordia schomburgkii
Cordia canescens Willd. ex Roem. & Schult. =Cordia polystachya
Cordia carnosa Rusby =Cordia bicolor
Cordia collococa Aubl. non L. =Cordia nodosa
Cordia cordifolia Kunth =Cordia tetrandra
Cordia cordifolia Willd. ex Roem. & Schuh. [nom. superfl.] =Cordia tetrandra
Cordia coriacea Sagot ex Benoist =Cordia sagotii
Cordia corymbosa Willd. ex Roem. & Schult. [nom. superfl.] =Cordia polycephala
Cordia corymbosa (Desv.) G. Don =Cordia polycephala
Cordia dasycephala (Desv.) Kunth =Cordia bullata subsp. humilis
Cordia dentata Vahl =Cordia dentata
Cordia exaltata var. melanoneura (Klotzsch) I.M. Johnst. =Cordia exaltata
Cordiaflavescens Aubl. [name of uncertain application, referable to Lauraceae]
Cordiaformicarum Willd. ex Roem. & Schult. =Cordia nodosa
Cordia globosa (Jacq.) Kunth =Cordia bullata subsp. humilis
Cordia globosa var. humilis (Jacq.) I.M. Johnst. =Cordia bullata subsp. humilis
Cordia globosa subsp. humilis (Jacq.) Borh. =Cordia bullata subsp. humilis
Cordia guianensis Klotzsch [nomen nudum] =Cordia fallax
Cordia gujanens is (Desv.) Roem. & Schult. =Cordiafallax
Cordia hebecarpa DC. =Cordia toqueve
Cordia heterophylla Willd. ex Roem. & Schult. [nom. superfl.] =Cordia toqueve
Cordia heterophylla Poir. =Cordia toqueve
Cordia hirta I.M. Johnst. =Cordia sagotii
Cordia hispidissima DC. =^Cordia nodosa
Cordia humilis (Jacq.) G. Don =Cordia bullata subsp. humilis
Cordia ierensis Britton =Cordia sericicalyx
Cordia lockhartii Kuntze [nom. inval.] =Cordia bicolor
Cordia lucida Splitg. ex Pulle [nomen nudum] =Cordia schomburgkii
Cordia macrostachya (Jacq.) Roem. & Schult. =Cordia curassavica
Cordia melanoneura Klotzsch [nomen nudum] =Cordia exaltata
Cordia miranda DC. =Cordia nodosa
226 Checklist of the Plants of the Guiana Shield
Cordia monosperma (Jacq.) Roem. & Schult. =^Cordia polycephala
Cordia opaca Rusby =Cordia sericicalyx
Cordia pubescens Willd. ex Roem. & Schult. =Cordia toqueve
Cordia scabrida Mart. =Cordia exaltata
Cordia scabrida auct. non Fresen. 1857. =Cordia naidophila
Cordia scandens Poir. (Dichapetalaceae) =Dichapetalum rugosum
Cordia spinescens L. [not in flora area]
Cordia tetraphylla Auhl. (Combretaceae) =Biichenavia tetraphylla
Cordia tobagensis Urb. =Cordia schomburgkii
Cordia tobagensis var. broadwayi Urb. =Cordia schomburgkii
Cordia umbraculifera DC. =Cordia tetrandra
Cordia viridis (Rusby) I.M. Johnst. =Cordia sericicalyx
Cordia volubilis Pittier =Cordia nodosa
Crematomia ciimanensis (Loefl.) Miers =Bourreria cumanensis
Heliotropiiim foliosum Roem. & Schult. =Heliotropiiim polyphyllum
Heliotropium fiimana (Fresen.) Gurke ^Heliotropiiim ternatum
Heliotropiiim ottonis Lehm. =Heliotropiiim ternatum
Heliotropium scorpioides Kunth =Heliotropium angiospermum
Heliotropium trinitensis Urb. =Heliotropium lagoense
Lithocardium sprucei (Mez) Kuntze =Cordia sprucei
Messerschmidia candidula Miers =Tournefortia candidula
Schleideniafumana Fresen. =Heliofropium ternatum
Schleidenia lagoensis Warm. =Heliotropium lagoense
Schleidenia polyphylla (Lehm.) Fresen. ^Heliotropium polyphyllum
Tournefortia aubletii J.F. Macbr. =Toiirnefortia bicolor
Tournefortia bicolor var. bicolor =Tournefortia bicolor
Tournefortia bicolor var. calycosa J.D. Sm. =Tournefortia bicolor
Tournefortia hirsutissima auct. non L.: sensu Pulle =Tournefortia cuspidata
Tournefortia hirsutissima L. f. bicolor (Sw.) D.L. Nash =Tournefortia bicolor
Tournefortia syringifolia Vahl ^Tournefortia maculata
Tournefortia tenuiflora Willd. ex Roem. & Schult. (Rubiaceae) =Declieuxia tenuiflora
Varronia bullata L. =Cordia bullata
Varronia corymbosa Desv. [nomen nudum] =Cordia polycephala
Varronia curassavica Jacq. =Cordia curassavica
Varronia dasycephala Desv. =Cordia bullata subsp. humilis
Varronia globosa Jacq. "^Cordia bullata subsp. humilis
Varronia grandiflora Desv. =Cordia grandiflora
Varronia guianensis Desv. =Cordia curassavica
Varronia humilis Jacq. =Cordia bullata subsp. humilis
Varronia lineata auct. non L. [nom. illeg.] =Cordia polycephala
Varronia macrostachya Jacq. =Cordia curassavica
Varronia monosperma Jacq. =Cordia polycephala
Varronia polycephala Lam. =Cordia polycephala
References
Diane N., H. Forther & H.H. Hilger. 2002. A systematic analysis of Heliotropium, Tournefortia, and allied taxa of the
Heliotropiaceae (Boraginales) based on ITSl sequences and morphological data. Am. J. Bot. 89: 287-295.
Feuillet, C. 2002. Boraginaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
145-151.
Johnston, I.M. 1928. Studies in the Boraginaceae VII. 1. The South American species of Heliotropium. Contrib. Gray Herb. 81
:
3-72.
Johnston, I.M. 1935. Studies in Boraginaceae X. The Boraginaceae of northeastern South America. J. Am. Arb. 16: 1-64.
Johnston, I.M. 1936. Boraginaceae. In A. Pulle, Flora of Suriname 4(1): 306-333.
Miller, J.S., J. Gaviria, R. Gomez, & G. Rodriguez. 1997. Boraginaceae. In J.A. Steyermark, PE. Berry, & B.K. Hoist, Flora of
the Venezuelan Guayana 3: 527-547.
Uittien, H. 1937. Boraginaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 496-497.
Brassicaceae
J.C. Lindeman, L.J. Raechal
Brassicajimcea (L.) Czem. & Coss. [introduced] FG
Cardamineflexuosa With. BO
Coronopus didymus (L.) Sm. [probably introduced] FG
Checklist of the Plants of the Guiana Shield 227
su
su
su
FG
FG
Lepidiiim virginicum L. [introduced]
Rorippa dubia (Pers.) H. Hara [introduced] [=Rorippa indica (L.) Hiem] GU
Rorippa nasturtium-aquaticum (L.) Hayek, [introduced] [=Nastnrtinm
officinale R. Br.]
Rorippa palustris (L.) Besser [introduced] [=Rorippa islandica (Leder)
Borbas]
Sinapis arvensis L. [introduced] GU
Synonyms and Misidentifications
Lepidiiim micropterum Miq. =Lepidiiim virginicum
Nasturtium pahistre (L.) DC., non Crantz =Rorippa palustris
Rorippa heterophylla (Blume) R.O. Williams =Rorippa dubia
References
Al-Shehbaz, I. 1988. Cruciferae. In R.A. Howard, Flora of the Lesser Antilles 4: 276-292.
Mitchell, J.D. 2002 Brassicaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
151.
Raechal, L.J. 1997. Brassicaceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 547-548.
Brunelliaceae
J.S. Miller
Brunellia comocladifolia Bonpl. BO
Brunellia comocladifolia suhsp. ptariana (Steyerm.) Cuatrec. [Mt. Roraima] BO
Brunellia neblinensis Steyerm. & Cuatrec. AM
Synonyms and Misidentifications
Brunellia integrifolia Szyszyl. var. ptariana Steyerm. =Brunellia comocladifolia subsp. ptariana
References
Cuatrecasas, J. 1970. Brunelliaceae. Flora Neotropica Monogr. 2: 1-188.
Miller, J.S. 1997. Brunelliaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 676-688.
BURSERACEAE
D.C. Daly
Bursera karsteniana Engl.
Bursera simaruba (L.) Sarg.
Bursera tomentosa (Jacq.) Triana & Planch.
Commiphora leptophloeos (Mart.) J.B. Gillett
Crepidospermum goudotianum (Tul.) Triana & Planch.
Crepidospermum prancei Daly
Crepidospermum rhoifolium (Benth.) Triana & Planch.
Dacryodes belemensis Cuatrec.
Dacryodes chimantensis Steyerm. & Maguire
Dacryodes cuspidata (Cuatrec.) Daly
Dacryodes glabra (Steyerm.) Cuatrec.
Dacryodes microcarpa Cuatrec.
Dacryodes negrensis Daly & M.C. Martinez
Dacryodes nitens Cuatrec.
Dacryodes peruviana (Loes.) H.J. Lam
Dacryodes roraimensis Cuatrec.
Dacryodes sclerophylla Cuatrec.
Dacryodes sp. A
Dacryodes steyermarkii Sandwith
Protium altsonii Sandwith
Protiitm amazonicum (Cuatrec.) Daly
Protium apiculatum Swart
Protium aracouchini (Aubl.) Marchand
Protium boomii Daly
Protium carolense Daly
Protium crassipetalum Cuatrec.
Protium crenatum Sandwith
Protium cuneatum Swart
BO
AM BO
BO
AM BO
AM
AM BO
AM BO
AM
AM BO
AM BO
AM BO
AM BO
AM BO
BO
BO
AM BO
BO
AM BO
AM
A \/l
BO
AM BO
BO
AM BO
AM BO
BO
GU FG
GU FG
GU SU
su?
FG
GU
FG
DA
GU SU FG
GU FG
GU
GU
GU SU FG
FG
SU FG
GU SU FG
GU
GU SU?
GU
GU SU? FG
228 Checklist of the Plants of the Guiana Shield
Protium decandrum (Aubl.) Marchand
Prothim demerarense Swart
Protium divaricatum Engl.
Protium divaricatum subsp. divaricatum
Protium divaricatum suhsp.fumarium Daly
Protium elegans Engl.
Protium ferrugineum Engl.
Protium gallicum Daly
Protium giganteum Engl.
Protium giganteum var. crassifolium (Engl.) Daly
Protium giganteum var. giganteum
Protium glabrescens Swart
Protium grandifolium Engl.
Protium guianense (Aubl.) Marchand
Protium heptaphyllum (Aubl.) Marchand
Protium heptaphyllum subsp. heptaphyllum
Protium heptaphyllum subsp. ulei (Swart) Daly
Protium inodorum Daly
Protium llanorum Cuatrec.
Protium macrocarpum Cuatrec.
Protium macrosepalum Swart
Protium melinonis Engl.
Protium morii Daly
Protium nitidifolium (Cuatrec.) Daly
Protium nodulosum Swart
Protium occultum Daly
Protium opacum Swart
Protium opacum subsp. exaggeratum Daly
Protium opacum subsp. opacum
Protium opacum subsp. rabelianum Daly
Protium pallidum Cuatrec.
Protium paniculatum Engl.
Protium paniculatum var. A
Protium paniculatum var. paniculatum
Protium paniculatum var. riedelianum (Engl.) Daly
Protium pilosum (Cuatrec.) Daly
Protium plagiocarpium Benoist
Protium poeppigianum Swart
Protium polybotryum (Turcz.) Engl.
Protium polybotryum subsp. blackii (Swart) Daly
Protium polybotryum subsp. polybotryum
Protium ptarianum Steyerm.
Protium pullei Swart [unverified name]
Protium reticulatum (Engl.) Engl.
Protium robiistum (Swart) D.M. Porter
Protium sagotianum Marchand
Protium sp. A
Protium sp. B
Protium sp. C
Protium sp. D
Protium sp. E
Protium sp. F
Protium spruceanum (Benth.) Engl.
Protium strumosum Daly
Protium subserratum (Engl.) Engl.
Protium subserratum subsp. A
Protium subserratum subsp. subserratum
Protium tenuifolium (Engl.) Engl.
Protium trifoliolatum Engl.
Protium unifoliolatum Spruce ex Engl.
Tetragastris altissima (Aubl.) Swart
Tetragastris hostmannii (Engl.) Kuntze
Tetragastris panamensis (Engl.) Kuntze
Trattinnickia boliviana (Swart) Daly
AM BO DA GU
GU
SU FG
FG
AM BO FG
AM BO FG
FG
AM FG?
AM BO
FG
AM GU SU
SU
FG
FG
AM GU SU
SU?
FG
AM BO FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM BO
FG
AM BO GU
AM BO
AM
AM BO
AM BO
AM BO
AM
AM
BO
AM BO
AM BO
AM BO
BO
AM
BO
AM
AM BO
AM
A A/T
AM
AM BO
AM BO
AM BO
BO
BO
AM BO
AM BO
DA
DA
DA
DA
GU
GU?
GU
GU
GU
SU
FG
FG
FG
FG
FG
FG
FG
FG
GU? SU? FG
FG
GU SU FG
GU
GU SU
SU
FG
GU SU FG
GU SU FG
GU FG
GU SU FG
SU FG
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU SU FG
FG?
Checklist of the Plants of the Guiana Shield 229
Trattinnickia biirserifoUa Mart. AM
Trattinnickia demerarae Sandwith
Trattinnickia glaziovii Swart
Trattinnickia rhoifolia Willd.
Synonyms and Misidentifications
Bursera aracouchili (Aubl.) Baill. =Protium aracouchini
Bursera bonairensis Bold. =Bursera simaruba
Bursera caudata Turcz. [nom. confus.] =Protium sagotianum
Bursera decandra (Aubl.) Baill. =Protium decandrum
Bursera gummifera L. =Bursera simaruba
Bursera integerrima (Tul.) Triana & Planch. ^Bursera simaruba
Bursera leptophloeos Mart. =Commiphora leptophloeos
Bursera martiana Engl. =Commiphora leptophloeos
Bursera orinocensis Engl. =Commiphora leptophloeos
Bursera subpubescens (Rose) Engl. =Bursera simaruba
Bursera trinitensis R.O. Williams =Bursera karsteniana
Bursera velutinifolia R.S. Cowan (Anacardiaceae) =Cyrtocarpa velutinifolia
Burseria gummifera (L.) Jacq. =Bursera simaruba
Dacryodes microcarpa var. lanceolata Cuatrec. =Dacryodes microcarpa
Dacryodes peruviana var. caroniensis Cuatrec. =Dacryodes peruviana
Elaphrium decandrum (Aubl.) Spreng. ex D. Dietr. =Protium decandrum
Elaphrium enneandrum Spreng. ex D. Dietr. =Protium decandrum
Elaphrium gujanense (Aubl.) Spreng. ex D. Dietr. =Protium guianense
Elaphrium heptaphyllum (Aubl.) Spreng. ex D. Dietr. =Protium heptaphyllum
Elaphrium heterophyllum (Willd.) Spreng. ex D. Dietr. =Protium aracouchini
Elaphrium integerrimum Tul. =Bursera simaruba
Elaphriumjacquinianum Kunth =Bursera tomentosa
Elaphrium simaruba (L.) Rose =Bursera simaruba
Elaphrium subpubescens Rose =Bursera simaruba
Elaphrium tomentosum Jacq. =Bursera tomentosa
Elaphrium trinitense (R.O. Williams) Rose =Bursera karsteniana
Hedwigia hostmannii Engl. =Tetragastris hostmannii
Hedwigia panamensis Engl. =Tetragastris panamensis
Hedwigia rhoifolia Benth. =Crepidospermum rhoifolium
Hedwigia rhoifolia (Willd.) Baill. =Trattinnickia rhoifolia
Hemicrepidospermum rhoifolium (Benth.) Swart =Crepidospermum rhoifolium
Idea altissima Aubl. =Tetragastris altissima
Idea aracouchini Aubl. =Protium aracouchini
Idea decandra Aubl. =Protium decandrum
Idea enneandra Aubl. =Protium decandrum
Idea guianensis Aubl. =Protium guianense
Idea heptaphylla Aubl. =Protium heptaphyllum
Idea heterophylla Willd. [nom. illeg.] =Protium aracouchini
Idea hostmanni Miq. =Protium guianense
Idea insignis Triana & Planch. =Protium sagotianum
Idea paniculata (Engl.) Villafranca =Protium paniculatum
Idea polybotiya Turcz. =Protium polybotryum
Idea pubescens Benth. =Protium unifoliolatum
Idea sprueeana Benth. ^Protium spruceanum
Idea surinamensis Miq. =Protium heptaphyllum
Idea viridiflora Lam. [nom. illeg.] =Protium guianense
Icicopsis caudata (Turcz.) Engl. =Protium sagotianum
Icicopsis ferruginea (Engl.) Engl. =Protium ferrugineum
Icicopsis insignis (Triana & Planch.) Engl. =Protium sagotianum
Icicopsis reticulata Engl. =Protium reticulatum
Icicopsis subserrata Engl. ^Protium subserratum
Icicopsis tenuifolia Engl. =Protium tenuifolium
Pachylobus peruviana Loes. =Daeryodes peruviana
Paraprotium amazonicum Cuatrec. =Protium amazonicum
Paraprotium ferrugineum (Engl.) Cuatrec. =Protium ferrugineum
Pistacia simaruba L. =Bursera simaruba
Protium almecega Marchand ^Protium spruceanum
Protium almecega var. ulei Swart =Protium heptaphyllum subsp. ulei
Protium angustifolium Swart =Protium heptaphyllum
BO
BO DA
GU SU FG
GU SU FG
FG
GU SU FG
230 Checklist of the Plants of the Guiana Shield
Protium aracouchini var. angustifolium Swart =Protium aracouchini
Protium araguense Cuatrec. [not in flora area]
Protium benthamii Swart =Protium unifoliolatum
Protium crassifolium Engl. =Protium giganteum var. crassifolium
Protiumfimbriatum Swart =Protium amazonicum
Protiumfirmum Swart =Protium apiculatum
Protium heptaphyllum war./Ioribundum Swart =Protium heptaphyllum
Protium heptaphyllum var. multiflorum (Engl.) Swart =Protium heptaphyllum
Protium heptaphyllum var. puberulum Engl. =Protium guianense
Protium heptaphyllum var. surinamense (Miq.) Swart =Protium heptaphyllum
Protium heptaphyllum var. unifoliolatum Swart =Protium heptaphyllum
Protium hostmannii (Miq.) Engl. =Protium guianense
Protium hostmannii var. brasiliensis Swart =Protium heptaphyllum
Protium insigne (Triana & Planch.) Engl. =Protium sagotianum
Protiumjoannis Cuatrec. =Protium trifoliolatum
Protium llewelyni J.F. Macbr. =Protium grandifolium
Protium martianum Engl. =Protium trifoliolatum
Protium multiflorum Engl. =Protium heptaphyllum
Protium neglectum Swart =Protium tenuifolium
Protium nitidum Engl. =Protium paniculatum var. riedelianum
Protium octandrum Swart =Protium heptaphyllum
Protium orinocense Rusby =Protium decandrum
Protium paniculatum var. pentamerum Swart =Protium paniculatum var. paniculatum
Protium paraense auct. non Swart 1966: sensu Cuatrec. =Protium altsonii
Protium pauciflorum Swart =Protium trifoliolatum
Protium puberulenta Steyerm. =Protium altsonii
Protium pubescens (Benth.) Engl. [nom. illeg.] =Protium unifoliolatum
Protium riedelianum Engl. =Protium paniculatum var. riedelianum
Protium schomburgkianum Engl. =Protium decandrum
Protium titubans J.F. Macbr. -Protium trifoliolatum
Protium unifoliolatum var. inconforme Swart =Protium unifoliolatum
Protium unifoliolatum var. macrophyllum Ruber =Protium unifoliolatum
Protium unifoliolatum var. puberulum Hoehne =Protium unifoliolatum
Protium unifoliolatum var. subserratum Engl. =Protium unifoliolatum
Tetragastris panamensis var. grandifolia Swart =Tetragastris panamensis
Tetragastris panamensis var. hirtella Swart =Tetragastris panamensis
Tetragastris paraensis Cuatrec. =Tetragastris panamensis
Tetragastris phanerosepala Sandwith =Tetragastris altissima
Tetragastris pilosa Cuatrec. =Protium pilosum
Tetragastris stevensonii Standi. =Tetragastris panamensis
Tetragastris trifoliolata (Engl.) Cuatrec. =Protium trifoliolatum
Tetragastris unifoliolata (Engl.) Cuatrec. =Protium unifoliolatum
Tingulonga almecega (Marchand) Kuntze =Protium spruceanum
Tingulonga aracuchini (Aubl.) Kuntze =Protium aracouchini
Tingulonga caudata (Turcz.) Kuntze =Protium sagotianum
Tingulonga divaricata (Engl.) Kuntze =Protium divaricatum
Tingulonga enneandra (Aubl.) Kuntze =Protium decandrum
Tingulongaferruginea (Engl.) Kuntze =Protium ferrugineum
Tingulonga grandifolia (Engl.) Kuntze =Protium grandifolium
Tingulonga guianensis (Aubl.) Kuntze =Protium guianense
Tingulonga heptaphylla (Aubl.) Kuntze =Protium heptaphyllum
Tingulonga hostmanni (Miq.) Kuntze =Protium guianense
Tingulonga insignis (Triana & Planch.) Kuntze =Protium sagotianum
Tingulonga martiana (Engl.) Kuntze =Protium trifoliolatum
Tingulonga multiflora (Engl.) Kuntze ^Protium heptaphyllum
Tingulonga nitida (Engl.) Kuntze =Protium paniculatum var. riedelianum
Tingulonga paniculata (Engl.) Kuntze =Protium paniculatum
Tingulonga riedeliana (Engl.) Kuntze =Protium paniculatum var. riedelianum
Tingulonga simplicifolia Kuntze =Protium unifoliolatum
Tingulonga spruceana (Benth.) Kuntze =Protium spruceanum
Tingulonga subserrata (Engl.) Kuntze =Protium subserratum
Tingulonga tenuifolia (Engl.) Kuntze =Protium tenuifolium
Tingulonga trifoliolata Kuntze =Protium trifoliolatum
Trattinnickia burserifolia subsp. obtusa Engl. =Trattinnickia burserifolia
Checklist of the Plants of the Guiana Shield 231
Trattinnickia burserifolia subsp. quinquejuga Engl. =Trattinnickia bursehfolia
Trattinnickia cuspidata Cuatrec. =Dac7yodes cuspidata
Trattinnickia glabra Steyerm. =Dacryodes glabra
Trattinnickia guianensis Klotzsch [nomen nudum] =Trattinnickia burserifolia
Trattinnickia lawrancei var. boliviana Swart =Trattinnickia boliviana
Trattinnickia rhoifolia var. lancifolia Cuatrec. =Trattinnickia boliviana
Trattinnickia rhoifolia subsp. sprucei Engl. =Trattinnickia rhoifolia
Trattinnickia rhoifolia subsp. willdenowii Engl. [nom. illeg.] =Trattinnickia rhoifolia
Trattinnickia schomburgkii Klotzsch [nomen nudum] =Trattinnickia burserifolia
Trattinnickia subchoripetala Swart =Trattinnickia burserifolia
References
Daly, D. 1987. Studies in neotropical Burseraceae I. A synopsis of the genus Crepidospermum. Brittonia 39: 51-58.
Daly, D. 1989. Studies in neotropical Burseraceae II. Generic limits in New World Protieae and Canarieae. Brittonia 41: 17-27.
Daly, D. 1992. Studies in neotropical Burseraceae VI. New taxa and combinations in Protium Burm.f. Brittonia 44: 280-299.
Daly, D.C. 1997. Burseraceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 688-728.
Daly, D.C. 2002. Burseraceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 151-
165.
Swart, J.J. 1942. A monograph of the genus Protium and some allied genera. Rec. Trav. Bot. Neerl. 34: 211-446.
Swart, J.J. 1951. Burseraceae. In A. Pulle, Flora of Suriname 3(2): 204-251.
BUXACEAE
K. Yatskievych
Buxus citrifolia (Willd.) Spreng. BO
Synonyms and Misidentifications
Tricera citrifolia Willd. =Buxus citrifolia
References
Yatskievych, K. 1997. Buxaceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 729-730.
Cabombaceae
J.H. Wiersema
Brasenia schreberi J.F. Gmel. [possibly extirpated from GU]
Cabomba aquatica Aubl. BO DA
Cabombafurcata Schuh. & Schult. f. AM BO
Cabomba haynesii Wiersema [expected in GU] BO
Synonyms and Misidentifications
Cabomba piauhyensis Gardner =Cabombafurcata
Cabomba piauhyensis var. albida Fassett =Cabomba haynesii
Cabomba piauhyensis vox. piauhyensis =Cabombafurcata
Cabomba warmingii Casp. =Cabombafurcata
References
Cramer, J. 1979. Cabombaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 385-389.
Fassett, N.C. 1953. A monograph of Cabomba. Castanea 18: 116-128.
Oergaard, M., H.W.E. van Bruggen, & P.J. van der Vlugt. 1992. Die Familie Cabombaceae (Cabomba und Brasenia). Aqua-
Planta sonderheft nr. 3.
Wiersema, J.H. 1 989. A new species of Cabomba from tropical America. Ann. Missouri Bot. Gard. 76(4): 1 1 67- 1168.
Wiersema, J.H. 1997. Cabombaceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 730-732.
Acanthocereus tetragonus (L.) Hummelinck
Cereus hexagonus (L.) Mill.
Cereus repandus (L.) Mill., non Haw. 1812
Disocactus amazonicus (K. Schum.) D.R. Hu
Guayana]
Epiphyllum phyllanthus (L.) Haw.
Hylocereus lemairei (Hook.) Britton & Rose
GU?
GU su FG
GU su FG
Cactaceae
.E. Leuenberger, B. Trujillo
BO
AM BO
BO
GU SU FG
>cpected in Venezuelan
AM BO DA GU SU FG
BO DA GU? SU? FG?
232 Checklist of the Plants of the Guiana Shield
Hylocereiis scandens (Salm-Dyck) Backeb. GU SU
Melocactus matanzamis Leon [cultivated] AM
Melocactus mazeliamis Riha AM BO
Melocactus neryi K. Schum. AM BO GU SU
Melocactus smithii (Alexander) Buining ex G.D. Rowley GU
Opuntia cochenillifera (L.) Mill, [cultivated only?] GU SU FG
Opuntia elatior Mill. BO
Pereskia aculeata Mill, [widely cultivated but apparently native in Guianas] BO GU SU FG
Pereskia guamacho F.A.C. Weber BO
Pilosocereus oligolepis (Vaupel) Byles & G.D. Rowley [described as having 5 BO GU SU FG
ribs]
i?/2z>5a/w Z?acc//era (J.S. Muell.) Steam AM BO DA GU SU FG
Rhipsalis floccosa Pfeiff. SU
Rhipsalis oblonga Loefgr. [possibly confused with R. occidentalis] SU?
Rhipsalis occidentalis Barthlott & Rauh [possibly confused with R. oblonga] SU?
Rhipsalis pachyptera Pfeiff. GU? FG?
Selenicereus extensus (Salm-Dyck ex DC.') Leuenb. FG?
Selenicereus megalantlnis (K. Schum. ex Vaupel) Moran [known for AM only AM
by vegetative plant (#4528) at Jardin Botanico Universitario de Maracay]
Selenicereus sp. [sect. ^a/m^vcAr/a] [expected in AM & BO] GU? SU? FG
Strophocactus wittii (K. Schum.) Britton & Rose AM
Synonyms and Misidentifications
Acanthocereus colombiamis Britton & Rose =Acanthocereus tetragonus
Acanthocereus pentagonus (L.) Britton & Rose =Acanthocereus tetragonus
Acanthocereus pitajaya (Jacq.) Dugand ex Croizat =Acanthocereus tetragonus
Cactus caripensis Kunth =Rhipsalis baccifera
Cactus hexagonus L. =Cereus hexagonus
Cactus neryi (K. Schum.) Britton & Rose =Melocactus neryi
Cactus pentagonus L. =Acanthocereus tetragonus
Cactus pereskia L. =Pereskia aculeata
Cactus phyllanthus L. =Epiphyllum phyllanthus
Cactus pitajaya Jacq. =Acanthocereus tetragonus
Cactus repandus L. ^Cereus repandus
Cactus smithii Alexander =Melocactus smithii
Cactus tetragonus L. =Acanthocereus tetragonus
Cephalocereus kanukuensis Alexander =Pilosocereus oligolepis
Cereus atroviridis Backeb. =Cereus repandus
Cereus caripensis (Kunth) DC. =Rhipsalis baccifera
Cereus curtisii Otto [Guianan records dubious]
Cereus grandiflorus Mill. [Guianan records dubious]
Cereus grenadensis Britton & Rose =Cereus repandus
Cereusjamacaru DC. [Guianan records dubious]
Cereus lemairei Hook. =Hylocereus lemairei
Cereus lepidotus Salm-Dyck =Cereus hexagonus
Cereus longiflorus Alexander =Cereus hexagonus
Cereus margaritensis J.R. Johnst. =Cereus repandus
Cereus megalanthus K. Schum. ex Vaupel =Selenicereus megalanthus
Cereus peruvianus auct. non (L.) Mill. [Guianan records dubious] =Cereus uruguayanus
Cereus pitajaya (Jacq.) DC. =Acanthocereus tetragonus
Cereus polyrhizus F.A.C. Weber =Hylocereus lemairei
Cereus triangularis (L.) Haw. [Guianan records dubious] =Hylocereus triangularis
Cereus trinitatensis Lem. & Herment =Hylocereus lemairei
Cereus wittii K. Schum. =Strophocactus wittii
Epiphyllum hookeri Haw. =Epiphyllum phyllanthus
Epiphyllum phyllanthus var. hookeri (Haw.) Kimnach =Epiphyllum phyllanthus
Hylocereus polyrhizus (F.A.C. Weber) Britton & Rose =Hylocereus lemairei
Hylocereus triangularis (L.) Britton & Rose [Guianan records dubious]
Hylocereus trinitatensis (Lem. & Herment) A. Berger =Hylocereus lemairei
Hylocereus venezuelensis Britton & Rose =Hylocereus lemairei
Mediocactus megalanthus (K. Schum. ex Vaupel) Britton & Rose =Selenicereus megalanthus
Melocactus guaricensis Croizat =Melocactus neryi
Melocactus schulzianus Buining & Brederoo =Melocactus neryi
Nopalea cochenillifera (L.) Salm-Dyck =Opuntia cochenillifera
Pereskia colombiana Britton & Rose =Pereskia guamacho
Checklist of the Plants of the Guiana Shield 233
Phyllocactus phyllanthits (DC.) Link =Epiphyllum phyllanthiis
Pilocereus repandus (L.) K. Schum. =Cereus repandus
Pilosocereiis kamikiiensis (Alexander) Leuenb. [described as having 4 ribs, illustrations in FVG contradictory] =Pilosocerens
oligolepis
Pilosocereus oligolepis subsp. kamikiiensis (Alexander) P.J. Braun & Esteves =Pilosocereus oligolepis
Rhipsalis cassutha Gaertn. =Rhipsalis baccifera
Rhipsalis minutiflora K. Schum. =Rhipsalis baccifera
Rhodocactus colombianus (Britton & Rose) P.M. Knuth =Pereskia guamacho
Rhodocactus guamacho (F.A.C. Weber) P.M. Knuth =Pereskia guamacho
Selenicereus nittii (K. Schum.) G.D. Rowley =Strophocactus wittii
Subpilocereus grenadensis (Britton & Rose) Backeb. =Cereus repandus
Subpilocereus margaritensis (J.R. Johnst.) Backeb. =Cereus repandus
Subpilocereus repandus (L.) Backeb. =Cereus repandus
Wilmattea venezuelensis (Britton & Rose) Croizat ^Hylocereus lemairei
Wittiocactus amazonicus (K. Schum.) Rauschert =Disocactus amazonicus
References
Hunt, D. 1992. CITES Cactaceae Checklist. Royal Botanic Gardens, Kew.
Leuenberger, B.E. 1987. A preliminary- list of Cactaceae from the Guianas and recommendations for future collecting and
preparation of specimens. Willdenowia 16: 497-510.
Leuenberger, B.E. 1989. Lectot>'pification of Cereus hexagonus (L.) Miller and taxonomic notes on related taxa from South
America. Bot. Jahrb. Syst. 111(2): 145-164.
Leuenberger, B.E. 1997. Cactaceae. In A.R.A. Gorts-van Rijn, Plora of the Guianas (series A) 31: 1-63.
Leuenberger, B.E. 2002. Cactaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
165-166.
Taylor, N.P. 1991. The genus Melocactus in Central and South America. Bradleya 9: 1-80.
Trujillo, B. 1997. Cactaceae. In J.A. Steyermark, RE. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 732-749.
Campanulaceae
T. Lammers, J.L. Luteyn, B.A. Stein
Burmeistera ramosa E. Wimm. AM
Centropogon cornutus (L.) Druce AM BO DA GU SU PG
Centropogon roraimanus E. Wimm. BO GU
Hippobroma longiflora (L.) G. Don [expected in FG] AM BO DA GU SU
Lobelia aquatica Cham. AM BO GU SU PG
Lobelia fastigiata Kunth BO
Siphocampylus reticulatus (Willd. ex Roem. & Schult.) Vatke AM BO
Siphocampylus sp. A AM BO
Siphocampylus tillettii Steyerm. AM
Wahlenbergia perrottetii (A. DC.) Thulin [native to Africa and Brazil]
Synonyms .^nd Misidentifications
Centropogon bonplandianus (Willd. ex Roem. & Schult.) C. Presl =Centropogon cornutus
Centropogon solanifolius Benth. [records based on misidentifications fide B. Stein]
Centropogon surinamensis (L.) C. Presl =Centropogon cornutus
Cephalostigma bahiense A. DC. =Wahlenbergia perrottetii
Isotoma longiflora (L.) C. Presl =Hippobroma longiflora
Laurentia longiflora (L.) Endl. =Hippobroma longiflora
Lobelia bonplandiana Willd. ex Roem. & Schult. =Centropogon cornutus
Lobelia cornuta L. =Centropogon cornutus
Lobelia longiflora L. =Hippobroma longiflora
Lobelia reticulata Willd. ex Roem. & Schult. =Siphocampylus reticulatus
Lobelia surinamensis L. =Centropogon cornutus
Siphocampylus tepuiensis Gleason =Siphocampylus reticulatus
References (incl. Lobeliaceae. See also Sphenocleaceae)
Gracie, C.A. 2002. Campanulaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. HofF, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
183-184.
Lanjouw, J. 1934. Campanulaceae. In A. Pulle, Flora of Suriname 4(1): 302-305.
Lanjouw, J. 1937. Campanulaceae (additions and corrections). In A. Pulle, Plora of Suriname 4(1 ): 494-495.
Stein, B.A. 1998. Campanulaceae. In J.A. Steyermark, P.P. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 122-129.
FG
234 Checklist of the Plants of the Guiana Shield
Canellaceae
Cinnamodendron temtifolium Uittien SU FG
References
Brown, J.L. & S.A. Mori. 2002. Canellaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76(2): 184-186.
Uittien, H. 1935. Canellaceae. In A. PuUe, Flora of Suriname 3(1): 304-305.
Wilson, T.K. 1960. The comparative morphology ofthe Canellaceae. I. Synopsis of genera and wood anatomy. Trop. Woods 112:
1-27.
Capparaceae
H.H. litis, J.C. Lindeman, T. Ruiz-Zapata
BO
BO
BO
BO
BO
DA GU? SU
BO
BO
BO
BO
BO
BO
GU
GU
GU
DA
Cappahs amplissima Lam.
Capparis detonsa Triana & Planch.
Capparis flexuosa (L.) L.
Capparis flexuosa subsp. polyantha (Triana & Planch.) H.H. litis
Capparis frondosa Jacq. [=Capparis baducca L.?] AM
Capparis grandipetala Maguire & Steyerm. [Serrania de la Neblina , expected
in AM]
Capparis guaguaensis Steyerm. [expected in GU]
Capparis hastata Jacq.
Capparis hastata f coccolobifolia (Mart, ex Eichler) H.H. litis
Capparis leprieurii Briq.
Capparis maroniensis Benoist
Capparis muco H.H. litis, L. Cumana, R. Delgado & Aymard
Capparis odoratissima Jacq.
Capparis osmantha Diels
Capparis pulcherrima Jacq.
Capparis sola J.F. Macbr. AM
Capparis sp. A AM
Capparis verrucosa Jacq.
Capparis yco (Mart.) Mart. & Eichler
Capparis yco var. guianensis Sagot
Cleome acideata L. AM
Cleomefriitescens Aubl. [=Cleome spinosa"?]
Cleome guianensis Aubl.
Cleome gynandra L. [naturalized]
Cleome latifolia Vahl ex DC.
Cleome parviflora Kunth
Cleome pilosa Benih. AM
Cleome rutidosperma DC. [naturalized]
Cleome serrata Jacq.
Cleome speciosa Raf.
Cleome spinosa Jacq. AM
Cleome stenophylla Klotzsch ex Urb.
Cleome viridiflora Schreb. AM
Crateva tapia L. AM
Morisonia americana L.
Steriphoma ellipticum (DC.) Spreng.
Synonyms and Misidentifications
Capparidastrum baducca (L.) Hutch. =Capparis frondosa
Capparis baducca L. [Capparisfrondosa Jacq. synonymized here by some authors] =Capparisfrondosa
Capparis cynophallophora L. =Capparis flexuosa
Capparis cynophallophora var. isthmensis (Eichler) H.H. litis ex Jans.-Jac. [nomen nudum] =Capparisflexuosa
Capparis flexuosa f. hastata (Jacq.) Dugand =Capparis hastata
Capparis guaguaensis Steyerm. =Capparis osmantha
Capparis isthmensis Eichler =Capparis flexuosa
Capparis macrophylla auct. non Kunth 1 82 1 . ^Capparis osmantha
Capparis montana (Aubl.) Lemee (Sapotaceae) [fide Lindeman] =Pradosia ptychandra
SU
SU
SU
SU
SU
FG
FG
FG
FG
FG?
BO SU FG
BO GU?
FG
FG
GU SU FG
FG
BO GU SU FG
BO SU FG
BO GU SU FG
BO DA GU
GU
SU
SU
FG
FG
GU SU FG
BO GU
GU
SU FG
FG
BO DA GU
GU
SU FG
BO
Checklist of the Plants of the Guiana Shield 235
Capparis nitida Ruiz & Pav. ex DC. =Capparis amplissima
Cappahs pendula Triana & Planch. =Capparis amplissima
Capparis perseifolia litis ex Jans.-Jac. [nomen nudum, fide Lindeman] =Capparis leprieurii
Capparis polyantha Triana & Planch. =Capparis flexuosa subsp. polyantha
Capparis sola var. longiracemosa Dugand =Capparis sola
Capparis surinamensis J.C. Went =Capparis maroniensis
Capparis trinitensis Urb. =Capparis verrucosa
Cleome amazonica Heilb. =Cleome viridiflora
Cleome brachypoda Briq. ^Cleome parviflora
Cleomefriesii Heilb. =Cleome viridiflora
Cleome gigantea L. =Cleome viridiflora
Cleome hostmanni Miq. =Cleome latifolia
Cleome hiimboldtii DC. =Cleome parviflora
Cleome kerberi Briq. ^Cleome viridiflora
Cleome micrantha Desv. ex Ham. [name of uncertain application] =Cleome parviflora
Cleome monochroma J.F. Macbr. =Cleome viridiflora
Cleome pentaphylla L. =Cleome gynandra
Cleome pittieri Briq. =Cleome pilosa
Cleome polygama L. =Cleome serrata
Cleome psoraleaefolia DC. =Cleome parviflora
Cleome pungens Willd. [nom. confus.] =Cleome spinosa
Cleome surinamensis Miq. =Cleome aculeata
Crateva acuminata DC. =Crateva tapia
Crateva benthamii Eichler =Crateva tapia
Crateva gynandra L. =Crateva tapia
Gynandropsis gynandra (L.) Briq. =Cleome gynandra
Gynandropsis pentaphylla (L.) DC. =Cleome gynandra
Gynandropsis speciosa (Kunth) DC. =Cleome speciosa
Melidiscus gigantea (L.) Raf. =Cleome viridiflora
Morisonia elliptica Rusby =Morisonia americana
Morisoniaflexuosa L. =Capparis flexuosa
Morisoniajohnstonii Urb. =Morisonia americana
Neocalyptrocalyx surinamensis (J.C. Went) Hutch. =Capparis maroniensis
Physostemon ambiguum Bong. =Cleome guianensis
Physostemon guianense (Aubl.) Malme =Cleome guianensis
Physostemon intermedium Moric. "^Cleome guianensis
Physostemon lanceolatum Mart. & Zucc. [fide Lindeman] =Cleome stenophylla
Quadrella cynophallophora (L.) Hutch. ^Capparis flexuosa
Quadrella odoratissima (Jacq.) Hutch. =Capparis odoratissima
Sinapistrum giganteum (L.) Moench. ^Cleome viridiflora
Tarenaya spinosa (Jacq.) Raf =Cleome spinosa
Voyara montana Aubl. (Sapotaceae) ^Pradosia ptychandra
References
litis, H.H. & S.A. Mori. 2002. Capparaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76(2): 186-188.
Jansen-Jacobs, M.J. 1976. Capparaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 512-
517.
Ruiz-Zapata, T. & H.H. litis. Capparaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4:
132-157.
Went, J.C. 1 937. Capparidaceae. In A. Pulle, Flora of Suriname 2( 1 ): 397-405.
Caprifoliaceae
RE. Berry, N. Ramirez
Sambucus canadensis L.
Sambucus canadensis var. laciniata A. Gray
Sambucus simpsonii Rehder [introduced] SU
Viburnum tinoides L. f
Viburnum tinoides var. roraimense (Killip & A.C. Sm.) Steyerm.
Synonyms and Misidentifications
Oreinotinus tinoides (L. f ) Oerst. =Viburnum tinoides
DA
DA
BO GU
BO GU
GU SU FG
GU SU FG
FG
FG
236 Checklist of the Plants of the Guiana Shield
Viburnum glabratum Kunth [not in flora area]
Viburnum leptophyllum Killip & A.C. Sm. ^Viburnum tinoides
Viburnum roraimense Killip & A.C. Sm. =Viburnum tinoides van roraimense
Viburnum tinoides var. venezuelense (Killip & A.C. Sm.) Steyerm. [not in flora area - Andes of Venezuela] =Viburnum
tinoides van roraimense
References
Killip, E.P. & A.C. Smith. 1929. The genus Viburnum in northwestern South America. Bull. Torrey Bot. Club 56: 265-274.
Ramirez, N. & RE. Berry. 1998. Caprifoliaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana
4: 158-161.
Steyermark, J.A. 1953. Caprifoliaceae. In J.A. Steyermark & collaborators, Contributions to the Flora of Venezuela. Fieldiana
Bot. 28(3): 618-619.
Caricaceae
J.S. Miller
Car/ca/7<3pflya L. [cultivated, naturalized] AM BO
Jacaratia spinosa (Aubl.) A. DC.
Vasconcellosia microcarpa (Jacq.) A. DC.
Vasconcellosia microcarpa subsp. heterophylla (Poepp. & Endl.) V.M. Badillo
Synonyms and Misidentifications
Carica microcarpa subsp. heterophylla (Poepp. & Endl.) V.M. Badillo =Vasconcellosia microcarpa subsp. heterophylla
Carica sativa Tussac -Carica papaya
Carica spinosa Aubl. =Jacaratia spinosa
Papaya spinosa A. DC. =Jacaratia spinosa
References
Badillo, V.M. 1971. Monografia de la familia Caricaceae. Publ. Asoc. Prof., Univ. Central de Venezuela. Maracay.
Lindeman, J.C. 1984. Caricaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 585-591.
Miller, J.S. 1998. Caricaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 162-163.
Mori, S.A. 2002. Caricaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 188-
190.
Anthodiscus mazarunensis Gilly
Anthodiscus obovatus Benth. ex Wittm.
Anthodiscus trifoliatus G. Mey.
Caryocar glabrum (Aubl.) Pers.
Caryocar glabrum subsp. album Prance & M.F. Silva
Caryocar glabrum subsp. glabrum
Caryocar gracile Wittm.
Caryocar microcarpum Ducke
Caryocar montanum Prance
Caryocar nuciferum L.
Caryocar pallidum A.C. Sm.
Caryocar villosum (Aubl.) Pers.
Synonyms and Misidentifications
Caryocar butyrosum (Aubl.) Willd. =Caryocar villosum
Caryocar coccineum Pilg. =Caryocar glabrum
Caryocar glabrum var. edule Wittm. =Caryocar microcarpum
Caryocar glabrum var. pilosum Wittm. =Caryocar microcarpum
Caryocar krukovii Gilly =Caryocar gracile
Caryocar riparium A.C. Sm. =Caryocar microcarpum
Caryocar tessmannii Pilg. =Caryocar glabrum
Caryocar tomentosum Willd. [nom. illeg.] =Caryocar nuciferum
Caryocar toxiferum Barb. Rodr. =Caryocar glabrum
Pekea butyrosa Aubl. =Caryocar villosum
Pekea tuberculosa Aubl. =Caryocar nuciferum
Pekea villosa (Aubl.) Poir. =Caryocar villosum
Rhizobolus butyrosus (Aubl.) J.F. Gmel. =Caryocar villosum
Caryocaraceae
GT. Prance
AM BO GU SU
AM BO
GU
AM DA GU
GU
GU
SU
SU
FG
FG
AM
AM BO
BO
DA GU
GU
SU FG
BO DA GU SU FG
AM BO GU FG?
AM FG
Checklist of the Plants of the Guiana Shield 237
AM BO GU SU FG
AM BO GU SU FG
AM BO DA GU SU
AM BO DA GU SU
AM BO DA
Rhizobolus pekea Gaertn. =Caryocar nuciferum
Rhizobolus tuberculosus (Aubl.) J.F. Gmel. =Caryocar nuciferum
Saouari glabra Aubl. =Caryocar glabrum
Saouari villosa Aubl. =Caryocar villosum
References
Granville, J.J. de. 2002. Caryocaraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76(2): 191-192.
Lanjouw, J. & P.F. van Heerdt. 1941. Caryocaraceae. In A. Pulle, Flora of Suriname 3(1): 366-372.
Prance, GT. 1972. Caryocaraceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 9. Mem. New York Bot.
Gard. 23: 1-832.
Prance, GT. 1998. Caryocaraceae. In J.A. Steyermark, P.E. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 164-170.
Prance, GT. &. M.F. da Silva. 1973. Caryocaraceae. Flora Neotropica Monogr. 12: 1-75.
Caryophyllaceae
G.A. Aymard C, N.L. Cuello A., R.A. DeFilippsf
Drymaria cordata (L.) Willd. ex Schult.
Drymaria cordata subsp. cordata
Polycarpaea corymbosa (L.) Lam.
Polycarpaea corymbosa var. brasiliensis (Cambess.) Chodat & Hassl.
Polycarpon apurense Kunth
Synonyms and Misidentifications
Holosteum cordatum L. =Drymana cordata
Polycarpaea atherophora Steud. =Polycarpaea corymbosa
Polycarpaea brasiliensis Cambess. =Polycarpaea corymbosa var. brasiliensis
References
Aymard C, G.A. & N.L. Cuello A. 1998. Caryophyllaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan
Guayana 4: 171-174.
DeFilipps, R.A. & S.L. Maina. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76(2): 193.
Ooststroom, S.J. van. 1934. Caryophyllaceae. In A. Pulle, Flora of Suriname 1(1): 150-153.
Casuarinaceae
Casuarina equisetifolia J.R. Forst. & G Forst. [Old World family, one species
widely cultivated]
References
Berg, C.C. 1992. Casuarinaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 11: 140.
Cecropiaceae
C.C. Berg
Cecropia angulata I.W. Bailey
Cecropia distachya Huber
Cecropia engleriana Snethl.
Cecropia ficifolia Warb. ex Snethl.
Cecropia granvilleana C.C. Berg
Cecropia kavanayensis Cuatrec.
Cecropia latiloba Miq.
Cecropia membranacea Trecul
Cecropia metens is Cuatrec.
Cecropia obtusa Trecul
Cecropia palmata Willd.
Cecropia peltata L.
Cecropia sciadophylla Mart.
Cecropia silvae C.C. Berg
Coussapoa angustifolia Aubl.
BO DA GU
AM BO FG
AM
AM
SU? FG
AM BO GU?
AM BO DA GU SU FG
AM
AM BO GU
GU SU
SU
FG
FG
AM BO DA GU SU FG
AM BO DA GU SU
SU
FG
FG?
GU SU FG
238 Checklist of the Plants of the Guiana Shield
Coussapoa argew/'ea Akkermans & C.C. Berg
Coussapoa asperifoUa Trecul AM
Coussapoa asperifoUa subsp. asperifoUa
Coussapoa asperifoUa subsp. magnifoUa (Trecul) Akkermans «fe C.C. Berg AM
Coussapoa asperifoUa subsp. rhamnoides (Standi.) Akkermans & C.C. Berg
Coussapoa crassivenosa Mildbr. AM
Coussapoaferruginea Trecul
Coussapoa latifoUa Aubl.
Coussapoa leprieurii Benoist
Coussapoa microcephala Trecul
Coussapoa orthoneura Standi. AM
Coussapoa parvifoUa Standi. AM
Coussapoa trinervia Spruce ex Mildbr. AM
Coussapoa viUosa Poepp. & Endl. AM
Coussapoa viridifoUa Cuatrec. AM
Pourouma acuminata Miq.
Pourouma bicolor Mart. ' AM
Pourouma bicolor subsp. bicolor AM
Pourouma bicolor subsp. digitata (Trecul) C.C. Berg & Heusden AM
Pourouma bolivarensis C.C. Berg
Pourouma cecropiifolia Mart. AM
Pourouma cucura Standi. & Cuatrec. AM
Pourouma guianensis Aub\. AM
Pourouma guianensis subsp. guianensis AM
Pourouma melinonii Benoist AM
Pourouma melinonii subsp. melinonii AM
Pourouma minor Benoist AM
Pourouma mollis Trecul
Pourouma mollis subsp. mollis
Pourouma ovata Trecul
Pourouma saulensis C.C. Berg & Kooy
Pourouma stipulacea C.C. Berg
Pourouma tomentosa Mart, ex Miq. AM
Pourouma tomentosa subsp. apiculata (Spruce ex Benoist) C.C. Berg & AM
Heusden
Pourouma tomentosa subsp. essequiboensis (Standi.) C.C. Berg & Heusden
Pourouma tomentosa subsp. maroniensis (Benoist) C.C. Berg & Heusden
Pourouma tomentosa subsp. tomentosa AM
Pourouma velutina Mart, ex Miq. AM
Pourouma villosa Trecul
BO GU?
BO DA GU SU FG
GU su FG
BO DA GU
BO
BO GU?
SU FG
GU su FG
FG
GU su FG
GU
GU
BO GU
BO
BO GU su FG
BO GU su FG
GU su FG
BO GU?
BO
BO GU su
BO DA GU su FG
BO DA GU su FG
BO GU su FG
BO GU su FG
BO GU su FG
BO GU su FG
BO GU su FG
BO
GU
FG
BO GU su FG
BO
BO
GU
GU
SU
su
su
FG
FG
FG
Synonyms and Misidentifications
Cecropia auyantepuiana Cuatrec. =Cecropia kavanayensis
Cecropia bureauiana V.A. Richt. =Cecropia palmata
Cecropia concolor subsp. engleriana (Snethl.) C.C. Berg & P. Franco =Cecropia engleriana
Cecropia dielsiana Snethl. =Cecropia peltata
Cecropiajuranyiana V.A. Richt. =Cecropia sciadophylla
Cecropia orinocensis Standi. =Cecropia latiloba
Cecropia paraensis Huber =Cecropia latiloba
Cecropia peltata var. Candida Velasquez =Cecropia metensis
Cecropia riparia Snethl. =Cecropia distachya
Cecropia sciadophylla var. decurrens Snethl. =Cecropia sciadophylla
Cecropia sciadophylla war. juranyiana (V.A. Richt.) Snethl. =Cecropia sciadophylla
Cecropia steyermarkii Cuatrec. =Cecropia kavanayensis
Cecropia surinamensis Miq. =Cecropia peltata
Cecropia tessmannii Mildbr. =Cecropia membranacea
Coussapoa cardonae Pittier =Coussapoa asperifoUa subsp. magnifoUa
Coussapoa cayennensis Hawkes =Coussapoa asperifoUa subsp. asperifoUa
Coussapoa cornifolia Standi. =Coussapoa parvifoUa
Coussapoa cuneata Miq. =Coussapoa microcephala
Coussapoa fagifolia Klotzsch =Coussapoa microcephala
Coussapoa hypochlora Standi. =Coussapoa asperifoUa subsp. magnifoUa
Coussapoa krukovii Standi. =Pourouma tomentosa subsp. apiculata
Coussapoa latifoUa var. obovata (Miq.) Miq. =Coussapoa latifoUa
Checklist of the Plants of the Guiana Shield 239
Coussapoa magnifolia Trecul =Coussapoa asperifolia subsp. magnifolia
Coussapoa microcephala subsp. cornifolia (Standi.) Akkermans & C.C. Berg =Coussapoa pai-vifolia
Coussapoa obovata Miq. =Coussapoa latifolia
Coussapoa prancei C.C. Berg =Coussapoa orthoneura
Coussapoa rhamnoides Standi. =Coussapoa asperifolia subsp. rhamnoides
Coussapoa steyermarkii Cuatrec. =Coussapoa crassivenosa
Coussapoa viridifolia van tenuifolia Cuatrec. =Coussapoa viridifolia
Coussapoa williamsii Cuatrec. =Coussapoa orthoneura
Pourouma acutiflora Trecul =Pourouma guianensis subsp. guianensis
Pourouma albistipulata Cuatrec. =Pourouma tomentosa subsp. tomentosa
Pourounia apaporiensis Cuatrec. =Pourouma melinonii subsp. melinonii
Pourouma apiculata Spruce ex Mildbr. =Pourouma tomentosa subsp. apiculata
Pourouma apiculata Spruce ex Benoist =Pourouma tomentosa subsp. apiculata
Pourouma aspera Trecul =Pourouma bicolor subsp. bicolor
Pourouma aurea Mildbr. =Pourouma minor
Pourouma camaratana Cuatrec. =Pourouma bicolor subsp. bicolor
Pourouma cuatrecasasii Standi. =Pourouma minor
Pourouma digitata Trecul =Pourouma bicolor subsp. digitata
Pourouma essequiboensis Standi. =Pourouma tomentosa subsp. essequiboensis
Pourouma garciana Cuatrec. =Pourouma cucura
Pourouma laevis Benoist =Pourouma villosa
Pourouma longipendula Ducke =Pourouma ovata
Pourouma maroniensis Benoist =Pourouma tomentosa subsp. maroniensis
Pourouma palmata Poepp. & Endl. =Pourouma guianensis subsp. guianensis
Pourouma populifolia Standi. =Pourouma acuminata
Pourouma radula Benoist =Pourouma guianensis subsp. guianensis
Pourouma steyermarkii Standi. & Cuatrec. =Pourouma velutina
Pourouma umbellifera W.C. Burger =Pourouma minor
References
Berg, C.C. 1992. Cecropiaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 11: 93-124.
Berg, C.C. 1998. Cecropiaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 174-190.
Berg, C.C. 2002. Cecropiaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
194-198.
Berg, C.C, R.W.A.P. Akkermans, & E.C.H. van Heusden. 1990. Cecropiaceae: Coussapoa and Pourouma. Flora Neotropica
Monogr. 51: 1-194.
Berg, C.C. & GR DeWolf. 1975. Moraceae. In J. Lanjouw & A.L. Stoffers, Flora of Suriname 5(1): 265-294.
Celastraceae
D.M. Keams
Elaeodendron xylocarpum (Vent.) DC. [possibly expected in GU]
Goupia glabra A\xh\. AM BO DA GU SU FG
Maytenus apiculata Steyerm.
Maytenus ficiformis Reissek AM BO GU
Maytenusfloribunda Reissek FG
Maytenus guyanensisYAoXzichQxKQissok AM BO DA GU SU FG
Maytenus huberi Steyerm.
Maytenus insculpta Steyerm.
Maytenus kanukuensis A.C. Sm. BO GU
Maytenus laevis Reissek
Maytenus longistipitata Steyerm.
Maytenus macrocarpa (Ruiz & Pav.) Briq.
Maytenus myrsinoides Reissek AM BO DA GU SU FG
Maytenus neblinae Steyerm. [Sierra de la Neblina]
Maytenus oblongata Reissek GU SU FG
Maytenus pittieriana Steyerm. BO
Maytenus planifolia A.C. Sm. GU
Maytenus pruinosa Reissek AM BO GU SU FG
Maytenus pustulata Steyerm. AM
Maytenus sp. A AM
Maytenus tetragona Griseb. FG
Zinowiewia aymardii Steyerm. BO DA
DA
BO
AM
AM
AM BO
BO
BO
AM
240 Checklist of the Plants of the Guiana Shield
Synonyms and Misidentifications
Cassine xylocarpa Vent. ^Elaeodendron xylocarpum
Celastrus macrocarpos Ruiz & Pav. =Maytenus macrocarpa
Celastrus myrtifolius L. (Rosaceae) =Prunus myrtifolia
Gonpia paraensis Huber =Goupia glabra
Goiipia tomentosa Aubl. =Goupia glabra
Maytenusfloribunda Pittier, non Reissek 1861 =Maytenus pittieriana
Maytenus guianensis Klotzsch [nomen nudum] -Maytenus guyanensis
Maytenus guianensis f. crenulata Steyerm. =Maytenus guyanensis
Maytenusjauaens is Steyerm. =Maytenus laevis
Maytenus reissekii Urb. =Maytenus myrsinoides
References
Keams, D.M. 1998. Celastraceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 190-197.
Loesener, T. 1942. Celastraceae. In A. Engler & K. Prantl, Die Naturlichen Pflanzenfamilien 20b: 87-197.
Mitchell, J.D. 2002. Celastraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, & J.D. Mitchell
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2)
199-202.
Ceratophyllaceae
RE. Berry
Ceratophyllum muricatum Cham. DA GU SU
Ceratophyllum muricatum subsp. australe (Griseb.) Les DA
Synonyms and Misidentifications
Ceratophyllum demersum var. cristatum (Spruce) K. Schum. =Ceratophyllum muricatum
Ceratophyllum lleranae Fassett ^Ceratophyllum muricatum
References
Les, D.H. 1 988. The evolution ofachene morphology in Ceratophyllum, II. Fruit variation and systematics ofthe "spiny-margined'
group. Syst. Bot. 13(1): 73-86.
Les, D.H. 1989. The evolution ofachene morphology in Ceratophyllum, IV. Summary ofproposed relationships and evolutional
trends. Syst. Bot. 14(2): 254-262.
Lowden, R.M. 1978. Studies on the submerged genus Ceratophyllum in the Neotropics. Aquatic Botany 4: 127-142.
Wilmot-Dear, 1985. Ceratophyllum revised, a study in leaf and fruit variation. Kew Bull. 40: 243-271.
Chenopodiaceae
R.A. DeFilippst, D.M. Keams
Chenopodium ambrosioides L. [naturalized] AM BO GU SU FG
References
Keams, D.M. 1998. Chenopodiaceae. In J.A. Steyermark, P.E. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 198.
Lanjouw, J. 1957. Chenopodiaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 291-292.
Chloranthaceae
C.A. Todzia
Hedyosmum intermedium Todzia AM BO GU?
Hedyosmum neblinae Todzia AM
Hedyosmum tepuiense Todzia AM BO GU
Synonyms and Misidentifications
Hedyosmum brasiliense Mart, [not in flora area]
Hedyosmum racemosum auct., non (Ruiz & Pav.) G. Don =Hedyosmum tepuiense
References
Todzia, C.A. 1988. Chloranthaceae: Hedyosmum. Flora Neotropica Monogr. 48: 1-138.
Todzia, C.A. 1993. New species of Hedyosmum from northern South America. Novon 3: 81-85.
Todzia, C.A. 1988. Chloranthaceae. In J.A. Steyermark, P.E. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 198-201
Checklist of the Plants of the Guiana Shield 241
Chrysobalanaceae
G.T. Prance
Acioa guianensis Aubl.
Acioa schultesii Maguire
Acioa somnolens Maguire
Chrysobalanus icaco L.
Chrysobalanus venezuelanus Prance
Couepia bracteosa Benth.
Couepia canescens (Gleason) Prance
Couepia canomemis (Mart.) Benth. ex Hook. f.
Couepia caryophylloides Benoist
Couepia chrysocalyx (Poepp.) Benth. ex Hook. f.
Couepia cognata (Steud.) Fritsch
Couepia cognata var. cognata
Couepia cognata var. major Prance
Couepia cognata var. membranacea Prance
Couepia comosa Benth.
Couepia elata Ducke
Couepia excelsa Ducke
i
Couepia exflexa Fanshawe & Maguire
Couepiafoveolata Prance
Couepia guianensis Aubl.
Couepia guianensis subsp. divaricata (Huber) Prance
Couepia guianensis subsp. glandulosa (Miq.) Prance
Couepia guianensis subsp. guianensis
Couepia habrantha Standi.
Couepiajoaquinae Prance
Couepia magnoliifolia Benth. ex Hook. f.
Couepia maguirei Prance
Couepia martinii Prance
Couepia multiflora Benth.
Couepia obovata Ducke
Couepia paraensis (Mart. & Zucc.) Benth.
Couepia paraensis subsp. glaucescens (Spruce ex Hook, f.) Prance
Couepia parillo DC.
Couepia racemosa Benth. ex Hook. f.
Couepia rankinae Prance
Couepia sandwithii Prance
Couepia steyermarkii Maguire
Exellodendron barbatum (Ducke) Prance
Exellodendron coriaceum (Benth.) Prance
Hirtella adderleyi Prance
Hirtella angustissima Sandwith
Hirtella araguariensis Prance
Hirtella bicornis Mart. & Zucc.
Hirtella bicornis var. bicornis
Hirtella bicornis var. pubescens Ducke
Hirtella brachystachya Spruce ex Hook. f.
Hirtella bullata Benth.
Hirtella caduca Fanshawe & Maguire
Hirtella castilloana Prance
Hirtella ciliata Mart. & Zucc.
Hirtella confertiflora Prance
Hirtella cordifolia Prance
Hirtella couepiiflora Prance
Hirtella cowanii Prance & Maguire
Hirtella davisii Sandwith
Hirtella deflexa Maguire [expected in GU]
Hirtella duckei Huber
Hirtella elongata Mart. & Zucc.
Hirtella eriandra Benth. [expected in the Guianas]
Hirtella glabrata Pilg.
Hirtella glandistipula Ducke
SU FG
AM
FG
BO DA GU SU FG
BO
GU SU FG
AM BO GU
AM GU
SU
FG
FG
AM
BO GU SU
BO GU
GU
GU
SU
BO GU
GU
GU
GU
FG
FG
BO GU
AM BO DA GU SU FG
AM DA GU SU FG
AM BO GU SU FG
AM BO GU SU FG
AM GU FG
FG
FG
AM BO
GU
FG
AM GU SU FG
AM BO GU
AM BO GU
AM GU SU FG
AM
BO
BO
GU
FG
BO GU SU FG
AM BO GU
AM
GU FG
FG
AM BO GU SU
SU
FG
FG
AM BO GU SU FG
AM
AM BO
BO
GU
GU
SU FG
AM
GU FG
AM
AM
FG
AM
BO DA GU FG
AM BO
AM GU
AM BO
AM BO
GU FG
242 Checklist of the Plants of the Guiana Shield
Hirtella glandidosa Spreng.
Hirtella guainiae Spruce ex Hook. f.
Hirtella guyanensis (Fritsch) Sandwith
Hirtella hispidida Miq.
Hirtella liesneri Prance
Hirtella longipedicellata Prance
Hirtella macrophylla Benth. ex Hook. f.
Hirtella macrosepala Sandwith
Hirtella margae Prance
Hirtella mucronata Prance
Hirtella obidensis Ducke
Hirtella orbicularis Prance
Hirtella paniciilata Sw.
Hirtella physophora Mart. & Zucc.
Hirtella pimichina Lasser & Maguire
Hirtella piresii Prance
Hirtella punctillata Ducke
Hirtella racemosa Lam.
Hirtella racemosa var. hexandra (Willd. ex Roem. & Schult.) Prance
Hirtella racemosa var. hispida Prance
Hirtella racemosa var. racemosa
Hirtella scabra Benth.
Hirtella schidtesii Prance
Hirtella silicea Griseb.
Hirtella sitbscandens Spruce ex Hook. f.
Hirtella siiffulta Prance
Hirtella tentaculata Poepp.
Hirtella teniiifolia Prance
Hirtella triandra Sw.
Hirtella triandra subsp. triandra
Hirtella ulei Pilg.
Licania affinis Fritsch
Licania alba (Bernoulli) Cuatrec.
Licania albiflora Fanshawe & Maguire
Licania amapaensis Prance
Licania apetala (E. Mey.) Fritsch
Licania apetala var. aperta (Benth.) Prance
Licania apetala var. apetala
Licania arachnoidea Fanshawe & Maguire
Licania boyanii Tutin
Licania biixifolia Sandwith
Licania canescens Benoist
Licania cardiophylla Prance
Licania candata Prance
Licania compacta Fritsch [Mt. Roraima]
Licania cordata Prance
Licania coriacea Benth.
Licania couepifolia Prance
Licania crassivenia Spruce ex Hook. f.
Licania criiegeriana Urb.
Licania ciiprea Sandwith
Licania cyathodes Benoist
Licania davillifolia Benoist
Licania densiflora Kleinhoonte
Licania discolor Pilg.
Licania divaricata Benth.
Licania egleri Prance
Licania elliptica Standi.
Licania emarginata Spruce ex Hook. f.
Licaniafanshawei Prance
Licaniafoldatsii Prance
Licaniafoveolata Prance
Licaniafurfuracea Prance
Licania glabriflora Prance
BO DA GU SU FG
AM
BO GU
AM BO
BO
DA GU SU FG
AM
AM FG
BO DA GU
GU
GU
GU
SU
SU
SU
SU
FG
AM
AM BO DA GU SU FG
AM GU SU FG
AM
SU FG
AM BO GU SU FG
AM BO DA GU SU FG
AM BO DA GU
GU
SU FG
FG
AM BO DA GU SU FG
AM BO GU
AM BO
BO DA GU SU FG
AM
BO GU
GU
FG
SU FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM
GU SU FG
BO DA GU SU FG
GU SU FG
FG
AM BO DA GU SU FG
AM BO GU SU FG
AM BO
BO
DA GU
GU
GU
GU
SU FG
AM BO GU SU FG
AM
GU
GU
FG
AM
AM BO
BO
BO
GU
GU SU
FG
SU FG
SU FG
AM BO DA GU SU FG
BO DA GU SU FG
DA GU SU FG
AM
SU FG
AM
DA GU SU FG
AM
BO
GU
BO GU SU FG
Checklist of the Plants of the Guiana Shield 243
Licania gracilipes Taub. AM
Licania granvillei Prance AM GU FG
Licania guianemis (Aubl.) Griseb. DA GU SU FG
Licania heteromorpha Benth. AM BO DA GU SU FG
Licania heteromorpha var. glabra (Mart, ex Hook, f.) Prance AM BO GU FG
Licania heteromorpha var. heteromorpha AM BO DA GU SU FG
Licania heteromorpha var. perplexans Sandwith GU
Licania heteromorpha var. subcordata Fritsch AM
Licania hirsuta Prance AM
Licania hispida Prance AM
Licania hitchcockii Maguire BO
Licania hypoleuca Benth. AM BO
Licania hypoleuca var. hypoleuca AM BO
Licania imbaimadaiensis Prance
Licania incana Aubl. AM BO
Licania intrapetiolaris Spruce ex Hook. f. AM BO
Licania irwinii Prance BO
Licaniajimenezii Prance
Licania kunthiana Hook. f. BO
Licania laevigata Prance AM
Licania lanceolata Prance AM
Licania lasseri Maguire BO
Licania lata J.F. Macbr. AM BO
Licania latifoUa Benth. ex Hook. f. AM BO
Licania latistipula Prance DA
Licania laxiflora Fritsch AM BO
Licania leptostachya Benth. BO
Licania leucosepala Griseb. AM BO DA
Licania licaniiflora (Sagot) S.F. Blake AM
Licania longistyla (Hook, f.) Fritsch AM DA
Licania macrophylla Benth. AM
Licania majuscula Sagot
Licania maxima Prance
Licania membranacea Sagot ex Laness.
Licania micrantha Miq. AM BO
Licania micrantha subsp. atabapoensis Prance AM
Licania micrantha subsp. micrantha AM BO
Licania microphylla Fanshawe & Maguire
Licania minutiflora (Sagot) Fritsch AM BO
L/cama wo/fo Benth. [US type from Br. Guiana! R. Schomb. 910] AM BO
Licania oblongifolia Standi. AM
Licania obtusifolia Fritsch [dubious species] FG
Licania occultans Prance [expected]
Licania octandra (Hoffmanns, ex Roem. & Schult.) Kuntze AM BO DA GU SU FG
Licania octandra subsp. octandra BO DA GU SU FG
Licania octandra subsp. pallida (Hook, f.) Prance AM GU
Licania orbicularis Spruce ex Hook. f. AM BO SU
Licania ovalifolia Kleinhoonte GU SU FG
Licania pakaraimensis Prance BO
Licania pallida Spruce e\ Sagot AM BO GU FG
Licania pai-viflora Benth. AM BO GU FG
Licania parvifolia Uuber AM GU
Licania parvifructa Fansha-we & Maguire AM BO GU FG
Licania persaudii Fanshawe & Maguire GU FG
Licania piresii Prance GU FG
Licania polita Spruce ex Hook. f. AM BO GU SU FG
Licania prismatocarpa Spruce ex Hook. f. AM
Licania pruinosa Benoist FG
Licania pyrifolia Griseb. AM BO DA
Licania reticulata Prance
Licania robusta Sagot
Licania rodriguesii Prance
Licania roraimensis Standi. [Mt. Roraima]
Licania rufescens Klotzsch ex Fritsch BO DA
GU SU FG
GU SU FG
GU
GU SU FG
GU
GU SU FG
GU SU
GU SU FG
GU SU FG
GU
FG
GU FG
FG
GU SU FG
GU SU FG
GU SU FG
GU SU FG
SU FG
GU SU FG
FG
GU FG
GU SU FG
GU SU FG
GU
GU SU FG
GU FG
SU FG
FG
GU
GU SU
244 Checklist of the Plants of the Guiana Shield
Licania sandwithii Prance
Licania savannarum Prance
Licania silvae Prance
Licania sp. A
Licania sprucei (Hook, f.) Fritsch
Licania steyermarkii Maguire
Licania stricta Kleinhoonte [dubious species]
Licania subarachnophylla Cuatrec.
Licania tepuiensis Prance
Licania triandra Mart, ex Hook. f.
Licania urceolaris Hook. f.
Licania vaupesiana Killip & Cuatrec.
Licania wurdackii Prance
Parinari campestris Aubl.
Pahnari excelsa Sabine
Parinari maguirei Prance
Parinari montana Aubl., emend. Ducke °
A \/f
GU
AM GU su FG
AM
AM BO GU FG
AM BO
SU
AM
BO
Parinari
Parinari
Parinari
pachyphylla Rusby
parvifolia Sandwith
rodolphii Huber
AM
AM
AM
AM
AM BO DA GU SU FG
AM BO
BO
DA GU
GU
SU FG
FG
AM BO
GU FG
BO DA GU SU FG
Parinari sprucei Hook. f. AM
Synonyms and Misidentifications
Balantium cordifolium Desv. ex Ham. =Parinari campestris
Chrysobalanus icaco var. pellocarpus (G. Mey.) Hook. f. =Chrysobalanus icaco
Chrysobalanus incanus (Aubl.) M. Gomez =Licania incana
Chrysobalanus montanus (Aubl.) M. Gomez =Parinari montana
Chrysobalanus pellocarpus G. Mey. =Chrysobalanus icaco
Chrysobalanus savannarum Britton =Chrysobalanus icaco
Chrysobalanus triandra (Sw.) Morales =Hirtella triandra
Couepia calophlebia Standi. =Couepia racemosa
Couepia divaricata Huber =Couepia guianensis subsp. divaricata
Couepia duckei Huber =Couepia paraensis subsp. glaucescens
Couepia glandulosa Miq. =Couepia guianensis subsp. glandulosa
Couepia glaucescens Spruce ex Hook. f. =Couepia paraensis subsp. glaucescens
Couepia leptostachya Benth. ex Hook. f. =Couepia guianensis subsp. guianensis
Couepia myrtifolia Benth. ex Hook. f. =Couepia guianensis subsp. glandulosa
Couepia pauciflora Huber =Couepia parillo
Couepia steudeliana Walp. =Couepia cognata
Couepia surinamensis Kleinhoonte =Couepia guianensis subsp. guianensis
Couepia thyrsiflora Hook. f. =Couepia guianensis subsp. glandulosa
Couepia versicolor Benoist =Couepia guianensis subsp. guianensis
Couepia villosa Fanshawe & Maguire =Couepia parillo
Ferolia amazonica Kuntze =Parinari excelsa
Ferolia coriacea (Benth.) Kuntze =Exellodendron coriaceum
Ferolia excelsa (Sabine) Kuntze =Parinari excelsa
Ferolia montana (Aubl.) Kuntze =Parinari montana
Hirtella aggregata Poir. =Hirtella paniculata
Hirtella americana Jacq. =Hirtella triandra
Hirtella americana L. [not in flora area]
Hirtella americana Aubl. =Hirtella racemosa
Hirtella americana var. hexandra (Willd. ex Roem. & Schult.) Hook. f. =Hirtella racemosa var. hexandra
Hirtella amplexicaulis Pilg. =Hirtella bullata
Hirtella apetala E. Mey. =Licama apetala
Hirtella aureohirsuta Pilg. =Hirtella bullata
Hirtella bracteata Mart. & Zucc. =Hirtella triandra
Hirtella bracteosa Steud. =Hirtella paniculata
Hirtella canomensis (Mart.) Spreng. =Couepia canomensis
Hirtella castanea DC. =Hirtella triandra
Hirtella caudata Kleinhoonte =Hirtella bicornis var. pubescens
Hirtella cauliflora Huber =Hirtella physophora
Hirtella cognata Steud. =Couepia cognata
Hirtella coppenamensis Kleinhoonte =Hirtella obidensis
Hirtella cotticaensis Kleinhoonte =Hirtella physophora
Checklist of the Plants of the Guiana Shield 245
Hirtella damaziana Beauverd =Hirtella glandulosa
Hirtella egemis Fritsch =Hirtella hispidula
Hirtellaformicaria Suess. =Hirtella duckei
Hirtella glaberrima Steud. (Leguminosae-Caesalpinioideae) =Crudia glaberrima
Hirtella hexandra Willd. ex Roem. & Schult. =Hirtella racemosa var. hexandra
Hirtella hirsuta Lam. =Hirtella paniculata
Hirtella hookeri Pilg. =Hirtella glandulosa
Hirtella indecora Schott =Hirtella paniculata
Hirtellajamaicensis Urb. =Hirtella triandra
Hirtella lanceolata Kleinhoonte =Hirtella obidensis
Hirtella manigera Kleinhoonte ^Hirtella silicea
Hirtella melinonii Benoist =Hirtella silicea
Hirtella multiflora Urb. =Hirtella triandra
Hirtella nemorosa Hoffmanns, ex Roem. & Schult. =Hirtella racemosa
Hirtella octandra Hoffmanns, ex Roem. & Schult. =Licania octandra
Hirtella paniculata Lam. =Hirtella triandra
Hirtella peruviana Pers. =Hirtella triandra
Hirtella praealta Sagot =Hirtella bicornis var. bicornis
Hirtella racemosa Ruiz & Pav. =Hirtella triandra
Hirtella rubra Benth. =Hirtella ciliata
Hirtella scandens Hoffmanns, ex Roem. & Schult. =Hirtella racemosa
Hirtella stipitadenia Rusby =Hirtella bullata
Hirtella strigulosa Steud. =Hirtella racemosa
Hirtella subsetosa Fanshawe & Maguire =Hirtella hispidula
Hirtella velutina Pilg. =Hirtella glandulosa
Hirtella violacea Steud. =Hirtella racemosa
Hirtella wachenheimii Benoist =Hirtella glandulosa
Licania anisophylla Standi. =Licania micrantha subsp. micrantha
Licania aperta Benth. =Licania apetala var. aperta
Licania apetala var. pendula (Benth.) Fritsch =Licania apetala var. apetala
Licania arachnites Standi. =Licania parvifolia
Licania aubletiana (Blume) Lemee =Acioa guianensis
Licania axilliflora (Sagot) Hochr. =Licania leptostachya
Licania benthamii Hook, f =Licania heteromorpha var. heteromorpha
Licania biglandulosa Griseb. ex Urb. =Licania heteromorpha var. heteromorpha
Licania bothynophylla Mart. =Licania octandra subsp. octandra
Licania campinensis Huber =Licania licaniiflora
Licania caracasana Klotzsch ex Hook. f. =Licania apetala var. aperta
Licania costata Spruce ex Hook. f. =Licania heteromorpha var. glabra
Licania crassifolia Benth. =Licania incana
Licania cymosa Fritsch [not in flora area]
Licania dahlgrenii Standi. =Licania apetala var. apetala
Licania egensis Fritsch ^Licania octandra subsp. pallida
Licania exiguifolia Standi. =Couepia cognata var. cognata
Licaniafloribunda Benth. =Licania apetala var. apetala
Licania galibica Benoist ^Licania membranacea
Licania glabra Mart, ex Hook, f =Licania heteromorpha var. glabra
Licania gracilis Kleinhoonte =Licania laxiflora
Licania grisea Kleinhoonte =Licania hypoleuca
Licania helvola Spruce ex Hook, f =Licania micrantha subsp. micrantha
Licania heteromorpha var. divaricata (Benth.) Fritsch =Licania divaricata
Licania hookeri Fritsch =Licania octandra subsp. pallida
Licania hookeri var. obtusa Huber =Licania octandra subsp. octandra
Licania hostmannii Fritsch =Licania majuscula
Licania huberiana Maguire =Licania licaniiflora
Licania hylaea Cuatrec. =Licania apetala var. apetala
Licania hypagyraea Malme =Licania kunthiana
Licania incana var. axilliflora Sagot =Licania leptostachya
Licania kanukuensis Standi. =Licania densiflora
Licania kuntzeana Urb. =Licania apetala var. aperta
Licania laurifolia Huber =Licania polita
Licania laxa Fanshawe & Maguire =Licania polita
Licania leptostachya var. axilliflora (Sagot) Fritsch =Licania leptostachya
Licania leptostachya var. crassifolia (Benth.) Benoist =Licania incana
246 Checklist of the Plants of the Guiana Shield
Licania longifolia Benoist =Licania alba
Licania hicida J.F. Macbr. =Licania mollis
Licania micrantha var. triandra (Mart, ex Hook, f.) Fritsch =Licania triandra
Licania microcarpa Hook. f. =Licania hypoleuca
Licania obovata Benth. ex Hook. f. =Licania latifolia
Licania obovatifolia Maguire =Licania licaniiflora
Licania pachystachya Kleinhoonte =Licania robusta
Licania pallida (Hook, f.) Kuntze ='Licania octandra subsp. pallida
Licania pallida Spruce ex Hook. f. =Licania pallida
Licania pallidula Standi. =Licania coriacea
Licania paniculata Fanshawe & Maguire =Licania elliptica
Licania parinarioides Huber =Licania licaniiflora
Licania parviflora (Blume) Lemee, nom. illeg. =Licania obtiisifolia
Licania parviflora var. condiiplicata Maguire =Licania hypoleuca
Licania parviflora var. membranacea Maguire =Licania kimthiana
Licania parviflora vox. pallida Hook. f. ^Licania pallida
Licania parvifolia Pittier =Licania hypoleuca
Licania pendula Benth. =Licania apetala var. apetala
Licania petrensis Prance =Licania emarginata
Licania poeppigii Fritsch =Licania polita
Licania pubiflora Benth. =Licania apetala var. aperta
Licania pulchravenia Killip & Cuatrec. =Licania triandra
Licania riparia (Gleason) Standi. =Licania minutiflora
Licania rondonii Pilg. =Licania minutiflora
Licania sclerophylla (Mart, ex Hook, f.) Fritsch [not in flora area]
Licania stahelii Kleinhoonte ^Licania ovalifolia
Licania stenocarpa Standi. =Licania octandra subsp. pallida
Licania stipata Fanshawe & Maguire [nomen nudum]
Licania takutuensis Standi. =Licania octandra subsp. octandra
Licania turiuva Cham. & Schltdl., nom. illeg. =Licania octandra subsp. octandra
Licania utilis (Hook, f.) Fritsch =Licania octandra subsp. octandra
Licania venosa Rusby =Licania alba
Licania wilson-brownei Maguire =Licania licaniiflora
Moquilea aubletiana Blume, nom. illeg. =Acioa guianensis
Moquilea bothynophylla (Mart.) Hook. f. =Licania octandra subsp. octandra
Moquilea canomensis Mart. =Couepia canomensis
Moquilea chrysocalyx Poepp. =Couepia chrysocalyx
Moquilea comosa (Benth.) Walp. =Couepia comosa
Moquilea couepia Steud. =Couepia guianensis
Moquileafloribunda (Benth.) Hook. f. =Licania apetala var. apetala
Moquilea glandulosa (Miq.) Walp. =Couepia guianensis subsp. glandulosa
Moquilea guianensis Aubl. =Licania guianensis
Moquilea kuntzeana (Urb.) R.O. Williams =Licania apetala var. aperta
Moquilea leucosepala (Griseb.) R.O. Williams ^Licania leucosepala
Moquilea licaniiflora Sagot =Licania licaniiflora
Moquilea longistyla Hook. f. =Licania longistyla
Moquilea macrocarpa Pittier =Licania pyrifolia
Moquilea minutiflora Sagot ^Licania minutiflora
Moquilea multiflora (Benth.) Walp. =Couepia multiflora
Moquilea orinocensis Rusby =Licania apetala var. apetala
Moquilea pallida Hook. f. =Licania octandra subsp. pallida
Moquilea paraensis Mart. & Zucc. =Couepia paraensis
Moquilea pendula (Benth.) Hook. f. =Licania apetala var. apetala
Moquilea pyrifolia (Griseb.) R.O. Williams =Licania pyrifolia
Moquilea riparia Gleason =Licania minutiflora
Moquilea sprucei Hook. f. =Licania sprucei
Moquilea turiuva (Cham. & Schltdl.) Hook. f. =Licania octandra
Moquilea utilis Hook. f. =Licania octandra subsp. octandra
Parinari barbatum Ducke =Exellodendron barbatum
Parinari brachystachya Benth. =Parinari excelsa
Parinari canescens Gleason =Couepia canescens
Parinari coriacea Benth.
Parinari glazioviana Warm. =Parinari excelsa
Parinari guyanense Fritsch =Hirtella guyanensis
Checklist of the Plants of the Guiana Shield 247
Parinari guyanensis (Aubl.) Lemee (Moraceae) =Brosimum rubescens
Parinari hostmannii Fritsch =Couepia cognata
Parinari krukovii Gleason =Couepia guianensis subsp. divaricata
Parinari laxiflora Ducke =Parinari excelsa
Parinari lucidissima Standi. =Parinari rodolphii
Parinari pilosa Standi. =Couepia canomensis
Parinari sylvestris Kuhlm. =Parinari excelsa
Petrocarya campestris (Aubl.) Willd. =Parinari campestris
Petrocarya excelsa (Sabine) Steud. =Parinari excelsa
Petrocarya montana (Aubl.) Willd. =Parinari montana
Salmasia guianensis (Aubl.) J.F. Gmel. =Hirtella racemosa
Salmasia racemosa Willd. =Hirtella racemosa
Tachibota guianensis Aubl. =Hirtella racemosa
References
Kleinhoonte, A. 1939. Rosaceae. In A. PuUe, Flora of Suriname 2(1): 426-456.
Prance, G.T. 1972. Chrysobalanaceae. Flora Neotropica Monogr. 9: 1-409.
Prance, G.T. 1981. Notes on Couepia and Hirtella. Brittonia 33: 347-356.
Prance, G.T. 1982. Chrysobalanaceae. Flora de Venezuela 14(2): 325-487.
Prance, G.T. 1986a. Chrysobalanaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 2: 1-146.
Prance, G.T. 1986b. Studies on the flora of the Guianas. 19. New taxa of Chrysobalanaceae for the Flora of the Guianas. Proc.
Kon. Ned. Akad. Wetensch., sen C. 89: 111-116.
Prance, G.T. 1989. Chrysobalanaceae (supplement). Flora Neotropica Monogr. 9S: 1-267.
Prance, G.T. 1992. New species and new records of neotropical Chrysobalanaceae. Kew Bull. 47: 633-646.
Prance, G.T. 1998. Chrysobalanaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 202-
246.
Prance, G.T. 2002. Chrysobalanaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 202-212.
Prance, GT. & A.R.A. Gorts-van Rijn. 1976. Chrysobalanaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora
ofSuriname 2(2): 524-555.
Clethraceae
RE. Berry, J.S. Miller
Clethra guyanensis Klotzsch ex Meisn. [Mt. Roraima] BO GU
References
Miller, J.S. & RE. Berry. 1998. Clethraceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4:
246-247.
Sleumer, H. 1967. Monographia Clethracearum. Bot. Jahrb. Syst. 87: 36-175.
Sleumer, H. 1971. Clethraceae. In T. Lasser, Flora de Venezuela 3(1): 171-183.
Clusiaceae
RE. Berry, N.L. Cuello A., B.K. Hoist, D.M. Keams, K. Kubitzki, J.J. Pipoly,
N.K.B. Robson, RF. Stevens, A.L. Weitzman
Calophyllum brasiliense Cambess.
Calophyllum longifolium Willd.
Calophyllum pachyphyllum Planch. & Triana
Caraipa ampla Ducke
Caraipa densifolia Mart.
Caraipa densifolia subsp. densifolia
Caraipa grandifolia Mart.
Caraipa grandifolia subsp. grandifolia
Caraipa heterocarpa Ducke
Caraipa llanorum Cuatrec.
Caraipa llanorum subsp. cordifolia Kubitzki
Caraipa llanorum subsp. llanorum
Caraipa longipedicellata Steyerm.
Caraipa parvielliptica Cuatrec.
Caraipa parvifolia Aubl.
Caraipa psilocarpa Kubitzki
Caraipa punctulata Ducke
AM
AM?
BO DA
AM BO
AM BO
AM BO
AM BO
AM
AM BO
AM
AM BO
AM BO
AM BO
BO
AM BO
GU su FG
GU su
EG
GU su FG
GU su FG
GU
GU
GU?
GU SU
FG
FG
248 Checklist of the Plants of the Guiana Shield
& Triana
Planch. & Triana) Hemsl.
Caraipa racemosa Cambess.
Caraipa richardiana Cambess.
Caraipa savannanim Kubitzki
Caraipa sp. A
Caraipa spuria Barb. Rodr.
Caraipa tereticaulis Tul.
Chrysochlamys membranacea Planch.
Chrysochlamys nicaraguensis (Oerst.,
Chrysochlamys pauciflora Steyerm. [may be synonymous with
Chrysochlamys weberbaueri]
Chrysochlamys weberbaueri Engl.
Clusia aishaltonensis Pipoly, sp. nov. ined.
Clusia amabilis Maguire [Section Clusiastrum]
Clusia amazonica Planch. & Triana [Section Criuvopsis^
Clusia annularis Maguire [Section Omphalanthera]
Clusia asymmetrica Pipoly [Section Criuva]
Clusia aymardii Pipoly
Clusia brachystyla Maguire
Clusia brachystyla van brachystyla
Clusia brongniartiana Planch. & Triana [dubious species]
Clusia candelabrum Planch. & Triana
Clusia capituliflora Pipoly, sp. nov. ined.
Clusia cardonae Maguire
Clusia cerroana Steyerm.
Clusia chiribiquetensis Maguire
Clusia cochlitheca Maguire
Clusia columnaris Engl.
Clusia comans (Meisn.) Pipoly
Clusia crassifolia Planch. & Triana
Clusia cuneata Benth.
Clusia duartei Maguire
Clusia duidae Gleason
Clusiafabiolae Pipoly
Clusiaflavida (Benth.) Pipoly
Clusia flaviflora Engl.
Clusiafockeana Miq.
Clusia gaudichaudii Choisy
Clusia grammadenioides Pipoly
Clusia grandiflora Splitg.
Clusia gratula Maguire
Clusia guayanae Pipoly
Clusia hammeliana Pipoly
Clusia hexacarpa Gleason
Clusia huberi Pipoly [Section Oedematopus]
Clusia imbricata Steyerm.
Clusia insignis Mart. [=Clusia grandiflora?]
Clusia kanukuana Maguire
Clusia leprantha Mart. [=Clusia purpurea!]
Clusia lopezii Maguire [Section Retinostemon]
Clusia lunanthera Maguire
Clusia macropoda Klotzsch ex Engl. [Section Clusia]
Clusia maguireana Pipoly [Section Criuva]
Clusia martiana Engl. [Section Criuvopsis]
Clusia martinii Sagot {=Clusia palmicidal]
Clusia melchiori Gleason [Section Criuva]
Clusia microstemon Planch. & Triana [Section Phloianthera]
Clusia minor L. [Section Retinostemon]
Clusia multilineata Pipoly [Section Criuva]
Clusia mutica Maguire
Clusia myriandra (Benth.) Planch. & Triana [Section Phloianthera]
Clusia nemorosa G. Mey. [Section Clusia]
Clusia obovata (Spruce ex Planch. & Triana) Pipoly
Clusia octandra (Poepp.) Pipoly
Clusia opaca Maguire [Section Criuva]
BO
AM BO
AM
AM
AM BO
AM BO
BO
BO
BO
AM
AM
AM
BO
AM BO
DA
AM
AM
AM
AM
AM
AM
AM
BO
BO
AM BO
AM BO
AM
AM BO
BO
AM
AM BO
BO
AM BO
BO
BO
BO
BO
BO
BO
DA
DA
DA
DA
DA
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
SU
su
SU
GU SU
GU
GU
GU
FG
FG
FG
FG
FG
GU
GU FG
GU
GU
SU
FG
GU SU FG
GU SU
FG
AM GU
GU
SU FG
FG?
AM GU SU FG
AM
SU
DA GU SU FG
BO GU
AM
GU
AM BO GU SU FG
AM DA GU SU
AM BO DA GU SU FG
AM
BO
GU
GU
BO DA GU SU FG
AM BO GU SU FG
AM BO GU FG
AM
AM BO
AM BO DA GU su FG
AM
BO
BO
DA GU
GU
su FG
GU
GU? SU
GU su
GU su
GU su
GU su
GU su
GU
GU
GU
GU su
Checklist of the Plants of the Guiana Shield 249
Clusia pachyphylla Gleason [?>QCi\on Anandrogyne]
Clusia palmicida Rich, ex Planch. & Triana [Section Clusia]
Clusia panapanari (Aubl.) Choisy [Section Phloianthera]
Clusia pai-vifolia Maguire [Section Clusiastnim]
Clusia parvula (Maguire) Pipoly
Clusia phelpsiae Lasser & Maguire [Section Anandrogyne] AM
Clusia phelpsiana Maguire [expected in GU] BO
Clusia picta Pipoly, sp. nov. ined.
Clusia platystigma Eyma FG
Clusia ptaritepuiemis (Steyerm.) Pipoly [Section Oedematopus] BO
Clusia pusilla Steyerm. AM BO
Clusia pusilla subsp. orinocensis Maguire AM BO
Clusia pusilla suhsp. pusilla [Section Clusiastnim] BO
Clusia radiata Maguire & K.D. Phelps [Section Criuva] AM
Clusia renggehoides Planch. & Triana [Section Phloianthera] AM
Clusia robusta Eyma
Clusia rosea Jacq. [^Clusia major L.?] BO SU FG
Clusia rotundifolia Gleason [Section Anandrogyne] AM
Clusia savannarum Maguire [Section Anandrogyne] BO
Clusia schomburgkiana (Planch. & Triana) Benth. ex Engl. [Section AM BO
Omphalanthera]
Clusia schomburgkii Vesque [Section Clusiastrum] BO
Clusia scrobiculata Benoist SU FG
Clusia sipapoana (Maguire) Pipoly [Section Quapoya] AM GU
Clusia sp. A [Section Oedematopus] AM
Clusia sp. B [Section Oedematopus] BO
Clusia sp. C [Section Oedematopus] AM BO
Clusia sp. D [Section Oedematopus] AM BO GU
Clusia spathulaefolia Engl. [Section Brachystemon] AM
Clusia steyermarkii Maguire [Section Clusiastrum] BO
Clusia stylosa Maguire
Clusia tabulamontana Maguire [Section Clusiastrum] BO
Clusia troncosii Maguire [Section uncertain] AM
Clusia viscida Engl. [Section Clusia] AM
Clusiella axillaris (Engl.) Cuatrec. AM
Clusiella elegans Planch. & Triana
Clusiella sp. A BO
Dystovomita sp.
Haploclathra leiantha (Benth.) Benth.
Hypericum marahuacanum N. Robson AM BO
Hypericum marahuacanum subsp. chimantaicum N. Robson [expected in GU] BO
Hypericum marahuacanum subsp. marahuacanum AM
Hypericum roraimense Gleason [Mt. Roraima] BO GU
Hypericum thesiifolium Kunth [expected in GU] BO
Lorostemon bombaciflorum Ducke GU
Lorostemon colombianus Maguire AM
Mahurea exstipulata Benth. AM BO
Mahurea exstipulata subsp. duckei (Huber) Kubitzki AM
Mahurea exstipulata subsp. exstipulata BO
Mahurea palustris Aubl.
Marila laxiflora Rusby AM BO
Marila saramaccana Pulle
Moronobea coccinea Aubl. BO
Moronobea intermedia Engl. BO
Moronobeajenmanii Engl. BO
Moronobea ptaritepuiana Steyerm. AM BO
Moronobea ptaritepuiana subsp. neblinensis Maguire AM
Moronobea ptaritepuiana suhsp. ptaritepuiana BO
Moronobea riparia (Spruce) Planch. & Triana AM GU
Neotatea duidae (Kobuski & Steyerm.) P.F. Stevens & A.L. Weitzman AM
Neotatea longifolia (Gleason) Maguire AM
Neotatea neblinae Maguire AM
Platonia insignis Mart. AM GU SU FG
Quapoya scandens k\xh\. FG
SU
GU su
GU su
GU
GU
GU
GU
GU
GU SU FG
GU su FG
GU
SU
FG
GU su FG
GU
GU FG'
GU
250 Checklist of the Plants of the Guiana Shield
Rheedia benthamiana Planch. & Triana
Rheedia brasiliensis (Mart.) Planch. & Triana \=Garcima brasiliensis Mart.]
Rheedia lateriflora L. [=Garcinia humilis (Vahl) C. Adams]
Rheedia macrophylla (Mart.) Planch. & Triana [=Garcinia macrophylla Mart.] AM
Rheedia madruno (Kunth) Planch. & Triana [=Garcinia madruno (Kunth) AM
Hammel]
Rheedia martinii Maguire [=Garcinia sp.]
Rheedia virens Planch. & Triana [=Garcinia sp.]
Symphonia globulifera L. f. AM
Thysanostemonfanshawei Maguire
Thysanostemon pakaraimae Maguire
Tovomita acutiflora M.S. Barros & G. Mariz
Tovomita albiflora A.C. Sm.
Tovomita atropurpurea Steyerm.
Tovomita brasiliensis (Mart.) Walp.
Tovomita aff. brasiliensis (Mart.) Walp. AM
Tovomita brevistaminea Engl.
Tovomita calodictyos Sandwith
Tovomita caloneura A.C. Sm.
Tovomita carinata Eyma AM
Tovomita choisyana Planch. & Triana
Tovomita clusiifolia G. Don
Tovomita divaricata Maguire
Tovomita eggersii Vesque AM
Tovomita elliptica Engl. AM
Tovomitafanshawei Maguire AM
Tovomita aff. glazioviana Engl. AM
Tovomita gracilipes Planch. & Triana AM
Tovomita grata Sandwith
Tovomita guianensis Aubl.
Tovomita longifolia (Rich.) Hochr. AM
Tovomita macrophylla (Poepp.) Walp. AM
Tovomita melinonis Vesque
Tovomita obovata Engl.
Tovomita obscura Sandwith
Tovomita pyrifolia Planch. & Triana
Tovomita rubella Spruce ex Planch. & Triana AM
Tovomita schomburgkii Planch. & Triana AM
Tovomita secunda Poepp. ex Planch. & Triana
Tovomita sp. A AM
Tovomita sp. B AM
Tovomita sp. C AM
Tovomita sp. D AM
Tovomita sp. E BO
Tovomita sp. F AM
Tovomita spruceana Planch. & Triana AM
Tovomita tenuiflora Benth. ex Planch. & Triana AM
Tovomita umbellata Benth. AM
Tovomita uniflora G. Don
Tovomita weddelliana Planch. & Triana
Tovomitidium speciosa (Ducke) Ducke [expected in FG]
Vismia baccifera (L.) Triana & Planch. AM
Vismia baccifera subsp. dealbata (Kunth) Ewan AM
Vismia cavalcantii M.E. Berg
Vismia cayennensis (Jacq.) Pers. AM
Vismia confertiflora Spruce ex Reichardt
Vismia gracilis Hieron.
Vismia guianensis (Aubl.) Choisy AM
Vismiajapurensis Reichardt AM
Vismia latifolia (Aubl.) Choisy
Vismia lauriformis (Lam.) Choisy [expected in GU]
Vismia laxiflora Reichardt AM
Vismia macrophylla Kunth AM
Vismia ramuliflora Miq.
GU SU FG
FG
GU
BO DA GU SU FG
BO GU
GU
GU
SU
SU
FG
BO DA GU
GU
GU
SU FG
BO GU
GU
BO GU
GU SU FG
BO DA GU
GU
GU
SU
SU
FG
GU SU FG
GU SU FG
FG
GU
BO GU
BO GU
BO GU?
GU SU FG
GU FG
BO DA GU
GU
GU
GU
GU
GU?
SU
SU?
FG
FG
BO GU SU FG
GU? SU FG
BO GU FG
BO GU
BO GU FG
FG
BO GU
BO SU? FG?
BO SU?
SU
FG?
BO DA GU SU FG
GU? SU FG
GU SU FG
BO DA GU SU FG
BO GU SU
GU SU FG
BO FG?
BO DA GU
BO DA GU SU FG
GU SU FG
Checklist of the Plants of the Guiana Shield 251
Vismia sandwithii Ewan
Vismia schultesii N. Robson
Vismia sessilifolia (Aubl.) Choisy
Vismia sprucei Sprague
Vismia steyermarkii N. Robson
Vismia tenuinervia (M.E. Berg) N. Robson
Synonyms and Misidentifications
Androstyliumfockeanum (Miq.) Miq. =Clusiafockeana
Arrudea purpurea Splitg. =Clusia leprantha
Calophyllum antillanum Britton =Calophyllum brasiliense
Calophyllum brasiliense var. antillanum (Britton) Standi. =Calophyllum brasiliense
Calophyllum lucidum Benth. =Calophyllum brasiliense
Calophyllum madruno Kunth =Rheedia madruno
Calophyllum piaroanum A. Castillo & C. Gil =Calophyllum brasiliense
Caopia acuminata (Lam.) Kuntze =Vismia guianensis
Caopia cayennensis (Jacq.) Kuntze =Vismia cayennensis
Caopia guianensis (Aubl.) Lyons =Vismia guianensis
Caopia latifolia (Aubl.) Kuntze =Vismia latifolia
Caopia laxiflora (Reichardt) Kuntze =Vismia laxiflora
Caopia sessilifolia (Aubl.) Kuntze =Vismia sessilifolia
Caraipa angustifolia Aubl. (Chrysobalanaceae) =Couepia sp.
Caraipaferruginea Steyerm. =Caraipa punctulata
Caraipa glabrata Mart, [not in flora area] =Caraipa grandifolia
Caraipa lacerdaei Barb. Rodr. [not in flora area] =Caraipa grandifolia
Caraipa latifolia Aubl. (Chrysobalanaceae) =Licania membranacea
Caraipa laxiflora Benth. =Caraipa densifolia
Caraipa leiantha Benth. =Haploclathra leiantha
Caraipa longifolia Aubl. (Chrysobalanaceae) =Licania alba
Caraipa paraensis Huber [not in flora area] =Caraipa grandifolia
Caraipa psidiifolia Ducke =Caraipa richardiana
Caraipa simplicior Sandwith ? =Caraipa tereticaulis
Caraipa variabilis Cambess. ? =Caraipa densifolia
Clusia axillaris Engl. =Clusiella axillaris
Clusia brachystyla var. guaiquinimensis Maguire =Clusia brachystyla
Clusia colorans Klotzsch ex Engl. =Clusia panapanari
Clusia columnaris var. magdalenensis Cuatrec. =Clusia columnaris
Clusia columnaris var. vaupesana Cuatrec. =Clusia columnaris
Clusia cucullata Klotzsch [nomen nudum]
Clusia galactodendron Desv. =Clusia minor
Clusia grandifolia Engl. =Clusia candelabrum
Clusia hexacarpa var. ptaritepuiana Steyerm. =Clusia hexacarpa
Clusia insignis var. Hoffmannseggiana Engl. =Clusia insignis
Clusiajenmanii Engl. =Clusia myriandra
Clusia longifolia Rich. =Tovomita longifolia
Clusia microphylla Klotzsch ex Engl. =Clusia panapanari
Clusia oedematopoidea Maguire =Clusia amazonica
Clusia parvicapsula Vesque =Clusia minor
Clusia planchoniana Engl. =Clusia schomburgkiana
Clusia planchoniana var. oblongifolia Vesque =Clusia schomburgkiana
Clusia purpurea (Splitg.) Engl. =Clusia leprantha
Clusia pusilla suhsp.Jauaensis Maguire =Clusia pusilla subsp. pusilla
Clusia pusilla suhsp. pakaraimae Maguire =Clusia pusilla subsp. pusilla
Clusia reducta Steyerm. =Clusia opaca
Clusia roraimana Steyerm. =Clusia schomburgkii
Clusia rosea var. colombiana Cuatrec. =Clusia rosea
Clusia scariosa Lasser & Maguire =Clusia pachyphylla
Clusia schomburgkiana var. lakei Maguire =Clusia schomburgkiana
Clusia sessilis Klotzsch ex Engl, [latter homonym of C. sessilis G. Forst.] ^Clusia cerroana
Clusia spruceana Planch. & Triana =Clusia schomburgkiana
Clusia stahelii Maguire =Clusiafockeana
Clusia uleana Engl. =Clusia martiana
Clusia venosa Jacq. =Clusia minor
Clusia williamsii Steyerm. =Clusia columnaris
Clusia wurdackiana Pipoly [nom. superfl.] =Clusia cerroana
AM GU su FG
AM GU?
AA/f
BO GU su FG
BO
AM
252 Checklist of the Plants of the Guiana Shield
Garcinia acuminata (Ruiz & Pav.) Hammel =Rheedia madnino
Garcinia benthamiana (Planch. & Triana) Pipoly [nomen nudum] =Rheedia benthamiana
Garcinia brasiliensis Mart. =Rheedia brasiliensis
Garcinia macrophylla Mart. =Rheedia macrophylla
Garcinia madnino (Kunth) Hammel =Rheedia madruno
Havetiaflavida Benth. =Clusiaflavida
Havetia octandra Poepp. =Chisia octandra
Havetiopsis flavida (Benth.) Planch. & Triana =Chisiaflavida
Havetiopsis flexilis Spruce ex Planch. & Triana =Clusiaflavida
Hypericum acuminatum Lam. =Vismia guianensis
Hypericum bacciferum L. =Vismia baccifera
Hypericum cayennensis Jacq. =Vismia cayennensis
Hypericum guianense Aubl. =Vismia guianensis
Hypericum latifolium Aubl. =Vismia latifolia
Hypericum lauriforme Lam. =Vismia lauriformis
Hypericum reticulatum Poir. =Vismia macrophylla
Hypericum rufescens Lam. =Vismia sessilifolia
Hypericum sessilifolium Aubl. =Vismia sessilifolia
Hypericum silenoides auct. non Juss. 1789. =Hypericum thesiifolium
Hypericum uliginosum Kunth =Hypericum thesiifolium
Mahurea casiquiarensis Benth. =Mahurea exstipulata
Mahurea duckei Huber =Mahurea exstipulata subsp. duckei
Mahurea linguiformis Tul. =Mahurea exstipulata subsp. exstipulata
Mahurea sororopantepuiana Steyerm. ="Mahurea exstipulata subsp. exstipulata
Mahurea tomentosa Ducke =Mahurea exstipulata subsp. duckei
Marialvaea brasiliensis Mart. =Tovomita brasiliensis
Marialvaea macrophylla Poepp. ^Tovomita macrophylla
Moronobeajenmanii war. fanshawei Maguire =Moronobeajenmanii
Moronobeajenmanii war. jenmanii =Moronobeajenmanii
Moronobea ptaritepuiana f. rosea Steyerm. =Moronobea ptaritepuiana
Moronobea riparia \ax.fimbrillata R.E. Schult. =Moronobea riparia
Moronobea riparia var. piraparanensis R.E. Schult. =Moronobea riparia
Oedematopus duidae Gleason =Clusia huberi
Oedematopus obovatus Spruce ex Planch. & Triana =Clusia obovata
Oedematopus octandrus (Poepp.) Planch. & Triana =Clusia octandra
Oedematopus parvulus Maguire =Clusia parvula
Oedematopus ptaritepuiensis Steyerm. =Clusia ptaritepuiensis
Oedematopus quadratus Maguire =Clusia obovata
Polythecandra schomburgkiana Planch. & Triana =Clusia schomburgkiana
Quapoya acuminata (Planch. & Triana) Kuntze =Clusia hammeliana
Quapoya bracteolata Sandwith =Clusia hammeliana
Quapoya panapanari Aubl. =Clusia panapanari
Quapoya peruviana (Poepp.) Kuntze =Clusia hammeliana
Quapoya peruviana var. guayanensis Maguire =Clusia hammeliana
Quapoya sipapoana Maguire =Clusia sipapoana
Renggeria comans Meisn. =Clusia comans
Renggeria montana Klotzsch [nomen nudum] =Clusia hammeliana
Rengifa acuminata Planch. & Triana ^^Clusia hammeliana
Rengifa peruviana Poepp. =Clusia hammeliana
Rengifa scandens (Aubl.) Planch. & Triana =Quapoya scandens
Rheedia acuminata (Ruiz & Pav.) Planch. & Triana =Rheedia madruno
Rheedia kappleri Eyma =Rheedia madruno
Rheedia sagotiana Engl. =Rheedia benthamiana
Symphonia coccinea (Aubl.) Oken =Moronobea coccinea
Symphonia microphylla R.E. Schult. =Symphonia globulifera
Symphonia utilissima R.E. Schult. =Symphonia globulifera
Tovomita angustata Steyerm. =Tovomita weddelliana
Tovomita cephalostigma Vesque =Tovomita umbellata
Tovomita duidae Steyerm. =Tovomita gracilipes
Tovomita guianensis Planch. & Triana, non Aubl. 1775 =Tovomita brevistaminea
Tovomitajenmanii Engl. =Tovomita brevistaminea
Tovomita myriandra Benth. =Clusia myriandra
Tovomita nigrescens Planch. & Triana =Tovomita umbellata
Tovomita richardiana Planch. & Triana [nom. illeg.] =Tovomita longifolia
Checklist of the Plants of the Guiana Shield 253
Tovomita stigmatosa Planch. & Triana =Tovomita umbellata
Tovomitopsis membranacea (Planch. & Triana) D'Arcy =Chrysochlamys membranacea
Tovomitopsis nicaraguensis Oerst., Planch. & Triana =Chrysochlamys nicaraguensis
Tovomitopsis standleyana (L.O. Williams) D'Arcy =Chrysochlamys weberbaueri
Verticillaria acuminata Ruiz & Pav. =Rheedia madruno
Vismia acuminata (Lam.) Pers. =Vismia guianensis
Vismia amazonica Ewan =Vismia gracilis
Vismia angusta Miq. =Vismia macrophylla
Vismia angustifolia Rusby =Vismia laxiflora
Vismia baccifera suhsp. ferruginea (Kunth) Ewan =Vismia guianensis
Vismia caparosa Kunth =^Vismia guianensis
Vismia cayennensis var. sessilifolia (Aubl.) M.E. Berg =Vismia sessilifolia
Vismia cayennensis var. tenuinervia M.E. Berg =Vismia tenuinervia
Vismia confertiflora auct. non Spruce ex Reichardt 1878: sensu Ewan =Vismia baccifera subsp. dealbata
Vismia dealbata Kunth =Vismia baccifera subsp. dealbata
Vismiafalcata Rusby =Vismia laxiflora
Vismiaferruginea Kunth =Vismia guianensis
Vismia glaziovii Ruhland = Vismia gracilis
Vismia guianensis var. acuminata (Lam.) M.E. Berg =Vismia guianensis
Vismia guianensis var. guianensis =Vismia guianensis
Vismia lindeniana Decne. [not in flora area]
Vismia pentagyna (Spreng.) Ewan [not in flora area]
Vismia reticulata (Poir.) Choisy = Vismia macrophylla
Vismia rufescens (Lam.) Pers. =Vismia sessilifolia
Vismia schomburgkiana Klotzsch ex M.R. Schomb. [nomen nudum]
Vismia tomentosa auct. non Ruiz & Pav. 1798 =Vismia schultesii
References
Berg, M.E. van der. 1979. Revisao das especies brasileiras do genero Rheedia L. Acta Amaz. 9(1): 43-74.
Ewan, J. 1962. Synopsis of the South American species of Vismia. Contrib. U.S. Nat. Herb. 35(5): 293-377.
Eyma, RJ. 1934. Guttiferae. In A. Pulle, Flora of Suriname 3(1): 65-118.
Keams, D.M., RE. Berry, RF. Stevens, N.L. Cuello A., J.J. Pipoly III, N.K.B. Robson, B.K. Hoist, K. Kubitzki, & A.L. Weitzman.
Clusiaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4. 1998. 248-329.
Kubitzki, K. 1978. Caraipa and Mahurea (Bonnetiaceae). In B. Maguire & collaborators, Botany of the Guayana Highland.
Mem. New York Bot. Gard. 29: 82-138.
Maguire, B. 1958. Guttiferae. In B. Maguire, J. Wurdack & collaborators. Botany of the Guayana Highland Part III. Mem. New
York Bot. Gard. 10(1): 49-61.
Maguire, B. 1961 . Guttiferae. In B. Maguire, J.J. Wurdack & collaborators. Botany of the Guayana Highland part IV. Mem. New
York Bot. Gard. 10(4): 21-32.
Maguire, B. 1964. Guttiferae subfam. Moronoboideae. In B. Maguire, J.J. Wurdack, & collaborators. Botany of the Guayana
Highland Part V. Mem. New York Bot. Gard. 10(5): 123-135.
Maguire, B. 1972. Clusiaceae - Quapoya and Renggeria. In B. Maguire & collaborators. Botany of the Guayana Highland Part
IX. Mem. New York Bot. Gard. 23: 192-195.
Pipoly, J.J. Ill & M.H.G. Gustafsson. 2002. Clusiaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M.
Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New
York Bot. Gard. 76 (2): 212-222.
Robson, N.K.B. 1987. Studies in the genus Hypericum L. (Guttiferae). 7. Section 29. Brathys (part 1). Bull. Br. Mus. Nat. Hist.
(Bot.) 16: 1-106.
Robson, N.K.B. 1990. Studies in the genus Hypericum L. (Guttiferae). 8. Sections 29. Brathys (part 2) and 30. Trigynobrathys.
Bull. Br. Mus. Nat. Hist. (Bot.) 20: 1-151.
Sandwith, N.Y 1936. Contributions to the flora of tropical America: XXVI. The genus Tovomita in British Guiana. Kew Bull.
1936:210-221.
Combretaceae
Alwan Al-Mayah, A.-R. A. & C.A. Stace. 1985. Resurrection of two Eighteenth Century names in American Combretaceae.
Nord. J. Bot. 5(5): 447-449.
Exell, A.W. 1935. Combretaceae. In A. Pulle, Flora of Suriname 3(1): 164-177.
Exell, A.W. 1953. The Combretum species of the New World. J. Linn. Soc. Lond. (Bot.) 55: 103-141.
Exell, A.W. & C.A. Stace. 1963. A revision of the genera Buchenavia and Ramatuella. Bull. Brit. Mus. (Nat. Hist.) 3: 1-46.
Exell, A.W. & C.A. Stace. 1966. Revision of the Combretaceae. Bol. Soc. Brot. (ser. 2) 40: 5-26.
Kawasaki, M.L. 2002. Combretaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 224-227.
Stace, C.A., A.-R. Alwan. 1998. Combretaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana
4: 329-352.
254 Checklist of the Plants of the Guiana Shield
COMBRETACEAE
A.-R. Alwan, C.A. Stace
Buchenavia amazonia Alwan & Stace FG?
Buchenavia congesta Ducke
Buchenaviafanshawei Exell & Maguire
Buchenavia grandis Ducke
Buchenavia guianensis (Aubl.) Alwan & Stace
Buchenavia macrophylla Eichler
Buchenavia megalophylla van Heurck & Miill. Arg.
Buchenavia nitidissima (Rich.) Alwan & Stace
Buchenavia ochroprumna Eichler
Buchenavia oxycarpa (Mart.) Eichler
Buchenavia pallidovirens Cuatrec.
Buchenavia pan'ifolia Ducke
Buchenavia pulcherrima Exell & Stace
Buchenavia reticulata Eichler
Buchenavia suaveolens Eichler
Buchenavia tetraphylla (Aubl.) R.A. Howard
Buchenavia viridiflora Ducke
Combretum assimile Eichler
Combretum brevistyhim Eichler
Combretum cacoucia Exell ex Sandwith
Combretum decandrum Jacq.
Combretumfrangulifolium Kunth
Combretum fruticosum (Loefl.) Stuntz
Combretum glabrum DC.
Combretum laurifolium Mart.
Combretum laxum Jacq.
Combretum llewelynii J.F. Macbr.
Combretum pyramidatum Desv.
Combretum rohrii Exell
Combretum rotundifolium Rich.
Combretum spinosum Bonpl.
Conocarpus erectus L. [expected in DA]
Laguncularia racemosa (L.) C.F. Gaertn.
Terminalia amazonia (J.F. Gmel.) Exell
Terminalia catappa L. [cultivated, naturalized]
Terminalia crispialata (Ducke) Alwan & Stace
Terminalia dichotoma G. Mey.
Terminalia guaiquinimae Maguire & Exell
Terminalia guyanensis Eichler
Terminalia lucida Hoffmanns, ex Mart.
Terminalia oblonga (Ruiz & Pav.) Steud.
Terminalia quintalata Maguire
Terminalia ramatuella Alwan & Stace AM
Terminalia steyermarkii Alwan & Stace BO
Terminalia tanibouca Rich. FG
Terminalia virens (Spruce ex Eichler) Alwan & Stace AM
Terminalia yapacana Maguire AM
Thiloa paraguariensis Eichler AM FG
Synonyms and Misidentifications
Buceras bucida Crantz [not in flora area] =Bucida buceras L.
Buchenavia capitata (Vahl) Eichler =Buchenavia tetraphylla
Buchenavia discolor Diels =Buchenavia ochroprumna
Buchenavia huberi Ducke =Buchenavia grandis
Buchenavia longibracteata Froes =Buchenavia congesta
Buchenavia ptariensis Steyerm. =Buchenavia tetraphylla
Buchenavia pterocarpa Exell & Stace =Buchenavia suaveolens
Buchenavia stellae Cuatrec. =Buchenavia macrophylla
Buchenavia vaupesana Cuatrec. =Buchenavia tetraphylla
Bucida buceras L. [not in flora area]
AM
AM GU
BO GU SU FG
FG
AM BO GU? SU? FG
GU SU? FG?
GU? FG
AM FG
AM
AM BO
AM GU
GU
FG
AM
AM
AM BO DA GU SU FG
AM GU FG
AM
AM
DA GU SU FG
BO
AM BO
AM BO DA GU
GU FG
AM BO DA GU FG
AM BO DA GU SU FG
AM GU?
AM BO GU SU FG
FG
AM
BO
DA
DA
GU SU FG
GU SU FG
DA GU SU FG
AM BO DA GU SU FG
GU SU FG
AM
DA GU SU FG
BO
BO DA GU SU FG
DA GU SU FG
BO GU FG
BO GU
Checklist of the Plants of the Guiana Shield 255
Biicida capitata Vahl =Buchenavia tetraphylla
Cacoucia coccinea Aubl. =Combretum cacoiicia
Chuncoa amazonia J.F. Gmel. =Terminalia amazonia
Combretum accedens Van Heurck & Miill. Arg. =Combretum laxum
Combretum aubletii DC. =Combretum rotiindifolium
Combretum brimnescens Gleason =Combrefiim laxum
Combretum elegans Kunth =Combretum rotimdifolium
Combretum ferrugineum Hoffmanns, ex Mart, [nomen nudum] =Combretum laxum
Combretum fulgens Gleason =Combretum laxum
Combretumfusiforme Gleason =Combretum glabrum
Combretum guanaiense Rusby =Combretum assimile
Combretum guianense Miq. =Combretum rotundifolium
Combretumjacquinii Griseb. =Combretum laxum
Combretum latipaniculatum Rusby =Combretum decandrum
Combretum laxum Aubl. =Combretum rotundifolium
Combretum loeflingii Eichler =Combretum fruticosum
Combretum magnificum Mart. =Combretum rotundifolium
Combretum oblongifolium Rusby =Combretum laxum
Combretum obtusifolium Rich. =Combretum laxum
Combretum phaeocarpum Mart. =Combretum pyramidatum
Combretum puberum Rich. =Combretum laxum
Combretum pulchellum Mart. ^Combretum laxum
Combretum punctatum Steud. =Combretum rotundifolium
Combretum sambuense Pittier =Combretum assimile
Combretum sprucei Eichler =Thiloa paraguariensis
Combretum terminalioides Steud. =Combretum laxum
Conocarpus racemosa L. =Laguncularia racemosa
Gaurafruticosa Loefl. =Combretum fruticosum
Gimbernatea oblonga Ruiz & Pav. =Terminalia oblonga
Gimbernatea obovata Ruiz & Pav. =Terminalia amazonia
Pamea guianensis Aubl. =Buchenavia guianensis
Ramatuela argentea Kunth =Terminalia ramatuella
Ramatuela crispialata Ducke =Terminalia crispialata
Ramatuela crispialata var. obtusa Maguire =Terminalia crispialata
Ramatuela latifolia Maguire =Terminalia virens
Ramatuela maguirei Exell & Stace =Terminalia virens
Ramatuela obtusa (Maguire) Exell & Stace =Terminalia crispialata
Ramatuela virens Spruce ex Eichler =Terminalia virens
Tanibouca guianensis Aubl. =Terminalia dichotoma
Terminaliafirma Mart. =Terminalia lucida
Terminalia nitidissima Rich. =Buchenavia nitidissima
Terminalia obovata (Ruiz & Pav.) Steud. =Terminalia amazonia
Terminalia odontoptera Van Heurck & Miill. Arg. =Terminalia amazonia
Terminalia opacifolia Maguire & Exell =Terminalia guaiquinimae
Terminalia oxycarpa Mart. =Buchenavia oxycarpa
Terminalia pamea DC. =Buchenavia guianensis
Terminalia virgata Maguire & Exell =Terminalia guaiquinimae
Thiloa inundata Ducke =Thiloa paraguariensis
I
COMPOSITAE
VA. Funk, H. Robinson, J.F. Pruski
Acanthospermum australe (Loefl) Kuntze
Acanthospermum hispidum DC.
Achnopogon steyermarkii Aristeg.
Achnopogon virgatus Maguire, Steyerm. & Wurdack
Achyroclineflaccida DC.
Achyrocline satureioides (Lam.) DC.
Achyrocline vargasiana DC.
Acmella brachyglossa Cass.
Acmella ciliata (Kunth) Cass.
Acmella radicans (Jacq.) R.K. Jansen
Acmella radicans var. debilis (Kunth) R.K. Jansen
AM BO GU SU FG
BO GU SU FG
BO
BO
GU
AM BO GU
AM BO GU
SU FG
AM BO GU FG?
AM
AM
256 Checklist of the Plants of the Guiana Shield
Acmella repens (Walter) Rich, ex Pers.
Acmella iiliginosa (Sw.) Cass.
Ageratina ibaguemis (Sch. Bip. ex Hieron.) R.M. King & H. Rob.
Ageratina roraimemis (N.E. Br.) R.M. King & H. Rob. [Mt. Roraima]
Ageratiim ballotifoUum (Maguire, Steyerm. & Wurdack) R.M. King & H. Rob.
Ageratitm conyzoides L. [introduced]
Ageratiim houstonianum Mill, [widely cultivated, possibly naturalized]
Ageratum scorpioideum Baker
Ambrosia crithmifolia DC.
Ambrosia microcephala DC.
Ambrosia peruviana Willd. [expected in the Guianas]
Austroeupatoriiim imdifolium (Kunth) R.M. King & H. Rob.
Ayapana amygdalina (Lam.) R.M. King & H. Rob.
Ayapana trinitensis (Kuntze) R.M. King & H. Rob.
Ayapana triplinervis (Vahl) R.M. King & H. Rob.
Baccharis brachylaenoides DC.
Baccharis brachylaenoides var. brachylaenoides
Baccharis brachylaenoides var. ligiistrina (DC.) Maguire & Wurdack
Baccharis densa (N.E. Br.) V.M. Badillo
Baccharis Hindi i DC.
Baccharis nitida (Ruiz & Pav.) Pers.
Baccharis rufescens Spreng.
Baccharis schomburgkii Baker
Baccharis trinervis (Lam.) Pers.
Baccharis varians Gardner
Baltimora geminata (Brandegee) Stuessy
Barrosoa metensis (B.L. Rob.) R.M. King & H. Rob.
Bidens alba (L.) DC.
Bidens alba var. radiata (Sch. Bip.) R.E. Ballard
Bidens cynapiifolia Kunth
Bidens pilosa L.
Bidens riparia Kunth
Bidens riparia var. refracta (Brandegee) O.E. Schulz
Blainvillea dichotoma (Murray) Cass.
Brickellia diffusa (Vahl) A. Gray
Calea abelioides S.F. Blake AM
Calea berteriana DC.
Calea caleoides (DC.) H. Rob. AM
Calea camani Maguire & K.D. Phelps AM
Calea divaricata Benth.
Calea esposi Maguire & K.D. Phelps AM
Calea kunhardtii Maguire AM
Calea linearifolia Maguire & Wurdack AM
Calea liicida Maguire & Wurdack
Calea lucidivenia Gleason & S.F. Blake
Calea lucidivenia var. cardonae (Maguire & Wurdack) Pruski
Calea lucidivenia var. lucidivenia
Calea lucidivenia var. orientalis (Maguire & Wurdack) Pruski
Calea lucidivenia var. punctata Maguire & Wurdack
Calea nana Maguire AM
Calea neblinensis (Maguire & Wurdack) Pruski AM
Calea nov. sp. 1 Pruski AM
Calea nov. sp. 2 Pruski AM
Calea nov. sp. 3 Pruski AM
Calea nov. sp. 4 Pruski
Calea nov. sp. 5 Pruski AM
Calea oliveri B.L. Rob. & Greenm.
Calea orbiculata Maguire & Aristeg. AM
Calea phelpsiae Lasser & Maguire AM
Calea politii Maguire AM
Calea punctata Maguire & Wurdack AM
Calea sipapoana Maguire AM
Calea solidaginea Kunth
Calea solidaginea subsp. deltophylla (R.S. Cowan) Pruski
FG
BO GU SU FG
BO
BO
BO
AM BO
BO
DA
DA
GU
GU
GU
su
SU
su
FG
FG
AM BO GU
AM BO
DA
GU su FG
FG
AM BO GU
AM BO GU
AM GU
AM BO
80
GU
GU
GU
AM BO GU
AM BO GU
BO GU su
BO GU
BO
GU
GU
FG
FG
AM BO DA GU su FG
BO DA GU
GU
GU
su FG
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
GU
GU
GU
GU
GU
GU
GU
FG
SU
BO
BO
GU
GU
SU
SU
FG
FG
AM
AM BO
AM
AM BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
AM
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM
Checklist of the Plants of the Guiana Shield 257
Galea sublantanoides V.M. Badillo
Galea tolimana Hieron.
Galea thcephala Maguire
Gentrathenim punctatiim Cass. DA GU FG
Ghaptalia integerrima (Veil.) Burkart GU
Ghaptalia nutans {L.)?o\. GU SU
Ghimantaea acopanensis Steyerm.
Ghimantaea cinerea (Gleason & S.F. Blake) Maguire, Steyerm. & Wurdack
Ghimantaea cinerea subsp. cinerea
Ghimantaea cinerea subsp. similis (Maguire, Steyerm. & Wurdack) Pruski &
O. Huber
Ghimantaea eriocephala Maguire, Steyerm. & Wurdack
Ghimantaea espeletoidea Maguire, Steyerm. & Wurdack
Ghimantaea huberi Steyerm.
Ghimantaea humilis Maguire, Steyerm. & Wurdack
Ghimantaea lanocaulis Maguire, Steyerm. & Wurdack
Ghimantaea lanocaulis subsp. lanocaulis
Ghimantaea lanocaulis subsp. murisipanensis Pruski & O. Huber
Ghimantaea mirabilis Maguire, Steyerm. & Wurdack
Ghimantaea rupicola Maguire, Steyerm. & Wurdack
Ghionolaena latifolia (Benth.) Baker
Ghromolaena bathyphlebia (B.L. Rob.) R.M. King & H. Rob.
Ghromolaena corymbosa (Aubl.) R.M. King & H. Rob. FG
Ghromolaena ivifolia (L.) R.M. King & H. Rob. GU
Ghromolaena laevigata (Lam.) R.M. King & H. Rob. GU SU FG
Ghromolaena maximilianii (Schrad. ex DC.) R.M. King & H. Rob. FG
C/2rowo/fle«a oJorato (L.) R.M. King & H. Rob. DA GU SU FG
Ghromolaena pharcidodes {B.L.Roh.) KM. King ScW-Kob. GU
Ghromolaena squalida{T>C.) R.M. Yimg&W.Roh. GU SU
Ghromolaena ternicapitulata Pruski
Ghromolaena thurnii (B.L. Rob.) R.M. King & H. Rob. BO GU
Ghromolaena tyleri (B.L. Rob.) R.M. King & H. Rob. AM
Glibadium armanii (Balbis) Sch. Bip. ex O.K. Schulz
Glibadium surinamense L.
Glibadium sylvestre (Aubl.) Baill.
Gondylidium iresinoides (Kunth) R.M. King & H. Rob.
Gonyza bonariensis (L.) Cronquist
Gonyza canadensis (L.) Cronquist [naturalized]
Gonyza laevigata (Rich.) Pruski
Gonyza primulaefolia (Lam.) Cuatrec. & Lourteig
Gosmos caudatus Kunth [widely cultivated, probably escaping]
Gyanthillium cinereum (L.) H. Rob. [naturalized]
Gyrtocymura scorpioides (Lam.) H. Rob.
Dasyphyllum vepreculatum (D. Don) Cabrera
Delilia biflora (L.) Kuntze
Duidaea marahuacensis Steyerm.
Duidaea pinifolia S.F. Blake
Duidaea rubriceps S.F. Blake
Duidaea tatei S.F. Blake
Eclipta prostrata (L.) L.
Egletesflorida Shinners
Egletes prostrata (Sw.) Kuntze
Egletes viscosa (L.) Less. BO
Elaphandra moriana Pruski FG
Elephantopus hirtiflorus DC.
Elephantopus mollis Kunth
Eleutheranthera ruderalis (Sw.) Sch. Bip.
Emilia coccinea (Sims) G. Don [African: cultivated, escaping]
Emiliafosbergii Nicolson
Emilia sonchifolia (L.) DC. ex Wight BO DA
Erechtites hieracifolius (L.) Raf. ex DC.
Erechtites valerianifolius (Link ex Spreng.) DC.
Erigeron karwinskianus DC.
Fleischmannia microstemon (Cass.) R.M. King & H. Rob. AM BO SU FG
GU SU FG
AM BO GU SU FG
AM
BO
GU
GU
SU FG
BO GU SU
SU
FG
FG
FG
BO GU SU
DA GU SU FG
BO DA GU SU FG
BO GU SU FG
BO
GU
AM
AM
AM
AM
AM BO DA GU SU FG
BO DA GU
GU
BO
AM BO
BO
AM BO
AM BO
AM BO
BO
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU SU FG
GU FG
258 Checklist of the Plants of the Guiana Shield
Gamochaeta amehcana (Mill.) Wedd.
Gamochaeta simplicicaiilis (Willd. ex Spreng.) Cabrera
Glossahon bilabiatiim (Maguire) Pruski
Glossarion rJwdanthum Maguire & Wurdack
Gnaphaliiim elegans Kunth
Gnaphalium polycaiilon Pers. [Mt. Roraima; Sierra de la Neblina, Brazil]
Gochnatia oUgocephala (Gardner) Cabrera [expected in GU]
Gongylolepis benthamiana R.H. Schomb.
Gongylolepis bracteata Maguire
Gongylolepis cortesii (S. Diaz) Pruski & S. Diaz [expected in AM]
Gongylolepis erioclada S.F. Blake
Gongylolepis fruticosa Maguire, Steyerm. & Wurdack
Gongylolepis glaberrima S.F. Blake
Gongylolepis hiiachamacari Maguire
Gongylolepis jauaensis (Aristeg., Maguire & Steyerm.) V.M. Badillo
Gongylolepis martiana (Baker) Steyerm. & Cuatrec.
Gongylolepis oblanceolata Pruski
Gongylolepis paniciilata Maguire & K.D. Phelps
Gongylolepis paniana Maguire
Gongylolepis pedunciilata Maguire
Gongylolepis yapacana Maguire
Guayania bulbosa (Aristeg.) R.M. King & H. Rob.
Guayania cerasifolia (Sch. Bip. ex Baker) R.M. King & H. Rob.
Guayania crassicaulis (Steyerm.) R.M. King & H. Rob.
Guayania penninervato (Wurdack) R.M. King & H. Rob.
Guayania roupalifolia (B.L. Rob.) R.M. King & H. Rob.
Guayania yaviana (Lasser & Maguire) R.M. King & H. Rob.
Hebeclinium macwphyllum (L.) DC.
Helerocondylus vitalbae (DC.) R.M. King & H. Rob.
Huberopappus maigualidoe Pruski
Ichthyofhere davidsei H. Rob.
Ichthyothere granvillei H. Rob.
Ichthyofhere terminalis (Spreng.) S.F. Blake
Imeria memorabilis (Maguire & Wurdack) R.M. King & H. Rob.
Imeria memorabilis var. densesericea Pruski [SW of Neblina]
Imeria memorabilis var. memorabilis
Koanophyllonfuscum (N.E. Br.) R.M. King & H. Rob.
Koanophyllon tatei (B.L. Rob.) R.M. King & H. Rob.
Lepidaploa bolivarensis (V.M. Badillo) H. Rob.
Lepidaploa clvysotricha (Alexander) H. Rob.
Lepidaploa cotoneasJer (Willd. ex Spreng.) H. Rob.
Lepidaploa ehretiifolia (Benth.) H. Rob.
Lepidaploa gracilis (Kunth) H. Rob.
Lepidaploa imeriensis (V.M. Badillo) Pruski
Lepidaploa pari (V.M. Badillo) H. Rob. [expected in the Guianas]
Lepidaploa remotiflora (Rich.) H. Rob.
Lepidaploa salzmannii (DC.) H. Rob. [Mt. Roraima]
Lessingianthus morilloi (V.M. Badillo) H. Rob.
Lycoseris triplinervia Less.
Melanthera nivea (L.) Small
Mikania amazonica Baker
Mikania banisteriae DC.
Mikania boomii Pruski
Mikania chaetoloba Pruski
Mikania chlorolepis Baker
Mikania congesta DC.
Mikania cordifolia (L. f.) Willd.
Mikania duidensis B.L. Rob.
Mikania gleasonii B.L. Rob.
Mikania guaco Bonpl.
Mikania hookeriana DC.
Mikania lindleyana DC.
Mikania lucida S.F. Blake [Mt. Roraima]
Mikania marahuacensis Steyerm. & Maguire
AM
BO GU
BO
BO GU
BO
BO GU
BO
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO DA
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
AM BO
BO
AM BO
AM BO
BO
BO
AM BO
AM BO
BO
AM BO
AM
DA
DA
DA
GU
GU
GU
GU
GU
GU
SU
SU
SU
SU
FG
FG
FG
FG
GU SU
GU
GU
GU
GU
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU SU FG
GU SU
Checklist of the Plants of the Guiana Shield 259
Mikania mazaruniensis W.C. Holmes & McDaniel
Mikania michelangeliana Steyerm.
Mikania micrantha Kunth
Mikania microptera DC.
Mikania neblinensis Aristeg.
Mikania nigropunctulata Hieron.
Mikania pannosa Baker
Mikania parviflora (Aubl.) H. Karst.
Mikania psilostachya DC.
Mikania rondonensis V.M. Badillo
Mikania simpsonii W.C. Holmes & McDaniel
Mikania solidinervia V.M. Badillo
Mikania sprucei Baker
Mikania trinitaria DC.
Mikania vitifolia DC.
Neblinaea promontohorum Maguire & Wurdack
Neurolaena lobata (L.) R. Br.
Oritrophium marahuacense Steyerm. & Maguire
Orthopappus angustifolius (Sw.) Gleason
Oyedaea scaberrima (Benth.) S.F. Blake
Oyedaea tepuiana (V.M. Badillo) Pruski [expected in the Guianas]
Oyedaea wurdackii Pruski
Pacourina edulis Aubl.
Parthenium hysterophorus L.
Pedis ciliaris L.
Pectis elongata Kunth
Pectis elongata var. elongata
Pectis elongata var. floribiinda (A. Rich.) Keil
Pectis humifusa Sw.
Pectis pygmaea Kunth
Pectis venezuelensis Steyerm. [expected in the Guianas]
Pentacaliafreemanii (Britton & Greenm.) Cuatrec.
Pentacalia neblinensis Pruski
Pentacalia phelpsiae (Cuatrec.) Cuatrec.
Pentacalia ptariana (Cuatrec.) Cuatrec.
Pentacalia yapacana (Aristeg.) Cuatrec.
Piptocarpha auyantepuiensis Aristeg.
Piptocarphajauaensis Aristeg. & Steyerm.
Piptocarpha leprosa (Less.) Baker
Piptocarpha opaca (Benth.) Baker
Piptocarpha opaca subsp. opaca
Piptocarpha poeppigiana (DC.) Baker
Piptocarpha polycephala Baker
Piptocarpha triflora (Aubl.) Benn. ex Baker
Piptocoma areolata (Wurdack) Pruski
Piptocoma roraimensis (Steyerm.) Pruski
Piptocoma schomburgkii (Sch. Bip.) Pruski
Piptocoma spruceana (Benth.) Pruski
Pluchea carolinensis (Jacq.) G. Don [probably cultivated only]
Pluchea odorata (L.) Cass.
Pluchea sagittalis (Lam.) Cabrera [expected in the Guianas]
Porophyllum ruderale (Jacq.) Cass.
Praxelis asperulacea (Baker) R.M. King & H. Rob.
Praxelis diffusa (Rich.) Pruski
Pseudelephantopus spicatus (Juss.) Rohr
Pseudogynoxys chenopodioides (Kunth) Cabrera [expected in the Guianas]
Pterocaulon alopecuroides (Lam.) DC.
Quelchia bracteata Maguire, Steyerm. & Wurdack
Quelchia cardonae Steyerm.
Quelchia conferta N.E. Br.
Quelchia eriocaulis Maguire, Steyerm. & Wurdack
Quelchia x grandifolia Maguire, Steyerm. & Wurdack
Riencourtia latifolia Gardner
Riencourtia oblongifolia Gardner
GU
AM
AM BO DA GU su FG
DA GU su FG
AM
AM BO GU su
AM BO GU
AM BO GU su FG
AM BO GU su FG
AM BO GU
AM
AM
AM BO GU
AM BO DA GU su FG
AM BO DA GU su
AM
BO GU su FG
AM
AM BO GU su FG
AM BO GU
AM BO
AM
GU su FG
GU su FG
BO
AM BO GU su FG
GU su FG
FG
DA GU
BO
AM BO
BO GU
AM
AM
BO
BO
BO
BO
BO GU
AM GU
AM
AM BO
BO
DA GU
GU
AM BO DA GU su FG
AM
BO GU
AM BO GU su FG
AM
BO
GU
GU
su
su
BO GU su FG
AM GU
AM BO GU su FG
AM
BO DA
GU su FG
BO GU su
BO
BO
BO GU
BO
BO
AM BO GU
SU
260 Checklist of the Plants of the Guiana Shield
Riencourtia pedunculosa {Rich.) Pruski AM BO
Rolandrafruticosa (L.) Kuntze AM BO DA
Salmea scandem (L.) DC. [expected in flora area]
Sciadocephala pakaraimae (Maguire & Wurdack) R.M. King & H. Rob.
[expected in BO]
Siapaea liesneri Pruski AM
Sigesbeckia orientalis L. [naturalized]
Sorichus asper (L.) Hill BO
Sphagneticola brachycarpa (Baker) Pruski AM BO
Sphagneticola thlobata (L.) Pruski AM BO
Spilanthes nervosa Chodat AM
Spilanthes urens Jacq. AM DA
Stenocephalum apiculatum (Mart.) Sch. Bip. BO
Stenopadus campestris Maguire & Wurdack AM
Stenopadus chimantensis Maguire, Steyerm. & Wurdack BO
Stenopadus colveei (Steyerm.) Pruski BO
Stenopadus connellii (N.E. Br.) S.F. Blake' BO GU
Stenopadus cucu/latus Maguire & Wurdack AM
Stenopadus huachamacari Maguire AM
Stenopadus jauensis Aristeg. BO
Stenopadus kunhardtii Maguire AM
Stenopadus megacephalus Pruski GU
Stenopadus obconicus Maguire & Wurdack AM
Stenopadus sericeus Maguire & Aristeg.
Stenopadus talaumifolius S.F. Blake
Stifftia cayennensis H. Rob. & Kahn
Stomatochaeta acuminata Pruski
Stomatochaeta condensata (Baker) Maguire & Wurdack
Stomatochaeta crassifolia (S.F. Blake) Maguire & Wurdack
Stomatochaeta cylindrica Maguire & Wurdack
Stomatochaeta cymbifolia (S.F. Blake) Maguire & Wurdack
Stomatochaeta steyermarkii Aristeg.
Struchium sparganophorum (L.) Kuntze
Symphyotrichum laeve (L.) A. Love & D. Love [cultivated, naturalized]
Synedrella nodiflora (L.) Gaertn.
Tagetes erecta L. [cultivated, naturalized]
Tessaria integrifolia Ruiz & Pav.
Trichogonia arguta (Kunth) Benth. & Hook. f. ex Klatt [Mt. Roraima]
Trichospira verticillata (L.) S.F. Blake
Tridax procumbens L.
Tuberculocarpus ruber (Aristeg.) Pruski
Tyleropappus dichotomus Greenm.
Unxia camphorata L. f.
Unxia suffruticosa (Baker) Stuessy
Verbesina angusta Maguire, Steyerm. & Wurdack
Verbesina columbiana B.L. Rob. [expected in the Guianas]
Verbesina guianensis Baker
Verbesina ligulata (Maguire & Wurdack) Pruski [expected in the Guianas.
Verbesina pilosa Maguire & Wurdack
Vernonanthura brasiliana (L.) H. Rob.
Wedelia ambigens S.F. Blake
Wedelia calycina Rich.
Wedelia villosa Gardner
Wulffia baccata (L.) Kuntze
Wulffia rubens Alexander
Xiphochaeta aquatica Poepp.
Youngiajaponica (L.) DC. [introduced]
Synonyms and Misidentifications
Acanthospermum brasilum Schrank =Acanthospermum australe
Acanthospermum humile var. hispidum (DC.) Kuntze =Acanthospermum hispidum
Achnopogon quelchioides Aristeg. =Achnopogon steyermarkii
Achyrocline satureioides var. vargasianum (DC.) Baker =Achyrocline vargasiana
Acmella brasiliensis Spreng. =Sphagneticola trilobata
GU su FG
GU su FG
GU?
GU
GU
GU
GU
GU su FG
BO GU
AM BO
BO
BO
GU
GU
FG
AM
BO
BO
BO
AM DA GU
GU
SU FG
FG
BO DA GU
GU
su
su
FG
BO DA
BO
BO DA GU FG
BO GU FG
AM
AM
AM BO
BO
BO
BO
BO
GU
GU
su FG
AM BO
BO
AM BO
BO
GU
GU
su FG
BO GU
GU
FG
AM BO DA GU
GU
su FG
AM BO DA GU
GU
su FG
Checklist of the Plants of the Guiana Shield 261
Acmella oppositifolia var. oppositifolia =Acmella repens
Acmella radicans var. radicans [not in flora area]
Adenostemma pakaraimae Maguire & Wurdack =Sciadocephala pakaraimae
Ageratum guianense Aubl. =Hebeclimitm macrophyllum
Alomia ballotifolia Maguire, Steyerm. & Wurdack =Ageratum ballotifolium
Ambrosia ciimanensis Kunth =Ambrosia peruviana
Ampherephis aristata Kunth =Centratherum punctatum
Ampherephis mutica Kunth =Centratherum punctatum
Aspilia bolivarana V.M. Badillo =Wulffia baccata
Aspilia rubra Aristeg. =Tuberculocarpus ruber
Aster laevis L. =Symphyotrichum laeve
Augusta benthamiana (R.H. Schomb.) Kuntze =Gongylolepis benthamiana
Augusta condensata (Baker) Kuntze =Stomatochaeta condensata
Augusta martiana (Baker) Kuntze =Gongylolepis martiana
Baccharis brasiliana L. =Vernonanthura brasiliana
Baccharis daphnoides subsp. schomburgkii (Baker) Malag. =Baccharis schomburgkii
Baccharis erioptera Benth. =Pterocaulon alopecuroides
Baccharis ligustrina DC. =Baccharis brachylaenoides var. ligustrina
Baccharis maguireana Malag. =Baccharis brachylaenoides var. brachylaenoides
Baccharis meridensis Steyerm. =Baccharis brachylaenoides
Baccharis ptariensis Steyerm. =Baccharis brachylaenoides var. ligustrina
Baccharis rhexioides Kunth =Baccharis trinervis
Baccharis roraimae M.R. Schomb. ^Baccharis brachylaenoides
Baccharis rufescens var. varians (Gardner) Baker =Baccharis varians
Baccharis schomburgkii subsp. occidentalis Steyerm. & Maguire =Baccharis schomburgkii
Baccharis trinervis var. rhexioides (Kunth) Baker =Baccharis trinervis
Baccharis trinitensis Kuntze =Ayapana trinitensis
Baccharis venulosa DC. =Baccharis brachylaenoides
Baccharis venulosa var. cuspidibracteata Steyerm. =Baccharis brachylaenoides var. brachylaenoides
Baccharis vitis-idaea Oliv. ex Thum, non Kuntze =Baccharis brachylaenoides var. ligustrina
Baccharis wurdackiana Malag. =Baccharis brachylaenoides var. ligustrina
Baccharoides brachylepis (Sch. Bip. ex Baker) Kuntze =Centratherum punctatum
Baccharoides muticum (Kunth) Kuntze =Centratherum punctatum
Baccharoides punctatum (Cass.) Kuntze =Centratherum punctatum
Baillieria aspera Aubl. =Clibadium surinamense
Baillieria barbascoYMnXh. =Clibadium sylvestre
Baillieria sylvestris Aubl. =Clibadium sylvestre
Baltimora ovata Rusby =Baltimora geminata
Bidens bipinnata var. cynapiifolia (Kunth) M. Gomez =Bidens cynapiifolia
Bidens caudata (Kunth) Sch. Bip. =Cosmos caudatus
Bidens leucantha var. pilosa (L.) Griseb. =Bidens pilosa
Bidens nivea L. =Melanthera nivea
Bidens pilosa f. indivisa Sherff =Bidens pilosa
Bidens verticillata L. =Trichospira verticillata
Buphthalmum strigosum Spreng. =Sphagneticola trilobata
Cacalia cinerea (L.) Kuntze =Cyanthillium cinereum
Cacalia cordifolia L. f. =Mikania cordifolia
Cacalia porophyllum L. =Porophyllum ruderale
Cacalia ruderalis (Jacq.) Sw. =Porophyllum ruderale
Cacalia sonchifolia L. =Emilia sonchifolia
Calea aspera Jacq. =Melanthera nivea
Calea cardonae Maguire & Wurdack =Calea lucidivenia var. cardonae
Calea cardonae subsp. neblinensis Maguire & Wurdack ^Calea neblinensis
Calea cardonae var. orientalis Maguire & Wurdack =Calea lucidivenia var. orientalis
Calea clematidifolia Steyerm. =Calea solidaginea subsp. deltophylla
Calea coUina Klatt =Calea berteriana
Calea congesta Rusby =Calea solidaginea
Calea deltophylla R.S. Cowan =Calea solidaginea subsp. deltophylla
Calea divaricata var. diffusa Maguire & Wurdack =-Calea divaricata
Calea herbert-smithii Rusby =Calea solidaginea
Calea holtonii Rusby =Calea berteriana
Calea huberana V.M. Badillo =Calea tolimana
Calea lobata (L.) Sw. =Neurolaena lobata
Calea lucidivenia f. lanceolata Steyerm. =Calea lucidivenia
262 Checklist of the Plants of the Guiana Shield
Galea luetzelburgii Suess. =Calea oliveri
Galea membranacea Maguire & Wurdack =Galea lucidivenia
Galea oliveri var. taeniotricha Steyerm. =Galea oliveri
Galea ovalis subsp. sabanensis Cuatrec. =Galea tolimana
Galea perforata Klatt ^Galea solidaginea
Galea prunifolia Kunth [not in flora area]
Galea scandens Griseb. =Galea solidaginea
Galea suffruticosa Maguire & Wurdack =Galea divaricata
Galea suriani Cass. =Neurolaena lobata
Galea ternifolia Oliv. ex Thum ^Galea oliveri
GardonaeajauaensisAhstQg., Maguire & Steyerm. =Gongylolepis jauaensis
Garphobolus latifolius Sch. Bip. =Pip1ocarpha opaca subsp. opaca
Gentratherum brachylepis Sch. Bip. ex Baker =Gentratherum punctatum
Gentratherum muticum (Kunth) Less. =Gentratheriim punctatum
Gentratherum punctatum var. parviflorum Baker =Gentratherum punctatum
Gentrospermum xanthioides Kunth =Acanthospermum australe
Geratocephalus acmelia var. depauperata Runtze =AcmeUa uliginosa
Geratocephalus urens (Jacq.) Kuntze =Spilanthes urens
Ghaptalia integrifolia (Cass.) Baker =Ghaptalia integerrima
Ghatiakella platyglossa Cass. =Wulffia baccata
Ghatiakella stenoglossa Cass. =Wulffia baccata
Ghimantaea cinerea f. glabra Steyerm. =Ghimantaea cinerea
Ghimantaea similis Maguire, Steyerm. & Wurdack =Ghimantaea cinerea var. similis
Ghionolaena breweri Steyerm. & Maguire =Ghionolaena latifolia
Ghuquiraga vepreculata D. Don =Dasyphyllum vepreculatum
Ghylodia sarmentosa Rich, ex Cass. =Wulffia baccata
Glibadium asperum (Aubl.) DC. =Glibadium surinamense
Glibadium barbasco (Kunth) DC. =Glibadium sylvestre
Glibadium caracasanum DC. =Glibadium surinamense
Glibadium pedunculosum (Rich.) DC. =Riencourtia pedunculosa
Glibadium rotundifolium DC. =Glibadium armanii
Glibadium schomburgkii Sch. Bip. ex M.R. Schomb. =Glibadium sylvestre
Glibadium surinamense var. asperum (Aubl.) Baker =Glibadium surinamense
Glibadium surinamense var. macrophyllum Steyerm. =Glibadium surinamense
Glibadium vargasii DC. =Glibadium sylvestre
Goleosanthus dijfusus (Vahl) Kuntze =Brickellia diffusa
Gomplaya trilobata (L.) Strother =Sphagneticola trilobata
Gonoclinium cerasifolium Sch. Bip. ex Baker =Guayania cerasifolia
Gonyza alopecuroides Lam. =Pterocaulon alopecuroides
Gonyza calophylla Willd. ex Sch. Bip. =Gochnatia oligocephala
Gonyza cayennensis Spreng. [name of uncertain application]
Gonyza chilensis Spreng. =Gonyza primulaefolia
Gonyza cinerea L. =Gyanthillium cinereum
Gonyza cotoneaster Willd. ex Spreng. ^Lepidaploa cotoneaster
Gonyza lobata L. =Neurolaena lobata
Gonyza riparia Kunth ^Tessaria integrifolia
Gonyza sagittalis Lam. =Pluchea sagittalis
Gonyza scorpioides Lam. =Gyrtocymura scorpioides
Gonyza trinervis Lam. =Baccharis trinervis
Goreopsis baccata L. =Wulffia baccata
Gotula viscosa L. =Egletes viscosa
Dermatophyllum schomburgkii Sch. Bip. ex M.R. Schomb. =Stomatochaeta condensata
Diazeuxis latifolia D. Don =Lycoseris triplinervia
Echinocephalum discoideum Baker =Melanthera nivea
Echinops fruticosus L. =Rolandrafruticosa
Echinops nodiflorus Lam. =Rolandrafruticosa
Eclipta alba (L.) Hassk. =Eclipta prostrata
Eclipta erecta L. [nom. illeg. superfl.] =Eclipta prostrata
Egletes commixta Shinners ^Egletes florida
Elephantopus angustifolius Sw. =Orthopappus angustifolius
Elephantopus carolinianus Willd. [not in flora area]
Elephantopus carolinianus var. mollis (Kunth) Beurl. =Elephantopus mollis
Elephantopus decemsquamatus V.M. Badillo =Elephantopus hirtiflorus
Elephantopus martii Graham ^Elephantopus mollis
Checklist of the Plants of the Guiana Shield 263
Elephantopus nudiflonis Willd. =Orthopappus angustifolius
Elephantopus pilosus Philipson =Elephantopus mollis
Elephantopus quadriflora Less. =Orthopappus angustifolius
Elephantopus scaber var. tomentosus Sch. Bip ex Baker =Elephantopus mollis
Elephantopus spicatus B. Juss. =Pseudelephantopus spicatus
Elephantopus tomentosus L. =Elephantopus mollis
Elephantopus virgatus Ham. [name of uncertain application]
Emiliajavanica (Burm.) C.B. Rob. [not in flora area]
Emilia sagittata (Vahl) DC. [not in flora area]
Erigeron bonariensis L. =Conyza bonahensis
Erigeron canadensis L. =Conyza canadensis
Erigeron chilensis (Spreng.) G. Don =Conyza primulaefolia
Erigeron laevigatus Rich. =Conyza laevigata
Erigeron mucronatus DC. =Erigeron karwinskianus
Ethulia sparganophora L. =Struchium sparganophorum
Eupator,
Eupatori
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator,
Eupator.
Eupator
Eupator,
Eupator,
Eupator.
Eupator.
Eupator.
Eupator.
Eupator.
Eupator.
Eupator
Eupator
Eupator.
Eupator
Eupator
Eupator.
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupator
Eupatori
Eupator
Eupaton
Eupatori
Eupatori
Eupator,
um amarum Vahl =Mikania parviflora
um amygdalinum Lam. =Ayapana amygdalina
um angulicaule Lasser & Maguire =Guayania yaviana
um asperulaceum Baker =Praxelis asperulacea
um bathyphlebium B.L. Rob. =Chromolaena bathyphlebia
um bulbosum Aristeg. =Guayania bulbosa
um cerasifolium Sch. Bip. ex Baker =Guayania cerasifolia
um conyzoides Vahl =Chromolaena odorata
um corymbosum Aubl. =Chromolaena corymbosa
um crassicaule Steyerm. =Guayania crassicaulis
um denticulatum Vahl =Mikania cordifolia Mikania micrantha
um dichotomum Sch. Bip. =Chromolaena odorata
um dijfusum Vahl =Brickellia diffusa
um duidense V.M. Badillo =Koanophyllonfuscum
umfuscum N.E. Br. =Koanophyllonfuscum
um ibaguense Sch. Bip. ex Hieron. =Ageratina ibaguensis
um ibaguense var. indiscretum Maguire, Steyerm. & Wurdack =Ageratina ibaguensis
um inulifolium Kunth =Austroeupatorium inulifolium
um iresinoides Kunth =Condylidium iresinoides
um ivifolium L. =Chromolaena ivifolia
um ivifolium var. gracillima Baker =Chromolaena ivifolia
um Ixodes Benth. =Ayapana amygdalina
um kaniaiense V.M. Badillo =Praxelis asperulacea
um kavanayense V.M. Badillo =Koanophyllonfuscum
um laevigatum Lam. =Chromolaena laevigata
um macrophyllum L. =Hebeclinium macrophyllum
um martiusii DC. =Chromolaena squalida
um memorabile Maguire & Wurdack =Imeria memorabilis
um metense B.L. Rob. =Barrosoa metensis
um microstemon Cass. =Fleischmannia microstemon
um microstemon var. albiflorum Kuntze -Fleischmannia microstemon
um microstemon var. lilacinum Kuntze =Fleischmannia microstemon
um morichalanum Aristeg. ^Barrosoa metensis
um obscurum DC. =Chromolaena ivifolia
um odoratum L. =Chromolaena odorata
um parviflorum Aubl. =Mikania parviflora
um pauciflorum Kunth =Praxelis diffusa
um penninervatum Wurdack =Guayania penninervata
um pharcidodes B.L. Rob. =Chromolaena pharcidodes
um pharcidodes var. pentanthum S.F. Blake =Chromolaena pharcidodes
um pharcidodes var. sororopanensis Steyerm. =^Chromolaena pharcidodes
um psiadiifolium DC. =Chromolaena laevigata
um ptaretepuiense V.M. Badillo ^^Ageratina ibaguensis
um roraimensis N.E. Br. =Ageratina roraimensis
um roraimensis f. albisetum Steyerm. =Ageratina roraimensis
um roupalifolium B.L. Rob. =Guayania roupalifolia
um roupalifolium f. latifolium Steyerm. =Guayania roupalifolia
um roupalifolium var. subintegrifolium Steyerm. =Guayania roupalifolia
um schomburgkii Benth. =Baccharis brachylaenoides
\
264 Checklist of the Plants of the Guiana Shield
Eiipatorium sieberianiim DC. =Condylidiiim iresinoides
Eupatorium sqiialidum DC. =Chwmolaena squalida
Eupatorium sqiialidum var. siibvehitiniim (DC.) Baker =Chromolaena squalida
Eupatorium squiresii Rusby ex B.L. Rob. =Ayapana trinilensis
Eupatorium subvelutinum DC. =Chromolaena squalida
Eupatorium surinamense Sch. Bip. ex Miq. =Ayapana trinitensis
Eupatorium tatei B.L. Rob. =Koanophyllon tatei
Eupatorium tepuianum Steyerm. =Guayania roupalifolia
Eupatorium thurnii B.L. Rob. =Chromolaena thurnii
Eupatorium triflorum Aubl. ^Piptocarpha triflora
Eupatorium trinitense (Kuntze) Rusby & B.L. Rob. =Ayapana trinitensis
Eupatorium triplinerve Vahl =Ayapana triplinervis
Eupatorium tyleri B.L. Rob. =Chromolaena tyleri
Eupatorium urticifolium L. f. =Praxelis diffusa
Eupatorium vitalbae DC. =Heterocondylus vitalbae
Eupatorium wageneri Hieron. =Condylidium iresinoides
Eupatorium yavianum Lasser & Maguire =Guayania yaviana
Eurydochus bracteatus Maguire & Wurdack =Gongylolepis wurdackii
Eurydochus cortesii S. Diaz = Gonglolepis cortesii (S.Diaz) Pruski & S. Diaz
Gamochaeta purpurea (L.) Cabrera =Gamochaeta simplicicaulis
Geissopappus caleoides (DC.) Benth. ex Baker =Calea caleoides
Glossarion rhodanthum var. bracteatum Maguire =Glossarion rhodanthum
Gnaphalium americanum Mill. =Gamochaeta americana
Gnaphalium purpureum var. americanum (Mill.) Klatt =Gamochaeta americana
Gnaphalium purpureum var. simplicicaule (Willd. ex Spreng.) Klatt =Gamochaeta simplicicaulis
Gnaphalium satureioides Lam. =Achyrocline satureioides
Gnaphalium satureioides var. vargasianum (DC.) Kuntze =Achyrocline vargasiana
Gnaphalium schomburgkii Sch. Bip. [nomen nudum] =Gnaphalium polycaulon
Gnaphalium simplicicaule Willd. ex Spreng. =Gamochaeta simplicicaulis
Gnaphalium spicatum Lam. =Gamochaeta americana
Gnaphalium suaveolens Veil. =Pluchea sagittalis
Gochnatia calophylla (Sch. Bip.) V.M. Badillo =Gochnatia oligocephala
Gongylolepis benthamiana var. pubescens Maguire & Wurdack =Gongylolepis benthamiana
Gongylolepis cabreraei V.M. Badillo =Gongylolepis benthamiana
Gongylolepis huachamacari subsp. neblinensis Maguire & Wurdack =Gongylolepis huachamacari
Gongylolepis maroana V.M. Badillo ^Gongylolepis martiana
Gongylolepis paniculata var. umbellata Maguire & K.D. Phelps =Gongylolepis paniculata
Gongylolepis terramarae Steyerm. =Gong}'lolepis paniculata
Greenmania boladorensis Hieron. =Unxia camphorata
Guaicaia bilabiata Maguire =Glossarion bilabiatum
Guayania davidsei R.M. King & H. Rob. =Guayania crassicaulis
Heterothalamus densus N.E. Br. =Baccharis densa
Ichthyothere cunabi Mart. =Ichthyothere terminalis
Ichthyothere suffruticosa var. pilosa Aristeg. =Ichthyothere terminalis
Imeria serratifolia V.M. Badillo =Imeria memorabilis var. memorabilis
Inula primulifolia Lam. =Conyza primulaefolia
Kleinia ruderalis Jacq. =Porophyllum ruderale
Koanophyllon kavanayense (V.M. Badillo) R.M. King & H. Rob. =Koanophyllonfuscum
Kuhnia arguta Kunth =Trichogonia arguta
Lagascea mollis Cav. [not in flora area]
Latreillea glabrata Benth. =Ichthyothere terminalis
Lepidaploa auyantepuiensis (Aristeg.) H. Rob. =Lepidaploa ehretiifolia
Leria integrifolia Cass. =Chaptalia integerrima
Leucopholis latifolia Benth. =Chionolaena latifolia
Lipochaeta scaberrima Benth. =Oyedaea scaberrima
Lucilia breweri (Steyerm. & Maguire) V.M. Badillo =Chionolaena latifolia
Lycoseris latifolia (D. Don) Benth. =Lycoseris triplinervia
Lycoseris oblongifolia Rusby =Lycoseris triplinervia
Marsea bonariensis (L.) V.M. Badillo =Conyza bonariensis
Melampodium australe Loefl. =Acanthospermum australe
Melampodium bonairense Bold. =Baltimora geminata
Melampodium camphoratum (L. f.) Baker =Unxia camphorata
Melampodium geminata Brandegee =Baltimora geminata
Melampodium ruderale Sw. =Eleutheranthera ruderalis
Checklist of the Plants of the Guiana Shield 265
Melampodium suffruticosum Baker =Unxia suffruticosa
Melanthera aspera (Jacq.) Steud ex Small =Melanthera nivea
Mikania acrensis B.L. Rob. =Mikania lindleyana
Mikania amara (Vahl) Willd. =Mikania parviflora
Mikania amara var. guaco (Humb. & Bonpl.) Baker =Mikania guaco
Mikania anzoatiguensis V.M. Badillo =Mikania vitifolia
Mikania argyrostigma Miq. ^Mikania guaco
Mikania aschersonii sensu Pruski, non Hieron. =Mikama chlorolepis
Mikania aspera Miq. =Mikania guaco
Mikania atriplicifolia Sch. Bip. ex Miq. =Mikania congesta
Mikania auyantepuensis S.F. Blake =Mikania sprucei
Mikania auyantepuensis var. brevipedicellata Steyerm. =Mikania sprucei
Mikania banisteriae var. gabrieli (Baker) G.M. Barroso =Mikania banisteriae
Mikania bolivarensis V.M. Badillo =Mikania pannosa
Mikania capayensis V.M. Badillo =Mikania trinitaria
Mikania denticulata (Vahl) Willd. =Mikania micrantha
Mikania divaricata Poepp. =Mikania parviflora
Mikania fJorida Aristeg. =Mikania lucida
Mikaniafockeana Miq. ^Mikania psilostachya
Mikania gabrieli Baker =Mikania banisteriae
Mikania gonoclada DC. =Mikania cordifolia
Mikania gracilipes B.L. Rob. [not in flora area]
Mikania gracilis Sch. Bip. ex Miq. =Mikania vitifolia
Mikania hexagona B.L. Rob. =Mikania microptera
Mikania hirsutissima DC. =Mikania banisteriae
Mikania hookeriana var. crassicaulis Steyerm. =Mikania hookeriana
Mikania hookeriana var. platyphylla (DC.) B.L. Rob. =Mikania hookeriana
Mikania hostmannii Miq. =Mikania cordifolia
Mikaniajavitens is V.M. Badillo =Mikania nigropunctulata
Mikania karuaiensis V.M. Badillo =Mikania psilostachya
Mikania kavanayensis V.M. Badillo =Mikania banisteriae
Mikania lanata Sch. Bip. [not in flora area]
Mikania manaosensis Hieron. =Mikania sprucei
Mikania micrantha var. congesta (DC.) B.L. Rob. =Mikania congesta
Mikania orinocensis Kunth =Mikania micrantha
Mikania parkeriana DC. =Mikania congesta
Mikania phelpsii Maguire & Steyerm. =Mikania lucida
Mikania platyphylla DC. =Mikania hookeriana
Mikania psilostachya var. racemulosa (Benth.) Baker =Mikania psilostachya
Mikania psilostachya var. scabra (DC.) Baker =Mikania psilostachya
Mikania ptaretepuiens is V.M. Badillo =Mikania banisteriae
Mikania punctata Klatt =Mikania vitifolia
Mikania racemulosa Benth. =Mikania psilostachya
Mikania roraimensis B.L. Rob. ^Mikania sprucei
Mikania ruiziana Poepp. =Mikania banisteriae
Mikania santamensis V.M. Badillo =Mikania trinitaria
Mikania scabra DC. =Mikania psilostachya
Mikania scandens var. congesta (DC.) Baker =Mikania congesta
Mikania scandens var. microptera (DC.) Baker =Mikania microptera
Mikania serratifolia Sieber ex DC. =Condylidium iresinoides
Mikania sprucei var. roraimensis (B.L. Rob.) Aristeg. =Mikania sprucei
Mikania stictophora Steyerm. =Mikania sprucei
Mikania stipitata Sch. Bip. ex Miq. =Mikania parviflora
Mikania surinamensis Miq. =Mikania cordifolia
Mikania tepuiensis V.M. Badillo =Mikania nigropunctulata
Mikania tillettii R.M. King & H. Rob. =Mikania gleasonii
Mikania vitrea B.L. Rob. =Mikania hookeriana
Monanthemum cruegeri Griseb. =Piptocarpha triflora
Neurolaena suriani (Cass.) Cass. =Neurolaena lobata
Ogiera ruderalis (Sw.) Griseb. =Eleutheranthera ruderalis
Oliganthes areolata Wurdack =Piptocoma areolata
Oliganthes roraimensis Steyerm. =Piptocoma roraimensis
Oliganthes schomburgkii Sch. Bip. =Piptocoma schomburgkii
Oliganthes spruceana Benth. =Piptocoma spruceana
266 Checklist of the Plants of the Guiana Shield
Ooclinium clavatum Benth. =Chromolaena ivifolia
Ooclinium villosum (Cass.) DC. =Praxelis diffusa
Oyedaea blakeana Steyerm. =Oyedaea scaberrima
Oyedaea rusbyi S.F. Blake [not in flora area]
Pedis linifolia var. linifolia [not in flora area]
Pentacalia gramabanemis (Aristeg.) Cuatrec. ^Pentacaliafreemanii
Pentacalia nigella (V.M. Badillo) Cuatrec. =Pentacaliafreemanii
Piptocarpha cruegeri (Griseb.) Baker =Piptocarpha triflora
Piptocarphafoliosa Cuatrec. =Piptocarpha poeppigiana
Piptocarpha kavanayensis V.M. Badillo =Pip1ocarpha polycephala
Piptocarpha opaca var. latifolia (Sch. Bip.) Baker =Piptocarpha opaca subsp. opaca
Piptocarpha paraensis Cabrera =Piptocarpha poeppigiana
Piptocarpha sprucei Baker =Piptocarpha leprosa
Piptocarpha tereticaulis (DC.) Baker ^'Piptocarpha poeppigiana
Piptocarpha upatensis V.M. Badillo =Gochnatia oligocephala
Pluchea purpurascens (Sw.) DC. =Phichea odorata
Pluchea suaveolens (Veil.) Kuntze =Phich^a sagittalis
PoUalesta areolata (Wurdack) Aristeg. =Piptocoma areolata
Pollalestafaiistiana Stutts =Piptocoma schomhurgkii
PoUalesta milleri (Johnst.) Aristeg. [not in flora area]
Pollalesta roraimensis (Steyerm.) Aristeg. =Piptocoma roraimensis
Pollalesta schomburgkii (Sch. Bip.) Aristeg. =Piptocoma schomburgkii
Pollalesta spruceana (Benth.) Aristeg. =Piptocoma spruceana
Polymnia stenocarpha S.F. Blake =Unxia siiffruticosa
Polymnia suffniticosa (Baker) V.M. Badillo =Unxia suffruticosa
Porophyllum ellipticum (L.) Cass. =Porophyllum ruderale
Porophyllum latifolium Benth. =Porophyllum ruderale
Porophyllum macrocephalum DC. ^Porophyllum ruderale
Porophyllum porophyllum (L.) Kuntze ^Porophyllum ruderale
Porophyllum ruderale subsp. macrocephalum (DC.) R.R. Johnson =Porophyllum ruderale
Praxelis karuaiensis (V.M. Badillo) R.M. King & H. Rob. =Praxelis asperulacea
Praxelis pauciflora (Kunth) R.M. King & H. Rob. =Praxelis diffusa
Praxelis villosa Cass. =Praxelis diffusa
Pseudelephantopus spiralis (Less.) Cronquist [not in flora area]
Pseudobaccharis densa (N.E. Br.) V.M. Badillo =Baccharis densa
Pseudobaccharis ligustrina (DC.) I.L. Teodoro =Baccharis brachylaenoides var. ligustrina
Pseudobaccharis rhexioides (Kunth) V.M. Badillo =Baccharis trinervis
Pseudobaccharis trinervis (Lam.) V.M. Badillo =Baccharis trinervis
Psila brachylaenoides (DC.) Aristeg. =Baccharis brachylaenoides
Psila brachylaenoides var. ligustrina (DC.) Aristeg. =Baccharis brachylaenoides var. ligustrina
Psila densa (N.E. Br.) V.M. Badillo =Baccharis densa
Psila ligustrina (DC.) Cabrera =Baccharis brachylaenoides var. ligustrina
Psila trinervis (Lam.) Cabrera =Baccharis trinervis
Pterocaulon virgatum var. alopecuroides (Lam.) Griseb. =Pterocaulon alopecuroides
Riencourtia glomerata Cass. ^Riencourtia pedunculosa
Riencourtia ovata S.F. Blake =Riencourtia latifolia
Riencourtia pittieri S.F. Blake =Riencourtia latifolia
Rolandra argentea Rottb. =Rolandrafruticosa
Rolandra septans Willd. ex Less. =Trichospira verticillata
Rolandra terminalis Spreng. =Ichthyothere terminalis
Schomburgkia caleoides DC. =Calea caleoides
Senecio chenopodioides Kunth =Pseudogynoxys chenopodioides
Senecio confusus Britten =Pseudogynoxys chenopodioides
Senecio freemanii Britton & Greenm. =Pentacaliafreemanii
Senecio gransabanensis Aristeg. =Pentacaliafreemanii
Senecio hieracifolius L. =Erechtites hieracifolius
Senecio nigellus V.M. Badillo =Pentacaliafreemanii
Senecio phelpsiae Cuatrec. =Pentacalia phelpsiae
Senecio ptarianus Cuatrec. =Pentacalia ptariana
Senecio valerianifolius Wolf =Erechtites valerianifolius
Senecio yapacanus Aristeg. =Pentacalia yapacana
Seruneum scaberrimum (Benth.) Kuntze =Wedelia calycina
Silphium trilobatum L. =Sphagneticola trilobata
Sonchus oleraceus L. [not in flora area]
Checklist of the Plants of the Guiana Shield 267
Sonchus oleraceus var. asper L. =Sonchus asper
Sparganophorus sparganophora (L.) C. Jeffrey =Struchium sparganophorum
Sparganophorus vaillantii Crantz =Struchium sparganophorum
Spilanthes acmella (L.) Murray =Acmella sp.
Spilanthes acmella var. uliginosa (Sw.) Baker =Acmella uliginosa
Spilanthes barinensis Aristeg. =Spilanthes nervosa
Spilanthes ciliata Kunth =Acmella ciliata
Spilanthes debilis Kunth ^Acmella radicans var. debilis
Spilanthes oppositifolia (Lam.) D'Arcy =Acmella repens
Spilanthes paniculata Wall, [not in flora area] =Acmella sp.
Spilanthes radicans Jacq. =Acmella radicans
Spilanthes repens Spreng. ex DC. =Spilanthes urens
Spilanthes tenella Kunth =Acmella radicans var. debilis
Spilanthes uliginosa Sw. =Acmella uliginosa
Spilanthes uliginosa var. discoidea Aristeg. =Acmella uliginosa
Spilanthes urens var. megalophylla S.F. Blake =Spilanthes urens
Spilanthes urens var. nervosa (Chodat) Hassl. =Spilanthes nervosa
Spiracantha cornifolia Kunth [not in flora area]
Stenopadus affinis Maguire, Steyerm. & Wurdack =Stenopadus talaumifolius
Stenopadus cardonae Maguire & Lasser =Stenopadus talaumifolius
Stenopadus cinereus Gleason & S.F. Blake =Chimantaea cinerea
Stenopadus cinereus f. glabra Steyerm. =Chimantaea cinerea
Stenopadus condensatus (Baker) S.F. Blake =Stomatochaeta condensata
Stenopadus crassifolius S.F. Blake [not in flora area]
Stenopadus cymbifolius S.F. Blake =Stomatochaeta cymbifolia
Stenopadus eurylepis S.F. Blake [not in flora area]
Stenopadus guaiquinimensis V.M. Badillo =Stomatochaeta condensata
Stenopadus kunhardtii var. grandifolia Maguire =Stenopadus kunhardtii
Stenopadus neblinensis Maguire & Wurdack =Stenopadus talaumifolius
Stenopadus steyermarkii Aristeg. =Stenopadus talaumifolius
Stenopadus stipitatus Maguire & Wurdack ^Stenopadus kunhardtii
Stenopadus talaumifolius var. magnifolius Steyerm. =Stenopadus talaumifolius
Stenopadus variabilis S.F. Blake =Stomatochaeta condensata
Stenopadus ventricosus Maguire & Aristeg. ^Stenopadus sericeus
Stifftia benthamiana (R.H. Schomb.) Baker =Gongylolepis benthamiana
Stifftia condensata Baker =Stomatochaeta condensata
Stifftia connellii N.E. Br. =Stenopadus connellii
Stifftia martiana Baker =Gongylolepis martiana
Stilpnopappus aquaticus (Poepp.) M.O. Dillon =Xiphochaeta aquatica
Stilpnopappus viridis Benth. ex Baker =Xiphochaeta aquatica
Stomatochaeta colveei Steyerm. =Stenopadus colveei
Tephrothamnus calophyllus Sch. Bip. =Gochnatia oligocephala
Tessaria dentata Ruiz & Pav. =Tessaria integrifolia
Thelechitonia brachycarpa (Baker) H. Rob. & Cuatrec. =Sphagneticola brachycarpa
Thelechitonia muricata Cuatrec. =Sphagneticola brachycarpa
Thelechitonia trilobata (L.) H. Rob. & Cuatrec. =Sphagneticola trilobata
Tilesia baccata var. baccata =Wulffia baccata
Tilesia capitata G. Mey. =Wulffia baccata
Tilesia rubens (Alexander) Pruski =Wulffia rubens
Trichospira biaristata Less. =Trichospira verticillata
Trichospira menthoides Kunth =Trichospira verticillata
Trichospira prieurei DC. =Trichospira verticillata
Trichospira pulegium Mart, ex DC. =Trichospira verticillata
Trinchinettia caleoides (DC.) Endl. =Calea caleoides
Trixis pedunculosa Rich. =Riencourtia pedunculosa
Tussilago integerrima Veil. =Chaptalia integerrima
Tussilago nutans L. =Chaptalia nutans
Unxia kubitzkii H. Rob. =Unxia suffruticosa
Verbesina alata L. [not in flora area]
Verbesina alba L. =Eclipta prostrata
Verbesina nodiflora L. =Synedrella nodiflora
Verbesina prostrata L. =Eclipta prostrata
Verbesina schomburgkii subsp. ligulata Maguire & Wurdack =Verbesina ligulata
Verbesina schomburgkii subsp. schomburgkii =Verbesina guianensis
268 Checklist of the Plants of the Guiana Shield
Verbesina weberbaueri Sagast. =Verbesina columbiana
Vernonia apicidata Mart, ex DC. =Stenocephalum apiculatum
Vernonia auyantepuiensis Aristeg. =LepidapJoa ehretiifolia
Vernonia bolivarensis V.M. Badillo =Lepidaploa bolivarensis
Vernonia brasiliana (L.) Druce =Vernonanthura brasiliana
Vernonia cardonae Aristeg. =Lepidaploa ehretiifolia
Vernonia chrysotricha Alexander =Lepidaploa chrysotricha
Vernonia cinerea (L.) Less. =Cyanthillium cinereum
Vernonia cotoneaster (Willd. ex Spreng.) Less, [misident. of Lepidaploa gracilis H. Rob.] =Lepidaploa cotoneaster
Vernonia decantha Sch. Bip. ^Lepidaploa ehretiifolia
Vernonia ehretiifolia Benth. =Lepidaploa ehretiifolia
Vernonia glandiilosa V.M. Badillo =Lepidaploa bolivarensis
Vernonia gracilis Kunth =Lepidaploa gracilis
Vernonia giiianensis V.M. Badillo =Lepidaploa salzmannii
Vernonia imeriensis V.M. Badillo =Lepidaploa imeriensis
Vernonia leprosa Less. =Piptocarpha leprosa
Vernonia miersiana Gardner =Lepidaploa salzmannii
Vernonia morilloi V.M. Badillo =Lessingianthiis morilloi
Vernonia moritziana Sch. Bip. =Lepidaploa gracilis
Vernonia odoratissima Kunth -Vernonanthura brasiliana
Vernonia opaca Benth. =Piptocarpha opaca
Vernonia pari V.M. Badillo =Lepidaploa pari
Vernonia poeppigiana DC. =Piptocarpha poeppigiana
Vernonia remotiflora Rich. =Lepidaploa remotiflora
Vernonia scahra Pers. =Vernonanthura brasiliana
Vernonia schomburgkiana Sch. Bip. =Lepidaploa ehretiifolia
Vernonia scorpioides (Lam.) Pers. =Cyrtocymiira scorpioides
Vernonia sericea Rich, [not in flora area] =Lepidaploa sericea (Rich.) H. Rob.
Vernonia sessilijlora Willd. =Lepidaploa remotiflora
Vernonia tereticaiilis DC. =Piplocarpha poeppigiana
Vernonia tricephala auct. non Gardner =Lepidaploa pari
Vernonia tricholepis DC. ^Lepidaploa remotiflora
Wedelia brachycarpa Baker =Sphagneticola brachycarpa
Wedelia brasiliensis (Spreng.) S.F. Blake [nom. illeg.] =Sphagneticola trilobata
Wedelia brasiliensis van villosa (Baker) S.F. Blake =Sphagneticola brachycarpa
Wedelia caracasana DC. =Wedelia calycina
Wedelia discoidea Less. =Eleutheranthera ruderalis
Wedeliafrutescens Jacq. =Wedelia calycina
Wedelia friiticosa Jacq. [not in flora area. Misident. of W calycina Rich.]
Wedeliajacquini O.E. Schulz =Wedelia calycina
Wedelia pahidosa DC. =Sphagneticola trilobata
Wedelia pahidosa var. villosa Baker ? =Sphagneticola brachycarpa
Wedelia parviflora Rich. =Wedelia calycina
Wedelia psammophila Poepp. =Eclipta prostrata
Wedelia rubra (Aristeg.) B.L. Turner =Tuberciilocarpus ruber
Wedelia scaberrima Benth. -Wedelia calycina
Wedelia trilobata (L.) Hitchc. =Sphagneticola trilobata
Willoughbya amazonica (Baker) Kuntze =Mikania amazonica
Willoughbya argyrostigma (Miq.) Kuntze =Mikania guaco
Willoughbya aspera (Miq.) Kuntze =Mikania guaco
Willoughbya banisteriae (DC.) Kuntze =Mikania banisteriae
Willoughbya chlorolepis (Baker) Kuntze =Mikania chlorolepis
Willoughbya cordifolia (L. f.) Kuntze =Mikania cordifolia
Willoughbya divaricata (Poepp.) Kuntze =Mikania parviflora
Willoughbya gabrieli (Baker) Kuntze =Mikania banisteriae
Willoughbya gracilis (Sch. Bip. ex Miq.) Kuntze =Mikania vitifolia
Willoughbya guaco (Humb. & Bonpl.) Kuntze =Mikania guaco
Willoughbya hirsutissima (DC.) Kuntze =Mikania banisteriae
Willoughbya lindleyana (DC.) Kuntze =Mikania lindleyana
Willoughbya micrantha (Kunth) Rusby =Mikania micrantha
Willoughbya pannosa (Baker) Kuntze =Mikania pannosa
Willoughbya parviflora (Aubl.) Kuntze =Mikania parviflora
Willoughbya parviflora var. guaco (Bonpl.) Kuntze =Mikania guaco
Willoughbya platyphylla (DC.) Kuntze =Mikania hookeriana
Checklist of the Plants of the Guiana Shield 269
Willoughbya psilostachya (DC.) Kuntze =Mikama psilostachya
Willoughbya scabra (DC.) Kuntze ^Mikania psilostachya
Willoughbya scandens var. congesta (DC.) Kuntze =Mikania congesta
Willoughbya scandens var. orinocensis (Kunth) Kuntze =Mikania micrantha
Willoughbya sprucei (Baker) Kuntze =Mikania sprucei
Willoughbya stipitata (Sch. Bip. ex Miq.) Kuntze ^Mikania parviflora
Willoughbya trinitaria (DC.) Kuntze =Mikania trinitaria
Willoughbya vitifolia (DC.) Kuntze =Mikania vitifolia
Wulffia capitata (G. Mey.) Sch. Bip. =Wulffia baccata
Wulffia maculata DC. = Wulffia baccata
Wulffia platyglossa (Cass.) DC. =Wulffia baccata
Wulffiia stenoglossa (Cass.) DC. =Wulffiia baccata
Zexmenia scaberrima (Benth.) Baker =Oyedaea scaberrima
Zexmenia tepuiana V.M. Badillo =Oyedaea tepuiana
References
Aristeguieta, L. 1964. Compositae. In T. Lasser, Flora de Venezuela 10: 1-947.
Badillo, V.M. 1994 (1995). Enumeracion de las Compuestas (Asteraceae) de Venezuela. Revista de la Facultad de Agronomia
(Maracay) 45: 1-191.
Bremer, K. 1994. Asteraceae: Cladistics & Classification. Portland, Oregon: Timber Press.
Funk, V.A. 1991. The Compositae of the Guianas, I: Heliantheae (Heliantheae, Tageteae, Coreopsideae). Rhodora 93(875): 256-
267.
King, R.M., and H. Robinson. 1987. The genera ofthe Eupatorieae (Asteraceae). Monogrographs in Systematic Botany Missouri
Botanical Garden 22: 1-581.
Koster, J.T. 1938. Compositae. In A. Pulle, Flora of Suriname 4(2): 87-165.
Pruski, J.F. 1991 (1993). Compositae of the Guayana Highland - V. The Mutisieae of the lost world of Brazil, Colombia, and
Guyana. Bol. Mus. Paraense Emilio Goeldi (Bot.) 7: 335-392.
Pruski, J.F. 1994. Compositae ofthe Guayana Shield, with special reference to the tribe Mutisieae. Page 28 (abstract) in Compositae:
Systematics, Biology, Utilization: Papers and Abstracts, coordinator D. J. N. Hind. Kew: Royal Botanic Gardens.
Pruski, J.F. 1997. Asteraceae. In J.A. Steyermark, P.E. Berry, & B.K. Hoist, Flora of the Venezuelan Guayana 3: 177-393.
Pruski, J.F. 2002. Asteraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 94-
116.
Robinson, H. 1992. The Asteraceae of the Guianas, III: Vemonieae and restoration of the genus Xiphochaeta. Rhodora 94(880):
348-361.
CONNARACEAE
E. Forero, E. Santana
Cnestidium guianense (G. Schellenb.) G. Schellenb.
Connanis cordatus L.A. Vidal, Carbono & Forero
Connarus cohaceus G. Schellenb.
Connarus ehanthus Benth. ex Baker
Connarus ehanthus var. stipitatus Forero
Connarusfasciculatus (DC.) Planch.
Connarusfasciculatus subsp. fasciculatus
Connarus incomptus Planch.
Connarus lambertii (DC.) Sagot
Connarus megacarpus S.F. Blake
Connarus patrisii (DC.) Planch.
Connarus perrottetii (DC.) Planch.
Connarus perrottetii var. perrottetii
Connarus perrottetii var. rufus Forero
Connarus punctatus Planch.
Connarus rigidus Forero
Connarus ruber (Poepp.) Planch.
Connarus ruber var. ruber
Connarus ruber var. sprucei (Baker) Forero
Connarus venezuelanus Baill.
Connarus venezuelanus var. orinocensis Forero
Connanis venezuelanus var. venezuelanus
Pseudoconnarus macrophyllus (Poepp.) Radlk.
Pseudoconnarus subtriplinervis (Radlk.) G. Schellenb.
Rourea amazonica (Baker) Radlk.
AM
AM
AM
AM
BO DA
BO
GU su FG
BO
BO
BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
BO
DA
BO DA
DA
DA
DA
GU SU FG
GU su FG?
GU su FG?
su FG
su FG
GU
GU
GU
GU su FG
GU su FG
GU su FG
GU
GU su FG
AM
GU
GU
GU?
SU
270 Checklist of the Plants of the Guiana Shield
Rourea cuspidata Benth. ex Baker
Rourea cuspidata van cuspidata
Rourea cuspidata var. multijuga Forero
Rourea cuspidata var. pedicellata Baker
Roureaforeroi Aymard & P.E. Berry
Roureafrutescens Aubl.
Rourea glabra Kunth
Rourea glabra var. glabra
Rourea grosourdyana Baill.
Rourea grosourdyana var. grosourdyana
Rourea kappleri Lanj.
Rourea krukovii Steyerm.
Rourea ligulata Baker
Rourea neglecta G. Schellenb.
Rourea neglecta var. neglecta
Rourea pubescens (DC.) Radlk.
Rourea pubescens var. pubescens
Rourea pubescens var. spadicea (Radlk.) Forero
Rourea revoluta Planch.
Rourea revoluta var. glabra Baker
Rourea revoluta var. revoluta
Rourea sprucei G. Schellenb.
Rourea sprucei var. sprucei
Rourea sprucei var. subcoriacea Forero
Rourea surinamensis Miq.
Synonyms and Misidentifications
Bernardinia guianensis G. Schellenb. =Cnestidium guianense
Connarus araucanus Cuatrec. =Connarus venezuelanus var. orinocensis
Connarus confertiflorus Baker =Connarus patrisii
Connarus fecundus Baker =Pseudoconnarus macrophyllus
Connarus floribundus Planch. =Connarus perrottetii
Connarus guianensis Lamb, ex DC. =Connarus lambertii
Connarus incomptus var. subcordata Baker =Connarus incomptus
Connarus macrophyllus Poepp. =Pseudoconnarus macrophyllus
Connarus neglectus G Schellenb. =Connarus coriaceus
Connarus opacus G Schellenb. =Connarus coriaceus
Connarus pubescens DC. =Rourea pubescens
Connarus schomburgkii auct. non Planch. 1850: sensu Pittier =Connarus ruber var. sprucei
Connarus schomburgkii Planch. =Connarus lambertii
Connarus sprucei auct. non Baker 1871 : sensu Pittier =Connarus lambertii
Connarus sprucei Baker =Connarus ruber var. sprucei
Omphalobiumfasciculatum DC. =Connarus fasciculatus
Omphalobium lambertii DC. =Connarus lambertii
Omphalobium patrisii DC. =Connarus patrisii
Omphalobium perrottetii DC. =Connarus perrottetii
Omphalobium rubrum Poepp. =Connarus ruber
Pseudoconnarusfecundus (Baker) Radlk. =Pseudoconnarus macrophyllus
Pseudoconnarus krukovii Steyerm. =Pseudoconnarus macrophyllus
Robergiafrutescens (Aubl.) Willd. =Roureafrutescens
Rourea amazonica Huber =Rourea amazonica
Rourea cardonae Lasser & Maguire =Roureafrutescens
Rourea glabra Kunth var. amazonica Baker =Rourea amazonica
Rourea glabra var. coriacea Baker =R. neglecta var. neglecta p.p., R. surinamensis p.p.
Rourea glabra var. parviflora Baker =Rourea cuspidata var. cuspidata
Rourea granatensis Cuatrec. =Rourea glabra var glabra
Rourea induta var. concinna Baker =Roureafrutescens
Rourea macrophylla (Poepp.) Baker =Pseudoconnarus macrophyllus
Rourea spadicea Radlk. =Rourea pubescens var. spadicea
Rourea subtriplinervis Radlk. =Pseudoconnarus subtriplinervis
Santalodes cuspidatum (Benth. ex Baker) Kuntze =Rourea cuspidata
Santalodesfrutescens (Aubl.) Kuntze =Roureafrutescens
Santalodes macrophyllum (Poepp.) Kuntze ^Pseudoconnarus macrophyllus
Santalodes revolutum (Planch.) Kuntze =^Rourea revoluta
AM
AM
AM
AM
AM
BO DA GU SU FG
AM BO
AM BO
BO
BO
BO
GU
GU
GU?
SU
FG
AM FG
AM FG
DA GU SU FG
DA FG
GU SU FG
GU
GU
GU
AM BO
AM
AM BO
BO DA GU SU FG
Checklist of the Plants of the Guiana Shield 271
References
Forero, E. 1983. Connaraceae. Flora Neotropica Monogr. 36: 1-207.
Forero, E. 2002. Connaraceae. In Mori, S.A., G. Cremers, C. Oracle, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 227-
230.
Forero, E. & A.R.A. Gorts-van Rijn. 1976. Connaraceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of
Suriname 2(2): 654-657.
Forero, E. & E. Santana. 1998. Connaraceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4:
365-376.
Lanjouw, J. 1940. Connaraceae. In A. Pulle, Flora of Suriname 2(2): 332-340.
CONVOLVULACEAE
D.F. Austin
Aniseia cernua Choisy
Aniseia martinicensis (Jacq.) Choisy
Bonamia apurensis D.F. Austin
Bonamia holtii O'Donell
Bonamia maripoides Hallier f.
Calycobolus glaber (Kunth) House
Convolvulus nodiflorus Desr.
Dicranostyles ampla Ducke
Dicranostyles ampla var. ampla
Dicranostyles ampla var. castanea D.F. Austin
Dicranostyles globostigma D.F. Austin
Dicranostyles guianensis Mennega
Dicranostyles holostyla Ducke
Dicranostyles Integra Ducke
Dicranostyles longifolia Ducke
Dicranostyles scandens Benth.
Dicranostyles sericea Gleason
Dicranostyles villosus Ducke
Evolvulus alsinoides (L.) L.
Evolvulus cardiophyllus Schltdl.
Evolvulus convolvuloides (Willd.) Steam
Evolvulus elegans Moric.
Evolvulus filipes Mart.
Evolvulusfrankenioides Moric.
Evolvulus glomeratus Nees & Mart.
Evolvulus nummularius (L.) L.
Evolvulus ovatus Femald
Evolvulus paniculatus (Bonpl.) Spreng.
Evolvulus pterocaulon Moric.
Evolvulus sericeus Sw.
Evolvulus sericeus var. holosericeus (Kunth) Ooststr.
Evolvulus sericeus var. sericeus
Evolvulus tenuis Mart, ex Choisy
Evolvulus tenuis subsp. sericatus (House) Ooststr.
Evolvulus villosissimus Ooststr.
Ipomoea alba L.
Ipomoea angulata Mart, ex Choisy
Ipomoea anisomeres B.L. Rob. & Bartlett
Ipomoea aquatica Forssk.
Ipomoea argentea Meisn.
Ipomoea asarifolia (Desr.) Roem. & Schult.
Ipomoea batatas (L.) Lam. [cuhivated and frequently escaping]
Ipomoea batatoides Choisy
Ipomoea bignonioides Lindl.
Ipomoea cairica (L.) Sweet
Ipomoea carnea Jacq.
Ipomoea carnea subsp. fistulosa (Mart, ex Choisy) D.F. Austin
Ipomoea discolor (Kunth) G. Don
Ipomoea distans Choisy
Ipomoeafimbriosepala Choisy
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
DA
DA
DA
GU SU FG
GU SU EG
GU SU FG
GU SU FG
GU SU?
GU SU?
GU SU?
GU SU FG
GU?
FG
GU?
SU FG
GU SU FG
GU SU FG
GU SU? FG'
GU
GU
GU
GU
GU
SU
SU
SU
SU
SU
FG
FG
FG
I
BO DA GU SU FG
FG?
GU SU FG?
GU SU FG
BO GU SU? FG?
BO GU?
BO DA GU SU FG
GU SU FG
FG
GU
BO DA GU SU?
BO DA GU SU?
BO
DA
FG
FG?
272 Checklist of the Plants of the Guiana Shield
Ipomoea gabrieli Choisy FG
Ipomoea hederifolia L.
Ipomoea horsfalliae Hook.
Ipomoea imperati (Vahl) Griseb.
Ipomoea incarnata (Vahl) Choisy
Ipomoea indica (Burm.) Merr.
Ipomoea killipiana O'Donell
Ipomoea leprieurii D.F. Austin
Ipomoea mauritiana Jacq.
Ipomoea meyeri (Spreng.) G. Don
Ipomoea minutiflora (M. Martens & Galeotti) House
Ipomoea nil (L.) Roth
Ipomoea pes-caprae (L.) R. Br.
Ipomoea phyllomega (Veil.) House
Ipomoea piurens is O'Donell
Ipomoea quamoclit L. ^.
Ipomoea reticulata O'Donell
Ipomoea rubens Choisy
Ipomoea schomburgkii Choisy
Ipomoea setifera Poir.
Ipomoea squamosa Choisy
Ipomoea subrevoluta Choisy
Ipomoea tiliacea (Willd.) Choisy
Ipomoea trifida (Kunth) G. Don
Ipomoea turbinata Lag.
Ipomoea violacea L. [expected in DA]
Iseia luxurians (Moric.) O'Donell
Jacquemontia agrestis (Choisy) Meisn.
Jacquemontia ciliata Sandwith
Jacquemontia densiflora (Meisn.) Hallier f.
Jacquemontia gracillima (Choisy) Hallier f.
Jacquemontia guyanensis (Aubl.) Meisn.
Jacquemontia pentantha (Jacq.) G Don
Jacquemontia sphaerostigma (Cav.) Rusby
Jacquemontia tamnifolia (L.) Griseb.
Jacquemontia velutina Choisy
Lysiostyles scandens Benth.
Maripa densiflora Benth.
Maripa elongata Ducke
Maripa glabra Choisy
Maripa longifolia Sagot ex Hallier f
.
Maripa paniculata Barb. Rodr.
Maripa pauciflora D.F. Austin
Maripa repens Rusby
Maripa reticulata Ducke
Maripa reticulata var. reticulata
Maripa reticulata var. rugosa (Ducke) D.F. Austin
Maripa scandens Aubl.
Maripa violacea (Aubl.) Ooststr. ex Lanj. & Uittien
Merremia aegyptia (L.) Urb.
Merremia aturensis (Kunth) Hallier f.
Merremia cissoides (Lam.) Hallier f.
Merremia digitata (Spreng.) Hallier f.
Merremia dissecta (Jacq.) Hallier f.
Merremia macrocalyx (Ruiz & Pav.) O'Donell
Merremia maypurensis Hallier f.
Merremia quinquefolia (L.) Hallier f.
Merremia ternifoliola Pittier
Merremia umbellata (L.) Hallier f.
Merremia wurdackii D.F. Austin & Staples
Odonellia hirtiflora (M. Martens & Galeotti) K.R. Robertson
Operculina hamiltonii (G Don) D.F. Austin & Staples
Operculina sericantha (Miq.) Ooststr.
BO GU SU
SU
DA GU SU FG
BO
BO DA GU?
FG
FG
AM BO
BO
BO
DA GU SU FG
AM BO GU SU FG
DA GU SU FG
AM BO DA GU SU FG
AM BO GU SU FG
BO GU SU FG
BO
DA GU SU FG
AM BO GU SU FG
DA GU SU FG
AM BO DA GU SU
DA GU SU FG
BO DA GU SU FG
AM BO
BO
GU?
GU SU FG
BO DA GU?
AM BO GU
DA GU SU FG
BO
BO
AM BO DA GU SU FG
AM BO
BO GU
FG
AM BO DA GU
GU?
SU
SU?
FG
BO DA GU SU FG
AM BO GU SU FG
AM
GU SU FG
FG
AM GU SU? FG
AM
BO DA
AM GU SU FG
AM GU SU FG
AM
AM BO DA GU SU FG
AM BO GU SU FG
GU SU FG
AM BO GU SU FG
AM BO DA GU
GU
SU FG
BO GU SU FG
AM BO DA GU SU FG
AM
BO
BO
GU
GU
BO DA GU SU FG
AM BO GU SU FG
AM BO
BO GU SU FG
BO DA GU SU FG
Checklist of the Plants of the Guiana Shield 273
Operculina turpethum (L.) Silva Manso [var. turpethum?] [introduced, native SU FG
to Asia]
Operculina turpethum var. ventricosa (Bertero) Staples & D.F. Austin SU
[introduced, native to Asia]
Tetralocularia pennellii O ' Donell FG
Synonyms and Misidentifications
Aniseia cernua Moric. =Aniseia martinicensis
Aniseia ensifolia Choisy =Aniseia martinicensis
Aniseia ensifolia var. minor Choisy =Aniseia cernua
Aniseia gracillima Choisy =Jacquemontia gracillima
Aniseia hastata Meisn. =Ipomoeafimbriosepala
Aniseia minor (Choisy) J.A. McDonald =Aniseia cernua
Aniseia trichantha Pittier =Iseia luxurians
Batatas crassicaulis Benth. =Ipomoea carnea suhsp. fistulasa
Batatas edulis (Thunb. ex Murray) Choisy =Ipomoea batatas
Batatas villosa Choisy =Ipomoea argentea
Calonyction aculeatum (L.) House =Ipomoea alba
Calonyction bona-nox (L.) Bojer =Ipomoea alba
Calonyction muricatum (L.) G. Don. =Ipomoea turbinata
Calonyction tuba (Schltdl.) Colla =Ipomoea violacea
Calycobolus pulchellus Willd. ex Roem. & Schult. =Calycobolus glaber
Calycobolus spectabilis (Choisy) House =Bonamia maripoides
Calystegia setifera (Poir.) Meisn. ^Ipomoea setifera
Cladostyles paniculatus Bonpl. =Evolvulus paniculatus
Convolvulus aculeatus L. =Ipomoea alba
Convolvulus acuminatus Vahl =Ipomoea indica
Convolvulus agrestis Choisy =Jacquemontia agrestis
Convolvulus alatus Ham. =Operculina hamiltonii
Convolvulus alsinoides L. =Evolvulus alsinoides
Convolvulus asarifolius Desr. =Ipomoea asarifolia
Convolvulus aturensis Kunth =Merremia aturensis
Convolvulus azureus Desr. =Jacquemontia pentantha
Convolvulus batatas L. =Ipomoea batatas
Convolvulus batatilla Kunth =Ipomoea carnea subsp. fistulosa
Convolvulus caracassanus Roem. & Schult. =Merremia umbellata
Convolvulus cissoides Lam. =Merremia cissoides
Convolvulus discolor Kunth =Ipomoea discolor
Convolvulus edulis Thunb. ex Murray =Ipomoea batatas
Convolvulus glaber Aubl. =Merremia macrocalyx
Convolvulus guyanensis Aubl. —Jacquemontia guyanensis
Convolvulus imperati Vahl =Ipomoea imperati
Convolvulus incarnatus Vahl =Ipomoea incarnata
Convolvulus indicus Burm. =Ipomoea indica
Convolvulus littoralis L. =Ipomoea imperati
Convolvulus macrocalyx Ruiz & Pav. =Merremia macrocalyx
Convolvulus martinicensis Jacq. =Aniseia martinicensis
Convolvulus meyeri Spreng. =Ipomoea meyeri
Convolvulus minutiflorus M. Martens & Galeotti =Ipomoea minutiflora
Convolvulus mollis Kunth =Ipomoea indica
Convolvulus muricatus L. =Ipomoea turbinata
Convolvulus nil L. =Ipomoea nil
Convolvulus nummularius L. =Evolvulus nummularius
Convolvulus paniculatus L. =Ipomoea mauritiana
Convolvulus pentanthus Jacq. =Jacquemontia pentantha
Convolvulus pes-caprae L. =Ipomoea pes-caprae
Convolvulus phillomega Veil. =Ipomoea phyllomega
Convolvulus pinnatus Desr. =Ipomoea quamoclit
Convolvulus quinquefolius (L.) L. =Merremia quinquefi)lia
Convolvulus riparius Kunth =Merremia cissoides
Convolvulus ruber Vahl =Ipomoea setifera
Convolvulus sphaerostigma Cav. =Jacquemontia sphaerostigma
Convolvulus stoloniferus Cirillo =Ipomoea imperati
Convolvulus tamnifolius (L.) G. Mey. =Jacquemontia tamnifolia
274 Checklist of the Plants of the Guiana Shield
Convolvulus tiliaceus Willd. =Ipomoea tiliacea
Convolvulus trifidus Kunth =Ipomoea thfida
Convolvulus umbellotus L. =Merremla umbellata
Convulvulus dissectus Jacq. =Merremia dissecta
Dicranosnles imafacensis Steyerm. =Dicranosnles guianensis
Dufourea glabra Kunth =Calycobolus glaber
Evolvulus alsinoides var. debilis (Kunth) Ooststr. ^Evolvulus alsinoides
Evolvulus angustissimus Kunth =Evolvulus sericeus var. sericeus
Evolvulus arenicola J.R. Johnst. =Evolvulus sericeus var. holosericeus
Evolvulus commelinifolius Femald =Evolvulus cardiophyllus
Evolvulus debilis Kunth =Evolvulus alsinoides
Evolvulus glomeratus f. strictus (Benth.) Ooststr. =Evolvulus glomeratus
Evolvulus holosericeus Kunth =Evolvulus sericeus var. holosericeus
Evolvulus linifolius auct. non L. 1762. =Evolvulus filipes
Evolvulus sericatus House =Evolvulus tenuis subsp. sericatus
Evolvulus strictus Benth. =Evolvulus glomeratus
Ipomoea acuminata (Vahl) Roem. & Schult. =Ipomoea indica
Ipomoea altissima Mart. e.\ Choisy =Operculina hamiltonii
Ipomoea alulata Miq. =Operculina hamiltonii
Ipomoea aturensis (Kunth) G. Don =Merremia aturensis
Ipomoea batatilla (Kunth) G. Don =Ipomoea carnea subsp. fistulosa
Ipomoea bona-nox L. =Ipomoea alba
Ipomoea cardiosepala Meisn. =Ipomoea phyllomega
Ipomoea cathartica Poir. =Ipomoea indica
Ipomoea cissoides f. viscidula Meisn. -Merremia cissoides
Ipomoea coccinea auct. non L. 1753. =Ipomoea hederifolia
Ipomoea comosa House =Ipomoea argentea
Ipomoea crassicaulis (Benth.) B.L. Rob. ^Ipomoea carnea subsp. fistulosa
Ipomoea demerariana Choisy =Ipomoea phyllomega
Ipomoea digitata L. [not in flora area]
Ipomoea digitata auct. non L. 1753. =Ipomoea mauritiana
Ipomoeafistulosa Mart, ex Choisy =Ipomoea carnea subsp. fistulosa
Ipomoeafragrans (Bojer) Bojer ex Choisy =Ipomoea rubens
Ipomoea glabra (Aubl.) Choisy =Merremia macrocalyx
Ipomoea guyanensis (Aubl.) Choisy =Jacquemontia guyanensis
Ipomoea hamiltonii G. Don =Operculina hamiltonii
Ipomoea hirtiflora M. Martens & Galeotti =Odonellia hirtiflora
Ipomoea hostmanni Meisn. =.Merremia macrocalyx
Ipomoea indica var. acuminata (Vahl) Fosberg =Ipomoea indica
Ipomoeajuncea Choisy =Merremia aturensis
Ipomoea littoralis (L.) Boiss. -Ipomoea imperati
Ipomoea luxurians Moric. =Iseia luxurians
Ipomoea macrantha Roem. & Schult. -Ipomoea violacea
Ipomoea macrocalyx (Ruiz & Pav.) Choisy =Merremia macrocalyx
Ipomoea martinicensis (Jacq.) G Mey. "^Aniseia martinicensis
Ipomoea mollicoma Miq. =Merremia umbellata
Ipomoea mollis (Kunth) G. Don =Ipomoea indica
Ipomoea muricata (L.) Jacq. =Ipomoea turbinata
Ipomoea paniculata (L.) R. Br. =Ipomoea mauritiana
Ipomoea parkeri Choisy =Ipomoea rubens
Ipomoea parkeri var. subsericea Meisn. =Ipomoea rubens
Ipomoea periyana Duchass. & Walp. =OdoneUia hirtiflora
Ipomoea polyanthes Roem. & Schult. -Merremia umbellata
Ipomoea pterodes Choisy =Operculina hamiltonii
Ipomoea quinquefolia L. ^Merremia quinquefolia
Ipomoea reptans (L.) Poir. =Ipomoea aquatica
Ipomoea riedelii Meisn. ^Ipomoea batatoides
Ipomoea riparia G. Don =Ipomoea rubens
Ipomoea rubra (Vahl) Millsp. =Ipomoea setifera
Ipomoea sericantha Miq. =Operculina sericantha
Ipomoea solanifolia L. [misapplied name, not in flora area]
Ipomoea squamisepala O'Donell =Ipomoea angulata
Ipomoea stolonifera (Cirillo) J.F. Gmel. =Ipomoea imperati
Checklist of the Plants of the Guiana Shield 275
Ipomoea tamnifolia L. =Jacquemontia tamnifolia
Ipomoea trichocarpa auct. non Elliott 1817. =Ipomoea trifida
Ipomoea triloba auct. non L. 1753. =Ipomoea trifida
Ipomoea tuba (Schltdl.) G. Don =Ipomoea violacea
Ipomoea tuberosa L. [not in flora area] =Merremia tuberosa (L.) Rendle
Ipomoea villosa (Choisy) Meisn. =Ipomoea argentea
Ipomoea villosa var. argentea (Meisn.) Hallier f. =Ipomoea argentea
Jacquemontia azurea (Desr.) Choisy =Jacqiiemontia pentantha
Jacquemontia densiflora Rusby =Jacquemontia densiflora
Jacquemontia hirtiflora (M. Martens & Galeotti) O'Donell =Odonellia hirtiflora
Jacquemontia lactescens Seem. =Odonellia hirtiflora
Jacquemontia luxurians (Moric.) Hallier f. =Iseia luxurians
Jacquemontia nodiflora (Desr.) G. Don =Convolvulus nodiflorus
Jacquemontia rusbyana Standi. =Jacquemontia densiflora
Jacquemontia violacea (Vahl) Choisy =Jacquemontia pentantha
Jacquemontia violacea var. densiflora Meisn. =Jacquemontia densiflora
Kuhlmaniella holostyla (Ducke) Barroso =Dicranostyles holostyla
Lysiostyles cayennensis (Meisn.) Roberty =Maripa scandens
Lysiostyles epiphytica Pittier =Dicranostyles scandens
Lysiostyles pubescens Gleason =Lysiostyles scandens
Lysiostyles scandens var. latifolia Benth. =Lysiostyles scandens
Maripa cayennensis Meisn. =Maripa scandens
Maripa passifloroides Benth. ex Meisn. =Maripa violacea
Maripa rugosa Ducke =Maripa reticulata var. rugosa
Maripa spectabilis Choisy =Bonamia maripoides
Maripa volubilis Pittier =Odonellia hirtiflora
Merremia glabra (Aubl.) Hallier f. =Merremia macrocalyx
Merremia parviflora Pittier =Merremia quinquefolia
Mouroucoa violacea Aubl. =Maripa violacea
Murucoa cayennensis (Meisn.) Kuntze =Maripa scandens
Murucoa densiflora (Benth.) Kuntze =Maripa densiflora
Murucoa scandens (Aubl.) Kuntze =Maripa scandens
Operculina alata (Ham.) Urb. =Operculina hamiltonii
Operculina altissima (Mart, ex Choisy) Meisn. =Operculina hamiltonii
Operculina passifloroides (Benth. ex Meisn.) Ducke =Maripa violacea
Operculina pterodes (Choisy) Meisn. =Operculina hamiltonii
Operculina surinamensis Meisn. =Operculina sericantha
Operculina triquetra (Vahl) Hallier f. =Operculina turpethum
Operculina ventricosa Peter ^Operculina turpethum var. ventricosa
Operculina violacea (Aubl.) Barb. Rodr. =Maripa violacea
Pharbitis acuminatus (Vahl) Choisy =Ipomoea indica
Pharbitis cathartica (Poir.) Choisy =Ipomoea indica
Pharbitis mollis (Kunth) Choisy =Ipomoea indica
Prevostea amazonicus Choisy =Calycobolus glaber
Prevostea spectabilis (Choisy) Meisn. =Bonamia maripoides
Quamoclit coccinea var. hederifolia (L.) House =Ipomoea hederifolia
Quamoclit hederifolia (L.) G. Don =Ipomoea hederifolia
Quamoclit pinnata (Desr.) Bojer =Ipomoea quamoclit
Quamoclit quamoclit (L.) Britton =Ipomoea quamoclit
Quamoclit vulgaris Choisy ^Ipomoea quamoclit
Thyella tamnifolia (L.) Raf. =Jacquemontia tamnifolia
References
Austin, D.F. 1982. Convolvulaceae. In T. Lasser, Flora de Venezuela.
Austin, D.F. 1982. Convolvulaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 377-424.
Austin, D.F. 2002. Convolvulaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
231-236.
Austin, D.F. & P.B. Cavalcante. 1982. Convolvulaceas da Amazonia. Publ. Avulsas Mus. Paraense E. Goeldi 36: 1-134.
Austin, D.F. & G.W. Staples III. 1981. Convolvulaceae. In B. Maguire & collaborators, Botany of the Guayana Highland Part XI.
Mem. N.Y. Bot. Gard. 32: 1-391.
Ooststroom, S.J. van. 1932. Convolvulaceae. In A. Pulle, Flora of Suriname 4(1): 66-102.
Ooststroom, S.J. van. 1937. Convolvulaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 468-471.
276 Checklist of the Plants of the Guiana Shield
Crassulaceae
D.M. Keams
Kalanchoe crenata (Andrews) Haw. [cultivated or escaped from cultivation] GU
A:a/a«c/7oe/7/««a/a (Lam.) Pers. [cultivated, naturalized] AM BO GU SU FG
Synonyms and Misidentifications
BryophyUum calycinum Salisb. =Kalanchoe pinnata
Biyophyllum pinnatum (Lam.) Kurz [nom. Illeg.] =Kalanchoe pinnata
Cotyledon pinnata Lam. =Kalanchoe pinnata
Kalanchoe brasiliensis Cambess. =Kalanchoe crenata
References
Keams, D.M. 1998. Crassulaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 430-431,
Stoffers, A.L. 1984. Crassulaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname V(l): 573-575.
CUCURBITACEAE
C. Jeffrey, D.M. Keams, M. Nee
Cayaponia angustiloba (Cogn.) Cogn.
Cayaponia botryocarpa C. Jeffrey
Cayaponia coriacea Cogn.
Cayaponia cruegeri (Naudin) Cogn.
Cayaponia granaiensis Cogn.
Cayaponia guianensis C. Jeffrey
Cayaponiajenmanii C. Jeffrey
Cayaponia macrocalyx Harms [Sierra de la Neblina]
Cayaponia metensis Cuatrec.
Cayaponia ophthalmica R.E. Schult.
Cayaponia racemosa (Mill.) Cogn.
Cayaponia hgida (Cogn.) Cogn.
Cayaponia selysioides C. Jeffrey
Cayaponia sp. A
Cayaponia tessmannii Harms
Cayaponia triangularis (Cogn.) Cogn.
Ceratosanthes palmata (L.) Urb.
Citrullus lanatus (Thunb.) Matsum. & Nakai [cultivated, occasionally
escaping]
Coccinia grandis (L.) Voigt [cultivated, occasionally escaping]
Cucumis anguria L. [cultivated, naturalized]
Cuciimis anguria var. anguria
Cucumis melo L. [cultivated, occasionally escaping]
Cucurbita moschata Duchesne [cultivated, occasionally escaping]
Doyerea emetocathartica Grosourdy
Fevillea cordifolia L.
Fevillea trilobata L.
Gurania acuminata Cogn.
Gurania bignoniacea (Poepp. & Endl.) C. Jeffrey
Gurania brevipedunculata Cogn.
Gurania huberi Cogn.
Gurania lobata (L.) Pmski
Gurania nigrescens C. Jeffrey
Gurania oxyphylla C. Jeffrey
Gurania reticulata Cogn.
Gurania robusta Suess.
Gurania simplicifolia (Steyerm.) C. Jeffrey
Gurania spruceana Cogn.
Gurania subumbellata (Miq.) Cogn.
Gurania trialata Cogn.
Helmontia cardiophylla Harms
Helmontia leptantha (Schltdl.) Cogn.
Kedrostis foetidissima (Jacq.) Cogn. [naturalized?]
Lagenaria siceraria (Molina) Standi, [cultivated, occasionally escaping]
Lujfa cylindrica (L.) M. Roem. [cultivated, occasionally escaping]
Lujfa sepium (G. Mey.) C. Jeffrey
BO GU SU
BO
AM GU SU FG
AM BO DA GU SU FG
AM BO
AM BO DA GU
BO GU SU FG
BO
BO DA
DA GU SU FG
AM BO DA GU SU FG
GU SU FG
AM BO GU SU FG
AM? BO?
FG?
GU SU FG
BO GU SU FG
AM GU
GU
SU FG
FG
DA GU? SU FG
DA
GU? SU FG?
GU SU FG
BO
DA FG
FG?
AM BO GU SU FG
AM BO GU
GU
SU FG
FG
AM BO GU SU FG
AM BO
BO
DA GU
GU
SU FG
GU SU FG
FG
GU? SU FG
BO GU
AM
BO DA GU SU FG
AM
FG
AM BO GU SU
SU
FG
AM BO GU SU FG
AM BO DA GU SU FG
BO GU SU FG
Checklist of the Plants of the Guiana Shield 277
Melothria ciicumis Veil.
Melothria pendiila L.
Melothria trilobata Cogn.
Momordica charantia L. [cultivated, naturalized]
Posadaea sphaerocarpa Cogn.
Psigiiria racemosa C. Jeffrey
Psiguria triphylla (Miq.) C. Jeffrey
Psigiiria umbrosa (Kunth) C. Jeffrey
Rytidostylis amazonica (C. Mart, ex Cogn.) Spruce ex Kuntze
Selysia pnmifera (Poepp. & Endl.) Cogn.
Siolmatra pentaphylla Harms
Wilbrandia hibiscoides Silva Manso
Synonyms and Misidentifications
Anguria bignoniacea Poepp. & Endl. =Giirania bignoniacea
Anguria cissoides Benth. =Gurania bignoniacea
Anguria elliptica Britton =Psiguria triphylla
Anguria guianensis Klotzsch ex Schltdl. =Gurania bignoniacea
Anguria kegeliana Schltdl. =Gurania lobata
Anguria leptantha Schltdl. =Helmontia leptantha
Anguria leptantha var. brevipes Schltdl. =Helmontia leptantha
Anguria lobata L. =Gurania lobata
Anguria longispica Sagot [name of uncertain application fide C. Jeffrey]
Anguria multiflora Miq. =Gurania lobata
Anguria simplicifolia Steyerm. =Gurania simplicifolia
Anguria spinulosa Poepp. & Endl. =Gurania lobata
Anguria subumbellata Miq. =Gurania subumbellata
Anguria treslingiana PuUe =Psiguria triphylla
Anguria trilobata sensu Pulle non (L.) Jacq. =Gurania lobata
Anguria triphylla Miq. =Psiguria triphylla
Anguria umbrosa Kunth =Psiguria umbrosa
Anguria umbrosa auct. non Kunth =Psiguria triphylla
Anguria vogeliana Suess. ^Psiguria triphylla
Anguriopsis margaritensis J.R. Johnst. =Doyerea emetocathartica
Apodanthera spp. [no records verified for Guianas]
Arkezostis granatensis (Cogn.) Kuntze =Cayaponia granatensis
Bryonia guadalupensis Spreng. =Melothria pendula
Bryonia latebrosa Alton ^Cayaponia racemosa
Bryonia racemosa Mill. =Cayaponia racemosa
Cayaponia andreana Cogn. =Cayaponia cruegeri
Cayaponia bonplandii (Cogn.) Cogn. =Cayaponia coriacea
Cayaponia calycina Cogn. =Cayaponia cruegeri
Cayaponia latebrosa (Alton) Cogn. =Cayaponia racemosa
Cayaponia sclerocarpa Sagot [name of uncertain application fide C. Jeffrey]
Cayaponia simplicifolia (Naudin) Cogn. [not in flora area]
Ceratosanthes corniculata (Lam.) Cogn. =Ceratosanthes palmata
Ceratosanthes latiloba var. acutiloba Cogn. ^Ceratosanthes palmata
Ceratosanthes tuberosa J.F. Gmel. =Ceratosanthes palmata
Citrullus vulgaris Schrad. ex Eckl. & Zeyh. =Citrullus lanatus
Corallocarpus emetocatharticus (Grosourdy) Cogn. =Doyerea emetocathartica
Cucumis sepium G Mey. =Luffa sepium
Cucurbita lagenaria L. =Lagenaria siceraria
Cucurbita siceraria Molina =Lagenaria siceraria
Doyerea angosturensis Grosourdy =Doyerea emetocathartica
Elaterium amazonicum Mart, ex Cogn. =Rytidostylis amazonica
Elaterium carthaginense Jacq. [not in flora area] =Rytidostylis carthaginensis (Jacq.) Kuntze
Fevilleajavilla Kunth =Fevillea cordifolia
Fevillea karstenii Cogn. =Fevillea cordifolia
Gurania breviflora Cogn. =Gurania bignoniacea
Gurania cissoides (Benth.) Cogn. =Gurania bignoniacea
Gurania diffusa Cogn. =Gurania huberi
Gurania diversifolia Cogn. =Gurania acuminata
Gurania durandii Cogn. =Gurania lobata
Gurania guianensis (Klotzsch ex Schltdl.) Cogn. =Gurania bignoniacea
Gurania kegeliana (Schltdl.) Cogn. =Gurania lobata
GU?
AM DA GU su FG
AM BO DA GU su
AM BO DA GU su EG
AM DA GU
GU?
su FG
AM BO
BO
DA GU su FG
AM BO FG
GU? su FG
GU
GU?
278 Checklist of the Plants of the Guiana Shield
Gurania klotzschiana Cogn. =Gurania subumbellata
Gurania levyana var. glabra Cogn. =Gurania robusta
Gurania multiflora (Miq.) Cogn. =Gurania lobata
Gurania sagotiana Cogn. =Gurania reticulata
Gurania spimdosa (Poepp. & Endl.) Cogn. =Gurania lobata
Helmontia simplicifolia Cogn. ^Helmontia leptantha
Lagenaria lagenaria (L.) Cockerell =Lagenaria siceraria
Lagenaria vulgaris Sen =Lagenaria siceraria
Luffa aegyptiaca Mill. =Luffa cylindrica
Luffa operculata (L.) Cogn. [in S. Am. Records =1. sepium. L. operculata valid taxon for Cent. Am. & Mex. Species] =Luffa
sepium
Luffa purgans (Mart.) Mart. =Luffa sepium
Melothriafluminensis Gardner =Melothria pendula
Melothria guadalupensis (Spreng.) Cogn. =Melothria pendula
Melothria heteroloba Pittier =Melothria trilobata
Momordica balsamina L. [dubious, probably based on misidentification fide C. Jeffrey]
Momordica cylindrica L. =Luffa cylindrica
Momordica lanata Thunb. =Citrullus lanatus
Momordica operculata L. =Luffa sepium
Momordica purgans Mart. =Lujfa sepium
Nhandiroba cordifolia (L.) Kuntze =Fevillea cordifolia
Perianthopodus bonplandii Cogn. =Cayaponia coriacea
Rytidostylis carthaginensis (Jacq.) Kuntze [not in flora area]
Trianosperm rigida Cogn. =Cayaponia rigida
Trianosperma angustiloba Cogn. =Cayaponia angustiloba
Trianosperma cruegeri Naudin =Cayaponia cruegeri
Trianosperma triangularis Cogn. =Cayaponia triangularis
Trichosanthes corniculata Lam. =Ceratosanthes palmata
Trichosanthes palmata L. =Ceratosanthes palmata
References
Jeffrey, C. 1978. Further notes on Cucurbitaceae: IV. Some New World taxa. Kew Bull. 33: 347-380.
Jeffrey, C. 1984. Cucurbitaceae. In A.L. StotTers & J.C. Lindeman, Flora of Suriname 5(1): 457-518.
Jeffrey, C. & B. Trujillo. 1992. Cucurbitaceae. In G. Morillo, Flora de Venezuela 5(1): 11-201.
Keams, D.M. 1998. Cucurbitaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 431-461.
Nee, M. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hofif, & J.D. Mitchell, eds. 2002. Guide to
the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 236-246.
CUNONIACEAE
R Berry, J.C. Bradford
Weinmannia balbisiana Kunth
Weinmannia brachystachya Willd. ex Engl, [expected in GU][Mt. Roraima]
Weinmannia corocoroensis J.C. Bradford & RE. Berry
Weinmannia elliptica Kunth
Weinmanniafagaroides Kunth [Mt. Roraima]
Weinmannia glabra L. f. [Mt. Roraima]
Weinmannia guyanensis Klotzsch ex Engl.
Weinmannia guyanensis var. guyanensis [Mt. Roraima]
Weinmannia ilutepuiensis RE. Berry & J.C. Bradford
Weinmannia laxiramea Killip & A.C. Sm. [Mt. Roraima]
Weinmannia pubescens Kunth
Weinmannia sorbifolia Kunth
Weinmannia sorbifolia var. sorbifolia
Weinmannia sp. A [Distinct from Weinmanniafagaroides]
Weinmannia velutina O.C. Schmidt
Synonyms and Misidentifications
Weinmannia baccariniana var. caracasana Pamp. =Weinmanniafagaroides
Weinmannia balbisiana var. moritzii (Engl.) Cuatrec. =Weinmannia balbisiana
Weinmannia balbisiana war. ptariana Cuatrec. =Weinmannia elliptica
Weinmannia balbisiana var. roraimensis Bemardi =Weinmannia elliptica
Weinmannia cardonae Cuatrec. =Weinmannia elliptica
Weinmannia caripensis Kunth =Weinmannia glabra
AM BO
BO
GU'
AM
AM BO
AM BO GU
AM BO GU
BO GU
BO GU
BO
BO GU
AM
AM
AJVl
AM
AM BO GU
Checklist of the Plants of the Guiana Shield 279
Weinmannia glabra var. caripensis (Kunth) Engl. =Weinmanma glabra
Weinmannia microphylla Kunth =Weinmanriia brachystachya
Weinmannia microphylla var. caracasana (Pamp.) Cuatrec. =Weinmanniafagaroides
Weinmannia microphylla war. fagaroides (Kunth) Cuatrec. =Weinmanniafagaroides
Weinmannia moritzii Engl. =Weinmannia balbisiana
Weinmannia neblinensis Maguire & Steyerm. ^Weinmannia pubescens
Weinmannia ovalis var. elliptica (Kunth) Cuatrec. =Weinmannia elliptica
Weinmannia ovalis var. roraimensis Pamp. =Weinmannia elliptica
Weinmannia pinnata L. =Weinmannia glabra
Weinmannia pinnata var. caripensis (Kunth) Cuatrec. =Weinmannia glabra
Weinmannia pinnata var. ptaritepuiana Cuatrec. =Weinmannia glabra
Weinmannia roraimensis Cuatrec. =Weinmanniafagaroides
Weinmannia steyermarkii Cuatrec. =Weinmannia velutina
Weinmannia tamana Cuatrec. =Weinmanniafagaroides
Weinmannia velutina f. steyermarkii (Cuatrec.) Bemardi =Weinmanma velutina
References
Bradford, J.C. & P.E. Berry. 1998. Cunoniaceae. In J.A. Steyermark, P.E. Berry & B.K. Hoist, Flora of the Venezuelan Guayana
4: 462-469.
Killip, E.P. & A.C. Smith. 1929. The genus Weinmannia in northern South America. Bull. Torrey Bot. Club 56: 361-377.
CUSCUTACEAE
D.F. Austin
Cuscuta americana L.
Cuscuta campestris Yunck.
Cuscuta costaricensis Yunck.
Cuscuta indecora Choisy
Cuscuta umbellata Kunth
BO
BO
BO
su
GU SU
References
Austin, D.F. 1982. Convolvulaceae. In Flora de Venezuela.
Austin, D.F. 1998. Cuscutaceae. In J.A. Steyermark, P.E. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 469-470.
Yuncker, T.G 1923. Revision of the South American species of Cuscuta. I. Am. J. Bot. 9: 557-575; II. Am. J. Bot. 10: 1-17.
Cyrilla racemiflora L.
Purdiaea nutans Planch.
Cyrillaceae
J.S. Miller
AM BO
AM BO
GU SU FG
Synonyms and Misidentifications
Alloiosepalum weberbaueri Gilg =Purdiaea nutans
Cyrilla antillana Michx. =CyriUa racemiflora
Cyrilla brevifolia N.E. Br. =Cyrilla racemiflora
Cyrilla perrottetii Briq. =Cyrilla racemiflora
Cyrilla racemiflora var. brevifolia (N.E. Br.) Maguire & Wurdack ex Steyerm. =Cyrilla racemiflora
Itea cyrilla L'Her. =Cyrilla racemiflora
Purdiaea weberbaueri (Gilg) Mattick =Purdiaea nutans
References
Miller, J.S. 1998. Cyrillaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 663-665.
Thomas, J.L. 1960. A monographic study of the Cyrillaceae. Contr. Gray Herb. 186: 1-114.
DiCHAPETALACEAE
G.T. Prance
Dichapetalum froesii Prance
Dichapetalum latifolium Baill.
Dichapetalum odoratum Baill.
Dichapetalum pedunculatum (DC.) Baill.
Dichapetalum rugosum (Vahl) Prance
Dichapetalum schulzii Prance
Tapura amazonica Poepp.
AM
AM
AM BO
AM BO
BO
AM
GU SU FG
GU SU
SU
FG
GU SU FG
280 Checklist of the Plants of the Guiana Shield
AM GU SU FG
AM BO DA GU su FG
AM BO DA GU SU FG
FG
Tapura amazonica van amazonica
Tapura capitulifera Spruce ex Baill.
Tapura guianensis Aubl.
Tapura singularis Ducke
SyMONYMS and MiSIDENTlFlCATlONS
Chailletiaflavicans Klotzsch =Dichapetalum rugosum
Chailletia parkeri Planch, ex Triana & Planch, [nomen nudum] =Dichapetalum rugosum
Chailletia pedunculata DC. =Dichapetalum pedunculatum
Chailletia sessiliflora DC. =Tapura guianensis
Chailletia vestita Benth. =Dichapetaliim rugosum
Dichapetalum amazonicum K. Krause =Dichapetalum latifolium
Dichapetalum flavicans (Klotzsch) Engl. =Dichapetalum rugosum
Dichapetalum glabrum (Vahl) Prance =Dichapetalum pedunculatum
Dichapetalum scandens (Poir.) I.M. Johnst. =Dichapetalum rugosum
Dichapetalum scandens var. cinerascens (Baill.) J.F. Macbr. =Dichapetalum latifolium
Dichapetalum vestitum (Benth.) Baill. ^Dichapetalum rugosum
Dichapetalum vestitum var. cinerascens Baill. =Dichapetalum latifolium
Rohria petioliflora Willd. =Tapura guianensis
Rohria schreberi J.F. Gmel. =Tapura guianensis
Rohria tapura J.F. Gmel. =Tapura guianensis
Symphyllanthus glaber Vahl =Dichapetalum pedunculatum
Symphyllanthus rugosus Vahl =Dichapetalum rugosum
Tapura amazonica var. ciliata (Gardner) Baill. =Tapura amazonica
Tapura amazonica var. cuspidata Baill. -Tapura amazonica
Tapura amazonica var. dasyphylla Baill. =Tapura amazonica
Tapura amazonica var. sublanceolata Baill. =Tapura amazonica
Tapura ciliata Gardner =Tapura amazonica
Tapura cucuUata Benth. =Tapura guianensis
Tapura negrensis Suess. =Tapura guianensis
References
Prance, G.T. 1972. Dichapetalaceae. Flora Neotropica Monogr. 10: 1-84.
Prance, G.T. 1979. Anew species of Dichapetalum from Suriname. Bull. Torrey Bot. Club 106(4): 309-312.
Prance, G.T. 1983. Additions to Neotropical Dichapetalaceae. Brittonia 35(1): 49-54.
Prance, G.T. 1998. Dichapetalaceae. In J.A. Steyermark, P.E. Berry & B.K. Hoist, Flora of the Venezuelan Guayana4: 666-671.
Prance, G.T. 2002. Dichapetalaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
247-249.
Stafleu, F.A. 1951. Dichapetalaceae. In A. Pulle, Flora of Suriname 3(2): 1 66- 1 72.
DiLLENIACEAE
G.A. Aymard C, C.L. Kelloff
Curatella americana L.
Davilla alata (Vent.) Briq.
Davilla kunthii A. St.-Hil.
Davilla lacunosa Mart.
Davilla nitida (Vahl) Kubitzki
Davilla rugosa Poir.
Davilla rugosa var. rugosa
Davilla steyermarkii Kubitzki [expected in GU]
Doliocarpus amazonicus Sleumer
Doliocarpus amazonicus subsp. amazonicus
Doliocarpus areolatus Kubitzki
Doliocarpus brevipedicellatus Garcke
Doliocarpus brevipedicellatus subsp. brevipedicellatus
Doliocarpus carnevaliorum Aymard
Doliocarpus dentatus (Aubl.) Standi.
Doliocarpus dentatus subsp. dentatus
Doliocarpus dentatus subsp. esmeraldae (Steyerm.) Kubitzki
Doliocarpus dentatus suhsp. ferrugineus (Rusby) Kubitzki
Doliocarpus dentatus subsp. lanceolatus Kubitzki
Doliocarpus dentatus subsp. latifolius Kubitzki
AM BO DA GU SU FG
GU SU FG
AM BO DA GU SU FG
FG
AM BO GU SU FG
AM BO DA GU SU FG
AM BO
BO
DA GU SU
SU
SU
FG
AM
AM BO DA GU SU FG
AM BO DA GU SU FG
AM
AM BO DA GU SU FG
AM BO DA GU SU FG
AM BO GU SU FG
AM
AM
AM BO
Checklist of the Plants of the Guiana Shield 281
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO DA
BO
BO
Doliocarpus dentatus subsp. obovatus Aymard
Doliocarpus dentatus subsp. undulatus (Eichler) Kubitzki
Doliocarpus gracilis Kubitzki
Doliocarpus guianensis (Aubl.) Gilg
Doliocarpus hispidobaccatus Aymard
Doliocarpus leiophyllus Kubitzki
Doliocarpus liesneri Aymard
Doliocarpus macrocarpus Mart, ex Eichler
Doliocarpus major J.F. Gmel.
Doliocarpus major subsp. major
Doliocarpus multiflorus Standi.
Doliocarpus novogranatensis Kubitzki
Doliocarpus ortegae Aymard
Doliocarpus paracusis Sleumer
Doliocarpus paucinervis Kubitzki
Doliocarpus prancei Kubitzki
Doliocarpus sagotianus Kubitzki
Doliocarpus savannarum Sandwith
Doliocarpus spraguei Cheesman
Doliocarpus verruculosus Kubitzki '
Neodillenia venezuelana Aymard
Pinzona coriacea Mart. & Zucc.
Tetracera asperula Miq.
Tetracera costata Mart, ex Eichler
Tetracera costata subsp. costata
Tetracera costata subsp. rotundifolia (Sm.) Kubitzki
Tetracera maguirei Aymard & B.M. Boom
Tetracera surinamensis Miq.
Tetracera tigarea DC.
Tetracera volubilis L.
Tetracera volubilis subsp. volubilis
Tetracera willdenowiana Steud.
Tetracera willdenowiana subsp. emarginata Kubitzki
Tetracera willdenowiana subsp. willdenowiana
Synonyms and Misidentifications
Calinea scandens Aubl. =Doliocarpus spraguei
Curatella americana var. pentagyna Donn. Sm. =Curatella americana
Curatella cambaiba A. St.-Hil. =Curatella americana
Curatella grisebachiana Eichler =Curatella americana
Davilla aspera (Aubl.) Benoist =Davilla kunthii
Davilla multiflora (DC.) A. St.-Hil. =Davilla nitida
Davilla pilosa Miq. =Davilla rugosa
Davilla surinamensis Miq. ^Davilla kunthii
Davilla tenuis Eichler =Davilla kunthii
Davilla wormiaefolia Baill. =Davilla alata
Doliocarpus belizensis Lundell =Pinzona coriacea
Doliocarpus brevipedicellatus var. densiflorus Eichler =Doliocarpus dentatus subsp. undulatus
Doliocarpus calinea J.F. Gmel. =Doliocarpus spraguei
Doliocarpus coriaceus (Mart. & Zucc.) Gilg =Pinzona coriacea
Doliocarpus densiflorus (Eichler) Steyerm. =Doliocarpus dentatus subsp. undulatus
Doliocarpus esmeraldae Steyerm. =Doliocarpus dentatus subsp. esmeraldae
Doliocarpus ferrugineus Rusby =Doliocarpus dentatus suhsp. ferrugineus
Doliocarpus ptariensis Steyerm. =Doliocarpus savannarum
Doliocarpus rolandri J.F. Gmel. =Doliocarpus major
Doliocarpus scandens Poir. =Doliocarpus major
Doliocarpus scandens (Aubl.) Gilg =Doliocarpus spraguei
Doliocarpus surinamensis Lanj. =Doliocarpus paracusis
Doliocarpus undulatus Eichler =Doliocarpus dentatus subsp. undulatus
Doliocarpus virgatus Sagot =Doliocarpus macrocarpus
Soramia guianensis Aubl. =Doliocarpus guianensis
Tetracera aspera (Aubl.) Willd. =Tetracera tigarea
Tetracera multiflora DC. =Davilla nitida
Tetracera nitida Vahl =Davilla nitida
GU
SU
su
GU SU
FG
FG
GU SU FG
GU SU FG
GU SU FG
FG
FG
AM BO GU SU FG
AM BO GU SU FG
AM
AM
AM BO DA GU SU FG
GU SU FG
BO GU SU FG
BO GU SU FG
BO GU
GU
FG
AM DA GU SU FG
BO DA GU SU FG
BO DA GU SU FG
BO DA GU SU FG
AM BO GU SU FG
AM FG
AM BO GU SU
282 Checklist of the Plants of the Guiana Shield
Tetracera ovalifolia auct. non DC. =Tetracera asperula
Tetracera ovalifolia DC. =Tetracera costata subsp. rotundifolia
Tetracera rotundifolia auct. non Sm. =Tetracera asperula
Tetracera rotundifolia Sm. =Tetracera costata subsp. rotundifolia
Tetracera trinitensis Sprague & R.O. Williams =Tetracera costata
Tigarea aspera Aubl. =Tetracera tigarea
Tigarea dentata Aubl. =Doliocarpus dentatus
References
Aymard, G. 1998. Dilleniaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 671-685.
Aymard, G. & J. Miller. 1994. Dilleniaceae novae Neotropicae III. Sinopsis y adiciones a las Dilleniaceae del Peru. Candollea
49(1): 169-182.
Aymard, G. & S.A. Mori. 2002. Dilleniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 250-253.
Kubitzki, K. 1970. Die gattung Tetracera. Mitt. Bot. Munch. 8: 1-98.
Kubitzki, K. 1971. Doliocarpus, Davilla uhd verwandte gattungen. Mitt. Bot. Miinch. 9: 1-105.
Lanjouw, J. & RF. van Heerdt. 1941. Dilleniaceae. In A. Pulle, Flora of Suriname 3(1): 386-408.
Droseraceae
A. Culham, R.D. de Stefano
Drosera arenicola Steyerm.
Drosera biflora Willd. ex Roem. & Schult.
Drosera capillaris Poir.
Drosera cayennensis Sagot ex Diels
Drosera communis A. St.-Hil.
Drosera esmeraldae (Steyerm.) Maguire & Wurdack
Drosera felix Steyerm. & L.B. Sm.
Drosera hirticalyx Duno & Culham
Drosera intermedia Hayne
Drosera kaieteurensis Brumm.-Ding.
Drosera meristocaulis Maguire & Wurdack
Drosera roraimae (Klotzsch ex Diels) Maguire «fe J.R. Laundon
Drosera sessilifolia A. St.-Hil.
Drosera yutajensis Duno & Culham
Synonyms and Misidentifications
Drosera americana Willd. =Drosera intermedia
Drosera arenicola var. occidentalis Maguire & Wurdack =Drosera arenicola
Drosera colombiana A. Fernandez ^Drosera cayennensis
Drosera dentata Benth. =Drosera sessilifolia
Drosera montana var. robusta Diels =Drosera roraimae
Drosera montana var. roraimae Klotzsch ex Diels =Drosera roraimae
Drosera panamensis M.D. Correa & A.S. Taylor =Drosera cayennensis
Drosera pusilla Kunth =Drosera biflora
Drosera sanariapoana Steyerm. =Drosera cayennensis
Drosera tenella Willd. ex Roem. & Schult. =Drosera capillaris
Drosera tenella var. esmeraldae Steyerm. =Drosera esmeraldae
References
Brummer-Dinger, C.A. 1955. Notes on Guiana Droseraceae. Acta Bot. Neerl. 4(1): 136-138.
Duno de Stefano, R. & A. Culham. 1998. Droseraceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan
Guayana 4: 697-703.
Maguire, B. & J.J. Wurdack. 1957. Droseraceae. In B. Maguire, J.J. Wurdack, & collaborators. Botany of the Guayana Highland
part 2. Mem. New York Bot. Gard. 9(3): 331-336.
AM BO
AM GU SU
AM BO GU SU FG
AM BO GU FG
AM BO
AM
BO
GU
AM
BO GU SU
BO GU
AM
AM BO GU
AM BO GU
AM
Ebenaceae
RE. Berry, C. Sothers
Diospyros artanthifolia Mart, ex Miq.
Diospyros capreifolia Mart, ex Hiem
Diospyros carbonaria Benoist
Diospyros cauliflora Mart.
AM
BO DA
GU SU FG
GU SU FG
FG
FG
GU SU FG
GU? FG
GU
GU SU FG
GU SU FG
SU FG
GU SU FG
Checklist of the Plants of the Guiana Shield 283
Diospyros cauligera F. White AM
Diospyros cavalcantei Sothers FG
Diospyros cayennensis A. DC. AM BO DA GU SU? FG
Diospyros dichroa Sandwith GU SU FG
Diospyros discolor Willd. [cultivated; naturalized?] FG
Diospyros duckei Sandwith FG?
Diospyros glomerata Spruce ex Hiem AM FG
Diospyros guianensis (Aubl.) Giirke AM BO DA
Diospyros guianensis subsp. akaraiensis (A.C. Sm.) F. White
Diospyros guianensis subsp. coriacea F. White BO
Diospyros guianensis subsp. guianensis AM DA
Diospyros inconstans Jacq. BO
Diospyros lissocarpoides Sandwith BO DA
Diospyros martinii Benoist ex Amshoff
Diospyros matheriana A.C. Sm.
Diospyros poeppigiana A. DC. AM
Diospyros ropourea B. Walln. FG
Diospyros santaremnensis Sandwith [perhaps expected in FG]
Diospyros sericea A. DC. GU
Diospyros sp. A i BO
Diospyros spruce i Hiem AM
Diospyros tenuiflora A.C. Sm. AM
Diospyros tetrandra Hiern SU FG
Diospyros vestita Benoist BO SU FG
Synonyms and Misidentifications
Diospyros akaraiensis A.C. Sm. =Diospyros guianensis subsp. akaraiensis
Diospyros anzoateguiensis Steyerm. =Diospyros inconstans
Diospyros delgadoi Standi. =Diospyros inconstans
Diospyros emarginata Hiem =Diospyros poeppigiana
Diospyros ierensis Britton =Diospyros cayennensis
Diospyros liriosmoides A.C. Sm. ^Diospyros poeppigiana
Diospyros longifolia (Spruce ex Engl.) Sleumer & F. White =Diospyros tenuiflora
Diospyros melinonii (Hiem) A.C. Sm. =Diospyros capreifolia
Diospyros paralea Steud. =Diospyros guianensis
Diospyros polyandra Spruce ex Hiem =Diospyros artanthifolia
Diospyros praetermissa Sandwith -Diospyros vestita
Diospyros subrotata Hiern =Diospyros artanthifolia
Maba inconstans (Jacq.) Griseb. =Diospyros inconstans
Maba mellinoni Hiem =Diospyros capreifolia
Paralea guianensis Aubl. =Diospyros guianensis
Ropourea guianensis Aubl. =Diospyros ropourea
References
Cavalcante, P.B. 1963. Contribu9ao ao conhecimento do genero Diospyros Dalech. na Amazonia. Bol. Mus. Goeldi n.s. 20: 1-
53.
Sothers, C. & RE. Berry. 1998. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 704-712.
Wallnofer, B. 1999. Neue Diospyros-Arten (Ebenaceae) aus Siidamerika. Ann. Naturhist. Mus. Wien, B, 101: 565-592.
Wallnofer, B. 2000. Neue Diospyros-Arten (Ebenaceae) aus Sudamerika - II. Ann. Naturhist. Mus. Wien, B, 102: 417-433.
Wallnofer, B. 2001. Lectotypification of Diospyros cayennensis A. DC. (Ebenaceae). Taxon 50: 887-889 (see Erratum in Taxon
50(4): 1319).
Wallnofer, B. 2001. The biology and systematics of Ebenaceae: a review. Ann Naturhist. Mus. Wein 103B: 485-512.
Wallnofer, B. 2003. Anew species of Diospyros from southwestem Amazonia. Ann. Naturhist. Mus. Wien, B, 104: 563-566.
Wallnofer, B. 2004. Ebenaceae. In: Kubitzki, K. (ed.): The families and genera of vascular plants. Vol. 6: 125-130. Springer
Verlag, Berlin.
Wallnofer, B. 2004. Lissocarpaceae. In: Kubitzki, K. (ed.): The families and genera of vascular plants. Vol. 6: 236-238. Springer
Verlag, Berlin.
Wallnofer, B. 2004. A revision of Lissocarpa Benth. (Ebenaceae subfam. Lissocarpoideae (Gilg in Engler) B. Walln.). Ann.
Naturhist. Mus. Wien, B, 105: 515-564.
Wallnofer, B. & S.A. Mori. 2002. Ebenaceae. In Mori S.A., Cremers G, Gracie C.A., Granville J.-J. de, Heald S.V, Hoff M. &
Mitchell J.D., eds: Guide to the vascular plants of central French Guiana. Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 254-257.
White, F. 1981. Ebenaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 1 1 . Mem. New York Bot. Gard.
32: 1-391.
284 Checklist of the Plants of the Guiana Shield
Elaeocarpaceae
L.J. Dorr, D.A. Smith, J.A. Steyermarkt
Sloanea bolivarensis Steyerm.
Sloanea brevipes Benth.
Sloanea carrenoi Steyerm.
Sloanea cataniapensis Steyerm.
Sloanea caudata Steyerm.
Sloanea cavicola Steyerm.
Sloanea crassifolia Earle Sm.
Sloanea davidsei Steyerm.
Sloanea durissima Spruce ex Benth.
Sloanea echinocarpa Uittien
Sloanea eichleri K. Schum.
Sloaneafloribunda Spruce ex Benth.
Sloanea garckeana K. Schum.
Sloanea gracilis Uittien
Sloanea grandiflora Sm.
Sloanea guianensis (Aubl.) Benth.
Sloanea latifolia (Rich.) K. Schum.
Sloanea lauhfolia (Benth.) Benth.
Sloanea laxiflora Spruce ex Benth.
Sloanea longiahstata Steyerm.
Sloanea macrophylla Benth. ex Turcz.
Sloanea megacarpa Steyerm. & Marc.-Berti
Sloanea merevariensis Pittier ex Steyerm.
Sloanea nitida G. Don
Sloanea obtusifolia (Moric.) K. Schum.
Sloanea parviflora Planch, ex Benth.
Sloanea parvifructa Steyerm.
Sloanea picapica Standi.
Sloanea picapica var. hirsuta C.E. Sm.
Sloanea pittieriana Steyerm.
Sloanea ptariana Steyerm.
Sloanea piibescens Benth.
Sloanea robusta Uittien
Sloanea rufa Planch, ex Benth.
Sloanea schomburgkii Spruce ex Benth.
Sloanea sinemariensis Aubl.
Sloanea sipapoana Steyerm.
Sloanea sp. A
Sloanea sp. B [Sierra de la Neblina]
Sloanea sp. C
Sloanea sp. D
Sloanea spathulata Earle Sm.
Sloanea steyermarkii Earle Sm.
Sloanea steyermarkii subsp. autanae (Steyerm.) Steyerm.
Sloanea steyermarkii swbsp.jauaensis (Steyerm.) Steyerm.
Sloanea steyermarkii subsp. steyermarkii
Sloanea stipitata Spruce ex Benth.
Sloanea subpsilocarpa Steyerm.
Sloanea synandra Spruce ex Benth.
Sloanea terniflora (M09. & Sesse ex DC.) Standi.
Sloanea trichosticha R.O. Williams & Sandwith
Sloanea tuerckheimii Donn. Sm.
Sloanea venezuelana Steyerm.
Sloanea wurdackii Steyerm.
Synonyms and Misidentifications
Ablania guianensis Aubl. =Sloanea guianensis
Adenobasium obtusifolium Moric. =Sloanea obtusifolia
Courimari guianensis Aubl. [name of uncertain application] =Sloanea sp.
Dasynema laurifolium Benth. =Sloanea laurifolia
Dasynema pubescens Poepp. =Sloanea pubescens
BO
AM
BO
GU su? FG
AM
BO
BO
BO GU
AM
GU
GU su EG
BO GU su FG
AM
GU su
su
FG
BO DA GU su FG
AM BO DA GU
GU
su FG
FG
AM GU FG?
AM BO GU su FG
AM
AM
BO
DA
GU
GU
su
FG?
FG
AM DA FG?
GU su FG
AM
BO
BO
GU
GU
AM BO su? FG
AM BO GU su
AM GU
GU
su FG
FG
AM
AM
BO
GU su FG
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO DA
AM
AM
BO GU
GU
su
FG
FG
AM GU su FG
BO
AM BO
Checklist of the Plants of the Guiana Shield 285
Sloanea aciitiflora Uittien =Sloanea laxiflora
Sloanea amplifrons Johnst. =Sloanea grandiflora
Sloanea autanae Steyerm. =Sloanea steyermarkii subsp. autanae
Sloanea brachytepala Ducke [considered distinct by D.A. Sm. and Steyerm., in AM] =Sloanea laurifolia
Sloanea bracteosa Ducke =Sloanea nitida
Sloanea breviseta Steyerm. =Sloanea guianensis
Sloanea corymbiflora DC. =Sloanea latifolia
Sloanea dentata L. [not in flora area]
Sloanea eriostemon Sprague & L. Riley =Sloanea tuerckheimii
Sloaneajauaensis Steyerm. =Sloanea steyermarkii subsp. jauaensis
Sloaneajauaensis van minor Steyerm. =Sloanea steyermarkii subsp. jauaensis
Sloanea maroana Steyerm. =Sloaneafloribunda
Sloanea massoni auct. non Sw. 1788: sensu Pittier et al. =Sloanea macrophylla
Sloanea obtusa (Splitg.) K. Sebum. =Sloanea parviflora
Sloanea oppositifolia Spruce ex Benth. =Sloanea laurifolia
Sloanea paniculata Spruce ex Benth. =Sloaneafloribunda
Sloanea sinemariensis auct. non Aubl. sensu Pittier et al. (as Sloanea surinamensis) =Sloanea eichleri
Sloanea usurpatrix Sprague & L. Riley =Sloanea tuerckheimii
Sloanea yapacanae Steyerm. =Sloanea steyermarkii subsp. jauaensis
References
Heald, S.V., A. Vicentini, & D.A. Smith. 2002. Elaeocarpaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V.
Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons.
Mem. New York Bot. Gard. 76 (2): 258-261.
Jansen-Jacobs, M.J. 1986. Elaeocarpaceae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 3(1-
2): 301-305.
Smith, C.E., Jr. 1954. The New World species of Sloanea. Contr. Gray Herb. 175: 1-114.
Smith, D.A. & J.A. Steyermark. 1998. Elaeocarpaceae. In J.A. Steyermark, P.E. Berry & B.K. Hoist, Flora of the Venezuelan
Guayana4: 712-730.
Uittien, H. 1932. Elaeocarpaceae. In A. Pulle, Flora of Suriname 3(1): 58-64.
Eremolepidaceae
J. Kuijt
Antidaphne antidaphneoides (Rizzini) Kuijt BO
Antidaphne viscoidea Poepp. & Endl. BO GU
Eubrachion gracile Kuijt BO
Synonyms and Misidentifications
Antidaphnefendleri (Tiegh.) Engl. =Antidaphne viscoidea
Antidaphne fendleri var. pseudoviscoidea Rizzini =Antidaphne viscoidea
Antidaphne orbicularis Kuijt =Antidaphne viscoidea
Antidaphne viscoidea var. alpestris Rizzini =Antidaphne viscoidea
Eremolepis mirocalyx Ernst =Antidaphne viscoidea
Ixidium antidaphneoides Rizzini =Antidaphne antidaphneoides
Stachyphyllum fendleri Tiegh. =Antidaphne viscoidea
References
Kuijt, J. 1988. Monograph of the Eremolepidaceae. Syst. Bot. Monogr. 18: 1-60.
Kuijt, J. 1988. Eremolepidaceae. In J.A. Steyermark, P.E. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 731-734.
Ericaceae
J.L. Luteyn
Agarista duckei (Huber) Judd [expected in GU]
Bejaria aestuans L. [primarily Andean taxon]
Bejaria imthurnii 'N.E. Br. [Mt. Roraima]
Bejaria neblinensis Maguire, Steyerm. & Luteyn
Bejaria sprucei Meisn.
Cavendishia callista Donn. Sm.
Cavendishia neblinae Maguire, Steyerm. & Luteyn
Cavendishia aff pubescens (Kunth) Hemsl.
Ceratostema glandulifera Maguire, Steyerm. & Luteyn
Disterigma cuspidatum (Planch, ex Wedd.) Nied.
AM BO
AM BO
BO
AM
AM BO
AM BO
AM
AM
GU?
GU
GU
GU
su FG
AM
286 Checklist of the Plants of the Guiana Shield
Disterigma hiimboldtii (Klotzsch) Nied.
Gaiiltheria alnifolia (Dunal) A.C. Sm.
Gauhheria alnifolia var. alnifolia
Gaitltheria erecta Vent. [Mt. Roraima]
Gaiiltheria setiilosa^.E. Br. [Mt. Roraima]
Gaylussacia buxifolia Kunth
Ledothamnus atroadenus Maguire, Steyerm. & Luteyn
Ledothamniis decumbem Maguire, Steyerm. &. Luteyn
Ledothamnus guyanensis Meisn. [Mt. Roraima]
Ledothamnus jauaensis Maguire, Steyerm. & Luteyn
Ledothamnus luteus Maguire, Steyerm. & Luteyn
Ledothamnus parviflorus Gleason
Ledothamnus sessiliflonis N.E. Br. [Mt. Roraima]
Mycerinus chimantensis Maguire, Steyerm. & Luteyn
Mycerinus sclewphyllus A.C. Sm.
Mycerinus viridiflorus Steyerm. & Magujre
Notopora auyantepuiensis Steyerm.
Notopora cardonae A.C. Sm.
Notopora chimantensis Steyerm. & Maguire
Notopora schomburgkii Hook. f.
Notopora smithiana Steyerm. & Maguire
Orthaea apophysata (Griseb.) A.C. Sm.
Orthaea crinita A.C. Sm.
Orthaea hispida A.C. Sm.
Orthaea merumensis Maguire, Steyerm. & Luteyn
Orthaea paruensis Maguire, Steyerm. & Luteyn
Orthaea thibaiidioides Maguire, Steyerm. & Luteyn
Orthaea venamensis Maguire, Steyerm. & Luteyn
Orthaea wurdackii Maguire, Steyerm. & Luteyn
Pernettya marginata N.E. Br. [Mt. Roraima]
Psammisia guianensis Klotzsch
Psammisia hookeriana Klotzsch
Psammisia urichiana (Britton) A.C. Sm.
Satyria carnosiflora Lanj.
Satyria cerander (Dunal) A.C. Sm.
Satyria panui-ensis (Benth. ex Meisn.) Benth. & Hook. f. ex Nied.
Sphyrospermum buxifolium Poepp. & Endl.
Sphyrospermum cordifolium Benth.
Sphyrospermum ellipticum Sleumer
Sphyrospermum klotzschianum (Mansf.) A.C. Sm.
Tepuia cardonae A.C. Sm.
Tepuia intermedia Steyerm.
Tepuia multiglandulosa Steyerm. & Maguire
Tepuia speciosa A.C. Sm.
Tepuia tatei Camp
Tepuia vareschii Steyerm.
Tepuia venusta Camp
Thibaudia breweri Steyerm. & Maguire
Thibaudia carrenoi Steyerm. & Maguire
Thibaudia cupatensis Huber
Thibaudia dolichandra Maguire, Steyerm. & Luteyn
Thibaudiaformosa (Klotzsch) Hoerold
Thibaudia glandulifera A.C. Sm.
Thibaudia involiicrata A.C. Sm.
Thibaudia longipes Maguire, Steyerm. & Luteyn
Thibaudia nutans Klotzsch ex Mansf.
Thibaudia smithiana Maguire, Steyerm. & Luteyn
Thibaudia truncata A.C. Sm.
Vaccinium chimantense Maguire, Steyerm. & Luteyn
Vaccinium corymbodendron Dunal
Vaccinium euryanthum A.C. Sm.
Vaccinium puberulum Klotzsch ex Meisn.
Vaccinium puberulum var. cardonae (A.C. Sm.) Maguire, Steyerm. & Luteyn
Vaccinium puberulum var. hitchcockii (Camp) Maguire, Steyerm. & Luteyn
AM BO
BO
BO
AM BO
AM BO
AM
BO
BO
AM BO
BO
BO
AM
BO
BO
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
AM
AM
BO
AM
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
BO
BO
BO
BO
BO
BO
BO
AM
BO
AM
AM BO
AM
AM
A \/f
BO
AM
AM
AM
BO
AM
BO
BO
BO
AM BO
BO
GU
GU
GU?
GU?
GU?
GU
GU
GU
GU
GU
GU
GU
GU
GU SU
SU? FG
GU SU FG
GU FG
GU SU FG
GU
GU
GU SU
GU
GU
GU
AM
BO
BO GU
BO
BO GU
BO GU
BO
BO
Checklist of the Plants of the Guiana Shield 287
Vaccinium puberulum yar.jauaeme Maguire, Steyerm. & Luteyn
Vaccinium puberulum van puberulum AM
Vaccinium puberulum var. spathulatum (Camp & A.C. Sm.) Maguire, Steyerm.
& Luteyn
Vaccinium puberulum var. subcrenulatum (Klotzsch ex Meisn.) Maguire, AM
Steyerm. & Luteyn
Vaccinium puberulum var. tepuiense (Camp & A.C. Sm.) Maguire, Steyerm. & AM
Luteyn
Vaccinium roraimense N.E. Br. [Mt. Roraima] AM
Vaccinium steyermarkii Luteyn
Synonyms and Misidentifications
Befaria spp. =Bejaria spp., nom. cons. prop. "Befaria"
Bejaria coarctata Bonpl. =Bejaria aestuans
Bejaria glauca Bonpl. =Bejaria aestuans
Bejaria glauca var. coarctata (Bonpl.) Mansf. & Sleumer =Bejaria aestuans
Bejaria glauca var. setosa Mansf. & Sleumer =Bejaria aestuans
Bejaria glauca var. tomentella Mansf. & Sleumer =Bejaria aestuans
Bejaria guianensis Klotzsch ex M.R. Schomb. [nomen nudum] =Bejaria sprucei
Bejaria guianensis Klotzsch ex N.E. Br. =Bejaria sprucei
Bejaria hispida Poepp. & Endl. =Bejaria aestuans
Bejaria puberula Klotzsch ex Mansf. & Sleumer =Bejaria sprucei
Bejaria schomburgkiana Klotzsch ex M.R. Schomb. [nomen nudum] -^Bejaria sprucei
Bejaria schomburgkiana Klotzsch ex Mansf. & Sleumer =Bejaria sprucei
Bejaria tatei Gleason =Bejaria imthurnii
Bejaria variabilis A.C. Sm. =Bejaria sprucei
Brossea lindeniana (Planch.) Kuntze =Gaultheria alnifolia var. alnifolia
Brossea roraimae (Klotzsch ex Meisn.) Kuntze =Gaultheria erecta
Cavendishia duidae A.C. Sm. =Cavendishia callista
Cavendishia gleasoniana A.C. Sm. =Cavendishia callista
Cavendishia phelpsiae Camp =Cavendishia callista
Cavendishia salicifolia Maguire, Steyerm. & Luteyn =Cavendishia callista
Cavendishia urichiana Britton =Psammisia urichiana
Ceratostema ulei Mansf. =Thibaudia nutans
Chupalon pubescens (Kunth) Kuntze =Cavendishia aff. pubescens
Epigaea cordifolia Sw. [not in flora area] =Gaultheria domingensis
Gaultheria cordifolia Kunth =Gaultheria erecta
Gaultheria lepida A.C. Sm. =Gaultheria erecta
Gaultheria lindeniana Planch. =Gaultheria alnifolia var. alnifolia
Gaultheria odorata Bredem. ex Willd. =Gaultheria erecta
Gaultheria odorata Kunth =Gaultheria erecta
Gaultheria roraimae Klotzsch ex Meisn. =Gaultheria erecta
Gaultheria scabra Willd. =Gaultheria erecta
Gaultheria tatei A.C. Sm. =Gaultheria erecta
Gaylussacia cacuminis A.C. Sm. =Gaylussacia buxifolia
Ledothamnus guyanensis var. nitidus A.C. Sm. =Ledothamnus guyanensis
Ledothamnus guyanensis subsp. tatei (A.C. Sm.) Maguire, Steyerm. & Luteyn =Ledothamnus guyanensis
Ledothamnus stenopetalus Maguire, Steyerm. & Luteyn =Ledothamnus guyanensis
Ledothamnus steyermarkii A.C. Sm. =Ledothamnus guyanensis
Ledothamnus steyermarkii subsp. longisetus Maguire, Steyerm. & Luteyn ^Ledothamnus guyanensis
Ledothamnus tatei A.C. Sm. =Ledothamnus guyanensis
Ledothamnus tatei var. puberulus A.C. Sm. =Ledothamnus guyanensis
Leucothoe duckei Huber =Agarista duckei
Leucothoe venezuelensis A.C. Sm. =Agarista duckei
Orthaea pendula Tutin =Orthaea apophysata
Orthaea pubifolia Maguire, Steyerm. & Luteyn =Orthaea hispida
Proclesia pubescens (Kunth) Klotzsch =Cavendishia aff. pubescens
Psammisia breweri Steyerm. & Maguire =Psammisia urichiana
Psammisia coriacea N.E. Br. =Thibaudia nutans
Psammisiaformosa Klotzsch ^Thibaudiaformosa
Psammisia leucostoma Benth. ex Meisn. =Psammisia guianensis
Riedelia panurensis (Benth. ex Meisn.) Kuntze =Satyria panurensis
Satyria maguirei Camp =Satyria carnosiflora
Sophoclesia cordifolia (Benth.) Klotzsch =Sphyrospermum cordifolium
288 Checklist of the Plants of the Guiana Shield
Sophoclesia klotzschiana Mansf. =Sphyrospermum klotzschianum
Sophoclesia major (Griseb.) Benth. & Hook. f. =Sphyrosperfniim cordifolium
Sophoclesia nummulahaefolia Klotzsch =Sphyrospermum buxifolium
Sophoclesia ovata Klotzsch =Sphyrospermum cordifolium
Sophoclesia subscandens Klotzsch =Sphyrospermum cordifolium
Sphyrospermum majus Griseb. =Sphyrospermum cordifolium
Sphyrospermum roraimae Klotzsch =Sphyrospermum buxifolium
Tepuiafloribunda Camp =Tepuia venusta
Tepuia tatei var. chimantana Steyerm. & Maguire =Tepuia tatei
Thibaudia alnifolia Dunal =Gaultheria alnifolia
Thibaudiajauaensis Steyerm. & Maguire =Thibaudia involucrata
Thibaudia pamirensis Benth. ex Meisn. =Satyria panurensis
Thibaudia pubescens Kunth =Cavendishia aff pubescens
Thibaudia roraimae Mansf. =Thibaudiaformosa
Thibaudia ulei (Mansf.) A.C. Sm. =Thibaudia nutans
Vaccinium cardonae A.C. Sm. =Vaccinium puberulum var. cardonae
Vaccinium cuspidatum Planch, ex Wedd. =Disterigma cuspidatum
Vaccinium hitchcockii Camp = Vaccinium puberulum var. hitchcockii
Vaccinium humboldtii Klotzsch =Disterigma humboldtii
Vaccinium spathulatum Camp & A.C. Sm. =Vaccinium puberulum var. spathulatum
Vaccinium subcrenulatum Klotzsch ex Meisn. =Vaccinium puberulum var. subcrenulatum
Vaccinium tepuiense Camp & A.C. Sm. =Vaccinium puberulum var. tepuiense
References
Lanjouw, J. 1934. Ericaceae. In A. Pulle, Flora of Suriname 4(1): 299-301.
Lanjouw, J. 1937. Ericaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 492-493.
Luteyn, J.L. 1983. Ericaceae Part I. Cavendishia. Flora Neotrop. Monogr. 35: 1-290.
Luteyn, J.L. 1987. Orthaea: new species and redefinition of the genus. Nord. J. Bot. 7(1): 31-37.
Luteyn, J.L. (ed.). 1995. Ericaceae - Part 11. The Superior-Ovaried Genera (Monotropoideae, Pyroloideae, Rhododendroideae,
and Vaccinioideae p.p.). Flora Neotropica Monogr. 66: 1-560.
Luteyn, J.L. 1998. Ericaceae. In J. A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 735-769.
Luteyn, J.L. 2002. Ericaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. HofF, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 261-
263.
Maguire, B., J.A. Steyermark, & J.L. Luteyn. 1978. Ericaceae. In B. Maguire & collaborators. Botany of the Guayana Highland.
Mem. New York Bot. Gard. 29: 139-203.
Sleumer, H.O. 1936. Die Arten der Gattung Vaccinium L. in Zentral- und Siidamerika. Notizbl. Bot. Gart. Berlin-Dahlem 13:
111-140.
Smith, A.C. 1932. The American species of Thibaudieae. Contr. U.S. Nat. Herb. 28: 31 1-547.
Smith, A.C. 1933. The genera Sphyrospermum and Disterigma. Brittonia 1: 203-232.
Erythroxylaceae
RE. Berry, T. Plowmant
Eiythroxylum amazonicum Peyr.
Erythroxylum amplum Benth.
Erythroxylum brevipes DC.
Erythroxylum campestre A. St.-Hil.
Erythroxylum cataractarum Spruce ex Peyr.
Erythroxylum citrifolium A. St.-Hil.
Erythroxylum coelophlebium Mart.
Erythroxylum cumanense Kunth
Erythroxylum divaricatum Peyr.
Erythroxylumfimbriatum Peyr.
Erythroxylum foetidum Plowman
Erythroxylum gracilipes Peyr.
Erythroxylum grisebachii Peyr.
Erythroxylum guanchezii Plowman
Erythroxylum havanense Jacq.
Erythroxylum hypoleucum Plowman
Erythroxylum impressum O.E. Schulz
Erythroxylum kapplerianum Peyr.
Erythroxylum lenticellosum Huber
Erythroxylum leptoneurum O.E. Schulz
AM BO GU SU EG
GU SU EG
GU?
GU?
AM
AM BO GU SU
SU
FG
GU SU EG
AM BO GU
A \/f
BO FG
AM
AM BO DA GU SU
SU
FG
AM
BO GU
BO GU
BO GU SU FG
FG
AM su FG
AM GU su
AM BO GU su FG
AM BO GU su FG
AM BO GU su FG
AM BO
AM BO GU su FG
AM BO GU
AM BO GU
AM BO GU su FG
AM BO
BO
GU
GU
FG
AM
AM BO
BO
GU su FG
BO GU FG
BO
AM BO GU FG
Checklist of the Plants of the Guiana Shield 289
Etythroxylum Ugiistrhnim DC.
Ery-throxylum lindemanii Plowman
Etythroxylum Uneolatiim DC.
Erythwxylum macrophyllum Cav.
Erythroxylum macrophyllum var. maa-ophyllum
Erythroxylum macrophyllum var. savannarum Plowman
Erythroxylum mucronatum Benth.
Erythroxylum oreophilum (O.E. Schulz ex Pilg.) Steyerm. & Maguire
Erythroxylum orinocense Kunth
Erythroxylum roraimae Klotzsch ex O.E. Schulz
Erythroxylum rufum Cav.
Erythroxylum schomburgkii Peyr.
Erythroxylum spruceanum Peyr.
Erythroxylum squamatum Sw.
Erythroxylum steyermarkii Plowman
Erythroxylum suberosum A. St.-Hil.
Erythroxylum ajf. subrotundum A. St.-Hil.
Erythroxylum vernicosum O.E. Schulz
Erythroxylum williamsii Standi, ex Plowman AM BO
Synonyms and Misidentifications
Erythroxylum acutifolium Steud. ex Peyr. ^Erythroxylum citrifolium
Erythroxylum albertianum Kuhlm. & W.A. Rodrigues =Erythroxylum mucronatum
Erythroxylum areolatum Veil. =Etythroxylum suberosum
Erythroxylum aristigerum Peyr. =E}ythroxylum squamatum
Erythroxylum aristigerum var. bahiense (Peyr.) O.E. Schulz =Erythroxylum squamatum
Erythroxylum bahiense Peyr. =E}ythroxylum squamatum
Erythroxylum christii O.E. Schulz [nomen nudum] =Erythroxylum ajf. subrotundum
Erythroxylum coelophlebium var. grisebachii (Peyr.) O.E. Schulz =Ejythroxylum grisebachii
Erythroxylum comosum O.E. Schulz =Erythroxylum mucronatum
Erythroxylum cuatrecasasii W.A. Gentner =Erythroxylum gracilipes
Erythroxylum duckei Huber =Erythroxylum citrifolium
Erythroxylum ectinocalyx Klotzsch ^Erythroxylum divaricatum
Erythroxylum filipes Huber =Eiythroxylum macrophyllum
Erythroxylumfloribundum Mart. =Erythroxylum macrophyllum
Erythroxylum gomphioides Planch. & Linden ex Triana & Planch. =Erythroxylum citrifolium
Erythroxylum havanense var. continentis O.E. Schulz =E}ythroxylum havanense
Erythroxylum kirkianum O.E. Schulz =Efythroxylum mucronatum
Erythroxylum laurinum Planch. & Linden ex Triana & Planch. =E}ythroxylum macrophyllum
Erythroxylum ligustrinum var. grandifolium Sagot =E}ythroxylum kapplerianum
Erythroxylum lucidum Kunth =Erythroxylum macrophyllum
Erythroxylum mapuerae Huber =Erythroxylum kapplerianum
Erythroxylum micranthum Bong, ex Peyr. =Erythroxylum citrifolium
Erythroxylum novogranatense var. macrophyllum O.E. Schulz =Erythroxylum gracilipes
Erythroxylum obtusum DC. =Erythroxylum havanense
Erythroxylum ovatum Cav. =E}ythroxylum havanense
Erythroxylum paraense Peyr. =Erythroxylum citrifolium
Erythroxylum recurrens Huber =Er}-throxylum gracilipes
Erythroxylum squamatum var. emarginatum Peyr. =Erythroxylum vernicosum
Erythroxylum squamatum var. microcarpum O.E. Schulz =Efythroxylum squamatum
Erythroxylum squamatum var. orinocense Kuntze ^Erythroxylum impressum
Erythroxylum squarrosum Klotzsch [nomen nudum] =Erythroxylum rufum
Erythroxylum testaceum Peyr. =Erythroxylum suberosum
Erythroxylum trinerve Huber =Erythroxylum squamatum
Erythroxylum venezuelense Steyerm. =Erythroxylum mucronatum
Erythroxylum vernicosum var. oreophilum O.E. Schulz ex Pilg. =Eiythroxylum oreophilum
References
Heald, S.V. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide
to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 263-265.
Plowman, T. & RE. Berry. 1999. Erythroxylaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 5: 59-71.
Schulz, O.E. 1907. Erythroxylaceae. In A. Engler, Das Pflanzenreich IV(134): 1-176.
Westhoff, V. 1942. Erythroxylaceae. In A. PuIIe, Flora of Suriname 3(2): 1-12.
290 Checklist of the Plants of the Guiana Shield
EUPHORBIACEAE
W.S. Armbruster, RE. Berry, H.-J. Esser, L.J. Gillespie, W.J. Hayden,
S.V. Heald, G.A. Levin, W. Punt, R. de S. Secco, G.L. Websterf
Acalypha alopeciiroides Jacq.
Acalypha arvensis Poepp.
Acalypha cuneata Poepp.
Acalypha diversifolia Jacq.
Acalypha indica L. [naturalized]
Acalypha lanceolata Willd. [naturalized]
Acalypha macrostachya Jacq.
Acalypha poiretii Spreng.
Acalypha scandens Benth.
Acalypha schiedeana Schltdl.
Acalypha setosa A. Rich.
Acalypha villosa Jacq.
Acidoton nicaraguensis (Hemsl.) G.L. Webster
Actinostemon amazoniciis Pax & K. HotYm.
Actinostemon schonibiirgkii (Klotzsch) Hochr.
Adelia triloba (Miill. Arg.) Hemsl.
Adenophaedra grandifolia (Klotzsch) Miill. Arg.
Alchornea castaneifolia (Willd.) A. Juss.
Alchornea discolor Poepp.
Alchornea jliiviatilis Secco
Alchornea glandulosa Poepp.
Alchornea grandiflora Mull. Arg.
Alchornea triplinervia (Spreng.) Miill. Arg.
Alchorneopsis floribunda (Benth.) Miill. Arg.
Amanoa aimerindae Leal
Amanoa congesta W.J. Hayden
Amanoa ciipatensis Huber
Amanoa glaiicophylla Miill. Arg.
Amanoa guianensis Aubl.
Amanoa neglecta W.J. Hayden
Amanoa oblongifolia Miill. Arg.
Amanoa steyermarkii Jabl.
Antidesma ghaesembilla Gaertn. [cultivated, naturalized]
Aparisthmium cordattim (A. Juss.) Baill.
Astrococcus cornutits Benth.
Bernardia amazonica Croizat [specimens not seen]
Bernardia sidoides (Klotzsch) Miill. Arg.
Caperonia castaneifolia (L.) A. St.-Hil.
Caperonia corchoroides Mull. Arg. [=Caperonia palustrisl]
Caperonia pahidosa Klotzsch [=Caperonia castaneifolia in FVG]
Caperonia palustris (L.) A. St.-Hil.
Caperonia stenophylla Miill. Arg.
Celianella montana Jabl.
Chaetocarpus schomburgkianiis (Kuntze) Pax & K. Hoffm.
Cnidoscolus wens (L.) Arthur
Conceveiba guianensis Aubl.
Conceveiba latifolia Benth.
Conceveiba martiana Baill.
Conceveiba ptariana (Steyerm.) Jabl.
Croton argenteus L.
Croton argyrophylloides Miill. Arg.
Croton argyrophyllus Kunth
Croton bolivarensis Croizat
Croton cajucara Benth.
Croton conduplicatus Kunth
Croton cuneatus Klotzsch
Croton aff draconoides Miill. Arg.
Croton essequiboensis Klotzsch [=Croton populifolius P. Mill.?]
Croton fragrans Kunth
Croton galeopsifolius Lanj.
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
AM BO
AM BO
AM
AM
AM
AM BO
AM
AM BO
AM
\ \ A
BO
AM BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
DA
DA
DA
DA
DA
DA
DA
DA
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
GU SU
GU SU
GU SU
GU
GU
GU
GU
GU
GU
GU
SU
FG
FG
FG
FG
GU
GU
GU SU? FG
GU FG
GU SU FG
GU
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
Checklist of the Plants of the Guiana Shield 291
Croton gossypiifolius Vahl BO
Croton guaiquinimae Steyerm. BO
Croton guianemis A\xh\. BO GU SU FG
Croton hirtus L'Her. AM BO DA GU SU FG
Croton hostmannii Miq. ex Schltdl. GU SU FG
Croton icabarui Jabl. BO
Croton kaieteuri Jabl. [probably conspecific with Croton cimeatus] GU
Croton kavayanensis Steyerm. FG?
Croton lobatiis L. BO DA GU SU FG
Croton longiradiatus Lanj. SU FG
Croton macradenis Gorts & Punt GU SU FG
Croton matourensis Auh\. AM BO GU SU FG
Croton megalodendron Miill. Arg. BO
Croton mollis Benth. AM
Croton neblinae Jabl. AM
Croton nepetifolius Baill. BO
Croton nuntians Croizat DA
Croton orinocensis Mull. Arg. AM
Croton palanostigma Klotzsch AM BO
Croton parodianus Croizat AM
Croton polypleurus Croizat
Croton potaroensis Lanj. AM BO
Croton pullei Lanj
.
AM
Croton pullei var. glabrior Lanj.
Croton pullei var. pullei
Croton rhamnifolius Kunth
Croton roraimensis Croizat BO
Croton roraimensis var. roraimensis
Croton rottlerifolius Baill.
Croton schiedeanus Schltdl. BO
GU FG
GU
GU FG
GU
GU
GU
GU? SU FG
SU FG
GU? SU
GU
GU
GU
FG?
GU SU FG
Croton scutatus RE. Berry & J.F. Gaskin AM
Croton sipaliwinensis Lanj. AM BO GU SU
Croton sp. A AM
Croton sp. B AM
Croton sp. C AM
Croton sp. D AM
Croton sp. E BO
Croton sp. F AM
Croton sp. G AM
Croton sp. H BO
Croton spiraeifolius Jabl. AM
Croton spruceanus Benth. AM
Croton stahelianus Lanj. SU
Croton subcoriaceus Jabl. AM
Croton subincanus Miill. Arg. BO GU SU FG
Croton subserratus Jabl. AM BO
Croton trinitatis Millsp. AM BO DA GU SU FG
Croton umbratilis Kunth BO
Croton vergarenae (Jabl.) L.J. Gillespie BO
Croton yavitensis Croizat AM
Dalechampia affinis Miill. Arg. AM BO GU SU FG
Dalechampia attenuistylus Armbr. BO SU FG
Dalechampia brevicolumna Armbr. SU? FG
Dalechampia brownsbergens is G.L. Webster & Armbr. BO DA SU
Dalechampia aff. cissifolia Poepp. GU
Dalechampia dioscoreifolia Poepp. AM BO FG
Dalechampiafragrans Armbr. SU FG
Dalechampia heterobractea Armbr. [often confused with Dalechampia BO GU SU FG
parvibracteata^
Dalechampia liesneri Huft AM
Dalechampia magnoliifolia Miill. Arg. AM BO GU FG
Dalechampia megacarpa Armbr. AM BO
Dalechampia micrantha Poepp. BO SU FG
Dalechampia papillistigma Armbr. BO
292 Checklist of the Plants of the Guiana Shield
Dalechampia parvibracteata Lanj
.
Dalechampia scandens L.
Dalechampia stipidacea Miill. Arg.
Dalechampia tenuiramea Mull. Arg.
Dalechampia tiliifolia Lam.
Dalechampia aff. triphylla Lam.
Dendrothrix multiglandidosa Esser
Dendrothrix yiitajensis (Jabl.) Esser
Discocarpus essequeboensis Klotzsch [="essequiboensis"]
Discocatpiis gentryi S.M. Hayden
Discocarpus spruceanus Miill. Arg.
Ditaxis polygama (Jacq.) L.C. Wheeler
Dodecastigma amazonicum Ducke
Dodecastigma integhfoliiim (Lanj.) Lanj. & Sandwith
Drypetesfanshawei Sandwith
Drypetes monachinoi Jabl.
Drypetes variabilis Uittien
Euphorbia bahiensis (Klotzsch & Garcke) Boiss.
Euphorbia blodgettii Engelm. [adventitious]
Euphorbia comosa Veil.
Euphorbia cotinifolia L.
Euphorbia cotinifolia subsp. cotinifolia
Euphorbia cotinifolia subsp. cotinoides (Miq.) Christenh.
Euphorbia dioeca Kunth
Euphorbia graminea Jacq. [adventitious]
Euphorbia heterophylla L.
Euphorbia heterophylla var. cyathophora (Murray) Griseb. [native]
Euphorbia heterophylla var. heterophylla [native]
Euphorbia hirta L.
Euphorbia humayensis Brandegee
Euphorbia hypericifolia L.
Euphorbia hyssopifolia L.
Euphorbia hyssopifolia var. pubescenticocca Christenh.
Euphorbia aff. laurifolia Juss.
Euphorbia mesembrianthemifolia Jacq. [adventitous]
Euphorbia neriifolia L. [cultivated, naturalized]
Euphorbia oerstediana (Klotzsch & Garcke) Boiss.
Euphorbia prostrata Alton
Euphorbia pulcheririma Willd. ex Klotzsch [cultivated, naturalized]
Euphorbia serpens Kunth
Euphorbia spruceana Boiss.
Euphorbia thymifolia L.
Euphorbia tirucalli L. [cultivated, naturalized]
Gavarretia terminalis Baill.
Glycydendron amazonicum Ducke
Gymnanthes hypoleuca Benth.
Haematostemon coriaceus (Baill.) Pax & K. Hoffm.
Haematostemon guianensis Sandwith GU
Hevea benthamiana Miill. Arg. AM
Hevea brasiliensis (Willd. ex A. Juss.) Miill. Arg. [widely cultivated, possibly FG?
indigenous in FG]
Hevea guianensis Aubl.
Hevea guianensis var. guianensis
Hevea guianensis var. lutea (Spruce ex Benth.) Ducke & R.E. Schult.
Hevea microphylla Ule
Hevea pauciflora (Spruce ex Benth.) Miill. Arg.
Hevea pauciflora var. coriacea Ducke
Hieronyma alchorneoides Allemao
Hieronyma alchorneoides var. alchorneoides
Hieronyma alchorneoides var. stipulosa Franco
Hieronyma oblonga (Tul.) Miill. Arg.
Hura crepitans L.
Jablonskia congesta (Benth. ex Miill. Arg.) G.L. Webster
Jatropha curcas L. [cultivated, naturalized]
BO GU
AM BO DA GU su FG
FG
AM
AM BO GU
su
FG
FG
AM
AM BO
AM GU su FG
AM
AM
BO
BO
DA
GU
GU
GU
su
FG
BO DA GU su FG
FG
BO su?
AM BO GU su FG
AM BO GU su FG
BO GU su FG
BO GU FG
FG
AM BO GU su
su
FG
FG
AM BO GU su
AM BO
BO
DA GU su FG
BO GU su FG
AM BO DA GU
GU
GU
GU
su
su
FG
FG'
AM BO
BO
GU
GU
su FG
FG
AM BO GU su FG
AM BO GU
AM GU su FG
AM GU FG
AM
AM BO GU SU FG
AM BO GU su FG
AM BO GU su FG
AM
AM BO GU su
AM BO GU su
AM BO DA GU su FG
GU su FG
DA FG
AM BO GU su FG
BO GU su FG
AM GU su FG?
BO GU su FG
AM BO DA GU
GU
AM
AM
GU
AM
AM
AM
AM BO GU
AM BO
AM
GU
AM BO DA GU
BO GU
AM BO
AM BO
AM BO
BO
BO
su FG
su FG
GU
FG
GU
GU su FG
GU su FG
FG
GU su FG
GU su FG
GU su FG
Checklist of the Plants of the Guiana Shield 293
Jatropha gossypiifoliaL. SU FG
Mabea angularis Hollander
Mabea anomala Miill. Arg.
Mabea arenicola Esser
Mabea biglandulosa Baill. ex Miill. Arg.
Mabeafrutescens Jabl.
Mabea linearifolia Jabl.
Mabea longibracteata Esser
Mabea montana Miill. Arg.
Mabea nitida Spruce ex Benth.
Mabea occidentalis Benth. [not in flora area]
Mabea pakaraimaea Esser, sp. nov. ined.
Mabea piriri Aubl.
Mabea pulcherrima Miill. Arg.
Mabea rubicunda Jabl.
Mabea salicoides Esser
Mabea sp. A AM
Mabea sp. B [expected in BO. Base of Roraima on Guyana side]
Mabea speciosa Miill. Arg.
Mabea speciosa subsp. A
Mabea speciosa subsp. B
Mabea speciosa subsp. speciosa
Mabea subsessilis Pax & K. Hoffm.
Mabea taquari Aubl. DA
Mabea trianae Pax
Manihot anomala Pohl
Manihot anomala subsp. pubescens D.J. Rogers & Appan
Manihot brachyloba MiiW. Arg. AM DA GU SU FG
Manihot carthaginensis (Jacq.) Miill. Arg. BO
Manihot esculenta Crantz [cultivated, naturalized]
Manihot aff. quinquepartita Huber ex D.J. Rogers & Appan
Manihot sprucei Pax
Manihot surinamensis D.J. Rogers & Appan
Manihot tristis Miill. Arg.
Manihot tristis subsp. saxicola (Lanj.) D.J. Rogers & Appan
Manihot tristis subsp. tristis [not in flora area]
Maprounea amazonica Esser
Maprounea guianensis Aubl.
Margaritaria nobHis L. f.
Micrandra brownsbergensis Lanj . [probably conspecific with Micrandra
elata]
Micrandra glabra (R.E. Schuh.) R.E. Schult.
Micrandra gleasoniana (Croizat) R.E. Schult.
Micrandra inundata RE. Berry & Wiedenhoeft
Micrandra lopezii R.E. Schult.
Micrandra minor Benth.
Micrandra rossiana R.E. Schult.
Micrandra siphonioides Benth.
Micrandra sp. A
Micrandra spruceana (Baill.) R.E. Schult. [record probably misident.]
Micrandra sprucei (Mull. Arg.) R.E. Schult.
Microstachys corniculata (Vahl) Griseb.
Omphalea diandra L.
Pausandrafordii Secco
Pausandra martinii Baill.
Pausandra trianae (Mull. Arg.) Baill. [specimens not seen]
Pedilanthus tithymaloides (L.) Poit.
Pedilanthus tithymaloides subsp. tithymaloides [native to GU, SU; cultivated
in all]
Pera bicolor (Klotzsch) Miill. Arg.
Pera citriodora Baill.
Pera decipiens (Mull. Arg.) Miill. Arg.
Pera distichophylla (Mart.) Baill.
Pera glabrata (Schott) Poepp. ex Baill.
GU SU FG
FG
FG
AM BO GU su FG
AM BO GU su FG
BO GU su FG
AM BO
AM
AM BO DA GU su FG
AM BO DA GU su FG
GU su FG
BO GU
GU
su
AM
GU?
AM BO
AM BO GU
AM BO GU FG
AM
AM GU? su
AM GU
AM BO GU su FG
DA GU su FG
FG
AM BO DA GU su FG
FG
BO GU su FG
BO GU su FG
AM BO GU su FG
AM
AM BO GU su
AM BO
AM BO DA GU su FG
GU su
GU su
AM BO DA GU su FG
AM
AM BO
BO
GU su FG
AM BO
BO GU su FG
BO DA GU su
BO DA
DA
GU su FG
FG
BO DA GU su FG
FG
BO
BO
GU su FG
AM
BO DA
DA
GU
GU
su
AM BO su? FG
AM
BO
BO GU FG
BO
GU FG
294 Checklist of the Plants of the Guiana Shield
Pera tomentosa (Benth.) Miill. Arg. AM
Phyllanthus acuminatus Vahl
Phyllanthus adiantoides Klotzsch
Phyllanthus amarus Schumach. & Thonn.
Phyllanthus atabapoensis Jabl.
Phyllanthus attenuatus Miq.
Phyllanthus bolivarensis Steyerm.
Phyllanthus borjaensis Jabl.
Phyllanthus brasiliensis (Aubl.) Poir.
Phyllanthus caribaeus Urb.
Phyllanthus caroliniensis Walter
Phyllanthus caroliniensis subsp. caroliniensis
Phyllanthus caroliniensis subsp. guianensis (Klotzsch) G.L. Webster
Phyllanthus caroliniensis subsp. stenopterus (Miill. Arg.) GL. Webster
Phyllanthus carrenoi Steyerm.
Phyllanthus chimantae Jabl.
Phyllanthus debilis Klein ex Willd. [naturalized]
Phyllanthus duidae Gleason
Phyllanthus elsiae Urb.
Phyllanthus fluitans Benth. ex Miill. Arg.
Phyllanthus gallinetae Jabl. [included in Phyllanthus myrsinites?]
Phyllanthus hyssopifolioides Kunth
Phyllanthus jablonskianus Steyerm. & Luteyn
Phyllanthus jauaensis Jabl.
Phyllanthusjuglandifolius Wi 1 1d
.
Phyllanthus juglandifolius subsp. cornifolius (Kunth) G.L. Webster
Phyllanthus juglandifolius subsp . juglandifolius
Phyllanthus lediformis Jabl. AM
Phyllanthus lindbergii Mull. Arg. AM
Phyllanthus longistylus Jabl. AM BO
Phyllanthus maguirei Jabl. AM
Phyllanthus majus Steyerm. AM BO GU
Phyllanthus microphyllus Kunth AM BO
Phyllanthus minutifolius Jabl. AM
Phyllanthus minutulus Mull. Arg. AM BO GU
Phyllanthus myrsinites Kunth AM GU
Phyllanthus myrsinites subsp. francaviIlanus (Miill. Arg.) G.L. Webster AM
Phyllanthus myrsinites subsp. myrsinites [=includes Phyllanthus gallinetae?] AM GU
Phyllanthus neblinae Jabl. AM
Phyllanthus niruri L. SU FG
Phyllanthus obfalcatus Lasser & Maguire AM
Phyllanthus orbiculatus Rich. AM BO SU FG
Phyllanthus orinocensis Steyerm. AM
Phyllanthus paezensis Jabl.
Phyllanthus paraqueensis Jabl. AM
Phyllanthus piscatorum Kunth [specimens not seen] AM
Phyllanthus pseudoconami Miill. Arg.
Phyllanthus pycnophyllus Miill. Arg.
Phyllanthus rupestris Kunth AM
Phyllanthus stipulatus iRaf.)G.L. Webster AM BO DA GU SU FG
Phyllanthus strobilaceus Jabl. AM
Phyllanthus subapicalis Jabl. AM
Phyllanthus subapicalis Jabl. subsp. A AM
Phyllanthus subapicalis Jabl. subsp. sequoiifolius (Jabl.) G.L. Webster BO
Phyllanthus subapicalis Jabl. subsp. subapicalis AM
Phyllanthus subglomeratus Poir. [cultivated only?] GU FG
Phyllanthus tenellus Roxb. [naturalized]
Phyllanthus tepuicola Steyerm. AM
Phyllanthus urinaria L. [naturalized] DA GU SU FG
Phyllanthus vacciniifolius (Miill. Arg.) Miill. Arg. AM
Phyllanthus ventuarii Jabl. AM
Piranhea longepedunculata Jabl.
Piranhea trifoliata Baill. AM
Plukenetia brachybotrya Miill. Arg. [group, vide also P. buchtienii, P.
loretensis]
BO
BO GU?
GU
BO GU
BO
BO
BO GU SU
su
BO GU
BO DA
BO GU?
GU?
Checklist of the Plants of the Guiana Shield 295
Plukenetia loretensis Ule
Plukenetia multiglandulosa Jabl.
Plukenetia penninervia Miill. Arg.
Plukenetia polyadenia Miill. Arg.
Plukenetia supraglandulosa L.J. Gillespie
Plukenetia verrucosa Sm.
Plukenetia volubilis L.
Podocalyx loranthoides Klotzsch
Pogonophora schomburgkiana Miers ex Benth.
Richeria grandis Vahl
Ricinus communis L. [cultivated, naturalized]
Sagotia brachysepala (Miill. Arg.) Secco
Sagotia racemosa Baill.
Sandwithia guyanensis Lanj.
Sandwithia heterocalyx Secco
Sapium argutum (Miill. Arg.) Huber
Sapium ciliatum Hemsl.
Sapium contortum Croizat
Sapium glandulosum (L.) Morong
Sapiumjenmanii Hemsl. '
Sapium paucinervium Hemsl.
Savia sessiliflora (Sw.) Willd.
Sebastiania bidentata (Mart.) Pax
Sebastiania guianensis (Miill. Arg.) Miill. Arg.
Senefeldera inclinata Miill. Arg.
Senefeldera macrophylla Ducke
Senefelderopsis chiribiquetensis (R.E. Schult. & Croizat) Steyerm.
Senefelderopsis croizatii Steyerm.
Tetrorchidium rubrivenium Poepp.
Tragiafallax Mull. Arg.
Tragiafendleri Miill. Arg.
Tragia guayanensis L.J. Gillespie
Tragia lessertiana (Baill.) Miill. Arg.
Tragia tabulaemontana L.J. Gillespie
Tragia volubilis L.
Synonyms and Misidentifications
Acalypha cuneata var. obovata (Benth.) Miill. Arg. =Acalypha cuneata
Acalypha diversifolia var. carpinifolia (Miill. Arg.) Miill. Arg. =Acalypha diversifolia
Acalypha diversifolia var. leptostachya (Kunth) Miill. Arg. =Acalypha diversifolia
Acalypha hirsutissima Willd. =Acalypha macrostachya
Acalypha karsteniana Pax & K. Hoffm. =Acalypha villosa
Acalypha leptostachya Kunth =Acalypha diversifolia
Acalypha leptostachya var. carpinifolia Miill. Arg. =Acalypha diversifolia
Acalypha macrophylla Kunth =Acalypha macrostachya
Acalypha macrostachya var. hirsutissima (Willd.) Miill. Arg. =Acalypha macrostachya
Acalypha macrostachya var. macrophylla (Kunth) Miill. Arg. =Acalypha macrostachya
Acalypha macrostachya var. tristis (Poepp.) Miill. Arg. =Acalypha macrostachya
Acalypha obovata Benth. =Acalypha cuneata
Acalypha persimilis var. corchorifolia (A. Rich.) Miill. Arg. [not in flora area]
Acalypha samydifolia Poepp. =Acalypha diversifolia
Acalypha schiedeana var. angustifolia (Mull. Arg.) Pax & K. Hoffm. =Acalypha schiedeana
Acalypha schiedeana f. angustifolia Miill. Arg. ^^^Acalypha schiedeana
Acalypha tristis Poepp. =Acalypha macrostachya
Acalypha villosa var. intermedia Miill. Arg. =^Acalypha villosa
Acalypha villosa war.paniculata Miill. Arg. =Acalypha villosa
Acalypha villosa var. tomentosa Mull. Arg. =Acalypha villosa
Acidoton venezolanus (Pax & K. Hoffm.) G.L. Webster =Acidoton nicaraguensis
Actinostemon depauperatus Pax & K. Hoffm. =Actinostemon schomburgkii
Actinostemon guyanensis Pax & K. Hoffm. ? ^Sandwithia guyanensis
Actinostemon parvifolius Pittier =Actinostemon schomburgkii
Alchornea brachygyne Pax & K. Hoffm. =Alchornea discolor
Alchornea cordata (A. Juss.) Miill. Arg. =Aparisthmium cordatum
Alchornea glandulosa yar.floribunda Benth. =Alchorneopsis floribunda
Alchornea orinocensis Croizat =Aparisthmium cordatum
AM BO GU
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GU su FG
GU su FG
AM GU su
AM BO GU
AM BO GU su FG
AM BO GU FG
AM BO DA GU su FG
BO GU su FG
AM BO GU su FG
AM BO DA GU su FG
AM BO
GU su
su
FG
FG
AM BO
BO DA GU su FG
BO DA GU su
BO DA GU su FG
BO
GU
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GU su FG
AM
AM BO GU
AM BO
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FG
296 Checklist of the Plants of the Guiana Shield
Alchornea schomburgkii Klotzsch =Alchornea discolor
Alchorneopsis trimera Lanj. =AIchomeopsis floribimda
Alectoroctonum caracasanitm Klotzsch & Garcke ^Euphorbia cotinifolia
Amanoa grandiflora (Mull. Arg.) Miill. Arg. =Amanoa guianensis
Amanoa guianensis van grandiflora Miill. Arg. =Amanoa guianensis
Amanoa pubescens Steyerm. =Amanoa almerindae
Angosty'les tabulamontana Croizat [dubious species of uncertain placement]
Antidesma triplinervia Spreng. =Alchornea triplinervia
Apodandra brachybotrya (Mull. Arg.) J.F. Macbr. =Plukenetia brachybotiya
Apodandra loretensis (Ule) Pax & K. Hoffm. =Plukenetia loretensis
Argythamnia polygama (Jacq.) Kuntze =Ditaxis polygama
Astrococcus coriaceus Baill. =Haematostemon coriaceus
Ateramnus hypoleucus (Benth.) Rothm. =Gymnanthes hypoleuca
Chaetocarpus stipularis Gleason =Chaetocarpus schomburgkianus
Chaetocarpus williamsii Steyerm. =Chaetocarpus schomburgkianus
Chamaesyce adenoptera (Bertol.) '^mdA\ i=Euphorbia dioeca
Chamaesyce dioeca (Kunth) Millsp. ^Euphorbia dioeca
Chamaesyce glomerifera Millsp. =Euphorbia hypericifolia
Chamaesyce hirta (L.) Millsp. ^Euphorbia hirta
Chamaesyce hypericifolia (L.) Millsp. =Euphorbia hypericifolia
Chamaesyce hyssopifolia (L.) Small = Euphorbia hyssopifolia (L.)
Chamaesyce mesemhrianthemifolia (Jacq.) Dugand =Euphorbia mesembrianthemifolia
Chamaesyce prosfrata (Alton) Small =Euphorbia prostrata
Chamaesyce serpens (Kunth) Small =Euphorbia serpens
Chamaesyce fhymifolia (L.) Millsp. =Euphorbia thymifolia
Cleidion sp. [not in flora area]
Cleidion? nicaraguense Hemsl. =Acidoton nicaraguensis
Clusiophyllum sprucei Miill. Arg. =Micrandra sprucei
Conami brasiliensis Aubl. =Phyllanthus brasiliensis
Conceveiba cordatum A. Juss. =Aparisthmium cordatum
Conceveiba hostmannii Benth. =Conceveiba guianensis
Conceveiba terminalis (Baill.) Miill. Arg. =Gavarretia terminalis
Conceveibaslrum martianum (Baill.) Pax & K. Hoffm. =Conceveiba martiana
Conceveibastrum ptarianum Steyerm. =Conceveiba ptariana
Conceveibum cordatum A. Juss. =Aparisthmium cordatum
Croton barlettii Lanj. =Croton potaroensis
Croton bartlettii Lanj. =Croton potaroensis
Croton benthamianus Miill. Arg. =Croton palanostigma
Croton cardonei Croizat =Croton potaroensis
Croton castaneifolius L. =Caperonia castaneifolia
Croton crenatus Miill. Arg. =Croton mollis
Croton glabellus L. [not in flora area]
Croton glandulosus L. [not in flora area]
Croton glandulosus subsp. hirtus (L'Her.) Croizat =Croton hirtus
Croton glandulosus var. hirtus (L'Her.) Miill. Arg. =Croton hirtus
Croton glandulosus subsp. hirtus (L'Her.) Croizat, Bull. =Croton hirtus
Croton kavanayensis Steyerm. =Croton matourensis
Croton lanjouwensis Jabl. =Croton matourensis
Croton matourensis var. benthamianus Mull. Arg. =Croton matourensis
Croton micans Sw. =Croton argyrophyllus
Croton miquelensis Ferguson ex Lanj. ^Croton trinitatis
Croton miquelianus Ferguson =Croton trinitatis
Croton monachinoensis Jabl. =Croton cuneatus
Croton nervosus Klotzsch [nom. illeg.] =Croton argyrophyllus
Croton pakaraimae Jabl. =Croton matourensis
Croton polygamus Jacq. =Ditaxis polygama
Croton populifolius Mill, [see Croton essequiboensis]
Croton populifolius var. essequiboensis (Klotzsch) Mull. Arg. =Croton essequiboensis
Croton roraimensis var. subinteger Steyerm. =Croton roraimensis
Croton sessiliflorum Sw. =Savia sessiliflora
Croton surinamensis Miill. Arg. =Croton cuneatus
Croton tafelbergicus Croizat =Croton guianensis
Croton xanthochloros Croizat =Croton megalodendron
Cunuria ? casiquiarensis Croizat =Podocalyx loranthoides
Checklist of the Plants of the Guiana Shield 297
Cumiria crassipes Miill. Arg. =Micrandra sprucei
Cunuha glabra R.E. Schult. ex Baldwin & R.E. Schult. =Micrandra glabra
Cumiria gleasoniana Croizat =Micrandra gleasoniana
Cumiria spruceana Baill. =Micrandra spruceana
Dactylostemon schomburgkii Klotzsch =Actinostemon schomburgkii
Dalechampia brevipes Briq. =Dalechampia scandens
Dalechampia colorata L. f. =Dalechampia scandens
Dalechampia cynanchoides S. Moore =Dalechampia tenuiramea
Dalechampia dioscoreifolia Pulle [latter homonym] =Dalechampia affinis
Dalechampia heterophylla Vahl =Dalechampia tiliifolia
Dalechampia latifolia Lam. =Dalechampia scandens
Dalechampia mollis Vahl =Dalechampia scandens
Dalechampia peruviana Lam. =Dalechampia tiliifolia
Dalechampia pruriens Griseb. =Dalechampia tiliifolia
Dalechampia roezliana var. amazonica Ule =Dalechampia magnoliifolia
Dalechampia rubiformis Spreng. =Dalechampia scandens
Dalechampia ruboides Kunth =Dalechampia scandens
Dalechampia sidaefolia Kunth =Dalechampia scandens
Dalechampia spathulata (Scheidw.) Baill. [not in flora area]
Dalechampia villosa Lam. ^Dalechampia scandens
Discocarpus brasiliensis Klotzsch ex Miill. Arg. =Discocarpus essequeboensis
Discocarpus mazarunensis Croizat =Chaetocarpus schomburgkianus
Dodecastigma mazarunense Croizat =Dodecastigma integrifolium
Diypetes maguireana Monach. =Chaetocarpus schomburgkianus
Drypetes spruceana Mull. Arg. =Chaetocarpus schomburgkianus
Elaeophora abutifolia Ducke =Plukenetia polyadenia
Elaeophora polyadenia (Miill. Arg.) Ducke =Plukenetia polyadenia
Euphorbia adenoptera Bertol. =Euphorbia dioeca
Euphorbia brasiliensis Lam. =Euphorbia hyssopifolia
Euphorbia buxifolia Lam. =Euphorbia mesembrianthemifolia
Euphorbia caracasana (Klotzsch & Garcke) Boiss. =Euphorbia cotinifolia
Euphorbia cotinoides Miq. =Euphorbia cotinifolia subsp. cotinoides
Euphorbia cyathophora Murray =Euphorbia heterophylla var. cyathophora
Euphorbia globulifera Kunth =Euphorbia hirta
Euphorbia morisoniana Klotzsch =Euphorbia heterophylla var. heterophylla
Euphorbia multinodis Urb. [not in flora area]
Euphorbia myrtillifolia L. [not in flora area]
Euphorbia surinamensis Lanj. =Euphorbia oerstediana
Euphorbia tenella Kunth =Euphorbia serpens
Euphorbia tithymaloides L. =Pedilanthus tithymaloides
Gaedawakka schomburgkiana Kuntze =Chaetocarpus schomburgkianus
Gaedawakka schomburgkianus Kuntze =Chaetocarpus schomburgkianus
Gitara venezolana Pax & K. Hoffm. =Acidoton nicaraguensis
Glochidion vacciniifolium Miill. Arg. =Phyllanthus vacciniifolius
Guarania laurifolia Baill. =Richeria grandis
Gymnanthes guyanensis Mull. Arg. =Sebastiania guianensis
Hermesia castaneifolia Willd. =Alchornea castaneifolia
Hevea confusa Hemsl. =Hevea pauciflora var. coriacea
Hevea lutea (Spruce ex Benth.) Miill. Arg. =Hevea guianensis var lutea
Hevea minor Hemsl. =Hevea pauciflora var. coriacea
Hevea paludosa Ule =Hevea pauciflora var. coriacea
Hieronyma laxiflora (Tul.) Miill. Arg. =Hieronyma alchorneoides var. alchorneoides
Hippomane biglandulosa L. [nom. illeg.] =Sapium glandulosum
Hippomane glandulosa L. ^Sapium glandulosum
Jatropha carthaginensis Jacq. =Manihot carthaginensis
Jatropha manihot L. =Manihot esculenta
Jatropha urens L. =Cnidoscolus urens
Julocroton argenteus (L.) Didr. =Croton argenteus
Julocroton vergarenae Jabl. =Croton vergarenae
Mabea argutissima Croizat =Mabea subsessilis
Mabea atroviridis Pax & K. Hoffm. =Mabea montana
Mabea caudata Pax & K. Hoffm. =Mabea speciosa subsp. speciosa
Mabea caudata var. concolor Lanj. =Mabea speciosa subsp. A
Mabea ciliata Gleason =Mabea montana
298 Checklist of the Plants of the Guiana Shield
Mabea costata Pax & K. Hoffm. =Mabea montana
Mabea eximia Ducke =Mabea pulcherrima
Mabeafloribimda Jabl. =Mabea montana
Mabea longepedicellata Pittier =Mabea montana
Mabea lucida Pax & K. Hoffm. =Mabea montana
Mabea maynensis Spruce ex Miill. Arg. =Mabea piriri
Mabea montana subsp. biglandulosa (Miill. Arg.) Hollander =Mabea biglandulosa
Mabea montana subsp. lucida (Pax & K. Hoffm.) Hollander =Mabea montana
Mabea muricata Jabl. =Mabea nitida
Mabea nitida var. albiflora Miill. Arg. =Mabea nitida
Mabea occidentalis var. laevigata (Miill. Arg.) Miill. Arg. =Mabea biglandulosa
Mabea occidentalis var. obovata (Miill. Arg.) Miill. Arg. =Mabea speciosa subsp. B
Mabea occidentalis var. purpurascens (Miill. Arg.) Miill. Arg. =Mabea speciosa subsp. B
Mabea orbiculata Jabl. =Mabea frutescens
Mabea pallida Mull. Arg. =Mabea occidentalis
Mabea parguazae Jabl. =Mabea montana,
Mabea parvifolia Pax & K. Hoffm. =Mabea trianae
Mabea piriri var. laevigata Miill. Arg. =Mabea biglandulosa
Mabea piriri var. obovata Miill. Arg. =Mabea speciosa subsp. B
Mabea piriri var. purpurascens Miill. Arg. =Mabea speciosa subsp. B
Mabea saramaccensis Croizat =Mabea speciosa subsp. A
Mabea schomburgkii Benth. =Mabea taquari
Mabea speciosa subsp. concolor (Miill. Arg.) Hollander =Mabea piriri
Mabea taquari var. angustifolia Miill. Arg. =Mabea taquari
Mabea verrucosa Pax & K. Hoffm. =Mabea montana
Manihot diffusa Pohl =Manihot esculenta
Manihot dulcis Pax =Manihot esculenta
Manihot glaziovii Miill. Arg. [not in flora area]
Manihot maguireana D.J. Rogers & Appan =Mamhot tristis subsp. saxicola
Manihot manihot (L.) Cockerell =Manihot esculenta
Manihot melanobasis Miill. Arg. [not in flora area]
Manihot orinocensis Croizat =Manihot tristis subsp. tristis
Manihot rusbyi Britton =Manihot brachyloba
Manihot saxicola Lanj. =Manihot tristis subsp. saxicola
Manihot utilissima Pohl ^Manihot esculenta
Maprounea guianensis var. guianensisf obtusata Miill. Arg. =Maprounea amazonica
Maprounea guianensis var. obtusata (Miill. Arg.) Mull. Arg. =Maprounea amazonica
Meborea guianensis Aubl. [name of uncertain application] =Phyllanthus sp.
Megalostylis poeppigii S. Moore =Dalechampia micrantha
Micrandra cunuri Baill. ex Miill. Arg. =Micrandra spruceana
Micrandra elata (Didr.) Miill. Arg. [see Micrandra brownsbergensis]
Micrandra siphonioides var. minor (Benth.) Mull. Arg. =Micrandra minor
Microstachys guianensis Klotzsch =Microstachys corniculata
Moeroris stipulata Raf. =Phyllanthus stipulatus
Omphalea elaeophoroides Steyerm. ex Standi. =Omphalea diandra
Omphalea megacarpa Hemsl. =Omphalea diandra
Pausandraflagellorachis Lanj. =Pausandra martinii
Pausandraflagellorhachis Lanj. =Pausandra martinii
Pausandra integrifolia Lanj . =Dodecastigma integrifolium
Pera echinocarpa Baill. [not in flora area]
Peraferruginea (Schott) Miill. Arg. =Pera glabrata
Pera nitida (Benth.) Jabl. =Pera decipiens
Pera obovata Baill. [not in flora area]
Pera schomburgkiana (Klotzsch) Mull. Arg. =Pera bicolor
Peridium bicolor Klotzsch =Pera bicolor
Peridium bicolor var. nitidum Benth. =Pera decipiens
Peridium bicolor var. tomentosum Benth. =Pera tomentosa
Peridium decipiens Miill. Arg. =Pera decipiens
Peridium ferrugineum Schott =Pera glabrata
Peridium glabratum Schott =Pera glabrata
Peridium schomburgkianum Klotzsch =Pera bicolor
Phyllanthus adenophyllus Miill. Arg. =Phyllanthus myrsinites
Phyllanthus anadenus Jabl. =Phyllanthus subapicalis subsp. subapicalis
Checklist of the Plants of the Guiana Shield 299
Phyllanthus brachycladus Miill. Arg. =Phyllanthus rupestris
Phyllanthus conami Sw. =Phyllanthus brasiliensis
Phyllanthus congestus Benth. ex Miill. Arg. =Jablonskia congesta
Phyllanthus corcovadensis Miill. Arg. =Phyllanthus tenellus
Phyllanthus cornifolius Kunth =Phyllanthus juglandifolius subsp. cornifolius
Phyllanthus croizatii Steyerm. =Phyllanthus urinaria
Phyllanthus delicatissimus Jabl. =Phyllanthus rupestris
Phyllanthus diffusus Klotzsch =Phyllanthus stipulatus
Phyllanthus dinizii Huber =Phyllanthus myrsinites
Phyllanthus epiphyllanthus L. [not in flora area]
Phyllanthusfrancavillanus Miill. Arg. =Phyllanthus myrsinites subsp. francavillanus
Phyllanthus glaucoviridis Jabl. =Phyllanthus myrsinites
Phyllanthus grandifolius L. [not in flora area]
Phyllanthus guianensis Klotzsch =Phyllanthus caroliniensis subsp. guianensis
Phyllanthus guyanensis Miill. Arg. =Phyllanthus attenuatus
Phyllanthus ichthyomethius Rusby =Phyllanthus piscatorum
Phyllanthus lathyroides Kunth [not in flora area]
Phyllanthus microcarpoides Croizat =Phyllanthus rupestris
Phyllanthus monocladus Urb. =Phyllanthus hyssopifolioides
Phyllanthus nobilis (L. f.) Miill. Arg. =Margaritaria nobilis
Phyllanthus pentrandrus Schumach. & Thonn. [not in flora area]
Phyllanthus pimichinianus Jabl. =Phyllanthus myrsinites subsp. francavillanus
Phyllanthus pseudoconami Miill. Arg. =Phyllanthus piscatorum
Phyllanthus pseudoconami var. pubescens Miill. Arg. =Phyllanthus piscatorum
Phyllanthus schomburgkianus Miill. Arg. ^Phyllanthus caroliniensis subsp. guianensis
Phyllanthus sequoiifolius Jabl. =Phyllanthus subapicalis subsp. sequoiifolius
Phyllanthus vacciniifolius subsp. vinillaensis Steyerm. =Phyllanthus vacciniifolius
Plukenetia abutifolia (Ducke) Pax & K. Hoffm. =Plukenetia polyadenia
Plukenetia buchtienii Pax =Plukenetia brachybotrya
Plukenetia macrostyla Ule =Plukenetia volubilis
Pogonophora schomburgkiana f. elliptica Pax & K. Hoffm. =Pogonophora schomburgkiana
Pogonophora schomburgkiana van longifolia Miers ex Benth. =Pogonophora schomburgkiana
Poinsettia heterophylla (L.) Klotzsch & Garcke =Euphorbia heterophylla var. heterophylla
Poinsettia oerstediana Klotzsch & Garcke =Euphorbia oerstediana
Poraresia anomala Gleason =Pogonophora schomburgkiana
Pseudosenefeldera inclinata (Miill. Arg.) Esser =Senefeldera inclinata
Rhodothyrsus macrophyllus (Ducke) Esser =Senefeldera macrophylla
Rhopalostylis buettnerioides Klotzsch ex Baill. =Dalechampia micrantha
Richeria grandis var. divaricata Miill. Arg. =Richeria grandis
Richeria grandis var. genuina Miill. Arg. =Richeria grandis
Richeria grandis var. obovata Mull. Arg. =Richeria grandis
Richeria grandis var. racemosa Miill. Arg. =Richeria grandis
Richeria laurifolia (Baill.) Baill. =Richeria grandis
Richeria loranthoides (Klotzsch) Miill. Arg. =Podocalyx loranthoides
Richeria obovata (Miill. Arg.) Pax & K. Hoffm. =Richeria grandis
Richeria racemosa (Miill. Arg.) Pax & K. Hoffm. =Richeria grandis
Ricinella triloba Miill. Arg. =Adelia triloba
Sagotia racemosa var. brachysepala Miill. Arg. =Sagotia brachysepala
Sagotia racemosa var. genuina Miill. Arg. =Sagotia racemosa
Sagotia racemosa var. ligularis Miill. Arg. =Sagotia racemosa
Sagotia racemosa var. microsepala Miill. Arg. =Sagotia racemosa
Sagotia tafelbergii Croizat =Sagotia racemosa
Sapium albomarginatum Pax & K. Hoffm. =Sapium glandulosum
Sapium aubletianum (Miill. Arg.) Huber [nom. illeg.] =Sapium glandulosum
Sapium aucuparium Jacq. [nom. illeg.] [see Sapium glandulosum] =Sapium glandulosum
Sapium biglandulosum (L.) Mull. Arg. [nom. illeg.] [see Sapium glandulosum] =Sapium glandulosum
Sapium biglandulosum var. aubletianum Miill. Arg. [nom. illeg.] =Sapium glandulosum
Sapium biglandulosum var. klotzschianum Miill. Arg. =Sapium glandulosum
Sapium biglandulosum var. lanceolatum Miill. Arg. =Sapium glandulosum
Sapium cladogyne Hutch. =Sapium jenmanii
Sapium ellipticum Hemsl. [see Sapium glandulosum]
Sapium guaricense Pittier =Sapium glandulosum
Sapium hemsleyanum Huber =Sapium glandulosum
300 Checklist of the Plants of the Guiana Shield
Sapium hippomane G. Mey. [nom. illeg.] =Sapium glandulosiim
Sapium klotzschiamim (Mull. Arg.) Huber =Sapiiim glandiilosum
Sapium lanceolatiim (Miill. Arg.) Huber =Saphim glandulosum
Sapium microdentatum Lanj . =Sapium paucinervium
Sapium montanum Lanj. =Sapium argutum
Sapium moritzianum Klotzsch =Sapium glandulosum
Sapium naiguatense Pittier =Sapium glandulosum
Sapium obtusilobum Miill. Arg. =Sapium glandulosum
Sapium paucistamineum Pittier =Sapium glandulosum
Sapium prunifolium Klotzsch =Sapium glandulosum
Sapium yutajense Jabl. =Dendwthhx yutajensis
Sebastiania corniculata (Vahl) Miill. Arg. =Microstachys corniculata
Sebastiania corniculata var. guyanensis (Klotzsch) Pax & K. Hoffm. =Microstachys corniculata
Sebastiania linearifolia Lanj. =Sebastiania bidentata
Sebastiania micrantha (Benth.) Lanj. =Microstachys corniculata
Securinega congesta (Benth. ex Mull. Arg.) Miill. Arg. =Jablonskia congesta
Senefeldera chiribiquetensis R. Schult. & Croizat =Senefelderopsis chiribiquetensis
Senefeldera karsteniana Pax & K. Hoffm. =Senefeldera inclinata
Senefeldera multiflora var. acutifolia Miill. Arg. =Senefeldera inclinata
Senefeldera yutajensis (Jabl.) G.L. Webster =Dendrothrix yutajensis
Senefelderopsis sipapoensis Jabl. =Senefelderopsis croizatii
Senefelderopsis venamoensis Jabl. =Senefelderopsis croizatii
Siphonia lutea Spruce ex Benth. ^Hevea guianensis var. lutea
Siphonia pauciflora Spruce ex Benth. =Hevea pauciflora
Spixia distichophylla Mart. =Pera distichophylla
Stilaginella laxiflora Tul. =Hieronyma alchorneoides var. alchorneoides
Stilaginella oblonga Tul. ^Hieronyma oblonga
Traganthus sidoides Klotzsch =Bernardia sidoides
Tragia corniculata Vahl =Microstachys corniculata
Tragia grandifolia Klotzsch =Adenophaedra grandifolia
Tragiajapurensis Mull. Arg. =Tragia fendleri
Tragiopsisfruticulosa H. Karst =Microstachys corniculata
Veconcibea latifolia (Benth.) Pax & K. Hoffm. =Conceveiba latifolia
References
Burger, W. & M. Huft. Euphorbiaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 36: 1-169.
Den Hollander, G. & C.C. Berg. 1986. Studies on the flora of the Guianas 21. Mabea species of the Guianas - a precursor. Proc.
Kon. Ned. Akad. Wet. ser. C, 89(2): 147-157.
Franco, P. 1990. The genus Hyeronima in South America. Bot. Jahrb. Syst. Ill: 297-346.
Gillespie, L.J. 1993. Euphorbiaceae of the Guianas: Annotated species checklist and key to the genera. Brittonia 45(1): 56-94.
Gillespie, L.J. 1993. A synopsis of the Neotropical Plukenetia including two new species. Syst. Bot. 18(4): 575-592.
Gillespie, L.J. 1994. A new section and two new species of Tragia from Venezuelan Guayana and French Guiana. Novon 4(4):
330-338.
Gillespie, L.J. 2002. Euphorbiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
266-298.
Gillespie, L.J. & W.S. Armbruster. 1997. A contribution to the Guianan Flora: Dalechampia, Haematostemon, Omphalea, Pera,
Plukenetia, and Tragia, with notes on subfamily Acalyphoideae. Smithsonian Contributions to Botany 86.
Gorts-van Rijn, A.R.A. 1 976. Euphorbiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2):
387-424.
Hayden, W.J. 1990. Notes on Neotropical Amanoa. Brittonia 42(4): 260-270.
Jablonski, E. 1965. Euphorbiaceae. In B. Maguire & collaborators. Botany of the Guayana Highland Part 6. Mem. New York
Bot. Gard. 12(3): 1-285.
Jablonski, E. 1967. Euphorbiaceae. In B. Maguire & collaborators. Botany of the Guayana Highland Part 7. Mem. New York
Bot. Gard. 17(1): 1-439.
Lanjouw, J. 1932. Euphorbiaceae. In A. PuUe, Flora of Suriname 2(1): 1-101.
Lanjouw, J. 1939. Euphorbiaceae (additions and corrections). In A. PuUe, Flora of Suriname 2(1): 457-470.
Rogers, D.J. & S.G. Appan. 1973. Manihot and Manihotoides. Flora Neotropica Monogr. 13: 1-272.
Webster, G.L. & W.S. Armbruster. 1991. A synopsis of the neotropical species of Dalechampia. J. Linn. Soc. Bot. 105: 137-177.
Webster, G.L., RE. Berry, W.S. Armbruster, H.-J. Esser, L.J. Gillespie, W.J. Hayden, G.A. Levin, R. de S. Secco, & S.V. Heald.
1999. Euphorbiaceae.In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 5: 72-
228.
Checklist of the Plants of the Guiana Shield 301
EUPHRONIACEAE
L. Marcano-Berti, J.A. Steyermarkt
Euphronia acuminatissima Steyerm. AM BO
Euphronia guianensis (R.H. Schomb.) Hallier f. {=Euphronia hirtelloides s.l.] BO GU
Euphronia hirtelloides Mart, ex Mart. & Zucc. AM
Synonyms and Misidentifications
Euphronia licanioides (Spruce ex Warm.) Hallier f. =Euphronia hirtelloides
Lightia guianensis R.H. Schomb. =Euphronia guianensis
Lightia licanioides Spruce ex Warm. =Euphronia hirtelloides
References
Lleras, E. 1976. Revision and taxonomic position of the genus Euphronia Martius ex Martius & Zuccarini (Vochysiaceae). Acta
Amaz. 6: 43-47.
Marcano-Berti, L. 1989. Euphroniaceae: una nueva familia. Pittieria 18: 15-17.
Marcano-Berti, L. 1998. Euphroniaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 21: 45-48.
Steyermark, J.A. & L. Marcano-Berti. 1999. Euphroniaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora
of the Venezuelan Guayana 5: 228-230.
Flacourtiaceae
G.A. Aymard C, RE. Berry, R.W. Kiger, M. Olson
Banara guianensis Aubl.
Banara nitida Spruce ex Benth.
Banara orinocensis (Cuatrec.) Sleumer
Carpotroche crispidentata Ducke
Carpotroche grandiflora Spruce ex Benth.
Carpotroche longifolia (Poepp.) Benth.
Carpotroche surinamensis Uittien
Casearia aculeata Jacq.
Casearia acuminata DC.
Casearia arborea (Rich.) Urb.
Casearia celastroides Klotzsch [nomen nudum]
Casearia combaymensis Tul. [Guianan material often det. C bracteifera or C. AM
singularis'\
Casearia commersoniana Cambess.
Casearia decandra Jacq.
Casearia duckeana Sleumer
Caseariafasciculata (Ruiz & Pav.) Sleumer
Casearia grandiflora Cambess.
Casearia guianensis (Aubl.) Urb.
Casearia hirsuta Sw.
Casearia javitensis Kunth
Casearia mariquitensis Kunth
Casearia mollis Kunth
Casearia neblinae Sleumer
Casearia negrensis Eichler
Casearia pitumba Sleumer
Casearia prunifolia Kunth
Casearia resinifera Spruce ex Eichler
Casearia rusbyana Briq.
Casearia spinescens (Sw.) Griseb.
Casearia spruceana Benth. ex Eichler
Casearia sylvestris Sw.
Casearia tremula (Griseb.) Griseb. ex Wright
Casearia ulmifolia Vahl ex Vent.
Casearia zizyphoides Kunth
Euceraea nitida Mart.
Euceraea rheophytica P.E. Berry & M. Olson
Euceraea sleumeriana Steyerm. & Maguire
Hasseltiafloribunda Kunth
Hecatostemon completus (Jacq.) Sleumer
Homalium guianense (Aubl.) Oken
AM BO GU su FG
BO DA
AM
AM
AM
BO DA
su
FG
GU su FG
GU su FG
AM BO DA GU su
GU? su FG
AM BO GU
GU
su FG
DA GU su FG
AM BO DA GU su FG
AM BO DA GU
GU
GU?
su FG
AM BO GU su FG
BO DA GU su FG
AM BO su
AM BO GU su FG
AM BO
BO
GU su FG
AM
su FG
BO DA GU su FG
AM FG
FG
BO DA GU su FG
BO DA GU
AM
AM BO
BO
DA GU
GU
su FG
AM BO GU su FG
AM BO GU su FG
AM BO GU su
AM
BO
BO GU
FG?
BO DA GU su FG
302 Checklist of the Plants of the Guiana Shield
Homalium racemosum Jacq.
Laetia americana L.
Laetia coriacea Spruce ex Benth.
Laetia cupulata Spruce ex Benth.
Laetia procera (Poepp.) Eichler
Laetia suaveolens (Poepp.) Benth.
Lindackeria latifolia Benth.
Lindackeria paludosa (Benth.) Gilg
Lunania parviflora Spruce ex Benth.
Mayna grandifolia (H. Karst.) Warb.
Mayna odorata Aubl.
Neoptychocarpiis apodanthiis (Kuhlm.) Buchheim
Neoptychocarpus killipii (Monach.) Buchheim
Neospnicea paterna Alford, sp. nov. ined.
Prockia criicis P. Browne ex L. [expected in the Guianas]
Ryania angustifolia (Turcz.) Monach.
Ryania dentata (Kunth) Miq.
Ryania dentata van dentata
Ryania pyrifera (Rich.) Uittien & Sleumer
Ryania speciosa Vahl
Ryania spruceana Monach.
Xylosma benthamii (Tul.) Triana & Planch.
Xylosma ciliatifoliiim (Clos) Eichler
Xylosma tessmannii Sleumer
Zuelania giiidonia (Sw.) Britton & Millsp.
AM BO
BO
DA GU SU FG
AM
AM
BO
GU
AM
AM
AM
BO DA GU
GU
GU
SU
SU
FG
AM
AM
BO GU
GU
FG?
GU SU
SU
FG
FG
FG
GU FG?
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
GU SU FG
GU SU FG
GU SU FG
GU FG?
GU FG
Synonyms and Misidentifications
Athenaea guianensis (Aubl.) J.F. Gmel. =Casearia guianensis
Azara umhellata C. Presl =Laetia americana
Banara glandiilosa (Desv. ex Ham.) Speg. =Banara guianensis
Banara guianensis auct. non Aubl. 1775 =Banara nitida
Banara guianensis var. mollis (Poepp.) Eichler =Banara guianensis
Banara mollis (Poepp.) Tul. =Banara guianensis
Banara sacupanensis Aristeg. =Banara orinocensis
Carpotroche amazonica Mart. =Carpotroche grandiflora
Carpotroche amazonica auct. non Mart. 1871 =Mayna grandifolia (p.p.)
Carpotroche laxiflora (Benth.) Walp. =Lindackeria paludosa
Carpotroche paludosa (Benth.) Walp. =Lindackeria paludosa
Carpotroche zuliana Pittier =Mayna grandifolia
Casearia aculeata auct. non Jacq. 1 760 ^Casearia spinescens
Casearia arguta auct. non Kunth 1821 [1823], sensu Pittier et al. =Casearia mollis
Casearia arguta Kunth [not in flora area]
Casearia arguta auct. non Kunth 1821 [1823], sensu Steyerm. =Casearia guianensis
Casearia bracteifera Sagot =Casearia combaymensis
Casearia cambessedesii Eichler =Casearia ulmifolia
Casearia carpinifolia Benth. =Casearia sylvestris
Casearia celtidifolia Poepp. ex Eichler non Kunth 1823 =Casearia ulmifolia
Casearia celtidifolia Kunth =Casearia ulmifolia
Casearia cotticensis Uittien =Casearia ulmifolia
Casearia densiflora Benth. =Casearia commersoniana
Casearia dentata (Aubl.) Sesse & M09. =Casearia commersoniana
Casearia guidonia (Sw.) Lundell non Benth. =Zuelania guidonia
Casearia inaequilatera auct. non Cambess. 1829 [1830] =Casearia sylvestris
Casearia laevigata Willd. ex Klotzsch [nomen nudum] =Casearia zizyphoides
Casearia lasiophylla auct. non Eichler 1871 =Casearia mollis
Casearia laurifolia Benth. =Casearia commersoniana
Casearia lingua Cambess. =Casearia sylvestris
Casearia macrophylla Vahl =Casearia pitumba
Casearia mollis auct. non Kunth 1821 [1823] =Casearia hirsuta
Casearia negrensis auct. non Eichler 1871 =Laetia suaveolens
Casearia nitida Jacq. [not in flora area]
Casearia petraea Benth. =Casearia ulmifolia
Casearia singularis Eichler =Casearia combaymensis
Casearia spinescens auct. non (Sw.) Griesb. 1863, sensu Pittier et al. =Casearia guianensis
Checklist of the Plants of the Guiana Shield 303
Casearia sylvesths var. carpinifolia (Benth.) Briq. =Casearia sylvestris
Caseaha sylvestris var. lingua (Cambess.) Eichler =Casearia sylvestris
Casearia sylvestris var. sylvestris =Casearia sylvestris
Casinga procera (Poepp.) Griseb. =Laetia procera
Casinga sitaveolens (Poepp.) Griseb. ex Benth. ^Laetia suaveolens
Chaetocraterjavitensis (Kunth) Raf. =Casearia javitensis
Dendrostylis grandifolia H. Karst. =Mayna grandifolia
Flacourtia benthamii Tul. =Xylosma benthamii
Flacourtia nitida Benth. =Xylosma benthamii
Guidonia apetala (Jacq.) Kuntze =Laetia americana
Guidonia calophylla (Eichler) Kuntze =Laetia suaveolens
Guidonia coriacea (Spruce ex Benth.) Kuntze =Laetia coriacea
Guidonia cupulata (Spruce ex Benth.) Kuntze =Laetia cupulata
Guidonia hirsuta (Sw.) M. Gomez =Casearia hirsuta
Guidonia procera (Poepp.) Kuntze =Laetia procera
Guidonia spinescens (Sw.) Griseb. =Casearia spinescens
Guidonia suaveolens (Poepp.) Kuntze ^Laetia suaveolens
Guidonia tremula (Griseb.) Maza =Casearia tremula
Guidonia ulmifolia (Vahl ex Vent.) Baill. ^Casearia ulmifolia
Hecatostemon dasygynus S.F. Blake =Hecatostemon completus
Hecatostemon guazumifolius (Kunth) Sleumer =Hecatostemon completus
Hisingera benthamii (Tul.) Clos =Xylosma benthamii
Homalium anzoateguiense Steyerm. =^Homalium racemosum
Homalium densiflorum Spruce ex Benth. =Homalium guianense
Homalium eleuthrostylum S.F. Blake =Homalium racemosum
Homalium eurypetalum S.F. Blake =Homalium racemosum
Homalium mituense Cuatrec. =Homalium racemosum
Homalium napimoga Spreng. =Homalium guianense
Homalium pedicellatum Spruce ex Benth. =Homalium racemosum
Homalium pittieri S.F. Blake =Homalium racemosum
Homalium puberulum Klotzsch =Homalium guianense
Homalium racoubea Sw. =Homalium guianense
Iroucana guianensis Aubl. =Casearia guianensis
Kuhlia mollis Poepp. =Banara guianensis
Laetia acuminata Bonpl. ex Triana & Planch. =Laetia americana
Laetia apetala Jacq. =Laetia americana
Laetia calophylla Eichler =Laetia suaveolens
Laetia casearioides Sagot ex Benth. =Laetia procera
Laetia completa Jacq. =Hecatostemon completus
Laetia guazumaefolia Kunth =Hecatostemon completus
Laetia guidonia Sw. =Zuelania guidonia
Laetia hirtella Kunth =Hecatostemon completus
Laetia obtusifolia Eichler ex Benoist [nomen nudum] =Laetia procera
Lindackeria latifolia auct. non Benth. 1851 =Lindackeria paludosa
Lindackeria maynensis var. laxiflora (Benth.) Mart, ex Pittier et al. =Lindackeria paludosa
Mayna denticulata Benth. =Mayna odorata
Mayna grandiflora (Spruce ex Benth.) R.E. Schult. =Carpotroche grandiflora
Mayna latifolia (Benth.) Benth. =Lindackeria latifolia
Mayna laxiflora Benth. =Lindackeria paludosa
Mayna paludosa Benth. =Lindackeria paludosa
Mayna zuliana (Pittier) A. Robyns =Mayna grandifolia
Myroxylon benthamii (Tul.) Kuntze =Xylosma benthamii
Napimoga guianensis Aubl. =Homalium guianense
Oncoba latifolia (Benth.) Eichler =Lindackeria latifolia
Oncoba maynensis var. laxiflora (Benth.) Eichler =Lindackeria paludosa
Patrisia acuminata (Spruce ex Eichler) Kuntze =Ryania angustifolia
Patrisia affinis Kunth =Ryania dentata
Patrisia bicolor DC. =Ryania speciosa
Patrisia dentata Kunth =Ryania dentata
Patrisia parviflora DC. =Ryania speciosa
Patrisia pyrifera Rich. =Ryania pyrifera
Patrisia speciosa (Vahl) Monach. =Ryania speciosa
Patrisia spruceana (Monach.) Lemee [not in flora area] =Ryania spruceana
Patrisia tomentosa (Miq.) M. Roem. =Ryania speciosa
304 Checklist of the Plants of the Guiana Shield
Piparea dentata Aubl. =Casearia commersoniana
Pitumba gidanensis Aubl. =Casearia pitumba
Prockia ohnocensis Cuatrec. =Banara orinocensis
Ptychocarpus apodanthus Kuhlm. =Neoptychocarpus apodanthus
Racoubea gidanensis Aubl. =HomaUum gidanense
Ryania acuminata Spruce ex Eichler =Ryania angusdfolia
Ryania bicolor (DC.) Eichler =Ryania speciosa
Ryania candollei Miq. =Ryama speciosa
Ryania casiquiarensis Steyerm. =Ryania speciosa
Ryania kunthii Miq. =Ryania dentata
Ryania mansoana auct. non Eichler 1 87 1 =Ryania speciosa
Ryania parviflora (DC.) Griseb. =Ryania speciosa
Ryania pyrifera auct. non (Rich.) Uitten & Sleumer 1935, sensu J.F. Macbr. =Ryania speciosa
Ryania pyrifera var. subuliflora Sandwith =Ryania speciosa
Ryania pyrifera var. tomentosa (Miq.) Sleumer =Ryania speciosa
Ryania sagotiana Eichler =Ryania pyrifera
Ryania sagotina auct. non Eichler 1871 =Ryania dentata
Ryania speciosa var. bicolor (A. DC.) Monach. =Ryania speciosa
Ryania speciosa var. minor Monach. =Ryania speciosa
Ryania speciosa var. stipularis (Linden & Planch.) Monach. =Ryania speciosa
Ryania speciosa var. subuliflora (Sandwith) Monach. =Ryama speciosa
Ryania speciosa var. tomentosa (Miq.) Monach. =Ryania speciosa
Ryania stipularis auct. non Linden & Planch. 1863 =Ryania spruceana
Ryania stipularis Linden & Planch. =Ryania speciosa
Ryania tomentosa Miq. =Ryania speciosa
Samyda arborea Rich. =Casearia arborea
Samyda guidonia L. (Meliaceae) =Guarea guidonia
Samyda hirsuta (Sw.) Poir. =Casearia hirsuta
Samyda petiolaris Spruce ex Eichler =Laetia suaveolens
Samyda pitumba Poir. =Casearia pitumba
Samyda procera Poepp. =Laetia procera
Samyda silvestris (Sw.) Poir. =Casearia sylvestris
Samyda spinescens Sw. =Casearia spinescens
Samyda suaveolens Poepp. =Laetia suaveolens
Tetracoryne angusdfolia Turcz. =Ryania angusdfolia
Xyladenius gladulosus Desv. ex Ham. =Banara guianensis
Xylosma niddum (Hell.) A. Gray ex Griseb. [not in flora area]
Xylosma pallidifolium Sleumer ex Steyerm. & O. Huber =Xylosma benthamd
Zuelania tremula Griseb. =Casearia tremula
References
Mori, S.A. & B. Fischer. 2002. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 319-
328.
Olson, M., RE. Berry, & G.A. Aymard C. 1999. Flacourtiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 5: 434-474.
Sleumer, H.O. 1980. Flacourtiaceae. Flora Neotropica Monogr. 22: 1-499.
Uittien, H. & H.O. Sleumer, 1935. Flacourtiaceae. In A. Pulle, Flora of Suriname 3(1): 282-303.
Gentianaceae
V.A. Albert, M.J. Jansen-Jacobs, P.J.M. Maas, O. Pihlar, L.Struwe
Adenolisianthus arboreus (Spruce ex Progel) Gilg
Celiantha bella Maguire & Steyerm.
Celiantha chimantensis (Steyerm. & Maguire) Maguire
Celiantha imthurniana (Oliv.) Maguire [Mt. Roraima]
Chelonanthus alatus (Aubl.) Pulle
Chelonanthus albus (Spruce ex Progel) V.M. Badillo
Chelonanthus angusdfolius (Kunth) Gilg
Chelonanthus longistylus (J.G.M. Pers. & Maas) Struwe & V.A. Albert
Chelonanthus purpurascens (Aubl.) Struwe, S. Nilsson & V.A. Albert
Chelonanthus schomburgkii (Griseb.) Gilg
Chelonanthus viridiflorus (Mart.) Gilg
Chorisepalum carnosum Ewan
AM
BO
BO GU
AM BO DA GU SU FG
AM BO GU
BO GU SU FG
AM BO GU SU FG
AM
BO
GU
GU
GU
Checklist of the Plants of the Guiana Shield 305
Chorisepalum ovatum Gleason
Chorisepalum psychotrioides Ewan BO GU
Chorisepalum rotundifolium Ewan BO GU
Chorisepalum sipapoanum (Maguire) Stmwe & V.A. Albert AM SU
Coutoubea humilis Sandwith GU
Coutoubea minor Kunth
Coutoubea ramosa A\xh\. AM BO DA GU SU FG
Coutoubea reflexaBQnXh. AM BO GU
Coutoubea spicata A\xh\. AM BO GU SU FG
Curtia conferta (Mart.) Knobl.
Curtia obtusifolia (Benth.) Knobl.
Curtia tenuifolia{Avih\.) Kiiohl. AM BO GU SU? FG
Enicostema verticillatum (L.) Engl, ex Gilg BO GU
Irlbachia cardonae (Gleason) Maguire AM BO GU
Irlbachia nemorosa (Willd. ex Roem. & Schult.) Merr. AM BO GU
Irlbachia phelpsiana Maguire
Irlbachia plantaginifolia Maguire
Irlbachia poeppigii (Griseb.) L. Cobb & Maas
Irlbachia pratensis (Kunth) L. Cobb & Maas
Irlbachia pumila (Benth.) Maguire
Irlbachia tatei (Gleason) Maguire AM BO GU
Macrocarpaea autanae Weaver
Macrocarpaea ayangannae J.R. Grant, Stmwe & J.K. Boggan [Mt. GU
Ayanganna]
Macrocarpaea marahuacae Struwe & V.A. Albert
Macrocarpaea neblinae Maguire & Steyerm.
Macrocarpaea piresii Maguire
Macrocarpaea rugosa Steyerm. BO GU
Neblinantha neblinae Maguire
Neblinantha parvifolia Maguire
Neurotheca loeselioides (Spruce ex Progel) Baill. AM GU SU FG
Neurotheca loeselioides subsp. loeselioides GU SU
Potalia amara Aubl. SU FG
Potalia elegans Struwe & V.A. Albert
Potalia maguireorum Struwe & V.A. Albert
Potalia resinifera Mart.
Rogersonanthus arboreus (Britton) Maguire & B.M. Boom
Rogersonanthus coccineus Steyerm. ex Struwe, S. Nilsson & V.A. Albert
Rogersonanthus quelchii (N.E. Br.) Maguire & B.M. Boom AM GU
Saccifolium bandeirae Maguire & Pires
Schidtesia benthamiana Klotzsch ex Griseb. BO GU SU?
Schultesia brachyptera Cham. AM BO GU SU FG
Schultesia guianensis (Aubl.) Malme BO GU SU FG
Schultesia pohliana Progel AM BO GU SU
Schultesia subcrenata Klotzsch ex Griseb. BO GU
Sipapoantha ostrina Maguire & B.M. Boom
Symbolanthus argyreus (Maguire) Struwe & K. Gould
Symbolanthus aureus Struwe & V.A. Albert
Symbolanthus camanensis Maguire & B.M. Boom
Symbolanthus elisabethae (M.R. Schomb.) Gilg AM BO GU
Symbolanthus huachamacariensis Steyerm.
Symbolanthus rosmarinifolius Struwe & V.A. Albert
Symbolanthus sessilis Steyerm. & Maguire
Symbolanthus yaviensis Steyerm.
Tachia gracilis Benth. AM GU
Tachia grandiflora Maguire & Weaver FG
Tachia grandifolia Maguire & Weaver
Tachia grandifolia van grandifolia
Tachia grandifolia van orientalis Maguire & Weaver
Tachia guianensis Aub\. GU SU FG
Tachia longipes L. Cobb & Maas SU
Tachia schomburgkiana Benth. BO GU
Tachia smithii Maguire & Weaver GU
Tapeinostemon breweri Steyerm. & Maguire
AM BO
AM
AM
AM
AM
BO
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM BO
AM
AM
AM
AM
BO
AM
AM
AM
BO
AM BO
AM BO
AM BO
AM BO
AM BO
306 Checklist of the Plants of the Guiana Shield
Tapeinostemonjauaensis Steyerm. & Maguire BO
Tapeinostemon longiflorum Maguire & Steyerm. AM
Tapeinostemon longiflorum var. australe Maguire & Steyerm. [Sierra de la AM
Neblina]
Tapeinostemon longiflorum var. longiflorum [Sierra de la Neblina] AM
Tapeinostemon rugosum Maguire & Steyerm. [Macizo del Chimanta] BO
Tapeinostemon sessiliflorum (Humb. & Bonpl. ex Schult.) Pruski & S.F. Sm. AM
Tapeinostemon spenneroides Benth.
Tetrapollinia caerulescens (Aubl.) Maguire & B.M. Boom
Voyria acuminata Benth.
Voyria aphylla (Jacq.) Pers.
Voyria aurantiaca Splitg.
Voyria caerulea Aubl.
Voyria chionea Benth.
Voyria clavata Splitg.
Voyria corymbosa Splitg.
Voyria corymbosa subsp. alba (Standi.) Ruyters & Maas
Voyria corymbosa subsp. corymbosa
Voyriaflavescens Griseb.
Voyria pittieri (Standi.) L.O. Williams
Voyria rosea Aubl.
Voyria spruceana Benth.
Voyria tenella Hook.
Voyria tenuiflora Griseb.
Voyriella parviflora (Miq.) Miq.
Synonyms and Misidentifications
Adenolisianthus virgatus (Progel) Gilg =Chelonanthus alatus
Apophragma tenuifolium (Aubl.) Griseb. =Curtia tenuifolia
Brachycodon pumilus (Benth.) Progel =Irlbachia pumila
Brachycodon ramosissimus (Benth.) Progel =Irlbachia poeppigii
Calolisianthus imthurnianus (Oliv.) Gleason =Celiantha imthurniana
Calolisianthus tatei Gleason =Irlbachia tatei
Calolisianthus tepuiensis Gleason ^Rogersonanthus arboreus
Chelonanthus acutangulus (Ruiz & Pav.) Gilg =Chelonanthus alatus
Chelonanthus arboreus Britton =Rogersonanthus arboreus
Chelonanthus bifidus (Kunth) Gilg =Chelonanthus angustifolius
Chelonanthus campanuloides (Spruce ex Benth.) Gilg =Irlbachia pratensis
Chelonanthus cardonae Gleason =Irlbachia cardonae
Chelonanthus chelonoides (L. f.) Gilg =Chelonanthus alatus
Chelonanthus fruticosus Maguire & B.M. Boom =Adenolisianthus arboreus
Chelonanthus pyriformis Gleason =Chelonanthus angustifolius
Chelonanthus uliginosus (Griseb.) Gilg =Chelonanthus purpurascens
Chorisepalum acuminatum Steyerm. =Chorisepalum psychotrioides
Chorisepalum breweri Steyerm. & Maguire =Chorisepalum ovatum
Chorisepalum ovatum var. sipapoanum Maguire =Chorisepalum sipapoanum
Chorisepalum psychotrioides var. acuminatum (Steyerm.) Maguire =Chorisepalum psychotrioides
Claytonia nemorosa Willd. ex Roem. & Schult. =Irlbachia nemorosa
Coutoubea racemosa G. Mey. =Coutoubea ramosa
Coutoubea ramosa var. longifolia Benth. =Coutoubea ramosa
Coutoubea ramosa f. racemosa (G. Mey.) Jonker =Coutoubea ramosa
Coutoubea ramosa var. racemosa (G. Mey.) Benth. =Coutoubea ramosa
Curtia quadrifolia Maguire =Curtia conferta
Curtia tenella (Mart.) Cham. =Curtia tenuifolia
Curtia tenuifolia subsp. tenella (Mart.) Grothe & Maas =Curtia tenuifolia
Ditomaga nemorosa (Willd. ex Roem. & Schult.) Raf. =Irlbachia nemorosa
Exacum guianense Aubl. =Schultesia guianensis
Exacum racemosum (G Mey.) Roem. & Schult. =Coutoubea ramosa
Exacum spicatum (Aubl.) Vahl =Coutoubea spicata
Exacum tenuifolium Aubl. =Curtia tenuifolia
Gentiana aphylla Jacq. =Voyria aphylla
Gentiana verticillata L. =Enicostema verticillatum
Helia acutangula (Ruiz & Pav.) Kuntze ^Chelonanthus alatus
Helia alata (Aubl.) Kuntze =Chelonanthus alatus
BO GU su
BO GU
GU
su FG
BO DA GU su EG
BO DA GU su FG
BO GU su FG
GU su FG
BO GU su FG
BO GU su FG
BO GU su FG
BO su FG
BO DA GU
GU su FG
DA GU su FG
BO GU su FG
GU su FG
BO GU su FG
Checklist of the Plants of the Guiana Shield 307
Helia alba (Spruce ex Progel) Kuntze =Chelonanthus albus
Helia angustifolia (Kunth) Kuntze =Chelonanthus angustifolius
Helia arborea (Spruce ex Progel) Kuntze =Adenolisianthus arboreus
Helia bifida (Kunth) Kuntze =Chelonanthus angustifolius
Helia caerulescens (Aubl.) Kuntze =Tetrapollinia caerulescens
Helia campanuloides (Spruce ex Benth.) Kuntze =Irlbachia pratensis
Helia chelonoides (L. f.) Kuntze =Chelonanthus alatus
Helia elisabethae (M.R. Schomb.) Kuntze =Symbolanthus elisabethae
Helia nemorosa (Willd. ex Roem. & Schult.) Kuntze =Irlbachia nemorosa
Helia pratensis (Kunth) Kuntze =Irlbachia pratensis
Helia pumila (Benth.) Kuntze =Irlbachia pumila
Helia purpurascens (Aubl.) Kuntze =Chelonanthus purpurascens
Helia ramosissima (Benth.) Kuntze =Irlbachia poeppigii
Helia schomburgkii (Griseb.) Kuntze =Chelonanthus schomburgkii
Helia trifidus (Kunth) Kuntze =Chelonanthus alatus
Helia uliginosa (Griseb.) Kuntze =Chelonanthus purpurascens
Irlbachia alata (Aubl.) Maas =Chelonanthus alatus
Irlbachia alata subsp. alata =Chelonanthus alatus
Irlbachia alata subsp. alba (Spruce ex Progel) J.G.M. Pers. & Maas =Chelonanthus albus
Irlbachia alata subsp. angustifolia (Kunth) J.G.M. Pers. & Maas =Chelonanthus angustifolius
Irlbachia alata subsp. arborea (Spruce ex Progel) J.G.M. Pers. & Maas =Adenolisianthus arboreus
Irlbachia alata subsp. longistyla J.GM. Pers. & Maas =Chelonanthus longistylus
Irlbachia alata subsp. viridifora (Mart.) J.G.M. Pers. & Maas =Chelonanthus viridiflorus
Irlbachia bonplandiana Fenzl =Irlbachia nemorosa
Irlbachia caerulescens (Aubl.) Griseb. =Tetrapollinia caerulescens
Irlbachia paruana Maguire =Irlbachia cardonae
Irlbachia purpurascens (Aubl.) Maas =Chelonanthus purpurascens
Irlbachia quelchii (N.E. Br.) Maas =Rogersonanthus quelchii
Irlbachia ramosissima (Benth.) Maguire =Irlbachia poeppigii
Irlbachia recurva (Benth.) Progel =Irlbachia nemorosa
Irlbachia subcordata (Benth.) Progel =Irlbachia nemorosa
Leiothamnus elisabethae M.R. Schomb. ^Symbolanthus elisabethae
Leiphaimos alba Standi. =Voyria corymbosa subsp. alba
Leiphaimos aphylla (Jacq.) Gilg =Voyria aphylla
Leiphaimos aurantiaca (Splitg.) Miq. =Voyria aurantiaca
Leiphaimos calycina (Splitg. ex A. DC.) Miq. =Voyria caerulea
Leiphaimos chionea (Benth.) Gilg =Voyria chionea
Leiphaimosflavescens (Griseb.) Gilg =Voyriaflavescens
Leiphaimos leucantha (Miq.) Gilg =Voyria tenuiflora
Leiphaimos montana Jonker =Voyria corymbosa subsp. alba
Leiphaimos obconica auct., non (Progel) Gilg =Voyria tenella
Leiphaimos pittieri Standi. -Voyria pittieri
Leiphaimos spruceana (Benth.) Gilg =Voyria spruceana
Leiphaimos tenella (Hook.) Miq. =Voyria tenella
Leiphaimos tenuiflora (Griseb.) Miq. =Voyria tenuiflora
Lisianthus acutangulus Ruiz & Pav. =Chelonanthus alatus
Lisianthus acutilobus Steyerm. =Irlbachia cardonae
Lisianthus alatus Aubl. =Chelonanthus alatus
Lisianthus albus Spruce ex Progel =Chelonanthus albus
Lisianthus angustifolius Kunth =Chelonanthus angustifolius
Lisianthus arboreus Spruce ex Progel =Adenolisianthus arboreus
Lisianthus bifidus Kunth =Chelonanihus angustifolius
Lisianthus caerulescens Aubl. =Tetrapollinia caerulescens
Lisianthus campanuloides Spruce ex Benth. =Irlbachia pratensis
Lisianthus cardonae (Gleason) Steyerm. =Irlbachia cardonae
Lisianthus chelonoides L. f. =Chelonanthus alatus
Lisianthus chimantensis Steyerm. & Maguire ^'Celiantha chimantensis
Lisianthus elisabethae (M.R. Schomb.) Griseb. =Symbolanthus elisabethae
Lisianthus fistulosus Poir. =Chelonanthus alatus
Lisianthus grandiflorus Aubl. =Chelonanthus alatus
Lisianthus imthurnianus 01 i v. =Celiantha imthurniana
Lisianthusjauaensis Steyerm. & Maguire =Irlbachia tatei
Lisianthus pratensis Kunth =Irlbachia pratensis
Lisianthus pumilus Benth. =Irlbachia pumila
308 Checklist of the Plants of the Guiana Shield
Lisianthus purpurascens Aubl. =Chelonanthus purpurascens
Lisianthus pyriformis (Gleason) Steyerm. =Chelonanthus angustifolius
Lisianthus quelchii N.E. Br. =Rogersonanlhus qiielchii
Lisianthus ramosissimus Benth. =Irlbachia poeppigii
Lisianthus recwvus Benth. =Irlbachia nemorosa
Lisianthus scabridulus Steyerm. =lrlbachia tatei
Lisianthus schomburgkii Griseb. =Chelonanthus schomburgkii
Lisianthus subcordatus Benth. =Irlbachia nemorosa
Lisianthus tatei (Gleason) Steyerm. =Irlbachia tatei
Lisianthus trijidus Kunth =Chelonanthus alatus
Lisianthus uliginosus Griseb. =CheIonanthus purpurascens
Lisianthus virgatus Progel ='Chelonanthus alatus
Lisyanthus alatus Aubl. =Chelonanthus alatus
Lisyanthus angustifolius (Kunth) Griseb. =Chelonanthus angustifolius
Lisyanthus bifidus (Kunth) Griseb. =Chelonanthus angustifolius
Lisyanthus caerulescens Aubl. =Tetrapollinia caerulescens
Lisyanthus purpurascens Aubl. =Chelonanthus purpurascens
Lisyanthus trifidus (Kunth) Griseb. =Chelonanthus alatus
Lisyanthus uliginosus Griseb. =Chelonanthus purpurascens
Macrocarpaea arborea (Britton) Ewan =Rogersonanthus arboreus
Macrocarpaea cerronis Ewan =Rogersonanthus arboreus
Macrocarpaea quelchii (N.E. Br.) Ewan =Rogersonanthus quelchii
Macrocarpaea salicifolia Ewan =Rogersonanthus arboreus
Macrocarpaea tepuiensis (Gleason) Steyerm. =Rogersonanthus arboreus
Octopleura loeselioides Spruce ex Progel =Neurotheca loeselioides
Pagaea nemorosa (Willd. ex Roem. & Schult.) Gilg =Irlbachia nemorosa
Pagaea poeppigii Griseb. =Irlbachia poeppigii
Pagaea pumila (Benth.) Gilg =Irlbachia pumila
Pagaea ramosissima (Benth.) Ewan =Irlbachia poeppigii
Pagaea recurva (Benth.) Benth. & Hook. f. =Irlbachia nemorosa
Pagaea subcordata (Benth.) Benth. & Hook. f. =Irlbachia nemorosa
Rogersonanthus tepuiensis (Gleason) Maguire & B.M. Boom =Rogersonanthus arboreus
Schuebleria conferta Mart. =Curtia conferta
Schuebleria obtusifolia Benth. =Curtia obtusifolia
Schuebleria tenella Mart. =Curtia tenuifolia
Schuebleria tenuifolia (Aubl.) G. Don =Curtia tenuifolia
Schultesia brachyptera f. heterophylla (Miq.) Jonker =Schultesia brachyptera
Schultesia heterophylla Miq. =Schultesia brachyptera
Schultesia schomburgkiana Progel ^Schultesia benthamiana
Stahelia spenneroides (Benth.) Jonker =Tapeinostemon spenneroides
Stahelia surinamensis Jonker =Tapeinostemon spenneroides
Symbolanthus aracamuniensis Steyerm. =Symbolanthus argyreus
Symbolanthus calygonus (Ruiz & Pav.) Griseb. [s.l.] =Symbolanthus elisabethae
Symbolanthus elisabethae subsp. occidentalis Maguire & B.M. Boom =Symbolanthus huachamacariensis
Symbolanthus quelchii (N.E. Br.) Gleason =Rogersonanthus quelchii
Symbolanthus yutajensis Maguire & B.M. Boom =Symbolanthus yaviensis
Tapeinostemon borrerioides Benth. ex Knobl. [nomen nudum] =Tapeinostemon sessiliflorum
Tapeinostemon capitatum Benth. =Tapeinostemon sessiliflorum
Tapeinostemon ptariense Steyerm. =Tapeinostemon spenneroides
Tapeinostemon surinamensis (Jonker) Steyerm. =Tapeinostemon spenneroides
Voyria araguensis H. Karst. =Voyria aphylla
Voyria parviflora Miq. =Voyriella parviflora
Voyriella oxycarpha Sandwith = Voyriella parviflora
Wurdackanthus argyreus Maguire =Symbolanthus argyreus
References
Grothe, E., and P.J.M. Maas. 1984. A scanning electron microscopy study of the seed coat structure of Curtia Chamisso and
Schlechtendahl and Hockinia Gardner (Gentianaceae). Proceedings Koninklijke Nederlandse Akademie van Wetenschappen.
Series C. 87: 33^2.
Jonker, F.R 1936. Gentianaceae. In A. Pulle, Flora of Suriname 4(1): 400-427.
Kuntze, C.E.O. 1891. Revisio Generum Plantarum par. 2. Leipzig.
Leeuwenberg, A.J.M. & PW. Leenhouts. 1980. Taxonomy. Pages 8-96 in: Engler and Prantl's Die Naturlichen Pflanzenfamilien,
Angiospermae: Ordnung Gentianales, Fam. Loganiaceae. Vol. 28b (1). Leeuwenberg, A. J. M., ed. Duncker and Humblot,
Berlin
Checklist of the Plants of the Guiana Shield 309
Maas-van de Kamer, H. & P. Maas. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 328-334.
Maas, P.J.M. 1985. Nomenclatural notes on neotropical Lisyantheae. Proc. Kon. Ned. Akad. Wetensch. (ser. c) 88(4): 405-412.
Maas, P.J.M. , S. Nilsson, A.M.C. Hollants, B.J.H. ter Welle, H. Persoon, & E.C.H. van Heusden. 1984. Systematic studies in
neotropical Gentianaceae - the Lisianthus complex. Acta Bot. Neerl. 32: 371-374.
Maas, P.J.M. & P. Ruyters. 1986. Voyria and Voyriella (Saprophytic Gentianaceae). Flora Neotropica Monogr. 41: 1-93.
Machado, I.C.S., I. Sazima, and M. Sazima. 1998. Bat pollination of the terrestrial herb Irlbachia alata (Gentianaceae) in
northeastern Brazil. Plant Systematics and Evolution 209: 231-237.
Maguire, B. 1981. Gentianaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 1 1 . Mem. N.Y. Bot. Gard.
32: 1-391.
Maguire, B. & B.M. Boom. 1989. Gentianaceae (part 3). In B. Maguire & collaborators. Botany of the Guayana Highland part
13. Memoirs ofthe New York Bot. Garden 51: 1-127.
Pringle, J.S. 1995. Gentianaceae. Pages 1-131 in: Flora of Ecuador. Vol. 53. G. Harling and L. Andersson, eds. Department of
Systematic Botany, Goteborg University, Goteborg.
Struwe, L., and V.A. Albert. 1998. Lisianthius (Gentianaceae), its probable homonym Lisyanthus, and the priority of Helia over
Irlbachia as its substitute. Harvard Papers in Botany 3: 63-71.
Struwe, L., V.A. Albert, and B. Bremer. 1994 (1995). Cladistics and family level classification ofthe Gentianales. Cladistics 10:
175-206.
Struwe, L., P.J.M. Maas, and V.A. Albert. 1997. Aripuana cullmaniorum, a new genus and species of Gentianaceae from white-
sands of southeastern Amazonas, Brazil. Harvard Papers of Botany 2: 235-253.
Struwe, L., P.J.M. Maas, O. Pihlar, & V.A. Albert. 1999. Gentianaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K.
Hoist, Flora ofthe Venezuelan Guayana 5: 474-542.
Struwe, L., M. Thiv, J. W. Kadereit, A. S-R. Pepper, T.J. Motley, RJ. White, J.H.E. Rova, K. Potgieter, & V.A. Albert. 1998.
Saccifolium (Saccifoliaceae), an endemic of Sierra de la Neblina on the Brazilian-Venezuelan frontier, is related to a temperate-
alpine lineage of Gentianaceae. Harvard Papers in Botany 3: 199-214.
Gesneriaceae
C. Feuillet, L.E. Skog, J.A. Steyermarkf
Alloplectus cristatus (L.) Mart.
Alloplectus cristatus var. epirotes Leeuwenb.
Alloplectus savannarum C.V Morton
Besleriaflavovirens Nees & Mart.
Besleria gibbosa (Poepp.) Hanst.
Besleria insolita C.V. Morton
Besleria laxiflora Benth.
Besleria parviflora L.E. Skog & Steyerm.
Besleria patrisii DC.
Besleria penduliflora Fritsch
Besleria saxicola C.V. Morton
Besleria sp. A AM
Besleria sp. B AM
Besleria sp. C AM
Chrysothemis dichroa Leeuwenb. AM
Chrysothemis pulchella (Donn ex Sims) Decne.
Chrysothemis rupestris (Benth.) Leeuwenb.
Chysothemis villosa (Benth.) Leeuwenb.
Codonanthe calcarata (Miq.) Hanst.
Codonanthe crassifolia (H. Focke) C.V. Morton
Codonanthopsis dissimulata (H.E. Moore) Wiehler
Codonanthopsis ulei Mansf.
Columnea calotricha Donn. Sm.
Columnea guianensis C.V. Morton
Columnea microsepala (C.V. Morton) Kvist & L.E. Skog
Columnea oerstediana Klotzsch ex Oerst.
Columnea sanguinea (Pers.) Hanst.
Corytoplectus congestus (Linden ex Hanst.) Wiehler
Corytoplectus deltoideus (C.V. Morton) Wiehler
Cremersia platula Feuillet & L.E. Skog
Diastema racemiferum Benth.
Drymonia antherocycla Leeuwenb.
Drymonia coccinea (Aubl.) Wiehler
Drymonia psilocalyx Leeuwenb. SU FG
GU
BO GU SU
BO GU SU FG
BO
GU FG
BO GU SU FG
BO GU
GU SU FG
BO GU
GU
BO DA GU
GU
GU
SU
SU
FG
AM BO GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM
GU
SU FG
AM
FG
AM BO GU? SU FG
AM
BO GU
FG
AM
SU FG
AM BO GU SU FG
310 Checklist of the Plants of the Guiana Shield
Drymonia serrulata (Jacq.) Mart. AM
Episciafimbriata Fritsch AM
Episcia reptans Mart. AM
Episcia sp. A AM
Episcia sp. B AM
Episcia sphalera Leeuwenb.
Episcia xantha Leeuwenb. AM
Gloxinia perennis (L.) Fritsch [cultivated or escaped from cultivation]
Gloxinia purpurascens (Rusby) Wiehler
Koellikeria erinoides (DC.) Mansf.
Kohleria hirsuta (Kunth) Regel AM
Lampadaria rupestris Feuillet & L.E. Skog
Lembocarpus amoenus Leeuwenb.
Napeanthus angustifolius Feuillet & L.E. Skog
Napeanthm Jelskii Fritsch FG
Napeanthus macrostoma Leeuwenb.
Napeanthus rupicola Feuillet & L.E. Skog
Nautilocalyx adenosiphon (Leeuwenb.) Wiehler
Nautilocalyx arenarius L.E. Skog & Steyerm. AM
Nautilocalyx bryogeton (Leeuwenb.) Wiehler
Nautilocalyx cataractarum Wiehler
Nautilocalyx chimantensis L.E. Skog & Steyerm.
Nautilocalyx coccineus Feuillet & L.E. Skog
Nautilocalyx cordatus (Gleason) L.E. Skog
Nautilocalyxfasciculatus L.E. Skog & Steyerm.
Nautilocalyx kohlerioides (Leeuwenb.) Wiehler
Nautilocalyx maguirei L.E. Skog & Steyerm.
Nautilocalyx mimuloides (Benth.) C.V. Morton
Nautilocalyx pallidus (Sprague) Sprague
Nautilocalyx pemphidius L.E. Skog
Nautilocalyx pictus (Hook.) Sprague
Nautilocalyx porphyrotrichus (Leeuwenb.) Wiehler
Nautilocalyx punctatus Wiehler
Nautilocalyx resioides (Leeuwenb.) Wiehler
Nautilocalyx sp. A
Nautilocalyx sp. B
Nautilocalyx sp. C
Nautilocalyx sp. D
Nautilocalyx sp. E
Nautilocalyx sp. F
Nautilocalyx sp. G
Nautilocalyx villosus (Kunth & Bouche) Sprague
Paradrymonia anisophylla Feuillet & L.E. Skog
Paradrymonia barbata Feuillet & L.E. Skog
Paradrymonia campostyla (Leeuwenb.) Wiehler
Paradrymonia ciliosa (Mart.) Wiehler
Paradrymonia densa (C.H. Wright) Wiehler
Paradrymonia longifolia (Poepp.) Wiehler
Paradrymonia maculata (Hook, f.) Wiehler
Paradrymonia metamorphophylla (J.D. Sm.) Wiehler
Paradrymonia sp. A
Paradrymonia sp. B
Paradrymonia sp. C
Paradrymonia sp. D
Paradrymonia sp. E
Rhoogeton cyclophyllus Leeuwenb.
Rhoogeton viviparus Leeuwenb.
Sinningia elatior (Kunth) Chautems
Sinningia incarnata (Aubl.) D.L. Denham
Sinningia schomburgkiana (Kunth & Bouche) Chautems
Tylopsacas cuneatum (Gleason) Leeuwenb.
Synonyms and Misidentifications
Achimenes erinoides DC. =Koellikeria erinoides
Alloplectus calotrichus (J.D. Sm.) Steam =Columnea calotricha
BO DA GU SU FG
GU su FG
SU FG
GU su FG
GU su FG
GU FG
BO
BO GU
GU
su FG
su FG
FG
su FG
GU
BO FG
BO
GU
BO
BO
GU
AM BO GU
AM
BO
GU
GU
GU
su
FG
FG
AM GU FG
AM
GU su FG
BO GU
AM
BO
BO
GU
AM
AM
BO
AM
AM
GU SU? FG
GU
GU
SU FG
GU SU FG
GU SU FG
FG
GU FG
AM
AM
AM
AM
AM
AM
GU
AM BO GU
AM BO
BO GU
GU
AM BO GU
SU FG
Checklist of the Plants of the Guiana Shield 311
AUoplectus circinatus Mart. =Diymonia coccinea
Alloplectus coccineus (Aubl.) Mart. =Drymonia coccinea
AUoplectus congestus Linden ex Hanst. =Corytoplectus congestus
Alloplectus deltoideus C.V. Morton =Corytoplectus deltoideus
AUoplectus meridensis Klotzsch ex Hanst. [not in flora area]
AUoplectus metamoi-phophyllus J.D. Sm. =Paradrymoma metamorphophylla
Alloplectus microsepalus C.V. Morton =Columnea microsepala
Alloplectus pallidus Sprague =Nautilocalyx pallidus
Alloplectus patrisii DC. =D}ymonia coccinea
Besleria bivalvis L. f. (Mendonciaceae) =Mendoncia bivalvis
Besleria bivalvis L. f. (Mendonciaceae) =Mendoncia bivalvis
Besleria coccinea Auh\. =Diymonia coccinea
Besleria incarnata Aubl. =Sinningia incarnata
Besleria lutea L. [not in flora area]
Besleria maasii Wiehler =Besleria patrisii
Besleria pulchella Donn ex Sims =Chrysothemis pulchella
Besleria sanguinea Pers. =Columnea sanguinea
Besleria serrulata Jacq. =Drymonia serrulata
Besleria spectabilis Kunth =Drymonia serrulata
Besleria verecunda C.V. Morton =Besleria patrisii
Besleria violacea Aubl. (Bignoniaceae) =Schlegelia violacea
Brachyloma rhodomallos (Hanst.) Hanst. =Kohleria hirsuta
Centrosolenia densa (C.H. Wright) Sprague =Paradrymonia densa
Centrosolenia hirsuta Benth. =Nautilocalyx cordatus
Codonanthe bipartita L.B. Sm. =Codonanthe calcarata
Codonanthe carnosa Hanst. [not in flora area]
Codonanthe confusa Sandwith =Codonanthe crassifolia
Codonanthe dissimulata H.E. Moore =Codonanthopsis dissimulata
Codonanthe ulei (Mansf.) H.E. Moore =Codonanthopsis ulei
Codonanthopsis ulei Fritsch [not in flora area]
Columnea affinis C.V. Morton =Columnea sanguinea
Columnea aureonitens Hook. =Columnea sanguinea
Columnea calotricha var. austroamericana C.V. Morton =Alloplectus savannarum
Columnea coccinea (Aubl.) Kuntze =Drymonia coccinea
Columnea scandens L. [not in flora area]
Columnea steyermarkii C.V. Morton =Alloplectus savannarum
Crantzia patrisii (DC.) Fritsch =Dtymonia coccinea
Dalbergaria aureonitens (Hook.) Wiehler =Columnea sanguinea
Dalbergaria sanguinea (Pers.) Steud. =Columnea sanguinea
Drymonia calcarata Mart. —Drymonia serrulata
Drymonia campostyla Leeuwenb. =Paradrymonia campostyla
Diymonia cristata Miq. =Drymonia serrulata
Drymonia sarmentosula Lem. =Drymonia serrulata
Drymonia turrialvae Hanst. [not in flora area]
Episcia adenosiphon Leeuwenb. =Nautilocalyx adenosiphon
Episcia bryogeton Leeuwenb. =Nautilocalyx bryogeton
Episcia ciliosa (Mart.) Hanst. =Paradrymonia ciliosa
Episcia cordata Gleason =Nautilocalyx cordatus
Episcia cuneata Gleason =Tylopsacas cuneatum
Episcia densa C.H. Wright =Paradrymonia densa
Episcia hirsuta (Benth.) Hanst. =Nautilocalyx cordatus
Episcia maculata Hook. f. =Paradtymonia maculata
Episcia mimuhides Benth. =Nautilocalyx mimuloides
Episcia porphyrotricha Leeuwenb. =Nautilocalyx porphyrotrichus
Episcia resioides Leeuwenb. =Nautilocalyx resioides
Gesneria caracasana Otto & Dietr. =Sinningia incarnata
Gesneria elatior Kunth =Sinningia elatior
Gesneria gollmeriana Hanst. =Sinningia incarnata
Gesneria hirsuta Kunth =Kohleria hirsuta
Gesneria lindeniana Brongn. =Sinningia incarnata
Gesneria rubricaulis Kunth & Bouche =Kohleria hirsuta
Gesneria schomburgkiana Kunth & Bouche =Sinningia schomburgkiana
Gesneria vargasii DC. =Sinningia incarnata
Gloxinia trichantha Miq. =Gloxinia perennis
312 Checklist of the Plants of the Guiana Shield
Hypocyrta crassifolia H. Focke =Codonanthe crassifolia
Hypocyrta gibbosa Poepp. =Besleria gibbosa
Isoloma pycnosuzygium Donn. Sm. =Kohleria hirsuta
Kohleria longipedunciilata Hanst. =Kohleria hirsuta
Kohleria pycnosuzygium (Donn. Sm.) V.M. Badillo =Kohleria hirsuta
Kohleria rhodomallos (Hanst.) Hanst. =Kohleria hirsuta
Kohleria rubricaulis (Kunth & Bouche) Hassk. =Kohleria hirsuta
Nautilocalyx lacteus Sandwith =Nautilocalyx pictus
Nematanthus calcaratus Miq. =Codonanthe calcarata
Pentadenia microsepala (C.V. Morton) Wiehler =Columnea microsepala
Rechsteineria caracasana (Otto & Dietr.) Kuntze =Sinningia incarnata
Rechsteineria gollmeriana (Hanst.) Kuntze =Sinningia incarnata
Rechsteineria incarnata (Aubl.) Leeuwenb. =Sinningia incarnata
Rechsteineria schomburgkiana (Kunth & Bouche) Leeuwenb. =Sinningia schomburgkiana
Rhoogeton leeuwenbergianus C.V. Morton =Rhoogeton viviparus
Tussacia pulchella (Donn ex Sims) Rchb. ex Walp. =Chrysothemis pulchella
Tussacia rupestris Benth. =Chrysothemis rupestris
Tylosperma cuneatum (Gleason) Leeuwenb. =Tylopsacas cuneatum
References
Feuillet, C. & L.E. Skog. 2002. Gesneriaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, &
J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot.
Gard. 76 (2): 334-344.
Feuillet, C. & L.E. Skog. 2002 (2003). Novae Gesneriaceae Neotropicarum XL New genera and species from the Guianas.
Brittonia 54 (4): 344-351.
Feuillet, C. & L.E. Skog. 2002 (2003). Novae Gesneriaceae Neotropicarum XII. New species of Gesneriaceae from the Guianas.
Brittonia 54 (4): 352-361.
Feuillet, C. & J.A. Steyermark. 1999. Gesneriaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 5: 542-573.
Leeuwenberg, A.J.M. 1958. The Gesneriaceae of Guiana. Acta Bot. Neerl. 7: 291-444.
Leeuwenberg, A.J.M. 1984. Gesneriaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 592-650.
Morton, C.V 1939. A revision of Besleria. Contr. U.S. Nat. Herb. 26: 395-474.
Wiehler, H.J. 1978. The genera Episcia, Alsobia, Nautilocalyx, and Paradrymonia. Selbyana 5: 11-60.
Haloragaceae
L.J. Raechal
Laurembergia tetrandra (Schott ex Spreng.) Kanitz BO
Laurembergia tetrandra subsp. tetrandra [expected in the Guianas]
Myriophyllum aquaticum (Veil.) Verde. GU? FG?
Synonyms and Misidentifications
Haloragis tetrandra Schott ex Spreng. =Laurembergia tetrandra
Myriophyllum proserpinacoides Gillies ex Hook. & Am. =Myriophyllum aquaticum
References
Orchard, A.E. 1 98 1 . A revision of South American Myriophyllum, and its repercussions on some Australian and North American
species. Brunonia4: 27-65.
Raechal, L.J. 1999. Haloragaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan Guayana
5:581-582.
Hernandiaceae
RE. Berry, J.S. Miller
Hernandia guianensis Aubl.
Sparattanthelium aruakorum Tutin
Sparattanthelium guianense Sandwith
Sparattanthelium tupiniquinorum Mart. AM
Sparattanthelium uncigerum (Meisn.) Kubitzki
Sparattanthelium wonotoboense Kosterm.
Synonyms and Misidentifications
Hernandia sonora Kosterm. ^Hernandia guianensis
Hernandia sonora L. [not in flora area - Asian]
BO DA GU SU
GU SU
GU
BO DA GU SU
BO GU SU
GU SU
FG
FG?
FG
Checklist of the Plants of the Guiana Shield 313
Sparattanthelium botociidorum Mart, [not in flora area]
Sparattanthelium botociidorum var. uncigerum Meisn. =Sparattanthelium uncigerum
Sparattanthelium macusiorum A.C. Sm. =Sparattanthelium wonotoboeme
Sparattanthelium melinonis Baill. ex Lemee [nomen nudum] =Sparattanthelium uncigerum
References
Kostermans, A.J.G.H. 1937. Hemandiaceae. In A. Pulle, Flora of Suriname 2(1): 338-344.
Kubitzki, K.U. 1969. Monographic der Hemandiaceen. Bot. Jahrb. Syst. 89: 78-205.
Kubitzki, K.U. 1976. Hemandiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 485-
486.
Kubitzki, K.U. 1982. Hemandiaceae. In Z. Luces de Febres, Flore de Venezuela 4(2): 317-324.
Miller, J.S. & RE. Berry. 1999. Hemandiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 5: 592-593.
Mori, S.A. & J.L. Brown. 2002. Hemandiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, &
J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot.
Gard. 76 (2): 344-347.
HiPPOCRATEACEAE
A.R.A. Gorts-van Rijn, J.R Hedin, A.M.W. Mennegaj
Anthodon decussatum Ruiz & Pav.
Cheiloclinium anomalum Miers
Cheiloclinium belizense (Standi.) A.C. Sm.
Cheiloclinium cognatum (Miers) A.C. Sm.
Cheiloclinium diffusiflorum (Miers) A.C. Sm.
Cheiloclinium habropodum A.C. Sm.
Cheiloclinium hippocrateoides (Peyr.) A.C. Sm.
Cheiloclinium klugii A.C. Sm.
Cheiloclinium obtusum A.C. Sm.
Cuervea kappleriana (Miq.) A.C. Sm.
Elachypterafloribunda (Benth.) A.C. Sm.
Hippocratea volubilis L.
Hylenaea comosa (Sw.) Miers
Hylenaeapraecelsa (Miers) A.C. Sm. [expected in the Guianas]
Hylenaea unguiculata Mennega
Peritassa compta Miers
Peritassa dulcis (Benth.) Miers
Peritassa huanucana (Loes.) A.C. Sm.
Peritassa laevigata (Hoffmanns, ex Link) A.C. Sm.
Peritassa peruviana (Miers) A.C. Sm.
Peritassa pruinosa (Seem.) A.C. Sm.
Prionostemma aspera (Lam.) Miers
Pristimera caudata Mennega
Pristimera holdeniana (A.C. Sm.) A.C. Sm.
Pristimera nervosa (Miers) A.C. Sm.
Pristimera tenuiflora (Mart, ex Peyr.) A.C. Sm.
Pristimera verrucosa (Kunth) Miers
Salacia amplectens A.C. Sm.
Salacia cordata (Miers) Mennega
Salacia duckei A.C. Sm.
Salacia elliptica (Mart, ex Schult.) G Don
Salacia gigantea Loes.
Salacia impressifolia (Miers) A.C. Sm.
Salacia insignis A.C. Sm.
Salaciajuruana Loes.
Salacia kanukuensis A.C. Sm.
Salacia maburensis Mennega
Salacia macrantha A.C. Sm.
Salacia miqueliana Loes.
Salacia multiflora (Lam.) DC.
Salacia multiflora subsp. mucronafa (Rusby) Mennega
Salacia multiflora subsp. multiflora
Salacia opacifolia (J.F. Macbr.) A.C. Sm.
Salacia pachyphylla (Miers) Peyr. GU
AM BO FG
AM BO DA GU su
AM BO GU su FG
BO DA GU su FG
AM GU su FG
AM
AM BO GU su FG
AM BO
BO
AM BO DA GU su FG
AM DA GU su FG
AM BO DA GU su FG
AM BO DA GU su
su
FG
AM BO GU
GU
su
AM BO GU su FG
AM BO DA GU su FG?
AM BO
AM BO GU
AM BO DA GU
GU
su
su
FG
AM BO DA GU su FG
BO DA su FG
BO GU
AM BO GU su FG
AM BO GU su
su? FG
AM BO GU su FG
AM
AM BO GU su
su
FG
FG
AM BO GU su FG
AM BO
BO
GU
GU
su
su
FG?
FG
AM BO DA GU su FG
BO DA GU su FG
AM BO GU su FG
AM
314 Checklist of the Plants of the Guiana Shield 4
Tontelea attenuata Miers
Tontelea cohacea A.C. Sm.
Tontelea corymbosa (Huber) A.C. Sm.
Tontelea cylindrocarpa (A.C. Sm.) A.C. Sm.
Tontelea glabra A.C. Sm.
Tontelea laxiflora (Benth.) A.C. Sm.
Tontelea mauritioides (A.C. Sm.) A.C. Sm.
Tontelea nectandrifolia (A.C. Sm.) A.C. Sm.
Tontelea ovalifolia (Miers) A.C. Sm.
Tontelea ovalifolia subsp. ovalifolia
Tontelea ovalifolia subsp. richardii (Peyr.) Gorts & Mennega
Tontelea sandwithii A.C. Sm.
Synonyms and Misidentifications
Amphizoma laxiflora (Benth.) Miers ='Tontelea laxiflora
Anthodon grandiflorus Benth. =Salacia impressifolia
Anthodon laxiflorus Benth. =Tontelea laxiflora
Anthodus ellipticus Mart, ex Schult. =Salacia elliptica
Cheiloclinium divaricata A.C. Sm. =Cheiloclinium hippocrateoides
Cheiloclinium glaziovii A.C. Sm. =Cheilocliniiim anomalum
Cheiloclinium gleasoniamim (A.C. Sm.) A.C. Sm. =Cheiloclinium hippocrateoides
Cheiloclinium jenmanii A.C Sm. =Cheiloclinium belizense
Cheiloclinium krukovii (A.C. Sm.) A.C. Sm. =Cheiloclinium hippocrateoides
Cheiloclinium lineolatum (A.C. Sm.) A.C. Sm. =Cheiloclinium cognatum
Cheiloclinium lucidum A.C. Sm. =Cheiloclinium diffusiflorum
Cheiloclinium meianthemum A.C. Sm. =Cheiloclinium belizense
Cheiloclinium minutiflorum (A.C. Sm.) A.C. Sm. [not in flora area]
Cheiloclinium parviflorum (Miers) A.C. Sm. =Cheiloclinium diffusiflorum
Cheiloclinium podostemmum (Sandwith) A.C. Sm. =Cheiloclinium belizense
Cuervea granatensis Miers =Cuervea kappleriana
Cuervea mitchellae (I.M. Johnst.) A.C. Sm. =Cuervea kappleriana
Cuervea ovalifolia Miers =Tontelea ovalifolia
Hippocratea aggregata Peyr. =Cuervea kappleriana
Hippocratea ambigua Peyr. [insufficiently known taxon]
Hippocratea aspera Lam. =Prionostemma aspera
Hippocratea comosa Sw. =Hylenaea comosa
Hippocratea crinita Pittier =Hylenaea comosa
Hippocratea emarginata Rudge ^Salacia multiflora subsp. multiflora
Hippocrateaflava Gleason =Cuervea kappleriana
Hippocrateafloribunda Benth. =Elachypterafloribunda
Hippocratea granatensis Peyr. =Cuervea kappleriana
Hippocratea holdeniana A.C. Sm. =Pristimera holdeniana
Hippocratea huanucana Loes. =Peritassa huanucana
Hippocratea inundata Peyr. =Cuervea kappleriana
Hippocratea kappleriana Miq. =Cuervea kappleriana
Hippocratea laevigata Rich. =Hippocratea volubilis
Hippocratea lancifolia Lundell =Elachyptera floribunda
Hippocratea lutea Gleason =Cuervea kappleriana
Hippocratea malpighiifolia Rudge =Prionostemma aspera
Hippocratea micrantha Cambess. =Elachypterafloribunda
Hippocratea mitchellae I.M. Johnst. =Cuervea kappleriana
Hippocratea multiflora Lam. =Salacia multiflora
Hippocratea obovata Rich. =Salacia multiflora subsp. multiflora
Hippocratea obovata var. guianense Peyr. =Salacia multiflora subsp. multiflora
Hippocratea opacifolia J.F. Macbr. =Salacia opacifolia
Hippocratea schomburgkii Klotzsch [nomen nudum] =Prionostemma aspera
Hippocratea tenuiflora Mart, ex Peyr. =Pristimera tenuiflora
Hippocratea verrucosa Kunth =Pristimera verrucosa
Hylenaea capilliflora Miers =Hylenaea comosa
Kippistia cognata Miers =Cheiloclinium cognatum
Kippistia diffusiflora Miers =Cheiloclinium diffusiflorum
Peritassa cuneata Mennega =Peritassa huanucana
Peritassa granulata (Urb.) A.C. Sm. =Peritassa laevigata
Peritassa laevigata subsp. obtusa Peyr. =Peritassa compta
AM GU FG
BO GU SU
SU?
FG
AM SU FG
GU SU FG
AM BO
BO
GU
GU
SU FG
GU SU FG?
AM BO DA GU SU FG?
AM BO DA GU
GU
SU FG?
FG
FG
Checklist of the Plants of the Guiana Shield 315
Peritassa rorida Miers =Peritassa laevigata
Peritassa rubicimda Miers =Peritassa laevigata
Peritassa smaragdina Miers =Peritassa laevigata
Prionostemma malpighiifolia (Rudge) Miers =Prionostemma aspera
Pristimera apiculata Miers =Cuervea kappleriana
Phstimera caribaea (Urb.) A.C. Sm. [not in flora area]
Raddia elliptica (Mart, ex Schult.) Miers =Salacia elliptica
Raddia grandiflora (Benth.) Miers =Salacia impressifolia
Raddia impressifolia Miers =Salacia impressifolia
Raddia lacunosa Miers =Salacia elliptica
Raddia pachyphylla Miers =Salacia pachyphylla
Salacia acreana A.C. Sm. =Salacia cordata
Salacia affinis Peyr. =Cheiloclinium cognatum
Salacia anomala (Miers) Peyr. =Cheiloclinium anomalum
Salacia anomalum (Miers) Peyr. =Cheiloclimum anomalum
Salacia attenuata (Miers) Peyr. =Tontelea attenuata
Salacia belizensis Standi. =Cheiloclinium belizense
Salacia brevistaminea Pittier =Anthodon decussatum
Salacia catalinensis Rusby =Prionostemma aspera
Salacia cognata (Miers) Peyr. =Cheiloclinium cognatum
Salacia cognata var. egensis Peyr. =Cheiloclinium cognatum
Salacia crassifolia Pittier =Salacia multiflora subsp. multiflora
Salacia cylindrocarpa A.C. Sm. =Tontelea cylindrocarpa
Salacia diffusiflora (Miers) Peyr. =Cheiloclimum diffusiflorum
Salacia dulcis Benth. =Peritassa dulcis
Salacia elliptica var. apiculata Peyr. =Salacia elliptica
Salaciafluminensis Peyr. =Tontelea ovalifolia subsp. richardii
Salacia gleasoniana A.C. Sm. =Cheiloclinium hippocrateoides
Salacia gracilis A.C. Sm. =Salacia opacifolia
Salacia grandiflora Peyr. =Salacia impressifolia
Salacia granulata Urb. =Peritassa laevigata
Salacia guianensis Klotzsch [nomen nudum] =Salacia elliptica
Salacia guianensis (Aubl.) Lemee [incertae sedis; excluded from Hippocrateaceae by A.C. Smith] =Macahanea guianensis
Aubl.
Salacia hippocrateoides Peyr. =Cheiloclinium hippocrateoides
Salacia krukovii A.C. Sm. =Cheiloclinium hippocrateoides
Salacia lacunosa (Miers) Peyr. =Salacia elliptica
Salacia laevigata (Hoffmanns, ex Link) DC. =Peritassa laevigata
Salacia laevigata subsp. obtusa Peyr. =Peritassa compta
Salacia laxiflora (Benth.) Peyr. =Tontelea laxiflora
Salacia laxiflora var. amygdalifolia Peyr. =Tontelea laxiflora
Salacia laxiflora var. oblongifolia Peyr. =Tontelea laxiflora
Salacia lineolata A.C. Sm. -Cheiloclinium cognatum
Salacia macrophylla Miq. =Salacia miqueliana
Salacia megistophylla Standi. =Salacia cordata
Salacia mucronata Rusby =Salacia multiflora subsp. mucronata
Salacia nectandrifolia A.C. Sm. =Tontelea nectandrifolia
Salacia obovata (Rich.) Peyr. =Salacia multiflora subsp. multiflora
Salacia obovata var. guianensis Peyr. =Salacia multiflora subsp. multiflora
Salacia parviflora Sagot ex Peyr. =Cheiloclinium diffusiflorum
Salacia patens Triana & Planch. =Tontelea ovalifolia
Salacia pittieriana A.C. Sm. =Salacia multiflora subsp. multiflora
Salacia podostemma Sandwith =Cheiloclinium belizense
Salacia pruinosa Seem. =Peritassa pruinosa
Salacia richardii Peyr. =Tontelea ovalifolia subsp. richardii
Salacia rugulosa Sagot [nomen nudum] =Salacia multiflora subsp. multiflora
Salacia scabra DC. =Prionostemma aspera
Salacia sphaerocarpa Rusby =Cheiloclinium cognatum
Sicyomorpha nervosa Miers =Pristimera nervosa
Sicyomorpha peruviana Miers =Peritassa peruviana
Sicyomorpha pruinosa (Seem.) Miers =Peritassa pruinosa
Thermophila cordata Miers =Salacia cordata
Thermophila emarginata (Rudge) Miers =Salacia multiflora subsp. multiflora
Thermophila macrophylla (Miq.) Miers =Salacia miqueliana
316 Checklist of the Plants of the Guiana Shield
Thermophila obovata (Rich.) Miers =Salacia multiflora subsp. multiflora
Thermophila nigulosa Miers =Salacia multiflora subsp. multiflora
Tonsella laevigata Hoffmanns, ex Linic =Peritassa laevigata
Tonsella malpighiifolia (Rudge) G. Mey. =Prionostemma aspera
Tontelea aubletiana Miers =Tontelea sp.
Tonteleafloribunda (Benth.) Miers =Elachypterafloribunda
Tonteleafluminensis (Peyr.) A.C. Sm. =Tontelea ovalifolia subsp. richardii
Tontelea longifolia Miers =Tontelea attenuata
Tontelea parviflora Miers =Cheiloclimum diffusiflorum
Tontelea richardii (Peyr.) A.C. Sm. =Tontelea ovalifolia subsp. richardii
Tontelea scandens Aubl. [nomen nudum]
References
Gorts-van Rijn, A. 2002. Hippocrateaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 347-353.
Gorts-van Rijn, A.R.A. & A.M.W. Mennega. 1994. Hippocrateaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A)
16: 1-157.
Loesener, T. 1942. Hippocrateaceae. In A. Engler & K. Prantl, Die Natiirlichen Pflanzenfamilien 20b: 198-231.
Mennega, A.M.W. & J. P. Hedin. 1999. Hippocrateaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych &.B.K. Hoist, Flora of
the Venezuelan Guayana 5: 594-617.
Smith, A.C. 1940. The American species of Hippocrateaceae. Brittonia 3: 341-355.
HUGONIACEAE
P.E. Berry, A. Jardim, N. Ramirez, D. Sabatier
AM BO DA GU su
AM GU
AM
AM GU su
AM GU
AM BO su
BO GU su
FG?
FG
Hebepetalum humiriifolium (Planch.) Benth. SU FG
Hebepetahim roraimense Secco & S.M.B. Silva
Hebepetalum sp. A
Roucheria calophylla Planch.
Roucheria columbiana Hallier f.
Roucheria laxiflora H. Winkl.
Roucheria schomburgkii Planch.
Roucheria sp. A AM
Synonyms and Misidentifications
Hebepetalum parviflorum Ducke =Roucheria calophylla
Hebepetalum punctatum Ducke =Roucheria columbiana
Hebepetalum schomburgkii (Planch.) Ducke =Roucheria schomburgkii
Roucheria angulata Gleason =Hebepetalum humiriifolium
Roucheria humeriifolia Planch. =Hebepetalum humiriifolium
Roucheria humiriifolia Planch. =Hebepetalum humiriifolium
Roucheria parviflora (Ducke) Ducke =Roucheria calophylla
Roucheria punctata (Ducke) Ducke =Roucheria columbiana
References
Bakhuizen van den Brink f, R.C. 1941. Linaceae. In A. Pulle, Flora of Suriname 3(1): 409-411.
Ramirez, N., P.E. Berry, & A. Jardim. 1999. Hugoniaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of
the Venezuelan Guayana 5: 618-623.
Sabatier, D. 2002. Hugoniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
353-354.
Secco, R. de S. & S.M.B. Silva. 1990. Materiaes para a flora amazonica - VIII. Contribu9ao a sistematica das Linaceae da
Amazonia Brasileira. Bol. Mus. Paraense Emilio Goeldi (Bot.) 6(1): 113-135.
HUMIRIACEAE
J. Cuatrecasast, O. Huber, D. Sabatier
Endopleura uchi (Huber) Cuatrec.
Humiria balsamifera Aubl.
Humiria balsamifera var. balsamifera
Humiria balsamifera var. coriacea Cuatrec.
Humiria balsamifera var. floribunda (Mart.) Cuatrec.
Humiria balsamifera var. guaiquinimana Cuatrec.
AM GU FG
AM BO GU SU FG
AM BO GU su FG
AM BO GU SU
AM BO
BO
GU su FG
BO
BO GU
AM BO
BO
GU
AM BO
BO
GU
AM BO
AM BO GU
AM
AM
AM
AM BO GU
Checklist of the Plants of the Guiana Shield 317
Hitmiha balsamifera var. giiianensis (Benth.) Cuatrec. AM GU SU
Humiria balsamifera var. iluana Cuatrec.
Hiimiha balsamifera var. imbaimadaiensis Cuatrec.
Humiria balsamifera var. laurina (Urb.) Cuatrec.
Humiria balsamifera vox. pilosa (Steyerm.) Cuatrec.
Humiria balsamifera var. savannarum (Gleason) Cuatrec.
Humiria balsamifera var. stenocarpa Cuatrec.
Humiria balsamifera var. subsessilis (Urb.) Cuatrec.
Humiria crassifolia Mart, ex Urb.
Humiriafruticosa Cuatrec.
Humiria wurdackii Cuatrec.
Humiriastrum colombianum Cuatrec.
Humiriastrum cuspidatum (Benth.) Cuatrec.
Humiriastrum excelsum (Ducke) Cuatrec. FG
Humiriastrum liesneri Cuatrec. AM
Humiriastrum obovatum (Benth.) Cuatrec. BO GU
Humiriastrum ottohuberi Cuatrec.
Humiriastrum piraparanense Cuatrec.
Humiriastrum subcrenatum (Benth.) Cuatrec.
Sacoglottis amazonica Mart.
Sacoglottis ceratocarpa Ducke
Sacoglottis cydonioides Cuatrec.
Sacoglottis guianensis Benth.
Sacoglottis guianensis var. guianensis
Sacoglottis maguirei Cuatrec.
Sacoglottis mattogrossensis Malme [Guianas specimens represent a new
species, fide Sabatier]
Schistostemon auyantepuiensis Cuatrec.
Schistostemon densiflorum (Benth.) Cuatrec.
Schistostemon dichotomum (Urb.) Cuatrec.
Schistostemon fernandezii Cuatrec.
Schistostemon oblongifolium (Benth.) Cuatrec.
Schistostemon retusum (Ducke) Cuatrec.
Schistostemon sylvaticum Sabatier
Vantanea guianensis Aubl.
Vantanea minor Benth.
Vantanea ovicarpa Sabatier
Vantanea parviflora Lam.
Vantanea parviflora var. parviflora
Synonyms and Misidentifications
Houmiri balsamifera Aub\. =Humiria balsamifera
Humiria balsamifera J. St.-Hil. =Humiria balsamifera Aubl.
Humiria cassiquiari Suess. & Bergdoh =Humiria balsamifera var. guianensis
Humiria densiflora Benth. ^Schistostemon densiflorum
Humiriafloribunda (Mart.) Urb. =Humiria balsamifera \ar.floribunda
Humiriafloribunda var. guianensis Urb. =Humiria balsamifera var. guianensis
Humiriafloribunda var. laurina Urb. =Humiria balsamifera var. laurina
Humiriafloribunda var. spathulata Gleason ^Humiria balsamifera var. laurina
Humiriafloribunda var. subsessilis Urb. =Humiria balsamifera var. subsessilis
Humiria guianense Benth. =Humiria balsamifera var. guianensis
Humiria pilosa Steyerm. =Humiria balsamifera var. pilosa
Humiria savannarum Gleason =Humiria balsamifera var. savannarum
Humiria subcrenata Urb. =Humiriastrum subcrenatum
Humirium balsamiferum Benth. =Humiria balsamifera
Humirium cuspidatum Benth. =Humiriastrum cuspidatum
Humirium floribundum Mart. =Humiria balsamifera \ar. floribunda
Humirium oblongifolium Benth. =Schistostemon oblongifolium
Humirium obovatum Benth. =Humiriastrum obovatum
Humirium surinamense Miq. =Humiria balsamifera var. guianensis
Lemnisciafloribunda Willd. =Vantanea guianensis
Lemniscia guianensis (Aubl.) J.F. Gmel. =Vantanea guianensis
Myrodendrum amplexicaule Willd. =Humiria balsamifera
Myrodendrum balsamiferum Raeusch. [comb, illeg.] =Humiria balsamifera
AM
AM
FG
DA GU FG?
AM BO
BO DA GU SU FG
AM BO GU SU FG
GU SU FG
AM
AM BO
BO
GU
GU SU
GU? SU FG
BO
AM
AM
FG
AM
BO
GU
GU
SU FG
FG
AM GU SU
SU
FG
FG
318 Checklist of the Plants of the Guiana Shield
Sacoglottis amazonica Benth. =Sacoglottis guianensis
Sacoglottis cuspidata (Benth.) Urb. =Humiriastrum cuspidatum
Sacoglottis densiflora (Benth.) Urb. =Schistostemon densiflorum
Sacoglottis dichotoma Urb. =Schistostemon dichotomum
Sacoglottis excelsa var. colombiana Cuatrec. =Humihastrum colombianum
Sacoglottis guianensis f. dolichocarpa Ducke =Sacoglottis guianensis
Sacoglottis guianensis var. hispidula Cuatrec. =Sacoglottis guianensis
Sacoglottis guianensis var. major Ducke =Sacoglottis guianensis
Sacoglottis guianensis f. sphaerocarpa Ducke =Sacoglottis mattogrossensis
Sacoglottis kaboeriensis Bakh. v.d. Brink f. =Schistostemon densiflorum
Sacoglottis oblongifolia (Benth.) Urb. =Schistostemon oblongifolium
Sacoglottis obovata (Benth.) Urb. =Humiriastrum obovatum
Sacoglottis retusa Ducke =Schistostemon retusum
Sacoglottis uchi Huber =Endopleura uchi
Vantanea cupularis Huber =Vantanea parviflora
References
Bakhuizen van den Brink f., R.C. 1941. Humiriaceae. In A. Pulle, Flora of Suriname 3(1): 412-421.
Cuatrecasas, J. 1961. A taxonomic revision of the Humiriaceae. Contrib. U.S. Nat. Herb. 35: 25-214.
Cuatrecasas, J. & O. Huber. 1999. Humiriaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 5: 623-641.
Sabatier, D. 1987. Studies on the Flora of the Guianas 12. Quelques nouveautes chez les Humiriaceae. Proc. Kon. Ned. Akad.
Wet. ser.C, 90(2): 203-210.
Sabatier, D. 2002. Humiriaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
355-357.
Hydrophyllaceae
J.S. Miller
AM BO DA GU
FG
AM BO DA GU su FG
GU su FG
Hydrolea elatior Schott
Hydrolea palustris (Aubl.) Raeusch. [naturalized?]
Hydrolea spinosa L.
Hydrolea spinosa var. spinosa
Synonyms and Misidentifications
Hydrolea minima Brand =Hydrolea elatior
Hydrolea multiflora Mart, ex Choisy =Hydrolea elatior
Nama spinosa (L.) Kuntze =Hydrolea spinosa
Sagonea palustris Aubl. -Hydrolea palustris
References
Constance, L. 1982. Hydrophyllaceae. In Z. Luces de Febres, Flora de Venezuela 8(3): 7-14.
Davenport, L.J. 1988. A monograph of Hydrolea. Rhodora 90 (862): 169-208.
Miller, J.S. 1999. Hydrophyllaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 5: 644-646.
ICACINACEAE
A.C. de Roon, R. A. Howard, R.D. de Stefano
Casimirella ampla (Miers) R.A. Howard
Dendrobangia boliviana Rusby
Discophora froesii Pires
Discophora guianensis Miers
Emmotum acuminatum (Benth.) Miers [Guianan records erroneous, but
possibly expected]
Emmotum celiac R.A. Howard
Emmotum conjunctum R.A. Howard
Emmotum fagifolium Ham.
Emmotum floribundum R.A. Howard
Emmotumfulvum R.A. Howard [Mt. Roraima]
Emmotum glabrum Benth. ex Miers
Emmotum orbiculatum (Benth.) Miers [Guianan records erroneous, but
possibly expected]
Emmotum yapacanum R.A. Howard
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
GU
GU
GU
GU
GU?
GU
SU
SU
FG
FG
FG
SU FG
Checklist of the Plants of the Guiana Shield 319
GU SU FG
GU FG
SU FG
GU
GU SU FG
GU SU FG
Leretia cordata Veil. DA
Pleurisanthes artocarpi Baill.
Pleurisanthes emarginata Tiegh. AM
Pleurisanthesflava Sandwith
Pleurisanthes parviflora (Ducke) R.A. Howard
Pleurisanthes sp. A AM
Poraqueiba guianensis Aubl.
Poraqueiba paraens is Ducke AM BO
Poraqueiba sericea Tul. AM
Synonyms and Misidentifications
Asterolepidion elatum Ducke =Dendrobangia boliviana
Clavapetalum elatum (Ducke) Ducke =Dendrobangia boliviana
Clavapetalum surinamense Pulle =Dendrobangia boliviana
Dendrobangia tenuis Ducke =Dendrobangia boliviana
Discophora panamensis Standi. =Discophora guianensis
Emmotum argenteum Gleason =Emmotum glabrum
Emmotum nudum R.A. Howard =Emmotum acuminatum
Emmotum ptarianum Steyerm. =Emmotum glabrum
Humirianthera ampla (Miers) Baehni =Casimirella ampla
Humirianthera duckei Huber =Casimirella ampla
Kummeria brasiliensis Mart. =Discophora guianensis
Leretia ampla Miers =Casimirella ampla
Leretia nitida Miers ^Leretia cordata
Leretia parviflora Ducke ^Pleurisanthes parviflora
Leretia poeppigiana (Baill.) Sleumer =Leretia cordata
Leretia vellozii Miers -Leretia cordata
Mappia ampla (Miers) Engl. =Casimirella ampla
Mappia cordata (Veil.) Engl. =Leretia cordata
Mappia nitida (Miers) Engl. ^Leretia cordata
Mappia pai-viflora (Ducke) Baehni =Pleurisanthes parviflora
Mappia poeppigiana Baill. =Leretia cordata
Pleurisanthes brasiliensis Tiegh. [not in flora area]
Pleurisanthes simpliciflora Sleumer [not in flora area]
Pogopetalum acuminatum Benth. =Emmotum acuminatum
Pogopetalum acutum Benth. =Emmotum fagifolium
Pogopetalum orbiculatum Benth. =Emmotum orbiculatum
Poraqueiba acuminata Miers =Poraqueiba sericea
Poraqueiba siirinamensis Miers =Poraqueiba guianensis
References
Howard, R.A. 1942. Studies in the Icacinaceae, III. A revision of Emmotum. J. Am. Arb. 23: 479-494.
Howard, R.A. 1942. Studies of the Icacinaceae IV. Considerations of the New World genera. Contrib. Gray Herb. 142: 3-60.
Howard, R.A. 1978. Icacinaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 10. Mem. New York Bot.
Gard. 29: 62-68.
Howard, R.A. 1992. A revision of Casimirella, including Humirianthera. Brittonia 44(2): 166-172.
Howard, R.A. & R.D. de Stefano. 1999. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 5: 646-658.
Jansen-Jacobs, M.J. 1979. Icacinaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 344-355.
Roon, A.C.de. 1994. Icacinaceae. In A. R.A. Gorts-van Rijn, Flora of the Guianas (series A) 16: 1-157.
Roon, A.C. de. & S.A. Mori. 2002. Icacinaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, &
J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot.
Gard. 76 (2): 358-362.
Sleumer, H. 1942. Icacinaceae. In A. Engler & K. Prantl, Die Naturlichen Ptlanzenfamilien 20b: 322-396.
IXONANTHACEAE
RE. Berry, N. Ramirez, D. Sabatier
Cyrillopsis micrantha (Steyerm.) P.E. Berry & N. Ramirez
Cyrillopsis paraensis Kuhlm.
Ochthocosmus attenuatus Steyerm. & Luteyn
Ochthocosmus barrae Hallier f.
Ochthocosmus berry i Steyerm.
Ochthocosmusfloribundus Gleason
Ochthocosmus longipedicellatus Steyerm. & Luteyn
AM
AM
BO
BO
BO
BO
GU
GU
GU?
GU
FG
320 Checklist of the Plants of the Guiana Shield
Ochthocosmus multiflorus Ducke AM
Ochthocosmus multiflorus van angustifolius Steyerm. & Luteyn AM
Ochthocosmus multiflorus var. canaripoensis Steyerm. & Luteyn AM
Ochthocosmus multiflorus var. multiflorus AM
Ochthocosmus parvifolius Hallier f. [incertae sedis] GU?
Ochthocosmus roraimae Benth. BO GU
Ochthocosmus roraimae var. grandifolius (Steyerm.) Steyerm. & Luteyn BO
Ochthocosmus roraimae var. roraimae BO GU
Synonyms and Misidentifications
Ochthocosmus grandifolius Steyerm. =Ochthocosmus roraimae var. grandifolius
Ochthocosmus micranthus Steyerm. =Cyrillopsis micrantha
Ochthocosmus roraimae var. parvifolius (Hallier f.) Steyerm. & Luteyn =Ochthocosmus parvifolius
References
Forman, L.L. 1965. A new genus of Ixonanthaceae with notes on the family. Kew Bull. 19: 517-526.
Kuhlmann, J.G. 1925. Cyrillaceae, in Contribui9ao para o conhecimento de algumas plantas novas. Arq. Jard. Bot. Rio de
Janeiro 4: 356-358.
Ramirez, N. & RE. Berry. 1999. Ixonanthaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 5: 664-671.
Steyermark, J.A. & J.L. Luteyn. 1980. Revision of the genus Ochthocosmus. Brittonia 32: 128-143.
Secco, R. de S. & S.M.B. Silva. 1990. Materiaes para a flora amazonica - VIII. Contribu9ao a sistematica das Linaceae da
Amazonia Brasileira. Bol. Mus. Paraense Emilio Goeldi (Bot.) 6(1): 113-135.
JUGLANDACEAE
Juglans sp. [probably based on misidentifications] FG
Synonyms and Misidentifications
Juglans brasiliensis Dode [not in flora area] =Juglans australis Griseb.
References
Manning, W.E. 1960. The genus Juglans in South America and the West Indies. Brittonia 12: 1-26.
Krameriaceae
L. Brako, B.B. Simpson
Krameria ixine Loefl.
Krameria spartioides Klotzsch ex O. Berg
Krameria tomentosa A. St.-Hil.
Synonyms and Misidentifications
Krameria arida O. Berg =Krameria ixine
Krameria evolvuloides Triana & Planch, ex Bennett =Krameria spartioides
Krameria ixina L. =Krameria ixine
Krameria lanceolata O. Berg [nom. illeg.] =Krameria ixine
Krameria linifolia Willd. ex Schult. & Schult. f. =Krameria ixine
References
Brako, L. 1999. Krameriaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
5: 674-676.
Simpson, B.B. 1989. Krameriaceae. Flora Neotropica Monogr. 49: 1-108.
Simpson, B.B. 1998. Krameriaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 21: 61-65.
Lacistemataceae
L.J. Raechal
Lacistema aggregatum (P.J. Bergius) Rusby AM BO DA
Lacistema grandifolium W. Schnizl.
Lacistema polystachyum W. Schnizl.
Synonyms and Misidentifications
Lacistema aggregatum var. elongatum Maguire =Lacistema aggregatum
Lacistema coriaceum A. DC. ^Lacistema aggregatum
GU
GU
GU?
GU su FG
GU su FG
GU su FG
Checklist of the Plants of the Guiana Shield 321
Lacistema elongatum Schnizl. =Lacistema aggregatum
Lacistema orinocense Baehni =Lacistema aggregatum
References
Mori, S.A. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide
to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 362-363.
Raechal, L.J. 1999. Lacistemaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 5: 676-677.
Sleumer, H.O. 1980. Flacourtiaceae. Flora Neotropica Monogr. 22: 1-499.
Vaandrager, G. 1938. Lacistemaceae. In A. PuIIe, Flora of Suriname 1(1): 258-261.
Lamiaceae
R.M. Harley
Anisomeles indica (L.) Kuntze [South Asian exotic]
Eriope crassipes Benth.
Eriope crassipes subsp. crassipes
Hypenia salzmannii (Benth.) Harley
Hyptidendron arboreum (Benth.) Harley
Hyptis atrorubens Poit.
Hyptis brachiata Briq.
Hyptis brevipes Poit. [frequently confused with H. lanceolata, fide Harley]
Hyptis capitata Jacq.
Hyptis colubrimontis Epling & Jativa [=Hyptis pyriformis var. yutajeensis]
Hyptis conferta Pohl ex Benth.
Hyptis conferta var. angustata (Briq.) Pool & Harley
Hyptis dilatata Benth.
Hyptis guanchezii Harley AM
Hyptis hirsuta Kunth
Hyptis huberi Harley AM
Hyptis laciniata Benth. AM
Hyptis lacustris A. St.-Hil. ex Benth. AM
Hyptis lanceolata Poir. AM
Hyptis lantanifolia Poit. AM
Hyptis luticola Epling AM
Hyptis microphylla Pohl ex Benth.
Hyptis mutabilis (Rich.) Briq. AM
Hyptis pachycephala Epling
Hyptis parkeri Benth. AM
Hyptis pectinata (L.)?oit.
Hyptis pulegioides Pohl ex Benth.
Hyptis pyriformis Epling & Jativa AM
Hyptis pyriformis var. pyriformis AM
Hyptis pyriformis var. yutajeensis Harley AM
Hyptis recurvata Poit. AM
Hyptis spicigera Lam.
Hyptis suaveolens (L.) Poit. AM
Leonotis nepetifolia (L.) W.T. Alton
Leonurus japonicus Houtt. [naturalized]
Marsypianthes chamaedrys (Vahl) Kuntze
Ocimum americanum L.
Ocimum campechianum Mill. AM
Ocimum gratissimum L. [naturalized]
Scutellaria leucantha Loes.
Scutellaria purpurascens Sw.
Scutellaria uliginosa A. St.-Hil. ex Benth.
Synonyms and Misidentifications
Ballota suaveolens L. =Hyptis suaveolens
Clinopodium chamaedrys Vahl =Marsypianthes chamaedrys
Eriope nudiflora Kunth ex Benth. =Eriope crassipes
Hyptis americana (Aubl.) Urb. =Hyptis spicigera
Hyptis arborea Benth. =Hyptidendron arboreum
Hyptis belizensis Lundell =Hyptis conferta var. angustata
AM
AM
BO
BO
AM BO
AM BO
BO
BO
AM
AM BO
AM BO
AM BO
BO
BO
DA
GU
GU
GU
GU SU
su
GU
GU SU
FG
FG
FG
FG
FG?
BO GU
BO GU
BO DA GU SU FG
BO GU SU FG
BO GU SU
BO SU FG
FG
BO GU SU FG
BO DA GU
GU
SU? FG
BO
BO
BO GU
GU
SU FG
FG
BO DA GU FG
DA GU SU FG
DA GU SU FG
BO DA GU
GU
SU
SU
FG
BO DA GU SU
SU
FG
FG
FG
322 Checklist of the Plants of the Guiana Shield
Hyptis brevipes var. serrata Briq. =Hyptis brevipes
Hyptis canescens Kunth =Hyptis mutabilis
Hyptis conferta var. angustifolia Benth. ex Epling [nom. illeg.] =Hyptis conferta var. angustata
Hyptis constricta Briq. "^Hyptis conferta var. angustata
Hyptis crenata var. angustifolia Benth. ex Seem. =Hyptis dilatata
Hyptis excelsa M. Martens & Galeotti =Hyptis conferta var. angustata
Hyptisferruginea Benth. =Hyptis conferta var. angustata
Hyptisjurgensenii Briq. =Hyptis conferta var. angustata
Hyptisjurgensenii var. angustata Briq. =Hyptis conferta var. angustata
Hyptis lanicephala Epling & Jativa =Hyptis dilatata
Hyptis lundii Benth. =Hyptis conferta var. angustata
Hyptis micrantha Pohl ex Benth. =Hyptis mutabilis
Hyptis mutabilis var. canescens (Kunth) Briq. =Hyptis mutabilis
Hyptis parkeri var. verbenifolia (J.A. Schmidt ex Mart.) Epling =Hyptis parkeri
Hyptis polystachya Kunth =Hyptis mutabilis
Hyptis radiata Willd. [not in flora area] =Hyptis alata (Raf.) Shinners
Hyptis recurvata var. megacephala Benth. =Hyptis recurvata
Hyptis salzmannii Benth. =Hypenia salzmannii
Hyptis spicata Poit. =Hyptis mutabilis
Hyptis verbenifolia J.A. Schmidt ex Mart. ^Hyptis parkeri
Leonurus heterophyllus Sweet =Leonurus japonicus
Leonurus sibiricus auct. non L. 1753 '=Leonurus japonicus
Marsypianthes hyptoides Mart, ex Benth. =Marsypianthes chamaedrys
Mesosphaerum atrorubens (Poit.) Kuntze =Hyptis atrorubens
Mesosphaerum brevipes (Poit.) Kuntze =Hyptis brevipes
Mesosphaerum capitatum (Jacq.) Kuntze =Hyptis capitata
Mesosphaerum confertum (Pohl ex Benth.) Kuntze =Hyptis conferta
Mesosphaerum dilatatum (Benth.) Kuntze =Hyptis dilatata
Mesosphaerum excelsum (M. Martens & Galeotti) Kuntze =Hyptis conferta var. angustata
Mesosphaerum hirsutum (Kunth) Kuntze =Hyptis hirsuta
Mesosphaerum laciniatum (Benth.) Kuntze =Hyptis laciniata
Mesosphaerum lacustre (St.-Hil. ex Benth.) Kuntze =Hyptis lacustris
Mesosphaerum lanceolatum (Poir.) Kuntze =Hyptis lanceolata
Mesosphaerum lantanifolium (Poit.) Kuntze =Hyptis lantanifolia
Mesosphaerum microphyllum (Pohl ex Benth.) Kuntze =Hyptis microphylla
Mesosphaerum mutabile (Rich.) Kuntze =Hyptis mutabilis
Mesosphaerum parkeri (Benth.) Kuntze =Hyptis parkeri
Mesosphaerum pectinatum (L.) Kuntze =Hyptis pectinata
Mesosphaerum recurvatum (Poit.) Kuntze =Hyptis recurvata
Mesosphaerum suaveolens (L.) Kuntze =Hyptis suaveolens
Nepeta mutabilis Rich. =Hyptis mutabilis
Nepeta pectinata L. =Hyptis pectinata
Ocimum micranthum Willd. =Ocimum campechianum
Phlomis nepetifolia L. =Leonotis nepetifolia
References
Epling, C. 1935-1937. Synopsis of the South American Labiatae. Repert. Sp. Nov. Beih. 85: 1-341.
Epling, C. 1949. Revision del genero Hyptis (Labiatae). Rev. Mus. La Plata (Sect. Bot.) 7: 153-497.
Harley, R.M. 1988. Revision of generic limits in Hyptis Jacq. (Labiatae) and its allies. Bot. J. Linn. Soc. 98: 87-95.
Harley, R.M. 1999. Lamiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
5: 678-700.
Harley, R.M. 1999. Lamiaceae. In Mori, S.A., G. Cremers, C. Grade, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
363-369.
Kostermans, A.J.G.H. 1936. Labiatae. In A. Pulle, Flora of Suriname 4(1): 334-353.
Kostermans, A.J.G.H. 1937. Labiatae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 498.
Lauraceae
J.G Rohwer, H. van der Werff
Aioiiea benthamiana Mez
Aiouea guianensis Aubl.
Aiouea impressa (Meisn.) Kosterm.
Aiouea laevis (Mart.) Kosterm.
AM BO
BO
BO
GU
GU
su
su
FG
FG
FG
AM GU
AM
AM BO GU su?
AM
BO DA GU
GU
su FG
BO DA GU
AM
AM GU su FG
AM GU
GU
su
su
FG
GU su FG
DA GU su FG
AM BO GU su FG
AM BO GU
GU
GU
su
su
su
su
FG
FG
AM GU su FG
AM BO GU su FG
FG
AM GU? su FG
BO DA GU
FG
AM BO DA GU su FG
GU su FG
Checklist of the Plants of the Guiana Shield 323
Aioiiea longipetiolata van der Werff GU FG
Aiouea maguireana (C.K. Allen) S.S. Renner AM
Aioiiea myristicoides Mez BO
Aiouea opaca van der Werff FG
Aniba affmis (Meisn.) Mez
Aniba burchellii Kosterm.
Aniba canelilla (Kunth) Mez
Aniba cinnamomiflora C.K. Allen
Aniba cithfolia (Nees) Mez
Aniba cylindriflora Kosterm.
Aniba excelsa Kosterm.
Anibaferniginea Kubitzki
Aniba guianensis Aubl.
Aniba hostmanniana (Nees) Mez
Aniba hypoglaiica Sandwith
Anibajenmanii Mez
Aniba kappleri Mez
Aniba megaphylla Mez
Aniba paniirensis (Meisn.) Mez
Aniba pai^iflora (Meisn.) Mez
Aniba percoriacea C.K. Allen
Aniba permoUis (Nees) Mez
Aniba hparia (Nees) Mez
Aniba rosaeodora Ducke
Aniba taubertiana Mez
Aniba terminalis Ducke
Aniba nilliamsii O.C. Schmidt
Beilschmiedia cwviramea (Meisn.) Kosterm.
Beilschmiedia hexanthera van der Werff
Cassytha filiformis L.
Chlorocardium rodiei (R.H. Schomb.) Rohwer, H.G Richt. & van der Werff
Cinnamomum cinnamomifolium (Kunth) Kosterm. [recorded from FG in error,
but perhaps expected in GU]
Cinnamomum semecarpifolium (Meisn.) Kosterm. AM
Cinnamomum triplinei-ve (Ruiz & Pav.) Kosterm.
Ciyptocaiya guianensis Meisn.
Cryptocaiya mawniensis Benoist
Dicypellium caiyophyllaceum (Mart.) Nees
Endlicheria anomala (Nees) Mez AM
Endlicheria arunciflora (Meisn.) Mez AM
Endlicheria bracteolata (Meisn.) C.K. Allen AM
Endlicheria canescens Chanderbali
Endlicheria chalisea Chanderbali AM
Endlicheria dictifarinosa C.K. Allen AM
Endlicheria directonervia C.K. Allen AM
Endlicheria gracilis Kosterm. AM
Endlicheria macrophylla (Meisn.) Mez AM
Endlicheria multiflora (Miq.) Mez AM
Endlicheria nilssonii C.K. Allen
Endlicheria paradoxa Mez
Endlicheria punctulata (Mez) C.K. Allen
Endlicheria pyriformis (Nees) Mez
Endlicheria reflectens (Nees) Mez
Endlicheria rubriflora Mez AM
Endlicheria sericea Nees
Endlicheria aff. szyszylowiczii Mez AM
Endlicheria szyszylowiczii Mez GU?
Endlicheria vinotincta C.K. Allen AM
Kubitzkia macrantha (Kosterm.) van der Werff BO
Kubitzkia mezii (Kosterm.) van der Werff GU SU FG
Licaria armeniaca (Nees) Kosterm. AM GU
Licaria aurea (Huber) Kosterm. GU SU
Licaria aureosericea van der Werff GU
Licaria brasiliensis (Nees) Kosterm. AM BO SU?
DA FG
FG
FG
FG?
BO GU SU FG
BO GU su FG
BO GU
GU
SU
FG
BO
BO GU su FG
BO DA GU su FG
BO
su?
GU su FG
GU su FG
BO GU
su
FG
FG
FG
324 Checklist of the Plants of the Guiana Shield
Licaria camara (R.H. Schomb.) Kosterm.
Licaria cannella (Meisn.) Kosterm.
Licaria cannella subsp. tenuicarpa (Kosterm. ex Rodrigues) Kurz
Licaria chrysophylla (Meisn.) Kosterm.
Licaria debilis (Mez) Kosterm.
Licaria dolichantha Kurz
Licaria guianensis Aubl.
Licaria macrophylla (A.C. Sm.) Kosterm.
Licaria martiniana (Mez) Kosterm.
Licaria multiflora (Kosterm.) Kosterm.
Licaria oppositifolia (Nees) Kosterm.
Licaria pachycarpa (Meisn.) Kosterm.
Licaria polyphylla (Nees) Kosterm.
Licaria rufotomentosa van der Werff
Licaria subbullata Kosterm.
Licaria tomentosa van der Werff
Licaria trinervis van der Werff
Licaria vernicosa (Mez) Kosterm.
Mezilaurus itauba (Meisn.) Taub. ex Mez
Mezilaurus lindaviana Schwacke & Mez
Mezilaurus sprucei (Meisn.) Taub. ex Mez
Mezilaurus thoroflora van der Werff
Nectandra amazonum Nees
Nectandra aurea Rohwer
Nectandra cissiflora Nees
Nectandra cuspidata Nees
Nectandrafulva Rohwer
Nectandra globosa (Aubl.) Mez
Nectandra hihua (Ruiz & Pav.) Rohwer
Nectandra lineata (Kunth) Rohwer
Nectandra martinicensis Mez
Nectandra paucinervia Coe-Teix.
Nectandra pearcei Mez
Nectandra pichurim (Kunth) Mez
Nectandra pulverulenta Nees
Nectandra purpurea (Ruiz & Pav.) Mez
Nectandra reticulata (Ruiz & Pav.) Mez
Nectandra ruforamula Rohwer
Nectandra sanguinea Rol. ex Rottb.
Ocotea aciphylla (Nees & Mart, ex Nees) Mez
Ocotea adenotrachelium (Nees) Mez
Ocotea aff. amazonica (Meisn.) Mez [name of uncertain application]
Ocotea argyrophylla Ducke
Ocotea atrata C.K. Allen
Ocotea basirecurva C.K. Allen
Ocotea bijuga (Rottb.) Bemardi [=Ocotea neesianal]
Ocotea bofo Kunth
Ocotea bracteosa (Meisn.) Mez
Ocotea canaliculata (Rich.) Mez
Ocotea caracasana (Nees) Mez
Ocotea ceanothifolia (Nees) Mez
Ocotea cernua (Nees) Mez [s.l.]
Ocotea cinerea van der Werff
Ocotea commutata (Nees) Mez
Ocotea cowaniana C.K. Allen
Ocotea crassifolia (Nees) Mez [Mt. Roraima]
Ocotea cujumary Mart.
Ocotea cymbarum Kunth
Ocotea debilis Mez
Ocotea depauperata C.K. Allen
Ocotea diffusa van der Werff
Ocotea discrepens C.K. Allen
Ocotea divaricata (Nees) Mez
Ocotea duidensis Moldenke
GU FG
AM DA GU
GU
su FG
BO GU su FG
AA/T
DA GU su FG
AiVl
ATVT
GU su FG
AiVl
GU su FG
GU
BO GU
GU FG
AM
BO
GU su FG
GU su FG
BO
AM
BO GU su FG
AM
BO
GU?
GU
su FG
AM
BO
GU
GU su
GU su FG
AM BO DA GU su
AM
AM BO DA GU su FG
AM DA GU su FG
AM
BO
GU
BO GU su
AM BO DA
FG
FG
AM BO GU su FG
AM
BO GU su
AM BO GU su FG
AM
AM BO GU?
su
FG
FG
AM
AM
su
AM BO
AM BO
AM BO DA GU
GU
su FG
BO GU FG
AM BO DA GU su FG
AM
AM
AM
AM
BO
BO
BO
BO
BO
DA
GU?
GU
GU SU
GU SU
FG
FG
FG
FG
FG
FG
FG
BO GU SU FG
BO FG
AM
AM BO
BO
DA GU SU FG
AM BO
AM BO DA GU SU FG
AM GU
AM BO GU SU FG
GU SU FG
GU SU
GU
Checklist of the Plants of the Guiana Shield 325
Ocotea endlicheriopsis Mez GU SU FG
Ocotea erectifolia (C.K. Allen) van der Werff BO
Ocotea esmeraldana Moldenke AM
Ocoteafascicidata (Nees) Mez
Ocotea fendleri (Meisn.) Rohwer
Ocoteaflavantha van der Werff
Ocoteafloribimda (Sw.) Mez
Ocotea glabra van der Werff
Ocotea glaiicophylla Moldenke
Ocotea glomerata (Nees) Mez
Ocotea gracUis (Meisn.) Mez
Ocotea guianensis Aubl.
Ocotea guianensis var. guianensis
Ocotea guianensis var. subsericea Kosterm.
Ocotea hemsleyana Mez
Ocotea huberi van der Werff
Ocotea indirectinervia C.K. Allen FG
Ocotea javitensis (Kunth) Pittier
Ocoteajulianii van der Werff
Ocotea laticostata C.K. Allen
Ocotea leucoxylon (Sw.)Laness. AM BO DA GU SU FG
Ocotea liesneri van der Werff
Ocotea longifolia Kunth
Ocotea megacarpa van der Werff
Ocotea mucronata (Poir.) Kosterm. FG
Ocotea myriantha (Meisn.) Mez
Ocotea neblinae C.K. Allen
Ocotea neesiana (Miq.) Kosterm. [see also Ocotea bijuga]
Ocotea nigra Benoist
Ocotea nilssonii C.K. Allen
Ocotea oblonga (Meisn.) Mez
Ocotea pauciflora (Nees) Mez
Ocotea percurrens Vicentini
Ocotea perrobusta (C.K. Allen) Rohwer BO
Ocotea petalanthera (Meisn.) Mez
Ocotea puberula (Rich.) Nees
Ocotea revoluta Moldenke
Ocotea rhodophylla Vicentini
Ocotea roseopedunculata van der Werff
Ocotea rubrinervis Mez
Ocotea rufovestita Ducke
Ocotea sanariapensis Lasser
Ocotea sandwithii Kosterm.
Ocotea scabrella van der Werff
Ocotea scandens Kosterm.
Ocotea schomburgkiana (Nees) Mez BO DA
Ocotea sp. A
Ocotea splendens (Meisn.) Baill. AM
Ocotea subterminalis van der Werff
Ocotea tillettsiana C.K. Allen BO
Ocotea tomentella Sandwith DA
Ocotea tomentosa van der Werff
Ocotea venosa Gleason
Ocotea wurdackiana C.K. Allen
Ocotea yutajensis C.K. Allen
Persea americana Mill, [cultivated, occasionally escaping] AM BO DA GU SU FG
Persea areolatocostae (C.K. Allen) van der Werff
Persea caerulea (Ruiz & Pav.) Mez
Persea croatii van der Werff
Persea croizatii van der Werff
Persea fastigiata Kopp
Persea Jhiviatilis van der Werff
Persea grandiflora Kopp
Perseajenmanii Mez AM BO GU
AM BO
AM
BO
AM
AM
AM
AM
AM
AM
AM
BO
AM BO
AM
AM BO
AM
AM BO
BO
AM BO
AM BO
BO
GU SU FG
GU SU FG
GU SU FG
GU
SU FG
SU FG
GU SU FG
GU? FG
FG
GU
FG
FG
GU SU FG
GU SU FG
FG
GU
GU FG
AM
AM
BO
AM
AM
BO
AM
AM
AM BO
AM
AM
FG
FG
GU su FG
FG
GU
GU
GU su FG
GU su FG
GU FG
FG
su FG
GU FG
326 Checklist of the Plants of the Guiana Shield
Perseajulianae van der Werff SU
Persea maguirei Kopp AM
Persea nivea Mez GU SU FG
Persea perseiphylla (C.K. Allen) van der Werff AM
Persea pseudofasciculata Kopp AM
Persea steyermarkii C.K. Allen AM
Pleurothyrium amapaense C.K. Allen AM SU
Rhodostemonodaphne celiana (C.K. Allen) Rohwer AM
Rhodostemonodaphne crenaticupula Madrinan
Rhodostemonodaphne elephantopus Madrinan
Rhodostemonodaphne kunthiana (Nees) Rohwer AM BO
Rhodostemonodaphne leptoclada Madrinan
Rhodostemonodaphne miranda (Sandwith) Rohwer
Rhodostemonodaphne mirecolorata (C.K. Allen) Rohwer
Rhodostemonodaphne morii Madriiian
Rhodostemonodaphne praeclara (Sandwich) Madrinan AM BO DA
Rhodostemonodaphne revolutifolia Madrinan
Rhodostemonodaphne riifovirgata Madrinan
Rhodostemonodaphne saiilensis Madrinan
Rhodostemonodaphne scandens Madriiian
Rhodostemonodaphne steyermarkiana (C.K. Allen) van der Werff BO
Sextonia rubra {Mez) van dQxWQxff BO GU SU FG
Synonyms and Misidentifications
Acrodiclidium anacardioides Meisn. =Mezilaurus itauba
Acrodiclidium aubletii Kosterm. =Licaria guianensis
Acrodiclidium brasiliense Nees =Licaria brasiliensis
Acrodiclidium canella (Meisn.) Mez =Licaria cannella
Acrodiclidium cayennense (Meisn.) Mez =Licaria cannella
Acrodiclidium chrysophyllum Meisn. =Licaria chrysophylla
Acrodiclidium debile Mez =Licaria debilis
Acrodiclidium guianense Nees =Licaria polyphylla
Acrodiclidium guianense var. caudatum Meisn. =Licaria polyphylla
Acrodiclidium itauba Meisn. =Mezilaurus itauba
Acrodiclidium kunthiana Nees =Rhodostemonodaphne kunthiana
Acrodiclidium macrophyllum A.C. Sm. =Licaria macrophylla
Acrodiclidium martinianum Mez =Licaria martiniana
Acrodiclidium meissneri Mez =Licaria polyphylla
Acrodiclidium oppositifolium Nees =Licaria oppositifolia
Acrodiclidium rigidum Kosterm. =Licaria chrysophylla
Acrodiclidium sprucei Meisn. =Mezilaurus sprucei
Aiouea demerarensis Kosterm. =Aiouea guianensis
Aiouea densiflora Nees =Aiouea laevis
Aiouea pernitida C.K. Allen =Aiouea myristicoides
Aiouea perrobusta C.K. Allen =Ocotea perrobusta
Aiouea schomburgkii Meisn. =Aiouea guianensis
Ampelodaphne arunciflora Meisn. =Endlicheria arunciflora
Ampelodaphne dasyantha Meisn. =Endlicheria multiflora
Ampelodaphne macrophylla Meisn. =Endlicheria macrophylla
Aniba castanea C.K. Allen =AnibajenmaniI
Aniba duckei Kosterm. =Aniba rosaeodora
Anibafirmula (Nees & Mart.) Mez [not in flora area]
Aniba gonggrijpii Kosterm. =Aniba panurensis
Aniba koumaroucapa Kosterm. =Aniba megaphylla
Aniba mas Kosterm. =Aniba panurensis
Aniba muca (Ruiz &. Pav.) Mez [not in flora area]
Aniba muelleriana Mez =Aniba citrifolia
Aniba murcana C.K. Allen =Aniba williamsii
Aniba ovalifolia Mez =Aniba hypoglauca
Aniba pittieri O.C. Schmidt =Aniba hostmanniana
Aniba riparia auct. non (Nees) Mez.: sensu C.K. Allen, Mem. New York Bot. Gar =Aniba kappleri
Aniba salicifolia (Nees) Mez =Aniba guianensis
Aniba sericea (Nees) Brooks =Endlicheria sericea
Aniba simulans C.K. Allen =Aniba taubertiana
Aniba trinitatis (Meisn.) Mez =Aniba citrifolia
Checklist of the Plants of the Guiana Shield 327
Aydendron affine Meisn. ^Aniba affinis
Aydendron cannella Meisn. =Licaria cannella
Aydendron citrifolium Nees =Aniba citrifolia
Aydendron curvirameum Meisn. =Beilschmiedia curviramea
Aydendron hostmannianum Nees =Aniba hostmanniana
Aydendron panurense Meisn. =Aniba panurensis
Aydendron trinitatis Meisn. =Aniba citrifolia
Cassytha americana Nees =Cassythafdiformis
Cassytha baccifera Sol. ex J.S. Muell. (Cactaceae) =Rhipsalis baccifera
Cinnamomum areolatocostae (C.K. Allen) Kosterm. =Persea areolatocostae
Cinnamomumfilamentosum (C.K. Allen) Kosterm. =Cinnamomum triplinerve
Clinostemon maguireanum (C.K. Allen) Kurz =Mezilaurus thoroflora
Cryptocarya canelilla Kunth =Aniba canelilla
Cryptocarya laevis Mart. =Aiouea laevis
Cryptocarya moschata Mart, [not in flora area]
Cryptocarya mucronata (Poir.) Spreng. =Ocotea mucronata
Cryptocarya pachycarpa Gleason =Aniba citrifolia
Endlicheria endlicheriopsis (Mez) Kosterm. =Ocotea endlicheriopsis
Endlicheria grandis Mez =Rhodostemonodaphne praeclara
Endlicheria impressa Mez =Aiouea impressa
Endlicheria levelii C.K. Allen =Endlicheria multiflora
Endlicheria maguireana C.K. Allen =Aiouea maguireana
Endlicheria melinonii Benoist =Endlicheria multiflora
Endlicheria wurdackiana C.K. Allen =Endlicheria rubriflora
Evonymodaphne armeniaca Nees =Licaria armeniaca
Goeppertia anomala Nees =Endlicheria anomala
Goeppertia multiflora Miq. =Endlicheria multiflora
Goeppertia reflectens Nees ^Endlicheria reflectens
Goeppertia sericea var. bracteolata Meisn. =Endlicheria bracteolata
Gymnobalanusfendleri Meisn. =Ocoteafendleri
Laurus caerulea Ruiz & Pav. =Persea caerulea
Laurus canaliculata Rich. =Ocotea canaliculata
Laurusfloribunda Sw. =Ocoteafloribunda
Laurus globosa Aubl. =Nectandra globosa
Laurus hihua Ruiz & Pav. =Nectandra hihua
Laurusjavitensis Kunth =Ocoteajavitensis
Laurus leucoxylon Sw. =Ocotea leucoxylon
Laurus puberula Rich. =Ocotea puberula
Laurus reticulata Ruiz & Pav. =Nectandra reticulata
Laurus triplinervis Ruiz & Pav. =Cinnamomum triplinerve
Licaria appelii (Mez) Kosterm. =Licaria guianensis
Licaria cayennensis (Meisn.) Kosterm. =Licaria cannella
Licaria cymbarum (Kunth) Pittier =Ocotea cymbarum
Licariafoveolata (Mez) Lemee =Licaria martiniana
Licaria maguireana C.K. Allen =Mezilaurus thoroflora
Licaria martiniana auct. non (Mez) Kosterm.; sensu C.K. Allen, Mem. New York Bot. G =Licaria oppositifolia
Licaria meissneri (Mez) Kosterm. =Licaria polyphylla
Licaria rigida (Kosterm.) Kosterm. =Licaria chrysophylla
Licaria schultesii C.K. Allen =Licaria macrophylla
Licaria simulans C.K. Allen =Licaria vernicosa
Licaria wilhelminensis C.K. Allen =Licaria subbullata
Mespilodaphne ceanothifolia Nees =Ocotea ceanothifolia
Mespilodaphne myriantha Meisn. =Ocotea myriantha
Mespilodaphne oblonga Meisn. =Ocotea oblonga
Mezilaurus anacardioides (Meisn.) Taub. ex Mez =Mezilaurus itauba
Mezilaurus maguireana C.K. Allen =Mezilaurus sprucei
Mezilaurus wurdackiana C.K. Allen =Mezilaurus lindaviana
Misanteca maguireana (Allen) Lundell =Mezilaurus thoroflora
Nectandra ambigua Meisn. =Nectandra amazonum
Nectandra angustifolia (Schrad.) Nees [not in flora area]
Nectandra barcellensis Meisn. =Ocotea cymbarum
Nectandra bredemeyeriana Nees =Nectandra hihua
Nectandra canescens Nees [not in flora area]
Nectandra cymbarum (Kunth) Nees =Ocotea cymbarum
328 Checklist of the Plants of the Guiana Shield
Nectandra glandulifoUa Lasser =Nectandra martinicensis
Nectandra grandis (Mez) Kosterm. =Rhodostemonodaphne praeclara
Nectandra guianensis Meisn. =Nectandra sanguinea
Nectandra kaburiensis Kosterm. ^Nectandra pearcei
Nectandra kunthiana (Nees) Kosterm. ^Rhodostemonodaphne kunthiana
Nectandra laurel Nees [not in flora area]
Nectandra leucantha var. attenuata Meisn. =Nectandra hihua
Nectandra leucantha var. guianensis Meisn. =Nectandra hihua
Nectandra lucida Nees (1833) [not in flora area] =Endlicheria paniculata (Spreng.) Macbr.
Nectandra lucida Nees (1836) =Nectandra hihua
Nectandra magnoliifolia Meisn. [not in flora area] =Nectandra canescens Nees
Nectandra maguireana C.K. Allen [not in flora area]
Nectandra meyeriana Lasser =Rhodostemonodaphne kunthiana
Nectandra miranda Sandwith =Rhodostemonodaphne miranda
Nectandra mollis (Kunth) Nees =Nectandra reticulata
Nectandra neesiana Miq. =Ocotea neesiana
Nectandra olivacea Lasser =Nectandra cuspidata
Nectandra pallida Nees =Nectandra amazonum
Nectandra pisi Miq. =Nectandra globosa
Nectandra pittieri Lasser =Nectandra reticulata
Nectandra polyphylla Nees =Licaria polyphylla
Nectandra praeclara Sandwith =Rhodostemonodaphne praeclara
Nectandra rhynchophylla Meisn. =Ocotea aciphylla
Nectandra rubra (Mez) C.K. Allen =Sextonia rubra
Nectandra schomburgkii Meisn. ^Nectandra hihua
Nectandra spumea Kubitzki =Ocotea esmeraldana
Nectandra surinamensis Mez ^Nectandra cissiflora
Nectandra vaga Meisn. =Nectandra globosa
Nectandra viburnoides Meisn. [not in flora area]
Nectandra willdenoviana Nees =Nectandra hihua
Ocotea abbreviata Schwacke & Mez [not in flora area]
Ocotea aciphylla var chimantaensis C.K. Allen =Ocotea aciphylla
Ocotea acutangula (Miq.) Mez =Ocotea leucoxylon
Ocotea barcellensis (Meisn.) Mez =Ocotea cymbarum
Ocotea budowskiana Bernardi =Ocotea erectifolia
Ocotea castanea C.K. Allen =Ocotea schomburgkiana
Ocotea caudata (Nees) Mez =Ocotea cernua
Ocotea celiana C.K. Allen =Rhodostemonodaphne celiana
Ocotea costulata (Nees) Mez =Ocotea aciphylla
Ocotea cuprea (Meisn.) Mez =Ocotea oblonga
Ocotea directoramea C.K. Allen =Ocotea debilis
Ocotea dissimilis C.K. Allen =Ocotea splendens
Ocotea duotincta C.K. Allen =Ocotea leucoxylon
Ocotea fallax (Miq.) Mez =Ocotea neesiana
Ocoteaferruginea auct. non (Meisn.) Mez 1889: sensu Bernardi, CandoUea 30: 261. =Rhodostemonodaphne steyermarkiana
Ocoteafinium C.K. Allen =Ocotea aciphylla
Ocoteafirmula Mart, [nomen nudum] [not in flora area] ^Anibafirmula (Nees) Mez
Ocoteaflorulenta (Meisn.) Mez =Ocotea neesiana
Ocoteafulvifolia C.K. Allen "^Ocotea aciphylla
Ocotea glandulosa Lasser =Ocoteafendleri
. Ocotea glaucina (Meisn.) Mez [not in flora area] =Ocotea notata (Nees) Mez
Ocotea globifera Mez =Ocotea splendens
Ocotea indecora (Schott) Mez [not in flora area]
Ocoteajenmanii Mez [nomen nudum]
Ocotea kunthiana (Nees) Mez =Rhodostemonodaphne kunthiana
Ocotea lasseriana C.K. Allen =Ocotea leucoxylon
Ocotea laxiflora (Meisn.) Mez =Ocotea pauciflora
Ocotea lineata Kunth =Nectandra lineata
Ocotea maguireana C.K. Allen =Ocotea aciphylla
Ocotea marowynensis (Miq.) Mez =Ocotea cernua
Ocotea martiniana (Nees) Mez =Ocotea puberula
Ocotea mirecolorata C.K. Allen =Rhodostemonodaphne mirecolorata
Ocotea opifera Mart. =Ocotea longifolia
Ocotea pendula (Poepp.) Eng\. [nomen nudum]
Checklist of the Plants of the Guiana Shield 329
Ocotea perseiphylla C.K. Allen =Persea perseiphylla
Ocotea persulcata C.K. Allen =Ocotea leucoxylon
Ocotea pichurim Kunth =Nectandra pichurim
Ocotea pubescens (Nees) Mez [not in flora area]
Ocotea punctulata Mez =EndUcheria punctulata
Ocotea punctulata auct. non Mez. 1889: sensu C.K. Allen, Mem. New York Bot. Card. ^Ocotea cernua
Ocotea rhynchophylla (Meisn.) Mez =Ocotea aciphylla
Ocotea rodiei (R.H. Schomb.) Mez ^Chlorocardium rodiei
Ocotea roraimae auct. non Mez 1920 =Licaria tomentosa
Ocotea roraimae Mez =Ocotea aciphylla
Ocotea rubra Mez =Sextonia rubra
Ocotea sericiflora C.K. Allen =Ocotea aciphylla
Ocotea simulans C.K. Allen =Endlicheria anomala
Ocotea steyermarkiana C.K. Allen =Rhodostemonodaphne steyermarkiana
Ocotea subalveolata C.K. Allen =Ocotea schomburgkiana
Ocotea subglabra Benoist =Ocotea puberula
Ocotea urophylla (Meisn.) Mez =Ocotea cernua
Ocotea vernicosa Mez =Licaria vernicosa
Ocotea wachenheimii Benoist =Ocotea floribunda
Oreodaphne aciphylla Nees & Mart, ex Nees =Ocotea aciphylla
Oreodaphne adenotrachelium Nees ^Ocotea adenotrachelium
Oreodaphne amazonica Meisn. =Ocotea amazonica
Oreodaphne bracteosa Meisn. =Ocotea bracteosa
Oreodaphne caudata Nees =Ocotea cernua
Oreodaphne cernua Nees =Ocotea cernua
Oreodaphne costulata Nees =Ocotea aciphylla
Oreodaphne crassifolia Nees =Ocotea crassifolia
Oreodaphne diospyrifolia Meisn. var. incompta Meisn. =Ocotea pauciflora
Oreodaphnefasciculata Nees =Ocoteafasciculata
Oreodaphne florulenta Meisn. =Ocotea neesiana
Oreodaphne glomerata Nees =Ocotea glomerata
Oreodaphne gracilis Meisn. =Ocotea gracilis
Oreodaphne guianensis var. aurea Meisn. =Ocotea guianensis
Oreodaphne hostmanniana Miq. =Ocotea puberula
Oreodaphne martiniana Nees =Ocotea puberula
Oreodaphne pauciflora Nees =Ocotea pauciflora
Oreodaphne schomburgkiana Nees =Ocotea schomburgkiana
Oreodaphne semecarpifolia Meisn. =Cinnamomum semecarpifolium
Oreodaphne splendens Meisn. =Ocotea splendens
Persea cinnamonifolia Kunth =Cinnamomum triplinerve
Persea coriacea Kosterm. [type based on mixed collection fide van der Werff]
Persea fastigiata var. pilosa Kopp =Persea fastigiata
Persea lignitepala Lasser =Persea caerulea
Persea sprucei Kosterm. =Cinnamomum semecarpifolium
Phoebe areolatocostae C.K. Allen =Persea areolatocostae
Phoebe cinnamomifolia (Kunth) Nees ^Cinnamomum triplinerve
Phoebe erectifolia C.K. Allen =Ocotea erectifolia
Phoebefilamentosa C.K. Allen =Cinnamomum triplinerve
Phoebe semecarpifolia (Meisn.) Mez =Cinnamomum semecarpifolium
Phoebe steyermarkiana C.K. Allen =Ocotea julianii
Pleurothyrium cowaniana C.K. Allen =Rhodostemonodaphne kunthiana
Pleurothyrium cowanianum C.K. Allen ^Rhodostemonodaphne kunthiana
Rhodostemonodaphne grandis (Mez) Rohwer =Rhodostemonodaphne praeclara
Strychnodaphne puberula var. truncata Meisn. =Ocotea puberula
Systemonodaphne geminiflora Mez =Kubitzkia mezii
Systemonodaphne macrantha Kosterm. =Kubitzkia macrantha
Systemonodaphne mezii Kosterm. ^Kubitzkia mezii
References
Allen, C.K. 1964. Lauraceae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part 5. Mem. New
York Bot. Card. 10(5): 1-278.
Allen, C.K. 1965. Lauraceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 6. Mem. New York Bot.
Card. 12(3): 1-285.
330 Checklist of the Plants of the Guiana Shield
Jansen-Jacobs, M.J. 1976. Lauraceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 451-
484.
Kostermans, A.J.G.H. 1936-1938. Revision of the Lauraceae. Rec. Trav. Bot. Neerl. 33: 719-757 (I), 34: 500-609 (II), 35: 56-129
(III), 35: 834-931 (V).
Kostermans, A.J.G.H. 1936. Lauraceae. In A. Pulle, Flora of Suriname 2(1): 244-337.
Kostermans, A.J.G.H. 1939. Lauraceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 481-487.
Kubitzki, K. & S. Renner. 1982. Lauraceae I (Aniba and Aiouea). Flora Neotropica Monogr. 31: 1-124.
Rohwer, J.G. 1986. Prodromus einer Monographic der Gattung Ocotea Aubl., sensu lato. Mitt. Inst. Allg. Bot. Hamburg 20: 3-
278.
Rohwer, J.G. 1993. Lauraceae: Nectandra. Flora Neotrop. Monogr. 60: 1-332.
Rohwer, J.G, H.G Richter, & H. van der Werff. 1991. Two new genera of neotropical Lauraceae and critical remarks on the
generic delimitation. Ann. Missouri Bot. Gard. 78: 388-400.
Werff, H. van der. 1986. Kubitzkia van der Werff, a new name for a genus of neotropical Lauraceae. Taxon 35(1): 164-166.
Werff, H. van der. 1987. A revision of Mezilaurus. Ann. Missouri Bot. Gard. 74: 153-182.
Werff, H. van der. 1991. A key to the genera of Lauraceae in the New World. Ann. Missouri Bot. Gard. 78: 377-387.
Werff, H. van der. 1995. Two new species C)f Lauraceae from central French Guiana. Brittonia 47(4): 372-375.
Werff, H. van der. 1997. Sextonia, a new genus of Lauraceae from South America. Novon 7(4):436-439.
Werff, H. van der. 2002. Lauraceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
370-384.
Werff, H. van der & J.G. Rohwer. 1999. Lauraceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 5: 700-750.
Lecythidaceae
S.A. Mori, GT. Prance
Allantoma lineata (Mart, ex O. Berg) Miers
Asteranthos brasiliensis Desf.
Bertholletia excelsa Bonpl.
Corythophora amapaensis Pires ex S.A. Mori & Prance
Corythophora labriculata (Eyma) S.A. Mori & Prance
Corythophora hmosa W.A. Rodrigues
Corythophora rimosa subsp. rubra S.A. Mori
Couratari calycina Sandwith
Couratari gloriosa Sandwith
Couratari guianensis Aubl.
Couratari multiflora (Sm.) Eyma
Couratari oblongifolia Ducke & R. Knuth
Couratari oligantha A.C. Sm.
Couratari riparia Sandwith
Couratari sandwithii Prance
Couratari stellata A.C. Sm.
Couratari tenuicarpa A.C. Sm.
Couroupita guianensis Aubl.
Eschweilera alata K.C. Sm.
Eschweilera apiculata (Miers) A.C. Sm.
Eschweilera boltenii S.A. Mori
Eschweilera bracteosa (Poepp. ex O. Berg) Miers
Eschweilera chartaceifolia S.A. Mori
Eschweilera collina Eyma
Eschweilera congestiflora (Benoist) Eyma
Eschweilera coriacea (DC.) S.A. Mori
Eschweilera decolorans Sandwith
Eschweilera eperuetorum Sandwith
Eschweilerafanshawei Sandwith
Eschweilera grandiflora (Aubl.) Sandwith
Eschweilera laevicarpa S.A. Mori
Eschweilera micrantha (O. Berg) Miers
Eschweilera neblinensis S.A. Mori
Eschweilera paniculata (O. Berg) Miers
Eschweilera parviflora (Aubl.) Miers
Eschweilera parvifolia Mart, ex DC.
Eschweilera pedicellata (Rich.) S.A. Mori
Eschweilera piresii S.A. Mori
AM
AM
AM GU
GU
SU
su
SU
su
FG
FG
FG
FG
FG
GU su FG
AM BO DA GU su FG
BO DA GU su FG
GU su FG
AM
BO
GU
GU
su
AM GU su
su
FG
BO DA GU su FG
BO GU
GU
su
su
FG
FG
AM
FG
AM BO DA GU su FG
GU? su FG
AM BO DA GU su FG
AM BO DA GU
GU
GU
su
su
FG
FG
AM BO FG
AM BO GU su FG
AM
AM
AM BO DA GU su FG
AM BO GU
AM BO GU su FG
FG
Checklist of the Plants of the Guiana Shield 331
Eschweilera piresii subsp. viridipetala S.A. Mori FG
Eschweilera potaroensis Sandwith GU
Eschweilera praeclara Sandwith GU
Eschweilera revoluta S.A. Mori AM
Eschweilera rionegrense S.A. Mori AM
Eschweilera roraimensis S.A. Mori AM
Eschweilera sagotiana Miers
Eschweilera simiorum (Benoist) Eyma
Eschweilera squamata S.A. Mori
Eschweilera subglandulosa (Steud. ex O. Berg) Miers
Eschweilera tenuifolia (O. Berg) Miers
Eschweilera wachenheimii (Benoist) Sandwith
Gustavia acuminata S.A. Mori
Gustavia augusta L.
Gustavia coriacea S.A. Mori
Gustavia gigantophylla Sandwith
Gustavia hexapetala (Aubl.) Sm.
Gustavia poeppigiana O. Berg
Gustavia pulchra Miers
Lecythis alutacea (A.C. Sm.) S.A. Mori
Lecythis brancoensis (R. Knuth) S.A. Mori
Lecythis chartacea O. Berg
Lecythis confertiflora (A.C. Sm.) S.A. Mori
Lecythis corrugata Poit.
Lecythis corrugata subsp. corrugata
Lecythis corrugata subsp. rosea (Spruce ex O. Berg) S.A. Mori
Lecythis holcogyne (Sandwith) S.A. Mori
Lecythis idatimon Aubl.
Lecythis persistens Sagot
Lecythis persistens subsp. aurantiaca S.A. Mori
Lecythis persistens subsp. persistens
Lecythis pneumatophora S.A. Mori
Lecythis poiteaui O. Berg
Lecythis schomburgkii O. Berg
Lecythis zabucajo Aubl.
Synonyms and Misidentifications
Allantoma fagifolia (Miq. ex O. Berg) Miers =Couratari multiflora
Bertholletia nobilis Miers =Bertholletia excelsa
Cercophora anomala Miers =Lecythis chartacea
Chytroma amara (Aubl.) Miers =Lecythis idatimon
Chytroma apiculata Miers =Eschweilera apiculata
Chytroma brancoensis R. Knuth =Lecythis brancoensis
Chytroma chartacea (O. Berg) Miers =Lecythis chartacea
Chytroma collina (Eyma) R. Knuth =Eschweilera collina
Chytroma congestiflora (Benoist) R. Knuth =Eschweilera congestiflora
Chytroma corrugata (Poit.) R. Knuth =Lecythis corrugata
Chytromafoetida R. Knuth =Lecythis poiteaui
Chytroma holcogyne (Sandwith) R. Knuth ^Lecythis holcogyne
Chytroma idatimon (Aubl.) Nied. =Lecythis idatimon
Chytroma labriculata (Eyma) R. Knuth =Corythophora labriculata
Chytroma laevicula (Miers) R. Knuth =Lecythis chartacea
Chytroma marawynensis (O. Berg) Miers =Lecythis chartacea
Chytroma perspicua Miers =Eschweilera pedicellata
Chytroma praeclara (Sandwith) Kunth =Eschweilera praeclara
Chytroma rosea (Spruce ex O. Berg) Miers =Lecythis corrugata subsp. rosea
Chytroma rubiflora Miers ^Lecythis corrugata subsp. corrugata
Chytroma salebrosa (O. Berg) Miers ^Lecythis corrugata subsp. corrugata
Chytroma schomburgkiana (O. Berg) Miers =Lecythis schomburgkii
Chytroma simiorum (Benoist) R. Knuth =Eschweilera simiorum
Chytroma simiorum van latifolia (Eyma) R. Knuth =Eschweilera simiorum
Couratari coriacea Mart, ex O. Berg =Couratari multiflora
Couratari fagifolia (Miq. ex O. Berg) Eyma =Couratari multiflora
Couratari lineata Mart, ex O. Berg =Allantoma lineata
Couratari pulchra Sandwith =Couratari guianensis
GU SU FG
GU SU
SU
FG
FG
AM BO DA GU SU FG
AM BO
GU SU FG
AM
AM BO DA GU SU FG
AM
GU
AM BO GU SU FG
AM BO DA GU
AM
AM BO GU
GU
SU FG
AM BO DA GU SU FG
GU SU FG
AM BO DA GU SU FG
DA GU SU FG
AM BO GU
GU SU
SU
FG
FG
GU SU? FG
FG
GU SU?
SU
FG
FG
FG
GU
AM BO DA GU SU FG
332 Checklist of the Plants of the Guiana Shield
Couratari reticulata A.C. Sm. =Couratari stellata
Couroupita guianensis var. surinamensis (Mart, ex O. Berg) Eyma =Couroupita guianensis
Couroupita surinamensis Mart, ex O. Berg =Couroupita guianensis
Couroupita venezuelensis R. Knuth =Couroupita guianensis
Eschweilera alutacea A.C. Sm. =Lecythis alutacea
Eschweilera amara (Aubl.) Nied. =Lecythis idatimon
Eschweilera chaffanjonii (Benoist) R. Knuth =Eschweilera parvifolia
Eschweilera chartacea (O. Berg) Eyma =Lecythis chartacea
Eschweilera conduplicata A.C. Sm. =Lecythis corrugata subsp. rosea
Eschweilera confertiflora A.C. Sm. =Lecythis confertiflora
Eschweilera confusa (Benoist) R. Knuth [nomen nudum]
Eschweilera corrugata (Poit.) Miers =Lecythis corrugata
EschweileraJIaccida Miers =Eschweilera pedicellata
Eschweilerafloribunda Eyma =Eschweilera micrantha
Eschweilera gracilipes (Sagot) R. Knuth =Eschweilera micrantha
Eschweilera grata Sandwith =Eschweilera parviflora
Eschweilera holcogyne Sandwith =Lecythis holcogyne
Eschweilera idatimon (Aubl.) Nied. =Lecythis idatimon
Eschweilera idatimonoides (O. Berg) Miers =Eschweilera pedicellata
Eschweilerajenmanii R. Knuth =Lecythis corrugata subsp. corrugata
Eschweilerajucunda (Benoist) R. Knuth =Eschweilera sagotiana
Eschweilera labriculata Eyma =Corythophora labriculata
Eschweilera longipes (Poit.) Miers ^Eschweilera pedicellata
Eschweilera lutea (Aubl.) Miers =Lecythis idatimon
Eschweilera macrophylla (O. Berg) Miers =Eschweilera pedicellata
Eschweilera marawynensis (O. Berg) Nied. =Lecythis chartacea
Eschweilera melinonis (Sagot) Knuth =Eschweilera sagotiana
Eschweilera montana A.C. Sm. =Eschweilera parviflora
Eschweilera obtecta (Miers) Nied. ^Eschweilera tenuifolia
Eschweilera odora (Poepp. ex O. Berg) Miers =Eschweilera coriacea
Eschweilera ovalis R. Knuth =Lecythis persistens subsp. persistens
Eschweilera pachycarpa Mart, [nomen nudum]
Eschweilera pallida Miers =Eschweilera coriacea
Eschweilera patrisii R. Knuth =Lecythis corrugata subsp. corrugata
Eschweilera persaudii R. Knuth =Lecythis alutacea
Eschweilera platycarpa (Poit.) Miers =Eschweilera pedicellata
Eschweilera poiteaui (O. Berg) Nied. =Lecythis poiteaui
Eschweilera retroflexa (Benoist) R. Knuth =Eschweilera coriacea
Eschweilera salebrosa (O. Berg) Nied. =Lecythis corrugata subsp. corrugata
Eschweilera schomburgkii (O. Berg) Nied. =Lecythis schomburgkii
Eschweilera simiorum var. latifolia Eyma =Eschweilera simiorum
Eschweilera tapuya (Benoist) R. Knuth =Eschweilera sagotiana
Grias aubletiana Miers =Gustavia augusta
Grias tetrapetala (Aubl.) Nied. =Gustavia augusta
Gustavia acuta S.A. Mori =Gustavia pulchra
Gustavia augusta var. guianensis O. Berg =Gustavia augusta
Gustavia augusta var. verrucosa Mart, ex O. Berg =Gustavia augusta
Gustaviafastuosa Willd. =Gustavia hexapetala
Gustaviafastuosa var. angustisepala O. Berg =Gustavia hexapetala
Gustaviafastuosa var. latisepala O. Berg =Gustavia hexapetala
Gustavia tetrapetala (Aubl.) DC. =Gustavia augusta
Gustavia urceolata Poit. =Gustavia augusta
Gustavia yaracuyensis Pittier =Gustavia poeppigiana
Japarandiba augusta (L.) Kuntze =Gustavia augusta
Japarandibafastuosa (Willd.) Kuntze =Gustavia hexapetala
Japarandiba hexapetala (Aubl.) Kuntze =Gustavia hexapetala
Jugastrum christii Pittier =Eschweilera tenuifolia
Jugastrum obtectum Miers =Eschweilera tenuifolia
Jugastrum poiteaui (O. Berg) Miers =Lecythis poiteaui
Jugastrum sifontesii Pittier =Eschweilera tenuifolia
Lecythis alba S.A. Mori [nomen nudum] =Lecythis persistens subsp. persistens
Lecythis amara Aubl. =Lecythis idatimon
Lecythis bracteosa Poepp. ex O. Berg =Eschweilera bracteosa
Lecythis chaffanjonii Benoist =Eschweilera parvifolia
Checklist of the Plants of the Guiana Shield 333
Lecythis chartacea var. calyce Sagot =Lecythis chartacea
Lecythis cognata Miers =Lecythis corrugata subsp. corrugata
Lecythis congestiflora Benoist =Eschweilera congestiflora
Lecythis coriacea DC. =Eschweilera coriacea
Lecythis crassinoda Miers =Lecythis zabucajo
Lecythis davisii Sandwith =Lecythis zabucajo
Lecythis davisii var. gracilipes Eyma =Lecythis zabucajo
Lecythis fagifolia Miq. ex O. Berg =Couratari multijlora
Lecythis gracilipes Sagot =Eschweilera micrantha
Lecythis grandiflora Aubl. =Eschweilera grandiflora
Lecythisjucunda Benoist =Eschweilera sagotiana
Lecythis karuaiensis Steyerm. =Lecythis alutacea
Lecythis laevicula Miers =Lecythis chartacea
Lecythis longipes Poit. =Eschweilera pedicellata
Lecythis lutea Aubl. =Lecythis idatimon
Lecythis macrophylla O. Berg =Eschweilera pedicellata
Lecythis marawynensis O. Berg =Lecythis chartacea
Lecythis martinii O. Berg =Lecythis corrugata subsp. corrugata
Lecythis melinonis Sagot =Eschweilera sagotiana
Lecythis micrantha O. Berg =Eschweilera micrantha
Lecythis multiflora Sm. =Couratari multijlora
Lecythis odora Poepp. ex O. Berg =Eschweilera coriacea
Lecythis paniculata O. Berg =Eschweilera paniculata
Lecythis parvijlora Aubl. =Eschweilera parvijlora
Lecythis parviflora Sagot ex Miers =Eschweilera sagotiana
Lecythis pedicellata Rich. =Eschweilera pedicellata
Lecythis platycarpa Poit. =Eschweilera pedicellata
Lecythis racemiflora Sagot =Lecythis poiteaui
Lecythis retroflexa (Benoist) R. Knuth =Eschweilera coriacea
Lecythis rosea Spruce ex O. Berg =Lecythis corrugata subsp. rosea
Lecythis rubicunda Miers =Lecythis corrugata subsp. corrugata
Lecythis sagotiana (Miers) Sagot =Eschweilera sagotiana
Lecythis salebrosa O. Berg =Lecythis corrugata subsp. corrugata
Lecythis simiorum Benoist =Eschweilera simiorum
Lecythis subglandulosa Steud. ex O. Berg =Eschweilera subglandulosa
Lecythis tapuya Benoist =Eschweilera sagotiana
Lecythis tenuifolia O. Berg =Eschweilera tenuifolia
Lecythis tumefacta Miers =Lecythis zabucajo
Lecythis venusta Miers =Lecythis corrugata subsp. corrugata
Lecythis wachenheimii Benoist =Eschweilera wachenheimii
Lecythis wullschlaegeliana O. Berg =Eschweilera pedicellata
Pirigara hexapetala Aubl. =Gustavia hexapetala
Pirigara tetrapetala Aubl. =Gustavia augusta
References
(for a more extensive bibliography, see Mori & Prance 1993)
Eyma, P.J. 1934. Lecythidaceae. In A. Pulle, Flora of Suriname 3(1): 119-155.
Mori, S.A. 1999. Lecythidaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
385-397.
Mori, S.A. & collaborators. 1987. The Lecythidaceae of a lowland neotropical forest: La Fumee Mountain, French Guiana.
Mem. New York Bot. Gard. 44: 1-190.
Mori, S.A. & G.T. Prance. 1990. Lecythidaceae - part II. Flora Neotropica Monogr. 21(11): 1-373.
Mori, S.A. & G.T. Prance. 1993. Lecythidaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 12: 1-144.
Mori, S.A. & G.T. Prance. 1999. Lecythidaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 5: 750-779.
Prance, G.T. & S.A. Mori. 1979. Lecythidaceae - part I. Flora Neotropica Monogr. 21(1): 1-270.
Leguminosae-Caesalpinioideae
G.A. Aymard C, R.C. Bamebyt, RE. Berry, R.S. Cowant, M.F. da Silva, D.M. Keams, A. Sprada Tavares,
B. Stergios, D. Velazquez, R.P. Wunderlin, N. Xena, J.L. Zarucchi
Apuleia leiocarpa (Vogel) J.F. Macbr. AM BO
Barnebydendron riedelii (Tul.) J.H. Kirkbr. AM
334 Checklist of the Plants of the Guiana Shield
Batesiafloribimda Spruce ex Benth. FG
Bauhinia aciileata L. BO GU
Bauhinia bicuspidata Benth.
Bauhinia breviloba Benth.
Bauhinia cinnamomea DC.
Bauhinia conferliflora Benth.
Bauhinia coronata Benth.
Bauhinia cupreonitens Ducke
Bauhinia cupulata Benth.
Bauhinia eilertsii Pulle
Bauhinia glabra Jacq.
Bauhinia guianensis Aubl.
Bauhinia kunthiana Vogel
Bauhinia longicuspis Spruce ex Benth.
Bauhinia microstachya (Raddi) J.F. Macbr.
Bauhinia outimouta Aubl. [^Bauhinia guianensis'?]
Bauhinia poiteauana Vogel [=Bauhinia smilacina?]
Bauhinia richardiana DC.
Bauhinia nitilans Spruce ex Benth.
Bauhinia scala-simiae Sandwith
Bauhinia siqueiraei Ducke
Bauhinia smilacina (Schott) Steud.
Bauhinia surinamensis Amshoff [=Bauhinia guianensis?]
Bauhinia ungulata L.
Bocoa alterna (Benth.) R.S. Cowan
Bocoa prouacensis Aubl.
Bocoa viridiflora (Ducke) R.S. Cowan
Brownea ariza Benth.
Brownea coccinea Jacq.
Brownea coccinea subsp. capitella (Jacq.) D. Velazq. & Agostini
Brownea grandiceps Jacq.
Brownea guianensis Klotzsch
Brownea longipedicellata Huber
Brownea similis R.S. Cowan
Caesalpinia bonduc (L.) Roxb.
Caesalpinia coriaria (Jacq.) Willd.
Caesalpinia granadillo Pittier
Caesalpinia pulcherrima (L.) Sw. [cultivated; naturalized?]
Campsiandra angustifolia Spruce ex Benth.
Campsiandra angustifolia var. angustifolia
Campsiandra angustifolia var. rosea (Poepp.) Stergios
Campsiandra aymardii Stergios
Campsiandra casiquiarensis Stergios
Campsiandra chigo-montero Stergios
Campsiandra comosa var. comosa
Campsiandra comosa var. surinamensis (Kleinhoonte) Stergios
Campsiandra curaara Stergios
Campsiandra emonensis Stergios
Campsiandraferruginea Stergios
Campsiandra gomez-alvareziana Stergios
Campsiandra gomez-alvareziana var. gomez-alvareziana
Campsiandra gomez-alvareziana var. pusilliflora Stergios
Campsiandra guayanensis Stergios
Campsiandra implexicaulis Stergios
Campsiandra laurifolia Benth.
Campsiandra macrocarpa R.S. Cowan
Campsiandra macrocarpa var. alveolata Stergios
Campsiandra macrocarpa var. grandifolia Stergios
Campsiandra macrocarpa var. macrocarpa
Campsiandra nutans Stergios
Campsiandra pasibensis Stergios
Campsiandra steyermarkiana Stergios
Campsiandra taphornii Stergios
Campsiandra velutina Stergios
su
su
GU su FG
GU
FG
AM GU su FG
AM BO
BO GU su
AM BO DA GU su
AM BO DA GU su FG
AM BO DA GU su FG
AM FG
AM GU
AM BO GU su FG
FG
FG
AM BO GU?
BO DA GU
BO DA GU
su
FG
FG
DA GU su FG
AM BO GU
GU
GU su FG
GU su FG
AM
AM BO DA GU
AM BO DA GU
GU?
GU
AM BO
AM BO
GU su FG
BO
BO
GU su FG
AM BO
AM BO
AM BO
AM
AM
AM GU
GU
GU
su
su
su
AM
AM
AM
AM
AM
AM
AM BO
AM BO DA
AM GU
AM
AM
AM
AM
AM BO
AM
AM
AM BO
Checklist of the Plants of the Guiana Shield 335
Campsiandra wurdackiana Stergios
Cassia cowanii H.S. Irwin & Bameby
Cassia cowanii var. cowanii
Cassia cowanii var. guianensis (Sandwith) H.S. Irwin & Bameby
Cassiafastuosa Willd. ex Benth.
Cassiafastuosa var. fastuosa
Cassia fistula L. [widely cultivated, occasionally escaping]
Cassia grandis L. f.
Cassia moschata Kunth
Cassia spruceana Benth.
Chamaecrista adiantifolia (Spruce ex Benth.) H.S. Irwin & Bameby
Chamaecrista adiantifolia var. adiantifolia
Chamaecrista adiantifolia war. pteridophylla (Sandwith) H.S. Irwin &
Bameby [expected in SU, FG]
Chamaecrista apoucouita (Aubl.) H.S. Irwin & Bameby
Chamaecrista calycioides (DC. ex Collad.) Greene
Chamaecrista calycioides var. calycioides
Chamaecrista desvauxii (Collad.) Killip
Chamaecrista desvauxii var. brevipes (Benth.) H.S. Irwin & Bameby
Chamaecrista desvauxii var. mollissima (Benth.) H.S. Irwin & Bameby
Chamaecrista desvauxii var. piptostegia H.S. Irwin & Bameby
Chamaecrista desvauxii var. saxatilis (Amshoff) H.S. Irwin & Bameby
Chamaecrista desvauxii var. triumviralis H.S. Irwin & Bameby
Chamaecrista diphylla (L.) Greene
Chamaecristafagonioides (Vogel) H.S. Irwin & Bameby
Chamaecristafagonioides var. fagonioides
Chamaecristaflexuosa (L.) Greene
Chamaecristaflexuosa var. flexuosa
Chamaecrista glandulosa (L.) Greene
Chamaecrista glandulosa var. crystallina H.S. Irwin & Bameby
Chamaecrista glandulosa var. rapidarum H.S. Irwin & Bameby
Chamaecrista hispidula (Vahl) H.S. Irwin & Bameby
Chamaecrista hymenaeifolia (Benth.) H.S. Irwin & Bameby
Chamaecrista kunthiana (Schltdl. & Cham.) H.S. Irwin & Bameby
Chamaecrista negrensis (H.S. Irwin) H.S. Irwin & Bameby
Chamaecrista negrensis var. negrensis
Chamaecrista nictitans (L.) Moench
Chamaecrista nictitans var. disadena (Steud.) H.S. Irwin & Bameby
Chamaecrista nictitans subsp. disadena (Steud.) H.S. Irwin & Bameby
Chamaecrista nictitans var. glabrata (Vogel) H.S. Irwin & Bameby [subsp.
patellaria] [widely naturalized]
Chamaecrista nictitans var. paraguariensis (Chodat & Hassl.) H.S. Irwin &
Barneby [subsp. patellaria]
Chamaecrista nictitans var. patellaria Kartesz & Gandhi
Chamaecrista nictitans suhsp. patellaria (Collad.) H.S. Irwin & Barneby
Chamaecrista nictitans var. pilosa (Benth.) H.S. Irwin & Bameby
Chamaecrista nictitans var. praetexta (Vogel) H.S. Irwin & Barneby [subsp.
patellaria] [expected in BO]
Chamaecrista orenocensis (Spruce ex Benth.) H.S. Irwin & Barneby
Chamaecrista pilosa (L.) Greene
Chamaecrista pilosa var. pilosa
Chamaecrista polystachya (Benth.) H.S. Irwin & Barneby
Chamaecrista ramosa (Vogel) H.S. Irwin & Barneby
Chamaecrista ramosa var. ramosa
Chamaecrista ramosa var. ventuarensis (H.S. Irwin) H.S. Irwin & Barneby
Chamaecrista roraimae (Benth.) Gleason
Chamaecrista rotundifolia (Pers.) Greene [common weed expected in the
Guianas]
Chamaecrista rotundifolia var. grandiflora (Benth.) H.S. Irwin & Barneby
Chamaecrista rotundifolia var. rotundifolia
Chamaecrista serpens (L.) Greene
Chamaecrista serpens var. grandiflora (Benth.) H.S. Irwin & Barneby
Chamaecrista serpens var. serpens
Chamaecrista viscosa (Kunth) H.S. Irwin & Bameby
AM BO
BO DA GU SU FG
GU SU FG
BO DA GU SU
AM SU FG
AM
GU
SU FG
FG
BO GU SU FG
AM DA GU
FG
AM GU
AM
AM GU
GU SU FG
BO GU FG
BO
AM BO GU SU FG
AM BO GU SU? FG
AM BO
BO
GU SU?
SU
SU
FG?
FG
AM BO
AM BO GU SU FG
BO GU SU FG
BO
AM BO DA GU SU FG
AM BO DA
AM BO
AM BO
AM BO
BO GU SU FG
AM
AM BO
AM
AM
AM BO GU SU FG
BO GU SU FG
BO GU SU FG
AM GU
BO GU
BO GU SU FG
AM BO GU SU
SU
FG
GU SU FG
AM
BO
BO
BO
DA
DA
GU
AM BO GU SU FG
AM BO
AM
BO GU
AM BO GU
AM BO GU
AM BO
AM BO GU
AM BO
BO GU
BO GU SU
336 Checklist of the Plants of the Guiana Shield
Chamaecrista viscosa var. major (Benth.) H.S. Irwin & Bameby
Chamaecrista viscosa var. viscosa [expected in GU?]
Chamaecrista zygophylloides (Taub.) H.S. Irwin & Bameby
Chamaecrista zygophylloides var. deamii (Britton & Rose) H.S. Irwin &
Bameby
Copaifera camibar Poveda, N. Zamora & RE. Sanchez
Copaifera epunctata Amshoff
Copaifera guyanensis Desf.
Copaifera langsdorffii Desf.
Copaifera officinalis (Jacq.) L.
Copaifera pubiflora Benth.
Copaifera reticulata Ducke
Crudia aequalis Ducke
Crudia amazonica Spruce ex Benth.
Crudia aromatica (Aubl.) Wiild.
Crudia bracteata Benth.
Crudia glaberrima (Steud.) J.F. Macbr.
Crudia oblonga Benth.
Crudia spicata (Aubl.) Willd.
Crudia tomentosa (Aubl.) J.F. Macbr.
Cynometra bauhiniifolia Benth.
Cynometra bauhiniifolia var. bauhiniifolia
Cynometra hostmanniana Tul.
Cynometra marginata Benth.
Cynometra marginata var. guianenis Dwyer
Cynometra marginata var. marginata
Cynometra parvifolia Tul.
Cynometra spruceana Benth.
Cynometra spruceana var. procera Benth.
Cynometra spruceana var. spruceana
Delonix regia (Bojer ex Hook.) Raf. [cultivated only?]
Dialium guianense (Aubl.) Sandwith
Dicorynia guianensis Amshoff
Dicorynia paraensis Benth.
Dicorynia paraensis var. breviflora (Benth.) R.C. Koeppen
Dicorynia paraensis var. paraensis
Dicymbe altsonii Sandwith
Dicymbe bernardii R.S. Cowan
Dicymbe corymbosa Spmce ex Benth.
Dicymbe duidae R.S. Cowan
Dicymbefraterna R.S. Cowan
Dicymbe hymenaea Bameby
Dicymbejenmanii Sandwith
Dicymbe mollis Bameby
Dicymbe neblinensis R.S. Cowan
Dicymbe paruensis R.S. Cowan
Dicymbe pharangophila R.S. Cowan
Dicymbe praeruptorum Bameby
Dicymbe uaiparuensis R.S. Cowan [expected in GU]
Dicymbe yutajensis R.S. Cowan
Dimorphandra conjugata (Splitg.) Sandwith
Dimorphandra cuprea Sprague & Sandwith
Dimorphandra cuprea subsp. cuprea
Dimorphandra cuprea subsp. ferruginea (Ducke) M.F. Silva
Dimorphandra davisii Sprague & Sandwith
Dimorphandra dissimilis R.S. Cowan
Dimorphandra macrostachya subsp. congestiflora (Sprague & Sandwith)
Silva
Dimorphandra macrostachya subsp. glabrifolia (Ducke) M.F. Silva
Dimorphandra macrostachya subsp. macrostachya
Dimorphandra multiflora Ducke
Dimorphandra pennigera Tul.
Dimorphandra polyandra Benoist
Dimorphandra pullei Amshoff
BO GU SU
BO
BO GU
BO GU
AM
SU
GU SU FG
GU
BO GU?
AM BO GU
SU
AM DA
FG
GU SU FG
FG
DA GU SU FG
AM BO DA GU FG
SU FG
GU FG
AM BO GU SU FG
AM BO GU SU FG
GU SU FG
AM BO GU
GU
SU
SU
FG
GU SU
SU
FG
FG
AM BO GU
AM BO
GU
AM BO GU SU FG
AM BO GU SU FG
GU SU FG
AM FG?
AM
AM
BO
GU
GU
GU
AM
BO GU SU
BO
GU
AM
AM
AM GU
AM GU
AM
BO
AM
GU SU
AM GU
GU
AM
BO
BO
GU
GU
.¥. AM BO GU
AM BO GU FG
AM BO GU
GU
SU FG
FG
AM
AM GU SU
SU
FG
FG
GU
GU
FG?
GU su
GU
GU su
GU
GU
GU
GU su
su? FG?
GU su FG
FG
BO GU su FG
GU FG?
GU su FG
su FG
GU FG
BO GU su? FG
BO GU su? FG
BO GU
Checklist of the Plants of the Guiana Shield 337
Dimorphandra unijuga Tul. AM
Dimorphandra vernicosa Spruce ex Benth. AM
Dimorphandra williamii M.F. Silva
Dinizia excelsa Ducke [expected in FG]
EUzabetha bicolor Ducke
Elizabetha coccinea M.R. Schomb. ex Benth.
EUzabetha coccinea var. coccinea
Elizabetha coccinea var. oxyphylla (Harms) R.S. Cowan
Elizabetha durissima Ducke
Elizabethafanshawei R.S. Cowan
Elizabetha grahamiae R.S. Cowan
Elizabetha leiogyne Ducke AM
Elizabetha macrostachya Benth. AM
Elizabetha paraensis Ducke
Elizabetha princeps M.R. Schomb. ex Benth. AM
Eperua bijuga Mart, ex Benth.
Eperuafalcata Aubl. AM
Eperua glabra R.S. Cowan
Eperua grandiflora (Aubl.) Benth. [referable to misidentified E.jenmanii ssp.
sandwithii in Venezuela]
Eperua grandiflora subsp. grandiflora
Eperua grandiflora subsp. guyanensis R.S. Cowan
Eperuajenmanii Oliv.
Eperuajenmanii suhsp. jenmanii
Eperuajenmanii subsp. sandwithii R.S. Cowan
Eperua leucantha Benth. AM
Eperua obtusata R.S. Cowan AM
Eperua purpurea Benth. AM
Eperua rubiginosa Miq.
Eperua rubiginosa var. grandiflora Pulle
Eperua rubiginosa var. rubiginosa
Eperua schomburgkiana Benth.
Eperua sp. A AM
Eperua venosa R.S. Cowan [expected in GU] BO
Exostyles sp.
Haematoxylum campechianum L. [cultivated only?]
Heterostemon conjugatus Spruce ex Benth. AM
Heterostemon ellipticus Mart, ex Benth. AM
Heterostemon ingifolius Sandwith
Heterostemon mazarunensis Sandwith
Heterostemon mimosoides Desf. AM
Heterostemon mimosoides var. complanatus R.S. Cowan
Heterostemon mimosoides var. mimosoides AM
Heterostemon mimosoides \ar. pacimoniensis R.S. Cowan AM
Heterostemon otophorus Sandwith
Hymenaea courbaril L. AM BO
Hymenaea courbaril var. courbaril AM BO
Hymenaea oblongifolia Huber
Hymenaea oblongifolia var. davisii (Sandwith) Lee & Langenh.
Hymenaea oblongifolia var. oblongifolia
Hymenaea oblongifolia var. palustris (Ducke) Lee & Langenh. [expected in
SU, FG]
Hymenaea parvifolia Huber BO
Jacqueshuberia brevipes Bameby GU
Jacqueshuberia pustulata Stergios & RE. Berry AM
Jacqueshuberia splendens Stergios & RE. Berry AM
Lecointea amazonica Ducke AM
Macrolobium acaciifolium (Benth.) Benth. AM
Macrolobium acrothamnos R.S. Cowan [expected in GU]
Macrolobium amplexans (Amshoff) R.S. Cowan
Macrolobium angustifolium (Benth.) R.S. Cowan AM
Macrolobium anomalum R.S. Cowan AM
Macrolobium bifolium (Aubl.) Pers. AM
Macrolobium campestre Huber
GU SU FG
GU su
GU su FG
GU su
su
FG
GU su FG
GU
GU
GU
GU su
su
FG
GU su FG
GU
GU su FG
GU su FG
GU FG?
GU
GU FG?
GU
BO DA GU SU FG
BO
BO
DA GU SU
su
FG
BO GU su FG
BO DA GU su FG
BO GU su FG
338 Checklist of the Plants of the Guiana Shield
Macrolobiiim campestre var. arboreum R.S. Cowan GU SU FG
Macrolobium canaliciilatum Spruce ex Benth. AM
Macrolobiiim canaliculatum var. canalicitlatum AM
Macrolobium canaliculatum var. strigulosum R.S. Cowan AM
Macrolobium cataractarum R.S. Cowan AM
Macrolobium discolor Benth. AM
Macrolobium discolor var. discolor AM
Macrolobium discolor var. egranulosum R.S. Cowan AM
Macrolobium evenulosum R.S. Cowan AM
Macrolobium exfoliatum R.S. Cowan AM
Macrolobium furcatum Ducke [expected in GU]
Macrolobium gracile Spruce ex Benth. AM
Macrolobium gracile var. confertum (Gleason) R.S. Cowan AM
Macrolobium gracile var. gracile AM
Macrolobium guianense (Aubl.) Pulle
Macrolobium huberiamim Ducke
Macrolobium huberianum var. huberianum
Macrolobium huberiamim var. pubirachis Amshoff
MacrolobiumJenmanii (Gleason) Sandwith
Macrolobium limbatum Spruce ex Benth.
Macrolobium limbatum war. fanshawei (R.S. Cowan) R.S. Cowan
Macrolobium limbatum var. limbatum
Macrolobium longeracemosum Amshoff
Macrolobium longipes R.S. Cowan
Macrolobium microcalyx Ducke
Macrolobium molle (Benth.) R.S. Cowan
Macrolobium montanum Ducke
Macrolobium montanum var. potaroanum R.S. Cowan
Macrolobium multijugum (DC.) Benth.
Macrolobium multijugum var. multijugum
Macrolobium pendulum Willd. ex Vogel
Macrolobium punctatum Spruce ex Benth.
Macrolobium rubrum R.S. Cowan
Macrolobium savannarum R.S. Cowan
Macrolobium savannarum \ar.foldatsii (R.S. Cowan) RE. Berry
Macrolobium savannarum var. savannarum
Macrolobium schinifolium R.S. Cowan
Macrolobium spectabile R.S. Cowan
Macrolobium stenopetalum Amshoff
Macrolobium suaveolens Spruce ex Benth.
Macrolobium suaveolens var. pakarimense R.S. Cowan
Macrolobium suaveolens var. suaveolens
Macrolobium taxifolium Spruce ex Benth.
Macrolobium unifoliolatum R.S. Cowan
Macrolobium unifoliolatum var. unifoliolatum
Macrolobium venulosum Benth. [possibly expected in GU]
Macrolobium wurdackii R.S. Cowan
Martiodendron elatum (Ducke) Gleason
Martiodendron elatum var. elatum
Martiodendron excelsum (Benth.) Gleason
Martiodendron mediterraneum (Mart, ex Benth.) R.C. Koeppen
Martiodendron parviflorum (Amshoff) R. Koeppen
Mora excelsa Benth.
Mora gonggrijpii (Kleinhoonte) Sandwith
Paloue brasiliensis Ducke
Paloue guianensis Aubl.
Paloue induta Sandwith
Paloue riparia Pulle
Paloveopsis emarginata R.S. Cowan
Parkinsonia aculeata L. [cultivated only?]
Peltogynefloribunda (Kunth) Pittier
Peltogyne lecointei Ducke
Peltogyne paniculata Benth.
Peltogyne paniculata subsp. paniculata
SU FG
GU FG
GU FG
GU
GU
AM GU
GU
AM
BO GU SU
AM
AM
AM
GU
GU
AM BO GU SU FG
AM BO GU SU FG
GU SU FG
AM
AM
AM
AM
AM
AM
AM
AM GU
SU
AM
AM
GU
GU
AM
AM
AM
AM
AM
AM
AM
AM GU
AM?
SU FG
BO DA GU SU
BO DA GU SU
SU FG
GU SU FG
GU
GU SU FG
GU
BO SU
BO DA GU
FG?
AM BO GU SU FG
AM BO GU SU FG
AM BO
AM
AM
AM
AM BO
AM BO
AM
GU SU FG
GU su FG
GU SU FG
GU
GU su FG
BO GU SU FG
BO GU SU FG
BO GU SU FG
BO GU SU FG
BO GU SU FG
FG
GU SU FG
GU SU FG
BO DA GU
GU?
SU FG
Checklist of the Plants of the Guiana Shield 339
PeItog}-}ie paniculaTa suhsp.piibescens {Ber\Xh.)M.¥. S\\\a. GU SU FG
Peltogy-ne parvifolia Spruce ex Benth.
Peltogy-ne parvifoUa subsp. glabra M.F. Silva
Peltogy-ne parvifolia subsp. pai-vifolia
Peltogyne venosa (Vahl) Benth.
Peltogy-ne venosa subsp. densiflora (Spruce ex Benth.) M.F. Silva
Peltogy-ne venosa subsp. venosa
Peltophorum dubiiim (Spreng.) Taub.
Peltophonim pterocarpum (DC.) Backer ex K. Heyne [cultivated only?] BO
Recordoxylon pulcherrimiim Bameby AM
Recordoxylon speciosum (Benoist) Gazel ex Bameby GU FG
Schizolobium parahyba (Veil.) S.F. Blake BO
Schizolobium parahyba var. amazonicum (Huber ex Ducke) Bameby BO
Senna aculeata (Benth.) H.S. Irwin & Bameby BO
Senna alata{h.)Roxh. AM BO GU SU FG
Senna atomaria (L.) H.S. Irwin & Bameby BO
Senna bacillaris (L. f.) H.S. Irwin & Bameby AM
Senna bacillaris var. bacillaris
Senna bacillaris var. benthamiana (J.F. Macbr.) H.S. In\ in & Bameby AM
Senna bicapsularis (L.) Roxb.
Senna bicapsularis var. bicapsularis
Senna chrysocarpa (Desv.) H.S. Irwin & Bameby
Senna cobanensis (Britton) H.S. Irwin & Bameby AM
Senna georgica H.S. Irwin & Bameby
Senna hirsuta (L.) H.S. Irwin & Bameby
Senna hirsuta var. hirsuta
Senna latifolia (G. Mey.) H.S. In\in & Bameby AM
Senna ligustrina (L.) H.S. Irwin & Bameby
Senna lourteigiana H.S. Irwin & Bameby SU? FG
Senna macranthera (DC. ex Collad.) H.S. Irwin & Bameby [cultivated, BO
perhaps escaping]
Senna macranthera var. quadrifoliolata (Pittier) H.S. Irwin & Bameby
Senna macrophylla (Kunth) H.S. Irwin & Bameby AM
Senna macrophylla var. gigantifolia (Britton & Killip) H.S. Irwin & Bameby AM
Senna multijuga (Rich.) H.S. Irvvin & Bameby AM
Senna multijuga subsp. lindleyana (Gardner) H.S. Irwin & Bameby
Senna multijuga var. multijuga AM
Senna multijuga subsp. multijuga AM
Senna multijuga var. peregrinatrix H.S. Irwin & Bameby [ssp. lindleyana]
Senna obtusifolia (L.) H.S. Irwin & Bameby AM
Senna occidentalis (L.) Link AM
Senna pallida (Vahl) H.S. Irwin & Bameby [expected]
Senna pallida var. pallida [expected in GU]
Senna pendula (Humb. & Bonpl. ex Willd.) H.S. InA'in & Bameby
Senna pendula var. indecora (Kunth) Luckow
Senna pendula var. meticola H.S. Irwin & Bameby [expected in BO]
Senna pendula var. tenuifolia (Benth.) H.S. Irwin & Bameby
Senna pilifera (Vogel) H.S. Irwin & Bameby
Senna pilifera var. subglabra (S. Moore) H.S. Irwin & Bameby [expected in
the Guianas]
Senna polyphylla (Jacq.) H.S. Irwin & Bameby [cultivated, perhaps escaping]
Senna quinquangulata (Rich.) H.S. Irwin & Barneby AM
Senna quinquangulata var. quinquangulata AM
Senna reticulata (Willd.) H.S. Irwin & Bameby AM
Senna robiniifolia (Benth.) H.S. Irwin & Bameby
Senna rupununiensis H.S. Irwin & Bameby [expected in BO]
Senna sandwithiana H.S. Irwin & Bameby
Senna siamea (Lam.) H.S. Iru'in & Bameby [cuhivated, occasionally
escaping]
Senna silvestris (Veil.) H.S. Irwin & Bameby AM
Senna silvestris var. silvestris [expected in FG] AM
Senna sophera (L.) Roxb.
Senna spectabilis (DC.) H.S. Irwin & Bameby
Senna spectabilis var. spectabilis
BO
BO
BO
BO DA GU SU FG
BO
BO DA GU SU FG
BO DA GU SU FG
BO
BO DA GU SU FG
BO DA GU SU FG
BO DA GU
BO DA
GU
BO
BO
BO GU SU FG
BO
BO GU SU FG
BO
GU
BO GU
BO GU SU FG
BO GU
GU
SU
BO DA
BO DA
AM BO DA
AM BO DA
GU
AM BO GU
AM GU
AM BO
BO DA GU
BO
AM BO
AM
AM BO SU FG
SU FG
SU
SU FG
SU FG
SU FG
SU FG
FG
SU FG
340 Checklist of the Plants of the Guiana Shield
Senna spinescens (Vogel) H.S. Irwin & Bameby
Senna spinescens var. spinescens
Senna splendida (Vogel) H.S. Irwin & Bameby [native? Widely cultivated]
Senna iindiilata (Benth.) H.S. Irwin & Bameby
Senna velutina (Vogel) H.S. Irwin & Bameby
Senna wurdackii H.S. Irwin & Bameby
Tachigali albiflora (Benoist) Zarucchi & Herend. SU FG
Tachigali amplifolia (Ducke) Bameby FG
Tachigali amplifolia (Ducke) Bameby FG
Tachigali bracteolata Dwyer FG
Tachigali bracteosa (Harms) Zarucchi & Pipoly SU?
Tachigali cavipes (Spruce ex Benth.) J.F. Macbr. AM FG?
Tachigali chiysophylla (Poepp.) Zamcchi &. Herend.
Tachigali davidsei Zarucchi & Herend.
Tachigali dwyeri (R.S. Cowan) Zamcchi & Herend.
Tachigali guianensis (Benth.) Zarucchi &:UerQnd. DA GU
Tachigali melinonii (Harms) Zamcchi & Herend.
Tachigali micropetala (Ducke) Zarucchi & Pipoly GU
Tachigali myrmecophila (Ducke) Ducke
Tachigali odoratissima (Spruce ex Benth.) Zarucchi & Herend. AM
Tachigali paniciilata Auh\. AM BO DA GU
Tachigali panicidata var. angiistifolia (Miq.) Dwyer
Tachigali paniculata var. panicidata
Tachigali paniculata var. sulcata (Benoist) Dwyer
Tachigali paraensis (Huber) Bameby
Tachigali physophora (Huber) Zarucchi & Herend. AM
Tachigali pimichinensis (R.S. Cowan) Zamcchi & Herend. AM
Tachigali plwnbea Ducke AM
Tachigali ptychophysca Spruce ex Benth. [expected in Venezuelan Guayana]
Tachigali piibiflora Benth. GU
Tachigali piilch?-a Dwyer BO
Tachigali reticulosa (Dwyer) Zarucchi & Herend. AM
Tachigali rigida Ducke AM
Tachigali rusbyi Harms AM BO DA GU
Tachigali schultesiana Dwyer [expected in Venezuelan Guayana]
Tachigali tinctoria (Benth.) Zamcchi & Herend. FG
Tamarindus indica L. [cultivated] BO DA GU
Vouacapoua americana Aubl. GU? SU FG
Vouacapoua macropetala Sandwith GU
Synonyms and Misidentifications
Adipera bicapsularis (L.) Britton & Rose =Senna bicapsularis
Adipera indecora (Kunth) Britton & Rose =Senna pendula var. indecora
Adipera spiciflora Pittier =Senna bicapsularis
Amphymenium mediterraneum Mart, ex Benth. =Martiodendron mediterraneum
Apalatoa spicata Aubl. =Crudia spicata
Apoleya leiocarpa (Vogel) Gleason =Apuleia leiocarpa
Apoleya molaris (Spruce ex Benth.) Gleason =Apuleia leiocarpa
Apuleia molaris Spruce ex Benth. =Apuleia leiocarpa
Apuleia praecox Mart. =Apuleia leiocarpa
Arouna divaricata Willd. =Dialium guianense
Arouna guianensis Aubl. =Dialium guianense
Bauhinia bahiensis Bong. =Bauhinia microstachya
Bauhinia benthamiana Taub. =Bauhinia ungulata
Bauhinia brachystachya (Benth.) Walp. =Bauhinia glabra
Bauhinia bredemeyeri Vogel ^Bauhinia aculeata
Bauhinia chalkos R.S. Cowan =Bauhinia cupreonitens
Bauhinia chrysophylla Vogel =Bauhinia guianensis
Bauhinia coronata auct. non Benth.: sensu Pittier et al. =Bauhinia outimouta
Bauhinia cumanensis Kunth =Bauhinia glabra
Bauhinia emarginata Mill. =Bauhinia aculeata
Bauhinia excisa (Griseb.) Hemsl. =Bauhinia guianensis
Bauhinia glandulosa (Kunth) DC. ^Bauhinia aculeata
Bauhinia guianensis var. kunthiana (Vogel) Wunderlin =Bauhinia kunthiana
Checklist of the Plants of the Guiana Shield 341
Bauhinia guianensis var. splendens (Y^unih) A.ras\iofi=Bauhinia guianensis
Bauhinia heterophylla Kunth =Bauhinia glabra
Bauhinia langsdorffiana Bong. =Bauhinia microstachya
Bauhinia langsdorffiana var. bahiensis (Bong.) Benth. =Bauhinia microstachya
Bauhinia longipetala (Benth.) Walp. =Bauhinia glabra
Bauhinia macrostachya Benth. =Bauhinia ungulata
Bauhinia microstachya var. bahiensis (Bong.) J.F. Macbr. =Bauhinia microstachya
Bauhinia miranda Pittier =Bauhinia aculeata
Bauhinia mollicella S.F. Blake =Bauhinia aculeata
Bauhinia mollifolia Pittier =Bauhinia aculeata
Bauhinia platycalyx Benth. =Bauhinia guianensis
Bauhinia punctata Jacq. [nom. illeg.] =Bauhinia glabra
Bauhinia riparia Splitg. ex Benth. =Bauhinia outimouta
Bauhinia rotundata Mill. =Bauhinia aculeata
Bauhinia rubiginosa Bong. =Bauhinia outimouta
Bauhinia schultzei Harms =Bauhinia aculeata
Bauhinia speciosa Vogel, non Roxb. =Bauhinia outimouta
Bauhinia spicata Vogel =Bauhinia microstachya
Bauhinia splendens Kunth =Bauhinia guianensis
Bauhinia splendens var. latifolia Benth. =Bauhinia guianensis
Bauhinia sprucei var. acuminata Benth. =Bauhinia guianensis
Bauhinia superba Steud. [nom. illeg.] =Bauhinia outimouta
Bauhinia ungula Jacq. =Bauhinia aculeata
Bauhinia vogeliana Steud. =Bauhinia microstachya
Binaria cumanensis (Kunth) Raf. =Bauhinia glabra
Brownea capitella Jacq. =Brownea coccinea subsp. capitella
Brownea caurensis Pittier [nomen nudum] =Brownea similis
Brownea caurensis var. major Pittier [nomen nudum] =Brownea similis
Brownea latifolia Jacq. =Brownea coccinea subsp. capitella
Brownea racemosa Jacq. =Brownea coccinea subsp. capitella
Caesalpiniaferruginea Decne. =Peltophorum pterocarpum
Caesalpinia inermis Roxb. [nomen nudum] =Peltophorum pterocarpum
Caesalpinia parahyba (Veil.) Allemao =Schizolobium parahyba
Campsiandra comosa Benth. [Campsiandra chigo-montero from Venezuela possibly misid. as this] =Campsiandra chigo-
montero
Campsiandra comosa var. laurifolia (Benth.) Cowan =Campsiandra laurifolia
Campsiandra rosea Poepp. =Campsiandra angustifolia var. rosea
Cansenia ungulata (L.) Raf. =Bauhinia ungulata
Cassia aculeata Benth. =Senna aculeata
Cassia adiantifolia Spruce ex Benth. =Chamaecrista adiantifolia
Cassia adiantifolia var. pteridophylla (Sandwith) Ducke =Chamaecrista adiantifolia var. pteridophylla
Cassia aeschinomene Collad. =Chamaecrista nictitans var. glabrata
Cassia alata L. =Senna alata
Cassia ampliflora Steud. =Senna multijuga
Cassia apoucouita Aubl. =Chamaecrista apoucouita
Cassia arenaria Kunth =Chamaecristaflexuosa
Cassia atomaria L. ^Senna atomaria
Cassia bacillaris L. f. =Senna bacillaris
Cassia bauhiniifolia Kunth =Chamaecrista rotundifolia var. grandifJora
Cassia bicapsularis L. =Senna bicapsularis
Cassia bicapsularis var. indecora (Kunth) Benth. =Senna pendula var. indecora
Cassia bicapsularis var. tenuifolia Humb. & Bonpl. ex Willd. =Senna pendula var. tenuifolia
Cassia biflora L. [ambiguous name, probably applied to Senna pallida]
Cassia brevipes DC. ex Collad. =Chamaecrista desvauxii var. brevipes
Cassia calliantha G. Mey. =Senna multijuga
Cassia calycioides DC. ex Collad. =Chamaecrista calycioides
Cassia chrysocarpa Desv. =Senna chrysocarpa
Cassia chrysogyne Miq. =Senna chrysocarpa
Cassia cobanensis (Britton) Lundell =Senna cobanensis
Cassia cultrifolia Kunth =Chamaecrista diphylla
Cassia desvauxii Collad. =Chamaecrista desvauxii
Cassia desvauxii var. brevipes Benth. =Chamaecrista desvauxii var. brevipes
Cassia desvauxii var. mollissima Benth. =Chamaecrista desvauxii var. mollissima
Cassia diphylla L. =Chamaecrista diphylla
342 Checklist of the Plants of the Guiana Shield
Cassia disadena Steud. =Chamaecrista nictitans subsp. disadena
Cassia elliptica Kunth =Senna atomaria
Cassia emarginata auct. non L. 1753 nee Isandrina emarginata auct. non (L.) Britton & Rose ex Britton & P. Wilson 1924
=Senna atomaria
Cassiafagonioides Vogel =Chamaecristafagonioides
Cassiafilipes Benth. =Chamaecrista rotundifolia var. grandiflora
Cassiaflavicoma var. paraguariensis Chodat & Hassl. =Chamaecris1a nictitans var. paraguariensis
Cassiaflexuosa L. =Chamaecrista flexuosa
Cassiafruticosa Mill, [not in flora area] ^Sennafniticosa (Mill.) Irwin & Bameby
Cassiafruticosa var. benthamiana J.F. Macbr. =Senna bacillaris var. benthamiana
Cassia glandulosa L. =Chamaecrista glandulosa
Cassia glandulosa var. swartzii (Wikstr.) J.F. Macbr. [not in flora area] =Chamaecrista glandulosa var. swartzii
Cassia gracilis Kunth =Chamaecrista desvauxii var. triumviralis
Cassia hirsuta L. =Senna hirsuta
Cassia hispidula Vahl =Chamaecrista hispidula
Cassia hoffmannseggii Mart, ex Benth. =Senna latifolia
Cassia hymenaeifolia Benth. =Chamaecrista hymenaeifolia
Cassia indecora Kunth ^Senna pendula var. indecora
Cassia indecora var. advena Willd. ex Vogel =Senna pendula var. indecora
Cassia kunthiana Schltdl. & Cham. =Chamaecrista kunthiana
Cassia latifolia G. Mey. =Senna latifolia
Cassia leiandra Benth. [Guianan records erroneous, but possibly expected]
Cassia leiandra var. guianensis Sandwith =Cassia cowanii var. guianensis
Cassia leiantha Benth. =Chamaecrista hispidula
Cassia ligustrina L. =Senna ligustrina
Cassia lindleyana Gardner =Senna multijuga subsp. lindleyana
Cassia lotoides Kunth =Chamaecrista hispidula
Cassia lucens Vogel =Senna silvestris var. silvestris
Cassia macranthera DC. ex Collad. =Senna macranthera
Cassia macrophylla Kunth =Senna macrophylla
Cassia maritima Willd. ex Vogel =Senna pilifera var. subglabra
Cassia mimosoides L. [not in flora area] =Chamaecrista mimosoides
Cassia multijuga Rich. =Senna multijuga
Cassia negrensis H.S. Irwin =Chamaecrista negrensis
Cassia nictitans L. =Chamaecrista nictitans
Cassia obtusifolia L. =Senna obtusifolia
Cassia occidentalis L. =Senna occidentalis
Cassia orenocensis Spruce ex Benth. '=Chamaecrista orenocensis
Cassia parahyba Veil. =Schizolobium parahyba
Cassia paramariboensis Miq. (Leguminosae-Faboideae) =Aeschynomene sensitiva
Cassia patellaria Collad. =Chamaecrista nictitans subsp. patellaria
Cassia patellaria var. glabrata Vogel =Chamaecrista nictitans var. glabrata
Cassia patellaria var. longifolia Benth. =Chamaecrista nictitans subsp. patellaria
Cassia patellaria var. ramosa Vogel =Chamaecrista nictitans var. patellaria
Cassia pendula Humb. & Bonpl. ex Willd. =Senna pendula
Cassia pennelliana Amshoff [not in flora area] =Chamaecrista nictitans var. paraguariensis
Cassia pilifera Vogel =Senna pilifera
Cassia pilifera var. subglabra S. Moore =Senna pilifera var. subglabra
Cassia pilosa L. =Chamaecrista pilosa
Cassia polyphylla Jacq. =Senna polyphylla
Cassia polystachya Benth. =Chamaecrista polystachya
Cassia praetexta Vogel =Chamaecrista nictitans var. praetexta
Cassia procumbens Mill. =Chamaecrista hispidula
Cassia prostrata Humb. & Bonpl. ex Willd. =Chamaecrista serpens
Cassia pteridophylla Sandwith =Chamaecrista adiantifolia var. pteridophylla
Cassia pulchra Kunth =Chamaecrista desvauxii var. mollissima
Cassia quinquangulata Rich. =Senna quinquangulata
Cassia racemosa Mill, [not in flora area] =Senna racemosa (Mill.) Irwin & Bameby
Cassia ramosa Vogel =Chamaecrista ramosa
Cassia reticulata Willd. =Senna reticulata
Cassia richardiana Kunth =Senna multijuga
Cassia robiniifolia Benth. =Senna robiniifolia
Cassia roraimae Benth. =Chamaecrista roraimae
Cassia rotundifolia Pers. =Chamaecrista rotundifolia
Checklist of the Plants of the Guiana Shield 343
Cassia rotimdifolia var. bauhiniifolia (Kunth) Benth. =Chamaecrista rotundifolia var. grandiflora
Cassia rotundifolia var. grandiflora Benth. =Chamaecrista rotundifolia var. grandiflora
Cassia sagotiana Benth. =Cassia spruceana
Cassia savannarum Miq. =Chamaecrista ramosa
Cassia saxatilis (Amshoff) H.S. Irwin =Chamaecrista desvauxii var. saxatilis
Cassia serpens L. =Chamaecrista serpens
Cassia serpens var. grandiflora Benth. =Chamaecrista serpens var. grandiflora
Cassia siamea Lam. =Senna siamea
Cassia silvestris Veil. =Senna silvestris
Cassia simplex Standi. =Chamaecrista nictitans var. paraguariensis
Cassia spectabilis DC. =Senna spectabilis
Cassia spinescens Vogel =Senna spinescens
Cassia stenocarpa Vogel =Chamaecrista nictitans var. disadena
Cassia tagera authors., auct. non L. 1753: sensu American =Chamaecrista kunthiana
Cassia tarantan Kunth =Senna reticulata
Cassia tenella Kunth =Chamaecrista serpens var. grandiflora
Cassia tetraphylla Martyn =Chamaecrista hispidula
Cassia tetraphylla Desv. =Chamaecrista desvauxii
Cassia tetraphylla var. aurivilla H.S. Irwin =Chamaecrista desvauxii var. brevipes
Cassia tetraphylla var. mollissima (Benth.) H.S. Irwin =Chamaecrista desvauxii var. mollissima
Cassia tetraphylla var. ramosa (Vogel) Amshofi =Chamaecrista ramosa
Cassia tetraphylla var. ventuarensis H.S. Irwin =Chamaecrista ramosa var. ventuarensis
Cassia tora authors, auct. non L. 1762: sensu American nee Emelista tora auct. non (L.) Britton & Rose 1924: sensu
American authors =Senna obtusifolia
Cassia tora L. [not in flora area]
Cassia undulata Benth. =Senna undulata
Cassia uniflora Spreng. [dubious name, not in Guianas]
Cassia uniflora var. ramosa (Vogel) Benth. =Chamaecrista ramosa
Cassia velutina Vogel =Senna velutina
Cassia viminea L. [not in flora area] =Senna viminea (L.) Irwin & Bameby
Cassia virgata Rich. ^Senna chrysocarpa
Cassia viscosa Kunth =Chamaecrista viscosa
Cassia viscosa var. major Benth. =Chamaecrista viscosa var. major
Cassia viscosopilosa Steud. =Chamaecrista viscosa var. major
Cassia zygophylloides Taub. =Chamaecrista zygophylloides
Cassia zygophylloides var. deamii (Britton & Rose) H.S. Irwin & Bameby =Chamaecrista zygophylloides var. deamii
Cathartocarpus moschatus (Kunth) G. Don =Cassia moschata
Caulotretus heterophyllus (Kunth) Warb. =Bauhinia glabra
Chamaecrista aeschinomene (Collad.) Greene =Chamaecrista nictitans var. glabrata
Chamaecrista bauhiniifolia (Kunth) Gleason =Chamaecrista rotundifolia var. grandiflora
Chamaecrista brevipes (DC. ex Collad.) Greene =Chamaecrista desvauxii var. brevipes
Chamaecrista cultrifolia (Kunth) Britton & Rose ex Britton & Killip =Chamaecrista diphylla
Chamaecrista gracilis (Kunth) Pittier ^Chamaecrista desvauxii var. triumviralis
Chamaecrista lucesiae Pittier =Chamaecrista nictitans var. praetexta
Chamaecrista nictitans var. ramosa (Vogel) H.S. Irwin & Bameby =Chamaecrista nictitans var. patellaria
Chamaecrista patellaria (Collad.) Greene =Chamaecrista nictitans subsp. patellaria
Chamaecrista pulchra (Kunth) Britton & Rose =Chamaecrista desvauxii var. mollissima
Chamaecrista ramulosa Killip & Pittier =Chamaecrista ramosa
Chamaecrista simplex Standi. ^Chamaecrista nictitans var. paraguariensis
Chamaecrista tenella (Kunth) Pittier et al. =Chamaecrista serpens var. grandiflora
Chamaefistula barbinervis Pittier =Senna macrophylla var. gigantifolia
Chamaefistulafruticosa auct. non Cassiafruticosa Mill. 1768. =Senna bacillaris
Chamaefistula gigantifolia Britton & Killip =^Senna macrophylla var. gigantifolia
Chamaefistula indecora (Kunth) G. Don =Senna pendula var. indecora
Chamaefistula quadrifoliolata Pittier =Senna macranthera var. quadrifoliolata
Chamaefistula quinquangulata (Rich.) Pittier =Senna quinquangulata
Chamaefistula undulata (Benth.) Pittier =Senna undulata
Chamaesenna aculeata (Benth.) Pittier =Senna aculeata
Copaiba officinalis Adans. =Copaifera officinalis
Copaifera jacquinii Desf. =Copaifera officinalis
Copaifera martii war. pubifiora (Benth.) Benth. =Copaifera pubifiora
Copaiva officinalis Jacq. =Copaifera officinalis
Crudia obliqua Griseb. =Crudia glaberrima
Crudia parivoa DC. =Crudia tomentosa
344 Checklist of the Plants of the Guiana Shield
Crudia pubescens Spruce ex Benth. =Crudia oblonga
Cynometra bauhiniifolia var. grandiflora Dwyer =Cynometra bauhiniifolia var. bauhiniifolia
Cynometra marginata var. giiianensis Ducke =Cynometra marginata
Cynometra marginata var. Jaevis Ducke ^Cynometra marginata
Cynometra microflora R.S. Cowan =Cynometra bauhiniifolia var. bauhiniifolia
Dialium divaricatum (Willd.) Vahl =Dialium guianense
Dicorynia breviflora Benth. =Dicorynia paraensis var. breviflora
Dicorynia spruceana Benth. =Dicorynia paraensis var. breviflora
Dimorphafalcata (Aubl.) Sm. =Eperua falcata
Dimorphandra congestiflora Sprague & Sandwith =Dimorphandra macrostachya subsp. congestiflora
Dimorphandra excelsa (Benth.) Baill. =Mora excelsa
Dimorphandraferruginea Ducke ^Dimorphandra cuprea suhsp. ferruginea
Dimorphandra glabrifolia Ducke =Dimorphandra macrostachya subsp. glabrifolia
Dimorphandra gonggrijpii Kleinhoonte =Mora gonggrijpii
Dimorphandra guianensis (R.H. Schomb. ex Benth. & Hook, f.) Baill. =Mora excelsa
Dimorphandra hohenkerkii Sprague & Safndwith =Dimorphandra polyandra
Dimorphandra lat[folia Tul. =Dimorphandra conjugata
Dimorphandra macrostachya auct. non Benth.: sensu Ducke =Dimorphandra macrostachya subsp. glabrifolia
Dimorphandra mora Benth. & Hook. f. =Mora excelsa
Ditremexa occidentalis (L.) Britton & Rose =Senna occidentalis
Echinocassia aculeata (Benth.) Britton & Rose =Senna aculeata
Elizabetha oxyphylla Harms =Elizabetha coccinea var. oxyphylla
Eperua kouroiiensis Benoist =Epenia grandiflora
Eperita stipulata Kleinhoonte =Eperuajenmanii subsp. sandwithii
Grimaldia baileyonim Britton & Rose ex L.H. Bailey ^Chamaecristafagonioides
Grimaldia deamii Britton & Rose =Chamaecrista zygophylloides var. deamii
Grimaldia orenocensis (Spruce ex Benth.) Pittier =Chamaecrista orenocensis
Hermesias ariza (Benth.) Kuntze =Brownea ariza
Hermesias capitella (Jacq.) Kuntze =Brownea coccinea subsp. capitella
Hermesias latifolia (Jacq.) Kuntze =Bro\\'nea coccinea subsp. capitella
Hermesias racemosa (Jacq.) Kuntze =BroM'nea coccinea subsp. capitella
Heterostemon cauliflorus Pittier [nomen nudum] =Heterostemon conjugatus
Hymenaea courbaril var. obtusifolia Ducke =Hymenaea courbaril
Hymenaea davisii Sandwith =Hymenaea oblongifolia var. davisii
Hymenaeafloribunda Kunth =Peltogyne floribiinda
Hymenaea latifolia Hayne =Peltogynefloribunda
Hymenaea multiflora Kleinhoonte =Hymenaea courbaril
Hymenaea palustris Ducke =Hymenaea oblongifolia var. palustris
Hymenaea resinifera Salisb. =Hymenaea courbaril
Hymenaea venosa Vahl =Peltogyne venosa
Leptolobium leiocarpum Vogel =Apuleia leiocarpa
Libidibia granadillo (Pittier) Pittier =Caesalpinia granadillo
Macrolobium acaciifolium var. vestitum Sandwith =Macrolobium acaciifolium
Macrolobium bifolium var. amplexans Amshoff =Macrolobium amplexans
Macrolobium chrysostachyum (Miq.) Benth. =Macrolobium angustifolium
Macrolobium confertum Gleason =Macrolobium gracile var. confertum
Macrolobium elegans Miq. =Macrolobium bifolium
Macrolobium flexuosum var. molle Benth. ^Macrolobium molle
Macrolobium foldatsii R.S. Cowan =Macrolobium savannarum war. foldatsii
Macrolobium hymenifolium Pittier =Macrolobium angustifolium
Macrolobium hymenioides Willd. =Macrolobium bifolium
Macrolobium simira (Aubl.) J.F. Gmel. (incertae sedis) =Vouapa simira
Macrolobium stamineum G. Mey. =Macrolobium bifolium
Macrolobium tenue Ducke =Macrolobium gracile var. gracile
Macrolobium unijugum (Poepp. & Endl.) R.S. Cowan =Macrolobium limbatum
Macrolobium unijugum war.fanshawei R.S. Cowan =Macrolobium limbatum var. fanshawei
Martia excelsa (Benth.) Benth. =Martiodendron excelsum
Martia parvifolia (Benth.) Benth. ^Martiodendron mediterraneum
Martiodendron pai-vifolium (Benth.) Gleason =Martiodendron mediterraneum
Martiusia elata Ducke =Martiodendron elatum
Martiusia excelsa Benth. =Martiodendron excelsum
Martiusia parviflora Amshoff =Martiodendron parviflorum
Martiusia parvifolia Benth. ^Martiodendron mediterraneum
Melanoxylon speciosum Benoist =Recordoxylon speciosum
Checklist of the Plants of the Guiana Shield 345
Mora giiianensis R.H. Schomb. ex Benth. & Hook. f. =Mora excelsa
Oiitea acaciifolia Benth. =Macrolobium acaciifolium
Outea guianertsis Aubl. =Macrolobium guianense
Outea multijuga DC. =Macrolobium multijugum
Panzerafalcata (Aubl.) Willd. =Eperuafalcata
Parivoa grandiflora Aubl. =Eperiia grandiflora
Parivoa tomentosa Aubl. =Cnidia tomentosa
Paidetia glandidosa Kunth =Bauhinia acideata
Paidetia ungulata (L.) A. Schmitz =Bauhinia ungulata
Peiranisia macwchlamys Pittier =Senna robiniifolia
Peiranisia robiniifolia (Benth.) Britton & Rose =Senna robiniifolia
Peltogyne amplissima Pittier =Peltogyne paniculata subsp. pubescens
Peltogyne densiflora Spruce ex Benth. =Peltogyne venosa subsp. densiflora
Peltogyne micrantha Ducke =Peltogyne parvifolia subsp. parvifolia
Peltogyne paraensis Huber =Peltogyne venosa subsp. densiflora
Peltogyne porphyrocardia Griseb. ex Benth. [nomen nudum] =Peltogynefloribunda
Peltogyne pubescens Benth. =Peltogyne paniculata subsp. pubescens
Peltogyne venosa var. densiflora (Spruce ex Benth.) Amshoff =Pe//ogy«e venosa subsp. densiflora
Peltophorum ferrugineum (Decne.) Benth. =Peltophorum pterocarpum
Peltophorum inerme (Roxb.) Naves ex Fem.-Vill. =Peltophorum pterocarpum
Phyllocarpus riedelii Tul.
Poinciana coriaria Jacq. =Caesalpinia coriaria
Poinciana pulcherrima L. =Caesalpinia pulcherrima
Poinciana regia Bojer ex Hook. =Delonix regia
Pseudocassia spectabilis (DC.) Britton & Rose =Senna spectabilis
Recordoxylon amazonicum (Ducke) Ducke =Recordoxylon speciosum
Schizolobium amazonicum Huber ex Ducke =Schizolobium parahyba var. amazonicum
Schizolobium excelsum Vogel =Schizolobium parahyba
Schnella bicomata Pittier =Bauhinia guianensis
Schnella brachystachya Benth. =Bauhinia glabra
Schnella cumanensis (Kunth) Britton & Rose =Bauhinia glabra
Schnella excisa Griseb. =Bauhinia guianensis
Schnella glabra (Jacq.) Dugand =Bauhinia glabra
Schnella heterophylla (Kunth) Benth. =Bauhinia glabra
Schnella longipetala Benth. =Bauhinia glabra
Schnella microstachya Raddi =Bauhinia microstachya
Schnella rubiginosa (Bong.) Benth. =Bauhinia outimouta
Schnella splendens (Kunth) Benth. =Bauhinia guianensis
Sclerolobium albiflorum Benoist =Tachigali albiflora
Sclerolobium amplifolium Ducke
Sclerolobium aureum var. grandiflorum Dwyer =Tachigali davidsei
Sclerolobium bracteosum Harms =Tachigali bracteosa
Sclerolobium chrysophyllum Poepp. =Tachigali chrysophylla
Sclerolobium dwyeri R.S. Cowan =Tachigali dwyeri
Sclerolobium guianense Benth. =Tachigali guianensis
Sclerolobium guianense var. guianense =Tachigali guianensis
Sclerolobium melinonii Harms =Tachigali melinonii
Sclerolobium micropetalum Ducke =Tachigali micropetala
Sclerolobium odoratissimum var. latifolium Dwyer =Tachigali odoratissima
Sclerolobium paraensis Huber =Tachigali paraensis
Sclerolobium physophorum Huber =Tachigali physophora
Sclerolobium pimichinense R.S. Cowan =Tachigali pimichinensis
Sclerolobium reticulosum Dwyer =Tachigali reticulosa
Sclerolobium roraimae R.H. Schomb. [nomen nudum? Mt. Roraima]
Sclerolobium tinctorium Benth. =Tachigali tinctoria
Senna lucens Vogel =Senna silvestris
Senna pendula var. advena (Willd.) H.S. Irwin & Bameby =Senna pendula var. indecora
Senna spruceana Benth. =Cassia spruceana
Tachia paniculata (Aubl.) Pers. [orthographic variant] =Tachigali paniculata
Tachigali carinata Gleason =Tachigali paniculata
Tachigali glauca Tul. =Tachigali cavipes
Tachigali paniculata var. cavipes Benth. =Tachigali cavipes
Tachigali paniculata var. cavipes Spruce ex Benth. ^Tachigali cavipes
Tachigali purpurea Rich. (Leguminosae-Faboideae) =Diplotropis purpurea
346 Checklist of the Plants of the Guiana Shield
Tachigali richardiana Tul. ex Lemee [apparently a nomen nudum]
Tachigali sulcata Benoist =Tachigali paniculata var. sulcata
Tachigali trigona Aubl. =Tachigali paniculata
Touchiroa aromatica Aubl. =Crudia aromatica
Vogelocassia cobanensis Britton =Senna cobanensis
Vouapa acaciifolia (Benth.) Baill. =Macrolobium acaciifolium
Vouapa angustifolia Benth. =Macrolobium angustifolium
Vouapa bifolia Aubl. =Macrolobium bifolium
Vouapa chrysostachya Miq. =Macrolobium angustifolium
Vouapa discolor (Benth.) Taub. =Macrolobium discolor
Vouapa gracile (Spruce) Taub. =Macrolobium gracile
Vouapa guianensis (Aubl.) Taub. =Macrolobium guianense
Vouapajenmani Gleason =Macrolobiumjenmanii
Vouapa limbata (Spruce) Taub. =Macrolobium limbatum
Vouapa multijuga (DC.) Taub. =Macrolobium multijugum
Vouapa punctata (Spruce) Taub. ^^Macroldbium punctatum
Vouapa staminea (G. Mey.) DC. =Macrolobium bifolium
Vouapa suaveolens (Spruce) Taub. =Macrolobium suaveolens
Vouapa taxifolia (Spruce) Taub. =Macrolobium taxifolium
References
Amshoff, GJ.H. 1939. Papilionaceae. In A. Pulle, Flora of Suriname 2(2): 1-257.
Aymard C. 1998. Caesalpiniaceae. In J.A. Steyermark, RE. Berry & B.K. Hoist, Flora of the Venezuelan Guayana 4: 1-121.
Bameby, R.C. & S.V. Heald. 2002. Caesalpiniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff,
& J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York
Bot. Gard. 76 (2): 167-183.
Bameby, R.C, B. Stergios, R.S. Cowan, RE. Berry, J.L. Zarucchi, R.R Wunderlin, D.M. Keams, M.R da Silva, A. Sprada
Tavares, Velazquez, N. Xena, & G.A. Cowan, R.S. 1961. Leguminosae. In B. Maguire, J.J. Wurdack, & collaborators.
Botany of the Guayana Highland part IV. Mem. New York Bot. Gard. 10(4): 65-87.
Cowan, R.S. 1968. Swartzia. Flora Neotropica Monogr. 1: 1-228.
Cowan, R.S. & J.C. Lindeman. 1989. Caesalpiniaceae p.p. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 7: 1-167.
Dwyer, J.D. 1951. The Central American, West Indian, and South American species of Copaifera (Caesalpiniaceae). Brittonia
7(3): 143-172.
Dwyer, J.D. 1954. The tropical American genus Tachigalia Aubl. (Caesalpiniaceae). Ann. Missouri Bot. Gard. 41: 223-260.
Dwyer, J.D. 1957. The tropical American genus Sclerolobium Vogel (Caesalpiniaceae). Lloydia 20: 67-118, suppl. 266-267.
Dwyer, J.D. 1958. The New World species of Cynometra. Ann. Missouri Bot. Gard. 45: 313-345.
Fonseca Vaz, A.M.S. da. 1979. Considera9oes sobre a taxonomia do genero Bauhinia L. sect. Tylotaea Vogel (Leguminosae -
Caesalpinioideae) do Brasil. Rodriguesia 31(51): 127-234.
Gorts-van Rijn, A.R.A., K.U. Kramer, & J.C. Lindeman. 1976. Papilionaceae (additions and corrections). In J. Lanjouw & A.L.
Stoffers, Flora of Suriname 2(2): 556-610.
Irwin, H.S. & R.C. Bameby. 1982. The American Cassiinae. A synoptical revision of Leguminosae tribe Cassieae subtribe
Cassiinae in the New World. Mem. New York Bot. Gard. 35: 1-918.
Koeppen, R.C. 1967. Revision of Dicorynia. Brittonia 19(1): 42-61.
Lee, YT. & J.H. Langenheim. 1975. Systematics of the genus Hymenaea L. Univ. Calif. Publ. Bot. 69: 1-109.
Polhill, R.M. & P.H. Raven (ed.). 1981. Advances in Legume Systematics. Royal Botanical Garden, Kew, England.
Silva, M.F. da. 1976. Revisao taxonomica do genero Peltogyne Vog. Acta Amaz. 6(1) suppl.: 1-61.
Silva, M.F. da. 1986. Dimorphandra. Flora Neotropica Monogr. 44: 1-127.
Stergios, B. 1996. Contributions to South American Caesalpinaceae. II. A taxonomic Update of Campsiandra (Caesalpinaceae).
Novon 6(4): 434-459.
Velasquez, D. 1981. Revision Taxonomica del Genero Brownea Jacq. (Leguminosae-Caesalpinioideae) para Venezuela. Thesis-
Univ. Central de Venezuela.
Leguminosae-Faboideae
G.A. Aymard C, RE. Berry, R.S. Cowanf, Nidia L. Cuello A., A. Delgado Salinas, PR. Fantz, R.H. Maxwell, K.M.
Redden,V.E. Ruddt, M. Sousa, D.R. Wind
Abrus fruticulosus Wall, ex Wight & Am. [introduced]
Abrus precatorius L. [cultivated, naturalized]
Abrus precatorius subsp. africanus Verde.
Abrus pulchellus Wall, ex Thwaites
Abrus pulchellus subsp. tenuiflorus (Benth.) Verde.
Acosmium nitens (Vogel) Yakovlev
Acosmium praeclarum (Sandwith) Yakovlev
SU
BO DA GU su
BO DA GU
BO
BO
BO GU
GU
SU
FG
FG
Checklist of the Plants of the Guiana Shield 347
Acosmium sp. A
Aeschynomene americana L.
Aeschynomene americana var. americana
Aeschynomene brasiliana (Poir.) DC.
Aeschynomene brasiliana var. brasiliana
Aeschynomene brasiliana var. carichanica Rudd
Aeschynomene evenia C. Wright
Aeschynomene evenia var. serrulata Rudd
Aeschynomenefilosa Mart, ex Benth.
Aeschynomenefluminensis Veil.
Aeschynomenefluminensis var. fluminensis
Aeschynomene histrix Poir.
Aeschynomene histrix var. densiflora (Benth.) Rudd
Aeschynomene histrix var. histrix
Aeschynomene interrupta Benth.
Aeschynomene paniculata Willd. ex Vogel
Aeschynomene pratensis Small
Aeschynomene pratensis var. caribaea Rudd
Aeschynomene riidis Benth.
Aeschynomene scabra G. Don
Aeschynomene sensitiva Sw.
Aeschynomene sensitiva var. sensitiva
Aid
Ald^
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
Aid
na aurea R.S. Cowan
na berryi R.S. Cowan & Steyerm.
na discolor Spruce ex Benth.
na elliptica R.S. Cowan
na heterophylla Spruce ex Benth.
na insignis (Benth.) Endl.
na insignis var. insignis
na insignis var. retusa R.S. Cowan
na kunhardtiana R.S. Cowan
na latifolia Spruce ex Benth.
na latifolia var. auyantepuiensis Pittier ex H.S. Irwin
na latifolia var. latifolia [expected in GU]
na latifolia var. pubescens R.S. Cowan [expected in GU]
na macrophylla Spruce ex Benth.
na macrophylla var. macrophylla
na macrophylla yav. yapacanensis (R.S. Cowan) Stergios
na paulberryi Aymard
na petiolulata R.S. Cowan
na reticulata R.S. Cowan
na sp. A
na sp.
na sp.
na sp.
na sp.
na sp.
na sp.
Alexa bauhiniiflora Ducke
Alexa canaracunensis Pittier
Alexa confusa Pittier
Alexa cowanii Yakovlev
Alexa grandiflora Ducke
Alexa herminiana N. Ramirez
Alexa imperatricis (R.H. Schomb.) Baill.
Alexa surinamensis Yakovlev
Alexa wachenheimii Benoist
Alysicarpus vaginalis (L.) DC. [introduced]
Andira coriacea Pulle
Andira grandistipula Amshoff
Andira inermis (W. Wright) DC.
Andira inermis subsp. inermis
Andira sp. A
Andira surinamensis (Bondt) Splitg. ex Amshoff
AM BO GU
BO DA GU FG
BO DA
AM BO GU SU FG
AM BO
BO
AM BO DA GU?
AM BO DA
GU
BO GU SU FG
BO
AM BO DA GU
GU
SU FG
AM BO DA GU
GU
SU FG
AM BO GU SU
BO GU FG
BO GU FG
BO GU
BO
BO DA GU SU FG
BO DA
AM
BO
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
GU
GU
GU
GU?
AM BO
BO
GU
GU
pn'
AM
rU
BO DA GU
GU
SU?
SU
GU SU FG
BO DA GU SU FG
GU SU FG
GU
BO DA GU SU FG
BO DA GU SU FG
AM
AM BO DA GU SU FG
348 Checklist of the Plants of the Guiana Shield
Andira fervequinata R.T. Penn., Aymard & Cuello BO
Andira thfoliolata Ducke AM
Andira villosa Kleinhoonte SU
Barbieha pinnata (Pers.) Baill. BO
Bowdichia virgilioides Kunth AM BO GU SU
Cajamts cajan (L.) Millsp. [cultivated, naturalized] BO DA GU SU FG
Calopogoniiim caeruleum (Benth.) C. Wright AM BO DA GU SU FG
Calopogonhim mucitnoides Desv. AM BO DA GU SU FG
Calopogoniiim vehttinum (Benth.) Amshoff GU SU FG
Canavalia brasiliensis Mart, ex Benth. BO GU SU FG
Canavalia dictyota Piper AM BO DA GU SU FG
Canavalia dictyota var. dictyota GU SU FG
Canavalia dictyota war. ferruginea Aymard & Cuello SU
Canavalia ensiformis (L.) DC. [escaped from cultivation] GU? SU FG
Canavalia gladiata (Jacq.) DC. SU
Canavalia gladiata var. gladiata [escaped'from cultivation] SU
Canavalia grandiflora Benth. AM BO GU FG
Canavalia plagiospenna Piper [probably escaped from cultivation] GU SU
Canavalia rosea (Sw.) DC. DA GU SU FG
Canavalia sericophylla Ducke AM
CandoUeodendron brachystachyum (DC.) R.S. Cowan SU FG
Centrolobium paraenseJ\x\. BO GU FG
Centrolobium paraense var. orinocense Benth. BO
Centrolobiitm paraense \ar. paraense BO GU
Centrosema angiistifolium (Kunth) Benth. non (Kunth) Benth. in Mart. AM BO GU SU
Centrosema brasilianum {L.)Ber\th. AM BO DA GU SU FG
Centrosema capitatum (Rich.) Amshoff GU SU FG
Centrosema macrocarpiim Benth. AM BO GU
Centrosema molle Mart, ex Benth. [specimens previously misidentified as AM BO DA GU SU FG
Centrosema pubescens]
Centrosema pasciioriim Mart, ex Benth. BO GU
Centrosema plitmieriiTurpin ex Pers.) Benth. AM BO DA GU SU? FG?
Centrosema tetragonolobum Schultze-Kraft & R.J. Williams AM BO
Centrosema triqiietriim (Hoffmanns, ex Benth.) Benth. AM DA GU
Centrosema venosiim Mart, ex Benth. AM BO
Centrosema vexillatiim Benth.
Centrosema virginianum (L.) Benth. AM BO
Chaetocalyx brasiliensis (Vogel) Benth.
Chaetocalyx scandens (L.) Urb,
Chaetocalyx scandens var. pubescens (DC.) Rudd
Clathrotropis brachypetala (Tul.) Kleinhoonte AM BO DA
Clathrotropis brimnea Amshoff
Clathrotropis glaiicophylla R.S. Cowan
Clathrotropis macrocarpa Ducke
Clathrotropis nitida (Benth.) Harms
Clathrotropis paradoxa Sandwith
Clitoria arborescens R. Br. BO DA
Clitoria brachycalyx Harms
Clitoria canescens Pittier ex Fantz AM
Clitoria cavalcantei Fantz BO
Clitoria coriacea Schery
Clitoria dendrina Pittier
Clitoriafalcata Lam. AM BO GU SU FG
Clitoriafalcata war. falcata AM BO GU SU FG
Clitoria guianensis {Anhl.) Benth. AM BO GU SU FG
Clitoria guianensis var. guianensis AM BO GU SU FG
Clitoria javitensis (Kunth) Benth.
Clitoriajavitensis var. grandifolia (Ducke) Fantz
Clitoria javitensis var. javitensis
Clitoria kaieteiirensis Fantz GU
Clitoria laiirifolia Voir. AM BO DA GU SU FG
Clitoria leptostachya Benth. GU
Clitoria leptostachya var. leptostachya GU
Clitoria pendens Fantz SU
BO
BO
AM BO
AM BO
AM BO
GU SU
GU? SU FG?
SU FG?
GU SU FG
GU SU
GU?
GU
GU
GU SU FG
GU
BO
BO
BO
AM BO GU SU FG
BO GU
GU
SU
FG
AM
AM
BO
SU
SU
FG
FG
AM BO DA GU
GU FG
BO DA GU
GU SU
AM BO GU
AM BO
AM BO
AM BO GU SU FG
AM GU
GU
AM BO GU
GU
SU FG
BO DA GU SU FG
AM GU
Checklist of the Plants of the Guiana Shield 349
Clitoha sagotii Fantz
Clitoria sagotii var. canaliculata Fantz
Clitoha sagotii var. sagotii
Clitoria sagotii var. sprucei Fantz
Clitoria simplicifolia (Kunth) Benth.
Clitoria steyermarkii Fantz
Clitoria ternatea L.
Clitoria ternatea var. ternatea [cultivated or escaped from cultivation]
Coursetiaferruginea (Kunth) Lavin
Crotalaria goreensis Guill. & Perr. [introduced]
Crotalaria incana L.
Crotalariajuncea Willd. [introduced]
Crotalaria maypurensis Kunth
Crotalaria maypurensis var. depauperata (Mart.) Windier & S.G. Skinner
Crotalaria maypurensis var. maypurensis
Crotalaria micans Link
Crotalaria nitens Kunth
Crotalaria pallida Alton [introduced]
Crotalaria pilosa Mill, non Crotalaria pilosa Thunb. 1800, nee Crotalaria
pilosa Roxb. ex Mart. 1820.
Crotalaria quinquefolia L. [introduced]
Crotalaria retusa L.
Crotalaria sagittalis L.
Crotalaria spectabilis Roth
Crotalaria stipularia Desv.
Crotalaria ajf. velutina Benth.
Crotalaria verrucosa L. [introduced]
Crotalaria zanzibarica Benth. [introduced]
Cymbosema roseum Benth. [expected in the Guianas]
Dalbergia amazonica (Radlk. ex Kopff) Ducke
Dalbergia ecastaphyllum (R Browne ex L.) Taub.
Dalbergiafoliosa (Benth.) A.M. Carvalho
Dalbergiafrutescens (Veil.) Britton
Dalbergiafrutescens var. frutescens
Dalbergia hygrophila (Mart, ex Benth.) Hoehne
Dalbergia intermedia A.M. Carvalho
Dalbergia inundata Spruce ex Benth.
Dalbergia monetaria L. f.
Dalbergia riedelii (Benth.) Sandwith
Dalbergia sp. A
Dalbergia sp. B
Dalbergia sp. C
Dalbergia sp. D
Dalbergia spruceana Benth.
Dalbergia subcymosa Ducke
Derris amazonica Killip
Derris negrensis Benth.
Derris pterocarpus (DC.) Killip
Desmodium adscendens (Sw.) DC.
Desmodium affine Schltdl.
Desmodium axillare (Sw.) DC.
Desmodium axillare var. acutifolium (Kunze) Urb.
Desmodium axillare var. stoloniferum (Rich, ex Poir.) B.G. Schub.
Desmodium barbatum (L.) Benth.
Desmodium cajanifolium (Kunth) DC.
Desmodium campyloclados Hemsl.
Desmodium distortum (Aubl.) J.F. Macbr.
Desmodium guianense DC.
Desmodium hickenianum Burkart
Desmodium incanum DC.
Desmodium intortum (Mill.) Urb. GU?
Desmodium molliculum (Kunth) DC. GU
Desmodium orinocense (DC.) Cuello AM
Desmodium pachyrrhizum Wogel BO
BO
BO GU SU FG
BO
GU
GU
SU
AM BO DA
AM BO GU FG
AM BO DA GU SU FG
AM BO
BO
BO
GU
GU
GU
SU FG
AM BO DA
AM BO GU
AM GU
AM BO DA GU SU FG
AM BO
BO
BO
GU SU FG
AM
AM
AM BO
DA SU FG
AM GU SU FG
AM BO DA
BO GU SU FG
AM BO
BO
DA
DA
GU SU FG
AM BO GU SU FG
AM BO GU SU FG
BO GU SU FG
AM BO DA GU SU FG
AM BO GU SU
AM
AM BO
BO
GU SU FG
FG
AM BO DA GU SU FG
350 Checklist of the Plants of the Guiana Shield
Desmodiiim procumbens (Mill.) Hitchc.
Desmodium procumbens var. procumbens
Desmodium sclerophyllum Benth.
Desmodium scorpiurus (Sw.) Desv.
Desmodium tortuosum (Sw.) DC. [introduced]
Desmodium triflorum (L.) DC. [introduced]
Desmodium velutinum DC.
Desmodium wydlerianum Urb.
Dioclea albiflora R.S. Cowan
Dioclea apurensis Kunth
Dioclea coriacea Benth.
Dioclea guianensis Benth.
Dioclea holtiana Pittier ex R.H. Maxwell
Dioclea huberi Ducke
Dioclea macrantha Ruber
Dioclea macrocarpa Huber
Dioclea malacocarpa Ducke
Dioclea megacarpa Rolfe
Dioclea reflexa Hook. f.
Dioclea rigida R.S. Cowan
Dioclea ruddiae R.H. Maxwell
Dioclea scabra (Rich.) R.H. Maxwell
Dioclea scabra var. brownii R.H. Maxwell
Dioclea scabra var. scabra
Dioclea scabra var. schulzii R.H. Maxwell
Dioclea steyermarkii R.H. Maxwell
Dioclea violacea Mart, ex Benth.
Dioclea virgata (Rich.) Amshoff
Dioclea virgata var. crenata R.H. Maxwell
Dioclea virgata var. virgata
Dioclea wilsonii Standi.
Diplotropis brasiliensis (Tul.) Benth.
Diplotropis martiusii Benth.
Diplotropis purpurea (Rich.) Amshoff
Diplotropis purpurea var. leptophylla (Kleinhoonte) Amshoff
Diplotropis purpurea var. purpurea
Diplotropis racemosa (Hoehne) Amshoff
Diplotropis racemosa var. racemosa
Diplotropis rigidocarpa H.C. Lima
Diplotropis sp. A
Diplotropis strigulosa R.S. Cowan
Diplotropis triloba Gleason
Dipteryx magnifica (Ducke) Ducke
Dipteryx odorata (Aubl.) Willd.
Dipteryx punctata (S.F. Blake) Amshoff
Dipteiyx rosea Spruce ex Benth.
Dussia discolor (Benth.) Amshoff
Eriosema crinitum (Kunth) G. Don
Eriosema crinitum var. crinitum
Eriosema crinitum var. stipulare (Benth.) Fortunato
Eriosema rufum (Kunth) G. Don
Eriosema rufum var. rufum
Eriosema simplicifolium (Kunth) G. Don
Eriosema simplicifolium var. simplicifolium
Eriosema violaceum (Aubl.) G Don
Erythrina amazonica Krukoff
Erythrinafusca Lour.
Erythrina mitis Jacq.
Erythrina poeppigiana (Walp.) Cook
Erythrina rubrinervia Kunth
Etaballia dubia (Kunth) Rudd
Fissicalyxfendleri Benth.
Flemingia strobilifera (L.) R. Br. [introduced]
Galactia dubia DC.
BO DA GU su
BO DA GU su
AM BO GU su
AM BO DA GU FG
AM BO DA su
AM BO GU su FG
FG
AM BO GU su
AM BO
BO su
su
AM BO DA GU su FG
AM
FG?
BO GU su FG
AM BO DA GU su FG
AM BO DA GU
GU
su FG
FG?
AM BO DA GU su FG
AM
AM BO
AM BO GU su FG
AM
AM BO GU
GU
su FG
AM
GU? su? FG?
AM BO DA GU su FG
AM GU su FG
AM BO DA GU su FG
GU su FG
AM
AM GU su FG
AM BO DA GU su
su
FG
FG
AM BO DA GU su FG
AM GU su
AM GU
GU
su
AM
AM
AM
AM
AM BO DA GU su FG
AM BO GU su FG
AM BO
GU su FG
AM BO DA GU su FG
AM BO GU su FG
AM BO DA GU su
BO DA GU su
BO DA GU su
AM BO DA GU su FG
GU su FG
AM BO GU su FG
GU su FG
BO DA GU su FG
AM BO
AM BO
AM
AM BO GU
AM BO GU
SU
FG?
Checklist of the Plants of the Guiana Shield 351
Galactia gracillima Benth.
Galactiajussiaeana Kunth
Galactiajiissiaeana var. jussiaeana
Galactia latisiliqiia Desv.
Galactia longifolia (Jacq.) Benth.
Galactia striata (Jacq.) Urb.
Gliricidia sepiitm (Jacq.) Kunth ex Walp. [widely cultivated & naturalized]
Hymenolobium excelsum Ducke
Hymenolobiumflavum Kleinhoonte
Hymenolobium heterocarpum Ducke
Hymenolobium nitidum Benth.
Hymenolobium petraeum Ducke
Hymenolobium pulcherrimum Ducke
Indigofera bongardiana (Kuntze) Burkart
Indigofera endecaphylla Jacq.
Indigoferafruticosa L. [escaped from cultivation]
Indigofera hirsuta L. [cultivated and escaped from cultivation]
Indigofera lespedezioides Kunth
Indigofera microcarpa Desv.
Indigofera spicata Forssk. '
Indigofera suffruticosa Mill, [cultivated only?]
Indigofera tinctoria L. [cultivated, perhaps escaped]
Lablab purpureus (L.) Sweet [widely cultivated]
Lonchocarpus chrysophyllus Kleinhoonte
Lonchocarpus crassispermus Poppend.
Lonchocarpus crucisrubierae Pittier
Lonchocarpus densiflorus Benth.
Lonchocarpus dipteroneurus Pittier
Lonchocarpus fendleri Benth.
Lonchocarpus floribundus Benth.
Lonchocarpus hedyosmus Miq.
Lonchocarpus heptaphyllus (Poir.) DC.
Lonchocarpus imatacensis Poppend.
Lonchocarpus longifolius (Benth.) Ducke
Lonchocarpus martynii A.C. Sm.
Lonchocarpus pictus Pittier
Lonchocarpus punctatus Kunth
Lonchocarpus rariflorus Mart, ex Benth.
Lonchocarpus rufescens Benth.
Lonchocarpus sericeus (Poir.) Kunth ex DC. [expected in the Guianas]
Lonchocarpus spruceanus Benth.
Lonchocarpus tubicalyx Pittier ex Poppend.
Lonchocarpus urucu Killip & A.C. Sm.
Lonchocarpus utilis A.C. Sm.
Luetzelburgia sp.
Machaerium acuminatum Kunth
Machaerium acutifolium Vogel
Machaerium acutifolium var. pseudacutifolium (Pittier) Rudd
Machaerium affine Benth.
Machaerium altiscandens Ducke
Machaerium amazonense Hoehne
Machaerium aristulatum (Spruce ex Benth.) Ducke
Machaerium aureiflorum Ducke
Machaerium dubium (Kunth) Rudd
Machaerium ferox (Mart, ex Benth.) Ducke
Machaerium floribundum Benth.
Machaerium floribundum var. floribundum
Machaerium guaremalense Pittier
Machaerium inundatum (Mart, ex Benth.) Ducke
Machaerium isadelphum (E. Mey.) Amshoff
Machaerium kegelii Meisn.
Machaerium leiophyllum (DC.) Benth.
Machaerium leiophyllum var. crista-castrense (Mart, ex Benth.) Rudd
Machaerium leiophyllum var. latifolium (Benth.) Rudd
BO su?
AM DA GU
GU
GU
su
su
FG
FG
AM BO GU
AM BO GU su FG
FG?
GU su FG
AM BO
GU
su
AM BO GU su FG
FG
AM
GU
GU
FG?
AM BO GU FG
AM BO GU
BO GU su FG
FG
BO DA GU su FG
BO GU FG
GU? su FG
GU su FG
BO
BO
AM BO
BO
DA
DA
GU
AM BO GU su FG
AM BO GU su FG
BO DA GU su FG
BO DA
su?
BO GU
BO DA GU su FG
BO
GU
GU
BO DA
DA
FG
AM
AM BO DA GU
GU
su FG
AM
BO
BO
BO GU
BO su FG
AM BO
AM BO
BO
GU FG
AM DA GU su
AM DA GU FG
AM
BO
DA GU FG
AM BO DA GU su
su
FG
AM BO DA GU su FG
AM BO GU
GU
su FG
AM BO GU FG
352 Checklist of the Plants of the Guiana Shield
Machaerium leiophyllum var. leiophyllum
Machaerium lunatum (L. f.) Ducke
Machaerium macrophylhim Benth.
Machaerium macrophyllum var. macrophylhim
Machaerium madeirense Pittier
Machaerium multifoliolatum Ducke
Machaerium mutisii Killip ex Rudd
Machaerium myrianthum Spruce ex Benth.
Machaerium paraense Ducke
Machaerium polyphyllum (Poir.) Benth.
Machaerium quinatum (Aubl.) Sandwith
Machaerium quinatum war. parviflorum (Benth.) Rudd
Machaerium quinatum var. quinatum
Machaerium robiniifolium (DC.) Vogel
Machaerium sp. A
Machaerium tovarense Pittier
Machaerium trifoliolatum Ducke
Macroptilium atropurpureum (Sesse & Moc. ex DC.) Urb.
Macroptilium gracile (Poepp. ex Benth.) Urb.
Macroptilium lathyroides (L.) Urb.
Macroptilium lathyroides var. lathyroides
Macroptilium longipedunculatum (Mart, ex Benth.) Urb.
Macroptilium monophyllum (Benth.) Marechal & Baudet
Monopteryx angustifolia Spruce ex Benth.
Monopteryx inpae W.A. Rodrigues
Monopteryx sp. A
Monopteryx uaucu Spruce ex Benth.
Mucuna pruriens (L.) DC. [cultivated, naturalized]
Mucuna pruriens var. utilis (Wall, ex Wight) Baker ex Burck
Mucuna rostrata Benth.
Mucuna sloanei Fawc. & Rendle
Mucuna urens (L.) Medik.
Muellerafrutescens (Aubl.) Standi.
Myrocarpus venezuelensis Rudd
Myrospermum frutescens Jacq.
Myroxylon balsamum (L.) Harms [cultivated?]
Myroxylon peruiferum L. f.
Ormosia amazonica Ducke
Ormosia bolivarensis (Rudd) Stirton
Ormosia cinerea Benoist
Ormosia coarctata Jacks.
Ormosia coccinea (Aubl.) Jacks.
Ormosia costulata (Miq.) Kleinhoonte
Ormosia coutinhoi Ducke
Ormosia discolor Spruce ex Benth.
Ormosiaflava (Ducke) Rudd
Ormosia grandiflora (Tul.) Rudd
Ormosia guyanensis C.H. Stirt., sp. nov. ined.
Ormosia lignivalvis Rudd
Ormosia macrocalyx Ducke
Ormosia macrophylla Benth.
Ormosia maguireorum Rudd
Ormosia melanocarpa Kleinhoonte
Ormosia nobilis Tul.
Ormosia nobilis var. nobilis
Ormosia paraensis Ducke
Ormosia smithii Rudd
Ormosia sp. A
Ormosia sp. B
Ormosia steyermarkii Rudd
Ormosia stipularis Ducke
Ormosia subsimplex Spruce ex Benth.
Ormosia williamsii Rudd
Pachyrhizus erosus (L.) Urb.
DA
AM BO
AM BO
AM BO
AM
AM
BO
AM BO
AM BO
AM BO
BO
BO
BO
GU SU FG
GU su FG
GU SU FG
GU su
GU su
GU su
GU su
su FG
su FG
GU su FG
GU su FG
GU su FG
GU
GU SU FG
FG
AM BO DA GU FG
AM BO DA GU su FG
AM BO DA
AM BO GU su FG
AM BO
AM
AM
BO
FG
AM
AM BO su FG
AM BO su
BO DA FG
GU su FG
AM BO DA GU su FG
DA GU su FG
AM BO
BO
GU
GU
SU
SU FG
BO GU su FG
AM BO GU su FG
BO GU su FG
GU su FG
AM
su FG
AM
GU
BO GU FG
AM
AM
AM
su FG
AM BO GU su FG
AM BO GU su FG
AM BO
BO
GU
GU
su FG
AM
BO
GU su FG
AM BO
AM
BO FG
FG
GU FG
GU su
su?
FG
GU su FG
FG
GU FG
GU su FG
su FG
Checklist of the Plants of the Guiana Shield 353
Paramachaerium ormosioides (Ducke) Ducke
Paramachaehum schomburgkii (Benth.) Ducke
Phaseolus lunatus L. [cultivated, naturalized] AM
Phaseohis reptans Ducke
Phaseolus vulgaris L. [probably cultivated only] BO
Phaseolus vulgaris var. vulgaris BO
Piscidia carthaginensis Jacq. BO
Platymiscium filipes Benth.
Platymiscium pinnatum (Jacq.) Dugand BO
Platymiscium trinitatis Benth. BO
Platymiscium ulei Harms ex Harms
Platypodium elegans Vogel BO
Platypodium elegans subsp. maxoniamim (Pittier) H.C. Lima BO
Poecilanthe amazonica (Ducke) Ducke AM
Poecilanthe ejfusa (Huber) Ducke SU FG
Poecilanthe hostmannii (Benth.) Amshoff AM GU SU FG
Poecilanthe ovalifolia Kleinhoonte SU
Poiretia punctata (Willd.) Desv. BO
Pterocarpus acapulcensis Rose BO
Pterocarpus amazonum (Benth.) Amshoff AM BO DA GU
Pterocarpus ojficinalis Jacq. BO DA GU SU FG
Pterocarpus officinalis subsp. officinalis GU SU FG
Pterocarpus rohriiVdihX AM BO DA GU SU FG
Pterocarpus santalinoides L'Her. ex DC. AM BO DA GU SU FG
Pwerana;7/2fl5'eo/o/d/e5 (Roxb.) Benth. [naturalized] GU SU FG
Rhynchosia edulis Griseb. BO
Rhynchosia melanocarpa Grear [perhaps expected in the Guianas] BO
Rhynchosia minima (L.) DC. DA GU SU FG
Rhynchosia phaseoloidesiS^'.) DC. BO DA GU SU FG
Rhynchosia reticulata (Sw.) DC. BO GU SU FG
Rhynchosia schomburgkii Benth. BO GU
Sesbania emerus {Aubl)\JTh. BO DA GU SU FG
Sesbania exasperata Kunth AM DA GU SU FG
Sesbania grandiffora (L.) Pers. [cultivated, occasionally escaping]
Sesbania sericea {WxWd.) DC. [mXxo(lviCQd~\ GU SU FG
Sesbania sesban (L.) Merr. [introduced] SU
Singana guianensis Aub\. [name of uncertain application] FG
Soemmeringia sempeifforens Mart. BO GU FG?
Spirotropislongifolia {DC.)Bm\\. BO GU SU FG
Spirotropis sp. A BO
Stylosanthes angustifolia Vogel BO GU SU FG
Stylosanthes capitata Vogel BO
Stylosanthes cayennensis Mohlenbr. [=Stylosanthes guianensis?] FG
Stylosanthes guianensis (Aubl.) Sw. AM BO GU SU FG
Stylosanthes guianensis var. gracilis (Kunth) Vogel AM BO GU SU FG
Stylosanthes guianensis var. guianensis AM BO GU SU FG
Stylosanthes hamata (L.) Taub. BO
Stylosanthes hispida Rich. [=Stylosanthes guianensis?] GU SU FG
Stylosanthes humilis Kunth AM BO
Stylosanthes suffruticosa Mohlenb.
Stylosanthes viscosa (L.) Sw. BO
Swartzia alato-sericea Bameby [expected in AM]
Swartzia amshoffiana R.S. Cowan
Swartzia angustifoliola Schery AM
Swartzia anomala R.S. Cowan
Swartzia apiculata R.S. Cowan
Swartzia aptera DC. FG
Swartzia arborescens (Aubl.) Pittier AM BO GU SU FG
Swartzia argentea Spruce ex Benth. AM
Swartzia argentea var. argentea AM
Swartzia aymardii Bameby BO
Swartzia bannia Sandwith GU SU
Swartzia barnebyana Cuello AM
Swartzia benthamiana Miq. AM GU SU FG
GU
GU SU FG
SU FG
GU
GU
354 Checklist of the Plants of the Guiana Shield
Swartzia benthamiana var. benthamiana
Swartzia benthamiana war. yatuensis R.S. Cowan
Swartzia brachyrachis Harms
Swartzia brachyrachis var. glabrata R.S. Cowan
Swartzia buntingii R.S. Cowan
Swartzia cardiosperma Spruce ex Benth.
Swartzia caudata R.S. Cowan
Swartzia coarctata Jacks.
Swartzia conferta Spruce ex Benth.
Swartzia conferta var. conferta
Swartzia cowanii Steyerm.
Swartzia cupavenensis R.S. Cowan
Swartzia cuspidata Spruce ex Benth.
Swartzia davisii Sandwith
Swartzia dipetala Willd. ex Vogel
Swartzia eriocarpa Benth.
Swartziafanshawei R.S. Cowan
Swartziafloribunda Spruce ex Benth.
Swartzia grandifoHa Bong, ex Benth.
Swartzia gitianensis (Aubl.) Urb.
Swartzia hostmannii Benth.
Swartzia invenusta Bameby
Swartziajenmanii Sandwith
Swartzia kaieteurensis (R.S. Cowan) B.M. Torke, sp. nov. ined.
Swartzia katawa R.S. Cowan
Swartzia laevicarpa Amshoff
Swartzia lamellata Ducke
Swartzia lamellata var. kaieteurensis R.S. Cowan
Swartzia lamellata var. lamellata
Swartzia latifolia Benth.
Swartzia latifolia var. latifolia
Swartzia latifolia var. sylvestris R.S. Cowan
Swartzia laitrifolia Benth. [expected in GU]
Swartzia laxiflora Bong, ex Benth. [expected in the Guianas]
Swartzia leblondii R.S. Cowan
Swartzia leiocalycina Benth.
Swartzia leiogyne (Sandwith) R.S. Cowan
Swartzia leptopetala Benth.
Swartzia longicarpa Amshoff
Swartzia longipedicellata Sandwith
Swartzia macrocarpa Spruce ex Benth.
Swartzia magiiirei R.S. Cowan
Swartzia microcarpa Spruce ex Benth.
Swartzia oblanceolata Sandwith
Swartzia oblonga Benth. [expected in GU]
Swartzia oedipiis Bameby
Swartzia pachyphylla Harms [Mt. Roraima]
Swartzia palustris Bameby
Swartzia panacoco (Aubl.) R.S. Cowan
Swartzia panacoco var. altsonii (Sandwith) R.S. Cowan
Swartzia panacoco var. cardonae (R.S. Cowan) R.S. Cowan
Swartzia panacoco var. kamarangensis R.S. Cowan
Swartzia panacoco var. panacoco
Swartzia panacoco var. polyanthera (Steud.) R.S. Cowan
Swartzia panacoco var. sagotii (Sandwith) R.S. Cowan
Swartzia panacoco var. sandwithiana (Cowan) R.S. Cowan
Swartzia panacoco var. tepuiensis (Schery) R.S. Cowan [expected in GU]
Swartzia parvifolia Schery [expected in GU]
Swartzia pendula Spruce ex Benth.
Swartzia piarensis R.S. Cowan [expected in GU]
Swartzia picta Spruce ex Benth.
Swartzia picta var. bolivarensis R.S. Cowan [expected in GU, possibly in
Swartzia picta var. picta
Swartzia pinnata (Vahl) Willd.
AM GU SU FG
AM
BO GU su
BO GU SU
AM
AM
AM
T?/~'
AM BO GU
FG
AM BO
BO
GU
AM
AM
GU
AM BO GU
GU
GU
AM
AM BO GU su FG
AM
BO
GU
GU
GU
GU
GU
su
su
FG
FG
AM BO GU
GU
GU
GU
GU
GU
GU
su
su
su
FG
AM
AM
AM
AM
AM
AM
BO
GU
GU
GU
SU
FG
FG
SU]
GU SU FG
AM
BO
AM
AM BO GU
GU
su FG
AM BO GU
GU
FG
GU su FG
GU? FG
GU
BO
AM BO
AM
BO
AM BO
BO
AM BO
AM
Checklist of the Plants of the Guiana Shield 355
Swartzia pittieri Schery
Swartzia polyphylla DC.
Swartzia recurva Poepp. [expected in GU?]
Swartzia rediviva R.S. Cowan
Swartzia remiger Amshoff
Swartzia reticulata Ducke
Swartzia roraimae Sandwith [Mt. Roraima]
Swartzia schomburgkii Benth.
Swartzia sericea Vogel
Swartzia sericea var. sericea
Swartzia sp. A
B
C
D
AM
AM
BO
DA GU SU FG
Swartzia sp.
Swartzia sp.
Swartzia sp.
Swartzia sp.
Swartzia sp.
Swartzia sprucei Benth.
Swartzia sprucei var. sprucei
Swartzia sprucei var. tessellata R.S. Cowan
Swartzia steyermarkii R.S. Cowan
Swartzia stipellata R.S. Cowan [expected in GU]
Swartzia tessmannii Harms [expected in GU]
Swartzia tillettii R.S. Cowan
Swartzia triptera Bameby
Swartzia vaupesiana R.S. Cowan
Swartzia wurdackii R.S. Cowan [expected in GU]
Swartzia xanthopetala Sandwith
Taralea cordata Ducke
Taralea cordata var. cordata
Taralea cordata var. rigida (Schery) H.C. Lima
Taralea crassifolia (Spruce ex Benth.) Ducke
Taralea nudipes (Tul.) Ducke
Taralea oppositifolia Aubl.
Taralea reticulata (Benth.) Ducke
Tephrosia adunca Benth.
Tephrosia Candida DC. [introduced]
Tephrosia cinerea (L.) Pers.
Tephrosia purpurea (L.) Pers.
Tephrosia senna Kunth
Tephrosia sessiliflora (Poir.) Hassl.
Tephrosia sinapou (Buc'hoz) A. Chev.
Teramnus labialis Spreng. [introduced?]
Teramnus uncinatus (L.) Sw.
Teramnus uncinatus subsp. uncinatus
Vatairea erythrocarpa (Ducke) Ducke
Vatairea guianensis Aubl.
Vatairea paraensis Ducke
Vataireopsis speciosa Ducke [records may represent misidentifications]
Vataireopsis surinamensis H.C. Lima
Vigna adenantha (G Mey.) Marechal, Mascherpa & Stainier
Vigna Candida (Veil.) Marechal, Mascherpa & Stainier
Vigna hosei (Craib) Backer [cultivated, naturalized]
Vignajuruana (Harms) Verde.
Vigna lasiocarpa (Mart, ex Benth.) Verde.
Vigna linearis (Kunth) Marechal, Mascherpa & Stainier
Vigna linearis var. linearis
Vigna longifolia (Benth.) Verde.
Vigna luteola (Jacq.) Benth.
Vigna peduncularis (Kunth) Fawc. & Rendle
Vigna peduncularis var. peduncularis
Vigna trichocarpa (C. Wright) A. Delgado-Salinas
Vigna unguiculata (L.) Walp. [cultivated, naturalized]
Vigna vexillata (L.) A. Rich.
Vigna vexillata var. vexillata
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
SU
GU SU FG
GU SU
GU
GU SU
GU? FG
GU? FG
GU SU
GU
GU
GU
GU
SU? FG
GU
AM BO GU SU FG
AM GU SU FG
AM BO
AM BO GU SU FG
FG
AM
BO
GU
GU
SU FG
BO GU SU
SU
FG
AM BO
BO
DA GU
GU
SU
SU
FG
AM BO GU SU FG
AM BO GU
GU
SU FG
BO GU SU FG
BO GU SU FG
FG
AM DA GU SU FG
BO DA SU FG
AM GU SU FG
AM GU SU FG
AM BO DA GU? SU FG
AM BO
SU FG
AM BO DA GU SU FG
AM BO GU SU FG
AM BO GU SU FG
AM BO GU SU FG
AM DA FG
DA GU SU FG
AM BO SU FG
AM BO
GU SU FG
GU SU FG
AM DA GU SU FG
AM DA GU SU FG
356 Checklist of the Plants of the Guiana Shield
BO
AM BO
AM BO
BO
BO
BO
BO
AM BO
AM BO
BO
AM BO
AM BO
GU
GU
GU
GU
GU
Zollernia grandifolia Schery BO
Zollernia paraensis Huber AM
Zornia brasiliensis Vogel
Zornia curvata Mohlenbr.
Zornia diphylla (L.) Pers.
Zornia gemella (Willd.) Vogel
Zornia glabra Desv.
Zornia gracilis DC.
Zornia giianipensis Pittier
Zornia herbacea Pittier
Zornia lasiocarpa A.R. Molina
Zornia latifolia Sm.
Zornia latifolia var. latifolia
Zornia myriadena Benth.
Zornia pardina Mohlenbr.
Zornia pardina var. vichadana Killip ex Mohlenbr.
Zornia reticulata Sm.
Zornia sericea Moric.
Synonyms and Misidentifications
Abrus minor Desv. =Abnis precatoriits subsp. africanus
Abriis tenuiflorus Benth. ^Abriis piilchelliis subsp. tenuifloriis
Acouroa violacea Aubl. ? =Dalbergia sp.
Aeschynomene biflora Fawc. & Rendle =Aeschynomene brasiliana
Aeschynomene conferta Benth. =Aeschynomene histrix var. histrix
Aeschynomene densiflora Benth. =Aeschynomene histrix var. densiflora
Aeschynomene emeriis Aubl. =Sesbania emerus
Aeschynomene gracilis Miq. ^Aeschynomene paniculata
Aeschynomene guarica Pittier =Aeschynomene brasiliana
Aeschynomene hispida Willd. [not in flora area] =Aeschynomene virginica (L.) Britton, Poggenb. & Stems.
Aeschynomene viscidida Michx. [not in flora area]
Agati grandiflora (L.) Desv. =Sesbania grandiflora
Aldina yapacanensis R.S. Cowan =Aldina macrophylla var. yapacanensis
Alexa imperatricis auct. non R.H. Schomb. 1845, nee Baill. 1870. =Alexa confusa
Alexa leiopetala Sandwith =Alexa imperatricis
Alexa superba R.S. Cowan =Alexa confusa
Alexandra imperatricis R.H. Schomb. =^Alexa imperatricis
AHania insignis Benth. =Aldina insignis
Amerimnon brownei Jacq. =Dalbergia ecastaphyllum
Amerimnon latifolium Willd. =Lonchocarpus heptaphyllus
Amerimnon pinnatum Jacq. =Platymiscium pinnatum
Amphiodon effusus Huber =Poecilanthe effusa
Andira retusa (Poir.) Kunth =Andira surinamensis
Anila bongardiana Kuntze =Indigofera bongardiana
Barbieria polyphylla (Poir.) DC. =Barbieria pinnata
Bowdichia brasiliensis (Tul.) Ducke =Diplotropis brasiliensis
Bowdichia guianensis Ducke =Diplotropis purpurea
Bowdichia martiusii (Benth.) Ducke =Diplotropis martiusii
Bowdichia racemosa Hoehne =Diplotropis racemosa
Bradburya angustifolium (Kunth) Kuntze =Centrosema brasilianum
Bradburya brasilianum (L.) Kuntze =Centrosema brasilianum
Bradburya macrocarpa (Benth.) Kuntze =Centrosema macrocarpum
Bradburya pascuora (Mart, ex Benth.) Kuntze =Centrosema pascuorum
Bradburya plumieri (Turpin ex Pers.) Kuntze =Centrosema plumieri
Bradburya triquetra (Hoffmanns, ex Benth.) Kuntze =Centrosema triquetrum
Bradburya venosum (Mart, ex Benth.) Kuntze =Centrosema venosum
Bradburya virginianum (L.) Kuntze =Centrosema virginianum
Cajanus indicus Spreng. =CaJanus cajan
Canavalia bracteolata Merr. =Dioclea macrantha var. bracteolata
Canavaliafendleri Piper =Canavalia brasiliensis
Canavalia maritima Thouars =Canavalia rosea
Canavalia obtusifolia DC. =Canavalia rosea
Centrolobium orinocense (Benth.) Pittier =Centrolobium paraense var. orinocense
Centrolobium patinense Pittier =Centrolobium paraense var. orinocense
SU
su
SU
su
FG
FG
FG
FG
GU SU FG
GU su FG
GU
GU
GU
Checklist of the Plants of the Guiana Shield 357
Centrosema angustifolium (Kunth) Benth. in Mart., 1859 =Centrosema brasilianum
Centrosema brasilianum var. angustifolium (Kunth) Amshoff=Ce«?ro5ewa brasilianum
Centrosema latissimum Ducke =Centrosema triquetrum
Centrosema macrocarpon auct. [orthographic error] =Centrosema macrocarpum
Centrosema platycarpum Benth. [not in flora area]
Centrosema pubescens auct. non Benth. 1837 [sensu Benth. in Mart., fide Fantz] =Centrosema molle
Chaetocalyxfissa Pittier =Chaetocalyx scandens var. pubescens
Chaetocalyx magniflora Pittier =Chaetocalyx scandens var. pubescens
Chaetocalyx nigrescens Pittier =Chaetocalyx scandens var. pubescens
Chaetocalyx paucifolia Pittier =Chaetocalyx scandens var. pubescens
Chaetocalyx perglandulosa Pittier =Chaetocalyx scandens var. pubescens
Chaetocalyx pubescens DC. =Chaetocalyx scandens var. pubescens
Chaetocalyx vestita Standi. =Chaetocalyx scandens var. pubescens
Clathrotropis colombiana Yakovlev =Clathrotropis brunnea
Clathrotropisflava Ducke =Ormosiaflava
Clathrotropis surinamensis Kleinhoonte =Ormosiaflava
Clitoria amoena Miq. =Clitoria arborescens
Clitoria angustifolia Kunth =Centrosema brasilianum
Clitoria angustifolia auct. non Kunth 1823 [1824] =Centrosema brasilianum
Clitoria arborea Mart, ex Benth. [not in flora area]
Clitoria brasiliana L. =Centrosema brasilianum
Clitoria brasiliensis L. =Centrosema brasilianum
Clitoria cajanifolia (C. Presl.) Benth. =Clitoria laurifolia
Clitoria capitata Rich. =Centrosema capitatum
Clitoria cerifera R.S. Cowan =Clitoria coriacea
Clitoria glycinoides DC. =Clitoriafalcata
Clitoria grandifolia Ducke =Clitoriajavitensis var. grandifolia
Clitoria guyanensis Benth. =Clitoria guianensis
Clitoria guyanensis Aubl. ex Desv. =Clitoria laurifolia
Clitoria javitensis var. glabra Sagot =Clitoria sagotii
Clitoria parvifolia Pittier [nomen nudum] =Clitoria laurifolia
Clitoria pinnata (Pers.) R.H. Sm. & G.P. Lewis ^^Barbieria pinnata
Clitoria plumieri Turpin ex Pers. =Centrosema plumieri
Clitoria poitaei DC. =Clitoria arborescens
Clitoria poiteai DC. =Clitoria arborescens
Clitoria poiteai Benth. =Clitoria arborescens
Clitoria polyphylla Poir. =Barbieria pinnata
Clitoria rubiginosa Juss. ex Pers. =Clitoriafalcata
Clitoria virginiana L. ^Centrosema virginianum
Collaea rosea Benth. =Galactiajussiaeana
Coronilla scandens L. =Chaetocalyx scandens
Coronilla sericea Willd. ^Sesbania sericea
Coublandiafrutescens Aubl. =Muellerafrutescens
Coumarouna magnifica Ducke =Dipteryx magnifica
Coumarouna odorata Aubl. =Dipteryx odorata
Coumarouna oppositifolia (Aubl.) Taub. =Taralea oppositifolia
Coumarouna punctata S.F. Blake =Dipteryx punctata
Coursetia arborea Griseb. =Coursetiaferruginea
Crotalaria anagyroides Kunth =Crotalaria micans
Crotalaria anagyroides var. pauciflora Griseb. =Crotalaria maypurensis
Crotalaria bracteata Schltdl. & Cham. =Crotalaria nitens
Crotalaria cuneifolia (Forssk.) Schrank =Crotalaria retusa
Crotalaria depauperata Mart. =Crotalaria maypurensis var depauperata
Crotalaria gajanensis Desv. =Clitoria laurifolia
Crotalaria guianensis Aubl. =Clitoria guianensis
Crotalaria leptophylla Benth. =Crotalaria maypurensis
Crotalaria paramariboensis Miq. =Crotalaria verrucosa
Crotalaria pilosa var. skutchii Senn =Crotalaria pilosa
Crotalaria pterocaula Desv. =Crotalaria pilosa
Crotalaria retusifolia Stokes ^Crotalaria retusa
Crotalaria saltiana Andrews =Crotalaria pallida
Crotalaria schiedeana Steud. =Crotalaria nitens
Cyclolobium amazonicum Ducke ^^Poecilanthe amazonica
Cyclolobium hostmannii Benth. =Poecilanthe hostmannii
358 Checklist of the Plants of the Guiana Shield
Cymbosema apurense (Kunth) Pittier =Dioclea apuremis
Cymbosema apurense (Kunth) Pittier, sensu Pitter Bol. Soc. Venez. Ci. Nat. 7: 154. =Cymbosema roseum
Cytisus cajan L. =Cajanus cajan
Cytisus sessiliflorus Poir. =Tephrosia sessiliflora
Cytisus violaceous Aubl. =Ehosema violaceum
Cytisus violaceus Aubl. =Eriosema violaceum
Dalbergia atropurpurea Ducke =Dalbergiafoliosa
Dalbergia aturensis Pittier =Dalbergia inundata
Dalbergia brownei (Jacq.) Urb. =Dalbergia ecastaphyllum
Dalbergia cucullata Pittier (Leguminosae-Caesalpinioideae) =Apuleia leiocarpa
Dalbergia glauca (Desv.) Amshoff =Dalbergiafoliosa
Dalbergia heptaphylla Poir. =Lonchocarpus heptaphyllus
Dalbergia pachycarpa Ducke =Dalbergia riedelii
Dalbergia pentaphylla Poir. =Lonchocarpus heptaphyllus
Dalbergia tomentosa (Spruce ex Benth.) Ducke =Dalbergia intermedia
Dalbergia variabilis Vogel =Dalbergiafrutescens
Deguelia scandens Aubl. =Derris pterocarpus
Derris ernestii (Harms) Ducke =Lonchocarpus hedyosmus
Derris floribundus (Benth.) Ducke =Lonchocarpus floribundus
Derris glabrescens (Benth.) J.F. Macbr. =Lonchocarpus densiflorus
Derris guianensis Benth. =Derris pterocarpus
Derris guyanensis Benth. =Derris pterocarpus
Derris hedyosma (Miq.) J.F. Macbr. =Lonchocarpus hedyosmus
Derris latifolia (Kunth ex DC.) Ducke =Lonchocarpus heptaphyllus
Derris longifolia Benth. =Lonchocarpus longifolius
Derris nicou (Aubl.) J.F. Macbr. =Lonchocarpus utilis
Derris scandens (Aubl.) Pittier =Derris pterocarpus
Derris urucu (Killip & A.C. Sm.) J.F. Macbr. =Lonchocarpus urucu
Derris utilis (A.C. Sm.) Ducke =Lonchocarpus utilis
Desmodium adscendens var. orinocense DC. =Desmodium orinocense
Desmodium asperum (Poir.) Desv. =Desmodium distortum
Desmodium axillare var. sintenisii Urb. =Desmodium axillare var. stoloniferum
Desmodium canum (J.F. Gmel.) Schinz & Thell. =Desmodium incanum
Desmodium dubium Pittier =Desmodium campyloclados
Desmodiumflexuosum Pittier =Desmodium campyloclados
Desmodium frutescens (Jacq.) Schindl. =Desmodium incanum
Desmodium molle (Vahl) DC. [not in flora area] =Desmodium glabrum (Mill.) DC.
Desmodium spectabilis Miq. =Desmodium distortum
Desmodium supinum DC. =Desmodium incanum
Dibrachion brasiliensis Tul. =Diplotropis brasiliensis
Dibrachion guianense Tul. =Diplotropis purpurea
Dibrachion riparium Spruce ex Benth. =Diplotropis martiusii
Dioclea broadwayana Pittier [nomen nudum] =Dioclea guianensis
Dioclea elliptica R.H. Maxwell [nomen nudum] =Dioclea scabra var. scabra
Dioclea glabra auct. non Benth. 1 837 =Dioclea scabra var. scabra
Dioclea guianensis var. villosior Benth. =Dioclea guianensis
Dioclea holtiana Pittier [nomen nudum] =Dioclea holtiana Pittier ex R.H. Maxwell
Dioclea lasiocarpa Mart. =Dioclea virgata
Diplotropis brachypetala Tul. =Clathrotropis brachypetala
Diplotropis grandiflora Tul. =Ormosia grandiflora
Diplotropis guianensis Benth. =Diplotropis purpurea
Diplotropis leptophyIla Kleinhoonte =Diplotropis purpurea var. leptophylla
Diplotropis nitida Benth. =Clathrotropis nitida
Diplotropis racemosa var. kaieteurensis Amshoff =Diplotropis racemosa var. racemosa
Dipteryx applanata Benth. =Taralea oppositifolia
Dipteryx cordata (Ducke) R.S. Cowan =Taralea cordata
Dipteryx crassifolia Spruce ex Benth. =Taralea crassifolia
Dipteryx nudipes Tul. =Taralea nudipes
Dipteryx oppositifolia (Aubl.) Willd. =Taralea oppositifolia
Dipteryx oppositifolia var. parviflora Benth. =Taralea oppositifolia
Dipteryx phaeophylla Steyerm. =Spirotropis longifolia
Dipteryx reticulata Benth. =Taralea reticulata
Dipteryx rigida (Schery) R.S. Cowan =Taralea cordata var. rigida
Dipteryx trifoliolata Ducke =Dipteryx punctata
Checklist of the Plants of the Guiana Shield 359
Dolicholus kuntzei Harms ex Kuntze =Rhynchosia reticulata
Dolicholus phaseoloides (Sw.) DC. =Rhynchosia phaseoloides
Dolicholus pittieri Standi. =Rhynchosia schomburgkii
Dolichos altissmus Jacq. =Mucuna urens
Dolichos cuneifolius Forssk. =Crotalaria retusa
Dolichos ensiformis L. =Canavalia ensiformis
Dolichos erosus L. =Pachyrhizus erosus
Dolichos gladiatus Jacq. =Canavalia gladiata
Dolichos lablab L. =Lablab purpureus
Dolichos luteolus Jacq. =Vigna luteola
Dolichos maritimus Aubl. =Canavalia rosea
Dolichos minimus L. =Rhynchosia minima
Dolichos pruriens L. =Mucuna pruriens
Dolichos roseus Sw. =Canavalia rosea
Dolichos scabra Rich. =Dioclea scabra
Dolichos uncinatus L. =Teramnus uncinatus
Dolichos unguiculatus L. =Vigna unguiculata
Dolichos urens L. =Mucuna urens
Dolichos virgatus Rich. =Dioclea virgata
Drepanocarpus aristulatus Spruce ex Benth. =Machaerium aristulatum
Drepanocarpus crista-castrensis van latifolius Benth. =Machaerium leiophyllum var. latifolium
Drepanocarpus dubius Kunth =Machaerium dubium
Drepanocarpus ferox Mart, ex Benth. =Machaerium ferox
Drepanocarpusferox J3 macrophyllum Benth. =Machaerium ferox
Drepanocarpusfrondosus Mart, ex Benth. =Machaerium leiophyllum var. latifolium
Drepanocarpus inundatus Mart, ex Benth. =Machaerium inundatum
Drepanocarpus isadelphus E. Mey. =Machaerium isadelphum
Drepanocarpus lunatus (L. f.) G. Mey. =Machaerium lunatum
Drepanocarpus macrophyllus Ducke =Machaerium macrophyllum
Drepanocarpus ? ovalifolius Pittier =Machaerium floribundum
Drepanocarpus paludicola Standi. ^Dalbergia inundata
Drepanocarpus venezuelensis Pittier =Machaeriumfloribundum
Dussia cayennensis Harms =Dussia discolor
Ecastaphyllum amazonicum Radlk. ex Kopff =Dalbergia amazonica
Ecastaphyllum benthamianum Miq. =Dalbergia monetaria
Ecastaphyllum foliosum Benth. =Dalbergia foliosa
Ecastaphyllum glaucum Desv. =Dalbergiafoliosa
Ecastaphyllum hygrophilum Mart, ex Benth. =Dalbergia hygrophila
Ecastaphyllum monetaria (L. f.) DC. =Dalbergia monetaria
Ecastaphyllum monetaria var. riedelii Benth. =Dalbergia riedelii
Ecastaphyllum pubescens DC. =Dalbergia subcymosa
Ecastaphyllum riedeli (Benth.) Radlk. =Dalbergia riedelii
Ecastaphyllum tomentosum Spruce ex Benth. =Dalbergia intermedia
Eriosema edule (Griseb.) Burkhart =Rhynchosia edulis
Eriosemaflaviflorum Miq. =Eriosema violaceum
Eriosemafusiforme Rusby [not in flora area] =Eriosema crinitum var. fusiforme (Rusby) Grear
Eriosema lanceolatum Benth. =Eriosema simplicifolium
Eriosema simplicifolium var. micranthum Grear =Eriosema simplicifolium
Eriosema stipulare Benth. =Eriosema crinitum var. stipulare
Eriosema volubile Micheli =Rhynchosia edulis
Erythrina corallodendrum L. [not in flora area]
Erythrina glauca Willd. =Erythrinafusca
Etaballia guianensis Benth. =Etaballia dubia
Galactia angustifolia Kunth =Galactiajussiaeana
Galactia camporum Sprague =Galactiajussiaeana
GalactiaJussiaeana var. angustifolia (Kunth) Burkart =Galactiajussiaeana
Galactia pinnata Pers. =Barbieria pinnata
Galactia striata var. tenuiflora (Klein ex Willd.) Burkart =Galactia striata
Galactia tenuiflora (Klein ex Willd.) Wight & Am. pro parte, sensu Benth. in Mart. =Galactia striata
Galactia velutina Benth. ^Galactia latisiliqua
Galega cathartica Sesse & Moc. =Tephrosia senna
Galega cinerea L. =Tephrosia cinerea
Galega sinapou Buc'hoz =Tephrosia sinapou
Galega toxicaria Sw. =Tephrosia sinapou
360 Checklist of the Plants of the Guiana Shield
Geoffroea discolor Benth. =Dussia discolor
Geoffroea inermis W. Wright =Andira inermis
Geoffroea pubescens Rich. =Andira surinamensis
Geoffroea retusa Poir. =Andira surinamensis
Geoffroea surinamensis Bondt =Andira surinamensis
Geoffroea violacea (Aubl.) Pers. [=Dalbergia sp.?] =Acouroa violacea
Glycine crinita Kunth =Eriosema crinitum
Glycine phaseoloides Sw. =Rhynchosia phaseoloides
Glycine punctata Willd. =Poiretia punctata
Glycine reflexa Nutt. =Rhynchosia minima
Glycine reticulata Sw. =Rhynchosia reticulata
Glycine rufa Kunth =Eriosema rufum
Glycine simplicifolia Kunth =Eriosema simplicifolium
Glycine striata Jacq. =Galactia striata
Glycine tenuiflora Klein ex Willd. =Galactia striata
Hecastophyllum dubium Kunth =Etaballia dubia
Hedysarum adscendens auct. non Sw. 1788 =Desmodium orinocense
Hedysarum adscendens Sw. =Desmodium adscendens
Hedysarum asperum Poir. =Desmodium distortum
Hedysarum axillare Sw. =Desmodium axillare
Hedysarum barbatum L. =Desmodium barbatum
Hedysarum brasilianum Poir. =Aeschynomene brasiliana
Hedysarum cajanifolium Kunth =Desmodium cajanifolium
Hedysarum cajanifoliun Kunth =Desmodium cajanifolium
Hedysarum canum J.F. Gmel. =Desmodium incanum
Hedysarum diphyHum L. =Zornia diphylla
Hedysarum distortum Aubl. =Desmodium distortum
Hedysarum ecastaphyllum P. Browne ex L. =Dalbergia ecastaphyllum
Hedysarum gemellum Willd. =Zornia gemella
Hedysarum hamatum L. =Stylosanthes hamata
Hedysarum hamatum var. viscosum L. =Stylosanthes viscosa
Hedysarum incanum Sw. =Desmodium incanum
Hedysarum procumbens Mill. =Desmodium procumbens
Hedysarum scorpiurus Sw. =Desmodium scorpiurus
Hedysarum stoloniferum Rich, ex Poir. =Desmodium axillare var. stoloniferum
Hedysarum tortuosum Sw. =Desmodium tortuosum
Hedysarum triflorum L. =Desmodium triflorum
Hedysarum vaginale L. =Alysicarpus vaginalis
Humboldtiellaferruginea (Kunth) Harms =Coursetiaferruginea
Indigofera anil L. =Indigofera suffruticosa
Indigofera pascuorum Benth. =Indigofera lespedezioides
Leptolobium costulatum Miq. =Ormosia costulata
Leptolobium nitens Vogel =Acosmium nitens
Leptolobium nitidum Miq. =Acosmium nitens
Lonchocarpus benthamianus Pittier =Lonchocarpus punctatus
Lonchocarpus boliviensis Pittier =Lonchocarpus densiflorus
Lonchocarpus discolor Huber =Lonchocarpus heptaphyllus
Lonchocarpus ernestii Harms ^Lonchocarpus hedyosmus
Lonchocarpus fendleri suhsp. pubescens Pittier =Lonchocarpus fendleri
Lonchocarpus glabrescens Benth. =Lonchocarpus densiflorus
Lonchocarpus guaricensis Pittier =Piscidia carthaginensis
Lonchocarpus killipii Ducke =Derris negrensis
Lonchocarpus latifolius Kunth ex DC. =Lonchocarpus heptaphyllus
Lonchocarpus margaritensis Pittier =Lonchocarpus hedyosmus
Lonchocarpus negrensis Benth. non Derris negrensis Benth. 1860 =Derris amazonica
Lonchocarpus nicou (Aubl.) DC. =Lonchocarpus utilis
Lonchocarpus nicou auct. non (Aubl.) DC. sensu Killip & A.C. Sm. 1825 ^Lonchocarpus utilis
Lonchocarpus nitidulus Benth. =Lonchocarpus floribundus
Lonchocarpus nitidus Spruce ex Benth. =Lonchocarpus floribundus
Lonchocarpus paniculatus Ducke =Lonchocarpus hedyosmus
Lonchocarpus pentaphyllus (Poir.) DC. =Lonchocarpus heptaphyllus
Lonchocarpus pterocarpus DC. ^Derris pterocarpus
Lonchocarpus sanctae-marthae Pittier =Lonchocarpus fendleri
Lonchocarpus scandens (Aubl.) Ducke =Derris pterocarpus
Checklist of the Plants of the Guiana Shield 361
Lonchocarpiis stenopteris Pittier =Lonchocarpus dipteroneurus
Lonchocarpus stenunis Pittier =Lonchocarpus fendleri
Lonchocarpiis violaceiis (Jacq.) DC. =Lonchocarpus punctatus
Machaerium aculeatiim Raddi [not in flora area]
Machaerium bracteatum Benth. =Machaerium kegelii
Machaerium caicareme Pittier ^Machaerium dubium
Machaerium compressicaule Ducke =Machaerium madeirense
Machaerium compressicaule var. manoense Ducke =Machaerium madeirense
Machaerium decorticans Ducke =Machaerium floribundum
Machaerium eggersii Hoehne =Machaerium robiniifolium
Machaerium ferrugineum (Willd.) Pers. =Machaerium quinatum
Machaerium ferrugineum fi parviflorum Benth. =Machaerium quinatum var. parviflorum
Machaeriumfrondosum (Mart, ex Benth.) Ducke =Machaerium leiophyllum var. latifolium
Machaerium latifolium var. manoense (Ducke) Hoehne =Machaerium madeirense
Machaerium longistipitatum Pittier =Machaerium floribundum
Machaerium nervosum Vogel =Machaerium quinatum var. parviflorum
Machaerium oblongifolium Vogel [not in flora area]
Machaerium pachyphyllum Pittier =Machaerium kegelii
Machaerium pseudacutifolium Pittier =Machaerium acutifolium var. pseudacutifolium
Machaerium rectipes Pittier =Machaerium affine
Machaerium rosescens Standi. ^Machaeriumfloribundum
Machaerium schomburgkii Benth. =Paramachaerium schomburgkii
Machaerium sieberi Benth. =Machaerium robiniifolium
Machaerium steinbachianum, Hoehne =Machaerium kegelii
Machaerium surinamense Miq. =Machaerium leiophyllum
Machaerium venezuelense (Pittier) Hoehne =Machaerium floribundum
Machaerium woodworthii Standi. =Machaerium floribundum
Meibomia axillaris (Sw.) Kuntze =Desmodium axillare
Meibomia axillaris var. acutifolia Kunze =Desmodium axillare var. acutifolium
Meibomia campyloclada (Hemsl.) Kuntze =Desmodium campyloclados
Micropteryx poeppigiana Walp. =Erythrina poeppigiana
Monopteryxjahnii Pittier =Fissicalyxfendleri
Moutouchi suberosa Aubl. =Pterocarpus officinalis
Mucuna altissima (Jacq.) DC. =Mucuna urens
Mucuna hassjoo (Piper & Tracy) Mansf. =Mucuna pruriens var. utilis
Mucuna utilis Wall, ex Wight =Mucuna pruriens var utilis
Muellera moniliformis L. f. =Muellerafrutescens
Neurocarpum cajanifolium C. Presl =Clitoria laurifolia
Neurocarpum guianense (Aubl.) Desv. =Clitoria guianensis
Neurocarpum javitense auct. non Kunth =Clitoria sagotii
Neurocarpum javitense Kunth =Clitoriajavitensis
Neurocarpum simplicifolium Kunth =Clitoria simplicifolia
Nicolsonia orinocensis (DC.) Schindl. ^Desmodium orinocense
Nissolia acuminata (Kunth) DC. =Machaerium acuminatum
Nissoliaferruginea Willd. =Machaerium quinatum
Nissolia leiophylla DC. =Machaerium leiophyllum
Nissolia polyphylla Poir. =Machaerium polyphyllum
Nissolia quinata Aubl. =Machaerium quinatum
Nissolia robiniifolia DC. ^Machaerium robiniifolium
Ormosia chlorocalyx Ducke =Ormosia macrocalyx
Ormosia coccinea var. subsimplex (Spruce ex Benth.) Rudd ^Ormosia subsimplex
Ormosia costulata var. trifoliata (Huber) Amshoff =Ormosia costulata
Ormosia crassicarpa Pierce ex Pittier =Ormosia paraensis
Ormosia cuneata Ducke =Ormosia coarctata
Ormosia euneura Harms =Ormosia amazonica
Ormosia fastigiata Tul. [not in flora area]
Ormosia heterophylla Pires =Ormosia paraensis
Ormosia micrantha Ducke =Ormosia discolor
Ormosia microsperma Pittier =Ormosia steyermarkii
Ormosia nobilis var. bolivarensis Rudd =Ormosia bolivarensis
Ormosia pacimonensis Spruce ex Benth. =Vatairea guianensis
Ormosia toledana Standi. =Ormosia macrocalyx
Ormosia trifoliata Huber =Ormosia costulata
Ormosiopsisflava Ducke =Ormosiaflava
362 Checklist of the Plants of the Guiana Shield
Phaseolus adenanthus G. Mey. =Vigna adenantha
Phaseolus appendiculatus Benth. =Vigna Candida
Phaseolus campestris Mart, ex Benth. =Macroptilium longipedunculatum
Phaseolus candidus Veil. =Vigna Candida
Phaseolus diversifolius Pittier =Vigna lasiocarpa
Phaseolus gracilis Poepp. ex Benth. =Macroptilium gracile
Phaseolus juruanus Harms ^Vignajuruana
Phaseolus lasiocarpus Mart, ex Benth. =Vigna lasiocarpa
Phaseolus lathyroides L. =Macroptilium lathyroides
Phaseolus linearis Kunth =Vigna linearis
Phaseolus longifolius Benth. =Vigna longifolia
Phaseolus longipedunculatus Mart, ex Benth. =Macroptilium longipedunculatum
Phaseolus monophyllus Benth. =Macroptilium monophyllum
Phaseolus peduncularis Kunth = Vigna peduncularis
Phaseolus pilosus Kunth =Vigna lasiocarpa
Phaseolus schottii Benth. =Vigna longifoliu
Phaseolus surinamensis Miq. =Vigna adenantha
Phaseolus trichocarpus C. Wright =Vigna trichocarpa
Phaseolus truxillensis Kunth =Vigna adenantha
Phaseolus vexillatus L. = Vigna vexillata
Phellocarpus amazonum Mart, ex Benth. =Pterocarpus amazonum
Piscidia guaricensis (Pittier) Pittier =Piscidia carthaginensis
Platymiscium duckei Huber =Platymiscium trinitatis
Platymiscium duckei var. nigrum (Ducke) Ducke =Platymiscium trinitatis
Platymiscium nigrum Ducke =Platymiscium trinitatis
Platymiscium polystachyum Benth. =Platymiscium pinnatum
Platymiscium polystachyum var. fendleri Benth. =Platymiscium pinnatum
Platypodium maxonianum Pittier =Platypodium elegans subsp. maxonianum
Platysema triquetra Hoffmanns, ex Benth. =Centrosema triquetrum
Possira arborescens Aubl. =Swartzia arborescens
POSsira triphylla Sw. =Swartzia arborescens
Pterocarpus amazonicus Huber =Pterocarpus santalinoides
Pterocarpus draco L. ^Pterocarpus officinalis
Pterocarpus esculentus Schumach. & Thonn. =Pterocarpus santalinoides
Pterocarpus frutescens Veil. =Dalbergiafrutescens
Pterocarpus grandis R.S. Cowan =Pterocarpus santalinoides
Pterocarpus lunatus L. f. =Machaerium lunatum
Pterocarpus magnicarpa Schery =Pterocarpus rohrii
Pterocarpus magnicarpus Schery =Pterocarpus rohrii
Pterocarpus podocarpus S.F. Blake =Pterocarpus acapulcensis
Pterocarpus rohrii var. rubiginosus Schery =Pterocarpus rohrii
Pterocarpus rufescens Benth. =Pterocarpus rohrii
Pterocarpus rupestris Pittier ^Pterocarpus rohrii
Pterocarpus suberosa (Aubl.) Pers. =Pterocarpus officinalis
Pterocarpus ulei Harms =Pterocarpus amazonum
Pterocarpus vernalis Pittier =Pterocarpus acapulcensis
Rhadinocarpus brasiliensis Vogel =Chaetocalyx brasiliensis
Rhynchosia caliensis Harms =Rhynchosia schomburgkii
Rhynchosia crinita (Kunth) DC. =Eriosema crinitum
Rhynchosia kuntzei Harms ex Kuntze =Rhynchosia reticulata
Rhynchosia melanosticta Griseb. =Rhynchosia edulis
Rhynchosia phaseoloides var. erecta Micheli =Rhynchosia melanocarpa
Rhynchosia reticulata var. kuntzei (Harms ex Kuntze) Grear =Rhynchosia reticulata
Rhynchosia rufa (Kunth) DC. =Eriosema rufum
Rhynchosia simplicifolia (Kunth) DC. =Eriosema simplicifolium
Rhynchosia violacea (Aubl.) DC. =Eriosema violaceum
Rittera dodecandra Vahl =Swartzia arborescens
Rittera pinnata Vahl =Swartzia pinnata
Robinia coccinea Aubl. =Ormosia coccinea
Robiniaferruginea Kunth =Coursetiaferruginea
Robinia nicou Aubl. =Lonchocarpus utilis
Robinia panacoco Aubl. =Swartzia panacoco
Robinia sepium Jacq. =Gliricidia sepium
Robinia sericea Poir. =Lonchocarpus sericeus
Checklist of the Plants of the Guiana Shield 363
Robinia tomentosa Willd. =Swartzia panacoco
Robinia violacea Jacq. [nom. dubium] =Lonchocarpus punctatus
Sesbania aegyptica Poir. =Sesbania sesban
Sphinctolobium glaucescens Miq. =Muellerafrutescens
Spirotropis candoUei Tul. =Spirotropis longifolia
Stenolobium brachycarpiim (Desv.) Benth. "^Calopogonium mucunoides
Stenolobium coendeum Benth. =Calopogonium caeruleum
Stenolobium velutinum Benth. =Calopogomum velutinum
Stizolobium altissimum (Jacq.) Pers. =Mucuna urens
Stizolobium aterrimun Piper & Tracy =Mucuna pruriens var. utUis
Stizolobium hassjo Piper & Tracy =Mucuna pruriens var. utilis
Stizolobium pruriens (L.) Medik. =Mucuna pruriens
Stylosanthes gracilis Kunth =Stylosanthes guianensis var. gracilis
Swartzia acuminata Willd. ex Vogel =Swartzia polyphylla
Swartzia acuminata war. puberula (Taub.) Glaz. =Swartzia polyphylla
Swartzia acuminata var. tridynamia Huber =Swartzia polyphylla
Swartzia alata Willd. =Swartzia guianensis
Swartzia alterna Benth. (Leguminosae-Caesalpinioideae) =Bocoa alterna
Swartzia apetala Raddi [not in flora area]
Swartzia apetala var. acuminata Amshoff =5'>var/z/a amshojfiana
Swartzia bifida Steud. =Swartzia arborescens
Swartzia brachystachya DC. =Candolleodendron brachystachyum
Swartzia corrugata Benth. [not in flora area]
Swartzia dicarpa Moric. ex Meisn. =Swartzia dipetala
Swartzia discolor Poepp. =Swartzia leptopetala
Swartzia dodecandra (Vahl) Willd. =Swartzia arborescens
Swartzia dolichopoda R.S. Cowan =Swartzia vaupesiana
Swartziafugax Spruce ex Benth. =Swartzia leptopetala
Swartzia grandifolia var. leiogyne Sandwith =Swartzia leiogyne
Swartzia longifolia DC. =Spirotropis longifolia
Swartzia melanoxylon Ducke =Swartzia leptopetala
Swartzia microstylis Benth. =Swartzia dipetala
Swartzia minutiflora Kleinhoonte (Leguminosae-Caesalpinioideae) =Bocoa prouacensis
Swartzia opacifolia J.F. Macbr. =Swartzia polyphylla
Swartzia parviflora DC. =Swartzia arborescens
Swartzia platygyne Ducke =Swartzia polyphylla
Swartzia polyanthera Steud. =Swartzia panacoco var. polyanthera
Swartzia polycarpa Ducke =Swartzia laxiflora
Swartzia prouacensis (Aubl.) Amshoff (Leguminosae-Caesalpinioideae) =Bocoa prouacensis
Swartzia rariflora Hoehne =Swartzia arborescens
Swartzia rotundata R.S. Cowan =Swartzia leptopetala
Swartzia schomburgkii var. guayanensis R.S. Cowan =Swartzia schomburgkii
Swartzia similis Benoist =Swartzia panacoco
Swartzia simplex (Sw.) Spreng. [not in flora area]
Swartzia tepuiensis Schery =Swartzia panacoco var. tepuiensis
Swartzia tomentosa (Willd.) DC. =Swartzia panacoco
Swartzia tomentosa var. altsoni Sandwith =Swartzia panacoco var. altsonii
Swartzia tomentosa var. cardonae R.S. Cowan =Swartzia panacoco var. cardonae
Swartzia tomentosa \ar. polyanthera (Steud.) Sandwith =Swartzia panacoco var. polyanthera
Swartzia tomentosa var. sagotii Sandwith =Swartzia panacoco var. sagotii
Swartzia tomentosa var. sandwithiana R.S. Cowan =Swartzia panacoco var. sandwithiana
Swartzia triphyllata Willd. =Swartzia arborescens
Swartzia urubuensis Ducke =Swartzia polyphylla
Swartzia vaupesiana var. glauca R.S. Cowan =Swartzia vaupesiana
Swartzia viridiflora Ducke (Leguminosae-Caesalpinioideae) =Bocoa viridiflora
Sweetia nitens (Vogel) Benth. =Acosmium nitens
Sweetia praeclara Sandwith =Acosmium praeclarum
Taralea casiquiarensis Pittier =Dipteryx magnifica
Taralea rigida Schery =Taralea cordata var. rigida
Taralea steyermarkii Schery =Taralea reticulata
Tephrosia brevipes Benth. =Tephrosia sessilifora
Tephrosia cathartica (Sesse & Moc.) Urb. =Tephrosia senna
Tephrosia cinerea var. littoralis (Jacq.) Benth. =Tephrosia cinerea
Tephrosia littoralis (Jacq.) Pers. =Tephrosia cinerea
364 Checklist of the Plants of the Guiana Shield
Tephrosia toxicaha (Sw.) Pers. =Tephrosia sinapou
Ternatea arborescens (W.T. Alton) Kuntze =Clitoria arborescens
Ternateajavitensis (Kunth) Kuntze =Clitoriajavitensis
Ternatea simplicifolia (Kunth) Kuntze =Clitoria simplicifolia
Toluifera balsamum L. =Myroxylon balsamum
Toiiantea latifolia (Benth.) Taub. =Swartzia latifolia
Touantea sericea (Vogel) Kuntze =Swartzia sericea
Tounatea acuminata (Willd. ex Vogel) Taub. =Swartzia polyphylla
Tounatea acuminata var. puberula Taub. =Swartzia polyphylla
Tounatea aptera (DC.) Taub. =Swartzia aptera
Tounatea arborescens (Aubl.) Brltton =Swartzia arborescens
Tounatea benthamiana (Mlq.) Taub. ^Swartzia benthamiana
Tounatea dodecandra (Vahl) Taub. =Swartzia arborescens
Tounatea guianensis Aubl. =Swartzia guianensis
Tounatea hostmanni (Benth.) Taub. =Swartzia hostmannii
Tounatea laxi/lora (Bong, ex Benth.) Taub. =Swartzia laxiflora
Tounatea laxiflora (Bong, ex Benth.) Kuntze =Swartzia laxiflora
Tounatea schomburgkii (Benth.) Taub. =Swartzia schomburgkii
Trifolium guianense Aubl. =Stylosanthes guianensis
Vatairea surinamensis Kleinhoonte =Vatairea guianensis
Vicia littoralis Jacq. =Tephrosia cinerea
Vigna repens (L.) Kuntze =Vigna luteola
Zollernia grandiflora Schery =Zollernia paraensis
Zollernia kanukuensis R.S. Cowan =Zollernia paraensis
Zollernia ulei Harms =Zollernia paraensis
Zornia marajoara Huber [nomen nudum] =Zornia guanipensis
Zornia pubescens Kunth =Zornia latifolia var. latifolia
Zornia surinamensis Miq. =Zornia latifolia var. latifolia
Zornia tenuifolia var. latifolia Benth. =Zornia guanipensis
References
Amshoff, G.J.H. 1939. Papilionaceae. In A. Pulle, Flora of Suriname 2(2): 1-257.
Aymard C, G. & N. Cuello A. 1991. Dos nuevas especies Neotropicales del genero Canavalia (Papilionoideae - Phaseoleae -
Diocleinae). Biollania 8: 87-92.
Aymard C, G.A., Nidia L. Cuello A., RE. Berry, V.E. Rudd, R.S. Cowan, RR. Fantz, R.H. Maxwell, C.H. Stirton, H.-H.
Poppendieck, H. Cavalcante de Lima, R.H. Fortunato, Basil Stergios, N. Xena de Enrich, D.A. Neill, R.T. Pennington, & C.
Gil. 1999. Fabaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 5: 231-
433.
Bameby, R.C. & S.V. Heald. 2002. Fabaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hofif, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 298-319.
Burkart, A. 1971. El genero Galactia (Legum.-Phaseoleae) en Sudamerica. Darwiniana 16: 663-796.
Cowan, R.S. 1961. Leguminosae. In B. Maguire, J.J. Wurdack, & collaborators. Botany of the Guayana Highland part IV. Mem.
New York Bot. Gard. 10(4): 65-87.
Ducke, A. 1942. Lonchocarpus, subgenus Phacelanthus Pittier, in Brazilian Amazonia. Trop. Woods 69: 2-7.
Dwyer, J.D. 1980. Leguminosae subfamily Papilionoideae (conclusion). In R.E. Woodson, Jr., R.W. Schery, & collaborators,
Flora of Panama. Ann. Missouri Bot. Gard. 67: 523-818.
Gorts-van Rijn, A.R.A., K.U. Kramer, & J.C. Lindeman. 1976. Papilionaceae (additions and corrections). In J. Lanjouw & A.L.
Stoffers, Flora of Suriname 2(2): 556-610.
Grear, J.W 1971. A revision of the American species of Eriosema. Mem. New York Bot. Gard. 20(3): 1-98.
Grear, J.W. 1978. A revision of the New World species of Rhynchosia. Mem. New York Bot. Gard. 31(1): 1-168.
Krukoff, B.A. 1969. Supplementary notes on the American species of Erythrina. III. Phytologia 19: 113-175.
Lima, H.C. de. 1982. Revisao taxonomica do genero Vatairea Aublet (Leguminosae - Faboideae). Arquiv. Jard. Bot. Rio de
Janeiro 26: 173-213.
Lima, H.C. de. 1985. Diplotropis Bentham. Estudo dos Taxons infragenericos. Acta Amaz. 15: 61-75.
Maxwell, R.H. 1970. The genus Cymbosema (Leguminosae): notes and distribution. Ann. Missouri Bot. Gard. 57(2): 252-257.
Mohlenbrock, R.H. 1958. A revision of the genus Stylosanthes. Ann. Missouri Bot. Gard. 44: 299-355.
Mohlenbrock, R.H. 1961. A monograph of the leguminous genus Zornia. Webbia 16(1): 1-141.
Mohlenbrock, R.H. 1963. Further considerations in Stylosanthes. Rhodora 65: 245-258.
Polhill, R.M. & P.H. Raven (ed.). 1981. Advances in Legume Systematics. Royal Botanical Garden, Kew, England.
Ramirez, N. 1995. Revision taxonomica del genero Alexa Moq. (Fabaceae, Sophoreae). Ann. Missouri Bot. Gard. 82(4): 549-
569.
Rojo, J. P. 1972. Pterocarpus. Revised for the world. Phanerog. Monogr. 5: 1-119.
Rudd, V.E. 1955. The American species ofAeschynomene. Contr. U.S. Nat. Herb. 32: 1-172.
Checklist of the Plants of the Guiana Shield 365
Rudd, V.E. 1955. A revision of the genus Chaetocalyx. Contr. U.S. Nat. Herb. 32: 247-277.
Rudd, V.E. 1963. The genus Dussia (Leguminosae). Contr. U.S. Nat. Herb. 32: 247-277.
Rudd, V.E. 1965. The American species of Ormosia. Contr. U.S. Nat. Herb. 32: 279-384.
Rudd, V.E. 1981. Two new species of Paramachaerium and a brief resume of the genus. Brittonia 33(3): 435-440.
Rudd, V.E. 1987. Studies in Machaerium (Leguminosae) VI. Phytologia 62: 282-302.
Rudd, V.E. 1989. Studies in Machaerium VII. Section II, Lineata. Part I. Species with wingless fruit. Phytologia 64: 1-12.
Sauer, J. 1964. Revision of Canavalia. Brittonia 16: 106-181.
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Biology, Agronomy, and Utilization.
Verdcourt, B. 1970. Studies in the Leguminosae - Papilionoideae for the 'Flora of Tropical East Africa': IV. Kew Bull. 24: 507-
569.
Williams, R.J. & R.R. Clements. 1990. Taxonomy of Centrosema. In R. Schultze-Kraft & R.J. Clements (ed.), Centrosema:
Biology, Agronomy, and Utilization.
Yakovlev, G.P. 1969. A review of Sweetia and Acosmium. Notes Royal Bot. Card. Edinb. 29: 347-355.
Leguminosae-Mimosoideae
R.C. Bameby, RE. Berry, D. Brunner, L. Cardenas, E. Forero, J.W. Grimes, H.C.F. Hopkins,
E.M. de Lamare Occhioni, G De Martino, O. Poncy
Abarema acreana (J.F. Macbr.) L. Rico
Abarema adenophora (Ducke) Bameby & J.W. Grimes
Abarema barbouriana (Standi.) Bameby & J.W. Grimes
Abarema barbouriana var. arenaria (Ducke) Bameby & J.W. Grimes
Abarema barbouriana var. barbouriana
Abarema commutata Bameby & J.W. Grimes
Abarema curvicarpa (H.S. Irwin) Bameby & J.W. Grimes
Abarema curvicarpa var. curvicarpa
Abaremaferruginea (Benth.) Pittier
Abaremafloribunda (Spmce ex Benth.) Bameby & J.W. Grimes
Abarema gallorum Bameby & J.W. Grimes
Abaremajupunba (Willd.) Britton & Killip
Abaremajupunba yds. jupunba
Abaremajupunba var. trapezifolia (Vahl) Bameby & J.W. Grimes
Abarema laeta (Benth.) Bameby & J.W. Grimes
Abarema leucophylla (Spruce ex Benth.) Bameby & J.W. Grimes AM
Abarema leucophylla var. leucophylla AM
Abarema levelii (R.S. Cowan) Bameby & J.W. Grimes AM
Abarema longipedunculata (H.S. Irwin) Bameby & J.W. Grimes [expected in BO
GU]
Abarema mataybifolia (Sandwith) Barneby & J.W. Grimes GU SU FG
Abarema microcalyx (Spmce ex Benth.) Bameby & J.W. Grimes AM
Abarema microcalyx var. microcalyx AM
Abarema villifera (Ducke) Barneby & J.W. Grimes AM BO SU
Acacia alemquerensis Huber [expected in the Guianas] BO
Acacia articulata Ducke BO SU
Acaciafarnesiana (L.) Wind. BO GU SU FG
Acaciafarnesiana var. farnesiana GU
Acaciafloribunda (Spruce ex Benth.) Barneby & J.W. Grimes GU
Acacia glomerosa Benth. BO
Acacia macracantha Humb. & Bonpl. ex Willd. [introduced] BO GU FG
Acacia podadenia (Britton & KiWip) L. Cardenas BO
Acacia polyphylla DC. GU
Acacia tamarindifolia (L.) Willd. BO
Acacia tenuifolia (L.) Willd.
Acacia tenuifolia var. producta J.W. Grimes
Acacia tenuifolia var. tenuifolia
Adenanthera pavonina L. [cultivated, naturalized]
Albizia barinensis L. Cardenas
Albizia glabripetala (H.S. Irwin) G.P Lewis & RE. Owen AM
Albizia lebbeck (L.) Benth. [cultivated, naturalized]
Albizia niopoides (Spruce ex Benth.) Burkart
Albizia subdimidiata (Splitg.) Barneby & J.W. Grimes
Albizia subdimidiata var. minor Bameby & J.W. Grimes
Albizia subdimidiata var. subdimidiata
AM BO
AM
AM
AM
AM
BO
BO
BO
GU
GU
GU
GU
GU
GU
GU?
GU?
SU
SU
FG
FG
FG
FG
FG
AM GU SU FG
AM FG
GU SU FG
AM SU FG
BO GU SU
SU
FG
FG
GU SU
SU
FG
BO GU
BO GU
GU SU FG
GU
BO DA GU
GU
GU
SU
SU
FG
366 Checklist of the Plants of the Guiana Shield
FG
FG
FG
GU
GU
GU
GU SU FG
GU su FG
GU FG
AM BO
AM BO
AM BO
AM
FG
FG
FG
Anadenanthera peregrina (L.) Speg. [natumlized?] AM BO GU SU
Anadenanthera peregrina var. peregrina AM BO GU SU
Balizia pedicellaris (DC.) Bameby& J.W. Grimes AM BO DA GU SU
Calliandra calothyrsus Meisn. [cultivated, perhaps escaping]
Calliandra coriacea (Humb. & Bonpl. ex Willd.) Benth. [expected in FG] AM GU SU
Calliandra cruegeri Griseb. AM BO GU
Calliandra glomerulata H. Karst. AM BO GU
Calliandra glomerulata var. glomerulata AM BO GU
Calliandra guildingii Benth. AM
Calliandra hymeniodes (Rich.) Benth. GU SU FG
Calliandra laxa (WiM.) Benth. AM BO GU
Calliandra laxa var. laxa BO
Calliandra laxa var. stipulacea (Benth.) Bameby AM BO GU
Calliandra laxa var. urimana Bameby BO
Calliandra pakaraimensis R.S. Cowan BO
Calliandra rigida Benth. • BO
Calliandra riparia Pittier BO
Calliandra surinamensis Benth. AM BO
Calliandra tergemina (L.) Benth. BO
Calliandra trinervia Benth. AM
Calliandra trinervia var. pilosifolia (R.S. Cowan) Bameby AM
Calliandra trinervia var. trinervia AM GU
Calliandra tsugoides R.S. Cowan AM BO
Calliandra vaupesiana R.S. Cowan AM
Calliandra vaupesiana var. oligandra Bameby AM
Cedrelinga cateniformis (Ducke) Ducke AM BO GU? SU
Chloroleucon acacioides (Ducke) Bameby & J.W. Grimes SU
Chloroleucon eurycyclum Bameby & J.W. Grimes BO
Chloroleucon mangense (Jacq.) Britton & Rose BO
Chloroleucon mangense var. tetrazyx Bameby & J.W. Grimes BO
Desmanthus pernambucanus (L.) Thell.
Desmanthus virgatus (L.) Willd. [expected in FG]
Dichrostachys glomerata (Forssk.) Chiov. [naturalized]
Entada polyphylla Benth. AM
Entada polystachya (L.) DC. BO
Enterolobium barinense L. Cardenas & H. Rodr. AM BO DA
Enterolobium barnebianum Mesquita & M.F. Silva [expected in Venezuelan
Guayana]
Enterolobium cyclocarpum (Jacq.) Griseb. [expected in Venezuelan Guayana]
Enterolobium oldemanii Bameby & J.W. Grimes
Enterolobium schomburgkii (Benth.) Benth.
Hydrochorea corymbosa (Rich.) Bameby & J.W. Grimes
Hydrochorea gonggrijpii (Kleinhoonte) Bameby & J.W. Grimes
Hydrochorea marginata (Spruce ex Benth.) Bameby & J.W. Grimes
Hydrochorea marginata var. panurensis (Spmce ex Benth.) Bameby & J.W. AM
Grimes
Hydrochorea marginata var. scheryi Bameby & J.W. Grimes AM
Inga acreana Harms
Inga acrocephala Steud.
Inga alata Benoist
Inga alba (Sw.) Willd. AM
Inga albicoria Poncy
Inga auristellae Harms AM
Inga bijuga Schery
Inga bourgonii (Aubl.) DC. AM
Inga brachystachys Ducke
Inga brevipes Benth.
Inga calanthoides Amshoff
Inga capitata Desv. AM
Inga cayennensis Sagot ex Benth. AM
Inga cinnamomea Spmce ex Benth.
Inga crassiflora Ducke
Inga cylindrica Mart.
Inga densiflora Benth. SU
GU SU
GU SU
FG
GU SU FG
GU SU FG
GU?
GU SU?
FG
GU SU FG
GU SU FG
GU SU FG?
SU FG
GU SU FG
GU FG
BO DA GU
GU
SU FG
FG
GU SU FG
BO
DA GU
GU
SU
SU
SU
SU
FG
FG
BO GU SU FG
BO GU SU
SU
FG
FG
FG
Checklist of the Plants of the Guiana Shield 367
Inga disticha Benth. GU SU FG
Inga ediilis Mart. AM BO DA GU SU FG
Ingafanchoniana Poncy GU? FG
Ingafastuosa (Jacq.) Willd. AM BO DA GU SU FG
Ingaflageinform is Mart. FG
Inga graciUflora Benth. GU SU FG
Inga gracilifolia Ducke AM BO DA GU FG
Inga heterophylla Willd. AM BO GU SU FG
Inga huberi Ducke DA GU SU FG
Inga inflata Ducke AM
Inga ingoides (Rich.) Willd. AM BO DA GU SU FG
IngaJava Pittier GU FG
Ingajenmanii Sandwith GU FG
Inga lateriflora Miq. AM BO DA GU SU FG
Inga laurina (Sw.) Willd. BO DA GU SU FG
Inga leiocalycina Benth. AM BO DA GU SU FG
Inga leptingoides Amshoff SU
Inga lomatophylla (Benth.) Pittier AM GU FG
Inga longipedunculata Ducke FG
Inga macrophylla Humb. & Bonpl. ex Willd. AM GU SU FG
Inga marginata Willd. AM GU SU FG
Inga melinonis Sagot DA GU FG
Inga micradenia Spruce ex Benth. AM BO
Inga multijuga Benth. BO
Inga multijuga subsp. multijuga BO
Inga neblinensis L. Cardenas & De Martino AM
Inga nobilis Willd. AM BO DA GU SU FG
Inga nobilis subsp. nobilis AM BO DA GU SU FG
Inga noiiragensis Poncy FG
Inga nubium Poncy FG
Inga nuda Salzm. ex Benth. GU SU FG
Inga paraensis Ducke BO GU FG
Inga pedunculata Ducke FG
Inga pezizifera Benth. AM BO DA GU SU FG
Inga pilosula (Rich.) J.F. Macbr. AM BO GU SU FG
Inga poeppigiana Benth. FG
Inga retinocarpa Poncy FG
Inga rhynchocalyx Sandwith GU FG
Inga rubiginosa (Rich.) DC. AM BO DA GU SU FG
Inga salicifolia T.D. Penn. [expected in AM, Serra da Neblina]
Inga sapindoides Willd. BO DA FG
Inga sarmentosa Glaz. ex Harms GU SU FG
Inga sertulifera DC. AM BO GU SU FG
Inga sertulifera subsp. leptopus (Benth.) T.D. Penn. GU
Inga sertulifera subsp. sertulifera AM BO GU SU FG
Inga splendens Willd. AM BO DA GU SU FG
Inga stenocalyx Spruce ex Benth. GU FG
Inga stenoptera Benth. AM BO DA GU
Inga stipularis DC. AM BO GU SU FG
Inga striata Benth. [=1. nuda?] FG
Inga suaveolens Ducke FG
Inga thibaudiana DC. AM BO GU SU FG
Inga thibaudiana subsp. thibaudiana AM BO GU SU FG
Inga tubiformis Benoist FG
Inga ulei Harms AM
Inga umbellifera (Vahl) Steud. ex DC. AM BO DA GU SU FG
Inga umbratica Poepp. BO FG
Inga venusta Standi. SU
Inga vera Willd. AM BO DA GU SU FG
Inga vera subsp. affinis (DC.) T.D. Penn. AM BO GU SU FG
Inga vera subsp. vera BO DA
Inga virgultosa (Vahl) Desv. SU FG
Leucaena leucocephala (Lam.) de Wit [cuhivated, naturalized] BO GU SU FG
Leucaena leucocephala subsp. glabrata (Rose) S. Zarate BO
368 Checklist of the Plants of the Guiana Shield
Lysiloma bahamensis Benth. [introduced, dubiously persisting]
Macrosamanea amplissima (Ducke) Bameby & J.W. Grimes
Macrosamanea consanguinea (R.S. Cowan) Bameby & J.W. Grimes
Macrosamanea discolor (Humb. & Bonpl. ex Willd.) Britton & Rose ex
Britton & Killip
Macrosamanea kegelii (Meisn.) Kleinhoonte
Macrosamanea pubiramea (Steud.) Bameby & J.W. Grimes
Macrosamanea pubiramea van lindsaeifolia (Spmce ex Benth.) Bameby &
J.W. Grimes
Macrosamanea simabifolia (Spmce ex Benth.) Pittier
Macrosamanea spruceana (Benth.) Killip
Mimosa annularis Benth.
Mimosa annularis var. odora Bameby
Mimosa annularis var. xinguensis (Ducke) Bameby
Mimosa arenosa (Willd.) Poir.
Mimosa arenosa var. arenosa
Mimosa bimucronata (DC.) Kuntze [cuhivated or escaped from cultivation]
Mimosa brachycarpoides Bameby
Mimosa camporum Benth.
Mimosa casta L. [possibly introduced in FG]
Mimosa colombiana Britton & Killip
Mimosa corynadenia Britton & Rose [expected in SW Venezuelan Guayana]
Mimosa debilis Humb. & Bonpl. ex Willd.
Mimosa debilis var. debilis
Mimosa diplotricha C. Wright ex Sauv.
Mimosa dormiens Humb. & Bonpl. ex Willd.
Mimosa guanchezii Bameby
Mimosa guilandinae (DC.) Bameby
Mimosa guilandinae var. duckei (Huber) Bameby
Mimosa guilandinae var. guilandinae
Mimosa hirsutissima Mart.
Mimosa huberi Bameby
Mimosa invisa Mart, ex Colla [expected in BO]
Mimosa invisa var. spiciflora (H. Karst.) Bameby [expected in BO]
Mimosa microcephala Humb. & Bonpl. ex Willd.
Mimosa microcephala var. cataractae (Ducke) Bameby
Mimosa microcephala var. communis Bameby
Mimosa microcephala var. lumaria Bameby
Mimosa microcephala var. microcephala
Mimosa myriadenia (Benth.) Benth.
Mimosa myriadenia var. dispersa Bameby
Mimosa myriadenia var. myriadenia
Mimosa orinocoensis Bameby
Mimosa orthocarpa Spmce ex Benth.
Mimosa pigra L.
Mimosa piscatorum Bameby
Mimosa polydactyla Humb. & Bonpl. ex Willd.
Mimosa pudica L.
Mimosa pudica var. hispida Brenan
Mimosa pudica var. pastoris Bameby
Mimosa pudica var. tetrandra (Humb. & Bonpl. ex Willd.) DC.
Mimosa pudica var. unijuga (Walp. & Duchass.) Griseb.
Mimosa quadrivalvis L.
Mimosa quadrivalvis var. leptocarpa (DC.) Bameby
Mimosa rufescens Benth. [expected in SU, FG]
Mimosa rufescens var. amnis-nigri Bameby
Mimosa rufescens var. rufescens
Mimosa scabrella Benth. [occasionally escapes cultivation]
Mimosa schomburgkii Benth.
Mimosa schrankioides Benth.
Mimosa schrankioides var. sagotiana (Benth.) Bameby
Mimosa schrankioides var. schrankioides
Mimosa sensitiva L.
Mimosa somnians Humb. & Bonpl. ex Willd.
AM
AM
AM
AM
AM
AM
BO
BO
GU
DA
SU
SU
FG
FG
FG
GU SU FG
GU SU FG
FG
BO
BO
BO
AM BO DA
DA
GU SU FG?
FG
AM BO
BO GU? SU FG
BO
GU SU FG
AM
BO DA GU?
SU
SU
FG
FG
FG
AM BO
AM BO
GU
GU
SU?
SU?
AM BO GU SU
SU
FG
FG
AM BO
AM BO GU FG?
AM
AM BO DA GU SU
SU
FG
FG
GU SU FG
AM DA
AM BO
AM BO
BO
DA GU SU FG
AM BO DA GU SU FG
AM BO GU
GU
GU
SU
SU
FG
AM BO GU SU FG
AM BO GU SU FG
AM BO DA GU FG
AM BO DA GU FG
AM
A \/f
BO DA GU
AM
AM BO DA
BO
BO GU FG
AM BO GU SU FG
AM BO GU SU FG
BO GU
AM BO
AM BO GU FG
Checklist of the Plants of the Guiana Shield 369
Mimosa somnians var. deminuta Bameby
Mimosa somnians var. somnians
Mimosa somnians var. viscida (Willd.) Bameby
Mimosa surumuensis Harms
Mimosa tarda Bameby
Mimosa velloziana Mart, [weedy, of doubtful native range]
Mimosa xanthocentra Mart, [expected in BO]
Mimosa xanthocentra var. xanthocentra [expected in BO]
Neptunia oleracea Lour, [expected in BO]
Neptunia plena (L.) Benth.
Parkia barnebyana H.C. Hopkins
Parkia decussata Ducke
Parkia discolor Spmce ex Benth.
Parkia gigantocarpa Ducke
Parkia igneiflora Ducke
Parkia nitida Miq.
Parkia panurensis Benth. ex H.C. Hopkins
Parkia pendula (Willd.) Benth. ex Walp.
Parkia reticulata Ducke
Parkia truncata R.S. Cowan
Parkia ulei (Harms) Kuhlm.
Parkia ulei var. surinamensis Kleinhoonte
Parkia velutina Benoist [expected in GU]
Pentaclethra macroloba (Willd.) Kuntze
Piptadeniafloribunda Kleinhoonte
Piptadenia imatacae Bameby [expected in GU]
Piptadenia leucoxylon Bameby & J.W. Grimes
Piptadenia moniliformis Benth. [expected in the Guianas]
Piptadenia uaupensis Spmce ex Benth.
Piptadenia viridiflora (Kunth) Benth.
Pithecellobium consanguineum R.S. Cowan
Pithecellobium dulce (Roxb.) Benth. [cultivated, naturalized]
Pithecellobium roseum (Vahl) Bameby & J.W. Grimes
Pithecellobium unguis-cati (L.) Benth. [cultivated, naturalized]
Plathymenia reticulata Benth.
Pseudopiptadenia psilostachya (DC.) GR Lewis & M.R Lima
Pseudopiptadenia suaveolens (Miq.) J.W. Grimes
Pseudosamanea guachapele (Kunth) Harms
Samanea inopinata (Harms) Bameby & J.W. Grimes
Samanea soman (Jacq.) Mem [cuUivated, perhaps escaping]
Stryphnodendron guianense (Aubl.) Benth.
Stryphnodendron guianense subsp. guianense
Stryphnodendron levelii R.S. Cowan
Stryphnodendron microstachyum Poepp.
Stryphnodendron moricolor Bameby & J.W. Grimes
Stryphnodendron polystachyum (Miq.) Kleinhoonte
Stryphnodendron pulcherrimum (Willd.) Hochr.
Zapotecaformosa (Kunth) H.M. Hem.
Zapotecaformosa snhs^. formosa
Zapoteca portoricensis (Jacq.) H.M. Hem.
Zapoteca portoricensis suhsp.flavida (Urb.) H.M. Hem.
Zygia ampla (Benth.) Pittier
Zygia basijuga (Ducke) Bameby & J.W. Grimes
Zygia cataractae (Kunth) L. Rico
Zygia claviflora (Spmce ex Benth.) Bameby & J.W. Grimes
Zygia collina (Sandwith) Bameby & J.W. Grimes
Zygia eperuetorum (Sandwith) Barneby & J.W. Grimes
Zygia inaequalis (Humb. & Bonpl. ex Willd.) Pittier
Zygia inundata (Ducke) H.C. Lima
Zygiajuruana (Harms) L. Rico
Zygia latifolia (L.) Fawc. & Rendle
Zygia latifolia var. communis Bameby & J.W. Grimes
Zygia latifolia var. lasiopus (Benth.) Bameby & J.W. Grimes
Zygia latifolia var. latifolia
AM BO
BO GU FG
GU FG
BO GU
AM
AM
AM
BO DA
BO
GU
GU
SU
su
FG
FG
SU FG
AM
GU
AM GU
AM BO DA GU SU FG
AM
AM BO GU su FG
FG
AM
DA GU SU FG
DA GU SU FG
BO SU FG
BO DA GU SU
SU
FG
FG
BO
BO GU
BO
FG?
GU SU? FG
BO GU
BO GU
SU
FG
GU SU FG
AM BO
BO
BO
DA GU
GU
GU
SU FG
BO GU SU FG
BO GU FG
AM
AM BO DA GU
GU
SU FG
FG
BO DA GU SU FG
BO GU SU FG
BO
BO
BO DA GU SU FG
BO DA GU
GU
BO
AM BO DA GU SU FG
AM GU SU FG
AM BO DA GU
GU
AM BO GU
GU
SU FG
FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU FG
DA GU SU FG
370 Checklist of the Plants of the Guiana Shield
Zygia palustris Bameby & J.W. Grimes
Zygia potawensis Bameby & J.W. Grimes
Zygia racemosa (Ducke) Bameby & J.W. Grimes
Zygia ramiflora (Benth.) Barneby & J.W. Grimes
Zygia sabatieri Bameby & J.W. Grimes
Zygia tetragona Bameby & J.W. Grimes
Zygia unifoUolata (Benth.) Pittier [expected in GU]
GU
BO GU
BO GU SU FG
FG
FG
AM BO
Synonyms and Misidentifications
Abarema claviflora (Spruce ex Benth.) Kleinhoonte =Zygia claviflora
Abarema racemosa (Ducke) Kleinhoonte =Zygia racemosa
Abarema trapezifolia (Vahl) Pittier =Abaremajupunba var. trapezifolia
Acacia alvaroi Cardenas & De Martino =Acacia alemquerensis
Acacia arenosa Willd. =Mimosa arenosa
Acacia chrysantha DC. [name of uncertain application]
Acaciaflexuosa Humb. & Bonpl. ex Willd. =Acacia macracantha
Acaciaformosa Kunth =Zapotecaformosa
Acacia glandulosa (Vahl) DC. [nomen nudum]
Acacia guachapele Kunth =Pseiidosamanea guachapele
Acacia guianensis (Aubl.) Willd. =Stryphnodendron guianense
Acaciajupunba Willd. =Abaremajupunba
Acacia laxa Willd. =Calliandra laxa
Acacia leucocephala (Lam.) Link =Leucaena leucocephala
Acacia lutea (Mill.) Britton =Acacia macracantha
Acacia macroloba Willd. =Pentaclethra macroloba
Acacia ouyrarema DC. [nomen nudum]
Acacia paniculata Willd. =Acacia tenuifolia
Acacia paniculiflora Steud. =Mimosa myriadenia
Acacia psilostachya DC. =Pseudopiptadenia psilostachya
Acacia pulcherrima Willd. =Stryphnodendron pulcherrimum
Acacia subdimidiata Splitg. =AIbizia subdimidiata
Acacia viridiflora Kunth ^Piptadenia viridiflora
Acuan virgatum (L.) W. Medicus =Desmanthus virgatus
Albizia caribaea (Urb.) Britton & Rose =Albizia niopoides
Albizia corymbosa (Rich.) GP. Lewis & RE. Owen =Hydrochorea corymbosa
Albizia guachapele (Kunth) Dugand =Pseudosamanea guachapele
Albizia inopinata (Harms) GP. Lewis =Samanea inopinata
Albizia longepedata (Pittier) Britton & Rose =Pseudosamanea guachapele
Anneslia calothyrsa (Meisn.) Kleinhoonte =Calliandra calothyrsus
Annesliafasciculata (Willd.) Kleinhoonte =Calliandra surinamensis
Anneslia tergemina (L.) Britton & Rose =Calliandra tergemina
Arthrosamanea corymbosa (Rich.) Kleinhoonte =Hydrochorea corymbosa
Arthrosamanea gonggrijpii (Kleinhoonte) Kleinhoonte =Hydrochorea gonggrijpii
Arthrosamanea multiflora (Kunth) Kleinhoonte [sensu Kleinhoonte] =Albizia subdimidiata
Arthrosamanea panurensis (Spmce ex Benth.) Pittier =Hydrochorea marginata var. panurensis
Arthrosamanea pullei (Kleinhoonte) Kleinhoonte =Hydrochorea gonggrijpii
Calliandra affinis Pittier =Calliandra cruegeri
Calliandra anthoniae J.W. Grimes =Calliandra coriacea
Calliandra caripensis Willd. =Calliandra tergemina
Calliandraflavida Urb. =Zapoteca portoricensis subsp. flavida
Calliandraformosa (Kunth) Benth. =Zapotecaformosa
Calliandra latifolia (L.) Griseb. =Zygia latifolia
Calliandra marginata Griseb. ex R.O. Williams =Zapotecaformosa subsp. formosa
Calliandra patrisii Sagot ? ^Calliandra hymeniodes
Calliandra pilosifolia R.S. Cowan =Calliandra trinervia var. pilosifolia
Calliandra portoricensis (Jacq.) Benth. =Zapoteca portoricensis
Calliandra resupina R.S. Cowan =Calliandra pakaraimensis
Calliandra stipulacea Benth. =Calliandra laxa var. stipulacea
Calliandra tenuiflora Benth. =Calliandra surinamensis
Calliandra trijugata Schery =Calliandra laxa var. laxa
Cathormion corymbosum (Rich.) Burkart =Hydrochorea corymbosa
Entada gigas Fawc. & Rendle [not in flora area]
Entada myriadenia Benth. =Mimosa myriadenia
Entadopsis polyphylla (Benth.) Britton =Entada polyphylla
Checklist of the Plants of the Guiana Shield 371
Entadopsis polystachya (L.) Britton =Entada polystachya
Enterolobium racemosum Ducke [apparently a nomen nudum]
Feuillea sapindoides (Willd.) Kuntze =Inga sapindoides
Feuilleea cayennensis (Sagot ex Benth.) Kuntze =Inga cayennensis
Feuilleea edulis (Mart.) Kuntze =Inga edulis
Feuilleea fagifolia (L.) Kuntze =Inga laurina
Feuilleea heterophylla (Willd.) Kuntze =Inga heterophylla
Feuilleea ingoides (Rich.) Kuntze =Inga ingoides
Feuilleea laurina (Sw.) Kuntze =Inga laurina
Feuilleea lindeniana (Benth.) Kuntze =Inga sapindoides
Feuilleea marginata (Willd.) Kuntze =Inga marginata
Feuilleea myriantha (Poepp.) Kuntze =Inga umbellifera
Feuilleea pezizifera (Benth.) Kuntze =Inga pezizifera
Feuilleea pilosula (Rich.) Kuntze =Inga pilosula
Feuilleea sciadon (Steud.) Kuntze =Inga umbellifera
Feuilleea setifera (DC.) Kuntze =Inga pilosula
Feuilleea splendens (Willd.) Kuntze =Inga splendens
Feuilleea umbellifera (Vahl) Kuntze ^Inga umbellifera
Feuilleea umbratica (Poepp.) Kuntze =Inga umbratica
Folianthera guianensis (Aubl.) Raf. =Stryphnodendron guianense
Hydrochorea acreana (J.F. Macbr.) Bameby & J.W. Grimes =Abarema acreana
Inga acuminata Benth. [material misidentified ] = /. rhynchocalyx
Inga adiantifoHa Kunth =Macrosamanea discolor
Inga affinis DC. =Inga vera subsp. affinis
Inga affinis Steud. =Inga pilosula
Inga aggregata G. Don =Inga alba
Inga amazonica L. Cardenas =Inga lomatophylla
Inga amazonica var. bracteifera (Ducke) L. Cardenas =Inga lomatophylla
Inga amazonica var. lomatophylla (Benth.) L. Cardenas [nom. superfl.] =Inga lomatophylla
Inga amazonica var. membranacea (Ducke) L. Cardenas =Inga lomatophylla
Inga angustifolia Willd. [not in flora area]
Inga antioquensis Britton & Killip =Inga sapindoides
Inga brachyptera Benth. =Inga macrophylla
Inga bracteosa Benth. =Inga macrophylla
Inga calocephala Poepp. =Inga macrophylla
Inga camuriensis Pittier =Inga sapindoides
Inga capitata Miq. ex Benth. =Abarema laeta
Inga caracasana Pittier =Inga sapindoides
Inga carachensis Pittier =Inga alba
Inga cataractae Kunth =Zygia cataractae
Inga cauliflora Willd. =Zygia latifolia var. communis
Inga commewijnensis Miq. =Inga nuda
Inga conglomerata Benoist =Inga nobilis
Inga coriacea Humb. & Bonpl. ex Willd. =Calliandra coriacea
Inga coriacea (Pers.) Desv. =Inga sertulifera subsp. sertulifera
Inga corymbifera Benth. =Inga nobilis
Inga crevauxii Sagot =Inga disticha
Inga cyclocarpa Ducke =Inga melinonis
Inga discolor Humb. & Bonpl. ex Willd. =Macrosamanea discolor
Inga disticha var. negrensis (Spruce ex Benth.) Ducke =Inga micradenia
Inga dysantha Benth. =Inga cayennensis
Inga fagifolia (L.) Willd. ex Benth. =Inga laurina
Ingafloribunda Benth. =Inga splendens
Ingaforfex Kunth =Pithecellobium roseum
Inga gladiata Desv. =Inga thibaudiana subsp. thibaudiana
Inga glomerata DC. =Zygia cataractae
Inga grandifolia Pittier =Inga sapindoides
Inga guaremalensis Pittier =Inga fastuosa
Inga inaequalis Humb. & Bonpl. ex Willd. =Zygia inaequalis
Inga inga (Veil.) J. Moore =Inga edulis
Inga laeta (Benth.) Poepp. =Abarema laeta
Inga latifolia (L.) Willd. =Zygia latifolia
Inga lawrenceana Britton & Killip =Inga umbellifera
Inga lindeniana Benth. =Inga sapindoides
372 Checklist of the Plants of the Guiana Shield
Inga longiflora Spruce ex Benth. [not in flora area]
Inga macradenia Mart, ex Benth. [nomen nudum] =Inga thibaudiana subsp. thibaudiana
Inga macrophylla Hook. =Inga pilosula
Inga meissneriana Miq. =Inga vera subsp. affinis
Inga microphylla Humb. & Bonpl. ex Willd. =Pithecellobium unguis-cati
Inga multiflora Benth. =Inga leiocalycina
Inga myriantha Poepp. =Inga iimbellifera
Inga negrensis Spruce ex Benth. =Inga micradenia
Inga niopo Humb. & Bonpl. ex Willd. =Anadenanthera peregrina
Inga nitida Willd. =Inga pilosula
Inga panurensis Spruce ex Benth. [not in flora area]
Inga pedicellaris DC. =Balizia pedicellaris
Inga pendula Willd. =Parkia pendula
Inga perrottetii Sagot =Inga nuda
Inga pilosiuscula Desv. =Inga pilosula
Inga prieurei Sagot =Inga nuda
Inga protracta Steud. =Inga hetewphylla
Inga pterocarpa DC. (Leguminosae-Caesalpinioideae) =Peltophonim pterocarpum
Inga pubescens DC. =Pithecellobium roseum
Inga pubiramea Steud. =Macrosamanea pubiramea
Inga racemiflora Ducke =Inga ulei
Inga ramiflora Steud. =Zygia latifolia var. lasiopus
Inga ricardorum Bemardi & Spichiger [Guianas material identified as IngaJava]
Inga riparia Pittier =Inga vera subsp. vera
Inga rosea (Vahl) Steud. =Pithecellobium roseum
Inga scabriuscula Benth. =Inga edulis
Inga sciadion Steud. =Inga umbellifera
Inga sciandion Steud. =Inga umbellifera
Inga semialata (Veil.) Mart. =Inga marginata
Inga setifera DC. =Inga pilosula
Inga speciosa Spruce ex Benth. =Inga lomatophylla
Inga speciosa var. bracteifera Ducke =Inga lomatophylla
Inga speciosa var. lomatophylla Benth. =Inga lomatophylla
Inga speciosa var. membranacea Ducke =Inga lomatophylla
Inga spuria Humb. & Bonpl. ex Willd. =Inga vera subsp. vera
Inga strigillosa Spruce ex Benth. =Inga leiocalycina
Inga tenuiflora Salzm. ex Benth. =Inga thibaudiana subsp. thibaudiana
Inga umbellata G. Don =Inga heterophylla
Inga unguis-cati (L.) Willd. =Pithecellobium unguis-cati
Inga unijuga Poepp. & Endl. (Leguminosae-Caesalpinioideae) =Macrolobium limbatum var. limbatum
Inga urnifera Kleinhoonte =Inga pezizifera
Inga velutina (Poir.) Willd. [not in flora area]
Inga venosa Griseb. ex Benth. =Ingafastuosa
Inga vera Kunth =Inga edulis
Jupunba trapezifolia (Vahl) Moldenke =AbaremaJupunba var. trapezifolia
Klugiodendron laetum (Benth.) Britton & Killip =Abarema laeta
Leptoglottis leptocarpa (DC.) Standi. =Mimosa quadrivalvis var. leptocarpa
Leucaena glabrata Rose =Leucaena leucocephala subsp. glabrata
Leucaena glauca (L.) Benth. [not in flora area]
Leucaena ulei Harms =Parkia ulei
Lysiloma guachapele (Kunth) Benth. =Pseudosamanea guachapele
Macrosamanea aquatica Pittier =Macrosamanea discolor
Macrosamanea basijuga (Ducke) Dugand =Zygia basijuga
Macrosamanea pedicellaris (DC.) Kleinhoonte =Balizia pedicellaris
Manganaroa alemquerensis (Huber) Speg. =Acacia alemquerensis
Manganaroa articulata (Ducke) Speg. =Acacia articulata
Marmaroxylon claviflorum (Benth.) L. Rico =Zygia claviflora
Marmaroxylon collinum (Sandwith) L. Rico =Zygia collina
Marmaroxylon eperuetorum (Sandwith) L. Rico =Zygia eperuetorum
Marmaroxylon racemosum (Ducke) Killip ex Record =Zygia racemosa
Marmaroxylon ramiflorum (Benth.) L. Rico =Zygia ramiflora
Mimosa acacioides Benth. [cuhivated] =Anadenanthera peregrina (L.) Speg.
Mimosa adhaerens Kunth =Mimosa debilis
Mimosa alba Sw. =Inga alba
Checklist of the Plants of the Guiana Shield 373
Mimosa asperata auct. non L. 1759. =Mimosa pigra
Mimosa asperata L. [not in flora area]
Mimosa baiihiniaefolia H. Karst. =Mimosa colombiana
Mimosa bipinnata Aubl. =Entada polystachya
Mimosa bonrgoni Aubl. =Inga boiirgonii
Mimosa bourgonii Aubl. =Inga bourgonii
Mimosa brevispica Harms =Mimosa invisa var. spiciflora
Mimosa cataractae Ducke =Mimosa microcephala var. cataractae
Mimosa cauliflora (Willd.) Poir. =Zygia latifolia var. communis
Mimosa ceratonia L. [not in flora area]
Mimosa coriacea Pers. =Inga sertulifera subsp. sertulifera
Mimosa corymbosa Rich. =Hydrochorea corymbosa
Mimosafagifolia L. =Inga laurina
Mimosafagifolia Jacq. =Inga laurina
Mimosafarnesiana L. =Acaciafarnesiana
Mimosafasciculata var. ernestiana Kuntze =Mimosa arenosa
Mimosafastuosa Jacq. =Ingafastuosa
Mimosaflavescens Splitg. =Mimosa camporum
Mimosa flaviseta Benth. =Mimosa camporum
Mimosaflexuosa (Humb. & Bonpl. ex Willd.) Poir. =Acacia macracantha
Mimosa guianensis Aubl. =Stryphnodendron guianense
Mimosa hispidida Kunth =Mimosa pudica var. tetrandra
Mimosa hostmannii Benth. =Mimosa debilis
Mimosa humilis Humb. & Bonpl. ex Willd. =Mimosa dormiens
Mimosa hymenaeodes Rich. =Calliandra hymeniodes
Mimosa inga Veil. =Inga edulis
Mimosa inga L. =Inga vera
Mimosa ingoides Rich. =Inga ingoides
Mimosa intermedia Kunth =Mimosa dormiens
Mimosajiramenensis H. Karst. =Mimosa velloziana
Mimosa latifolia L. =Zygia latifolia
Mimosa laurina Sw. =Inga laurina
Mimosa leucocephala Lam. =Leucaena leucocephala
Mimosa lucida Vahl =Inga pilosula
Mimosa lutea Mill. =Acacia macracantha
Mimosa macracantha (Humb. & Bonpl. ex Willd.) Poir. =Acacia macracantha
Mimosa macrophylla (Humb. & Bonpl. ex Willd.) Poir. =Inga macrophylla
Mimosa mangensis Jacq. =Chloroleucon mangense
Mimosa martensis Britton & Killip =Mimosa camporum
Mimosa micracantha Benth. =Mimosa rufescens var. amnis-nigri
Mimosa micrantha var. plurijuga Ducke =Mimosa rufescens var. rufescens
Mimosa microcephala Humb. & Bonpl. ex Willd. =Mimosa microcephala var. microcephala
Mimosa multijuga (Benth.) Kuntze =Inga multijuga
Mimosa natans L. f. =Neptunia oleracea
Mimosa nitida (Willd.) Poir. =Inga pilosula
Mimosa notata Steud. =Mimosa debilis
Mimosa obovata Benth. [not in flora area] =Mimosa laticifera Rizzini & Mattos Filho
Mimosa palpitans Humb. & Bonpl. ex Willd. =Mimosa somnians
Mimosa panamensis (Benth.) Standi. =Mimosa debilis
Mimosa paniculata (Willd.) Poir. =Acacia tenuifolia
Mimosa paniculata Benth. =Mimosa annularis var. odora
Mimosa parae Poir. =Inga heterophylla
Mimosa pellita Humb. & Bonpl. ex Willd. =^Mimosa pigra
Mimosa peregrina L. =Anadenanthera peregrina
Mimosa pigra auct. non L. 1755. =Mimosa pigra
Mimosa pilosula Rich. =Inga pilosula
Mimosa plena L. =Neptunia plena
Mimosa plumiifolia Kleinhoonte =Mimosa microcephala var. cataractae
Mimosa polystachia L. =Entada polystachya
Mimosa portoricensis Jacq. =Zapoteca portoricensis
Mimosa prostrata Lam. [nom. illeg.] =Neptunia oleracea
Mimosa pulcherrima (Willd.) Poir. =Stryphnodendron pulcherrimum
Mimosa pusilla Benth. =Mimosa camporum
Mimosa rosea Vahl =Pithecellobium roseum
374 Checklist of the Plants of the Guiana Shield
Mimosa rubiginosa Rich. =Inga nibiginosa
Mimosa sagotiana Benth. =Mimosa schrankioides var. sagotiana
Mimosa saman Jacq. =Samanea saman
Mimosa santanderensis Britton & Rose ex Britton & Killip =Mimosa schrankioides var. sagotiana
Mimosa sapindoides (Willd.) Poir. =Inga sapindoides
Mimosa somniculosa Kunth ^'Mimosa somnians
Mimosa spiciflora H. Karst. =Mimosa invisa var. spiciflora
Mimosa splendens (Willd.) Poir. =Inga splendens
Mimosa spuria (Humb. & Bonpl. ex Willd.) Poir. =Inga vera subsp. vera
Mimosa tamarindifolia L. =Acacia tamarindifolia
Mimosa tergemina L. =Calliandra tergemina
Mimosa tetrandra Humb. & Bonpl. ex Willd. =Mimosa pudica var. tetrandra
Mimosa tobagensis Urb. =Mimosa somnians
Mimosa tomentosa Humb. & Bonpl. ex Willd. =Mimosa hirsutissima
Mimosa trapezifolia Vahl =Abaremajupimba var. trapezifolia
Mimosa umbellifera Vahl =Inga umbellifera
Mimosa unguis-cati L. =Pithecellobiiim unguis-cati
Mimosa unijuga Walp. & Duchass. =Mimosa pudica var. unijuga
Mimosa velloziana war.jiramenensis (H. Karst.) Benth. =Mimosa velloziana
Mimosa viva Veil. [nom. illeg.] =Mimosa velloziana
Mimosa viva L. [not in flora area]
Mimosa xantholasia Benth. =Mimosa arenosa
Mimosa xinguensis Ducke =Mimosa annularis var. xinguensis
Neptunia natans (L. f.) Druce =Neptunia oleracea
Neptunia prostrata (Lam.) Baill. =Neptuma oleracea
Neptunia surinamensis Steud. =Neptunia plena
Newtonia psilostachya (DC.) Brenan =Pseudopiptadenia psilostachya
Newtonia suaveolens (Miq.) Brenan =Pseudopiptadenia suaveolens
Niopa peregrina (L.) Britton & Rose =Anadenanthera peregrina
Parkia auriculata Spruce ex Benth. =Parkia discolor
Parkia oppositifolia Spruce ex Benth. =Parkia nitida
Parkia pectinata auct. non Benth. 1875: sensu Benth. =Parkia panurensis
Pentaclethrafilamentosa Benth. =Pentaclethra macroloba
Piptadenia biuncifera Benth. =Piptadenia viridiflora
Piptadenia cateniformis Ducke =Cedrelinga cateniformis
Piptadenia guianensis (Aubl.) Benth. =Stryphnodendron guianense
Piptadenia niopo (Humb. & Bonpl. ex Willd.) Spruce =Anadenanthera peregrina
Piptadenia peregrina (L.) Benth. =Anadenanthera peregrina
Piptadenia polystachya Miq. =Stryphnodendron polystachyum
Piptadenia psilostachya (DC.) Benth. =Pseudopiptadenia psilostachya
Piptadenia rubescens Pittier ^Piptadenia viridiflora
Piptadenia speciosa Britton & Killip =Piptadema viridiflora
Piptadenia suaveolens Miq. =Pseudopiptadenia suaveolens
Piptadenia tocantina Ducke =Stryphnodendron polystachyum
Pithecellobium acacioides Ducke =Chloroleucon acacioides
Pithecellobiiim adiantifolium (Kunth) Benth. [not in flora area] =Macrosamanea discolor
Pithecellobium amplissimum Ducke =Macrosamanea amplissima
Pithecellobium aquaticum (Pittier) R.S. Cowan ^Macrosamanea discolor
Pithecellobium arenarium Ducke =Abarema barbouriana var. arenaria
Pithecellobium arenicola R.S. Cowan =Macrosamanea discolor
Pithecellobium barbourianum Standi. =Abarema barbouriana
Pithecellobium basijugum Ducke [not in flora area] =Zygia basijuga
Pithecellobium bijugatum Schery =Hydrochorea marginata var. scheryi
Pithecellobium catenaeformis (Ducke) L. Cardenas =Cedrelinga cateniformis
Pithecellobium cauliflorum (Willd.) Mart. =Zygia latifolia var. communis
Pithecellobium collinum Sandwith =Zygia collina
Pithecellobium corymbosum (Rich.) Benth. =Hydrochorea corymbosa
Pithecellobium curvicarpum H.S. Irwin =Abarema curvicarpa
Pithecellobium dinizii Ducke =Zygia ramiflora
Pithecellobium discolor (Humb. & Bonpl. ex Willd.) J.F. Macbr. =Macrosamanea discolor
Pithecellobium divaricatum Benth. =Zygia cataractae
Pithecellobium elegans Ducke [not in flora area] ^Balizia elegans (Ducke) Bameby & Grimes
Pithecellobium eperuetorum Sandwith =Zygia eperuetorum
Pithecellobiumfanshawei Sandwith =Abarema barbouriana
Checklist of the Plants of the Guiana Shield 375
Pithecellobiutn fernigineiim Benth. =Abaremaferniginea
PUhecellobium floribundiim Spruce ex Benth. =Abarema floribunda
Pithecellobiiim forfex (Kunth) Benth. =PitheceUobhim roseum
Pithecellobium glabripetahm H.S. Irwin =Albizia glabripetala
PUhecellobium glomeratum (DC.) Benth. ^Zygia cataractae
Pithecellobium gonggrijpii Kleinhoonte =Hydrochorea gonggrijpii
Pithecellobium guachapele (Kunth) J.F. Macbr. =Pseudosamanea guachapele
Pithecellobium guaricense Pittier =PitheceUobium roseum
Pithecellobium huberi Ducke =Zygia latifolia
Pithecellobium inaequale (Humb. & Bonpl. ex Willd.) Benth. =Zygia inaequalis
Pithecellobium inopinatum (Harms) Ducke =Samanea inopinata
Pithecellobiumjupunba (Willd.) Urb. =Abaremajupunba
Pithecellobium kegelii Meisn. =Macrosamanea kegelii
Pithecellobium laetum Benth. =Abarema laeta
Pithecellobium laetum Benth. =Abarema laeta
Pithecellobium lanceolatum Benth. [not in flora area]
Pithecellobium larensis L. Cardenas =PitheceUobium unguis-cati
Pithecellobium lasiopus Benth. =Zygia latifolia var. lasiopus
Pithecellobium latifoliimt (L.) Benth. =Zygia latifolia
Pithecellobium leucophyllum Spruce ex Benth. =Abarema leucophylla
Pithecellobium levelii R.S. Cowan =Abarema levelii
Pithecellobium lindseifolium Spruce ex Benth. =Macrosamanea pubiramea var. lindsaeifolia
Pithecellobium longepedatum Pittier =Pseudosamanea guachapele
Pithecellobium longipedunculatum H.S. Irwin =Abarema longipedunculata
Pithecellobium mangense (Jacq.) J.F. Macbr. [not in flora area] =Chloroleucon mangense (Jacq.) Britton & Rose
Pithecellobium marginatum Spruce ex Benth. =Hydrochorea marginata
Pithecellobium mataybifolium Sandwith =Abarema mataybifolia
Pithecellobium microcalyx Spruce ex Benth. =Abarema microcalyx
Pithecellobium microchlamys Pittier =PitheceUobium unguis-cati
Pithecellobium microphyllum (Humb. & Bonpl. ex Willd.) Benth. =PitheceUobium unguis-cati
Pithecellobium multiflorum (Kunth) Benth. [sensu Benth. 1944] =Albizia subdimidiata
Pithecellobium niopoides (Benth.) Spruce ex Benth. =Albizia niopoides
Pithecellobium miriense H.S. Irwin =Samanea inopinata
Pithecellobium panurense Spruce ex Benth. =Hydrochorea marginata var. panurensis
Pithecellobium parxiflorum Pittier =Pithecellobium roseum
Pithecellobium pedicellare (DC.) Benth. =Balizia pedicellaris
Pithecellobium pilosulum Pittier =Zygia inaequalis
Pithecellobium polycephalum Benth. [not in flora area] =Albizia polycephala
Pithecellobium pubescens (DC.) Benth. =Pithecellobium roseum
Pithecellobium pulchellum Pittier =Pithecellobium unguis-cati
Pithecellobium pullei Kleinhoonte =Hydrochorea gonggrijpii
Pithecellobium racemosum Ducke =Zygia racemosa
Pithecellobium ramiflorum Benth. =Zygia ramiflora
Pithecellobium saman (Jacq.) Benth. =Samanea saman
Pithecellobium samanigua (Pittier) J.F. Macbr. =Pseudosamanea guachapele
Pithecellobium schomburgkii Benth. =Enterolobium schomburgkii
Pithecellobium simabifolium Spruce ex Benth. =Macrosamanea simabifolia
Pithecellobium spruceanum Benth. =Macrosamanea spruceana
Pithecellobium subaquaticum Schery =Macrosamanea discolor
Pithecellobium trapezifolium (Vahl) Benth. =Abaremajupunba var. trapezifolia
Pithecellobium unifoliolatum Benth. =Zygia unifoliolata
Pithecellobium villiferum Ducke =Abarema villifera
Pithecolobium acreana J.F. Macbr. =Abarema acreana
Pithecolobium adenophorum Ducke =Abarema adenophora
Pithecolobium basijugum Ducke =Zygia basijuga
Pithecolobium cauliflorum (Willd.) Mart. =Zygia latifolia var. communis
Pithecolobium claviflorum Spruce ex Benth. =Zygia claviflora
Pithecolobium racemosum Ducke =Zygia racemosa
Pityrocarpa viridiflora (Kunth) Brenan =Piptadenia viridiflora
Poponax canescens Britton =Acacia macracantha
Poponaxfarnesiana (L.) Raf. =Acacia farnesiana
Poponaxflexuosa (Humb. & Bonpl. ex Willd.) Britton & Rose =Acacia macracantha
Poponax lutea (Mill.) Britton & Rose =Acacia macracantha
Poponax macracantha (Humb. & Bonpl. ex Willd.) Killip =Acacia macracantha
376 Checklist of the Plants of the Guiana Shield
Samanea adiantifolia (Kunth) Pittier =Macrosamanea discolor
Samanea corymbosa (Rich.) Pittier =Hydrochorea coiymbosa
Samanea pedicellaris (DC.) Killip ex Record =Balizia pedicellaris
Samanea polycephala (Benth.) Pittier [not in flora area] =Albizia polycephala
Samanea samanigiia Pittier =Pseudosamanea guachapele
Samanea simabifolia (Spruce ex Benth.) Pittier =Macrosamanea simabifolia
Schrankia leptocarpa DC. =Mimosa quadrivalvis var. leptocarpa
Senegalia glomerosa (Benth.) Britton & Rose =Acacia glomerosa
Senegalia panicidata (Willd.) Killip =Acacia tenuifoUa
Senegalia podadenia Britton & Killip =Acacia podadenia
Senegalia tamahndifolia (L.) Britton & Rose =Acacia tamarindifolia
Serianthes inopinata Harms =Samanea inopinata
Stryphnodendron angustum Benth. =Stryphnodendron pulcherhmum
Stryphnodendron flammatiim Kleinhoonte ? =Dimorphandra sp.
Stryphnodendron floribiindum Benth. =Stryphnodendron pulcherrimum
Stryphnodendron guianense i.floribunduih (Benth.) Ducke =Stryphnodendron pulcherrimum
Stryphnodendron inaequale Benth. =Stiyphnodendron microstachyum
Stryphnodendron melinonis Sagot =St}yphnodendron pulcherrimum
Stryphnodendron purpureum Ducke =Stfyphnodendron microstachyum
Vachelliafarnesiana (L.) Wight & Am. =Acaciafarnesiana
Zygia cauliflora (Willd.) Killip ex Record '=Zygia latifolia
Zygia divaricata (Benth.) Pittier =Zygia cataractae
Zygia glomerata (DC.) Pittier =Zygia cataractae
Zygia trunciflorum (Ducke) L. Rico =Zygiajuruana
Zygia unguis-cati (L.) Sudw. =Pithecellobium unguis-cati
References
Bameby, R.C. 1991. Sensitivae Censitae. A description of the genus Mimosa (Mimosaceae) in the New World. Mem. New York
Bot. Card. 65: 1-835.
Bameby, R.C. & J. Grimes (in press). Mimosaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A).
Bameby, R.C, J.W. Grimes, P.E. Berry, D. Bmnner, E. Forero, L. Cardenas, G. De Martino, H.C.F. Hopkins & E.M. de Lamare
Occhioni. 2001 . Mimosaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan Guayana
6: 580-686.
Cowan, R.S. 1961. Leguminosae. In B. Maguire, J.J. Wurdack, & collaborators, Botany of the Guayana Highland part IV. Mem.
New York Bot. Gard. 10(4): 65-87.
Grimes, J.W. 2002. Mimosaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
484-510.
Hopkins, H.C.F. 1986. Parkia. Flora Neotropica Monogr. 43: 1-123.
Jansen-Jacobs, M.J. 1976. Mimosaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 611-
653.
Kleinhoonte, A. 1940. Mimosaceae. In A. Pulle, Flora of Suriname 2(2): 258-331.
Polhill, R.M. & P.H. Raven (ed.). 1981. Advances in Legume Systematics. Royal Botanical Garden, Kew, England.
Poncy, O. 1985. Le genre Inga (Legumineuses, Mimosoideae) en Guyane fran^aise. Systematique, morphologic des formes
juveniles, ecologie. Mem. Mus. Nat. Hist. Nat. (n.s.), Sen B 31: 1-153.
Poncy, O. 1991. Le genre Inga dans les Guyanes: notes phytogeographiques. C.R. Soc. Biogeogr. 67(2): 95-111.
Poncy, O. 1991. Deux nouvelles especes de Inga (Mimosaceae) et notes nomenclaturales et taxonomiques sur trois autres especes.
Bull. Mus. natl. Hist, nat., Paris, sen 4, 13, sect. B, Adansonia, 3-4: 147-154.
Lentibulariaceae
RG. Taylor
Genliseafiliformis A. St.-Hil.
Genlisea glabra P. Taylor
Genlisea guianensis N.E. Bn
Genlisea pygmaea A. St.-Hil.
Genlisea repens Benj
.
Genlisea roraimensis N.E. Br.
Genlisea sanariapoana Steyerm.
Utricularia adpressa Salzm. ex A. St.-Hil. & Girard
Utricularia alpina Jacq.
Utricularia amethystina Salzm. ex A. St.-Hil. & Girard
Utricularia aureomacidata Steyerm.
Utricularia benjaminiana Oliv.
AM BO
BO
GU SU
AM BO GU
AM BO GU SU
AM BO GU SU
AM BO GU
AM
BO GU SU
AM BO GU SU
AM BO
BO
GU SU
AM BO GU SU
FG
FG
FG
FG
FG
Checklist of the Plants of the Guiana Shield 377
Utriciilaria breviscapa C. Wright ex Griseb.
Uthcidaria calycifida Benj.
Utriciilaria campbelliana Oliv.
Utriciilaria chiribiquitensis A. Fern.
Utriciilaria choristotheca P. Taylor
Utriciilaria costata P. Taylor
Utriciilaria ciicullata A. St.-Hil. & Girard
Utriciilaria determannii P. Taylor
Utricularia erectiflora A. St.-Hil. & Girard
Utriciilariafimbriata Kunth
Utriculariafoliosa L.
Utricularia gibba L.
Utricularia guyanensis A. DC.
Utricularia heterochroma Steyerm.
Utricularia hispida Lam.
Utricularia humboldtii R.H. Schomb.
Utricularia hydrocarpa Vahl
Utriculariajamesoniana Oliv.
Utriculariajuncea Vahl
Utricularia lloydii Merl
Utricularia longeciliata A. DC.
Utricularia mirabilis P. Taylor
Utricularia myriocista A. St.-Hil. & Girard
Utricularia nana A. St.-Hil. & Girard
Utricularia naviculata P. Taylor
Utricularia neottioides A. St.-Hil. & Girard
Utricularia nervosa Weber ex Benj.
Utricularia olivacea C. Wright ex Griseb.
Utricularia oliveriana Steyerm.
Utricularia pubescens Sm.
Utricularia pusilla Vahl
Utricularia quelchii N.E. Br.
Utricularia sandwithii P. Taylor
Utricularia schultesii A. Fern.
Utricularia simulans Pilg.
Utricularia spruceana Benth. ex Oliv.
Utricularia steyermarkii P. Taylor
Utricularia subulata L.
Utricularia tenuissima Tutin
Utricularia trichophylla Spruce ex Oliv.
Utricularia tricolor A. St.-Hil.
Utricularia triloba Benj.
Utricularia viscosa Spruce ex Oliv.
Synonyms and Misidentifications
Akentra inflata Benj. =Utricularia benjaminiana
Aranellafimbriata (Kunth) Barnhart =Utricularia fimbriata
Avesicaria neottioides (A. St.-Hil. & Girard) Barnhart =Utricularia neottioides
Biovularia olivacea (C. Wright ex Griseb.) Kamienski =Utricularia olivacea
Calpidisca amethystina (Salzm. ex A. St.-Hil. & Girard) Barnhart =Utricularia amethystina
Calpidisca calycifida (Benj.) Gleason =Utricularia calycifida
Calpidisca hispida (Lam.) Barnhart =Utricularia hispida
Calpidisca humboldtii (R.H. Schomb.) Gleason =Utricularia humboldtii
Calpidisca modesta (A. DC.) Barnhart =Utricularia amethystina
Calpidisca roraimensis (N.E. Br.) Gleason ^Utricularia amethystina
Genlisea anfractuosa Tutin =Genlisea filiformis
Genlisea esmeraldae Steyerm. =Genlisea pygmaea
Genlisea nigrocaiilis Steyerm. [based on mixed collection] =Genlisea pygmaea and Genlisea repens
Genlisea pulchella Tutin =Genlisea repens
Lecticula spruceana (Benth. ex Oliv.) Barnhart =Utricularia spruceana
Orchyllium alpinum (Jacq.) Barnhart ^Utricularia alpina
Orchyllium humboldtii (R.H. Schomb.) Barnhart =Utricularia humboldtii
Orchyllium quelchii (N.E. Br.) Gleason =Utricularia quelchii
Orchyllium schimperi (Schenck) Barnhart =Utriculariajamesoniana
Polypompholyx laciniata Benj. =Utricularia simulans
AM BO GU
AM BO GU SU FG
AM BO GU
AM BO
BO
SU FG
AM BO GU SU
SU
FG
BO GU SU
AM BO
AM BO DA GU SU FG
BO DA GU SU FG
AM BO
BO
GU SU FG
AM BO GU SU FG
AM BO GU
AM BO GU SU FG
AM BO GU SU FG
AM BO GU SU FG
AM SU FG
AM BO GU SU FG
AM
AM BO GU SU FG
AM BO GU SU FG
AM
AM BO
AM BO
AM BO GU SU
AM BO
AM BO GU FG
BO GU SU FG
AM BO GU
AM BO GU SU
AM
AM BO GU SU FG
AM
BO
AM BO GU SU FG
AM BO GU SU FG
BO GU SU FG
AM
AM BO GU SU FG
AM BO GU SU FG
378 Checklist of the Plants of the Guiana Shield
Polypompholyx schomburgkii Klotzsch =Utriciilaria longeciliata
Setiscapella pusilla (Vahl) Bamhart ex Britton & P. Wilson =Utricularia pusilia
Setiscapella subiilata (L.) Bamhart =Utricularia subulata
Stomoisiajuncea (Vahl) Bamhart =Utriculariajuncea
Stomoisia spicata (Sylven) Gleason =Utricularia erectiflora
Utricularia alutacea Tutin =Utricularia amethystina
Utricularia amazonasana Steyerm. =Utricularia hydrocarpa
Utricularia amethystina f. alutacea (Tutin) Steyerm. =Utricularia amethystina
Utricularia angulosa Poir. ^Utriculariajuncea
Utricularia angiistifolia Benj. =Utricularia hispida
Utricularia arenicola Tutin =Utricularia nana
Utricularia arenicola var. kavanayena Steyerm. =Utricularia nana
Utricularia aureolimba Steyerm. =Utricularia adpressa
Utricularia ayacuchae Steyerm. =Utricularia cucullata
Utricularia bolivarana Steyerm. =Utricularia amethystina
Utricularia campbelliana var. minor Steyerm. =Utricularia campbelliana
Utricularia coccinea Benj. =Utricularia hydrocarpa
Utricularia concinna N.E. Br. =Utriculariajamesoniana
Utricularia congesta Steyerm. =Utricularia simulans
Utricularia congesta f. deminutiva Steyerm. =Utricularia simulans
Utricularia connellii^.E. Br. =Utricularia pubescens
Utricularia cuspidata Steyerm. ^Utricularia calycifida
Utricularia dubia Benj. =Utricularia nana
Utriculariaflaccida auct. non A. DC: sensu P. Taylor, Mem. New York Bot. Gard. 17(1) =Utricularia nervosa
Utriculariaflaccida A. DC. [not in flora area]
Utriculariafockeana Miq. =Utricularia hydrocarpa
Utricularia guianensis Splitg. ex de Vries =Utricularia foliosa
Utricularia hirtella A. St.-Hil. & Girard ^Utricularia amethystina
Utricularia humboldtii f. cuneata Steyerm. =Utricularia humboldtii
Utricularia inflata Walter [sensu stricto, not in flora area]
Utricularia kaieteurensis Steyerm. =Utricularia amethystina
Utricularia leptantha Benj. =Utricularia pusilla
Utricularia magnifica Pilg. =Utricularia myriocista
Utricularia maguirei Steyerm. =Utricularia calycifida
Utricularia modesta A. DC. =Utricularia amethystina
Utricularia montana Jacq. =Utricularia alpina
Utricularia muscosa Benj. ^Utricularia calycifida
Utricularia obovata Miq. =Utricularia amethystina
Utricularia obtusa Sw. =Utricularia gibba
Utricularia oligosperma A. St.-Hil. =Utriculariafi)liosa
Utricularia orinocensis Steyerm. =Utricularia simulans
Utricularia parkeriana A. DC. =Utricularia gibba
Utricularia peckii S.F. Blake =Utricularia guyanensis
Utricularia pectinata Splitg. ex de Vries =Utricularia longeciliata
Utricularia peltata Spruce ex Oliv. =Utricularia pubescens
Utricularia praelonga A. St.-Hil. & Girard [not in flora area]
Utricularia punctifolia Benj. =Utricularia amethystina
Utricularia purpurea Walter [not in flora area]
Utricularia resupinata P. Taylor ^Utricularia spruceana
Utricularia roraimensis N.E. Br. =Utricularia amethystina
Utricularia rubricaulis Tutin =Utricularia guyanensis
Utricularia schimperi Schenck =Utriculariajamesoniana
Utricularia sciaphila Tutin =Utricularia pubescens
Utricularia spatulata Splitg. ex de Vries =Utricularia amethystina
Utricularia spatulifolia Pilg. =Utricularia amethystina
Utricularia spicata Sylven =Utricularia erectiflora
Utricularia stricta G. Mey. =Utriculariajuncea
Utricularia subpeltata Steyerm. =Utricularia pubescens
Utricularia tenuifolia Benj. =Utricularia gibba
Utricularia tenuiscapa Pilg. =Utricularia subulata
Utricularia tepuiana Steyerm. ^Utricularia amethystina
Utricularia turumiquirensis Steyerm. ^Utricularia amethystina
Utricularia venezuelana Steyerm. =Utricularia pubescens
Utricularia versicolor Benj. =Utricularia amethystina
Checklist of the Plants of the Guiana Shield 379
Vesiciilina cucullata (A. St.-Hil. & Girard) Barnhart =Utricularia cucullata
References
Smith, N.P. 2002. Lentibulariaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
397-398.
Taylor, P. 1967. Lentibulariaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 7. Mem. New York Bot.
Gard. 17(1): 1-439.
Taylor, P. 1989. The genus Utricularia - a taxonomic monograph. Kew Bull. Add. Ser. XIV: 1-724.
Taylor, P. 1999. Lentibulariaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
5: 782-803.
LiSSOCARPACEAE
P.E. Berry
Lissocarpa benthamii Giirke AM GU
Lissocarpa guianensis Gleason GU
Lissocarpa stenocarpa Steyerm. AM
References
Berry, P.E. 2001 . Lissocarpaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
6: 19-21.
LOASACEAE
J.S. Miller
Klaprothia mentzelioides Kunth AM
Mentzelia aspera L. FG?
References
Miller, J.S. 2001. Loasaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 6:
22.
LOGANIACEAE
P.E. Berry, A.E. Brant, A.J.M. Leeuwenberg
Antomaovata?o\A AM BO GU SU FG
Bonyiinia aqiiatica Ducke
Bonyiinia minor N.E. Br.
Bonyimia siiperba M.R. Schomb. ex Progel [Mt. Roraima]
Mitreola petiolata (J.F. Gmel.) Torr. & A. Gray SU FG?
Mostuea surinamensis Benth. [expected in FG]
Polypremmim procumbens L.
SpigeliaanthelmiaL. BO DA GU SU FG
Spigeliaflemmingiana Cham. & Schltdl.
Spigelia gracilis A. DC.
Spigelia guianensis (Aubl.) Lemee [incertae sedis; excluded from Speigelia AM BO GU FG
and Loganiaceae by Hurley]
Spigelia hamelioides Kunth FG
Spigelia multispica Steud. BO GU SU FG
Spigelia polystachya Klotzsch ex Progel BO GU
Strychnos barnhartiana Krukoff
Strychnos brachiata Ruiz & Pav.
Strychnos bredemeyeri (Schult. & Schult. f ) Sprague & Sandwith BO DA GU
Strychnos castelnaeana Wedd.
Strychnos cayennensis Krukoff& Barneby
Strychnos cogens Benth. AM BO GU
Strychnos davidsei Krukoff & Barneby
Strychnos diaboli Sandwith AM BO GU
Strychnos divaricans Ducke
Strychnos erichsonii M.R. Schomb. ex Progel AM BO GU
Strychnos eugeniifolia Monach.
Strychnos fendleri Sprague & Sandwith BO
Strychnos glabra Sagot ex Progel GU SU FG
AM
AM BO
BO
GU
GU
SU
SU?
GU
AM
AM
BO
BO
SU FG
FG?
FG
SU FG
SU FG
SU FG
SU FG
380 Checklist of the Plants of the Guiana Shield
Strychnos giiianensis (Aubl.) Mart.
Strychnos hirsuta Spruce ex Benth.
Strychnos jobertiana Baill.
Strychnos mattogrossensis S. Moore
Strychnos medeola Sagot ex Progel
Strychnos melinoniana Baill.
Strychnos mitscherlichii M.R. Schomb.
Strychnos mitscherlichii var. mitscherlichii
Strychnos oiapocensis Froes
Strychnos panamensis Seem.
Strychnos panurensis Sprague & Sandwith
Strychnos peckii B.L. Rob.
Strychnos ramentifera Ducke
Strychnos rondeletioides Spruce ex Benth.
Strychnos subcordata Spruce ex Benth. [expected in FG]
Strychnos tomentosa Benth.
Strychnos toxifera R.H. Schomb. ex Benth.
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
DA
AM
AM
BO
BO DA
GU SU FG
GU FG
FG
SU FG
GU SU FG
GU SU FG
GU SU FG
SU FG
GU FG
FG
GU SU FG
SU FG?
SU FG
GU
GU SU FG
GU SU FG
Synonyms and Misidentifications
Antonia ovata var. excelsa Paula =Antonia ovata
Antonia ovata war. pilosa (Hook.) Progel =Antonia ovata
Antonia pilosa Hook. -Antonia ovata
Antonia pubescens Bong. =Antonia ovata
Bonyunia cinchonoides Gleason & Standi. =Bonyunia minor
Lasiostoma bredemeyeri Schult. & Schult. f. =Strychnos bredemeyeri
Lasiostoma curare Kunth ^Strychnos giiianensis
Montira guianensis Aubl. [excluded from Loganiaceae by Hurley] =Spigelia guianensis
Mostuea brasiliensis Huber =Mostuea surinamensis
Rouhamon bredemeyeri (Schult. & Schult. f.) A. DC. ^Strychnos bredemeyeri
Rouhamon guianensis Aubl. =Strychnos guianensis
Rouhamon pedunculatum A. DC. =Strychnos bredemeyeri
Spigelia filipes Rusby =Spigelia multispica
Spigelia humboldtiana Cham. & Schltdl. [not in flora area] =Spigelia scabra Cham. & Schlechtend.
Spigelia humilis Benth. =Spigelia guianensis
Spigelia nervosa Steud. ^Spigelia anthelmia
Spigelia schomburgkiana Benth. =Spigeliaflemmingiana
Strychnos blackii Ducke ^Strychnos brachiata
Strychnos bovetiana Pires =Strychnos erichsonii
Strychnos curare (Kunth) Benth. =Strychnos guianensis
Strychnos darienensis Seem. =Strychnos bredemeyeri
Strychnos lanceolata Spruce ex Benth. =Strychnos guianensis
Strychnos pedunculata (A. DC.) Benth. =Strychnos bredemeyeri
Strychnos rouhamon Benth. =Strychnos guianensis
Strychnos ruizii Sprague & Sandwith =Strychnos brachiata
Strychnos schomburgkiana Klotzsch ex M.R. Schomb. [nomen nudum] =Strychnos bredemeyeri
Strychnos smilacina Benth. =Strychnos mitscherlichii
Strychnos smithiana Krukoff ^S/ryc/zwo^ rondeletioides
Strychnos solerederi Gilg =Strychnos mitscherlichii var. mitscherlichii
Strychnos syntoxica Sprague & Sandwith =Strychnos toxifera
Strychnos trichostyla Ducke =Strychnos jobertiana
Strychnos trinitensis Griseb. =Strychnos bredemeyeri
Strychnos urbanii Barb. Rodr. =Strychnos erichsonii
References
Bordenave, B. 2002. Loganiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
398-405.
Hurley, H.H. 1967. A taxonomic revision of the genus Spigelia. George Washington University, Washington, DC. (Unpublished
Ph.D. thesis)
Krukoff, B.A. & J. Monachino. 1942. The American species of Strychnos. Brittonia 4: 248-322. (See also the subsequent series
of supplementary notes in various journals.)
Leeuwenberg, A.J.M. 1961. The Loganiaceae ofAfrica II. A revision of Mostuea Didr. Meded. Landb. Wag. 61(4): 1-31.
Leeuwenberg, A.J.M. 1969. Notes on American Loganiaceae. III. Revision of Bonyunia Rich. Schomb. Acta Bot. Neerl. 18(1):
152-158.
Leeuwenberg, A.J.M. 1971. Notes on American Loganiaceae. V. Key to the genera. Acta Bot. Neerl. 20(5): 539-542.
Checklist of the Plants of the Guiana Shield 381
Leeuwenberg, A.J.M. 1975. The Loganiaceae of Africa XII. A revision of Mitreola L. Meded. Landb. Wag. 74(23): 1-28.
Leeuwenberg, A.J.M. (ed.). 1980. Loganiaceae. In Engler & Prantl, Die Natiirlichen Pflanzenfamilian 28bl: 1-255.
Leeuwenberg, A.J.M., RE. Berry, A.E. Brant. 2001 . Loganiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 6: 22-36.
Raalte, M.H. van. 1932. Loganiaceae. In A. Pulle, Flora of Suriname 4(1): 103-110.
Smith, L.B. 1960. Notes on South American phanerogams - III. Wrightia 2(2): 90-102.
Uittien, H. 1937. Loganiaceae (additions and corrections). In A. Pulle, Flora of Suriname 4(1): 472-474.
LORANTHACEAE
J. Kuijt
Cladocolea elliptica Kuijt
Cladocolea intermedia (Rizzini) Kuijt
Cladocolea micrantha (Eichler) Kuijt
Cladocolea nitida Kuijt
Cladocolea roraimensis (Steyerm.) Kuijt
Cladocolea sandwithii (Maguire) Kuijt
Gaiadendron punctatum (Ruiz & Pav.) G. Don
Ixocactus clandestinus (Mart.) Kuijt
Oryctanthus alveolatus (Kunth) Kuijt
Oryctanthus florulentus (Rich.) Tiegh.
Oryctanthus occidentalis (L.) Eichler
Oryctanthus occidentalis subsp. continentalis Kuijt
Oryctanthus phthirusoides Rizzini
Oryctanthus spicatus (Jacq.) Eichler
Oryctina myrsinites (Eichler) Kuijt
Phthirusa bisexualis Rizzini
Phthirusa coarctata A.C. Sm.
Phthirusa disjectifolia (Rizzini) Kuijt
Phthirusa guyanens is Eichler
Phthirusa monetaria Sandwith
Phthirusa nitens (Mart.) Eichler
Phthirusa podoptera (Cham. & Schltdl.) Kuijt
Phthirusa pycnostachya Eichler
Phthirusa pyrifolia (Kunth) Eichler
Phthirusa rufa (Mart.) Eichler
Phthirusa stelis (L.) Kuijt
Phthirusa stenophylla Eichler
Psittacanthus acinarius (Mart.) Mart.
Psittacanthus biternatus (Hoffmanns.) Blume
Psittacanthus brachynema Eichler
Psittacanthus cinctus (Mart.) Mart.
Psittacanthus clusiifolius (Willd.) Eichler
Psittacanthus collum-cygni Eichler
Psittacanthus cordatus (Hoffmanns.) Blume
Psittacanthus corynocephalus Eichler
Psittacanthus cucullaris (Lam.) Blume
Psittacanthusjulianus Rizzini
Psittacanthus lamprophyllus Eichler
Psittacanthus lasianthus Sandwith
Psittacanthus leptanthus Sandwith
Psittacanthus melinonii (Tiegh.) Engl.
Psittacanthus montis-neblinae Rizzini
Psittacanthus peronopetalus Eichler
Psittacanthus plagiophyllus Eichler
Psittacanthus redactus Rizzini
Psittacanthus rhynchanthus (Benth.) Kuijt
Psittacanthus robustus (Mart.) Mart.
Struthanthus concinnus Mart.
Struthanthus confertus Mart.
Struthanthus dichotrianthus Eichler
Struthanthus eichlerianus Rizzini
Struthanthus elegans Mart.
Struthanthus gracilis (Gleason) Steyerm. & Maguire
SU
BO
BO GU
GU
SU EG
BO
GU
AM BO GU
GU
AM BO DA GU SU FG
AM BO DA GU SU EG
AM
BO
GU
GU
GU
SU
SU
AM GU SU FG
AM
GU
AM BO GU SU
AM GU
GU
SU FG
AM BO GU
AM
GU FG
AM BO DA GU
GU
SU
SU
FG
AM BO DA GU SU FG
AM GU
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO DA
GU SU?
BO SU
GU FG
FG
GU? SU FG
FG
GU SU
GU
FG
FG
GU SU FG
FG
BO
GU
GU SU
SU?
BO DA GU SU FG
BO
GU
BO GU
382 Checklist of the Plants of the Guiana Shield
BO
BO
BO
GU
GU
GU
GU
Struthanthus marginatiis (Desr.) Blume
Struthanthus phyllyraeoides (Kunth) Blume
Struthanthus syringifolius (Mart.) Mart.
Struthanthus vulgaris Eichler
Tripodanthus acutifolius (Ruiz & Pav.) Tiegh. [expected in the Guianas]
Synonyms and Misidentifications
Apodina cucuUaris (Lam.) Tiegh. =Psittacanthus cucullaris
Cladocolea apiculata Kuijt =Cladocolea intermedia
Cladocolea clandestina (Mart.) Kuijt =Ixocactus clandestinus
Furarium disjectifolium Rizzini ^Phthirusa disjectifolia
Gaiadendron lanceolatum (Ruiz & Pav.) Baehni =Gaiadendron punctatum
Gaiadendron poasense Donn. Sm. =Gaiadendron punctatum
Gaiadendron punctatum (Ruiz & Pav.) G. Don \ar. puracense (Kunth) Steyerm. =Gaiadendron punctatum
Gaiadendron puracensis (Kunth) G. Don =Gaiadendron punctatum
Gaiadendron revolutum Rizzini =Gaiadendron punctatum
Gaiadendron tagua (Kunth) G. Don =Gaiadendron punctatum
Gaiadendron tagua var. reductum Rizzini =Gaiadendron punctatum
Gaiadendron tagua var. revolutum (Rizzini) Rizzini =Gaiadendron punctatum
Loranthus acinarius Mart. =Psittacanthus acinarius
Loranthus acutifolius Ruiz & Pav. =Tripodanthus acutifolius
Loranthus aduncus G Mey. =Phthirusa stelis
Loranthus alveolatus Kunth =Oryctanthus alveolatus
Loranthus amplexicaulis Kunth =0}yctanthus alveolatus
Loranthus biternatus Hoffmanns. =Psittacanthus biternatus
Loranthus cinctus Mart. =Psiltacanthus cinctus
Loranthus conduplicatus Kunth ^Phthirusa stelis
Loranthus cucullaris Lam. =Psittacanthus cucullaris
Loranthus cupulifer Kunth =Psittacanthus cucullaris
Loranthus eugenioides Kunth =Tripodanthus acutifolius
Loranthus falcifrons Mart. =Psittacanthus cucullaris
Loranthus florulentus Rich. =Oiyctanthus florulentus
Loranthusformosus Cham. & SchUdl. =Psittacanthus robustus
Loranthus guadalupensis DC. (Rubiaceae) =Notopleura guadalupensis
Loranthus lanceolatum Ruiz & Pav. =Gaiadendron punctatum
Loranthus magdalenae Cham. & Schhdl. =Phthirusa stelis
Loranthus nitens Mart. =Phthirusa nitens
Loranthus occidentalis L. =Oiyctanthus occidentalis
Loranthus orinocensis Spreng. =Phthirusa stelis
Loranthus paniculatus Kunth =Phthirusa stelis
Loranthus patrisii DC. (Combretaceae) =Combretum sp.
Loranthus podopterus Cham. & Schltdl. =Phthirusa podoptera
Loranthus pterygopus Mart. =Phthirusa podoptera
Loranthus punctatus Ruiz & Pav. =Gaiadendron punctatum
Loranthus puracensis Kunth ^Gaiadendron punctatum
Loranthus pyrifolius Kunth =Phthirusa pyrifolia
Loranthus retroflexus Ruiz & Pav. =Phthirusa stelis
Loranthus rhynchanthus Benth. =Psittacanthus rhynchanthus
Loranthus robustus Mart. =Psittacanthus robustus
Loranthus ruficaulis Poepp. & Endl. =Oryctanthus florulentus
Loranthus speciosus Pohl =Psittacanthus robustus
Loranthus stelis L. =Phthirusa stelis
Loranthus syringifolius Mart. =Struthanthus syringifolius
Loranthus tagua Kunth =Gaiadendron punctatum
Notanthera lanceolata (Ruiz & Pav.) G. Don =Gaiadendron punctatum
Oryctanthus alveolatus var. amplexicaulis (Kunth) Rizzini =Oryctanthus alveolatus
Oryctanthus alveolatus var. kuijtii Rizzini =Oryctanthus alveolatus
Oryctanthus amplexicaulis (Kunth) Eichler =Oryctanthus alveolatus
Oryctanthus botryostachys Eichler ^Oryctanthus alveolatus
Oryctanthus laceratus Kuijt =Oryctanthus alveolatus
Oryctanthus ruficaulis (Poepp. & Endl.) Eichler =Oryctanthus florulentus
Passovia suaveolens Karst. =Phthirusa stelis
Passowia guyanensis (Eichler) Tiegh. =Phthirusa guyanensis
Passowia micrantha (Eichler) Tiegh. =Cladocolea micrantha
SU FG
Checklist of the Plants of the Guiana Shield 383
Passowia myrsinites (Eichler) Tiegh. =Oiyctina myrsinites
Passowia orinocensis (Spreng.) Tiegh. =Phthirusa stelis
Passowia pyrifolia (Kunth) Tiegh. =Phthiriisa pyrifoUa
Phoradendron karuaiamim Steyerm. =Dendrophthora kaniaiana
Phrygilanthus acutifolhis (Ruiz & Pav.) Eichler =Tripodanthus acutifolius
Phygilanthiis acutifolius subsp. coriaceus Rizzini =Tripodanthus acutifolius
Phiygilanthus acutifolius subsp. eugenioides lateovatus Rizzini =Thpodanthus acutifolius
Phiygilanthus acutifolius subsp. eugenioides vulgaris Rizzini =Tripodanthus acutifolius
Phrygilanthus lanceolatum (Ruiz & Pav.) Eichler =Gaiadendron punctatum
Phiygilanthus megathermicus Rizzini =Phthirusa stenophylla
Phrygilanthus punctatus (Ruiz & Pav.) Eichler =Gaiadendron punctatum
Phthirusa adunca (G. Mey.) Maguire =Phthirusa stelis
Phthirusa adunca var. magdalenae (Cham. & Schltdl.) Rizzini =Phthirusa stelis
Phthinisa adunca var. orinocensis (Spreng.) Steyerm. =Phthirusa stelis
Phthir'usa adunca var. rigidifolia Rizzini =Phthirusa stelis
Phthirusa anastyla Rizzini =Phthirusa stelis
Phthir'usa angulata K. Krause =Phthirusa stelis
Phthirusa bernar'diana Rizzini =Cladocolea micrantha
Phthirusa calloso-albida Rizzini =Phthirusa guyanensis
Phthir'usa castillana Rizzini =Phthirusa guyanensis
Phthinisa cothur-nata Rizzini =Phthirusa stelis
Phthir-usa gracilis Gleason =Struthanthus gracilis
Phthirusa hippocrateoides Steyerm. & Maguire =Struthanthus syringifolius
Phthirusa huberi Rizzini =Phthirusa stenophylla
Phthirusa intermedia Rizzini =Cladocolea intermedia
Phthirusa maculata Rizzini =Oryctanthus phthirusoides
Phthirusa maritima Rizzini =Phthirusa stelis
Phthirusa megathermica (Rizzini) Rizzini =Phthirusa stenophylla
Phthir-usa micrantha Eichler =Cladocolea micrantha
Phthir-usa micrantha var. bolivariensis Rizzini =Cladocolea micrantha
Phthir-usa myrsinites Eichler =Oryctina myrsinites
Phthirusa myrsinites var. savannarum (Maguire) Rizzini =Oryctina myrsinites
Phthir-usa orinocensis (Spreng.) Eichler =Phthirusa stelis
Phthir-usa paniculata (Kunth) J.F. Macbr. =Phthirusa stelis
Phthir-usa pauciflora Eichler [not in flora area] =Dendropemon pauciflorus (Swartz) Tiegh.
Phthirusa percrassa Rizzini =Phthirusa stenophylla
Phthirusa perdivergens Rizzini =Phthirusa stelis
Phthir-usa perforata Rizzini =Phthirusa guyanensis
Phthir-usa phaeocladus Eichler =Phthirusa guyanensis
Phthirusa ptariana Steyerm. =Struthanthus syringifolius
Phthirusa punctata Gleason =Phthir-usa stelis
Phthir-usa pyramidalis Rizzini =Phthir-usa stelis
Phthir-usa retr-oflexa (Ruiz & Pav.) Kuijt =Phthirusa stelis
Phthirusa r-oraimense Steyerm. =Cladocolea roraimensis
Phthir-usa rubr-omicans Rizzini =Phthirusa stelis
Phthir-usa sandwithii Maguire =Cladocolea sandwithii
Phthirusa savannar-um Maguire =Oryctina myrsinites
Phthir-usa seitzii Krug & Urb. =Phthirusa stelis
Phthir-usa squamulosa (Pohl) Klotzsch ex Eichler =Phfhir-usa r-ufa
Phthirusa theobromae (Willd.) Eichler =Phthirusa stelis
Psittacanthus calyculatus var. wurdackii Rizzini =Psittacanthus r-hynchanthus
Psittacanthus chrismarii Urb. =Psittacanthus rhynchanthus
Psittacanthus cinctus var. guainiae Rizzini =Psittacanthus cinctus
Psittacanthus clusiifolius var. pseudojulianus Rizzini =Psittacanthus clusiifolius
Psittacanthus cupulifer (Kunth) Klotzsch =Psittacanthus cucullaris
Psittacanthus falcifi'ons (Mart.) Mart. =Psittacanthus cucullaris
Psittacanthus intermedius Rizzini =Psittacanthus robustus
Psittacanthus r-hynchanthus subsp. wur-dackii (Rizzini) Kuijt ^Psittacanthus rhynchanthus
Psittacanthus semiarticulatus Rizzini =Psittacanthus r-hynchanthus
Psittacanthus war-mingii Eichler =Psittacanthus acinarius
Struthanthus chimantensis Steyerm. & Maguire =Struthanthus gracilis
Struthanthus cupulifer Rizzini =Sfr-uthanthus gr-acilis
Struthanthus dichotrianthus var. dichotrianthus =Str-uthanthus dichotrianthus
Struthanthus dichotrianthus var. lasserianus Rizzini =Str-uthanthus dichotrianthus
384 Checklist of the Plants of the Guiana Shield
Struthanthus dissimilis Rizzini =Struthanthus dichotrianthus
Struthanthus gracilis var. mucronatus (Steyerm.) Rizzini =Struthanthus gracilis
Struthanthus granulatus Rizzini =Struthanthus dichotrianthus
Struthanthus micranthus (Eichler) Baehni & J.F. Macbr. =Cladocolea micrantha
Struthanthus mucronatus Steyerm. =Struthanthus gracilis
Struthanthus nitens (Mart.) Mart. =Phthirusa nitens
Struthanthus pseudolepidotus Rizzini =Phthirusa nitens
Struthanthus pterygopus (Mart.) Mart. =Phthirusa podoptera
Struthanthus retroflexa (Ruiz & Pav.) Kuijt =Phthirusa stelis
Struthanthus squamulosus Klotzsch ex Eichler =Phthirusa rufa
Struthanthus terniflorus Eichler =Struthanthus dichotrianthus
Struthanthus translucens Rizzini =Struthanthus dichotrianthus
Struthanthus umbellatus Kuijt =Phthirusa pycnostachya
Struthanthus yavitensis Rizzini =Struthanthus dichotrianthus
Taguaria puracense (Kunth) Raf. =Gaiadendron punctatum
Tripodanthus eugenioides (Kunth) TxQgh. -Tripodanthus acutifoUus
References
Krause, K. 1932. Loranthaceae. In A. Pulle, Flora of Suriname 1(1): 4-24.
Kuijt, J. 1976. Revision of the genus Oryctanthus. Bot. Jahrb. Syst. 95: 478-534.
Kuijt, J. 1985. The genus Cladocolea. J. Am. Arb. 56: 265-335.
Kuijt, J. 2001 . Loranthaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 6:
37-59.
Kuijt, J. 2002. Loranthaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 405-
408.
Lindeman, J.C. & A.R.A. Gorts-van Rijn. 1968. Loranthaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of
Suriname 1(2): 295-300.
Rizzini, C.T. 1978. Loranthaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 10. Mem. New York Bot.
Gard. 29: 23-36.
Rizzini, C.T. 1982. Loranthaceae. In Z. Luces de Febres, Flora de Venezuela 4(2): 7-316.
Lythraceae
A. Lourteigt
Ammannia auriculata Willd. [expected in the Guianas]
Ammannia coccinea Rottb. [expected in the Guianas]
Ammannia latifolia L.
Crenea maritima Aubl.
Cuphea anagalloidea A. St.-Hil.
Cuphea anisoclada Lourteig
Cuphea annulata Koehne
Cuphea antisyphilitica Kunth
Cuphea antisyphilitica var. acutifolia Benth.
Cuphea antisyphilitica var. antisyphilitica
Cuphea blackii Lourteig
Cuphea bolivariensis Lourteig
Cuphea cardonae Lourteig
Cuphea carthagenensis (Jacq.) J.F. Macbr.
Cuphea cataractarum Spruce ex Koehne
Cuphea curiosa Lourteig
Cuphea curiosa var. curiosa
Cuphea curiosa var. oresbia Lourteig
Cuphea dactylophora Koehne
Cuphea distichophylla Lourteig
Cuphea elliptica Koehne
Cuphea galeato-calcarata Lourteig
Cuphea gracilis Kunth
Cuphea insolita Lourteig
Cuphea maigualidensis Lourteig
Cuphea melvilla Lindl.
Cuphea micrantha Kunth
Cuphea odonellii Lourteig
Cuphea pleiantha Lourteig
AM BO
AM BO
AM BO
AM BO
AM BO
BO
BO
BO
AM
AM BO
AM BO
AM BO
AM BO
AM
AM BO
AM
AM
AM BO
AM BO
BO
AM BO
AM
DA
DA
DA
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
su
FG
FG
SU FG
SU
SU
FG
FG
FG
FG
BO GU
BO
GU
BO DA GU
BO
BO GU
BO DA
Checklist of the Plants of the Guiana Shield 385
Ciiphea repens Koehne [expected in the Guianas]
Cuphea rhodocalyx Lourteig AM
Cuphea hgidula Benth.
Cuphea tenuissima Koehne
Lafoensia vandelliana Cham. & Schhdl. [expected in the Guianas]
Lagerstroemia speciosa (L.) Pers. [cultivated or escaped from cultivation]
Lawsonia inermis L. SU
Pehria compacta (Rusby) Sprague
Rotala mexicana Cham. & Schltdl. AM SU FG
Rotala ramosior (L.) Koehne [expected in the Guianas]
Synonyms and Misidentifications
Ammannia ramosior L. =Rotala ramosior
Crenea surinamensis (L. f.) Koehne =Crenea maritima
Cuphea antisyphilitica f. gracillima Koehne =Cuphea antisyphilitica var. acutifolia
Cuphea antisyphilitica f. subhirsuta Koehne =Cuphea antisyphilitica
Cuphea balsamona Cham. & Schltdl. =Cuphea carthagenensis
Cuphea gracilis auct. non Kunth 1824, nee Seem. 1854. =Cuphea antisyphilitica
Cuphea gracilis var. major Koehne =Cuphea antisyphilitica var. acutifolia
Cuphea gracilis Kunth var. media Koehne =Cuphea gracilis
Cuphea mimuloides Cham. & Schltdl. var. guianensis Koehne =Cuphea anagalloidea
Cuphea multicaulis Koehne =Cuphea gracilis
Cuphea pauciflora Koehne =Cuphea gracilis
Cuphea rhodocalyx var. setosa Lourteig =Cuphea cardonae
Cuphea speciosa (A. Anderson) Kuntze =Cuphea melvilla
Dodecas maritimus (Aubl.) Griseb. =Crenea maritima
Dodecas surinamensis L. f. =Crenea maritima
Grislea compacta Rusby =Pehria compacta
Grislea secunda Loefl. =Pehria compacta
Hypobrichia spruceana Benth. =Rotala mexicana
Hypobrichia spruceana var. tenuifolia Griseb. =Rotala mexicana
Lafoensia pacari A. St.-Hil. [not in flora area]
Lawsonia alba Lam. =Lawsonia inermis
Lawsonia speciosa L. =Lawsoma inermis
Lythrum carthagenense Jacq. =Cuphea carthagenensis
Melvilla speciosa A. Anderson [nomen nudum] =Cuphea melvilla
Rotala apetala F. Muell. =Rotala mexicana
Rotala mexicana var. apetala (F. Muell.) Lourteig =Rotala mexicana
Rotala mexicana var. spruceana (Benth.) Koehne =Rotala mexicana
References
Cook, C.D.K. 1979. A revision of the genus Rotala. Boissiera 29: 1-156.
Graham, S.A. 1985. A revision ofAmmannia in the western hemisphere. J. Arnold Arb. 56: 265-335.
Jonker, F.R 1941. Lythraceae. In A. Pulle, Flora of Suriname 3(1): 422-432.
Kawasaki, M.L. 2002. Lythraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
408-409.
Lourteig, A. 1964. Lythraceae Austroamericanae, addenda et corrigenda. Sellowia 16: 119-162.
Lourteig, A. 1 986. Revision del genero Lafoensia Vandelli (Litraceas). Mem. Soc. Cienc. Nat. La Salle 45( 1 23): 115-157.
Lourteig, A. 1986. Revision de dos secciones del genero Cuphea P. Browne. Phytologia 60(1): 17-55.
Lourteig, A. 1987. Lythraceae Austroamericanae, addenda et corrigenda II. Sellowia 39: 15-48.
Lourteig, A. 2001 . Lythraceae. In J. A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 6:
59-80.
Magnoliaceae
RE. Berry, J. S. Miller
Dugandiodendron chimantense (Steyerm. & Maguire) Lozano BO
Dugandiodendron ptaritepuianum (Steyerm.) Lozano [Mt. Roraima] BO GU
Synonyms and Misidentifications
Magnolia chimantensis Steyerm. & Maguire =Dugandiodendron chimantense
Magnolia ptaritepuiana Steyerm. =Dugandiodendron ptaritepuianum
Magnolia roraimae Steyerm. =Dugandiodendron ptaritepuianum
386 Checklist of the Plants of the Guiana Shield
References
Berry, RE. and J.S. Miller. 2001. Magnoliaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 6: 80-82.
Lozano Contreras, G. 1994. Dugandiodendron y Talauma (Magnoliaceae) en el Neotropica. Academia de Ciencias Exactas,
Fisicas y Naturales Coleccion Jorge Alvarez Lleras No. 3.
Steyermark, J.A. 1951. Magnoliaceae. In Botanical Exploration in Venezuela - 1. Fieldiana Bot. 28(1): 233-234.
Malpighiaceae
C. Anderson, W.R. Anderson
Banisteriopsis caapi (Griseb.) C.V Morton AM
Banisteriopsis Carolina W.R. Anderson FG
Banisteriopsis cinerascens (Benth.) B. Gates
Banisteriopsis cristata (Griseb.) Cuatrec.
Banisteriopsis krukoffii B. Gates
Banisteriopsis hicida (Rich.) Small
Banisteriopsis lyrata B. Gates
Banisteriopsis maguirei B. Gates
Banisteriopsis martiniana (A. Juss.) Cuatrec.
Banisteriopsis martiniana var. martiniana
Banisteriopsis martiniana var. subenervia Cuatrec.
Banisteriopsis mathiasiae (W.R. Anderson) W.R. Anderson
Banisteriopsis muricata (Cav.) Cuatrec.
Banisteriopsis nummifera (A. Juss.) B. Gates
Banisteriopsis pulcherrima (Sandwith) B. Gates
Banisteriopsis schwannioides (Griseb.) B. Gates
Banisteriopsis wiirdackii B. Gates
Blepharandra angiistifolia (Kunth) W.R. Anderson
Blepharandrafimbriata MacBryde
Blepharandra heteropetala W.R. Anderson
Blepharandra hypoleuca (Benth.) Griseb.
Bunchosia apiculata Huber
Bunchosia argentea (Jacq.) DC.
Bunchosia armeniaca (Cav.) DC.
Bunchosia decussiflora W.R. Anderson
Bunchosia glandulifera (Jacq.) Kunth
Bunchosia mollis Benth.
Bunchosia petraea W.R. Anderson
Bunchosia polystachia (Andrews) DC.
Burdachia prismatocarpa A. Juss.
Burdachia sphaerocarpa A. Juss.
Byrsonima aerugo Sagot
Byrsonima altissima DC.
Byrsonima arthropoda A. Juss.
Byrsonima baccae W.R. Anderson
Byrsonima basiliana W.R. Anderson
Byrsonima bronweniana W.R. Anderson
Byrsonima carraoana Steyerm.
Byrsonima chalcophylla Nied.
Byrsonima christianeae W.R. Anderson [expected in BO]
Byrsonima chrysophylla Kunth
Byrsonima coccolobifolia Kunth
Byrsonima concinna Benth.
Byrsonima coniophylla A. Juss.
Byrsonima cowanii W.R. Anderson
Byrsonima crassifolia (L.) Kunth
Byrsonima crispa A. Juss.
Byrsonima cuprea Griseb.
Byrsonima densa (Poir.) DC.
Byrsonima dubia W.R. Anderson
Byrsonima duidana W.R. Anderson
Byrsonima eugeniifolia Sandwith
Byrsonima euryphylla Pilg.
Byrsonimafanshawei W.R. Anderson
GU
BO GU su
AM BO
AM BO
BO
DA
DA
GU su FG
AM BO
AM BO DA GU su FG
BO DA GU su FG
AM
GU
BO GU FG
FG
BO GU
FG
BO FG
AM
BO GU
AM
AM BO GU
FG
BO GU su? FG
BO DA
AM GU FG
AM su FG
BO GU su?
AM
FG
AM BO
AM GU
BO DA GU
GU
GU
su
su
FG
FG
AM
AM
AM
BO
BO
GU
GU
GU
AM BO su
BO DA GU su
AM BO GU
AM
AM
AM BO DA GU su FG
AM
A \/t
BO FG
AM
GU su FG
BO GU
AM
GU
GU
su
su
GU
GU SU? FG?
GU
GU
GU SU FG
Checklist of the Plants of the Guiana Shield 387
Byrsonima fernandezii Cuatrec. AM
Byrsonima flexipes W.R. Anderson AM
Byrsonimaformosa W.R. Anderson
Byrsonimafrondosa A. Juss. AM
Byrsonima gymnocalycina A. Juss.
Byrsonima huberi W.R. Anderson AM
Byrsonima incarnata Sandwith
Byrsonimajapurensis A. Juss. AM BO
Byrsonima kariniana W.R. Anderson AM
Byrsonima krukoffii W.R. Anderson
Byrsonima laevigata (Poir.) DC.
Byrsonima laevis Nied. AM
Byrsonima leucophlebia Griseb. BO
Byrsonima linguifera Cuatrec. AM
Byrsonima luetzelburgii Steyerm. AM
Byrsonima macrostachya W.R. Anderson AM
Byrsonima maguirei W.R. Anderson [Sierra de la Neblina] AM
Byrsonima nitidissima Kunth AM BO
Byrsonima pachypoda^.K. Anderson BO GU
Byrsonima poeppigiana A. ]\jiss. AM
Byrsonima punctulata A. Juss. AM
Byrsonima rubrobracteata W.R. Anderson [expected in BO]
Byrsonima schomburgkiana Benth.
Byrsonima sericea DC.
Byrsonima spicata (Cav.) DC.
Byrsonima steyermarkii W.R. Anderson
Byrsonima stipulacea A. Juss.
Byrsonima surinamensis W.R. Anderson
Byrsonima tillettii W.R. Anderson
Byrsonima verbascifolia (L.) DC.
Byrsonima wurdackii W.R. Anderson
Camarea affinis A. St.-Hil.
Clonodia complicata (Kunth) W.R. Anderson
Coleostachys genipifolia A. Juss.
Diacidia cordata (Maguire) W.R. Anderson [Sierra de la Neblina]
Diacidiaferruginea (Maguire & K.D. Phelps) W.R. Anderson
Diacidia galphimioides Griseb.
Diacidia glaucifolia (Maguire) W.R. Anderson
Diacidia hypoleuca (Maguire) W.R. Anderson
Diacidia kunhardtii (Maguire) W.R. Anderson
Diacidia rufa (Maguire) W.R. Anderson [Sierra de la Neblina]
Diacidia steyermarkii (Maguire) W.R. Anderson
Diacidia stipidaris (Maguire & K.D. Phelps) W.R. Anderson
Diacidia vestita (Benth.) B.D. Jacks.
Dicellajulianii (J.F. Macbr.) W.R. Anderson [expected in AM; Serra da
Neblina]
Diplopterys cabrerana (Cuatrec.) B. Gates AM
Diploptetys pauciflora {G.Mey.)^\Qd. GU FG
Excentradenia adenophora (Sandwith) W.R. Anderson BO DA GU
Excentradenia propinqua {V^.K. Andtrson) '^.K. Anderson GU SU FG
Glandonia williamsii Steyerm. AM
Heteropterys alata (W.R. Anderson) W.R. Anderson AM BO
Heteropterys atabapensis W.R. Anderson AM
Heteropterys ayacuchensis W.R. Anderson AM
Heteropterys biglandidosa A. Juss. FG
Heteropterys cristata Benth. AM BO GU
Heteropterys cuatrecasasii W.R. Anderson AM
Heteropterys dichromocalyx W.R. Anderson [expected in AM; Pico da
Neblina]
Heteroptetys guianensis W.R. Anderson
Heteropterys hoffmanii W.R. Anderson
Heteropterys huberi W.R. Anderson
Heteropterys leona (Cav.) Exell
Heteropterys macradena (DC.) W.R. Anderson
GU
AM BO GU
FG
AM BO DA GU SU FG
AM BO
BO DA GU SU
SU
FG
BO GU
AM BO GU SU FG
AM
GU SU
AM BO DA
FG
AM
AM
AM BO
AM
ATV/tAJYl
AM
AM
BO
AM
AM
GU
GU
AM
DA GU SU FG
AM BO GU SU FG
AM BO
BO
BO
BO
GU
GU
GU
SU
SU
FG
AM
AM GU SU FG
AM
GU
AM BO
AM BO
AM GU FG
AM
GU
GU
DA GU SU FG
AM
AM
BO GU SU FG
AM BO DA GU SU FG
AM BO
GU SU FG
FG
GU
FG
AM
FG
388 Checklist of the Plants of the Guiana Shield
Heteropterys macroslachya A. Juss.
Heteropterys maguirei W.R. Anderson
Heteropterys mathewsana A. Juss.
Heteropterys megaptera A. Juss.
Heteropterys molesta W.R. Anderson
Heteropterys neblinemis W.R. Anderson [Sierra de la Neblina]
Heteropterys nervosa A. Juss.
Heteropterys oblongifolia Gleason
Heteropterys oligantha W.R. Anderson
Heteropterys orinocensis (Kunth) A. Juss.
Heteropterys quetepensis Steyerm.
Heteropterys siderosa Cuatrec.
Heteropterys steyermarkii W.R. Anderson
Heteropterys subhelicina Nied.
Heteropterys velutina W.R. Anderson
Hiraea affwis Miq.
Hiraea apaporiensis Cuatrec.
Hiraea celiana W.R. Anderson
HiraeafagifoUa (DC.) A. Juss.
Hiraeafaginea (Sw.) Nied.
Hiraeafimbriata W.R. Anderson
Hiraea gracieana W.R. Anderson
Hiraea longipes W.R. Anderson
Hiraea longipilifera W.R. Anderson
Hiraea morii W.R. Anderson
Hiraea neblinensis W.R. Anderson
Hiraea quapara (Aubl.) Sprague
Hiraea steyermarkii W.R. Anderson
Hiraea tepuiensis Steyerm.
Hiraea ternifolia (Kunth) A. Juss.
Jiibelina grisebachiana W.R. Anderson
Jubelina magnifica W.R. Anderson
Jubelina riparia A. Juss. FG
Jubelina rosea (Miq.) Nied. SU FG
Lophanthera longifo/ia (Kunth) Griseb.
Lophopterys euryptera Sandwith BO DA GU
Lophopterys inpana W.R. Anderson
Lophopterys splendens A. Juss. FG
Lophopteiys surinamensis (Kosterm.) Sandwith SU
Malpighia emarginata DC. BO DA FG
Malpighia glabra L.
Mascagnia arenicola C. Anderson GU SU FG
Mascagnia castanea (Cuatrec.) W.R. Anderson
Mascagnia cynanchifolia Griseb.
Mascagnia dissimilis C.V. Morton & Moldenke
Mascagnia divaricata (Kunth) Nied. DA SU FG
Mascagnia eggersiana (Nied.) W.R. Anderson
Mascagnia glandulifera Cuatrec.
Mascagnia guianensis W.R. Anderson GU SU FG
Mascagnia liesneri W.R. Anderson
Mascagnia macrodisca (Triana & Planch.) Nied.
Mascagnia microcarpa (Sandwith) W.R. Anderson
Mascagnia ovatifolia (Kunth) Griseb.
Mascagnia poeppigiana (A. Juss.) W.R. Anderson AM GU
Mascagnia schunkei W.R. Anderson AM FG
Mascagnia sepium (A. Juss.) Griseb. AM BO GU SU FG
Mascagnia sinemariensis (Aubl.) Griseb. AM BO GU FG
Mascagnia stannea (Griseb.) Nied.
Mascagnia surinamensis (Kosterm.) W.R. Anderson AM GU SU
Mezia angelica W.R. Anderson [expected in SU] GU FG
Mezia curranii W.R. Anderson
Mezia huberi W.R. Anderson
Mezia includens (Benth.) Cuatrec. AM BO GU FG
Mezia rufa W.R. Anderson
BO
AM BO
AM BO
AM
AM
AM
AM
BO
AM
AM
AM
AM BO
AM
AM
AM BO
BO
AM
AM
AM
GU SU
GU
Checklist of the Plants of the Guiana Shield 389
GU
BO GU
BO DA
DA
DA
GU SU FG
DA GU
GU?
SU FG
GU SU FG
FG
DA GU SU FG
AM BO DA GU SU FG
GU SU FG
GU
BO
GU SU FG
AM BO DA GU SU FG
BO DA GU SU FG
AM BO GU SU
SU
FG
FG
AM
BO GU
GU
SU?
AM BO
BO
BO
BO
GU
GU
GU
GU
SU FG
AM BO GU SU FG
Pterandraflavescens Maguire AM
Pterandra gidanensis W.R. Anderson [expected in BO]
Pterandra sericea W.R. Anderson
Spachea elegans (G. Mey.) A. Juss.
Stigmaphyllon adenodon A. Juss.
Stigmaphyllon adenodon var. adenodon
Stigmaphyllon bannisterioides (L.) C.E. Anderson
Stigmaphyllon ciliatum (Lam.) A. Juss.
Stigmaphyllon convolvulifolium A. Juss. [expected in DA]
Stigmaphyllon palmatum (Cav.) A. Juss.
Stigmaphyllon puberum (Rich.) A. Juss.
Stigmaphyllon sinuatum (DC.) A. Juss.
Tetrapterys acutifolia Cav.
Tetrapterys anomala W.R. Anderson
Tetrapterys aristeguietae W.R. Anderson
Tetrapterys calophylla A. Juss.
Tetrapterys crispa A. Juss.
Tetrapterys discolor (G. Mey.) DC.
Tetrapterysfimbripetala A. Juss.
Tetrapterys glabrifolia (Griseb.) Small
Tetrapterys gracilis W.R. Anderson
Tetrapterys maranhamensis A. Juss.
Tetrapterys megalantha W.R. Anderson
Tetrapterys mucronata Cav.
Tetrapterys oleifolia (Benth.) Griseb.
Tetrapterys pusilla Steyerm.
Tetrapterys rhodopteron Oliv.
Tetrapterys styloptera A. Juss.
Synonyms and Misidentifications
Alcoceratothrix rugosa (Benth.) Nied. ^Byrsonima stipulacea
Alcoceratothrix stipulacea (A. Juss.) Nied. =Byrsonima stipulacea
Banisteria argentea (Kunth) Spreng. =Banisteriopsis muricata
Banisteria caapi Griseb. =Banisteriopsis caapi
Banisteria cristata Griseb. =Banisteriopsis cristata
Banisteria fagifolia DC. =Hiraea fagifolia
Banisteria heterophylla Willd. [non Stigmaphyllon heterophyllum Hook. 1 843] =Stigmaphyllon sinuatum
Banisteria leona Cav. =Heteropterys leona
Banisteria leptocarpa Benth. =Banisteriopsis martiniana
Banisteria lucida Rich. =Banisteriopsis lucida
Banisteria macradena DC. =Heteropterys macradena
Banisteria martiniana A. Juss. =Banisteriopsis martiniana
Banisteria metallicolor A. Juss. =Banisteriopsis muricata
Banisteria mossii C.V. Morton =Heteropterys nervosa
Banisteria multiflora DC. =Heteropterys leona
Banisteria muricata Cav. =Banisteriopsis muricata
Banisteria orbicularis Nied. =Banisteriopsis cristata
Banisteria orinocensis Kunth =Heteropterys orinocensis
Banisteria ovata Cav. =Stigmaphyllon bannisterioides
Banisteria pubera Rich. =Stigmaphyllon puberum
Banisteria quapara Aubl. =Hiraea quapara
Banisteria reticulata (Poir.) C.B. Rob. =Heteropterys leona
Banisteria schomburgkiana Benth. =Banisteriopsis muricata
Banisteria sinemariensis Aubl. =Mascagnia sinemariensis
Banisteria sinuata DC. =Stigmaphyllon sinuatum
Banisteria splendens DC. [nom. superfl.] =Stigmaphyllon sinuatum
Banisteriopsis argentea (Kunth) C.B. Rob. =Banisteriopsis muricata
Banisteriopsis cabrerana Cuatrec. =Diplopterys cabrerana
Banisteriopsis elegans subsp. cordata (Nied.) Sandwith [not in flora area]
Banisteriopsis elegans war. pulcherrima Sandwith ^Banisteriopsis pulcherrima
Banisteriopsis gardneriana (A. Juss.) W.R. Anderson & B. Gates [reported from FG in error, fide W. Anderson]
Banisteriopsis inebrians C.V. Morton ^Banisteriopsis caapi
Banisteriopsis leptocarpa (Benth.) R.O. Williams =Banisteriopsis martiniana
Banisteriopsis martiniana var. laevis Cuatrec. =Banisteriopsis martiniana var. subenervia
390 Checklist of the Plants of the Guiana Shield
Banistehopsis metallicolor (A. Juss.) O'Donell & Lourteig =Banisteriopsis muhcata
Banisteriopsis schombwgkiana (Benth.) C.B. Rob. =Banisteriopsis muricata
Blepharandra cretacea (Gleason) Steyenn. =Blepharandra hypoleuca
Blepharandra ptariana Steyerm. =Blepharandra hypoleuca
Brachypterys borealis A. Juss. [nom. superfl.] =Stigmaphyllon bannisterioides
Brachypterys ovata (Cav.) Small =StigmaphyUon bannisterioides
Bunchosia maritima (Veil.) J.F. Macbr. [not in flora area]
Bunchosia nitida (Jacq.) Rich, [not in flora area]
Bunchosia rhombifolia Turcz. =Bunchosia mollis
Bunchosia schomburgkiana Nied. ^Bunchosia mollis
Bunchosia squarrosa Griseb. =Tetrapterys styloptera
Bunchosia tuberculata DC. [dubious name of uncertain status, fide W. Anderson]
Burdachia atractoides Nied. =Burdachia sphaerocarpa
Burdachia prismatocarpa var. sphaerocarpa (A. Juss.) Nied. =Burdachia sphaerocarpa
Burdachia sphaerocarpa var. glandifera (Gleason) W.R. Anderson =Burdachia sphaerocarpa
Burdachia williamsii Steyerm. =Burdachla prismatocarpa
Byrsonima angustifolia Kunth =Blepharandra angustifolia
Byrsonima aubletii Kosterm. =Byrsonima altissima
Byrsonima barkleyana Cuatrec. =Byrsonima punctulata
Byrsonima bolivarana Steyerm. =Byrsonima carraoana
Byrsonima bracteolaris Benth. =Byrsonima concinna
Byrsonima carmeniana Cuatrec. =Byrsonima crispa
Byrsonima chalcophylla var. carraoana (Steyerm.) W.R. Anderson =Byrsonima carraoana
Byrsonima coriacea (Sw.) DC. [not in flora area]
Byrsonima coriacea var. spicata (Cav.) Nied. [nom. superfl.] =Byrsonima spicata
Byrsonima cretacea Nied. =Byrsonima luetzelburgii
Byrsonima cretacea Gleason =Blepharandra hypoleuca
Byrsonima cydoniifolia A. Juss. [not in flora area]
Byrsonima discolor Pilg. =Byrsonima altissima
Byrsonimaferruginea Kunth =Byrsonima crassifolia
Byrsonimaferruginea var. macrophylla Benth. =Byrsonima aerugo
Byrsonima fluminensis Nied. =Byrsonima japurensis
Byrsonima inundata Benth. =Byrsonimajapurensis
Byrsonimajapurensis subsp. fluminensis (Nied.) Cuatrec. =Byrsonimajapurensis
Byrsonimajapurensis subsp. silvatica Cuatrec. =Byrsonimajapurensis
Byrsonima laurifolia Kunth =Byrsonima crassifolia
Byrsonima obversa Miq. =Byrsonima laevigata
Byrsonima propinqua Benth. =Byrsonima spicata
Byrsonima reticulata (Poir.) DC. =Heteropterys leona
Byrsonima rugosa Benth. =Byrsonima stipulacea
Byrsonima sessilifolia Benth. =Byrsonima coccolobifolia
Byrsonima uvulifera Spruce ex Nied. =Byrsonimajapurensis
Byrsonima verbascifolia var. denudata Cuatrec. =Byrsonima verbascifolia
Clonodia racemosa var. orinocensis Nied. =Clonodia complicata
Coleostachys hypoleuca Benth. ^^Blepharandra hypoleuca
Coleostachys vestita Benth. =Diacidia vestita
Diacidia duckeana Maguire =Diacidia galphimioides
Diacidia parvifolia Cuatrec. =Diacidia galphimioides
Dicella amazonica Fires =Dicellajulianii
Diplopterys includens (Benth.) Nied. =Mezia includens
Diplopterys involuta (Turcz.) Nied. =Mezia includens
Diplopterys involuta var. ovata Nied. =Mezia curranii
Diplopterys microcarpa Sandwith =Mascagnia microcarpa
Diplopterys riparia (A. Juss.) Nied. =Jubelina riparia
Diplopterys rosea (Miq.) Nied. =Jubelina rosea
Diplopterys spruceana (Nied.) Nied. [not in flora area] =Mascagnia bracteosa Griseb.
Dolichopterys surinamensis Kosterm. =Lophopterys surinamensis
Galphimia longifolia Kunth =Lophanthera longifolia
Heteropterys acutifolia A. Juss. =Heteropterys orinocensis
Heteropterys aflicana A. Juss. [nom. superfl.] =Heteropterys leona
Heteropterys argentea Kunth =Banisteriopsis muricata
Heteropterys beecheyana var. alata W.R. Anderson =Heteropterys alata
Heteropterys carinata Benth. =Heteropterys cristata
Heteropterys castanea Cuatrec. =Mascagma castanea
Checklist of the Plants of the Guiana Shield 391
Heteropterys catoptera W.R. Anderson =Heteropterys subhelicina
Heteropterys helicina Griseb. =Heteropterys orinocensis
Heteropterys lasseri W.R. Anderson =Heteropterys megaptera
Heteropterys mossii (C.V. Morton) Cuatrec. =Heteropterys nervosa
Heteropterys multiflora (DC.) Hochr. =Heteropterys leona
Heteropterys oleifolia (Benth.) Griseb. =Tetrapterys oleifolia
Heteropterys reticulata (Poir.) Nied. =Heteropterys leona
Heteropterys stannea Griseb. =Mascagnia stannea
Heteropterys suberosa Griseb. =Heteropterys nervosa
Hiraea adenophora Sandwith =Excentradenia adenophora
Hiraea bifurcata W.R. Anderson =Hiraea ternifolia
Hiraea chrysophylla A. Juss. =Hiraeafaginea
Hiraea complicata Kunth =Clonodia complicata
Hiraea demerarensis (A. Juss.) Nied. =Hiraeafaginea
Hiraea divaricaa Kunth =Mascagma divaricata
Hiraeafulgens A. Juss. =Hiraeafaginea
Hiraeafulgens van demerarensis A. Juss. =Hiraeafaginea
Hiraea gaudichaudiana A. Juss. [not in flora area]
Hiraea gracilis Benth. =Tetrapterys maranhamensis
Hiraea laurifolia A. Juss. =Mascagnia stannea
Hiraea macrodisca Triana & Planch. =Mascagnia macrodisca
Hiraea multiradiata A. Juss. =Hiraea quapara
Hiraea nitida Kunth =Clonodia complicata
Hiraea oblongifolia DC. =Mascagnia divaricata
Hiraea oleifolia Benth. =Tetrapterys oleifolia
Hiraea ovatifolia Kunth =Mascagma ovatifolia
Hiraea platytriphylla Hochr. =Hiraea ternifolia
Hiraea poeppigiana A. Juss. =Mascagma poeppigiana
Hiraea propinqua W.R. Anderson =Excentradenia propinqua
Hiraea schizoptera Turcz. =Mascagnia sinemariensis
Hiraea sepium A. Juss. =Mascagnia sepium
Hiraea swartziana A. Juss. [nom. superfl.] =Hiraeafaginea
Jubelina bracteosa auct. non Mascagnia bracteosa Griseb. =Jubelina grisebachiana
Lophopterys splendens var. obovata Nied. =Lophopterys euryptera
Malpighia altissima Aubl. =Byrsonima altissima
Malpighia argentea Jacq. =Bunchosia argentea
Malpighia armeniaca Cav. =Bunchosia armeniaca
Malpighia bannisterioides L. =Stigmaphyllon bannisterioides
Malpighia crassifolia L. =Byrsonima crassifolia
Malpighia elegans G. Mey. =Spachea elegans
Malpighiafaginea Sw. =Hiraeafaginea
Malpighia glandulifera Jacq. =Bunchosia glandulifera
Malpighia moureila Aubl. =Byrsonima crassifolia
Malpighia punicifolia L. =Malpighia glabra
Malpighia punicifolia auct. non L. 1762 =Malpighia emarginata
Malpighia reticulata Poir. =Heteropterys leona
Malpighia spicata Cav. =Byrsonima spicata
Malpighia ternifolia Kunth =Hiraea ternifolia
Malpighia verbascifolia L. =Byrsonima verbascifolia
Malpighia volubilis Sims =Mascagnia sinemariensis
Mascagnia anisopetala (A. Juss.) Griseb. [not in flora area]
Mascagnia anisopetala var. macrodisca (Triana & Planch.) Kosterm. =Mascagma macrodisca
Mascagnia benthamiana (Griseb.) W.R. Anderson =Mascagnia stannea
Mascagnia complicata (Kunth) Nied. =Clonodia complicata
Mascagnia cordifolia var. peruviana J.F. Macbr. ^Mascagnia schunkei
Mascagnia hondensis C.V. Morton =Mascagnia sinemariensis
Mascagnia lehmanniana Nied. =Clonodia complicata
Mascagnia leucanthele Griseb. [not in flora area]
Mascagnia materdei Cuatrec. =Mascagnia poeppigiana
Mascagnia multiglandulosa Nied. [not in flora area]
Mascagnia multiglandulosa var. surinamensis Kosterm. =Mascagnia surinamensis
Mascagnia nervosa Nied. =Mascagnia ovatifolia
Mascagnia nitida (Kunth) Nied. =Clonodia complicata
Mascagnia pauciflora (G. Mey.) Cuatrec. =Diplopterys pauciflora
392 Checklist of the Plants of the Guiana Shield
Mascagnia schizoptera (Turcz.) Cuatrec. =Mascagnia sinemahensis
Mascagnia sericans Nied. =Mascagnia stannea
Mascagnia subsericea Cuatrec. =Mascagma poeppigiana
Mascagnia tenuifolia var. eggersiana Nied. =Mascagnia eggersiana
Mascagnia volubilis (Sims) Nied. =Mascagnia sinemahensis
Sipapoa cordata Maguire =Diacidia cordata
Sipapoaferruginea Maguire & K.D. Phelps ==Diacidiaferruginea
Sipapoa glaiicifolia Maguire =Diacidia glaucifolia
Sipapoa hypoleuca Maguire =Diacidia hypoleuca
Sipapoa kiinhardtii Maguire =Diacidia kitnhardtii
Sipapoa rufa Maguire =Diacidia rufa
Sipapoa steyermarkii Maguire =Diacidia steyermarkii
Sipapoa stipularis Maguire & K.D. Phelps =Diacidia stipularis
Sipapoa vestita (Benth.) Maguire =Diacidia vestita
Skoliopterys lehmanniana (Nied.) Cuatrec. =Clonodia complicata
Stenocalyx involutus Turcz. =Mezia includens
Stigmaphyllon angustilobum A. Juss. [not in flora area]
Stigmaphyllon brachiatum Triana & Planch. =StigmaphyUon sinuatum
Stigmaphyllon fulgens A. Juss. [nom. superfl.] =Stigmaphyllon sinuatum
Stigmaphyllon grenadense Nied. =Stigmaphyllon adenodon
Stigmaphyllon hypoleucum Miq. =Stigmaphyllon sinuatum
Stigmaphyllon latifolium Benth. =Stigmaphyllon convolvulifolium
Stigmaphyllon martianum A. Juss. ^Stigmaphyllon sinuatum
Stigmaphyllon mathiasiae W.R. Anderson =Banisteriopsis mathiasiae
Stigmaphyllon monancistrum Nied. =Stigmaphyllon sinuatum
Stigmaphyllon ovatum (Cav.) Nied. =Stigmaphyllon bannisterioides
Stigmaphyllon periplocifolium A. Juss. [not in flora area]
Stigmaphyllon purpureum Benth. ^Stigmaphyllon sinuatum
Stigmaphyllon sagittatum A. Juss. =^Stigmaphyllon palmatum
Stigmaphyllon splendens Cuatrec. [nom. superfl.] =Stigmaphyllon sinuatum
Stigmaphyllon tiliifolium Nied. [not in flora area]
Tetrapodenia glandifera Gleason =Burdachia sphaerocarpa
Tetrapterys benthamiana Griseb. =Mascagnia stannea
Tetrapteiys boliviensis Nied. =Tetrapterys styloptera
Tetraptetys crebriflora A. Juss. =Tetraptetys mucronata
Tetrapterys glaberrima Benth. =Tetrapterys mucronata
Tetrapteiys huachamacariensis W.R. Anderson =Tetrapterys fimbripetala
Tetrapterys includens Benth. -Mezia includens
Tetrapterys julianii J.F. Macbr. =Dicellajulianii
Tetrapterys mucronata var. crebriflora (A. Juss.) J.F. Macbr. =Tetrapterys mucronata
Tetrapterys mucronata subsp. crebriflora (A. Juss.) Kosterm. =Tetrapterys mucronata
Tetrapterys phylladenophora Maguire & Steyerm. =Tetraptetys rhodopteron
Tetrapterys poeppigiana (A. Juss.) Griseb. =Mascagnia poeppigiana
Tetraptetys silvatica Cuatrec. =Tetrapterys mucronata
Tetrapteiys squarrosa (Griseb.) Griseb. =Tetrapterys styloptera
Triopterys discolor G. Mey. =Tetrapterys discolor
References
Anderson, C. 1993. The identities of the sericeous-leaved species of Stigmaphyllon in the Amazon region. Contr. Univ. Mich.
Herb. 19: 393-413.
Anderson, C. 1997. Monograph of Stigmaphyllon (Malpighiaceae). Systematic Botany Monographs 51: 1-313.
Anderson, W.R. 1981. Malpighiaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 1 1. Mem. N.Y. Bot.
Gard. 32: 1-391.
Anderson, W.R. 1990. The taxonomy of Jubelina. Contr. Univ. Mich. Herb. 17: 21-37.
Anderson, W.R. 1993. Notes on Neotropical Malpighiaceae - IV. Contr. Univ. Mich. Herb. 19: 355-392.
Anderson, W.R. 1994. New species of Hiraea from the Guianas and adjacent Brazil. Brittonia 46(2): 126-133.
Anderson, W.R. 1993. Notes on Neotropical Malpighiaceae - VI. Contr. Univ. Mich. Herb. 21: 37-84.
Anderson, W.R. 2001. Malpighiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 6: 82-185.
Anderson, W.R. 2001. Malpighiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 410-428.
Gates, B. 1982. Banisteriopsis, Diplopterys. Flora Neotropica Monogr. 30: 1-237.
Gorts-van Rijn, A.R.A. & M.J. Jansen-Jacobs. 1976. Malpighiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers,
Flora of Suriname 2(2): 445-450.
Checklist of the Plants of the Guiana Shield 393
Jonker, F.P. 1939. Malpighiaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 478-480.
Kostermans, A.J.G.H. 1936. Malpighiaceae. In A. Pulle, Flora of Suriname 2(1): 146-243.
Niedenzu, F. 1928. Malpighiaceae. In A. Engler, Das Pflanzenreich IV. 141: 1-870.
BO
Malvaceae
L.J. Dorr, P.A. Fryxell, M.J. Jansen-Jacobs
Abelmoschus moschatus Medik. [cultivated] BO
Abelmoschus spp. [cultivated]
Abutilon indicum (L.) Sweet [cultivated]
Allosidastrum pyramidatum (Desp. ex Cav.) Krapov., Fryxell & D.M. Bates
Bastardia viscosa (L.) Kunth [introduced]
Briquetia spicata (Kunth) Fryxell
Cienfuegosia ajfinis (Kunth) Hochr.
Cienfuegosia heterophylla (Vent.) Garcke
Gaya gaudichaudiana A. St.-Hil.
Gaya occidentalis (L.) Sweet
Gossypium barbadense L. [cultivated only?]
Gossypium hirsutum L. [cultivated only?]
Hehssantia crispa (L.) Brizicky
Hibiscus bifurcatus Cav.
Hibiscus dimidiatus Schrank
Hibiscus furcellatus Desr.
Hibiscus guianemis Aubl. [name of uncertain application]
Hibiscus mutabilis L. [cultivated?]
Hibiscus peruvianus R.E. ¥r. AM
Hibiscus phoeniceus Jacq. [cultivated?]
Hibiscus prunifolius F. Dietr.
Hibiscus radiatus Cav. [cultivated]
Hibiscus rosa-sinensis L. [cultivated]
Hibiscus sabdariffa L. [cultivated]
Hibiscus schizopetalus (Dyer) Hook. f. [cultivated]
Hibiscus sororius L.
Hibiscus sthatus Cav.
Hibiscus striatus subsp. lambertianus (Kunth) O.J. Blanch, ex Proctor
Hibiscus trilobus Aubl.
Hibiscus trilobus var. ingratus (Miq.) O.J. Blanch. & Fryxell
Hibiscus varians Splitg.
Hibiscus verbasciformis Klotzsch ex Hochr.
Malachra alceifolia Jacq.
Malachrafasciata Jacq.
Malachra palmata Moench
Malachra radiata (L.) L.
Malvastrum coromandelianum (L.) Garcke
Pavonia angustifolia Benth.
Pavonia cancellata (L.) Cav.
Pavonia castaneifolia A. St.-Hil. & Naudin
Pavonia dasypetala Turcz. AM
Pavoniaflavispina Miq.
Pavoniafruticosa (Mill.) Fawc. & Rendle
Pavonia geminiflora Mode.
Pavonia imatacensis Steyerm.
Pavonia malacophylla (Link & Otto) Garcke AM
Pavonia paludicola Nicolson ex Fryxell
Pavonia paniculata Cav.
Pavonia piptocalyx Krapov.
Pavonia schiedeana Steud.
Pavonia sidifolia Kunth
Peltaea riedelii (Giirke) Standi.
Peltaea sessiliflora (Kunth) Standi. AM
Peltaea speciosa (Kunth) Standi. AM
Peltaea surumuensis (Ulbr.) Krapov. & Cristobal [possibly expected in the BO
Guianas]
Peltaea trinervis (C. Presl) Krapov. & Cristobal BO
DA
GU SU FG
GU SU
SU
FG?
BO GU SU FG
BO GU
BO DA
BO
GU
BO DA GU SU FG
BO GU SU FG
BO GU
BO DA GU
GU
SU FG
BO DA GU SU FG
SU
FG
GU FG
GU?
BO GU SU
GU SU FG
BO GU SU FG
GU SU FG
DA GU SU FG
DA GU SU
DA GU SU
BO SU FG
BO
GU?
SU FG
GU SU FG
BO DA GU SU FG
DA GU SU FG
BO GU
DA GU
GU
SU
FG
BO GU
BO GU SU FG
BO GU SU FG
FG?
BO DA GU
GU
SU FG
BO
BO
DA GU
GU
SU FG
FG
GU SU FG
BO
GU
GU
SU
BO GU SU
GU SU
BO
AM BO
BO
BO
DA GU
GU
SU FG
BO DA GU
SU
BO DA GU SU FG
AM BO
BO
GU SU FG
AM BO DA GU
GU
SU FG
AM BO DA GU
GU
SU FG
AM BO GU SU FG
AM BO DA GU
GU
GU
SU FG
FG
GU SU FG
BO GU SU
BO GU
GU SU FG
DA GU SU FG
DA GU
GU
SU
SU
SU
FG
394 Checklist of the Plants of the Guiana Shield
Pseudabutilon umbellatum (L.) Fryxell
Sida acuta Burm. f.
Sida aggregata C. Presl
Sida angustissima A. St.-Hil.
Sida cilia?!s L.
Sida cordata (Burm. f.) Borss. Waalk.
Sida cordifolia L.
Sida glomerata Cav.
Sidajamaicensis L.
Sida linifolia Juss. ex Cav.
Sida poeppigiana (K. Schum.) Fryxell
Sida rhombifolia L.
Sida santaremensis Monteiro
Sida serrata Willd. ex Spreng.
Sida setosa Mart, ex Colla
Sida spinosa L.
Sida surumuensis Ulbr.
Sida urens L.
Sida viarum A. St.-Hil.
Sidastrum micranthum (A. St.-Hil.) Fryxell
Sidastrum quinquenerviiim (Duchass. ex Triana & Planch.) Baker f.
Talipariti tiliaceum (L.) Fryxell
Talipariti tiliaceum \ar. pernambucense (Arruda) Fryxell
Thespesia lampas (Cav.) Dalzell & Gibson [cultivated]
Thespesia populnea (L.) Sol. ex Correa [cultivated or escaped from
cultivation]
Urena lobata L. AM GU SU FG
Urena sirtuata L.
Wissadula contracta (Link) R.E. Fr.
Wissadula excelsior {CdM.)C.?rQs\ BO GU SU
Wissadula hernandioides (L'Her.) Garcke
Wissadula hirsuta C. Presl
Wissadula penplocifolia{L.)C.?XQs\QxT\m?k\XQS, BO GU SU
Synonyms and Misidentifications
Abutilon crispum (L.) Medik. =Herissantia crispa
Abutilon patens A. St.-Hil. =Wissadula excelsior
Abutilon spicatum Kunth =Briquetia spicata
Althaea racemosa Sw. =Pavonia paludicola
Bogenhardia crispa (L.) Kearney =Herissantia crispa
Cienfuegosia phlomidifolia (A. St.-Hil.) Garcke =Cienfuegosia affinis
Fugosia campestris Benth. =Cienfuegosia affinis
Fugosia guianensis Klotzsch ex M.R. Schomb. =Cienfuegosia affinis
Fugosia phlomidifolia A. St.-Hil. =Cienfuegosia affinis
Fugosia retusa Turcz. ^Cienfuegosia affinis
Gayoides crispum (L.) Small =Herissantia crispa
Gossypium latifolium Murr. =Gossypium hirsutum
Gossypium peruvianum Cav. =Gossypium barbadense
Gossypium punctatum Schumach. & Thonn. =Gossypium hirsutum
Gossypium religiosum L. =Gossypium hirsutum
Gossypium vitifolium Lam. =Gossypium barbadense
Hibiscus abelmoschus L. =Abelmoschus moschatus
Hibiscus affinis Kunth =Cienfuegosia affinis
Hibiscus bracteosus DC. =Talipariti tiliaceum var. pernambucense
Hibiscus brasiliensis L. [not in flora area]
Hibiscus cancellatus L. =Pavonia cancellata
Hibiscus cubensis A. Rich. =Hibiscus striatus subsp. lambertianus
Hibiscus elatus Sw. ^Talipariti tiliaceum
Hibiscus esculentus L. [cultivated] =Abelmoschus esculentus (L.) Moench
Hibiscus fraternus L. =Hibiscus sabdariffa
Hibiscus ingratiis Miq. =Hibiscus trilobus var. ingratus
Hibiscus lambertianus Kunth =Hibiscus striatus subsp. lambertianus
Hibiscus lampas Cav. ^Thespesia lampas
Hibiscus moschatus (Medik.) Salisb. =Abelmoschus moschatus
Hibiscus pernambucensis Arruda =Talipariti tiliaceum var. pernambucense
AM BO DA
BO GU
DA
GU
Checklist of the Plants of the Guiana Shield 395
Hibiscus salviifoUus A. St.-Hil. =Hibisciis striatus subsp. lambertianus
Hibiscus spathulatus Garcke =Hibiscus verbasciformis
Hibiscus tiliaceus L. =TaIipariti tiliaceum
Lopimia dasypetala (Turcz.) Standi. =Pavoma dasypetala
Lopimia malacophylla (Link & Otto) Mart. =Favonia malacophyUa
Malache chiapensis Standi. =Pavonia sidifolia
Malache scabra B. Vogel ^Pavonia pahidicola
Malachra alceifolia vav.fasciata (Jacq.) A. Robyns =Malachrafasciata
Malachra alceifolia var. rotundifolia (Schrank) Guerke =Malachra alceifolia
Malachra capitata L. =Malachra alceifolia
Malachra heptaphylla Fischer =Malachra palmata
Malachra kegeliana Garcke =Malachrafasciata
Malachra trinervis C. Presl =Peltaea trinervis
Pavonia chiapensis (Standi.) Standi. =Pavoma sidifolia
Pavonia guanacastensis Standi. =Pavonia cancellata
Pavoniajulianae Uittien =Peltaea riedelii
Pavonia racemosa (Sw.) Sw. =Pavonia paludicola
Pavonia rosea Schltdl. =Pavonia schiedeana
Pavonia scabra (B. Vogel) Cif. =Pavoma paludicola
Pavonia sessiliflora Kunth =Peltaea sessilifora
Pavonia speciosa Kunth =Peltaea speciosa
Pavonia spicata Cav. [nom. illeg.] =Pavonia paludicola
Pavonia spinifex Cav. [not in flora area]
Pavonia surumuensis Ulbr. =Peltaea surumuensis
Pavonia trinervis (C. Presl) A. Robyns =Peltaea trinei-vis
Pavonia typhalaea Cav. =Pavoniafruticosa
Pavonia ulbrichiana Kearney =Hibiscus dimidiatus
Peltaea aliculata Krapov. & Cristobal =Peltaea surumuensis
Pseudabutilon spicatum (Kunth) R.E. Fr. =Briquetia spicata
Redutea heterophylla Vent. =Cienfuegosia heterophylla
Sida carpinifolia L. f. =Sida acuta
Sida contracta Link =Wissadula contracta
Sida crispa L. =Herissantia crispa
Sida excelsior Cav. =Wissadula excelsior
Sidafruticosa Mill. =Pavonia fruticosa
Sida guianensis K. Schum. =Sidastrum quinquenervium
Sida hernandioides L'Her. =Wissadula hernandioides
Sida malacophylla Link & Otto =Pavonia malacophylla
Sida micrantha A. St.-Hil. =Sidastrum micranthum
Sida periplocifolia L. =Wissadula periplocifolia
Sida quinquenervia Duchass. ex Triana & Planch. =Sidastrum quinquenei-vium
Sida radiata L. =Malachra radiata
Sida savannarum K. Schum. =Sida aggregata
Sida setifera C. Presl =Sida aggregata
Sida surinamensis Miq. =Sida setosa
Sida umbellata L. =Pseudabutilon umbellatum
Sida veronicifolia Lam. =Sida cordata
Urena lobata var. sinuata (L.) Hochr. [s.l.] =Urena sinuata
Urena moenchii Gomez =Malachra palmata
Urena reticulata Cav. =Urena lobata
Wissadula amplissima (L.) R.E. Fr. [non Sida amplissima L.] =Wissadula hernandioides
Wissadula patens (A. St.-Hil.) Garcke "^Wissadula excelsior
Wissadula spicata (Kunth) C. Presl =Briquetia spicata
Wissadula zeylanica Medik. =Wissadula periplocifolia
Rl f CRENCES
Clement, I.D. 1957. Studies in Sida. 1. Contrib. Gray Herb. 180: 1-91.
Dorr, L.J. 2002. Malvaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. HofT, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 428-
431.
Fryxell, PA. 2001. Malvaceae. In J. A. Steycrmark, PE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
6: 186-219.
Jansen-Jacobs, M.J. 1986. Malvaceae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 3(1-2):
259-276.
Kearney, T.H. 1957. A tentative key to the South American species of Hibiscus L. Leafl. W. Bot. 8(7): 161-168.
396 Checklist of the Plants of the Guiana Shield
Kearney, T.H. 1958. A tentative key to the South American species of Pavonia Cav. Leafl. W. Bot. 8(10): 225-248.
Kearney, T.H. 1958. A tentative key to the South American species of Sida L. Leafl. W. Bot. 8(1 1): 249-270.
Krapovickas, A. & C.L. Cristobal. 1965. Revision del genero Peltaea. Kurtziana 2: 135-216.
Uittien, H. 1932. Malvaceae. In A. Pulle, Flora of Suriname 3(1): 1-25.
Uittien, H. 1941. Malvaceae (additions and corrections). In A. Pulle, Flora of Suriname 3(1): 433-435.
Marcgraviaceae
A.C. de Roon, S. Dressier
Marcgravia coriacea Vahl
Marcgravia eichleriana Wittm.
Marcgravia magnibracteata Lanj. & Heerdt
Marcgravia magiiirei de Roon
Marcgravia patellulifera de Roon
Marcgravia pedimculosa Triana & Planch.
Marcgravia piinctifolia S. Dressier
Marcgravia purpurea I.W. Bailey
Marcgravia sororopaniana Steyerm.
Marcgravia sprucei (Wittm.) Gilg
Marcgraviastrum mixtum (Triana & Planch.) Bedell ex de Roon & S. Dressier AM
Marcgraviastrum pendulum (Lanj. & Heerdt) Bedell
Norantea guianensis Aubl.
Norantea guianensis subsp. guianensis
Norantea guianensis subsp. japurensis (Mart.) Bedell
Sarcoperaflannnifera de Roon & Bedell
Sarcopera tepuiensis (de Roon) Bedell
Sarcopera tepuiensis subsp. coccinea de Roon & S. Dressier
Sarcopera tepuiensis subsp. tepuiensis
Souroubea dasystachya Gilg & Werderm.
Souroubea guianensis Aubl.
Souroubea guianensis subsp. cylindrica (Wittm.) de Roon
Souroubea guianensis subsp. guianensis
Synonyms and Misidentifications
Marcgravia cuyuniensis I.W. Bailey =Marcgravia coriacea
Marcgravia dasystachya Gilg ex Ule [nomen nudum]
Marcgravia dubia Kunth =(Araceae) Monstera dubia
Marcgravia gracilis Sagot =Marcgravia pedunculosa
Marcgravia parviflora Rich, ex Wittm. ^Marcgravia pedunculosa
Marcgravia parviflora var. macrophylla Wittm. =Marcgravia pedunculosa
Marcgravia parviflora var. sprucei Wittm. =Marcgravia sprucei
Marcgravia picta Willd. [dubious species, misident. fide de Roon]
Marcgravia roraimae Gilg & Werderm. =Marcgravia coriacea
Marcgravia roraimensis Gilg ex Ule [nomen nudum]
Marcgravia steyermarkii de Roon =Marcgravia sororopaniana
Marcgravia umbellata L. [Guianan records misident., fide de Roon]
Norantea guianensis var. gracilis Wittm. =Norantea guianensis subsp. japurensis
Noranteajapurensis Mart. =Norantea guianensis subsp. japurensis
Norantea mixta Triana & Planch. =Marcgraviastrum mixtum
Norantea paracusis Mart. =Norantea guianensis subsp. japurensis
Norantea peduncularis Poepp. ex Wittm. =Marcgraviastrum mixtum
Norantea pendula Lanj. & Heerdt =Marcgraviastrum pendulum
Norantea tepuiensis de Roon =Sarcopera tepuiensis
Souroubea guianensis var. cylindrica Wittm. =Souroubea guianensis subsp. cylindrica
Souroubea guianensis var. tomentella Steyerm. =Souroubea guianensis subsp. cylindrica
References
Bailey, I.W. 1922. The pollination of Marcgravia: a classical case of omithophily? Amer. J. Bot. 9: 370-384.
De Roon, A.C. 1970. Anew species of Marcgravia (Marcgraviaceae) from Venezuela. Acta Bot. Neerl. 19 (6): 799-803.
De Roon, A.C. & S. Dressier. 1999. New taxa of Sarcopera and Marcgraviastrum (Marcgraviaceae) from the Guayana Shield.
Novon 9: 248-252.
Dressier, S. 2001
.
Marcgraviaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan Guayana
6: 248-260.
Heald, S.V, A.C. De Roon, & S. Dressier. Marcgraviaceae. 1999. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V
Heald, M. Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons.
Mem. New York Bot. Card. 76 (2): 431-437.
AM BO DA GU SU FG
AM
GU su FG
AM BO FG
AM
AM
BO
GU SU FG
AM BO GU su FG
BO GU FG?
AM
AM BO GU su FG
AM BO DA GU su FG
AM BO DA GU su FG
AM BO
AM BO GU?
AM BO GU
AM
BO GU
AM BO GU
AM BO DA GU su FG
AM BO DA GU
GU su FG
Checklist of the Plants of the Guiana Shield 397
Lanjouw, J. & P.F. van Heerdt. 1941. Marcgraviaceae. In A. Pulle, Flora of Suriname 3(1): 373-385.
Melastomataceae
F. Almeda, RE. Berry, A. Freire-Fierro, A. Groger, B.K. Hoist, N.G. Luckana, F.A. Michelangeli,
T. Morley, D. Penney, S.S. Renner, O.R. Robinson, J.J. Wurdackj, K. Yatskievych
Acanthella piilchra Gleason
Acanthella sprucei Hook. f.
Aciotis acuminifolia (Mart, ex DC.) Triana
Aciotis annua (Mart, ex DC.) Triana [expected in DA]
Aciotis circaeifolia (Bonpl.) Triana
Aciotis indecora (Bonpl.) Triana
Aciotis olivieriana Freire-Fierro
Aciotis ornata (Miq.) Gleason
Aciotis polystachya (Bonpl.) Triana
Aciotis purpurascens (Aubl.) Triana
Aciotis rubricaulis (Schrank & Mart, ex DC.) Triana
Aciotis viscida (Benth.) Freire-Fierro
Acisanthera bivalvis (Aubl.) Cogn.
Acisanthera crassipes (Naudin) Wurdack
Acisanthera hedyotidea (C. Presl) Triana
Acisanthera limnobios (DC.) Triana
Acisanthera nana Ule
Acisanthera quadrata Pers.
Acisanthera rosulans Huber
Acisanthera tetraptera (Cogn.) Gleason
Acisanthera uniflora (Vahl) Gleason
Adelobotrys adscendens (Sw.) Triana
Adelobotrys ayangannensis Wurdack
Adelobotrys barbata Triana
Adelobotrys ciliata (Naudin) Triana
Adelobotrys duidae (Gleason) Wurdack
Adelobotrysfruticosa Wurdack
Adelobotrys linearifolia L. Uribe
Adelobotrys monticola Gleason
Adelobotrys permixta Wurdack
Adelobotrys rotundifolia Triana
Adelobotrys saxosa Wurdack
Adelobotrys scandens (Aubl.) DC.
Adelobotrys spruceana Cogn.
Adelobotrys stenophylla Wurdack
Appendicularia thymifolia (Bonpl.) DC.
Bellucia grossularioides (L.) Triana
Bellucia huberi (Wurdack) S.S. Renner
Bellucia pentamera Naudin
Bertolonia repens F. Michelangeli, sp. nov. ined.
Bertolonia venezuelensis Wurdack
Blakea rosea (Ruiz & Pav.) D. Don
Boyania ayangannae Wurdack
Centronia neblinae Wurdack
Chaetolepis anisandra Naudin
Chaetolepis phelpsiae Gleason
Chaetolepis phelpsiae subsp. chimantensis Wurdack
Chaetolepis phelpsiae subsp. phelpsiae
Clidemia acurensis Wurdack
Clidemia alternifolia Wurdack
Clidemia andersonii Wurdack
Clidemia anisophylla DC.
Clidemia aphanantha (Naudin) Sagot
Clidemia attenuata (Naudin) Cogn.
Clidemia ayangannensis Wurdack
Clidemia bernardii Wurdack
Clidemia bullosa DC.
Clidemia buntingii Wurdack
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM
BO
AM BO
BO
BO
AM BO
AM
AM BO
BO
AM
BO
DA
DA
DA
GU su FG
GU su FG
GU FG
GU su FG
GU su FG
GU su FG
su FG
GU
GU su FG
GU FG
GU su
GU su
GU
su FG
su FG
GU su FG
GU su FG
GU
GU SU FG
AM
AM
AM BO
AM BO
BO GU
SU
AM BO
AM
SU FG
AM BO DA FG
AM
GU SU FG
AM BO GU SU FG
AM
AM BO DA GU
AM
AM
AM BO
GU
AM
AM BO GU
AM BO
BO
AM
DA
AM
AM
GU su FG
AM BO su FG
AM GU
GU
FG
AM BO
AM BO
BO
DA GU
GU
su FG
398 Checklist of the Plants of the Guiana Shield
CUdemia capillipes (Triana) Cogn.
Clidemia capitata Benth.
CUdemia capitellata (Bonpl.) D. Don
Clidemia capitellata var. capitellata
Clidemia capitellata var. dependem (D. Don) J.F. Macbr.
Clidemia capitellata var. levelii Wurdack
Clidemia charadrophila Tutin
Clidemia conglomerata DC.
Clidemia debilis Crueg. [expected in GU]
Clidemia dentata D. Don
Clidemia duidae Gleason
Clidemia epibatehiim DC.
Clidemia epiphytica (Triana) Cogn.
Clidemia granvillei Wurdack
Clidemia heptamera Wurdack
Clidemia heteroneiira (DC.) Cogn.
Clidemia hirta (L.) D. Don
Clidemia hirta var. elegans (Aubl.) Griseb.
Clidemia hirta var. hirta
Clidemia hirta var. tiliifolia (DC.) J.F. Macbr.
Clidemia invohicrata DC.
Clidemia japiirensis DC.
Clidemiajapurensis var. heterobasis (DC.) Wurdack
Clidemiajapiirensis var. japurensis
Clidemiajiiruensis (Pilg.) Gleason
Clidemia happieri Cogn.
Clidemia laevifolia Gleason
Clidemia linearis (Gleason) Wurdack
Clidemia marahiiacensis Wurdack
Clidemia micrantha Sagot
Clidemia microthyrsa R.O. Williams
Clidemia minutiflora (Triana) Cogn.
Clidemia morichensis Wurdack
Clidemia neblinae Wurdack
Clidemia novemnervia (DC.) Triana
Clidemia octona (Bonpl.) L.O. Williams
Clidemia octona subsp. guayanensis Wurdack
Clidemia octona subsp. octona
Clidemia ostentata Wurdack
Clidemia pakaraimae Wurdack
Clidemia petiolata DC.
Clidemia piperifolia Gleason
Clidemia piistulata DC.
Clidemia pycnaster Tutin
Clidemia pycnaster subsp. pycnaster
Clidemia pycnaster subsp. robusta Wurdack
Clidemia rubra (Aubl.) Mart.
Clidemia sandwithii Wurdack
Clidemia saiilensis Wurdack
Clidemia septuplinervia Cogn.
Clidemia sericea D. Don
Clidemia sessiliflora (Naudin) Cogn.
Clidemia siapensis Wurdack
Clidemia silvicola Gleason
Clidemia simpsonii Wurdack
Clidemia stellipilis (Gleason) Wurdack
Clidemia strigillosa (Sw.) DC.
Clidemia tepuiensis Wurdack
Clidemia tococoidea (DC.) Gleason
Clidemia trinitensis (Crueg.) Griseb.
Clidemia ulei Pilg.
Clidemia urceolata DC.
Clidemia venosa (Gleason) Wurdack
Comolia ayangannae Wurdack
BO
BO
BO
BO
BO
BO
BO
BO
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
AM BO
AM
AM
BO
AM
BO
AM BO
AM BO
AM
BO
AM
AM BO
DA
DA
DA
DA
DA
DA
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU?
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU?
GU
GU
GU
GU
GU
GU
GU
GU
SU FG
SU FG
SU FG
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
SU
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
SU
FG
FG
AM BO
AM BO
BO
AM BO
AM
AM
AM BO
BO
AM BO
AM
AM
AM BO
AM BO
BO
BO
AM
AM
AM
AM BO
AM
AM BO
AM
AM BO
AM BO
BO
AM BO
AM
AM BO
BO
AM
AM
AM
AM
AM
AM
AM BO
AM
AM BO
AM BO
BO
BO
GU
GU su
GU su FG
GU su FG
su FG
GU
su FG
GU su? FG
su FG
su
GU su FG
su FG
FG
Checklist of the Plants of the Guiana Shield 399
Comolia coriacea Gleason BO
Comolia latifolia (Aubl.) Cogn. [dubious species] FG
Comolia leptophylla (Bonpl.) Naudin
Comolia microphylla Benih. GU SU FG
Comolia montana Gleason
Comolia mimmulahoides (Bonpl.) Naudin
Comolia prostrata Wurdack
Comolia serpyllacea Wurdack
Comolia smithii Wurdack
Comolia vernicosa (Benth.) Triana
Comolia villosa (Aubl.) Triana
Comoliopsis neblinae Wurdack [Sierra de la Neblina]
Conostegia superba Naudin [Sierra de la Neblina]
Desmoscelis villosa (Aubl.) Naudin
Ernestia blackii Brade & Markgr.
Ernestia cataractae Tutin
Ernestia confertiflora Wurdack
Ernestia cordifolia O. Berg ex Triana AM
Ernestia glandiilosa Gleason
Ernestia granvillei Wurdack
Ernestia maguirei Wurdack
Ernestia minor Gleason
Ernestia pullei Gleason
Ernestia rubra Pulle
Ernestia subglabra Wurdack
Ernestia tenella (Bonpl.) DC.
Ernestia tenella var. sprucei Cogn.
Ernestia tenella var. tenella
Grajfenrieda caiyophyllea Triana GU
Graffenrieda caudata Wurdack GU
Grajfenrieda cinnoides Gleason
Grajfenrieda fantastica R.E. Schult. & L.B. Sm.
Graffenriedafruticosa Wurdack
Grajfenrieda hitchcockii Gleason
Grajfenrieda intei-media Triana GU
Graffenrieda invinii Wurdack GU
Graffenriedajauana Wurdack
Graffenrieda kralii Wurdack
Graffenrieda lanceolata Gleason
Graffenrieda miconioides Naudin
Graffenrieda obliqua Triana GU
Graffenrieda patens Triana
Graffenrieda pedunculata Gleason
Graffenrieda polymera Gleason
Graffenrieda polymera subsp. neblinensis Wurdack
Graffenrieda polymera subsp. polymera
Grajfenrieda reticulata Wurdack
Graffenrieda rotundifolia (Bonpl.) DC.
Grajfenrieda rufa Wurdack
Graffenrieda rupestris Ducke
Graffenrieda sessilifolia Triana GU
Graffenrieda sessilifolia subsp. A
Graffenrieda sessilifolia subsp. cardonae Wurdack
Grajfenrieda sessilifolia subsp. occidentalis Wurdack AM
Graffenrieda sessilifolia subsp. sessilifolia BO
Graffenrieda sipapoana Wurdack AM
Graffenrieda steyermarkii Wurdack BO
Graffenrieda tricalcarata Gleason AM
Grajfenrieda versicolor Gleason AM
Graffenrieda weddellii Naudin AM BO GU
Henriettea granulata O. Berg ex Triana AM BO GU
Henriettea horridula Pilg. AM
Henriettea maroniensis Sagot BO GU SU FG
Henriettea martiusii (DC.) Naudin AM
400 Checklist of the Plants of the Guiana Shield
Henriettea mucronata (Gleason) Renner
Henhettea multiflora Naudin
Henriettea patrisiana DC.
Henriettea ramiflora (Sw.) DC.
Henriettea spriiceana Cogn.
Henriettea stellaris O. Berg ex Triana
Henriettea succosa (Aubl.) DC.
Henriettella caudata Gleason
Henriettella diickeana Hoehne
Henriettellaflavescens (Aubl.) Triana
Henriettella heteroneura Gleason
Henriettella ininiensis Wurdack
Henriettella ovata Cogn.
Henriettella prancei Wurdack
Henriettella steyermarkii Wurdack
Leandra agrestis (Aubl.) Raddi
Leandra aristigera (Naudin) Cogn.
Leandra candelabrum (J.F. Macbr.) Wurdack
Leandra chaetodon (DC.) Cogn.
Leandra clidemioides (Naudin) Wurdack
Leandra cremersii Wurdack
Leandra cuneata (Mart.) Cogn.
Leandra divaricata (Naudin) Cogn.
Leandra edentida Gleason
Leandrafrancavillana Cogn.
Leandra glandulifera (Triana) Cogn.
Leandra gorzulae Wurdack
Leandra intermedia (DC.) Cogn.
Leandra lindeniana (Naudin) Cogn.
Leandra longisepala Wurdack
Leandra magiiirei Wurdack [Sierra de la Neblina]
Leandra micrantha Wurdack
Leandra micropetala (Naudin) Cogn.
Leandra neblinensis Wurdack
Leandra paleacea Wurdack
Leandra phelpsiae Gleason
Leandra polyadena Ule
Leandra procumbens Ule
Leandra purpurea Gleason
Leandra rufescens (DC.) Cogn.
Leandra sanguinea Gleason
Leandra sanguinea subsp. sanguinea
Leandra sanguinea subsp. tepuiensis Wurdack
Leandra secunda (D. Don) Cogn.
Leandra solenifera Cogn.
Leandra steyermarkii Wurdack
Leandra verticillata (DC.) Cogn.
Loreya arborescens (Aubl.) DC.
Loreya mespiloides Miq.
Loreya ovata O. Berg ex Triana
Loreya spruceana Benth. ex Triana
Loreya strigosa Gleason
Loreya subrotundifolia (Wurdack) S.S. Renner
Macairea axilliflora Wurdack
Macairea cardonae Wurdack
Macairea chimantensis Wurdack
Macairea duidae Gleason
Macairea lanata Gleason
Macairea lasiophylla (Benth.) Wurdack
Macairea linearis Gleason
Macairea maroana Wurdack
Macairea multinervia Benth.
Macairea neblinae Wurdack
Macairea pachyphylla Benth.
AM
BO DA GU su FG
BO GU su FG
BO GU su FG
BO
BO GU su FG
BO GU su FG
BO GU su FG
BO FG
su FG
AM
AM BO
AM BO
AM BO
AM BO
BO
BO
BO
AM BO
WA
BO
BO
BO
AM
AM
AM
AM BO
BO
BO
BO
AM BO
BO
AM BO
AM
AM BO
AM
AM
AM BO
AM
AM BO
AM
AM
BO
BO
AM
AM
AM BO
AM
AM
BO
AM
AM BO
DA
DA
GU
GU
GU
GU
GU
GU
GU
GU?
GU
GU
GU
GU
GU
GU SU
FG
FG
FG
FG
SU FG
FG
GU SU FG
su FG
GU
GU
GU FG
GU su FG
GU su
GU
GU su
GU su FG
FG
GU su? FG
GU su FG
FG
Checklist of the Plants of the Guiana Shield 401
Macairea parvifolia Benth.
Macairea rigida Benth.
Macairea rufescens DC.
Macairea spruceana O. Berg ex Triana
Macairea stylosa Triana
Macairea thyrsiflora DC.
Macrocentrum anfractum Wurdack
Macrocentrum angustifolium Gleason
Macrocentrum anychioides Gleason
Macrocentrum brevipedicellatum Wurdack
Macrocentrum chimantense Wurdack
Macrocentrum cristatum (DC.) Triana
Macrocentrum cristatum var. cristatum
Macrocentrum cristatum var. microphyllum Cogn.
Macrocentrum cristatum war. parviflorum (DC.) Cogn.
Macrocentrum droseroides Triana
Macrocentrum fasciculatum (Rich, ex DC.) Triana
Macrocentrum fruticosum Gleason
Macrocentrum gesneriaceum Sandwith
Macrocentrum gracile Wurdack
Macrocentrum huberi Wurdack
Macrocentrum latifolium Wurdack
Macrocentrum longidens (Gleason) Wurdack
Macrocentrum maguirei Wurdack
Macrocentrum minus Gleason
Macrocentrum neblinae Wurdack
Macrocentrum parvulum Gleason
Macrocentrum repens (Gleason) Wurdack
Macrocentrum rubescens Gleason
Macrocentrum steyermarkii Wurdack
Macrocentrum stipulaceum Wurdack
Macrocentrum vestitum Sandwith
Maguireanthus ayangannae Wurdack
Maieta guianensis Aubl.
Maieta neblinensis Wurdack
Maieta poeppigii Mart, ex Cogn.
Mallophyton chimantense Wurdack
Marcetia taxifolia (A. St.-Hil.) DC.
Meriania broccha Wurdack
Meriania crassiramis (Naudin) Wurdack
Meriania neblinensis Wurdack
Meriania ornata Wurdack
Meriania sclerophylla (Naudin) Triana
Meriania urceolata Triana
Miconia abbreviata Markgr.
Miconia abysmophila Wurdack
Miconia acinodendron (L.) Sweet
Miconia acuminata (Steud.) Naudin
Miconia acutifolia Ule
Miconia aeruginosa Naudin
Miconia affinis DC.
Miconia aguitensis Gleason
Miconia alata (Aubl.) DC.
Miconia albicans (Sw.) Triana
Miconia alborufescens Naudin
Miconia aliquantula Wurdack
Miconia alternans Naudin
Miconia amacurensis Wurdack
Miconia ampIa Triana
Miconia aplostachya (Bonpl.) DC.
Miconia argyrophylla DC.
Miconia argyrophylla subsp. argyrophylla
Miconia argyrophylla subsp. gracilis Wurdack
Miconia aristata Gleason AM
BO GU
AM
AM
AM
AM
AM
AAA
BO GU
GU
AJVl
AM
BO
BO
GU
BO GU SU FG
GU SU FG
GU SU FG
BO GU SU FG
AM BO GU
GU SU FG
GU SU FG
GU
GU
BO
FG
AM
AM
BO GU
AM
BO
GU
GU
SU
AM
BO
GU
GU
GU
FG
AM BO GU SU FG
AM
AM BO
BO
GU SU FG
AM BO GU
AM
BO GU
AM
AM
BO GU
AM BO GU
AM BO GU
AM
AM GU SU
SU
FG
FG
AM BO GU
GU?
AM BO DA GU SU FG
AM BO
AM BO GU SU FG
AM BO GU SU FG
AM BO GU
SU FG
AM DA GU SU FG
BO DA GU FG
AM BO GU SU FG
AM BO DA GU SU FG
AM BO GU SU FG
GU SU FG
BO GU SU FG
402 Checklist of the Plants of the Guiana Shield
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon,
Micon,
Micon,
Micon,
Micon
Micon
Micon
Micon
Micon
Micon
Micon
Micon,
Micon,
Micon,
Micon
AM BO
AM
AM BO
AM
AM BO
AM BO
AM BO
AM
BO
AM BO
AM
AM BO
AM BO
AM BO
GU
GU su FG
GU su
GU
su FG
GU
GU su
GU
GU su? FG
GU su FG
FG
GU su FG
FG
GU su FG
GU su FG
a biglandulosa Gleason
a biglomerata (Bonpl.) DC.
a borjensis Wurdack DA
a brachybotrya Triana
a bracteata {DC.) JndiWdi DA
a brevipes Benth.
a bubalina (D. Don) Naudin
a buntingii Wurdack
a cacatin (Aubl.) S.S. Renner
a cacumina Wurdack
a campestris (Benth.) Triana
a carassana Cogn.
a cautis Wurdack
a centrodesma Naudin
a ceramicarpa (DC.) Cogn. DA
a ceramicarpa var. candolleana Cogn.
a ceramicarpa var. ceramicarpa AM BO DA
a ceramicarpa var. crozieriae Cremers & C.V. Martin
a chysophyllaiKxch.) \]rh. AM BO DA
a ciliata (Rich.) DC. AM BO
a cowanii Wurdack [perhaps expected in FG]
a crassinervia Cogn. Cogn. AM BO GU
a curta (Gleason) Wurdack AM BO
a curta subsp. curta AM BO
a curta subsp. ptariensis Wurdack BO
a decurrens Cogn. AM BO
a diaphanea Gleason
a dioica Wurdack BO
a dispar Benth. AM BO
a dodecandra Cogn. AM BO
a dolichorrhyncha Naudin AM BO
a egensis Cogn. AM BO
a elaeoides Naudin AM
a elata (Sw.) DC. AM BO
a eriodonta DC.
a eugenioides Triana AM BO
afaUaxDC. AM BO DA
afanshawei Wurdack
afragilis Naudin BO DA
afragrans Cogn. AM BO
afrancavillana Cogn.
a gratissima Benth. ex Triana AM BO
a grossidentata Wurdack AM BO
a guaiquinimae Wurdack BO
a holosericea (L.) DC. AM BO
a huberi Wurdack BO
a hypoleuca (Benth.) Triana BO
a ibaguensis (Bonpl.) Triana AM BO
a iluensis Wurdack BO
a impetiolaris (Sw.) D. Don ex DC. AM BO
a impetiolaris var. spruceana Cogn. AM BO
a inaequalifolia Triana AM
a kappleri Naudin SU FG
a kavanayensis Wurdack BO
a laevigata (L.) DC. BO
a lappacea (DC.) Triana GU SU FG
a lasseri Q\Q2iSon AM BO GU SU
a lateriflora Cogn. BO DA GU SU FG
a lateriflora snUp. lateriflora BO DA GU SU FG
a lateriflora subsp. monticellensis Wurdack BO GU SU
a lepidota DC. AM BO GU SU FG
a longifolia (Auhl) DC. AM BO DA GU SU FG
a longispicata Triana AM GU FG
a loreyoides Triana GU
GU SU FG
FG
GU su
GU
GU su FG
FG
FG
GU SU
GU
GU FG
GU
GU SU? FG
GU su FG
GU su FG
GU su FG
GU su
GU
GU
Checklist of the Plants of the Guiana Shield 403
Miconia lourteigiana Wurdack
Miconia macrothyrsa Benth.
Miconia macrotis Cogn.
Miconia maguirei Gleason
Miconia marginata Triana
Miconia mariae Wurdack
Miconia maroana Wurdack
Miconia matthaei Naudin
Miconia melinonis Naudin
Miconia minutiflora (Bonpl.) DC.
Miconia mirabilis (Aubl.) L.O. Williams
Miconia mituana Wurdack
Miconia mutabilis (DC.) Triana
Miconia myriantha Benth.
Miconia navioensis Wurdack [expected in FG]
Miconia neblinensis Wurdack [expected in AM, Serra da Neblina]
Miconia nervosa (Sm.) Triana
Miconia oldemanii Wurdack
Miconia perobscura Wurdack [Sierra de la Neblina]
Miconia perturbata Wurdack
Miconia phaeophylla Triana
Miconia pileata DC.
Miconia pilgeriana Ule
Miconia platypoda Gleason
Miconia plukenetii Naudin
Miconia poeppigii Triana
Miconia polita Gleason
Miconia prasina (Sw.) DC.
Miconia pseudoaplostachya Cogn.
Miconia pseudocapsularis Wurdack
Miconia pubipetala Miq.
Miconia punctata (Desr.) D. Don ex DC.
Miconia pyrifolia Naudin
Miconia racemosa (Aubl.) DC.
Miconia radulaefolia (Benth.) Naudin
Miconia regelii Cogn.
Miconia rhytidophylla Naudin
Miconia roraimensis Ule [Mt. Roraima]
Miconia rubiginosa (Bonpl.) DC.
Miconia rufescens (Aubl.) DC.
Miconia ruficalyx Gleason
Miconia rugosa Triana
Miconia rupestris Ule
Miconia rupticalyx Wurdack
Miconia sagotiana Cogn.
Miconia sarmentosa Cogn.
Miconia sastrei Wurdack
Miconia serialis DC.
Miconia serrulata (DC.) Naudin
Miconia silicicola Gleason
Miconia solmsii Cogn.
Miconia splendens (Sw.) Griseb.
Miconia sprucei Triana
Miconia stenostachya DC.
Miconia stephananthera Ule
Miconia steyermarkii Gleason
Miconia strigosa (Triana) Wurdack
Miconia superba Ule
Miconia tetraspermoides Wurdack
Miconia theizans (Bonpl.) Cogn.
Miconia tillettii Wurdack
Miconia tinifolia Naudin
Miconia tomentosa (Rich.) D. Don ex DC.
Miconia traillii Cogn.
AM
AM
AM
BO
BO
BO
BO
BO
GU
GU
GU
GU
SU FG
AM BO GU SU FG
FG
AM BO GU SU FG
AM BO DA GU SU FG
AM
FG
AM BO DA GU
GU?
SU FG
AM BO DA GU SU FG
FG
AM
AM BO
AM BO GU SU FG
AM BO FG
AM BO
GU SU
BO DA GU SU FG
AM BO
BO
GU?
GU
SU FG
AM BO DA GU SU FG
AM BO GU FG
AM BO GU
AM BO DA GU SU FG
AM BO GU SU FG
BO DA GU SU FG?
BO DA GU SU FG
AM GU
GU FG?
AM GU? SU?
AM BO GU?
AM BO GU SU
AM BO GU SU FG
AM BO DA GU SU FG
AM BO GU SU
AM BO GU
AM GU
GU SU
SU
FG
FG
FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM BO GU
AM BO
AM BO DA GU SU FG
AM
AM BO GU SU
AM BO GU SU FG?
AM
AM
BO GU
BO GU SU FG
AM BO GU SU FG
AM BO GU SU FG
AM BO GU
AM BO GU SU FG
AM GU SU FG
AM BO
AM
AM BO
AM
AM
AM BO
AM
BO
AM BO
BO
AM
AM BO
AM
AM
GU SU FG
su FG
404 Checklist of the Plants of the Guiana Shield
Miconia trimera Wurdack SU FG
Miconia trinervia (Sw.) D. Don ex Loudon FG
Miconia truncata Triana
Miconia tschudyoides Cogn. SU FG
Miconia imdata Triana
Miconia wittii Ule
Miconia yatuensis Wurdack
Microlicia benthamiana Triana ex Cogn. GU
Microlicia giianayana Wurdack
Monochaetum bonplandii (Kunth) Naudin
Mouriri acutiflora Naudin
Mouriri anguUcosta Morley
Mouriri angiistifolia Spruce ex Triana
Mouriri brevipes Hook. GU SU
Mouriri cauliflora Mart, ex DC.
Mouriri coUocarpa Ducke
Mouriri crassifolia Sagot
Mouriri densifoliata Ducke
Mouriri duckeana Morley
Mouriri dumetosa Cogn.
Mouririficoides Morley
Mouririfrancavillana Cogn.
Mouriri grandiflora DC.
Mouriri guianensis Aubl.
Mouriri huberi Cogn.
Mouriri latihila Morley
Mouriri longifolia (Kunth) Morley
Mouriri myrtifolia Spruce ex Triana [expected in GU]
Mouriri myrtilloides (Sw.) Poir.
Mouriri myrtilloides subsp. orinocensis Morley
Mouriri nervosa Pilg.
Mouriri nigra (DC.) Morley
Mouriri oligantha Pilg.
Mouriri pauciflora Spruce ex Cogn.
Mouriri rhizophorifolia (DC.) Triana
Mouriri sagotiana Triana
Mouriri sideroxylon Sagot ex Triana
Mouriri subumbellata Triana
Mouriri uncitheca Morley & Wurdack
Mouriri vernicosa Naudin
Mouriri viridicosta Morley
Myriaspora egensis DC.
Neblinanthera cumbrensis Wurdack
Nepsera aquatica (Aubl.) Naudin
Ochthephilus repentinus Wurdack
Opisthocentra clidemioides Hook. f. AM
Ossaea coarctiflora Wurdack FG
Ossaea mavacana Wurdack
Ossaea micrantha (Sw.) Macfad.
Ossaea quadrisulca (Naudin) Wurdack
Pachyloma coriaceum DC.
Pachyloma huberioides (Naudin) Triana
Pachyloma pusillum Wurdack
Pachyloma setosum Wurdack
Phainantha laxiflora (Triana) Gleason
Phainantha maguirei Wurdack
Phainantha myrteoloides Wurdack
Phainantha steyermarkii Wurdack
Poteranthera pusilla Bong.
Pterogastra divaricata (Bonpl.) Naudin
Pterogastra minor Naudin
Pterolepis glomerata (Rottb.) Miq.
Pterolepis trichotoma (Rottb.) Cogn.
Rhynchanthera dichotoma (Desr.) DC.
GU SU FG
GU? SU FG
AM
GU SU FG
FG
AM
FG
AM GU SU FG
AM BO DA GU SU FG
BO DA GU SU FG
AM
AM
AM BO
AM BO
AM BO
BO SU FG
AM BO GU SU FG
FG
AM
BO
FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM FG
AM
AM GU SU FG
FG
AM BO GU SU FG
AM
BO DA GU
GU
SU FG
AM
AM
GU
FG
AM
AM
AM
BO
BO
BO
BO
GU
GU
GU
AM BO
AM BO GU
AM BO
BO GU SU FG
AM BO GU SU FG
DA GU FG
GU?
GU SU
GU SU FG
GU SU FG
GU SU FG
Checklist of the Plants of the Guiana Shield 405
Rhynchanthera grandiflora (Auhl) DC. AM BO GU SU FG
Rhynchanthera hispida Naudin GU? SU
Rhynchantheraserrulata (Rich.) DC. BO GU SU FG
Salpingo glandulosa (Gleason) Wurdack [expected in GU]
Salpingo maguirei Gleason
Salpingo paleacea Wurdack GU
Salpingo pusilla (Gleason) Wurdack
Salpingo secundo Schrank & Mart, ex DC.
Sandemonia hoehnei (Cogn.) Wurdack [Sierra de la Neblina]
Siphonthera cordifolia (Benth.) Gleason AM BO GU
Siphonthero cowonii Wurdack
Siphonthera dawsonii Wurdack
Siphonthera duidae (Gleason) Wurdack
Siphontherafasciculoto (Gleason) Almeda & O.R. Rob.
Siphontherafoliosa (Naudin) Wurdack
Siphonthera hostmannii Cogn.
Toteanthus duidae Gleason
Tibouchino aspero Aubl.
Tibouchino aspero var. aspero
Tibouchino aspero var. asperrima Cogn.
Tibouchino bipenicilloto (Naudin) Cogn.
Tibouchino cothorinoe Pittier
Tibouchino cryptodeno Gleason
Tibouchino dissitifloro Wurdack
Tibouchino duidae Gleason
Tibouchinofroterno N.E. Br. AM BO GU
Tibouchinofroterno suhsp. froterno AM BO GU
Tibouchinofroterna subsp. poruano Wurdack
Tibouchino geitneriana (Schltdl.) Cogn. AM BO GU
Tibouchino gracilis (Bonpl.) Cogn. BO GU
Tibouchino huberi Wurdack
Tibouchino kunhordtii Gleason
Tibouchino llonorum Wurdack
Tibouchino longifolio (Vahl) Baill. BO GU
Tibouchino multifloro (Gardner) Cogn. FG
Tibouchino sipapoona Gleason
Tibouchino spruceono Cogn. [expected in BO]
Tibouchino steyermarkii Wurdack
Tibouchino striphnocalyx (DC.) Gleason
TococooristatoBtnih. BO GU SU FG?
Tococo bolivarensis Gleason AM BO GU
Tococo bolivarensis subsp. bolivarensis AM BO GU
Tococo bolivarensis subsp. occidentolis Wurdack
Tococo bullifero Triana
Tococo coryophyllea (DC.) S.S. Renner
Tococo cilioto Triana
Tococo cinnomomeo Triana
Tococo cordoto O. Berg ex Triana
Tococo coronoto Benth. AM BO GU
Tococo desiliens Gleason GU
Tococo erythrophyllo (Ule) Wurdack AM BO GU
Tococa guionensis Aubl. AM BO DA GU SU FG
Tococo hirto O. Berg ex Triana
Tococo loncifolio Spruce ex Triana
Tococo liesneri Wurdack
Tococa macrophysco Spruce ex Triana
Tococa mocrospermo Mart. AM BO GU
rococcrw/'/ewj' (Benth.) Triana AM BO GU
Tococa obovato Gleason
Tococo obovato subsp. neblinensis Wurdack
Tococa obovato subsp. obovato
Tococa oligontho Gleason
Tococo pachystochya Wurdack
Tococo pouciflora Spruce ex Triana
AM BO
AM
AM
AM
AM
AM
BO
AM
AM
BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
AM
AM BO
AM
AM BO
AM
AM BO
AM
BO
AM
AM
BO
AM
AM BO
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM BO
AM
AM BO
AM BO
406 Checklist of the Plants of the Guiana Shield
Tococa wtundifolia (Triana) Wurdack
Tococa subciliata (DC.) Triana
Tococa tepuiensis Wurdack
Tococa tepuiensis subsp. glabrata Wurdack
Tococa tepuiensis subsp. tepuiensis
Topobeaferruginea Gleason [perhaps expected in GU]
Topobea parasitica Aubl.
Tryssophyton merumense Wurdack
Votomita guianensis Aubl.
Votomita aff. monadelpha (Ducke) Morley
Votomita ohnocensis Morley
Votomita ventuarensis Morley
AM
AM GU
AM BO
AM
BO
AM BO
GU
su
GU su
su
FG
FG
AM
AM
Synonyms and Misidentifications
Acanthella montana Gleason =Acanthella sprucei
Acanthella plicata Gleason =AcantheUa pjilchra
Aciotis acutiflora (Mart.) Triana =Aciotis indecora
Aciotis acutiflora var. parvifolia Cogn. =Aciotis indecora
Aciotis aequatorialis Cogn. =Aciotis acuminifolia
Aciotis alata (Beurl.) Almeda =Aciotis rubricaulis
Aciotis amazonica Cogn. =Aciotis acuminifolia
Aciotis amazonica var. radicans Cogn. =Aciotis acuminifolia
Aciotis anomala Brade =Aciotis indecora
Aciotis aristata Ule =Aciotis rubricaulis
Aciotis asplundii Wurdack =Aciotis rubricaulis
Aciotis caulialata (Ruiz & Pav.) Triana =Aciotis rubricaulis
Aciotis circaeifolia var. major Cogn. =Aciotis purpurascens
Aciotis cordata J.F. Macbr. [not in flora area]
Aciotis dichotoma (Benth.) Cogn. =Aciotis acuminifolia
Aciotis dichotoma var. anomala (Miq.) Cogn. =Aciotis acuminifolia
Aciotis dichotoma var. longifolia S. Moore =Aciotis acuminifolia
Aciotis discolor D. Don =Aciotis purpurascens
Aciotis dysophylla (Benth.) Triana =Aciotis annua
Aciotis fragilis (Rich, ex DC.) Cogn. =Aciotis purpurascens
Aciotis fragilis var. lancifolia (Steud. ex Cogn.) Cogn. =Aciotis purpurascens
Aciotis herbacea Steud. ex Cogn. =Aciotis ornata
Aciotis indecora var. glabrescens Cogn. =Aciotis viscida
Aciotis indecora var. macrophylla Cogn. =Aciotis indecora
Aciotis indecora var. sagotiana Cogn. =Aciotis indecora
Aciotis laxa (DC.) Cogn. =Aciotis indecora
Aciotis laxa var. A =Aciotis viscida
Aciotis laxa var. B =Aciotis indecora
Aciotis laxa var. kappleriana (Naudin) Cogn. =Aciotis indecora
Aciotis laxa var. laxa =Aciotis indecora
Aciotis laxa var. robusta Cogn. =Aciotis indecora
Aciotis levyana Cogn. =Aciotis rubricaulis
Aciotis longifolia (DC.) Triana =Aciotis circaeifolia
Aciotis longifolia var. glabra Huber ^Aciotis purpurascens
Aciotis martinicensis (Naudin) Urb. =Aciotis purpurascens
Aciotis pellucida (Rich, ex DC.) Pittier et al. =Aciotis purpurascens
Aciotis purpurascens var. alata (Beurl.) Cogn. =Aciotis rubricaulis p.p.; Aciotis purpurascens p.p.
Aciotis purpurascens var. longifolia Cogn. =Aciotis purpurascens
Aciotis purpurascens var. martinensis J.F. Macbr. =Aciotis rubricaulis
Aciotis purpurascens var. pellucida (Rich, ex DC.) Cogn. =Aciotis purpurascens
Aciotis rostellata (Naudin) Triana =Aciotis indecora
Aciotis sieberi (Steud.) Triana =Aciotis purpurascens
Aciotis sileniflora (Bonpl.) Triana =Aciotis purpurascens
Aciotis sphaeranthera (Naudin) Cogn. =Aciotis acuminifolia
Aciotis trinitensis Cogn. =Aciotis purpurascens
Aciotis uliginosa (Bonpl. ex Naudin) Triana =Aciotis purpurascens
Aciotis viscosa (Bonpl. ex Naudin) Triana =Aciotis indecora
Acisanthera adscendens C. Wright =Acisanthera quadrata
Acisanthera beccabunga (Mart, ex Naudin) Triana =Acisanthera bivalvis
Acisanthera boissieriana Cogn. =Acisanthera hedyotidea
Checklist of the Plants of the Guiana Shield 407
Acisanthera brevifolia (DC.) Griseb. =Acisanthera bivalvis
Acisanthera erecta Gleason =Macairea lasiophylla
Acisanthera gracilis Ule =Acisanthera crassipes
Acisanthera inundata (DC.) Triana [not in flora area]
Acisanthera inundata var. pusilla Cogn. =Acisanthera limnobios
Acisanthera lasiophylla (Benth.) Gleason =Macairea lasiophylla
Acisanthera pellucida Wright ex Griseb. =Acisanthera limnobios
Acisanthera quadrata Juss. ex Poir. =Acisanthera quadrata
Acisanthera recurva (Rich.) Griseb. =Acisanthera uniflora
Acisanthera salzmannii (Naudin) Cogn. =Acisanthera hedyotidea
Acisanthera trivalvis (Aubl.) Cogn. =Acisanthera bivalvis
Adelobotrys guianensis (DC.) Gleason =Adelobotrys adscendens
Adelobotrys laxiflora Triana =Phainantha laxiflora
Adelobotrys monticola subsp. occidentalis Wurdack =Adelobotrys monticola
Apatitia blakeoides Desv. =Bellucia grossularioides
Arthrostema mimmularioides (Bonpl.) DC. =Comolia nummularioides
Arthrostemma angusturenese (Rich.) Cogn. ^Pterolepis glomerata
Arthrostemma caulialatum Ruiz & Pav. =Aciotis rubricaulis
Arthrostemma glomerata (Rottb.) Cham. =Pterolepis glomerata
Arthrostemma glomeratum (Rottb.) Cham. =Pterolepis glomerata
Arthrostemma ladanoides (Rich.) DC. =Pterolepis trichotoma
Arthrostemma latifolium D. Don [dubious species, type not found]
Arthrostemma villosum (Aubl.) DC. =Comolia villosa
Axinanthera macrophylla H. Karst. =Bellucia pentamera
Bellucia arborescens (Aubl.) Baill. =Loreya arborescens
Belinda aricuaizensium Pittier =Bellucia pentamera
Bellucia aubletii Naudin =BeUucia grossularioides
Bellucia brasiliensis Naudin =Bellucia grossularioides
Bellucia cacatin (Aubl.) Sagot =Miconia cacatin
Bellucia circumscissa Spruce ex Cogn. =BeUucia grossularioides
Bellucia hostmannii Naudin =Bellucia grossularioides
Bellucia mespiloides (Miq.) J.F. Macbr. =Loreya mespiloides
Bellucia nervosa Raf. =Bellucia grossularioides
Bellucia quinquenervia (Aubl.) H. Karst. =Bellucia grossularioides
Bellucia subrotundifolia Wurdack =Loreya subrotundifolia
Blakea cacatin (Aubl.) D. Don ex DC. =Miconia cacatin
Blakea calycanthus Markgr. =Blakea rosea
Blakea caudata Triana =Blakea rosea
Blakea quinquenervia Aubl. =Bellucia grossularioides
Blakea triplinervia L. f. =Bellucia grossularioides
Brachypremna petiolata Gleason =Ernestia pullei
Calophysa tococoidea DC. =Clidemia tococoidea
Capitellaria benthamii Naudin =Clidemia capitata
Centronia crassiramis (Naudin) Triana =Meriania crassiramis
Chaenophoraferruginea Crueg. =Miconia hypoleuca
Chaenopleuraferruginea (Crueg.) Griseb. =Miconia hypoleuca
Chaenopleurafulva (Rich.) Griseb. =Miconia chrysophylla
Chaenopleura hypoleuca Benth. =Miconia hypoleuca
Chaenopleura longifolia (Aubl.) Griseb. =Miconia longifolia
Chaetogastra callichaeta Benth. =Pterolepis glomerata
Chaetogastra geitneriana Schltdl. =Tibouchina geitneriana
Chaetogastra glomerata (Rottb.) Benth. =Pterolepis glomerata
Chaetogastra hypericoides DC. =Desmoscelis villosa
Chaetogastra lasiophylla Benth. =Macairea lasiophylla
Chaetogastra rhynchanthera Benth. =Monochaetum bonplandii
Chaetolepis citrina Gleason =Chaetolepis anisandra
Charianthus spp. [Caribbean taxon, none in flora area]
Chitonia bubalina D. Don =Miconia bubalina
Chitonia fothergilla (Desr.) D. Don =Miconia mirabilis
Clidemia affinis (Naudin) Cogn. =Clidemia pustulafa
Clidemia agrestis (Aubl.) D. Don =Leandra agrestis
Clidemia aristigera Naudin =Leandra aristigera
Clidemia benthamiana Miq. =Clidemia hirta var hirta
Clidemia berbiceana DC. =Miconia acinodendron
408 Checklist of the Plants of the Guiana Shield
Clidemia biformis Gleason =Clidemia anisophylla
Clidemia bonplandii Cogn. =Clidemia debilis
Clidemia bracteata DC. =Micoma bracteata
Clidemia campestris Benth. =Miconia campesths
Clidemia campestris var. pauciflora Benth. =Miconia biglomerata
Clidemia candolleana Cogn. [nom. illeg.] =Clidemia bullosa
Clidemia ceramicarpa DC. =Miconia ceramicarpa
Clidemia chaetodon (DC.) Naudin =Leandra chaetodon
Clidemia coccinea DC. =Miconia ceramicarpa var. ceramicarpa
Clidemia coriacea (Naudin) Cogn. =Clidemia tepuiensis
Clidemia depaiiperata DC. =Leandra agrestis
Clidemia dependens D. Don =Clidemia capitellata var. dependens
Clidemia desmantha Benth. =Miconia rhytidophylla
Clidemia divaricata Naudin =Leandra divaricata
Clidemia drosera Sagot ? =Clidemia micrantha
Clidemia elegans (Aubl.) D. Don =Clidemia hirta var. elegans
Clidemia glabrata Steud. =Miconia ibaguensis
Clidemia heterobasis DC. =Clidemiajapurensis var. heterobasis
Clidemia heteroclita Naudin =Clidemiajapurensis wax. japurensis
Clidemia lappacea DC. =Miconia lappacea
Clidemia lutescens var. lindeniana Naudin =Leandra lindeniana
Clidemia maculata Benth. =Miconia ibaguensis
Clidemia manoacensis Kraenzl. =Clidemia bullosa
Clidemia martii Naudin =Miconia centrodesma
Clidemia miconioides Benth. =Miconia ibaguensis
Clidemia mutabilis DC. =Miconia mutabilis
Clidemia naevula (Naudin) Triana =Clidemiajapurensis var. heterobasis
Clidemia neglecta D. Don - Clidemia capitellata (Bonpl.) D.Don
Clidemia novemnervia var. affinis (Naudin) Wurdack =Clidemia pustulata
Clidemia novemnervia var. depauperata Pittier =Clidemia bullosa
Clidemia petiolata var. angustifolia Cogn. =Clidemia petiolata
Clidemia petiolata var. colorata Cogn. =Clidemia petiolata
Clidemia radulaefolia Benth. =Miconia radulaefolia
Clidemia rariflora Benth. =Clidemia bullosa
Clidemia rhamnifolia var. macrodon Naudin =Leandra divaricata
Clidemia rubra var. biacuta (Naudin) Cogn. =Clidemia sericea
Clidemia secunda D. Don =Leandra secunda
Clidemia smithii Gleason ? =Clidemia laevifolia x Clidemia hirta
Clidemia spicata (Aubl.) DC. =Clidemia capitellata var. dependens
Clidemia surinamensis Miq. =Clidemia novemnervia
Clidemia tiliifolia DC. =Clidemia hirta var. tiliifolia
Clidemia trichodes DC. =Miconia bracteata
Clidemia tristis (Naudin) Sagot =Miconia bracteata
Clidemia umbonata DC. =Clidemia strigillosa
Clidemia violacea DC. =Miconia ceramicarpa var. ceramicarpa
Clidemia virgata Pittier =Miconia ibaguensis
Comolia affinis Hoehne =Comolia villosa
Comolia angustifolia Gleason =Comolia villosa
Comolia anomala Pittier =Acisanthera uniflora
Comolia hirtella Naudin =Comolia villosa
Comolia hoehnei Cogn. =Sandemania hoehnei
Comolia kuhlmannii Hoehne =Comolia villosa
Comolia lythrarioides Naudin =Comolia villosa
Comolia neglecta Cogn. =Comolia villosa
Comolia purpurea Miq. =Comolia villosa
Comolia surinamensis Miq. =Comolia villosa
Comolia tetraptera Cogn. =Acisanthera tetraptera
Comolia veronicaefolia Benth. =Comolia villosa
Conostegia acuminata Steud. =Miconia acuminata
Conostegia icosandra (Sw.) Urb. [not in flora area]
Conostegia petiolata Gleason [not in flora area] =Conostegia montana (Sw.) DC.
Copedesma nitens Gleason =Miconia ruficalyx
Coryphadenis abnormis (Naudin) Morley =Votomita guianensis
Cremanium trinitatis Crueg. =Miconia minutiflora
Checklist of the Plants of the Guiana Shield 409
Davya adscendens (Sw.) Griseb. =Adelobotrys adscendens
Davya ciliata Naudin =^Adelobotrys ciliata
Davya crassiramis Naudin =Mehania crassiramis
Davya guianemis DC. =Adelobotrys adscendens
Davya scandens (Aubl.) Naudin =Adelobotrys scandens
Davya sclewphylla Naudin ^Meriania sclerophylla
Decaraphefloribunda Miq. =Micoma acinodendron
Decaraphefockeana Miq. =Miconia acinodendron
Decaraphe hostmannii Steud. =Miconia serrulata
Desmoscelis mollis Pittier =Desmoscelis villosa
Dicrananthera hedyotidea C. Presl =Acisanthera hedyotidea
Dicrananthera salzmannii Naudin =Acisanthera hedyotidea
Diolena longidens Gleason =Macrocentrum longidens
Diolena repens Gleason '=Macrocentrum repens
Diplochita bubalina (D. Don) DC. =Miconia bubalina
Diplochitaflorida DC. =Miconia mirabilis
Diplochita fothergilla (Desr.) DC. =Miconia mirabilis
Diplochita leucocephala DC. =Miconia serrulata
Diplochita mucronata (Desr.) DC. =Miconia holosericea
Diplochita parviflora Benth. =Miconia pubipetala
Diplochita serrulata DC. =Miconia serrulata
Diplochita serrulata var. latifolia DC. =Miconia serrulata
Diplochita tomentosa (Rich.) Griseb. =Miconia tomentosa
Ernestia lata Gleason =Ernestia tenella var. sprucei
Emychaenia punctata (Desr.) Griseb. =Miconia punctata
Farringtonia fasciculata Gleason =Siphantherafasciculata
Fothergilla guianensis Aubl. =Miconia mirabilis
Fothergilla mirabilis Aubl. =Miconia mirabilis
Glossocentrum collinum Crueg. =Miconia minutiflora
Grajfenrieda candelabrum J.F. Macbr. =Leandra candelabrum
Grajfenrieda curta Gleason =Miconia curta
Graffenriedafloribunda Triana =Graffenrieda miconioides
Grajfenrieda hitchcockii var. par-vifolia Wurdack =Graffenrieda hitchcockii
Graffenrieda ovalifolia Naudin =Graffenrieda weddellii
Graffenrieda stenopetala Ule =Graffenrieda intermedia
Guildingia psidioides Hook. =Mouriri rhizophorifolia
Hartigia barbigera (DC.) Miq. =Miconia ciliata
Hartigia oblongifolia Miq. =Miconia racemosa
Hartigia spectabilis Miq. =Miconia racemosa
Henriettea brunnescens Standi. =Loreya mespiloides
Henriettea oblongifolia Naudin =Henriettea martiusii
Henriettea orinocensis Naudin =Henriettea martiusii
Henriettea succosa var. guianensis Gleason =Henriettea ramiflora
Henriettea succosa var. kappleriana Steud. ex Naudin ^Henriettea succosa
Henriettea surinamensis Miq. =Henriettea ramiflora
Henriettea trinervia Naudin =Henriettea ramiflora
Henriettella longistyla Ule =Henriettella ovata
Henriettella micrantha Gleason =Henriettella ovata
Henriettella patrisiana (DC.) Naudin =Henriettea patrisiana
Henriettella ramiflora (Sw.) Naudin =Henriettea ramiflora
Henriettella venosa Gleason =Clidemia venosa
Heterotrichum novemnervium DC. =Clidemia novemnervia
Heterotrichum strigosum Triana =Miconia strigosa
Hormocalyx hirsutus Gleason =Tococa macrosperma
Jucunda tomentosa (Rich.) Benth. =Miconia tomentosa
Lasiandra bipenicillata Naudin =Tibouchina bipenicillata
Lasiandra tibouchina (Desr.) Naudin =Tibouchina aspera var. aspera
Leandra anisophylla (DC.) Cogn. =Clidemia anisophylla
Leandra chimantensis Wurdack =Leandra procumbens
Leandra cuspidata (Triana) Cogn. =Miconia eugenioides
Leandra fendleri Cogn. =Leandra lindeniana
Leandra hylophila Gleason =Leandra clidemioides
Leandra linearis Gleason =Clidemia linearis
Leandra longicoma Cogn. [not in flora area]
410 Checklist of the Plants of the Guiana Shield
Leandra montana Gleason =Aciotis indecora
Leandra pulverulenta (DC.) Cogn. =Micoma aliquantula
Leandra rhamnifolia var. macrodon (Naudin) Cogn. =Leandra divaricata
Leandra rufescens var. asperiuscula (DC.) Cogn. =Leandra rufescens
Leandra rufescens (DC.) Cogn. var. glandulosa Cogn. =Leandra rufescens
Leandra scabra DC. [not in flora area] =Leandra melastomoides Raddi
Leandra stellipilis Gleason =Clidemia stellipilis
Leiostegia vernicosa Benth. =Comolia vernicosa
Loevigia sericea H. Karst. & Triana =Monochaetum bonplandii
Loreya acutifolia O. Berg ex Triana =Loreya arborescens
Loreya brunnescens (Standi.) Gleason =Loreya mespiloides
Loreya coUatata Wurdack ^Loreya spruceana
Loreyaflavescens (Aubl.) Naudin =Henriettellaflavescens
Loreya huberi Wurdack =BeUucia huberi
Loreya maguirei Wurdack =Loreya arborescens
Loreya minor Cogn. =Loreya ovata
Loreya mucronata Gleason =Henriettea mucronata
Loreya quadrifolia Gleason =Loreya strigosa
Macairea albiflora Cogn. =Macairea thyrsiflora
Macairea aspera N.E. Br. =Macairea thyrsiflora
Macairea calvescens Naudin =Macairea pachyphylla
Macairea lanata subsp. eglandulosa Wurdack =Macairea lanata
Macairea maguirei Wurdack =Macairea thyrsiflora
Macairea schultesii Wurdack =Macairea thyrsiflora
Macrocentrum glandulosum Gleason =Salpinga glandulosa
Macrocentrum montanum Gleason =Macrocentrum cristatum var. microphyllum
Macrocentrum pusilium Gleason =Salpinga pusilla
Maieta dispar Miq. =Maieta guianensis
Maieta glandulifera Standi. =Tococa guianensis
Maietajuruensis Pilg. =Clidemiajuruensis
Marcetia andicola Naudin =Marcetia taxifolia
Marcetia cordigera var. andicola (Naudin) Cogn. =Marcetia taxifolia
Marcetiajuniperina (Humb.) DC. =Marcetia taxifolia
Meisneria cordifolia Benth. =Siphanthera cordifolia
Meisneria microlicioides Naudin =Siphanthera cordifolia
Melastoma acinodendron L. =Miconia acinodendron
Melastoma adscendens Sw. =Adelobotrys adscendens
Melastoma agrestis Aubl. =Leandra agrestis
Melastoma alatum Aubl. =Miconia alata
Melastoma albicans Sw. =Miconia albicans
Melastoma aplostachyum Bonpl. =Miconia aplostachya
Melastoma aquaticum Aubl. =Nepsera aquatica
Melastoma arborescens Aubl. =Loreya arborescens
Melastoma aromaticum Vahl =Tibouchina aspera var. aspera
Melastoma biglomeratum Bonpl. =Miconia biglomerata
Melastoma bivalve Aubl. =Acisanthera bivalvis
Melastoma bivalvis Aubl. =Acisanthera bivalvis
Melastoma brunnea Vahl =Henriettea succosa
Melastoma cacatin Aubl. =Miconia cacatin
Melastoma capitellatum Bonpl. ^Clidemia capitellata
Melastoma chrysophyllum Rich. =Miconia chrysophylla
Melastoma ciliatum Rich. =Miconia ciliata
Melastoma dichotomum Desr. =Rhynchanthera dichotoma
Melastoma diffusum Desr. =Acisanthera uniflora
Melastoma dodecandrum Desr. =Miconia dodecandra
Melastoma elatum Sw. =Miconia elata
Melastoma elegans Aubl. =Clidemia hirta var. elegans
Melastomaflavescens Aubl. =Henriettellaflavescens
Melastomafothergillum Desr. =Miconia mirabilis
Melastomafulvum Rich. =Miconia chrysophylla
Melastoma grandiflorum Aubl. =Rhynchanthera grandiflora
Melastoma grossularioides L. =Bellucia grossularioides
Melastoma heteroneurum DC. =Clidemia heteroneura
Melastoma hirtum L. =Clidemia hirta
Checklist of the Plants of the Guiana Shield 411
Melastoma holosericea L. =Miconia holosericea
Melastoma ibaguense Bonpl. =Miconia ibaguensis
Melastoma impetiolaris Sw. =Miconia impetiolaris
Melastoma laevigatum L. =Miconia laevigata
Melastoma laevigatum Aubl. [nom. illeg., non M. laevigata L.]
Melastoma longifoliiim Aubl. =Miconia longifolia
Melastoma maietiim Desr. =Maieta guianensis
Melastoma micranthum Sw. =Ossaea micrantha
Melastoma minutiflonim Bonpl. =Miconia minutiflora
Melastoma mucronatum Desr. =Micoma holosericea
Melastoma nervosum Sm. =Miconia nervosa
Melastoma octonum Bonpl. ^CUdemia octona
Melastoma parviflorum Aubl. =Miconia prasina
Melastoma physophorum Vahl =Tococa guianensis
Melastoma pleurocarpum E. Mey. =Micoma acinodendron
Melastoma prasinum Sw. =Miconia prasina
Melastoma punctatum Desr. =Miconia punctata
Melastoma purpurascens Aubl. =Aciotis purpurascens
Melastoma purpurea (Aubl.) Willd. =Aciotis purpurascens
Melastoma racemosum Aubl. =Miconia racemosa
Melastoma ramiflorum Sw. =Henriettea ramiflora
Melastoma rubiginosum Bonpl. =Miconia rubiginosa
Melastoma rubnim Aubl. =Clidemia rubra
Melastoma rufescens Aubl. =Miconia riifescens
Melastoma scandens Aubl. =Adelobotrys scandens
Melastoma scorpioides Schltdl. & Cham. =Miconia trinervia
Melastoma spicatum Aubl. =Clidemia capitellata var. dependens
Melastoma splendens Sw. =Miconia splendens
Melastoma strigillosum Sw. =Clidemia strigillosa
Melastoma succosum Aubl. =Henriettea succosa
Melastoma tamonea Sw. =Miconia mirabilis
Melastoma theizans Bonpl. =Miconia theizans
Melastoma tibouchinum Desr. =Tibouchina aspera var. aspera
Melastoma tococo Desr. =Tococa guianensis
Melastoma tomentosum Rich. =Miconia tomentosa
Melastoma trinervium Sw. =Miconia trinervia
Melastoma trivalvis Aubl. =Acisanthera bivalvis
Melastoma villosum Aubl. =Desmoscelis villosa
Meliandra monadelpha (Ducke) Morley =Votomita monadelpha
Meriania duidae Gleason =Adelobotrys duidae
Merianiaferruginea Suess. =Adelobotrys barbata
Meriania paraensis Ducke =Meriania urceolata
Miconia agrestis (Aubl.) Baill. =Leandra agrestis
Miconia amplexans (Crueg.) Cogn. =Miconia tomentosa
Miconia anceps Naudin ^Miconia trinervia
Miconia argyrophylla var. attenuata Cogn. =Miconia argyrophylla subsp. argyrophylla
Miconia arirambae Huber =Miconia alborufescens
Miconia attenuata DC. =Miconia prasina
Miconia attenuata var. subquintuplino-via DC. =Miconia prasina
Miconia axilliflora Naudin =Miconia acuminata
Miconia barbigera DC. =Miconia ciliata
Miconia berbiceana (DC.) Naudin =Miconia acinodendron
Miconia bifrons Naudin =Miconia lepidota
Miconia bracteolaris Naudin =Miconia eriodonta
Miconia bracteolaris var. cardiophora Naudin ^Miconia eriodonta
Miconia buchtienii Cogn. =Miconia centrodesma
Miconia cachimbensis Brade =Miconia alborufescens
Miconia catyophyllaea (DC.) Triana =Tococa caryophyllea
Miconia caudiculata Pittier =Miconia trinervia
Miconia centrandra Ule =Miconia centrodesma
Miconia ceramicarpa var. navioensis Wurdack =Miconia ceramicarpa var. candolleana
Miconia ceramicarpa var. violacea (DC.) Cogn. =Miconia ceramicarpa var. ceramicarpa
Miconia chamissonis Naudin [not in flora area]
Miconia clidemioides Steud. =Miconia acinodendron
412 Checklist of the Plants of the Guiana Shield
Miconia compacta Gleason =Micoma carassana
Miconia congesta Cogn. =Miconia poeppigii
Miconia cristulata Naudin =Miconia prasina
Miconia cruegeriana Naudin =Miconia hypoleuca
Miconia demerarensis Gleason =Micoma bracteata
Miconia disparilis (Standi.) R.O. Williams =Miconia lateriflora subsp. lateriflora
Miconia eriodonta var. oblongifolia DC. =Miconia eriodonta
Miconia erioneura Cogn. =Tococa guianensis
Miconia eriophora Miq. =Miconia ibaguensis
Miconia eriopoda Benth. =Miconia acinodendron
Miconia erythrophylla Ule =Tococa erythrophylla
Miconia erythropila Steud. =Miconia ceramicarpa var. ceramicarpa
Miconiafendleriana Cogn. =Miconia phaeophylla
Miconiaflorida (DC.) Naudin =Miconia mirabilis
Miconiafockeana Miq. =Miconia nervosa
Miconiafothergilla (Desr.) Naudin =Mic&nia mirabilis
Miconiafulva (Rich.) DC. ^Miconia chrysophylla
Miconiafulva var. angustifolia Cogn. =Miconia chrysophylla
Miconiafulva var. aubletiana (Naudin) Cogn. =Miconia chrysophylla
Miconia globidiflora (Rich.) Cham, [not in flora area]
Miconia glossocentra Naudin =Miconia minutiflora
Miconia giiaiquinimae subsp. angustifolia Wurdack =Miconia guaiquinimae
Miconia guianensis (Aubl.) Cogn. =Miconia mirabilis
Miconia heterochroa Miq. =Miconia albicans
Miconia holosericea var. mucronata (Desr.) Cogn. ^Miconia holosericea
Miconia holosericea var. subquintuplinervia Cogn. =Miconia holosericea
Miconia hypargyrea Miq. =Miconia stenostachya
Miconia ibaguensis var. glabrata (Steud.) Cogn. =Miconia ibaguensis
Miconia impetiginosa DC. =Miconia rufescens
Miconia involucrata Donn. Sm. =Miconia ampla
Miconia iodopila Steud. =Miconia ceramicarpa var. ceramicarpa
Miconia killipi Gleason =Miconia loreyoides
Miconia lambertiana DC. =Miconia longifolia
Miconia leschenaultiana (Naudin) Sagot =Miconia ceramicarpa var. ceramicarpa
Miconia leucocephala (DC.) Naudin =Miconia serrulata
Miconia longifolia var. aubletiana Naudin =Miconia chrysophylla
Miconia longistyla Steud. =Miconia argyrophylla subsp. argyrophylla
Miconia macrophylla Steud. =Miconia prasina
Miconia macrophylla var. hostmannii (Steud.) Cogn. =Miconia serrulata
Miconia macrophylla var. latifolia (DC.) Cogn. =Miconia serrulata
Miconia macrophylla var. leucocephala (DC.) Cogn. =Miconia serrulata
Miconia macrophylla var. serrulata (DC.) Cogn. =Miconia serrulata
Miconia macrostachya DC. =Miconia aplostachya
Miconia megaphylla Gleason =Miconia ampla
Miconia micrantha Pilg. =Miconia wittii
Miconia microcarpa DC. =Miconia affinis
Miconia miqueliana Naudin =Miconia acinodendron
Miconia mucronata (Desr.) Naudin =Miconia holosericea
Miconia mucronulata Ule =Miconia prasina
Miconia multinervulosa Cogn. =Miconia theizans
Miconia naudiniana Miq. =Miconia lepidota
Miconia nemoralis Naudin =Miconia prasina
Miconia nitens Benth. -Tococa nitens
Miconia obovalis Naudin =Miconia splendens
Miconia pachypoda Gleason =Miconia superba
Miconia pallida Gleason =Miconia elaeoides
Miconia palmetorum Pittier =Miconia stephananthera
Miconia panicularis Gleason =Miconia egensis
Miconia parviflora (Benth.) Cogn. =Miconia pubipetala
Miconia planinervia Naudin =Miconia affinis
Miconia plebeia Naudin =Miconia brevipes
Miconia plumosa Gleason =Miconia rugosa
Miconia prasina var. attenuata (DC.) Cogn. =Miconia prasina
Miconia pseudorubiginosa Cogn. =Miconia macrothyrsa
Checklist of the Plants of the Guiana Shield 413
Miconia ptewphora Miq. =Miconia alata
Miconia pteropoda Benth. =Miconia prasina
Miconia purpurascens (Aubl.) DC. =Aciotis purpurascens
Miconia racemosa var. barbigera (DC.) Cogn. =Miconia ciliata
Miconia racemosa var. ciliata (Rich.) Griseb. =Miconia ciliata
Miconia revoluta Benth. =Miconia prasina
Miconia robusta (Sagot) Cogn. =Miconia tschudyoides
Miconia rubiginosa var. kuhlmannli Hoehne =Miconia brevipes
Miconia rubiginosa var. platyura Naudin =Micoma rubiginosa
Miconia rufescens var. grandifolia Cogn. =Miconia rufescens
Miconia schomburgkii Benth. =Miconia lepidota
Miconia scorpioides (Schltdl. & Cham.) Naudin =Miconia trinervia
Miconia scrobiculata Cogn. =Miconia radulaefolia
Miconia semota Markgr. =Miconia carassana
Miconia septuplinervis Pittier =Miconia nervosa
Miconia setimarginata Pittier =Tococa oligantha
Miconia spondylantha DC. =Miconia nervosa
Miconia spondylantha var. leschenaultiana Naudin =Miconia ceramicarpa var. ceramicarpa
Miconia subciliata DC. =Tococa subciliata
Miconia subtriloba Gleason =Miconia centrodesma
Miconia surinamensis Gleason =Miconia poeppigii
Miconia sylvestris Gleason =Miconia centrodesma
Miconia tessmannii Markgr. =Miconia lateriflora subsp. lateriflora
Miconia tinifolia var. roraimensis Wurdack =Miconia tinifolia
Miconia tomentella Cogn. =Miconia serialis
Miconia trichodes (DC.) Triana =Miconia bracteata
Miconia trinitatis (Crueg.) Naudin =Miconia minutiflora
Miconia verticilliflora Steud. ex Naudin =Miconia pubipetala
Miconia virgulata Gleason =Miconia ciliata
Microlicia bivalvis (Aubl.) DC. =Acisanthera bivalvis
Microlicia brevifolia DC. =Acisanthera bivalvis
Microlicia bryanthoides Oliv. =Microlicia benthamiana
Microlicia limnobios DC. =Acisanthera limnobios
Microlicia recurva (Rich.) DC. =Acisanthera uniflora
Microlicia steyermarkii Gleason =Microlicia benthamiana
Microphysca rotundifolia Triana =Tococa rotundifolia
Miocarpus paludosus Naudin =Acisanthera limnobios
Monochaetum sericeum Naudin ex Linden =Monochaetum bonplandii
Mouriri abnormis Naudin =Votomita guianensis
Mouriri acutiflora var. amblyodon Naudin =Mouriri acutiflora
Mouriri acutiflora var. oligantha Gleason =Mouriri collocarpa
Mouriri anomala Pulle =Mouriri crassifolia
Mouriri pendulifolia Cogn. =Mouriri acutiflora
Mouriri plasschaerti Pulle =Mouriri nigra
Mouriri plasschaertii Pulle =Mouriri nigra
Mouriri polyantha Miq. =Mouriri guianensis
Mouriri princeps Naudin =Mouriri grandiflora
Mouriri ulei Pilg. =Mouriri guianensis
Mouriri weddellii Naudin =Mouriri guianensis
Myriaspora decipiens Naudin =Myriaspora egensis
Myriaspora surinamensis Steud. '=Myriaspora egensis
Myrmidone lanceolata Cogn. =Tococa macrosperma
Myrmidone macrosperma (Mart.) Mart. =Tococa macrosperma
Noterophila beccabunga Mart, ex Naudin =Acisanthera bivalvis
Noterophila brevifolia (DC.) Naudin =Acisanthera bivalvis
Noterophila limnobios (DC.) Mart. =Acisanthera limnobios
Noterophila pusilla Naudin =Acisanthera limnobios
Octopleura ciliata Triana =Miconia lateriflora subsp. lateriflora
Octopleura micrantha (Sw.) Griseb. =Ossaea micrantha
Olisbea rhizophorifolia DC. ^'Mouriri rhizophorifolia
Onoctonia calcarata Naudin =Acisanthera crassipes
Onoctonia crassipes Naudin =Acisanthera crassipes
Onoctonia pauciflora Naudin =Acisanthera crassipes
Osbeckia aubletiana Spreng. =Rhynchanthera grandiflora
4 1
4
Checklist of the Plants of the Guiana Shield
Osbeckia glomerata (Rottb.) DC. =Pterolepis glomerata
Osbeckia striphnocalyx DC. =Tibouchina striphnocalyx
Ossaea ciliata (Triana) Cogn. =Miconia lateriflora subsp. lateriflora
Ossaea disparilis Standi. =Miconia lateriflora subsp. lateriflora
Ossaea duckeana Hoehne =Clidemia epibaterium
Ossaea epibaterium (DC.) Triana =Clidemia epibaterium
Ossaeaflavescens (Aubl.) DC. =Henriettellaflavescens
Ossaea trichopoda Gleason =Leandra chaetodon
Ossaea trinitensis Crueg. =Clidemia trinitensis
Oxymeris agrestis (Aubl.) Triana =Leandra agrestis
Oxymeris anisophylla (DC.) Triana =Clidemia anisophylla
Oxymeris aristigera (Naudin) Triana =Leandra aristigera
Oxymeris asperiuscula (DC.) Triana =Leandra rufescens
Oxymeris chaetodon (DC.) Triana =Leandra chaetodon
Oxymeris cuspidata Triana ^Miconia eugenioides
Oxymeris glandulifera Triana =Leandra ghndulifera
Oxymeris lindeniana (Naudin) Triana =Leandra lindeniana
Oxymeris pulverulenta (DC.) Triana =Miconia aliquantula
Oxymeris rufescens (DC.) Triana =Leandra rufescens
Oxymeris seciinda (D. Don) Triana =Leandra secunda
Pachyloma nanum Wurdack =Pachyloma pusillum
Pachyloma scandens Ducke =Pachyloma huberioides
Petaloma mouriri Sw. =Mouriri guianensis
Petaloma myrtilloides Sw. =Mouriri myrtilloides
Phyllopus martiusii DC. =Henriettea martiusii
Platycentrum clidemioides Naudin =Leandra clidemioides
Pleroma bipenicillata (Naudin) Triana =Tibouchina bipenicillata
Pleroma tibouchinum (Desr.) Triana =Tibouchina aspera var. aspera
Pogonorhynchus amplexans Crueg. =Miconia tomentosa
Poteranthera calcarata (Naudin) Triana =Acisanthera crassipes
Poteranthera crassipes (Naudin) Triana =Acisanthera crassipes
Poteranthera diiidae Gleason ^Siphanthera duidae
Poterantherafoliosa (Naudin) Cogn. =Siphantherafoliosa
Poteranthera minor Gleason =Siphantherafoliosa
Poteranthera pauciflora (Naudin) Triana =Acisanthera crassipes
Pterogastra glabra Gleason =Pterogastra divaricata
Pterogastra major Triana =Pterogastra divaricata
Pterolepis hispida (Rich.) Gleason =Pterolepis trichotoma
Pterolepis hostmannii Steud. =Pterolepis glomerata
Pterolepis ladanoides (Rich.) Triana =Pterolepis trichotoma
Pterolepis lasiophylla (Benth.) Triana =Macairea lasiophylla
Pterolepis pumila (Bonpl.) Cogn. =Pterolepis trichotoma
Pterolepis pumila YSir. procera Cogn. =Pterolepis trichotoma
Pterolepis pusilla Miq. =Pterolepis trichotoma
Pterolepis striphnocalyx (DC.) Cogn. =Tibouchina striphnocalyx
Rhexia acisanthera L. =Acisanthera quadrata
Rhexia angusturensis Rich. =Pterolepis glomerata
Rhexia aquatica (Aubl.) Sw. =Nepsera aquatica
Rhexia aspera (Aubl.) Willd. =Tibouchina aspera var. aspera
Rhexia bicornis Rottb. =Acisanthera uniflora
Rhexia bivalvis (Aubl.) Vahl =Acisanthera bivalvis
Rhexia bonplandii Kunth =Monochaetum bonplandii
Rhexia canescens Bonpl. =Monochaetum bonplandii
Rhexia divaricata Bonpl. =Pterogastra divaricata
Rhexia glomerata Rottb. =Pterolepis glomerata
Rhexia gracilis Bonpl. =Tibouchina gracilis
Rhexia hispida Rich. =Pterolepis trichotoma
Rhexia hypericoides Willd. =Desmoscelis villosa
Rhexia incana Spreng. =Monochaetum bonplandii
Rhexia indecora Bonpl. =Aciotis indecora
Rhexiajuniperina Humb. =Marcetia taxifolia
Rhexia ladanoides Rich. =Pterolepis trichotoma
Rhexia latifolia Aubl. ^Comolia latifolia
Rhexia leptophylla Bonpl. =Comolia leptophylla
Checklist of the Plants of the Guiana Shield 415
Rhexia longifolia Vahl =Tibouchina longifolia
Rhexia mimmularioides Bonpl. =Comolia nummularioides
Rhexia polystachya Bonpl. =Aciotis polystachya
Rhexia piimila Bonpl. =Pterolepis trichotoma
Rhexia recurva Rich. =Acisanthera iiniflora
Rhexia rotundifolia Bonpl. =Grajfenrieda rotundifolia
Rhexia serrulata Rich. =Rhynchanthera serrulata
Rhexia sileniflora Bonpl. =Aciotis purpurascens
Rhexia strigosa Willd. ex Sprang. =Pterolepis trichotoma
Rhexia taxifolia A. St.-Hil. =Marcetia taxifolia
Rhexia tenella Bonpl. =Ernestia tenella
Rhexia thymifolia Bonpl. =Appendicidaria thymifolia
Rhexia trichotoma Rottb. ^Pterolepis trichotoma
Rhexia trivalvis (Aubl.) Vahl =Acisanthera bivalvis
Rhexia uniflora Vahl =Acisanthera uniflora
Rhexia villosa Aubl. =Comolia villosa
Rhexia villosissima Rich. =Desmoscelis villosa
Rhynchanthera acuminata Benth. =Rhynchanthera grandiflora
Rhynchanthera acuminata var. sublaevis Cogn. =Rhynchanthera grandiflora
Rhynchanthera adenophora Miq. =Acisanthera uniflora
Rhynchanthera ambigua Naudin =Rhynchanthera grandiflora
Rhynchanthera betulifolia Cogn. =Rhynchanthera grandiflora
Rhynchanthera cardonae Wurdack =Rhynchanthera grandiflora
Rhynchanthera grandiflora var. microphylla Naudin =Rhynchanthera grandiflora
Rhynchanthera intermedia Ule =Rhynchanthera grandiflora
Rhynchanthera latifolia Cogn. [not in flora area]
Rhynchanthera microphylla Gleason =Rhynchanthera grandiflora
Rhynchanthera modesta Naudin =Rhynchanthera serrulata
Rhynchanthera orinocensis Sprague =Rhynchanthera grandiflora
Rhynchanthera parviflora Naudin =Rhynchanthera serrulata
Rhynchanthera virgata Wurdack =Rhynchanthera hispida
Sagraea aphanantha (Naudin) Triana =Clidemia aphanantha
Sagraea capillipes Triana =Clidemia capillipes
Sagraea capitata (Benth.) Triana =Clidemia capitata
Sagraea cognata Steud. =Clidemia rubra
Sagraea coriacea (Naudin) Triana ^Clidemia tepuiensis
Sagraea minutiflora Triana =Clidemia minutiflora
Sagraea neurocarpa Naudin =Ossaea micrantha
Sagraea petiolata (DC.) Triana =Clidemia petiolata
Sagraea rubra (Aubl.) Triana =Clidemia rubra
Salpinga cristata DC. =Macrocentrum cristatum
Salpingafasciculata DC. =Macrocentrum fasciculatum
Salpinga parviflora DC. =Macrocentrum cristatum var. parviflorum
Sandemania glandulosa Wurdack =Sandemania hoehnei
Sarmentaria decora Naudin =Adelobotrys adscendens
Siphanthera capitata Gleason =Siphanthera hostmannii
Siphanthera cordifolia var. glomerata Gleason =Siphanthera cordifolia
Siphantherajenmanii Gleason =Siphanthera cordifolia
Siphanthera microlicioides (Naudin) Cogn. =Siphanthera cordifolia
Siphanthera paraensis Huber =Siphanthera hostmannii
Siphanthera tatei Gleason =Siphanthera hostmannii
Siphantheropsis williamii Brade =Macairea lasiophylla
Spennera acuminifolia Mart, ex DC. =Aciotis acuminifolia
Spennera acutiflora Mart. =Aciotis indecora
Spennera alata Beurl. =Aciotis rubricaulis
Spennera annua Mart, ex DC. =Aciotis annua
Spennera anomala Miq. =Aciotis acuminifolia
Spennera aquatica (Aubl.) Mart, ex DC. =Nepsera aquatica
Spennera caulialata (Ruiz & Pav.) Naudin =Aciotis rubricaulis
Spennera chaetodon DC. ^Leandra chaetodon
Spennera dichotoma Benth. =Aciotis acuminifolia
Spennera dichotoma var. lanceolata Naudin =Aciotis acuminifolia
Spennera dysophylla Benth. =Aciotis annua
Spennerafragilis Rich, ex DC. =Aciotis purpurascens
416 Checklist of the Plants of the Guiana Shield
Spennera grandifolia Miq. =Aciotis purpurascens
Spennera hydwphila Miq. =Nepsera aqualica
Spennera indecora (Bonpl.) DC. =Aciotis indecora
Spennera kappleriana Naudin =Aciotis indecora
Spennera lancifolia Steud. ex Cogn. =Aciotis purpurascens
Spennera laxa DC. =Aciotis indecora
Spennera longifolia DC. =Aciotis circaeifolia
Spennera martinicensis Naudin =Aciotis purpurascens
Spennera ornata Miq. =Aciotis ornata
Spennera pellucida Rich, ex DC. =Aciotis purpurascens
Spennera polystachya (Bonpl.) Mart, ex DC. =Aciotis polystachya
Spennera quintuplinervia Miq. =Clidemiajapurensis var.japurensis
Spennera rostellata Naudin =Aciotis indecora
Spennera rubricaulis Schrank & Mart, ex DC. =Aciotis rubricaulis
Spennera sieberi Steud. =Aciotis purpurascens
Spennera sileniflora (Bonpl.) DC. =Aciotis purpurascens
Spennera sphaeranthera Naudin =Aciotis acuminifolia
Spennera tetraptera Miq. =Aciotis annua
Spennera uliginosa Bonpl. ex Naudin =Aciotis purpurascens
Spennera viscida Benth. =Aciotis viscida
Spennera viscosa Bonpl. ex Naudin =Aciotis indecora
Staphidiastrum aphananthum Naudin =Clidemia aphanantha
Staphidiastrum attenuatum Naudin =Clidemia attenuata
Staphidiastrum cognatum (Steud.) Naudin =Clidemia rubra
Staphidiastrum coriaceum Naudin =Clidemia tepuiensis
Staphidiastrum pulverulentum (DC.) Naudin =Miconia aliquantula
Staphidiastrum rubrum (Aubl.) Naudin =Clidemia rubra
Staphidiastrum rubrum var. biacutum Naudin =Clidemia sericea
Staphidium affine Naudin =Clidemia pustulata
Staphidium agreste (Aubl.) Naudin =Leandra agrestis
Staphidium benthamianum (Miq.) Naudin =Clidemia hirta var hirta
Staphidium bracteosum Naudin =Clidemia capitellata var. dependens
Staphidium conglomeratum (DC.) Naudin =Clidemia conglomerata
Staphidium elegans (Aubl.) Naudin =Clidemia hirta var elegans
Staphidium glabratum (Steud.) Naudin =Miconia ibaguensis
Staphidium hostmannii Naudin =Clidemia dentata
Staphidium invohicratum (DC.) Naudin =Clidemia involucrata
Staphidium lappaceum (DC.) Naudin =Miconia lappacea
Staphidium mutabile (DC.) Naudin =Miconia mutabilis
Staphidium petiolatum (DC.) Naudin =Clidemia petiolata
Staphidium sessiliflorum Naudin =Clidemia sessiliflora
Staphidium spicatum (Aubl.) Naudin =Clidemia capitellata var. dependens
Staphidium triste Naudin =Miconia bracteata
Tamonea guianensis Aubl. =Miconia mirabilis
Tetrameris trivalvis Naudin =Macairea multinervia
Tetrameris villosa (Aubl.) Naudin =Comolia villosa
Tetrazygia discolor (L.) DC. [not in flora area]
Tibouchina adscendens (C. Wright) M. Gomez =Acisanthera quadrata
Tibouchina brachyanthera Pittier =Tibouchina aspera var. asperrima
Tibouchina impressa Gleason =Tibouchina geitneriana
Tibouchina lasiophylla (Benth.) Cogn. =Macairea lasiophylla
Tibouchina multinervia (Benth.) Baill. =Macairea multinervia
Tibouchina pseudomollis Gleason =Tibouchina geitneriana
Tibouchina yavitensis Pittier =Tibouchina striphnocalyx
Tococa acuminata Benth. =Tococa guianensis
Tococa aubletii D. Don =Tococa guianensis
Tococa cardiophylla Naudin =Tococa guianensis
Tococa castrata Naudin =Tococa coronata
Tococa coriaceae S. Moore =Tococa guianensis
Tococa didymophysca Naudin =Tococa guianensis
Tococa discolor Pilg. =Tococa guianensis
Tococa egensis Naudin =Tococa coronata
Tococa erioneura (Cogn.) Wurdack =Tococa guianensis
Checklist of the Plants of the Guiana Shield 417
Tococa erythrostachya Spruce ex Triana =Tococa guianensis
Tococaferniginea G. Nicholson =Tococa guianensis
Tococa glandulosa Gleason =Tococa coronata
TococaJuruensis Pilg. =Tococa guianensis
Tococa lasiostyla Cogn. =Tococa subciliata
Tococa longisepala Cogn. =Tococa coronata
Tococa loretensis Ule =Tococa guianensis
Tococa maieta D. Don =Maieta guianensis
Tococa montana Gleason =Tococa obovata subsp. obovata
Tococa occidentalis Naudin =Tococa guianensis
Tococa planifolia Benth. =Tococa subciliata
Tococa rareimi Benth. ['Roraimae'] =Tococa guianensis
Tococa spruceana Cogn. =Tococa bullifera
Tococa subnuda Benth. =Tococa coronata
Tococa traillii Cogn. =Tococa coronata
Tococa ulei Pilg. =Tococa bullifera
Tococa wedellii Naudin =Tococa nitens
Topobea "guianensis Aubl." [invalid name from herbarium records, = Topobea parasitica]
Topobea cuspidata Gleason =Topobea parasitica
Tricentrum leptophyllum (Bonpl.) DC. =Comolia leptophylla
Truncaria caryophyllea DC. =Tococa caryophyllea
Tschudya asperiuscula DC. =Leandra rufescens
Tschudya berbiceana (DC.) Griseb. =Miconia acinodendron
Tschudya ibaguensis (Bonpl.) Griseb. =Miconia ibaguensis
Tschudya pulverulenta DC. =Miconia aliquantula
Tschudya robusta Sagot =Miconia tschudyoides
Tschudya rufescens DC. =Leandra rufescens
Tschudya spondylantha (DC.) Griseb. ^Miconia nervosa
Tschudya strigillosa Griseb. =Leandra clidemioides
Tschudya verticillata DC. =Leandra verticillata
Tulasnea foliosa Naudin =Siphantherafoliosa
Uranthera dicranophora Naudin =Acisanthera quadrata
Uranthera hostmannii Naudin =Acisanthera uniflora
Uranthera recurva (Rich.) Naudin =Acisanthera uniflora
Urodesmium huberioides Naudin =Pachyloma huberioides
Valdesia rosea Ruiz & Pav. =Blakea rosea
Webera quinquenervia (Aubl.) J.F. Gmel. =Bellucia grossularioides
References
(for a more extensive bibliography, see Wurdack et al. 1993)
Berry, P.E., A. Groger, B.K. Hoist, T. Morley, F.A. Michelangeli, N.G Luckana, F. Almeda, S.S. Renner, A. Freire-Fierro, O.R.
Robinson, & K. Yatskievych. 2001. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 6: 263-528.
Freire-Fierro, A. 2002. Monograph of Aciotis (Melastomataceae). Systematic Botany Monographs 62: 1-99.
Gleason, H.A. 1932. A synopsis of the Melastomataceae of British Guiana. Brittonia 1: 127-184.
Gleason, H.A. 1935. Melastomaceae. In A. Pulle, Flora of Suriname 3(1): 178-281.
Gorts-van Rijn, A.R.A. & M.J. Jansen-Jacobs. 1986. Melastomataceae (additions and corrections). In A.L. Stoffers & J.C.
Lindeman, Flora of Suriname 3(2): 356-429.
Morley, T. 1976. Memecyleae (Melastomataceae). Flora Neotropica Monogr. 15: 1-295.
Renner, S.S. 1989. Systematic studies in the Melastomataceae: Bellucia, Loreya, and Macairea. Mem. New York Bot. Gard. 50:
1-112.
Wurdack, J.J. 1961. Melastomataceae. In B. Maguire, J.J. Wurdack, & collaborators. Botany of the Guayana Highland part IV.
Mem. New York Bot. Gard. 10(4): 32-47.
Wurdack, J.J. 1 973. Melastomataceae. In T. Lasser, Flora de Venezuela 8: 1-819. (Mouriri & Votomita contributed by T. Morley)
Wurdack, J.J. & S. Renner. 2002. Melastomataceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff,
& J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York
Bot. Gard. 76 (2): 437-465.
Wurdack, J.J.,T. Morley, & S. Renner. 1993. Melastomataceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 13: 1-
424.
Meliaceae
K.S. Edwards, T.D. Pennington
Azadirachta indica A. Juss. [cultivated, naturalized]
418 Checklist of the Plants of the Guiana Shield
Cabralea canjerana (Veil.) Mart.
Cabralea canjerana subsp. canjerana
Carapa giiianensis Aubl.
Carapa procera DC.
Cedrela fissilis Veil.
Cedrela odorata L.
Guarea carinata Ducke
Guarea cinnamomea Harms
Guarea costata A. Juss.
Guarea glabra Vahl
Guarea gomma Pulle
Guarea grandifolia DC.
Guarea guidonia (L.) Sleumer
Guarea humaitensis T.D. Penn.
Guareajuglandiformis T.D. Penn.
Guarea kunthiana A. Juss.
Guarea macrophylla Vahl
Guarea macrophylla subsp. pachycarpa (C. DC.) T.D. Penn.
Guarea macrophylla suhsp. pendulispica (C. DC.) T.D. Penn.
Guarea michel-moddei T.D. Penn. & S.A. Mori
Guarea pubescens (Rich.) A. Juss.
Guarea pubescens subsp. pubescens
Guarea pubescens subsp. pubiflora (A. Juss.) T.D. Penn.
Guarea scabra A. Juss.
Guarea silvatica C. DC.
Guarea sprucei C. DC.
Guarea trunciflora C. DC.
Guarea velutina A. Juss.
Melia azedarach L. [cultivated, naturalized]
Swietenia macrophylla King
Trichilia acuminata (Humb. & Bonpl. ex Roem. & Schult.) A. DC. & C. DC
Trichilia areolata T.D. Penn.
Trichilia cipo (A. Juss.) C. DC.
Trichilia elegans A. Juss.
Trichilia elegans subsp. elegans
Trichilia euneura C. DC.
Trichilia gamopetala T.D. Penn.
Trichilia inaequilatera T.D. Penn.
Trichilia lecointei Ducke
Trichilia lepidota Mart.
Trichilia lepidota subsp. leucastera (Sandwith) T.D. Penn.
Trichilia martiana C. DC.
Trichilia mazanensis J.F. Macbr. [expected in the Guianas]
Trichilia micrantha Benth.
Trichilia micropetala T.D. Penn.
Trichilia moschata Sw.
Trichilia pallida Sw.
Trichilia pleeana (A. Juss.) C. DC.
Trichilia quadrijuga Kunth
Trichilia quadrijuga subsp. quadrijuga
Trichilia rubra C. DC.
Trichilia schomburgkii C. DC.
Trichilia schomburgkii subsp. javariensis T.D. Penn.
Trichilia schomburgkii subsp. schomburgkii
Trichilia septentrionalis C. DC.
Trichilia surinamensis (Miq.) C. DC.
Trichilia surumuensis C. DC.
Trichilia tetrapetala C. DC. BO
Synonyms and Misidentifications
Carapa surinamensis Miq. =Carapa procera
Cedrela brownii Loefl. ex Kuntze =Cedrela odorata
Cedrela guianensis A. Juss. =Cedrela odorata
Granatum surinamensis (Miq.) Kuntze =Carapa procera
Guarea adenocarpa Pittier =Trichilia pleeana
GU
GU
BO DA GU su FG
GU? su FG
BO DA
BO DA GU su
su
su
FG
FG
FG?
FG
BO DA GU su FG?
BO GU su FG
AM GU su FG
AM BO DA GU
GU
GU
su FG
AM GU su FG
AM BO GU su FG
AM BO su? FG
AM BO GU
FG
AM BO DA GU su FG
AM BO DA GU su FG
AM BO su FG?
GU su FG
AM GU su FG
AM
BO GU
GU
AM BO
BO
DA GU su
su
FG
FG
BO GU su? FG
BO GU
BO GU
GU? su FG
AM
AM BO
GU su
BO GU su
BO GU su
GU? su FG
AM BO
AM GU
GU
su
su
FG
FG
AM BO GU su FG
BO DA GU
AM BO DA GU su FG
AM BO DA GU su FG
AM DA GU
AM BO DA GU su FG
AM BO
AM BO DA GU su FG
AM BO GU su FG
AM GU
GU
su FG
Checklist of the Plants of the Guiana Shield 419
Guarea affinis A. Juss. =Guarea piibescens subsp. pubescens
Giiarea aubletii A. Juss. =Guarea giiidonia
Guarea concinna Sandwith =Guarea pubescens subsp. pubescens
Guarea davisii Sandwith =Guarea pubescens subsp. pubescens
Guarea delgadoi Pittier =Trichilia pleeana
Guarea guara (Jacq.) P. Wilson =Guarea guidonia
Guarea pallida C. DC. =Guarea pubescens
Guarea pedicellata C. DC. =Guarea silvatica
Guarea pendulispica C. DC. =Guarea macrophylla subsp. pendulispica
Guarea puberula Pittier =Guarea guidonia
Guarea pubiflora A. Juss. [not in flora area] =Guarea pubescens (Rich.) A.Juss. subsp. pubiflora
Guarea richardiana A. Juss. =Guarea pubescens subsp. pubescens
Guarea schomburgkii C. DC. =Guarea glabra
Guarea surinamensis Miq. ex C. DC. [nomen nudum] =Guarea guidonia
Guarea trichilioides L. =Guarea guidonia
Guarea trichilioides var. pachycarpa C. DC. =Guarea macrophylla subsp. pachycarpa
Moschoxylum cipo A. Juss. =Trichilia cipo
Moschoxylum pleeanum A. Juss. =Trichilia pleeana
Moschoxylum propinquum Miq. =Trichilia quadrijuga
Moschoxylum surinamense Miq. =Trichilia surinamensis
Odontandra acuminata Humb. & Bonpl. ex Roem. & Schult. =Trichilia acuminata
Odontocandra karstenii Triana & Planch. =Trichilia acuminata
Portesia echinocarpa de Vriese =Trichilia pallida
Surenus brownii (Loefl. ex Kuntze) Kuntze =Cedrela odorata
Trichilia acariaeantha Harms =Trichilia micrantha
Trichilia alternans C. DC. =Trichilia surinamensis
Trichilia brachystachya Klotzsch ex C. DC. =Trichilia pallida
Trichilia compacta A.C. Sm. =Trichilia quadrijuga subsp. quadrijuga
Trichilia cuneifolia Pulle =Trichilia schomburgkii
Trichilia davisii Sandwith =Trichilia pallida
Trichilia echinocarpa (de Vriese) Walp. =Trichilia pallida
Trichiliafuscescens Radlk. =Trichilia martiana
Trichilia grandis Lasser & Maguire =Trichilia septentrionalis
Trichilia guianensis Klotzsch ex C. DC. =Trichilia rubra
Trichilia karstenii (Triana & Planch.) A. DC. & C. DC. =Trichilia acuminata
Trichilia lanceolata Pittier [nom. illeg.] =Trichilia septentrionalis
Trichilia leucastera Sandwith =Trichilia lepidota subsp. leucastera
Trichilia maynasiana C. DC. =Trichilia surinamensis
Trichilia montana var. fendleriana C. DC. =Trichilia pallida
Trichilia moritzii C. DC. =Trichilia septentrionalis
Trichilia poeppigii C. DC. =Trichilia schomburgkii
Trichilia propinqua (Miq.) C. DC. =Trichilia quadrijuga subsp. quadrijuga
Trichilia pubescens Rich. =Guarea pubescens
Trichilia roraimana C. DC. =Trichilia micrantha
Trichilia smithii C. DC. =Trichilia pleeana
Trichilia stelligera Radlk. =Trichilia euneura
Trichilia subsessilifolia C. DC. =Trichilia schomburgkii
Trichilia subsimplex Steyerm. =Trichilia pallida
Trichilia trinitensis A. Juss. =Trichilia pallida
References
Pennington, T.D. 2002. Meliaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. HofT, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Card. 76 (2):
465-472.
Pennington, T.D., B.T. Styles, & D.A.H. Taylor. 1981. Meliaceae. Flora Neotropica Monogr. 28: 1-470.
Pennington, T.D. & K.S. Edwards. 2001 In J.A. Steyermark, PE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 6: 528-549.
Pennington, T.D. & A.R.A. Gorts-van Rijn. 1984. Meliaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 519-
569.
IVIendonciaceae
D. Wasshausen
Mendoncia aspera (Ruiz & Pav.) Nees GU SU FG
420 Checklist of the Plants of the Guiana Shield
BO GU SU FG
AM BO
BO
FG
GU FG
BO DA GU SU FG
AM
AM
AM BO
AM BO
GU SU FG
BO
AM
Mendoncia bivalvis (L. f.) Merr.
Mendoncia cardonae Leonard
Mendoncia coriacea Wassh.
Mendoncia crenata Lindau
Mendoncia glabra (Poepp. & Endl.) Nees
Mendoncia hoffmannseggiana Nees
Mendoncia neblinensis Wassh.
Mendoncia obovata Lindau
Mendoncia phalacra Leonard
Mendoncia spriicei Lindau
Mendoncia sqiiamuligera Nees
Mendoncia steyermarkii Wassh.
Mendoncia williamsii Wassh.
Synonyms and Misidentifications
Mendoncia hirsiita (Poepp. & Endl.) Nee§ =Mendoncia bivalvis
Mendoncia perrottetiana Nees ^Mendoncia bivalvis
Mendoncia pubescens Hoffsgg. ex Nees =Mendoncia hoffmannseggiana
Mendoncia schomburgkiana Nees ^Mendoncia hoffmannseggiana
Mendozia hirsuta Poepp. & Endl. =Mendoncia bivalvis
References
Bremekamp, C.E.B. 1938. Acanthaceae. In A. PuUe, Flora of Suriname 4(2): 166-252.
Wasshausen, D.C. 1989. New species of Mendoncia from the Venezuelan Guayana. Ann. Missouri Bot. Gard. 76: 1118-1124.
Wasshausen, D.C. 2001 . Mendonciaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 6: 550-554.
Wasshausen, D.C. 2002. Mendonciaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 472-474.
Menispermaceae
R.C. Bamebyt, R Hiepko
Abula barbata Miers
Abiita bullata Moldenke
Abuta candollei Triana & Planch.
Abuta grandifolia (Mart.) Sandwith
Abuta grisebachii Triana & Planch.
Abuta imene (Mart.) Eichler
Abuta obovata Diels
Abuta pahnii (Mart.) Krukoff& Bameby
Abuta rufescens Aubl.
Abuta sandwithiana Krukoff& Bameby
Abuta solimoesensis Krukoff& Bameby
Abuta velutina Gleason
Anomospermum chloranthum Diels
Anomospermum chloranthum subsp. chloranthum
Anomospermum chloranthum subsp. confusum Krukoff& Bameby
Anomospermum grandifolium Eichler
Anomospermum steyermarkii Krukoff& Bameby
Borismenejapurensis (Mart.) Bameby
Caryomenefoveolata Bameby & Krukoff
Caryomene glaucescens (Moldenke) Bameby & Krukoff
Caryomene olivascens Bameby & Krukoff
Cissampelos andromorpha DC.
Cissampelosfasciculata Benth.
Cissampelos ovalifolia DC.
Cissampelos pareira L.
Cissampelos tropaeolifolia DC.
Curarea candicans (Rich, ex DC.) Bameby & Krukoff
Curarea toxicofera (Wedd.) Bameby & Krukoff
Disciphania ernstii Eichler
Disciphania ernstii var. ernstii
Disciphania moriorum Bameby
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM
AM BO
AM BO
BO
BO
AM BO
AM
AM?
AM?
DA
DA
DA
GU SU FG
GU FG
GU SU FG
GU SU FG
FG?
GU SU FG
GU SU FG
GU SU FG
GU SU FG
FG
GU?
FG
FG
FG
GU
SU?
SU
FG
FG
FG
GU SU FG
GU SU FG?
GU SU
GU SU
GU SU FG
FG
SU FG
AM BO
BO
GU su FG
AM
AM BO DA GU SU FG
AM BO DA GU su FG
DA GU su FG
GU su FG
AM
FG
AM BO DA GU su FG
AM FG
AM GU su FG
FG
FG
GU su FG
AM
FG
AM GU
GU
su
su
FG
Checklist of the Plants of the Guiana Shield 421
Disciphania unilateralis Bameby
Elephantomene eburnea Bameby & Krukoff
Hyperbaena domingensis (DC.) Benth.
Odontocarya mallosperma Bameby
Odontocarya sp. A
Odontocarya tamoides (DC.) Miers
Odontocarya tamoides van canescens (Miers) Bameby
Odontocarya tamoides var. tamoides
Odontocarya wullschlaegelii (Eichler) Bameby
Orthomene hirsuta (Krukoff& Moldenke) Bameby & Krukoff
Orthomene prancei Bameby & Krukoff
Orthomene schomburgkii (Miers) Bameby & Krukoff
Orthomene verruculosa (Krukoff& Bameby) Bameby & Krukoff
Sciadotenia cayennensis Benth.
Sciadotenia duckei Moldenke
Sciadotenia eichleriana Moldenke
Sciadotenia sagotiana (Eichler) Diels
Sciadotenia sprucei Diels in Engl.
Telitoxicum duckei (Diels) Moldenke
Telitoxicum inopinatum Moldenke
Telitoxicum krukovii Moldenke
Telitoxicum minutiflorum (Diels) Moldenke GU? FG?
Synonyms and Misidentifications
Abuta amara Aubl. (Aristolochiaceae) =Aristolochia amara
Abuta candicans Rich, ex DC. =Curarea candicans
Abuta concolor Poepp. & Endl. =Abuta grandifolia
Abuta guyanensis Eichler =Abuta grandifolia
Abuta klugii Moldenke =Abuta imene
Abuta limaciifolia Diels =Curarea candicans
Abuta macrocarpa Moldenke =Anomospermum grandifolium
Abuta pullei Diels =Curarea candicans
Abuta splendida Krukoff & Moldenke =Abuta rufescens
Abuta tomentosa Sagot =Sciadotenia sagotiana
Abuta umbellata Sagot =Sciadotenia cayennensis
Abuta verruculosa Krukoff & Bameby =Orthomene verruculosa
Abuta wilson-brownei R.S. Cowan =Abuta rufescens
Anelasma domingense (DC.) Miers =Hyperbaena domingensis
Anomospermum hirsutum Krukoff & Moldenke =Orthomene hirsuta
Anomospermum japurense (Mart.) Eichler =BorismeneJapurensis
Anomospermum schomburgkii Miers =Orthomene schomburgkii
Chondrodendron candicans (Rich, ex DC.) Sandwith =Curarea candicans
Chondrodendron limaciifolium (Diels) Moldenke =Curarea candicans
Chondrodendron toxicoferum (Wedd.) Krukoff& Moldenke =Curarea toxicofera
Cissampelos ramiflora Miers =Cissampelos andromorpha
Cocculus domingensis DC. ^Hyperbaena domingensis
Cocculus grandifolius Mart. =Abuta grandifolia
Cocculus imene Mart. =Abuta imene
Cocculusjapurensis Mart. =Borismenejapurensis
Cocculus pahnii Mart. ^Abuta pahnii
Cocculus pauper Griseb. =Odontocarya tamoides var. canescens
Cocculus tamoides DC. =Odontocarya tamoides
Cocculus toxicoferus Wedd. =Curarea toxicofera
Hyperbaena cuatrecasasii Moldenke =Borismenejapurensis
Hyperbaena graciliflora Miers =Hyperbaena domingensis
Odontocarya hederifolia var. canescens Miers =Odontocarya tamoides var. canescens
Odontocarya paupera (Griseb.) Diels =Odontocarya tamoides var. canescens
Pachygone domingensis (DC.) Eichler =Hyperhaena domingensis
Pachygone graciliflora (Miers) Sagot =Hyperbaena domingensis
Sciadotenia candicans Diels =Curarea candicans
Somphoxylon wullschlaegelii Eichler =Odontocarya wullschlaegelii
Stephania elliptica DC. (Capparaceae) =Steriphoma ellipticum
References
Bameby, R.C. 1970. Revision of Neotropical Menispermaceae tribe Tinosporeae. Mem. New York Bot. Gard. 20(2): 81-158.
422 Checklist of the Plants of the Guiana Shield
Bameby, R.C. 2001. Menispermaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 6: 554-578.
Bameby, R.C. 2002. Menispermaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 474-483.
Bameby, R.C. & B.A. Krukoff. 1971. Supplementary notes in American Menispermaceae. VIII. A generic survey of the American
Triclisieae and Anomospermeae. Mem. New York Bot. Gard. 22(2): 1-89.
Diels, L. 1934. Menispermaceae. In A. Pulle, Flora of Suriname 2(1): 123-131.
Jansen-Jacobs, M.J. 1976. Menispermaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2):
430-440.
Jonker, F.P. 1939. Menispermaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 476-477.
Krukoff, B.A. & R.C. Bameby. 1938. Studies ofAmerican Menispermaceae, with special reference to species used in preparation
of arrow-poisons. Brittonia 3: 1-74. (See also numerous articles by R.C. Bameby, B.A. Krukoff, and H.N. Moldenke in a
series of papers published from 1941 through 1971 in Bull. Torrey Bot. Club and Mem. New York Bot. Gard.)
Menyanthaceae
K. Yatskievych
Nymphoides indica (L.) Kuntze BO GU SU FG
Synonyms and Misidentifications
Menyanthes indica L. =Nymphoides indica
Nymphoides humboldtiana (Kunth) Kuntze =Nymphoides indica
Nymphoides indica subsp. occidentalis A. Raynal =Nymphoides indica
Villarsia humboldtiana Kunth =Nymphoides indica
References
Jonker, F.P. 1936. Menyanthaceae. In A. Pulle, Flora of Suriname 4(1): 428-430.
Yatskievych, K. 2001. Menyanthaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 6: 578-579.
MOLLUGINACEAE
.A. DeFilippst, J.A. Steyermark!
AM BO DA GU FG
AM BO DA GU SU FG
Glinus radiatus (Ruiz & Pav.) Rohrb.
Mollugo verticillata L.
Trianthema portulacastnmi L. [pantropical weed, expected]
Synonyms and Misidentifications
Mollugo radiata Ruiz & Pav. =Glinus radiatus
References
Burger, W. 1983. Aizoaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 199-212.
Eyma, P.J. 1934. Aizoaceae. In A. Pulle, Flora of Suriname 1(1): 158-161.
Nevling, L.I. Jr. 1961 . Aizoaceae. In R.E. Woodson Jr., R.W. Schery, & collaborators. Flora ofPanama. Ann. Missouri Bot. Gard.
48(1): 80-85.
Steyermark, J.A. 2001. Molluginaceaein J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 6: 686-688.
Mollinedia grazielae Peixoto
Mollinedia killipii J.F. Macbr.
Mollinedia ovata Ruiz & Pav.
Mollinedia repanda Ruiz & Pav.
MONIMIACEAE
RE. Berry, A.L. Peixoto, S.S. Renner
AM GU
AM
AM BO GU
AM BO GU
SU
SU FG
Synonyms and Misidentifications
=Mollinedia ovataMollinedia glabricaulis Maguire & Steyerm
Mollinedia laurina Tul. =Mollinedia ovata
Mollinedia neblinensis Maguire & Steyerm. =Mollinedia ovata
Mollinedia ptariensis Steyerm. =Mollinedia ovata
Mollinedia roraimensis Steyerm. =Mollinedia repanda
Mollinedia yaviensis Lasser & Maguire ^Mollinedia repanda
Checklist of the Plants of the Guiana Shield 423
References
Berry, RE. & A.L. Peixoto. 2001. Monimiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 6: 689-691.
Jansen-Jacobs, M.J. 1976. Monimiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 425-
426.
Lanjouw, J. 1939. Monimiaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 471-472.
Mitchell, J.D. 2002. Monimiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
510-515.
Peixoto, A.L. 1987. Nota sobre as especies de Mollinedia (Monimiaceae, Monimioideae) das Guianas. Bradea 4(41): 331-336.
Perkins, J. & E. Gilg. 1901. Monimiaceae. In A. Engler, Das Pflanzenreich IV(lOl): 1-122.
Petter, H.F.M. 1932. Monimiaceae. In A. Pulle, Flora of Suriname 2(1): 107-112.
Steyermark, J.A. 1951. Monimiaceae. In Botanical Exploration in Venezuela - 1. Fieldiana Bot. 28(1): 235-236.
MONOTACEAE
J.S. Miller
Pakaraimaea dipterocarpacea Maguire & RS. Ashton
Pakaraimaea dipterocarpacea subsp. dipterocarpacea
Pakaraimaea dipterocarpacea subsp. nitida Maguire & Steyerm.
BO
BO
GU
GU
References
Maguire, B. & RS. Ashton. 1977. Pakaraimoideae, Dipterocarpaceae of the Western Hemisphere. II. Systematic, geographic,
and phyletic considerations. Taxon 26(4): 343-368.
Maguire, B. & RS. Ashton. 1980. Pakaraimaea dipterocarpacea II. Taxon 29(2/3): 225-231.
Maguire, B. & J.A. Steyermark. 1981. Pakaraimaea dipterocarpacea - III. In B. Maguire & collaborators. Botany of the Guayana
Highland part 11. Mem. New York Bot. Gard. 32: 1-391.
Miller, J.S. 2001. Monotaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
6: 691-692.
MORACEAE
C.C. Berg
Bagassa guianensis Aubl.
Batocarpus amazonicus (Ducke) Fosberg
Brosimum acutifolium Huber
Brosimum acutifolium subsp. acutifolium
Brosimum acutifolium subsp. obovatum (Ducke) C.C Berg
Brosimum alicastrum Sw.
Brosimum alicastrum subsp. bolivarense (Pittier) C.C. Berg
Brosimum guianense (Aubl.) Huber
Brosimum lactescens (S. Moore) C.C. Berg
Brosimum melanopotamicum C.C. Berg
Brosimum parinarioides Ducke
Brosimum parinarioides subsp. parinarioides
Brosimum potabile Ducke
Brosimum rubescens Taub.
Brosimum utile (Kunth) Pittier
Brosimum utile subsp. longifolium (Ducke) C.C. Berg
Brosimum utile subsp. ovatifolium (Ducke) C.C. Berg
Castilla elastica Sesse
Castilla elastica subsp. elastica
Clarisia biflora Ruiz & Pav.
Clarisia ilicifolia (Spreng.) Lanj. & Rossberg
Clarisia racemosa Ruiz & Pav.
Dorstenia brasiliensis Lam.
Dorstenia cayapia subsp. asaroides (Hook.) C.C. Berg [expected in SU]
Dorstenia contrajerva L. [expected in GU]
Ficus albert-smithii Standi.
Ficus amazonica (Miq.) Miq.
Ficus americana Aubl.
Ficus blepharophylla Vazq. Avila [Mt. Roraima]
Ficus broadwayi Urb.
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
DA
DA
GU SU FG
FG
GU SU FG
SU FG
GU FG
GU SU? FG
GU
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU SU FG
GU SU FG
GU
GU
GU
GU
GU
GU
GU
SU
SU
SU
SU
SU
SU
SU
FG
FG
FG
FG
FG
FG
424 Checklist of the Plants of the Guiana Shield
Ficus cabalUna Standi.
Ficus catappifolia Kunth & Bouche
Ficus cremersii C.C. Berg
Ficus crocata (Miq.) Miq.
Ficus dendwcida Kunth
Ficus donnell-smithii Standi.
Ficus eximia Schott
Ficus gomelleira Kunth & Bouche
Ficus greiffwna Dugand
Ficus guianensis Desv. ex Ham.
Ficus hebetifolia Dugand
Ficus insipida Willd.
Ficus insipida subsp. insipida
Ficus insipida subsp. scabra C.C. Berg
Ficus krukovii Standi.
Ficus leiophylla C.C. Berg
Ficus malacocarpa Standi.
Ficus maroniensis Benoist
Ficus mathewsii (Miq.) Miq.
Ficus matiziana Dugand
Ficus maxima Mill.
Ficus moUicula Pittier
Ficus nymphaeifolia Mill.
Ficus obtusifolia Kunth
Ficus padifolia Kunth
Ficus pakkensis Standi.
Ficus paludica Standi.
Ficus panurensis Standi.
Ficus paraensis (Miq.) Miq.
Ficus pertusa L. f.
Ficus piresiana Vazq. Avila & C.C. Berg
Ficus pulchella Schott
Ficus roraimensis C.C. Berg
Ficus schumacheri (Liebm.) Griseb.
Ficus sphenophylla Standi.
Ficus subapiculata (Miq.) Miq.
Ficus tepuiensis C.C. Berg & Simonis [expected in GU]
Ficus trigona L. f.
Ficus trigonala L.
Ficus yoponensis Desv.
Helicostylis elegans (J.F. Macbr.) C.C. Berg
Helicostylis pedunculata Benoist
Helicostylis scabra (J.F. Macbr.) C.C. Berg
Helicostylis tomentosa (Poepp. & Endl.) Rusby
Madura tinctoria (L.) D. Don ex Steud.
Madura tinctoria subsp. tinctoria
Maquira calophylla (Poepp. & Endl.) C.C. Berg
Maquira coriacea (H. Karst.) C.C. Berg
Maquira guianensis Aubl.
Maquira guianensis subsp. costaricana (Standi.) C.C. Berg
Maquira guianensis subsp. guianensis
Maquira sderophylla (Ducke) C.C. Berg
Naudeopsis glabra Spruce ex Pittier
Naudeopsis guianensis (Mildbr.) C.C. Berg
Naudeopsis oblongifolia (Kuhlm.) Carauta
Naudeopsis ulei subsp. ulei [expected in FG]
Perebea guianensis Aubl.
Perebea guianensis subsp. guianensis
Perebea mollis (Poepp. & Endl.) Huber
Perebea mollis subsp. mollis
Perebea rubra (Trecul) C.C. Berg
Poulsenia sp.
Pseudolmedia laevigata Trecul
Pseudolmedia laevis (Ruiz & Pav.) J.F. Macbr.
AM BO DA GU SU FG
BO DA GU SU FG
FG
BO
DA
AM BO GU SU FG
AM DA GU
AM BO DA GU
GU
SU FG
FG
AM BO DA GU SU FG
AM FG
BO DA GU SU FG
BO
BO DA GU SU FG
AM SU
SU
FG
FG
AM BO DA GU SU FG
GU SU FG
AM BO GU SU
AM GU FG?
AM BO DA GU SU FG
AM BO
AM BO DA GU SU FG
AM GU?
GU?
BO GU SU FG
DA GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM GU
GU
FG
FG?
FG?
AM BO DA GU SU FG
AM BO
GU? SU? FG
BO
AM BO
BO
DA GU
GU?
SU
SU?
FG
AM BO
SU FG
AM
AM BO DA GU SU FG
BO GU SU FG
BO GU
AM FG
AM BO SU
BO GU SU FG
BO
BO GU SU FG
AM GU SU
SU
FG
FG
GU SU FG
AM
AM GU SU FG
AM GU SU
SU
SU
SU
FG
FG
FG
AM BO GU FG
AM BO DA GU SU FG
AM BO GU SU FG
AM BO GU SU
AM BO SU FG
AM BO GU SU
AM
AM BO
BO
BO
GU
GU
GU
SU
AM BO GU SU FG
GU SU FG
Checklist of the Plants of the Guiana Shield 425
Sorocea muriculata Miq.
Sorocea muriculata subsp. muriculata
Sorocea muriculata subsp. uaupensis (Baill.) C.C. Berg
Sorocea pubivena Hemsl.
Sorocea pubivena subsp. hirtella (Mildbr.) C.C. Berg
Sorocea pubivena subsp. oligotricha (Akkermans & C.C. Berg) C.C. Berg
Sorocea sprucei (Baill.) J.F. Macbr.
Sorocea sprucei subsp. sprucei
Trymatococcus amazonicus Poepp. & Endl.
Trymatococcus oligandrus (Benoist) Lanj.
Synonyms and Misidentifications
Acanthinophyllum ilicifolia (Spreng.) W. Burger ^Clarisia ilicifolia
Acanthinophyllum spruceana (Lanj.) W. Burger =Clarisia ilicifolia
Acanthinophyllum strepitans Allemao =Clarisia ilicifolia
Acanthosphaera amara (Ducke) Ducke =Naucleopsis ulei
Alicastrum guianense (Aubl.) Kuntze =Brosimum guianense
Aliteria sagotii Benoist =Clarisia ilicifolia
Anonocarpus amazonicus Ducke =Batocarpus amazonicus
Bagassa sagotiana Bureau ex Benth. & Hook. f. =Bagassa guianensis
Bagassa tiliifolia (Ham.) Benoist =Bagassa guianensis
Brosimopsis acutifolia (Huber) Ducke =Brosimum acutifolium
Brosimopsis lactescens S. Moore =Brosimum lactescens
Brosimopsis obovata Ducke =Brosimum acutifolium subsp. obovatum
Brosimum angustifolium Ducke =Brosimum rubescens
Brosimum aubletii Poepp. & Endl. =Brosimum guianense
Brosimum bolivarense (Pittier) Romero ^Brosimum alicastrum subsp. bolivarense
Brosimum discolor Schott =Brosimum guianense
Brosimum lanciferum Ducke =Brosimum rubescens
Brosimum latifolium Standi. =Brosimum alicastrum subsp. bolivarense
Brosimum lemeei (Benoist) Lemee =Brosimum guianense
Brosimum longifolium Ducke =Brosimum utile subsp. longifolium
Brosimum mello-barretoi Standi. =Naucleopsis oblongifolia
Brosimum myristicoides Standi. =Brosimum potabile
Brosimum ovatifolium Ducke =Brosimum utile subsp. ovatifolium
Brosimum palmarum Standi. =Brosimum guianense
Brosimum paraense Huber =Brosimum rubescens
Brosimum rigidum Ducke ^Brosimum utile subsp. ovatifolium
Brosimum rotundatum Standi. =Brosimum guianense
Brosimum velutinum (S.F. Blake) Ducke =Brosimum guianense
Chlorophora tinctoria (L.) Gaudich. ex Benth. & Hook. f. =Maclura tinctoria
Clarisia ilicifolia var. cuyunensis (Lanj.) Lanj. & Rossberg =Clarisia ilicifolia
Clarisia ilicifolia var. guianensis (Lanj.) Lanj. & Rossberg ^Clarisia ilicifolia
Clarisia spruceana Lanj. =Clarisia ilicifolia
Clarisia strepitans (Allemao) Lanj. =Clarisia ilicifolia
Clarisia strepitans var. cuyunensis Lanj. =Clarisia ilicifolia
Clarisia strepitans var. guianensis Lanj. =Clarisia ilicifolia
Dorstenia sabanensis Cuatrec. =Dorstenia brasiliensis
Dorstenia tubicina Ruiz & Pav. =Dorstenia brasiliensis
Exoecaria ilicifolia Spreng. =Clarisia ilicifolia
Ferolia guianensis Aubl. =Brosimum rubescens
Ficus acarouaniensis Benoist =Ficus gomelleira
Ficus anguina Benoist =Ficus nymphaeifolia
Ficus angustifolia (Miq.) Miq. =Ficus amazonica
Ficus arctocarpa Standi. =Ficus panurensis
Ficus arukensis Standi. =Ficus paraensis
Ficus citrifolia Mill, [sensu stricto, not in flora area]
Ficus clusiifolia Schott =Ficus guianensis
Ficus corpulenta Pittier =Ficus guianensis
Ficus ernstiana Pittier =Ficus trigona
Ficus erratica Standi. =Ficus guianensis
Ficus erythrosticta (Miq.) Miq. =Ficus pertusa
Ficus expansa Pittier =Ficus eximia
Ficus fagifolia (Miq.) Miq. =Ficus trigona
426 Checklist of the Plants of the Guiana Shield
Ficusfanshawei Standi. =Ficus trigona
Ficus foveata Pittier ^Ficus eximia
Ficus foveolata Pittier =Ficus eximia
Ficus gardneriana (Miq.) Miq. [not in flora area] =Ficus obtusifolia
Ficus gemina (Miq.) Miq. =Ficus pertusa
Ficus gigantea Kunth [not in flora area] =Ficus citrifolia
Ficus glabrata Kunth =Ficus insipida
Ficus glandulosa Pittier =Ficus eximia
Ficus glaucescens (Liebm.) Miq. =Ficus maxima
Ficus gleasonii Standi. =Ficus mathewsii
Ficus gracilis Pittier (Aquifoliaceae) =Ilex divahcata
Ficus guanarensis Pittier =Ficus eximia
Ficus hartii Warb. =Ficus trigonata
Ficus hydrophila Pittier =Ficus paracusis
Ficus involuta (Liebm.) Miq. =Ficus obtusifolia
Ficus jacquiniifolia A. Rich. =Ficus americana
Ficus kanukuensis Standi. =Ficus pertusa
Ficus lanjouwii DeWolf =F/cw5 maroniensis
Ficus leucosticta (Miq.) Miq. =Ficus paludica
Ficus longifolia Schott [sensu stricto, not in flora area]
Ficus longistipula Pittier =Ficus insipida subsp. insipida
Ficus maguirei Standi. =Ficus panurensis
Ficus manicariarum Standi. =Ficus paludica
Ficus maroana Pittier =Ficus guianensis
Ficus martinii Miq. =Ficus guianensis
Ficus mendelsonii Britton =Ficus broadwayi
Ficus mensalis Standi. =Ficus guianensis
Ficus orinocensis Pittier =Ficus paracusis
Ficus pallida Vahl [not in flora area]
Ficus palmicida Pittier =Ficus pertusa
Ficus parkeri Miq. =Ficus maxima
Ficus parkeriana (Miq.) Sandwith ^Ficus guianensis
Ficus pascuorum Pittier =Ficus obtusifolia
Ficus perforata L. =Ficus americana
Ficus plicato-ostiolata Pittier =Ficus trigona
Ficus prinoides Humb. & Bonpl. ex Willd. [not in flora area] =Ficus pallida Vahl
Ficus radula Humb. & Bonpl. ex Willd. =Ficus maxima
Ficus ramiflora Standi. =Ficus caballina
Ficus rolanderi (Liebm.) Miq. =Ficus pertusa
Ficus savannarum Standi. =Ficus broadwayi
Ficus scabrida Pittier =Ficus malacocarpa
Ficus surinamensis Miq. =Ficus amazonica
Ficus tamatamae Pittier =Ficus caballina
Ficus thelephora Benoist =Ficus paraensis
Ficus tigrensis Pittier =Ficus mollicula
Ficus turbinata Pittier =Ficus eximia
Ficus urbaniana Warb. =Ficus nymphaeifolia
Ficus velutina Willd. [not in flora area]
Ficus vulpina Benoist =Ficus trigona
Galactodendrum utile Kunth =Brosimum utile
Greeneina affinis (Miq.) Kuntze =Helicostylis tomentosa
Helicostylis affinis Steud. ex Miq. =Helicostylis tomentosa
Helicostylis asperifolia Ducke =Helicostylis scabra
Helicostylis bolivarensis Pittier =Brosimum alicastrum subsp. bolivarense
Helicostylis lancifolia Ducke =Helicostylis elegans
Helicostylis obtusifolia Standi. =Helicostylis tomentosa
Helicostylis poeppigiana (Mart.) Trecul =Helicostylis tomentosa
Lanessania oligandra Benoist =Trymatococcus oligandrus
Lanessania turbinata Baill. =Trymatococcus amazonicus
Laurea tiliifolia Gaudich. =Bagassa guianensis
Maquira costaricana (Standi.) C.C. Berg =Maquira guianensis subsp. costaricana
Morus tinetoria L. =Maclura tinetoria
Naucleopsis amara Ducke =Naucleopsis ulei subsp. ulei
Naucleopsis mello-barretoi (Standi.) C.C. Berg =Naucleopsis oblongifolia
Checklist of the Plants of the Guiana Shield 427
Noyera mollis (Poepp. & Endl.) Ducke =Perebea mollis
Noyera rubra Trecul =Perebea rubra
Ogcodeia amara (Ducke) Ducke =Naucleopsis ulei subsp. ulei
Ogcodeia glabra (Spruce ex Pittier) Mildbr. =Naucleopsis glabra
Ogcodeia guianensis Mildbr. =Naucleopsis guianensis
Ogcodeia oblongifolia Kuhlm. =Naucleopsis oblongifolia
Olmedia asperula Standi. =Helicostylis tomentosa
Olmedia calophylla Poepp. & Endl. =Maquira calophylla
Olmedia caurensis Pittier =Pseudolmedia laevis
Olmedia guianensis (Aubl.) Trecul =Maquira guianensis
Olmedia laevis Ruiz & Pav. =Pseudolmedia laevis
Olmedia maquira Steud. =Maquira guianensis
Olmedia maxima Ducke =Maquira coriacea
Olmedia mollis Poepp. & Endl. =Perebea mollis
Olmedia obliqua Huber =Maquira coriacea
Olmedia polycephala Pittier =Helicostylis tomentosa
Olmedia tomentosa Poepp. & Endl. =Helicostylis tomentosa
Olmedia virgata Pittier =Sorocea sprucei subsp. sprucei
Olmedioperebea calophylla (Poepp. & Endl.) Ducke =Maquira calophylla
Olmedioperebea sclerophylla Ducke =Maquira sclerophylla
Olmediophaena maxima (Ducke) Ducke =Maquira coriacea
Olmediophaena obliqua (Huber) Ducke ^Maquira coriacea
Perebea costaricana Standi. =Maquira guianensis subsp. costaricana
Perebea elegans J.F. Macbr. =Helicostylis elegans
Perebea laurifolia Trecul =Maquira guianensis subsp. guianensis
Perebea lecointei Huber =Perebea mollis subsp. mollis
Perebea mollis subsp. rubra (Trecul) C.C. Berg =Perebea rubra
Perebea paraensis Huber =Perebea mollis subsp. mollis
Perebea xinguana Standi. =Maquira sclerophylla
Pharmacosycea guyanensis Miq. =Ficus maxima
Pharmacosycea parkeriana Miq. =Ficus guianensis
Piratinera acutifolia (Huber) Pittier =Brosimum acutifolium
Piratinera guianensis Aubl. =Brosimum guianense
Piratinera lancifera (Ducke) Benoist =Brosimum rubescens
Piratinera lemeei Benoist =Brosimum guianense
Piratinera paraensis (Huber) Benoist =Brosimum rubescens
Piratinera scabridula S.F. Blake =Brosimum guianense
Piratinera velutina S.F. Blake =Brosimum guianense
Pseudolmedia coriacea H. Karst. =Maquira coriacea
Pseudolmedia multinervis Mildbr. =Pseudolmedia laevis
Pseudolmedia sagotii Benoist =Pseudolmedia laevis
Pseudolmedia scabra J.F. Macbr. =Helicostylis scabra
Pseudosorocea sprucei Baill. =Sorocea sprucei
Pseudosorocea uaupensis Baill. =Sorocea muriculata subsp. uaupensis
Sorocea amazonica Miq. =Sorocea muriculata subsp. muriculata
Sorocea dentata Huber =Sorocea muriculata subsp. muriculata
Sorocea guayanensis W.C. Burger =Sorocea muriculata subsp. uaupensis
Sorocea hirtella Mildbr. =Sorocea pubivena subsp. hirtella
Sorocea hirtella subsp. o//gorr/c/7a Akkermans & C.C. Berg =Sorocea pubivena subsp. oligotricha
Sorocea opima J.F. Macbr. =Sorocea pubivena subsp. hirtella
Sorocea uaupensis (Baill.) J.F. Macbr. =Sorocea muriculata subsp. uaupensis
Trymatococcus paraensis Ducke =Trymatococcus amazonicus
Trymatococcus turbinatus (Baill.) Ducke =Trymatococcus amazonicus
Urostigma amazonicum Miq. =Ficus amazonica
Urostigma angustifolium Miq. ^^Ficus amazonica
Urostigma crocatum Miq. =Ficus crocata
Urostigma erythrostictum Miq. =Ficus pertusa
Urostigmafagifolium Miq. =Ficus trigona
Urostigma gardnerianum Miq. =Ficus obtusifolia
Urostigma geminum Miq. =Ficus pertusa
Urostigma involutum Liebm. =Ficus obtusifolia
Urostigma leucostictum Miq. =Ficus paludica
Urostigma mathewsii Miq. =Ficus mathewsii
Urostigma paraense Miq. =Ficus paraensis
428 Checklist of the Plants of the Guiana Shield
Urostigma rolanderi Liebm. =Ficus pertusa
Urostigma schumacheri Liebm. =Ficus schumacheri
References (see also Cecropiaceae)
Berg, C.C. 1972. Olmedieae and Brosimeae (Moraceae). Flora Neotropica Monogr. 7: 1-229.
Berg, C.C. 1992. Moraceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 11: 10-92.
Berg, C.C. 2001. Moraceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 6:
693-729.
Berg, C.C. 2002. Moraceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 515-
525.
Berg, C.C. & R.W.A.R Akkermans. 1985. Studies on the flora of the Guianas 14. New taxa and combinations in Sorocea and a
key to its species. Proc. Kon. Ned. Akad. Wet. ser. C, 88(4): 381-394.
Berg, C.C. & G.R DeWolf 1975. Moraceae. In J. Lanjouw & A.L. Stoffers, Flora of Suriname 5(1): 173-265.
Berg, C.C, M. Vasquez Avila, & F. Kooy. 1984. Ficus species of Brazilian Amazonia and the Guianas. Acta Amaz. 14(1/2)
suppl: 159-194.
Lindeman, J.C. 1984. Moraceae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 652-654.
MORINGACEAE
J.S. Miller
Moringa oleifera Lam. [Old World family, one species widely cultivated] BO
Synonyms and Misidentifications
Moringa pterygosperma Gaertn. =Moringa oleifera
References
Miller, J.S. 2001. Moringaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
6: 729-730.
Stoffers, A.L. 1984. Moringaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 570-572.
MUNTINGIACEAE
RE. Berry
Muntingia calabura L. BO GU SU FG
References
(alt. Tiliaceae)
Myoporaceae
Bontia daphnoides L. [cultivated; dubiously native or naturalized] GU SU FG
References
Lanjouw, J. 1938. Myoporaceae. In A. Pulle, Flora of Suriname 4(2): 255-256.
MVRICACEAE
P.E. Berry, N. Ramirez
Morella rotundata (Steyerm. & Maguire) Parra-Os. BO
Synonyms and Misidentifications
Myrica rotundata Steyerm. & Maguire = Morella rotundata
References
Ramirez, N. & P.E. Berry. 2001 . Myrsinaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan
Guayana 6: 733-734.
MVRISTICACEAE
G. Aymard, P.E. Berry, W.A. Rodrigues
Compsoneura debilis (Spruce ex A. DC.) Warb. AM GU
Compsoneura sprucei (A. DC.) Warb. AM SU
AM BO
AM BO
AM BO
AM BO
AM
AM
AM BO
AM BO
AM BO
AM
AM
AM BO
AM
AM
AM
AM
AM
AM BO
AM
AM
AM BO
AM BO
AM BO
AM
GU FG
GU su FG
GU su FG
GU su
GU
GU su FG
GU su FG
su FG?
GU
GU su FG
GU su
GU su FG
FG
GU su FG
Checklist of the Plants of the Guiana Shield 429
Compsoneura itlei Warb.
Iryanthera hostmannii (Benth.) Warb.
Iryantherajuruensis Warb.
Iryanthera laevis Markgr.
Iryanthera lancifolia Ducke DA
Iryanthera macrophylla (Benth.) Warb.
Iryanthera obovata Ducke
Iryanthera paradoxa (Schwacke) Warb.
Iryanthera paraensis Huber
Iryanthera sagotiana (Benth.) Warb.
Iryanthera tessmannii Markgr.
Iryanthera tricornis Ducke BO
Iryanthera ulei Warb.
Osteophloeum platyspermum (Spruce ex A. DC.) Warb.
Virola calophylla (Spruce) Warb.
Virola duckei A..C. Sm.
F/>o/a e/o«ga?(3 (Benth.) Warb. DA
Virola kwatae Sabatier
Virola michelii Heckel
Virola minutiflora Ducke
Virola multicostata Ducke FG
Virola parvifolia Ducke
Virola pavonis (A. DC.) A.C. Sm.
Virola rugulosa (Spruce) Warb.
Virola sebifera A\M. DA GU SU FG
Virola sp. A
Virola sp. B
Virola steyermarkii W.A. Rodrigues
Virola surinamensis{Ko\.QX '^oiih.)'^2Lxh. DA GU SU FG
Virola theiodora (Spruce ex Benth.) Warb.
Virola venosa (Benth.) Warb. SU
Synonyms and Misidentifications
Compsoneura costaricensis Warb. =Compsoneura sprucei
Iryanthera congestiflora J.F. Macbr. =Iryanthera hostmannii
Iryanthera elongata Huber =Iryanthera paraensis
Iryanthera leptoclada Markgr. =Iryanthera hostmannii
Iryanthera longiflora Ducke =Iryanthera paradoxa
Iryanthera porcata A.H. Gentry ^Iryanthera lancifolia
Iryanthera sessilis Markgr. =Iryanthera paraensis
Myristica calophylla Spruce =Virola calophylla
Myristica cuspidata Spruce ex Benth. =Virola elongata
Myristica debilis Spruce ex A. DC. =Compsoneura debilis
Myristica elongata Benth. =Virola elongata
Myristicafatua auct. non Houtt. 1774(7): sensu Sw. Prodr. 96. 1788 =Virola surinamensis
Myristica hostmannii Benth. =Iryanthera hostmannii
Myristica laurifolia Spruce ex A. DC. =Compsoneura sprucei
Myristica longicuspis Spruce ex Warb. =Virola elongata
Myristica macrophylla Benth. =Iryanthera macrophylla
Myristica melinonii Benoist =Virola michelii
Myristica membranacea Poepp. ex A. DC. =Virola elongata
Myristica mexicana Hemsl. =Compsoneura sprucei
Myristica mocoa A. DC. ^Virola sebifera
Myristica panamensis Hemsl. =Virola sebifera
Myristica paradoxa Schwacke =Iryanthera paradoxa
Myristica pavonis A. DC. =Virola pavonis
Myristica platysperma Spruce ex A. DC. =Osteophloeum platyspermum
Myristica punctata Spruce ex Benth. =Virola elongata
Myristica rugulosa Spruce = Virola rugulosa
Myristica sebifera (Aubl.) Sw. =Virola sebifera
Myristica sebifera var. cordifolia A. DC. ^Virola sebifera
Myristica sebifera var. curvinervia A. DC. =Virola sebifera
Myristica sebifera var. longifolia Lam. =Virola surinamensis
Myristica sprucei A. DC. =Compsoneura sprucei
430 Checklist of the Plants of the Guiana Shield
Myristica surinamensis Rol. ex Rottb. =Virola surinamensis
Myristica theiodora Spruce ex Benth. =Virola theiodora
Myristica uaiipensis Spruce ex A. DC. =Virola elongata
Myristica venosa Benth. =Virola venosa
Myristica venosa war. pavonis (A. DC.) Warb. =Virola pavonis
Myristica venosa \av. poeppigii A. DC. =Virola venosa
Osteophloeiim sulcatum Little =Osteophloeum platyspermum
Palala calophylla (Spruce) Kuntze =Virola calophylla
Palala cuspidata (Spruce ex Benth.) Kuntze =Virola elongata
Palala debilis (Spruce ex A. DC.) Kuntze =Compsoneura debilis
Palala elongata (Benth.) Kuntze =Virola elongata
Palala macrophylla (Benth.) Kuntze =Iryanthera macrophylla
Palala membranacea (Poepp. ex A. DC.) Kuntze =Virola elongata
Palala mexicana (Hemsl.) Kuntze =Compsoneura sprucei
Palala mocoa (A. DC.) Kuntze =Virola sebifera
Palala pavonis (A. DC.) Kuntze =Virola pavonis
Palala platysperma (Spruce ex A. DC.) Kuntze =Osteophloeum platyspermum
Palala punctata (Spruce ex Benth.) Kuntze =Virola elongata
Palala rugulosa (Spruce) Kuntze =Virola rugulosa
Palala sprucei (A. DC.) Kuntze =Compsoneura sprucei
Palala surinamensis (Rol. ex Rottb.) Kuntze =Virola surinamensis
Palala theiodora (Spruce ex Benth.) Kuntze =Virola theiodora
Palala uaupensis (Spruce ex A. DC.) Kuntze =Virola elongata
Palala venosa (Benth.) Kuntze =Virola venosa
Virola boliviensis Warb. =Virola sebifera
Virola cuspidata (Spruce ex Benth.) Warb. =Virola elongata
Virola cuspidata van membranacea (Poepp. ex A. DC.) Warb. =Virola elongata
Virola elliptica A.C. Sm. =Virola pavonis
Virola elongata van longicuspis Warb. =Virola elongata
Virola elongata van punctata (Spruce ex Benth.) Warb. =Virola elongata
Virola elongata van subcordata Warb. =Virola elongata
Virola glaziovii Warb. =Virola surinamensis
Virola incolor Warb. = Virola calophylla
Virola melinonii (Benoist) A.C. Sm. = Virola michelii
Virola mocoa (A. DC.) Warb. =Virola sebifera
Virola mycetis Pulle -Virola sebifera
Virola panamensis (Hemsl.) Warb. =Virola sebifera
Virola peruviana van tomentosa Warb. =Virola sebifera
Virola rufula Warb. =Virola theiodora
Virola venezuelensis Warb. =Virola sebifera
Virola venosa var. poeppigii (A. DC.) Warb. =Virola venosa
Virola warburgii Pittier =Virola sebifera
References
Jansen-Jacobs, M.J. 1976. Myristicaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2):
427-429.
Mitchell, J.J. 2002. Myristicaceae. In Mori, S.A., G. Cremers, C. Oracle, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
525-532.
Ooststroom, S.J. van. 1934. Myristicaceae. In A. Pulle, Flora of Suriname 2(1): 113-122.
Ooststroom, S.J. van. 1939. Myristicaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 473-475.
Rodrigues, W.A. 1980. Revisao taxonomica das especies de Virola Aublet do Brasil. Acta Amaz. 10(1) suppl. 1-127.
Rodrigues, W.A., G Aymard & RE. Berry. 2001. Myristicaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 6: 734-747.
Smith, A.C. & R.R Wodehouse. 1937. The American species of Myristicaceae. Brittonia 2(5): 393-510.
Myrsinaceae
J.J. Pipoly, III, J.M. Ricketson
Ardisia elliptica Thunb.
Ardisia guianensis (Aubl.) Mez
Ardisia semicrenata Mart.
Ctenardisia stenobotrys (Standi.) Lundell & Pipoly
Cybianthus agostinianus Pipoly
AM
AM
AM
BO DA
GU
GU SU FG
FG?
Checklist of the Plants of the Guiana Shield 431
Cybianthus amplus (Mez) G. Agostini
Cybianthus apiculatiis (Steyerm.) G. Agostini
Cybianthus breweri G. Agostini
Cybianthus cardonae G. Agostini
Cybianthus crotonoides (M.R. Schomb. ex Mez) G. Agostini
Cybianthus deltatus Pipoly
Cybianthus detergens Mart.
Cybianthus diiidae (Gleason & Moldenke) G. Agostini
Cybianthusfabiolae Pipoly
Cybianthus fulvopidverulentus (Mez) G. Agostini
Cybianthus fulvopulverulentus suhsp. fulvopulverulentus
Cybianthus fulvopulverulentus subsp. magnoliifolius (Mez) Pipoly
Cybianthus fuscus Mart.
Cybianthus gardneri (A. DC.) G. Agostini
Cybianthus grandifolius (Mez) G. Agostini
Cybianthus guyanensis (A. DC.) Miq.
Cybianthus guyanensis subsp. guyanensis
Cybianthus guyanensis subsp. multipunctatus (A. DC.) Pipoly
Cybianthus guyanensis suhsp. pseudoicacoreus (Miq.) Pipoly
Cybianthus holstii Pipoly
Cybianthus huberi Pipoly
Cybianthus julianii Pipoly
Cybianthus lepidotus (Gleason) G Agostini
Cybianthus leprieurii G Agostini
Cybianthus liesneri Pipoly & Ricketson
Cybianthus lineatus (Benth.) Pipoly
Cybianthus longifolius Miq.
Cybianthus maguirei G Agostini ex Pipoly
Cybianthus microbotrys A. DC.
Cybianthus multicostatus Miq.
Cybianthus penduliflorus Mart.
Cybianthus piresii Pipoly
Cybianthus plowmanii Pipoly
Cybianthus potiaei (Mez) G Agostini [=Cybianthus microbotrys?]
Cybianthus prevostae Pipoly
Cybianthus prieurii A. DC.
Cybianthus ptariensis (Steyerm.) Pipoly
Cybianthus punctatus (Mez) G Agostini
Cybianthus quelchii (N.E. Br.) G. Agostini
Cybianthus resinosus Mez [not in flora area]
Cybianthus reticulatus (Benth. ex Mez) G Agostini
Cybianthus roraimae (Steyerm.) G. Agostini
Cybianthus sipapoensis Pipoly & G Agostini
Cybianthus sp. nov. 1
Cybianthus sp. nov. 2
Cybianthus spathulifolius G. Agostini ex Pipoly
Cybianthus spicatus (Kunth) G Agostini
Cybianthus steyermarkianus (G Agostini) G Agostini
Cybianthus surinamensis (Spreng.) G. Agostini
Cybianthus venezuelanus Mez
Cybianthus wurdackii G. Agostini ex Pipoly
Myrsine coriacea (Sw.) R. Br. ex Roem. & Schult.
Myrsine coriacea subsp. coriacea
Myrsine coriacea subsp. reticulata (Steyerm.) Pipoly
Myrsine guianensis (Aubl.) Kuntze
Myrsine macrocarpa Pipoly
Myrsine maguireana Pipoly
Myrsine minima (Steyerm.) Pipoly
Myrsine nitida (Mez) Pipoly
Myrsine pellucida (Ruiz & Pav.) Spreng.
Myrsine perpauciflora Pipoly
Myrsine picturata Pipoly
Myrsine resinosa (A.C. Sm.) Pipoly [Mt. Roraima]
Stylogyne atra Mez
AM BO GU FG
AM
BO
GU
AM BO
BO GU
AM
GU
AM BO GU
GU
AM BO DA GU su FG
AM BO GU su
AM BO DA GU su FG
su FG?
FG
BO GU FG
AM BO GU su FG
AM GU FG
BO GU su FG
AM GU su FG
AM
BO
AM
AM BO GU
FG
AM
AM BO GU
AM
AM
GU su FG
AM
AM
AM
AM
BO , GU su FG
FG
AM BO
BO
BO
GU
GU
GU
su FG
AM BO GU
AM
AM
AM
BO GU
GU
GU
AM
AM BO
BO
GU
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AM BO
BO
GU su FG
AM BO GU
AM BO
BO
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AM
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GU
GU
AM
AM BO
BO GU
AM SU
432 Checklist of the Plants of the Guiana Shield
Stylogyne depaiiperata Mez
Stylogyne incognita Pipoly
Stylogyne lasseri (Lundell) Pipoly
Stylogyne longifolia (Mart, ex Miq.) Mez
Stylogyne micrantha (Kunth) Mez
Stylogyne orinocensis (Kunth) Mez
Stylogyne schombiirgkiana (A. DC.) Mez
Stylogyne viridis (Lundell) Ricketson & Pipoly
AM
AM
BO
BO
BO
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GU su FG
GU FG
Synonyms and Misidentifications
Ardisia acuminata Willd. ^Ardisia guianensis
Ardisia acuminifolia Lundell =Ardisia guianensis
Ardisia amanuensis Lundell =Ardisia guianensis
Ardisia cookii Lundell ^Ardisia guianensis
Ardisia hostmannii Miq. =Stylogyne orinocensis
Ardisia lasseri Lundell =Stylogyne lasseri
Ardisia longicaudata Lundell ^Ardisia guianensis
Ardisia micrantha Kunth =Stylogyne micrantha
Ardisia nigricans Miq. =Stylogyne atra
Ardisia nigricans Miq. =Stylogyne orinocensis
Ardisia orinocensis Kunth =Stylogyne orinocensis
Ardisia pseudoicacorea Miq. =Cybianthus guyanensis subsp. pseudoicacoreus
Ardisia quelchii N.E. Br. =Cybianthus quelchii
Ardisia schomburgkiana (A. DC.) Miq. =Stylogyne schomburgkiana
Ardisia stenobotiys Standi. =Ctenardisia stenobotrys
Ardisia surinamensis Miq. =Stylogyne orinocensis
Badula schomburgkiana A. DC. =Stylogyne schomburgkiana
Caballeriaferruginea Ruiz & Pav. =Myrsine coriacea subsp. coriacea
Caballeria pellucida Ruiz & Pav. ^Myrsine pellucida
Conomorpha ampla Mez =Cybianthus amplus
Conomorpha apiculata Steyerm. -Cybianthus apiculatus
Conomorpha candolleana Mez =Cybianthus spicatus
Conomorpha crotonoides M.R. Schomb. ex Mez =Cybianthus crotonoides
Conomorpha curvivenia Gleason =Cybianthus lepidotus
Conomorpha depressa Steyerm. =Cybianthus quelchii
Conomorpha duidae Gleason & Moldenke =Cybianthus duidae
Conomorphafulvopulverulenta Mez =Cybianthus fulvopulverulentus
Conomorpha glaucorubens Mez =Cybianthus spicatus
Conomorpha gracilis A.C. Sm. =Cybianthus spicatus
Conomorpha grandiflora Mez =Cybianthus spicatus
Conomorpha guyanensis A. DC. =Cybianthus guyanensis
Conomorpha heterantha Benth. ex Miq. =Cybianthus guyanensis subsp. guyanensis
Conomorpha latifolia (Miq.) Mez =Cybianthus spicatus
Conomorpha laxiflora A. DC. =Cybianthus spicatus
Conomorpha laxiflora var. latifolia Miq. =Cybianthus spicatus
Conomorpha lepidota Gleason =Cybianthus lepidotus
Conomorpha lepidota f. acutata Steyerm. =Cybianthus lepidotus
Conomorpha leprosa Steyerm. =Cybianthus fulvopulverulentus subsp. fulvopulverulentus
Conomorpha lithophyta R.E. Schult. =Cybianthus fulvopulverulentus subsp. fulvopulverulentus
Conomorpha macrophylla Mart. =Cybianthus amplus
Conomorpha madeirensis A.C. Sm. =Cybianthus spicatus
Conomorpha magnoliifolia Mez =Cybianthus fulvopulverulentus subsp. magnoliifolius
Conomorpha multipunctata (A. DC.) Miq. =Cybianthus guyanensis subsp. pseudoicacoreus
Conomorpha peruviana var. guyanensis (A. DC.) Mez =Cybianthus guyanensis
Conomorpha ptariensis Steyerm. =Cybianthus duidae
Conomorpha punctata Mez =Cybianthus punctatus
Conomorpha reticulata Benth. ex Miq. =Cybianthus reticulatus
Conomorpha rigida Mez =Cybianthus fulvopulverulentus subsp. magnoliifolius
Conomorpha riparia R.E. Schult. =Cybianthus fulvopulverulentus subsp. fulvopulverulentus
Conomorpha roraimae Steyerm. =Cybianthus roraimae
Conomorpha sessilis A.C. Sm. =Cybianthus punctatus
Conomorpha spicata (Kunth) Mez =Cybianthus spicatus
Conomorpha steyermarkiana G. Agostini =Cybianthus steyermarkianus
Conomorpha utiarityi Hoehne =Cybianthus amplus
Checklist of the Plants of the Guiana Shield 433
Ctenardisia stenobotrys (Standi.) Fonnegra & S.L. Jung [nom. superfl.] =Ctenardisia stenobotrys
Cybianthus brownii Gleason =Cybianthus venezuelanus
Cybianthus comatus Mez =Cybianthus prieurii
Cybianthus egemis Mez =Cybianthus venezuelanus
Cybianthus foliosus Rusby =Stylogyne micrantha
Cybianthus gracillimus Mez [GU records misident., fide Pipoly]
Cybianthus lithophytus (R.E. Schult.) G. Agostini =Cybianthus fulvopulverulentus subsp. fulvopulverulentus
Cybianthus magnoliifolius (Mez) G Agostini =Cybianthus fulvopulvendentus subsp. magnoliifolius
Cybianthus multipunctatus A. DC. =Cybianthus guyanensis subsp. multipunctatus
Cybianthus nitidus Miq. =Cybianthus prieurii
Cybianthus pseudoicacoreus (Miq.) G Agostini =Cybianthus guyanensis subsp. pseudoicacoreus
Cybianthus subspicatus Benth. ex Miq. =Cybianthus prieurii
Cybianthus sylvaticus (Gleason) G. Agostini =Cybianthus grandifolius
Cybianthus viridiflorus A.C. Sm. =Cybianthus prieurii
Grammadenia lineata Benth. =Cybianthus lineatus
Grammadenia ptariensis Steyerm. =Cybianthus ptariensis
Grammadenia ptariensis subsp. auyantepuiensis G Agostini =Cybianthus ptariensis
Icacorea acuminata (Willd.) Lundell [not in flora area] =Ardisia guianensis
Icacorea acuminifolia (Lundell) Lundell =Ardisia guianensis
Icacorea amanuensis (Lundell) Lundell =Ardisia guianensis
Icacorea cookii (Lundell) Lundell =Ardisia guianensis
Icacorea guianensis Aubl. =Ardisia guianensis
Icacorea longicaudata (Lundell) Lundell =Ardisia guianensis
Manglillaferruginea (Ruiz & Pav.) Roem. & Schult. =Myrsine coriacea subsp. coriacea
Manglilla pellucida (Ruiz & Pav.) Roem. & Schult. =Myrsine pellucida
Myrsineferruginea (Ruiz & Pav.) Spreng. =Myrsine coriacea subsp. coriacea
Myrsine myricoides Schltdl. =Myrsine coriacea subsp. coriacea
Myrsine ovalifolia Miq. =Myrsine guianensis
Myrsine roraimensis (A.C. Sm.) Pipoly =Myrsine nitida
Myrsine schomburgkiana Miq. (Sapotaceae) =Elaeoluma schomburgkiana
Myrsine spicata Kunth =Cybianthus spicatus
Parathesis viridis Lundell =Stylogyne viridis
Peckia guyanensis (A. DC.) Kuntze =Cybianthus guyanensis
Peckia longifolia (Miq.) Kuntze =Cybianthus longifolius
Peckia penduliflora (Mart.) Kuntze =Cybianthus penduliflorus
Peckia prieureii (A. DC.) Kuntze =Cybianthus prieurii
Peckia purpurea Rusby =Cybianthus venezuelanus
Peckia surinamensis (Spreng.) Kuntze =Cybianthus surinamensis
Rapanea ambigua Mez =Myrsine coriacea subsp. coriacea
Rapanea coriacea (Sw.) Mez =Myrsine coriacea
Rapaneaferruginea (Ruiz & Pav.) Mez =Myrsine coriacea subsp. coriacea
Rapanea guianensis Aubl. =Myrsine guianensis
Rapanea minima Steyerm. =Myrsine minima
Rapanea myricoides (Schltdl.) Lundell =Myrsine coriacea subsp. coriacea
Rapanea nitida Mez =Myrsine nitida
Rapanea oblonga Pohl ex Miq. =Myrsine guianensis
Rapanea ovalifolia (Miq.) Mez =Myrsine guianensis
Rapanea pellucida (Ruiz & Pav.) Mez =Myrsine pellucida
Rapanea perforata Mez =Myrsine pellucida
Rapanea resinosa A.C. Sm. =Myrsine resinosa
Rapanea reticulata Steyerm. =Myrsine coriacea subsp. reticulata
Rapanea roraimensis A.C. Sm. =Myrsine nitida
Salvadora surinamensis Spreng. =Cybianthus surinamensis
Samara coriacea Sw. =Myrsine coriacea
Stylogyne amazonica Mez =Stylogyne orinocensis
Stylogyne kappleri Mez ^Stylogyne orinocensis
Stylogyne latifolia A.C. Sm. =Stylogyne micrantha
Stylogyne micans Mez =Stylogyne orinocensis
Stylogyne sprueeana Mez =Stylogyne atra
Stylogyne surinamensis (Miq.) Mez =Stylogyne orinocensis
Stylogyne turbacensis Mez [reported from FG in error, fide Pipoly]
Stylogyne venezuelana Mez =Stylogyne micrantha
Tinus guianensis (Aubl.) Kuntze =Ardisia guianensis
Tinus hostmannii (Miq.) Kuntze =Stylogyne orinocensis
434 Checklist of the Plants of the Guiana Shield
Tinus orinocensis (Kunth) Kuntze =Stylogyne orinocensis
Tinus schomburgkiana (A. DC.) Kuntze =Stylogyne schomburgkiana
Tinus surinamensis (Miq.) Kuntze =Stylogyne orinocensis
Walleria laxiflora Mart. =Cybianthiis spicatus
Weigeltia grandifolia Mez =Cybianthus grandifolius
Weigeltia grandifolius Mez =Cybianthus grandifolius
Weigeltia longifolia (Miq.) Benth. =Cybianthus longifolius
Weigeltia microbotrys (DC.) Mez =Cybianthus microbotrys
Weigeltia parviflora Mez =Cybianthus leprieurii
Weigeltia potiaei Mez =Cybianthus potiaei
Weigeltia quelchii (N.E. Br.) Mez =Cybianthus quelchii
Weigeltia surinamensis (Spreng.) Mez =Cybianthus surinamensis
Weigeltia sylvatica Gleason =Cybianthus grandifolius
References
Agostini, G. 1980. Una nueva clasificacion del genero Cybianthus (Myrsinaceae). Acta Biol. Venez. 10: 129-185.
Bottelier, H.P. 1936. Myrsinaceae. In A. Pulle, Flora of Suriname 4(1): 431-442.
Pipoly, J.J. 1983. Contributions toward a monograph of Cybianthus, III: a revision of subgenus Laxiflorus. Brittonia 35(1): 61-
80.
Pipoly, J.J. 1992. The genus Cybianthus subgenus Conomorpha in Guayana. Ann. Missouri Bot. Gard. 79(4): 908-957.
Pipoly, J.J. 2002. Myrsinaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 532-
539.
Pipoly, J.J. & J.M. Ricketson. 2001. Myrsinaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 6: 748-775.
Myrtaceae
F. Grifo, P. Hiepko, B.K. Hoist, L. Landrum
Blepharocalyx eggersii (Kiaersk.) Landrum
Calycolpus alternifolius (Gleason) Landrum
Calycolpus bolivarensis Landrum
Calycolpus calophyllus (Kunth) O. Berg
Calycolpus cochleatus McVaugh
Calycolpus goetheanus (DC.) O. Berg
Calycolpus revolutus (Schauer) O. Berg
Calycolpus roraimensis Steyerm.
Calycolpus schomburgkianus O. Berg
Calycolpus surinamensis McVaugh
Calycorectes batavorum McVaugh
Calycorectes bergii Sandwith
Calycorectes enormis McVaugh
Calycorectes grandifolius O. Berg
Calycorectes guyanensis Mattos
Calycorectes maximus McVaugh
Calyptranthes amshoffae McVaugh
Calyptranthes bipennis O. Berg
Calyptranthes bracteata M.L. Kawas. & B. Hoist
Calyptranthes clusiifolia O. Berg
Calyptranthes conduplicata B. Hoist
Calyptranthes crebra McVaugh
Calyptranthes fasciculata O. Berg
Calyptranthes forsteri O. Berg
Calyptranthes lepida McVaugh
Calyptranthes lucida Mart, ex DC.
Calyptranthes macrophylla O. Berg
Calyptranthes multiflora O. Berg
Calyptranthes nigrescens B. Hoist
Calyptranthes ovoidea McVaugh
Calyptranthes pulchella DC.
Calyptranthes pullei Burret ex Amshoff
Calyptranthes pullei var. immaculata McVaugh
Calyptranthes ruiziana O. Berg
Calyptranthes smithii McVaugh
AM BO
AM BO
BO
AM
BO
BO
DA
DA
GU
BO DA
GU
GU SU FG
GU SU FG
GU
GU
GU SU
SU
GU SU FG
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AM SU
AM
AM BO DA GU
AM
GU
AM BO GU SU
AM BO GU SU
AM BO
AM
FG
FG
FG
GU
Checklist of the Plants of the Guiana Shield 435
Calyptranthes speciosa Sagot
Calyptranthes speciosa var. speciosa
Calyptranthes spicata Amshoff
Calyptranthes vexata McVaugh
Campomanesia aromatica (Aubl.) Griseb.
Campomanesia grandiflora (Aubl.) Sagot
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Eugen.
Eugen.
Eugen.
Eugen.
Eugen.
Eugen
Eugen
Eugen
Eugen
Eugen
Eugen
Eugen
Eugen
Eugen
Eugen
Eugen
Eugen
Eugen
Eugen
a albicans (O. Berg) Urb.
a amblyosepala McVaugh
a amshojfae McVaugh
a anastomosans DC.
a arawakorum Sandwith
a argyrophylla B. Hoist & M.L. Kawas.
a armeniaca Sagot
a baileyi Britton
a biflora (L.) DC.
a brownsbergii Amshoff
a cachoeirensis O. Berg
a callichroma McVaugh
a chysophylhim Poir.
a citrifolia Poir.
a coffeifolia DC.
a conduplicata B. Hoist
a conjuncta Amshoff
a cowanii McVaugh
a cribrata McVaugh
a cuaoensis McVaugh
a cucullata Amshoff
a cupulata Amshoff
a cupulata var. cupulata
a denigrata McVaugh
a dentata (O. Berg) Nied.
a diplocampta Diels
a dittocrepis O. Berg
a egensis DC.
a emarginata (Kunth) DC.
a essequiboensis Sandwith
a eurycheila O. Berg
a exaltata Rich, ex O. Berg
a excelsa O. Berg
a fasciculiflora O. Berg
afeijoi O. Berg
a ferreiraeana O. Berg
aflavescens DC.
a florida DC.
afoetida Pers. [unverified name]
a gomesiana O. Berg
a gongylocarpa M.L. Kawas. & B. Hoist
a griseiflora McVaugh
a incanescens Benth.
a kaieteurensis Amshoff
a lambertiana DC.
a lambertiana var. hispidula McVaugh
a lambertiana var. lambertiana
a latifolia Aubl.
a ligustrina (Sw.) Willd.
a ligustrina var. hebecarpa Amshoff
a ligustrina var. ligustrina
a longiracemosa Kiaersk.
a luciae Amshoff
a macrocalyx (Rusby) McVaugh
a magna B. Hoist
a marlierioides Rusby
a marowynensis Miq.
a mimus McVaugh
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
AM
BO
AM BO
BO
BO
AM
AM
AM
BO
AM
AM
AM BO
AM
AM BO
AM BO
BO
AM BO
DA
DA
DA
DA
DA
DA
GU su FG
GU su
su
FG
GU
GU su FG
GU su FG
GU su FG
GU
GU su FG
GU
FG
GU
su
FG
GU su FG
su FG
su FG
su FG
GU su FG
GU
GU su FG
GU'?
GU su FG
su FG
su FG
FG
FG
GU SU
GU SU
SU
FG'
GU
GU
FG
GU SU FG
GU su FG
GU su FG
GU su FG
GU su FG
GU su FG
FG
FG
GU
BO GU
AM BO DA GU SU
SU
FG
AM BO DA GU SU FG
GU SU FG
BO GU
GU
su
SU
BO GU
AM
GU su FG
BO GU su FG
BO GU
AM
GU
su FG
FG
436 Checklist of the Plants of the Guiana Shield
Eugenia minuscida McVaugh
Eugenia monticola (Sw.) DC.
Eugenia morii B. Hoist & M.L. Kawas.
Eugenia moritziana H. Karst.
Eugenia muricata DC.
Eugenia omissa McVaugh
Eugenia pachystachya McVaugh
Eugenia patens Poir.
Eugenia patrisii Vahl
Eugenia polystachya Rich.
Eugenia polystachyoides Amshoff
Eugenia producta DC.
Eugenia protenta McVaugh
Eugenia pseudopsidium Jacq.
Eugenia pubescens (Kunth) DC.
Eugenia punicifolia (Kunth) DC.
Eugenia quadriovulata Amshoff
Eugenia ramiflora Desv.
Eugenia roseiflora McVaugh
Eugenia sigillata McVaugh
Eugenia sp. A
Eugenia sp. B
Eugenia spruceana O. Berg
Eugenia stipitata McVaugh [cultivated in southern Amazonas]
Eugenia tafelbergica Amshoff
Eugenia tapacumensis O. Berg
Eugenia tepuiensis Steyerm.
Eugenia tetramera (McVaugh) M.L. Kawas. & B. Hoist
Eugenia trinervia Vahl
Eugenia tumulescens McVaugh
Eugenia umbonata McVaugh
Eugenia undulata Aubl.
Eugenia uniflora L. [cultivated; naturalized?]
Eugenia wullschlaegeliana Amshoff
Eugenia yatuae (McVaugh) B. Hoist
Marlierea bipennis (O. Berg) McVaugh
Marlierea biptera Amshoff
Marlierea buxifolia Amshoff
Marlierea caesariata McVaugh
Marlierea cana McVaugh
Marlierea caudata McVaugh
Marlierea convexivenia B. Hoist
Marlierea cuprea Amshoff
Marlierea ensiformis McVaugh [Serra da Neblina]
Marliereaferruginea (Poir.) McVaugh
Marliereafoveolata B. Hoist
Marlierea guildingiana (Griseb.) Krug & Urb.
Marlierea insignis McVaugh
Marlierea karuaiensis (Steyerm.) McVaugh
Marlierea ligustrina McVaugh [probably restricted to Venezuela,
Marlierea lituatinervia (O. Berg) McVaugh
Marlierea macrophylla Amshoff
Marlierea maguirei McVaugh
Marlierea mcvaughii B. Hoist
Marlierea montana (Aubl.) Amshoff
Marlierea multiglomerata Amshoff
Marlierea pudica McVaugh
Marlierea rugosior McVaugh
Marlierea salticola Amshoff
Marlierea schomburgkiana O. Berg
Marlierea scytophylla Diels
Marlierea sp. A
Marlierea spruceana O. Berg
Marlierea subcordata B. Hoist
BO
BO
AM BO
AM BO
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
fide Hoist] AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
DA
DA
BO DA
BO
BO
DA
su
GU su FG?
FG
su FG
GU su FG
FG
GU su FG
GU FG
GU su
FG
GU
GU su FG
GU su FG
GU su
su FG
GU?
GU
GU
SU
FG
GU SU
GU SU FG
GU SU FG
GU FG
SU FG
SU
SU FG
GU
GU SU FG
GU
GU
GU?
GU
GU
GU su FG
su FG
GU
GU su
Checklist of the Plants of the Guiana Shield 437
Marlierea siiborbiculahs McVaugh
Marlierea siibidata McVaugh
Marlierea summa McVaugh
Marlierea umbraticola (Kunth) O. Berg
Marlierea uniflora McVaugh
Marlierea ventnarensis B. Hoist
Marlierea vicina McVaugh
Melaleuca leucadendron L. [cultivated; naturalized?]
Melaleuca quinquenervia (Cav.) S.T. Blake [cultivated? naturalized?]
AM
AM
AM BO GU
AM BO GU
AM
AM
Myrc
Myrc
Myrc
Myrc
Myrc
Myrc,
Myrc.
Myrc.
Myrc.
Myrc,
Myrc.
Myrc
Myrc,
Myrc
Myrc
Myrc
Myrc
Myrc
Myrc
Myrc
Myrc
Myrc
Myrc
Myrc
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc.
Myrc
Myrc.
Myrc
Myrc
a albidotomentosa (Amshoff) McVaugh
a aliena McVaugh
a amazonica DC.
a bolivarensis (Steyerm.) McVaugh
a bonnetiasylvestris (Steyerm.) Steyerm.
a bracteata (Rich.) DC.
a calycampa Amshoff
a citrifolia (Aubl.) Urb.
a clausa McVaugh [expected in GU]
a clusiifolia (Kunth) DC.
a compta McVaugh
a coumeta (Aubl.) DC.
a crispa McVaugh
a cuprea (O. Berg) Kiaersk.
a decorticans DC.
a deflexa (Poir.) DC.
a dichasialis McVaugh
a ehrenbergiana (O. Berg) McVaugh
a exploratoris McVaugh
a extranea McVaugh
afallax (Rich.) DC.
afenestrata DC.
afenzliana O. Berg
a gentryi B. Hoist
a gigas McVaugh
a gonini McVaugh
a graciliflora Sagot
a grandis McVaugh
a guianensis (Aubl.) DC.
a guianensis var. guianensis
a imperfecta McVaugh
a inaequiloba (DC.) D. Legrand
a induta McVaugh
a intonsa (McVaugh) B. Hoist
a kylistophylla B. Hoist
a liesneri B. Hoist
a lucida McVaugh
a magnoliifolia DC.
a minutiflora Sagot
a multiflora (Lam.) DC.
a myriantha McVaugh
a nigrescens DC.
a nubicola McVaugh
a paivae O. Berg
a pistrinalis McVaugh
a platyclada DC.
a porphyrea McVaugh
a ptariensis (Steyerm.) McVaugh
a pyrifolia (Desv. ex Ham.) Nied.
a quifarensis (Benth.) Sagot
a revolutifolia McVaugh
a roHindata (Amshoff) McVaugh
a rotimdata var. atrans McVaugh
a rotundata var. rotundata
a rufipila McVaugh
AM BO
AM
AM BO
AM BO
BO
AM BO
BO
BO
BO
AM
AM
AM BO
GU
SU
FG
SU FG?
FG
GU SU
GU SU FG
GU
GU SU FG
GU SU FG?
GU SU FG
FG
SU FG
BO DA GU SU FG
AM BO GU SU FG
AM BO
BO GU
AM BO GU
GU
SU
AM BO
BO
DA GU
GU
SU FG
FG
AM
SU
FG
GU SU FG
AM GU
AM BO DA GU SU FG
AM BO DA GU
GU
SU FG
AM BO GU SU FG
AM
BO
BO
AM
AM BO
AM BO GU SU FG
AM BO GU SU FG
AM BO GU
GU
GU
SU FG
AM BO GU
AM BO GU SU
AM BO SU
BO GU
GU
SU FG
BO
AM BO
BO
GU
GU
SU FG
AM BO
AM BO
BO
GU
AM BO GU
GU SU FG
438 Checklist of the Plants of the Guiana Shield
Myrcia rupta M.L. Kawas. & B. Hoist FG
Myrcia salticola (Steyerm.) McVaugh BO
Myrcia saxatilis (Amshoff) McVaugh SU FG
Myrcia servataMcWdoxgh AM BO DA GU SU FG
Myrcia sessiliflora McVaugh
Myrcia sipapensis McVaugh
Myrcia sororopanensis Steyerm. AM BO GU
Myrcia sp. A
Myrcia sp. B
Myrcia sp. C
Myrcia sp. D
Myrcia speciosa (Amshoff) McVaugh
Myrcia splendens (Sw.) DC. AM BO DA
Myrcia subobliqua (Benth.) Nied.
Myrcia subsessilis O. Berg
Myrcia subsessilis var. ovalis (O. Berg) McVaugh
Myrcia subsessilis var. subsessilis
Myrcia sylvatica (G. Mey.) DC.
Myrcia tafelbergica Amshoff
Myrcia tepuiensis Steyerm.
Myrcia tomentosa (Aubl.) DC.
Myrcianthesfragrans (Sw.) McVaugh
Myrcianthes prodigiosa McVaugh
Myrcianthes rhopaloides (Kunth) McVaugh
Myrciaria amazonica O. Berg [=Myrciariafloribunda, fide Sobral]
Myrciaria cordata O. Berg
Myrciaria dubia (Kunth) McVaugh
Myrciariafloribunda (West ex Willd.) O. Berg AM BO DA
Myrciaria puberulenta B. Hoist
Myrciaria tenella (DC.) O. Berg
Myrciaria vismeifolia (Benth.) O. Berg
Myrteola nummularia (Poir.) O. Berg
Plinia costata Amshoff
Plinia involucrata (O. Berg) McVaugh
Plinia pinnata L.
Plinia rivularis (Cambess.) Rotman
Plinia sp. A
Plinia sp. B
Pseudanamomis umbellulifera (Kunth) Kausel
Psidium acutangulum DC. AM BO DA
Psidium cinereum DC.
Psidium densicomum DC.
Psidium grandifolium Mart, ex DC.
Psidium guajava L. [cultivated, naturalized]
Psidium guineense Sw.
Psidium laruotteanum Cambess.
Psidium maribense DC.
Psidium salutare (Kunth) O. Berg
Psidium sartorianum (O. Berg) Nied.
Psidium striatulum DC.
Siphoneugena dussii (Krug & Urb.) Proen9a
Syzygium cumini (L.) Skeels [cultivated; naturalized?] AM BO DA
Syzygiumjambos (L.) Alston [cultivated, frequently escaping]
Syzygium malaccense (L.) Merr. & Perry [cultivated, frequently escaping] AM DA
Ugni myricoides (Kunth) O. Berg AM BO GU
Synonyms and Misidentifications
Acrandra guianensis Amshoff =Eugenia amshoffae
Aulomyrcia albidotomentosa Amshoff =A/yrc/a albidotomentosa
Aulomyrcia amshoffiana Steyerm. =Myrcia albidotomentosa
Aulomyrcia bolivarensis Steyerm. =Myrcia bolivarensis
Aulomyrcia chilensis O. Berg =Myrcia aliena
Aulomyrcia citrifolia (Aubl.) Amshoff =7l^rc/a citrifolia
Aulomyrcia divaricata O. Berg =Myrcia rufipila
AM BO
AM
AM
AM
BO
BO
AM BO
AM
AM BO
AM BO
AM BO
BO
BO
AM
BO
BO
AM BO
AM
BO
BO
AM
AM BO
AM BO
AM
AM
AM
BO
BO
AM BO
BO
AM BO
BO
BO
AM BO
BO
AM
AM BO
BO
GU
GU SU FG
GU SU
FG
FG
GU SU FG
GU SU
GU SU FG
GU SU
GU? SU FG
GU
GU
GU SU FG
GU SU FG
GU SU
GU SU
SU?
GU
GU SU FG
GU
GU
SU
GU SU FG
GU SU FG
GU
GU SU
GU SU FG
GU SU
GU SU
GU SU
GU SU FG
Checklist of the Plants of the Guiana Shield 439
Aulomyrcia dumosa O. Berg =Myrcia platyclada
Aidomyrcia ediilis O. Berg =Myrcia platyclada
Aulomyrcia grandiflora O. Berg [not in flora area]
Aulomyrcia hostmanniana O. Berg =Myrcia amazonica
Aulomyrcia inaequiloba var. paniculata (O. Berg) Amshoff =7\^rc/(3 inaequiloba
Aulomyrcia karuaiensis Steyerm. =Marlierea karuaiensis
Aulomyrcia obtusa (Schauer) O. Berg =^Myrcia guianensis var. gidanensis
Aulomyrcia paniculata O. Berg =Myrcia inaequiloba
Aulomyrcia parvifolia Steyerm. =Myrcia rotundata var. rotundata
Aulomyrcia pirarensis O. Berg =Myrcia inaequiloba
Aulomyrcia platyclada (DC.) Amshoff=Myrcia platyclada
Aulomyrcia platyclada var. kaieteurensis Amshoff =Myrcia platyclada
Aulomyrcia ptariensis Steyerm. =Myrcia ptariensis
Aulomyrcia pyrifolia (Desv. ex Ham.) O. Berg =Myrcia pyrifolia
Aulomyrcia roraimae (Oliv. ex Thum) Steyerm. =Myrcia guianensis var. guianensis
Aulomyrcia roraimensis O. Berg =Myrcia guianensis var. guianensis
Aulomyrcia rosulans O. Berg =Myrcia tomentosa
Aulomyrcia rotundata Amshoff =Myrcia rotundata
Aulomyrcia salticola Steyerm. =Myrcia salticola
Aulomyrcia schaueriana (Miq.) Amshoff =MFrc/a graciliflora
Aulomyrcia spp. =Myrcia spp.
Aulomyrcia spruceana O. Berg =Myrcia amazonica
Aulomyrcia tetramera Amshoff? =Myrcia decorticans
Aulomyrcia tomentosa (Aubl.) Amshoff =Myrcia tomentosa
Aulomyrcia triflora O. Berg =Myrcia citrifolia
Aulomyrcia vinacea Steyerm. =Myrcia multiflora
Calycampe angustifolia O. Berg =Myrcia calycampa
Calycampe latifolia O. Berg =Myrcia calycampa
Calycolpus calophyllus var. angustifolius Steyerm. =Calycolpus calophyllus
Calycolpus chnoiophyllus L. Riley =Calycolpus revolutus
Calycolpus glaber (Benth.) O. Berg =Calycolpus goetheanus
Calycolpus pyrifer L. Riley =Calycolpus revolutus
Calycorectes macrocalyx Rusby =Eugenia macrocalyx
Calycorectes yatuae McVaugh =Eugenia yatuae
Calyptranthes apoda McVaugh [nom. illeg.] =Calyptranthes vexata
Calyptranthes florifera McVaugh =Calyptranthes multiflora
Calyptranthes obtusa Benth. =Marlierea montana
Calyptranthes sericea Griseb. ^Calyptranthes fasciculata
Campomanesia coaetanea O. Berg =Campomanesia aromatica
Campomanesia glabra Benth. =Calycolpus goetheanus
Campomanesia tetramera McVaugh =Eugenia tetramera
Catinga aromatica Aubl. =Eugeniafeijoi
Catinga moschata Aubl. =Calycorectes bergii
Eugenia acetosans Poir. =Myrcia citrifolia
Eugenia adenocalyx DC. =Eugenia citrifolia
Eugenia atropunctata Steud. =Eugeniaflorida
Eugenia benthamii O. Berg =Eugenia punicifolia
Eugenia brachybotrya DC. (Melastomataceae) =Mouriri guianensis
Eugenia bracteata Rich. =Myrcia bracteata
Eugenia caurensis Steyerm. =Eugenia biflora
Eugenia comete Aubl. =Myrcia coumeta
Eugenia compta O. Berg =Eugenia pseudopsidium
Eugenia congestissima Diels =Eugenia dittocrepis
Eugenia coriacea O. Berg =Eugenia polystachyoides
Eugenia cryptadena var. gracilis Amshoff =Eugenia cowanii
Eugenia cumini (L.) Druce =Syzygium cumini
Eugenia deflexa Poir =Myrcia deflexa
Eugenia fallax Rich. =Myrcia fallax
Eugeniaferruginea Poir. =Marliereaferruginea
Eugeniaflavescens var. guianensis Sagot =Eugenia flavescens
Eugeniafloribunda West ex Willd. =Myrciariafloribunda
Eugeniafulvipes Sagot =Eugenia ramiflora
Eugenia guianensis Aubl. =Myrcia guianensis
Eugenia helinovis Sagot =Eugenia coffeifolia
440 Checklist of the Plants of the Guiana Shield
Eugenia inaeqiiiloba DC. =Myrcia inaequiloba
Eugeniajambos L. =Syzygiumjambos
Eugenia kunthiana DC. =Eugenia punicifolia
Eugenia lingua O. Berg =Eugenia cachoeirensis
Eugenia Uewelynii Steyerm. =Eugenia chrysophyllum
Eugenia malaccensis L. =Syzygium malaccense
Eugenia melinonis Sagot =Eugenia coffeifolia
Eugenia wm/ Aubl. =Eugenia biflora
Eugenia montana Aubl. =Marlierea montana
Eugenia monticola var. latifoUa Krug & Urb. =Eugenia monticola
Eugenia monticola var. racemosa Amshoff =Eugenia monticola
Eugenia multiflora Lam. =Myrcia multiflora
Eugenia nigra DC. (Melastomataceae) =Mouriri nigra
Eugenia nitida Benth. =Myrcia inaequiloba
Eugenia ochra O. Berg =Eugenia tapacumensis
Eugenia ottonis O. Berg =Eugenia pubeseens
Eugenia peregrina McVaugh =Eugenia baileyi
Eugenia prieurii Jacq. =Eugenia pseudopsidium
Eugenia producta auct. non DC. 1828: sensu McVaugh =Eugenia sp. B
Eugenia prosoneura O. Berg =Eugenia gomesiana
Eugenia protracta O. Berg [non Steud. 1843] =Eugenia protenta
Eugenia protracta Steud. =Myrciariafloribunda
Eugenia psidioides DC. =Eugenia pseudopsidium
Eugenia ptariensis Steyerm. =Eugenia anastomosans
Eugenia punicifolia var. brachypoda (DC.) Krug & Urb. =Eugenia ramiflora
Eugenia pyrifolia Desv. ex Ham. =Myrcia pyrifolia
Eugenia quitarensis Benth. =Myrcia quitarensis
Eugenia racemifera Sagot =Eugeniaflorida
Eugenia ramiflora var. montana Amshoff =Eugeniaferreiraeana
Eugenia rivularis Cambess. =Plinia rivularis
Eugenia rondonensis Steyerm. =Myrcianthes fragrans
Eugenia roraimana O. Berg =Eugenia tapacumensis
Eugenia salzmannii Benth. =Myrciariafloribunda
Eugenia schomburgkii Benth. =Eugenia lambertiana var. lambertiana
Eugenia sinemariensis Aubl. [name of uncertain application] =Eugenia sp.
Eugenia steyermarkii Standi. =Myrcianthes fragrans
Eugenia subalterna Benth. =Eugenia punicifolia
Eugenia subobliqua Benth. =Myrcia subobliqua
Eugenia tenella DC. =Myrciaria tenella
Eugenia tomentosa Aubl. =Myrcia tomentosa
Eugenia turumiquirensis Steyerm. =Myrcianthes fragrans
Eugenia vismeaefolia Benth. =Myrciaria vismeifolia
Eugenia wentii Amshoff =Eugenia macrocalyx
Gomidesia bonnetiasylvestris Steyerm. =Myrcia bonnetiasylvestris
Gomidesia lindeniana O. Berg =Myrciafenzliana
Jambosa vulgaris DC. =Syzygium jambos
Krugiaferruginea (Poir.) Urb. =Marliereaferruginea
Marlierea acuminata O. Berg =Marliereaferruginea
Marlierea dussii Krug & Urb. =Siphoneugena dussii
Marlierea gleasonii McVaugh =Myrcia decorticans
Marlierea intonsa McVaugh =Myrcia intonsa
Marlierea obtusa (Benth.) O. Berg ^Marlierea montana
Marlierea uaupensis O. Berg =Marlierea spruceana
Mitranthes sartoriana O. Berg =Psidium sartorianum
Myrcia acuminata (Kunth) DC. =Myrciafallax
Myrcia aguitensis Gleason =Myrciafallax
Myrcia berberis DC. =Myrcia fallax
Myrcia bipennis (O. Berg) McVaugh =Marlierea bipennis
Myrcia divaricata O. Berg =Myrcia rufipila
Myrcia dumosa (O. Berg) Krug & Urb. =Myrcia platyclada
Myrcia edulis (O. Berg) Krug & Urb. =Myrcia platyclada
Myrciaferruginea (Poir.) DC. =Marliereaferruginea
Myrcia grandiflora (O. Berg) Lemee [not in flora area] =Aulomyrcia grandiflora
Myrcia granulata R.O. Williams =Plinia rivularis
Checklist of the Plants of the Guiana Shield 441
Myrcia hirtellaefolia Gleason =Myrcia bracteata
Myrcia hostmanniana Kiaersk. =Myrcia amazonica
Myrcia humboldtiana DC. ^Myrcia deflexa
Myrcia kegeliana O. Berg =Myrcia fallax
Myrcia laevis O. Berg =Myrcia lucida
Myrcia leptoclada DC. =Myrcia amazonica
Myrcia lucida var. attenuata McVaugh =Myrcia lucida
Myrcia obtusa Schauer =Myrcia guianensis var. guianensis
Myrcia paniculata (Jacq.) Krug & Urb. =Myrcia citrifolia
Myrcia planifolia McVaugh =Myrcia revolutifolia
Myrcia roraimae Oliv. ex Thura =Myrcia guianensis var. guianensis
Myrcia schomburgkiana O. Berg [nom. illeg.] =Myrcia servata
Myrcia spruceana O. Berg =Myrciafallax
Myrcia subsessilis 13 ovalis O. Berg =Myrcia subsessilis var. ovalis
Myrcia subsessilis var. subcordata O. Berg =Myrcia subsessilis var. subsessilis
Myrcia tetramera Amshoff ? =Myrcia decorticans
Myrcia triflora Cambess. =Myrcia citrifolia
Myrcianthes umbelhdifera (Kunth) Alain =Pseudanamomis umbellulifera
Myrciaria bipennis O. Berg =Marlierea bipennis
Myrciaria caurensis Steyerm. =Myrciaria dubia
Myrciaria ehrenbergiana O. Berg =Myrcia ehrenbergiana
Myrciaria involucrata O. Berg =Plinia involucrata
Myrciaria lituatinervia O. Berg =Marlierea lituatinervia
Myrciaria nitida (Benth.) O. Berg =Myrcia inaequiloba
Myrciaria paraensis O. Berg ^Myrciaria dubia
Myrciaria protracta (Steud.) O. Berg =Myrciariafloribunda
Myrciaria quitarensis (Benth.) O. Berg =Myrcia quitarensis
Myrciaria uliginosa O. Berg =Myrciariafloribunda
Myrciaria verticillata O. Berg ^Myrciariafloribunda
Myrtus alternifolia Gleason =Calycolpus alternifolius
Myrtus biflora L. =Eugema biflora
Myrtus calophylla Kunth =Calycolpus calophyllus
Myrtus citrifolia Aubl. =Myrcia citrifolia
Myrtus clusiifolia Kunth =Myrcia clusiifolia
Myrtus cumini L. =Syzygium cumini
Myrtus deflexa (Poir.) Kunth =Myrcia deflexa
Myrtus emarginata Kunth =Eugenia emarginata
Myrtus erythroxyloides Kunth =Pseudanamomis umbellulifera
Myrtus fragrans Sw. =Myrcianthes fragrans
Myrtus goetheana DC. =Calycolpus goetheanus
Myrtus latifolia (O. Berg) V.M. Badillo =Myrcia calycampa
Myrtus ligustrina Sw. =Eugenia ligustrina
Myrtus longifolia Kunth (Melastomataceae) =Mouriri longifolia
Myrtus monticola Sw. =Eugenia monticola
Myrtus myricoides Kunth =Ugni myricoides
Myrtus myricoides var. roraimensis (N.E. Br.) Steyerm. =Ugni myricoides
Myrtus myricoides var. stenophylla (Oliv.) Steyerm. =Ugni myricoides
Myrtus myricoides var. turumiquirensis Steyerm. =Ugni myricoides
Myrtus nummularia Poir =Myrteola nummularia
Myrtus pubescens Kunth =Eugenia pubescens
Myrtus punicifolia Kunth ^Eugenia punicifolia
Myrtus rhopaloides Kunth '=Myrcianthes rhopaloides
Myrtus roraimensis N.E. Br. =Ugni myricoides
Myrtus salutaris Kunth =Psidium salutare
Myrtus splendens Sw. =Myrcia splendens
Myrtus stenophylla Oliv. =Ugni myricoides
Myrtus sylvatica G. Mey. =Myrcia sylvatica
Myrtus umbellata Desv. ex Ham. (Melastomataceae) =Mouriri guianensis
Myrtus umbellulifera Kunth =Pseudanamomis umbellulifera
Myrtus umbraticola Kunth =Marlierea umbraticola
Plinia dussii (Krug & Urb.) Urb. =Siphoneugena dussii
Pliniafruticosa Steyerm. =Siphoneugena dussii
Psidium aquaticum Benth. =Psidium striatulum
Psidium araca Raddi =Psidium guineense
442 Checklist of the Plants of the Guiana Shield
Psidium aromaticum Aubl. =Campomanesia aromatica
Psidium australe Cambess. =Psidium cinereum
Psidium benthamianum O. Berg =Psidium guineeme
Psidium dubium Kunth =Myrciaria dubia
Psidium glaucescens O. Berg =Psidium laruotteanum
Psidium grandiflorum Aubl. =Campomanesia grandiflora
Psidium guianense sensu Amshoff, not Pers. =Psidium acutangulum
Psidium guildingianum Griseb. =Marlierea guildingiana
Psidium guyanense Pers. =Psidium guineeme
Psidium ovatifolium O. Berg =Psidium densicomum
Psidium ovatifolium van glabrum Amshoff =Psidium densicomum
Psidium parviflorum Benth. =Psidium striatulum
Psidium persoonii McVaugh [If distinct, known from BO, GU, SU, FG] =Psidium acutangulum
Psidium quinquedentatum Amshoff =Psidium laruotteanum
Psidium sprucei O. Berg =Psidium guineense
Psidium submetrale McVaugh =Psidium dnereum
Siphoneugena densiflora auct. non O. Berg 1857: McVaugh =Siphoneugena dussii
Siphoneugena densiflora van salicifolia McVaugh =Siphoneugena dussii
Siphoneugena densiflora van salicifolia McVaugh =Siphoneugena dussii
Siphoneugena densiflora van tepuiensis Steyerm. =Siphoneugena dussii
Siphoneugena dussii van salicifolia (McVaugh) Proen^a =Siphoneugena dussii
Stenocalyx patrisii (Vahl) O. Berg =Eugenia patrisii
Ugni angustifolia Burret =Ugni myricoides
Ugni myricoides f. bifaria McVaugh =Ugni myricoides
Ugni myricoides f. grossa McVaugh =Ugni myricoides
Ugni myricoides f. oligandra McVaugh =Ugni myricoides
Ugni myricoides van roraimensis (N.E. Bn) McVaugh =Ugni myricoides
Ugni myricoides f. stenophylla (Oliv.) McVaugh =Ugni myricoides
Ugni roraimensis (N.E. Bn) Burret =Ugni myricoides
Ugni stenophylla (Oliv.) Burret =Ugni myricoides
Ugni vaccinioides Burret =Ugni myricoides
References
Amshoff, G.J.H. 1951. Myrtaceae. In A. Pulle, Flora of Suriname 3(2): 56-158.
Hoist, B.K. & M.L. Kawasaki. 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 539-551.
Hoist, B.K., L, Landrum, & F. Grifo. 2003. Myrtaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 7:1-99.
Kawasaki, M.L. & B.K. Hoist. 1994. New species and a new combination in Myrtaceae from northeastern South America.
Brittonia 46(2): 137-143.
Landrum, L.R. 1986. Campomanesia, Pimenta, Blepharocalyx, Legrandia, Acca, Myrrhinium, and Luma. Flora Neotropica
Monogn45: 1-178.
McVaugh, R. 1958. Myrtaceae (Calyptranthes and Marlierea). In B. Maguire, J.J. Wurdack, & collaborators, Botany of the
Guayana Highland part 3. Mem. New York Bot. Gard. 10(1): 1-156.
McVaugh, R. 1969. Myrtaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 8. Mem. New York Bot.
Gard. 18(2): 1-290.
Sobral, M. 1993. Sinopse de Myrciaria. Napaea 9: 13-41.
Nelumbonaceae
Nelumbo nucifera Gaertn. [cultivated, naturalized] GU SU FG?
References
Cramer, J. 1979. Nelumbonaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 390-392.
Nyctaginaceae
GA. Aymard C, R.A. DeFilippsf, J.A. Steyermarkf
Boerhavia coccinea Mill. AM? BO
Boerhavia diffusa L. AM BO DA GU SU FG
Bougainvillea buttiana Holttum & Standi. BO
Bougainvillea spectabilis Willd. BO
Checklist of the Plants of the Guiana Shield 443
AM
AM
BO
BO
BO
BO
AM BO
BO
AM
BO
AM BO
AM
BO
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM BO
BO
AM
AM BO
DA GU SU
DA GU SU FG
DA
GU
GU SU FG
DA GU SU FG
GU SU FG
Bougainvillea spp. [several species cultivated]
Guapira amaciirensis Steyerm. BO
Guapira bolivarensis Steyerm. BO
Guapira cuspidata (Heimerl) Lundell AM BO
Guapira eggersiana (Heimerl) Lundell BO
Guapiraferruginea (Klotsch ex Choisy) Lundell BO
Guapirafragrans (Dum.-Cours.) Little AM BO
Guapira glabriflora Steyerm. AM
Guapira kanukuensis (Standi.) Lundell
Guapira marcano-bertii Steyerm. BO DA
Guapira microphylla (Heimerl) Lundell BO
Guapira neblinensis Maguire & Steyerm. AM
Guapira rusbyana (Heimerl ex Standi.) Lundell BO DA
Guapira salicifolia (Heimerl) Lundell
Guapira sancarlosiana Steyerm.
Guapira sipapoana Steyerm.
Mirabilisjalapa L. [cultivated, escaping and naturalized]
Neea amaruayensis Steyerm.
Neea bernardii Steyerm.
Neea bracteosa Steyerm.
Neea brevipedunculata Steyerm.
Neea cedenensis Steyerm.
Neea clarkii Steyerm.
Neea constricta Spruce ex J.A. Schmidt GU FG
Neea davidsei Steyerm. DA
Neeafloribunda Poepp. & Endl.
Neea grandis Maguire & Steyerm.
Neea guaiquinimae Steyerm.
Neea huachamacarae Steyerm.
Neea ignicola Steyerm.
Neea liesneri Steyerm.
Neea mapourioides Steyerm.
Neea marahuacae Steyerm.
Neea mollis Spruce ex J.A. Schmidt GU FG
Neea neblinensis Maguire & Steyerm.
Neea obovata Spruce ex Heimerl
Neea ovalifolia Spruce ex J.A. Schmidt
Neea parimensis Steyerm.
Neea robusta Steyerm.
Neea sebastianii Steyerm.
Neea spruceana Heimerl GU FG
Neea subglabrata Steyerm. GU
Neea tepuiensis Steyerm.
Neea tristis Heimerl
Pisonia aculeata L.
Pisonia macranthocarpa (Donn. Sm.) Donn. Sm. GU
Synonyms and Misidentifications
Boerhavia caribaea Jacq. =Boerhavia coccinea
Boerhavia paniculata Rich. ^Boerhavia diffusa
Boerhavia surinamensis Miq. =Boerhavia diffusa
Guapira ayacuchae Steyerm. =Guapira cuspidata
Guapira davidsei Steyerm. =Guapiraferruginea
Guapira glabra (Heimerl) Steyerm. =Guapira eggersiana
Guapira graciliflora sensu Mennega et al. (1988), non (Mart.) Lundell =Guapira salicifolia
Guapira guianensis Aubl. ? =Guapira eggersiana
Guapira heimerliana (Standi.) Lundell =Guapira salicifolia
Guapira olfersiana (Link, Klotzsch & Otto) Lundell [not in flora area]
Guapira pacurero (Kunth) Little [not in flora area]
Guapira riedeliana (Fisch. ex Heimerl) Lundell [not in flora area]
Guapira schomburgkiana (Heimerl) Lundell =Guapira cuspidata
Neea cauliflora Heimerl =Neeafloribunda
Neea nigricans (Sw.) Heimerl [not in flora area]
Pisonia aculeata var. macranthocarpa Donn. Sm. =Pisonia macranthocarpa
GU SU FG
444 Checklist of the Plants of the Guiana Shield
Pisonia albiflora Heimerl =Guapira salicifolia
Pisonia cuspidata Heimerl =Giiapira cuspidata
Pisonia eggersiana Heimerl =Guapira eggersiana
Pisonia eucalyptifolia Kunth ^Gnapirafragrans
Pisoniaferruginea Klotzsch ex Choisy =Guapiraferruginea
Pisoniafragrans Dum.-Cours. =Guapirafragrans
Pisonia glabra Heimerl =Guapira eggersiana
Pisonia guianensis (Aubl.) R.A. Howard ? =Guapira eggersiana
Pisonia inermis Jacq. [not in flora area] =Guapirafragrans
Pisonia microphylla Heimerl =Guapira microphylla
Pisonia nitida Mart, ex J.A. Schmidt =Guapira eggersiana
Pisonia olfersiana sensu Maguire (1948), non Link, Klotzsch & Otto =Guapira eggersiana
Pisonia pacurero sensu Pulle (1906), non Kunth =Guapira eggersiana
Pisonia salicifolia Heimerl =Guapira salicifolia
Pisonia schomburgkiana Heimerl =Guapira cuspidata
Torrubia cepalantha Standi, ex Pittier =Guapira ferruginea
Torrubia cuspidata (Heimerl) Standi. =Guapira cuspidata
Torrubiaferruginea (Klotsch ex Choisy) Standi. =Guapiraferruginea
Torrubiafragrans (Dum.-Cours.) Standi. =Guapirafragrans
Torrubia heimerliana Standi. =Guapira salicifolia
Torrubia kanukiiensis Standi. ^Guapira kanukuensis
Torrubia laurina Standi. =Guapira eggersiana
Torrubia microphylla (Heimerl) Standi. =Guapira microphylla
Torrubia olfersiana (Link, Klotzsch & Otto) Standi, [not in flora area]
Torrubia rusbyana Heimerl ex Standi. =Guapira rusbyana
Torrubia salicifolia (Heimerl) Standi. =Guapira salicifolia
Torrubia schomburgkiana (Heimerl) Standi. =Guapira cuspidata
References
DeFilipps, R.A. & S.L. Maina. 2002. Nyctaginaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M.
Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New
YorkBot. Gard. 76 (2): 551-553.
DeFilipps, R.A. & S.L. Maina. 2003. Phytolaccaceae, (+ remarks on Achatocarpaceae), Nyctaginaceae, Aizoaceae,
Chenopodiaceae, Amaranthaceae, Portulacaceae, Basellaceae, Molluginaceae, Caryophyllaceae. In M.J. Jansen-Jacobs, ed..
Flora of the Guianas, Fascicle 22 (series A). Royal Botanic Gardens, Kew, U.K.
Steyermark, J.A. 1987. Nyctaginaceae. In Flora of the Venezuelan Guayana - III. Ann. Missouri Bot. Gard. 74(3): 614-635.
Standley, P.C. 1931. The Nyctaginaceae of northwestern South America. Field Mus. Bot. 11: 73-114.
Steyermark J.A. & GA. Aymard C. 2003. Nyctaginaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of
the Venezuelan Guayana 7: 101-118.
Nymphaeaceae
J.H. Wiersema
Nymphaea amazonum Mart. & Zucc.
Nymphaea amazonum subsp. amazonum
Nymphaea ampla (Salisb.) DC.
Nymphaea conardii Wiersema
Nymphaea gardneriana Planch.
Nymphaea glandulifera Rodschied
Nymphaeajamesoniana Planch.
Nymphaea lingulata Wiersema
Nymphaea lotus L. [cultivated, naturalized]
Nymphaea odorata Alton
Nymphaea potamophila Wiersema AM
Nymphaea pulchella DC.
Nymphaea rubra Roxb. ex Salisb. [cuhivated, naturalized]
Nymphaea rudgeana G. Mey.
Nymphaea tenerinervia Casp.
Victoria amazonica (Poepp.) Sowerby
Synonyms and Misidentifications
Castalia blanda (G Mey.) G. Lawson ^Nymphaea glandulifera
Castalia spp. =Nymphaea spp.
Leuconymphaea amazonum (Mart. & Zucc.) Kuntze =Nymphaea amazonum
DA GU SU FG
DA GU
GU
SU
SU
FG
BO GU
BO GU
BO DA GU
GU
GU
GU
GU
SU
SU
FG
BO DA GU
DA GU
GU
SU
SU
FG
BO DA GU
GU?
GU
SU FG
Checklist of the Plants of the Guiana Shield 445
Leuconymphaea blanda (G. Mey.) Kuntze ^Nymphaea glandulifera
Nymphaea ampla auct. non (Salisb.) DC: sensu South American authors =Nymphaea pulchella
Nymphaea blanda G. Mey. [nom. superfl.] =Nymphaea glandulifera
Nymphaea blanda var. amazonum (Mart. & Zucc.) Planch. =Nymphaea amazonum
Nymphaea odorata var. gigantea Tricker =Nymphaea odorata
Nymphaea parkeriana Lehm. =Nymphaea odorata
Nymphaea rudgeana var. amazonum (Mart. & Zucc.) Griseb. =Nymphaea amazonum
Victoria regia Lindl. ^Victoria amazonica
References
(see also Cabombaceae, Ceratophyllaceae, Nelumbonaceae)
Conard, H.S. 1905. The Waterlilies: A Monograph of the Genus Nymphaea. Carnegie Institution of Washington.
Cramer, J. 1979. Nymphaeaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 370-384.
Mori, S.A. 2002. Nymphaeaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
554.
Wiersema, J.H. 1987. A monograph ofNymphaea subgenus Hydrocallis. Syst. Bot. Monogr. 16: 1-112.
Wiersema, J.H. 2003. Nymphaeaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana7: 118-124.
OCHNACEAE
C. Sastre
Adenanthe bicarpellata Maguire, Steyerm. & Wurdack
Adenanthe ciliata Sastre
Adenarake macrocarpa Sastre
Adenarake muriculata Maguire & Wurdack
Blastemanthus gemmiflorus (Mart.) Planch.
Blastemanthus gemmiflorus subsp. gemmiflorus
Blastemanthus gemmiflorus subsp. sprucei (Tiegh.) Sastre
Cespedesia spathulata (Ruiz & Pav.) Planch.
Elvasia brevipedicellata Ule
Elvasia calophyllea DC.
Elvasia canescens (Tiegh.) Gilg
Elvasia elvasioides (Planch.) Gilg
Elvasia essequibensis Engl.
Elvasia macrostipularis Sastre & Lescure
Elvasia quinqueloba Spruce ex Engl.
Elvasia sphaerocarpa R.S. Cowan
Ouratea acuminata (DC.) Engl.
Ouratea angulata Tiegh.
Ouratea aquatica (Kunth) Engl.
Ouratea arbobrevicalyx Sastre
Ouratea articulata Sastre
Ouratea asisae Maguire & Steyerm.
Ouratea attenuata Tiegh.
Ouratea bipartita Sastre
Ouratea brevicalyx Maguire & Steyerm.
Ouratea brevipedicellata Maguire & Steyerm.
Ouratea candollei (Planch.) Tiegh.
Ouratea cardiosperma (Lam.) Engl.
Ouratea castaneifolia (DC.) Engl.
Ouratea cataractarum Sandwith
Ouratea cerebroidea Sastre
Ouratea cernuiflora Sandwith
Ouratea chaffanjonii (Tiegh.) Sastre
Ouratea clarkii Sastre
Ouratea coccinea Mart, ex Engl.
Ouratea croizatii Maguire & Steyerm.
Ouratea culminicola Maguire & Steyerm.
Ouratea davidsii Sastre
Ouratea decagyna Maguire
Ouratea deminuta Maguire & Steyerm.
Ouratea discophora Ducke
BO GU
GU
AM
AM GU
AM
A A/t
GU
AM BO DA GU
BO GU
A A/t
BO? GU?
AM
AM
BO GU
GU
GU
GU
GU?
su
su
AM BO DA GU? su
BO
AM
AM
AM
AM
AM
AM
AM BO DA
AM
A \A
BO
AM
A \A
BO
BO
AM
AM BO
FG
FG
FG
GU
GU
su FG
GU? su FG
GU su
GU
GU FG
GU
GU? SU FG
446 Checklist of the Plants of the Guiana Shield
Ouratea duidae Steyerm.
Ouratea engleri Tiegh.
Ouratea erecta Sastre
Ouratea evoluta Maguire & Steyerm.
Ourateafasciculata Maguire & Steyerm.
Ourateaferruginea Engl.
Ourateafrancinae Sastre
Ourateafrontium Sleumer
Ourateafusiformis Sastre
Ouratea gigantophylla (Ehrh.) Engl.
Ouratea gillyana (Dwyer) Sandwith & Maguire
Ouratea grandiflora (DC.) Engl.
Ouratea grosourdyi (Tiegh.) Steyerm.
Ouratea guaiquinimensis Sastre
Ouratea guianensis Aubl.
Ouratea guildingii (Planch.) Urb.
Ouratea guriensis Sastre
Ouratea heterobracteata Sastre
Ouratea huberi Maguire & Steyerm.
Ouratea impressa (Tiegh.) Lemee
Ouratea kamikuensis Sastre
Ouratea lajaensis Sastre
Ouratea leblondii (Tiegh.) Lemee
Ouratea leprieurii Tiegh.
Ouratea liesneri Sastre
Ouratea longifolia (Lam.) Engl.
Ouratea longistyla Maguire & Steyerm.
Ouratea maasorum Sastre
Ouratea macrocarpa Sastre
Ouratea maguirei Sastre
Ouratea maigualidae Sastre
Ouratea marahuacensis Maguire & Steyerm.
Ouratea mazaruniensis A.C. Sm. & Dwyer
Ouratea medinae Maguire & Steyerm.
Ouratea megaphylla Sastre
Ouratea melinonii (Tiegh.) Lemee
Ouratea microcalyx (Engl.) Sastre
Ouratea multibracteata Steyerm. ex Sastre
Ouratea obovata Sastre
Ouratea occultinervis Sastre
Ouratea oligantha Steyerm. ex Sastre
Ouratea olivaeformis (A. St.-Hil.) Engl, [unverified name]
Ouratea orisina Sastre
Ouratea ornata Maguire & Steyerm.
Ouratea papillata Maguire & Steyerm.
Ouratea paratatei Sastre
Ouratea paruensis Maguire & Steyerm.
Ouratea pendula (Poepp.) Engl.
Ouratea pisiform is Engl.
Ouratea polyantha (Triana & Planch.) Engl.
Ouratea pseudoguildingii Sastre
Ouratea pseudotatei Maguire & Steyerm.
Ouratea ptaritepuiensis Steyerm.
Ouratea pulverulenta Sastre
Ouratea ramosissima Maguire & Steyerm.
Ouratea rigida Engl.
Ouratea riparia Sleumer
Ouratea roraimae Engl.
Ouratea rorida Sastre
Ouratea rotundipetala Maguire & Steyerm.
Ouratea rupununiensis Klotzsch ex Engl.
Ouratea saldariagae Sastre
Ouratea sastrei Zarucchi, nom. nov. ined.
Ouratea saiilensis Sastre
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
DA
BO DA
BO
BO
BO
AM BO
AM BO
BO
BO
AM
AM BO
BO
AM BO
AM BO
AM
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
AM
GU
GU
GU
GU
GU
GU
FG
SU FG
FG
FG
SU
SU FG
FG
SU FG
FG
FG
SU FG
SU FG
SU FG
SU FG
SU FG
FG
GU su
GU su FG
GU su FG
GU su FG
GU
GU su?
Checklist of the Plants of the Guiana Shield 447
Oiiratea schomburgkii (Planch.) Engl.
Ouratea scottii Sastre
Ouratea scottii subsp. scottii
Ouratea sculpta (Tiegh.) Sastre
Ouratea sipapoensis Maguire & Steyerm. AM
Ouratea soderstromii Sastre BO
Ouratea spruceana Engl. AM BO
Ouratea squamata Sastre BO
Ouratea steyermarkii Sastre AM
Ouratea subamplexicaulis Maguire & Steyerm. BO
Ouratea superba Engl AM BO GU SU
Ouratea tatei Gleason BO GU
Ouratea thyrsoidea Engl. AM
Ouratea timehriensis Sastre GU
Ouratea vaccinioides (A. St.-Hil. & Tul.) Engl. GU?
Ouratea verruculosa Engl. FG
Ouratea yapacanae Sastre
Perissocarpa steyermarkii (Maguire) Steyerm. & Maguire
Perissocarpa umbellifera Steyerm. & Maguire
Philacra auriculata Dwyer
Philacra duidae (Gleason) Dwyer
Philacra longifolia (Gleason) Dwyer
Philacra steyermarkii Maguire
Poecilandra pumila Steyerm.
Poecilandra retusa Tul.
Poecilandra retusa var. retusa
Poecilandra retusa var. sclerophylla (Ule) Sastre
Sauvagesia aliciae Sastre
Sauvagesia aliciae subsp. aliciae
Sauvagesia aliciae subsp. aratayensis Sastre
Sauvagesia amoena Ule
Sauvagesia angustifolia Ule
Sauvagesia deflexifolia Gardner
Sauvagesia elata Benth.
Sauvagesia erecta L.
Sauvagesia erecta var. erecta
Sauvagesia erioclada Maguire & K.D. Phelps
Sauvagesiafalcisepala Sastre
Sauvagesiafruticosa Mart.
Sauvagesia guianensis (Eichler) Sastre
Sauvagesia guianensis subsp. guianensis
Sauvagesia imthurniana (Oliv.) Dwyer
Sauvagesia linearifolia A. St.-Hil.
Sauvagesia linearifolia subsp. venezuelensis Maguire & Wurdack
Sauvagesia longifolia Eichler
Sauvagesia longipes Steyerm.
Sauvagesia nudicaulis Maguire & Wurdack
Sauvagesia pulchella Planch.
Sauvagesia ramosa (Gleason) Sastre
Sauvagesia ramosissima Spruce ex Eichler
Sauvagesia roraimensis Ule
Sauvagesia rubiginosa A. St.-Hil.
Sauvagesia sprengelii A. St.-Hil.
Sauvagesia tafelbergensis Sastre
Sauvagesia tenella Lam.
Tyleria apiculata Sastre
Tyleria aristata Maguire & Wurdack
Tyleria breweriana Steyerm.
Tyleriafloribunda Gleason
Tyleria grandiflora Gleason
Tyleria linearis Gleason
Tyleria pendula Maguire & Wurdack
Tyleria phelpsianaMdLguwQ &.S>iQyQvm. BO GU
Tyleria silvana Maguire AM
AM
AM GU
AM BO
AM
AM
AM
BO
BO GU
AM BO GU
AM BO
BO
GU
AM FG
AM
BO GU?
FG
BO GU SU
AM
AM DA GU SU FG
AM BO DA GU su FG
AM BO DA GU su FG
AM
AM
AM BO
AM BO GU
AM BO GU
AM BO GU
AM
AM
AM BO
BO
GU
GU
FG
AM
GU su
AM BO
AM BO GU su FG
AM BO GU su
AM BO GU su FG
BO GU su
su
FG
FG
AM BO GU su FG
AM BO
AM
BO
AM BO
AM
AM
AM
448 Checklist of the Plants of the Guiana Shield
Tyleria spathulata Gleason AM
Tyleria spectabilis Maguire & Wurdack AM
Tyleria terrae-humilis Sastre AM
Tyleria tremuloidea Maguire & Wurdack AM
Wallacea insignis Spruce ex Benth. & Hook. f. AM
Wallacea multiflora Ducke AM
Synonyms and Misidentifications
Blastemanthus densiflorus Hallier =Blastemanthus gemmiflorus subsp. sprucei
Blastemanthus paniculatus Tiegh. =Blastemanthus gemmiflorus subsp. gemmiflorus
Blastemanthus sprucei Tiegh. =Blastemanthus gemmiflorus subsp. sprucei
Camptouratea leblondii Tiegh. =Ouratea leblondii
Camptouratea sagotii Tiegh. =Ouratea leblondii
Cercouratea chaffanjonii Tiegh. =Ouratea chaffanjonii
Cercourateaferruginea (Engl.) Tiegh. =Ourateaferruginea
Cercouratea grosourdyi Tiegh. =Ouratea grosourdyi
Cercouratea laxa Tiegh. =Ouratea superba
Cespedesia amazonica Huber =Cespedesia spathulata
Cespedesia bonplandii Goudot =Cespedesia spathulata
Cespedesia sprucei Tiegh. =Cespedesia spathulata
Elvasia caurensis Pittier =Elvasia elvasioides
Elvasia hostmannia Planch. =Elvasia elvasioides
Elvasia sagotii (Tiegh.) Lemee ^'Elvasia elvasioides
Elvasia schomburgkii Tiegh. =Elvasia calophyllea
Godoya gemmiflora Mart. =Blastemanthus gemmiflorus
Godoya spathulata Ruiz & Pav. ^Cespedesia spathulata
Gomphia aquatica Kunth =Ouratea aquatica
Gomphia candollei Planch. =Ouratea candollei
Gomphia castaneifolia DC. =Ouratea castaneifolia
Gomphia grandiflora DC. =Ouratea grandiflora
Gomphia polyantha Triana & Planch. =Ouratea polyantha
Gomphia surinamensis Planch. =Ouratea cardiosperma
Hostmannia elvasioides Planch. =Elvasia elvasioides
Hostmannia sagotii Tiegh. =Elvasia elvasioides
Kaieteurea gillyana Dwyer =Ouratea gillyana
Leitgebia colombiana R. Schult. =Sauvagesiafruticosa
Leitgebia gleasoniana Lasser =Sauvagesia guianensis subsp. guianensis
Leitgebia guianensis Eichler =Sauvagesia guianensis
Leitgebia imthurniana Oliv. =Sauvagesia imthurniana
Luxemburgia duidae Gleason =Philacra duidae
Luxemburgia longifolia Gleason =Philacra longifolia
Ouratea ayacuchae Maguire & Steyerm. =Ouratea chaffanjonii
Ouratea coccinea auct. non Engl. 1876: sensu Maguire & Steyerm. =Ourateafusiformis p.p., Ouratea maguirei p.p.
Ouratea crenata Spruce ex Tiegh. =Ouratea pisiform is
Ouratea discophora subsp. pervenulosa Maguire & Steyerm. =Ouratea discophora
Ouratea elongata Sastre [non O. elongata (Oliv.) Engl.] =Ouratea sastrei
Ouratea guianensis auct. non Aubl. 1775: sensu Maguire & Steyerm. ^Ouratea rorida p.p.
Ouratea inconformis Maguire & Steyerm. =Ouratea superba
Ouratea integrifolia (DC.) Sagot [nomen nudum]
Ouratea inundata Spruce ex Engl. =Ouratea superba
Ouratea longifolia auct. non (Lam.) Engl.: sensu Maguire & Steyerm. =Ouratea guianensis
Ouratea macurucoensis Maguire & Steyerm. =Ouratea steyermarkii
Ouratea odora Poepp. ex Engl. =Ouratea superba
Ouratea pittieri Sleumer =Ouratea grosourdyi
Ouratea polygyna Engl, [not in flora area]
Ouratea propingua Maguire & Steyerm. =Ouratea ptaritepuiensis
Ouratea piimila Maguire & Steyerm. =Ouratea aquatica
Ouratea pumila var. heterodoxa Maguire & Steyerm. =Ouratea aquatica
Ouratea racemiformis Ule =Ouratea schomburgkii
Ouratea sagotii (Tiegh.) Lemee =Ouratea leblondii
Ouratea surinamensis (Planch.) Wehlb. =Ouratea cardiosperma
Ouratea vasivae Spruce ex Engl. =Ouratea grandiflora
Ouratea venezuelensis Steyerm. =Ouratea thyrsoidea
Ouratea wurdackiana Maguire & Steyerm. =Ouratea superba
Pentaspatella ramosa Gleason =Sauvagesia ramosa
Checklist of the Plants of the Guiana Shield 449
Poecilandra sclerophylla Ule =Poecilandra retusa var. sclerophylla
Roraimanthus imthurniamis (Oliv.) Gleason =Sauvagesia imthurniana
Roraimanthus imthiirnianus (Oliv.) Gleason =Sauvagesia imthurniana
Sauvagesia defwiens A.C. Sm. ^Sauvagesia roraimensis
Sauvagesia diickei Sleumer =Sauvagesia ramosa
Sauvagesia duidae auct. non Steyerm. 1952: sensu Steyerm. =Sauvagesia aliciae
Sauvagesia duidae Steyerm. =Sauvagesia erioclada
Sauvagesia elata subsp. occidentalis Maguire & Wurdack =Sauvagesia elata
Sauvagesia erecta Aubl. [non L.] ^Sauvagesia sprengelii
Sauvagesia erecta var. rubiginosa (A. St.-Hil.) Eichler =Sauvagesia rubiginosa
Sauvagesia erioclada var. grandiflora Maguire & Wurdack =Sauvagesia erioclada
Sauvagesia guianensis subsp. gleasoniana (Lasser) Steyerm. =Sauvagesia guianensis subsp. guianensis
Sauvagesia guianensis subsp. guaiquinimensis Steyerm. =Sauvagesia guianensis subsp. guianensis
Sauvagesia guianensis subsp. sipapoensis Steyerm. =Sauvagesia guianensis subsp. guianensis
Sauvagesia imthurniana subsp. chimantensis Maguire, Steyerm. & Wurdack =Sauvagesia imthurniana
Sauvagesia inconspicua Dwyer =Sauvagesia tenella
Sauvagesia kappleri Miq. =Sauvagesia sprengelii
Sauvagesia laxa Mart. =Sauvagesia rubiginosa
Sauvagesia marahuacensis Steyerm. =Sauvagesia guianensis subsp. guianensis
Sauvagesia miniata Steyerm. =Sauvagesia ramosissima
Sauvagesia rosacea Ule =Sauvagesiafruticosa
Sauvagesia salicifolia Ging. =Sauvagesia erecta
Sauvagesia serpyllifolia Mart. =Sauvagesia sprengelii
Sauvagesia smithiana Dwyer =Sauvagesia roraimensis
Sauvagesia striata Gleason & Dwyer =Sauvagesia angustifolia
Sauvagesia surinamensis Miq. =Sauvagesia rubiginosa
Trichovaselia canescens Tiegh. =Elvasia canescens
Vaselia quinqueloba (Spruce ex Engl.) Tiegh. =Elvasia quinqueloba
Villouratea spiciformis Tiegh. =Ouratea rigida
References
Allorge, L. & C. Sastre. 1990. Distribution geographique dans les Guyanes, d'Apocynaceae et d'Ochnaceae. Essai de mise en
evidence de centres de speciation. C.R. Soc. Biogeogr. 67(2): 77-94.
Dwyer, J.D. 1943. The taxa of the monogeneric tribe Elvasieae (Ochnaceae). Bull. Torrey Bot. Club 70: 42-49.
Dwyer, J.D. 1945. The taxonomy of the genus Sauvagesia (Ochnaceae). Bull. Torrey Bot. Club 72(6): 521-540.
Maguire, B., & J.J. Wurdack. 1961. Ochnaceae. In Botany of the Guayana Highland part IV. Mem. New York Bot. Gard. 10(4):
6-21.
Maguire, B. & J.A. Steyermark. 1989. Ouratea in Guayana and adjacent Amazonian hylaea. In B. Maguire & collaborators,
Botany of the Guayana Highland part XIII. Mem. New York Bot. Gard. 51 : 56-102.
Sastre, C. 1971. Distribution geographique des especes de Sauvagesia L. C.R. Soc. Biogeogr. 421: 47-59.
Sastre, C. 1984. Etude sur le genre Sauvagesia L. en Guyane fran9aise. Saussurea 15: 111-118.
Sastre, C. 1986. Studies on the Flora of the Guianas 23. Cinq especes nouvelles d'Ouratea Aublet (Ochnaceae) des Guyanes.
Bull. Mus. Nat. Hist. ser. 4, 8, sect. B, Adansonia 4: 413-422.
Sastre, C. 1987. Studies on the Flora of the Guianas 30. Considerations phytogeographiques sur les Ochnacees guyanaises. C.R.
Soc. Biog. 63(3): 89-97.
Sastre, C. 1988. Synopsis generis Ouratea Aubl. Bull. Mus. natn. Hist. nat. Paris, ser. 4, 10, sect. B, Adansonia 1: 47-67.
Sastre, C. 1991. Duae novae species Ourateae (Ochnaceae) inGuianiscollectae. Studies on the Flora of the Guianas. 59. CandoUea
46: 395-398.
Sastre, C. 1994. Three new taxa of Ouratea (Ochnaceae) from South America. Brittonia 46(4): 309-313.
Sastre, C. 1995. Novelties in the Neotropical genus Ouratea Aublet (Ochnaceae). Novon 5(2): 193-200.
Sastre, C. 1998. Novelties in Sauvagesioideae (Ochnaceae) from Venezuela and Guyana. Novon 8(3): 301-303.
Sastre, C. 2001. New Ouratea species (Ochnaceae) from Venezuela and adjacent countries. Novon 1 1(1): 105-1 18.
Sastre, C. 2001. Ochnaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 7:
124-161.
Sastre, C. 2002. Ochnaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 554-
559.
Wehlburg, C. 1 937. Ochnaceae. In A. Pulle, Flora of Suriname 3( I ): 328-34 1
.
Olacaceae
p. Hiepko, J.M. MacDougal
Aptandra liriosmoides Spruce ex Miers AM GU
Aptandra tubicina (Poepp.) Benth. ex Miers BO
450 Checklist of the Plants of the Guiana Shield
Brachynema axillare R. Duno & RE. Berry
Cathedra acuminata (Benth.) Miers
Chaunochiton angustifolium Sleumer
Chaunochiton kappleri (Sagot ex Engl.) Ducke
Chaunochiton loranthoides Benth.
Dulacia Candida (Poepp.) Kuntze
Dulacia crassa (Monach.) Sleumer [expected in BO]
Dulacia cyanocarpa Sleumer
Dulacia guianensis (Engl.) Kuntze
Dulacia inopiflora (Miers) Kuntze
Dulacia macwphylla (Benth.) Kuntze
Dulacia redmondii Steyerm.
Dulacia tepuiensis (Steyerm.) Sleumer & Steyerm.
Heisteria acuminata (Bonpl.) Engl.
Heisteria barbata Cuatrec.
Heisteria cauliflora Sm.
Heisteria densifrons Engl.
Heisteria duckei Sleumer
Heisteria insculpta Sleumer
Heisteria maguirei Sleumer
Heisteria maytenoides Spruce ex Engl.
Heisteria ovata Benth.
Heisteria pentandra (Benth. ex Reissek) Engl.
Heisteria scandens Ducke
Heisteria spruceana Engl.
Maburea trinervis Maas
Minquartia guianensis Aubl.
Ptychopetalum olacoides Benth.
Schoepfia brasiliensis A. DC.
Schoepfia clarkii Steyerm.
Schoepfia lucida Pulle
Schoepfia tepuiensis Steyerm.
Ximenia americana L.
Ximenia americana var. americana
Synonyms and Misidentifications
Aptandra benthamiana Miers =Aptandra liriosmoides
Aptandra spruceana Miers =Aptandra tubicina
Cathedra caurensis Pittier =Cathedra acuminata
Cathedra crassifolia Benth. ex Miers =Cathedra acuminata
Cathedra guianensis Sandwith =Cathedra acuminata
Chaunochiton mouriroides A.C. Sm. ^Aptandra tubicina
Chaunochiton tubicinum (Poepp.) Lemee =Aptandra tubicina
Diplocrater acuminatus Benth. =Cathedra acuminata
Dulacia adhaerens var. stenopoda Cuatrec. =Dulacia Candida
Heisteria acuta Engl. =Heisteria pentandra
Heisteria coccinea Jacq. [not in flora area]
Heisteriafatoensis Standi. =Heisteria scandens
Heisteriaflexuosa auct. non Engl. 1892: sensu V.M. Badillo =Heisteria pentandra
Heisteriaflexuosa Engl. =Heisteria ovata
Heisteria guianensis V.M. Badillo =Heisteria acuminata
Heisteria kappleri Sagot ex Engl. =Chaunochiton kappleri
Heisteria longifolia Spruce ex Engl. (Ebenaceae) -Diospyros tenuiflora
Heisteria longipes Standi. =Heisteria acuminata
Heisteria microcalyx Sagot =Heisteria densifrons
Heisteria microcarpa Spruce ex Engl. =Heisteria densifrons
Heisteria nitida auct. non Spruce ex Engl.: sensu V.M. Badillo =Heisteria ovata
Heisteria surinamensis Amshoff =Heisteria ovata
Heisteria tubicina Poepp. =Aptandra tubicina
Heymassoli inermis Aubl. =Ximenia americana
Heymassoli spinosa Aubl. =Ximenia americana
Liriosma Candida Poepp. =Dulacia Candida
Liriosma crassa Monach. =Dulacia crassa
Liriosma guianensis Engl. "^Dulacia guianensis
Liriosma inopiflora Miers [not in flora area] =Dulacia inopiflora
AM
AM BO
AM BO
AM
AM BO
AM
AM BO
AM
AM
AM
BO
AM BO
AM
AM
AM
AM
BO
AM
AM BO
AM
AM BO
AM
AM BO
BO
AM
AM
BO
AM BO
AM BO
DA GU FG
GU SU FG
GU
GU SU FG
GU
DA
DA
FG
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
GU
GU
SU
SU
SU
FG
FG
Checklist of the Plants of the Guiana Shield 451
Liriosma macrophylla (Benth.) Roem. =Dulacia macrophylla
Liriosma micrantha Spruce ex Engl, [nomen nudum] =Dulacia inopiflora
Liriosma tepuiensis Steyeim. =Dulacia tepiiiensis
Olax macrophylla Benth. =Dulacia macrophylla
Olax schomburgkii Klotzsch [nomen nudum] =Dulacia guianensis
Rhaptostylum acuminatum Bonpl. =Heisteria acuminata
Rhaptostylum pentandrum Benth. ex Reissek =Heisteria pentandra
Secretania loranthacea Miill. Arg. =Minquartia guianensis
References
Amshoff, G.J.H. 1938. Olacaceae. In A. Pulle, Flora of Suriname 1(1): 262-272.
Hiepko, R 1993. Olacaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 14: 3-35.
Hiepko, R 2002. Olacaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 559-
562.
MacDougal, J.M. 2003. Olacaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana7: 162-186.
Sleumer, H.O. 1984. Olacaceae. Flora Neotropica Monogr. 38: 1-159.
Oleaceae
J.S. Miller, G.A. Aymard C.
Chionanthus compactus Sw. AM BO
Chionanthus guianensis (Aubl.) Pers. FG
Chionanthus implicatus (Rusby) RS. Green AM
Jasminum spp. [cultivated, perhaps escaping]
Synonyms and Misidentifications
Chionanthus caribea Jacq. =Chionanthus compactus
Chionanthus guianensis (Aubl.) Steam [comb, superfl.]
Chionanthus mayepea Vahl =Chionanthus guianensis
Chionanthus tetrandra Vahl =Chionanthus guianensis
Linociera caribea (Jacq.) Knobl. =Chionanthus compactus
Linociera tetrandra (Vahl) Roem. & Schult. =Chionanthus guianensis
Mayapea implicata Rusby =Chionanthus implicatus
Mayepea caribea (Jacq.) Kuntze =Chionanthus compactus
Mayepea guianensis Aubl. =Chionanthus guianensis
References
Green, RS. 1994. A revision of Chionanthus in S. America and the description of Priogymnanthus, gen. nov. Kew Bull. 49(2):
261-286.
Miller, J.S. & G.A. Aymard C. 2003. Oleaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 7: 186-187.
Steam, W.T. 1976. Union of Chionanthus and Linociera. Ann. Missouri Bot. Gard. 63: 355-357.
Onagraceae
RH. Raven, E.M. Zardini
Ludwigia affinis (DC.) H. Hara
Ludwigia decurrens Walter
Ludwigia densiflora (Micheli) H. Hara
Ludwigia dodecandra (DC.) Zardini & RH. Raven
Ludwigia elegans (Cambess.) H. Hara
Ludwigia erecta (L.) H. Hara
Ludwigiafoliobracteolata (Munz) H. Hara
Ludwigia helminthorrhiza (Mart.) H. Hara
Ludwigia hyssopifolia (G. Don) Exell
Ludwigia inclinata (L. f ) M. Gomez
Ludwigia latifolia (Benth.) H. Hara
Ludwigia leptocarpa (Nutt.) H. Hara
Ludwigia mexiae (Munz) H. Hara
Ludwigia nervosa (Poir.) H. Hara
Ludwigia octovalvis (Jacq.) RH. Raven
Ludwigia peruviana (L.) H. Hara
AM BO DA GU SU FG
AM BO
BO
DA
DA
GU
GU
su
SU
FG
FG
AM BO DA GU su FG
BO DA GU FG
BO DA GU su
AM BO DA GU su FG
AM BO GU su
AM BO DA GU su FG
AM BO DA GU su
su
FG
AM BO GU su FG
AM BO
BO
DA GU su FG
452 Checklist of the Plants of the Guiana Shield
BO GU
BO GU
BO DA GU
GU
BO DA GU
Ludwigia quadrangularis (Micheli) H. Hara
Iwfifwzg/a r/g/V/o (Miq.) Sandwith AM SU
Ludwigia sedoides (Bonpl.) H. Hara
Ludwigia sericea (Cambess.) H. Hara
Ludwigia torulosa {Am.) Yi.WdiXSi SU FG
Synonyms and Misidentifications
Jussiaea affinis DC. =Ludwigia affinis
Jussiaea affinis var. dodecandra (DC.) Munz =Ludwigia dodecandra
Jussiaea affinis var. midtifiora Jonker =Ludwigia affinis
Jussiaea aluligera Miq. =Ludwigia leptocarpa
Jussiaea decurrens (Walter) DC. =Ludwigia decurrens
Jussiaea densiflora Micheli =Ludwigia densifiora
Jussiaea dodecandra DC. =Ludwigia dodecandra
Jussiaea elegans Cambess. =Ludwigia elegans
Jussiaea erecta L. =Ludwigia erecta
Jussiaea erecta var. sebana DC. =Ludwigia erecta
Jussiaeafoliobracteolata Munz =Ludwigia foliobracteolata
Jussiaea helminthorrhiza Mart. =Ludwigia helminthorrhiza
Jussiaea hexamera Miq. =Ludwigia affinis
Jussiaea hyssopifolia G. Don =Ludwigia hyssopifolia
Jussiaea incUnata L. f. ^Ludwigia inclinata
Jussiaea latifolia Benth. =Ludwigia latifolia
Jussiaea leptocarpa Nutt. =Ludwigia leptocarpa
Jussiaea leptocarpa var. aluligera (Miq.) Jonker =Ludwigia leptocarpa
Jussiaea leptocarpa var. gemiina Munz =Ludwigia leptocarpa
Jussiaea leptocarpa var. meyeriana (Kuntze) Munz =Ludwigia leptocarpa
Jussiaea linifolia auct. non Vahl: sensu Hutch. & Dalziel =Ludwigia erecta
Jussiaea linifolia Vahl =Ludwigia hyssopifolia
Jussiaea lithospermifolia Micheli =Ludwigia rigida
Jussiaea lithospermifolia var. pubescens Munz =Ludwigia rigida
Jussiaea lithospermifolia var. typica Munz =Ludwigia rigida
Jussiaea maypurensis Kunth =Ludwigia nervosa
Jussiaea mexiae Munz =Ludwigia mexiae
Jussiaea micrantha Kunze =Liidwigia hyssopifolia
Jussiaea mollis Kunth =Ludwigia peruviana
Jussiaea natans Bonpl. =Ludwigia helminthorrhiza
Jussiaea nervosa Poir. =Ludwigia nervosa
Jussiaea nervosa var. pubescens Micheli =Ludwigia nervosa
Jussiaea nervosa var. typica Munz ^Ludwigia nervosa
Jussiaea octonervia Lam. =Ludwigia octovalvis
Jussiaea palustris G. Mey. =Ludwigia decurrens
Jussiaea peruviana L. =Ludwigia peruviana
Jussiaea peruviana var. glaberrima J.D. Sm. =Ludwigia peruviana
Jussiaea peruviana var. typica Munz =Ludwigia peruviana
Jussiaea pterophora Miq. =Ludwigia decurrens
Jussiaea quadrangularis Micheli =Ludwigia quadrangularis
Jussiaea rigida Miq. =Ludwigia rigida
Jussiaea sedoides Bonpl. =Ludwigia sedoides
Jussiaea suffruticosa L. =Ludwigia octovalvis
Jussiaea surinamensis Miq. =Ludwigia leptocarpa
Jussiaea torulosa Am. =Ludwigia torulosa
Jussiaea variabilis var. affinis (DC.) Kuntze =Ludwigia affinis
Jussiaea variabilis var. meyeriana Kuntze =Ludwigia leptocarpa
Ludwigia affinis var. dodecandra (DC.) Munz =Ludwigia dodecandra
Ludwigia lithospermifolia (Micheli) H. Hara =Ludwigia rigida
Ludwigia micrantha (Kunze) H. Hara =Ludwigia hyssopifolia
Oenothera octovalvis Jacq. =Ludwigia octovalvis
Oocarpon torulosum (Am.) Urb. =Ludwigia torulosa
References
Jonker, F.P. 1942. Oenotheraceae. In A. Pulle, Flora of Suriname 3(2): 13-34.
Kawasaki, M.L. 2002. Onagraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
562-564.
Checklist of the Plants of the Guiana Shield 453
Zardini, E.M. & P.H. Raven. 1991. Onagraceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 10: 1-44.
Zardini, E.M. & RH. Raven. 2003. Onagraceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 7: 188-197.
Opiliaceae
R Hiepko, J.S. Miller
Agonandra brasiliensis Miers ex Benth. & Hook, f
Agonandra brasiliensis subsp. brasiliensis
Agonandra brasiliensis subsp. racemigera Hiepko
Agonandra silvatica Ducke
AM BO
AM BO
BO
AM
DA
DA
GU
GU
GU
SU
SU
SU FG
References
Hiepko, R 1993. Opiliaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 14: 36-39.
Miller, J.S. 2003. Opiliaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 7:
198-199.
OXALIDACEAE
G.A. Aymard C, RE. Berry, A. Lourteigt
Biophytum calophyllum (Progel) Guillaumin AM
Biophytum cardonaei Pittier
Biophytum kayae Aymard & RE. Berry AM
Biophytum lourteigiae Aymard & RE. Berry AM
Biophytum ottohuberi Aymard & RE. Berry
Oxalis barrelieri L.
Oxalis corniculata L. [naturalized]
Oxalis debilis Kunth
Oxalis debilis var. corymbosa (DC.) Lourteig
Oxalisfrutescens L. AM
Oxalisfrutescens subsp. borjensis (Kunth) Lourteig AM
Oxalisfrutescens subsp. frutescens
Oxalis juruensis Diels AM
Oxalis mucronulata Norlind
Ruptiliocarpon aff caracolita Hammel & N. Zamora [referred to segregate
family Lepidobotryaceae by Hammel & Zamora]
BO
BO
BO
BO
BO
BO
BO
GU
GU SU FG
GU SU FG
GU
GU
GU SU FG
GU SU FG
GU
SU
Synonyms and Misidentifications
Acetosella schomburgkiana (Progel) Kuntze =Oxalis frutescens
Biophytum casiquiarense R. Knuth =Biophytum calophyllum
Biophytum passargei R. Knuth =Biophytum calophyllum
Oxalis amazonica Progel =Oxalis barrelieri
Oxalis borjensis Kunth =Oxalis frutescens subsp. borjensis
Oxalis calophylla Progel =Biophytum calophyllum
Oxalis pentantha Jacq. =Oxalis frutescens
Oxalis schomburgkiana Progel =Oxalis frutescens
Oxalis schomburgkiana var. lasiocarpa Progel =Oxalis frutescens
Oxalis schomburgkiana var. leiocarpa Progel =Oxalis frutescens
Oxalis sepium A. St.-Hil. [not in flora area]
References
(incl. Lepidobotryaceae)
Aymard C., G.A. & RE. Berry. 2003. Oxalidaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 7: 619-625.
Hammel, B.E. & N.A. Zamora. 1993. Ruptiliocarpon (Lepidobotryaceae): a new arborescent genus and tropical American link
to Africa, with a reconsideration of the family. Novon 3(4): 408-417.
Jonker, RR 1942. Oxalidaceae. In A. Pulle, Flora of Suriname 3(2): 44-55.
Lourteig, A. 1980. Oxalidaceae. In R.E. Woodson, Jr., R.W. Schery, & collaborators. Flora of Panama. Ann. Missouri Bot. Gard.
67: 823-850.
Lourteig, A. 1994. Oxalis L. subgenero Thamnoxys (Endl.) Reiche emend. Lourt. Bradea7(l): 1-199.
Mitchell, J.D. 2002. Oxalidaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
564-566.
454 Checklist of the Plants of the Guiana Shield
Papaveraceae
R.W. Kiger
Argemone mexicana L. [cultivated, naturalized] SU FG
References
Stoffers, A.L. 1979. Papaveraceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 331-334.
Passifloraceae
C. Feuillet, S.S. Tillett
Ancistrothyrsiis hirteUiis A.H. Gentry AM
Ancistrothyrsus cf. tessmannii Harms
Dilkea acuminata Mast.
Dilkeajohannesii Barb. Rodr.
Dilkea parvi/lora Killip
Dilkea retiisa Mast.
Dilkea sp. A Feuillet, ined.
Dilkea sp. B Feuillet, ined.
Dilkea wallisii Mast.
Mitostemma glaziovii Mast.
Mitostemmajenmanii Mast.
Passiflora acuminata DC.
Passiflora adenopoda DC.
Passiflora aimae Annonay & Feuillet
Passiflora ambigua Hemsl.
Passiflora amicorum Wurdack
Passiflora amoena L.K. Escobar
Passiflora arta Feuillet, ined.
Passiflora ascidia Feuillet
Passiflora auriculata Kunth
Passiflora balbis Feuillet
Passiflora bicornis Houst. ex Mill.
Passiflora biflora Lam.
Passiflora bomareifolia Steyerm. & Maguire
Passiflora Candida (Poepp. & Endl.) Mast.
Passiflora capparidifolia Killip
Passiflora cardonae Killip
Passiflora cauliflora Harms
Passiflora cerasina Annonay & Feuillet
Passiflora ceratocarpa F. Silveira
Passiflora cirrhiflora Juss.
Passiflora citrifolia (Juss.) Mast.
Passiflora coccinea Aubl.
Passiflora compar Feuillet, ined.
Passiflora coriacea Juss.
Passiflora costata Mast.
Passiflora crenata Feuillet & Cremers
Passiflora cuneata Willd.
Passiflora cyanea Mast.
Passiflora deficiens Mast.
Passiflora edulis Sims
Passiflora edulis i.flavicarpa Degener [cultivated, escaping]
Passiflora exura Feuillet
Passiflorafanchonae Feuillet
Passiflora filipes Benth.
Passiflorafoetida L.
Passiflorafoetida mqx. foetida
Passiflorafoetida var. gossypiifolia (Desv. ex Ham.) Mast.
Passiflorafoetida var. hispida (DC.) Killip
Passiflorafoetida var.. orinocensis (Killip) Feuillet, ined.
Passiflora fuchsiiflora Hemsl.
Passiflora gabriellana Vanderplank
Passiflora garckei Mast.
GU FG
AM BO DA
GU
GU
GU
GU
GU
SU
SU
FG
FG
FG
FG
BO?
DA
GU FG
FG
AM
BO GU
GU SU FG
GU
BO GU
AM BO DA GU SU FG
AM BO
BO
DA
GU
AM BO
BO GU SU FG
BO GU
AM BO GU
AM
GU
SU FG
FG
BO GU SU FG
FG
AM BO DA
DA
GU
GU
GU
SU FG
FG
AM BO
BO
BO
DA
DA
GU
GU
SU FG
FG
AM BO DA GU SU FG
AM BO
BO?
DA GU
GU
SU FG
FG
FG
AM BO DA GU SU FG
AM BO
BO DA
GU SU FG
AM BO DA GU SU FG
AM BO DA
GU SU FG
FG
BO GU SU FG '
GU FG
GU SU FG
GU su FG
FG
GU
GU
SU
GU su FG
su FG
Checklist of the Plants of the Guiana Shield 455
Passiflora glandulosa Cav. AM BO GU SU FG
Passiflora gleasonii Killip BO? GU
Passiflora gracilis J. Jacq. ex Link BO
Passiflora giiazumaefolia Juss. BO
Passiflora holtii Killip AM
Passiflora involucrata (Mast.) A.H. Gentry AM
Passiflora kawensis Feuillet
Passiflora laurifolia L. AM BO DA
Passiflora leptopoda Harms
Passiflora longicitspis Vanderplank & S. Vanderplank
Passiflora longiracemosa Ducke AM BO
Passiflora maguirei Killip AM
Passiflora maliformis L. [cultivated]
Passiflora micropetala Mast.
Passiflora misera Kunth BO
Passiflora moritziana Planch, [expected in GU] BO
Passiflora multiformis Jacq. AM
Passiflora nitida Kunth AM GU SU FG
Passiflora nuriensis Steyerm. BO
Passiflora ovata Martin ex DC.
Passiflora pachyantha Killip
Passiflora pedata L.
Passiflora pedata var. pedata
Passiflora pedata var. stipiilaris (Killip) Killip
Passiflora phaeocaiila Killip
Passiflora phellos Feuillet
Passiflora picturata Ker Gawl.
Passiflora plumosa Feuillet & Cremers
Passiflora pulchella Kunth
Passiflora pyrrhantha Harms
Passiflora qiiadrangularis L. [cultivated or escaped from cultivation]
Passiflora quadriglandulosa Rodschied
Passiflora quelchii N.E. Br.
Passiflora retipetala Mast.
Passiflora riparia Mart, ex Mast.
Passiflora rubra L.
Passiflora rufostipulata Feuillet
Passiflora saulensis Feuillet
Passiflora sclerophylla Harms
Passiflora securiclata Mast.
Passiflora seemannii Griseb.
Passiflora serrato-digitata L.
Passiflora serndata Jacq.
Passiflora sp. A
Passiflora sp. B
Passiflora s^. C
Passiflora sp. D Feuillet & J.M. MacDougal, ined. GU
Passiflora sp. E Feuillet, ined. FG
Passiflora sp. F Feuillet, ined. FG
Passiflora sp. G Feuillet, ined. FG
Passiflora sp. H Feuillet & J.M. MacDougal, sp. nov. ined. GU SU FG
Passiflora spicata Mast. GU
Passiflora spinosa (Poepp. & Endl.) Mast. BO
Passiflora stipulata Aubl. FG
Passiflora suberosa L. BO DA SU
Passiflora subpeltata Ortega DA
Passiflora trialata Feuillet & J.M. MacDougal FG
Passiflora tiiberosa Jacq. DA GU
Passiflora variolata Poepp. & Endl. AM BO DA FG
Passiflora vespertilioL. AM BO DA GU SU FG
Passiflora vitifolia Kunth FG?
Synonyms and Misidentifications
Dilkeajohannesii var. parvifolia Hoehne =Dilkea acuminata
Dilkea magniflca Steyerm. =Dilkea acuminata
GU SU
AM BO DA GU
GU
su
su
AM BO GU
AM
AM
BO su
FG
AM BO
AM BO
DA GU su FG
AM BO DA GU
GU
BO GU su FG
AM GU
GU su FG
FG
FG
AM BO GU
AM BO GU
AM
GU su FG
BO
AM BO
456 Checklist of the Plants of the Guiana Shield
Passiflora capsularis L. [not in flora area]
Passiflorafoetida van moritziana (Planch.) Killip =Passiflora moritziana
Passiflora gossypifolia Desv. ex Ham. =Passiflorafoetida var. gossypiifolia
Passiflora hispida DC. ex Triana & Planch. =Passiflorafoetida var. hispida
Passiflora kegeliana Garcke =Passiflora auriculata
Passiflora lonchophora Harms =Passiflora retipetala
Passiflora murucuia L. [not in flora area]
Passiflora pallida L. =Passiflora suberosa
Passiflora pedata subsp. stipularis Killip =Passiflora pedata var. stipularis
Passiflora retrorsa Killip =Passiflora securiclata
Passiflora rotundifolia L. [not in flora area]
Passiflora serratifolia L. [not in flora area]
Tacsonia spinosa Poepp. & Endl. =Passiflora spinosa
References
Escobar, L.K. 1994. Two new species and a key to Passiflora subg. Astrophea. Syst. Bot. 19(2): 203-210.
Feuillet, C. 1986. Deux Passifloraceae nouvelles et quelques especes rares en Guyane fran^aise. Candollea 41: 173-178.
Feuillet, C. 1989. Diversity and distribution on Guianan Passifloraceae. In L.B. Holm-Nielsen, l.C. Nielsen & H. Balslev,
Tropical Forests, Botanical Dynamics, Speciation and Diversity. Academic Press, London.
Feuillet, C. 1994. Two new species of Passiflora (Passifloraceae) from French Guiana. Novon 4(3): 236-241.
Feuillet, C. 2002. Anew series and three new species of Passiflora subgenus Astrophea from the Guianas. Brittonia 54(1): 18-29.
Feuillet, C. 2004. Passifloraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
566-570.
Feuillet, C. 2004. Passiflora phellos, a new species of subgenus Passiflora (Passifloraceae) from northern South America. Novon
14 (3): 285-287.
Feuillet, C. & G. Cremers. 1984. Passifloraceae nouvelles ou meconnues de Guyane franfaise. Proc. Kon. Ned. Akad. Wet. ser.
C, 87: 377-386.
Gentry, A.H. 1981. Distributional patterns and an additional species of the Passiflora vitifolia complex: Amazonian species
diversity due to edaphically differentiated communities. PI. Syst. Evol. 137: 95-105.
Killip, E.P. 1937. Passifloraceae. In A. Pulle, Flora of Suriname 3(1): 306-327.
Killip, E.P. 1938. The American species of Passifloraceae. Publ. Field Mus. Nat. Hist., Bot. Ser., 19: 1-613.
Tillett, S.S. 1988. Passionis passifloris II. Terminologia. Emstia 48: 1-40.
Tillett. S.S. 2003. Passifloraceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
7: 625-667.
Pedaliaceae
J.S. Miller
Craniolaria annua L. [expected in SU] AM BO GU
Sesamum orientale L. [cultivated & escaping] AM BO GU SU FG
Synonyms and Misidentifications
Sesamum indicum L. =Sesamum orientale
References
(incl. Martyniaceae)
Eseltine, GP. van. 1929. A preliminary study of the unicorn plants (Martyniaceae). N.Y. State Ag. Expt. Sta. (Geneva) Tech.
Bull. 149: 1-41.
Miller, J.S. 2003. Pedaliaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 7:
667-670.
Pulle, A. 1932. Pedaliaceae. In A. Pulle, Flora of Suriname 4(1): 111-112.
Seegeler, C.J. P. 1989. Sesamum orientale L. (Pedaliaceae): sesame's correct name. Taxon 38(4): 656-659.
Peridiscaceae
B.K. Hoist
Peridiscus lucidus Benth. AM BO
Whittonia guianensis Sandwith GU
References
Hoist, B.K. 2003. Peridiscaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
7:670-671.
Sandwith, N.Y. 1962. Contributions to the flora of tropical America: LXIX. A new genus of Peridiscaceae. Kew Bull. 15: 467-
471.
Checklist of the Plants of the Guiana Shield 457
Phytolaccaceae
G.A. Aymard C, N.L. Cuello A., R.A. DeFilippsj
DA GU SU FG
GU SU
BO DA GU SU FG
BO DA GU SU FG
BO
BO GU FG
BO DA GU SU FG
BO DA
BO FG
GU SU FG
BO DA GU FG
DA GU FG?
Microtea debilis Sw. AM
Microtea glochidiata Moq. [Serra Neblina, expected in AM]
Microtea maypurensis (Kunth) G Don AM
Petiveria alliacea L.
Phytolacca rivinoides Kunth & Bouche AM
Phytolacca rugosa A. Braun & Bouche
Phytolacca thyrsiflora Fenzl ex J.A. Schmidt
Rivina humilis L.
Rivina humilis var. humilis
Seguieria aculeata Jacq.
Seguieria americana L.
Seguieria macrophylla Benth.
Trichostigma octandrum (L.) H. Walter
Synonyms and Misidentifications
Ancistrocarpus maypurensis Kunth =Microtea maypurensis
Ceratoccoca maypurensis (Kunth) Willd. ex Schult. =Microtea maypurensis
Microtea maypurensis var. angustifolia Moq. =Microtea maypurensis
Petiveriafoetida Salisb. =Petiveria alliacea
Phytolacca americana L. [not in flora area]
Rivina octandra L. =Trichostigma octandrum
Seguieria cordata Britton =Seguieria macrophylla
Seguieria foliosa Benth. =Seguieria americana
References
Aymard C, G.A. & N.L. Cuello A. Phytolaccaceae. 2003. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of
the Venezuelan Guayana 7: 672-680.
Burger, W. 1983. Phytolaccaceae. In W. Burger, Flora Costaricensis. Fieldiana Bot. (n.s.) 13: 199-212.
DeFilipps, R.A. & S.L. Maina. 2003. Phytolaccaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M.
Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New
York Bot. Gard. 76 (2): 571-574.
Raeder, K. 1961. Phytolaccaceae. In R.E. Woodson Jr., R.W. Schery, & collaborators. Flora of Panama. Ann. Missouri Bot.
Gard. 48(1): 66-79.
Rohwer, J. 1982. Ataxonomic revision of the genera Seguieria Loefl. and Gallesia Casar. Mitt. Bot. Staats. Munchen 18: 231-
288.
Stoffers, A.L. 1957. Phytolaccaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 209-217.
Walter, H. 1909. Phytolaccaceae. In A. Engler, Das Pflanzenreich 4(83): 1-154.
PiCRAMNIACEAE
J.R. Pirani
Picramnia guianensis (Aubl.) Jans.-Jac. [expected in BO]
Picramniajuniniana J.F. Macbr.
Picramnia latifolia Tul.
Picramnia magnifolia J.F. Macbr.
Picramnia nuriensis Steyerm.
Picramnia pentandra Sw.
Picramnia sellowii Planch.
Picramnia spruceana Engl.
Picrasma excelsa (Sw.) Planch.
Synonyms and Misidentifications
Picramnia longissima Tul. =Picramnia latifolia
Picramnia macrostachys Klotzsch ex Engl. =Picramnia latifolia
Picramnia platystachya Killip & Cuatrec. =Picramnia magnifolia
Picramnia sellowii subsp. spruceana (Engl.) Pirani =Picramnia spruceana
Picramnia tariri DC. =Picramnia guianensis
Picramnia tristamina Steyerm. =Picramnia juniniana
Tariri guianensis Aubl. =Picramnia guianensis
AM
AM
AM
AM
BO
BO
BO
BO
DA
GU SU FG
GU
GU SU FG
GU
GU
GU SU FG
GU SU FG
FG?
458 Checklist of the Plants of the Guiana Shield
PiPERACEAE
R. Callejas-Posada, A.R.A. Gorts-van Rijn, J.A. Steyermarkt
Peperomia acuminata Ruiz & Pav. [Mt. Roraima]
Peperomia alata Ruiz & Pav.
Peperomia alpina (Sw.) A. Dietr. [Mt. Roraima]
Peperomia angularis C. DC.
Peperomia angustata Kunth
Peperomia blanda (Jacq.) Kunth
Peperomia celiae Yunck.
Peperomia circinnata Link
Peperomia cladara Yunck.
Peperomia delascioi Steyerm. [Mt. Roraima]
Peperomia dendrophila Schltdl. & Cham.
Peperomia distachya (L.) A. Dietr. [expected in GU]
Peperomia duidana Trel. ex Gleason
Peperomia elongata Kunth
Peperomia elongata var. elongata
Peperomia elongata var. guianensis Yunck.
Peperomia elongata var. piliramea Trel. & Yunck.
Peperomia emarginella (Sw. ex Wikstr.) C. DC.
Peperomia enantiostachya C. DC.
Peperomiafundacionens is Steyerm.
Peperomia galioides Kunth [Mt. Roraima]
Peperomia gentryi Steyerm.
Peperomia glabella (Sw.) A. Dietr.
Peperomia glabella var. glabella
Peperomia glabella var. nervulosa (C. DC.) Yunck.
Peperomia gracieana Gorts
Peperomia guaiquinimana Trel. & Yunck.
Peperomia haematolepis Trel.
Peperomia hernandiifolia (Vahl) A. Dietr.
Peperomiajamesoniana C. DC.
Peperomia lanceolato-peltata C. DC.
Peperomia lancifolia Hook.
Peperomia loxensis Kunth
Peperomia macrostachya (Vahl) A. Dietr.
Peperomia macrostachya var. macrostachya
Peperomia macrostachya var. nematostachya (Link) Trel. & Yunck.
Peperomia maculosa (L.) Hook.
Peperomia magnoliifolia (Jacq.) A. Dietr.
Peperomia maguirei Yunck.
Peperomia manarae Steyerm.
Peperomia marahuacensis Steyerm.
Peperomia maypurensis Kunth
Peperomia neblinana Yunck.
Peperomia obtusifolia (L.) A. Dietr.
Peperomia obtusifolia var. obtusifolia
Peperomia ouabianae C. DC.
Peperomia pellucida (L.) Kunth
Peperomia pernambucensis Miq.
Peperomia pilicaulis C. DC.
Peperomia purpurinervis C. DC. [Mt. Roraima]
Peperomia quadrangularis (J.V. Thomps.) A. Dietr.
Peperomia quadrifolia (L.) Kunth
Peperomia quaesita Trel.
Peperomia rhombea Ruiz & Pav.
Peperomia rotundata Kunth [Mt. Roraima]
Peperomia rotundata var. rotundata
Peperomia rotundata var. trinervula (C. DC.) Steyerm. [Mt. Roraima]
Peperomia rotundifolia (L.) Kunth
Peperomia schomburgkii C. DC.
Peperomia serpens (Sw.) Loudon
Peperomia spruceana Benth. ex C. DC.
AM BO GU
AM BO
BO
GU
GU
SU FG
FG
AM BO DA GU FG
BO GU? SU
AM
SU?
AM BO
AM BO GU
AM BO
AM SU FG
AM BO GU
AM BO DA GU SU FG
GU SU FG
AM BO DA GU FG
AM GU SU
AM BO DA GU FG
AM
AM
AM BO
AM BO
AM BO DA GU SU FG
BO DA GU SU FG
AM BO
BO
DA GU SU FG
FG
AM
AM BO GU
AM BO
BO
AM BO GU
AM
AM BO DA GU SU FG
AM BO DA GU SU FG
BO GU FG
AM BO
AM BO DA GU SU FG
GU SU FG
GU
AM
AM BO
AM BO
AM BO DA GU SU FG
GU SU FG
AM BO GU SU FG
AM BO DA GU SU FG
BO GU
GU
SU FG
FG
BO GU? FG
AM BO GU
GU
SU FG
AM BO
GU
AM BO
AM
AM BO
AM BO DA GU
GU
SU FG
AM BO DA GU SU FG
AM
AM BO
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
AM
AM
AM
AM
AM BO
AM BO
BO
AM
AM BO
AM BO
BO
FG
GU SU FG
GU SU FG
Checklist of the Plants of the Guiana Shield 459
Peperomia striata Ruiz & Pav.
Peperomia tenella (Sw.) A. Dietr. GU
Peperomia tenella var. tenella GU
Peperomia tenella var. tyleri (Trel.) Steyerm.
Peperomia tetraphylla (G. Forst.) Hook. & Am. GU?
Peperomia transparens Miq. SU FG
Peperomia uaupesensis Yunck.
Peperomia urocarpa Fisch. & C.A. Mey. GU
Peperomia venezueliana C. DC.
Peperomia venusta Yunck.
Peperomia yatuensis Steyerm.
Peperomia yutajensis Steyerm.
Piper abellinum Trel. & Yunck.
Piper adenandrum (Miq.) C. DC.
Piper aduncum L.
Piper aduncum var. aduncum
Piper aduncum var. cordulatum (C. DC.) Yunck.
Piper aduncum var. garcia-barrigae Trel. & Yunck.
Piper aequale Vah\ DA GU SU FG
Piper alatabaccum Trel. & Yunck. GU SU FG
Piper amalago L. GU SU FG
Piper amapense Yunck. GU? FG
Piper amplectenticaule Trel. & Yunck. FG
Piper amplum Kunth FG
Piper angustifolium Lam. FG
Piper anonifolium (Kunth) C. DC. BO DA GU SU FG
Piper anonifolium var. anonifolium BO DA GU SU FG
Piper anonifolium YSLT.parkerianum (CDC.) Steyerm. BO GU SU FG
Piper arboreum Auhl AM BO DA GU SU FG
Piper arboreum var. arboreum AM BO DA GU SU FG
Piper arboreum var. hirtellum Yunck. [subsp. arboreum^ AM BO GU SU FG
Piper augustumRudge AM BO GU SU FG
Piper augustum var. augustum GU SU FG
Piper augustum var. pubinerve Trel. & Yunck. GU FG
Piper auritum Kunth SU FG
Piper avellanum (Miq.) CDC. AM BO GU SU FG
Piper avellanum var. angustifolium Trel. & Yunck. SU
Piper avellanum var. avellanum GU SU FG
Piper baccans (Miq.) C. DC. AM GU
Piper baccans var. sancarlosianum (C. DC.) Steyerm. AM
Piper bartlingianum (Miq.) CDC AM BO GU SU FG
Piper bolivaranum Yunck. BO GU
Piper brasiliense C DC. GU
Piper brownsbergense Yunck. SU FG
Piper cernuum Veil. AM BO GU? FG
Piper cernuum var. cernuum AM BO GU? FG
Piper cernuum var. perlongispicum (Yunck.) Steyerm. BO
Piper cililimbum Yunck. AM BO GU
Piper ciliomarginatum Gorts & Christenh. GU
Piper consanguineum (Kunth) CDC BO GU SU FG
Piper coruscans Kunth AMBODAGU FG
Piper crassinervium Kunth AM BO FG
Piper cumanense Kunth BO
Piper cuyunianum Steyerm. BO GU
Piper cyrtopodon (Miq.) C. DC. GU SU FG
Piper demeraranum (Miq.) C. DC. AM BO DA GU SU FG
Piper dilatatum Rich. BO DA GU SU FG
Piper divaricatum G. Mey. DA GU SU FG
Piper duckei C. DC. GU
Piper dumosum Rudge GU SU FG
Piper dunstervilleorum Steyerm. BO
Piper eucalyptifolium Rudge GU FG
Piperfanshawei Yunck. GU
Piperflexicaule (Miq.) C. DC. GU
460 Checklist of the Plants of the Guiana Shield
Piperfoveolatum Kunth ex C. DC.
Piperfrancovilleaniim C. DC.
Piperfuligineum Kunth
Piper gabrielianum (Miq.) C. DC.
Piper gentryi Steyerm.
Piper glabrescens (Miq.) C. DC.
Piper glabrescens var. caparomim (C. DC.) Yunck.
Piper glabrescens var. glabrescens
Piper giiianense (Klotzsch) C. DC.
Piper hermannii Trel. & Yunck.
Piper hippocrepiforme Steyerm.
Piper hirtilimbum Trel. & Yunck.
Piper hispidum Sw.
Piper holtii Trel. & Yunck.
Piper hostmannianum (Miq.) C. DC.
Piper Immistratum Gorts & K.U. Kramer
.
Piper inaequale C. DC.
Piper insigne (Kunth) C. DC.
Piperjauaense Steyerm.
Piperjavitense Kunth
Piperjitlianii Callejas
Piper kappleri C. DC.
Piper kegelianum (Miq.) C. DC.
Piper kegelii C. DC.
Piper laevigatiim Kunth
Piper lemaense Yunck.
Piper liesneri Steyerm.
Piper marginatum Jacq.
Piper marginatum var. marginatum
Piper microstachyum Vahl [=Piper angustifoliuml]
Piper modestum (Miq.) C. DC. [insufficiently known taxon]
Piper morilloi Steyerm.
Piper mosaicum Steyerm.
Piper neblinanum Yunck.
Piper nematanthera (Miq.) C. DC.
Piper nematanthera var. nematanthera
Piper nematanthera var. sagotii (C. DC.) Trel. & Yunck.
Piper nigrispicum C. DC.
Piper nigrum L. [cultivated, perhaps escaping]
Piper obliquum Ruiz & Pav.
Piper oblongifolium (Klotzsch) C. DC.
Piper oblongifolium var. glabrum C. DC.
Piper oblongifolium var. oblongifolium
Piper otto-huberi Steyerm.
Piper otto-huberi var. eciliatum Steyerm.
Piper otto-huberi var. otto-huberi
Piper ovatum Vahl [expected in GU]
Piper papilliferum Steyerm.
Piper paramaribense C. DC.
Piper parapeltobryon Trel.
Piper peltatum L. [=Pothomorphe peltata; =Lepianthes peltatum]
Piper perciliatum Trel. & Yunck.
Piper perstipulare Steyerm.
Piper piscatorum Trel. & Yunck.
Piper politii Yunck.
Piper politii subsp. politii
Piper politii subsp. sipapoense Steyerm.
Piper politii subsp. toronoense Steyerm.
Piper pseudoacreanum Steyerm.
Piper pseudoglabrescens Trel. & Yunck.
Piper pubivaginatum Steyerm.
Piper pulleanum Yunck.
Piper reticulatum L.
Piper rudgeanum (Miq.) C. DC.
AM BO GU su FG
AM BO
GU su FG
FG
BO
GU
GU
GU
su
su
AM GU FG'
AM
BO
GU
AM BO DA GU su FG
AM
AM BO DA GU
GU
su
su
FG
FG
FG
GU su FG
AM BO
AM
BO
GU SU FG
BO GU SU
SU
FG
AM BO
rU
AM GU
AM BO DA GU SU FG
AM BO DA
FG
FG
AM
AM
AM BO
AM
AM
AM
AM
AM
BO
BO
DA
GU SU FG
GU SU FG
FG
GU SU FG
GU SU FG
GU SU FG
GU
GU SU FG
GU SU FG
AM
AM BO DA GU SU FG
AM
AM BO GU
AM BO GU FG
AM BO
AM BO
AM
BO
AM
AM BO DA GU
AM
GU SU FG
BO DA GU su
su
FG
FG?
BO DA
BO
BO
GU
BO GU
BO DA GU
BO GU
BO GU
BO
BO
BO GU
Checklist of the Plants of the Guiana Shield 461
Piper nipuminianum Trel. & Yunck. AM GU
Piper sabanaense Yunck. BO
Piper salicifolium Vahl SU
Piper schiedeanum Steud. GU? SU?
Piper steyermarkii Yunck. BO
Piper subalpinum Yunck. BO
Piper subciliatuni C. DC. FG
Piper subduidaense Yunck. AM
Piper tamayoanum Steyerm. BO
Piper tectonifolium Kunth GU? FG
Piper tenue Kunth
Piper tepuiense Steyerm.
Piper toronotepuiense Steyerm.
Piper tortivemdosum Yunck. AM
Piper trichoneuron (Miq.) C. DC. SU FG
Piper trigonum C. DC. AM
Piper tuberculatum isicq. SU FG
Piper umbellatum L. [=Pothomorphe umbellata; -Lepianthes umbellatum]
Piper venamoense Steyerm.
Piper vitaceum Yunck.
Piper vitaceum var. venezuelense Steyerm.
Piper wachenheimii Trel. SU FG
Synonyms and Misidentifications
Acrocarpidium majus Miq. =Peperomia urocarpa
Acrocarpidium tenellum (Sw.) Miq. =Peperomia tenella
Acrocarpidium urocarpum (Fischer & C. Mey.) Miq. =Peperomia urocarpa
Artanthe adenandra Miq. =Piper adenandrum
Artanthe aduncum (L.) Miq. =Piper aduncum
Artanthe amazonica Miq. =Piper coruscans
Artanthe anonifolia (Kunth) Miq. =Piper anonifolium
Artanthe apiculata Klotzsch =Piper anonifolium
Artanthe asperifolia (Ruiz & Pav.) Miq. =Piper hispidum
Artanthe augusta (Rudge) Miq. =Piper augustum
Artanthe avellana Miq. =Piper avellanum
Artanthe baccans Miq. =Piper baccans
Artanthe bartlingianum Miq. =Piper bartlingianum
Artanthe berbicense Miq. =Piper hostmannianum
Artanthe caripensis (Kunth) Miq. =Piper aequale
Artanthe caudata (Vahl) Miq. =Piper marginatum
Artanthe consanguinea (Kunth) Miq. =Piper consanguineum
Artanthe demerarana Miq. =Piper demeraranum
Artanthe eximia (Kunth) Miq. =Piper cernuum
Artanthe flagellaris (Kunth) Miq. =Piper obliquum
Artanthe guianensis Klotzsch =Piper guianense
Artanthe hirsuta (Sw.) Miq. =Piper hispidum
Artanthe hostmanniana Miq. =Piper hostmannianum
Artanthe javitensis (Kunth) Miq. =Piperjavitense
Artanthe kegeliana Miq. =Piper kegelianum
Artanthe luschnathiana (Kunth) Miq. =Piper arboreum var. hirtellum
Artanthe magnifica Miq. [nom. illeg.] =Piper obliquum
Artanthe marginata (Jacq.) Miq. -Piper marginatum
Artanthe obrumbata Miq. =Piper arboreum var. hirtellum
Artanthe parkeriana Miq. =Piper anonifolium var. parkerianum
Artanthe peduncularis Miq. =Piper pedunculare
Artanthe pseudochurumayu (Kunth) Miq. =Piper coruscans
Artanthe ramiflora Miq. ex C. DC. =Piper hostmannianum
Artanthe schomburgkii Klotzsch =Piper pedunculare
Artanthe spectabilis Miq. =Piper cernuum
Artanthe superba Miq. =Piper obliquum
Enckea dubia Kunth =Piperfoveolatum
Heckeria peltata (L.) Kunth =Piper peltatum
Heckeria umbellata (L.) Kunth =Piper umbellatum
Lepianthes peltatum (L.) Raf. ^Piper peltatum
462 Checklist of the Plants of the Guiana Shield
Lepianthes umbellatum (L.) Raf. =Piper umbellatum
Ottonia ovata (Vahl) Trel. =Piper ovatum
Peperomia alata var. angustifolia C. DC. =Peperomia alata
Peperomia angulata Kunth =Peperomia quadrangularis
Peperomia atabapoensis Steyerm. =Peperomia quadrangularis
Peperomia bartlettii C. DC. =Peperomia rotundifolia
Peperomia blanda var. langsdorffii (Miq.) Henschen =Peperomia blanda
Peperomia boomii Steyerm. =Peperomia lanceolato-peltata
Peperomia caulibarbis Miq. =Peperomia glabella var. glabella
Peperomia chimantana Yunck. =Peperomiajamesoniana
Peperomia choroniana var. puberulenta Yunck. =Peperomia hernandiifolia
Peperomia ciliata Kunth =Peperomia blanda
Peperomia cladara f. ciliata Yunck. =Peperomia cladara
Peperomia cladara f. perglabra Yunck. =Peperomia cladara
Peperomia cryptotricha Trel. ex V.M. Badillo [nomen nudum] =Peperomia tetraphylla
Peperomia cuneata Miq. =Peperomia obwsifolia
Peperomia decipiens C. DC. =Peperomia blanda
Peperomia diffusa C. DC. =Peperomia loxensis
Peperomia distachya f. geminispica (Trel. & Yunck.) Steyerm. =Peperomia distachya
Peperomia emarginata Ruiz & Pav. =Peperomia obtusifolia
Peperomia emarginella var. glabrior C. DC. =Peperomia emarginella
Peperomia epilobioides Trel. & Yunck. =Peperomia rotundata var. rotundata
Peperomia erasmiaeformis Trel. =Peperomia lancifolia
Peperomia ernstiana C. DC. =Peperomia venezueliana
Peperomiafoveolata Steyerm. =Peperomia distachya
Peperomia geminispica Trel. & Yunck. =Peperomia distachya
Peperomia glabella var. melanostigma (Miq.) Dahlst. ^^^Peperomia glabella var. nervulosa
Peperomia guaiquinimana Trel. ex V.M. Badillo [nomen nudum] =Peperomia guaiquinimana
Peperomia hederacea Miq. =Peperomia urocarpa
Peperomia horrescens Trel. ex V.M. Badillo [nomen nudum] =Peperomia venezueliana
Peperomia ionophylla Griseb. =Peperomia serpens
Peperomia lancifolia subsp. erasmiaeformis (Trel.) Steyerm. =Peperomia lancifolia
Peperomia lanjouwii Yunck. =Peperomia rotundifolia
Peperomia longemucronata C. DC. =Peperomia alpina
Peperomia longemucronata f. longirostrata (Yunck.) Steyerm. =Peperomia alpina
Peperomia longifolia C. DC. =Peperomia pernambucensis
Peperomia longirostrata Yunck. =Peperomia alpina
Peperomia macaroana Trel. ex V.M. Badillo [nomen nudum] =Peperomia blanda
Peperomia magnoliifolia var. emarginulata C. DC. =Peperomia obtusifolia
Peperomia major C. DC. =Peperomia urocarpa
Peperomia melanostigma var. nervulosa C. DC. =Peperomia glabella var. nervulosa
Peperomia microreticulata Steyerm. ^Peperomia alata
Peperomia moritzii C. DC. =Peperomia rotundata var. trinervula
Peperomia myosuroides (Rudge) A. Dietr. =Peperomia macrostachya
Peperomia myriocarpa Miq. =Peperomia elongata var. elongata
Peperomia nematostachya Link =Peperomia macrostachya var. nematostachya
Peperomia nilssonii Yunck. =Peperomia alata
Peperomia nummularifolia Kunth =Peperomia rotundifolia
Peperomia obtusifolia var. cuneata (Miq.) Griseb. =Peperomia obtusifolia
Peperomia obtusifolia var. emarginata (Ruiz & Pav.) Dahlst. =Peperomia obtusifolia
Peperomia obtusifolia var. emarginulata (C. DC.) Trel. & Yunck. =Peperomia obtusifolia
Peperomia omnicola C. DC. =Peperomia striata
Peperomia omnicola var. oblanceolata Trel. =Peperomia striata
Peperomia ornata Yunck. =Peperomia maypurensis
Peperomia paniculata Regel =Peperomia pernambucensis
Peperomia parkeriana Miq. ^Peperomia macrostachya var. macrostachya
Peperomia peltata (L.) A. Dietr. =Piper peltatum
Peperomia pennellii Trel. & Yunck. =Peperomia dendrophila
Peperomia pseudojamesoniana Steyerm. =Peperomiajamesoniana
Peperomia purpurascens Moritz ex C. DC. =Peperomia rotundata var. trinervula
Peperomia pyrifolia Kunth =Peperomia acuminata
Peperomia reflexa (L. f.) A. Dietr. [nom, illeg., non Kunth] =Peperomia tetraphylla
Peperomia reflexa f. americana Miq. =Peperomia tetraphylla
Peperomia reflexa var. americana (Miq.) Dahlst. =Peperomia tetraphylla
Checklist of the Plants of the Guiana Shield 463
Peperomia replans C. DC. =Peperomia serpens
Peperomia roraimana C. DC. =Peperomia rotundata var. trinervula
Peperomia rotundifolia f. ovata Dahlst. =Peperomia rotundifoUa
Peperomia rotundifolia var. ovata (Dahlst.) C. DC. =Peperomia rotundifolia
Peperomia saginans Trel. ex V.M. Badillo [nomen nudum] =Peperomia quadrangularis
Peperomia salvaje C. DC. ^Peperomia blanda
Peperomia silvestris C. DC. =Peperomia ouabianae
Peperomia surinamensis C. DC. =Peperomia elongata var. elongata
Peperomia tafelbergensis Yunck. =Peperomia ouabianae
Peperomia tetraphylla var. americana (Miq.) Yunck. =Peperomia tetraphylla
Peperomia trifolia (L.) A. Dietr. [not in flora area]
Peperomia trinervula C. DC. =Peperomia rotundata var. trinervula
Peperomia trinervula var. suboppositifolia Trel. & Yunck. =Peperomia rotundata var. rotundata
Peperomia tyleri Trel. =Peperomia tenella var. tyleri
Peperomia umbellata (L.) Kunth =Piper umbellatum
Peperomia victoriana C. DC. =Peperomia angustata
Peperomia victoriana var. marginata C. DC. =Peperomia angustata
Peperomia viridispica Trel. =Peperomia angustata
Peperomia wurdackii Yunck. =Peperomia ouabianae
Piper acarouanyanum C. DC. =Piper avellanum
Piper acutissimum Trel. =Piper trigonum
Piper adenophorum C. DC. =Piper dumosum
Piper aduncum var. brachyarthrum (Trel.) Yunck. =Piper aduncum var. cordulatum
Piper aequale var. santa-martanum (C. DC.) Trel. & Yunck. =Piper aequale
Piper affine C. DC. =Piper paramaribense
Piper aggregatum P.J. Bergius (Lacistemataceae) =Lacistema aggregatum
Piper agostinii Yunck. =Piper kegelianum
Piper alpinum Sw. =Peperomia alpina
Piper amalago f. ceanothifolium (Kunth) Steyerm. [var. medium] =Piper amalago
Piper amalago var. medium (Jacq.) Yunck. =Piper amalago
Piper amazonasense Trel. & Yunck. =Piper baccans var. sancarlosianum
Piper amazonicum (Miq.) C. DC. =Piper coruscans
Piper amplectens (Miq.) C. DC. =Piperfuligineum
Piper angremondii DC. =Piper hispidum
Piper angustatum (Kunth) Poir. =Peperomia angustata
Piper angustifolium var. cordulatum C. DC. =Piper aduncum var. cordulatum
Piper angustum Rudge ex Spreng. [nom. superfl.] =Piper augustum
Piper anisatum Kunth =Piper marginatum
Piper anonifolium Kunth [nom. invalid] =Piper anonifolium
Piper arboreum \dx. falcifolium (Trel.) Yunck. =Piper arboreum
Piper arboreum var. latifolium (C. DC.) Yunck. =Piper arboreum var. hirtellum
Piper arboreum subsp. tuberculatum (Jacq.) Tebbs =Piper tuberculatum
Piper arieianum C. DC. =Piper trigonum
Piper arieianum var. chimantanum (Yunck.) Steyerm. =Piper trigonum
Piper arieianum var. duidaense (Trel.) Steyerm. ^Piper trigonum
Piper aristeguietae Yunck. =Piper obliquum
Piper asperifolium Rich. =Piper hispidum
Piper asperifolium Ruiz & Pav. =Piper hispidum
Piper bartlingianum var. parimanum (Trel.) Steyerm. =Piper bartlingianum
Piper benoistii Trel. ex Lemee [apparently a nomen nudum]
Piper blandum Jacq. =Peperomia blanda
Piper brachystachyum C. DC. =Piper eucalyptifolium
Piper cabellense C. DC. =Piper cernuum
Piper caracasanum Bredem. =Piper obliquum
Piper caripense Kunth =Piper aequale
Piper carniconnectivum C. DC. [not in flora area]
Piper Caudatum Vahl =Piper marginatum
Piper cauranum Trel. =Piper demeraranum
Piper ceanothifolium Kunth =Piper amalago
Piper celtidifolium Kunth =Piper aduncum
Piper chimantanum Yunck. =Piper trigonum
Piper citrifolium Lam. =Piper anonifolium var. anonifolium
Piper citrifolium auct. non Lam.: sensu Trel. & Yunck. =Piper anonifolium
Piper citrifolium var. parkerianum (Miq.) Trel. & Yunck. =Piper anonifolium var. parkerianum
464 Checklist of the Plants of the Guiana Shield
Piper colubrimim Link =Piper divaricatum
Piper consanguineum Kunth [pub. as synonym] -Piper consanguineum
Piper coruscans var. membranaceum (C. DC.) Steyerm. =Piper coruscans
Piper corylifolium Kunth =Piper dilatatum
Piper crassum Veil. =Piper divaricatum
Piper demeraranum var. rubescens (C. DC.) Trel. & Yunck. =Piper demeraranum
Piper demimitum Yunck. =Piper bolivaranum
Piper diospyrifoliiim Kunth [not in flora area]
Piper distachyon L. =Peperomia distachya
Piper diiidaense Trel. =Piper trigonum
Piper duidaense var. eciliatum Trel. & Yunck. =Piper trigonum
Piper elongatum var. brachyarthrum Trel. =Piper aduncum var. cordulatum
Piper elongatum var. cordulatum (C. DC.) C. DC. =Piper aduncum var. cordulatum
Piper emarginatum (Ruiz & Pav.) Vahl =Peperomia obtusifolia
Piper emarginellum Sw. ex Wikstr. =Peperomia emarginella
Piper esmeraldanum Trel. =Piper demerar'anum
Piper eximium Kunth =Piper cernuum
Piperfalcifolium Trel. =Piper arboreum
Piperflagellare Kunth =Piper obliquum
Piperflexuosum Rudge =Piperflexicaule
Piperfulgidum Yunck. [not in flora area]
Piperfunckii C. DC. =Piper aequale
Piper geayi Trel. ex Lemee [apparently a nomen nudum]
Piper geniculatum Sw. ^Piper arboreum
Piper geniculatum var. latifolium C. DC. =Piper arboreum var. hirtellum
Piper geniculatum i. puberula C. DC. =Piper tuberculatum
Piper geniculatum var. B C. DC. =Piper arboreum var. hirtellum
Piper gigantifolium C. DC. =Piper cernuum
Piper glabellum (Miq.) C. DC. [nom. illeg., non Piper glabellum Sw.] =Piper divaricatum
Piper glabellum Sw. =Peperomia glabella
Piper glabrilimbum C. DC. =Piper divaricatum
Piper gleasonii Yunck. =Piper trichoneuron
Piper godebertiamim Trel. ex Lemee [apparently a nomen nudum]
Piper hernandiifolium Vahl =Peperomia hernandiifolia
Piper hirsutum Sw. =Piper hispidum
Piper hispidum Kunth [nom. illeg., non Piper hispidum Sw.] =Piper hispidum
Piper hispidum var. obliquum Trel. & Yunck. =Piper hispidum
Piper hohenackeri C. DC. =Piper anonifolium
Piper hostmannianum var. berbicense (Miq.) C. DC. =Piper hostmannianum
Piper hostmannianum var. glabrirameum Trel. & Yunck. =Piper hostmannianum
Piper hostmannianum var. ramiflorum C. DC. -Piper hostmannianum
Piper insipiens Trel. & Yunck. =Piper divaricatum
Piperjohnstonii C. DC. =Piper aequale
Piper lapathifolium Steud. [not in flora area]
Piper latum Kunth =Piper reticulatum
Piper lenormandianum C. DC. =Piper kappleri
Piper leprieuri Pulle =Piper consanguineum
Piper loxense (Kunth) Poir. =Peperomia loxensis
Piper lucaeanum var. magnifolium C. DC. =Piper trigonum
Piper macrophyllum Sw. =Piper arboreum
Piper macrostachyon Vahl =Peperomia macrostachya
Piper maculosum L. =Peperomia maculosa
Piper magnoliifolium Jacq. =Peperomia magnoliifolia
Piper maguirei Yunck. ='Piper paramaribense
Piper marginatum var. anisatum (Kunth) C. DC. =Piper marginatum
Piper marowinianum C. DC. =Piper kappleri
Piper marowynense C. DC. =Piper avellanum
Piper maypurense (Kunth) Poir. =Peperomia maypurensis
Piper medium Jacq. =Piper amalago
Piper miquelii C. DC. =Piper eucalyptifolium
Piper mollipilosum C. DC. [not in flora area]
Piper moraballianum Trel. =Piper avellanum
Piper myosuroides Rudge =Peperomia macrostachya
Piper nitidum Vahl [nom. illeg., non Piper nitidum Sw] =Piper divaricatum
Checklist of the Plants of the Guiana Shield 465
Piper nitidum Sw. =Piper arboreum
Piper mimmidarifolium Sw. =Peperomia rotundifolia
Piper obliquum var. eximium (Kunth) C. DC. =Piper cernuum
Piper obriimbatum (Miq.) C. DC. =Piper arboreum var. hirtellum
Piper obtusifolium L. =Peperomia obtusifolia
Piper oparumaense Yunck. =Piper obliquum
Piper orenocanum C. DC. =Piper coruscans
Piper orenocanum var. koriaboense Trel. & Yunck. =Piper coruscans
Piper parakegelianum Sagot ex Lemee [apparently a nomen nudum]
Piperparimanum Trel. =Piper bartlingianum
Piperparkerianum (Miq.) C. DC. =Piper anonifolium var. parkerianum
Piper pavasense Steyerm. =Piperjulianii
Piper pedunculare (Miq.) C. DC. [comb, illeg.]
Piper pellucidum L. =Peperomia pellucida
Piper perlongispicum Yunck. =Piper cernuum var. perlongispicum
Piper perlongivillosum Yunck. =Piperfrancovilleanum
Piper piresii Yunck. [not in flora area]
Piperpoiteanum Kunth =Piperfoveolatum
Piperpruinosum Kunth =Piper peltatum
Piper pseudochurumayu var. membranaceum C. DC. =Piper coruscans
Piper pseudomarowynense Trel. ex Lemee [apparently a nomen nudum]
Piper purpurascens Desf. ? =Piper angustifolium
Piper quadrangulare J.V. Thomps. =Peperomia quadrangularis
Piper reflexum L. f. =Peperomia tetraphylla
Piper rhododendrifolium Kunth =Piper anonifolium
Piper rhynchostachyum C. DC. =Piper eucalyptifolium
Piper rio-paraguanum Trel. ex V.M. Badillo [nomen nudum] =Piper hostmannianum
Piper romboutsii Yunck. =Piper divaricatum
Piper rotundifolium L. =Peperomia rotundifolia
Piper rubescens C. DC. =Piper demeraranum
Piper rugosum Vahl =Piper anonifolium
Piper ruizianum D. Dietr. =Piper obliquum
Piper sagoti C. DC. =Piper nematanthera var. sagotii
Piper saltuum C. DC. =Piper trigonum
Piper sancarlosianum C. DC. =Piper baccans var. sancarlosianum
Piper santa-martanum C. DC. =Piper aequale
Piper saramaccanum Yunck. =Piper obliquum
Piper scabrum Sw. =Piper hispidum
Piper serpens Sw. =Peperomia serpens
Piper smilacifolium Kunth =Piper reticulatum
Piper speciosum Kunth =Piper peltatum
Piper statarium Trel. & Yunck. =Piper avellanum
Piper subcrassifolium Yunck. =Piper hostmannianum
Piper submelanostictum var. amelanostictum Yunck. =Piper obliquum
Piper submelanostictum var. submelanostictum =Piper obliquum
Piper surinamense C. DC. =Piper consanguineum
Piper tenellum Sw. =Peperomia tenella
Piper tetraphyllum G. Forst. =Peperomia tetraphylla
Piper tiliifolium Ham. (Moraceae) =Bagassa guianensis
Piper tonoroanum Trel. & Yunck. =Piper avellanum
Piper tuberculatum var. minus C. DC. =Piper tuberculatum
Piper tuberculatum var. scandens Trel. & Yunck. =Piper tuberculatum
Piper variegatum Kunth =Piper cumanense
Piper verrucosum Sw. =Piper arboreum
Piper warakabacoura (Miq.) C. DC. =Piper bartlingianum
Piper wurdackii Yunck. =Piper coruscans
Piper zediogodiense Trel. ex L. Williams =Piper aequale
Pothomorphe peltata (L.) Miq. =Piper peltatum
Pothomorphe umbellata (L.) Miq. =Piper umbellatum
Quebitea guianensis Aubl. =Piper humistratum
Schilleria aequalis (Vahl) Kunth =Piper aequale
Schilleria caudata (Vahl) Kunth =Piper marginatum
Schilleria marginata (Jacq.) Kunth =Piper marginatum
Steffensia adunca (L.) Kunth =Piper aduncum
466 Checklist of the Plants of the Guiana Shield
Steffensia anonifolia Kunth =Piper anonifolium
Steffensia consanguinea Kunth =Piper consanguineum
Steffensia eximia Kunth =Piper cernuum
Steffensia hirsuta (Sw.) Kunth =Piper hispidum
Steffensia insignis Kunth =Piper insigne
Steffensia luschnathiana Kunth =Piper arboreum var. hirtellum
Steffensia obliqua (Ruiz & Pav.) Kunth =Piper obliquum
Steffensia pseudochiirumayu Kunth =Piper coruscans
References
Kramer, K.U. & A.R.A. Gorts-van Rijn. 1968. Piperaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname
1(2): 415-421.
Gorts-van Rijn, A. 2002. Piperaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 574-584.
Steyermark, J.A. 1984. Piperaceae. Flora de Venezuela 11(2): 1-619.
Steyermark, J.A. & R. Callejas-Posada. 2003. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan
Guayana 7: 681-738.
Tebbs, M.C. 1989. Revision of Piper in the New World. 1. Review of characters and taxonomy of Piper section Macrostachys.
Bull. Brit. Mus. Nat. Hist. (Bot.) 19: 117-158.
Tebbs, M.C. 1990. Revision of Piper in the New World. 2. The taxonomy of Piper section Churumayu. Bull. Brit. Mus. Nat. Hist.
(Bot.) 20(2): 193-236.
Tebbs, M.C. 1993. Revision of Piper in the New World. 3. The taxonomy of Piper sections Lepianthes and Radula. Bull. Nat.
Hist. Mus. 23(1): 1-50.
Trelease, W. & T.G. Yuncker. 1950. The Piperaceae of Northern South America. Urbana, Illinois: University of Illinois Press.
Yuncker, T.G. 1957. Piperaceae. In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 218-290.
Yuncker, T.G. 1972. The Piperaceae of Brazil. Hoehnea 2: 19-366.
Plantaginaceae
G. Rodriguez
Plantago lanceolata L. [introduced]
Plantago major L. [introduced] BO DA GU SU
FG
FG
References
D'Arcy, W.G 1971. Plantaginaceae. In R.E. Woodson, Jr., R.W. Schery, & collaborators, Flora of Panama. Ann. Missouri Bot.
Gard. 58(3): 363-369.
Lanjouw, J. 1938. Plantaginaceae. In A. Pulle, Flora of Suriname 4(2): 253-254.
Rodriguez, G. 2003. Plantaginaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 7: 739.
Plumbaginaceae
C.A. Vargas
Plumbago auriculata Lam. [widely cultivated, perhaps escaping]
Plumbago zeylanica L. [naturalized] BO DA GU? SU FG
Synonyms and Misidentifications
Plumbago scandens L. ^Plumbago zeylanica
References
Vargas, C.A. 2003. Plumbaginaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 7: 740-741.
PODOSTEMACEAE
RE. Berry
Apinagia arminensis P. Royen
Apinagia brevicaulis Mildbr.
Apinagia corymbosa (Tul.) Engl.
Apinagia corymbosa var. corymbosa
Apinagia crispa P. Royen
Apinagia digitata P. Royen
Apinagia divertens Went
SU FG
BO
BO GU SU FG
BO GU SU
SU
FG
FG
SU FG
Checklist of the Plants of the Guiana Shield 467
Apinagia exilis (Tul.) P. Royen
Apinagiaflexuosa (Tul.) P. Royen
Apinagia goejei Went
Apinagia guyanensis (Pulle) P. Royen
Apinagia hulkiana (Went) P. Royen
Apinagia imthurnii (Goebel) P. Royen
Apinagia itanensis Schnell
Apinagia kochii (Engl.) P. Royen
Apinagia latifolia (Goebel) P. Royen
Apinagia leptophylla (Goebel) P. Royen
Apinagia longifolia (Tul.) P. Royen
Apinagia marowynensis (Went) P. Royen
Apinagia midtibranchiata (Matthiesen) P. Royen
Apinagia nana Went [=Apinagia pilgeril]
Apinagia penicillata (P. Royen) P. Royen
Apinagia petiolata Hollander
Apinagia pilgeri Mildbr.
Apinagia pusilla Tul
.
Apinagia richardiana (Tul.) P. Royen
Apinagia ruppioides (Kunth) Tul.
Apinagia secundiflora (Tul.) Pulle
Apinagia staheliana (Went) P. Royen
Apinagia suritmuensis (Engl.) P. Royen
Apinagia treslingiana (Went) P. Royen
Apinagia versteegiana (Went) P. Royen
Jenmaniella ceratophylla Engl.
Jenmaniella ceratophylla var. ceratophylla
Jenmaniella ceratophylla var. hexandra Engl.
Jenmaniella ceratophylla var. parva P. Royen
Jenmaniella guianensis Engl.
Jenmaniella isoetifolia P. Royen
Jenmaniellajenmanii (Engl.) P. Royen
Jenmaniella tridactylitifolia Engl.
Jenmaniella varians Engl.
Marathrum aeruginosum P. Royen
Marathrum capillaceum (Pulle) P. Royen
Marathrum pauciflorum Tul.
Marathrum pauciflorum var. heterophyllum P. Royen
Marathrum pauciflorum var. pauciflorum
Marathrum squamosum Wedd.
Marathrum squamosum var. squamosum
Marathrum utile Tul.
Mourerafluviatilis Aubl.
Oserya flabellifera Tul. & Wedd. [not in flora area?]
Oserya minima P. Royen
Oserya perpusilla (Went) P. Royen
Oserya sphaerocarpa Tul. & Wedd.
Podosiemum dichotomum Klotzsch [dubious species]
Rhyncholacis apiculata P. Royen
Rhyncholacis applanata K.I. Goebel
Rhyncholacis applanata var. applanata
Rhyncholacis applanata var. laxipinnata P. Royen
Rhyncholacis brassicifolia P. Royen
Rhyncholacis brevistamina P. Royen
Rhyncholacis coronata P. Royen
Rhyncholacis cristata P. Royen
Rhyncholacis dentata P. Royen
Rhyncholacis divaricata Matthiesen
Rhyncholacis flagellifolia P. Royen
Rhyncholacis guyanensis P. Royen
Rhyncholacis hydrocichorium Tul.
Rhyncholacisjenmanii Engl.
Rhyncholacis linearis Tul.
Rhyncholacis macrocarpa Tul. [dubious species]
BO GU su
GU su
su
EG
BO
GU
su
su
EG
GU su EG?
EG
BO
GU
GU
EG
AM BO DA GU su EG
su EG?
AM BO
GU
su
su
su
su
AM BO DA GU su EG
AM BO
GU su EG
AM BO
BO
BO
GU
GU
GU
GU
GU
GU
GU
GU
GU
,
GU
su
su
su
AM
BO
GU
GU
GU
su EG
AM FG
AM
AM
AM BO GU su
su
su
EG
BO GU
GU
GU
GU
su FG
AM BO GU
AM BO
BO
GU
GU
GU?
GU
GU
su
su
AM BO
BO
BO
GU
GU
GU
GU
GU su?
468 Checklist of the Plants of the Guiana Shield
Rhyncholacis oligandra Wedd.
Rhyncholacis oligandra var. oligandra
Rhyncholacis oligandra var. tenella P. Royen
Rhyncholacis palmettifolia P. Royen
Rhyncholacis palmettifolia P. Royen var. palmettifolia
Rhyncholacis palmettifolia var. rosea P. Royen
Rhyncholacis penicillata Matthiesen
Tristicha trifaria (Bory ex Willd.) Spreng.
Weddellina sqiiamulosa Tul.
AM
AM
AM
AM
BO
BO DA
BO
GU
GU
GU
GU
GU
GU
GU
GU SU
FG
FG
Synonyms and Misidentifications
Apinagiaflabellifera (Tul. & Wedd.) Went =Oserya flabellifera
Apinagia perpusilla Went =Oserya perpusilla
Dufourea trifaria Bory ex Willd. =Tristicha trifaria
Ligeaflexuosa Tul. =Apinagiaflexuosa
Ligea longifolia (Tul.) Tul. =Apinagia longifolia
Ligea richardiana Tul. =Apinagia richardiana
Ligea richardiana var. corymbosa Tul. =Apinagia corymbosa
Ligea richardiana var. exilis Tul. =Apinagia exilis
Ligea secundiflora Tul. =Apinagia secundiflora
Lophogyne capillacea Pulle =Marathrum capillaceum
Marathrum squamosum var. spruceanum Wedd. =Marathrum squamosum var. squamosum
Neolacis corymbosa (Tul.) Wedd. =Apinagia corymbosa
Neolacis corymbosa var. exilis Tul. =Apinagia exilis
Neolacis secundiflora (Tul.) Wedd. =Apinagia secundiflora
Oenoneflexuosa (Tul.) Wedd. =Apinagiaflexuosa
Oenone guyanensis Pulle =Apinagia guyanensis
Oenone hulkiana Went =Apinagia hulkiana
Oenone imthurnii Goebel =Apinagia imthurnii
Oenone kochii Engl. =Apinagia kochii
Oenone longifolia Tul. =Apinagia longifolia
Oenone marowynensis Went =Apinagia marowynensis
Oenone multibranchiata Matthiesen =Apinagia multibranchiata
Oenone penicillata P. Royen =Apinagia penicillata
Oenone richardiana (Tul.) Warm. =Apinagia richardiana
Oenone secundiflora (Tul.) Engl. =Apinagia secundiflora
Oenone staheliana Went =Apinagia staheliana
Oenone treslingiana Went =Apinagia treslingiana
Oenone versteegiana Went =Apinagia versteegiana
Podostemum ruppioides Kunth =Apinagia ruppioides
Tristicha hypnoides (A. St.-Hil.) Spreng. =Tristicha trifaria
References
Berry, RE. 2004. Podostemaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan Guayana
8:301-316.
Den Hollander, G. & C.C. Berg. 1983. Studies on the flora oftheGuianas 4. Anew species ofApinagia and a key to the Apinagia
species in Suriname. Acta Bot. Neerl. 32(3): 223-226.
Philbrick, C.T. 2002. Podostemaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 585.
Royen, R van. 1951-1954. The Podostemaceae of the New World. Med. Bot. Mus. Herb. Rijks Univ. Utrecht 107: 1-151 (1951);
115: 1-21(1953); 119:215-263.
Schnell, R. 1969. Contribution a I'etude des Podostemacees de Guyane. Adansonia ser. 2 9(2): 249-271.
POLYGALACEAE
G.A. Aymard C., P.E. Berry, B. Eriksen
Barnhartiafloribunda Gleason
Bredemeyera altissima (Poepp.) A.W. Benn.
Bredemeyera bracteata Klotzsch ex Hassk.
Bredemeyera densiflora A.W. Benn.
Bredemeyera densiflora var. densiflora
Bredemeyera densiflora var. glabra A.W. Benn.
Bredemeyerafloribunda Willd.
BO GU SU
BO GU
GU
BO GU SU
GU SU?
BO GU SU
BO GU
FG
Checklist of the Plants of the Guiana Shield 469
Bredemeyera lucida (Benth.) Klotzsch ex Hassk.
Bredemeyera myrtifolia K^N. Benn.
Bredemeyera sp. A
Diclidanthera bolivarensis Pittier
Diclidanthera octandra Gleason
Diclidanthera sp. A
Diclidanthera wurdackiana Aymard & RE. Berry
Monnina cacumina N.E. Br.
Moutabea aculeata (Ruiz & Pav.) Roepp. & Endl.
Moutabea aff. chodatiana Huber
Moutabea guianensis Aubl.
Moutabea sp. A
Moutabea sp. B
Polygala adenophora DC.
Polygala appressa Benth.
Polygala asperuloides Kunth
Polygala blakeana Steyerm.
Polygala brevialata Chodat
Polygala caracasana Kunth
Polygala coelosioides Mart.
Polygala echinosperma Gorts
Polygala galioides Poir.
Polygala glochidiata Kunth
Polygala glochidiata var. glochidiata
Polygala hebeclada DC.
Polygala hygrophila Kunth [expected in FG]
Polygala longicaulis Kunth
Polygala martiana A.^. Benn.
Polygala membranacea (Miq.) Gorts
Polygala microspora S.F. Blake
Polygala paniculata L.
Polygala planellasi Molinet & M. Gomez
Polygala pseudocoelosioides Chodat
Polygala sanariapoana Steyerm.
Polygala savannarum Chodat
Polygala sipapoana Wurdack
Polygala spectabilis DC.
Polygala spruceana A.W. Benn.
Polygala subtilis Kunth
Polygala tenella Willd.
Polygala timoutoides Chodat
Polygala timoutoides var. maguirei (Wurdack) Marques
Polygala timoutou Aubl.
Polygala trichosperma L.
Polygala violacea Aubl., emend. Marques
Securidaca bialata Benth.
Securidaca cacumina Wurdack
Securidaca cayennensis S.F. Blake
Securidaca coriacea Bonpl.
Securidaca divaricata Nees & Mart.
Securidaca diversifolia (L.) S.F. Blake
Securidaca fruticans Wurdack
Securidaca lateralis A.W. Benn.
Securidaca longifolia Poepp.
Securidaca macrophylla (Benth.) Wurdack
Securidaca maguirei Wurdack
Securidaca marginata Benth.
Securidaca paniculata Rich.
Securidaca paniculata var. lasiocarpa Oort
Securidaca paniculata var. paniculata
Securidaca pendula Bonpl.
Securidaca prancei Wurdack
Securidaca pubescens DC.
Securidaca pubiflora Benth.
AM BO DA GU su FG
AM
BO
AM BO
AM
AM
AM BO
AM BO GU
AM BO GU FG
AM BO
AM BO GU su FG
AM BO DA
AM
AM BO GU su FG
AM BO
BO
GU su FG
AM BO GU su FG
AM BO
BO
GU?
su FG
BO GU su FG
BO GU
BO GU
GU?
AM BO GU su
AM BO GU su
su
FG
FG
FG
AM BO
BO
GU
GU
GU
FG'
AM
AM BO GU
AM
AM BO GU su FG
AM BO GU
AM BO GU su
AM BO GU su FG
AM BO GU
AM BO GU
AM BO GU su FG
AM BO GU su FG
AM
AM
BO
BO DA
GU su FG
AM BO
FG
AM BO DA GU su
AM GU su FG
AM BO DA GU su FG
AM
AM
AM
BO
GU
GU
GU
FG
AM BO DA GU su FG
AM BO DA GU su FG
BO DA GU su FG
AM BO GU
AM BO
AM BO GU
GU
su FG
470 Checklist of the Plants of the Guiana Shield
Securidaca pyramidalis Sprague ex Sandwith
Securidaca retusa Benth.
Securidaca savannarum Wurdack
Securidaca scandens Jacq.
Securidaca speciosa Wurdack
Securidaca spinifex Sandwith
Securidaca tenuifolia Chodat
Securidaca tomentosa A. St.-Hil. & Moq.
Securidaca uniflora Oort
Securidaca volubilis L., emend. Oort
Securidaca warmingiana Chodat
Synonyms and Misidentifications
Acosta aculeata Ruiz & Pav. =Moutabea aculeata
Bredemeyera lucida (Benth.) A.W. Benn. =Bredemeyera lucida
Bredemeyera parviflora A.W. Benn. =Bred^meyera myrtifolia
Catocoma altissima Poepp. & Endl. ^Bredemeyera altissima
Catocomafloribunda Benth. =Bredemeyerafloribunda
Catocoma lucida Benth. =Bredemeyera lucida
Diclidanthera orinocensis Pittier =Diclidanthera bolivarensis
Monnina cacumina var. duidae (S.F. Blake) Steyerm. =Monnina cacumina
Monnina duidae S.F. Blake =Monnina cacumina
Monnina membranacea Miq. =Polygala membranacea
Monnina uaipanensis Wurdack =Monnina cacumina
Moutabea longifolia Poepp. & Endl. =Moutabea aculeata
Polygala adenophora var. gracilis Chodat =Polygala adenophora
Polygala adenophora var. robusta Chodat =Polygala adenophora
Polygala alopecurus Chodat =Polygala tenella
Polygala angustifolia Kunth =Polygala violacea
Polygala angustifolia var. latifolia A. St.-Hil. ex Chodat =Polygala violacea
Polygala appressa subsp. cumbrensis Wurdack =Polygala appressa
Polygala appressa var. gracillima (S. Watson) A.W. Benn. =Polygala appressa
Polygala appressa var. kavanayena Steyerm. =Polygala appressa
Polygala aristeguietae Wurdack =Polygala brevialata
Polygala brizoides A. St.-Hil. =Polygala violacea
Polygala camporum Benth. =Polygala violacea
Polygala columbica Chodat =Polygala caracasana
Polygala diversifolia L. =Securidaca diversifolia
Polygalafendleri Chodat =Polygala exigua yar. fendleri
Polygalafunckii Chodat =Polygala tenella
Polygala gracilis Kunth =Polygala tenella
Polygala incarnata Aubl. =Polygala adenophora
Polygala leptocaulis Torr. & A. Gray =Polygala tenella
Polygala maguirei Wurdack =Polygala timoutoides var. maguirei
Polygala mollis Kunth =Polygala violacea
Polygala monticola Kunth =Polygala violacea
Polygala monticola var. brizoides (A. St.-Hil.) Steyerm. =Polygala violacea
Polygala paludosa A. St.-Hil. & Moq. [not in flora area] =Polygala tenella
Polygala peduncularis A. Rich. =Polygala planellasi
Polygala retifera S.F. Blake =Polygala caracasana
Polygala sororia Miq. =Polygala trichosperma
Polygala stipulata Chodat =Polygala membranacea
Polygala subsecunda S.F. Blake =Polygala brevialata
Polygala variabilis Kunth =Polygala trichosperma
Polygala violacea Vahl, non Aubl. =Polygala martiana
Securidaca complicata Kunth =Securidaca pendula
Securidaca cordata J.R. Johnst. =Securidaca scandens
Securidaca corytholobium A.W. Benn. =Securidaca macrophylla
Securidaca densiflora Oort =Securidaca pubescens
Securidaca ecristata Wurdack =Securidaca cacumina
Securidaca fendleri Chodat =Securidaca diversifolia
Securidaca hostmannii Miq. =Securidaca paniculata var. lasiocarpa
Securidaca latifolia Benth. =Securidaca pubiflora
Securidaca major Sagot ex A.W. Benn. =Securidaca pubescens
AM BO GU
AM BO GU
AM
BO
AM BO
GU SU
FG
FG
AM
AM
AM
BO
BO
FG?
GU SU FG
GU SU FG
GU?
Checklist of the Plants of the Guiana Shield 471
Securidaca mollis Kunth =Securidaca coriacea
Secuhdaca orinocensis Rusby =Securidaca pendula
Securidaca rivinifolia A. St.-Hil. & Moq. =Securidaca divaricata
Securidaca rivinifolia van parvifolia A.W. Benn. =Securidaca divaricata
Securidaca venosa Rusby =Securidaca diversifolia
References
Aymard C.,G.A, RE. Berry, and B. Eriksen. 2004. Polygalaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 8: 316-347
Chodat, R. 1893. Monographia Polygalacearum. Mem. Soc. phys. et d'hist. nat. Geneve 31 part 2(2): 1-500.
Gorts-van Rijn, A.R.A. 1974. Notes on Polygalaceae from Suriname. Acta Bot. Neerl. 23(2): 189-191.
Gorts-van Rijn, A.R.A. 1976. Polygalaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2):
518-523.
Jacobs-Brouwer, A. 2002. Polygalaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 585-589.
Marques, M.d.C.M. 1979. Revisao das especies do genero Polygala L. (Polygalaceae) do estado do Rio de Janeiro. Rodriguesia
31(48): 69-339.
Marques, M.d.C.M. 1980. Revisao das especies do genero Bredemeyera Willd. (Polygalaceae) do Brasil. Rodriguesia 32(54):
269-321.
Oort,A.J.R 1939. Polygalaceae. In A. Pulle, Flora of Suriname 2(1): 406-425.
POLYGONACEAE
Aymard C., GA. J.Brandbyge, R.A. Howard
Antigonon leptopus Hook. & Am. [cultivated, naturalized]
Coccoloba acuminata Kunth
Coccoloba ascendens Duss ex Lindau
Coccoloba caracasana Meisn.
Coccoloba charitostachya Standi.
Coccoloba conduplicata Maguire
Coccoloba coronata Jacq.
Coccoloba cruegeri Lindau
Coccoloba declinata (Veil.) Mart.
Coccoloba densifrons Mart, ex Meisn.
Coccoloba dugandiana A. Fernandez
Coccoloba excelsa Benth.
Coccoloba fallax Lindau
Coccoloba gymnorrhachis Sandwith
Coccoloba latifolia Poir.
Coccoloba llewelynii R.A. Howard
Coccoloba lucidula Benth.
Coccoloba marginata Benth.
Coccoloba mollis Casar.
Coccoloba obtusifolia Jacq.
Coccoloba ochreolata Wedd.
Coccoloba orinocana R.A. Howard
Coccoloba ovata Benth.
Coccoloba padiformis Meisn.
Coccoloba savannarum Standi.
Coccoloba schomburgkii Meisn.
Coccoloba sp. A
Coccoloba sp. B
Coccoloba sp. C
Coccoloba sp. D
Coccoloba spruceana Lindau
Coccoloba striata Benth.
Coccoloba uvifera (L.) L.
Coccoloba wiirdackii R.A. Howard
Polygonum acuminatum Kunth
Polygonumferrugineum Wedd.
Polygonum glabrum Willd.
Polygonum hydropiperoides Michx.
Polygonum meisnerianum Cham. & Schltdl.
AM BO
BO
DA GU
GU
SU FG
AM BO DA GU SU FG
AM
BO
BO
GU
SU
GU SU
AM BO
BO
DA
GU
SU FG
AM BO
AM BO DA GU SU FG
AM BO DA
BO GU SU FG
AM BO DA GU SU FG
AM BO
BO GU SU FG
AM BO DA GU SU FG
AM BO SU FG
AM BO
AM BO
AM BO
AM BO GU
GU?
GU
SU FG
AM BO GU
AM
AM
AM
BO
AM DA
AM BO GU
AM GU SU FG
AM
AM BO DA
DA
GU SU FG
FG
AM DA GU SU FG
AM DA GU SU FG
FG
FG
GU SU FG
GU su FG
GU
BO GU
AM BO
BO DA GU
AM BO DA GU SU
AM BO
BO
GU su
472 Checklist of the Plants of the Guiana Shield
Polygonum meisnerianum var. beyrichianum (Cham. & Schltdl.) Meisn.
Polygonum punctatum Elliott AM DA
Ruprechtia brachysepala Meisn.
Ruprechtia brachystachya Meisn.
Ruprechtia cniegeh Griseb. ex Lindau [expected in Guyana]
Ruprechtia howardiana Aymard & RE. Berry BO
Ruprechtia laxiflora Meisn. AM
Ruprechtia ramiflora (Jacq.) C.A. Mey.
Ruprechtia tangarana Standi.
Ruprechtia tenuiflora Benth.
Symmeria paniculata Benth.
Triplaris americana L. SU FG
Triplaris caracasana Cham.
Triplaris cumingiana Fisch. & C.A. Mey. ex C.A. Mey. AM
Triplaris weigeltiana (Rchh.) Kuntze BO DA GU SU FG
Synonyms and Misidentifications
Blochmannia weigeltiana Rchb. =Triplaris weigeltiana
Coccoloba bolivarana LI. Williams [nomen nudum] =Coccoloba llewelynii
Coccoloba bracteolosa Meisn. =Coccoloba excelsa
Coccoloba caribaea Urb. =Coccoloba coronata
Coccoloba caurana Standi. =Coccoloba fallax
Coccoloba cyclophylla S.F. Blake =Coccoloba caracasana
Coccoloba excoriata L. [not in flora area] =Coccoloba tenuifolia L.
Coccoloba grandifolia Jacq. [not in flora area] =Coccoloba pubescens L.
Coccoloba grandis Benth. =Coccoloba latifolia
Coccoloba guianensis Meisn. ^Coccoloba marginata
Coccoloba micropunctata Eyma =Coccoloba excelsa
Coccoloba novogranatensis Lindau =Coccoloba coronata
Coccoloba paraensis Meisn. =Coccoloba excelsa
Coccoloba parimensis Benth. =Coccoloba excelsa
Coccoloba parimensis var. schomburgkii Meisn. ^Coccoloba excelsa
Coccoloba pittieri R. Knuth ex Pittier =Coccoloba striata
Coccoloba polystachya Wedd. =Coccoloba mollis
Coccoloba sagotii Lindau =Coccoloba lucidula
Coccoloba sparsifolia Lindau =Coccoloba ascendens
Coccoloba trinitatis Lindau =Coccoloba marginata
Persicaria acuminata (Kunth) M. Gomez =Polygonum acuminatum
Persicaria densiflora (Meisn.) Moldenke [s.l.] =Polygonum glabrum
Persicaria glabra (Willd.) M. Gomez =Polygonum glabrum
Persicaria hydropiperoides (Michx.) Small =Polygonum hydropiperoides
Persicaria meisneriana (Cham. & Schltdl.) M. Gomez =Polygonum meisnerianum
Persicaria punctata (Elliott) Small =Polygonum punctatum
Polygonum acuminatum var. glabrecens Meisn. =Polygonum acuminatum
Polygonum acuminatum var. weddelii Meisn. ^Polygonum acuminatum
Polygonum declinatum Veil. =Coccoloba declinata
Polygonum densiflorum Meisn. =Polygonum glabrum
Polygonum portoricense Bertero ex Small [nom. illeg.] =Polygonum glabrum
Ruprechtia amentacea Meisn. =Ruprechtia tenuiflora
Ruprechtia concina Pittier =Ruprechtia ramiflora
Ruprechtia coriacea (H. Karst.) S.F. Blake =Ruprechtia ramiflora
Ruprechtia hamanii S.F. Blake =Ruprechtia ramiflora
Ruprechtia marowynensis Eyma =Ruprechtia brachysepala
Triplaris americana Aubl., non L. =Triplaris weigeltiana
Triplaris caracasana var. vargasii Meisn. =Triplaris caracasana
Triplaris coriacea H. Karst. =Ruprechtia ramiflora
Triplaris felipensis Wedd. =Triplaris americana
Triplaris pavonii Meisn. =Triplaris americana
Triplaris ramiflora Jacq. =Ruprechtia ramiflora
Triplaris schomburgkiana Benth. =Triplaris americana
Triplaris surinamensis Cham. =Triplaris weigeltiana
Triplaris surinamensis var. benthamiana Meisn. =Triplaris weigeltiana
Triplaris surinamensis var. chamissoana Meisn. =Triplaris weigeltiana
Triplaris surinamensis var. crassifolia Benth. =Triplaris weigeltiana
Checklist of the Plants of the Guiana Shield 473
Triplaris vahliana Fisch. & C.A. Mey. ex C.A. Mey. =Triplaris weigeltiana
References
Aymard C.,G.A. and R.A. Howard. 2004. Polygonaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of
the Venezuelan Guayana 8: 347-370.
Brandbyge, J. 1986. A revision of the genus Triplaris (Polygonaceae). Nord. J. Bot. 6(5): 545-570.
Brandbyge, J. 2002. Polygonaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
590-592.
Cocucci, A.E. 1961. Revision del genero Ruprechtia (Polygonaceae). Kurtziana 1: 217-269.
Eyma, P.J. 1934. Polygonaceae. In A. Pulle, Flora of Suriname 1(1): 49-71.
Howard, R.A. 1960. Studies in the genus Coccoloba, IX. A conspectus of the South American species. J. Am. Arb. 41: 213-258,
357-390.
Howard, R.A. 1961. Studies in the genus Coccoloba, X. New species and a summary of distribution in South America. J. Am.
Arb. 42: 87-95.
Lindeman, J.C. & A.R.A. Gorts-van Rijn. 1968. Polygonaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of
Suriname 1(2): 303-309.
Wilson, K.L. 1990. Some widespread species of Persicaria and their allies. Kew Bull. 45: 621-636.
J.A. Steyermarkt
AM
BO
BO
BO
BO
AM BO DA GU SU FG
AM BO DA GU su? FG
AM BO DA
AM
GU
FG?
AM BO DA GU SU FG
AM BO DA FG?
AM
BO
BO
GU
GU?
GU
su
su?
FG
FG
Portulaca elutior Mart, ex Rohrb.
Portulaca halimoides L.
Portulaca insignis Steyerm.
Portulaca mucronata Link
Portulaca oleracea L. [introduced]
Portulaca pilosa L.
Portulaca pusilla Kunth
Portulaca pygmaea Steyerm.
Portulaca quadrifida L. [introduced?]
Portulaca rubricaulis Kunth
Portulaca sedifolia N.E. Br.
Portulaca teretifolia Kunth
Portulaca umbraticola Kunth
Talinum fruticosum (L.) Juss.
Talinum paniculatum (Jacq.) Gaertn.
Synonyms and Misidentifications
Portulaca cayennensis D. Legrand =Portulaca sedifolia
Portulaca cinerea Poelln. =Portulaca teretifolia
Portulacafruticosa L. =Talinum fruticosum
Portulaca lanata Rich. =Portulaca pilosa
Portulaca marginata Kunth =Portulaca oleracea
Portulaca paniculata Jacq. =Talinum paniculatum
Portulaca portidacastrum L. (Aizoaceae) =Sesuvium portulacastrum
Portulaca triangularis Jacq. =Talinum fruticosum
Talinum triangulare (Jacq.) Willd. =Talinumfruticosum
References
Legrand, CD. 1962. Las especies americanas de Portulaca. An. Mus. Hist. Nat. Montevideo (ser. 2) 7: 1-147.
Ooststroom, S.J. van. 1943. Portulacaceae. In A. Pulle, Flora of Suriname 1(1): 486-491.
Steyermark, J.A. 2005. Portulacaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Holsl, Flora of the Venezuelan
Guayana 8: 376-383.
Primulaceae
PE. Berry, J.S. Miller
Anagallis pumila Sw. AM
Synonyms and Misidentifications
Centunculus pentandrus R. Br. ^Anagallis pumila
Centunculus pumilus (Sw.) Kuntze =Anagallis pumila
Micropyxis pumila (Sw.) Duby =Anagallis pumila
BO
474 Checklist of the Plants of the Guiana Shield
References
Miller, J.S. and RE. Berry. 2004. Primulaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan
Guayana 8: 383-383.
Proteaceae
J.A. Steyermarkt
BO
BO
AM
BO
AM BO GU su FG
BO GU su FG
BO
BO
AM
BO GU
AM BO DA GU su FG
FG
AM GU FG
AM
BO
su
AM BO GU
Euplassa chimantensis Steyerm. BO
Euplassa glaziovii (Mez) Steyerm. [Mt. Roraima]
Euplassa pinnata {Ldim.)\.M. ]o)\n?,\.. GU SU FG
Euplassa venezuelana Steyerm.
Panopsis ornatinervia Steyerm.
Panopsis parimensis Steyerm.
Panopsis ptariana Steyerm.
Panopsis rubescens (Pohl) Rusby
Panopsis sessilifolia (Rich.) Sandwith
Panopsis tepuiana Steyerm.
Roupala chimantensis Steyerm.
Roupala ghotii Steyerm.
Roupala minima Steyerm.
Roupala montana Aubl.
Roupala nitida Rudge
Roupala obtusata Klotzsch
Roupala paruensis Steyerm.
Roupala schulzii Mennega
Roupala sororopana Steyerm.
Roupala suaveolens Klotzsch
Synonyms and Misidentifications
Adenostephanus guianensis Meisn. =Euplassa pinnata
Andriapetalum cayennensis Klotzsch ex Meisn. =Panopsis sessilifolia
Andriapetalum rubescens Pohl ^Panopsis rubescens
Andriapetalum sessilifolium (Rich.) Klotzsch =Panopsis sessilifolia
Euplassa meridionalis Salisb. =Euplassa pinnata
Panopsis cuaensis Steyerm. =Panopsis rubescens
Panopsis hameliifolia (Rudge) Salisb. =Panopsis sessilifolia
Panopsis rubescens var. simulans J.F. Macbr. =Panopsis rubescens
Panopsis schomburgkii (Klotzsch) Pittier =Roupala suaveolens
Roupala complicata Kunth =Roupala montana
Roupala dentata R. Br. =Roupala montana
Roupala dissimilis Pittier =Roupala montana
Roupala hameliifolia Rudge =Panopsis sessilifolia
Roupala laurifolia Salisb. & Knight =Roupala nitida
Roupala macropoda Klotzsch & H. Karst. =Roupala montana
Roupala media R. Br. =Roupala montana
Roupala montana B complicata (Kunth) Griseb. =Roupala montana
Roupala montana var. dentata (R. Br.) Sleumer =Roupala montana
Roupala pinnata Lam. =Euplassa pinnata
Roupala pullei Mennega =Roupala suaveolens
Roupala pyrifolia Salisb. & Knight =Roupala montana
Roupala schomburgkii Klotzsch =Roupala suaveolens
Roupala sessilifolia Rich. =Panopsis sessilifolia
References
Mennega, A.M.W. 1934. Proteaceae. In A. Pulle, Flora of Suriname 1(1): 154-157.
Mennega, A.M.W. 1968. Proteaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of Suriname 1(2): 315-320.
Sleumer, H.O. 1954. Proteaceae americanae. Bot. Jahrb. 76: 139-211.
Plana, V. 2002. Proteaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 592-
594.
Steyermark, J.A. 2004. Proteaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 8: 384-393.
GU?
GU su FG
GU su FG
Checklist of the Plants of the Guiana Shield 475
QUIINACEAE
J.V. Schneider, G. Zizka
Froesia gereaitana J.V. Schneid. & Zizka AM AM
Froesia thcarpa Pires AM
Froesia veneznelensis Steyerm. & G.S. Bunting
Lacunaria crenata (Tul.) A.C. Sm. [expected in lowlands ofBO or AM]
Lacunariajenmanii (Oliv.) Ducke AM
Lacunaria macrostachya (Tul.) A.C. Sm. AM BO
Lacunaria oppositifolia Pires AM
Lacunaria panamensis (Standi.) Standi. SU
Lacunaria umbonata Pires FG
Quiina amazonica A.C. Sm.
Quiina attenuata J.V. Schneid & Zizka
Quiina brevensis Pires
Quiina congesta R.S. Cowan
Quiina cruegeriana Griseb.
Quiina florida Tul.
Quiina guianensis Aubl.
Quiina indigofera Sandwith
Quiina integrifolia Pulle
Quiina leptoclada Tul.
Quiina longifolia Spruce ex Planch. & Triana
Quiina macrophylla Tul.
Quiina obovata Tul.
Quiina oiapocensis Pires
Quiina parvifolia Lanj. & Heerdt
Quiina pteridophylla (Radlk.) Pires
Quiina rhytidopus Tul.
Quiina sessilis Choisy
Quiina tinifolia Planch. & Triana
Quiina wurdackii Pires
Quiina yatuensis J.V. Schneid. & Zizka
Touroulia amazonica Pires & A.S. Foster [Sierra de la Neblina]
Touroulia guianensis Aubl.
Synonyms and Misidentifications
Lacunaria silvatica (Pulle) A.C. Sm. =Lacunariajenmanii
Lacunaria spruceana (Engl.) Pires =Lacunaria oppositifolia
Quiina acutangula Ducke =Quiina pteridophylla
Quiina albiflora A.C. Sm. =Quiina obovata
Quiina crenata Tul. =Lacunaria crenata
Quiina decaisneana Planch. & Triana =Lacunaria sp. jenmanii?
Quiina gracilis A.C. Sm. =Quiina tinifolia
Quiina macrostachya Tul. =Lacunaria macrostachya
Quiina oblanceolata Sandwith =Quiina obovata
Quiina poeppigiana Tul. =Quiina florida
Quiina silvatica Pulle =Lacunariajenmanii
Quiina spruceana Engl. ^Lacunaria oppositifolia
Tourouliajenmanii Oliv. "^Lacunariajenmanii
Touroulia pteridophylla Radlk. =Quiina pteridophylla
References
Lanjouw, J. & P.F. van Heerdt. 1941. Quiinaceae. In A. Pulle, Flora of Suriname 3(1): 355-365.
Mori, S.A. 2002. Quiianaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. HotT, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 594-
598.
Schneider, J. and G. Zizka. 1997. Two new species of Quiianaceae (Quiina, Froesia) from the Venezuelan Guayana, and some
remarks on the genus Froesia Pires.
Schneider, J.V. & G. Zizka. 2004. Quiinaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 8: 393-407.
GU?
AM
GU
SU
AM BO GU SU FG
AM
BO DA GU SU FG
BO DA GU
SU
SU
FG
FG
AM
AM FG
BO GU SU FG
AM BO GU
SU
FG
AM BO GU SU FG
AM BO GU SU
FG
AM
AM BO
AM AM
AM
AM BO GU SU FG
476 Checklist of the Plants of the Guiana Shield
Rafflesiaceae
W. Meijerf, G. Yatskievych
Apodanthes caseariae Poit. BO GU SU FG
Apodanthes surinamensis Pulle SU
Synonyms and Misidentifications
Apodanthesflacourtiae H. Karst. =Apodanthes caseariae
Apodanthes roraimae Vattimo-Gil =Apodanthes caseariae
References
Gentry, A.H. 1973. Rafflesiaceae. In R.E. Woodson et al., Flora of Panama. Ann. Missouri Bot. Gard. 60: 17-21.
Gomez, L.D. 1983. Rafflesiaceae. In Flora Costaricensis, edited by W. Burger. Fieldiana, Bot. n.s. 13: 89-93.
Meijer, W. 1993. Rafflesiaceae. In The Families and Genera ofVascular Plants, Vol. 2, edited by K. Kubitzi, J.G. Rohwer, and V.
Bittrich, pp. 557-563. Berlin: Springer-Verlag.
Mori, S.A. 2002. Rafflesiaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
598.
Vattimo-Gil, I. de. 1956. Notes on Apodanthes caseariae Poit. and Pilostyles calliandrae (Gardn.) R. Br. Not. Syst. 15(2): 225-
229.
Vattimo-Gil, 1. de. 1971. Contribui9ao ao conhecimento de tribo Apodanthea (sic) R. Br. Parte I - Conspecto das especies.
Rodriguesia 26(38): 37-62.
Vattimo-Gil, I. de. 1973. Notas sobre o genero Apodanthes Poit. com descrifao de duas novas especies. Rev. Brasil. Biol. 33:
135-141.
Vattimo-Gil, I. de. 1978. Uma nova especie de Apodanthes Pot. (Rafflesiaceae). Rodriguesia 29: 47-51. 1978.
Yatskievych, G. & W. Meijer. 2004. Rafflesiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 8: 407-410.
Ranunculaceae
RE. Berry
Clematis dioica var. brasiliana (DC.) Eichler [not in flora area, expected in FG]
Clematis guadeloupae Pers. BO
Clematis guadeloupae var. guadeloupae [Expected in GU] BO
Clematis populifolia Turcz. BO
Synonyms and Misidentifications
Clematis caracasana Humb. & Bonpl. ex DC. =Clematis guadeloupae var. guadeloupae
Clematis caripensis Kunth =Clematis guadeloupae var guadeloupae
Clematis floribunda Triana & Planch. =Clematis populifolia
Clematis goudotiana Triana & Planch. =Clematis populifolia
References
Berry, RE. 2004. Ranunculaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan Guayana
8:410-413.
Lourteig, A. 1952. Ranunculaceas de Sudamerica templada. Darwiniana 9: 397-609.
Lourteig, A. 1971. Ranunculaceae. In T. Lasser, Flora de Venezuela 3(1): 29-51.
Rhabdodendraceae
Rhabdodendron amazonicum (Spruce ex Benth.) Huber GU SU FG
Synonyms and Misidentifications
Lecostemon amazonicum Spruce ex Benth. =Rhabdodendron amazonicum
Lecostemon sylvestre Gleason =Rhabdodendron amazonicum
Lecostemon terniflorum M09. & Sesse ex DC. (Elaeocarpaceae) =Sloanea terniflora
Rhabdodendron crassipes (Spruce ex Benth.) Huber =Rhabdodendron amazonicum
Rhabdodendron crassipes var. cayenense Benth. =Rhabdodendron amazonicum
Rhabdodendron sylvestre (Gleason) Maguire =Rhabdodendron amazonicum
References
Mori, S.A. 2002. Rhabdodendraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 598-600.
Prance, G.T. 1972. Rhabdodendraceae. Flora Neotropica Monogr. 11: 1-22.
Checklist of the Plants of the Guiana Shield 477
AM
AM
AM
AM
AM
BO GU su FG
BO GU
GU
BO DA GU su FG
FG
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
GU
DA GU
GU
SU
SU
FG
FG
FG?
FG
Rhamnaceae
RE. Berry, J.A. Steyermarkt
Ampelozizyphus amazonicus Ducke
Colubrina glandulosa Perkins
Colubrina glandulosa var. glandulosa
Gouania blanchetiana Miq.
Gouania canescens Rich.
Gouania colurnifolia Reissek
Gouania cornifolia Reissek
Gouania discolor Benth.
Gouaniafrangulifolia (Willd. ex Roem. & Schult.) Radlk.
Gouania lupuloides (L.) Urb. [s.l.]
Gouania polygama (Jacq.) Urb.
Gouania striata Rich.
Gouania velutina Reissek
Gouania virgata Reissek ["Guiana"]
Gouania wurdackii Steyerm.
Rhamnus acuminata Maguire & Steyerm. [Sierra de la Neblina]
Rhamnus chimantensis Steyerm. & Maguire
Rhamnus longipes Steyerm.
Rhamnus marahuacensis Steyerm. & Maguire
Rhamnus neblinensis Maguire & Steyerm. [Sierra de la Neblina]
Rhamnus psilocarpa Maguire & Steyerm. [Sierra de la Neblina]
Rhamnus sipapoensis Steyerm.
Rhamnus ulei Pilg. [Mt. Roraima]
Ziziphus cinnamomum Triana & Planch.
Ziziphusjujuba Mill, [cultivated; records may be referable to Ziziphus
mauritiana]
Ziziphus mauritiana Lam. [cultivated, naturalized]
Ziziphus saeri Pittier
Synonyms and Misidentifications
Gouania hypochroa Reissek =Gouania discolor
Gouania pubescens Poir. =Gouania polygama
Gouania tomentosa Jacq. =Gouania polygama
Gouania ulei Pilg. =Gouania discolor
Rhamnus cumanensis Loefl. (Boraginaceae) =Bourreria cumanensis
Rhamnus polygamus Jacq. =Gouania polygama
Trisecusfrangulaefolius Willd. ex Roem. & Schult. =Gouania frangulifolia
ZiziphusJujuba (L.) Lam., non Mill. =Ziziphus mauritiana
References
Heald, S.V. 2002. Rhamnaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. HofT, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
600-602.
Johnston, M.C. 1971. Revision of Colubrina. Brittonia 23: 2-53.
Johnston, M.C. & L.V.A. Johnston. 1978. Rhamnus. Flora Neotropica Monogr. 20: 1-96.
Lanjouw, J. 1 932. Rhamnaceae. In A. Pulle, Flora of Suriname 2( 1 ): 1 02- 1 06.
Steyermark, J.A. and RE. Berry. 2004. Rhamnaceae. In J.A. Steyermark, RE. Berry, K. Yatskicvych & B.K. Hoist, Flora of the
Venezuelan Guayana 8: 473-484.
Suessenguth, K. 1953. Rhamnaceae. In A. Engler & K. Prantl, Die Natiirlichen Pflanzenfamilien 2()d: 7-173.
BO GU?
GU FG
GU SU FG
BO GU su FG
BO
Rhizophoraceae
J.A. Steyermarkt
Cassipourea elliptica (Sw.) Poir. [=Cassipourea guianensis'?]
Cassipourea guianensis Aubl.
Cassipourea lasiocalyx Alston
Cassipourea peruviana Alston
Cassipourea serrata Benth.
Rhizophora x harrisonii Leechm. [putative hybrid species]
Rhizophora mangle L.
AM BO
SU FG
DA GU
GU
SU FG
GU su
GU
DA GU su FG
DA GU su FG
478 Checklist of the Plants of the Guiana Shield
Rhizophora racemosa G. Mey.
Sterigmapetalum chrysophyllum Aymard & Cuello
Sterigmapetaliim colombianum Monach.
Sterigmapetalum exappendiculatum Steyerm. & Liesner [perhaps expected in
GU]
Sterigmapetalum guianense Steyerm.
Sterigmapetalum guianense subsp. guianense [expected in GU]
Sterigmapetalum guianense subsp. ichunense Steyerm. & Liesner
Sterigmapetalum resinosum Steyerm. & Liesner
DA GU
AM
AM
AM
BO
BO
BO
BO
BO
su
su
su
su
FG
GU
Synonyms and Misidentifications
Rhizophora mangle var. racemosa (G. Mey.) Engl. =Rhizophora racemosa
References
(see also Anisophylleaceae)
Alston, A.H.G. 1925. Revision of Cassipourea. Kew Bull. 1925: 241-276.
Breteler, F.J. 1969. The Atlantic species of Rhizophora. Acta Bot. Neerl. 18: 434-444.
Ding Hou. 1960. A review of the genus Rhizophora. Blumea 10: 625-634.
Jonker, F.R 1942. Rhizophoraceae. In A. Pulle, Flora of Suriname 3(2): 35-43.
Juncosa, A.M. & RB. Tomlinson. 1988. A historical and taxonomic synopsis of Rhizophoraceae and Anisophylleaceae. Ann.
Missouri Bot. Gard. 75: 1278-1295.
Mori, S.A. 2002. Rhizophoraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
603-604.
Prance, GT. et al. 1975. Revisao taxonomica das especies amazonicas de Rhizophoraceae. Acta Amazonica 5: 5-22.
Steyermark, J.A. 2004. Rhizophoraceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 8: 484-490.
Steyermark, J.A. & R. Liesner. 1983. Revision of the genus Sterigmapetalum. Ann. Missouri Bot. Gard. 70: 179-193.
ROSACEAE
GA. Aymard, N.L. Cuello A.
BO
BO
BO GU SU FG
AM
BO
BO
BO GU SU FG
BO GU SU FG
BO
AM BO
BO GU
AM BO
Hesperomeles obtusifolia (Pers.) Lindl.
Hesperomeles obtusifolia var. obtusifolia
Prunus accumulans (Koehne) C.L. Li & Aymard
Prunus amplifolia Pilg.
Prunus espinozana C.L. Li
Prunus lichoana Aymard
Prunus myrtifolia (L.) Urb.
Primus myrtifolia var. myrtifolia
Prunus wurdackii C.L. Li
Rubus floribundus Kunth
Rubus guyanensis Focke
Rubus urticifolius Poir.
Synonyms and Misidentifications
Crataegus obtusifolia Pers. =Hesperomeles obtusifolia
Hesperomeles heterophylla (Ruiz & Pav.) Hook. =Hesperomeles obtusifolia
Mespilus heterophylla Ruiz & Pav. ^Hesperomeles obtusifolia
Prunus myrtifolia var. accumulans Koehne =Prunus accumulans
Prunus sphaerocarpa Sw. =Prunus myrtifolia
Rubus robustus C. Presl =Rubus floribundus
Rubus schomburgkii Klotzsch [nomen nudum] =Rubus guyanensis
References
Aymard, G.A. & N.L. Cuello A. 2004. Rosaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 8: 490-496.
Kleinhoonte, A. 1939. Rosaceae. In A. Pulle, Flora of Suriname 2(1): 426-456.
Mitchell, J.D. 2002. Rosaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 605-
606.
Checklist of the Plants of the Guiana Shield 479
RUBIACEAE
E. Araujo da Anuncia9ao, C. Bestetti Costa, R. Cortes, P.G. Delprete, J.H. Kirkbride,
C. Persson, J.A. Steyermarkt, CM. Taylor, A. Vincentini, D. Zappi
Alibertia bertierifolia K. Schum.
AUbertia edidis (Rich.) A. Rich, ex DC.
Alibertia edulis var. edulis
Alibertia edulis var. obtusiuscula (Steyerm.) Delprete & C. Perss.
Alibertia latifolia (Benth.) K. Schum.
Alibertia latifolia var. latifolia
Alseis floribunda Schott
Alseis labatioides H. Karst. ex K. Schum.
Alseis longifolia Ducke
Alseis longifolia var. pentamera Bremek.
Alseis smithii Standi.
Amaioua brevidentata Steyerm.
Amaioua corymbosa Kunth
Amaioua guianensis Aubl.
Amaioua guianensis var. guianensis
Amaioua guianensis var. macrantha Steyerm. [Sierra de la Neblina]
Amphidasya neblinae Steyerm. [Sierra de la Neblina]
Antirhea triflora J.H. Kirkbr.
Aphanocarpus steyermarkii (Standi.) Steyerm.
Bathysa bathysoides (Steyerm.) Delprete
Bertiera diversiramea Steyerm.
Bertiera guianensis Aubl.
Bertiera parviflora Spruce ex K. Schum.
Borreria alata (Aubl.) DC.
Borreria bolivarensis Steyerm.
Borreria capitata (Ruiz & Pav.) DC.
Borreria capitata var. capitata
Borreria capitata var. suaveolens (G. Mey.) Steyerm.
Borreria capitata var. tenella (Kunth) Steyerm.
Borreria cataractarum Steyerm.
Borreria confertifolia Steyerm.
Borreria densiflora DC.
Borreria hispida Spruce ex K. Schum.
Borreria intricata Steyerm.
Borreriajangouxii Steyerm.
Borreria latifolia (Aubl.) K. Schum.
Borreria macrocephala Standi. & Steyerm.
Borreria ovalifolia M. Martens & Galeotti
Borreria prostrata (Aubl.) Miq.
Borreria pygmaea Spruce ex K. Schum.
Borreria remota (Lam.) Bacigalupo & E.L. Cabral
Borreria repens DC.
Borreria verticillata (L.) G Mey.
Borreria wurdackii Steyerm.
Borreria xanthophylla Bremek.
Bothriospora corymbosa (Benth.) Hook. f.
Botryarrhena pendula Ducke
Botryarrhena venezuelensis Steyerm.
Calycophyllum obovatum (Ducke) Ducke
Calycophyllum venezuelense Steyerm.
Capirona decorticans Spruce
Chalepophyllum guianense Hook. f.
Chimarrhis brevipes Steyerm.
Chimarrhis cymosa Jacq.
Chimarrhis microcarpa Standi.
Chimarrhis turbinata DC.
Chiococca alba (L.) Hitchc.
Chiococca alba subsp. alba
Chiococca auyantepuiensis Steyerm.
Chiococca lucens Standi. & Steyerm.
AM BO GU
AM BO GU su FG
AM BO GU su FG
AM BO GU
AM BO DA GU su FG
GU su FG
FG
BO DA GU
GU su FG
GU su FG
GU
GU? FG
AM BO GU su FG
AM BO DA GU su FG
AM BO DA GU su FG
AM
AM
BO
FG
AM BO
BO GU su FG
AM BO DA GU su FG
AM
AM
BO
GU su FG
AM BO GU su FG
AM BO GU
GU
su
su
FG
AM BO
BO
GU
GU
su FG
AM BO GU
BO GU su FG
AM BO GU su FG
AM
AM
AM BO DA GU su FG
AM
GU
AM BO DA GU su FG
AM BO
AM BO GU su FG
AM BO DA GU su FG
AM BO DA GU su FG
AM
AM BO
AM BO
AM
AM
BO
BO
BO
BO
BO
GU
DA
su
FG
GU
GU
GU SU FG
GU
GU
GU su FG
GU su FG
GU FG
GU FG
480 Checklist of the Plants of the Guiana Shield
Chiococca miiltipedunculata Steyerm.
Chiococca nitida Benth. AM
Chiococca nitida var. amazonica Miill. Arg.
Chiococca nitida var. chimantensis Steyerm.
Chiococca nitida var. nitida AM
Chiococca pubescens Humb. & Bonpl. ex Roem. & Schult. [expected in GU]
Chomelia caurensis (Standi.) Steyerm.
Chomelia delascioi Steyerm.
Chomelia glabricalyx Steyerm.
Chomelia malaneoides Miill. Arg. AM
Chomelia monachinoi Steyerm.
Chomelia obtusa Cham. & Schltdl. [possibly misident. of C. angustifolia]
Chomelia polyantha S.F. Blake
Chomelia schomburgkii Steyerm.
Chomelia splitgerberi Bremek.
Chomelia stergiosii Steyerm.
Chomelia tenuiflora Benth. AM
Chomelia volubilis (Standi.) Steyerm. AM
Cinchonopsis amazonica (Standi.) L. Andersson AM
Coccochondra laevis (Steyerm.) Rauschert AM
Coccochondra laevis subsp. laevis AM
Coccochondra laevis subsp. maigualidae J.H. Kirkbr.
Coccocypselum aiireiim (Spreng.) Cham. & Schltdl.
Coccocypselum condalia Pers.
Coccocypselum guianense (Aubl.) K. Schum. AM
Coccocypselum herbaceum DC.
Coccocypselum hirsutum Bartl. ex DC.
Coccocypselum huberi Steyerm.
Coccocypselum lanceolatum (Ruiz & Pav.) Pers. AM
Coffea arabica L. [cultivated or escaped from cultivation] AM
Coffea liberica W. Bull ex Hiem [cultivated or escaped from cultivation]
Cordiera myrciifolia (Spruce ex K. Schum.) C. Perss. & Delprete AM
Cordiera thflora A. Rich. AM
Coryphothammts auyantepuiensis (Steyerm.) Steyerm.
Cosmibuena grandiflora (Ruiz & Pav.) Rusby
Coussarea amapaensis Steyerm.
Coussarea brevicaulis K. Krause [nomen nudum] AM
Coussarea evoluta Steyerm. [Sierra de la Neblina] AM
Coussareafanshawei Steyerm. GU
Coussarea grandis Miill. Arg. AM
Coussarea granvillei Delprete & B.M. Boom
Coussarea hallei Steyerm.
Coussarea hirticalyx Standi. AM
Coussarea hirticalyx var. glabrior Steyerm.
Coussarea krukovii Standi.
Coussarea lasseri Steyerm.
Coussarea leptoloba (Spreng. ex Benth. & Hook, f.) Miill. Arg. AM
Coussarea leptophragma Miill. Arg. AM
Coussarea liesneri Steyerm. AM
Coussarea machadoana Standi.
Coussarea mapourioides Bremek.
Coussarea micrococca Bremek.
Coussarea ovalis Standi. AM
Coussarea paniculata (Vahl) Standi. AM
Coussarea racemosa A. Rich.
Coussarea revoluta Steyerm. AM
Coussarea sp. A [Sierra de la Neblina] AM
Coussarea surinamensis Bremek.
Coussarea violacea Aubl. AM
Coutarea hexandra (Jacq.) K. Schum. AM
Coutarea hexandra var. hexandra
Declieuxiafruticosa (Willd. ex Roem. & Schult.) Kuntze AM
Declieuxia tenuiflora (Willd. ex Roem. & Schult.) Steyerm. & J.H. Kirkbr. AM
Dendrosipanea revoluta Steyerm. AM
GU
BO
BO
GU
GU
FG
BO
BO DA
GU EG
FG
BO
BO
BO
GU su FG
BO
BO
GU
GU
su
su?
su
rJU
GU su FG
VlC\
GU?
rJU
BO
BO
BO GU FG
BO
BO DA GU
GU
su FG
BO GU FG
BO
BO GU
BO GU su FG
DA GU FG
BO GU su FG
GU su FG
BO
BO DA GU
GU
su FG?
FG
FG?
FG
FG
GU? FG?
GU? FG?
GU
BO
GU FG
GU?
GU FG
GU? SU FG
GU su FG
BO GU su FG
GU su FG
GU?
su
GU su FG
BO GU su FG
GU su FG
BO GU su FG
BO GU
Checklist of the Plants of the Guiana Shield 481
Dendrosipanea spigelioides Ducke AM
Didymochlamys connellii N.E. Br.
Diodia apicidata (Willd. ex Roem. & Schult.) K. Schum.
Diodia hyssopifolia (Willd. ex Roem. & Schult.) Cham. & Schltdl.
Diodia kuntzei K. Schum.
Diodia multiflora DC.
Diodia ocymifolia (Willd. ex Roem. & Schult.) Bremek.
Diodia pulchristipula Bremek.
Diodia radula (Willd. & Hoffmamis. ex Roem. & Schult.) Cham. & Schltdl.
Diodia radula subsp. venezuelensis Steyerm.
Diodia sarmentosa Sw.
Diodia spicata Miq.
Diodia teres Walter
Duidania montana Standi.
Duroia amapana Steyerm.
Duroia aquatica (Aubl.) Bremek.
Duroia bolivarensis Steyerm.
Duroia eriopila L. f.
Duroia eriopila var. eriopila
Duroia eriopila var. tafelbergensis Steyerm.
Duroia fusifera Hook. f. ex K. Schum.
Duroia genipoides Hook. f. ex K. Schum.
Duroia gransabanens is Steyerm.
Duroia kotchubaeoides Steyerm.
Duroia longiflora Ducke
Duroia maguirei Steyerm.
Duroia maguirei var. maguirei
Duroia maguirei var. patentinervia Steyerm.
Duroia martiniana (Miers) Bremek.
Duroia merumensis Steyerm.
Duroia micrantha (Ladbr.) Zarucchi & J.H. Kirkbr.
Duroia nitida Steyerm.
Duroia paruensis Steyerm.
Duroia plicata Benoist FG
Duroia retrorsipila Steyerm. BO
Duroia saccifera (Mart, ex Roem. & Schult.) Hook. f. ex K. Schum. AM
Duroia spraguei Wemham GU
Duroia strigosa Steyerm. BO
Duroia velutina Hook. f. ex K. Schum. [expected in AM]
Elaeagia maguirei Standi.
Elaeagia maguirei var. maguirei
Elaeagia maguirei var. pubens Steyerm.
Emmeorhiza umbellata (Spreng.) K. Schum.
Emmeorhiza umbellata subsp. septentrionalis Steyerm.
Faramea angustifolia Spruce ex Miill. Arg.
Faramea anisocalyx Poepp. & Endl.
Faramea berryi Steyerm.
Faramea boomii Steyerm. [Sierra de la Neblina]
Faramea brevipes Steyerm.
Faramea calyciflora A. Rich, ex DC.
Faramea capillipes Miill. Arg. [specimens in Guianas may represent
misidentifications]
Faramea cardonae Standi. & Steyerm.
Faramea corymbosa Aubl.
Faramea costata Steyerm.
Faramea crass ifolia Benth.
Faramea cyanea Miill. Arg.
Faramea egregia Sandwith
Faramea fansha\vei Steyerm.
Faramea guianensis (Aubl.) Bremek.
Faramea invinii Steyerm.
Faramea lourteigiana Steyerm.
Faramea maguirei Steyerm.
Faramea morilloi Steyerm.
BO GU
AM BO GU SU FG
AM BO
BO
DA
DA
GU su FG
AM BO
BO
BO
DA GU
GU
SU
su
FG
AM BO DA GU su FG
AM GU su FG
AM BO GU su FG
AM
GU su
su
FG
FG
BO GU?
AM BO DA GU su FG
AM BO DA GU su
su
FG
AM BO GU
AM BO
BO
GU su FG
AM
GU su FG
AM
AM
AM
GU
FG
AM BO DA GU su FG
AM
AM
AM BO GU SU
AM BO GU SU
AM BO
BO GU SU FG
BO GU
AM
AM BO FG
AM BO
AM
GU
FG
AM BO
BO
GU su FG
GU SU
su
FG
FG
BO GU
GU?
BO GU
GU FG
GU SU FG
GU su
su?
FG
FG
BO GU
AM BO GU
AM BO GU su FG
AM
AM BO GU su FG
GU su FG
AM BO
FG
AM BO
GU
GU su
su
su
su
FG
FG
FG
FG
FG
AM GU
GU
su FG
482 Checklist of the Plants of the Guiana Shield
Faramea multiflora A. Rich, ex DC.
Faramea neblinae Steyerm. [Sierra de la Neblina]
Faramea occidentalis (L.) A. Rich.
Faramea occidentalis subsp. occidentalis
Faramea orinocensis Standi.
Faramea paniculata (Aubl.) Benth. \=Coussarea sp.?]
Faramea parvibractea Steyerm.
Faramea polytriadophora Bremek.
Faramea quadricostata Bremek., emend. Steyerm.
Faramea qiiinqueflora Poepp. & Endl.
Faramea sessiliflora Aubl.
Faramea sessiliflora var. peduncidata Bremek.
Faramea sessiliflora var. sessiliflora
Faramea sessilifolia (Kunth) DC.
Faramea spathacea Miill. Arg. ex Standi.
Faramea stenopetala Mart.
Faramea tamberlikiana Miill. Arg.
Faramea tamberlikiana subsp. tamberlikiana
Faramea tinguana Miill. Arg.
Faramea torqiiata Miill. Arg.
Faramea yavitensis Steyerm.
Faramea yutajensis Steyerm.
Ferdinandiisa boomii Steyerm. [Sierra de la Neblina]
Ferdinandusa goudotiana K. Schum.
Ferdinandusa goudotiana var. eciliata Steyerm.
Ferdinandusa goudotiana var. goudotiana [Mt. Roraima]
Ferdinandusa goudotiana var. psilocarpa Steyerm.
Ferdinandusa guainiae Spruce ex K. Schum.
Ferdinandusa neblinensis Steyerm. [Sierra de la Neblina]
Ferdinandusa paraensis Ducke
Ferdinandusa rudgeoides (Benth.) Wedd.
Ferdinandusa sprucei K. Schum.
Ferdinandusa uaupensis Spruce ex K. Schum.
Galium hypocarpium (L.) Endl. ex Griseb.
Galium surinamense Dempster
Genipa americana L.
Genipa americana var. americana
Genipa americana var. caruto (Kunth) K. Schum.
Genipa spruceana Steyerm. [=Genipa americana L.?]
Geophila cordifolia Miq.
Geophila cordifolia var. cordifolia
Geophila gracilis (Ruiz & Pav.) DC.
Geophila macropoda (Ruiz & Pav.) DC.
Geophila orbicularis (Miill. Arg.) Steyerm.
Geophila orbicularis var. neblinae Steyerm. [Sierra de la Neblina]
Geophila orbicularis var. orbicularis
Geophila repens (L.) I.M. Johnst.
Geophila tenuis (Mull. Arg.) Standi.
Gleasonia duidana Standi.
Gleasonia duidana var. duidana
Gleasonia duidana var. latifolia Steyerm.
Gonzalagunia bunchosioides Standi.
Gonzalagunia cornifolia (Kunth) Standi.
Gonzalagunia dicocca Cham. & Schltdl. [var. guianensis Steyerm.]
Gonzalagunia spicata (Lamb.) M. Gomez
Gonzalagunia surinamensis Bremek.
Guettarda acreana K. Krause
Guettarda argentea Lam.
Guettarda aromatica Poepp. & Endl.
Guettarda divaricata (Humb. & Bonpl. ex Roem. & Schult.) Standi.
Guettardafanshawei Steyerm.
Guettarda macrantha Benth.
Guettarda malacophylla Standi. BO
Guettarda pohliana MiiW. Arg. SU
AM
AM SU
AM
FG?
AM BO GU?
AM
AM
AM
AM BO GU
AM BO GU
AM BO
BO GU
AM GU
AM
GU SU FG
GU SU FG
AM
AM
BO
GU
GU
SU
AM BO DA GU su FG
AM BO DA SU
AM BO DA GU su FG
AM BO GU su FG
AM BO GU su FG
AM BO GU
GU
GU?
su FG
FG?
AM BO
AM
AM BO
AM BO DA GU su FG
AM GU su FG
AM BO
AM BO
AM
GU
GU FG
BO GU su FG
GU su FG
DA GU su FG
AM BO DA GU
GU
su FG
FG
GU su FG
AM BO GU
GU FG
AM GU su FG
AM
AM BO
AM
AM
AM
BO
BO
AM
AM BO
BO
AM BO
AM BO
AM
AM
AM BO
Checklist of the Plants of the Guiana Shield 483
Guettarda roupalifolia Rusby FG
Guettarda spruceana MxiW.Axg. BO GU SU FG
Guettarda viburnoides Cham. & Schltdl. GU
Hamelia axillaris Sw. FG
Hamelia patens Jacq. DA GU
Henriqueziajenmanii K. Schum. GU
Henriquezia nitida Spruce ex Benth.
Henriquezia nitida var. nitida
Henriquezia verticillata Spruce ex Benth.
Hillia foldatsii Steyerm.
Hillia illustris (Veil.) K. Schum. DA
Hillia maguirei Steyerm.
Hillia parasitica Jacq.
Hillia psammophila Steyerm.
Hillia rivalis CM. Taylor AM
Hillia ulei K. Krause
Hoffmannia megistophylla Standi.
Holstianthus barbigularis Steyerm. AM
Ibetralia surinamensis Bremek.
Isertia bullata K. Schum., emend. Delprete '
Isertia coccinea (Aubl.) J.F. Gmel.
Isertia hypoleuca Benth.
Isertia parviflora Yahl DA
Isertia rosea Spruce ex K. Schum.
Isertia spiciformis DC.
Isertia verrucosa (Bonpl.) Standi.
Isertia wilhelminensis Steyerm.
Ixora acuminatissima Miill. Arg.
Ixora aluminicola Steyerm.
Ixora coccinea L. [cultivated or escaped from cultivation]
Ixora davisii Sandwith
Ixoraferrea (Jacq.) Benth.
Ixora graciliflora Benth.
Ixora intensa K. Krause
Ixora intropilosa Steyerm. BO
Ixora panurensis Miill. Arg. AM
Ixora parviflora Vahl
Ixora piresii Steyerm.
Ixora potaroensis Steyerm.
Ixora pubescens Willd. ex Schult. & Schult. f.
Ixora sandwithiana Steyerm.
Ixora schomburgkiana Benth. BO
Ixora ulei K. Krause BO
Ixora versteegii Bremek.
Ixora yavitensis Steyerm. AM
Kutchubaea insignis Fisch. ex DC. GU FG
Kutchubaea longiloba Steyerm. BO
Kutchubaea micrantha Steyerm.
Kutchubaea morilloi Steyerm.
Kutchubaea neblinensis Steyerm. [Sierra de la Neblina]
Kutchubaea sericantha Standi.
Ladenbergia amazonensis Ducke
Ladenbergia lambertiana (A. Braun ex Mart.) Klotzsch
Ladenbergia oblongifolia (Mutis) L. Andersson
Ladenbergia sp. A
Limnosipanea palustris (Seem.) Hook. f.
Limnosipanea spruceana Hook. f.
Machaonia brasiliensis (Hoffmanns, ex Humb.) Cham. & Schltdl.
Maguireocharis neblinae Steyerm. [Sierra de la Neblina]
Maguireothamnus speciosus (N.E. Br.) Steyerm.
Maguireothamnus speciosus subsp.jauaensis Steyerm.
Maguireothamnus speciosus subsp. speciosus
Maguireothamnus tatei (Standi.) Steyerm. AM
Malanea auyantepuiensis Steyerm. BO
GU SU FG
GU FG
GU SU
GU FG
SU FG
SU
GU SU FG
GU SU FG
GU SU FG
SU FG
GU SU
SU
FG
GU FG
GU SU FG
GU SU FG
GU FG
GU
GU SU FG
SU FG
GU
GU
GU
SU FG
GU
SU
FG
FG
GU FG
GU SU FG
SU FG
AM
AM BO
AM GU
AM
AM
AM BO DA GU
AM
AM
AM BO GU
AM BO
BO
GU
AM
BO GU
BO
BO GU
AM
BO
AM
AM
AM BO
AM BO
AM
AM
AM
AM
AM BO
BO
AM BO
AM
GU su
GU
GU
GU su
GU
484 Checklist of the Plants of the Guiana Shield
Malanea chimantensis Steyerm. BO
Malanea ciliolata Steyerm. GU
Malanea cruzii Steyerm. GU
Malanea gabrielensis Miill. Arg. AM BO GU
Malanea guaiquinimensis Steyerm. BO
Malanea hypoleuca Steyerm. BO GU FG
Malaneajauaensis Steyerm. BO
Malanea macrophylla GrisQh. BO DA GU SU FG
Malanea macrophylla var. macrophylla GU SU FG
Malanea microphylla Standi. & Steyerm. BO
Malanea obovata Hochr. BO GU
Malanea ptariensis Steyerm. AM BO
Malanea sarmentosa Auhl AM BO SU FG
Malanea schomburgkii Steyerm.
Malanea setulosa Steyerm. BO
Malanea sipapoensis Steyerm. « AM
Malanea tafelbergensis Steyerm.
Malanea ueiensis Steyerm.
Malanea ursina Standi.
M^wema a/Z)fir (Aubl.) Wemham GU SU FG
Manettia calycosa Griseb.
Manettia calycosa var. karsteniana K. Schum.
Manettia coccinea (Aubl.) Willd.
Manettia reclinata Mutis ex L.
Merumea coccocypseloides Steyerm.
Merumea plicata Steyerm.
Mitracarpus diffusus (Willd. ex Roem. & Schult.) Cham. & Schltdl.
Mitracarpus frigidus (Willd. ex Roem. & Schult.) K. Schum.
Mitracarpus frigidus var.fruticosus (Standi.) Steyerm.
Mitracarpus frigidus var. orinocensis Steyerm.
Mitracarpus hirtus (L.) DC.
Mitracarpus humboldtianus Cham. & Schltdl.
Mitracarpus microspermus K. Schum.
Mitracarpus parvulus K. Schum.
Mitracarpus scabrellus Benth.
Mitracarpus sp. A
Morinda aurantiaca (K. Krause) Steyerm.
Morinda brachycalyx (Bremek.) Steyerm.
Morinda calycina (Benth.) Steyerm.
Morinda citrifolia L. [cultivated, naturalized]
Morinda debilis (Sandwith) Steyerm.
Morinda longiloba (Steyerm.) Steyerm.
Morinda longipedunculata Steyerm. [Sierra de la Neblina] AM
Morinda peduncularis Kunth AM
Morinda sparsiflora (Steyerm.) Steyerm.
Morinda surinamensis (Bremek.) Steyerm. [possibly =M. tenuiflora]
Morinda tenuiflora (Benth.) Steyerm. AM BO
Morinda venezuelensis (Steyerm.) Steyerm. BO
Neblinathamnus argyreus Steyerm. [Sierra de la Neblina] AM
Neblinathamnus brasiliensis Steyerm. AM
Neobertiera gracilis Wemham GU SU FG
Notopleura aligera (Steyerm.) CM. Taylor BO GU
Notopleura aneurophylla (Standi.) CM. Taylor GU?
Notopleura anomothyrsa (K. Schum. & Donn. Sm.) CM. Taylor FG
Notopleura crassa (Benth.) CM. Taylor [Mt. Roraima] AM BO GU
Notopleura guadalupensis (DC.) CM. Taylor GU FG
Notopleura macrophylla (Ruiz & Pav.) CM. Taylor GU FG
Notopleura merumensis (Steyerm.) CM. Taylor GU
Notopleura microbracteata (Steyerm.) CM. Taylor FG
Notopleura multiramosa (Steyerm.) CM. Taylor BO GU
Notopleura perpapillifera (Steyerm.) CM. Taylor BO GU
Notopleura sandwithiana (Steyerm.) CM. Taylor BO GU
Notopleura saiilensis (Steyerm.) CM. Taylor GU FG
Notopleura tapajozensis (Standi) Bremek. AM BO GU SU FG
GU SU FG
GU SU FG
GU
GU
GU
GU
GU SU FG
GU? SU? FG?
GU su FG
GU
GU
GU
GU su FG
GU
GU
GU
GU
GU
GU
su
GU su FG
Checklist of the Plants of the Guiana Shield 485
Notopleura thesceloantha (Steyerm.) CM. Taylor
Notopleura uliginosa (Sw.) Bremek.
Oldenlandia corymbosa L. [=Hedyotis corymbosa (L.) Lam.]
Oldenlandia fdicaulis K. Schum.
Oldenlandia lancifolia (Schumach.) DC. [naturalized] [=Hedyotis lancifolia
Schumacher]
Oldenlandia tenuis K. Schum.
Oldenlandiopsis callitrichoides (Griseb.) Terrell & W.H. Lewis [introduced]
Pagamea anisophylla Standi. & Steyerm.
Pagamea capitata Benth.
Pagamea capitata subsp. capitata
Pagamea capitata subsp. conferta (Standi.) Steyerm.
Pagamea capitata subsp. thibaudiifolia (Wemham) Steyerm.
Pagamea coriacea Spruce ex Benth.
Pagamea coriacea var. acuta Steyerm.
Pagamea coriacea var. coriacea
Pagamea diceras Steyerm.
Pagamea duidana Standi. & Steyerm.
Pagamea guianensis Aubl.
Pagamea guianensis var. guianensis
Pagamea hirsuta Spruce ex Benth.
Pagameajauaens is Steyerm.
Pagamea magniflora Steyerm.
Pagamea montana Gleason & Standi.
Pagamea pauciflora Standi. & Steyerm.
Pagamea plicata Spruce ex Benth.
Pagamea plicatiformis Steyerm.
Pagamea sessiliflora Spruce ex Benth.
Pagamea standleyana Steyerm.
Pagamea thyrsiflora Spruce ex Benth.
Pagamea velutina Steyerm.
Pagameopsis garryoides (Standi.) Steyerm.
Pagameopsis maguirei Steyerm.
Palicourea amapaensis Steyerm.
Palicourea anceps Standi.
Palicourea brachyloba (Mull. Arg.) B.M. Boom
Palicourea bracteosa Standi.
Palicourea calophylla DC.
Palicourea cardonae Steyerm.
Palicourea corymbifera (Miill. Arg.) Standi.
Palicourea crocea (Sw.) Roem. & Schult.
Palicourea croceoides Desv. ex Ham. [=Palicourea riparia]
Palicourea fastigiata Kunth
Palicourea foldatsii Steyerm.
Palicoureaformosa (A. Rich.) Lemee
Palicourea glabriflora Steyerm.
Palicourea grandiflora (Kunth) Standi.
Palicourea grandifolia (Willd. ex Roem. & Schult.) Standi.
Palicourea guianensis Aubl.
Palicourea guianensis subsp. guianensis
Palicourea huberi Steyerm.
Palicourea irwinii Steyerm.
Palicourea lancigera (Standi.) Steyerm.
Palicourea longiflora DC.
Palicourea longistipulata (Miill. Arg.) Standi.
Palicourea longistipulata subsp. chrysorrhachis (Bremek.) Steyerm.
Palicourea longistipulata subsp. longistipulata
Palicourea marcgravii A. St.-Hil.
Palicourea nitidella (Miill. Arg.) Standi.
Palicourea obtusata K. Krause
Palicourea ottohuberi J.H. Kirkbr.
Palicourea pedunculosa (A. Rich.) DC. ex Lemee [dubious name]
Palicourea pensilis J.H. Kirkbr.
Palicourea perquadrangularis Wemham
AM
BO DA GU su FG
AM BO DA GU su FG
AM
AM BO
BO
DA GU
GU
GU
su FG
AM
AM BO GU su
BO GU su
AM BO
AM
AM
AM
AM
AM
AM BO
BO
AM BO
BO
AM BO
AM
AM
BO
AM BO
AM
AM BO
AM BO
GU
GU
GU
GU
GU
GU
GU
SU
su
su
FG
FG
FG
GU
FG
BO
AM BO GU SU FG
AM
AM BO GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM BO GU
AM
AM
AM
FG
FG
AM BO FG?
AM BO GU SU FG
AM
AM
A X/f
BO GU SU FG
FG
AM
AM GU SU FG
AM BO GU su FG
BO GU su FG
AM
FG
AM BO
BO
BO
BO
GU
GU
FG
AM BO GU
486 Checklist of the Plants of the Guiana Shield
Palicourea perquadrangularis van guayanensis Steyerm.
Palicourea quadrifolia (Rudge) DC.
Palicourea quadrifolia subsp. quadrifolia
Palicourea rigida Kunth
Palicourea sp. A
Palicourea tepuicola Steyerm.
Palicourea triphylla DC.
Palicourea wurdackiana J.H. Kirkbr.
Patabea tenuiflora DC. \=Psychotria sp.]
Patima guianensis Aubl.
Perama dichotoma Poepp.
Perama dichotoma var. dichotoma
Perama dichotoma var. monocephala Steyerm.
Perama galioides (Kunth) Poir.
Perama galioides var. galioides
Perama hirsuta Aubl.
Perama plantaginea (Kunth) Hook. f.
Platycarpum decipiens Woodson & Steyerm.
Platycarpum duckei Steyerm.
Platycarpum eglandidosum Steyerm.
Platycarpum maguirei Steyerm. [Sierra de la Neblina]
Platycarpum negrense Ducke
Platycarpum negrense var. glaucum G.K. Rogers
Platycarpum orinocense Bonpl.
Platycarpum orinocense var. grandiflorum Steyerm.
Platycarpum orinocense var. orinocense
Platycarpum rhododactylum Woodson & Steyerm.
Platycarpum rugosum Steyerm.
Platycarpum schultesii Steyerm.
Platycarpum schultesii var. zarucchii G.K. Rogers
Posoqueria coriacea M. Martens & Galeotti
Posoqueria latifolia (Rudge) Roem. & Schult.
Posoqueria latifolia subsp. gracilis (Rudge) Steyerm.
Posoqueria latifolia subsp. latifolia
Posoqueria longiflora Aubl.
Posoqueria panamensis (Walp. & Duchass.) Walp.
Posoqueria trinitatis DC.
Posoqueria williamsii Steyerm.
Psychotria acuminata Benth.
Psychotria adderleyi Steyerm.
Psychotria adenophora Steyerm.
Psychotria aetantha (Sandwith) Steyerm.
Psychotria alba Ruiz & Pav.
Psychotria albert-smithii Standi.
Psychotria alloantha Steyerm.
Psychotria altsonii (Sandwith) Steyerm. GU
Psychotria amplectens Benth. AM SU
Psychotria anartiothrix Steyerm. AM
Psychotria anceps Kunth
Psychotria anceps var. anceps
Psychotria antennaeformis Steyerm.
Psychotria apoda Steyerm.
Psychotria appuniana Steyerm.
Psychotria astrellantha Wemham
Psychotria aubletiana Steyerm.
Psychotria ayangannensis Steyerm.
Psychotria barnebyana Steyerm.
Psychotria berryi Wingf.
Psychotria berteroana DC.
Psychotria blakei Standi. & Steyerm.
Psychotria bolivarensis (Standi. & Steyerm.) Steyerm.
Psychotria boliviana Standi.
Psychotria borjensis Kunth
Psychotria bostrychothyrsus Sandwith
AM BO GU
AM GU SU FG
AM GU SU FG
AM BO DA GU SU FG
AM
BO
AM BO GU SU FG
AM BO GU
GU
FG
FG
AM BO GU SU FG
AM BO GU SU FG
AM
AM BO GU
GU
AM BO GU SU FG
AM
AM
AM
GU
AM
AM
AM
AM BO
AM BO
AM BO
BO
BO GU
AM
AM
GU
AM BO DA GU SU FG
AM BO GU SU FG
AM BO DA GU SU FG
AM BO DA GU SU FG
AM BO DA GU FG
GU SU FG
AM
AM BO DA GU SU FG
AM
AM
GU
GU
AM BO GU SU
SU
FG
FG
AM BO GU SU FG
GU SU FG
FG
BO DA GU SU FG
GU
AM GU SU FG
AM BO GU
GU
FG
AM
AM
AM
AM
AM
BO
BO
BO
GU
AM BO GU FG
AM BO GU SU FG
BO GU SU FG
Checklist of the Plants of the Guiana Shield 487
Psychotria boyanii Steyerm. GU
Psychotria bracteocardia (DC.) Mull. Arg. AM BO GU SU FG
Psychotria brazaoi Steyerm. [expected in AM; Serra da Neblina]
Psychotria bremekampiana Steyerm. GU? SU FG
Psychotria brevipes DC. FG
Psychotria callithrix (Miq.) Steyerm. GU SU FG
Psychotria callithrix var. callithrix SU FG
Psychotria callithrix var. tontaneoides (Britton & Standi.) Steyerm. GU FG
Psychotria campyloneura Miill. Arg. AM BO DA GU SU FG
Psychotria campylopoda Standi. AM BO GU
Psychotria capitata Ruiz & Pav. AM BO GU SU FG
Psychotria capitata subsp. amplifolia (Raeusch.) Steyerm. GU SU FG
Psychotria capitata subsp. capitata GU
Psychotria capitata subsp. inundata (Benth.) Steyerm. AM BO GU SU FG
Psychotria carapichea (Poir.) Delprete FG
Psychotria cardiomorpha CM. Taylor & A. Pool AM GU SU FG
Psychotria carrenoi Steyerm. BO
Psychotria carthagenensis Jacq. AM BO GU SU FG
Psychotria casiquiaria Miill. Arg. AM BO
Psychotria celiac Steyerm. [expected in AM; Serra da Neblina]
Psychotria ceratantha Standi. AM BO GU
Psychotria cerronis Steyerm. BO
Psychotria colorata (Willd. ex Roem. & Schult.) Mull. Arg. AM BO GU SU FG
Psychotria concinna Oliv. [Mt. Roraima] AM BO GU
Psychotria cornigera Benth. AM BO GU SU FG
Psychotria coussareoides Standi. AM BO GU FG
Psychotria crocochlamys Sandwith BO GU
Psychotria ctenophora Steyerm. SU FG
Psychotria cupularis (Mull. Arg.) Standi. AM BO DA GU SU FG
Psychotria cypellantha Steyerm. FG
Psychotria deflexa DC. AM BO DA GU SU FG
Psychotria deinocalyx Sandwith BO GU SU FG
Psychotria duidana Standi. AM
Psychotria duricoria Standi. & Steyerm. AM
Psychotria duricoria var. duricoria AM
Psychotria duricoria var. longiloba Steyerm. AM
Psychotria durifolia Standi. AM
Psychotria egensis Miill. Arg. AM GU?
Psychotria ernestii K. Krause BO GU SU FG
Psychotria everardii Wemham AM BO GU
Psychotria everardii subsp. everardii GU
Psychotriafanshawei (Standi.) Steyerm. BO GU
Psychotriafarameoides Bremek. SU
Psychotriafranquevilleana Miill. Arg. AM
Psychotria galbaoensis Steyerm. GU? FG
Psychotria glandulicalyx Steyerm. AM BO GU
Psychotria gracilenta Mull. Arg. AM BO DA GU SU FG
Psychotria granvillei Steyerm. FG
Psychotria guanchezii Steyerm. AM
Psychotria hemicephaelis Wemham BO GU
Psychotria heteroneura Steyerm. [expected in AM; Serra da Neblina] GU
Psychotria hoffmannseggiana (Willd. ex Roem. & Schult.) Mull. Arg. AM BO DA GU SU FG
Psychotria horizontalis Sw. AM BO GU
Psychotria humboldtiana (Cham.) Mull. Arg. AM BO
Psychotria hyptoides Benth. GU
Psychotria imthurniana Oliv. BO GU
Psychotria iodotricha Mull. Arg. AM BO DA GU SU FG
Psychotria irwinii Steyerm. BO DA GU SU FG
Psychotriajauaensis Steyerm. AM BO
Psychotria kaieteurensis Sandwith GU
Psychotria kappleri (Miq.) Mull. Arg. ex Benoist GU SU FG
Psychotria lateralis Steyerm. FG FG
Psychotria leiantha Steyerm. [Sierra de la Neblina] AM
Psychotria ligularis (Rudge) Steyerm. GU SU FG
488 Checklist of the Plants of the Guiana Shield
Psychotria ligularis van carapichea (Poir.) Steyerm. SU FG
Psychotna ligularis van ligularis GU SU FG
Psychotria longicuspis Miill. Arg. AM GU
Psychotna lourteigiana Steyerm. AM BO
Psychotria hipulina Benth. AM BO GU SU FG
Psychotria lupulina subsp. lupulina AM BO GU SU FG
Psychotria lupulina subsp. rhodoleuca (Miill. Arg.) Steyerm. AM BO
Psychotna maguireorum Steyerm. AM GU
Psychotria mapourioides DC. BO GU SU FG
Psychotria mapourioides van mapourioides BO GU SU FG
Psychotria maturacensis Steyerm. [Sierra de la Neblina] AM GU
Psychotria mazaruniensis Standi. GU FG
Psychotria medusula Miill. Arg. AM GU FG
Psychotria microbotrys Ruiz ex Standi. AM BO GU SU FG
Psychotna microdon (DC.) Urb. BO GU
Psychotria morindoides (A. Rich.) Lemee
.
FG
Psychotria moroidea Steyerm. SU FG
Psychotria muscosa (Jacq.) Steyerm. GU SU FG
Psychotria muscosa subsp. breviloba Steyerm. GU SU FG
Psychotria nana K. Krause AM
Psychotria nematostachya Steyerm. [Sierra de la Neblina] AM
Psychotria nivea (Sandwith) Steyerm. GU
Psychotria oblita Wemham [Mt. Roraima] BO GU
Psychotria oblonga (DC.) Steyerm. AM BO GU SU FG
Psychotria Occidentalis Steyerm. DA
Psychotna officinalis (Aubl.) Raeusch. ex Sandwith AM BO GU SU FG
Psychotna officinalis subsp. officinalis GU SU FG
Psychotria officinalis subsp. wilhelminensis Steyerm. SU
Psychotria oiapoquensis Steyerm. FG
Psychotria pacimonica Miill. Arg. AM GU FG
Psychotria pakaraimensis Steyerm. GU
Psychotria pallidinervia Steyerm. [Sierra de la Neblina] AM
Psychotria paniculata (Aubl.) Raeusch. AM BO GU SU FG
Psychotria parimensis Steyerm. AM
Psychotria parvibractea Steyerm. BO GU
Psychotria paupertina Standi. & Steyerm. AM BO
Psychotna pectinata Steyerm. [Sierra de la Neblina] AM
Psychotria perferruginea Steyerm. FG
Psychotria phaneroloma Standi. & Steyerm. AM BO GU
Psychotria phaneroloma van phaneroloma GU
Psychotria phelpsiana Steyerm. BO
Psychotria pilosa Ruiz & Pav. AM
Psychotriaplatypoda DC. AM BO GU SU FG
Psychotria plocamipes Wemham GU
Psychotria podocephala (Miill. Arg.) Standi. AM BO GU? FG
Psychotria poeppigiana Miill. Arg. AM BO DA GU SU FG
Psychotria poeppigiana subsp. barcellana (Miill. Arg.) Steyerm. AM BO GU SU FG
Psychotria poeppigiana subsp. poeppigiana AM BO DA GU SU FG
Psychotria polycephala Benth. AM BO DA GU
Psychotria potaroensis (Sandwith) Steyerm. GU
Psychotriaprancei Steyerm. SU FG
Psychotriaprunifolia (Kunth) Steyerm. AM
Psychotria pseudinundata Wemham SU FG
Psychotriapsittacina Steyerm. GU
Psychotria puberulenta Steyerm. GU
Psychotria pullei Bremek. SU FG
Psychotriapungens Steyerm. FG
Psychotria racemosa Rich. AM BO DA GU SU FG
Psychotna remota Benth. AM GU SU FG
Psychotria rosea (Benth.) Mull. Arg. AM GU FG
Psychotria rosea var. rosea GU FG
Psychotria schlechtendaliana Miill. Arg. FG?
Psychotna schomburgkii Benth. AM GU
Psychotria sipapoensis Steyerm. AM
Checklist of the Plants of the Guiana Shield 489
Psychotr,
Psychotr
Psychotr^
Psychotr,
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr,
Psychotr.
Psychotr,
Psychotr,
Psychotr,
Psychotr,
Psychotr,
Psychotr,
Psychotr,
Psychotr,
Psychotr,
Psychotr,
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr
Psychotr,
Psychotr
Psychotr
Psychotr,
Psychotr,
Psychotr
Psychotr
a spadicea (Pittier) Standi. & Steyerm.
a spectabilis Steyerm.
a speluncae Standi. & Steyerm.
a spehmcae subsp. speluncae
a sphaerocephala Miill. Arg.
a spiciflora Standi.
a squamelligera Steyerm.
a stipulosa Mull. Arg.
a subundulata Benth.
a surinamensis Bremek.
a tapirapecoana Steyerm. [expected]
a tatei Standi.
a tepuiensis (Steyerm.) Steyerm.
a tetramera Steyerm.
a tillettii Steyerm.
a transtens Wemham
a tricholoba Miill. Arg.
a trichophoroides Miill. Arg.
a trichotoma M. Martens & Galeotti
a turbinella Miill. Arg.
a turbinella var. sororiella (Miill. Arg.) Steyerm.
a ulviformis Steyerm.
ia urceolata Steyerm.
ia urniformis Steyerm.
ia vareschii Steyerm.
ia variegata Steyerm.
ia vasivensis (Miill. Arg.) Standi.
ia vellosiana Benth.
ia venezuelensis Steyerm. [Sierra de la Neblina]
ia ventuariana Standi. & Steyerm.
ia venulosa Miill. Arg.
ia vichadensis Standi.
ia viridibractea Steyerm.
ia viridis Ruiz & Pav.
ia wessels-boeri Steyerm.
ia wilhelminensis Steyerm.
ia wonotobensis (Bremek.) Steyerm.
ia wurdackii Steyerm.
ia yapacanensis Steyerm.
ia yavitensis Steyerm.
Randia aculeata L.
Randia aculeata var. dasyphylla Steyerm.
Randia amazonasensis Steyerm.
Randia armata (Sw.) DC.
Randia asperifolia (Sandwith) Sandwith
Randia brevipes Steyerm.
Randia calycina Cham.
Randia dioica H. Karst.
Randia hebecarpa Benth.
Randia pubiflora Steyerm.
Randia venezuelensis Steyerm.
Remijia amazonica K. Schum.
Remijia amazonica var. amazonica
Remijia aracamuniensis (Steyerm.) CM. Taylor
Remijia argentea Steyerm.
Remijia delascioi Steyerm.
Remijia densiflora Benth.
Remijia densiflora subsp. densiflora
Remijia densiflora var. densiflora
Remijia densiflora var. minima Steyerm.
Remijia densiflora subsp. stenopetala (Standi. & Steyerm.) Steyerm.
Remijiafirmula (Mart.) Wedd.
Remijia globosa (Steyerm.) CM. Taylor [Sierra de la Neblina]
Remijia hispida Spruce ex K. Schum.
AM BO
AM BO
AM
AM
AM
AM BO
AM
AM BO
AM BO
AM
AM
AM
AM BO
AM
AM BO
AM
AM BO
AM
AM
AM BO
AM BO
GU
GU
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
AM BO
AM
AM
AM
AM
AM
AM BO
AM BO
BO
AM BO
AM
AM
AM
DA
GU
GU
GU
SU
FG
FG
GU SU FG
SU FG
GU
GU
GU
GU
GU?
SU FG
GU SU FG
FG
GU
GU SU FG
GU
GU FG
FG
GU? SU
SU
SU
GU SU
GU SU
GU?
GU
GU
FG
FG
FG
490 Checklist of the Plants of the Guiana Shield
Remijia longifolia Benth. ex Standi.
Remijia magidrei Steyerm. [Sierra de la Neblina]
Remijia marahuacensis Steyerm.
Remijia morilloi Steyerm.
Remijia pacimonica Standi.
Remijia pedunculata (H. Karst.) Flueck.
Remijia pilosinervula Steyerm.
Remijia reducta Steyerm.
Remijia roraimae (Benth.) K. Schum.
Remijia roraimae van adpressa Steyerm.
Remijia roraimae var. auyantepuiensis Steyerm.
Remijia roraimae var. guianensis Steyerm.
Remijia roraimae var. roraimae
Remijia sessilis Steyerm. [Sierra de la Neblina]
Remijia sipapoensis Steyerm.
Remijia steyermarkii Standi.
Remijia ulei K. Krause
Remijia uniflora CM. Taylor
Remijia wurdackii Steyerm.
Retiniphyllum chloranthum Ducke
Retiniphyllum concolor (Spruce ex Benth.) Mull. Arg., emend. Cortes
Retiniphyllum discolor (Spruce ex Benth.) Mull. Arg.
Retiniphyllum guianense Steyerm.
Retiniphyllum laxiflorum (Benth.) N.E. Br.
Retiniphyllum laxiflorum var. laxiflorum
Retiniphyllum maguirei Standi.
Retiniphyllum pauciflorum Kunth ex K. Krause
Retiniphyllum pilosum (Spruce ex Benth.) Miill. Arg.
Retiniphyllum scabrum Benth.
Retiniphyllum schomburgkii (Benth.) Miill. Arg.
Retiniphyllum schomburgkii subsp. schomburgkii
Retiniphyllum secundiflorum Bonpl.
Retiniphyllum tepuiense Steyerm.
Retiniphyllum truncatum Mull. Arg.
Richardia brasiliensis Gomes
Richardia scabra L.
Ronabea latifolia Aubl.
Rondeletia orinocensis Steyerm.
Rosenbergiodendron densiflorum (K. Schum.) Fagerl.
Rosenbergiodendronformosum (Jacq.) Fagerl.
Rosenbergiodendronformosum var. nitidum (K. Schum.) C. Gust.
Rosenbergiodendron longiflorum Fagerl.
Rudgea ayangannensis Steyerm.
Rudgea bremekampiana Steyerm.
Rudgea cardonae Steyerm.
Rudgea corniculata Benth.
Rudgea cornifolia (Kunth ex Roem. & Schult.) Standi.
Rudgea cornigera Bremek.
Rudgea crassiloba (Benth.) B.L. Rob.
Rudgea enervia Steyerm.
Rudgea graciliflora Standi.
Rudgea guyanensis (A. Rich.) Sandwith
Rudgea hostmanniana Benth.
Rudgea hostmanniana subsp. hostmanniana
Rudgea hostmanniana subsp. maypurensis (Standi.) Zappi
Rudgea klugii Standi.
Rudgea lanceifolia Salisb.
Rudgea merumensis Steyerm.
Rudgea oldemanii Steyerm.
Rudgea phaneroneura Steyerm.
Rudgea sandemanii Sandwith
Rudgea sclerocalyx (Miill. Arg.) Zappi
Rudgea stipulacea (DC.) Steyerm.
Rudgea tanaosepala Sandwith
AM
AM
AM GU
AM
AM
AM
BO GU
AM
AM BO GU
AM BO
BO
BO GU
AM BO GU
AM
AM
AM
AM
AM
AM
AM BO GU
AM BO GU
AM
GU
AM BO GU SU
AM BO GU su
SU
AM GU
AM
AM BO GU
AM BO GU
GU
su
su
AM
AM
AM GU
GU
BO DA GU FG
AM BO GU su FG
AM BO
AM BO DA GU su FG
AM BO DA GU?
AM BO DA
GU su FG
GU
FG
AM
FG
AM BO DA GU su FG
GU su FG
AM BO GU
GU
GU
su? FG
FG
AM BO DA GU su FG
AM BO DA GU su FG
AM
AM GU su FG
AM
BO
GU
GU
su FG
FG
FG
AM
AM BO GU
GU
su FG
FG
Checklist of the Plants of the Guiana Shield 491
Rudgea woronowii Standi.
Rudgea wurdackii Steyerm.
Sabicea amazonensis Wemham
Sabicea aspera Aubl.
Sabicea bariensis Steyerm.
Sabicea brachycalyx Steyerm.
Sabicea cinerea Aubl. [expected in GU]
Sabicea glabrescens (K. Schum.) Benth.
Sabicea grandifolia Steyerm.
Sabicea morillorum Steyerm.
Sabicea oblongifolia (Miq.) Steyerm.
Sabicea panamensis Wemham
Sabicea paraens is (K. Schum.) Wemham
Sabicea romboutsii Bremek.
Sabicea surinamensis Bremek.
Sabicea tillettii Steyerm.
Sabicea velutina Benth.
Sabicea velutina subsp. chimantensis Steyerm.
Sabicea velutina subsp. duidensis Steyerm.
Sabicea velutina subsp. velutina
Sabicea venezuelensis Steyerm.
Sabicea villosa Willd. ex Roem. & Schult.
Sabicea villosa var. adpressa (Wemham) Standi.
Sabicea villosa var. villosa
Schradera bipedunculata Steyerm.
Schradera bipendunculata Steyerm.
Schradera brevipes Steyerm.
Schradera hilliifolia Steyerm. [Sierra de la Neblina]
Schradera maguirei Steyerm.
Schradera marahuacensis Steyerm.
Schradera nilssonii Steyerm.
Schradera polycephala DC.
Schradera surinamensis Standi.
Schradera ternata Steyerm.
Schradera yutajensis Steyerm.
Semaphyllanthe intonsa (Steyerm.) L. Andersson
Semaphyllanthe merumensis (Steyerm.) L. Andersson
Semaphyllanthe spectabilis (Steyerm.) L. Andersson
Simira ignicola Steyerm.
Simira pisoniiformis (Baill.) Steyerm.
Simira rubescens (Benth.) Bremek. ex Steyerm.
Simira tinctoria Aubl.
Sipanea ayangannensis Steyerm.
Sipanea biflora (L. f.) Cham. & Schltdl.
Sipanea carrenoi Steyerm.
Sipanea cowanii Steyerm.
Sipanea galioides Wemham
Sipanea glaberrima (Bremek.) Steyerm.
Sipanea gleasonii Steyerm.
Sipanea glomerata Kunth
Sipanea hispida Benth. ex Wemham
Sipanea hispida var. hispida
Sipanea micrantha Sandwith
Sipanea ovalifolia Bremek.
Sipanea ovalifolia var. ovalifolia
Sipanea ovalifolia var. villosissima Steyerm.
Sipanea pratensis Aubl.
Sipanea pratensis var. dichotoma (Kunth) Steyerm.
Sipanea pratensis var. pratensis
Sipanea setacea Steyerm.
Sipanea stahelii Bremek.
Sipanea veris S. Moore
Sipanea wilson-brownei R.S. Cowan
Sipaneopsis cururuensis J.H. Kirkbr.
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AM
AM SU
GU su FG
AM
AM BO GU
SU FG
BO DA GU su FG
AM
AM
AM BO DA GU
GU
su
su
su
FG
FG?
FG
AM
AM BO
BO
GU su FG
AM
AM BO GU su FG
AM BO
AM BO DA GU su FG?
AM BO DA FG?
AM BO
BO
BO
DA GU su FG
AM
AM
AM
AM
BO GU
BO GU
GU
GU
su
su
FG
AM
BO
GU
GU
GU
AM
AM BO
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su
FG?
FG
BO DA GU su FG
BO
GU
BO GU
GU
su
AM
AM BO GU FG?
AM BO
GU
su
su
FG
FG
FG
AM BO DA GU su FG
AM BO DA GU su FG
GU su FG
BO
su FG?
AM BO GU
BO GU
GU
FG
492 Checklist of the Plants of the Guiana Shield
Sipaneopsis foldatsii Steyerm. AM
Sipaneopsis hubeh Steyerm. AM
Sipaneopsis maguirei Steyerm. AM
Sipaneopsis morichensis Steyerm. AM
Sipaneopsis pacimoniensis Steyerm. AM
Sipaneopsis rupicola (Spruce ex K. Schum.) Steyerm. AM
Spermacoce glabra Michx.
Spermacoce hexangularis Aubl. [name of uncertain application]
Spermacoce radicans Aubl. [=Diodia sp.?]
Sphinctanthus maculatus Spruce ex K. Schum.
Sphinctanthus striiflorus (DC.) Hook. f. AM
Stachyarrhena duckei Standi. AM
Stachyarrhena penduliflora K. Schum. AM
Stachyarrhena reticulata Steyerm. AM
Stachyarrhena spicata Hook. f. AM
Tobagoa maleolens Urb.
Tocoyena brevifolia Steyerm. AM
Tocoyenaformosa (Cham. & Schhdl.) K. Schum.
Tocoyena guianensis K. Schum. AM
Tocoyena longiflora Aubl.
Tocoyena neglecta N.E. Br.
Tocoyena orinocensis Standi. & Steyerm.
Tocoyena pendulina Spruce ex Standi.
Tocoyena surinamensis Bremek.
Uncaria guianensis (Aubl.) J.F. Gmel.
Uncaria tomentosa (Willd. ex Roem. & Schult.) DC.
Warszewiczia coccinea (Vahl) Klotzsch
Warszewiczia elata Ducke
Warszewiczia schwackei K. Schum.
Wittmackanthus stanleyamis (R.H. Schomb.) Kuntze
Synonyms and Misidentifications
Alibertia acuminata (Benth.) Sandwith =Alibertia edulis var. edulis
Alibertia acuminata var. acuminata =Alibertia edulis var. edulis
Alibertia acuminata var. obtusiuscula Steyerm. =Alibertia edulis var. obtusiuscula
Alibertia benensis Standi. =Cordiera triflora
Alibertia davidsei Steyerm. =Alibertia edulis var. obtusiuscula
Alibertia granulosa Rusby =Alibertia latifolia
Alibertia hexagyna H. Karst. ^Alibertia edulis var. edulis
Alibertia lanceolata (Cham.) Mennega [nomen nudum]
Alibertia latifolia var. pargueniana Steyerm. =Alibertia latifolia
Alibertia latifolia var. parvifolia K. Schum. =Alibertia latifolia
Alibertia longistipulata L. Riley =Alibertia edulis var. edulis
Alibertia myrciifolia Spruce ex K. Schum. =Cordiera myrciifolia
Alibertia myrciifolia var. myrciifolia =Cordiera myrciifolia
Alibertia myrciifolia var. tepuiensis Steyerm. =Cordiera myrciifolia
Alibertia nitidula (Bremek.) L. Andersson =Alibertia edulis var. edulis
Alibertia panamensis L. Riley =Alibertia edulis var. edulis
Alibertia steinbachii Standi. =Cordiera triflora
Alibertia stenantha Standi. =Alibertia bertierifolia
Alibertia surinamensis (Bremek.) Steyerm. =Ibetralia surinamensis
Alibertia tenuifolia K. Krause =Cordiera triflora
Alibertia tobagensis Sprague & R.O. Williams ^Alibertia edulis var. edulis
Alibertia triflora (A. Rich.) K. Schum. =Cordiera triflora
Alibertia triloba Steyerm. =Cordiera myrciifolia
Alibertia trinitatis Sprague & R.O. Williams =Alibertia edulis var. edulis
Alibertia tutumilla Rusby =Alibertia edulis var. edulis
Alibertia uniflora Standi. =Cordiera myrciifolia
Alibertia utilis A. Rich. =Alibertia edulis
Alseis leiantha S.F. Blake =Alseis labatioides
Alseis trichocarpa Standi. & Steyerm. =Alseis labatioides
Amaioua fusifera Spruce ex Benth. 8c Hook. f. [nomen nudum] =Duroiafusifera
Amaioua genipoides Spruce ex K. Schum. [nomen nudum] ^Duroia genipoides
Amaioua peruviana Desf. =Amaioua corymbosa
GU
BO
BO
BO
FG
FG
FG
FG?
GU?
GU
GU?
GU SU FG
FG
BO GU
AM BO
AM
SU
AM BO DA GU SU FG
AM DA GU SU FG
AM GU SU FG
AM
GU
Checklist of the Plants of the Guiana Shield 493
Amaioita saccifera Mart, ex Roem. & Schult. =Duroia saccifera
Amaioua iirsina Standi. =Duroia eriopila var. eriopila
Amaioua velutina Spruce ex Benth. & Hook, [nomen nudum] =Duroia velutina
Anisomeris caiirensis Standi. =Chomelia caurensis
Anisomeris malaneoides (Miill. Arg.) Standi. =Chomelia malaneoides
Anisomeris obtusa (Cham. & Schltdl.) K. Schum. =Chomelia obtusa
Anisomeris polyantha (S.F. Blake) Rusby =Chomelia polyantha
Anisomeris tenuiflora (Benth.) Pulle =Chomelia tenuiflora
Anisomeris volubilis Standi. =Chomelia volubilis
Antirhea surinamensis Bremek. =Guettarda acreana
Aphanocarpus steyermarkii f. elongatus Steyerm. =Aphanocarpus steyermarkii
Aphanocarpus steyermarkii f. glabrior Steyerm. =Aphanocarpus steyermarkii
Appunia angulata (Benth.) Baill. =Morinda peduncularis
Appunia aurantiaca (K. Krause) Sandwith =Morinda aurantiaca
Appunia brachycalyx (Bremek.) Steyerm. =Morinda brachycalyx
Appunia calycina (Benth.) Sandwith =Morinda calycina
Appunia debilis Sandwith =Morinda debilis
Appunia longiloba Steyerm. ^Morinda longiloba
Appunia sparsiflora Steyerm. =Morinda sparsiflora
Appunia surinamensis (Bremek.) Steyerm. =Morinda surinamensis
Appunia tenuiflora (Benth.) Jacks. & Hook. f. =Morinda tenuiflora
Appunia tenuiflora var. leiophylla Steyerm. =Morinda tenuiflora
Appunia venezuelensis Steyerm. =Morinda venezuelensis
Basanacantha asperifolia Sandwith =Randia asperifolia
Basanacantha dioica (H. Karst.) Hook. f. =Randia dioica
Basanacantha hebecarpa (Benth.) Hook. f. =Randia hebecarpa
Basanacantha spinosa (Jacq.) K. Schum. =Randia armata
Basanacantha spinosa f. macrophylla K. Schum. =Randia amazonasensis
Bellardia repens Willd. =Coccocypselum guianense
Bellardia tontanea (Kunth) Schult. & Schult. f. =Coccocypselum guianense
Bellynkxia angulata (Benth.) Miill. Arg. =Morinda peduncularis
Bellynkxia brachycalyx Bremek. =Morinda brachycalyx
Bellynkxia surinamensis Bremek. =Morinda surinamensis
Bertiera guianensis subsp. guianensis =Bertiera guianensis
Bertiera palustris A. Rich. =Sipanea wilson-brownei
Billardiera paniculata Vahl =Coussarea paniculata
Borojoa lanceolata (Cham.) Cuatrec. =Alibertia edulis var. edulis
Borreria assurgens (Ruiz & Pav.) Griseb. =Borreria remota
Borreria capitata i. ferruginea (A. St.-Hil.) Steyerm. =Borreria capitata var. capitata
Borreria capitata f. glabra Steyerm. =Borreria capitata var. capitata
Borreria exilis L.O. Williams =Borreria repens
Borreriaferruginea (A. St.-Hil.) DC. =Borreria capitata var. capitata
Borreriafockeana Miq. =Borreria latifolia
Borreria hispida var. glabrescens K. Schum. =Borreria hispida
Borreria hyssopifolia (Willd. ex Roem. & Schult.) Bacigalupo & E.L. Cabral =Diodia hyssopifolia
Borreria kappleriana Miq. =Borreria capitata var. capitata
Borreria laevis (Lam.) Griseb. [not in flora area] =Spermacoce tenuoir L.
Borreria laevis auct., non (Lam.) Griseb. =Borreria remota
Borreria latifolia var. fockeana (Miq.) Bremek. =Borreria latifolia
Borreria latifolia var. minor K. Schum. =Borreria latifolia
Borreria latifolia f. minor (K. Schum.) Steyerm. =Borreria latifolia
Borreria multiflora (DC.) Bacigalupo & E.L. Cabral =Diodia multiflora
Borreria ocimoides auct., non (Burm. f.) DC. =Borreria prostrata
Borreria ocymifolia (Willd. ex Roem. & Schult.) Bacigalupo & E.L. Cabral =Diodia ocymifolia
Borreria oligodonta Steyerm. =Borreria verticillata
Borreria spicata (Miq.) Bacigalupo & E.L. Cabral =Diodia spicata
Borreria splitgerberi Bremek. =Borreria latifolia
Borreria suaveolens G Mey. =Borreria capitata var. suaveolens
Borreria tenella (Kunth) Cham. & Schltdl. =Borreria capitata var. tenella
Borreria tenella war. platyphylla K. Schum. =Borreria capitata var. capitata
Borreria umbellata Spreng. =Emmeorhiza umbellata
Brignolia pubigera Benth. =Isertia parviflora
Bruinsmania isertioides Miq. =Isertia parviflora
Buchia plantaginea Kunth =Perama plantaginea
494 Checklist of the Plants of the Guiana Shield
Buena triflora Benth. =Cosmibuena grandiflora
Callicocca alba (Aubl.) J.F. Gmel. =Psychotria ulviformis
Callicocca purpurea (Aubl.) J.F. Gmel. =Psychotria variegata
Callicocca tomentosa (Aubl.) J.F. Gmel. =Psychotria poeppigiana
Calycophyllum intonsum Steyerm. =Semaphyllanthe intonsa
Calycophyllum merumense Steyerm. =Semaphyllanthe merumensis
Calycophyllum spectabile Steyerm. =SemaphyUanthe spectabilis
Capirona leiophloea Benoist =Capirona decorticans
Capirona surinamensis Bremek. =Capirona decorticans
Capirona wurdackii Steyerm. =Capirona decorticans
Carapichea guianensis Aubl. =Psychotria ligularis var. carapichea
Carapichea kappleri Miq. =Psychotria kappleri
Cascarilla roraimae (Benth.) Wedd. =Remijia roraimae
Cassupa verrucosa Bonpl. =Isertia verrucosa
Cephaelis aetantha Sandwith =Psychotria aetantha
Cephaelis alba (Aubl.) Willd. =Psychotria ulviformis
Cephaelis altsonii Sandwith =Psychotria altsonii
Cephaelis amoena Bremek. =Psychotria colorata
Cephaelis aneurophylla Standi. =Notopleura aneurophylla
Cephaelis axillaris Sw. =Psychotria aubletiana
Cephaelis barcellana (Miill. Arg.) Standi. =Psychotria poeppigiana subsp. barcellana
Cephaelis blepharophylla Standi. =Psychotria medusula
Cephaelis bolivarensis Standi. & Steyerm. =Psychotria bolivarensis
Cephaelis bracteocardia DC. =Psychotria bracteocardia
Cephaelis cacuminis Steyerm. =Psychotria aubletiana
Cephaelis callithrix Miq. =Psychotria callithrix
Cephaelis colorata Willd. ex Roem. & Schult. =Psychotria colorata
Cephaelis dichotoma Willd. ex Roem. & Schult. [nom. illeg.] =Psychotria hoffmannseggiana
Cephaelis dichotoma Rudge =Psychotria platypoda
Cephaelis duckei Standi. =Psychotria spectabilis
Cephaelis ernestii K. Krause =Psychotria transiens
Cephaelisfanshawei Standi. =Psychotriafanshawei
Cephaelis glabrescens (Mull. Arg.) Standi, [not in flora area] =Psychotria glabrescens Muell. Arg.
Cephaelis guianensis (Aubl.) Standi. =Psychotria ligularis var. carapichea
Cephaelis hemicephaelis (Wemham) Steyerm. =Psychotria hemicephaelis
Cephaelis hirsuta M. Martens & Galeotti =Psychotria poeppigiana
Cephaelis hirta Miq. =Psychotria bremekampiana
Cephaelis hoffmannseggiana Willd. ex Roem. & Schult. =Psychotria hoffmannseggiana
Cephaelis humboldtiana Cham, =Psychotria humboldtiana
Cephaelis humboldtiana var. caudata Miill. Arg. =Psychotria humboldtiana
Cephaelis humboldtiana var. ornata (Mull. Arg.) Standi. =Psychotria humboldtiana
Cephaelis jenmanii Wemham =Psychotria mazaruniensis
Cephaelis kaieteurensis Wemham =Psychotria appuniana
Cephaelis kappleri (Miq.) Standi. =Psychotria kappleri
Cephaelis ligularis (Rudge) A. Rich. =Psychotria ligularis
Cephaelis microcephala Willd. ex Roem. & Schult. =Psychotria prunifolia
Cephaelis microcephala Miq. [nom. illeg.] =Psychotria hoffmannseggiana
Cephaelis muscosa (Jacq.) Sw. =Psychotria muscosa
Cephaelis nivea Sandwith =Psychotria nivea
Cephaelis oblonga DC. =Psychotria oblonga
Cephaelis paraensis Standi. =Psychotria ulviformis
Cephaelis patrisii D. Dietr. =Psychotria sp.
Cephaelis potaroensis Sandwith =Psychotria potaroensis
Cephaelis prunifolia Kunth =Psychotria prunifolia
Cephaelis pubescens Hoffmanns, ex Roem. «fe Schult. =Psychotria bracteocardia
Cephaelis purpurea (Aubl.) Willd. =Psychotria variegata
Cephaelis rosea Benth. =Psychotria rosea
Cephaelis rubra Willd. ex Roem. & Schult. =Psychotria hoffmannseggiana
Cephaelis sandwithiana Steyerm. =Notopleura sandwithiana
Cephaelis sororiella (Miill. Arg.) Standi. =Psychotria turbinella var. sororiella
Cephaelis sphaerocephala (Mull. Arg.) B.L. Rob. =Psychotria sphaerocephala
Cephaelis stipulosa (Mull. Arg.) Standi. =Psychotria stipulosa
Cephaelis surinamensis Standi. =Psychotria variegata
Cephaelis sylvatica Bremek. =Psychotria ctenophora
Checklist of the Plants of the Guiana Shield 495
Cephaelis tatei Standi. =Psychotria crocochlamys
Cephaelis tepuiensis Steyerm. =Psychotria tepuiensis
Cephaelis tomentosa (Aubl.) Vahl =Psychotria poeppigiana
Cephaelis tontaneoides (Britton & Standi.) R.O. Williams & Cheesman =Psychotria callithrix var. tontaneoides
Cephaelis trichophoroides (Miill. Arg.) Standi. =Psychotria trichophoroides
Cephaelis violacea (Aubl.) Sw. =Psychotria apoda
Cephalodendron aracamuniensis Steyerm. =Remijia aracamuniensis
Cephalodendron globosum Steyerm. =Remijia globosa
Chalepophyllum cohaceum Gleason =Maguireothamnus speciosus subsp. speciosus
Chalepophyllum guianense var. cuneatum Steyerm. =Chalepophyllum guianense
Chalepophyllum latifolium Standi. =Maguireothamnus tatei
Chalepophyllum longilobum Steyerm. =Chalepophyllum guianense
Chalepophyllum pungens Standi. & Steyerm. (Melastomataceae) =Acanthella sprucei
Chalepophyllum speciosum N.E. Br. =Maguireothamnus speciosus
Chalepophyllum tatei Standi. =Maguireothamnus tatei
Chimarrhis bathysoides Steyerm. =Bathysa bathysoides
Chimarrhis longistipulata Bremek. =Chimarrhis microcarpa
Chiococca alba var. micrantha (J.R. Johnst.) Steyerm. =Chiococca alba
Chiococca alba subsp. parvifolia (Wullschl. ex Griseb.) Steyerm. =Chiococca alba
Chiococca alba f. pilosa Steyerm. =Chiococca alba
Chiococca brachiata Ruiz & Pav. =Chiococca alba
Chiococca cordata R.S. Cowan =Chiococca nitida var. amazonica
Chiococca erubescens Wemham ^Chiococca nitida var. nitida
Chiococca micrantha J.R. Johnst. =Chiococca alba
Chiococca parvifolia Wullschl. ex Griseb. =Chiococca alba
Chiococca racemosa L. =Chiococca alba
Chomelia angustifolia Benth. =Chomelia obtusa
Chomelia barbellata Standi. ^Chomelia tenuiflora
Chomelia glabriuscula Steyerm. ^Chomelia malaneoides
Chondrococcus laevis Steyerm. =Coccochondra laevis
Chytropsia astrellantha (Wemham) Bremek. =Psychotria astrellantha
Chytropsia guianensis Bremek. =Psychotria moroidea
Cinchona amazonica Standi. =Cinchonopsis amazonica
Cinchona brasiliensis Hoffmanns, ex Humb. =Machaonia brasiliensis
Cinchonafirmula Mart. =Remijiafirmula
Cinchona grandiflora Ruiz & Pav. =Cosmibuena grandiflora
Cinchona lambertiana A. Braun ex Mart. =Ladenbergia lambertiana
Cinchona magnifolia Ruiz & Pav. =Ladenbergia oblongifolia
Cinchona oblongifolia Mutis =Ladenbergia oblongifolia
Cinchona pedunculata H. Karst. =Remijia pedunculata
Cinchona roraimae Benth. =Remijia roraimae
Coccocypselum aureum var. capitatum (Benth.) Steyerm. =Coccocypselum aureum
Coccocypselum brevipetiolatum Steyerm. =Coccocypselum hirsutum
Coccocypselum canescens Willd. ex Schult. & Schult. f. ^Coccocypselum lanceolatum
Coccocypselum croatii Steyerm. =Coccocypselum condalia
Coccocypselum guianense var. patens Steyerm. =Coccocypselum guianense
Coccocypselum tontanea Kunth ^Coccocypselum guianense
Coffea crassiloba Benth. =Rudgea crassiloba
Coffea laurifolia Kunth [nom. illeg.] =Rudgea sclerocalyx
Coffea paniculata Aubl. =Faramea paniculata
Coffea sessilis Veil. =Psychotria vellosiana
Coffea stipulacea DC. =Rudgea stipulacea
Coffea subsessilis Benth. =Ronabea latifolia
Coffea tenuiflora Benth. =Morinda tenuiflora
Commianthus concolor Spruce ex Benth. =Retiniphyllum concolor
Commianthus discolor Spruce ex Benth. =Retiniphyllum discolor
Commianthus pilosus Spruce ex Benth. =Retiniphyllum pilosum
Commianthus schomburgkii Benth. =Retiniphyllum schomburgkii
Condalia lanceolata Ruiz & Pav. =Coccocypselum lanceolatum
Condalia repens Ruiz & Pav. =Coccocypselum condalia
Conosiphon aureus Poepp. & Endl. =Sphinctanthus striiflorus
Cordiera acuminata Benth. =Alibertia edulis var. edulis
Cordiera bertierifolia (K. Schum.) Kuntze =Alibertia bertierifolia
Cordiera edulis (Rich.) Kuntze =Alibertia edulis
496 Checklist of the Plants of the Guiana Shield
Cordiera hexagyna (H. Karst.) Kuntze =Alibertia edulis var. edulis
Cordiera latifolia Benth. =Alibertia latifolia
Cordiera trifolia Steud. =Cordiera triflora
Cosmibuena triflora (Benth.) Klotzsch =Cosmibuena grandiflora
Coiipoui aquatica Aubl. =Duroia aquatica
Coupoui martiniana (Miers) Wemham =Duroia martiniana
Coupoui micrantha Ladbr. =Diiroia micrantha
Coussarea multiflora (A. Rich.) Lemee =Faramea multiflora
Coussarea paniculata (Aubl.) Lemee, nom. illeg. =Faramea paniculata
Coussarea schomburgkiana (Benth.) Benth. & Hook. f. =Coussarea violacea
Coutarea campanilla DC. =Coiifarea hexandra
Coutarea hexandra var. campanilla (DC.) Steyerm. =Coutarea hexandra
Coutarea hexandra var. pubescens K. Schum. =Coutarea hexandra
Coutarea hexandra f. pubescens (Pohl) Steyerm. =Coutarea hexandra
Coutarea lindeniana Baill. =Coutarea hexandra
Coutarea pubescens Pohl =Coutarea hexatgdra
Coutarea speciosa Aubl. =Coutarea hexandra
Cupirana martiniana Miers =Duroia martiniana
Declieuxia alfredii Ernst =Declieuxiafruticosa
Declieuxia brasiliensis (Spreng.) Mtill. Arg. =Declieuxia tenuiflora
Declieuxia chiococcoides Kunth =Declieuxiafruticosa
Declieuxia chiococcoides var. guyanensis Miill. Arg. =Declieuxiafruticosa
Declieuxiafruticosa var. guyanensis (Miill. Arg.) Standi. =Declieuxiafruticosa
Declieuxiafruticosa var. mexicana (DC.) Standi. "^Declieuxia fruticosa
Declieuxia mexicana DC. =Declieuxiafruticosa
Declieuxia roraimensis Wemham ^Psychotria capitata subsp. inundata
Dendrosipanea wurdackii Steyerm. =Dendrosipanea spigelioides
Dicrobotryum divaricatum Humb. & Bonpl. ex Roem. & Schult. =Guettarda divaricata
Diodella rigida (Cham. & Schltdl.) Small =Diodia apiculata
Diodella teres (Walter) Small =Diodia teres
Diodia hyssopifolia var. articulata (Pohl ex DC.) Steyerm. =Diodia hyssopifolia
Diodia hyssopifolia f. glabriuscula Steyerm. =Diodia hyssopifolia
Diodia hyssopifolia var. linearis (Willd. ex Roem. & Schult.) Steyerm. =Diodia hyssopifolia
Diodia hyssopifolia f. psiloclada Steyerm. =Diodia hyssopifolia
Diodia ottonis K. Schum. ex Pittier =Tobagoa maleolens
Diodia prostrata Sw. =Diodia teres
Diodia rigida Cham. & Schltdl. =Diodia apiculata
Diodia setigera DC. =Diodia apiculata
Diodia surinamensis Miq. =Diodia hyssopifolia
Diodia teres var. angustata A. Gray =Diodia teres
Diodia teres subsp. angustata (A. Gray) Steyerm. =Diodia teres
Diodia teres f. latifolia Steyerm. =Diodia teres
Diodia teres f. latior Steyerm. =Diodia teres
Diodia teres f. leiocarpa Steyerm. =Diodia teres
Diodia teres subsp. prostrata (Sw.) Steyerm. =Diodia teres
Duroia eriopila f. glabra Steyerm. =Duroia eriopila var. eriopila
Duroia genipoides Spruce ex Hook. f. [nomen nudum] =Duroia genipoides
Duroia sprucei Rusby =Duroia micrantha
Euosmia corymbosa Benth. =Bothriospora corymbosa
Evea guianensis Aubl. =Faramea guianensis
Evea tomentosa (Aubl.) Standi. =Psychotria poeppigiana
Evea tontaneoides Britton & Standi. =Psychotria callithrix var. tontaneoides
Faramea anisocalyx var. pulchella (Miill. Arg.) Steyerm. =Faramea anisocalyx
Faramea leptoloba Benth. & Hook. f. =Coussarea leptoloba
Faramea longifolia Benth. =Faramea sessilifolia
Faramea multiflora var. epedunculata Steyerm. =Faramea multiflora
Faramea multiflora var. salicifolia (C. Presl) Steyerm. =Faramea multiflora
Faramea occidentalis var. meridionalis Steyerm. =Faramea occidentalis
Faramea pachydictyon Miill. Arg. =Faramea torquata
Faramea paludicola Steyerm. =Faramea torquata
Faramea pulchella Miill. Arg. =Faramea anisocalyx
Faramea salicifolia C. Presl =Faramea multiflora
Faramea schomburgkiana Benth. =Coussarea violacea
Faramea truncata DC. =Faramea corymbosa
Checklist of the Plants of the Guiana Shield 497
Faramea umbelliflora Bremek. =Faramea quinqueflora
Galium croceum Ruiz & Pav. ^Galium hypocarpium
Gamotopea alba (Aubl.) Bremek. =Psychotria ulviformis
Gamotopea callithhx (Miq.) Bremek. =Psychotria callithrix
Gamotopea oblonga (DC.) Lemee =Psychotria oblonga
Gamotopea purpurea (Aubl.) Bremek. =Psychotria variegata
Gamotopea surinamensis (Standi.) Bremek. =Psychotria variegata
Garapatica edulis (Rich.) H. Karst. =Alibertia edulis
Gardenia armata Sw. =Randia armata
Gardenia edulis (Rich.) Poir. =Alibertia edulis
Genipa caruto Kunth =Genipa americana var. caruto
Genipa edulis Rich. ^Alibertia edulis
Genipa ? striiflora DC. =Sphinctanthus striiflorus
Geophila herbacea (Jacq.) K. Schum. =Geophila repens
Geophila paraensis Huber ex Standi, [nomen nudum] =Psychotria ulviformis
Geophila picta Rolfe =Psychotria ulviformis
Geophila violacea (Aubl.) DC. ? =Coccocypselum sp.
Gleasonia duidana var. oblanceolata Steyerm. =Gleasonia duidana var. duidana
Gonzalagunia dicocca var. guianensis Steyerm. =Gonzalagunia dicocca
Gonzalagunia dicocca subsp. venezuelensis Steyerm. =Gonzalagunia dicocca
Guettarda coccinea Aubl. =Isertia coccinea
Guettarda grandiflora Steud. =Guettarda aromatica
Guettarda leiantha Steyerm. =Guettarda acreana
Guettarda ulei K. Krause =Guettarda acreana
Hamelia lutea Rohr ex Sm. =Hamelia axillaris
Hamelia patens var. glabra Oerst. =Hamelia patens
Hedyotis lancifolia Schumach. =Oldenlandia lancifolia
Hemidiodia ocymifolia (Willd. ex Roem. & Schult.) K. Schum. =Diodia ocymifolia
Henriquezia aturensis Standi. =Platycarpum orinocense var. orinocense
Henriquezia nitida var. oblonga (Spruce ex Benth.) Steyerm. =Henriquezia nitida var. nitida
Henriquezia oblonga Spruce ex Benth. =Henriquezia nitida var nitida
Hillia tubiflora Cham. =Hillia illustris
Houstoniafruticosa Willd. ex Roem. & Schult. =Declieuxiafruticosa
Isertia commutata Miq. =Isertia spiciformis
Isertia parviflora var. hirta Steyerm. =Isertia parviflora
Isertia pterantha Bremek. =Isertia spiciformis
Ixora alternifolia Jacq. (Solanaceae) =Cestrum alternifolium
Ixora angulata Benth. =Morinda peduncularis
Ixora duidae Standi. =Rudgea stipulacea
Ixorafloribunda (Rich.) Griseb. [not in flora area]
Ixora gleasonii Standi. =Ixora schomburgkiana
Ixora guianensis (Aubl.) Spreng. [see Coffea guianensis']
Ixora mazarunensis Standi. =Ixora davisii
Ixora occidentalis L. =Faramea occidentalis
Ixora orinocensis Spruce ex Standi, [nomen nudum]
Ixora podocarpa Benth. =Simira pisoniiformis
Ixora surinamensis Bremek. =Ixora graciliflora
Ixora tenuiflora (DC.) Lemee =Patabea tenuiflora
Ixora xantholoba Standi. =Ixora graciliflora
Knoxia brasiliensis Spreng. =Declieuxia tenuiflora
Ladenbergia lucens Standi. =Ladenbergia lambertiana
Ladenbergia magnifolia (Ruiz & Pav.) Klotzsch =Ladenbergia oblongifolia
Ladenbergia puberula Steyerm. =Ladenbergia lambertiana
Ladenbergia roraimae (Benth.) Klotzsch =Remijia roraimae
Ladenbergia schomburgkii Klotzsch =Ladenbergia lambertiana
Ladenbergia venamoensis Steyerm. =Ladenbergia lambertiana
Limnosipanea guaricensis Pittier =Limnosipanea spruceana
Limnosipanea kuntzei Standi. =Limnosipanea spruceana
Limnosipanea schomburgkii Hook. f. =Limnosipanea palustris
Limnosipanea ternifolia Pittier =Limnosipanea spruceana
Lonicera alba L. =Chiococca alba
Macrocnemum coccineum Vahl =lVarszewiczia coccinea
Maguireothamnus jauaensis (Steyerm.) Steyerm. =Maguireothamnus speciosus suhsp.jauaensis
Maguireothamnus jauaensis var. breweri Steyerm. =Maguireothamnus speciosus subsp. Jauaensis
498 Checklist of the Plants of the Guiana Shield
Maguireothamnus tatei var. latifolius (Standi.) Steyerm. =Maguireothamnus tatei
Malanea angustifolia Bartl. [nomen nudum] =Malanea obovata
Malanea macrophylla Bartl. ex Griseb. [nomen nudum] =Malanea macrophylla
Malanea wraimensis Wemham =Malanea obovata
Malanea sarmentosa f. tomentosa Steyerm. =Malanea sarmentosa
Manettia coccinea var. spraguei (Wemham) Steyerm. =Manettia coccinea
Manettia iiniflora Kunth =Manettia reclinata
Mapouria alba (Ruiz & Pav.) Miill. Arg. =Psychotna alba
Mapouria borjensis (Kunth) Miill. Arg. =Psychotria borjensis
Mapouria brownei (Spreng.) Lemee [not in flora area] =Psychotria brownei Spreng.
Mapouria chionantha (DC.) Miill. Arg. ^Psychotria mapourioides
Mapouria cupularis Miill. Arg. =Psychotria cupularis
Mapouriaficigemma (DC.) Lemee =Psychotria carthagenensis
Mapouriafockeana (Miq.) Bremek. =Psychotria carthagenensis
Mapouria guianensis Aubl. =Psychotria mapourioides
Mapouria herbacea var. orbicularis Miill. Arg. =Geophila orbicularis
Mapouria mahuriensis Bremek. =Psychotria cupularis
Mapouria microdon (DC.) Bremek. =Psychotria microdon
Mapouria opaca Bremek. =Psychotria mapourioides
Mapouria ornithophila Bremek. =Psychotria anceps
Mapouria patrisii (DC.) Lemee =Psychotria anceps
Mapouria podocephala Miill. Arg. =Psychotria podocephala
Mapouria remota (Benth.) Miill. Arg. =Psychotria remota
Mapouria sclerocalyx Miill. Arg. =Rudgea sclerocalyx
Mapouria sororia (DC.) Lemee [dubious name] =Psychotria sp.
Mapouria spruceana Miill. Arg. =Rudgea sclerocalyx
Mapouria subsessilis (Benth.) Miill. Arg. =Ronabea latifolia
Mapouria subsessilis var. [gamma] angustifolia Miill. Arg. =Ronabea latifolia
Mapouria tenuis Miill. Arg. =Geophila tenuis
Mapouria umbrosa Miill. Arg. =Psychotria borjensis
Mapouria vasivensis Miill. Arg. =Psychotria vasivensis
Mapouria wonotobensis Bremek. =Psychotria wonotobensis
Margaritopsis astrellantha (Wemham) L. Andersson =Psychotria astrellantha
Mattuschkea galioides Kunth =Perama galioides
Mattuschkea hispida Kunth =Perama galioides
Mitracarpus discolor Miq. =Mitracarpus humboldtianus
Mitracarpusfruticosus Standi. =Mitracarpusfrigidus war. fruticosus
Mitracarpus puberulus Benth. =Borreria prostrata
Mitracarpus rude Benth. =Mitracarpus hirtus
Mitracarpus villosus (Sw.) DC. =Mitracarpus hirtus
Morinda tenuiflora var. leiophylla (Steyerm.) Steyerm. =Morinda tenuiflora
Mussaendaformosa Jacq. =Rosenbergiodendronformosum
Mussaenda spinosa Jacq. =Randia armata
Nacibea alba Aubl. ^Manettia alba
Nacibea coccinea Aubl. =Manettia coccinea
Naletonia violacea (Aubl.) Bremek. =Psychotria capitata subsp. amplifolia
Nauclea tomentosa Willd. ex Roem. & Schult. =Uncaria tomentosa
Neblinathamnus aracamunianus Steyerm. =Neblinathamnus brasiliensis
Neblinathamnus glabratus Steyerm. =Neblinathamnus argyreus
Nonatelia grandiflora Kunth =Palicourea grandiflora
Nonatelia longiflora Aubl. =Palicourea longiflora
Nonatelia lutea Aubl. [=Palicourea longiflora!] =Palicourea lutea
Nonatelia lutea Aubl. ex Steud. =Palicourea longiflora
Nonatelia macrophylla Kunth =Palicourea grandifolia
Nonatelia officinalis Aubl. =Psychotria officinalis
Nonatelia paniculata Aubl. =Psychotria paniculata
Nonatelia racemosa Aubl. =Psychotria racemosa
Nonatelia secundiflora (Bonpl.) Spreng. =Retiniphyllum secundiflorum
Nonatelia violacea Aubl. =Psychotria capitata
Oldenlandia herbacea (L.) DC. [not in flora area]
Oribasia racemosa (Aubl.) J.F. Gmel. =Psychotria racemosa
Ourouparia guianensis Aubl. =Uncaria guianensis
Ourouparia tomentosa (Willd. ex Roem. & Schult.) K. Schum. =Uncaria tomentosa
Pagamea auyantepuiensis Steyerm. =Coryphothamnus auyantepuiensis
Checklist of the Plants of the Guiana Shield 499
Pagamea brevipedimcidata Steyerm. =Pagamea capitata
Pagamea capitata f. breviloba Steyerm. =Pagamea capitata subsp. conferta
Pagamea conferta Standi. =Pagamea capitata subsp. conferta
Pagamea coriacea var. piibescens Steyerm. =Pagamea coriacea
Pagamea gariyoides Standi. =Pagameopsis gariyoides
Pagamea plicata var. multinervia Steyerm. =Pagamea plicata
Pagamea reducta Steyerm. =Pagamea standleyana
Pagamea spadicea Pittier =Psychotria spadicea
Pagamea steyermarkii Standi. =Aphanocarpus steyermarkii
Pagamea uniflora Steyerm. =Pagamea standleyana
Pagameopsis maguirei var. angiistifoUus Steyerm. =Pagameopsis maguirei
Pagameopsis maguirei var. glabnis Steyerm. =Pagameopsis maguirei
Pagameopsis maguirei subsp. neblinensis Steyerm. =Pagameopsis maguirei
Pagameopsis maguirei subsp. pus iIlus Steyerm. =Pagameopsis maguirei
Palicourea "latifolia (Rudge) Roem. & Schult." [invalid name, error for Posoqueria latifolia]
Palicourea aculeifera Steyerm. =Palicourea longistipulata subsp. chrysorrhachis
Palicourea cayennensis Desv. =Palicourea pedunculosa
Palicourea chionantha DC. =Psychotria mapourioides
Palicourea chrysorrhachis Bremek. =Palicourea longistipulata subsp. chrysorrhachis
Palicourea crocea f. heterodoxa Steyerm. =Palicourea croceoides
Palicourea crocea var. riparia (Benth.) Griseb. =Palicourea croceoides
Palicourea divaricata Schltdl. =Psychotria schlechtendaliana
Palicourea elliptica Kunth =Psychotria ernestii
Palicourea expetens Standi. =Palicourea corymbifera
Palicourea grandifolia var. sprucei (Miill. Arg.) Steyerm. =Palicourea grandifolia
Palicourea guianensis f. glabra Steyerm. =Palicourea guianensis subsp. guianensis
Palicourea guianensis subsp. occidentalis Steyerm. =Palicourea guianensis subsp. guianensis
Palicourea guianensis var. tetramera Bremek. ^Palicourea guianensis subsp. guianensis
Palicourea guianensis var. trimera Bremek. =Palicourea guianensis subsp. guianensis
Palicourea longiflora (Aubl.) A. Rich. =Palicourea longiflora DC.
Palicourea nicotianifolia auct., not Cham. & Schltdl. =Palicourea calophylla
Palicourea rigida var. hirtibacca Steyerm. =Palicourea rigida
Palicourea riparia Benth. =Palicourea croceoides
Palicourea roraimae Standi. =Psychotria oblita
Palicourea sprucei {MnW. Arg.) K. Schum. =Palicourea grandifolia
Patabea alba Kunth =Psychotria lupulina subsp. rhodoleuca
Patabea coccinea Aubl. ^^Ixora davisii
Patima laxiflora Benth. =Retiniphylhim laxiflorum
Perama dichotoma var. hirsutula Steyerm. =Perama dichotoma var. dichotoma
Perama dichotoma var. scaposa (Gleason & Standi.) Steyerm. =Perama dichotoma var. dichotoma
Perama galioides f. cinera Steyerm. =Perama galioides
Perama galioides var. densipila Steyerm. =Perama galioides
Perama galioides var. hispida (Kunth) Steyerm. =Perama galioides
Perama galioides var. intermedia Steyerm. =Perama galioides
Perama galioides var. longiflora Steyerm. =Perama galioides
Perama galioides f. macrocephala Steyerm. =Perama galioides
Perama galioides f. quaternata Steyerm. =Perama galioides
Perama hirsuta var. stricta (Benth.) Bremek. =Perama hirsuta
Perama scaposa Gleason & Standi. =Perama dichotoma var. dichotoma
Perama stricta Benth. =Perama hirsuta
Petagomoa atricapilla Bremek. =Psychotria iodotricha
Petagomoa glabra Bremek. =Psychotria sp.
Petagomoa hirta (Miq.) Bremek. =Psychotria hremekampiana
Petagomoa nigricans Bremek. =Psychotria trichophoroides
Petagomoa tricholoba (Miill. Arg.) Bremek. =Psychotria tricholoba
Portlandia hexandra Jacq. =Coutarea hexandra
Posoqueria gracilis (Rudge) Roem. & Schult. =Posoqueria latifolia subsp. gracilis
Posoqueria intermedia Miq. =Posoqueria trinitatis
Posoqueria panamensis subsp. grandiflora (H. Karst.) Steyerm. =Posoqueria panamensis
Pseudochimarrhis difformis Benoist =Chimarrhis sp.
Pseudochimarrhis turbinata Benoist =Chimarrhis turhinata
Psychotria alboviridula K. Krause =Psychotria remota
Psychotria aligera Steyerm. =Notopleura aligera
Psychotria amazonica Miill Arg. =Palicourea grandifolia
500 Checklist of the Plants of the Guiana Shield
Psychotr,
Psychotr,
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a amplifolia Raeusch. =Psychotria capitata subsp. amplifolia
a amplinoda Steyerm. =Notopleura tapajozensis
a amplinoda van ayangannae Steyerm. =Notopleura tapajozensis
a aneurophylla (Standi.) Steyerm. =Notopleura aneurophylla
a anomothyrsa K. Schum. & Donn. Sm. =Notopleura anomothyrsa
a arcuata Benth. =Psychotria capitata subsp. inundata
a arenaria Standi. & Steyerm. =Psychotria ceratantha
a atabapoensis Standi. =Psychotria subundulata
a atricapilla Bremek. =Psychotria iodotricha
a aubletiana var. andina Steyerm. =Psychotria aubletiana
a aubletiana var. cacuminis (Steyerm.) Steyerm. =Psychotria aubletiana
a aubletiana f. tomentella Steyerm. =Psychotria aubletiana
a auyantepuiensis Standi. =Psychotria imthurniana
a avia Standi. & Steyerm. =Rudgea stipulacea
a axillaris Willd. =Ronabea latifolia
a bahiensis var. bahiensis (sensip Steyerm.) [P. bahiensis is endemic to Bahia, Brazil] =Psychotria acuminata
a bahiensis var. cornigera (Benth.) Steyerm. =Psychotria cornigera
a barbiflora DC. =Psychotria hoffmannseggiana
a barcellana Miill. Arg. =Psychotria poeppigiana subsp. barcellana
a berteroana subsp. venezuelica Steyerm. =Psychotria berteroana
a blepharophylla (Standi.) Steyerm. =Psychotria medusula
a brachybotrya Miill. Arg. =Psychotria gracilenta
a brachyloba Miill. Arg. =Palicourea brachyloba
a bracteata DC. =Psychotria pseudinundata
'a bracteata var. intermedia Miill. Arg. =Psychotria maguireorum
a bracteata var. latifolia Miill. Arg. =Psychotria maguireorum
a bracteata var. tenuifolia Miill. Arg. =Psychotria maguireorum
a breweri Steyerm. =Psychotria tatei
a cacuminis Steyerm. =Psychotria aubletiana
a cacuminis Standi. =Psychotria everardii
a calviflora Steyerm. ='Psychotria colorata
a capitata var. roraimensis (Wemham) Steyerm. =Psychotria capitata subsp. inundata
a capitata var. septentrionalis Steyerm. =Psychotria capitata subsp. inundata
a cardiomorpha suhsp. perpusilla (Steyerm.) CM. Taylor & A. Pool -Psychotria cardiomorpha
a chionantha DC. =Psychotria mapourioides
a chlorantha Benth. =Psychotria anceps
a chondroloma Standi. =Psychotria schomburgkii
a cincta Standi. =Psychotria longicuspis
'a cordifolia Kunth [nom. illeg.] =Psychotria cardiomorpha
a cordifolia subsp. perpusilla Steyerm. [nom. illeg.] =Psychotria cardiomorpha
a cornifolia Kunth ^'Rudgea cornifolia
a corymbifera Mull. Arg. =Palicourea corymbifera
a coussareoides var. ciliata Steyerm. =Psychotria coussareoides
a crassa Benth. =Notopleura crassa
a crassa f. alba Steyerm. =Notopleura crassa
a crassa f. angustior Steyerm. =Notopleura crassa
a crocea Sw. =Palicourea crocea
a cubitalis Standi. & Steyerm. =Psychotria capitata subsp. inundata
a cuspidata Bredem. ex Roem. & Schult. [not in flora area]
a davidsei Steyerm. [nom. illeg.] =Psychotria berryi
a deflexa subsp. campyloneura (Miill. Arg.) Steyerm. =Psychotria campyloneura
a deflexa subsp. venulosa (Mull. Arg.) Steyerm. =Psychotria venulosa
a dichotoma (Rudge) Bremek. [nom. illeg.] =Psychotria platypoda
a didymocarpos (A. Rich.) Lemee =Psychotria acuminata
a edaphothrix Steyerm. =Palicourea bracteosa
a erecta (Aubl.) Standi. & Steyerm. =Ronabea latifolia
a erecta f.fluctuans (Standi.) Steyerm. =Ronabea latifolia
a erythrophylla Miill. Arg. =Psychotria hoffmannseggiana
a everardii subsp. neblinensis Steyerm. =Psychotria everardii
a everardii subsp. sipapoensis Steyerm. =Psychotria duricoria var. duricoria
a everardii subsp. tatei (Standi.) Steyerm. =Psychotria tatei
afastigiata (Kunth) Spreng. =Palicourea fastigiata
afimbriata Benth. -Rudgea cornifolia
aflavovirens Suess. ^Psychotria lupulina subsp. rhodoleuca
Checklist of the Plants of the Guiana Shield 501
PsychotriaflexuosaWiWd. [nom. superfl.] =Psychotria paniculata
Psychotriafluctuans Standi. =Ronabea latifoUa
Psychothafluctuans var. angustifolia (Miill. Arg.) Standi. =Ronabea latifolia
Psychotriafockeana Miq. =Psychotria carthagenensis
Psychotria foetidiflora Standi. =Psychotria boliviana
Psychotriafoveolata Ruiz & Pav. =Psychotria carthagenensis
Psychotria glabrescens Miill. Arg. [not in flora area]
Psychotria glandulicalyx var. densipubens Steyerm. =Psychotria glandulicalyx
Psychotria glaucescens Kunth =Psychotria horizontalis
Psychotria grandifolia Willd. ex Roem. & Schult. =Palicourea grandifolia
Psychotria guadalupensis (DC.) R.A. Howard =Notopleura guadalupensis
Psychotria guianensis (Aubl.) Rusby [nom. illeg., non Psychotria guianensis (Aubl.) Raeusch.] =Palicourea guianensis
Psychotria guianensis (Aubl.) Raeusch. [nomen nudum] =Palicourea guianensis
Psychotria hancorniifolia Benth. =Psychotria vellosiana
Psychotria hancorniifolia var. angustifolia Miill. Arg. =Psychotria vellosiana
Psychotria hancorniifolia var. longifolia Miill. Arg. =Psychotria vellosiana
Psychotria heterocarpa Standi. & Steyerm. =Psychotria capitata subsp. inundata
Psychotria hoffmannseggiana var. erythrophylla (Miill. Arg.) Steyerm. =Psychotria hoffmannseggiana
Psychotria homoplastica Moore =Psychotria stipulosa
Psychotria horizontalis var. glaucescens (Kunth) Steyerm. =Psychotria horizontalis
Psychotria horizontalis var. psilophylla Steyerm. =Psychotria horizontalis
Psychotria humboldtiana var. ornata Miill. Arg. =Psychotria humboldtiana
Psychotria imthurniana i. ptariensis (Standi. & Steyerm.) Steyerm. =Psychotria imthurniana
Psychotria inundata K. Krause [nom. illeg.] =Rudgea cornifolia
Psychotria inundata Benth. =Psychotria capitata subsp. inundata
Psychotria involucrata Sw. [nom. superfl.] ^Psychotria officinalis
Psychotria iodotricha subsp. atricapilla (Bremek.) Steyerm. =Psychotria iodotricha
Psychotria kukenanensis Steyerm. =Psychotria boliviana
Psychotria lancigera Standi. =Palicourea lancigera
Psychotria longiflora Poir. [nom. illeg.] =Palicourea longiflora
Psychotria longiflora Willd. =Palicourea longiflora
Psychotria longistipula Benth. =Psychotria racemosa
Psychotria longistipulata Miill. Arg. =Palicourea longistipulata
Psychotria loretensis Standi. =Psychotria longicuspis
Psychotria lupulina var. maypurensis (Humb. & Bonpl. ex Roem. & Schult.) Steyerm. =Psychotria lupulina subsp.
rhodoleuca
Psychotria lupulina f. pubens Steyerm. =Psychotria lupulina subsp. rhodoleuca
Psychotria macrophylla Ruiz & Pav. =Notopleura macrophylla
Psychotria macrophylla subsp. anomothyrsa (K. Schum. & Donn. Sm.) Steyerm. =Notopleura anomothyrsa
Psychotria macropoda Ruiz & Pav. =Geophila macropoda
Psychotria maguirei Standi. =Rudgea stipulacea
Psychotria mapourioides var. chionantha (DC.) Steyerm. =Psychotria mapourioides
Psychotria mapourioides var. opaca (Bremek.) Steyerm. =Psychotria mapourioides
Psychotria mapuria Roem. & Schult. [nom. illeg.] =Psychotria mapourioides
Psychotria martiana Miill. Arg. =Psychotria platypoda
Psychotria maypurensis Humb. & Bonpl. ex Roem. & Schult. ^Psychotria lupulina subsp. rhodoleuca
Psychotria merumensis Steyerm. =Notopleura merumensis
Psychotria microbracteata Steyerm. =Notopleura microbracteata
Psychotria microcephala (Willd. ex Roem. & Schult.) Miill. Arg. [nom. illeg.] =Psychotria prunifolia
Psychotria microcephala (Miq.) Miq. =Psychotria hoffmannseggiana
Psychotria microcephala var. tripotamica Mull. Arg. =Psychotria prunifolia
Psychotria microdon var. meridionalis Steyerm. =Psychotria microdon
Psychotria multiramosa Steyerm. =Notopleura multiramosa
Psychotria necopinata Standi. ^Psychotria pacimonica
Psychotria nervosa Benth. =Psychotria lupulina subsp. lupulina
Psychotria nitida Willd. [nom. superfl.] =Psychotria mapourioides
Psychotria nitidella Mull. Arg. =Palicourea nitidella
Psychotria officinalis (Aubl.) Raeusch. [nomen nudum] ^Psychotria officinalis
Psychotria palicoureoides Miill. Arg. [nom. illeg.] =Psychotria microbotrys
Psychotria paradoxa Miill. Arg. =Psychotria amplectens
Psychotria patens Sw. [not in flora area]
Psychotria pedunculosa A. Rich. ^^Palicourea pedunculosa
Psychotria pendula subsp. trinitensis Urb. =Notopleura perpapillifera
Psychotria perpapillifera Steyerm. =Notopleura perpapillifera
502 Checklist of the Plants of the Guiana Shield
Psychotha persimilis Miill. Arg. =Psychotria lupiilina subsp. rhodolenca
Psychotria phaneroloma var. angustior Steyerm. =Psychotria phaneroloma
Psychotha phaneroneiira Standi. =Psychotria everardii
Psychotria ptariensis Standi. & Steyerm. =Psychotria imthurniana
Psychotria quadrifolia Rudge =Palicourea quadrifolia
Psychotria racemifera Standi. =Psychotria polycephala
Psychotria racemosa (Aubl.) Raeusch. [nom. illeg.] =Psychotria racemosa
Psychotria ramiflora Rusby [records due to misidentifications]
Psychotria rhodoleiica Miill. Arg. =Psychotria lupulina subsp. rhodoleuca
Psychotria rigida Bredem. ex Willd. ex Roem. & Schult. =Palicourea rigida
Psychotria romboutsii Bremek. =Psychotria ampJectens
Psychotria ronaldii Steyerm. =Psychotria boliviana
Psychotria roraimensis Wemham ^Psychotria capitata subsp. inundata
Psychotria rosea f. calvescens Steyerm. =Psychotria rosea
Psychotria rubra (Willd. ex Roem. & Schult.) Mull. Arg. =Psychotria hoffmannseggiana
Psychotria rudgei Bremek. =Psychotria lupulina subsp. lupulina
Psychotria sancti-caroli Standi. =Palicourea longistipulata subsp. longistipulata
Psychotria sandwithiana (Steyerm.) Steyerm. =Notopleura sandwithiana
Psychotria saiilensis Steyerm. =Notopleura saiilensis
Psychotria sessilifiora Humb. & Bonpl. ex Schult. (Gentianaceae) =Tapeinostemon sessiliflorum
Psychotria sessilis (Veil.) Miill. Arg. [nom. illeg.] =Psychotria vellosiana
Psychotria sessilis var. angustifolia (Miill. Arg.) Steyerm. [nom. illeg.] =Psychotria vellosiana
Psychotria setifera Benth. =Psychotria capitata subsp. inundata
Psychotria sororiella Miill. Arg. =Psychotria turbinella var. sororiella
Psychotria sororopanensis Standi. & Steyerm. =Psychotria vellosiana
Psychotria speluncae subsp. brevicalyx Steyerm. =Psychotria speluncae
Psychotria speluncae subsp. exserta Steyerm. =Psychotria speluncae
Psychotria spicata Miill. Arg. =Psychotria spiciflora
Psychotria sprucei Miill. Arg. =Palicourea grandifolia
Psychotria steinii Steyerm. =Psychotria leiantha
Psychotria subcuspidata Miill. Arg. =Psychotria cornigera
Psychotria tapajozensis Standi. =Notopleura tapajozensis
Psychotria thesceloantha Steyerm. =Notopleura thesceloantha
Psychotria triphylla (DC.) Miill. Arg. =Palicourea triphylla
Psychotria uliginosa Sw. =Notopleura uliginosa
Psychotria urophylla Schltdl. =Psychotria acuminata
Psychotria vaupesana Standi, ex Steyerm. =Psychotria spiciflora
Psychotria vernicifolia Standi. =Psychotria everardii
Psychotria verrucosa Miill. Arg. =Palicourea coiymbifera
Psychotria viburnoides Kunth =Psychotria anceps
Psychotria violacea Auh\. ? [name of uncertain application] =Coccocypselum sp.
Psychotria xanthocephala Miill. Arg. =Psychotria prunifolia
Psychotria yutajensis Steyerm. ^Psychotria ceratantha
Randia caracasana Standi. =Randia dioica
Randiaformosa (Jacq.) K. Schum. =Rosenbergiodendronformosum
Randiaformosa var. densiflora Bartl. ex K. Schum. =Rosenbergiodendron densiflorum
Randiaformosa var. formosa =Rosenbergiodendronformosum
Randiaformosa var. nitida K. Schum. =Rosenbergiodendronformosum var. nitidum
Randia orinocensis Rusby =Rosenbergiodendronformosum var. nitidum
Randia ruizana DC. =^Rosenbergiodendron longiflorum
Randia spinosa (Jacq.) H. Karst. =Randia armata
Relbunium croceum (Ruiz & Pav.) K. Schum. =Galium hypocarpium
Relbunium glaberrimum Standi. =Galium hypocarpium
Relbunium guadalupense (Spreng.) Urb. [not in flora area]
Relbunium hypocarpium (L.) Hemsl. =Galium hypocarpium
Relbunium hypocarpium subsp. hypocarpium =Galium hypocarpium subsp. hypocarpium
Relbunium hypocarpium subsp. nitidum (Kunth) Ehrend. =Galium hypocarpium
Relbunium nitidum (Kunth) K. Schum. =Galium hypocarpium
Remijia berryi Steyerm. =Remijia pedunculata
Remijia densiflora f glabricalyx Steyerm. =RemiJia densiflora var. densiflora
Remijia spruceana Benth. ex K. Schum. =Remijiafirmula
Retiniphyllum angustiflorum K. Krause =Retiniphyllum concolor
Retiniphyllum cearense Standi. =Retiniphyllum scabrum
Retiniphyllum erythranthum Standi. ^Retiniphyllum scabrum
Checklist of the Plants of the Guiana Shield 503
Retiniphylhim laxifloriim van longilobmn Steyerm. =RetimphyUum laxiflorum var. laxiflorum
RetiniphyUum magiiirei var. reticidatum Steyerm. "^Retiniphyllum maguirei
RetiniphyUum martianum Miill. Arg. =ReUniphyUiim concolor
RetiniphyUum pallidum Miill. Arg. =RetiniphyUum tnmcatum
RetiniphyUum scabrum var. ayangannense Steyerm. = RetinuphyUum scabrum
RetiniphyUum scabrum var. erythranthum (Standi.) Steyerm. =RetiniphyUum scabrum
RetiniphyUum schomburgkii var. angustiflorum Ruber =RetiniphyUum schomburgkii
RetiniphyUum schombui-gkii var. hirticalyx Steyerm. =RetiniphyUum schomburgkii
RetiniphyUum schomburgkii subsp. occidentale Steyerm. =RetiniphyUum schomburgkii
RetiniphyUum truncatum var. angustifoUum Steyerm. =RetiniphyUum truncatum
Richardsonia scabra (L.) A. St.-Hil. =Richardia scabra
Ronabea erecta Aubl. =Ronabea latifoUa
Ronabea latifoUa var. hispidida Bremek. =Ronabea latifolia
Ronabea morindoides A. Rich. =Psychotria morindoides
Rondeletia capitatum Benth. =Coccocypselum aureum
Rondeletia glomerulata Spreng. [dubious name]
Rondeletia microdon DC. =Psychotria microdon
Rondeletia repens L. =Geophila repens
Rondeletia rupicola Spruce ex K. Schum. =Sipaneopsis rupicola
Rondeletia rupicola var. chiribiquetana R.E. Schult. =Sipaneopsis rupicola
Rubia nitida Kunth =Galium hypocarpium
Rudgea avia (Standi. & Steyerm.) Steyerm. =Rudgea stipulacea
Rudgea berryi Steyerm. & Dwyer =Rudgea klugii
Rudgea bolivarensis Steyerm. =Rudgea hostmanniana subsp. hostmanniana
Rudgea Carolina Standi. & Steyerm. =Rudgea lanceifolia
Rudgea corocoroensis Steyerm. =Rudgea hostmanniana subsp. hostmanniana
Rudgea dasyantha Bremek. =Rudgea lanceifolia
Rudgea duidae (Standi.) Steyerm. =Rudgea stipulacea
Rudgeafimbriata (Benth.) Standi. =Rudgea cornifolia
Rudgeafissistipula Miill. Arg. =Rudgea lanceifolia
Rudgea laurifolia Steyerm. =Rudgea sclerocalyx
Rudgea maypurensis Standi. =Rudgea hostmanniana subsp. maypurensis
Rudgea morichensis Steyerm. =Rudgea sclerocalyx
Rudgea obscura Sandwith =Psychotria astrellantha
Rudgea ovalifolia Salisb. =Rudgea lanceifolia
Rudgea schomburgkiana Benth. =Rudgea crassiloba
Rudgea sipapoensis Steyerm. =Rudgea lanceifolia
Rudgea standleyana Steyerm. =Rudgea klugii
Rudgea tillettii Steyerm. =Psychotria coussareoides
Sabicea aspera var. glabrescens K. Schum. =Sabicea glabrescens
Sabicea aspera var. velutina (Benth.) K. Schum. =Sabicea velutina
Sabicea colombiana Wemham =Sabicea panamensis
Sabicea edulis (Rich.) Seem, ex B.D. Jackson =Alibertia edulis
Sabicea glabrescens var. oblongifolia (Miq.) Sandwith =Sabicea oblongifolia
Sabicea guianensis Wemham [nom. illeg.] =Sabicea velutina
Sabicea guianensis (Aubl.) Baill. =Patima guianensis
Sabicea hirsuta Kunth =Sabicea villosa
Sabicea hirsuta var. adpressa Wemham =Sabicea villosa var. adpressa
Sabicea leucotricha K. Krause =Sabicea velutina
Sabicea velutina var. oblongifolia Miq. =Sabicea oblongifolia
Saldanha illustris Veil. =Hillia illustris
Schradera ligularis Rudge =Psychotria ligularis
Schradera spicata Spruce ex Hook. f. [nomen nudum] =Stachyarrhena spicata
Schwenkfeldia aurea Spreng. =Coccocypselum aureum
SemaphyUanthe obovata (Ducke) L. Andersson =Calycophyllum ohovatum
Semaphyllanthe venezuelensis (Steyerm.) L. Andersson =CalycophyUum venezuelense
Sickingia orinocensis Spreng. =Plafycarpum orinocense var. orinocense
Sickingia pisoniiformis Baill. =Simira pisoniiformis
Sickingia tinctoria (Aubl.) Lemee =Simira tinctoria
Simira eiythroxylon var. saxicola Steyerm. =^Simira rubescens
Sipanea acinifolia Spruce ex Sprague =Sipanea veris
Sipanea colombiana Wemham =Sipanea pratensis var. dichotoma
Sipanea coppenamensis Bremek. =Sipanea pratensis var. pratensis
Sipanea dichotoma Kunth =Sipanea pratensis var. dichotoma
504 Checklist of the Plants of the Guiana Shield
Sipanea glowerata var. paucinervia Steyerm. =Sipanea glomerata
Sipanea palustris (A. Rich.) J.H. Kirkbr. =Sipanea wilson-brownei
Sipanea palustris Seem. [nom. illeg.] =Limnosipanea palustris
Sipanea pratensis f. brachycarpa Steyerm. =Sipanea pratensis var. dichotoma
Sipanea pratensis f. breviflora Steyerm. =Sipanea pratensis var. dichotoma
Sipanea pratensis f. glabriloba Steyerm. =Sipanea pratensis var. dichotoma
Sipanea pratensis f. glabrior Steyerm. =Sipanea pratensis var. dichotoma
Sipanea pubinoda Steyerm. =Sipanea pratensis var. dichotoma
Sipanea radicans Endl. =Sipanea biflora
Sipanea rupicola Spruce ex K. Schum. [nomen nudum] =Sipaneopsis rupicola
Sipanea spraguei Wemham =Sipanea veris
Sipaneopsis wurdackii Steyerm. =Sipaneopsis maguirei
Solena gracilis Rudge =Posoqueria latifolia subsp. gracilis
Solena latifolia Rudge =Posoqueria latifolia
Spermacoce alata Aubl. =Borreria alata
Spermacoce apiculata Willd. ex Roem. & Schult. =Diodia apiculata
Spermacoce aspera Aubl. =Mitracarpus hirtus
Spermacoce assurgens Ruiz & Pav. =Borreria remota
Spermacoce aturensis Kunth =Borreria capitata var. tenella
Spermacoce caeridescens Aubl. =Borreria latifolia
Spermacoce capitata Ruiz & Pav. =Borreria capitata
Spermacoce capitellata Willd. ex Roem. & Schult. =Borreria remota
Spermacoce densiflora (DC.) Alain =Borreria densiflora
Spermacoce diffusa Willd. ex Roem. & Schult. =Mitracarpus diffusus
Spermacoce exilis (L.O. Williams) CD. Adams =Borreria repens
Spermacoceferruginea A. St.-Hil. =Borreria capitata var. capitata
Spermacocefrigida Willd. ex Roem. & Schult. =Mitracarpus frigidus
Spermacoce guianensis Bremek. =Borreria remota
Spermacoce hirta L. =Mitracarpus hirtus
Spermacoce hyssopifolia Willd. ex Roem. & Schult. ^Diodia hyssopifolia
Spermacoce latifolia Aubl. =Borreria latifolia
Spermacoce linearis Willd. ex Roem. & Schult. =Diodia hyssopifolia
Spermacoce longifolia Aubl. =Mitracarpus hirtus
Spermacoce longifolia Aubl. ex Miq. =Diodia ocymifolia
Spermacoce mauritiana Gideon =Borreria repens
Spermacoce ocymifolia Willd. ex Roem. & Schult. =Diodia ocymifolia
Spermacoce ocymoides Burm. f. [not in flora area, neotropical specimens = Borreria ovalifolia or prostrata] =Borreria
prostrata
Spermacoce orinocensis Willd. ex Roem. & Schult. =Borreria capitata var. tenella
Spermacoce prostrata Aubl. =Borreria prostrata
Spermacoce radula Willd. & Hoffmanns, ex Roem. & Schult. =Diodia radula
Spermacoce remota Lam. =Borreria remota
Spermacoce rigida Kunth [nom. illeg.] =Diodia apiculata
Spermacoce rigida Willd. ex Roem. & Schult. [nom. illeg.] =Diodia apiculata
Spermacoce spinosa Sw. [nom. illeg.] =Borreria densiflora
Spermacoce tenella Kunth =Borreria capitata var. tenella
Spermacoce verticillata L. =Borreria verticillata
Spermacoce villasa Sw. =Mitracarpus hirtus
Sphinctanthus rupestris Benth. =Sphinctanthus striiflorus
Sprucea rubescens Benth. =Simira rubescens
Stachyarrhena longifolia Hook. f. =Stachyarrhena spicata
Stachyarrhena spicata var. multinervis K. Schum. =Stachyarrhena spicata
Stannia grandiflora H. Karst. =Posoqueria panamensis
Stannia panamensis Walp. & Duchass. =Posoqueria panamensis
Strempeliafimbriata (Benth.) Bremek. =Rudgea cornifolia
Strempelia guyanensis A. Rich. =Rudgea guyanensis
Synisoon schomburgkianum Baill. =Retiniphyllum laxiflorum var. laxiflorum
Tapogomea alba Aubl. =Psychotria ulviformis
Tapogomea carapichea Poir. =Psychotria ligularis var. carapichea
Tapogomea glabra Aubl. ^Psychotria medusula
Tapogomea purpurea Aubl. =Psychotria variegata
Tapogomea tomentosa Aub\. =Psychotria poeppigiana
Tapogomea violacea Aubl. =Psychotria apoda
Tetramerium sessifolium Kunth =Faramea sessilifolia
Checklist of the Plants of the Guiana Shield 505
Thieleodoxa lanceolata Cham. =Alibertia edulis var. edulis
Thieleodoxa uitidula Bremek. =Alibertia edulis var. edulis
Tocoyena guianensis var. communis Steyerm. =Tocoyena guianensis
Tocoyena guianensis var. glabriuscula Steyerm. =Tocoyena guianensis
Tontanea guianensis Aubl. =Coccocypselum guianense
Uragoga alba (Ruiz & Pav.) Kuntze =Psychotria alba
Uragoga alba (Aubl.) Kuntze =Psychotria ulviformis
Uragoga glabra (Aubl.) Kuntze =Psychotria medusula
Uragoga hirta (Miq.) Pulle =Psychotria bremekampiana
Uragoga kappleri (Miq.) Pulle =Psychotria kappleri
Uragoga oblonga (DC.) Kuntze =Psychotria oblonga
Uragoga purpurea (Aubl.) Kuntze =Psychotria variegata
Uragoga tomentosa (Aubl.) K. Schum. =Psychotria poeppigiana
Uragoga tomentosa (Aubl.) Kuntze =Psychotria poeppigiana
Uragoga uliginosa (Sw.) Kuntze =Notopleura uliginosa
Uragoga violacea (Aubl.) Pulle =Psychotria apoda
Valantia hypocarpia L. =Galium hypocarpium
Virecta biflora L. f. =Sipanea biflora
Warszewiczia obovata Ducke =Calycophyllum obovatum
Warszewiczia schomburgkiana Klotzsch =Warszewiczia coccinea
Yutajea liesneri Steyerm. =Isertia parviflora
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Persson, C. 1996. Phylogeny of the Gardenieae (Rubiaceae). Bot. J. Linn. Soc. 121: 91-109.
Persson, C. 2000a. Phylogeny of the neotropical Alibertia group (Rubiaceae), with emphasis on the genus Alibertia, inferred
from ITS and 5S ribosoma! DNA sequences. Amer. J. Bot. 87: 1018-1028.
Persson, C. 2000b. Phylogeny of the Gardenieae based on chloroplast DNA sequences from the rpsl6 intron and tmL(UAA)-
F(GAA) intergenic spacer. Nordic J. Bot. 20: 257-269.
Persson, C. 2003. Agouticarpa, a new neotropical genus of Tribe Gardenieae (Rubiaceae). Brittonia 55: 176-201.
Piesschaert, F. 2001 . Carpology and Morphology ofthe Psychotrieae (Rubiaceae-Rubioideae). Dissertation. Katholieke Universiteit
Leuven, Belgium.
Piesschaert, F., S. Jansen, I. James, E. Robbrecht, and E. Smets. 2001. Morphology, anatomy, and taxonomic position of
Pagameopsis (Rubiaceae-Rubioideae). Brittonia 33: 490-504.
Puff, C, L. Andersson, U. Rohrhofer, and A. Igersheim. 1993. The tribe Schradereae. Bot. Jahrb. Syst. 114: 449-479.
Rendle, A.B. 1936. Spermacoce remota Lam. J. Bot. 74: 10-12.
Robbrecht, E. 1993. Genera Rubiacearum. Opera Bot. Belg. 6: 173-196.
Rogers, G.K. 1984. Gleasonia, Henriquezia, and Platycarpum (Rubiaceae). Fl. Neotrop. Monogr. 39: 1-135.
Standley, P C. 1931. Studies ofAmerican plants-V. Publ. Field Columbian Mus., Bot. Ser. 8: 295-398.
Steyermark, J.A. 1963. Rubiaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 5. Mem. New York Bot.
Gard. 10(5): 1-278.
Steyermark, J.A. 1965. Rubiaceae. In B. Maguire & collaborators. Botany ofthe Guayana Highland part 6. Mem. New York Bot.
Gard. 12(3): 1-285.
Steyermark, J.A. 1967. Rubiaceae. In B. Maguire & collaborators, Botany ofthe Guayana Highland part 7. Mem. New York Bot.
Gard. 17(1): 1-439.
Steyermark, J.A. 1972. Rubiaceae. In B. Maguire & collaborators. Botany ofthe Guayana Highland part 9. Mem. New York Bot.
Gard. 23: 1-832.
Steyermark, J.A. 1974. Rubiaceae. In T. Lasser, Flora de Venezuela 9: 1-2070. Caracas: Instituto Botanico.
Steyermark, J.A. 1981. New species of Rubiaceae from French Guiana, Brazil, and Colombia. Brittonia 33(3): 385-400.
Steyermark, J.A. 1984. New Rubiaceae from French Guiana. Brittonia 36(2): 154-160.
Steyermark, J.A. 1984. Flora ofthe Venezuelan Guayana-I. Ann. Missouri Bot. Gard. 71: 297-340.
Steyermark, J.A. 1987. Flora ofthe Venezuelan Guayana-II. Ann. Missouri Bot. Gard. 74: 85-116.
Steyermark, J.A. 1988. Notes on Oldenlandia filicaulis and Oldenlandia tenuis (Rubiaceae). Ann. Missouri Bot. Gard. 75: 736-
738.
Taylor, CM. 1992. Revision of Cosmibuena (Rubiaceae). Ann. Missouri Bot. Gard. 79: 886-900.
Taylor, CM. 1993. Revision of Palicourea (Rubiaceae: Psychotrieae) in the West Indies. Moscosoa 7: 201-241.
Taylor, CM. 1994. Revision of Hillia (Rubiaceae). Ann. Missouri Bot. Gard. 81(4): 571-609.
Taylor, CM. 1996. Overview ofthe Psychotrieae (Rubiaceae) in the Neotropics. Opera Bot. Belg. 7: 261-270.
Taylor, CM. 1997. Conspectus ofthe genus Palicourea (Rubiaceae-Psychotrieae) with the description of some new species from
Ecuador and Colombia. Ann. Missouri Bot. Gard. 84: 224-262.
Taylor, CM. 1999. 162(20). Rubiaceae-Coussareae. In: G Hading and L. Andersson, eds., Flora of Ecuador 62: 245-320. Council
Checklist of the Plants of the Guiana Shield 507
for Nordic Publications in Botany, Copenhagen.
Taylor, CM. 2001a. Rubiaceae. In: W. D. Stevens, C. Ulloa Ulloa, A. Pool, and O. M. Montiel, eds., Flora de Nicaragua.
Monogr. Syst. Bot. Missouri Bot. Gard. 85. St. Louis: Missouri Botanical Garden Press.
Taylor, CM. 2001b. Overview of the neotropical genus Notopleura (Rubiaceae: Psychotrieae), with the description of some new
species. Ann. Missouri Bot. Gard. 88: 478-515.
Taylor, CM. 2002a. Rubiacearum Americanarum Magna Hama Pars X. New species and a new subspecies ofFaramea (Coussareae)
from Central and South America. Novon 12: 563-570.
Taylor, CM. 2002b. Rubiacearum Americanarum Magna Hama Pars XI. A new species ofAlseis (Calycophylleae) from Central
America and notes on the morphology of this neotropical genus. Novon 12: 571-574.
Taylor, CM. 2002c. Rubiacearum Americanarum Magna Hama Pars VI: New species of and morphological notes on Psychotria
subg. Psychotria (Psychotrieae) from Mesoamerica and western South America. Novon 12: 120-132.
Taylor, CM. 2004. The neotropical genus Ronabea (Rubiaceae, Lasiantheae). Syst. Geogr. PI. 74: 35-42.
Taylor, CM., and L. Andersson. 1999. 162(18). Rubiaceae-Psychotrieae (1). In: G. Harling and L. Andersson, eds.. Flora of
Ecuador 62: 134-241. Council for Nordic Publications in Botany, Copenhagen.
Taylor, CM., and J.L. Clark. 2001. Rubiacearum Americanarum Magna Hama V. Amphidasya in Mesoamerica and western
South America. Novon 1 1 : 489-497.
Taylor, CM., and D.H. Lorence. 1993. On the status of Randia armata (Sw.) DC. (Rubiaceae: Gardenieae). Taxon 42: 865-867.
Taylor, CM., and D.H. Lorence. 2001. Rubiacearum Americanarum Magna Hama Pars II. New and newly circumscribed taxa of
Guettarda (Guettardeae). Novon 11: 127-134.
Taylor, CM., J.A. Steyermark, P.G. Delprete, A. Vincentini, R. Cortes, D. Zappi, C Persson, C Bestetti Costa, & E. Araujo da
Anuncia9ao. 2004. Rubiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan Guayana
8: 497-847.
Verdcourt, B. 1976. Rubiaceae (Part 1). In: R. M. Polhill, ed., Flora of Tropical East Africa. 414pp. Crown Agents for Overseas
Governments and Administrations, London, United Kingdom.
Verdcourt, B. 1983. Notes on Mascarene Rubiaceae. Kew Bull. 37: 521-571.
Zappi, D. C, J. Semir, andN. I. Pierozzi. 1995. Genipa infundibuliformis sp. nov. and notes on Genipa americana (Rubiaceae).
Kew Bull. 50:761-771.
RUTACEAE
J.A. Kallunki
Adiscanthus fusciflonis Ducke AM
Amyris guianensis Aubl. FG
Angostura thfoliata (Willd.) T.S. Elias AM BO DA
Apocaidon carnosum R.S. Cowan AM
Citrus spp. [species and cultivars cultivated and perhaps escaping]
Conchocarpus acuminatus (Pilg.) Kallunki & Pirani
Conchocarpusfanshawei (Sandwith) Kallunki & Pirani
Conchocarpus guyanensis (Pulle) Kallunki & Pirani
Conchocarpus heterophyllus (A. St.-Hil.) Kallunki & Pirani
Conchocarpus longifolius (A. St.-Hil.) Kallunki & Pirani
Conchocarpus ramiflorus (Spruce ex Engl.) Kallunki & Pirani
Conchocarpus toxicarius (Engl.) Kallunki & Pirani
Conchocarpus ucayalinus (Huber) Kallunki & Pirani
Decagonocarpus cormitus R.S. Cowan
Decagonocarpus oppositifolius Spruce ex Engl.
Ertela trifolia (L.) Kuntze [=Monnieria trifolia L.]
Erythrochiton brasiliensis Nees & Mart.
Esenbeckia cowanii Kaastra
Esenheckia grandiflora Mart.
Esenbeckia grandiflora var. grandiflora
Esenbeckia pilocarpoides Kunth
Esenbeckia pilocarpoides subsp. pilocarpoides
Galipea davisii Sandwith
Galipea trifoliata Aubl.
Hortia coccinea Spruce ex Engl.
Hortia neblinensis Maguire & B. Boom
Hortia regia Sandwith BO GU
Hortia vandelliana Groppo AM
Metrodoreaflavida K. Krause
Murraya paniculata (L.) Jack [cultivated; naturalized?]
Neoraputia paraensis (Ducke) Emmerich
Pilocarpus demerarae Sandwith
Pilocarpus microphyllus Stapf ex Wardleworth
FG
GU
su FG
BO su
BO DA su FG
AM
GU FG
AM BO GU FG
AM
AM
AM BO GU su FG
AM BO GU su FG
FG
BO DA GU su FG
BO DA GU su FG
AM BO GU su FG
AM BO GU su FG
AM BO GU
su
FG
FG
AM
AM
SU
BO
BO DA
GU
GU
su
SU
FG
FG
508 Checklist of the Plants of the Guiana Shield
Pilocarpus racemosus Vahl
Pilocarpus racemosus subsp. racemosus
Raputia aromatica Aubl.
Raputia brevipedunculata Kallunki
Raputia maroana (R.S. Cowan) Kallunki
Raputia neblinensis (R.S. Cowan) Kallunki
Raputia szczerbanii (Steyerm.) Kallunki
Rauia resinosa Nees & Mart.
Rauia spicata Haye
Rauia subtruncata Steyerm.
Raveniopsis abyssicola R.S. Cowan
Raveniopsis breweri Steyerm.
Raveniopsis capitata R.S. Cowan
Raveniopsis cowaniana Steyerm. & Luteyn
Raveniopsis fraterna R.S. Cowan
Raveniopsis jauaensis Steyerm.
Raveniopsis linearis (Gleason) R.S. Cowan
Raveniopsis nubicola R.S. Cowan
Raveniopsis paruana (R.S. Cowan) R.S. Cowan
Raveniopsis peduncularis Pittier & Lasser
Raveniopsis ruellioides (Oliv.) R.S. Cowan
Raveniopsis sericea R.S. Cowan
Raveniopsis stelligera (R.S. Cowan) R.S. Cowan
Raveniopsis steyermarkii R.S. Cowan
Raveniopsis tomentosa Gleason
Raveniopsis trifoliolata R.S. Cowan
Rutaneblina pusilla Steyerm. & Luteyn
Spathelia ulei (Engl, ex Harms) R.S. Cowan & Brizicky
Spiranthera guianensis Sandwith
Spiranthera parviflora Sandwith
Ticoreafoetida Aubl.
Ticorea longiflora DC.
Ticorea pedicellata DC.
Triphasia trifolia (Burm. f.) R Wilson [cultivated and apparently escaped]
Zanthoxylum amapaense (Albuq.) RG Waterman
Zanthoxylum apiculatum (Sandwith) RG. Waterman
Zanthoxylum caribaeum Lam.
Zanthoxylum compactum (Huber ex Albuq.) RG Waterman
Zanthoxylum ekmanii (Urb.) Alain
Zanthoxylumfagara (L.) Sarg.
Zanthoxylum /Iavum Vahl
Zanthoxylum hermaphroditum Willd. [type "Guiana"]
Zanthoxylum huberi RG Waterman
Zanthoxylumjuniperinum Poepp.
Zanthoxylum martinicense (Lam.) DC.
Zanthoxylum nigrum Mart.
Zanthoxylum pentandrum (Aubl.) R.A. Howard
Zanthoxylum rhoifolium Lam.
Zanthoxylum sp. A
Zanthoxylum sp. B AM
Zanthoxylum syncarpum Tul. BO
Synonyms and Misidentifications
Almeidea guyanensis Pulle =Conchocarpus guyanensis
Amyris decandra (Aubl.) Willd. (Burseraceae) =Protium decandrum
Amyris enneandra (Aubl.) Willd. (Burseraceae) =Protium decandrum
Amyris guyanensis (Aubl.) Willd. (Burseraceae) =Protium guianense
Angostura acuminata (Pilg.) Albuq. =Conchocarpus acuminatus
Angostura cuspare Roem. & Schult. [nom. illeg.] =Angostura trifoliata
Angosturafanshawei (Sandwith) Albuq. ^Conchocarpus fanshawei
Angosturafontanesiana (A. St.-Hil.) Albuq. [not in flora area]
Angostura pentagyna (A. St.-Hil.) Albuq. =Conchocarpus longifolius
Angostura ramiflora (Spruce ex Engl.) Albuq. =Conchocarpus ramiflorus
Angostura toxicaria (Spruce ex Engl.) Albuq. =Conchocarpus toxicarius
BO FG
AM
BO FG
FG
FG
AM
BO
GU SU
BO SU
AA/f
BO GU
/\M
BO
AM BO
AM
BO
BO
AM
AM
AM
BO
AM BO
BO
GU
GU
AM
BO
BO
AM
AM
AM BO
BO
GU
GU
AM BO GU
GU SU
SU
FG
FG
GU SU FG
GU FG
BO DA FG
BO GU
BO
DA
GU
GU
BO DA GU SU FG
BO FG?
FG?
BO
DA SU
BO DA
SU?
BO DA GU SU FG
AM BO
BO
GU SU FG
Checklist of the Plants of the Guiana Shield 509
Angostura ucayalina (Huber) Albuq. =Conchocarpus ucayalinus
Angostura iilei (K. Krause) Gereau =Conchocarpus ucayalinus
Bonplandia angostura Spreng. =Angostura trifoliata
Bonplandia angostura Rich, ex B.D. Jacks. =Angostura trifoliata
Bonplandia trifoliata Willd. =Angostura trifoliata
Chalcas exotica (L.) Millsp. =Murraya paniculata
Chalcas paniculata L. =Murraya paniculata
Conchocarpus pentagynus (A. St.-Hil.) Kallunki & Pirani [nomen nudum] =Conchocarpus longifolius
Cusparia angostura (Rich, ex B.D. Jacks.) A. Lyons =Angostura trifoliata
Cuspariafanshawei Sandwith =Conchocarpusfanshawei
Cuspariafebrifuga Humb. =Angostura trifoliata
Cusparia officinalis (J. Hancock) Engl. =Angostura trifoliata
Cusparia pentagyna (A. St.-Hil.) Engl. =Conchocarpus longifolius
Cusparia ramiflora Spruce ex Engl. =Conchocarpus ramiflorus
Cusparia toxicaria Spruce ex Engl. =Conchocarpus toxicarius
Cusparia trifoliata (Willd.) Engl. =Angostura trifoliata
Cusparia ucayalina Huber =Conchocarpus ucayalinus
Decagonocarpus cornutus var. congestus R.S. Cowan =Decagonocarpus cornutus
Diommafruticosa Steyerm. =Spathelia ulei
Diomma ulei Engl, ex Harms =Spathelia ulei
Esenbeckia obovalifolia Pittier =Esenbeckia grandiflora var. grandiflora
Esenbeckia venezuelensis Engl. =Esenbeckia pilocarpoides subsp. pilocarpoides
Fagara amapaensis Albuq. =Zanthoxylum amapaense
Fagara apiculata Sandwith =Zanthoxylum apiculatum
Fagara caribaea (Lam.) Krug & Urb. =Zanthoxylum caribaeum
Fagara compacta Huber ex Albuq. =Zanthoxylum compactum
Fagara ekmanii Urb. =Zanthoxylum ekmanii
Fagaraflava Krug & Urb. =Zanthoxylumflavum
Fagara guianensis Lam. =Zanthoxylum hermaphroditum
Fagara martinicensis Lam. =Zanthoxylum martinicense
Fagara ocumarensis Pittier =Zanthoxylum juniperinum
Fagara pentandra Aubl. =Zanthoxylum pentandrum
Fagara perrottetii DC. =Zanthoxylum rhoifolium
Fagara pterota L. =Zanthoxylumfagara
Fagara rhoifolia (Lam.) Engl. =Zanthoxylum rhoifolium
Fagara syncarpa (Tul.) Engl. =Zanthoxylum syncarpum
Galipea aromatica (Aubl.) Spreng. =Raputia aromatica
Galipea cusparia A. St.-Hil. ex DC. [nom. illeg.] ^Angostura trifoliata
Galipeafebrifuga (Humb.) Baill. =Angostura trifoliata
Galipeafissa Miq. =Galipea trifoliata
Galipea heterophylla A. St.-Hil. =Conchocarpus heterophyllus
Galipea longifolia A. St.-Hil. =Conchocarpus longifolius
Galipea officinalis J. Hancock =Angostura trifoliata
Galipea pentagyna A. St.-Hil. =Conchocarpus longifolius
Galipea stelligera R.S. Cowan =Raveniopsis stelligera
Lubaria szczerbanii Steyerm. =Raputia szczerbanii
Monnieria trifolia L. =Ertela trifolia
Murraya exotica L. =Murraya paniculata
Myllanthus maroana R.S. Cowan =Raputia maroana
Myllanthus neblinensis R.S. Cowan =Raputia neblinensis
Neoraputia cowanii Emmerich =Neoraputia paraensis
Pilocarpus latifolius A. St.-Hil. =Pilocarpus racemosus subsp. racemosus
Raputia paraensis Ducke =Neoraputia paraensis
Rauia prancei W.A. Rodrigues & M.F. Silva [not in flora area]
Rauia racemosa Nees & Mart, [not in flora area]
Rauia ulei K. Krause =Conchocarpus ucayalinus
Ravenia linearis Gleason =Raveniopsis linearis
Ravenia paruana R.S. Cowan =Raveniopsis paruana
Ravenia ruellioides Oliv. =Raveniopsis ruellioides
Ravenia ruellioides var. ptariana Steyerm. =Raveniopsis ruellioides
Ravenia tatei Gleason =Raveniopsis ruellioides
Raveniopsis liesneri Steyerm. =Raveniopsis stelligera
Raveniopsis ruellioides var. angusta R.S. Cowan =Raveniopsis ruellioides
Raveniopsis stelligera var. plicata R.S. Cowan =Raveniopsis stelligera
510 Checklist of the Plants of the Guiana Shield
Schinusfagara L. =Zanthoxylumfagara
Sciuhs aromatica J.F. Gmel. =Raputia aromatica
Sciuris officinalis (J. Hancock) Oken =Angostura trifoliata
Spathelia chimantaensis R.S. Cowan & Brizicky =Spathelia ulei
Spatheliafruticosa (Steyerm.) R.S. Cowan & Brizicky =Spathelia ulei
Spatheliajauaensis R.S. Cowan =Spathelia ulei
Spathelia neblinaensis R.S. Cowan & Brizicky =Spathelia ulei
Zanthoxylum belizense Lundell [specimens re-determined as Zanthoxylum ekmanii]
Zanthoxylum lucidum A. St.-Hil. [specimens re-determined as Zanthoxylum pentandrum]
Zanthoxylum microcarpum Griseb. [specimens re-determined as Zanthoxylum rhoifolium]
Zanthoxylum ocumarense (Pittier) Steyerm. =Zanthoxylum juniperinum
Zanthoxylum perrottetii DC. =Zanthoxylum rhoifolium
Zanthoxylum pterota (L.) Kunth =Zanthoxylum fagara
Zanthoxylum ruizianum (Klotzsch ex Engl.) J.F. Macbr. =Zanthoxylum rhoifolium
References
Albuquerque, B.W.R de. 1977 ("1976"). Revisao taxonomica das Rutaceae do Estado do Amazonas. Acta Amazonica suppl.
6(3): 1-67.
Albuquerque, B.W.R de. 1981. Novas combina9oes em Angosture Roemer & Schultes (Rutaceae). Acta Amazonica 11 (4): 849-
852.
Cowan, R.S. 1960. Rutaceae. In B. Maguire & J.J. Wurdack, Botany of the Guayana Highland part IV. Mem. New York Bot.
Gard. 10(2): 24-37.
Cowan, R.S. 1967. Rutaceae of the Guayana Highland. Mem. New York Bot. Gard. 14(3): 1-14.
Elias, T.S. 1970. The correct name for the genus Cusparia (Rutaceae). Taxon 19(4): 573-575.
Emmerich, M. 1978. Contribu9ao ao conhecimento da tribo Cusparineae (sic) (Rutaceae). Rodriguesia 45: 223-307.
Kaastra, R.C. 1982. Pilocarpinae (Rutaceae). Flora Neotropica Monogr. 33: 1-197.
Kallunki, J.A. 1990. An emended description of and new combinations in Raputia Aublet. Brittonia 42: 175-177.
Kallunki, J.A. 1992. A revision of Erythrochiton s.l. Brittonia 44(2): 107-139.
Kallunki, J.A. 1994. Revision of Raputia Aubl. (Cuspariinae, Rutaceae). Brittonia 46(4): 279-295.
Kallunki, J.A. 2002. Rutaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 649-
656.
Kallunki, J.A. 2005. Rutaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
9: 1-39.
Porter, D.M. & T.S. Elias. 1979. Rutaceae. In R.E. Woodson Jr., R.W. Schery, & collaborators. Flora of Panama. Ann. Missouri
Bot. Gard. 66: 123-164.
Waterman, P.G. 1975. New combinations in Zanthoxylum L. (1753). Taxon 24: 361-366.
Sabiaceae
G.A. Aymard C, N.L. Cuello A.
Meliosma gentryi Aymard & Cuello
Meliosma herbertii Rolfe
Meliosma herbertii var. herbertii
Meliosma herbertii var. tepuiensis (Steyerm. & Maguire) Steyerm. & Gentry
Meliosma aff meridensis Lasser
Meliosma vasquezii Gentry AM
Ophiocaryon chironectes Bameby FG?
Ophiocaryon duckei Bameby AM
Ophiocaryon heterophyllum (Benth.) Urb. [records from Guyana probably GU?
represent Ophiocaryon maguirei]
Ophiocaryon maguirei Bameby GU
Ophiocaryon manausense (W.A. Rodrigues) Bameby AM
Ophiocaryon paradoxum R.H. Schomh. GU FG?
Ophiocaryon sp. A [expected in AM]
Synonyms and Misidentifications
Meliosma nubicola Steyerm. & Lasser =Meliosma meridensis
Meliosma tepuiensis Steyerm. & Maguire =Meliosma herbertii var. tepuiensis
Meliosma uberrima Idrobo & Cuatrec. =Meliosma meridensis
Phoxanthus manausensis W.A. Rodrigues =Ophiocaryon manausense
References
Aymard C, G.A. and N.L. Cuello A. 2005. Sabiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
BO DA GU
BO DA GU
BO
BO
Checklist of the Plants of the Guiana Shield 511
Venezuelan Guayana 9: 39-43.
Bameby, R.C. 1972. Meliosmaceae - Ophiocaryon. In B. Maguire & collaborators, Botany of the Guayana Highland part 9.
Mem. New York Bot. Gard. 23: 1-832.
Steyermark, J.A. & A.H. Gentry. 1992. Sabiaceae. In G. Morillo, Flora de Venezuela 5(1): 203-220.
Santalaceae
J.S. Miller
Thesium tepuiense Steyerm. BO GU
Synonyms and Misidentifications
Anstroamericum tepuiense (Steyerm.) Hendrych =Thesium tepuiense
References
Miller, J.S. 2005. Santalaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 9:
44-45.
Steyermark, J. 1951. Fieldiana, Botany 28. 1: 225
Sapindaceae
R Acevedo R., H.T. Beck
Allophylus amazonicus (Mart.) Radlk.
Allophylus edulis (A. St.-Hil.) Hieron. ex Niederl.
Allophylus latifolius Huber
Allophylus leucoclados Radlk.
Allophylus parimensis Steyerm.
Allophylus punctatus (Poepp. & Endl.) Radlk.
Allophylus racemosus Sw.
Allophylus rohustus Radlk.
Cardiospermum corindum L.
Cardiospermum grandiflorum Sw.
Cardiospermum halicacabum L.
Cardiospermum halicacabum var. halicacabum
Cardiospermum halicacabum var. microcarpum (Kunth) Blume
Cupania americana L. [s.l.]
Cupania cinerea Poepp. & Endl.
Cupania diphylla Vahl
Cupania hirsuta Radlk.
Cupania hispida Radlk.
Cupania kukenanica Steyerm.
Cupania rubiginosa (Poir.) Radlk.
Cupania scrobiculata Rich.
Cupania scrobiculata var. guianensis (Miq.) Uittien
Cupania scrobiculata var. scrobiculata
Dilodendron costaricense (Radlk.) Gentry & Steyerm.
Dilodendron elegans (Radlk.) Gentry & Steyerm.
Dodonaea viscosa Jacq.
Houssayanthus macrolophus (Radlk.) Hunz.
Matayba adenanthera Radlk.
Matayba arborescens (Aubl.) Radlk.
Matayba atropurpurea Radlk.
Matayba camptoneura Radlk.
Matayba elegans Radlk.
Matayba guianensis Aubl.
Matayba inelegans Spruce ex Radlk.
Matayba kavanayena (Steyerm.) Steyerm.
Matayba laevigata (Miq.) Radlk.
Matayba longipes Radlk.
Matayba macrostylis Radlk.
Matayba opaca Radlk.
Matayba peruviana Radlk.
Matayba ptariana Steyerm.
Matayba ptariana subsp. gimiquinimae Steyerm.
Matayba ptariana subsp. ptariana
AM
GU FG
FG
FG
AM
BO
BO
BO
BO
GU
GU FG
AM BO
BO
DA GU su FG
AM DA GU su FG
BO su FG
AM BO
su FG
BO DA GU su FG
AM
BO GU
AM BO GU
AM BO GU su FG
GU su FG
GU FG
BO
BO
BO
DA
GU su FG
AM BO
AM BO DA GU su FG
AM
AM BO GU su
AM BO GU
AM BO GU FG
AM BO DA GU su FG
AM BO
GU su FG
AM
AM GU
AM BO GU su FG
AM BO GU FG
AM BO
BO
GU
AM BO
512 Checklist of the Plants of the Guiana Shield
Matayba purgans (Poepp.) Radlk. GU FG
Matayba robusta Radlk. AM
Matayba spruceana (Benth.) Radlk. BO
Matayba stenodictya Radlk.
Matayba yiitajemis Steyerm. AM
Melicoccus bijugatus Jacq. [cultivated or escaped from cultivation] BO
Melicocciis oliviformis Kunth AM
Melicoccus oliviformis subsp. intermedius (Radlk.) Acev.-Rodr. AM
Melicoccus pedicellaris (Sagot ex Radlk.) Acev.-Rodr.
Nephelium lappaceum L. [cultivated]
Paullinia acuminata Uittien
Paullinia alata (Ruiz & Pav.) G. Don
Paullinia alsmithii Macbr. AM BO
Paullinia anisoptera Turcz. BO DA
Paullinia anodonta Radlk.
Paullinia anomophylla Radlk.
Paullinia arenicola Gleason
Paullinia bernhardi Uittien
Paullinia bracteosa Radlk. AM
Paullinia caloptera Radlk. AM BO
Paullinia cambessedesii Triana & Planch.
Paullinia capreolata (Aubl.) Radlk. AM BO DA
Paullinia clathrata Radlk.
Paullinia conduplicata Radlk.
Paullinia cupana Kunth AM
Paullinia cururu L. BO
Paullinia curvicuspis Radlk.
Paullinia dasygonia Radlk.
Paullinia demerarae Sandwith
Paullinia fibulata Rich, ex Juss.
Paulliniafuscescens Kunth BO DA
Paullinia grandifolia Benth. ex Radlk.
Paullinia hitchcockii Gleason
Paullinia imberbis Radlk. AM
Paullinia ingifolia Rich, ex Juss. AM BO
Paullinia isoptera Radlk.
Paullinia latifolia Benth. ex Radlk.
Paullinia leiocarpa Griseb. AM BO
Paullinia lingulata Acev.-Rodr.
Paullinia livescens Radlk.
Paullinia micrantha Cambess. BO DA
Paullinia novemalata Uittien
Paullinia nuriensis Steyerm. BO
Paullinia pachycarpa Benth. GU FG
Paullinia paullinioides Spruce ex Radlk. AM
Paullinia perlata Radlk.
Paullinia pinnataL. BO DA
Paullinia plagioptera Radlk.
Paullinia rubiginosa Cambess.
Paullinia rufescens Rich, ex Juss. AM BO DA
Paullinia rugosa Benth. ex Radlk.
Paullinia serjaniifolia Triana & Planch.
Paullinia sphaerocarpa Rich, ex Juss.
Paullinia spicata Benth.
Paullinia stellata Radlk.
Paullinia stenopetala Sagot SU
Paullinia subnuda Radlk. GU
Paullinia tetragona Aubl. FG
Paullinia triantennata F. Silveira
Paullinia tricornis Radlk.
Paullinia trilatera Radlk.
Paullinia venosa Radlk.
Paullinia verrucosa Radlk. BO
Paullinia vespertilio Sw.
FG
GU SU FG
GU
GU
SU FG
FG
SU FG
SU FG
GU FG
SU FG
FG
GU
SU
GU
GU FG
FG
GU SU FG
FG
GU FG
FG
GU SU FG
GU
FG
GU SU FG
GU SU
GU
FG
GU SU FG
GU
GU SU FG?
GU
FG
GU
GU SU
SU
FG
AM BO
AM
AM
GU
GU SU FG
GU SU FG
FG
GU FG
GU FG
GU? SU FG
SU FG
GU FG
GU SU
GU SU FG
SU FG
SU FG
GU SU FG
GU?
FG
GU FG
GU su FG
su FG
GU? su FG?
GU? su FG?
FG
GU
GU FG
GU FG
Checklist of the Plants of the Guiana Shield 513
Paullinia xestophylla Radlk. GU
Pentascyphus thyrsiflorus Radlk.
Porocystis toulicioides Radlk.
Pseudimafrutescens (Aubl.) Radlk.
Pseudima pallidum Radlk.
Sapindus saponaria L. AM BO DA
Sapindus saponaria var. inaequalis (DC.) Radlk.
Scyphonichium multiflorum (Mart.) Radlk.
Serjania adusta Radlk. BO
Serjania caracasana (Jacq.) Willd. BO
Serjania chartacea Radlk.
Serjania clematidea Triana & Planch. AM BO
Serjania communis Cambess. AM
Serjania communis var. communis AM
Serjania didymadenia Radlk. FG
Serjania exarata Radlk. GU
Serjania grandifolia Sagot ex Radlk. BO GU SU FG
Serjania membranacea Splitg. AM BO DA GU SU FG
Serjania oblongifolia Radlk. GU SU FG
Serjania paucidentata DC. BO DA GU SU FG
Serjania pedicellaris Radlk. SU FG
Serjania pyramidata Radlk. BO DA GU FG
Serjania rhombea Radlk. AM
Serjania setulosa Radlk. FG
Talisia acutifolia Radlk. AM
Talisia carinata Radlk. GU SU FG
Talisia caudata Steyerm. AM
Talisia chartacea Acev.-Rodr. AM BO
Talisia clathrata Radlk. FG
Talisia clathrata subsp. canescens Acev.-Rodr. FG
Talisia cupularis Radlk. FG
Talisia eximia K.U. Kramer SU
Talisiafirma Radlk. AM
Talisiafurfuracea Sandwith GU SU FG
Talisia guianensis Aubl. GU FG
Talisia hemidasyaRa&lk. BO DA GU SU FG
Talisia hexaphylla Yahl BO DA GU FG
Talisia laevigata Acev.-Rodr. BO
Talisia longifolia{Benth.) Radlk. GU SU FG
Talisia macrophylla (Mart.) Radlk. AM BO GU SU FG
Talisia megaphylla Sagot ex Radlk. GU SU FG
Talisia microphylla Uittien GU SU FG
Talisia mollis Kunth ex Cambess. GU SU FG
Talisia nervosa Radlk. AM BO
Talisia obovata A.C. Sm. AM
Talisia pachycarpa Radlk. FG
Talisia pilosula Sagot ex Radlk. FG
Talisia praealta {SagoX) Radlk. SU FG
Talisia retusaR.?>.Co^an AM BO GU SU
Talisia simaboides K.U. Kramer SU FG
Talisia squarrosa Radlk. GU I'G
Talisia sylvatica {huhl.) Radlk. GU SU FG
Thinouia myriantha Triana & Planch. DA GU SU IG
Toulicia elliptica Radlk. GU? SU FG?
Toulicia guianensis Aubl. AM BO DA GU SU FG
Toulicia patentinervis Radlk. GU SU
Toulicia petiolulata Radlk. AM
Toulicia pulvinata Radlk. AM BO GU SU FG
Urvillea ulmacea Y.\xn\h AM BO GU SU? FG
Vouarana anomala (Steyerm.) Acev.-Rodr. BO
Vouarana guianensis Aubl. GU SU FG
Synonyms and Misidentifications
Allophylus occidentalis (Sw.) Radlk. =Allophylus racemosus
514 Checklist of the Plants of the Guiana Shield
Cardiospermum macrolophum Radlk. =Houssayanthus macrolophus
Cardiospermum microcarpum Kunth =Cardiospermum halicacabum var. microcarpum
Cupania kavanayena Steyerm. =Matayba kavanayena
Cupania lachnocarpa Radlk. [not in flora area]
Cupania lanuginosa Sagot ex Radlk. =Cupania hirsuta
Cupania latifolia Kunth =Cupama amehcana s.l.
Cupania macrophylla Mart. =Talisia macrophylla
Cupania papillosa Radlk. =Cupania latifolia
Cupania roraimae Steyerm. =Cupania kukenanica
Cupania scrobiculata f. reticulata (Cambess.) Radlk. =Cupania scrobiculata var. scrobiculata
Cupania semidentata Radlk. ^Cupania latifolia
Cupania spniceana Benth. =Matayba spruceana
Cupania tepuiensis Steyerm. & Maguire =Cupania rubiginosa
Dilodendron bipinnatum var. elegans Radlk. =Dilodendron elegans
Dipterodendwn costahcense Radlk. =Dilodendwn costaricense
Dipterodendron elegans (Radlk.) Radlk. =Dilodendron elegans
Dipterodendwn venezuelense Steyerm. =Dilodendron elegans
Enourea capreolata Aubl. =Paullinia capreolata
Matayba affinis Steyerm. =Matayba opaca var. opaca
Matayba chimantensis Steyerm. & Maguire =Matayba peruviana
Matayba discolor var. atropurpurea Radlk. =Matayba atropurpurea
Mataybafallax Radlk. =Matayba opaca VRT.fallax
Mataybajauaensis Steyerm. =Matayba peruviana
Matayba longipes subsp. tepuiensis Steyerm. =Matayba longipes
Matayba macrolepis Radlk. =Matayba macrostylis
Matayba miquelii Uittien ^Matayba laevigata
Matayba oligandra Sandwith =Matayba peruviana
Matayba oligandra var. occidentalis Steyerm. =Matayba peruviana
Matayba oligandra \m. ptariana (Steyerm.) Steyerm. =Matayba peruviana
Matayba opaca var. fallax (Radlk.) Uittien =Matayba opaca
Matayba reducta Steyerm. =Matayba camptoneura
Matayba sororopaniana Steyerm. =Matayba inelegans
Matayba tovarensis Radlk. =Matayba longipes
Matayba venezuelana Steyerm. =Matayba longipes
Melicoccus bijuga (Jacq.) L. ^Melicoccus bijugatus
Melicoccus bijugata L. =Melicoccus bijugatus
Paullinia caracasana Jacq. =Serjania caracasana
Paulliniafasciculata Radlk. =Paullinia alata
Paullinia sorbilis Mart. ^Paullinia cupana
Racaria sylvatica Aubl. =Talisia sylvatica
Robinia rubiginosa Poir. =Cupania rubiginosa
Sapindus arborescens Aubl. =Matayba arborescens
Sapindus frutescens Aubl. =Pseudimafrutescens
Schmidelia amazonica Mart. =Allophylus amazonicus
Schmidelia occidentalis Sw. =Allophylus racemosus
Schmidelia punctata Poepp. & Endl. =Allophylus punctatus
Serjania ierensis Britton =Serjania caracasana
Serjania mariquitensis Triana & Planch. =Serjania caracasana
Serjania prancei Acev.-Rodr. [nomen nudum] =Serjania caracasana
Talisia alien ii Croat =Talisia macrophylla
Talisia amaruayana Steyerm. ^Talisia nervosa
Talisia cararensis Cuatrec. =Talisia hexaphylla subsp. hexaphylla
Talisia dwyeri Croat =Talisia nervosa
Talisia elephantipes Sandwith =Talisia macrophylla
Talisia glandulifera Steyerm. =Talisia hemidasya
Talisia grandifolia Cuatrec. =Talisia nervosa
Talisia heterodoxa Steyerm. =Talisia retusa
Talisia medrii Guarim =Talisia acutifolia
Talisia micrantha Radlk. =Talisia sylvatica
Talisia olivaeformis (Kunth) Radlk. =Melicoccus oliviformis
Talisia oliviformis (Kunth) Radlk. =Melicoccus oliviformis
Talisia panamensis Pittier =Talisia hexaphylla subsp. hexaphylla
Talisia pedicellaris Sagot ex Radlk. =Melicoccus pedicellaris
Talisia pentantha Steyerm. =Talisia macrophylla
Checklist of the Plants of the Guiana Shield 515
Talisia piilventlenta Radlk. =Melicoccus pedicellaris
Talisia reticulata Radlk. =Talisia sylvatica
Talisia sancarlosiana Steyerm. =Talisia macrophylla
Talisia tirirensis Steyerm. & Maguire =Talisia nervosa
Tapirocarpus talisia Sagot =Talisia mollis
Toulicia anomala Steyerm. =Voiiarana anomala
Trichilia caudata Killip & Cuatrec. =Matayba macrostylis
Trichilia ptariana Steyerm. =Matayba peruviana
Urvillea macrolopha (Radlk.) Radlk. =Houssayanthus macrolophus
References
Acevedo-R., R 2002. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds. 2002.
Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 656-669.
Acevedo-R., R & H.T. Beck. 2005. Sapindaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 46-89.
Kramer, K.U. 1973 (1972). Studies in the New World Sapindaceae. I. Notes on the genus Talisia Aublet. Acta Bot. Need. 21:
671-678.
Kramer, K.U. 1976. Sapindaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 487-51 1.
Radlkofer, L. 1931-1934. Sapindaceae. In A. Engler, Das Pflanzenreich IV,165 Bd. I: 1-1018, Bd. II: 1019-1539.
Uittien, H. 1937. Sapindaceae. In A. Pulle, Flora of Suriname 2(1): 345-396.
Sapotaceae
K.S. Edwards, T.D. Pennington
Chromolucuma baehniana Monach.
Chromolucuma rubriflora Ducke
Chrysophyllum amazonicum T.D. Penn.
Chrysophyllum argenteum Jacq.
Chrysophyllum argenteum subsp. auratum (Miq.) T.D. Penn.
Chrysophyllum argenteum subsp. nitidum (G. Mey.) T.D. Penn.
Chrysophyllum cainito L. [cultivated, naturalized]
Chrysophyllum cuneifoliiim (Rudge) A. DC.
Chrysophyllum eximium Ducke [expected in EG]
Chrysophyllum lucentifolium Cronquist
Chrysophyllum lucentifolium subsp. pachycarpum Pires & T.D. Penn.
Chiysophyllum manaosense (Aubrev.) T.D. Penn.
Chrysophyllum pomiferum (Eyma) T.D. Penn.
Chrysophyllum prieurii A. DC.
Chrysophyllum sanguinolentum (Pierre) Baehni
Chiysophyllum sanguinolentum subsp. balata (Ducke) T.D. Penn.
Chrysophyllum sanguinolentum subsp. sanguinolentum
Chrysophyllum sanguinolentum subsp. spurium (Ducke) T.D. Penn.
Chrysophyllum sparsiflorum Klotzsch ex Miq.
Chrysophyllum ucuquirana-branca (Aubrev. & Pellegr.) T.D. Penn.
Chrysophyllum venezuelanense (Pierre) T.D. Penn.
Diploon cuspidatum (Hoehne) Cronquist
Ecclinusa atabapoensis (Aubrev.) T.D. Penn.
Ecclinusa bullata T.D. Penn.
Ecclinusa dumetorum (Baehni) T.D. Penn.
Ecclinusa guianensis Eyma
Ecclinusa lanceolata (Mart. & Eichler) Pierre
Ecclinusa orinocoensis Aubrev.
Ecclinusa parviflora T.D. Penn.
Ecclinusa psilophylla Sandwith
Ecclinusa ramiflora Mart, [expected in GU]
Ecclinusa ulei (K. Krause) Gilly ex Cronquist [Ml. Roraima]
Elaeoluma crispa T.D. Penn.
Elaeoluma glabrescens (Mart. & Eichler) Aubrev.
Elaeoluma nuda (Baehni) Aubrev. [expected in GU]
Elaeoluma schomburgkiana (Miq.) Baill.
Manilkara bidentata (A. DC.) A. Chev.
Manilkara bidentata subsp. bidentata
Manilkara bidentata subsp. surinamensis (Miq.) T.D. Penn.
Manilkara bolivarensis T.D. Penn.
GU
AM
BO GU
AM BO DA GU su EG
AM BO DA GU su EG
GU su EG
GU su
su
su
EG
EG
AM BO GU FG
AM BO GU
GU su
EG
AM BO GU su FG
AM GU su FG
AM BO GU su FG
AM BO GU
AM GU su FG
AM
AM
BO
BO
GU SU? FG
FG
DA GU FG
AM
AM
SU FG?
AM BO DA GU SU FG
AM GU FG
AM
AM
GU SU
AM SU FG
AM
AM BO GU
AM BO SU
AM BO GU
AM BO DA GU su EG
BO DA GU su FG
AM BO
BO
DA GU su FG
51 Checklist of the Plants of the Guiana Shield
Manilkara excelsa (Ducke) Standi.
Manilkara hiiberi (Ducke) A. Chev.
Manilkara paraensis (Huber) Standi.
Manilkara pubicarpa Monach.
Manilkara zapota (L.) P. Royen [widely cultivated and escaping]
Micropholis acutangula (Ducke) Eyma
Micropholis casiquiarensis Aubrev.
Micropholis caudata T.D. Penn.
Micropholis cayennensis T.D. Penn.
Micropholis egensis (A. DC.) Pierre
Micropholis gardneriana (A. DC.) Pierre
Micropholis giiyanensis (A. DC.) Pierre
Micropholis giiyanensis subsp. duckeana (Baehni) T.D. Penn.
Micropholis guyanensis subsp. guyanensis
Micropholis humboldtiana (Roem. & Schult.) T.D. Penn.
Micropholis longipedicellata Aubrev.
Micropholis maguirei Aubrev.
Micropholis melinoniana Pierre
Micropholis mensalis (Baehni) Aubrev.
Micropholis obscura T.D. Penn.
Micropholis porphyrocarpa (Baehni) Monach.
Micropholis sanctae-rosae (Baehni) T.D. Penn.
Micropholis spectabilis (Steyerm.) T.D. Penn.
Micropholis splendens Gilly ex Aubrev.
Micropholis submarginalis Pires & T.D. Penn.
Micropholis suborbicularis Aubrev.
Micropholis venamoensis (Steyerm.) T.D. Penn.
Micropholis veniilosa (Mart. & Eichler) Pierre
Mimusops spp. [cultivated or escaped from cultivation]
Podoluma pacimoniensis Aubrev.
Pouteria ambelaniifolia (Sandwith) T.D. Penn.
Pouteria amygdalicarpa (Pittier) T.D. Penn.
Pouteria anomala (Pires) T.D. Penn.
Pouteria arcuata T.D. Penn.
Pouteria atabapoensis (Aubrev.) T.D. Penn.
Pouteria aubrevillei Bemardi
Pouteria bangii (Rusby) T.D. Penn. [expected in GU]
Pouteria benai (Aubrev. & Pellegr.) T.D. Penn.
Pouteria bilocularis (H. Winkl.) Baehni
Pouteria brachyandra (Aubrev. & Pellegr.) T.D. Penn.
Pouteria caimito (Ruiz & Pav.) Radlk.
Pouteria campanulata Baehni [possibly expected in GU]
Pouteria canaimaensis T.D. Penn.
Pouteria cayennensis (A. DC.) Eyma
Pouteria cicatricata T.D. Penn.
Pouteria cladantha Sandwith
Pouteria congestifolia Pilz
Pouteria coriacea (Pierre) Pierre
Pouteria crassiflora Pires & T.D. Penn. [possibly expected in FG]
Pouteria euspidata (A. DC.) Baehni
Pouteria euspidata subsp. euspidata
Pouteria euspidata subsp. dura (Eyma) T.D. Penn.
Pouteria euspidata subsp. robusta (Mart. & Eichler) T.D. Penn.
Pouteria decorticans T.D. Penn.
Pouteria deliciosa T.D. Penn.
Pouteria durlandii (Standi.) Baehni
Pouteria durlandii subsp. durlandii
Pouteria egregia Sandwith
Pouteria elegans (A. DC.) Baehni
Pouteria engleri Eyma
Pouteria eugeniifolia (Pierre) Baehni
Pouteria filipes Eyma
PouteriaJimbriata Baehni
Pouteriafranciscana Baehni
AM
AM
BO
BO
BO
BO
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
BO
DA
DA
DA
DA
DA
DA
DA
GU?
GU su FG
su FG
GU
FG
GU
GU
FG
GU? su FG
GU
GU su FG
GU su FG
GU su FG
GU
GU
GU
GU
DA GU
GU
GU
GU
GU
GU
GU
GU
GU
GU
DA GU
GU
GU
GU
GU
GU
GU
GU
GU
SU
su
su
GU SU
SU
SU
su
su
su
su
su
su
su
su
su
su
su
su
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
FG
SU FG
FG
FG
FG
FG
FG
Checklist of the Plants of the Guiana Shield 517
Pouteria gabhelemis (Gilly ex Aubrev.) T.D. Penn.
Pouteria glomerata (Miq.) Radlk.
Pouteria glomerata subsp. glomerata
Pouteria glomerata subsp. stylosa (Pierre) T.D. Penn.
Pouteria gomphiifolia (Mart, ex Miq.) Radlk.
Pouteria gongrijpii Eyma
Pouteria grandis Eyma
Pouteria guianensis Aubl.
Pouteria hispida Eyma
Pouteriajariensis Pires & T.D. Penn.
Pouteria kaieteurensis T.D. Penn.
Pouteria laevigata (Mart.) Radlk.
Pouteria latianthera T.D. Penn. [expected in FG]
Pouteria lucumifolia (Reissek ex Maxim.) T.D. Penn.
Pouteria macrocarpa (Mart.) Dietr.
Pouteria macrophylla (Lam.) Eyma
Pouteria maguirei (Aubrev.) T.D. Penn.
Pouteria manaosensis (Aubrev. & Pellegr.) T.D. Penn.
Pouteria melanopoda Eyma
Pouteria aff. minima T.D. Penn.
Pouteria oblanceolata Pires
Pouteria orinocoensis (Aubrev.) T.D. Penn.
Pouteria penicillata (Baehni) Aubrev.
Pouteria pimichinensis T.D. Penn.
Pouteria plicata T.D. Penn.
Pouteria procera (Mart.) T.D. Penn.
Pouteria pullei Eyma [imperfectly known species]
Pouteria putamen-ovi T.D. Penn.
Pouteria reticulata (Engl.) Eyma
Pouteria reticulata subsp. reticulata [expected in GU]
Pouteria reticulata subsp. surinamensis T.D. Penn.
Pouteria retinervis T.D. Penn.
Pouteria rigida (Mart. & Eichler) Radlk.
Pouteria rigida subsp. rigida
Pouteria rigida subsp. tomentosa (Aubrev.) T.D. Penn.
Pouteria rigidopsis (Monach.) T.D. Penn.
Pouteria rodriguesiana Pires & T.D. Penn.
Pouteria rostrata (Huber) Baehni
Pouteria sagotiana (Baill.) Eyma
Pouteria sapota (Jacq.) H.E. Moore & Steam [widely cultivated]
Pouteria scrobiculata Monach. ex T.D. Penn.
Pouteria singularis T.D. Penn.
Pouteria sipapoensis T.D. Penn.
Pouteria speciosa (Ducke) Baehni
Pouteria stipitata Cronquist
Pouteria surumuensis Baehni
Pouteria tenuisepala Pires & T.D. Penn.
Pouteria torta (Mart.) Radlk.
Pouteria torta subsp. glabra T.D. Penn.
Pouteria torta subsp. tuberculata (Sleumer) T.D. Penn.
Pouteria trigonosperma Eyma
Pouteria trilocularis Cronquist
Pouteria ucuqui Pires & R.E. Schult.
Pouteria venosa (Mart.) Baehni
Pouteria venosa subsp. amazonica T.D. Penn.
Pouteria virescens Baehni
Pouteria williamii (Aubrev. & Pellegr.) T.D. Penn.
Pradosia beardii (Monach.) T.D. Penn.
Pradosia caracasana (Pittier) T.D. Penn.
Pradosia cochlearia (Lecomte) T.D. Penn.
Pradosia cochlearia subsp. cochlearia
Pradosia grisebachii (Pierre) T.D. Penn.
Pradosia huberi (Ducke) Ducke
Pradosia ptychandra (Eyma) T.D. Penn.
AM
AM BO GU su FG
AM BO
BO su
FG
AM
su FG
GU su FG
AM BO GU su FG
BO DA GU
GU
su
su?
FG
FG
AM FG
AM
FG?
GU? SU FG
AM
su
GU su FG
BO
GU FG
AM BO
GU
AM
BO DA GU
su
FG
FG
AM BO DA GU su FG
AM BO DA
GU
su
su
FG
FG
AM BO GU
AM BO GU
AM BO GU
GU?
GU?
su FG
GU su FG
AM BO GU
FG
AM
GU su FG
BO
AM GU
FG
AM BO GU FG
AM BO GU
GU su
FG
FG
AM
AM
AM BO DA GU su FG
AM BO DA GU
GU
su FG
FG
FG
AM BO
BO
BO DA
GU
FG
FG
FG
GU su FG
518 Checklist of the Plants of the Guiana Shield
GU SU
GU SU
GU SU FG
FG
GU SU FG
GU SU FG
Pradosia schomburgkiana (A. DC.) Cronquist AM BO
Pradosia schomburgkiana subsp. schomburgkiana AM BO
Pradosia schomburgkiana subsp. sericea T.D. Penn. AM
Pradosia surinamensis (Eyma) T.D. Penn. BO
Pradosia verticillata Ducke
Sarcaulus brasiliensis (A. DC.) Eyma
Sarcaulus brasiliensis subsp. brasiliensis
Synonyms and Misidentifications
Achras balata Aubl. [described from cultivated material] =Mimusops sp.
Achras caimito Ruiz & Pav. =Pouteria caimito
Achras venosa Mart. =Pouteria venosa
Achras zapota L. =Manilkara zapota
Achrouteria pomifera Eyma =Chrysophyllum pomiferum
Amorphospermum schomburgkianum (Miq.) Baehni =Elaeoluma schomburgkiana
Bumelia amazonica K. Krause =Pouteria cuspidata subsp. cuspidata
Bumelia cuneifolia Rudge =Chrysophylhim cuneifolium
Bumelia micrantha (Kunth) Willd. (Myrsinaceae) =Stylogyne micrantha
Bumelia nervosa Vahl =Pouteria macrophylla
Calocarpum cooperi (Cronquist) Lundell =Pouteria torta subsp. tuberculata
Calocarpum mammosum (L.) Pierre =Pouteria sapota
Chrysophyllum alnifolium Engl., non Baker =Pouteria engleri
Chrysophyllum ambelaniifolium Sandwith =Pouteria ambelaniifolia
Chrysophyllum anomalum Pires =Pouteria anomala
Chrysophyllum aquaticum Kunth =Micropholis humboldtiana
Chrysophyllum auratum Miq. =Chrysophyllum argenteum subsp. auratum
Chrysophyllum auratum var. majus Miq. =Chrysophyllum argenteum subsp. auratum
Chrysophyllum beardii Monach. =Pradosia beardii
Chrysophyllum brasiliense A. DC. =Sarcaulus brasiliensis
Chrysophyllum brasiliense var. minus Miq. ^Sarcaulus brasiliensis subsp. brasiliensis
Chrysophyllum cayennense A. DC. =Pouteria cayennensis
Chrysophyllum cochlearium Lecomte =Pradosia cochlearia
Chrysophyllum cuspidatum Hoehne =Diploon cuspidatum
Chrysophyllum dumetorum Baehni =Ecclinusa dumetorum
Chrysophyllum emarginatum Klotzsch ex M.R. Schomb. [name of uncertain application]
Chrysophyllum ficoides Steyerm. =Ecclinusa ulei
Chrysophyllum guianense (Eyma) Baehni =Ecclinusa guianensis
Chrysophyllum guyanense Klotzsch ex Miq. =Chrysophyllum argenteum subsp. auratum
Chrysophyllum humbodltianum Roem. & Schult. =Micropholis humboldtiana
Chrysophyllum macoucou Aubl. [incertae sedis; excluded from Sapotaceae by Pennington] =Aubletella macoucou (Aubl.)
Pierre
Chrysophyllum macrophyllum Mart., non Lam. =Sarcaulus brasiliensis subsp. brasiliensis
Chrysophyllum macrophyllum Lam. =Pouteria macrophylla
Chrysophyllum melinonii Engl. =Micropholis guyanensis subsp. guyanensis
Chrysophyllum monachinoanum Steyerm. =Pradosia beardii
Chrysophyllum nitidum G. Mey. =Chrysophyllum argenteum subsp. nitidum
Chrysophyllum obtusatum Willd. ex Roem. & Schult. =Micropholis humboldtiana
Chrysophyllum psilophyllum A. DC. =Chrysophyllum argenteum subsp. nitidum
Chrysophyllum psilophyllum (Sandwith) Baehni =Ecclinusa psilophylla
Chrysophyllum pyriforme Willd. [incertae sedis; excluded from Sapotaceae by Pennington] =Aubletella macoucou (Aubl.)
Pierre
Chrysophyllum reticulatum Engl. =Pouteria reticulata
Chrysophyllum rufocupreum Ducke ^Chrysophyllum eximium
Chrysophyllum schomburgkianum A. DC. ^Pradosia schomburgkiana
Chrysophyllum sericeum A. DC. =Chrysophyllum argenteum subsp. auratum
Chrysophyllum sericeum var. acutifolium Miq. =Chrysophyllum argenteum subsp. auratum
Chrysophyllum sericeum var. obtusifolium Miq. =Chrysophyllum argenteum subsp. auratum
Chrysophyllum sessiliflorum Poir. =Pouteria cayennensis
Chrysophyllum steyermarkii Monach. =Chrysophyllum sparsiflorum
Chrysophyllum ulei K. Krause =Ecclinusa ulei
Cynodendron nitidum (G. Mey.) Aubrev. =Chrysophyllum argenteum subsp. nitidum
Diploon venezuelana Aubrev. =Diploon cuspidatum
Ecclinusa balata Ducke =Chrysophyllum sanguinolentum subsp. balata
Ecclinusa cochlearia (Lecomte) Aubrev. =Pradosia cochlearia
Checklist of the Plants of the Guiana Shield 519
Ecclinusa ciineifolia (Rudge) Aubrev. "^Chrysophyllum cuneifolium
Ecclinusa grisebachii Pierre =Pradosia grisebachii
Ecclinusa prieurii (A. DC.) Aubrev. =Chrysophyllum prieurii
Ecclinusa sanguinolenta (Pierre) Engl. =Chrysophyllum sanguinolentum
Ecclinusa spuria Ducke =Chysophyllum sanguinolentum subsp. spurium
Ecclinusa ucuquirana-branca Aubrev. & Pellegr. =Chrysophyllum ucuquirana-branca
Englerella macrocarpa Pierre =Pouteha speciosa
Englerella speciosa Ducke =Pouteria speciosa
Eremoluma sagotiana Baill. =Poutena sagotiana
Eremoluma williamii Aubrev. & Pellegr. =Pouteria williamii
Franchetella atabapoensis Aubrev. =Pouteria atabapoensis
Franchetella gongghjpii (Eyma) Aubrev. =Pouteria gongrijpii
Franchetella neblinaensis Aubrev. =Pouteria trilocularis
Glycoxylon maguirei Aubrev. =Pradosia schomburgkiana subsp. schomburgkiana
Glycoxylon rubrum Aubrev. =Pradosia beardii
Gomphiluma martiana Baill. =Pouteria gomphiifolia
Guapeba coriacea Pierre (Tectariaceae) =Poiiteria coriacea
Guapeba stylosa Pierre =Pouteria glomerata subsp. stylosa
Ichthyophora elegans (Miq.) Baehni =Pouteria elegans
Ichthyophora leptocarpa (Miq.) Baehni =Pouteria elegans
Labatia ? bilocularis H. Winkl. =Pouteria bilocularis
Labatia laevigata Mart. =Pouteria laevigata
Labatia parviflora Pittier =Pouteria glomerata subsp. glomerata
Labatia pedunculata Willd. =Pouteria guianensis
Labatia torta Mart. =Pouteria torta
Labatia tovarensis Engl. =Pouteria guianensis
Lucuma duckei Huber =Pouteria venosa subsp. amazonica
Lucuma glabrescens Mart. & Eichler =Elaeoluma glabrescens
Lucuma glomerata Miq. =Pouteria glomerata
Lucuma gomphiifolia Mart, ex Miq. =Pouteria gomphiifolia
Lucuma gomphiifolia var. blepharantha Mart. =Pouteria gomphiifolia
Lucumajenmanii Pittier =Pouteria torta subsp. glabra
Lucuma procera Mart. =Pouteria procera
Lucuma pulverulenta Mart. & Eichler =Pouteria cayennensis
Lucuma rigida Mart. & Eichler =Pouteria rigida
Lucuma rostrata Huber =Pouteria rostrata
Lucuma sagotiana (Baill.) Engl. =Pouteria sagotiana
Lucuma sericea K. Krause =Pouteria surumuensis
Lucuma speciosa Ducke =Pouteria speciosa
Lucuma temare Kunth =Pouteria caimito
Manilkara amazonica (Huber) A. Chev. =Manilkara bidentata subsp. surinamensis
Manilkara balata (Pierre) Dubard =Manilkara bidentata subsp. bidentata
Manilkara surinamensis (Miq.) Dubard =Manilkara bidentata subsp. surinamensis
Manilkara williamsii Standi. =^Manilkara bidentata subsp. bidentata
Meioluma guianensis Baill. =Micropholis venulosa
Micropholis burchelliana Pierre =Micropholis gardneriana
Micropholis calophylloides Pierre =Micropholis venulosa
Micropholis cowanii Aubrev. =Micropholis guyanensis subsp. guyanensis
Micropholis eugeniifolia Pierre =Pouteria eugeniifolia
Micropholis melinonii (Engl.) Mennega et al. =Micropholis guyanensis subsp. guyanensis
Micropholis mucronata Pierre =Micropholis venulosa
Micropholis paraens is Huber ex Eyma =Micropholis acutangula
Micropholis robusta (Mart. & Eichler) Mennega et al. =Pouteria cuspidata subsp. robusia
Micropholis steyermarkii Monach. =Pouteria eugeniifolia
Micropholis wurdackii Aubrev. =Micropholis egensis
Mimusops balata (Aubl.) C.F. Gaertn. [see Achras balata}
Mimusops balata var. gutta Pierre =Manilkara bidentata subsp. bidentata
Mimusops balata var. melinonis Pierre =Manilkara bidentata subsp. bidentata
Mimusops balata var. schomburgkii Pierre =Manilkara bidentata subsp. bidentata
Mimusops bidentata A. DC. =Manilkara bidentata
Mimusops huberi Ducke =Manilkara huberi
Mimusops paraensis Huber =Manilkara paraensis
Mimusops surinamensis Miq. =Manilkara bidentata subsp. surinamensis
Myrtiluma eugeniifolia (Pierre) Aubrev. =Pouteria eugeniifolia
520 Checklist of the Plants of the Guiana Shield
Nemaluma engleri (Eyma) Aubrev. & Pellegr. =Pouteria engleri
Neopometia ptychandra (Eyma) Aubrev. =Pradosia ptychandra
Neoxythece atabapoensis Aubrev. =Ecclinusa atabapoensis
Neoxythece cladantha (Sandwith) Aubrev. =Pouteria cladantha
Neoxythece dura (Eyma) Aubrev. & Pellegr. =Poiiteria cuspidata subsp. dura
Neoxythece gabrielensis Gilly ex Aubrev. =Pouteria gabrielensis
Neoxythece maguirei Aubrev. =Poiiteria pimichinensis
Neoxythece robusta (Mart. & Eichler) Aubrev. & Pellegr. =Poiiteria cuspidata subsp. robusta
Neoxythece roraimaensis Aubrev. =Pouteria rigida subsp. rigida
Neoxythece schulzii Aubrev. =Pouteria cuspidata subsp. dura
Neoxythece tomentosa Aubrev. =Pouteria rigida subsp. tomentosa
Oxythece ambelaniifolia (Sandwith) Cronquist =Pouteria ambelaniifolia
Oxythece caracasana Pittier =Pradosia caracasana
Oxythece dura (Eyma) Gilly =Pouteria cuspidata subsp. dura
Oxythece elegans (A. DC.) Urb. =Pouteria elegans
Oxythece guianensis Lecomte ? =Pouteria^guianensis
Oxythece leptocarpa Miq. =Pouteria elegans
Oxythece rigida (Mart. & Eichler) Monach. =Pouteria rigida
Oxythece robusta var. brevipetiolasta Monach. =Pouteria cuspidata subsp. robusta
Oxythece schomburgkiana (Miq.) Cronquist =Elaeoluma schomburgkiana
Oxythece scrobiculata Monach. [nom. inval.] =Pouteria scrobiculata
Oxythece steyermarkiana Monach. =Elaeoluma schomburgkiana
Paralabatia acutangula (Ducke) Baehni =Micropholis acutangula
Paralabatia casiquiarensis Aubrev. =Pouteria lucumifolia
Paralabatia orinocoensis Aubrev. =Pouteria orinocoensis
Paramicropholis acutangula (Ducke) Aubrev. & Pellegr. =Micropholis acutangula
Passaveria lanceolata Mart. & Eichler =Ecclinusa lanceolata
Planchonella elegans (A. DC.) Baehni =Pouteria elegans
Planchonella guianensis P. Royen =Chtysophyllum pomiferum
Planchonella melinonii (Engl.) Baehni =Micropholis guyanensis subsp. guyanensis
Platyluma calophylloides (Pierre) Baill. =Micropholis venulosa
Podoluma benai Aubrev. & Pellegr. =Pouteria benai
Pouteria acutangula (Ducke) Baehni =Micropholis acutangula
Pouteria auyantepuiensis Steyerm. =Elaeoluma nuda
Pouteria benoistii Aubrev. =Pouteria hispida
Pouteria burchelliana (Pierre) Baehni =Micropholis gardneriana
Pouteria caimito var. strigosa Monach. =Pouteria guianensis
Pouteria demerarae Sandwith =Pouteria guianensis
Pouteria duckeana Baehni =Micropholis guyanensis subsp. duckeana
Pouteria dura Eyma =Pouteria cuspidata subsp. dura
Pouteria echinocarpa W.A. Rodrigues =Pouteria torta subsp. tuberculata
Pouteria elegans var. micrantha (Miq.) Baehni =Pouteria cuspidata subsp. cuspidata
Pouteriajenmanii (Pittier) Sandwith =Pouteria torta subsp. glabra
Pouteria lucentifolia (Standi.) Baehni =Chrysophyllum venezuelanense
Pouteria melinoniana (Pierre) Baehni =Micropholis melinoniana
Pouteria melinonii (Engl.) Baehni =Micropholis guyanensis subsp. guyanensis
Pouteria mensalis Baehni =Micropholis mensalis
Pouteria minutiflora (Britton) Sandwith =Pouteria coriacea
Pouteria multiflora (A. DC.) Eyma [not in flora area]
Pouteria nuda Baehni =Elaeoluma nuda
Pouteria obidensis Huber =Pouteria guianensis
Pouteria pedunculata (Willd.) Poir. =Pouteria guianensis
Pouteria pittieri Baehni =Pouteria glomerata subsp. glomerata
Pouteria pomifera (Eyma) Baehni =Chrysophyllum pomiferum
Pouteria porphyrocarpa Baehni =Micropholis porphyrocarpa
Pouteria ptychandra Eyma =Pradosia ptychandra
Pouteria robusta (Mart. & Eichler) Eyma =Pouteria cuspidata subsp. robusta
Pouteria robusta var. longifolia Eyma =Pouteria cuspidata subsp. robusta
Pouteria rufa (Mart. & Eichler ex Mart.) Lemee =Micropholis guyanensis
Pouteria scrobiculata Monach. [nom. inval.] =Pouteria scrobiculata
Pouteria scytalophora Eyma =Pouteria bangii
Pouteria sericea (Alton) Baehni [Australian species, recorded from GU in error]
Pouteria spectabilis Steyerm. ^^Micropholis spectabilis
Pouteria spuria (Ducke) Pittier et al. =Chrysophyllum sanguinolentum subsp. spurium
Checklist of the Plants of the Guiana Shield 521
Pouteria steyermarkii Monach. =Pouteria eugeniifolia
Pouteria suffulta Baehni =Poiiteria durlandii subsp. durlandii
Pouteria siirinamensis Eyma =Pradosia surinamensis
Pouteria tovarensis Klotzsch & Karst. ex Engl. & Prantl =Pouteria guianensis
Pouteria venamoensis Steyerm. =MicrophoJis venamoensis
Pouteria venulosa (Mart. & Eichler) Baehni =Micropholis venulosa
Prieurella cuneifolia (Rudge) Pierre ex Aubrev. =ChrysophyUum cuneifolium
Prieurella maguirei Aubrev. =Pouteria maguirei
Prieurella prieurii (A. DC.) Aubrev. =Chiysophyllum prieurii
Pseudocladia minutiflora (Britton) Aubrev. =Pouteria coriacea
Pseudocladia neblinaensis Aubrev. =Pouteria bilocularis
Pseudocladia orinocoensis Aubrev. =Pouteria lucumifolia
Pseudocladia scytalophora (Eyma) Aubrev. =Pouteria bangii
Pseudolabatia filipes (Eyma) Aubrev. =Pouteria filipes
Pseudolabatia penicillata (Baehni) Aubrev. =Pouteria penicillata
Pseudoxythece ambelaniifolia (Sandwith) Aubrev. =Pouteria ambelaniifolia
Radlkoferella brachyandra Aubrev. & Pellegr. =Pouteria brachyandra
Radlkoferella grandis (Eyma) Aubrev. =Pouteria grandis
Radlkoferella trigonosperma (Eyma) Aubrev. =Pouteria trigonosperma
Radlkoferella venosa (Mart.) Aubrev. =Pouteria venosa
Ragala sanguinoleuta Pierre =Chrysophyllum sanguinolentum
Ragala ulei (K. Krause) Aubrev. =Ecclinusa ulei
Richardella cayennensis (A. DC.) Aubrev. =Pouteria cayennensis
Richardella cladantha (Sandwith) Baehni =Pouteria cladantha
Richardella macrophylla (Lam.) Aubrev. =Pouteria macrophylla
Richardella surumuensis (Baehni) Aubrev. =Pouteria surumuensis
Richardella trigonosperma (Eyma) Baehni =Pouteria trigonosperma
Sandwithiodoxa egregia (Sandwith) Aubrev. =Pouteria egregia
Sideroxylon acutangulum Ducke =Micropholis acutangula
Sideroxylon amygdalicarpum Pittier =Pouteria amygdalicarpa
Sideroxylon bangii Rusby =Pouteria bangii
Sideroxylon calophylloides (Pierre) Engl. =Micropholis venulosa
Sideroxylon cuspidatum A. DC. =Pouteria cuspidata
Sideroxylon cuspidatum var. crassifolium Miq. =Pouteria cuspidata subsp. dura
Sideroxylon cuspidatum var. ellipticum Miq. =Pouteria cuspidata subsp. cuspidata
Sideroxylon egense A. DC. =Micropholis egensis
Sideroxylon elegans A. DC. =Pouteria elegans
Sideroxylon elegans var. micranthum Miq. =Pouteria cuspidata subsp. cuspidata
Sideroxylon eugeniifolium (Pierre) Engl. =Pouteria eugeniifolia
Sideroxylon guianense (Baill.) Engl. =Micropholis venulosa
Sideroxylon guyanense A. DC. ^Micropholis guyanensis
Sideroxylon robustum Mart. & Eichler =Pouteria cuspidata subsp. robusta
Sideroxylon rufum Mart. & Eichler ex Mart. =Micropholis guyanensis subsp. guyanensis
Sideroxylon rugosum var. parviflorum Miq. =Micropholis melinoniana
Sideroxylon venulosum Mart. & Eichler =Micropholis venulosa
Syzygiopsis sericea Cronquist =Pouteria arcuata
Vitellaria pulverulenta (Mart. & Eichler) Radlk. =Pouteria cayennensis
Referrncks
(for a more extensive bibliography, see Pennington 1990)
Aubreville, A. 1972. Sapotaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part IX. Mem. Now >'ork Bot.
Card. 23: 1-832.
Eyma, RJ. 1936. Sapotaceae. In A. Pulle, Flora of Suriname 4( 1 ): 354-399.
Pennington, T.D. 1990. Sapotaceae. Flora Neotropica Monograph 52: 1-770.
Pennington, T.D. 2002. Sapotaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. dc Granville, S.V. Heald, M. HolT, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. (iard. 76 (2):
669-683.
Pennington, T.D. & K.S. Edwards. 2005. Sapotaceae. In J. A. Steyermark, PH. Berry, K. Yatskicvych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 89-137.
Sarraceniaceae
RE. Berry, K.M. Cameron, R. Riina, J. A. Steyermarkt
Heliamphora chimantensis Wistuba, Carow & Harbarth BO
522 Checklist of the Plants of the Guiana Shield
Heliamphora elongata Nerz
Heliamphora folliculata Wistuba, Harbarth & Carow
Heliamphora heterodoxa Steyerm.
Heliamphora heterodoxa var. exappendicidata Maguire & Steyerm.
Heliamphora heterodoxa var. heterodoxa
Heliamphora hispida Wistuba & Nerz
Heliamphora ionasii Maguire
Heliamphora minor Gleason
Heliamphora nutans Benth.
Heliamphora sarracenioides Carow, Wistuba & Harbarth
Heliamphora tatei Gleason
Heliamphora tatei var. neblinae (Maguire) Steyerm.
Heliamphora tatei var. tatei
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
BO
BO
BO
GU
Synonyms and Misidentifications
Heliamphora heterodoxa var. glabra Maguire =Heliamphora heterodoxa var. heterodoxa
Heliamphora macdonaldae Gleason =Heliamphora tatei var. tatei
Heliamphora neblinae Maguire =Heliamphora tatei var. neblinae
Heliamphora neblinae var. viridis Maguire =Heliamphora tatei var. neblinae
Heliamphora tatei var. macdonaldae (Gleason) Maguire ^Heliamphora tatei var. tatei
Heliamphora tatei var. parva Maguire =Heliamphora tatei var. neblinae
Heliamphora tyleri Gleason =Heliamphora tatei var. tatei
References
Berry, P.E., R. Riina, & J.A. Steyermark. 2005. Sarraceniaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 9: 138-144.
Maguire, B. 1978. Sarraceniaceae. In B. Maguire & collaborators. Botany of the Guayana Highland part 10. Mem. New York
Bot. Gard. 29: 36-62.
SCROPHULARIACEAE
N.H. Holmgren, K.A. Vincent
Achetaria guianensis Pennell
Achetaria ocimoides (Cham. & Schltdl.) Wettst.
Achetaria scutellarioides (Benth.) Wettst.
Alectra aspera (Cham. & Schltdl.) L.O. Williams
Alectra stricta Benth.
Angelonia angiistifolia Benth. [probably cultivated only]
Angelonia biflora Benth.
Angelonia salicariifolia Bonpl. [probably cultivated only]
Anisantherina hispidula (Mart.) Pennell
Bacopa aquatica Aubl.
Bacopa bacopoides (Benth.) PuUe
Bacopa callitrichoides (Kunth) Pennell
Bacopa connata (Pennell) Pennell
Bacopa gratioloides (Cham.) Chodat & Hassl.
Bacopa innominata (M. Gomez) Alain
Bacopa laxiflora (Benth.) Wettst. ex Edwall
Bacopa monnieri (L.) Wettst.
Bacopa monnierioides (Cham.) B.L. Rob.
Bacopa repens (Sw.) Wettst.
Bacopa reptans (Benth.) Wettst. ex Edwall
Bacopa salzmannii (Benth.) Wettst. ex Edwall
Bacopa serpyllifolia (Benth.) Pennell
Bacopa sessiliflora (Benth.) Edwall
Bacopa stricta (Schrad.) Wettst. ex Edwall
Benjaminia reflexa (Benth.) D'Arcy
Buchnera longifolia Kunth
Buchnera palustris (Aubl.) Spreng.
Buchnera pusilla Kunth
Buchnera rosea Kunth
Buchnera spruceana Philcox
Buchnera ternifolia Kunth
Buchnera weberbaueri Diels
Capraria biflora L.
BO GU SU FG
GU SU FG
BO GU FG
BO GU FG
AM BO
GU
SU FG
BO GU SU
AM BO GU SU FG
BO DA GU SU
SU
FG
AM BO
BO
BO DA
GU
GU
AM BO
BO
GU
GU
SU
FG
AM BO GU
AM GU
AM BO GU
AM BO GU
BO DA GU SU FG
BO GU
AM BO GU SU FG
AM BO GU SU FG
AM BO GU SU FG
AM BO GU
AM BO GU SU
AM
BO
AM BO DA GU
AM BO DA GU SU FG
BO
AM BO GU su FG
AM
AM BO
AM BO GU
AM BO DA GU su FG
AM BO GU su FG
AM DA GU su FG
DA GU su FG
AM BO DA FG
DA GU su
FG
AM BO
BO
DA GU
GU
GU
GU
GU
su
su
FG
FG
AM DA GU su
AM BO GU
Checklist of the Plants of the Guiana Shield 523
Castilleja arvemis Schltdl. & Cham.
Conobea aquatica Aubl.
Conobea scoparioides (Cham. & Schltdl.) Benth.
Escobedia grandiflora (L. f.) Kuntze
Lindernia brachyphylla Pennell
Lindernia Crustacea (L.) F. Muell.
Lindernia diffusa (L.) Wettst.
Lindernia dubia (L.) Pennell
Lindernia microcalyx Pennell & Stehle
Mecardonia procumbens (Mill.) Small
Micranthemum umbrosum S.F. Blake
Russelia sarmentosa Jacq.
Scoparia dulcis L.
Stemodia foliosa Benth.
Stemodia pratensis (Aubl.) C.P. Cowan [=Stemodiafoliosa ?]
Stemodia verticillata (Mill.) Hassl.
Toreniafournieri Linden ex E. Foum. [probably cultivated or escaped from
cultivation]
Torenia thouarsii (Cham. & Schltdl.) Kuntze [naturalized]
Vellosiella spathacea (Oliv.) Melch.
Synonyms and Misidentifications
Achetaria scutellarioides (Benth.) Kuntze [nom. illeg.] =Achetaria scutellarioides
Agalinis hispidula (Mart.) D'Arcy =Anisantherina hispidula
Alectra brasiliensis Benth. =A!ectra aspera
Alectrafluminensis (Veil.) Steam =Alectra aspera
Bacopa chamaedryoides (Kunth) Wettst. =Mecardoma procumbens
Bacopa ciliata (Pennell) Pennell =Bacopa serpyllifolia
Bacopa conferta (Pennell) Standi. =Bacopa sessiliflora
Bacopa domingensis (Spreng.) Pennell =Bacopa stricta
Bacopa punctata Pennell =Bacopa monnierioides
Bacopa reflexa (Benth.) Edwall =Benjaminia reflexa
Bacopa sessiliflora (Benth.) Pulle [nom. illeg.] =Bacopa sessiliflora
Benjaminia utriculariiformis Mart, ex Benj. =Benjaminia reflexa
Beyrichia ocimoides Cham. & Schltdl. =Achetaria ocimoides
Beyrichia scutellarioides Benth. =Achetaria scutellarioides
Bramia monnieri (L.) Drake =Bacopa monnieri
Buchnera capillaris Desv. =Lindernia Crustacea
Buchnera elongata Sw. [records may represent Buchnera pusilla] =Buchnera longifolia
Buchnera grandiflora L. f. =Escobedia grandiflora
Buchnera lithospermifolia Kunth =Buchnera ternifolia
Buchnera lobelioides Cham. & Schltdl. =Buchnera ternifolia
Buchnera macrocarpa Kunth =Buchnera longifolia
Buchnera virgata Kunth [name dubiously applied to Guianan specimens]
Caconapea auriculata (B.L. Rob.) Pennell =Bacopa laxiflora
Caconapea conferta Pennell =Bacopa sessiliflora
Caconapea debilis Pennell =Bacopa reptans
Caconapea gratioloides Cham. =Bacopa gratioloides
Caconapea sessiliflora (Benth.) Pennell =Bacopa sessiliflora
Caconapea stricta (Schrad.) Britton =Bacopa stricta
Capraria biflora f. hirsuta Loes. =Capraria biflora
Capraria Crustacea L. =Lindernia Crustacea
Capraria integerrima Miq. ( Pedal iaceae) =Sesamum orientale
Conobea innominata M. Gomez =Bacopa innominata
Curanga bivalvis (L. f.) Druce (Mendonciaceae) =Mendoncia hivalvis
Drupina cristata L. (Mendonciaceae) ^Mendoncia bivalvis
Erinus procumbens Mill. =Mecardonia procumbens
Escobedia parimensis Pennell =Escobedia grandiflora
Gerardia hispidula Mart. =Anisantherina hispidula
Globifera umbrosum J.F. Gmel. =Micranthemum umbrosum
Glossostylis aspera Cham. & Schltdl. =Alectra aspera
Gratiola dubia L. =Lindernia dubia
Gratiola repens Sw. =Bacopa repens
Herpestis auriculata B.L. Rob. =Bacopa laxiflora
Herpestis caprarioides Kunth =Mecardonia procumbens
524 Checklist of the Plants of the Guiana Shield
Herpestis chamaedryoides Kunth =Mecardoma procumbens
Herpestis ciliata Pennell =Bacopa serpyllifolia
Herpestis connata Pennell =Bacopa connata
Herpestis domingensis Spreng. =Bacopa striata
Herpestis laxiflora Benth. =Bacopa laxiflora
Herpestis procumbens (Mill.) Urb. =Mecardonia procumbens
Herpestis ranaria Benth. =Bacopa monnierioides
Herpestis reflexa Benth. =Benjaminia reflexa
Herpestis reptans Benth. =Bacopa reptans
Herpestis salzmanni Benth. =Bacopa salzmannii
Herpestis serpyllifolia Benth. =Bacopa serpyllifolia
Herpestis sessiliflora Benth. =Bacopa sessiliflora
Herpestis sessiliflora Benth. =Bacopa sessiliflora
Herpestis striata Schrad. =Baaopa striata
Hydranthelium aallitriahoides Kunth =Baaopa callitrichoides
Ilysanthes dubia (L.) Barahart =Lindernia flubia
Ilysanthes riparia Raf. =Lindernia dubia
Ilysanthes rotundifolia Benth. =Lindernia rotundifolia
Lindernia barrosorum L.B. Sm. =Lindernia brachyphylla
Lindernia rotundifolia (L.) Standi. & L.O. Williams [nom. illeg.] =Lindernia microcalyx
Maauillamia limosa Pennell =Bacopa repens
Maauillamia repens (Sw.) Pennell ^Baaopa repens
Matourea pratensis Aubl. =Stemodia pratensis
Melasma brasiliensis Chodat & Hassl. =Aleatra aspera
Melasma melampyroides (Rich.) Pennell =Alectra aspera
Melasma spathaceum Oliv. =Vellosiella spathaaea
Mella laxiflora (Benth.) Pennell =Bacopa laxiflora
Micranthemum orbiaidatum Michx. =Miaranthemum umbrosum
Micranthemum pilosum Ernst =Miaranthemum umbrosum
Monoaardia humilis Pennell =Baaopa salzmannii
Monocardia lilaaina Pennell =Baaopa salzmannii
Monoaardia violaaea Pennell =Bacopa salzmannii
Naiadothrix reflexa (Benth.) Pennell =Benjaminia reflexa
Nortenia thouarsii Cham. & Schltdl. =Torenia thouarsii
Pediaularis melampyroides Rich. =Aleatra aspera
Picria surinamensis Spreng. (Mendonciaceae) =Mendonaia bivalvis
Piripea palustris Aubl. =Buchnera palustris
Ranaria monnierioides Cham. =Bacopa monnierioides
Scrophulariafluminensis Veil. =Aleatra aspera
Sphaerotheca scoparioides Cham. & Schltdl. =Conobea scoparioides
Stemodia parviflora Alton =Stemodia vertiaillata
Stemodia pusilla Benth. [name apparently misapplied to S. pratensis^
Stemodia surinamensis Miq. [name of uncertain application]
Torenia Crustacea (L.) Cham. & Schltdl. =Lindernia Crustacea
Torenia thouarsii var. nivea Pennell =Torenia thouarsii
Vandellia Crustacea (L.) Benth. =Lindernia Crustacea
Vandellia diffusa L. =Lindernia diffusa
References
D'Arcy, W.G. 1979. Scrophulariaceae. In R.E. Woodson, Jr., R.W. Schery, & collaborators. Flora of Panama. Ann. Missouri Bot.
Card. 66: 173-272.
Diaz-Miranda, D. 1991. Scrophulariaceae en la flora de Venezuela I. Introduccion. Emstia 1(3): 95-108.
Holmgren, N.H. 2002. In Mori, S.A., G. Cremers, C. Oracle, J.J. de Granville, S.V Heald, M. Hoff, & J.D. Mitchell, eds. 2002.
Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76(2): 683-685.
Holmgren, N.H. & K.A. Vincent. 2005. Scrophulariaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of
the Venezuelan Guayana 9: 144-167.
Pennell, F.W. 1940. Some new species of Scrophulariaceae from Guiana. Not. Nat. Acad. Nat. Sci. Phila. 46: 1-6.
Philcox, D. 1965. Revision of the New World species of Buchnera L. Kew Bull. 18: 275-315.
Turner, B.L. & C.C. Cowan. 1993. Taxonomic overview of Stemodia for South America. Phytologia 75(4): 281-324.
SiMAROUBACEAE
E.V Franceschinelli, W.W. Thomas
Picrolemma sprucei Hook. f. AM GU FG
Checklist of the Plants of the Guiana Shield 525
Quassia amara L. [cultivated, naturalized]
Simaba cedron Planch, [cultivated, naturalized]
Simaba gidanensis Aubl.
Simaba guianensis subsp. ecaudata (Cronquist) Cavalcante
Simaba guianensis subsp. guianensis
Simaba guianensis subsp. huberi Francesch. & W. Thomas
Simaba monophylla (Oliv.) Cronquist
Simaba morettii Feuillet
Simaba obovata Spruce ex Engl.
Simaba orinocensis Kunth
Simaba paraensis Ducke
Simaba polyphylla (Cavalc.) W.W. Thomas
Simaba sp. A
Simarouba amara Aubl.
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
BO
GU SU EG
GU su EG
GU SU EG
GU su EG
GU su EG
DA
DA
DA
GU
GU
GU
GU
su
SU
FG
EG
FG
FG
Synonyms and Misidentificatioxs
Aeschhon excelsa (S\v.) Kuntze (Picramniaceae) =Picrasma excelsa
Aruba guianensis Aubl. =Simaba guianensis
Picrolemma pseudocoffea Ducke =Picrolemma spnicei
Quassia cedron (Planch.) Baill. ^Simaba cedron
Quassia cuspidata (Spruce ex Engl.) Noot. =Simaba guianensis
Quassia guianensis (Aubl.) Dietr. =Simaba guianensis
Quassia monophylla (Oliv.) Noot. =Simaba monophylla
Quassia multiflora (A. Juss.) Noot. =Simaba orinocensis
Quassia simarouba L. f. =Simarouba amara
Simaba alata Maguire =Simaba orinocensis
Simaba cuspidata Spruce ex Engl. =Simaba guianensis
Simaba guianensis subsp. polyphylla Cavalcante =Simaba polyphylla
Simaba multiflora A. Juss. =Simaba orinocensis
Simaba trichilioides A. St.-Hil. [not in flora area]
Simarouba monophylla Oliv. =Simaba monophylla
References
Cavalcante, P.B. 1983. Revisao taxonomica do genero Simaba Aublet na America do Sul. Publ. Avulsas Mus. Goeldi 37: 1-85.
Cronquist, A. 1944. Studies in the Simaroubaceae. II. The genus Simarouba. Bull. Torrey Bot. Club 71: 226-234.
Cronquist, A. 1944. Studies in Simaroubaceae. III. The genus Simaba. Lloydia 7: 81-92.
Cronquist, A. 1944. Studies in Simaroubaceae. IV. Resume of the American genera. Brittonia 5: 128-147.
Cronquist, A. 1945. Additional notes on the Simaroubaceae. Brittonia 5: 469-470.
Feuillet, C. 1983. Etudes sur les Simaroubaceae. II. Un Simaba nouveau de Guyane fran9aise dans la section Fioribundae Engl.:
S. morettii. Candollea 38(2): 745-750.
Jansen-Jacobs, M.J. 1979. Simaroubaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 319-330.
Nooteboom, H.P. 1962. Generic delimitation in Simaroubaceae tribus Simaroubeae and a conspectus of the genus Quassia L.
Blumea 11: 509-528.
Thomas, W.W. 1984. Anew species of Simaba (Simaroubaceae) from Para. Brazil, with a key to the species north of the Amazon
River. Brittonia 36(3): 244-247.
Thomas, W.W. 1 985. The Simaba guianensis complex in northern South America. Acta Amazonica 1 5( 1 -2) suppl.: 7 1 -79.
Thomas, W.W. 2002. Simaroubaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. HotT, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 685-689.
Thomas, W.W. & E.V. Eranceschinelli. 2005. Simaroubaceae. In J.A. Steyermark, PE. Berr>. K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 9: 168-176.
Siparuna bifida (Poepp. & Endl.) A. DC.
Siparuna cristata (Poepp. & Endl.) A. DC.
Siparuna cuspidata (Tul.) A. DC.
Siparuna decipiens (Tul.) A. DC.
Siparuna ficoides S.S. Renner & Hausner
Siparuna guianensis Aubl.
Siparuna obstipa J.F. Macbr.
Siparuna pachyantha A.C. Sm.
Siparuna poeppigii (Tul.) A. DC.
Siparuna reginae (Tul.) A. DC.
SlPARUNACEAE
G. Hausner, S.S. Renner
AM BO
AM BO GU SU FG
AM GU SU FG
AM BO
BO
DA GU SU FG
AM BO DA GU SU FG
AM
GU
GU su
FG
FG
AM BO GU su
526 Checklist of the Plants of the Guiana Shield
Synonyms and Misidentifications
Citrosma amazonica (Mart.) A. DC. =Siparuna poeppigii
Citrosma bifida Poepp. & Endl. =Siparuna bifida
Citrosma cristata Poepp. & Endl. =Siparuna chstata
Citrosma cuspidata Tul. =Siparuna cuspidata
Citrosma decipiens Tul. =Siparuna decipiens
Citrosma guianensis (Aubl.) Tul. =Siparima giiianensis
Citrosma reginae Tul. =Siparuna reginae
Conuleum guianense A. Rich. =Siparuna decipiens
Siparuna amazonica (Mart.) A. DC. =Siparuna poeppigii
Siparuna argyrochrysea Perkins =Siparuna cuspidata
Siparuna emarginata R.S. Cowan =Siparuna pachyantha
Siparuna micrantha A. DC. =Siparuna reginae
Siparuna sancheziana Steyerm. =Siparuna reginae
Siparuna sprucei A. DC. =Siparuna poeppigii
Siparuna surinamensis Lanj. = Siparuna vqquinae (Tul.) A.DC.
References
Jansen-Jacobs, M.J. 1976. Monimiaceae (additions and corrections). In J. Lanjouw & A.L. Stoffers, Flora of Suriname 2(2): 425-
426.
Lanjouw, J. 1939. Monimiaceae (additions and corrections). In A. Pulle, Flora of Suriname 2(1): 471-472.
Mitchell, J.D. 2002. Monimiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
510-515.
Perkins, J. & E. Gilg. 1901. Monimiaceae. In A. Engler, Das Pflanzenreich IV(lOl): 1-122.
Petter, H.F.M. 1932. Monimiaceae. In A. Pulle, Flora of Suriname 2(1): 107-112.
Renner, S.S. & G. Hausner. 2005. Siparunacae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 180-184.
Steyermark, J.A. 1951. Monimiaceae. In Botanical Exploration in Venezuela - 1. Fieldiana Bot. 28(1): 235-236.
SOLANACEAE
W.G D'Arcy, C. Benitez de Rojas, M.H. Nee
Brunfelsia guianensis Benth.
Brunfelsia imatacana Plowman
Brunfelsia martiana Plowman
Brunfelsia uniflora (Pohl) D. Don
Capsicum annuum L.
Capsicum annuum var. annuum [cultivated, frequently escaping]
Capsicum annuum var. glabriusculum (Dunal) Heiser & Pickersgi
Capsicumfrutescens L.
Capsicum spp.
Cestrum alternifolium (Jacq.) O.E. Schulz
Cestrum glabrescens (C.V. Morton) Steyerm. & Maguire
Cestrum latifolium Lam.
Cestrum multiflorum Roem. & Schult.
Cestrum neblinense D'Arcy & Benitez
Cestrum racemosum Ruiz & Pav.
Cestrum reflexum Sendtn.
Cestrum schlechtendahlii G. Don
Cestrum schulzianum Francey
Cestrum strigilatum Ruiz & Pav.
Cestrum subuniflorum Dunal
Cestrum tubulosum Sendtn.
Cyphomandra endopogon Bitter
Cyphomandra endopogon subsp. guianensis Bohs
Cyphomandra hartwegii (Miers) Walp.
Cyphomandra hartwegii subsp. hartwegii
Cyphomandra oblongifolia Bohs
Cyphomandra tegore (Aubl.) Walp.
Cyphomandra tobagensis Sandwith
Datura inoxia Mill.
Datura metel L. [cultivated or escaped from cultivation]
Datura stramonium L. [naturalized?]
GU SU FG
BO
GU SU FG
BO GU
AM BO DA GU
GU
SU FG
FG
AM BO DA GU SU
AM BO
BO
DA GU SU FG
AM BO
AM BO DA GU SU FG
FG
AM
AM
BO
GU? FG?
AM BO GU FG
AM
AM
AM
AM BO
FG
FG
GU SU FG
GU SU
SU
FG
SU FG
BO GU
BO
SU FG
FG?
AM GU su FG
AM BO GU su FG
AM BO
AM BO
BO
DA GU su FG
AM BO DA GU su FG
AM BO
BO
DA
DA
GU
GU
su FG
AM BO
AM BO DA GU su FG
AM BO GU su FG
AM BO GU FG
AM BO
AM BO GU su FG
AM
Checklist of the Plants of the Guiana Shield 527
Deprea orinocensis (Kunth) Raf. BO
Hawkesiophyton itlei (Dammer) Hunz. AM FG
Lycianthes geminata Bitter [expected in GU, SU] FG
Lycianthes heteroclita (Sendtn.) Bitter SU
Lycianthes nitida Bitter GU
Lycianthes pauciflora{Wdih\)B\XXQr AM BO GU SU FG
Markea coccinea Rich. GU SU FG
Markeaformicarum Dammer
Markea longiflora Miers
Markea reticulata Steyerm. & Maguire
Markea sessiliflora Ducke
Melananthus ulei Carvalho
Nicotiana tabacum L. [cultivated, perhaps escaping]
Physalts angulata L.
Physalts cordata Mill.
Physalis ignota Britton [probably cultivated or escaped from cultivation]
Physalts lagascae Roem. & Schult.
Physalis pubescens L.
Schwenckia americana L.
Schwenckia elegans Carvalho
Schwenckia glabrata Kunth
Schwenckia grandiflora Benth.
Schwenckia heterantha Carvalho
Schwenckia huberi Benitez
Schwenckia lateriflora (Vahl) Carvalho GU
Schwenckia micrantha Benth. BO DA
Schwenckia mollissima Nees & Mart.
Solandra grandiflora Sw.
Solandra longiflora Tussac
Solandra paraensis Ducke
Solanum accrescens Standi. & C.V. Morton
Solanum acerifolium Dunal
Solanum adhaerens Roem. & Schult.
Solanum agrarium Sendtn. [expected in SU, FG]
Solanum allophyllum (Miers) Standi.
Solanum altissimum Pittier ex. Benitez
Solanum americanum Mill.
Solanum anceps Ruiz & Pav.
Solanum arboreum Dunal
Solanum asperum Rich.
Solanum aturense Dunal
Solanum badilloi Benitez
Solanum bicolor Roem. & Schult.
Solanum campaniforme Roem. & Schult.
Solanum capsicoides All. [introduced]
Solanum caripense Dunal
Solanum circinatum Bohs
Solanum cordovense Sesse & Moc.
Solanum coriaceum Dunal
Solanum costatum M. Nee
Solanum crinitum Lam.
Solanum davidsei Carvalho
Solanum erythrotrichum Femald
Solanum extensum Bitter
Solanum fulvidum Bitter
Solanum hirtum Vahl
Solanum jamaicense Mill.
Solanum lanceifolium Jacq.
Solanum lepidotum Dunal
Solanum leucocarpon Dunal
Solanum mammosum L.
Solanum maroniense Poit.
Solanum monachophyllum Dunal
Solanum morii S. Knapp
SU
AM
AM GU su FG?
FG
FG?
AM BO GU
BO DA GU
GU
FG
BO
AM
BO GU su FG
GU su FG
AM BO DA
BO DA GU su FG
AM BO FG?
AM
AM BO GU
BO GU
GU su
FG
AM
GU
GU
BO GU su FG
GU su FG
AM BO
BO
BO
BO
GU
GU?
GU
GU
su FG
FG
BO DA GU su FG
BO DA
GU FG?
AM BO GU su FG
BO GU su FG
FG
AM BO DA GU
GU
su FG
FG
BO
BO GU su?
AM BO DA GU su FG
BO GU su FG
AM BO DA GU su FG
GU su FG
BO
AM BO GU su FG
AM BO GU su FG
BO GU su FG
AM BO DA
GU
FG
FG
FG
AM BO su FG
AM BO DA GU su FG
AM BO DA GU su FG
AM BO
BO
BO
GU FG
FG
FG
BO GU su FG
528 Checklist of the Plants of the Guiana Shield
Solanum nuricum M. Nee
Solarium oocarpum Sendtn.
Solanum oppositifolium Ruiz & Pav.
Solanum paludosumMoric
.
Solanum pensile Sendtn.
Solanum rubiginosum Vahl
Solanum rufistellatum Steyerm.
Solanum rugosum Dunal
Solanum schlechtendalianum Walp.
Solanum schomburgkii Sendtn.
Solanum seaforthianum Andrews
Solanum semotum M. Nee
Solanum sendtnerianum Van Heurck & Miill. Arg.
Solanum sessiliflorum Dunal [probably cultivated or escaped from cultivation]
Solanum stramoniifolium Jacq.
Solanum subinerme Jacq.
Solanum swartzianum Roem. & Schult.
Solanum tepuiense S. Knapp
Solanum torvum Sw.
Solanum tricuspidatum Rich, ex Dunal
Solanum umbratile I.M. Johnst.
Solanum vanheurckii Mtill. Arg.
Solanum velutinum Dunal
Solanum vestissimum Dunal AM
Witheringia solanacea L'Her. AM FG
Synonyms and Misidentifications
Bassovia macrophylla (Dunal) J. Coulter =Witheringia solanacea
Bassovia sylvatica Aubl. =Solanum anceps
Brachistus macrophyllus (Dunal) Miers =Witheringia solanacea
Bnmfelsia maritima Benth. [not in flora area] =Brunfelsia latifolia (Pohl) Benth.
Brunfelsia spruceana Benth. =Brunfelsia martiana
Capsicum conoide Mill. ? =Capsicum annuum
Capsicum curvipes Dunal ? ^Capsicum annuum
Capsicum hispidum var. glabruiusculum Dunal =Capsicum annuum var. glabriusculum
Capsicum macrophyllum (Dunal) Standi. =Witheringia solanacea
Capsicum solanaceum var. macrophylla (Dunal) Kuntze =Witheringia solanacea
Cestrum acuminatissimum Dunal =Cestrum schlechtendahlii
Cestrum alternifolium var. mitanthum O.E. Schulz =Cestrum alternifolium
Cestrum baenitzii Lingelsh. =Cestrum schlechtendahlii
Cestrum grande Pittier =Cestrum racemosum
Cestrum latifolium var. tenuiflorum (Kunth) O.E. Schulz =Cestrum latifolium
Cestrum megalophyllum Dunal =Cestrum schlechtendahlii
Cestrum oliganthum Dunal =Cestrum latifolium
Cestrum oliganthum var. latifolium Dunal =Cestrum latifolium
Cestrum prieurei Dunal =Cestrum latifolium
Cestrum racemosum var. grande (Pittier) Francey ^Cestrum racemosum
Cestrum silvaticum Francey [not in flora area] =Cestrum microcalyx Francey
Cestrum tenuiflorum Kunth =Cestrum latifolium
Cestrum tenuiflorum var. glabrescens C.V. Morton =Cestrum glabrescens
Cyphomandra allophylla (Miers) Hemsl. =Solanum allophyllum
Cyphomandra bolivarensis Steyerm. =Cyphomandra tobagensis
Daturafastuosa L. =Datura metel
Ectozoma ulei Dammer ^Hawkesiophyton ulei
Franciscea uniflora Pohl =Brunfelsia uniflora
Hawkesiophyton panamense (Standi.) Hunz. =Hawkesiophyton ulei
Lycianthes guianensis (Dunal) Bitter =Lycianthes pauciflora
Markea camponoti Ducke =Markea longiflora
Markea panamensis Standi. ^Hawkesiophyton ulei
Markea porphyrobaphes Sandwith =Markea sessiliflora
Physalis capsicifolia Dunal [name of uncertain application] ^Physalis sp.
Physalis hirsuta Dunal [name of uncertain application] =Physalis sp.
Physalis orinocensis Kunth =Deprea orinocensis
Physalis surinamensis Miq. ^Physalis pubescens
Checklist of the Plants of the Guiana Shield 529
Physalis swinamensis auct. non Miq.: sensu Steyerm. =Physalis pubescens
Piouandra allophylla Miers =Solaniim allophyllum
Saracha solanacea (L'Her.) Miers =Witheringia solanacea
Schwenckia americana var. angustifolia J.A. Schmidt =Schwenckia americana
Schwenckia americana var. hirta (Klotzsch ex Benth.) Carvalho =Schwenckia americana
Schwenckia giiianensis Benth. =Schwenckia americana
Schwenckia guineensis Schumach. & Thonn. =Schwenckia americana
Schwenckia hirta Klotzsch ex Benth. =Schwenckia americana
Schwenckia patens Kunth =Schwenckia glabrata
Schwenckia ulei Dammer ^Schwenckia grandiflora
Sicklera tetrandra A. Braun & Bouche =Witheringia solanacea
Solanum altissimum Pittier ex. Benitez [nomen nudum] =Solanum altissimum
Solanum americanum var. nodiflorum (Jacq.) Edmonds =Solanum americanum
Solanum antillarum O.E. Schulz =Solanum nudum
Solanum arcuatum Sendtn. =Solanum acerifolium
Solanum arecunarum Pittier et al. [nomen nudum] ^'Solanum sessiliflorum
Solanum asperrimum Bitter & O. Moritz =Solanum aturense
Solanum aubletii Pulle =Solanum anceps
Solanum boissieri Dunal =Solanum schomburgkii
Solanum bolivarense Pittier et al. [nomen nudum] =Solanum monachophyllum
Solanum brachyacanthum Dunal =Solanum lanceifolium
Solanum brachybotryon Dunal =Solanum velutinum
Solanum callicarpifolium Kunth & Bouche =Solanum bicolor
Solanum confusum Pittier et al. [nomen nudum] =Solanum velutinum
Solanum cuneifolium Dunal =Solanum jamaicense
Solanum cyananthum Dunal =Solanum crinitum
Solanum cyananthum war. jubatum (Roem. & Schult.) Dunal =Solanum crinitum
Solanum cyrrhosum Dunal =Solanum seaforthianum
Solanum demerarense Dunal =Solanum stramoniifolium
Solanum demerarense var. inerme Dunal =Solanum stramoniifolium
Solanum diphyllum L. [not in flora area]
Solanum dolichostylum O.E. Schulz =Solanum arboreum
Solanum donnellsmithii J. Coulter =Solanum adhaerens
Solanum echinocarpum Bredem. ex Roem. & Schult. =Solanum hirtum
Solanum enoplocalyx Dunal =Solanum adhaerens
Solanum flavescens Dunal =Solanum hirtum
Solanum flavescens var. albescens Dunal =Solanum hirtum
Solanum foetidum Ruiz & Pav. [not in flora area]
Solanumformosum Dunal =Solanum crinitum
Solanum fraternorum Pittier et al. [nomen nudum] =Solanum velutinum
Solanum geminifolium Schltdl. & Cham. =Solanum schlechtendalianum
Solanum globiferum Dunal =Solanum mammosum
Solanum granelianum D'Arcy =Solanum pensile
Solanum guianense Klotzsch [nomen nudum]
Solanum guianense Dunal =Lycianthes pauciflora
Solanum hastatum Sendtn. =Solanum acerifolium
Solanum heterophyllum Lam. =Solanum subinerme
Solanum hostmannii Dunal =Solanum schomburgkii
Solanum humboldtii Dunal =Solanum adhaerens
Solanum hyporrhodium Pittier et al. [nomen nudum] =Solanum vestissimum
Solanum intermedium Sendtn. [not in flora area]
Solanum ipomoea Sendtn. =Solanum pensile
Solanumjapurense Dunal =Lycianthes sp.
Solanumjavitense Kunth [nom. superfl.] =Solanum monachophyllum
Solanumjubatum Roem. & Schult. -Solanum crinitum
SolanumJuripeba Rich. =Solanum subinerme
Solanum laetum Miq. =Solanum pensile
Solanum lanceifolium var. brachyacanthum (Dunal) O.E. Schulz =Solanum lanceifolium
Solanum lanugiflorum Pittier et al. [nomen nudum] ^Solanum velutinum
Solanum longiflorum Vahl =Solanum subinerme
Solanum macranthum Dunal =Solanum crinitum
Solanum manaense Sagot ex Lemee [nomen nudum]
Solanum mesopliarthrum Pittier et al. [nomen nudum] =Solanum velutinum
Solanum mesopliarthrum Benitez & Steyerm. =Solanum velutinum
530 Checklist of the Plants of the Guiana Shield
Solarium miquelii C.V. Morton =Solanum pensile
Solarium molle Dunal =Solanum monachophyllum
Solarium nigrum L. [misident. for S. americanum fide Nee]
Solanum nodiflorum Jacq. =Solanum americanum
Solanum nudum Dunal [not in flora area]
Solanum obscurum herb. Willd. =Solanum lanceifolium
Solanum obtusifrons Pittier et al. [nomen nudum] =Solanum hirtum
Solanum ochraceo-ferrugineum (Dunal) Femald [not in flora area]
Solanum oleraceum Dunal ex Poir. =Solanum americanum
Solanum orenocense Dunal =Solanum hirtum
Solanum orientale Benitez =Solanum oocarpum
Solanum orinocense Dunal =Solanum stramoniifolium
Solanum pauciflorum Vahl =Lycianthes pauciflora
Solanum platanifolium Hook. =Solanum mammosum
Solanum platyphyllum Dunal =Solanum stramoniifolium
Solanum prehensile Pittier et al. [nomen nudum] =Solanum pensile
Solanum proteanthum Bohs =Cyphomandra oblongifolia
Solanum prunifolium Roem. & Schult. =Solanum seaforthianum
Solanum pterocaulon Dunal =Solanum americanum
Solanum puberuloba Steyerm. =Solanum oppositifolium
Solanum quinquangulare Roem. & Schult. =Solanum acerifolium
Solanum radula Vahl ^Solanum asperum
Solanum rhamphidacanthum Bitter =Solanum velutinum
Solanum richardii (Dunal) Lemee ? =Solanum anceps
Solanum ripivagum Pittier et al. [nomen nudum] =Solanum arboreum
Solanum sacupanense Rusby =Solanum monachophyllum
Solanum salignum Roem. & Schult. =Solanum seaforthianum
Solanum salviifolium auct. =Solanum asperum
Solanum salzmannii Dunal =Solanum paludosum
Solanum scabridum Dunal =Solanum vestissimum
Solanum scandens L. f. =Solanum pensile
Solanum scandens L. =Solanum pensile
Solanum secundum Bitter & O. Moritz =Solanum aturense
Solanum sempervirens Dunal =Solanum pensile
Solanum siparunoides Ewan ^Solanum aturense
Solanum stamineum Steud. =Solanum subinerme
Solanum stramoniifolium var. inerme (Dunal) Whalen =Solanum stramoniifolium
Solanum styracifolium Roem. & Schult. =Solanum subinerme
Solanum styracoides Rusby =Solanum pensile
Solanum surinamense Steud. =Solanum leucocarpon
Solanum sylvaticum (Aubl.) Bitter =Solanum anceps
Solanum tegore Aubl. =Cyphomandra tegore
Solanum timotense Pittier et al. [nomen nudum] =Solanum hirtum
Solanum tobagense (Sandwith) Bohs =Cyphomandra tobagensis
Solanum topirito Pittier et al. [nomen nudum] =Solanum lanceifolium
Solanum topiro Dunal =Solanum sessiliflorum
Solanum toxicarium Rich. =Solanum stramoniifolium
Solanum trichocarpum Miq. =Solanum stramoniifolium
Solanum triste var. crassipes Dunal =Solanum leucocarpon
Solanum umbrosum Dunal =Solanum lanceifolium
Solanum undecimangulare Roem. & Schult. =Solanum stramoniifolium
Solanum verbascifolium L. [not in flora area]
Witheringia macrophylla Dunal = Witheringia solanacea
References
Bemardello, L.M. & A.T. Hunziker. 1987. A synoptical revision of Solandra (Solanaceae). Nord. J. Bot. 7: 639-652.
Bitter, G. 1919. Die Gattung Lycianthes. Ahb. Naturwiss. Vereine Bremen 24(2): 292-520.
Bohs, L. 1994. Cyphomandra (Solanaceae). Flora Neotropica Monograph 63: 1-175.
Carvalho, L. d'A Freire de. 1978. O genero Schwenckia D. van Rooyen ex Linnaeus no Brasil - Solanaceae. Rodriguesia 44:
307-524.
Cuatrecasas, J. 1958. Notes on American Solanaceae. Feddes Repert. 61(1): 74-86.
D'Arcy, W.G. 1973. Solanaceae. In R.E. Woodson, Jr., R.W. Schery, & collaborators, Flora ofPanama. Ann. Missouri Bot. Gard.
60: 573-780.
D'Arcy, W.G.
, C. Benitez de Rojas, & M.H. Nee. 2005. Solanaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K.
Hoist, Flora of the Venezuelan Guayana 9: 194-246.
Checklist of the Plants of the Guiana Shield 531
Edmonds, J.M. 1972. A synopsis of the taxonomy of Solanum sect. Solanum (Maurella) in South America. Kew Bull. 27: 95-
114.
Francey, P. 1935-1936. Monographic du genre Cestrum L. Candollea 6: 46-398, 7: 1-132.
Hawkes, J.G., R.N. Lester, M. Nee, & N. Estrada-R. 1991. Solanaceae III. Taxonomy, Chemistry, Evolution. Royal Botanic
Gardens, Kew.
Nee, M. 2002. Solanaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 689-
700.
Plowman, T. 1979. The genus Brunfelsia: a conspectus of the taxonomy and biogeography. Pp. 475-491 in J.G. Hawkes, R.N.
Lester, & A.D. Skelding, The Biology and Taxonomy of the Solanaceae. Linnaean Society of London, Academic Press.
Whalen, M.D. 1984. Conspectus of species groups in Solanum subgenus Leptostemonum. Gentes Herb. 12(4): 179-282.
Whalen, M.D., D.E. Costich, & C.B. Heiser. 1981. Taxonomy of Solanum section Lasiocarpa. Gentes Herb. 12(2): 41-129.
Sphenocleaceae
B.A. Stein
Sphenoclea zeylanica Gaertn. [naturalized?] AM BO DA GU SU FG
References
Stein, B.A. 2005. Sphenocleaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
9: 247-248.
Sterculiaceae
P.E. Berry, C.L. Cristobal, L.J. Dorr, J.G. Saunders
Ayenia magna L. BO
Ayenia praeclara Sandwith
Ayenia tomentosa L.
Byttneria aristeguietae Cristobal
Byttneria aurantiaca Mildbr.
Byttneria caripensis Cristobal
Byttneria catalpifolia Jacq.
Byttneria catalpifolia subsp. catalpifolia
Byttneria cordifolia Sagot
Byttneria divaricata Benth.
Byttneria divaricata var. divaricata
Byttneria genistella Triana & Planch.
Byttneria morii L.C. Bamett & Dorr
Byttneria obliqua Benth.
Byttneria piresii Cristobal
Byttneria rhamnifolia Benth.
Byttneria scabra L.
Byttneria uaupensis Spruce ex K. Schum.
Guazuma ulmifolia Lam.
Guazuma ulmifolia var. tomentella K. Schum.
Guazuma ulmifolia var. ulmifolia
Helieteres baruensis Jacq.
Helicteres brevispira A. St.-Hil.
Helicteres guazumifolia Kunth
Helicteres heptandra L.B. Sm.
Helicteres pentandra L.
Herrania camargoana R.E. Schult.
Herrania kanukuensis R.E. Schult.
Herrania lemniscata (M.R. Schomb.) R.E. Schult.
Melochia arenosa Benth.
Melochia graminifolia A. St.-Hil.
Melochia lanceolata Benth.
Melochia manducata C. Wright
Melochia melissifolia Benth.
Melochia nodiflora Sw.
Melochia parvifolia Kunth
Melochia parvifolia var.fasciculata (Benth.) Hassl.
Melochia parvifolia var. parvifolia
GU SU
GU SU
AM BO
BO
FG
AM GU
AM GU
SU FG
DA GU SU FG
DA GU SU
AM BO GU
GU
FG
FG
AM
AM
AM BO
AM BO
BO
GU
GU
SU FG
AM BO DA GU SU FG
AM BO DA
AM BO
BO GU SU
AM GU
AM BO
BO
GU
BO GU? SU FG
AM
GU SU FG
BO DA GU SU FG
AM BO DA GU SU
AM BO GU
GU SU FG?
DA
GU SU FG
BO
AM BO DA GU
GU
SU?
SU?
AM BO DA GU
532 Checklist of the Plants of the Guiana Shield
Melochia pyramidata L.
Melochia pyramidata var. pyramidata
Melochia simplex A. St.-Hil.
Melochia tomentosa L.
Melochia tomentosa var. frutescens DC.
Melochia tomentosa var. tomentosa
Melochia ulmifolia Benth.
Melochia villosa (Mill.) Fawc. & Rendle
Melochia villosa var. tomentosa (K. Schum.) Goldberg
Melochia villosa var. villosa
Sterculia abbreviata E.L. Taylor ex Mondragon
Sterculia amazonica E.L. Taylor ex Mondragon
Sterculia apetala (Jacq.) H. Karst
Sterculiafrondosa Rich.
Sterculia guianensis Sandwith
Sterculia kayae RE. Berry
Sterculia lisae E.L Taylor, sp. nov. ined.
Sterculia multiovula E.L Taylor, sp. nov. ined.
Sterculia pruriens (Aubl.) K. Schum.
Sterculia pruriens var. glabrescens E.L. Taylor, var. nov. ined.
Sterculia pruriens var. pruriens
Sterculia rugosa R. Br.
Sterculia speciosa K. Schum.
Sterculia villifera Steud.
Theobroma bicolor Bonpl.
Theobroma cacao L.
Theobroma cacao subsp. sphaerocarpum (A. Chev.) Cuatrec.
Theobroma grandiflorum (Willd. ex Spreng.) K. Schum.
Theobroma microcarpum Mart.
Theobroma obovatum Klotzsch ex Bernoulli
Theobroma subincanum Mart.
Theobroma velutinum Benoist
Uladendron codesuri Marc.-Berti
Waltheria albicans Turcz.
Waltheria berteroi (Spreng.) J.G. Saunders
Waltheria carmensarae J.G. Saunders
Waltheria collina K. Schum.
Waltheria indica L.
Waltheria involucrata Benth.
Waltheria microphylla Cav. [considered part of Waltheria indica complex by
L.J. Dorr]
Waltheria operculata Rose
Waltheria paniculata Benth.
Waltheria viscosissima A. St.-Hil.
FG?
FG?
GU su
BO su FG?
BO
BO
BO GU su FG
AM BO
BO
DA GU su FG
AM BO DA
AM DA
AM BO
BO DA
AM
AM
AM
AM
AM
AM
AM
AM
BO
BO
BO
BO
BO
AM BO
DA
DA
GU
BO
AM BO GU
AM BO
BO
BO
AM BO DA GU
AM BO
BO
DA GU
GU
GU
AM BO GU
FG
SU FG
FG
GU FG
GU SU FG
GU
GU SU FG
GU
GU FG
GU? su FG
GU su FG
GU su FG
GU
GU
GU su FG
su FG
su FG
Synonyms and Misidentifications
Bubroma grandiflorum Willd. ex Spreng. =Theobroma grandiflorum
Buttneria scabra Aubl. =Byttneria scabra
Byttneria sparganiocarpa Uittien =Byttneria divaricata
Byttneria venezuelensis Steyerm. =Byttneria divaricata var. divaricata
Cacao guianensis Aubl. =Theobroma subincanum
Cacao sylvestris Aubl. =Theobroma subincanum
Helicteres apetala Jacq. =Sterculia apetala
Helicteres brachypetala Leane =Helicteres heptandra
Helicteres bracteosa Mart. =Helicteres pentandra
Helicteres pronifolia Rich. =Helicteres pentandra
Herrania mariae (Mart.) Decne. ex Goudot [not in flora area]
Ivira pruriens Aubl. =Sterculia pruriens
Lightia lemniscata M.R. Schomb. =Herrania lemniscata
Malachra berterii Spreng. =Waltheria berteroi
Melochia benthamii K. Schum. =Melochia ulmifolia
Melochia carpinifolia J.C. Wendl. =Melochia nodiflora
Melochia carthaginensis Willd. ex K. Schum. =Melochia ulmifolia
Melochia cinerea A. St.-Hil. & Naudin =Melochia arenosa
Checklist of the Plants of the Guiana Shield 533
Melochia concinna Miq. =Melochia melissifolia
Melochia conglobata Sesse & Moc. =Melochia nodiflora
Melochiafasciculata Benth. =Melochia parvifolia yar. fasciculata
Melochia glandulifera Standi. ^Melochia manducata
Melochia hirsuta Cav. =Melochia villosa
Melochia hirsuta var. tomentosa K. Schum. =^Melochia villosa var. tomentosa
Melochia llanorum Cuatrec. =Melochia arenosa
Melochia obovata Urb. =Melochia parvifolia var. parvifolia
Melochia ramuliflora R.E. Fr. =Melochia iilmifolia
Melochia scordiifolia Turcz. =Melochia parvifolia var. parvifolia
Melochia spicata (L.) Fryxell [not in flora area, or = M villosa]
Qiiamoclit rodriguesii J.I. Falcao =Melochia graminifolia
Riedlea dichotoma Turcz. =Melochia ulmifolia
Riedlea heterotricha Turcz. =Melochia villosa
Riedlea littoralis K. Schum. [nom. superfl.] =Melochia arenosa
Riedlea littoralis Poepp. & Endl. =Melochia arenosa
Riedlea ramuliflora Miq. =Melochia ulmifolia
Riedlea urticifolia Turcz. =Melochia nodiflora
Sida villosa Mill. =Melochia villosa
Sitella involucrata (Benth.) L.H. Bailey =Waltheria involucrata
Sterculia excelsa Mart, [not in flora area]
Sterculia parviflora (Ducke) E.L. Taylor [nom. illeg.] =Sterculia kayae
Sterculia pruriens var. parviflora Ducke =Sterculia kayae
Theobroma alba Bernoulli (Chrysobalanaceae) =Licania alba
Theobroma guianense (Aubl.) J.F. Gmel. =Theobroma subincanum
Theobroma mariae (Mart.) K. Schum. [not in flora area] =Herrania mariae (Mart.) Decne. ex Goud.
Theobroma speciosum Mart, [not in flora area]
Theobroma sphaerocarpa A. Chev. =Theobroma cacao subsp. sphaerocarpum
Theobroma sylvestre Mart., non Cacao sylvestris Aubl. [not in flora area]
Theobroma sylvestris (Aubl.) G. Don =Theobroma subincanum
Turnera valleana Standi. & L.O. Williams =Waltheria operculata
Waltheria americana L. =Waltheria indica
Waltheria hirsuta C.Presl ^Waltheria viscosissima
Waltheria machrisiantha L.B. Sm. =Waltheria viscosissima
Waltheria subcordata Standi. =Waltheria berteroi
Waltheria tubiflora Klotzsch = Waltheria viscosissima
Xylosterculia riigosa (R. Br.) Kosterm. =Sterculia rugosa
References
Bamett, L.C. & L.J. Dorr. 1990. Anew, arborescent species of Byttneria from French Guiana. Brittonia 42(4): 271-275.
Cristobal, C.L. 1976. Estudio taxonomico del genero Byttneria Loefl. Bonplandia 4: 1-428.
Cristobal, C.L., J.G Saunders, & RE. Berry. 2005. Sterculiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist,
Flora of the Venezuelan Guayana 9: 248-281.
Cuatrecasas, J. 1964. Cacao and its allies. Ataxonomic revision of the genus Theobroma. Contr. U.S. Nat. Herb. 35: 379-614.
Dorr, L.J. 2002. Sterculiaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. HofT, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 700-
706.
Goldberg, A. 1967. The genus Melochia L. Contrib. United States National Herbarium 34(5): 191-363.
Saunders, J.G. 1995. Systematics and evolution of Waltheria. Dissertation, University of Texas, Austin.
Uittien, H. 1932. Sterculiaceae. In A. Pulle, Flora of Suriname 3(1 ): 34-48.
Uittien, H. 1941. Sterculiaceae (additions and corrections). In A. Pulle, Flora of Suriname 3(1): 437.
Styracaceae
RW. Fritsch, D. Sabalier, J.A. Stcyermarkt
Styrax argenteus C. Presl
Styn-ax argenteus var. ramirezii (Greenm.) Gonsoulin
Styrax camporus Pohl
Styrax discolor M.F. Silva
Styrax duidae Steyerm.
Styrax duidae subsp. duidae
Styrax duidae subsp. paruae Maguire
Styrax glaber Sw.
Styrax glabratus Schott
AM
AM
AM
AM
BO
GU?
GU su
su
FG
FG
FG
FG
534 Checklist of the Plants of the Guiana Shield
Styrax guaiquinimae (Maguire & Steyerm.) P.W. Fritsch
Styrax guanayanus Maguire & K.D. Phelps
Styrax guyanens is A. DC.
Styrax hypochryseus Perkins
Styrax lancifolius Klotzsch ex Seub.
Styrax longifolius Standi.
Styrax longipedicellatiis Steyerm.
Styrax neblinae (Maguire) P.W. Fritsch
Styrax oblongiis (Ruiz & Pav.) A. DC.
Styrax paUidus A. DC.
Styrax sieberi Perkins
Styrax sipapoanus Maguire
Styrax tafelbergensis Maguire
Styrax tessmannii Perkins
Styrax wurdackiorum Steyerm.
Styrax yutajensis (Maguire) P.W. Fritsch .
AM
AM
AM
AM
AM
AM
BO
BO
BO GU
GU
GU
GU
SU
su
FG
FG
FG
FG
GU? su
GU
BO
Synonyms and Misidentifications
Strigilia glabrata (Schott) Miers =Styrax glabratiis
Strigilia guyanensis (A. DC.) Miers =Styrax guyanensis
Strigilia psilophylla (A. DC.) Miers =Styrax glabratiis
Styrax costanus Steyerm. =Styrax glaber
Styrax duidae subsp. neblinae Maguire =Styrax neblinae
Styraxfanshawei Sandwith =Styrax glaber
Styrax filii Maguire =Styrax longipedicellatiis
Styrax glaber var. micranthus Perkins =Styrax glaber
Styrax guanayanus var. yutajensis Maguire =Styrax yutajensis
Styrax guyanensis \ar.japurensis Seub. =Styrax guyanensis
Styraxjaiiaens is Steyerm. =Styrax guanayanus
Styrax laiiraceiis Perkins =Styrax glabratus
Styrax micrasterus Perkins =Styrax glaber
Styrax occidentalis Sw. ex Thunb. =Styrax glaber
Styrax psilophyllus A. DC. =Styrax glabratus
Styrax roraimae Perkins =Styrax glaber
Styrax squamulosus M.F. Silva ^Styrax glabratus
Styrax subleprosum Klotzsch [nomen nudum] =Styrax glaber
Styrax tepuiensis Steyerm. & Maguire =Styrax glaber
Styrax tepuiensis subsp. guaiquinimae Maguire & Steyerm. =Styrax guaiquinimae
Styrax tepuiensis subsp. huachamacarii Maguire & Steyerm. =Styrax longipedicellatus
References
Fritsch, P.W., S.A. Mori, and J.L. Brown. 2002. Styracaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald,
M. Hoff, & J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New
York Bot. Gard. 76 (2): 706-708.
P.W. Fritsch & J.A. Steyermark. 2005. Styracaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 283-289.
Maguire, B. & Yung-Chau Huang. 1978. Styracaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part 10.
Mem. New York Bot. Gard. 29: 204-223.
Perkins, J. 1 907. Styracaceae. In A. Engler, Das Pflanzenreich IV(24 1 ): 1 - 11 1
.
SURIANACEAE
Suriana maritima L. FG?
References
Howard, R.A. 1988. Surianaceae. In R.A. Howard, Flora of the Lesser Antilles 4(1): 574-575.
SVMPLOCACEAE
RE. Berry, D. Sabatier, J.A. Steyermarkj
Symplocos acananensis Steyerm.
Symplocos bolivarana Aristeg.
Symplocos chimantensis Steyerm. & Maguire
BO
BO
BO
Checklist of the Plants of the Guiana Shield 535
AM GU su FG
AM BO
BO GU FG
AM
AM
AM
GU FG
AM BO
BO
BO
GU
GU
AM
Symplocos guianensis (Aubl.) Giirke
Symplocosjauaemis Steyerm. & Maguire
Symplocos martinicensis Jacq.
Symplocos neblinae Maguire & Steyerm.
Symplocos nitens (Pohl) Benth.
Symplocos nitens van claussenii Brand
Symplocos pilosiuscida Brand
Symplocos pycnophylla Sleumer [Mt. Roraima]
Symplocos schomburgkii Klotzsch ex Brand [Mt. Roraima]
Symplocos ulei Brand
Symplocos yapacanensis Steyerm.
Synonyms and Misidentifications
Ciponima guianensis Aubl. =Symplocos guianensis
Stemmatosiphon nitens Pohl =Symplocos nitens
Symplocos roraimensis Steyerm. =Symplocos pycnophylla
References
Brand, A. 1901. Symplocaceae. In A. Engler, Das Pflanzenreich IV(242): 1-100.
Maguire, B. & Yung-Chau Huang. 1978. Symplocaceae. In B. Maguire & collaborators, Botany ofthe Guayana Highland. Mem.
New York Bot. Gard. 29: 223-230.
Mori, S.A. and J.L. Brown. 2002. Symplocaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, &
J.D. Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot.
Gard. 76 (2): 708-709.
Steyermark, J.A. & RE. Berry. 2005. Symplocaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 290-295.
Tepuianthaceae
J.S. Miller
Tepuianthus auyantepuiensis Maguire & Steyerm.
Tepuianthus sarisahnamensis Maguire & Steyerm.
Tepuianthus sarisarinamensis subsp. duidensis Maguire & Steyerm.
Tepuianthus sarisahnamensis subsp. sarisarinamensis
Tepuianthus savannensis Maguire & Steyerm.
Tepuianthus yapacanensis Maguire & Steyerm.
AM
AM
AM
AM
AM
BO
BO
BO
References
Maguire, B. & J.A. Steyermark. 1981. Tepuianthaceae. In B. Maguire & collaborators, Botany of the Guayana Highland part XI.
Mem. New York Bot. Gard. 32: 1-391.
Miller, J.S. 2005. Tepuianthaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
9: 297-299.
Ternstroemiaceae
RE. Berry, A.L. Weitzman
Freziera calophylla Triana & Planch.
Freziera carinata A.L. Weitzman
Freziera roraimensis Tul.
Ternstroemia brevistyla Kobuski
Ternstroemia browniana Kobuski [expected in BO]
Ternstroemia campinicola B.M. Boom
Ternstroemia candolleana Wawra
Ternstroemia carinata A.L. Weitzman [expected in GU]
Ternstroemia crass ifolia Benth.
Ternstroemia dehiscens Huber
Ternstroemia delicatula Choisy [expected in BO]
Ternstroemia dentata (Aubl.) Sw. [expected in BO]
Ternstroemia discoidea Gleason
Ternstroemia distyla Kobuski
Ternstroemia duidae Gleason
Ternstroemia dura Gleason
Ternstroemia gleasoniana Kobuski [expected in BO]
Ternstroemia grandiosa Kobuski [expected in BO]
AM
AM BO
BO
BO
GU
GU SU FG?
AM
AM
BO
GU
GU
SU
FG?
GU SU FG
GU SU FG
AM
AM
AM
AM
BO GU
GU
GU
536 Checklist of the Plants of the Guiana Shield
Ternstroemia guanchezii B.M. Boom
Ternstroemia laevigata Wawra
Ternstroemia longipes Klotzsch ex Wawra [expected in BO]
Ternstroemia maguirei B.M. Boom
Ternstroemia punctata (Aubl.) Sw.
Ternstroemia pungens Gleason
Ternstroemia retusifolia Kobuski
Ternstroemia roraimae Klotsch [Mt. Roraima]
Ternstroemia schombiirgkiana Benth. [expected in BO]
Ternstroemia sp. A
Ternstroemia sp. B
Ternstroemia tristyla Gleason
Ternstroemia verticillata Klotzsch ex Wawra
AM
BO GU
GU
AM
BO GU SU
AM BO
BO
GU?
SU
GU SU
AM
AM
AM
FG
BO GU SU
Synonyms and Misidentifications
Eroteiim calophyllum (Triana & Planch.) Kuntze =Freziera calophylla
Eroteum guianense (Klotzsch ex Wawra) Kuntze =Freziera roraimensis
Eiirya calophylla (Triana & Planch.) Szyszyl. =Freziera calophylla
Eurya guianensis (Klotzsch ex Wawra) Szyszyl. =Freziera roraimensis
Eurya roraimensis (Tul.) Szyszyl. =Freziera roraimensis
Freziera cuatrecasasii Kobuski =Freziera calophylla
Freziera esmeraldana Little =Freziera calophylla
Freziera guianensis Klotzsch ex Wawra =Freziera roraimensis
Freziera hieronymi Kobuski =Freziera calophylla
Lettsomia guianensis Klotzsch ex M.R. Schomb. [nomen nudum] =Freziera roraimensis
Taonabo dentata Aubl. =Ternstroemia dentata
Taonabo punctata Aubl. =Ternstroemia punctata
Ternstroemia candolleana var. rotundata Wawra =Ternstroemia candoUeana
Ternstroemia cayennensis Planch, [nomen nudum]
Ternstroemia circumscissilis Kobuski [not in flora area]
Ternstroemia corymbosa Sm. [name of uncertain application, not Ternstroemia]
Ternstroemia duidae f. latifolia Kobuski =Ternstroemia duidae
Ternstroemia monosperma Gleason =Ternstroemia discoidea
Ternstroemia paucifolia Gleason =Ternstroemia duidae
Ternstroemia revoluta Splitg. ^Ternstroemia punctata
References
Berry, RE. & A.L. Weitzman. 2005. Temstroemiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 300-308.
Jansen-Jacobs, M.J. & J.C. Lindeman. 1979. Theaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 356-366.
Kobuski, C.E. 1942. Studies on the Theaceae, XII. Notes on the South American species of Ternstroemia. J. Am. Arb. 23: 298-
343.
Weitzman, A.L., S. Dressier & P.F. Stevens. 2004. Temstroemiaceae. Families and Genera of Vascular Plants 6: 450-460.
Tetrameristaceae
RE. Berry, J.S. Miller
Pentamerista neotropica Maguire AM
References
Miller, J.S. & RE. Berry. 2005. Tetrameristaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 309-310.
Theaceae
A.L. Weitzman
Gordoniafruticosa (Schrad.) H. Keng AM BO GU SU FG
Synonyms and Misidentifications
Gordonia semiserrata (Nees) Spreng. =Gordoniafruticosa
Haemocharis caracasana Linden & Planch. =Gordoniafruticosa
Haemocharis pubescens (Planch. & Linden) Linden & Planch. =Gordoniafruticosa
Laplacea camelliaefolia Triana & Planch. =Gordoniafruticosa
Laplacea caracasana Klotzsch & H. Karst. ex Wawra [nom. illeg.] =Gordoniafruticosa
Laplaceafruticosa (Schrad.) Kobuski ^Gordoniafruticosa
Checklist of the Plants of the Guiana Shield 537
Laplacea fruticosa var. chimantae Steyerm. =Gordoniafruticosa
Laplacea pubescens Planch. & Linden ex Triana & Planch. =Gordoniafruticosa
Laplacea pubescens var. camelliaefolia (Triana & Planch.) Kobuski =Gordoniafruticosa
Laplacea pubescens var. minor Steyerm. =Gordoniafruticosa
Laplacea semiserrata (Nees) Cambess. =Gordoniafruticosa
Lindleya ?fruticosa Nees =Gordoniafruticosa
Lindleya semiserrata Nees [nomen nudum] =Gordomafruticosa
Wikstroemia camelliaefolia (Triana & Planch.) S.F. Blake =Gordomafruticosa
Wikstroemia fruticosa Schrad. =Gordoniafruticosa
References
(see also Bonnetiaceae)
Jansen-Jacobs, M.J. & J.C. Lindeman. 1979. Theaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 356-366.
Kobuski, C.E. 1942. Studies on the Theaceae, XII. Notes on the South American species of Temstroemia. J. Am. Arb. 23: 298-
343.
Weitzman, A.L. Theaceae. 2005. Theaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora ofthe Venezuelan
Guayana 9: 311-313.
Theophrastaceae
J.C. Lindeman, J.S. Miller, B. Stahl
AM BO GU su FG
AM BO GU su FG
GU su FG
GU
Clavija imatacae B. Stahl BO DA GU
Clavija lancifolia Desf
Clavija lancifolia subsp. chermontiana (Standi.) B. Stahl
Clavija lancifolia subsp. lancifolia
Clavija macrophylla (Link ex Roem. & Schult.) Miq.
Jacquinia armillaris Jacq. BO
Synonyms and Misidentifications
Clavija chermontiana Standi. =Clavija lancifolia subsp. chermontiana
Clavija lancifolia auct. non Desf. 1832: sensu Mez =Clavija imatacae
Jacquinia arborea auct. non Vahl: sensu Kunth =Jacquinia armillaris
Jacquinia revoluta Jacq. =Jacquima armillaris
References
Lindeman, J.C. 1979. Theophrastaceae. In A.L. Stoffers &. J.C. Lindeman, Flora of Suriname 5(1): 367-369.
Miller, J.S. & B. Stahl. 2005. Theophrastaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 325-329.
Stahl, B. 1991. A revision of Clavija. Opera Bot. 107: 1-77.
Stahl, B. 2002. Theophrastaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell,
eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2):
709-711.
Thymelaeaceae
P.E. Berry, Z.S. Rogers, J.A. Steyermarkt
Daphnopsis americana (Mill.) J.R. Johnst.
Daphnopsis americana subsp. caribaea (Griseb.) Nevling
Daphnopsis dircoides Steyerm.
Daphnopsis granitica Pruski & Barringer
Daphnopsis guaiquinimae Steyerm.
Daphnopsis longifolia Taub.
Daphnopsis longipedunculata Gilg ex Ule & Domke [=Funifera sp.?] [Ml.
Roraima]
Daphnopsis nevlingiana Steyerm.
Daphnopsis steyermarkii Nevling
Goodallia guianensis Benth.
Lasiadenia ottohuberi Plowman & Nevling
Lasiadenia rupestris Benth.
Lophostoma amoenum Nevling
Schoenobiblus amazonicus Steyerm.
Schoenobiblus daphnoides Mart.
Schoenobiblus panamensis Standi. & L.O. Williams
BO
BO
BO
BO
BO
BO
AM BO
AM
AM BO
AM
AM
FG
GU?
GU
GU
GU
GU
FG?
FG
538 Checklist of the Plants of the Guiana Shield
Synonyms and Misidentifications
Daphnopsis caribea Griseb. =Daphnopsis amehcana subsp. caribaea
Laurus americanus Mill. =Daphnopsis americana
References
Domke, W. 1934. Untersuchungen iiber die systematische und geographische Gliederung der Thymelaeaceen. Bibliotheca Botanica
111: 1-151.
Kawasaki, M.L. 2002. Thymelaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 711-712.
Nevling, L.I. 1959. A revision of the genus Daphnopsis. Ann. Missouri Bot. Gard. 46: 257-358.
Rogers, Z.S., RE. Berry, & J.A. Steyermark. 2005. Thymelaeaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K.
Hoist, Flora of the Venezuelan Guayana 9: 332-342.
TiLIACEAE
•L.J. Dorr, M.J. Jansen-Jacobs, W. Meijert
Apeiba albiflora Ducke AM BO DA
Apeiba glabra Aubl.
Apeiba intermedia Uittien
Apeiba macropetala Ducke
Apeiba membranacea Spruce ex Benth.
Apeiba petoiimo Auhl. BO DA
Apeiba schombiirgkii Szyszyl.
Apeiba tibourbou Aubl. AM BO DA
Apeiba uittienii Jans.-Jac. & Westra
Christiana africana DC.
Christiana mennegae (Jans.-Jac. & Westra) Kubitzki
Corchorus aestuans L.
Corchorus hirtus L.
Corchorus orinocensis Kunth
Corchorus siliquosus L.
Luehea alternifolia (Mill.) Mabb.
Luehea Candida (DC.) Mart.
Luehea cymulosa Spruce ex Benth.
Luehea paniculata Mart. & Zucc. SU
Lueheopsis althaeiflora (Spruce ex Benth.) Burret
Lueheopsis duckeana Burret
Lueheopsis rosea (Ducke) Burret
Lueheopsis rugosa (Pulle) Burret
Mollia glabrescens Benth.
Mollia globularis Meijer
Mollia grandiflora Meijer
Mollia lepidota Spruce ex Benth.
Mollia lepidota subsp. casiquiarensis (Baehni) Meijer
Mollia lepidota subsp. sphaerocarpa (Gleason ex Record) Meijer
Mollia sp. A
Mollia speciosa Mart. AM GU
Mollia tomentosa Spruce ex Benth.
Mollia ulei Burret
Triumfetta althaeoides Lam.
Triumfetta bogotensis DC.
Triumfetta lappula L.
Triumfetta rhomboidea Jacq.
Triumfetta semitriloba Jacq.
Vasivaea alchorneoides Baill.
Synonyms and Misidentifications
Alegria Candida DC. =Luehea Candida
Apeiba aspera Aubl. =Apeiba glabra
Apeiba burchellii Sprague =Apeiba glabra
Apeiba echinata Gaertn. =Apeiba petoumo
Apeiba echinata var. macropetala (Ducke) Ducke =Apeiba macropetala
Apeiba hypoleuca Steud. =Apeiba petoumo
Apeiba surinamensis Uittien =Apeiba schomburgkii
AM
AM BO
AM BO
AM BO
AM BO
BO
BO
BO
AM BO
BO
BO
AM
AM
BO
AM
AM BO
AM
AM
AM
AM
AM
AM BO
BO
BO
BO
BO
AM BO
GU SU FG
SU FG
SU
GU
GU
GU SU FG
GU SU
GU SU FG
GU
SU
GU SU FG
GU SU
GU
GU
GU SU FG
GU SU FG
GU SU FG
GU
GU
GU
GU
GU
GU SU FG
SU FG
GU FG
GU SU?
GU
Checklist of the Plants of the Guiana Shield 539
Apeiba tiboiirboii var. krukofii Uittien =Apeiba albiflora
Apeiba tibonrbou var. membranacea Lockh. ex Griseb. =Apeiba schomburgkii
Apeiba tibourbou var. nigosa Szyszyl. =Apeiba albiflora
Asterophorum mennegae Jans.-Jac. & Westra ^Christiana mennegae
Bartramia indica L. =Triumfetta rhomboidea
Carpodiptera ? schomburgkii Baill. =Christiana africana
Cedrus alternifolia Mill. =Luehea alternifolia
Christiana cordifolia Hook. f. "^Christiana africana
Christiana madagascariensis Baill. =Christiana africana
Corchorus acutangulus Lam. =Corchorus aestuans
Corchorus argutus Kunth =Corchorus orinocensis
Corchorus argutus var. benthamii K. Schum. =Corchorus orinocensis
Corchorus hirtus var. orinocensis (Kunth) K. Schum. =Corchorus orinocensis
Corchorus pilolobus Link =Corchorus hirtus
Guenetia macrosperma Sagot (Bombacaceae) =Catostemmafragrans
Luehea althaeiflora Spruce ex Benth. =Lueheopsis althaeiflora
Lueheaferruginea Turcz. =Luehea alternifolia
Lueheaflavescens Uittien =Lueheopsis rosea
Luehea maroniensis Benoist =Lueheopsis rugosa
Luehea nobilis Linden & Planch. =Luehea Candida
Luehea rosea Ducke =Lueheopsis rosea
Luehea speciosa Willd. =Luehea alternifolia
Luehea tessrnannii Burret =Luehea cymulosa
Lueheopsis flavescens (Uittien) Burret =Lueheopsis rosea
Lueheopsis violacea Standi. =Lueheopsis duckeana
Mollia lepidota var. casiquiarensis Baehni =Mollia lepidota subsp. casiquiarensis
Mollia scabra Meijer =Mollia ulei
Mollia speciosa var. alata Meijer [nomen nudum] =Mollia sp. A
Mollia sphaerocarpa Gleason ex Record =Mollia lepidota subsp. sphaerocarpa
Mollia stellaris Meijer =Mollia ulei
Speirostyla tiliifolia Baker =Christiana africana
Triumfetta bartramia L. [nom. illeg.] =Triumfetta rhomboidea
Triumfetta bartramia sensu Lay, non L. =Triumfetta rhomboidea
Triumfetta bolivariensis D.E. Gonzalez [nomen nudum] =Triumfetta bogotensis
Triumfetta hispida A. Rich. =Triumfetta bogotensis
Triumfetta hostmannii Miq. =Triumfetta lappula
Triumfetta macrophylla Vahl =Triumfetta althaeoides
Triumfetta rubricaulis Kunth =Triumfetta semitriloba
Triumfetta sinuosa Miq. =Triumfetta lappula
Triumfetta surinamensis Steud. =Triumfetta althaeoides
Triumfetta valenciensis R. Knuth [nomen nudum] =Triumfetta bogotensis
References
Dorr, L.J. 2002. Tiliaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. HolT, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 712-
716.
Dorr, L.J. & W. Meijer. 2005. Tiliaceae. In J.A. Steyermark, PE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 9: 343-362.
Jansen-Jacobs, M.J. 1986. Tiliaceae (additions and corrections). In A.L. Stoffers & J.C. Lindeman, Flora ofSurinamc 3(1-2):
293-300.
Jansen-Jacobs, M.J. & W. Meijer (in press). Tiliaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A).
Jansen-Jacobs, M.J. & L.Y.Th. Westra. 1 995. A new species ofApeiba (Tiliaceae) from the Venezuelan-Brazilian border. Brittonia
47(3): 335-339.
Ko Ko Lay 1950. The American species of Triumfetta L. Ann. Missouri Bot. Gard. 37(3): 315-395.
Kubitzki, K. 1995. Asterophorum and Tahitia congeneric with Christiana (Tiliaceae). Bot. Jahrb. Syst. 116: 537-542.
Meijer, W. 1978. Mollia (Tiliaceae). A revision of the South American species. In B. Maguire & collaborators. Botany of the
Guayana Highland - Part X. Mem. New York Bot. Gard. 29: 68-82.
Uittien, H. 1 932. Tiliaceae. In A. Pulle, Flora of Suriname 3( 1 ): 49-57.
Uittien, H. 1941. Tiliaceae (additions and corrections). In A. Pulle, Flora of Suriname 3(1): 438-441.
Trigoniaceae
P.E. Berry, E. Lleras-Perez
Trigonia candelabra Lleras SU
540 Checklist of the Plants of the Guiana Shield
GU su FG
GU FG
BO DA GU su FG
AM BO GU
AM BO
BO
GU
GU
AM BO
AM
GU
BO GU
GU
su FG
FG?
BO GU su FG
Trigonia coppenamensis Stafleu SU
Trigonia ehrendorferi Lleras BO
Trigonia hypoleuca Griseb.
Trigonia laevis Aubl.
Trigonia microcarpa Sagot ex Warm.
Trigonia nivea Cambess.
Trigonia nivea var. nivea
Trigonia reticulata Lleras
Trigonia sericea Kunth
Trigonia spruceana Benth. ex Warm.
Trigonia subcymosa Benth.
Trigonia villosa Aubl.
Trigonia villosa var. macrocarpa (Benth.) Lleras
Trigonia villosa var. villosa
Synonyms and Misidentifications
Trigonia hypoleuca var. pubescens Warm. =Trigonia hypoleuca
Trigonia kaieteurensis Maguire =Trigonia microcarpa
Trigonia laevis var. microcarpa (Sagot ex Warm.) Sagot =Trigonia microcarpa
Trigonia macrocarpa Benth. =Trigonia villosa var. macrocarpa
Trigonia villosa var. cuneata DC. =Trigonia villosa var. villosa
Trigonia villosa var. oblonga DC. =Trigonia villosa var. villosa
Trigonia villosa var. obtusata DC. ^Trigonia villosa var. villosa
Trigonia xanthopila Garcke =Trigonia hypoleuca
References
Berry, P.E. 2005. Trigoniaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
9: 364-368.
Lleras, E. 1978. Trigoniaceae. Flora Neotropica Monogr. 19: 1-73.
Lleras, E. 1998. Trigoniaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 21: 49-60.
Mori, S.A. 2002. Trigoniaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 716-
718.
Stafleu, F.A. 1951. Trigoniaceae. In A. PuUe, Flora of Suriname 3(2): 173-177.
I
Tropaeolaceae
RE. Berry, L.J. Raechal
Tropaeolum orinocense P.E. Berry AM
References
Bayer, C. & O. Appel. 2003. Families and Genera of Vascular Plants. 5: 400^404.
Berry, P.E. & L.J. Raechal. Tropaeolaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 9: 372.
TURNERACEAE
C. Feuillet, M. Hoff, M.-M. Arbo
Piriqueta cistoides (L.) Griseb.
Piriqueta cistoides subsp. caroliniana (Walter) Arbo
Piriqueta cistoides subsp. cistoides
Piriqueta guianensis N.E. Br.
Piriqueta guianensis subsp. guianensis
Piriqueta undulata Urb.
Piriqueta viscosa Griseb.
Piriqueta viscosa subsp. tovarensis Urb.
Piriqueta viscosa subsp. viscosa
Turnera acuta Willd. ex Schult.
Turnera annectens Arbo
Turnera argentea Arbo
Turnera aurantiaca Benth.
Turnera brasiliensis Willd. ex Roem. & Schult.
Turnera breviflora Moura
AM BO
AM BO
AM BO
AM BO
AM BO
BO
AM BO
AM BO
AM
AM
GU SU FG
GU
GU
GU
GU
GU
SU FG
SU FG
SU FG
BO
GU SU FG
GU su FG
GU
Checklist of the Plants of the Guiana Shield 541
Turnera caerulea M09. & Sesse ex DC. GU su
Turnera caerulea var. suhnamensis (Urb.) Arbo & A. Fernandez GU su
Turnera castilloi Arbo AM
Turnera cicatricosa Arbo AM BO GU
Turnera aurassavica Urb. GU?
Turnera glaziovii Urb. GU? su FG
Turnera guianensis Aubl. AM BO GU su FG
Turnera huberi Arbo AM
Turnera lineata Urb. AM BO GU
Turnera macrophylla Urb. AM
Turnera odorata Rich. AM BO GU su FG
Turnera paruana Arbo AM
Turnera pumilea L. BO GU FG
Turnera rupestris Aubl. GU su FG
Turnera rupestris vax.frutescens (Aubl.) Urb. GU su FG
Turnera rupestris var. rupestris GU su FG
Turnera scabra Millsp. AM BO GU FG?
Turnera schomburgkiana Urb. BO GU
Turnera steyermarkii Arbo AM
Turnera subulata Sm. GU su FG
Turnera velutina Benth. GU
Turnera venosa Urb. AM
Turnera waltherioides Urb. BO GU
Wormskioldia lobata Urb. [cultivated?] FG
Synonyms and Misidentifications
Piriqueta caroUniana (Walter) Urb. =Piriqueta cistoides subsp. caroliniana
Piriqueta caroliniana var. integrifolia Urb. =Piriqueta cistoides subsp. caroliniana
Piriqueta cistoides var. bracteolata Urb. =Piriqueta cistoides subsp. cistoides
Piriqueta cistoides var. foliosa (Garcke) Urb. =Piriqueta cistoides subsp. cistoides
Piriqueta cistoides var. genuina Urb. =Piriqueta cistoides subsp. cistoides
Piriqueta cistoides var. latifolia Urb. =Piriqueta cistoides subsp. cistoides
Piriquetafoliosa Garcke =Piriqueta cistoides subsp. cistoides
Piriqueta lanceolata Benth. =Piriqueta cistoides subsp. cistoides
Piriqueta stenophylla Klotzsch =Piriqueta cistoides subsp. cistoides
Piriqueta tomentosa Kunth =Piriqueta cistoides subsp. cistoides
Piriqueta villosa Aubl. =Piriqueta cistoides subsp. cistoides
Turnera aspera Poir. =Piriqueta cistoides subsp. cistoides
Turnera aturensis D. Dietr. =Piriqueta cistoides subsp. cistoides
Turnera benthamiana R.H. Schomb. =Turnera velutina
Turnera carpinifolia Kunth =Turnera acuta
Turnera chamaedrys Klotzsch [excluded taxon]
Turnera cistoides L. =Piriqueta cistoides
Turnera corchorifolia Willd. ex Schult. =Turnera odorata
Turnera corchoroides Klotzsch [excluded taxon]
Turnera desvauxii DC. =Piriqueta cistoides subsp. cistoides
Turnerafrutescens Aubl. =Turnera rupestris var.frutescens
Turnera grandifolia Bremek. =Turnera glaziovii
Turnera hexandra Spreng. =Turnera odorata
Turnera hirta Desv., non Willd. =Piriqueta cistoides subsp. cistoides
Turnera humboldtii Spreng. =Turnera guianensis
Turnera muricata Rusby ^Turnera macrophylla
Turnera opifera Mart. =Turnera caerulea var. surinamensis
Turnera parviflora Benth. =Turnera odorata
Turnera pilosula Urb. =Turnera acuta
Turnera refracta Klotzsch ^Turnera caerulea var. surinamensis
Turnera subglabra Klotzsch =Turnera aurantiaca
Turnera tomentosa Willd. ex Schult. =Piriqueta cistoides subsp. cistoides
Turnera tomentosa Kunth =Turnera odorata
Turnera ulmifolia var. elegans (Otto) Urb. =Turnera subulata
Turnera ulmifolia var. intermedia Urb. =Turnera scabra
Turnera ulmifolia f. serissima Killip =Turnera lineata
Turnera ulmifolia var. surinamensis Urb. =Turnera caerulea var. surinamensis
Waltheria caroliniana Walter =Piriqueta cistoides subsp. caroliniana
542 Checklist of the Plants of the Guiana Shield
References
Arbo, M.M. 1990. Turaeraceae: novedades para la Guayana Venezolana. Ann. Missouri Bot. Gard. 77: 340-352.
Arbo, M.M. 1995. Turneraceae - Parte I. Piriqueta. Flora Neotropica Monogr. 67: 1-156.
Berry, P.E. & L.J. Raechal. 2005. Turneraceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 373-384.
Bremekamp, E. 1941. Turneraceae. In A. Pulle, Flora of Suriname 3(1): 342-354.
Hoff, M. 2002. Turneraceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 718-
719.
Lemee, A. 1953. Flore de la Guyane fran9aise 3: 82. Paul Lechevalier, Paris.
Pulle, A. 1906. An enumeration of the Vascular Plants known from Surinam, together with their distribution and Synonymy.
Leiden.
Urban, I. 1 883a. Monographic des Familie der Tumeraceen. Jahrbuch des Koniglichen Botanischen Gartens und des botanischen
Museums zu Berlin, 2: 1-152.
Urban, I. 1883b. Turneraceae. In C.F.R Martins, Flora Brasiliensis 13(3): 88-170, tab. 31-48.
Ulmaceae
C.C. Berg, RE. Berry, J.S. Miller
AM BO GU SU FG
AM BO DA GU
GU
SU FG
AM GU SU
AM BO DA GU SU FG
Ampelocera edentula Kuhlm.
Celtis ignanaea (Jacq.) Sarg.
Celtis schippii Standi.
Trema integerrima (Beurl.) Standi.
Trema micrantha (L.) Blume
Synonyms and Misidentifications
Rhamnus iguanaeus Jacq. =CeUis igiianaea
Rhamnus micranthus L. =Trema micrantha
Sponia integerrima Beurl. =Trema integerrima
Sponia micrantha (L.) Decne. =Trema micrantha
References
Berg, C.C. 1992. Ulmaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 1 1 : 3-9.
Berg, C.C. 2002. Ulmaceae. In Mori, S.A., G Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 720-
722.
Miller, J.S. & RE. Berry. 2005. Ulmaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 9: 386-390.
Ooststroom, S.J. van. 1932. Ulmaceae. In A. Pulle, Flora of Suriname 1(1): 47-48.
Urticaceae
C.C. Berg, A. Pool
Boehmeria cylindrica (L.) Sw.
Boehmeria ramiflora Jacq.
Laportea aestuans (L.) Chew
Phenax sonneratii (Poir.) Wedd. [naturalized?]
Pilea daiiciodora Pav. ex Wedd.
Pilea fendleri Killip
Pilea imparifolia Wedd.
Pilea involucrata (Sims) Urb.
Pilea microphylla (L.) Liebm.
Pilea pubescens Liebm.
Pilea tabularis C.C. Berg
Pouzolzia occidentalis (Liebm.) Wedd.
Urera baccifera (L.) Gaudich. ex Wedd.
Urera caracasana (Jacq.) Gaudich. ex Griseb.
Urera laciniata Goudot ex Wedd.
SU FG
SU FG
GU SU FG
FG
AM
DA
GU SU FG
BO DA
BO GU SU FG
BO GU SU FG
FG
BO
BO GU SU FG
BO GU SU
SU
FG
FG
Synonyms and Misidentifications
Fleurya aestuans (L.) Gaudich. ex Miq. =Laportea aestuans
Leucococcus occidentalis Liebm. =Pouzolzia occidentalis
Parietaria microphylla L. =Pilea microphylla
Parietaria sonneratii Poir. =Phenax sonneratii
Checklist of the Plants of the Guiana Shield 543
Phenax vulgaris Wedd. =Phenax sonneratii
Pilea chrysosplenioides Wedd. =Pilea involucrata
Pilea dauciodora var. crenata Wedd. =Pileafendleri
Pilea dendrophila Miq. [not in flora area]
Pilea guyanensis Wedd. =Pilea pubescens
Pilea leptophylla Killip =Pilea fendleri
Pilea pubescens var. guyanensis (Wedd.) Wedd. =Pilea pubescens
Pilea pubescens var. involucrata (Sims) Wedd. =Pilea involucrata
Urtica aestuans L. =Laportea aestuans
Urtica alceifolia Poir. =Urera caracasana
Urtica baccifera L. =Urera baccifera
Urtica caracasana Jacq. ^Urera caracasana
Urtica cylindrica L. =Boehmeria cylindrica
Urtica involucrata Sims =Pilea involucrata
References
Berg, C.C. 1992. Urticaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 11: 125-139.
Berg, C.C. 2002. Urticaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V. Heald, M. Hoff, & J.D. Mitchell, eds.
2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard. 76 (2): 722-
725.
Pool, A. 2005. Urticaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 9:
391-402.
Rooij, M.J.M. de. 1975. Urticaceae. In J. Lanjouw & A.L. Stoffers, Flora of Suriname 5(1): 300-318.
Valerianaceae
N.X. de Enrich
Valeriana scandens L. AM BO
Valeriana scandens var. candolleana (Gardner) C.A. Miill. [Mt. Roraima] AM BO
Valeriana scandens var. subcordata C.A. Miill. BO
Synonyms and Misidentifications
Valeriana alpina Veil. =Valeriana scandens var. candolleana
Valeriana candolleana Gardner =Valeriana scandens var. candolleana
Valeriana phaseoli Braun =Valeriana scandens
Valeriana scandens var. dentata C.A. Miill. =Valeriana scandens var. candolleana
References
Xena de Enrech, N. 1992. Valerianaceae. In G Morillo, Flora de Venezuela 5(1): 221-267.
Xena de Enrech, N. 2005. Valerianaceae. In J.A. Steyermark, P.E. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 9: 402-404.
Verbenaceae
G.A. Aymard C, M.J. Jansen-Jacobs
Aegiphila bracteolosa Moldenke
Aegiphila deppeana Steud.
Aegiphila elata Sw.
Aegiphila elata var. elata
Aegiphila elata var. macrophylla (Kunth) Lopez-Pal.
Aegiphila filipes Mart. & Schauer ex Schauer
Aegiphila glandulifera Moldenke
Aegiphila glandulifera var. pyramidata Moldenke
Aegiphila gleasonii Moldenke [insufficiently known taxon]
Aegiphila hoehnei Moldenke
Aegiphila hoehnei var. venezuelensis Moldenke
Aegiphila hystricina Aymard & Cuello
Aegiphila integrifolia (Jacq.) B.D. Jacks.
Aegiphila intermedia Moldenke
Aegiphila laevis (Aubl.) J.F. Gmel.
Aegiphila laxiflora Benth.
Aegiphila lewisiana Moldenke
Aegiphila Ihotzkiana Cham.
Aegiphila longifolia Turcz.
AM
AM BO
BO
AM
AM BO
AM BO
AM BO
AM
AM
AM BO
AM BO
AM
AM BO
BO
BO
DA
GU
GU
GU
GU
GU
GU
GU
GU
FG
SU
SU
SU
SU
SU
FG
FG
FG
BO DA GU
AM BO GU
AM BO
AM BO
AM BO
AM BO
BO
AM
DA
BO GU
AM BO GU
AM
AM BO GU
AM BO GU
AM BO GU
GU FG
FG?
FG?
AM
A IV/f
BO DA GU su FG
AM
DA GU su FG
DA GU su FG
BO
BO
DA GU su FG
BO DA
AM
BO DA GU su FG
BO
544 Checklist of the Plants of the Guiana Shield
Aegiphila macramha Ducke SU FG
Aegiphila membranacea Turcz. SU FG
Aegiphila mollis Kunth
Aegiphila mollis var. intermedia Moldenke
Aegiphila mollis var. mollis
Aegiphila mollis var. surfaceana (Moldenke) Moldenke
Aegiphila parviflora Moldenke
Aegiphila pendula Moldenke
Aegiphila perplexa Moldenke
Aegiphila racemosa Veil. SU FG
Aegiphila roraimensis Moldenke
Aegiphila sprnceana Moldenke
Aegiphila venezuelensis Moldenke
Aegiphila venezuelensis var. serrata Moldenke
Aegiphila venezuelensis var. venezuelensis
Aegiphila villosa (Aubl.) J.F. Gmel.
Aegiphila vitelliniflora Klotzsch ex Walp.
Aegiphila vitelliniflora var. egleri Moldenke [insufficiently known taxon]
Amasonia campestris (Aubl.) Moldenke
Amasonia obovata Gleason
Avicennia germinans (L.) L.
Avicennia schaueriana Stapf & Leechm. ex Moldenke
Bouchea prismatica (L.) Kuntze
Bouchea prismatica var. longirostra Grenzeb.
Citharexyliim macrophyllum Poir.
Citharexylum poeppigii Walp.
Citharexylum sp. A
Citharexylum spinosum L.
Citharexylum venezuelense Moldenke
Clerodendrum aculeatum (L.) Schltdl.
Clerodendrum indicum (L.) Kuntze [cultivated, naturalized]
Clerodendrumjaponicum (Thunb.) Sweet [cultivated, naturalized]
Clerodendrum philippinum Schsiwev {widely c\x\i\\2iXQd'\ AM BO
Clerodendrum ternifolium Kunth BO
Clerodendrum thomsonae Balf. [cultivated, naturalized] AM BO DA
Cornutia pubescens C.F. Gaertn.
Cornutia pyramidata L.
Duranta erecta L. AM BO DA
Duranta obtusifolia Kunth BO
Duranta sp. A BO
Duranta sp. B AM
Lantana achyranthifolia Desf. AM BO
Lantana brasiliensis Link [expected in GU]
Lantana camaraL. AM BO DA GU SU FG
Lantana canescens Kunth BO DA GU
Lantana cujabensis Schauer AM BO
Lantanafucata Lindl. BO
Lantana glutinosa Poepp. BO DA
Lantana radula Sw. BO GU
Lantana ruiz-teranii Lopez-Pal. & Steyerm. BO
Lantana trifolia L. AM BO GU SU FG
Lippia alba {Mill) "N.E.Br. AM BO DA GU SU FG
Lippia aristata Schauer GU
Lippia micromera Schauer AM BO DA GU
Lippia origanoides Kunth BO GU
Petrea blanchetiana Schauer AM BO FG
Petrea bracteata Sieud. AM BO GU SU FG
Petrea macrostachya Benth. AM GU SU FG
Petrea pubescens Turcz. AM BO
Petrea sulphurea Jans.-Jac. FG
Petrea volubilis L. AM BO DA GU SU FG
Phyla betulifolia (Kunth) GreenQ AM BO DA GU SU FG
Priva lappulacea(L.) ?ers. BO DA GU SU FG
Stachytarpheta angustifolia(Mi\\.) Wahl BO GU SU FG
GU SU FG
GU SU FG
GU SU
GU SU FG
GU su
su
FG
FG
GU su FG
AM BO DA GU su FG
AM BO
BO
DA GU su FG
BO GU su FG
AM BO GU
BO GU su FG
GU su FG
AM
AM BO GU su
AM BO GU
GU
su FG
FG
AM
AM BO GU su FG
AM BO GU su FG
AM
BO GU
su FG
AM
BO DA GU su FG
AM GU su FG
Checklist of the Plants of the Guiana Shield 545
Stachytarpheta cayennensis (Rich.) Vahl
Stachytarphetajamaicensis (L.) Vahl
Stachytarpheta lopez-palacii Moldenke
Stachytarpheta mutabilis (Jacq.) Vahl
Stachytarpheta sprucei Moldenke
Tamonea spicata Aubl.
Vitex agnus-castus L. [cultivated, naturalized]
Vitex calothyrsa Sandwith
Vitex capitata Vahl
Vitex compressa Turcz.
Vitex guianensis Moldenke
Vitex klugii Moldenke
Vitex orinocensis Kunth
Vitex orinocensis var. multiflora (Miq.) Huber
Vitex orinocensis var. orinocensis
Vitex panshiniana Moldenke
Vitex schomburgkiana Schauer
Vitex sprucei Briq.
Vitex stahelii Moldenke
Vitex triflora Vahl
Synonyms and Misidentifications
Aegiphila cowanii Moldenke =Aegiphila venezuelensis
Aegiphila guianensis Moldenke =AegiphiIa integrifoUa
Aegiphila integrifoUa var. guianensis (Moldenke) Lopez-Pal. =Aegiphila integrifoUa
Aegiphila lutea Lam. =Aegiphila laevis
Aegiphila macrophylla Kunth =Aegiphila data var. macrophylla
Aegiphila martinicensis Jacq. [not in flora area]
Aegiphila steyermarkii Moldenke =Aegiphila roraimensis
Aegiphila steyermarkii var. macrophylla Moldenke =Aegiphila venezuelensis
Aegiphila steyermarkii var. obtusifolia Moldenke =Aegiphila roraimensis
Aegiphila surfaceana Moldenke =Aegiphila mollis var. surfaceana
Amasonia arborea Kunth =Amasonia campestris
Amasonia campestris var. surinamensis Moldenke =Amasonia campestris
Amasonia erecta L. f. =Amasonia campestris
Amasonia erecta var. punicea (Vahl) Pulle =Amasonia campestris
Amasonia integerrima Spreng. (Acanthaceae) =Bravaisia integerrima
Amasonia lasiocaulos Mart. & Schauer ex Schauer =Amasonia campestris
Amasonia punicea Vahl =Amasonia campestris
Amasonia spruceana Moldenke =Amasonia campestris
Avicennia germinans (L.) Steam [comb, superfl.] =Avicennia germinans
Avicennia germinans var. cumanensis (Kunth) Moldenke =Avicennia germinans
Avicennia germinans var. guayaquilensis (Kunth) Moldenke =Avicennia germinans
Avicennia germinans f. venezuelensis Moldenke =Avicennia germinans
Avicennia marina (Forssk.) Vierh. [not in flora area]
Avicennia nitida Jacq. =Avicennia germinans
Avicennia schaueriana f. candicans Moldenke =Avicennia schaueriana
Avicennia tomentosa var. cumanensis Kunth =Avicennia germinans
Avicennia tomentosa var. guayaquilensis Kunth =Avicennia germinans
Bontia germinans L. =Avicennia germinans
Callicarpa integrifoUa Jacq. =Aegiphila integrifoUa
Citharexylum amazonicum Moldenke =Citharexylum macrophyllum
Citharexylum fruticosum L. =Citharexylum spinosum
Citharexylum fruticosum var. brittonii Moldenke ^Citharexylum spinosum
Citharexylum poeppigii f. anomalum Moldenke =Citharexylum poeppigii
Citharexylum scandens Briq. & Spruce (Bignoniaceae) =Schlegelia scandens
Clerodendrum aculeatum var. guianense Moldenke =Clerodendrum aculcatum
Clerodendrum fragrans (Vent.) Willd. =Clcrodendrum philippinum
Clerodendrum fragrans var. pleniflora Schauer =Clerodendrum philippinum
Clerodendrumfragrans var. pleniflorum Schauer =Clerodendrum philippinum
Cornutia cayennensis Schauer =Cornutia puhescens
Cryptocalyx nepetifolia Benth. ^Lippia betulifolia
Duranta coriacea Hayek =Duranta obtusifolia
Duranta plumieri var. alba Mast. =Duranta erecta
546 Checklist of the Plants of the Guiana Shield
Duranta repens L. =Duranta erecta
Duranta repens var. canescens Moldenke =Diiranta erecta
Duranta repens var. lopez-palacii Moldenke =Duranta erecta
Ghinia spicata (Aubl.) Moldenke =Tamonea spicata
Lantana aculeata L. =Lantana camara
Lantana alba Mill. =Lippia alba
Lantana armata Schauer =Lantana camara
Lantana armata var. guianensis Moldenke =Lantana camara
Lantana camara var. aculeata (L.) Moldenke =Lantana camara
Lantana camara var. mista (L.) L.H. Bailey =Lantana camara
Lantana camara var. moritziana (Otto & A. Dietr.) Lopez-Pal. =Lantana camara
Lantana geminata (Kunth) Spreng. =Lippia alba
Lantana mista L. =Lantana camara
Lantana moritziana Otto & A. Dietr. =Lantana camara
Lantana trifolia f. hirsuta Moldenke =Lantana trifolia
Lantana trifolia f. oppositifolia Moldenke =l.antana trifolia
Lantana trifolia var. rigiduscula Briq. =Lantana trifolia
Lantana trifolia f. rubierensis Moldenke =Lantana trifolia
Lippia betiilifolia Kunth =Phyla betulifolia
Lippia geminata Kunth =Lippia alba
Lippia schomburgkiana Schauer =Lippia origanoides
Lippia stoechadifolia (L.) Kunth [not in flora area] =Phyla stoechadifolia (L.) Small
Manabea arborescens Aubl. =Aegiphila integrifolia
Manabea laevis Aubl. =Aegiphila laevis
Manabea villosa Aubl. =Aegiphila villosa
Petrea algentryi Moldenke =Petrea blanchetiana
Petrea arborea Kunth =Petrea volubilis
Petrea aspera Turcz. =Petrea volubilis
Petrea glandulosa Pittier =Petrea pubescens
Petrea kohautiana C. Presl =^Petrea volubilis
Petrea martiana Schauer ^Petrea bracteata
Petrea morii Moldenke =Petrea blanchetiana
Petrea peruviana Moldenke ^Petrea blanchetiana
Petrea pubescens var. albicalyx Moldenke =Petrea pubescens
Petrea rivularis Moldenke ^Petrea volubilis
Petrea rugosa Kunth [not in flora area]
Petrea schomburgkiana Schauer =Petrea bracteata
Petrea subserrata Cham, [not in flora area] =Petrea racemosa Nees
Phyla nodiflora (L.) Greene [not in flora area]
Priva lappulacea f. albiflora Moldenke =Priva lappulacea
Stachytarpheta angustifolia var. elatior Schrad. ex Schult. =Stachytarpheta angustifolia
Stachytarpheta angustifolia f. elatior (Schrad. ex Schult.) Lopez-Pal. =Stachytarpheta angustifolia
Stachytarpheta cayennensis f. albiflora Moldenke =Stachytarpheta cayennensis
Stachytarpheta dichotoma (Ruiz & Pav.) Vahl =Stachytarpheta cayennensis
Stachytarpheta dichotoma f. albiflora (Moldenke) Moldenke =Stachytarpheta cayennensis
Stachytarpheta elatior Schrad. ex Schult. =Stachytarpheta angustifolia
Stachytarpheta elatior war. jenmanii Moldenke =Stachytarpheta angustifolia
Stachytarpheta orubica (L.) Vahl [not in flora area]
Stachytarpheta pilosiuscula Kunth =Stachytarphetajamaicensis
Stachytarpheta roraimensis Moldenke =Stachytarpheta sprucei
Stachytarpheta roraimensis var. pubescens Moldenke =Stachytarpheta sprucei
Stachytarpheta surinamensis Miq. ex Pulle [nomen nudum] ^Stachytarpheta angustifolia
Taligalea campestris Aubl. =Amasonia campestris
Verbena angustifolia Mill. =Stachytarpheta angustifolia
Verbena cayennensis Rich. =Stachytarpheta cayennensis
Verbena dichotoma Ruiz & Pav. =Stachytarpheta cayennensis
Verbenajamaicensis L. =Stachytarphetajamaicensis
Verbena lappulacea L. =Priva lappulacea
Verbena mutabilis Jacq. ^Stachytarpheta mutabilis
Verbena prismatica L. =Bouchea prismatica
Verbena suffruticosa Aubl. [not in flora area] =Phyla stoechadifolia (L.) Small
Vitex appunii Moldenke =Vitex schomburgkiana
Vitex brittoniana Moldenke =Vitex compressa
Vitex multiflora Miq. =Vitex orinocensis var. multiflora
Checklist of the Plants of the Guiana Shield 547
Vitexpacimonensis Spruce ex Sandwith =Vitex calothyrsa
Vitex spongiocarpa Ducke =Vitex sprucei
Vitex spongiocarpa var. longidentata Moldenke = Vitex sprucei
Vitex sprucei var. longidentata Moldenke =Vitex sprucei
Vitex sprucei var. vaupesensis Moldenke =Vitex sprucei
Vitex triflora f. quinquefoliolata (Moldenke) Moldenke =Vitex triflora
Vitex triflora var. quinquefoliolata Moldenke =Vitex triflora
Vitex wittrockiana Moldenke = Vitex capitata
Volkameriafragrans Vent. =Clerodendrum philippinum
References
Aymard C, G.A. 2005. Verbenaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 9: 407-445.
Jansen-Jacobs, M.J. 1988. Verbenaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 4: 1-114.
Jansen-Jacobs, M.J. 2002. Verbenaceae. In Mori, S.A., G. Cremers, C. Gracie, J.J. de Granville, S.V Heald, M. Hoff, & J.D.
Mitchell, eds. 2002. Guide to the vascular plants of central French Guiana: Part 2. Dicotyledons. Mem. New York Bot. Gard.
76 (2): 725-728.
Lopez-Palacios, S. 1977. Verbenaceae. In Flora de Venezuela.
Moldenke, H.N. 1940a. Avicenniaceae. In A. Pulle, Flora of Suriname 4(2): 322-325.
Moldenke, H.N. 1940b. Verbenaceae. In A. Pulle, Flora of Suriname 4(2): 357-321.
Moldenke, H.N. 1971. A Fifth Summary of the Verbenaceae, Avicenniaceae, Stilbaceae, Dicrostylidaceae, Symphoremaceae,
Nyctanthaceae, and Eriocaulaceae of the World. H.N. Moldenke, Wayne, N.J.
Rueda, R.M. 1994. Systematics and evolution of the genus Petrea. Ann. Missouri Bot. Gard. 81(4): 610-652.
ViOLACEAE
A. Fernandez del Valle, W.H.A. Hekking
Amphirrhox latifolia Mart, ex Eichler [=Amphirrhox longifolia s.l.]
Amphirrhox longifolia (A. St.-Hil.) Spreng.
Amphirrhox surinamensis Eichler [=Amphirrhox longifolia s.l.]
Corynostylis arborea (L.) S.F. Blake
Corynostylis carthagenensis H. Karst.
Corynostylis pubescens S. Moore
Corynostylis volubilis L.B. Sm. & A. Fernandez
Gloeospermum sphaerocarpum Triana & Planch.
Hybanthus calceolaria (L.) Schulze-Menz
Hybanthus oppositifolius (L.) Taub.
Hybanthus phyllanthoides (Planch. & Linden) L.B. Sm. & A. Fernandez
Leonia cymosa Mart.
Leonia glycycarpa Ruiz & Pav.
Noisettia orchidiflora (Rudge) Ging.
Paypayrola confertiflora Tul.
Paypayrola grandiflora Tul.
Paypayrola guianensis Aubl.
Paypayrola hulkiana Pulle
Paypayrola longifolia Tul.
Rinorea amapensis Hekking
Rinorea bahiensis (Moric.) Kuntze
Rinorea brevipes (Benth.) S.F. Blake
Rinorea camptoneura (Radlk.) Melch.
Rinorea endotricha Sandwith
Rinoreafalcata (Mart, ex Eichler) Kuntze
Rinoreaflavescens (Aubl.) Kuntze
Rinorea guianensis Aubl.
Rinorea lindeniana (Tul.) Kuntze
Rinorea lindeniana var. lindeniana
Rinorea macrocarpa (Mart, ex Eichler) Kuntze
Rinorea melanodonta S.F. Blake
Rinorea neglecta Sandwith
Rinorea ovalifolia (Britton) S.F. Blake
Rinorea paniculata (Mart.) Kuntze
Rinorea pectino-squamata Hekking
Rinorea pubiflora (Benth.) Sprague & Sandwith
Rinorea pubiflora var. grandifolia (Eichler) Hekking
GU su FG
AM BO DA GU su FG
GU su FG
AM BO
BO
DA GU su FG
AM
AM
AM GU? su FG
BO GU su FG
AM BO
BO
DA GU su FG
AM BO GU
AM BO GU su FG
GU su FG
GU su FG
AM GU FG
AM GU su
su
FG
FG
AM BO DA GU su
su
FG
FG
FG
AM GU su FG
AM
BO DA GU
AM BO su FG
AM BO GU
GU
su FG
FG
AM BO GU su
AM BO GU su
AM BO
BO
GU
GU
su FG
GU su FG
AM BO
AM BO FG
FG
AM BO DA GU su FG
AM BO DA GU su FG
548 Checklist of the Plants of the Guiana Shield
AM BO GU SU FG
AM
AM BO DA GU su FG
AM
BO SU FG
Rinorea pubiflora \ar. pubiflora
Rinorea racemosa (Mart.) Kuntze
Rinorea riana Kuntze
Rinorea sprucei (Eichler) Kuntze
Rinoreocarpus ulei (Melch.) Ducke
Synonyms and Misidentifications
AJsodeia brevipes Benth. =Rinorea brevipes
Alsodeia camptoneura Radlk. =Rinorea camptoneura
Alsodeiafalcata Mart, ex Eichler =Rinoreafalcata
Alsodeiafalcata var. grandifolia Eichler =Rinorea pubiflora var. grandifolia
Alsodeiaflavescens (Aubl.) Spreng. ^Rinoreaflavescens
Alsodeia guianensis (Aubl.) Eichler =Rinorea pubiflora var. pubiflora
Alsodeia guianensis var. brevipes (Benth.) Eichler =Rinorea brevipes
Alsodeia guianensis var. laxiflora (Benth.) Eichler =Rinorea brevipes
Alsodeia guianensis var. lindeniana (Tul.) Eichler =Rinorea lindeniana
Alsodeia guianensis var. parviflora Eich\Qr°=Rinoreafalcata
Alsodeia laxiflora Benth. =Rinorea brevipes
Alsodeia lindeniana Tul. =Rinorea lindeniana
Alsodeia longiflora Oudem. =Amphirrhox longiflora
Alsodeia macrocarpa Mart, ex Eichler ^Rinorea macrocarpa
Alsodeia martini Turcz. =Rinorea pubiflora var. pubiflora
Alsodeia ovalifolia Britton =Rinorea ovalifolia
Alsodeia pallida Klotzsch ex M.R. Schomb. [nomen nudum]
Alsodeia paniculata Mart. =Rinorea paniculata
Alsodeia piparea Spreng. (Flacourtiaceae) =Casearia commersoniana
Alsodeia prunifolia Spreng. =Rinorea riana
Alsodeia pubiflora Benth. =Rinorea pubiflora
Alsodeia racemosa Mart. =Rinorea racemosa
Alsodeia riana (DC. ex Ging.) Turcz. =Rinorea riana
Alsodeia sprucei Eichler =Rinorea sprucei
Calyptrion nitidum Benth. =Corynostylis arborea
Conohoria aubletii D. Dietr. (Flacourtiaceae) =Casearia commersoniana
Conohoria brevipes (Benth.) Miq. =Rinorea brevipes
Conohoriaflavescens Aubl. =Rinorea flavescens
Conohoria laxiflora (Benth.) Miq. =Rinorea brevipes
Conohoria passoura DC. ex Ging. =Rinorea pubiflora var. pubiflora
Conohoria riana DC. ex Ging. [nom. illeg.] =Rinorea riana
Corynostylis hybanthus (L.) Mart. =Corynostylis arborea
Gloeospermum ulei Melch. =Rinoreocarpus ulei
Hybanthus circaeoides (Kunth) Baill. =Hybanthus oppositifolius
Hybanthus ipecacuanha (L.) Baill. =Hybanthus calceolaria
lonidium calceolaria (L.) Vent. =Hybanthus calceolaria
lonidium circaeoides Kunth =Hybanthus oppositifolius
lonidium ipecacuanha (L.) Vent. =Hybanthus calceolaria
lonidium oppositifolium (L.) Roem. & Schult. =Hybanthus oppositifolius
lonidium phyllanthoides Planch. & Linden =Hybanthus phyllanthoides
Leonia melinoniana (Baill.) Baill. ex Sagot =Leonia glycycarpa
Noisettia longifolia Kunth =Noisettia orchidiflora
Passoura guianensis Aubl. =Rinorea pubiflora var. pubiflora
Paypayrola ventricosa Tul. =Paypayrola grandiflora
Riana guianensis Aubl. =Rinorea riana
Rinorea laxiflora (Benth.) Melch. =Rinorea brevipes
Rinorea martini (Turcz.) S.F. Blake =Rinorea pubiflora var. pubiflora
Rinorea passoura (DC. ex Ging.) Kuntze [nom. illeg.] =Rinorea pubiflora var. pubiflora
Rinorea passoura Kuntze =Rinorea pubiflora var pubiflora
Rinorea passoura var. andersonii Sandwith ex Hekking =Rinorea pubiflora var. grandifolia
Rinorea pulleana Melch. =Rinorea pubiflora var. grandifolia
Rinorea riana (DC. ex Ging.) Kuntze [nom. illeg.] =Rinorea riana
Rinorea surinamensis Melch. ^Rinoreafalcata
Rinoreocarpus salmoneus Ducke =Rinoreocarpus ulei
Spathularia longifolia A. St.-Hil. =Amphirrhox longifolia
Viola arborea L. =Corynostylis arborea
Viola calceolaria L. =Hybanthus calceolaria
Checklist of the Plants of the Guiana Shield 549
Viola hybanthus Aubl. =Corynostylis arborea
Viola hybanthus L. =Corynostylis arborea
Viola ipecacuanha L. =Hybanthus calceolaria
Viola itoubou Aubl. =Hybanthus calceolaria
Viola oppositifolia L. =Hybanthus oppositifolius
References
Fernandez del Valle, A. 2005. Violaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan
Guayana 9: 445-464.
Hekking, W.H.A. 1988. Violaceae part I - Rinorea and Rinoreocarpus. Flora Neotropica Monogr. 46: 1-208.
ViSCACEAE
J. Kuijt
Dendrophthora crispula (Rizzini) Kuijt [expected in GU]
Dendrophthora decipiens Kuijt [Mt. Roraima]
Dendrophthora densifrons (Ule) Kuijt [expected in the Guianas]
Dendrophthora elliptica (Gardner) Krug & Urb.
Dendrophthorafanshawei (Maguire) Kuijt
Dendrophthorafendleriana (Eichler) Kuijt
Dendrophthora intermedia (Rizzini) Kuijt
Dendrophthorajauana Rizzini
Dendrophthora karuaiana (Steyerm.) Kuijt
Dendrophthora macbridei (Standi, ex J.F. Macbr.) Kuijt
Dendrophthora microsoma Rizzini
Dendrophthora nitidula (Rizzini) Kuijt
Dendrophthora obliqua (C. Presl) Wiens
Dendrophthora oligantha Kuijt
Dendrophthora perfurcata (Rizzini) Kuijt
Dendrophthora roraimae (Oliv.) Ule
Dendrophthora tenuiflora (Steyerm. & Maguire) Kuijt
Dendrophthora tepuiana (Steyerm.) Kuijt
Phoradendron aphyllum Steyerm.
Phoradendron bathyoryctum Eichler
Phoradendron berryi Rizzini
Phoradendron bilineatum Urb.
Phoradendron chrysocladon A. Gray
Phoradendron crassifolium (Pohl ex DC.) Eichler
Phoradendron diminutivum E.A. Kellogg
Phoradendron dipterum Eichler
Phoradendron exiguum Trel.
Phoradendron granvillei Kuijt
Phoradendron haughtii Kuijt
Phoradendron herbert-smithii Trel.
Phoradendron hexastichum (DC.) Griseb.
Phoradendron inaequidentatum Rusby
Phoradendron kelloggii Kuijt
Phoradendron krameri Kuijt
Phoradendron longipetiolatum Urb. [expected in GU]
Phoradendron mairaryense Ule
Phoradendron membranifolium Kuijt
Phoradendron microstachyum Kuijt
Phoradendron morsicatum Rizzini
Phoradendron mucronatum (DC.) Krug & Urb.
Phoradendron nervosum Oliv. [expected in flora area]
Phoradendron northropiae Urb.
Phoradendron obtusissimum (Miq.) Eichler
Phoradendron pellucidulum Eichler
Phoradendron perrottetii (DC.) Eichler
Phoradendron piperoides (Kunth) Trel.
Phoradendron platycaulon Eichler
Phoradendron poeppigii (Tiegh.) Kuijt
Phoradendron pteroneuron Eichler
Phoradendron pulleanum E.H.L. Krause
AM BO
AM BO
BO
GU
GU
AM BO GU
GU su?
AM BO
AM BO
AM BO
BO
AM
BO
BO
AM BO
BO
GU
AM BO
BO
GU
AM BO
BO
GU
GU
su
AM DA
AM
BO
GU
GU
su FG
AM BO GU su FG
AM
BO GU su
BO
FG
AM
BO DA GU
AM BO GU
AM BO DA GU FG
AM
BO
GU su
AM BO
BO
GU
AM
BO GU
AM BO GU
AM GU
AM BO GU su
AM BO GU
AM BO GU su FG
AM BO DA GU su FG
AM
GU
BO GU su
AM GU su FG
550 Checklist of the Plants of the Guiana Shield
Phoradendron quadrangulare (Kunth) Griseb.
Phoradendron racemositm (Aubl.) Krug & Urb.
Phoradendron scariosum Rizzini
Phoradendron schultesii Kuijt
Phoradendron steyermarkii Rizzini
Phoradendron strongyloclados Eichler
Phoradendron triflorum E.A. Kellogg
Phoradendron trinervium (Lam.) Griseb.
Phoradendron tunaeforme (DC.) Eichler
Phoradendron undulatum (Pohl ex DC.) Eichler
Synonyms and Misidentifications
Dendrophthora basiandra Kuijt =Dendrophthora macbridei
Dendrophthora elliptica var. stenophylla Krug & Urb. =Dendrophthora elliptica
Dendrophthora hylaeana Rizzini =Dendrophthora elliptica
Dendrophthora inaequidentata (Rusby) Trel. =Phoradendron inaequidentatum
Dendrophthora lacryma-jobi E.A. Kellogg -Dendrophthora crispula
Dendrophthora parvispicata Rizzini =Dendrophthora microsoma
Dendrophthora rubicunda Ule =Dendrophthora elliptica
Loranthus piperoides Kunth =Phoradendron piperoides
Loranthus quadrangulare Kunth =Phoradendron quadrangulare
Phoradendron acinacifolium Mart, ex Eichler =Phoradendron obtusissimum
Phoradendron acinacifolium var. surinamense Rizzini =Phoradendron obtusissimum
Phoradendron apertum Trel. =Phoradendron trinervium
Phoradendron appunii Trel. =Phoradendron trinervium
Phoradendron ayliffei Philcox =Phoradendron undulatum
Phoradendron baileyae Trel. =Phoradendron trinervium
Phoradendron baileyae var. exile (Rizzini) Rizzini =Phoradendron trinervium
Phoradendron bolivianum Trel. =Phoradendron hexastichum
Phoradendron caerulesens Trel. =Phoradendron mucronatum
Phoradendron caracasanum Urb. =Phoradendron mucronatum
Phoradendron caracasanum var. parvifolium Rizzini =Phoradendron mucronatum
Phoradendron carinatum Trel. ^Phoradendron bilineatum
Phoradendron cayennense Eichler =Phoradendron racemosum
Phoradendron chaguaramasarum Trel. =Phoradendron mucronatum
Phoradendron chrysocarpum var. stehlei Trel. & Stehle =Phoradendron undulatum
Phoradendron crenato-callosum Rizzini =Phoradendron herbert-smithii
Phoradendron crispulum Rizzini =Dendrophthora crispula
Phoradendron demerarae Trel. =Phoradendron dipterum
Phoradendron dendrophthora Rizzini =Dendrophthora elliptica
Phoradendron densifrons Ule =Dendrophthora densifrons
Phoradendron dimidiatum (Miq.) Eichler ^Phoradendron perrottetii
Phoradendron duidanum Trel. =Dendrophthorafendleriana
Phoradendron duidanum var. hymenaeifolium Rizzini =Dendrophthora obliqua
Phoradendron dunstervillorum Rizzini =Phoradendron perrottetii
Phoradendron ellipticum (Gardner) Eichler ^Dendrophthora elliptica
Phoradendron emarginatum Mart, ex Eichler =Phoradendron mucronatum
Phoradendron emarginatum var. truncato-retusum Rizzini =Phoradendron mucronatum
Phoradendron essequibense Trel. =Phoradendron strongyloclados
Phoradendron excisum Rizzini =Phoradendron northropiae
Phoradendron exile Rizzini =Phoradendron trinervium
Phoradendron fanshawei Maguire =Dendrophthorafanshawei
Phoradendronfendlerianum Eichler =Dendrophthorafendleriana
Phoradendron flavens (Sw.) Griseb. =Phoradendron racemosum
Phoradendronflavens var. longispicum Rizzini =Phoradendron chrysocladon
Phoradendron glauco-lutescens Rizzini =Phoradendron piperoides
Phoradendron granaticola Trel. =Phoradendron nervosum
Phoradendron granaticola var. taeniicaule (Rizzini) Rizzini =Phoradendron nervosum
Phoradendron guascanum Trel. =Dendrophthora obliqua
Phoradendron harmsianum Ule =Phoradendron perrottetii
Phoradendron herminieri Trel. =Phoradendron undulatum
Phoradendron huricola Trel. =Phoradendron quadrangulare
Phoradendron ichythystoma Rizzini =Phoradendron obtusissimum
Phoradendron insigne Steyerm. =Phoradendron pellucidulum
AM BO DA GU
AM BO
BO
DA GU
AM
BO
AM BO GU
AM BO
AM BO
BO
GU
AM BO
su
su
su
FG
FG
FG
Checklist of the Plants of the Guiana Shield 551
Phoradendron intermedium Rizzini =Dendrophthora intermedia
Phoradendronjenmanii Trel. =Phoradendron inaequidentatum
Phoradendron johnstonii Trel. =Phoradendron strongyloclados
Phoradendron knoopii Warb. ex Trel. =Phoradendron chrysocladon
Phoradendron leptarthrum Rizzini =Phoradendron herbert-smithii
Phoradendron linguiforme Rizzini =Phoradendron mairaryense
Phoradendron longiarticulatum Rizzini =Phoradendron dipterum
Phoradendron lyonii Trel. =Phoradendron mucronatum
Phoradendron macbridei Standi, ex J.F. Macbr. =Dendrophthora macbridei
Phoradendron martinicensis (DC.) Griseb. =Phoradendron crassifolium
Phoradendron nitidulum Rizzini =Dendrophthora nitidula
Phoradendron obliquum (C. Presl) Eichler =Dendrophthora obliqua
Phoradendron ottonis Eichler =Phoradendron mucronatum
Phoradendron ottonis var. oblongifolium Rizzini =Phoradendron mucronatum
Phoradendron pachyphyllum Trel. =Phoradendron chrysocladon
Phoradendron pennivenium (DC.) Eichler (Loranthaceae) =Oryctanthus florulentus
Phoradendron perfurcatum Rizzini =Dendrophthora perfurcata
Phoradendron prancei Rizzini =Phoradendron inaequidentatum
Phoradendron pseudomucronatum Rizzini ^Phoradendron pellucidulum
Phoradendron ptarianum Steyerm. =Phoradendron longipetiolatum
Phoradendron quadrangulare (Kunth) Krug & Urb. =Phoradendron quadrangulare
Phoradendron roraimae Oliv. ^Dendrophthora roraimae
Phoradendron rubrum var. longipedunculatum Rizzini =Phoradendron quadrangulare
Phoradendron rugulosum Urb. =Phoradendron pteroneuron
Phoradendron sanctae-martae Trel. =Phoradendron trinervium
Phoradendron scariosum Rizzini =Phoradendron perrottetii
Phoradendron schottii A. Gray =Phoradendron piperoides
Phoradendron semivenosum Rizzini =Phoradendron pellucidulum
Phoradendron semivenosum var. agostinii Rizzini =Phoradendron pellucidulum
Phoradendron semivenosum var. longipes Rizzini =Phoradendron pellucidulum
Phoradendron spectabile Rizzini =Dendrophthorafendleriana
Phoradendron spectabile var. altimontanum Rizzini =Dendrophthorafendleriana
Phoradendron sulfuratum Rizzini =Phoradendron mairaryense
Phoradendron surinamense PuUe =Phoradendron strongyloclados
Phoradendron taeniicaule Rizzini =Phoradendron nervosum
Phoradendron ? tatei Trel. =Dendrophthorafendleriana
Phoradendron tenuiflorum Steyerm. & Maguire =Dendrophthora tenuiflora
Phoradendron tepuianum Steyerm. =Dendrophthora tepuiana
Phoradendron theloneuron Rizzini =Phoradendron trinervium
Phoradendron tovarense Urb. =Phoradendron dipterum
Phoradendron treleasei Rizzini =Phoradendron trinervium
Phoradendron tubulosum Urb. =Phoradendron nervosum
Phoradendron venezuelense Trel. =Phoradendron quadrangulare
Phoradendron venezuelense var. fdispicum Rizzini =Phoradendron quadrangulare
Phoradendron verticillatum Fawc. & Rendle [nom. superfl.] =Phoradendron trinervium
Phoradendron verticillatum (L.) Druce (Vitaceae) =Cissus verticillata
Phoradendron williamsii Rizzini =Phoradendron pellucidulum
Phoradendron zuloagae Trel. =Phoradendron trinervium
Viscum crassifolium Pohl ex DC. =Phoradendron crassifolium
Viscum dichotomum Bert, ex Spreng. =Phoradendron piperoides
Viscum dimidiatum Miq. =Phoradendron perrottetii
Viscum ellipticum Gardner =Dendrophthora elliptica
Viscum flavens Sw. [nom. superfl.] =Phoradendron racemosum
Viscumfockeanum Miq. =Phoradendron piperoides
Viscum glandulosum Miq. =Phoradendron racemosum
Viscum hexastichum DC. =Phoradendron hexastichum
Viscum latifolium Lam. =Phoradendron racemosum
Viscum martinicensis DC. =Phoradendron crassifolium
Viscum mucronatum DC. =Phoradendron mucronatum
Viscum obliquum C. Presl =Dendrophthora obliqua
Viscum obtusissimum Miq. =Phoradendron ohtusissimum
Viscum perrottetii DC. =Phoradendron perrottetii
Viscum piperoides (Kunth) DC. =Phoradendron piperoides
Viscum quadrangulare (Kunth) DC. ^Phoradendron quadrangulare
552 Checklist of the Plants of the Guiana Shield
Viscum racemosum Aubl. =Phoradendron racemosum
Viscum trinervium Lam. =Phoradendron trinervium
Viscum timaeforme DC. =Phoradendron lunaeforme
Viscum undulatiim Pohl ex DC. =Phoradendron undulatum
Viscum verticillatum L. (Vitaceae) =Cissus verticillata
References
(see also Eremolepidaceae, Loranthaceae)
Krause, K. 1932. Loranthaceae. In A. Pulle, Flora of Suriname 1(1): 4-24.
Kuijt, J. 1961. A revision of Dendrophthora. Wentia 6: 1-145.
Kuijt, J. 2005. Viscaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana 9: 464-
492.
Lindeman, J.C. & A.R.A. Gorts-van Rijn. 1968. Loranthaceae (additions and corrections). In A. Pulle & J. Lanjouw, Flora of
Suriname 1(2): 295-300.
Rizzini, C.T. 1978. Loranthaceae. In B. Maguire & collaborators. Botany of the Guayana Highland, part 10. Mem. New York
Bot. Gard. 29: 23-36.
Rizzini, C.T. 1982. Loranthaceae. In Z. Luce's de Febres, Flora de Venezuela 4(2): 7-316.
Vitaceae
F.D. Chitty, J.A. Steyermarkj
Cissus alata Jacq.
Cissus descoingsii J.A. Lombardi
Cissus duarteana Cambess.
Cissus erosa Rich.
Cissus gongylodes (Baker) Planch.
Cissus guyanensis Desc.
Cissus haematantha Miq.
Cissus palmata Poir.
Cissus spinosa Cambess.
Cissus subrhomboidea (Baker) Planch.
Cissus surinamensis Desc.
Cissus trianae Planch.
Cissus ulmifolia (Baker) Planch.
Cissus venezuelensis Steyerm.
Cissus verticillata (L.) Nicolson & C.E. Jarvis
Cissus verticillata subsp. verticillata
Vitis tiliifolia Willd. ex Roem. & Schult.
Synonyms and Misidentifications
Cissus bonariensis Hook. & Am. =Cissus palmata
Cissus cervii Dunaiski =Cissus gongylodes
Cissus cordifolia L. ^Cissus verticillata subsp. verticillata
Cissus elongata Miq. =Cissus erosa
Cissus erosa var. salutaris (Kunth) Planch. =Cissus erosa
Cissus gibertii (Baker) Planch. =Cissus palmata
Cissus guaranitica Chodat & Hassl. =Cissus erosa
Cissus hassleriana Chodat =Cissus spinosa
Cissus lucida Poir. =Cissus erosa
Cissus martiniana Woodson & Seibert =Cissus trianae
Cissus microcarpa Vahl =Cissus erosa
Cissus ovata Lam. =Cissus verticillata subsp. verticillata
Cissus palmata var. balansaeana Planch. =Cissus palmata
Cissus paraguayensis Planch. =Cissus palmata
Cissus parkeri (Baker) Planch. =Cissus spinosa
Cissus pubescens Kunth =Cissus alata
Cissus quadrialata Kunth =Cissus erosa
Cissus rhombifolia Vahl =Cissus alata
Cissus salutaris Kunth =Cissus erosa
Cissus sexangularis Ernst =Cissus erosa
Cissus sicyoides L. =Cissus verticillata subsp. verticillata
Cissus smilacina Kunth =Cissus verticillata subsp. verticillata
Cissus tamoides Cambess. =Cissus verticillata subsp. verticillata
Cissus trianae var. martiniana (Woodson & Seibert) Steyerm. =Cissus trianae
BO DA
BO GU
FG
BO DA GU SU FG
FG
BO DA GU su FG
DA GU SU FG
DA GU
GU
su
su
su
su
FG
FG
BO GU
BO DA GU su FG
BO DA GU su FG
FG?
Checklist of the Plants of the Guiana Shield 553
Cissus tricuspis (Baker) Planch. =Cissus gongylodes
Cissus thfoliata (L.) L. ? =Cissus erosa
Cissus verticillata subsp. laciniata (Baker) J.A. Lombardi =Cissus verticillata subsp. verticillata
Vitis alata (Jacq.) Kuntze =Cissus alata
Vitis bakeh Herter =Cissiis palmata
Vitis erosa Baker =Cissus erosa
Vitis gibertii Baker =Cissus palmata
Vitis gongylodes Baker =Cissus gongylodes
Vitis miqiieliana Baker =Cissus erosa
Vitis palmata (Poir.) Baker =Cissus palmata
Vitis parkeri Baker =Cissus spinosa
Vitis rhombifolia (Vahl) Baker =Cissus alata
Vitis sicyoides (L.) Morales, [nom. illeg.] =Cissus verticillata subsp. verticillata
Vitis sicyoides (L.) Miq. =Cissus verticillata subsp. verticillata
Vitis sicyoides (L.) Baker
,
[nom. illeg.] =Cissus verticillata subsp. verticillata
Vitis sicyoides f. laciniata Baker =Cissus verticillata subsp. verticillata
Vitis sicyoides var. smilacina (Kunth) Baker =Cissus verticillata subsp. verticillata
Vitis spinosa (Cambess.) Baker =Cissus spinosa
Vitis tricuspis Baker =Cissus gongylodes
References
Descoings, B. 1991. Contribution a I'etude des Vitacees d'Amerique tropicale: deux Cissus nouveaux des Guyanes. Bull. Soc.
Bot. France, Lett. Bot. 138(3): 249-256.
Delascio Chitty, F. & J.A. Steyerrnark. 2005. Vitaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the
Venezuelan Guayana 9: 492-499.
Gorts-van Rijn, A.R.A. 1979. Vitaceae. In A.L. Stoffers & J.C. Lindeman, Flora of Suriname 5(1): 335-343.
Lombardi, J.A. 1995. Typification of names of South American Cissus (Vitaceae). Taxon 44(2): 193-206.
Suessenguth, K. 1953. Vitaceae. In A. Engler & K. Prantl, Die Natiirlichen Pflanzenfamilien 20d: 174-333.
VOCHYSIACEAE
L. Marcano-Berti
Erisma blancoa Marc.-Berti
Erisma calcaratum (Link) Warm.
Erismafloribundum Rudge
Erismafloribundum var. floribundum
Erismajapura Spruce ex Warm.
Erisma laurifolium Spruce ex Warm.
Erisma micranthum Spruce ex Warm.
Erisma nitidum DC.
Erisma splendens Stafleu
Erisma uncinatum Warm.
Qualea acuminata Spruce ex Warm.
Qualea amapaensis Balslev & S.A. Mori
Qualea coerulea Aubl.
Qualea cyanea Ducke
Qualea decorticans Ducke [expected in AM]
Qualea dinizii Ducke
Qualea grandiflora Mart.
Qualea homosepala Ducke
Qualea mori-boomii Marc.-Berti
Qualea paraensis Ducke
Qualea polychroma Stafleu
Qualea psidiifolia Spruce ex Warm.
Qualea pulcherrima Spruce ex Warm.
Qualea rosea Aubl.
Qualea rupicola Ducke
Qualea schomburgkiana Warm.
Qualea sprucei Warm.
Qualea suprema Ducke
Qualea themistoclesii Ducke
Qualea tricolor Benoist
Qualea tuberculata Stafleu
Qualea wurdackii Marc.-Berti
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
GU
GU
FG
FG
FG
BO DA GU SU FG
BO GU? FG
FG?
SU FG
BO DA GU SU FG
SU
BO
FG
BO GU
BO GU
FG
SU FG
BO GU
FG
554 Checklist of the Plants of the Guiana Shield
Ruizterania albiflora (Warm.) Marc.-Berti
Ruizterania cassiquiarensis (Spruce ex Warm.) Marc.-Berti
Ruizterania esmeraldae (Standi.) Marc.-Berti
Ruizteraniaferruginea (Steyerm.) Marc.-Berti
Ruizterania nitida (Stafleu) Marc.-Berti
Ruizterania obtusata (Briq.) Marc.-Berti
Ruizterania obtusata subsp. multivenosa Marc.-Berti
Ruizterania obtusata subsp. obtusata
Ruizterania retusa (Spruce ex Warm.) Marc.-Berti
Ruizterania rigida (Stafleu) Marc.-Berti
Ruizterania trichanthera (Spruce ex Warm.) Marc.-Berti
Salvertia convallariodora A. St.-Hil.
Vochysia angustifolia Ducke
Vochysia apopetala Ule
Vochysia bautistae Marc.-Berti
Vochysia calophylla Spruce ex Warm.
Vochysia cassiquiarensis Stafleu
Vochysia catingae Ducke
Vochysia cayennensis Warm.
Vochysia complicata Ducke
Vochysia costata Warm.
Vochysia crassifolia Warm.
Vochysia densiflora Spruce ex Warm.
Vochysia elegans Stafleu
Vochysia expansa Ducke
Vochysiaferruginea Mart.
Vochysia glaberrima Warm.
Vochysia grandis Mart.
Vochysia grandis var. uaupensis Warm.
Vochysia guianensis Aubl.
Vochysiajonkeri Marc.-Berti
Vochysia julianensis Marc.-Berti
Vochysia aff. laxiflora Stafleu
Vochysia liscanoi Marc.-Berti
Vochysia maguirei Marc.-Berti
Vochysia neyratii Normand
Vochysia obscura Warm.
Vochysia ortegae Marc.-Berti & Bautista
Vochysia pauciflora Steyerm.
Vochysia pinkusii A.C. Sm. [expected in BO]
Vochysia punctata Spruce ex Warm.
Vochysia rubiginosa Stafleu
Vochysia sabatieri Marc.-Berti
Vochysia saccata Stafleu
Vochysia schomburgkii Warm.
Vochysia spathiphylla Stafleu
Vochysia speciosa Warm.
Vochysia splendens Spruce ex Warm.
Vochysia steyermarkiana Marc.-Berti
Vochysia surinamensis Stafleu
Vochysia surinamensis var. inflata Stafleu
Vochysia surinamensis var. surinamensis
Vochysia tetraphylla (G. Mey.) DC.
Vochysia tilletii Marc.-Berti
Vochysia tomentosa (G. Mey.) DC.
Vochysia tucanorum Mart.
Vochysia tucanorum var. tucanorum
Vochysia venezuelana Stafleu
Vochysia vismiifolia Spruce ex Warm.
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
AM
BO
GU
GU
SU FG
BO
BO
GU
SU
BO
BO
BO
BO
BO
BO
BO
BO
GU
GU
GU
GU
GU
SU
SU
FG
FG
GU SU FG
SU
GU
FG
FG
AM
GU
AM
FG
AM
AM
AM BO
BO
DA GU SU FG
AM BO DA GU SU FG
BO DA GU SU FG
AM
AM BO GU SU
SU
SU
FG
AM BO
AM BO
Synonyms and Misidentifications
Cucullaria excelsa M. Vahl =Vochysia tomentosa
Cucullariaferruginea (Mart.) Spreng. ^Vochysiaferruginea
Cucullaria grandis (Mart.) Spreng. =Vochysia grandis
Checklist of the Plants of the Guiana Shield 555
Cucullaria tetraphylla G. Mey. =Vochysia tetraphylla
Cucullaria tomentosa G. Mey. =Vochysia tomentosa
Debraeafloribundum (Rudge) Roem. & Schult. =Ehsmafloribundum
Debraea violacea (Mart.) Steud. =Erisma calcaratum
Ditmariafloribunda (Rudge) Spreng. =Erismafloribundum
Ditmaria violacea (Mart.) Spreng. =Ehsma calcaratum
Erisma pallidiflorum Ducke =Erismafloribundum
Erisma parvifolium Gleason =Erismafloribundum
Erisma parvifolium yslt. pallidiflorum (Ducke) Ducke =Erismafloribundum
Erisma pulverulentum Poepp. ex Warm, [nomen nudum] =Erisma uncinatum
Erisma violaceum Mart. =Erisma calcaratum
Qualea albiflora Warm. =Ruizterania albiflora
Qualea apodocarpa Steyerm. =Ruizteraniaferruginea
Qualea calcarata Link. =Erisma calcaratum
Qualea cassiquiarensis Spruce ex Warm. =Ruizterania cassiquiarensis
Qualea esmeraldae Standi. =Ruizterania esmeraldae
Qualeaferruginea Steyerm. =Ruizteraniaferruginea
Qualea nitida Stafleu =Ruizterania nitida
Qualea obtusata Briq. =Ruizterama obtusata
Qualea retusa Spruce ex Warm. =Ruizterania retusa
Qualea retusa var. coriacea Ducke =Ruizterania obtusata
Qualea rigida Stafleu =Ruizterania rigida
Qualea rubiginosa Stafleu =Ruizteraniaferruginea
Qualea rubiginosa var. angustior Steyerm. =Ruizteraniaferruginea
Qualea speciosa Huber =Qualea acuminata
Qualea trichanthera Spruce ex Warm. =Ruizterania trichanthera
Qualea verruculosa Stafleu =Ruizterania esmeraldae
Ruizterania apodocarpa (Steyerm.) Marc.-Berti =Ruizteraniaferruginea
Ruizterania rubiginosa (Stafleu) Marc.-Berti =Ruizteraniaferruginea
Ruizterania rubiginosa var. angustior (Steyerm.) Marc.-Berti =Ruizteramaferruginea
Ruizterania verruculosa (Stafleu) Marc.-Berti =Ruizterania esmeraldae
Vochyaferruginea (Mart.) Standi. =Vochysiaferruginea
Vochysia arcuata Garcke =Vochysia tetraphylla
Vochysia curvata Klotzsch ex R.M. Schomb. [nomen nudum] =Vochysia crassifolia
Vochysia grandis var. genuina Briq. =Vochysia grandis
Vochysia inundata var. venosa Ducke =Vochysia saccata
Vochysia javitensis Stafleu = Vochysia angustifolia
Vochysia lucida Klotzsch ex R.M. Schomb. [nomen nudum] =Vochysia glaberrima
Vochysia racemosa Poir. [name of uncertain application, not Vochysia^
Vochysia tetraphylla (G. Mey.) H. Stone & W. Freeman =Vochysia tetraphylla
Vochysia vismiifolia var. robusta Ducke =Vochysia splendens
References
Marcano-Berti, L. 1969. Un nuevo genero de las Vochysiaceae. Pittieria 2: 3-27.
Marcano-Berti, L. 1989. Vochysiaceae: novedades y correciones. Pittieria 18: 5-14.
Marcano-Berti, L. 1998. Vochysiaceae. In A.R.A. Gorts-van Rijn, Flora of the Guianas (series A) 21 : 1-44.
Marcano-Berti, L. 2005. Vochysiaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist. Flora of Ihc Venc/iiclan
Guayana 9: 500-524.
Stafleu, F.A. 1948-1954. A monograph of the Vochysiaceae. 1. Salvertia & Vochysia. Rcc. Trav. Bot. Neerl. 41 : .V)7-540 (194S).
111. Qualea. Acta Bot. Neerl. 2: 144-217 (1953). IV. Erisma. Acta Bot. Neerl. 3: 459-480 ( 1954).
Stafleu, F.A. 1 95 1 . Vochysiaceae. In A. Pulle, Flora of Suriname 3(2): 1 78- 1 99.
Steyermark, J.A. 1987. Vochysiaceae: a reevaluation of the genus Euphronia. In J.A. Steyermark, l-lora of the Venc/uclan
Guayana - 11. Ann. Missouri Bot. Gard. 74( 1 ): 85- 1 1 6.
WiNTERACEAE
J.S. Miller
Drimys roraimensis (A.C. Sm.) Ehrend. & Gottsb.[Mt. Roraima] AM BO GU
Synonyms ani.i Misidf.ntifications
Drimys brasiliensis var. ptaritepuiensis Steyerm. ^Drimys roraimensis
Drimys brasiliensis var. roraimensis A.C. Sm. = Drimys roraimensis
References
Smith, A.C. 1 943. The American species of Drimys. J. Am. Arb. 24: 1 -33.
556 Checklist of the Plants of the Guiana Shield
Eherendorfer, F, I. Silberbauer-Gottsberger & G. Gottsberger. 1979. Variation on the population, racial, and species level in the
primitive relic angiosperm genus Drimys (Winteraceae) in South America. Plant Systematics and Evolution 132: 53-83.
Miller, J.S. 2005. Winteraceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of the Venezuelan Guayana
9: 524-525.
Zygophyllaceae
G.A. Aymard C, N.L. Cuello A., B.E. Leuenberger
BO
BO
BO
BO
BO
GU SU
GU SU
Guaiacum officinale L.
Kallstwemia maxima (L.) Hook. & Am. FG
KaUstroemia pubescens (G. Don) Dandy
Thbulus cistoides L.
Tribulus zeyheri Sond.
Tribulus zeyheri subsp. macranthiis (Hassk.) Hadidi
.
Synonyms and Misidentifications
Kallstwemia caribaea Rydb. =Kallstroemia pubescem
Tribulus macranthus Hassk. =Tribulus zeyheri subsp. macranthus
Tribulus maximus L. =Kallstroemia maxima
Tribulus pubescens G. Don =Kallstroemia pubescens
References
Aymard C, GA. & N.L. Cuello A. 2005. Zygophyllaceae. In J.A. Steyermark, RE. Berry, K. Yatskievych & B.K. Hoist, Flora of
the Venezuelan Guayana 9: 580-584.
Howard, R.A. 1988. Zygophyllaceae. In R.A. Howard, Flora of the Lesser Antilles 4(1): 551-555.
Lasser, T. 1971. Zygophyllaceae. In T. Lasser, Flora de Venezuela 3(1): 75-89. Stafleu, F.A. 1951. Zygophyllaceae. In A. Pulle,
Flora of Suriname 3(2): 200-203.
cm
Justicia wilhelmensis Wasshausen (Acanthaceae)
Checklist of the Plants of the Guiana Shield 557
Passiflora kawensis Feuillet (Passifloraceae)
558 Checklist of the Plants of the Guiana Shield
Stifftia uniflora Ducke amd Stifftia cayennensis H.Rob. & Kahn (Compositae)
Checklist of the Plants of the Guiana Shield 559
Talisia simaboides Kramer (Sapindaceae)
560 Checklist of the Plants of the Guiana Shield
ccfTULcea^
Pinzona coriacea Mart. & Zucc. (Dilleniaceae)
Checklist of the Plants of the Guiana Shield 561
Lacunahajenmanii (Oliv.) Ducke (Quiinaceae)
562 Checklist of the Plants of the Guiana Shield
Literature citation for illustrations
Page 4 Henriettea multiflora Naudin, In: Wurdack, J. J., Morley, T., and S. Renner 1993.
Melastomataceae. Flora of the Guianas: Series A: Phanerogams, Fascicle 13:
425. Koenigstein, Germany. Koeltz. Illustrator: Alice Tangerini
Page 16 Guapira eggersiana (Heimerl) Lundell, In: DeFilippe, R. (in review) Nyctaginaceae.
Flora of the Guianas: Series A: Phanerogams, Fascicle xx. Richmond, United
Kingdom. Royal Botanical Garden, Kew. Illustrator: Alice Tangerini.
Page 556 Justicia wilhelmensis Wasshausen, In: Wasshausen, D. C. 2006. Acanthaceae. Flora
ofthe Guianas: Series A: Phanerogams, Fascicle 23: 1-141. Richmond, United
Kingdom. Royal Botanical Garden, Kew. Illustrator: Alice Tangerini.
Page 557 Passiflora kawensis Feuillet, In: Feuillet, C. 1994. Two new species of Passiflora
(Passifloraceae) from French Guiana. Novon 4(3): 236-238. Illustrator: Cathy
Pasquale.
Page 558 Stifftia uniflora Ducke and Stifftia cayennensis H. Robinson & B. Kahn, In: Robinson,
H. and B. Kahn. 1985. A new species of Stifftia from Cayenne (Asteraceae:
Mutisieae) Phytologia 58(4): 248-251. Illustrator: Alice Tangerini.
Page 559 Talisia simaboides Kramer, In: Acevedo-Rodriquez, P. 2003. Melicocceae
(Sapindaceae): Melicoccus and Talisia. Flora Neotropica 87: 1-178. Illustrator:
Bobbi Angell.
Page 560 Pinzona coriacea Mart. & Zucc, In: Mori, S. A., G Cremers, C. A. Gracie, J. de
Granville, S. V. Heald, M. Hoff and J. D. Mitchell. 2002. Guide to the Vascular
plants of Central French Guiana, Pt. 2. Dicotyledons. Memoirs of the New York
Botanical Garden 76: 251. Illustrator: Bobbi Angell.
Page 561 Lacunaria jenmanii (Oliv.) Ducke, In: Mori, S. A., G Cremers, C. A. Gracie, J. de
Granville, S. V. Heald, M. Hoff and J. D. Mitchell. 2002. Guide to the Vascular
plants of Central French Guiana, Pt. 2. Dicotyledons. Memoirs of the New York
Botanical Garden 76: 596. Illustrator: Bobbi Angell.
Page 583 Rhynchospora cordatachenia M.T. Strong, In: Strong, M. T. 2005. Two new species
ofRhynchospora section Tenues (Cyperaceae) from the Guianas, South America.
Novon 15(3): 479-483. Illustrator: AHce Tangerini.
Checklist of the Plants of the Guiana Shield 563
APPENDIX
Individuals who provided plant identifications.
(t = deceased)
Dr. Pedro Acevedo R. (US)
Dr. Mac Alford (USMS)
Dr. Lucile Allorge (P)
Dr. Frank Almeda (CAS)
Dr. Christiane Anderson (MICH)
Dr. William R. Anderson (MICH)
Dr. A. Aptroot (U)
Dr. Scott Armbruster (ALA)
Dr. Daniel F. Austin (Arizona-Sonora
Desert Museum)
Dr. Franklin Ayala (AMAZ)
Dr. Gerardo A. Aymard C. (PORT)
Dr. Michael J. Balick (NY)
Dr. Harvey E. Ballard (OH)
Dr. Julie Barcelona (PNH)
Dr. Rupert C. Bameby t (NY)
Dr. Lisa C. Bamett (US)
Dr. S. Barrier (P)
Dr. Kerry Barringer (BKL)
Dr. Daniel Barthelemy (MPU)
Dr. Cornells C. Berg (BG)
Dr. Paul Berry (MICH)
Dr. M.H. Bischler (G)
Mr. John K. Boggan (US)
Dr. Brian Boom (NY)
Mr. Jason Bradford (MO)
Dr. John Brandbyge (AAU)
MS Arian Jacobs-Brouwer (U)
Dr. William R. Buck (NY)
Dr. Ricardo Callejas (HUA)
Dr. Koen Camelbeke (Stichting
Arboretum Wespelaar, RUG)
Dr. Kenneth K. Cameron (NY)
Lie. German Camevali (MO)
Dr. Paul M. Catling (DAOM)
Dr. Andre Chanderbali (FLAS)
Dr. Mark Chase (MICH)
Dr. Eric A. Christenson
Dr. John L. Clarke (US)
Dr. H. David Clarke (UNCA)
Dr. M.J.E. Coode (K)
Dr. Alain Coute (PC)
Dr. Richard Cowanf (PERTH)
Dr. George Cremers (P)
Dr. Thomas B. Croat (MO)
Dr. Jose Cuatrecasasf (US)
Dr. C. Cusset (P)
Dr. Douglas C. Daly (NY)
Dr. Chris Davidson (Boise Botanical
Garden)
Dr. Garrit Davidse (MO)
Dr. Robert DeFilippsj (US)
Dr. Alfonso Delgado-Salinas (MEXU)
Dr. Piero Delprete (NY)
Dr. Paula DePriest (US)
Dr. B.M. Descoings (MPU)
Mr. Ron Determann (Atlanta Botanical
Garden)
Dr. Pierre Detienne (CTFT)
Lie. Wilmer Diaz (GUYN)
Dr. Laurence J. Dorr (US)
Dr. Stefan Dressier (Leiden)
Dr. Renski Ek (U)
H.-J. Esser (U)
Dr. Robert B. Faden (US)
Dr. Paul R. Fantz (NCSC)
Dr. Marie L. Farr (BPI)
Dr. Christian Feuillet (US)
Dr J.J. Floret (P)
Dr. James P Folsom (HNT)
Dr. Enrique Forero (MO)
Dr. Alina Freire-Fierro (PH)
Dr. V.A. Funk (US)
Dr. Alwyn H. Gentry t (MO)
Dr. Lynn J. Gillespie (CAN)
Dr. Tom Givnish (WIS)
Dr. Paul Goetghebeur (GENT)
Dr. Aaron Goldberg (US)
Dr. Peter Goldblatt (MO)
Dr. A.R.A. Gorts-van Rijn (U)
Dr. Eric J. Gouda (U)
Dr. S.R. Gradstein (GOET)
Dr. Jason R. Grant (NEU)
Dr. Jean-Jacques de Granville (CAY)
Dr. James Grimes (NY at the time of
the work)
Dr. Susan Grose (WTU)
Dr. Micheline Guerlesquin (ANGUC)
Dr. J. Hagemann (B)
Dr. Eric Hagsater (AMO)
Dr. William J. Hahn (Georgetown
Univ.)
Dr. Madjit Hakki (B)
Dr. Bertel Hansen (C)
Dr. RayM. Harley (K)
Dr. Robert R. Haynes (UNA)
Dr. W.H.A. Hekkingt (U)
Dr. H.v. Helvoort(U)
Dr. Andrew Henderson (NY)
Dr. Terry Henkel (HSC)
Dr. E. Hennipman (U)
Dr. Patrick Herendeen (Geo.
Washington Univ.)
Dr. Paul Hiepko (B)
Dr. Steven Hill (CLEMS)
Dr. Michel Hoff(STR)
Mr. Bruce Hoffman (HAW)
Dr. Tom HoUowell (US)
Dr. Walter C. Holmes (BAYLU)
Dr. Noel H. Holmgren (NY)
Dr. Lauritz B. Holm-Nielsen (AAU)
Mr. Bruce Hoist (SEL)
Dr. Charles N. Horn (NBYC)
Dr. Otto Huber (Venezuela)
Dr. Marion J. Jansen-Jacobs (U)
Dr. C. Jeffrey (L)
Dr. J. Jeremie (P)
Dr. A.C. Jermy (BM)
Dr. S. Jovet-Ast (PC)
Dr. Emmet J. Judziewicz (UWSP)
Dr. F. Kahn (CAY)
Dr. Jacquelyn Kallunki (NY)
Dr. CarolL. Kelloff(US)
Dr. Helen Kennedy (UBC)
Dr. Robert W. Kiger (CM)
Dr. P Kirk (IMI)
Dr. Joseph H. Kirkbride (BARC)
Dr. B. B. Klitgaard (BM)
Dr. Sandy Knapp(BM)
Dr. Robert Krai (BRIT)
Dr. Karl U. Kramer (Z)
Dr. W. John Kress (US)
Dr. R. Kruijt (U)
Dr. Job Kuijt (UVIC)
Dr. Klaus Kubitzki (HBG)
Dr. Lars Peter Kvist (AAU)
Prof Dr. H.W. Lack (B)
Dr. Tom Lammers (OSH)
Dr. Leslie R. Landrum (ASU)
Dr. A. de Langc (AMD)
Dr. Anthony J.M. Lccuwen berg
(WAG)
Dr. David B. Lcllinger (US)
Dr. Blanca Leon (TEX)
Dr. Beat Ernst Leuenberger (B)
Dr. Gwilym Lewis (K)
Mr. Ron Liesner(MO)
Dr. H.C. deLima(RB)
Dr. Jan C. Lindeman (U)
Dr. Eduardo Lleras-Perez (CEN)
Dr. Pierre-Andre Loizeau (G)
Dr. Julio A. Lombardi (HRCB)
Ms. Ximena Londofio (Cali,
Colombia)
Dr. Alicia Lourteig t(P)
Dr. Carlyle Luer (MO)
564 Checklist of the Plants of the Guiana Shield
Dr. Eimear Nic Lughada (K) Dr.
Dr. James L. Luteyn (NY) Dr.
Mr. Harry E. Luther (SEL) Dr.
Dr. Paul J.M. Maas (U) Dr.
Dr. H. Maas-van de Kamer (U) Mr.
Dr. Bob Magill (MO) Dr.
Dr. Luis Marcano-Berti via (U) Dr.
Dr. Richard H. Maxwell (JEF) Dr.
Dr. Lucille M. McCook (MISS) Dr.
Dr. Tim McDowell (ETSU) Dr.
Mr. Greg McKee (US) Dr.
Dr. J.T. de Medeiros Costa (UFP)
Mr. Max E. Medley (DHL) Dr.
Dr. Alan W. Meerow (FTG) Dr.
Dr. Willem Meijerj (KY) Dr.
Dr. L. Melendez-Howell (PC) Dr.
Dr. A.M.W. Mennega t(U) Dr.
Dr. Fabian Michelangeli (NY) Dr.
Dr. James Miller (MO) Sr.
Dr. David Minter (IMI) Dr.
Dr. John D. Mitchell (NY) Dr.
Dr. Reinaldo Monteiro (HRCB) Dr.
Dr. Robbin Moran (NY) Dr.
Dr. Luis E. Mora-Osejo (COL) Dr.
Dr. Scott Mori (NY) Dr.
Dr. Gilberto Morillo (HUA) Dr.
Dr. Thomas Morley (MIN) Mr.
Dr. Joao Murca Pires (MG) Dr.
Dr. Jose C. Murillo (COL) Dr
Dr. Michael Nee (NY) Dr.
Dr. DavidA.Neill(MO) Dr.
Dr. Dan H. Nicolson (US) Dr.
Dr. James N. Norris (US) Dr.
Dr. B. Oellgaard (AAU) Dr.
Mr. Darin Penneys (FLAS) Dr.
Dr. R. Toby Pennington (E) Dr.
Dr. T.D. Pennington (K) Dr.
Dr. Paul Peterson (US) Dr.
Dr. John J. Pipoly (Florida) Dr.
Dr. Timothy Plowman j (MO) Dr.
Roger M. Polhill (K) Dr.
Odile Poncy (P) Dr.
Hans-Helmut Poppendieck (HBG) Dr.
Ghillean T. Prance (K) Dr.
John F. Pruski (MO) Dr.
W. Punt (U) Dr.
J.M. Rankin (BM) Dr.
Thomas Raus (B) Dr.
Peter H. Raven (MO) Dr.
Robert W. Read t (US) Mr.
Karen M. Redden (Geo. Dr.
Washington Univ.) Dr.
Susanne Renner (UMSL) Dr.
Harold Robinson (US) Dr.
Norman K.B. Robson (BM) Mr.
Andre Robyns (BR) Ms.
William A. Rodrigues (UPCB) Ms.
Jack D. Rogers (WSP)
Gustavo A. Romero (AMES) Dr.
A.C. de Roon (U) Dr.
Velva E. Ruddt (US) Dr.
Daniel Sabatier (MPA) Dr.
Gary G. Samuels (BPI) Dr.
Claude Sastre (P) Dr.
Jakob J. Schneller (Z) Dr.
Brian Schrire (K) Dr.
Matthew Sewell (NY) Dr.
M.F. da Silva (INPA) Dr.
David A. Simpson (K) Dr.
Beryl Simpson (TEX) Dr.
Laurence E. Skog (US) Dr.
Flemming Skov (AAU) Dr.
Alan R. Smith (UC) Dr.
Lyman B Smith t (US) Dr.
Steven F. Smith (US) Dr.
James Solomon (MO) Dr.
Mario Sousa Sanchez (MEXU) Dr.
Chelsea Specht (U.CA) Dr.
Rodolphe E. Spichiger (G) Dr.
Clive A. Stace (LTR) Dr.
Bertil Stahl (SUNIV)
Fred W. Stauffer (G)
Basil Stergios (PORT)
Dennis W. Stevenson (NY)
Donald E. Stone (DUKE)
Mark T. Strong (US)
D.L. Szlachetko (UGDA)
Charlotte M. Taylor (MO)
Peter Taylor (K)
Oswaldo Tellez Valdez (MEXU)
Y. Therezien (P)
W. Wayt Thomas (NY)
Sue Thompson (CM)
Pierre Tixier (PC)
Ben Torke (MO)
Erin A. Tripp (DUKE)
Amy Trojnar (Southern Illinois
University)
Gordon C. Tucker (EIU)
Yvonne Veyret (P)
Warren L. Wagner (US)
Warren H. Wagner Jr.f (MICH)
Bruno Wallnofer (W)
Dieter Wasshausen (US)
Grady L. Webster t (DAV)
Anna Weitzman (US)
Tom Wendt (TEX)
H. vanderWerff(MO)
Marga CM. Werkhoven (BBS)
John H. Wiersema (BARC)
Paulo G. Windisch (HASU)
Donald R. Windier (BALT)
Robert Wingfield (CORO)
John J. Wurdackt (US)
Kenneth Wurdack (US)
Thomas Zanoni (NY)
D. Zappi (K)
Elsa M. Zardini (MO)
James Zarucchi (MO)
Brigitte Zimmer (B)
Checklist of the Plants of the Guiana Shield 565
INDEX TO GENERA
Abarema 365, 370-372, 374-375
Abelmoschus 393-394
Aberemoa 193
Abildgaardia 88, 95-96, 98, 103
Ablania 284
Abolboda 178-181
Abrus 346, 356, 499
Abuta 420-421
Abutilon 393-394
Acacallis 118, 134
Acacia 365, 370, 372-376
Acalypha 290-295
Acanthella 397, 406, 495
Acanthinophyllum 425
Acanthocereus 231-232
Acanthospermum 255, 260, 262, 264
Acanthosphaera 425
Acetosella 453
Achetaria 522-523
Achimenes 310
Achlyphila 179
Achnopogon 255, 260
Achras 518-519
Achrouteria 5 1
8
Achyranthes 187-188
Achyrocline 255, 260, 264
Acianthera 134
Acicarpa 158
Acidoton 290, 295-297
Acineta 118
Acioa 241, 245-246
Aciotis 397, 406-407, 410-411, 413-
417
Acisanthera 397, 406-41 1, 413-417
Acmella 255-256, 260-262, 267
Acnida 188
Acoidium 134
Acopanea 223
Acosmium 346-347, 360, 363, 365
Acosta 470
Acouroa 356, 360
Acrandra 438
Acrocarpidium 461
Acroceras 149, 165, 167
Acrocomia 67, 69-70
Acrodiclidium 326
Acrostichum 50-51
Actinostachys 52
Actinostemon 290, 295, 297
Acuan 370
Ada 118
Adelia 290, 299
Adelobotrys 397, 407, 409-41 1, 41
5
Adenanthe 445
Adenanthera 365
Adenarake 445
Adeneleutherophora 134
Adenobasium 284
Adenocalymna 214, 217-218
Adenolisianthus 304, 306-307
Adenophaedra 290-300
Adenostemma 261
Adenostephanus 474
Adhatoda 185
Adiantopsis 22-23
Adiantum 22-23, 51
Adipe 134
Adipera 340
Adiscanthus 507
Aechmea 73, 78-81,83
Aegiphila 543-546
Aegopogon 149, 158
Aeranthus 134
Aeschrion 525
Aeschynomene 342, 347, 356, 360,
364
Agalinis 523
Aganisia 118, 134
Agarista 285, 287
Agati 356
Ageratina 256, 263
Ageratum 256, 261
Agonandra 453
Agrosticula 158
Agrostis 158
Ahouai 199
Aiouea 322-323, 326-327, 330
Aira 158
Akentra 377
Albizia 365, 370, 375-376
Alchomea 290, 295-297
Alchomeopsis 290, 295-296
Alcoceratothrix 389
Aldina 347, 356
Alectoroctonum 296
Alectra 522-524
Alegria 538
Alexa 347, 356
Alexandra 356
Alibertia 479, 492-493, 495-497, 503,
505-506
Alicastrum 425
Alisma 59
Aliteria425
Allamanda 195,202
Allania 356
AUantoma 330-331
Alloiosepalum 279
Allophylus 51 1,513-514
Alloplectus 309-3 11
Allosidastrum 393
Allosorus 30
Almeidea 508
Alocasia 59, 64
Alomia 261
Alophia 1 1
1
Alpinia 182
Alseis 479, 492
Alsodeia 548
Alsophila 26-28
Alstroemeria 112-113
Altemanthera 187-188
Althaea 394
Alysicarpus 347, 360
Amaioua 479, 492-493
Amanoa 290, 296
Amaranthus 187-188
Amaryllis 113
Amasonia 544-546
Amauropelta 54
Ambelania 195, 198-199,203
Amblyanthera 199
Ambrosia 256, 261
Amerimnon 356
Ammannia 384-385
Amomum 182
Amorphospermum 518
Ampelocera 542
Ampelodaphne 326
Ampelozizyphus 477
Ampherephis 261
Amphidasya 479,507
Amphidesmium 44
Amphilophium 214, 217-218
Amphineuron 54
Amphiodon 356
Amphiphyllum 173, 175
Amphirrhox 547-548
Amphiscopia 185
Amphizoma 314
Amphymenium 340
Amyris 507-508
Anacampta 199
Anacardium 189-190
Anacheilium 134
Anadenanthera 366, 372-374
Anagallis 473
Ananas 73, 79
Ananthacorus 51, 55
Anaphyllopsis 59, 64
Anartia 198-199
Anastrophus 158
Anathallis 134
Anatherum 158
Anaxagorea 190, 193-195
Ancistrocarpus 457
Ancistrothyrsus 454
Andira 347-348, 356, 360
Andriapetalum 474
Andropogon 149-150, 158-159, 161-
162
Androstylium 251
Anechites 199
Aneilema 85
566 Checklist of the Plants of the Guiana Shield
Anelasma 421
Anemia 45, 52
Anemopaegma 214, 217-219
Anetium 50, 55
Angadenia 199
Angelonia 522
Angostura 508-510
Angostyles 296
Angraecum 134-135
Anguria 277
Anguriopsis 277
Aniba 323, 326-328, 330
Anila 356
Anisacanthus 183, 185
Anisantherina 522-523
Aniseia 271, 273-274
Anisolobus 199
Anisomeles 321
Anisomeris 493
Anisophyllea 190
Anneslia 370
Annona 190, 193-194
Anomoctenium 217
Anomospermum 420-421
Anonocarpus 425
Anoplophytum 79
Anredera 2 1
3
Anthaenantia 159
Anthephora 150, 172
Anthereon 135
Anthericum 113
Anthodiscus 236
Anthodon 313-315
Anthodus 314
Anthurium 59-60, 64-66
Antidaphne 285
Antidesma 290, 296
Antigonon 47
1
Antirhea 479, 493
Antonia 379-380
Antrophyum 23, 55
Apalanthe 111
Apalatoa 340
Aparisthmium 290, 295-296
Apatitia 407
Apatostelis 135
Apeiba 538-539
Aphanocarpus 479, 493, 499
Aphelandra 183, 185-186
Apinagia 466-468
Apluda 159
Apocaulon 507
Apocynum 199
Apodandra 296
Apodanthera 277
Apodanthes 476
Apodina 382
Apoleya 340
Apophragma 306
Appendicularia 397, 415
Appunia 493
Aptandra 449-450
Apteria 83-84
Apuleia 333, 340, 344, 358
Aracamunia 118
Arachniodes31-33, 48, 54
Araeococcus 73
Aralia 11,207
Aranella 377
Aratitiyopea 80, 179
Arberella 150
Archytaea 223-224
Ardisia 430, 432-433
Argemone 454
Argythamnia 296
Aristida 150, 159-161, 171
Aristolochia 207-208, 421
Arkezostis 277
Arouna 340
Arrabidaea214-215,217-219
Arrhostoxylum 185
Arrudea 251
Artanthe461
Arthrosamanea 370
Arthrostema 407
Arthrostemma 407
Arthrostylidium 150, 159, 161
Aruba 525
Arum 64
Arunda 159
Arundinaria 159
Arundmella 150, 158-159, 169
Arundo 150, 159
Asclepias 208, 210-211
Ascolepis 88, 95, 99
Aspasia 118
Aspegrenia 135
Aspidium 32
Aspidogyne 118, 140, 144
Aspidosperma 195, 199-201
Aspilia 261
Asplenium 24-25, 32, 56
Asplundia 87
Astephanus 211
Aster 261
Asteranthos 330
Asterogyne 67, 69
Asterolepidion 319
Asterophorum 539
Astrocaryum 67, 69
Astrococcus 290, 296
Astronium 189
Asystasia 183, 185-186
Ateramnus 296
Athenaea 302
Athyrocarpus 85
Atractantha 150
Attalea 67, 69-72
Aubletella57, 518
Augusta 261
Aulizia 135
Aulomyrcia 438-440
Aulonemia 150, 159
Austroamericum 5 1
1
Austroeupatorium 256, 263
Avesicaria 377
Avicennia 544-545
Axinanthera 407
Axonopus 150-151, 158-161, 164-169
Ayapana 256, 261,263-264
Aydendron 327
Ayenia 531
Ayensua 73, 178
Azadirachta 417
Azara 302
Azolla51
Baccharis 256, 261-264, 266
Baccharoides 261
Bacopa 522-524
Bactris 67-72
Badula 432
Bagassa 423, 425-426, 465
Baillieria 261
Balantium 244
Balizia 366, 372, 374-376
Ballota321
Baltimora 256, 261,264
Bambusa 160
Banara 301-304
Banisteria 389
Banisteriopsis 386, 389-390, 392
Barbacenia 178
Barbieria 348, 356-357, 359
Barbosella 118, 135
Barjonia 208
Barleria 183, 185
Bamebydendron 333
Barnhartia 468
Barrosoa 256, 263
Bartramia 539
Basanacantha 493
Basella213
Baskervilla 118, 135
Bassovia 528
Bastardia 393
Batatas 273
Batemannia 118
Batesia 334
Bathmium 53
Bathysa 479, 495, 505
Batis213
Batocarpus 423, 425
Bauhinia 334, 340-341, 343, 345
Beadlea 135
Becquerelia 88, 95, 102
Befaria 287
Begonia 213-214
Beilschmiedia 323, 327
Bejaria 285, 287
Bellardia 493
Bellucia397,407, 410, 417
Bellynkxia 493
Beloglottis 118, 135
Beloperone 183, 185
Benitzia 84
Benjaminia 522-524
Bemardia 290, 300
Bemardinia 270
Bertholletia 330-331
Bertiera 479, 493
Bertolonia 397
Besleria 309, 311-312
Checklist of the Plants of the Guiana Shield 567
Beyrichia 523
Bidens 256, 261
Bifrenaria 118, 134-135, 148
Bignonia 217-218
Bihai 110
Billardiera 493
Billbergia 73, 79
Binaria 341
Biophytum 453
Biovularia 377
Bisboeckelera 88, 95, 98
Bixa 220
Blainvillea 256
Blakea397,407,417
Blastemanthus 445, 448
Blechnum 25-26, 47
Blechum 183, 185-186
Blepharandra 386, 390
Blepharocalyx 434
Blepharodon 208-209, 211-212
Bletia 118, 135
Blochmannia 472
Blotiella 29, 31,51
Blutaparon 187-188
Bocagea 193
Bocageopsis 191, 193-194
Bocoa 334, 363
Boehmeria 542-543
Boerhavia 442-443
Bogenhardia 394
Bolbitis 23, 42-43, 50-51
Bolbophyllaria 135
Bolboschoenus 95
Bollea 118, 135, 141
Bomarea 112-113
Bombacopsis 220-222
Bombax 221-222
Bonafousia 198-200
Bonamia271,273, 275
Bonatea 135
Bonnetia 223-224
Bonpartea 79
Bonplandia 509
Bontia 428, 545
Bonyunia 379-380
Borismene 420-421
Borojoa 493
Borreria 479, 493, 498, 504
Bothriochloa 151, 160
Bothriospora 479, 496
Botryarrhena 479
Bouchea 544, 546
Bougainvillea 442-443
Bourreria 224-226, 477
Boussingaultia 213
Bouteloua 151, 159
Bowdichia 348, 356
Boyania 397
Brachiaria 151, 160, 164-167, 172
Brachionidium 118
Brachistus 528
Brachycodon 306
Brachyloma 3 1
1
Brachynema 450
Brachypremna 407
Brachypterys 390
Brachystele 118, 140, 147
Bradburya 356
Braemia 118, 141, 146
Bramia 523
Brasenia 231
Brassavola 118, 135, 138
BrassiallS, 135, 138
Brassica 226
Bravaisia 183, 185-186,545
Bredemeyera 468-470
Brewcaria 73-74, 80
Brickellia 256, 262-263
Brignolia 493
Briquetia 393-395
Briza 151
Brocchinia 58, 74, 79
Bromelia 74, 79-80
Bromus 160
Brosimopsis 425
Brosimum 247, 423, 425-427
Brossea 287
Brownea 334, 341,344
Bruinsmania 493
Brunellia 227
Bmnfelsia 526, 528
Bryonia 277
Bryophyllum 276
Bubroma 532
Buceras 254
Buchenavia 226, 254-255
Buchia 493
Buchnera 522-524
Bucholzia 188
Bucida 254-255
Buena 494
Buforrestia 85
Bulbophyllum 118-119, 135, 137
Buibostylis 88, 95-102
Bulbulus 160
Bumelia518
Bunchosia 386, 390-391
Buphthalmum 261
Burdachia 386, 390, 392
Burlingtonia 135
Burmannia 83-84
Burmeistera 233
Bursera 227, 229
Burseria 229
Buttneria 532
Buxus 231
Byrsonima 386-387, 389-391
Byttneria 531-532
Caballeria432
Cabomba 231
Cabralea418
Cabrera 1 60
Cacalia 261
Cacao 532
Caconapea 523
Cacoucia 255
Cactus 232
Caesalpinia 334, 341, 344-345
Cajanus 348, 356, 358
Caladium 60, 64
Calathea 114-116, 182
Caldasia213
Calea 256-257, 261-262, 264, 266-267
Calinea281
Calliandra 366, 370-371, 373-374
Callicarpa 545
Callichlamys 215, 218-219
Callicocca 494
Callisia 85-86
Calocarpum 518
Caiolisianthus 306
Calonyction 273
Calophyllum247, 251
Calophysa 407
Calopogonium 348, 363
Calotropis209, 211
Calpidisca 377
Caluera 119
Calycampe 439
Calycobolus 271, 273-275
Calycolpus 434, 439,441
Calycophyllum 479, 494, 503, 505
Calycorectes 434, 439
Calyptranthes 434-435, 439, 442
Calyptrion 548
Calyptrocarya 88, 95
Calyptrostylis 95
Calystegia 273
Camarea 387
Camaridium 135
Cameraria 200
Campelia 85
Campnosperma 189
Campomanesia 435, 439, 442
Campsiandra 334-335, 341
Camptouratea 448
Campylocentrum 119, 134-135
Campyloneurum 45-49
Campylosiphon 83
Cananga 193
Canavalia 348, 356, 359
Candolleodendron 348, 363
Canna 84
Cansenia 341
Caopia 251
Caperonia 290, 296
Capirona 479, 494
Capiteilaria 407
Capparidastrum 234
Capparis 234-235
Capraria 522-523
Capriola 160
Capsicum 526, 528
Caraguata 79
Caraipa 247-248, 251
Carapa418
Carapichea 494
Caraxeron 1 88
Cardamine 226
Cardiopetalum 191, 193
Cardiospermum 511, 514
Cardonaea 262
568 Checklist of the Plants of the Guiana Shield
Carex 88-89, 95-96
Carica 236
Carludovica 87
Carolinea 222
Carphobolus 262
Carpodiptera 539
Carpotroche 301-303
Carptotepala 106
Caryocar 236-237
Caryochloa 160
Caryomene 420
Cascarilla 494
Casearia 301-304, 548
Casimirella 318-319
Casinga 303
Cassebeera 22, 24, 51
Cassia335, 341-343, 345
Cassine 240
Cassipourea 477
Cassupa 494
Cassytha 323, 327
Castalia 444
Castilla 423
Castilleja 523
Casuarina 237
Catasetum 119, 135-136, 143, 149
Catharanthus 196,201,203
Cathartocarpus 343
Cathedra 450
Cathormion 370
Catinga 439
Catocoma 470
Catopsis74, 79, 81-82, 182
Catostemma 220, 222, 539
Cattleyall9, 136, 149
Caularthron 136, 138-139
Caulotretus 343
Cavanillesia 220
Cavendishia 285, 287-288
Cayaponia 276-278
Cecropia 237-238
Cedrela 418-419
Cedrelinga 366, 374
Cedrus 539
Ceiba 220, 222
Celastrus 240
Celianella 290
Celiantha 304, 306-307
Celosia 187-188
Celtis 542
Cenchrus 151, 160
Centratherum 257, 261-262
Centrogenium 136
Centroglossa 136
Centrolobium 348, 356
Centronia 397, 407
Centropogon 233
Centrosema 348, 356-357, 362
Centrosolenia 3 1
1
Centrospermum 262
Centunculus 473
Cephaelis 494-495
Cephalocarpus 89, 96
Cephalocereus 232
Cephalodendron 495
Cephaloschoenus 96
Cephalostemon 173, 175
Cephalostigma 233
Ceradenia 35-38, 47-49
Ceratoccoca 457
Ceratocephalus 262
Ceratochilus 136
Ceratophyllum 240
Ceratophytum215, 218
Ceratopteris 45, 51
Ceratosanthes 276-278
Ceratostema 285, 287
Cerbera 200
Cercophora 331
Cercouratea 448
Cerdana 225
Cereus 231-233
Cespedesia 445, 448
Cestrum 497, 526, 528
Chaenanthe 136
Chaenophora 407
Chaenopleura 407
Chaetaria 160-161
Chaetocalyx 348, 357, 362
Chaetocarpus 290, 296-297
Chaetochlamys 185
Chaetochloa 161
Chaetocrater 303
Chaetogastra 407
Chaetolepis 407
Chaetospora 96
Chailletia 280
Chalcas 509
Chalepophyllum 479, 495
Chamaecrista 335-336, 341-344
Chamaedorea 70
Chamaefistula 343
Chamaesenna 343
Chamaesyce 296
Chamelophyton 136
Chamissoa 187-188
Chaptalia 257, 262, 264, 267
Charianthus 407
Chatiakella 262
Chaubardia 119, 136, 140
Chaubardiella 119, 136, 148
Chaunochiton 450
Cheilanthes22-24, 29, 50-51
Cheiloclinium 313-316
Cheiradenia 119, 136
Cheiroglossa 45
Chelonanthus 304, 306-308
Chelyorchis 120, 143
Chenopodium 240
Chevaliera 79
Chimantaea 257, 262, 267
Chimarrhis 479, 495, 499
Chiococca 495, 497
Chionanthus 45
1
Chionolaena 257, 262, 264
Chitonia 407
Chloris 151, 158, 161
Chlorocardium 7, 323, 329
Chloroleucon 366, 373-375
Chlorophora 425
Chomelia 480, 493, 495
Chondrococcus 495
Chondrodendron 42
1
Chorisepalum 304-306
Christella 54
Christiana 538-539
Chromolaena 257, 263-264, 266
Chromolucuma 515
Chrysobalanus241,244
Chrysochlamys 248, 253
Chrysophyllum 57, 515, 518-521
Chrysothemis 309, 311-312
Chuncoa 255
Chupalon 287
Chuquiraga 262
Chusquea 151, 161
Chylodia 262
Chytroma331
Chytropsia 495
Cienfuegosia 393-395
Cinchona 495
Cinchonopsis 480, 495
Cinnamodendron 234
Cinnamomum 323, 327, 329
Ciponima 535
Cipura 1 1
1
Cissampelos 420-421
Cissus 551-553
Citharexylum 544-545
Citrosma 526
Citrullus 276-278
Citrus 507
Cladium 89, 96
Cladobium 136
Cladocolea 381-384
Cladostyles 273
Clarisia 423, 425
Clathrotropis 348, 357-358
Clavapetalum 3 1
9
Clavija 537
Claytonia 306
Cleidion 296
Cleistes 120, 136, 141, 146
Clematis 476
Cleome 234-235
Clerodendmm 544-545, 547
Clethra 247
Clibadium 257, 261-262
Clidemia 397-398, 407-411, 414-416
Clinopodium 321
Clinostemon 327
Clitoria 348-349, 357, 361, 364
Clonodia 387, 390-392
Clowesia 120, 135-136
Clusia 248-249, 251-252
Clusiella249,251
Clusiophyllum 296
Clytostoma215, 217
Cnemidaria 26-28, 48
Cnestidium 269-270
Cnidoscolus 290, 297
Coccinia 276
Checklist of the Plants of the Guiana Shield 569
Coccochondra 480, 495
Coccocypselum 480, 493, 495, 497,
502-503, 505
Coccoloba 471-472
Cocculus 421
Cochleanthes 120, 136, 138
Cochlidium 35-37, 47-49, 51
Cochlospermum 220, 222
Cocos 68, 70
Codonanthe 309, 311-312
Codonanthopsis 309-311
Codonemma 200
Coelorachis 151, 161, 163-164, 170
Coeppertia not found
Coffea 480, 495, 497
Cohniella 136
Coix 151
Coleosanthus 262
Coleostachys 387, 390
Collaea 357
Collophora 200
Colocasia 60, 64
Colubrina 477
Columnea 309-312
Comanthera 106
Combretum 254-255, 382
Commelina 85
Commelinopsis 85
Commianthus 495
Commiphora 227, 229
Comocladia 189
Comolia 398-399, 407-408, 410, 414-
417
Comoliopsis 399
Comparettia 120
Complaya262
Compsoneura 428-430
Conami 296
Conceveiba 290, 296, 300
Conceveibastrum 296
Conceveibum 296
Conchocarpus 507-509
Condalia 495
Condylidium 257, 263-265
Condylocarpon 196, 199-202
Connarus 269-270
Connellia 74, 79-81
Conobea 523-524
Conocarpus 254-255
Conoclinium 262
Conohoria 548
Conomorpha 432
Conoria 200
Conosiphon 495
Conostegia 399, 408
Conuleum 526
Convolvulus 271, 273-275
Convulvulus 274
Conyza 257, 262-264
Copaiba 343
Copaifera 336, 343
Copaiva 343
Copedesma 408
Copemicia 68, 70
Coptophyllum 52
Corallocarpus 277
Corchorus 538-539
Cordia 224-226
Cordiera 480, 492, 495-496
Cordyline 58
Coreopsis 262
Comutia 544-545
Coronilla 357
Coronopus 226
Cortaderia 151, 159
Coryanthes 120, 136, 140
Corymborkis 120
Corynostylis 547-549
Corypha 70
Coryphadenis 408
Coryphothamnus 480, 498
Corythophora 330-332
Corytoplectus 309-3 11
Cosmibuena 480, 494-496
Cosmos 257, 261
Costus86, 182
Cottendorfia 79-80
Cotula 262
Coublandia 357
Couepia 241, 244-247, 251
Couma 196, 200,203
Coumarouna 357
Coupoui 496
Couratari 330-333
Courimari 284
Couroupita 330, 332
Coursetia 349, 357, 360, 362
Coussapoa 237-239
Coussarea 480, 482, 493, 496
Coutarea 480, 496, 499
Coutoubea 305-306
Cranichis 120, 136
Craniolaria 456
Crantzia 3 1
1
Crataegus 478
Crateva 234-235
Cremanium 4-8
Cremastosperma 191, 193
Cremastus 218
Crematomia 226
Cremersia 309
Crenea 384-385
Crepidospermum 227, 229
Crepinella 207
Crescentia 215
Crinum 112-113
Crotalaria 349, 357, 359
Croton 290-291, 296-297
Crudia 245, 336, 340, 343-346
Cryptangium 96
Cryptarrhena 120, 136
Cryptocalyx 545
Cryptocarya 323, 327
Cryptocentrum 120, 136
Cryptochloa 150, 161, 170
Cryptophoranthus 136
Cryptostegia 209
Ctenanthe 114, 116
Ctenardisia 430, 432-433
Ctenitis 32-33, 52-53
Ctenium 151
Ctenopteris 36
Cucullaria 554-555
Cucumis 276-277
Cucurbita 276-277
Cuervea 313-315
Culcita31
Cunuria 296-297
Cupania 511, 514
Cuphea 384-385
Cupirana 496
Curanga 523
Curarea 420-421
Curatella 280-281
Curculigo 112-113
Curtia 305-306, 308
Cuscuta 279
Cusparia 509
Cuspidaria 215
Cyanthillium 257, 261-262, 268
Cyathea 26-28, 48-49
Cyathoglottis 136
Cyathula 187-188
Cybianthus 205, 430-434
Cybistax215, 218
Cycas 56
Cyclanthus 87
Cyclodium 31-34, 49
Cyclolobium 357
Cyclopeltis 49, 52
Cyclopogon 120, 135-136, 144, 146-
147^
Cyclosorus 54
Cycnoches 120, 136
Cydista 215, 217-218
Cylindrosperma 200
Cymbidium 136
Cymbocarpa 83
Cymbopetalum 191, 193-194
Cymbopogon 151, 161
Cymbosema 349, 358
Cynanchum 209, 211-212
Cynodendron 518
Cynodon 151, 160, 165
Cynometra 336, 344
Cynomorium 213
Cynosurus 161
Cypella 1 1
1
Cyperus 89-90, 95-96, 98-99, 101-102
Cyphomandra 526, 528, 530
Cypripedium 136
Cyrilla 279
Cyrillopsis 319-320
Cyrtocarpa 189-190,229
Cyrtocymura 257, 262, 268
Cyrtopera 136
Cyrtopodium 120, 136
Cyrtospemia 60, 64-65
Cyrtospermum 189
Cytisus 358
Dacryodes227, 229, 231
Dactyloctenium 151, 161
570 Checklist of the Plants of the Guiana Shield
Dactylostegium 185
Dactylostemon 297
Dahlgrenia 70
Dalbergaria 3 1
1
Dalbergia 349, 356, 358-360, 362
Dalechampia 291-292, 297-299
Danaea 44, 50
Daphnopsis 537-538
Dasynema 284
Dasyphyllum 257, 262
Datura 526, 528
Davallia 29
Davalliopsis 39
Davilla 280-281
Davya 409
Debraea 555
Decagonocarpus 507, 509
Decaraphe 409
Declieuxia 226, 480, 496-497
Degranvillea 120
Deguelia 358
Delilia257
Delonix 336, 345
Dendrobangia 3 1 8-3 1
9
Dendrobium 136
Dendropanax 205, 207
Dendropemon 383
Dendrophthora 383, 549-551
Dendrosipanea 480-481, 496
Dendrostylis 303
Dendrothrix 292, 300
Dennstaedtia 29-31
Deprea 527-528
Dermatophyllum 262
Derris 349, 358, 360
Desmanthus 366, 370
Desmochaetal88
Desmodium 349-350, 358, 360-361
Desmoncus 68, 70
Desmoscelis 399, 407, 409, 411, 414-
415
Diacidia 387, 390, 392
Diacrium 136
Diadenium 120, 136
Dialium 336, 340, 344
Diandrochloa 161
Dianella 113
Dianthera 185
Diastema 309
Diazeuxis 262
Dibrachion 358
Dicella 387, 390, 392
Dichaea 120-121, 136-138, 141
Dichanthelium 1 5 1 , 1 64- 1 67
Dichanthium 151, 158
Dichapetalum 226, 279-280
Dichorisandra 85
Dichromena 96-97, 101, 121
Dichrostachys 366
Dicksonia 31
Diclidanthera 469-470
Diclinanona 191, 193-194
Dicliptera 183, 185
Dicorynia 336, 344
Dicrananthera 409
Dicranoglossum 46-47, 49
Dicranopteris 34
Dicranopygium 87
Dicranostyles 271, 274-275
Dicrobotryum 496
Dicrypta 137
Dictyocaryum 68, 70-71
Dictyostega 83-84
Dicymbe 336
Dicypellium 323
Didactyle 137
Didymiandrum 90, 96-97
Didymochlaena 23, 31-32
Didymochlamys 481
Didymoglossum 39
Didymopanax 207
Diectomis 161
Dieffenbachia 60, 64-66
Digitaria 151-152, 158, 161, 165-168,
170-172
Digomphia215, 218
Dikylikostigma 137
Dilkea 454-455
Dilodendron 511, 514
Dimerandra 121, 137, 139, 141, 143
Dimerocostus 86
Dimorpha 344
Dimorphandra 336-337,344
Dinizia 337
Dioclea350, 356, 358-359
Diodella 496
Diodia 481, 492-493, 496-497, 504
Diolena 409
Diomma 509
Dioscorea 103-104
Diospyros 282-283
Diothonea 137
Diphasium 18
Diplacrum 90, 97, 99, 102
Dipladenia 200
Diplasia 90, 97
Diplazium 25, 54-56
Diplochita 409
Diplocrater 450
Diploon515, 518
Diplopterygium 34
Diplopterys 387, 389-391
Diplotropis 345, 350, 356, 358
Dipteracanthus 185
Dipteranthus 121, 137, 144
Dipterodendron 514
Dipteryx 350, 357-358, 363
Disciphania 420-421
Discocarpus 292, 297
Discophora 318-319
Discyphus 121, 137, 147
Disocactus 231, 233
Disteganthus 74, 78-80,
Disterigma 285-286, 288
Distictella 215, 217-218
Distictis 215, 217-219
Ditassa209, 211
Ditaxis 292, 296
Ditmaria 555
Ditomaga 306
Ditremexa 344
Dodecas 385
Dodecastigma 292, 297-298
Dodonaea 511
Dolicholus 359
Dolichopterys 390
Dolichos 359
Doliocarpus 280-282
Dorstenia 423, 425
Doryopteris 22-24, 5
1
Doyerea 276-277
Dracontium 60-61, 65
Drejera 185
Drepanocarpus 359
Drepanospermum 189
Drimys 555
Drosera 282
Drupina 523
Dryadella 121
Drymaria 237
Drymonia 309-3 11
Drynaria 47
Dryopteris 32-33
Drypetes 292, 297
Duboisia 137
Duckea97, 101, 173, 175
Duckeella 121, 137
Dufourea 274, 468
Dugandiodendron 385-386
Duguetia 191, 193
Duidaea 257
Duidania481
Dulacia 450-451
Dunstervillea 121
Duranta 544-546
Duroia 481, 492-493, 496
Dussia 350, 359-360
Duvemoia 185
Dyckia 80
Dyspemptemorion 185
Dystovomita 249
Ebermaiera 185
Ecastaphyllum 359
Ecclinusa 515, 518-520
Eccremis 112-113
Echeandia 112-113
Echinella 137
Echinocassia 344
Echinocephalum 262
Echinochloa 152, 164-167
Echinodorus 59
Echinolaena 152, 160, 165
Echinops 262
Echinoschoenus 97
Echinosepala 137
Echites 200-201
Eclipta 257, 262, 267-268
Ectozoma 528
Egleria 90
Egletes 257, 262
Eichhomia 173
Elachyptera 313-314, 316
Checklist of the Plants of the Guiana Shield 571
Elaeagia 481
Elaeis 68
Elaeodendron 239-240
Elaeoluma433, 515, 518-520
Elaeophora 297
Elaphandra 257
Elaphoglossum 42-43, 45, 50-51
Elaphrium 229
Elaterium 277
Eleocharis 101-102
Elephantomene 421
Elephantopus 257, 262-263
Eleusine 152, 161
Eleutheranthera 257, 264-265, 268
Eleutherine 1 1
1
Elionurus 152, 158, 161, 163
Elizabetha 337, 244
Elleanthus 121, 134, 137, 140, 147
Elodea 110-111
Elopium 65
Eloyella 121, 144
Eltroplectris 121, 136, 143-144
Elvasia 445, 448-449
Elytraria 183, 186
Emilia 257, 261, 263
Emmeorhiza481,493
Emmotum 318-319
Encholirium 80
Enckea 46
1
Encyclia 121, 136-139, 141
Endlicheria 323, 326-329
Endopleura 316, 318
Englerella519
Enicostema 305-306
Enourea 514
Entada 366, 370-371,373
Entadopsis 370-371
Enterolobium 366, 371,375
Enteropogon 152, 161-162
Enterosora 35-37, 47-49
Eperua 337, 344-345
Ephedranthus 191, 193
Ephippiorhynchium 97
Epidendropsis 138
Epidendrum 121-123, 135, 137-139,
141, 143-144, 147
Epidryos 173
Epigaea 287
Epiphyllum 231-233
Episcia310-311
Epistephium 123, 139-140
Eragrostis 152, 161-164, 169-170
Eranthemum 183, 185
Erechtites 257, 266
Eremolepis 285
Eremoluma 519
Erianthus 162
Erigeron 257, 263
Erinus 523
Eriocaulon 104-107
Eriochloa 152, 162, 164
Eriochrysis 152, 158, 170
Eriope 321
Eriopsis 123, 140, 146
Eriosema 350, 358-360, 362
Eriosorus 23-24, 36
Eriotheca 220-222
Erisma 553, 555
Emestia399,407,409, 415
Erosion 162
Eroteum 536
Ertela 507, 509
Ervatamia201
Erycina 123, 139, 143, 146
Eryngium 195
Erythrina 350, 359, 361
Erythrochiton 507
Erythrodes 123, 140, 144
Erythroxylum 288-289
Eschatogramme 47
Eschweilera 330-333
Escobedia 523
Esenbeckia 507, 509
Esmeraldia 2 1
1
Etaballia 350, 359-360
Ethulia 263
Eubrachion 285
Euceraea 301
Euchlaena 162
Eugenia 435-436, 438-442
Eulophia 123, 136, 141
Eulophidium 140
Euosmia 496
Eupatorium 263-264
Euphorbia 292, 296-297, 299
Euphronia 301
Euplassa 474
Eurya 536, 542
Eurychaenia 409
Eurydochus 264
Eurystyles 123, 148
Euterpe 68, 70
Evea 496
Evelyna 140
Everardia 91, 97
Evodianthus 87-88
Evolvulus 271, 273-274
Evonymodaphne 327
Exacum 306
Excentradenia 387, 391
Exellodendron 241, 244, 246
Exochogyne91,97, 99
Exoecaria 425
Exostyles 337
Exothostemon 201
Fagara 509
Faramea 481-482, 495-497, 504
Farringtonia 409
Feea 39
Ferdinandusa 482
Femandezia 140
Ferolia 244, 425
Festuca 162
Feuillea 371
Feuilleea 371
Fevillea 276-278
Ficus 423-428
Fimbristylis 91, 97-99, 101-102
Fischeria209, 211
Fissicalyx 350, 361
Fittonia 185
Flacourtia 303
Fleischmannia 257, 263
Flemingia 350
Fleurya 542
Floscopa 85
Folianthera 371
Forsteronia 196, 199, 201, 203
Fothergilla 409
Fractiunguis 140
Franchetella 519
Franciscea 528
Freziera 535, 536
Froelichia 187-188
Froesia 475
Froesiochloa 152
Froesiodendron 193
Fugosia 394
Fuirena91,98, 102
Funastrum 2 1
1
Furarium 382
Furcraea 58
Fusaea 191, 193-194
Gaedawakka 297
Gaiadendron 381-384
Galactia 350-351, 357, 359-360
Galactodendrum 426
Galactophora 196, 199-200
Galeandra 123-124, 140, 144
Galega 359
Galeottia 124, 143, 148
Galipea 507, 509
Galium 482, 497, 502-503, 505
Galphimia 390
Gamochaeta 258, 264
Gamotopea 497
Garapatica 497
Garayella 140
Garcinia 250, 252
Gardenia 497
Garuga 1 89
Gaultheria 286-288
Gaura 255
Gavarretia 292, 296
Gaya 393
Gaylussacia 286-287
Gayoides 394
Geissopappus 264
Geissospermum 196
Genipa 482, 497, 507
Genlisea 376-377
Gentiana 306
Geoffroea 360
Geonoma 68, 70-7
1
Geophila 482, 497-498, 501, 503
Gerardia 523
Gesneria 31
1
Ghinia 546
Gibasis 85-86
Gilibertia 207
Gimbernatea 255
Gitara 297
572 Checklist of the Plants of the Guiana Shield
Glandonia 387
Glaphyropteris 54
Glaziostelma 211
Gleasonia 482, 497
Gleichenia 34
Glinus 422
Gliricidia351,362
Globifera 523
Glochidion 297
Gloeospermum 547-548
Glossarion 258, 264
Glossocentrum 409
Glossostylis 523
Gloxinia 310-311
Glycine 360
Glycoxylon 519
Glycydendron 292
Glyphotaenium 36
Gnaphalium 258, 264
Gnetum 57
Gochnatia 258, 262, 264, 266-267
Godmania 215, 217
Godoya 448
Goeppertia 327
Gomidesia 440
Gomphia 448
Gomphichis 124, 148
Gomphiluma 519
Gomphrena 187-188
Gongora 124, 140
Gongylolepis 258, 261-262, 264, 267
Goniophlebium 47
Goniopteris 54
Gonolobus209, 211
Gonzalagunia 482, 497
Goodallia 537
Goodyera 140
Gordonia 536-537
Gossypium 393-394
Gouania 477
Gouinia 152, 172
Goupia 239-240
Govenia 124
Graffenrieda399, 409, 415
Grammadenia 433
Grammitis 35-38, 47-49
Granatum 418
Gratiola 523
Gravisia 80
Greeneina 426
Greenmania 264
Grias 332
Grimaldia 344
Grislea 385
Guacamaya 173
Guadual52, 160, 162
Guaiacum 556
Guaicaia 264
Guanchezia 124, 135
Guapeba 519
Guapira 443-444
Guarania 297
Guarea 304, 418-419
Guatteria 191-194
Guatteriopsis 192-193
Guayania 258, 262-264
Guazuma 531
Guenetia 539
Guettarda 482-483, 493, 496-497, 507
Guidonia 303
Guildingia 409
Guilielma 71
Gurania 276-278
Gustavia 331-333
Guzmania 74, 79-82, 182
Gymnanthes 292, 296-297
Gymnobalanus 327
Gymnogramma 23
Gymnogramme 55
Gymnopogon 152, 161-162, 170
Gymnopteris 23
Gymnosiphon 83-84
Gynandropsis 235
Gynerium 152, 162, 170
Gynestum 71
Gynocraterium 183
Habenaria 124-125, 135, 140, 144
Hackelochloa 152, 160, 163
Haemadictyon 201
Haematostemon 292, 296
Haematoxylum 337
Haemocharis 536
Haloragis 312
Haloschoenus 98
Hamelia 483, 497
Hancornia 201
Handroanthus 218
Haploclathra249, 251
Haplolophium 215
Haplostylis 98
Hartigia 409
Hasseltia301
Havetia 252
Havetiopsis 252
Hawkesiophyton 527-528
Hebanthe 188
Hebeclinium 258, 261,263
Hebepetalum 316
Hecastophyllum 360
Hecatostemon 301-303
Hecistopteris 55
Heckeria 461
Hedwigia 229
Hedychium 182
Hedyosmum 240
Hedyotis 485, 497
Hedysarum 360
Heisteria 450-451
Helia 306-307
Heliamphora 521-522
Heliconia 110, 117
Helicostylis 424, 426-427
Helicteres 531-532
Heliotropium 225-226
Helmia 104
Helmontia 276-278
Helonoma 125, 135-136, 142, 147
Helopus 162
Helosis 212-213
Hemarthria 152, 170
Hemicarpha 98
Hemicrepidospermum 229
Hemidictyum 25, 56
Hemidiodia 497
Hemigraphis 183
Hemionitis 23, 51
Hemitelia 28
Henriettea 399-400, 409-411, 414
Henriettella 400, 409-41 1, 414
Henriquezia 483, 497
Hepetis 80
Herissantia 393-395
Hermesia 297
Hermesias 344
Hemandia 312
Herpestis 523-524
Herpetacanthus 183, 185
Herrania 531-533
Hesperomeles 478
Heteranthera 173
Heterocondylus 258, 264
Heteropetalum 192, 194
Heteropogon 152, 158, 162
Heteropsis 61, 65-66
Heteropterys 387-391
Heterostemon 337, 344
Heterotaxis 125, 137, 142-143
Heterothalamus 264
Heterotrichum 409
Hevea 292, 297, 300
Hexadesmia 140
Hexapterella 83-84
Hexisea 125, 137, 140
Heymassoli 450
Hibiscus 393-395
Hicriopteris 34
Hieronyma 292, 297, 300
Hillia483,497, 503
Himatanthus 196,201-202
Hippeastrum 112-113
Hippobroma 233
Hippocratea 313-314
Hippomane 297
Hiraea 388-389, 391
Hirtella 241-242, 244-247
Hisingera 303
Histiopteris 29, 51
Hoehneella 140
Hoffmannia 483
Hohenbergia 74, 79
Holcus 162
Holosteum 237
Holstianthus 483
Homalium 301-304
Homalocenchrus 162
Homalomena 61
Homoeotes 39
Homolepis 152-153, 164-166
Hoppia 98
Hormocalyx 409
Homschuchia 194
Hortia 507
Checklist of the Plants of the Guiana Shield 573
Hortsmania 201
Hostmannia 448
Houlletia 125, 141
Houmiri 317
Houssayanthus 511,514-515
Houstonia 497
Huberodendron 220
Huberopappus 258
Huebneria 141
Humboldtia 141
Humboldtiella 360
Humiria 316-317
Humirianthera 3 1
9
Humiriastrum 317-318
Humirium 3 1
7
Huntleya 125, 141
Huperzia 17-19
Hura 292
Hybanthus 547-549
Hybochilus 141
Hydranthelium 524
Hydrochorea 366, 370-371, 373-376
Hydrocleys 114
Hydrocotyle 195
Hydrolea318
Hydromystria 111
Hygrophila 183, 185
Hylaea 196,202
Hylaeanthe 114, 116
Hylaeorchis 125, 135, 142
Hylenaea 313-314
Hylocereus 231-233
Hymenachne 153, 162, 164-165
Hymenaea 337, 344
Hymenocallis 112-113
Hymenodium 43
Hymenolobium 351
Hymenophyllopsis 25, 40-41
Hymenophyllum 38-41
Hymenostachys 40
Hyospathe68, 71
Hyparrhenia 153, 158, 161, 172
Hypenia321,322
Hyperbaena 421
Hypericum 249, 252
Hypobrichia 385
Hypocyrta 312
Hypogynium 162
Hypolepis 23, 29
Hypolytrum 91-92, 98, 101-102
Hypoporum 98
Hypoxis 112-113
Hyptidendron 321
Hyptis 321-322
Ibatia212
Ibetralia 483, 492
Icacorea 433
Ichnanthus 153, 159, 162-166
Ichthyophora 519
Ichthyothere 258, 264, 266
Icica 229
Icicopsis 229
Ilex 203-205, 426
Illecebrum 188
Ilysanthes 524
Imeria 258, 263-264
Impatiens 2 1
3
Imperata 153, 163, 170
Indigofera351,356, 360
Inga 366-367, 371-374
Inula 264
lonidium 548
lonopsis 125, 139, 141
Ipomoea 271-275
Iresine 187-188
Iria 98
Iriartea 68,71
Iriartella68, 71
Iris 111
Irlbachia 305-308
Iroucana 303
Iryanthera 429-430
Isachne 153, 163, 166-167
Ischaemopogon 163
Ischaemum 153, 158-163
Ischnosiphon 114-116
Iseia 272-275
Isertia 483, 493-494, 497, 505
Isidrogalvia 112-113
Isochilus 125, 139,141
Isoetes 17
Isolepis 98
Isoloma 312
Isotoma 233
Itea 279
Ithycaulon 30
Ivira 532
Ixialll
Ixidium 285
Ixocactus 381-382
Ixophorus 153
Ixora483,497, 499
Jablonskia 292, 299-300
Jacaranda 215, 217-218
Jacaratia 236
Jacobinia 185
Jacquemontia 273-275
Jacqueshuberia 337
Jacquinia 537
Jacquiniella 125, 138-139
Jambosa 440
Jamesonia 23
Japarandiba 332
Jasarum 61
Jasminum 451
Jatropha 292-293, 297
Jenmaniella 467
Jessenia 71
Jubelina 388, 390-391
Jucunda 409
Jugastrum 332
Juglans 320
Julocroton 297
Juncus 111-112
Jupunba 372
Juruasia 185
Jussiaea 452
Justicia 183-186
Kaieteurea 448
Kalanchoe 276
Kallstroemia 556
Karatas 80
Kedrostis 276
Kefersteinia 125, 141
Kegelia 141
Kegeliella 125, 141
Kieseria 224
Kippistia 314
Klaprothia 379
Kleinia 264
Klugiodendron 372
Knoxia 497
Koanophyllon 258, 263-264
Koellensteinia 125, 136, 141-142, 146,
149
Koellikeria310
Koemickanthe 115-116
Kohleria 310-312
Kopsia 196
Koyamaea 92
Krameria 320
Krugia 440
Kubitzkia 329
Kuhlia 303
Kuhlmaniella 275
Kuhnia 264
Kummeria 3 1
9
Kunhardtia 173, 175
Kutchubaea 483
Kyllinga 92, 96, 98-99
Labatia519
Lablab351,359
Lacistema 320-321, 463
Lacmellea 196,201,203
Lacostea 40
Lacunaria 475
Ladenbergia 483, 495, 497
Laelia 141
Laetia 302-304
Lafoensia 385
Lagascea 264
Lagenaria 276-278
Lagenocarpus 92, 98-99, 102
Lagerstroemia 385
Laguncularia 254-255
Lampadaria 310
Lamprococcus 80
Landolphia 201
Lanessania 426
Langsdortria212-213
Lanium 141
Lanlana 544, 546
Laplacca 536-537
Laportea 542-543
Larentia 1 1
1
Lasiacis 153, 163-167
Lasiadenia 537
Lasiandra 409
Lasiolepis 107
Lasiostoma 380
Lastrea 54
Lastreopsis 32-33, 48, 52-53
574 Checklist of the Plants of the Guiana Shield
Latreillea 264
Laurea 426
Laurembergia 312
Laurentia 233
Laurus 327, 538
Lawsonia 385
Laxoplumeria 196
Leandra 400, 407-4 11,414-417
Leaoa 141
Lecanium 40
Lecointea 337
Lecostemon 476
Lecticula 377
Lecythis 193,331-333
Ledothamnus 286-287
Leersia 153, 162-164
Leiostegia 410
Leiothamnus 307
Leiothrix 105-107
Leiphaimos 307
Leitgebia 448
Lellingeria 35-38
Lembocarpus 310
Lemna 112
Lemniscia 317
Leochilus 125, 139
Leonia 547-548
Leonotis 321-322
Leonurus 321-322
Leopoldinia 68-69
Lepanthes 125, 141
Lepanthopsis 125, 144-145, 148
Lepianthes 460-462
Lepidagathis 184-186
Lepidaploa 258, 262, 264, 268
Lepidium 227
Lepidocaryum 69, 71
Lepidocordia 225
Lepidotis 18
Leptochilus 43
Leptochloa 153, 161-163
Leptocoryphium 153, 159, 165, 169
Leptoglottis 372
Leptolobium 344, 360
Leptorchis 141
Leptorhoeo 85
Leptoschoenus 99
Leptostachya 186
Leptostachys 163
Leretia319
Leria 264
Lessingianthus 258, 268
Lettsomia 536
Leucaena 367, 370, 372-373
Leucococcus 542
Leucohyle 125, 141, 148
Leuconymphaea 444, 445
Leucopholis 264
Leucothoe 287
Libidibia 344
Licania 242-246, 251,533
Licaria 323-324, 326-329
Ligea 468
Ligeophila 125, 140, 144
Lightia301,532
Limnobium51, 110-111
Limnocharis 114
Limnosipanea 483, 497, 504
Limodorum 141
Lindackeria 302-303
Lindemia 523-524
Lindleya 537
Lindleyella 141
Lindmania 74-75, 79-80
Lindsaea 23, 29-31
Linociera451
Liparis 125, 141, 144, 148
Lipocarpha 92, 95, 98-99, 101
Lipochaeta 264
Lippia 144, 545-546
Liriosma 450-451
Lisianthus 307-308
Lissocarpa 379
Lisyanthus 308
Lithachne 153, 164
Lithocardium 226
Litobrochia 51
Lobelia 233
Lochnera 201
Lockhartia 125-126, 138, 140-141
Loevigia 410
Lomagramma 43, 48
Lomaria 26
Lomaridium 45
Lomariopsis 26, 43
Lonchitis 30-31
Lonchocarpus 351, 356, 358, 360-364
Lonicera 497
Lophanthera 388, 390
Lophiaris 141
Lophidium 52
Lophogyne 468
Lophopterys 388, 390-391
Lophostoma 537
Lophotocaqjus 59
Lopimia 395
Loranthus 382, 550
Loreya 400
Lorostemon 249
Loxopterygium 189
Lubaria 509
Lucilia 264
Lucuma 5 1
9
Ludovia 87-88
Ludwigia 451-452
Lueddemannia 126, 136
Luehea 538-539
Lueheopsis 538-539
Luetzelburgia 351
Luffa 276-278
Lunania 302
Lundia 215-218
Luxemburgia 448
Luziola 153, 160, 163
Lycaste 126, 141-142
Lycianthes 527-530
Lycomormium 14
Lycopodiella 18-19
Lycopodium 18-19
Lycoseris 258, 262, 264
Lycurus 163
Lygodium 43-44
Lyroglossa 126, 147
Lysiloma 368, 372
Lysiostyles 272, 275
Lythrum 385
Maba 283
Mabea 293, 297-298
Maburea 450
Macaglia 201
Macahanea 57, 315
Macairea 400-401, 407, 410, 414-416
Macfadyena216, 218
Machaerina 92
Machaerium 351-352, 359, 361-362
Machaonia 483, 495
Madura 424-426
Macoubea 197,201-203
Macoucoua 205
Macradenia 126, 141, 147
Macrocarpaea 305, 308
Macrocentrum 401, 409-410, 415
Macrochordion 80
Macroclinium 126, 141, 143
Macrocnemum 497
Macroditassa 209
Macrolobium 57, 337-338, 344-346,
372
Macropharynx 197, 200
Macroptilium 352, 362
Macrosamanea 368, 371-372, 374-376
Macroscepis 212
Macuillamia 524
Madarosperma 212
Magnolia 385
Maguirea 65
Maguireanthus 401
Maguireocharis 483
Maguireothamnus 483, 495, 497-498
Mahurea 249, 252
Maieta401, 410-411, 417
Malache 395
Malachra 393, 395, 532
Malanea 483-484, 498
Malaxis 126, 142
Mallophyton401
Malmea 192, 194
Malouetia 197,200-202
Malpighia388, 391
Malvastrum 393
Manabea 546
Managa 57
Manaosella 216-217
Mandevilla 197, 199-202
Manettia 484, 498
Manganaroa 372
Mangifera 189
Manglilla 433
Manicaria 69, 7
1
Manihot 293, 297-298
Manilkara 515-516, 518-519
Manisuris 163
Checklist of the Plants of the Guiana Shield 575
Manniella 142
Mansoa216-219
Mapania 92, 97-99
Mapaniopsis 99
Mapouria 498
Mappia319
Maprounea 293, 298
Maquira 424, 426-427
Marahuacaea 173, 175
Maranta 115-116
Marathrum 467-468
Marattia 44
Marcetia 401, 410, 414-415
Marcgravia 396
Marcgraviastrum 396
Margaritaria 293, 299
Margaritopsis 498
Marginaria 47
Marialvaea 252
Marila 249
Maripa 272, 275
Mariscus 99
Markea 527-528
Markleya 71
Mariierea 436-437, 439-442
Marmaroxylon 372
Marsdenia 209-212
Marsea 264
Marsilea 44
Marsupiaria 142
Marsypianthes 321-322
Martia 344
Martinella 216, 218-219
Martiodendron 338, 340, 344
Martiusia 344
Mascagnia 388-392
Masdevallia 126, 142
Matayba 511-512, 514-515
Matelea 199,210-212
Matisia 220, 222
Matourea 524
Mattuschkea 498
Mauria 1 89
Mauritia 69, 71
Mauritiella 69, 71
Maxillaria 126-127, 135-137, 142-144,
146-147
Maximbignya 69, 71
Maximiliana 71
Maximilianea 220
Mayaca 117
Mayapea 451
Mayepea 451
Mayna 302-303
Maytenus 239-240
Meborea 298
Mecardonia 523-524
Mecodium 40
Medadenium 143
Mediocactus 232
Megalastrum 32-33, 49, 52-53
Megalostylis 298
Megaskepasma 184
Megastachya 163
Meibomia 361
Meioluma 519
Meisneria410
Melaleuca 437
Melampodium 264-265
Melananthus 527
Melanoxylon 344
Melanthera 258, 261-262, 265
Melasma 524
Melastoma 410-411
Melia418
Meliandra 411
Melicoccus 512, 514-515
Melidiscus 235
Melinis 153, 163
Meliosma 510
Mella 524
Melloa216, 218
Melocactus 232
Melochia 531-533
Melothria 277-278
Melpomene 35, 37-38
Melviila 385
Memora 216-219
Menadenium 143
Mendoncella 143
Mendoncia 311,41 9-420, 423
Mendozia 420
Meniscium 54
Mentzelia 379
Menyanthes 422
Meoschium 163
Meresaldia 212
Meriania401,407, 409, 411
Merizadenia 201
Merostachys 153
Merremia 272-275
Mertensia 34-35
Merumea 484
Mesadenella 127, 143
Mesechites 197,201
Mesosetum 153, 163-167
Mesosphaerum 322
Mespilodaphne 327
Mespilus 478
Messerschmidia 226
Metalepis 210
Metastelma212
Metaxya 27, 32, 44, 49
Metrodorea 507
Meyenia 186
Mezia388, 390, 392
Mezilaurus 324, 326-327
Mezobromelia 75, 80-83
Miconia 401-404, 407-414, 416-417
Micrandra 293, 296-298
Micranthemum 523-524
Microchloa 153
Microgonium 40
Microgramma 46-49, 51
Microlepia 30
Microlicia404, 413
Micropapyrus 99
Micropholis 516, 518-521
Microphysca413
Microplumeria 197, 199-201
Micropolypodium 35-37
Micropteryx 361
Micropyxis 473
Microstachys 293, 298, 300
Microtea 457
Miersiella 83-84
Mikania 258-259, 261, 263, 265, 268-
269
Milium 164
Miltoniopsis 127
Mimosa 368-370, 372-374, 376
Mimusops 516, 518-519
Minquartia 450-45
1
Miocarpus 413
Mirabilis 443
Misanteca 327
Mitostemma 454
Mitozus201
Mitracarpus 484, 498, 504
Mitranthes 440
Mitreola 379
Mitrospora 99
Mnasium 175, 177
Mnesithea 164
Moeroris 298
Mollia 538-539
Mollinedia 422
Mollugo 422
Molongum 197, 199, 201-202
Momordica 277-278
Monachanthus 143
Monanthemum 265
Monnieria 507, 509
Monnina 469-470
Monocardia 524
Monochaetum 404, 407, 410, 413-414
Monogramma 37
Monophyllanthe 115
Monopteryx 352, 361
Monotagma 115-116
Monotrema 173, 175
Monstera 61, 65-66, 396
Montira 380
Montrichardia 61, 64-65
Moquilea 246
Mora 174-175,338,344-345
Moraea 1 1
1
Morella428
Morinda 484, 493, 495, 497-498
Moringa 428
Morisonia 234-235
Morniodes 127
Moronobea 249, 252
Morus 426
Mosannona 192, 194
Moschoxylum 419
Mostuea 379-380
Mourera 467
Mouriri 404, 409, 413, 414, 417, 439-
441
Mouroucoa 275
Moutabea 469-470
576 Checklist of the Plants of the Guiana Shield
Moutouchi 361
Mucoa 197, 199
Mucuna352, 359, 361,363
Muellera 352, 357,361,363
Muntingia 428
Murdannia 85
Murraya 507, 509
Murucoa 275
Musa81, 110, 117
Mussaenda 498
Mussatia 216, 218-219
Myanthus 143
Mycerinus 286
Myllanthus 509
Myoxanthus 127, 137, 143, 145
Myrcia 437-441
Myrcianthes 438, 441
Myrciaria 438-442
Myriaspora 404, 413
Myrica 428
Myriocladus 153-154, 164
Myriophyllum 3 1
2
Myristica 429-430
Myrmidone 413
Myrocarpus 352
Myrodendrum 3 1
7
Myrosma 115-116
Myrospermum 352
Myroxylon 303, 352, 364
Myrsine 431-433
Myrteola438, 441
Myrtiluma 519
Myrtus441
Nacibea 498
Naiadothrix 524
Najas 117
Naletonia 498
Nama318
Nanodes 143
Nanopteris 37
Napeanthus 3 1
Napimoga 303
Nasturtium 227
Nauclea 498
Naucleopsis 424-427
Nautilocalyx 310-312
Navia 75-76, 80-81, 113
Neblinaea 259
Neblinantha 305
Neblinanthera 404
Neblinaria 224
Neblinathamnus 484, 498
Nectandra 324, 327-329
Neea 443
Neeragrostis 164
Nelsonia 184, 186
Nelumbo 442
Nemaluma 520
Nematanthus 312
Neobartlettia 143
Neobertiera 484
Neocalyptrocalyx 235
Neocouma 197, 199, 202-203
Neodillenia281
Neogleasonia 224
Neolacis 468
Neolehmannia 143
Neomarica 111
Neopometia 520
Neoptychocarpus 302, 304
Neoraputia 507, 509
Neoregelia 76, 81
Neosprucea 302
Neotatea 224, 249
Neottia 143
Neoxythece 520
Nepeta 322
Nephelea 28
Nephelium 512
Nephradenia 210-211
Nephrodium 33
Nephrolepis 32, 44-45, 49
Nepsera404, 410, 414-416
Neptunia 369, 373-374
Nerium 197
Neurocarpum 361
Neurodium 46-47
Neurogramme 5
1
Neurolaena 259, 261-262, 265
Neurolepis 154, 164
Neuromanes 40
Neurophyllum 40
Neurotheca 305, 308
Newtonia 374
Nhandiroba 278
Nicolsonia 361
Nicotiana 527
Nidema 127, 137-139, 143
Nidularium 81
Nietneria 112-113
Niopa 374
Niphidium 46-47
Nissolia 361
Noisettia 547-548
Nonatelia 498
Nopalea 232
Norantea 396
Nortenia 524
Notanthera 382
Noterophila413
Notholaena 23-24
Notopleura 382, 484-485, 494, 499-
502, 505
Notopora 286
Notylia 127-128, 137, 143
Noyera 427
Nymphaea 444-445
Nymphoides 422
Nypa 69
Ochroma 220, 222
Ochthephiius 404
Ochthocosmus 319-320
Ocimum 321-322
Ocotea 324-329
Octomeria 128, 135, 143-144
Octopleura308, 413
Odonellia 272, 274-275
Odontadenia 197, 202
Odontandra419
Odontocandra 419
Odontocarya 42
1
Odontoglossum 128, 143
Odontoloma 3
1
Odontomanes 40
Odontonema 184-186
Odontosoria 30
Oeceoclades 128, 134, 140
Oedematopus 252
Oenocaqjus 69, 71-72
Oenone 468
Oenothera 452
Ogcodeia 427
Ogiera 265
01ax451
Oldenlandia 485, 497-498
Oldenlandiopsis 485
Oleandra 32, 44-45
01fersia31,33
Oliganthes 265
01isbea413
Oliveriana 128
Olmedia 427
Olmedioperebea 427
Olmediophaena 427
Olyra 154, 164
Omphalea 293, 298
Omphalobium 270
Oncidium 128, 139, 143-144
Oncoba 303
Oncostylis 99
Ondidium 128
Onoctonia 413
Onychacanthus 186
Onychium 144
Oocarpon 452
Ooclinium 266
Operculina 272-275
Ophidion 128, 136
Ophiocaryon 510
Ophioglossum 45
Ophrys 144
Opisthocentra 404
Oplismenus 154, 164-167
Opuntia 232
Orbignya 72
Orchis 144
Orchyllium 377
Orectanthe 179, 181
Oreinotinus 235
Orelia 202
Orellana 220
Oreobolus 92, 99
Oreodaphne 329
Oreopanax 205-207
Oribasia 498
Oritrophium 259
Orleanesia 128, 141
Ormoloma 30-31
Ormopteris 5
1
Ormosia 352, 357-358, 360-362
Ormosiopsis 361
Omithidium 144
Checklist of the Plants of the Guiana Shield 577
Omithocephalus 128-129, 144
Orthaea 286-287
Orthiopteris 3
1
Orthoclada 154, 158, 164, 167
Orthomene421
Orthopappus 259, 262-263
Oryctanthus 381-383
Oryctina381,383
Oryza 154, 164
Osbeckia 413-414
Oserya 467-468
Osmunda 45
Ossaea 404, 411, 413-415
Osteophloeum 429-430
Otachyrium 154
Otoglossum 129, 143-144
Otostylis 129, 134, 141, 144, 148-149
Ottonia 462
Ouratea 445-449
Ourouparia 498
Outea 345
Oxalis 453
Oxandra 192-194
Oxycaryum 92, 101
Oxymeris 414
Oxypetalum210, 212
Oxythece 520
Oyedaea 259, 264, 266, 268-269
Pachira 220-223
Pachygone 421
Pachylobus 229
Pachyloma404, 414, 417
Pachyptera 2 1
8
Pachyrhizus 352, 359
Pachystachys 184, 186
Pacouria 198,201 203
Pacourina 259
Paepalanthus 105-109
Paesia 23, 30-31
Pagaea 308
Pagamea 485, 498-499
Pagameopsis 485, 499, 506
Pakaraimaea 423
Palala 430
Palhinhaea 19
Palicourea 485, 498, 502, 506
Palma 72
Palmorchis 129, 137, 143
Paloue 2, 338
Paloveopsis 338
Paltonium 47
Pamea 255
Panax 207
Pancratium 113
Panicum 154, 164, 167, 172
Panopsis 474
Panzera 345
Papaya 236
Paphinia 129, 141-142
Paphiopedilum 144
Pappophorum 155
Parabignonia 216
Paradrymonia 3 1 0-3 1
1
Paragonia 215, 218-219
Parahancomia 198, 200-203
Paralabatia 520
Paralea 283
Paramachaerium 353, 361
Paramansoa 2 1
8
Paramicropholis 520
Paraprotium 229
Parascheelea 72
Paratheria 155, 166
Parathesis 433
Pariana 155, 167-168
Parietaria 54
Parinari 244, 246-247
Parivoa 345
Parkeria 45
Parkia 369, 372, 374, 376
Parkinsonia 338
Parodiolyra 155, 164, 166, 170
Parthenium 259
Paspalidium 155, 164-165
Paspalum 155156- 167-169, 171-172
Passaveria 520
Passiflora 454-456
Passoura 548
Passovia 382
Passowia 382-383
Patabea 486, 497, 499
Patima 486, 499, 503
Patrisia 303
Pauletia 345
Paullinia 512-514
Pausandra 293, 298
Pavonia 393-395
Paypayrola 547-548
Peckia 433
Pecluma 46-49
Pectis 259, 266
Pedicularis 524
Pedilanthus 293, 297
Pehria 385
Peiranisia 345
Pekea 236
Pelexia 129, 144, 147-148
Pellaea 24
Peltaea 393, 395
Peltapteris 43
Peltogyne 338-339, 344-345
Peltophorum 339, 341, 345, 372
Peltophyllum 177
Pennisetum 156, 160, 166, 169
Pentacalia 259, 266
Pentaclethra 369-370, 374
Pentadenia 312
Pentamerista 536
Pentascyphus 513
Pentaspatella 448
Peperomia 458-459, 461-465
Pepinia77, 81
Pera 293-294, 298, 300
Perama 486, 493, 498-499
Perebea 424, 427
Pereskia 232-233
Perianthopodus 278
Peridiscus 456
Peridium 298
Perissocarpa 447
Peristeria 129, 141, 144
Peritassa 313-316
Pemettya 286
Persea 325-327, 329
Persicaria 472
Peschiera 198-199,202
Pessopteris 47
Petagomoa 499
Petaloma 414
Petastoma 218-219
Petiveria 457
Petrea 544, 546
Petrocarya 247
Pfaffia 187-188
Phaeosphaerion 85
Phainantha 404, 407
Pharbitis 275
Pharmacosycea 427
Pharus 156, 169
Phaseolus 353, 362
Phegopteris 53
Phellocarpus 362
Phelpsiella 173
Phenakospermum 176
Phenax 542-543
Philacra 447-447
Philodendron 61-66
Philodice 105
Philonotion 66
Philotria 1 1
1
Philoxerus 188
Phlebodium 46-47, 49
Phlegmariurus 19
Phlomis 322
Phoebe 329
Phoenix 69
Phoradendron 383, 549-552
Phoxanthus 510
Phragmipedium 129, 136, 144, 147
Phragmites 156, 159, 169
Phryganocydia 2 1 6, 2 1
9
Phrygilanthus 383
Phrynium 1 16
Phthirusa 383-384
Phyla 544, 546
Phyllanthus 294, 296-299
Phyllocactus 233
Phyllocarpus 345
Phyllodes 116
Phyliopus414
Phymatidium 144
Phymatodcs 47
Physalis 527-529
Physinga 144
Physosiphon 144
Physostemon 235
Physurus 144
Phylarrhiza 81
Phytolacca 457
Picramnia 457
Picrasma 525
Picria 524
578 Checklist of the Plants of the Guiana Shield
Picrolemma 524-525
Pilea 542-543
Pilocarpus 507-509
Pilocereus 233
Pilosocereus 232-233
Pilumna 144
Pinelia 144
Pinelianthe 129, 144
Pinus 56
Pinzona 281
Pionandra 529
Piparea 304
Piper 459-466
Piptadenia 369-370, 374-375
Piptatherum 169
Piptocarpha 259, 262,264-266, 268
Piptocoma 259, 265-266
Piranhea 294
Piratinera 427
Piresia 156, 161, 164, 170
Pirigara 333
Piripea 524
Piriqueta 540-541
Piscidia 353, 360, 362
Pisonia 443-444
Pistacia 229
Pistia 63
Pitcaimia 77, 80-81
Pithecellobium 369, 371-376
Pithecoctenium 216-219
Pithecolobium 375
Pitumba 304
Pityrocarpa 375
Pityrogramma 23-24, 45, 50-51
Plagiogyria 45
Planchonella 520
Plantago 466
Plathymenia 369
Platonia 249
Platyaechmea 8
1
Platycarpum 486, 497, 503
Platycentrum 414
Platylepis 99
Platyluma 520
Platymiscium 353, 356, 362
Platypodium 353, 362
Platysema 362
Platystele 129, 144-145, 148
Platythelys 129
Plectrophora 129, 144
Pleonotoma 216-218
Pleopeltis 37, 46-48
Pleroma414
Pleurisanthes 319
Pleurostachys 92
Pleurostima 178
Pleurothallis 129-131, 134-137, 139-
141, 143-148
Pleurothyrium, 326, 329
Plinia 438, 440-441
Pluchea 259, 262, 264, 266
Plukenetia 294-297, 299
Plumbago 466
Plumeria 198, 202
Plumeriopsis 202
Poa 169-170
Pochota 222
Podocalyx 295-296, 299
Podocarpus 56
Podoluma516, 520
Podostemum 467-468
Poecilandra 447, 449
Poecilanthe 353, 356-357
Pogonia 146
Pogonophora 295, 299
Pogonorhynchus 414
Pogonostylis 99
Pogopetalum 319
Pogospermum 81
Poinciana 345
Poinsettia 299
Poiretia 353, 360
PoUalesta 266
Polybotrya31,33,47
Polycarpaea 188,237
Polycarpon 237
Polycycnis 131, 146
Polygala 469-470
Polygonum 471-472
Polylychnis 184, 186
Polymnia 266
Polyotidium 131, 141
Polypodium 46-49
Polypogon 170
Polypompholyx 377-378
Polypremnum 379
Polystachya 131, 138, 144, 146, 148
Polystichopsis 33
Polystichum 31-34
Polytaenium 55
Polythecandra 252
Ponera 146
Pontederia 173
Ponthieva 131
Poponax 375
Poraqueiba 319
Poraresia 299
Porocystis 513
Porophyllum 259, 261, 264, 266
Portesia419
Portlandia 499
Portulaca 473
Posadaea 277
Posoqueria 486, 499, 504
Possira 362
Potalia 305
Potamoganos 216, 218
Potarophytum 173
Poteranthera 404, 414
Pothomorphe 460-461, 465
Pothos 66
Pothuava81
Poulsenia 424
Pourouma 238-239
Pourretia 8
1
Pouteria 205, 516-521
Pouzolzia 542
Pradosia 234-235, 517-521
Praxelis 259, 263-264, 266
Prescottia 131, 136, 146
Prestoea 69-70, 72
Prestonia 198,200-202
Prevostea 275
Prieurella 521
Prionostemma 313-316
Pristimera 313-315
Priva 544, 546
Prockia 302, 304
Proclesia 287
Promenaea 146
Prosthechea 131, 134, 137-139, 146
Protium 227-230, 508
Prunus 240, 478
Psadacocalymma 186
Psammisia 286-287
Pseudabutilon 394-395
Pseudanamomis 438, 441
Pseudechinolaena 156
Pseudelephantopus 259, 263, 266
Pseudephedranthus 192-193
Pseuderanthemum 184
Pseuderiopsis 146
Pseudima 513-514
Pseudobaccharis 266
Pseudobombax 221
Pseudocalymma 216-217, 219
Pseudocassia 345
Pseudochimarrhis 499
Pseudocladia 521
Pseudoconnarus 269-270
Pseudogynoxys 259, 266
Pseudolabatia 521
Pseudolmedia 424, 427
Pseudolycopodieila 19
Pseudomaxillaria 146
Pseudopaegma 219
Pseudopiptadenia 369-370, 374
Pseudosamanea 369-370, 372, 375-
376
Pseudosenefeldera 299
Pseudosorocea 427
Pseudoxandra 192-194
Pseudoxythece 521
Psidium 438, 440-442
Psiguria 277
Psila 266
Psilocarya 99
Psilochilus 131, 146
Psilogramme 24
Psilotum 17, 19
Psittacanthus 381-383
Psychopsis 131, 143-144, 146
Psychotria 486-489, 494-505
Psygmorchis 146
Pterandra 389
Pterichis 131
Pteridanetium 55
Pteridium 30-31, 51
Pteris 31, 50-51
Pterocarpus 353, 361-362
Pterocaulon 259, 261-262, 266
Pterogastra404, 414
Checklist of the Plants of the Guiana Shield 579
Pterolepis 404, 407, 414-415
Pteroscleria 99
Pterozonium 23-24, 51
Ptilophyllum 40
Ptychocarpus 304
Ptychomeria 84
Ptychopetalum 450
Pueraria 353
Pulchranthus 184, 186
Purdiaea 279
Puya77, 81
Pycreus 99
Pyrostegia216
Pyrrorhiza 109
Quadrella 235
Quadricasaea 202
Qualea 553, 555
Quamoclit 275
Quapoya 249, 252
Quararibea 221-222
Quassia 522
Quebitea 465
Quekettia 131, 146
Quelchia 259
Quesnelia 81
Quiina 475
Racaria 514
Racinaea77, 79, 81-82
Racoubea 304
Raddial56, 170-171,315
Raddiella 156, 164-165, 170
Radia 178
Radlkoferella521
Ragala521
Ragatelus 40
Raimondia 194
Ramatuela 255
Ranaria 524
Randia 489, 493, 497-498, 502
Rapanea 433
Rapatea 173-175
Raphia 72
Raputia 508-510
Rauia 508-509
Rauvolfla 198-199,202
Ravenala 176
Ravenia 509
Raveniopsis 508-509
Rechsteineria 312
Recordoxylon 339, 344-345
Redutea 395
Rehia 156, 160
Reichenbachanthus 131, 140, 146-147
Reimaria 170
Reimarochloa 156-158, 170
Relbunium 502
Remijia 489-490, 494-495, 497, 502
Remirea 92, 96, 99
Renealmia 116, 182
Renggeria 252
Rengifa 252
Restrepia 146
Restrepiella 146
Restrepiopsis 131, 146
Retiniphyllum 490, 495, 498-499, 502-
504
Reussia 173
Rhabdadenia 198,200
Rhabdodendron 476
Rhacodiscus 186
Rhadinocarpus 362
Rhamnus 477, 542
Rhaptostylum 45
1
Rheedia 250-253
Rhexia 414-415
Rhigospira 198-199,202
Rhipidia 202
Rhipidocladum 157, 159
Rhipidopteris 43
Rhipsalis 232-233, 327
Rhizobolus 236-237
Rhizophora 477-478
Rhodocactus 233
Rhodognaphalopsis 222-223
Rhodospatha 63, 66
Rhodostemonodaphne 326-329
Rhodothyrsus 299
Rhoeo 85
Rhoogeton310, 312
Rhopalostylis 299
Rhynchanthera 404-405, 410, 413, 415
Rhynchelytrum 157, 163, 170, 172
Rhynchocladium 92, 96
Rhyncholacis 467-468
Rhynchopera 146
Rhynchosia 353, 359-360, 362
Rhynchospora 92-103
Rhytachne 157, 161, 163-164, 170
Rhytiglossa 186
Riana 548
Richardella521
Richardia 490, 503
Richardsonia 503
Richeria 295, 297, 299
Ricinella 299
Ricinus 295
Riedelia 287
Riedlea 533
Riencourtia 259-260, 262, 266-267
Rinorea 547-549
Rinoreocarpus 548
Rittera 362
Rivina 457
Robergia 270
Robinia 362-363, 514
Rodriguezia 131, 135, 146
Roentgenia 216-217, 219
Rogersonanthus 305-308
Rohria 280
Rolandra 260, 262, 266
Rollinia 192-194
Ronabea 490, 495, 498, 500-501, 503
Rondeletia 490, 503
Rondonanthus 105, 107-109
Ropourea 283
Roraimanthus 449
Rorippa 227
Rosenbergiodendron 490, 498, 502
Roshevitzia 170
Rotala 385
Rottboellia 157, 170
Roucheria 316
Rouhamon 380
Roulinia 212
Rouliniella212
Roupala 474
Rourea 269-270
Roystonea 69, 72
Rubia 503
Rubus 478
Rudgea 490-491, 495, 497-498, 500-
501,503-504
Rudolfiella 131, 135, 141, 147
Ruellia 184-186
Ruizterania 554-555
Rumohra 29, 32-33, 47, 54
Ruppia 175
Ruprechtia 472-473
Ruptiliocarpon 453
Russelia 523
Rutaneblina 508
Ryania 302-304
Rytidostylis 277-278
Sabal 69, 72
Sabicea491,503
Saccharum 157, 162, 170
Saccifolium 305, 309
Sacciolepis 157, 162, 164-167, 170
Saccoloma 29-3
1
Sacoglottis 317-318
Sacoila 131, 141, 146-148
Sagittaria 59
Sagonea 3 1
8
Sagotia 295, 299
Sagraea 4 1
5
Salacia 313-316
Saldanha 503
Saldanhaea219
Salmasia 247
Salmea 260
Salpichlaena 26
Salpinctes 198,202
Salpinga405,410, 415
Salvadora 433
Salvertia 554-555
Salvinia 51
Samanea 369
Samara 433
Sambucus 235
Samyda 304
Sanchezia 184
Sandemania405,408, 415
Sandwithia 295
Sandwithiodoxa 521
Santalodcs 270
Saouari 237
Sapindus 513-514
Sapium 295, 297, 299-300
Saracha 529
Saranthe 1 1
6
Sarcaulus57, 518
Sarcoglottis 131, 143-144, 146-148
580 Checklist of the Plants of the Guiana Shield
Sarcopera 396
Sarcostemma 210-212
Sarmentaria 415
Satyria 286-288
Satyrium 146
Sauvagesia 447-449
Savia 295
Saxofridericia 174-175
Scaphosepalum 131, 142, 146
Scaphyglottis 131-132, 136, 140-141,
146-148
Scelochilus 132
Scheelea 69, 72
Schefflera 206-207
Schiekia 109
Schilleria 465
Schinus510
Schismatoglottis 63, 66
Schistostemon 3 1 7-3 1
8
Schizachyrium 157-159, 170
Schizaea 50, 52
Schizolobium 339, 341-342, 345
Schizoloma 31
Schizopsis 219
Schlechterella 147
Schlegelia216, 219, 311,545
Schleidenia 226
Schlimia 132
Schlumbergeria 81
Schmidelia 514
Schnella 345
Schoenobiblus 537
Schoenocephalium 174-175
Schoenoplectus 94-95, 101-102
Schoenus 101
Schoepfia 450
Schomburgkia 132
Schradera491,503
Schrankia 376
Schuebleria 308
Schultesia 305-306, 308
Schwenckia 527, 529
Schwenkfeldia 503
Sciadocephala 260-261
Sciadodendron 207
Sciadophyllum 207
Sciadotenia 421
Sciaphila 177
Scirpidium 101
Scirpus 101-102
Sciuris 510
Scleria 94-95, 98, 101-102
Sclerolobium 345
Scleronema 221-223
Scoparia 523
Scrophularia 524
Scutellaria 321
Scuticaria 132, 142
Scyphonichium 513
Sebastiania 295, 297, 300
Secondatia 198
Secretania 451
Securidaca 469-471
Securinega 300
Seguieria 457
Selaginella 18-21
Selenicereus 232-233
Selenipedium 132, 136, 147
Selenodesmium 40
Selysia 277
Semaphyllanthe 491, 494, 503
Senapea 57
Senecio 266
Senefeldera 295, 299-300
Senefelderopsis 295, 299-300
Senegalia 376
Senna 339-346
Serapias 147
Serianthes 376
§ericographis 186
Serjania 513-514
Serrastylis 147
Sertumera 188
Seruneum 266
Sesamum 456, 523
Sesbania 353, 356-357, 363
Sesuvium 187,473
Setaria 157, 160-161, 164-167, 170
Setiscapella 378
Sextonia 326, 328-329
Siapaea 260
Sickingia 503
Sicklera 529
Sicyomorpha 315
Sida 394-395
Sidastrum 395
Sideroxylon 521
Sievekingia 132, 147
Sigesbeckia 260
Sigmatostalix 132
Silphium 266
Simaba 525
Simarouba 252
Simira 491, 497, 503-504
Sinapis 227
Sinapistrum 235
Singana 353
Sinningia 310-312
Siolmatra 277
Sipanea 491, 493, 503-505
Sipaneopsis 491-492, 503-504
Sipapoa 392
Sipapoantha 305
Siparuna 525-526
Siphanthera 405, 409-410, 414-415,
417
Siphantheropsis 415
Siphocampylus 233
Siphoneugena 438, 440-442
Siphonia 300
Sisyrinchium 111
Sitella 533
Skoliopterys 392
Sloanea 284-285, 476
Smilax 175-176
Sobralia 132, 136, 147
Socratea 69, 71-72
Soemmeringia 353
Solandra 527
Solanum 527-530
Solena 504
Solenidium 132, 143
Somphoxylon 421
Sonchus 266-267
Sophia 223
Sophoclesia 287-288
Soramia 281
Sorghastrum 157-158, 170
Sorghum 157-158, 162, 170
Soridium 177
Sorocea 425, 427
Souroubea 396
Souza 1 1
1
Spachea389, 391
Sparattanthelium 312-313
Sparattosperma 216, 218
Sparganophorus 267
Spartina 157, 170
Spathanthus 174-175
Spathelia 508-510
Spathicarpa 66
Spathiger 147
Spathiphyllum 63, 66
Spathodea219
Spathularia 548
Specklinia 147
Speirostyla 539
Spennera 415-416
Spermacoce 492-403, 504, 506
Spermodon 102
Sphaeradenia 87-88
Sphaerocionium 40
Sphaeropteris 28
Sphaerotheca 424
Sphagneticola 260-262, 266-268
Spheneria 157, 169-170
Sphenoclea 531
Sphinctanthus 492, 495, 497, 504
Sphinctolobium 363
Sphyrospermum 286-288
Spicanta 26
Spigelia 379-380
Spilanthes 260, 262, 267
Spiracantha 267
Spiranthera 508
Spiranthes 132, 144, 147
Spirodela 112
Spirotropis 353, 358, 363
Spixia 300
Spondias 189
Spongiosperma 198-199, 201-202
Sponia 542
Sporobolus 157-158, 167, 170-172
Sprucea 504
Stachyarrhena 492, 503-504
Stachyphyllum 285
Stachytarpheta 544-546
Stahelia 308
Stanhopea 132, 136, 147
Stannia 504
Staphidiastrum 416
Staphidium416
Checklist of the Plants of the Guiana Shield 581
Staurogyne 184-186
Steffensia 465-466
Stegolepis 174-175
Steinchisma 157, 165-167
Steirachne 157, 161, 171
Steiropteris 54
Stelestylis 87
Stelis 132-133, 135, 139, 147-148
Stellilabium 133
Stemmadenia 198,202
Stemmatosiphon 535
Stemodia 523-524
Stemonacanthus 186
Stenia 133, 148
Stenocalyx 392, 442
Stenocephalum 260, 268
Stenochlaena 26
Stenocoryne 148
Stenolobium 363
Stenoloma 3
1
Stenomeria 2 1
Stenopadus 260, 267
Stenophyllus 102
Stenoptera 148
Stenorrhynchos 148
Stenosolen 198,202
Stenospermation 63, 65-66
Stenotaphrum 157, 171
Stephania421
Stephanotella212
Sterculia 532-533
Sterigmapetalum 478
Steriphoma234, 421
Steyerbromelia 77, 80-81
Steyermarkochloa 157, 167, 171
Sticherus 34-35
Stictophyllorchis 133, 141, 146, 148
Stictophyllum 148
Stifftia 260, 267
Stigmaphyllon 389-392
Stigmatopteris 31-34,48
Stilaginella 300
Stilpnopappus 267
Stipa 171
Stizolobium 363
Stizophyllum216, 218
Stomatochaeta 260-262, 267
Stomoisia 378
Stratiotes 114
Strempelia 504
Strephium 171
Streptachne 171
Streptocalyx 73, 81
Streptochaeta 157
Streptogyna 157, 171
Streptostachys 157, 164, 171
Strigilia 534
Stromanthe 115-116
Strophocactus 232-233
Struchium 260, 263, 267
Struthanthus 381-384
Struthiopteris 26
Strychnodaphne 329
Strychnos 379-380
Stryphnodendron 369-371, 373-374.
376
Sturmia 148
Stylogyne 431-434, 518
Stylosanthes 353, 360, 363-364
Styrax 533-534
Subpilocereus 233
Surenus419
Suriana 534
Swartzia 346, 353-355, 363-364
Sweetia 363
Swietenia418
Syagrus 69-70
Symbolanthus 305, 307-308
Symmeria 472
Symphonia 250, 252
Symphyllanthus 280
Symphyotrichum 260-261
Symplocos 534-535
Synedrella 260, 267
Syngonanthus 105-109
Syngonium 63, 66
Syngramma 24
Syngrammatopsis 24
Synisoon 504
Syntherisma 171
Systemonodaphne 329
Syzygiopsis 521
Syzygium 438-441
Tabebuia 216-219
Tabema 202
Tabemaemontana 198-203
Tacca 177
Tachia 305, 345
Tachibota 247
Tachigali 340, 345-346
Tacsonia 456
Taenitis 49
Tagetes 260
Taguaria 384
Taligalea 546
Talinum 473
Talipariti 394-395
Talisia 513-515
Tamarindus 340
Tamonea 416, 545-546
Tampoa 57
Tanaecium 217, 219
Tanibouca 255
Taonabo 536
Tapeinostemon 305-306, 308, 502
Tapirira 189-190
Tapirocarpus 5 1
5
Tapogomea 504
Tapura 279-280
Taralea 355, 357-358, 363
Tarenaya 235
Tariri 457
Tassadia210-212
Tateanthus 405
Tecoma217, 219
Tectaria 32-33, 49, 52-53
Telanthera 188
Teliostachya 184-186
Telitoxicum 42
1
Telminostelma 212
Tephrosia 355, 358-359, 363-364
Tephrothamnus 267
Tepuia 286, 288
Tepuianthus 535
Teramnus 355, 359
Terminalia 254-255
Tematea 364
Temstroemia 535-536
Terpsichore 35-38, 46-47
Tessaria 260, 262, 267
Tetracera 281-282
Tetracoryne 304
Tetragamestus 148
Tetragastris 148
Tetralocularia 273
Tetrameranthus 192
Tetrameris 416
Tetramerium 504
Tetrapodenia 392
Tetrapollinia 306-308
Tetrapterys 389-392
Tetrastichella219
Tetrazygia 4 1
6
Tetrorchidium 295
Teuscheria 133, 135, 148
Thalia 115-116
Thaumatophyllum 66
Thecophyllum 81
Thelechitonia 267
Thelypteris 23, 32-33, 37, 47-49, 51.
53-55
Thenardia 203
Theobroma 532-533
Thermophila 3 1 5-3 1
6
Thesium 51
1
Thespesia 394
Thevetia 198-200
Thibaudia 286-288
Thieleodoxa 505
Thiloa 254-255
Thinouia 513
Thismia 83
Thoa 57
Thoracocarpus 65, 87
Thrasya 157-158, 166, 168, 171
Thunbergia 185-186
Thurnia 177
Thyella275
Thymocarpus 1 16
Thyrsacanlhus 186
Thyrsanthus 203
Thyrsodium 189
Thysanostemon 250
Tibouchina 405, 407, 409-41 1, 414,
416
Ticorea 508
Tigarea 282
Tilesia 267
Tillandsia 77-79, 81-82, 182
Tinantia 85-86
Tingulonga 230
Tinus 433-434
582 Checklist of the Plants of the Guiana Shield
Tobagoa 492, 496
Tococa 405-406, 409-413, 416-417
Tocoyena 492, 505
Tofieldia 113
Toluifera 364
Tonina 106
Tonsella 316
Tontanea 505
Tontelea 314-316
Topobea406, 417
Torenia 523-524
Tomelia 66
Torrubia 444
Torulinium 102
Touantea 364
Touchiroa 346
Toulicia513, 515
Tounatea 364
Toumefortia 225-226
Touroulia 475
Tovomita 250-253
Tovomitidium 250
Tovomitopsis 253
Trachypogon 158-159, 162, 171-172
Tradescantia 85-86
Traganthus 300
Tragia 295, 300
Tragiopsis 300
Trattinnickia 228-231
Trema 542
Trianosperm 278
Trianosperma 278
Trianthema 422
Triaristella 148
Triaristellina 148
Tribulus 556
Tricentrum 4 1
7
Tricera 23
1
Trichachne 172
Trichanthera 185-186
Trichelostylis 102
Trichilia 418-419, 515
Trichipteris 28
Trichocentrum 133-134, 136, 138,
141, 143-144, 148
Trichochaeta 102
Trichogonia 260, 264
Tricholaena 172
Trichomanes 38-41
Trichopilia 133, 144, 148
Trichosalpinx 148
Trichosanthes 133, 141, 144-148, 278
Trichospira 260-261, 266-267
Trichostigma 457
Trichovaselia 449
Tricuspis 172
Tridax 260
Tridens 158, 172
Trifolium 364
Trigonia 539-540
Trigonidium 133, 148
Trigynaea 192, 194
Trilepis 95
Trimezia 111
Trinchinettia 267
Triodia 172
Triopterys 392
Triphasia 508
Triphora 133, 146
Triplaris 472-473
Triplophyllum 32-33, 53
Tripodanthus 382-384
Tripogandra 85-86
Tripogon 158, 160
Tripsacum 158, 172
Trisecus 477
Trisetella 133, 142, 148
Trismeria 24
Tristicha 468
Xrithrinax 72
Triumfetta 538-539
Triuris 177
Trixis 267
Trizeuxis 133, 148
Tropaeolum 540
Troschelia 109
Truncaria 417
Trymatococcus 425-427
Tryssophyton 406
Tschudya 417
Tuberculocarpus 260-261, 268
Tulasnea 417
Tumera 533, 540-541
Tussacia 82, 312
Tussilago 267
Tyleria 447-448
Tyleropappus 260
Tylopsacas 310-312
Tylosperma 312
Tynanthus217, 219
Typha 178
Typhonium 63, 66
Udora 1 1
1
Ugni 438, 441-442
Uladendron 532
Uleiorchis 133, 148
Ulleria 186
Uncaria 492, 498
Uncinia 95
Unona 194
Unonopsis 192, 194
Unxia 260, 264-267
Uragoga 505
Uralepis 172
Urania 176
Uranthera417
Urena 394-395
Urera 542-543
Urochloa 158, 167, 172
Urodesmium 417
Urospatha 63-64, 66
Urospathella 63-64, 66
Urostachys 19
Urostigma 427-428
Urtica 543
Urvillea513, 515
Utricularia 376-379
Uva 194
Uvaria 194
Vaccinium 286-288
Vachellia 376
Valantia 505
Valdesia 417
Valeriana 543
Valota 172
Vandellia 524
Vandenboschia 41
Vanilla 133-134, 148
Vantanea 317-318
Vargasiella 134
Varronia 226
Vasconcellosia 236
Vaselia 449
Vasivaea 538
Vatairea 355, 361,364
Vataireopsis 355
Veconcibea 300
Vellosiella 523-524
Vellozia 178
Verbena 546
Verbesina 260, 267-268
Vemonanthura 260-261, 268
Vernonia 268
Verticillaria 253
Vesiculina 379
Vetiveria 158-159
Veyretia 148
Viburnum 235-236
Vicia 364
Victoria 444-445
Vigna 355, 356, 359, 362, 364
Vilfa 172
Villarsia 422
Villouratea 449
Vinca 203
Vincetoxicum 212
Viola 548-549
Virecta 505
Virola 429-430
Viscum 551-552
Vismia 250-253
Vitellaria521
Vitex 545-547
Vitis 552-553
Vittaria51, 55
Vochya 555
Vochysia 554-555
Vogelia 84
Vogelocassia 346
Volkameria 547
Votomita 406, 408, 411, 413, 417
Vouacapoua 340
Vouapa 57, 344, 346
Vouarana 513, 515
Voyara 235
Voyria 306-309
Voyriella 306, 308
Vriesea75, 78, 81-83
Wachendorfia 109
Wahlenbergia 233
Wallacea 448
Walleria 434
Checklist of the Plants of the Guiana Shield 583
Waltheria 532-533, 541
Warszewiczia 492, 497, 505
Webera417
Websteria 95, 97, 101-102
Weddellina 469
Wedelia 260, 266, 268
Weigeltia 434
Weinmannia 278-279
Werauhia 78, 81-83
Whittonia 456
Wikstroemia 537
Wilbrandia 277
Willoughbya 268-269
Willughbeja 203
Wilmattea 233
Windsorina 175
Wissadula 394-395
Witheringia 528-530
Wittiocactus 233
Wittmackanthus 492
Wittmackia 83
Wolffiall2
Wolffiellall2
Wolffiopsis 112
Wormskioldia 541
Wulffia 260-262, 267, 269
Wullschlaegelia 134, 148
Wurdackanthus 308
Wurdackia 109
Xanthosoma 63-64, 66
Xerorchis 134, 139
Ximenia 450
Xiphidium 109, 111
Xiphochaeta 260,267
Xiphopteris 37-38
Xyladenius 304
Xylobium 134, 142, 148
Xylophragma217, 219
Xylopia 192-194
Xylosma 302-304
Xylosterculia 533
Xyris 179-181
Youngia 260
Yutajea 505
Yuyba 72
Zamia 56
Zantedeschia 66
Zanthoxylum 508-510
Zapoteca 369-370, 373
Zea 158, 162
Zephyranthes 113
Zeugites 158
Zexmenia 269
Zeyheria219
Zingiber 182
Zinowiewia 239
Ziziphus 477
Zollemia 356, 364
Zomia 356, 360, 364
Zostera 183
Zschokkea 203
Zuelania 302-304
Zygia 369-376
Zygopetalum 148-149
Zygophlebia 36-37, 49
Zygosepalum 134, 139, 143,148-149
Zygostates 134
Rhynchospora cordatachenia M.T. Strong (Cypearaceae)
584 Checklist of the Plants of the Guiana Shield
INDEX TO FAMILIES
Acanthaceae 183
Adiantaceae 22
Agavaceae 58
Aizoaceae 187
Alismataceae 59
Amaranthaceae 187
Anacardiaceae 189
Anisophylleaceae 190
Annonaceae 190
Apiaceae 195
Apocynaceae 195
Aquifoliaceae 203
Araceae 59
Araliaceae 205
Arecaceae 67
Aristolochiaceae 207
Asclepiadaceae 208
Aspleniaceae 24
Balanophoraceae 212
Balsaminaceae 213
Basellaceae 213
Bataceae 213
Begoniaceae 213
Bignoniaceae 214
Bixaceae 220
Blechnaceae 25
Bombacaceae 220
Bonnetiaceae 223
Boraginaceae 224
Brassicaceae 226
Bromeliaceae 73
Brunelliaceae 227
Burmanniaceae 83
Burseraceae 227
Buxaceae 231
Cabombaceae 23
1
Cactaceae 23
1
Campanulaceae 233
Canellaceae 234
Cannaceae 84
Capparaceae 234
Caprifoliaceae 235
Caricaceae 236
Caryocaraceae 236
Caryophyllaceae 237
Casuarinaceae 237
Cecropiaceae 237
Celastraceae 239
Ceratophyllaceae 240
Chenopodiaceae 240
Chloranthaceae 240
Chrysobalanaceae 241
Clethraceae 247
Clusiaceae 247
Combretaceae 254
Commelinaceae 85
Compositae 255
Connaraceae 269
Convolvulaceae 271
Costaceae 86
Crassulaceae 276
Cucurbitaceae 276
Cunoniaceae 278
Cuscutaceae 279
Cyatheaceae 26
Cycadaceae 56
Cyclanthaceae 87
Cyperaceae 88
Cyrillaceae 279
Davalliaceae 29
Dennstaedtiaceae 29
Dichapetalaceae 279
Dicksoniaceae 31
Dilleniaceae 280
Dioscoreaceae 103
Droseraceae 282
Dryopteridaceae 31
Ebenaceae 282
Elaeocarpaceae 284
Eremolepidaceae 285
Ericaceae 285
Eriocaulaceae 104
Erythroxylaceae 288
Euphorbiaceae 290
Euphroniaceae 301
Flacourtiaceae 301
Gentianaceae 304
Gesneriaceae 309
Gleicheniaceae 34
Gnetaceae 57
Grammitidaceae 35
Haemodoraceae 109
Haloragaceae 312
Heliconiaceae 110
Hemandiaceae 312
Hippocrateaceae 313
Hugoniaceae 316
Humiriaceae 316
Hydrocharitaceae 110
Hydrophyllaceae 318
Hymenophyllaceae 38
Hymenophyllopsidaceae 41
Icacinaceae 318
Iridaceae 111
Isoetaceae 17
Ixonanthaceae 3 1
9
Juglandaceae 320
Juncaceae 111
Krameriaceae 320
Lacistemataceae 320
Lamiaceae 321
Lauraceae 322
Lecythidaceae 330
Leguminosae-Caesalpinioideae
333
Leguminosae-Faboideae 346
Leguminosae-Mimosoideae 365
Lemnaceae 112
Lentibulariaceae 376
Liliaceae 112
Limnocharitaceae 114
Lissocarpaceae 379
Loasaceae 379
Loganiaceae 379
Lomariopsidaceae 42
Loranthaceae 381
Lycopodiaceae 17
Lygodiaceae 43
Lythraceae 384
Magnoliaceae 385
Malpighiaceae 386
Malvaceae 393
Marantaceae 114
Marattiaceae 44
Marcgraviaceae 396
Marsileaceae 44
Mayacaceae 117
Melastomataceae 397
Meliaceae 417
Mendonciaceae 419
Menispermaceae 420
Menyanthaceae 422
Metaxyaceae 44
Molluginaceae 422
Monimiaceae 422
Monotaceae 423
Moraceae 423
Moringaceae 428
Muntingiaceae 428
Musaceae 1 1
7
Myoporaceae 428
Myricaceae 428
Myristicaceae 428
Myrsinaceae 430
Myrtaceae 434
Najadaceae 1 17
Nelumbonaceae 442
Nyctaginaceae 442
Nymphaeaceae 444
Ochnaceae 445
Olacaceae 449
Oleaceae 451
Oleandraceae 44
Onagraceae 45
1
Ophioglossaceae 45
Opiliaceae 453
Orchidaceae 118
Osmundaceae 45
Oxalidaceae 453
Papaveraceae 454
Parkeriaceae 45
Passifloraceae 454
Pedaliaceae 456
Peridiscaceae 456
Phytolaccaceae 457
Picramniaceae 457
Pinaceae 56
Piperaceae 458
Plagiogyriaceae 45
Plantaginaceae 466
Plumbaginaceae 466
Poaceae 149
Podocarpaceae 56
Podostemaceae 466
Polygalaceae 468
Polygonaceae 471
Polypodiaceae 45
Pontederiaceae 173
Portulacaceae 473
Primulaceae 473
Proteaceae 474
Psilotaceae 17
Pteridaceae 50
Quiinaceae 475
RafHesiaceae 476
Ranunculaceae 476
Rapateaceae 173
Rhabdodendraceae 476
Rhamnaceae 477
Rhizophoraceae 477
Rosaceae 478
Rubiaceae 479
Ruppiaceae 175
Rutaceae 507
Sabiaceae 510
Salviniaceae 51
Santalaceae 5 1
1
Sapindaceae 511
Sapotaceae 515
Sarraceniaceae 521
Schizaeaceae 52
Scrophulariaceae 522
Selaginellaceae 19
Simaroubaceae 524
Siparunaceae 525
Smilacaceae 175
Solanaceae 526
Sphenocleaceae 531
Sterculiaceae 531
Strelitziaceae 176
Styracaceae 533
Surianaceae 534
Symplocaceae 534
Taccaceae 177
Tectariaceae 52
Tepuianthaceae 535
Temstroemiaceae 535
Tetrameristaceae 536
Theaceae 536
Thelypteridaceae 53
Theophrastaceae 537
Thumiaceae 177
Thymelaeaceae 537
Tiliaceae 538
Trigoniaceae 539
Triuridaceae 177
Tropaeolaceae 540
Tumeraceae 540
Typhaceae 178
Ulmaceae 542
Urticaceae 542
Valerianaceae 543
Velloziaceae 178
Verbenaceae 543
Violaceae 547
Viscaceae 549
Vitaceae 552
Vittariaceae 55
Vochysiaceae 553
Winteraceae 555
Woodsiaceae 55
Xyridaceae 178
Zamiaceae 56
Zingiberaceae 182
Zosteraceae 183
Zygophyllaceae 556

ttiiif
Nautilocalyx pemphidius