S M I T H S O N I A N C O N T R I B U T I O N S T O Z O O L O G Y ? N U M B E R 4 8 0
An Illustrated Checklist
of the American Crayfishes
(Decapoda: Astacidae, Cambaridae,
and Parastacidae)
Horton H. Hobbs, Jr.
SMITHSONIAN INSTITUTION
SMITHSONIAN INSTITUTION PRESS
Washington, D.C.
1989
A B S T R A C T
Horton H. Hobbs, Jr. An Illustrated Checklist of the American Crayfishes (Decapoda:
Astacidae, Cambaridae, and Parastacidae). Smithsonian Contributions to Zoology, number 480,
236 pages, 379 figures, 1989.?This alphabetical list consists of the names of all of the
American crayfishes, including two fossil species, that were described prior to 1 January 1988.
In the treatment accorded each specific or subspecific taxon there is a synonymy citing
alterations in generic and subgeneric assignments, erroneous spellings, and synonyms.
References are also made to recent descriptions, discussions, and keys. The composition and
locations of type series are cited, as are the type locality, range, and habitat of each. Nomina nuda
that have been neither validated nor synonymized with other crayfish taxa, together with two
names suppressed by the International Commission on Zoological Nomenclature, are listed
separately. Comparable illustrations, which are arranged in related species groups, are presented
for each of the 379 recognized extant species and subspecies. Appendix 1 provides an
alphabetical list of American crayfishes and the countries, states, and/or provinces in which they
live. Appendix 2 consists of an alphabetical list of American countries and states, or provinces,
and their crayfish faunas; also cited are references to studies that pertain either directly to them
or which should be helpful in studies of them. Appendix 3 includes information comparable to
that in the main body of the checklist on crayfishes described during 1988 and 1989.
OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is
recorded in the Institution's annual report, Smithsonian Year. SERIES COVER DESIGN: The coral
Montastrea cavernosa (Linnaeus).
Library of Congress Cataloging in Publication Data
Hobbs, Horton Holcombc, 1914-
An illustrated checklist of the American crayfishes (Decapods: Astacidae, Cambaridae, and Parastacidae).
(Smithsonian contributions to zoology ; no. 480)
Bibliography: p.
Includes index.
Supt. of Docs, no.: SI 1.27:480
1. Crayfish?United States. 2. Crustacea?United States. I. Title. II. Series
QI1.S54 no. 480 591 s [595.3'841] 88-600418
[QL444.M33]
Contents
Page
Introduction 1
Location of Types 2
Brief History of the Nomenclature of American Crayfishes 2
Synopsis of American Astacidae 4
Acknowledgments 4
Nomina Nuda neither Validated nor Synonymized with Other Crayfish Taxa . . . . 5
Suppressed Names 5
Species Inquirenda 5
The Checklist 6
Family Astacidae 6
Family Cambaridae 8
Family Parastacidae 79
Appendix 1: Alphabetical List of American Crayfishes with Their Distribution . . . 82
Appendix 2: Alphabetical List of American Countries and States, or Provinces, and
Their Crayfish Faunas 88
Appendix 3: Alphabetical List of American Crayfishes Described during 1987 and
1988 91
Literature Cited 93
Explanation of Illustrations I l l
Figures 1-379 112
Index 206
in

An Illustrated Checklist
of the American Crayfishes
(Decapoda: Astacidae, Cambaridae,
and Parastacidae)
Horton H. Hobbs, Jr.
Introduction
With the realization that some seventy-odd species had been
described since the appearance of the most recent checklist of
the North and Middle American crayfishes (Hobbs, 1974b), that
a revision of the genus Cambarellus had been completed by
Fitzpatrick (1983a), and that a similar undertaking by him
treating the genus Orconectes was in progress, it seemed
appropriate that preparation begin on updating the 1974
checklist. Soon after the project was initiated, it became
obvious that numerous modifications in the synonymies would
be essential; to the best of my knowledge, aided by the
assistance of those recognized in the "Acknowledgments"
section, such emendations and additions have been included.
Although Bahamonde (1951, 1958) and Holthuis (1956)
helped immeasurably in elucidating the nature of the parastacid
fauna of Chile, and Buckup and Rossi (1980) ably reviewed the
parastacids of Brazil and Uruguay, the only comprehensive
summary of the crayfishes of South America since Faxon's
(1914) compilation was that of Riek (1971). This work
constituted an important contribution, particularly in pointing
out the distinct nature of the two Chilean stocks, but it lacked
the benefit of the information provided in the study of Buckup
and Rossi in which species that had been declared synonyms by
Riek were resurrected and a previously unknown one was
described. With these additions to our knowledge of the South
American crayfish fauna, it seemed appropriate that the revised
checklist be expanded to encompass the American parastacids.
Unfortunately, unavailable at this time is a review of the
Chilean crayfish fauna by Dr. Ludwig Buckup, which is in
press.
All of the American crayfishes (including two fossil species)
Horton H. Hobbs, Jr., Department of Invertebrate Zoology, National
Museum of Natural History, Smithsonian Institution, Washington,
D.C. 20560.
that were described prior to 1 January 1987 are believed to be
accounted for in the following list While those few species that
have been described since that date do not appear in the list,
combinations involving older names have been updated to 1
October 1987, and all new taxa that have come to my attention
as of 1 January 1989 are included in Appendix 3.
The format of the 1974 checklist has been followed to the
extent possible. Thus a list of collections containing type
specimens of American crayfishes is followed by a brief
historical summary of nomcnclaiural changes affecting the
supraspecific taxa embracing these American crayfishes, and
appended to it is a list of those taxa adopted herein. Preceding
the alphabetical list of species the nomina nuda that have been
neither validated nor associated with an established crayfish
taxon are cited as are those names that have been suppressed by
the International Commission on Zoological Nomenclature.
For each of the specific or subspecific taxa listed, a synonomy
is provided in which are included alterations in generic and
subgencric assignments, erroneous spellings, and synonyms; in
only rare instances are references to misidentifications listed.
Citations are also made to recent descriptions, discussions,
illustrations, and keys. The locations of the type series are
pointed out, and, where possible, the sexes of the syntypes and
primary types (holotype, allotype, and morphotype) are noted.
In instances in which the type designations included a
holotypic male, form I, and a "paratypic male, form II" (the
latter clearly separated from the remainder of the paratypic
series), the paratypic male has been referred to here as
"morphotype," the name enclosed in quotation marks. In citing
the types of those taxa for which a morphotype was designated
by the describer, the quotation marks are omitted.
The type locality, range, and habitats are cited for each
species and subspecies. The designations "primary, secondary,
and tertiary burrowers" follow Hobbs (1942b:20): primary
burrowers are those largely restricted to burrows; secondary
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
burrowers are those that generally occupy burrows but often
wander into open water during times when the mouths of the
burrows are flooded; and tertiary burrowers are those that
burrow "only in periods of drought or occasionally [or usually],
but not necessarily, during the breeding season."
To the extent possible, the illustrations, which are arranged
in related species groups, are made from types of the species,
and the rather few instances in which types were not available,
specimens were chosen from, or as close to, the type locality as
possible (see "Explanation of Illustrations").
Appendix 1 consists of an alphabetical list of the families,
subfamilies, genera, species, and subspecies of the crayfishes
occurring in the Americas and the countries and states or
provinces in which each occurs. Appendix 2 contains a list of
the countries and states or provinces with references to the
crayfishes (listed in Appendix 1) that are known or have been
reported to occur within their boundaries. Appendix 3 consists
of an alphabetical list of American crayfishes described during
1987 and 1988, after this manuscript had been completed.
LOCATION OF TYPES.?The following abbreviations are used
for institutional collections and most personal collections in
which specimens of one or more type series have been
deposited.
Alabama Mus. Nat.
Hist.
AMNH
AMS
ANSP
BCPM
BMNH
BSNH
Chas. Mus.
CM
GM
FSM
FSBC
IBM
INHS
La. Polytechnic Insi.
MCZ
MHNP
MMNH
MMZ
MN.UFRJ
Mus. Washburn Coll.
Alabama Museum of Natural History, Tuscaloosa,
Alabama.
American Museum of Natural History, New York,
New York.
Australian Museum, Sydney, Australia.
Academy of Natural Sciences of Philadelphia,
Pennsylvania.
British Columbia Provincial Museum, Victoria,
British Columbia, Canada.
British Museum (Natural History), London, Eng-
land, (paratypes not listed in Hobbs (1974b)
subsequently received in exchange from USNM.)
Boston Society of Natural History, Boston, Massa-
chusetts.
Charleston Museum, South Carolina.
Carnegie Museum, Pittsburg, Pennsylvania.
Museum d'Histoire Naturelle,Geneva,Switzerland.
Florida Slate Museum, Gainesville, Florida.
Bureau of Marine Research, St. Petersburg, FLorida.
Institute de Biologi'a de la Universidad Nacional
Autonoma de Mexico, Mexico City, Mexico.
Illinois State Laboratory of Natural History, Urbana,
Illinois.
Louisiana Polytechnic Institute, Ruston, Louisiana.
Museum of Comparative Zoology, Harvard Univer-
sity, Cambridge, Massachusetts.
Museum National d'Histoire Naturelle, Paris,
France.
Mississippi Museum of Natural History, Jackson,
Mississippi
Museum of Zoology, University of Michigan, Ann
Arbor, Michigan
Museu Nacional da Universidade Federal do Rio de
Janeiro, Brazil.
Museum of Washbum College, now Department of
Biology, Washburn University, Topeka, Kansas.
Mus. Zool. Torino Institute e Museo de Zoologia, Delia Universita di
Torino, Italy.
NCSM North Carolina State Museum of Natural History,
Raleigh, North Carolina
NMV National Museum of Victoria, Melbourne, Australia.
OSM Ohio State Museum, Columbus, Ohio.
PM Peabody Museum of Natural History, Yale Univer-
sity, New Haven, Connecticut.
RNHL Rijksmuseum van Natuurlijke Historie, Leiden, The
Netherlands
RR Collection of Rendell Rhoades, Ashland College,
Ashland, Ohio (insofar as is known, all of
collection has now been deposited in OSM or
USNM).
TMM Texas Memorial Museum, Austin, Texas.
TU Department of Zoology, Tulane University, New
Orleans, Louisiana.
UF Department of Zoology, University of Florida,
Gainesville, Florida (most, if not all of the crayfish
types in this collection were transferred to FSBC).
USNM Collections of former United States National Mu-
seum, now in National Museum of Natural
History, Smithsonian Institution, Washington,
D.C.
UTM University of Tennessee Museum, Knoxville, Ten-
nessee
UZM Universitets Zoologiske Museum, Krystalgade,
K0benhavn 0 . Denmark.
Wurzburg Mus. Wiirzburg Museum, Wiirzburg, Germany.
ZBM Zoologisches Museum der Humboldt-Universitat,
Berlin, East Germany.
ZIAS Zoological Institute, Academy of Sciences, Lenin-
grad, U.S.S.R.
In addition, paratypic specimens of a few species have been
reported to be in the collections of the following: D.W.
Albaugh, address uncertain; Donald Batch, Eastern Kentucky
University, Richmond, Kentucky; Joe B. Black, Louisiana
College, Pineville, Louisiana; Coe College, Cedar Rapids,
Iowa; Troy C. Dorris, Oklahoma State University, Stillwater,
Oklahoma; H.H. Hobbs IH.WittenbergUniversity,Springfield,
Ohio; Leslie Hubricht, Meridian, Mississippi; Hugo L. James,
University of Bridgeport, Bridgeport, Connecticut; Max R.
Matteson, University of Illinois, Urbana, Illinois; Ronald D.
Oesch, Glendale, Missouri; Joseph F. Pickett, Sr., Jacksonville,
Florida; Rudolph Prins, Western Kentucky University, Bowl-
ing Green, Kentucky; Rollin D. Reimer, address uncertain;
H.W. Robison, Southern Arkansas University, Magnolia,
Arkansas; Jerry G. Walls, Hightstown, New Jersey.
Five large crayfish collections?those of the University of
Michigan Museum of Zoology, the University of Kansas
Museum of Natural History, personal collections of Raymond
W. Bouchard, Joseph F. Fitzpatrick, Jr., and the author?have
been transferred to the National Museum of Natural History,
Smithsonian Institution, and most, if not all, of the paratypic
specimens cited by previous authors as having been deposited
in those collections are now in the Smithsonian crayfish
collection.
BRIEF HISTORY OF THE NOMENCLATURE OF AMERICAN
CRAYFISHES.?North America and Middle America: Prior to
NUMBER 480
the publication of the first summary treatment of North
American crayfishes by Girard (1852), the eight recognized
species were assigned to the genus Astacus. Erichson (1846)
had proposed the subgenus Cambarus to receive the North
American species included in his study, and this taxon was
elevated to generic rank by Girard (1852) in his revision of the
nearctic crayfishes in which 20 species were recognized. With
the adoption of the genus Cambarus by Hagen (1870) in his
"Monograph of the North American Astacidae," in which a
clear delineation was made between Cambarus (occurring east
of the continental divide) and Astacus (largely restricted to the
western versant) a generic stability became established that
remained unchanged until 1933. Faxon (1884) presented the
first checklist of holarctic crayfishes and listed 60 North
American species. In the following year, his revision of the
family Astacidae appeared, but in it he made few changes in the
"species groups" that had been proposed by Hagen.
As recounted in my previous checklist (Hobbs, 1974:2-3),
with some reshuffling of species, Ortmann (1905c, 1905d, 1906a) proposed the
recognition of six subgenera of Cambarus: Cambarus, Bartonius, Faxonius,
Cambarellus, Procambarus, and Paracambarus. Fowler (1912), in reviewing
the Crustacea of New Jersey, pointed out that Ortmann (1905c) overlooked
Faxon's (1898) selection of Astacus Bartonii Fabricius as the type species of
Cambarus. Ortmann had chosen that crayfish as the type species of the
subgenus Bartonius, and Astacus Blandingii Harlan as that of his subgenus
Cambarus. Thus the name Bartonius was shown to be a junior objective
synonym of Erichson's Cambarus, and Ortmannicus was proposed to replace
Ortmann's subgenus Cambarus.
In his checklist of the crayfishes[,j Faxon (1914) included 110 species and
subspecies from North America but recognized no subgenera. An alteration in
the generic assignment of the North American crayfishes had its inception with
Creaser (1933a) who elevated Ortmann's subgenus Faxonius to generic rank.
Not until 1942, however, was there a major departure from the generic structure
that had been established by Girard (1852) and Hagen (1870).
Creaser (1933b) erected the monotypic subgenus Faxonella to recieve
Cambarus clypeatus Hay. This subgenus was questionably accorded generic
status by Creaser (1962) and was clearly so treated by Fitzpatrick (1963).
Lyle (1938) designated the subgenus Girardiella to encompass Faxon's
(1914) Group II.
Hobbs (1942a) proposed the elevation of Ortmann's subgeneric groupings of
Cambarus to generic rank, recognizing the subfamilies Astacinae and
Cambarinae, the latter comprising the following genera: Cambarus, Procam-
barus (including Fowler's Ortmannicus and Lyle's Girardiella), Orconectes (=
Ortmann's Faxonius), Cambarellus, Paracambarus, and Troglocambarus, the
latter described as new. Orconectes Cope (1872) was considered to be a
subjective senior synonym of Ortmann's Faxonius.
[Boa (1950), in revising the European crayfishes, proposed the genus
Pacifasiacus to receive the American astacids largely restricted to the Pacific
versant of North America, a group that had been assigned previously to the
genus Astacus].
Villalobos (1955) recognized the Cambaroidinae [comprising the eastern
Asiatic crayfishes] as a subfamily of the Astacidae, and Laguarda (1961)
designated a fourth subfamily, the Cambarellinae. Fitzpatrick and Payne (1968)
added the genus Hobbseus, and Hobbs (1969b), in revising the genus
Cambarus, proposed the genus FaUicambarus. In the latter revision, 10
subgenera of Cambarus were recognized. One of them, Barbicambarus, was
elevated to generic rank by Bouchard (1972a).
In 1972, Hobbs (1972a) recognized 16 subgenera of Procambarus,
resurrecting the subgenus Ortmannicus (Fowler, 1912) and Girardiella (Lyle,
1938) and returning Paracambarus to subgeneric status in Procambarus. Later
in the same year, Hobbs' (1972b) key to most of the crayfishes included
[t]herein was made available.
In a synopsis of the families and genera of crayfishes of the
world, Hobbs (1974a) proposed a revised structure of the
Astacuran infraorder Astacidea in which three superfamilies,
Nephropoidea, Parastacoidea, and Astacoidea were recognized
and diagnosed. Synonymies and diagnoses were included for
the families and genera of the last two. In that study, the
subfamily Cambarinae was elevated to familial rank, and the
proposal made that, in addition to the nominate subfamily, the
Cambaroidinae and Cambarellinae be included within it. The
most recent checklist of the North (and Middle) American
crayfishes was that of Hobbs (1974b) in which 294 species and
subspecies were recognized.
Bouchard and Hobbs (1976) added the monotypic subgenus
Exilicambarus to the genus Cambarus, and Hobbs (1977c)
proposed the monotypic genus Bouchardina for a disjunct
crayfish occurring in southwestern Arkansas. Bouchard
(1978b) erected the subgenus Hobbsastacus in the genus
Pacifastacus, and Hobbs (1981) recognized the then monotypic
subgenus Distocambarus of the genus Procambarus. The latter
subgenus was elevated to generic rank by Hobbs and Carlson
(1983). Hobbs (1983) proposed the subgenus Fitzcambarus to
receive an apparently disjunct member of the genus.
The genus Cambarellus was revised by Fitzpatrick (1983a)
who recognized within it three subgenera: Dirigicambarus and
Pandicambarus in addition to the nominate subgenus. The
most recent major revision was that of Fitzpatrick (1987a) who
proposed the recognition of 10 subgenera in the genus
Orconectes: Billecambarus, Buannuliflctus, Crockerinus, Fax-
onius, Gremicambarus, Hespericambarus, Orconectes, Pro-
cericambarus, Rhoadesius, and Tragulicambarus.
South America: Of those who described crayfishes occur-
ring in South America, the first five authors (Poeppig, 1835; H.
Milne Edwards, 1837; Nicolet, 1849; Martens 1869; and
Philippi, 1882, 1894) who described crayfishes occurring on
this continent employed the generic name Astacus. In a revision
of the genus, Erichson (1846) proposed that five subgenera of
Astacus be recognized, and assigned all of the American
crayfishes (including Astacus (C.) Chilensis H. Milne Edwards,
1837) to the subgenus Cambarus; presumably, he was unaware
of Poeppig's (1835) description of A. pugnax. Herklots (1861),
in abstracting the taxa treated by Dc Haan in his Fauna
Japonica, placed the same species in the subgenus Astacus. The
generic name Parastacus Huxley (1879) was not generally
applied to the South American crayfishes until Faxon
(1898:683) expanded the definition of the genus and presented
descriptions of five new species, two of which had been
previously described. The only supraspecific change that has
been suggested subsequent to the adoption of the generic name
Parastacus was the proposal of Riek (1971) that two of the
species that had been assigned to Parastacus be reassigned to
a new genus Samastacus, establishing the new combinations S.
araucanius (Faxon, 1914) and S. spinifrons (Philippi, 1882).
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
The most recent study adding or ressurecting taxa was that of
Buckup and Rossi (1980).
SYNOPSIS OF AMERICAN ASTACIDEA.?The following struc-
turing of the American Astacidea is modified from the
classification of the Astacidea presented by Hobbs (1974a,
1974b).
Infraorder ASTACIDEA
Supeifamily NEPHROPOIDEA Dana, 18S2 (lobsters and marine allies)
Superfamily ASTACOIDEA Latreille, 1802-1803
Family ASTACIDAE Latreille, 1802-1803
Genus Pacifastacus Bott, 1950
Subgenus Hobbsastacus Bouchard, 1978b
Subgenus Pacifastacus Bott, 19S0
Family CAMBARIDAE Hobbs, 1942a
Subfamily CAMBARELUNAE Laguarda, 1961
Genus Cambarellus Ortmann, 1905c
Subgenus Cambarellus Ortmann, 1905c
Subgenus Dirigicambarus Fitzpatrick, 1983a
Subgenus Pandicambarus Fitzpatrick, 1983a
Subfamily CAMBARINAE Hobbs, 1942a
Genus Barbicambarus Hobbs, 1969b
Genus Bouchardina Hobbs, 1977c
Genus Cambarus Erichson, 1846
Subgenus Avilicambarus Hobbs, 1969b
Subgenus Cambarus, Erichson, 1846
Subgenus Depressicambarus Hobbs, 1969b
Subgenus Erebicambarus Hobbs, 1969b
Subgenus Exilicambarus Bouchard and Hobbs, 1976
Subgenus Hiaticambarus Hobbs, 1969b
Subgenus Jugicambarus Hobbs, 1969b
Subgenus Lacunicambarus Hobbs, 1969b
Subgenus Puncticambarus Hobbs, 1969b
Subgenus Veticambarus Hobbs, 1969b
Genus Distocambarus Hobbs, 1981
Subgenus Distocambarus Hobbs, 1981
Subgenus Fitzcambarus Hobbs, 1983
Genus Fallicambarus Hobbs, 1969b
Subgenus Creaserinus Hobbs, 1973a
Subgenus Fallicambarus Hobbs, 1969b
Genus Faxonella Creaser, 1933b
Genus Hobbseus Fitzpatrick and Payne, 1968
Genus Orconectes Cope, 1872
Subgenus Billecambarus Fitzpatrick, 1987a
Subgenus Buannulifictus Fitzpatrick, 1987a
Subgenus Crockerinus Fitzpatrick, 1987a
Subgenus Faxonius Ortmann, 1905c
Subgenus Gremicambarus Fitzpatrick, 1987a
Subgenus Hespericambarus Fitzpatrick, 1987a
Subgenus Orconectes Cope, 1872
Subgenus Procericambarus Fitzpatrick, 1987a
Subgenus Rhoadesius Fitzpatrick, 1987a
Subgenus Tragulicambarus Fitzpatrick, 1987a
Genus Procambarus Ortmann, 1905d
Subgenus Acucauda Hobbs, 1972a
Subgenus Austrocambarus Hobbs, 1972a
Subgenus CapUlkambarus Hobbs, 1972a
Subgenus Girardiella Lyle, 1938
Subgenus Hagenides Hobbs, 1972a
Subgenus Leconticambarus Hobbs, 1972a
Subgenus Lonnbergius Hobbs, 1972a
Subgenus Mexicambarus Hobbs, 1972a
Subgenus Ortmannicus Fowler, 1912
Subgenus Paracambarus Ortmann, 1906a
Subgenus Pennides Hobbs, 1972a
Subgenus Procambarus Ortmann, 19O5d
Subgenus Remoticambarus Hobbs, 1972a
Subgenus Scapulicambarus Hobbs, 1972a
Subgenus Tenuicambarus Hobbs, 1972a
Subgenus Villalobosus Hobbs, 1972a
Genus Troglocambarus Hobbs, 1942a
Superfamily PARASTACOIDEA Huxley, 1879
Family PARASTACIDAE Huxley, 1879
Genus Parastacus Huxley, 1879
Genus Samastacus Rick, 1971
ACKNOWLEDGMENTS.?Again, I extend my thanks to all of
those persons listed under "Acknowledgments" in Hobbs
(1974b:4) for their assistance in obtaining information concern-
ing crayfish types and/or for their criticisms and suggestions
pertaining to one or more drafts of that manuscript, and for
corrections and improvements resulting from their assistance in
the preparation of the 1974 checklist that have been incorpo-
rated in this revision. In subsequent queries concerning a
number of items in the 1974 checklist, J.F. Fitzpatrick, Jr., has
been most helpful, and his careful reading, suggestions, and
corrections of the manuscript have resulted in a much more
reliable and complete final version. Invaluable also was the
assistance of Douglas G. Smith, of the University of Massachu-
setts, who provided me with meticulous responses to questions
related to type material housed in the Museum of Comparative
Zoology. Jacques Forest, of the Museum National d'Histoire
Naturelle, Paris, generously furnished me not only with
photographs of H. Milne Edwards' type of Astacus chilensis,
but lent me the other South American parastacids in the Paris
museum. Gustavo Augusto S. de Melo of the Museu de
Zoologia da Sa"o Paulo kindly lent me a specimen of
Parastacus laevigatus, which could not be obtained elsewhere.
A number of persons have assisted in checking the numerous
references cited herein or have read parts or the entire
manuscript, offering advice and criticisms; those assisting with
the present manuscript are Margaret A. Andolshek, formerly of
the Smithsonian Institution; three other Smithsonian col-
leagues, Thomas E. Bowman, Fenner A. Chace, Jr., and C.W.
Hart, Jr.; H.H. Hobbs III of Wittenberg University; Douglas G.
Smith, and Monica Ter-Minasian, of the Madeira School; and
my wife, Georgia B. Hobbs. Janice Clark and C.W. Hart, Jr.,
have been both patient and helpful in assisting me in organizing
the manuscript in the word processor, as has Molly Ryan in
offering advice involving the preparation of the figures.
Through the expertise of Carolyn S. Hahn of the Smithsonian
Library, many incomplete and/or difficult-to-find references
became available. To all of those persons mentioned above I
extend my thanks for their interest and help; unquestionably,
the compilation is much improved by their contributions.
Finally, I am indeed grateful to Barbara Spann and Don Fisher
for the skillful editing of this work.
NUMBER 480
Nomina Nuda neither Validated nor Synonymized with
Other Crayfish Taxa
Through the efforts of J.F. Fitzpatrick, Jr., the nomina nuda
created by Clay Lyle (1938), and listed by Hobbs (1974b:4) in
his checklist as not having been validated or synonymized with
other species, now have been associated with previously or
subsequently described taxa. The synonymies involved were
kindly furnished to me by Dr. Fitzpatrick (in litt., 3 March
1983). During his tenure on the faculty of Mississippi State
University, Dr. Fitzpatrick received from Dr. Lyle the
collection on which the latter had based his unpublished
dissertation and the abstract (1938) creating the nomina nuda.
Much of the collection was in rather poor condition, but
retrievable parts of it are now housed in the Smithsonian
Institution. The synonymies involved may be traced through
the index herein.
To my knowledge, only two nomina nuda applied to
American crayfishes, one inadvertently, have not been unques-
tionably synonymized with currently recognized crayfish taxa:
Cambarus schrnittii Penn, 1941:8 [questionably synonymized with Cambarel-
lus puer Hobbs (Hobbs, 1974b:4)].
Cambarus lucifugus Packard, 1888:82 (lapsus for Crangonyx lucifugus Hay,an
amphipod (Hobbs, 1974b:4)].
Suppressed Names
Astacus fossor Rafinesque, 1817:42 [Suppressed by Opinion 522 of the
International Commission on Zoological Nomenclature. (See Hemming,
1958:212.)]
Astacus oreganus Randall, 1840:138 [Suppressed by Opinion 855 of the
International Commission on Zoological Nomenclature. (See Melville and
China, 1968:84.)]
Species Inquirenda
With the discovery of additional fossils conspecific with
Cambarus primaevus Packard (1880:222) (= Procambarus
(Austrocambarus) primaevus (Feldmann et al., 1981:794)),
only the lack of an association of one taxon with a crayfish
seems to remain:
Procambarus wiegmanni (Erichson)
Astacus (Cambarus) Wiegmanni Erichson, 1846:99.
Cambarus Wiegmanni.?Hagen, 1870:54.
Cambarus wiegmanni.?Faxon, 1890:621.
Procambarus wiegmanni.?Hobbs, 1942a:342 [by implication]; 1974b:4.
Procambarus weigmanni.?Villalobos, 1949:338 [erroneous spelling].
Procambarus Weigmanni.?Villalobos, 1950:381 [erroneous spelling].
TYPES.?Type lost from BMNH.
TYPE LOCALITY.?Mexico.
RANGE.?Unknown.
HABITAT.?Unknown.
REMARKS.?Nothing further is known than was reported in
my previous checklist (Hobbs: 1974b:4-5):
The identity of this crayfish has not been recognized with certainty since it was
described. Villalobos (1950) reviewed its history and designated Erichson's
name a nomen nudum. Inasmuch as Erichson diagnosed and described the
species and stated that it occurs in Mexico, it cannot be considered a nomen
nudum.
The species was referred to the genus Procambarus on the basis of
Erichson's (1846:99) statement, "Des zweite Glied am dritten und am vierten
Beinpaar beim Mannchen mit einem hakenfdrmigen Fortsatz)." Only [three]
crayfish genera are known to occur in Mexico [Orconectes (G.) virilis has been
reported to have been introduced (Campos and Contreras, 1985:218)], and, of
the [three], this statement could apply only to a member of the genus
Procambarus.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
The Checklist
Family ASTACIDAE Latreille, 1802-1803
Genus Pacifastacus Bott
Astacus.?Dana, 1852:522.
Cambarus.?Girard, 1852:90.
Cambrus.?Girard, 1852:90 [erroneous spelling].
Potamobius.?Ortmann, 1902:286.
Cambaras.?Anonymous, 1927, explanation of figure [erroneous spelling].
Cambals.?Anonymous, 1927:258 [erroneous spelling].
Pacifastacus Boa, 1950:24. [Type species, by original designation, Astacus
Klamathensis Stimpson, 1857a: 87. Gender masculine.]
Pasifastacus.?Sanders, 1967:33 [erroneous spelling].
Pacificastacus.?LaCaze, 1970:2 [erroneous spelling].
Pasifasticus.?Anonymous, 1972, cover page [erroneous spelling].
Pacifastaculus.?Kamila, 1973:123 [erroneous spelling].
Gambelly.?Cabantous, 1976:50 [erroneous spelling].
Subgenus Hobbsastacus Bouchard, 1977b:38 [nomen nudum]; 1978b:431.
[Type species, by original designation, Cambrus [sic] Gambelii Girard,
1852:90 (= "Astacus Gambelii Girard, 1852:90"). Gender masculine.]
Pacifiastacus.?Mason, 1976:473 [erroneous spelling].
Pacifasticus.?Crenshaw, Lemons, and Russo, 1980:243 [erroneous spelling].
Subgenus Hobbsastacus Bouchard
Cambrus.?Girard, 1852:90 [lapsus for Cambarus].
Astacus.?Stimpson, 1857a:87.
Cambarus.?Hagen, 1870:31.
Subgenus Astacus.?Faxon, 1898:666.
Potamobius.?Ortmann, 1902:286.
Subgenus Potamobius.?Ortmann, 1902:286.
Pacifastacus Bott, 1950:24.
Gambelly.?Cabantous, 1976:50 [erroneous spelling].
Subgenus Hobbsastacus Bouchard, 1977b:38 [nomen nudum];1978b:431.
[Type species, by original designation, Cambrus [sic] Gambelii Girard,
1852:90 (= "Astacus Gambellii Girard, 1852:90." Gender masculine.]
A. chenoderma, suggesting that both A. breviforceps and A.
subgrundialis are probable synonyms. Since the types are lost
and accounts of specimens assignable to either have not
appeared since Miss Rathbun's review of the species, I have
followed her in including these species as probable synonyms
of Pacifastacus chenoderma. The species was transferred to the
genus Pacifastacus and referred to the subgenus Hobbsastacus
because the rostrum with as many as seven pairs of marginal
spines is a feature foreign to members of the Eurasian
Astacidae but strongly resembles that of four members of the
American astacid genus Pacifastacus: P. connectens, P. fortis,
P. gambelii, and P. nigrescens.
Pacifastacus (Hobbsastacus) connectens (Faxon)
FIGURE 1
Astacus gambelii connectens Faxon, 1914:360, pi. 7: figs. 6,10,pl. 10.
Pacifastacus gambelii connectens.?Bott, 1950:24 [by implication].
Pacifastacus gambeli connectens.?Riegel, 1959:43.
Pacifastacus gambelii connectans.?Holt, 1960:61 [erroneous spelling].
Pacifastacus connectens.?Hobbs, 1972b:22,figs. 12d, 14a; 1974b:5,fig. 1.
Pacifastacus (Hobbsastacus) connectens.?Bouchard, 1978b:431.
Pacifastacus conectens.?Fitzpatrick, 1983b: 155 [erroneous spelling].
TYPES.?Syntypes, USNM 23096 (2 males, 1 female),
USNM 44410 (1 female), MCZ 7385 (1 male) transferred from
USNM.
TYPE LOCALITY.?Snake River, Upper Salmon Falls, Twin
Falls County (not Logan County as cited by Hobbs, 1974b),
Idaho.
RANGE.?Idaho and Oregon.
HABITAT.?Streams.
Pacifastacus (Hobbsastacus) chenoderma (Cope)
Astacus chenoderma Cope, 1871:606.?Rathbun, 1926:130, pi. 29: fig. 7; pis.
31-34.
?Astacus subgrundialis Cope, 1871:605. [Types not extant Type locality,
"fresh water deposit in the Territory of Idaho, near Hot Springs Mountain."]
?Astacus breviforceps Cope, 1871:606. [Types not extant Type locality, same
as that for Astacus chenoderma, see below.]
Astacus chinoderma.?Brodsky, 1981:122 [erroneous spelling].
Pacifastacus chenoderma.?Hobbs, 1974b: 5.
Pacifastacus (Hobbsastacus) chenoderma.?Bouchard, 1978b:431.
TYPES.?Syntype, USNM 353343; other syntypes probably
lost
TYPE LOCALITY.?"Catherine's creek, Idaho." According to
an annotation in a reprint, this is "a local name for the upper
part of Castle Creek, Owyhee County." Previously reported to
be a Pliocene deposit, Feldmann et al. (1981:788) stated that
this fossil was "described from the Miocene Payette Formation
in Idaho and Oregon."
RANGE.?Miocene and Pliocene(?) deposits in Idaho and
Oregon.
REMARKS.?Rathbun (1926) redescribed and figured Cope's
Pacifastacus (Hobbsastacus) fortis (Faxon)
FIGURE 3
Astacus nigrescens fortis Faxon, 1914:360, pi. 7: figs. 5,9, pi. 9: fig. 2.
Pacifastacus nigrescens fortis.?Bott, 1950:24 [by implication].
Pacifastacus fortis.?Hobbs, 1972b:23, figs. 13c, 14c; 1974b:5, fig. 3.?Eng
and Daniels, 1982:197, figs. 1-6.
Pacifastacus (Hobbsastacus) fortis.?Bouchard, 1978b:431; 1980a:416.
TYPES.?Syntypes, USNM 44404 (1 male, 2 females),
USNM 44405 (3 females), MCZ 7383 (1 male, 1 female).
TYPE LOCALITY.?Fall River, Fall City Mills, Shasta
County, California.
RANGE.?Known only from Shasta County, California.
HABITAT.?Streams.
Pacifastacus (Hobbsastacus) gambelii (Girard)
FIGURE 2
Cambrus Gambelii Girard, 1852:90 [lapsus for Cambarus].
Astacus Gambelii.?Stimpson, 1857b:492.?Hagen, 1870:90, pi. 1: figs. 97,
98, pi. 3: fig. 170.pl. 11.
NUMBER 480
Astacus Gambeli.?Hagen, 1870:28 [erroneous spelling].
Cambarus Gambeli.?Hagen, 1870:31 [erroneous spelling].
Astacus (Astacus) Gambelii.?Faxon, 1884:152 [by implication].
Astacus Gambellii.?Underwood 1886:364 [erroneous spelling].
Astacus (Astacus) gambelii.?Faxon, 1898:666 [by implication].
Astacus gambeli.?Kingsley, 1899:822 [erroneous spelling].
Potamobius (Potamobius) gambeli.?Ortmann, 1902:286 [by implication;
erroneous spelling].
Astacus gambelii gambelii.?Faxon, 1914:360 [by implication].
Astacus gambelii.?Raihbun, 1926:133.
Pacifastacus gambelii gambelii.?Bott, 1950:24 [by implication].
Pacifastacus gambelii.?Kozloff, 1955:156.?Hobbs, 1972b:22, figs. 13a,
14b;1974b:5,fig. 2.
Pacifastacus gambeli.?Riegel, 1959:29,43-44, figs. 3c, 8.
Astacus gambelii.?Laguarda, 1962:284 [erroneous spelling].
Astacus gambelii gambelii.?Laguarda, 1962:280 [erroneous spelling].
Cambarus gambelii.?Hobbs, 1967a: 130 [erroneous spelling].
Pacifastacus gambelii gambelii.?Hobbs, 1967a: 130 [erroneous spelling].
Pacifastacus (Hobbsastacus) gambelii.?Bouchard, 1978b:431.
TYPES.?Syntypes, ANSP 1531 (10+, mostly fragments),
ANSP 1532 (5 specimens).
TYPE LOCALITY.?"California."
RANGE.?Pacific slope and in the Missouri River drainage in
California(?), Idaho, Montana, Nevada, Oregon, Utah, Wash-
ington, and Wyoming.
HABITAT.?Streams and lakes.
Pacifastacus (Hobbsastacus) nigrescens (Stimpson)
FIGURE 4
Astacus nigrescens Stimpson, 1857a:87; 1857b:492.?Hagen, 1870:92, pi. 3:
fig. 168.
Astacus (Astacus) nigrescens.?Faxon, 1884:152 [by implication].
Potamobius (Potamobius) nigrescens.?Ortmann, 1902:286 [by implication].
Astacus nigrescens nigrescens.?Faxon, 1914:360 [by implication].
Pacifastacus nigrescens nigrescens.?Bott, 1950:24 [by implication].
Pacifastacus nigrescens.?Riegel, 1959:44, figs. 3d, 9,10.?Hobbs, 1972b:23,
figs. 13b, 14d; 1974b:7,fig.4.?Bouchard, 1978c:419.
Pacifastacus nigresens.?Goldman, 1973:107 [erroneous spelling].
Gambelly nigrescens.?Cabantous, 1976:50 [erroneous spelling].
Pacifastacus (Hobbsastacus) nigrescens.?Bouchard, 1978b:431.
TYPES.?Not extant; "probably destroyed in Chicago fire in
1871" (Faxon, 1914:408).
TYPE LOCALITY.?Vicinity of San Francisco, California.
RANGE.?Vicinity of San Francisco, California.
HABITAT.?Streams.
REMARKS.?This crayfish is probably extinct.
Subgenus Pacifastacus Bott
Astacus.? Dana, 1852:522.
Subgenus Astacus.?Faxon, 1884:151 [by implication].
Potamobius.?Ortmann, 1902:286.
Subgenus Potamobius.?Ortmann, 1902:286.
Cambaras.?Anonymous, 1927,unnumbered figure [erroneous spelling].
Cambals.?Anonymous, 1927:258 [erroneous spelling].
Pacifastacus Bott, 1950:24. [Type species, by original designation, Astacus
Klamathensis Stimpson, 1857a:87. Gender masculine.]
Subgenus Pacifastacus.?Brinck, 1968:85.
Pasifasticus.?Anonymous, 1972, cover page [erroneous spelling].
Pacifastaculus.?Kamita, 1973:123 [erroneous spelling].
Pacifiastacus.?Mason, 1976:473 [erroneous spelling].
Pacifastacus (Pacifastacus) leniusculus klamathensis
(Stimpson)
FIGURE 6
Astacus Klamathensis Stimpson, 1857a:87.?Hagen, 1870:93, pi. 3: fig.
169.?Faxon, 1885a: 131, pi. 6: figs. 1,2.
C[ambarus] Klamathensis.?Hagen, 1870:102 [erroneous combination].
Astacus klaymathensis.?Huxley, 1880b:223 [erroneous spelling].
Astacus (Astacus) Klamathensis.?Faxon, 1884:151 [by implication].
Astacus klamathensis.?Faxon, 1890:634.
Astacus (Astacus) klamathensis.?Faxon, 1898:665.
Potamobius (Potamobius) klamathensis.?Ortmann, 1902:286.
Pacifastacus klamathensis.?Bott, 1950:24 [by implication].?Riegel,
1959:41. figs. 3b, 7,10.
Pacifastacus leniusculus klamathensis.?Miller and Van Hyning, 1970:77.?
Hobbs, 1974b:6, fig. 6 [not 5 as noted].
Pacifastacus leniusculus leniusculus.?Hobbs, 1974b:81 [figs. 5 and 6 are
interchanged].
Pacifastacus klamathenis.?Upson, 1977:43 [erroneous spelling].
Pacifastacus (Pacifastacus) leniusculus klamathensis.?Bouchard, 1978b:431.
Pacifastacus leniuscilus klamathensis.?Huner and Barr, 1980:145 [erroneous
spelling].
Pacifastacus leniusculus klamathensis.?Brodsky, 1981:188 [erroneous spell-
ing].
Pacifastacus lenisculus klamathensis.?Fitzpatrick, 1983b: 155 [erroneous
spelling].
TYPES.?Not extant; "probably destroyed in the Chicago fire
in 1871" (Faxon, 1914:409).
TYPE LOCALITY.?Lake Klamath, Klamath County, Oregon.
RANGE.?British Columbia and Idaho south to central
California.
HABITAT.?Lakes and cold, swift streams.
Pacifastacus (Pacifastacus) leniusculus leniusculus (Dana)
FIGURE 5
Astacus leniusculus Dana, 1852:522, 524; 1855, pi. 33: fig. la.b.?Hagen,
1870:94.
Astacus laeviusculus.?Lockington, 1878:304 [erroneous spelling].
Astacus leenisculus.?Huxley, 188Oa:305 [erroneous spelling].
Astacus (Astacus) leniusculus.?Faxon, 1884:151 [by implication]; 1885a:132,
pi. 6: fig. 4.
Potamobius (Potamobius) leniusculus.?Ortmann, 1902:286 [by implication].
Potamobius leniusculus.?Bowkiewicz, 1926:1.
Cambaras americanus Anonymous, 1927, unnumbered figures [nomen
nudum].
Cambals americunus Anonymous, 1927:258 [nomen nudum].
Pacifastacus leniusculus.?Bott, 1950:24 [by implication].?Riegel, 1959:39,
figs. ld-h,3a,6,10.
Pacifastacus lenuisculus.?Hopkins, 1967:53 [erroneous spelling].
Pacifastacus trowbridgeii.?Sanders, 1967:33 [figure: misidentification and
erroneous spelling].
"Astacus" (Pacifastacus) leniusculus.?Brinck, 1968:85.
Pacifastacus leniusculus leniusculus.?Miller and Van Hyning, 1970:77.?
Hobbs, 1972b:21,figs. 12a, 14e; 1974b:6,fig. 6 (not 5 as noted).
Pasifasticus leniusculus.?Anonymous, 1972, cover page [erroneous spelling].
Pacifastaculus leniuscus.?Kamita, 1973:123 [erroneous spellings].
Pacifastacus lenisculus.?Brodsky, 1973:52 [erroneous spelling].
P[acifastacus] I. lenusculus.?Hart and Hart, 1974:132 [erroneous spelling].
8 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Pacifastacus Uniusculus klamathensis.?Hobbs, 1974b: fig. 6 [figs. 5 and 6 are
interchanged].
Pacifastacus lenuisculus.?Schapiro, 1975:15 [erroneous spelling].
Pacifastacus (Pacifastacus) Uniusculus Uniusculus.?Bouchard, 1978b:431.
Pacifasticus Uniusculus.?Crenshaw, Lemons, and Russo, 1980:243 [errone-
ous spelling].
Pacifastacus Uniusculus.?Crenshaw, Lemons, and Russo, 1980:254 [errone-
ous spelling].
Pacifastacus Uniuskulus.?Brodsky, 1981:109 [erroneous spelling].
Pacifastacus I. Uniuaculus.?Brodsky, 1981:189 [erroneous spelling].
Pacifastacus I. Unisculus.?Fitzpalrick, 1983b: 155 [erroneous spelling].
Pacifastacus lenisuculus.?McGriff, 1983:238 [erroneous spelling].
Pacifastacus leniusculas.?Avault and Huner, 1985:43 [erroneous spelling].
TYPES.?Syntypes, USNM 2019 (1 male), USNM 2161 (1
male), USNM 44408 (1 male).
TYPE LOCALITY.?Columbia River and Puget Sound, Wash-
ington.
RANGE.?British Columbia, California, Idaho, Nevada,
Oregon, Utah, and Washington. Introduced into Sweden
(Svardson, 1965:92) and Japan (Kamita, 1970:140).
HABITAT.?Streams and lakes.
REMARKS.?Astacus leniusculus was placed on the Official
List of Specific Names in Zoology, name number 2244, in
Opinion 855 of the International Commission on Zoological
Nomenclature. (See Melville, 1968:84).
Pacifastacus (Pacifastacus) leniusculus trowbridgii
(Stimpson)
FIGURE 7
Astacus Trowbridgii Stimpson, 1857a:87; 1857b:493.?Hagen, 1870:93. pl.3:
fig. 171, pi. 10.
Astacus Trownbridgi.?Huxley, 1883:271 [erroneous spelling].
Astacus (Astacus) Trowbridgii.?Faxon, 1884:152 [by implication].
Astacus (Astacus) trowbridgii.?Faxon, 1898:666 [by implication].
Astacus trowbridgii.?Hay 1899b:958.
Potamobius (Potamobius) trowbridgei.?Ortmann, 1902:286 [by implication;
erroneous spelling].
Pacifastacus trowbridgii.?Bott, 1950:24 [by implication].?Riegel, 1959:30,
fig. lc-?.?Mason, 1970a:37, figs. 1-16; 1970b:969,figs. 1-7.
Astacus trbwbridgi.?Pennak, 1953:461.
Astacus trowbridgei.?Pilgrim, 1960:249 [erroneous spelling].
Pacifastacus Trowbridgii.?Haertel and Osterberg, 1967:464.
Pacifastacus trowbridgeii.?Sanders, 1967:33 [erroneous spelling].
Pasifastacus trowbridgeii.?Sanders, 1967:33 [erroneous spelling].
Pacifastacus leniusculus trowbridgii.?Miller and Van Hyning, 1970:77.?
Hobbs, 1972b:22, figs. 5a, 12b; 1974b:6, fig. 7.
Pacifastacus trowbridgL?Goldman, 1973:106 [erroneous spelling].
Pacifastaculus trowbridgii.?Kamita, 1973:123 [erroneous spelling].
Pacifastacus trawbridgii.?Miyakc, 1973:7 [erroneous spelling].
Pacifastacus leniuslus trowbridgii.?Mason, 1976:450 [erroneous spelling].
Pacifastacus trowbridgei.?Lake and Newcombe, 1975:211 [erroneous spell-
ing].
Pacifiastacus leniusculus trowbridgii.?Mason, 1976:473 [erroneous spelling].
Pacifasticus leniusculus trowbridgii.?Momot and Gowing, 1977:2043 [erro-
neous spelling].
Pacifastacus (Pacifastacus) Uniusculus trowbridgii.?Bouchard, 1978b:431.
Pacifastacus Unisculus trowbridgii.?Fitzpatrick, 1983b: 155 [erroneous spell-
ing].
TYPES.?Syntypes, USNM 2080 (2 males, 2 females), MCZ
3510(1 male),BSNH,PM.
TYPE LOCALITY.?Columbia River above Astoria, Oregon.
RANGE.?British Columbia, California, Idaho, Oregon,
Nevada and Washington. Introduced into Japan (Kamita,
1970:148).
HABITAT.?Streams and lakes.
Family CAMBARIDAE Hobbs, 1942a
Subfamily CAMBARELLINAE Laguarda, 1961
Genus Cambarellus Ortmann
Cambarus.?Saussure, 1857a: 102.
Subgenus Cambarellus Ortmann, 1905c:106. [Type species, by original
designation, Cambarus Montezumae Saussure, 1857a: 102. Gender: mascu-
line.].?Fitzpatrick, 1983a:267.
Cambarellus.?Hobbs, 1942a:350.
Camberellus.?Villalobos, 1953:357 [erroneous spelling].
Caambarus.?Villalobos, 1955:276 [erroneous spelling].
Cambarella.?Carlisle and Knowles, 1959:50 [erroneous spelling].
Subgenus Dirigicambarus Fitzpatrick, 1983a:267. [Type species, by original
designation, Cambarus Shufeldtii Faxon, 1884:134. Gender: masculine.]
Subgenus Pandicambarus Fitzpatrick, 1983a:268. [Type species, by original
designation, Cambarellus schmitti Hobbs, 1942b: 149. Gender: masculine.]
Subgenus Cambarellus Ortmann
Cambarus.-?Saussure, 1857a:102.
Subgenus Cambarellus Ortmann, 19O5c:106. [Type species, by original
designation, Cambarus Montezumae Saussure, 1857a:102. Gender mascu-
line.].?Fitzpatrick, 1983a:267.
Cambarellus.?Hobbs, 1942a:340.
Caambarus.?Villalobos, 1955:276 [erroneous spelling].
Cambarellus (Cambarellus) alvarezi Villalobos
FIGURE 9
Cambarellus alvarezi Villalobos, 1952:525, pi. 1: figs. 1-7, pi. 2: figs.
1-5.?Hobbs, 1972b:27, fig. 15f; 1974b:7.fig. 14.
Cambarellus (Cambarellus) alvarezi.?Fitzpatrick, 1983a:267,fig. 3L.
TYPES.?Holotype and morphotype, IBM 1-1952 (male I,
male II); paratypes, USNM.
TYPE LOCALITY.?Potosi, Nuevo Le6n, 23 km (14.4 mi)
northwest of Galena, Mexico.
RANGE.?Known only from the vicinity of the type locality.
HABITAT.?"Colecta realizada en un deposito de agua de una
cuenca cerrada, situada en la parte Norte de Valle Salado"
(Villalobos, 1952:531).
Cambarellus (Cambarellus) areolatus (Faxon)
FIGURE 12
Cambarus Montezumae var. areolata Faxon, 1885a: 123.
Cambarus montezumae areolatus.?Faxon, 1898:661, pi. 66: fig. 2.
Cambarus (Cambarellus) montezumae areolatus.?Ortmann, 19O6a:23.
Cambarellus montezumae areolatus.?Hobbs, 1942a:350 [by implication].?
Villalobos, 1955:276.
NUMBER 480
Cambarellus moniezumae var. areolata.?Villalobos, 1953:358.
Caambarus montezumae areolatus.?Villalobos, 1955:276 [erroneous spell-
ing].
Cambarellus areolatus.?Hobbs, 1972b:27,figs. 16b, 17b; 1974b:7,fig. 16.
Cambarellus (Cambarellus) areolatus.?Fitzpatrick, 1983a:267,fig. 3s.
TYPES.?Syntypes, MCZ 3650 (male 1,3 females, male II).
TYPE LOCALITY.?Near Parras, Coahuila, Mexico.
RANGE.?Known only from vicinity of type locality.
HABITAT.?Unknown.
Cambarellus (Cambarellus) chapalanus (Faxon)
FIGURE 10
Cambarus chapalanus Faxon, 1898:661, pi. 67: figs. 1,2.
Cambarus chaplanus.?Harris, 1903:82 [erroneous spelling].
Cambarus (Cambarellus) chapalanus.?Ortmann, 1905c:106.
Cambarus montezumae chapalanus.?Faxon, 1914:371.
Cambarus chapalamus.?Van S trad en, 1942:5 [erroneous spelling].
Cambarellus montezumae chapalanus.?Hobbs, 1942a:35O [by implication].
Cambarellus chapalanus.?Villalobos, 1955:277.?Hobbs, 1972b:28, figs.
16c,17c;1974b:7,fig. 15.
Cambarellus campalanus.?Villalobos, 1983:263 [erroneous spelling in
Hobbs' translation].
Cambarellus (Cambarellus) chapalanus.?Fitzpatrick, 1983a:267,fig. 3P.
TYPES.?Holotype, USNM 17698 (male I); paratypes,
USNM, MCZ.
TYPE LOCALITY.?Lake Chapala, State of Jalisco, Mexico.
RANGE.?Jalisco and Michoac&i, Mexico.
HABITAT.?Lentic situations.
Cambarellus (Cambarellus) chihuahuae Hobbs
FIGURE 11
Cambarellus chihuahuae Hobbs, 1980:194, fig. 1.
Cambarellus (Cambarellus) chihuahuae.?Fitzpairick, 1983a:267,fig. 3K.
Cambarellus chihauhuae.?Fitzpatrick, 1983b: 160 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
148895,148896,148897; paratypes, USNM, BMNH.RNHL.
TYPE LOCALITY.?Ojo de Carbonera, 4.3 kilometers (2.7
mi) south of Ejido Rancho Nuevo, approximately 57.6
kilometers (36 air miles) west of Villa Ahumada (30?35'N,
106?51'W), Chihuahua, Mexico.
RANGE.?Known only from the vicinity of the type locality.
HABITAT.?Streams.
Cambarellus (Cambarellus) montezumae (Saussure)
FIGURE 14
Cambarus Montezumae Saussure, 1857a: 102; 1858:459, fig. 22.
Cambarus Montezumae var. tridens von Martens, 1872:130. [Syntypes, ZBM.
Type locality: Puebla, Mexico.]
Cambarus Montezumae Montezumae.?von Martens, 1872:130 [by implica-
tion].
Cambarus montezumae.?Faxon, 1890:633.
Cambarus montezumae montezumae.?Faxon, 1898:660 [by implication].
Cambarus montezumae dugesii Faxon, 1898:660, pi. 66: fig. 1. [Syntypes,
USNM 16087 (4 male II, 12 female), MCZ. Type locality: State of
Guanajuato, Mexico.]
Cambarus montezumae dugesi?flay, 1899b:960.?Villalobos, 1953:345.
Cambarus (Cambarellus) montezumae.?Ortmann, 1905c: 106.
Cambarus (Cambarellus) montezumae dugesi.?Ortmann, 1906a:20.
Cambarus (Cambarellus) Montezumae.?Andrews, 1910:83 [by implication].
Cambarellus montezumae.?Hobbs, 1942a:340; 1972b:28, figs. 16f, 18b,d;
1974b:8,fig. 17.
Cambarellus montezumae montezumae.?Hobbs, 1942a:350 [by implica-
tion].?Villalobos, 1943:588, pi. 1; 1955:256; 1983:243.
Cambarellus montezumae dugesii.?Hobbs, 1942a:350 [by implication].
Cambarellus montezumae forma lermensis Villalobos, 1943:603, pL 2:
figs. 3, 5, 8, 11, 14, 18, 20. [Holotype, allotype, and "morphotype,"
IBM 50043(009) (male I, female, male II); paratypes, IBM. Type locality:
Lerma, Estado de Mexico, Mexico.]
Cambarellus Montezumae.?Villalobos, 1943:587.
Cambarellus montezumae lermensis.?Villalobos, 1953:345, 359; 1955:38,
262, pi. 59: figs. 3,5,8,11,14,18,20; 1983:40,248. pi. 59: figs. 3,5,8,11,
14,18,20.
Cambarellus montezumae var. tridens.?Villalobos, 1953:345.
Cambarellus montezumae dugesi.?Villalobos, 1953:345.
Cambarus montezumae var. tridens.?Villalobos, 1953:353.
Cambarellus montazumae.?Huner and Barr, 1980:140 [erroneous spelling].
Cambarellus (Cambarellus) moniezumae.?Fitzpatrick, 1983a:267,fig. 30.
TYPES.?Syntypes, USNM 20583 (male I), GM (uncata-
logued,dry, 13 specimens), BMNH 3587.
TYPE LOCALITY.?Swamps of the Valley of Mexico; ponds
in Chapultepec Park, Mexico.
RANGE.?Pacific slope and Valley of Mexico.
HABITAT.?Sluggish streams and lentic situations.
Cambarellus (Cambarellus) occidentalis (Faxon)
FIGURE 8
Cambarus montezumae occidentalis Faxon, 1898:661, pi. 66: figs. 3,4.
Cambarus (Cambarellus) montezumae occidentalis.?Ortmann, 1905c: 106 [by
implication].
Cambarellus montezumae occidentalis.?Hobbs, 1942a:350 [by implica-
tion].?Villalobos, 1953:358; 1955:276.
Cambarellus occidentalis.?Hobbs, 1972b:27, figs. 16a, 17a; 1974b:8,fig. 13.
Cambarellus (Cambarellus) occidentalis.?Fitzpatrick, 1983a:267,fig. 3M.
TYPES.?Syntypes, MCZ 3652 (2 male 1,2 male II, female
dry).
TYPE LOCALITY.?Mazatl?n, State of Sinaloa, Mexico.
RANGE.?Michoacan and Sinaloa, Mexico.
HABITAT.?Lentic situations.
Cambarellus (Cambarellus) patzcuarensis Villalobos
FIGURE 15
Cambarellus montezumae patzcuarensis Villalobos, 1943:607, pi. 2: figs. 1,6,
9,10,12,15,19,21.23; 1953:345; 1955:265,pl. 59: figs. 1,6,9,10,12,15,
19, 21, 23; 1983:252, pi. 59: figs. 1, 6, 9, 10, 12,15,19,21, 23.?Miyake,
1973, fig. 9.?Kamiu, 1973:123, figs. 1-4.
Cambarellus patzacuarensis.?Villalobos, 1953:345 [erroneous spelling].
Cambarellus patzcuarensis.?Villalobos, 1955:280.?Hobbs, 1972b:28, figs.
16d, 17d; 1974b: 8, fig. 18.
Cambarellus (Cambarellus) patzquarensis.?Fitzpatrick, 1983a:267 [errone-
ous spelling].
10 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambarellus (Cambarellus) patzcuareruis.?Fitzpatrick, 1983a:274, fig. 3R.
TYPES.?Holotype, allotype, and "morphotype," IBM
90040(005) (male I, female, male II); paratypes, IBM.
TYPE LOCALITY.?Lago de Patzcuaro, State of Michoacan,
Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Lake.
Cambarellus (Cambarellus) prottxus Villalobos and Hobbs
FIGURE 13
Cambarellus prolixus Villalobos and Hobbs, 1981:490, figs. 1-3.
Cambarellus (Cambarellus) prolixus.?Fitzpatrick, 1983a:267, fig. 3N.
TYPES.?Holotype, allotype, and morphotype, USNM
177206, 177207,177208 (male I, female, male II); paratypes,
IBM, RNHL, TMM, USNM, BMNH.
TYPE LOCALITY.?Lago de Chapala, 500 meters offshore
from north levee at Ajijic, Jalisco, Mexico.
RANGE.?Known only from Lago de Chapala.
HABITAT?Lentic situations.
Cambarellus (Cambarellus) zempoalensis Villalobos
FIGURE 16
Cambarellus montezumae forma zempoalensis Villalobos, 1943:601, pi. 2: figs.
2,4,7,13,16,17.22.
Cambarellus montezumae zempoalensis.?Villalobos, 1953:345; 1955:257, pi.
59: figs. 2,4,7,13,16,17,22; 1983:244, pL 59: figs. 2,4,7,13,16.17,22.
Cambarellus zempoalensis.?Hobbs, 1972b:28, figs. 16e, 18a,c; 1974b:9, fig.
19.
Cambarellus (Cambarellus) zempoalensis.?Fitzpatrick, 1983a:267, fig. 3Q.
TYPES.?Holotype, allotype, and "morphotype," IBM
100042(OOFA) (male I, female, male II); paratypes, IBM.
TYPE LOCALITY.?Lagunas de Zempoala, State of Morelos,
Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Crater lakes.
Subgenus Dirigicambarus Fitzpatrick
Cambar us.?Faxon, 1884:134.
Subgenus Cambarellus.?Ortmann, 1905c: 106.
Cambarellus.?Hobbs. 1942a:350.
Cambarella.?Carlisle and Knowles, 1959:50 [erroneous spelling].
Subgenus Dirigicambarus Fitzpatrick, 1983a:267. [Type species, by original
designation, Cambarus Shufeldtii Faxon, 1884:134. Gender masculine.]
Cambarellus (Dirigicambarus) shufeldtii (Faxon)
FIGURE 17
Cambarus Shufeldtii Faxon, 1884:134; 1885a:124,pL 7: fig. 1; pL 10: figs. 8,
8', 8a, 8a'.
Cambarus shufeldtii.?Underwood, 1886:372.
Cambarus shufeldti.?Ortmann, 1902:278.
Cambarus (Cambarellus) shufeldti.?Ortmann, 1905c:106.
Cambarellus shufeldtii.?Hobbs, 1942a:350 [by implication]; 1972b:25, fig.
15a; 1974b:9, fig. 8.?Perm, 1942:644; 1959:14, figs. 24, 45. 63. 77.
86.?Hobbs and Marchand. 1943:17. figs. 1. 11. 15. 21.?Albaugh and
Black, 1973. fig. 2.?Huner. 1977:10.?Bouchard. 1978d. cover photo-
graph.?Page, 1985b:363.figs. 90-93.?Pflieger. 1987a:29; 1987b:13.
Cambarellus shufeltdii.?Bick. Homuff, and Lambremont, 1953:227 [errone-
ous spelling].
Cambarellus shufeldti.?Pennak, 1953:462.
Cambarella shufeldti.?Carlisle and Knowles, 1959:50 [erroneous spelling.]
Cambarellus schufeldi.?Sprague and Couch, 1971:530 [erroneous spelling.]
Cambarellus shufeldtii.?Anonymous, 1976, caption for illustration on
unnumbered page [erroneous spelling].
Cambarellus shulfeldtii.?Jordan and Dunham, 1981:385 [erroneous spelling].
Cambarellus (Dirigicambarus) shufeldtii.?Fitzpatrick, 1983a:268. fig. 3c.
TYPES.?Syntypes, USNM 4860 (3 male 1,7 female), MCZ
3684 (male I, male 11,2 females).
TYPE LOCALITY.?Near New Orleans, Louisiana.
RANGE.?Mississippi River drainage system in southern
Illinois, Missouri, Kentucky, Tennessee, Arkansas, and in
Louisiana east of the Mississippi and Atchafalaya rivers; Red
River system in Louisiana and Texas; Brazos and Colorado
rivers in Texas; Pascagoula, Pearl, upper Tombigbee, and
Escatawpa river systems in Mississippi; and southwestern
Alabama. Introduced into Rapides, St. Bernard, and St.
Tammany parishes, Louisiana (Penn, 1950b:422), and into
Harris County, Georgia (Penn, personal communication). It has
not been collected subsequently in the last-mentioned area.
HABITAT.?Swamps, ditches, sloughs, lakes, ponds, and
sluggish streams. Burrows as water disappears from habitat.
Subgenus Pandicambarus Fitzpatrick
Cambarus.?Penn, 1941:8.
Cambarellus.?-Hobbs. 1942b: 149.
Subgenus Pandicambarus Fitzpatrick, 1983a:268. [Type species, by original
designation, Cambarellus schmitti Hobbs, 1942b: 149. Gender masculine.]
Cambarellus (Pandicambarus) blacki Hobbs
FIGURE 18
Cambarellus blacki Hobbs, 1980:201. fig. 2.
Cambarellus (Pandicambarus) blacki.?Fitzpatrick, 1983a:268, fig. 3E.
TYPES.?Holotype, allotype, and morphotype, USNM
148901,148902, and 148903; paratypes, USNM.
TYPE LOCALITY.?Small cypress pond at Oak Grove,
Escambia County, FLorida.
RANGE.?Known only from the type locality.
HABITAT.?Cypress pond.
Cambarellus (Pandicambarus) diminutus Hobbs
FIGURE 24
Cambarellus diminutus Hobbs, 1945b:467, figs. 1, 2, 5, 6. 10, 12, 13. 17, 18,
20.25,26; 1972b:26.fig. 15b; 1974b:7.fig. 12.
Cambarellus (Pandicambarus) diminutus.?Fitzpatrick, 1983a:268, fig. 3H.
TYPES.?Holotype, allotype, and "morphotype," USNM
NUMBER 480 11
81554 (male I, female, male II); paratypes, USNM, MCZ, CM,
TU.
TYPE LOCALITY.?Sand-bottomed stream, 3.5 miles (5.6
km) south of Irvington, Mobile County, Alabama.
RANGE.?Mobile County, Alabama, and George and Jack-
son counties, Mississippi.
HABITAT.?Pools, sluggish portions of streams and ditches.
Burrows as water disappears from habitat.
81556 (male I, female, male ID; paratypes, MCZ, ANSP, TU,
USNM.
TYPE LOCALITY.?Roadside ditch, 7 miles (11.2 km) west of
Dayton, Liberty County, Texas, on U.S. Highway 90.
RANGE.?From Brazos and Brazoria counties, Texas, east-
ward through the coastal plain to and in the Mississippi basin
from the lower part of the delta to Johnson County, Illinois.
HABITAT.?Swamps, ditches, sloughs, ponds, and sluggish
streams.
Cambarellus (Pandicambarus) lesUei Fitzpatrick and
Lining
FIGURE 20
Cambarellus schmiiii.?Uobbs, 1942b: 151 [in part].
Cambarellus lesliei Fitzpatrick, 1976b:55 [nomen nudum].?Fitzpatrick and
Laning. 1976:138, fig. 1.
Cambarellus leslei.?Bouchard, 1978b:451 [erroneous spelling].
Cambarellus (Pandicambarus) lesliei.?Fitzpatrick, 1983a:268,fig. 3F.
TYPES.?Holotype, allotype, and morphotype, USNM
146616, 146617, 146618 (male I, female, male II); paratypes,
USNM.TU.
TYPE LOCALITY.?0.5 mile (0.8 km) south of Alabama Port,
Mobile County, Alabama.
RANGE.?Baldwin, Mobile, and Washington counties, Ala-
bama, and George County, Mississippi.
HABITAT.?Streams and pools.
Cambarellus (Pandicambarus) ninae Hobbs
FIOURE23
Cambarellus ninae Hobbs, 195Oa:89, figs. 1-17; 1972b:26, fig. 15d; 1974b:8,
fig. 4.?Albaugh and Black, 1973:177, fig. 2.
Cambarellus (Pandicambarus) ninae.?Fitzpatrick, 1983a:268, fig. 3J.
TYPES.?Holotype, allotype, and morphotype, USNM
89768, 89770, 89769 (male I, female, male II); paratypes,
MCZ, ANSP, USNM, TU.
TYPE LOCALITY.?Borrow ditches in the Aransas Refuge
(along East Short Road), Aransas County, Texas.
RANGE.?Aransas, Calhoun, Refugio, and Victoria counties,
Texas.
HABITAT.?Lentic situations.
Cambarellus (Pandicambarus) puer Hobbs
FIGURE 21
?Cambarus schmittii Penn, 1941:8 [nomen nudum].
Cambarellus puer Hobbs, 1945b:469. figs. 7-9, 11, 14-16, 19, 21-24;
1972b:27,figs. 5c, 15e; 1974b:8,fig. 10.?Penn, 195Ob:423.?Albaugh and
Black, 1973:177, fig. 2.?Chambers, Payne, and Kennedy, 1979:39, figs.
1-5; 1980:169, figs. 1-4.?Page, 1985b:366, figs. 94-97.?Pflieger,
1987a:29; 1987b: 13.
Cambarellus (Pandicambarus) puer.?Fitzpatrick. 1983a:268, fig. 3G.
TYPES.?Holotype, allotype, and "morphotype," USNM
Cambarellus (Pandicambarus) schmitti Hobbs
FIGURE 19
Cambarellus schmitti Hobbs, 1942b: 149-153 [in part, not specimens from
Alabama], figs. 176-180, 286-295; 1972b:26. fig. 15c; 1974b:9, fig.
9.?Fitzpatrick and Laning, 1976:137.
Cambarellus (Pandicambarus) schmitti.?Fitzpatrick, 1983a:268,fig. 3D.
TYPES.?Holotype, allotype, and "morphotype," USNM
81291 (male I, female, male II); paratypes, USNM, MCZ,
Chas. Mus., TU, UF (now in USNM and FSBC).
TYPE LOCALITY.?Small spring flowing into the Suwannee
River at Branford, Suwannee County, Florida (Sec. 16, T. 6S,
R. 14E).
RANGE.?From Santa Rosa County, eastward to the Suwan-
nee River drainage system in Florida.
HABITAT.?Clear streams and springs.
Cambarellus (Pandicambarus) texanus Albaugh and Black
FIGURE 22
Cambarellus texanus Albaugh and Black, 1973:177, fig. 1.
Cambarellus (Pandicambarus) texanus.?Fitzpatrick, 1983a:268,3l.
TYPES.?Holotype, allotype, and morphotype, USNM
144191, 144192, 144193 (male I, female, male II); paratypes,
USNM, MCZ, TU, Texas A&M University, and Joe B. Black.
TYPE LOCALITY.?Ditch beside Farm Road 521, 1.5 mile
(2.4 km) south, 1 mile (1.6 km) west of Wadsworth, 12 miles
(19.2 km) south of Bay City, Matagorda County, Texas.
RANGE.?East of the Lavaca River and Bay to the Brazos
River drainage system, Texas.
HABITAT.?Roadside ditches.
Subfamily CAMBARINAE Hobbs, 1942a
CAMBARINAE Hobbs, 1942a:338.
CARBARINAE Eberly, 1955:283 [erroneous spelling].
Genus Barbicambarus Hobbs
Cambarus.?Faxon, 1884:120.
Subgenus Bartonius Ortmann, 1905c:120.
Subgenus Cambarus.?Fowler,1912:341 [by implication].
Subgenus Barbicambarus Hobbs, 1969b:98. [Type species, by original
designation, Cambarus cornutus Faxon, 1884:120. Gender, masculine.]
Barbicambarus.?Bouchard, 1972a:56.
12 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Barbicambarus cornutus (Faxon)
FIGURE 25
Cambarus cornutus Faxon, 1884:120; 1885*80, pi. 5: figs. 1,2; pi. 9: figs. 3,
3'.?Cooper and Cooper, 1968:19, illustrated.
Cambarus (Bartonius) cornutus.?Ortmann, 1905c: 120.
Cambarus (Cambarus) cornutus.?Fowler, 1912:341 [by implication].
Cambarus (Barbicambarus) cornutus.?Hobbs, 1969b:98, figs. 2d, 5,13k, 14j,
17b; 1974b:9,fig. 20.
Barbicambarus cornutus.?Bouchard, 1972a:56.
TYPES.?Holotype, MCZ 3566 (1 male I).
TYPE LOCALITY.?Green River, near Mammoth Cave,
Edmonson County, Kentucky.
RANGE.?Barren and Green river systems in Kentucky and
Tennessee.
HABITAT.?Streams.
Genus Bouchardina Hobbs
Bouchardina Hobbs, 1977c:734. [Type species, by original designation,
Bouchardina robisoni Hobbs, 1977c:734. Gender feminine.]
Bourchardina.?Anonymous, 1987:4 [erroneous spelling].
Bouchardina robisoni Hobbs
FIGURE 26
Bouchardina robisoni Hobbs, 1977c:734, fig. 1.
Bourchardina robinsoni.?Anonymous, 1987:4 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
147146,147147, 145743 (male I, female, male II); paratypes,
BMNH.USNM.
TYPE LOCALITY.?Backwaters of Bayou Bodcaw (Red
River Basin) in borrow ditch along Sunray Road, 4 miles (6.4
km) north of Lewisville off State Route 29, Lafayette County,
Arkansas (Sec. 14, T. 15S.R. 2W).
RANGE.?Bodcaw and Dorcheat bayou basins in southwest-
em Arkansas.
HABITAT.?Lentic to sluggish lotic situations.
Genus Cambarus Erichson
Aslacus.?Fabricius, 1798:407.
Subgenus Cambarus Erichson, 1846:97. [Type species, by subsequent
designation (Faxon, 1898:644), Astacus Bartonii Fabricius, 1798:407.
Proposed as subgenus of Aslacus. Gender masculine.]
Cambarus.?Girard, 1852:88.
Cambrarus.?Anonymous, 1861:27 [erroneous spelling].
Gambarus ? Huxley, 188Ob:81 [erroneous spelling].
Orconeaes Cope, 1881:881 [in part].
Camburus.?Faxon, 1885c:358 [erroneous spelling].
Camberus.?W.S. Miller, 1895:336 [erroneous spelling].
Camparus.?Williamson, 1899:47 [erroneous spelling].
S[ambarus].?Stecle, 1902:11 [erroneous spelling].
Bartonius Ortmann, 1905c:97. [Type species, by original designation, "C.
bartoni" Fabricius, 1798:407. Proposed as subgenus of Cambarus, treated
unintentionally as generic name by Williamson, 1907:749, and declared a
synonym of Cambarus by Fowler, 1912:341. Gender masculine.]
?Cambaraus.?Turner, 1926:43 (erroneous spelling].
Cambarug.?Bicrstein and Vinogradov, 1934:62 [erroneous spelling].
Bartontius.?Rioja, 1941:193 [erroneous spelling].
Cambaus.?Okada,1948:133 [erroneous spelling].
Canbarus.?McKeever, 1958:324 [erroneous spelling].
Subgenus Veticambarus Hobbs, 1969b:96. [Type species, by original designa-
tion, Cambarus pristinus Hobbs, 1965:268. Gender: masculine.]
Subgenus Aviticambarus Hobbs, 1969b:99. [Type species, by original
designation, Orconectes hamulatus Cope, 1881:881. Gender masculine.]
Subgenus Erebicambarus Hobbs, 1969b:99. [Type species, by original
designation, Cambarus bartoni tenebrosus Hay, 1902a:232. Gender
masculine.]
Subgenus Puncticambarus Hobbs, 1969b:101. [Type species, by original
designation, Cambarus extraneus Hagen, 1870:73. Gender: masculine.]
Subgenus Depressicambarus Hobbs, 1969b: 102. [Type species, by original
designation, Astacus latimanus LeConte, 1856:402. Gender: masculine.]
Subgenus Hiaticambarus Hobbs, 1969b:105. [Type species, by original
designation, Cambarus longulus Girard, 1852:90. Gender: masculine.]
Subgenus Jugicambarus Hobbs, 1969b: 106. [Type species, by original
designation, Cambarus bartonii asperimanus Faxon, 1914:391. Gender
masculine.]
Subgenus Lacunicambarus Hobbs, 1969b:110. [Type species, by original
designation, Cambarus diogenes Girard, 1852:88. Gender: masculine.]
Cambaroides.?Unestam, 1969:204 [lapsus calami].
cambarus.?Padgett, 1970:19 [lapsus calami].
Cambaras.?Bouchard, 1973a:106 [erroneous spelling].
Cambraus.?Peters, 1975:iii [erroneous spelling].
Subgenus Exilicambarus Bouchard and Hobbs, 1976:2. [Type species, by
original designation, Cambarus (Exilicambarus) cracens Bouchard and
Hobbs, 1976:2. Gender masculine.]
Carbarus.?Bouchard, 1976c:17 [erroneous spelling].
Cambarinus.?Spitzy, 1976:445 [erroneous spelling].
Camabarus.?Huner, 1977:12 [erroneous spelling].
Procambarus.?Wharton, 1978:46 [lapsus calami].
cabarus.?Root, 1979:K-2 [erroneous spelling].
Cambaurs.?Brodsky, 1983:465 [erroneous spelling].
Cabmarus.?Thome, 1987:421 [erroneous spelling].
Subgenus Aviticambarus Hobbs
Orconectes Cope, 1881:881 [in part].
Cambarus.?Faxon, 1884:145 [in part].
Subgenus Bartonius Ortmann, 1905c:97 [in part].
Subgenus Cambarus.?Fowler, 1912:341.
Subgenus Aviticambarus Hobbs, 1969b:99. [Type species, by original
designation, Orconectes hamulatus Cope, 1881:881. Gender masculine.]
Carbarus.?Bouchard, 1976c:17 [erroneous spelling].
Cambarus (Aviticambarus) hamulatus (Cope)
FIGURE 27
Orconectes hamulatus Cope, 1881:881, pi. 7: fig. la.b.
Cambarus hamulatus.?Faxon, 1884:145.?Hobbs and Ban-, 1960:17, figs.
1-10.
Cambarus (Bartonius) hamulatus.?Ortmann, 1905c:120.
Cambarus (Cambarus) hamulatus.?Fowler, 1912:341 [by implication].?
Ortmann, 1931:%.
Cambarus hamueatus.?Fleming, 1938:303 [erroneous spelling].
Cambarus (Aviticambarus) hamulatus.?Hobbs, 1969b:99, figs. 5, 13f, 14f,
17j;1974b:10,fig.21.
Cambarus hamuetus.?Bouchard, 1976a:573 [erroneous spelling].
Orconectes haninlatus.?Bmdsky, 1981:199 [erroneous spelling].
TYPES.?Syntypes, MCZ 3678 (male II, female).
TYPE LOCALITY.?Nickajack Cave, Marion County, Ten-
nessee.
RANGE.?From the upper Sequatchie Valley, Tennessee,
NUMBER 480 13
southward to Blount County, Alabama.
HABITAT.?Subterranean streams.
Cambarus (Aviticambarus) jonesi Hobbs and Barr
FIGURE 28
Cambarus jonesi Hobbs and Barr, 1960:19, figs. 11-20.
Cambarus (Aviticambarus) jonesi.?Hobbs, 1969b:99, figs. 5,17k; 1974b:10,
fig. 22.
Aviticambarus jonesi.?Cooper, 1975:54.?Culver, 1982:65.
Carbarus jonesi.?Bouchard, 1976c:17 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
104407, 104408, 104799 (male I, female, male II); paratypes:
TU.USNM.
TYPE LOCALITY.?Cave Spring Cave, 12.1 miles (19.4 km)
northwest of Valhermosa, Morgan County, Alabama.
RANGE.?Tennessee River basin between Florence and
environs of Guntersville, Alabama.
HABITAT.?Subterranean waters.
Subgenus Cambarus Erichson
Astacus.?Fabricius, 1798:407.
Subgenus Cambarus Erichson, 1846:97 [in part].?Fowler, 1912:341 [in
part].?Hobbs, 1969b:109. [Type species, by subsequent designation (Faxon
1898:644), Astacus Bartonii Fabricius, 1798:407. Gender: masculine.]
Cambarus.?Girard, 1852:88.
Subgenus Astacus.?Herklots, 1861:144.
Subgenus Bartonius Ortmann, 1905c:97 [in part].
Bartonius.?Williamson, 1907:749 [intended as subgeneric name].
Procambarus.?Estrada, 1972:73 [lapsus].
Cambaras.?Bouchard, 1973a:106 [erroneous spelling].
Cambarus (Cambarus) bartonii bartonii (Fabricius)
FIGURE 29
Astacus Bartonii Fabricius, 1798:407.
Astacus ciliaris Rafinesque, 1817:42 [Types not extant Type locality: brooks
near Fishkill, Dutchess County, and Newburgh, Orange County, New York.]
Astacus pusillus Rafinesque, 1817:42. [Types not extant. Type locality: brooks
in New York, near "Saratoga, Lake George, Lake Champlain, Ulica,
Oswego, Ac."]
Astacus bartonii.?Harian, 1835:230.
Astacus Bartoni.?Tellkampf, 1845:85.
Astacus (Cambarus) Bartonii.?Erichson, 1846:97.
Cambarus Bartonii.?Girard, 1852:88.
Cambarus montanus Girard, 1852:88. [Types destroyed in the Chicago fire in
1871; paratype(?) ANSP 322 (male II), from James River, Virginia. Type
locality, restricted by Ortmann, 1931:114, "Tributary of James River,
Rockbridge County, Virginia."]
Cambarus pusillus.?Girard, 1852:90.
Astacus (Astacus) Bartonii.?Herklots, 1861:144.
Cambarus Bartoni.?Hagen, 1870:30.
Cambarus bartonii.?Packard, 1880:222.
Cambarus Bartonii Bartonii.?Faxon, 1885a:61 [by implication].
Cambarus bartoni.?Shufeldt, 1898:227.
Cambarus bartonii bartonii.?Hay, 1899b:959 [by implication].
Cambarus bartoni bartoni.?Hay, 19O2b:435.?Crocker, 1957:42, pi. 1: figs.
2,3,6, pi. 2: figs. 5-7.
Cambarus (Bartonius) bartoni.?Ortmann, 1905c: 117,120.
Astacus bartoni.?Ortmann, 1906b:348.
Cambarus bartoni typicus.?Ortmann, 1906b:450.
Cambarus bartonius bartoni.?Chidester, 1908:710.
Cambarus (Cambarus) bartonii.?Fowler, 1912:341.
Cambarus bartonii montanus.?Faxon, 1914:385.
Cambarus montanus montanus.?Ortmann, 1931:106.
Cambarus (Cambarus) montanus montanus.?Ortmann, 1931:114.
Cambarus (Cambarus) bartoni bartoni.?Ortmann, 1931:130.
Cambarus bartoni montanus.?Ortmann, 1931:114.
Cambarus (Cambarus) bartonii bartonii.?Fleming, 1938:303.?
Hobbs,1969b:109.figs. 5,13,14.191; 1974b:10,fig. 23.
Cambarus bartonius.?Roberts, 1944:370 [erroneous spelling].
Canbarus bartoni.?Thompson, 1967:47 [erroneous spelling].
Cambarus bartonni.?Coleman, 1972:21 [erroneous spelling].
Procambarus bartoni.?Estrada, 1972:73 [lapsus calami].
Cambaras bartoni.?Bouchard, 1973a: 106 [erroneous spelling].
Cambarus (Cambarus) bartoni bartonii.?Hobbs III, 1975:298 [erroneous
spelling].
Cambarus (cambarus) bartonii bartonii.?Seaman, 1979:24 [lapsus].
Cambarus boutoni.?Blewett and Eble, 1979:39 [erroneous spelling].
TYPES.?Type, UZM (male II).
TYPE LOCALITY.?North America; "probably neighborhood
of Philadelphia, Pa." (Faxon,1914:423).
RANGE.?New Brunswick, Canada, to northern Georgia, and
eastern parts of Kentucky and Tennessee to the Atlantic Ocean;
restricted to the mountains and foothills in South Carolina and
Georgia.
HABITAT.?Mostly lotic habitats, but occasionally frequent-
ing ponds and often burrowing along streams and in seepage
areas.
Cambarus (Cambarus) bartonii carinirostris Hay
FIGURE 30
Cambarus bartonii carinirostris Hay, 1914:384.
Cambarus (Cambarus) bartoni carinirostris.?Ortmann, 1931:107.
Cambarus (Cambarus) bartonii carinirostris.?Hobbs, 1969b:109, fig. 19m;
1974b:ll,fig. 24.?Thoma, 1982:875.?Thoma and Jezerinac 1982:136.?
Jezerinac, 1983:4.
TYPES.?"Type" and paratypes, USNM 23962 (male I, 7
male II, 15 female); paratypes, MCZ 7399 (male I, male II,
female).
TYPE LOCALITY.?Gandy Creek at Osceola, Randolph
County, West Virginia.
RANGE.?Tributaries of the Cheat, Greenbrier, and Tygart
rivers in West Virginia and widespread in eastern Ohio. Exact
range not known.
HABITAT.?Small to moderate sized streams.
REMARKS.?Both Ortmann (1931) and Hobbs (1972b) have
remarked on the advisability of recognizing this subspecies,
and the latter indicated that "it is questionable that this
subspecies should be recognized." Bouchard (1976b:587)
recognized no subspecies of Cambarus (C.) bartonii and placed
Hay's C. (C.) bartonii carinirostris and C. (C.) bartonii
cavatus in synonomy with this species. Inasmuch as Jezerinac
(1983) and Thoma (1982), who are currently investigating the
subspecies of Cambarus bartonii, continue to recognize the
two non-nominate subspecies, the two taxa are provisionally
retained herein.
14 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambarus (Cambarus) bartonii cavatus Hay
FIGURE 31
Camparus bartonii.?Williamson, 1899:47.
Cambarus bartoni cavatus Hay, 1902b:435.
Cambarus (Bartonius) bartoni.?Ortmann 1905c: 120 [in part].
Cambarus (Cambarus) bartonii cava/m.?Fowler, 1912:341 [by implica-
tion].?Hobbs, 1969b:109,figs. 5,19n; 1974b:ll,fig. 25.
Cambarus bartonii cavatus.?Faxon, 1914:425.
Cambarus (Cambarus) bartoni cavatus.?Ortmann, 1931:127.
Cambarus (Cambarus) species L?Hobbs, 1969b: 109, fig. 5.
Cambarus species K.?Hobbs, 1969b:145 [not p. 109 and fig. 5].
TYPES.?Syntypes, USNM 25017 (male II, female, 2 juv.
male).
TYPE LOCALITY.?Powell River, Tazewell, Claiborne
County, Tennessee.
RANGE.?From Tennessee and northwestern Georgia to
southwestern Virginia and Ohio.
HABITAT.?Streams and springs.
REMARKS.?See "Remarks" under Cambarus (C.) bartonii
carinirostris.
Cambarus (Cambarus) howardi Hobbs and Hall
FIGURE 32
Cambarus (Cambarus) species K.?Hobbs, 1969b: 109, Fig. 5.
Cambarus (Cambarus) howardi Hobbs and Hall, 1969:281, figs. 1-12.?
Hobbs, 1974b:ll,fig. 26.
Cambarus howardi.?Anonymous, 1971:158.
Procambarus howardi.?Wharton, 1978:46 [lapsus calami].
TYPES.?Holotype, allotype, and morphotype, USNM
129866, 129867, 129868 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Sope Creek, tributary to the Chatta-
hoochee River at Paper Mill Road, 1.5 miles (2.4 km) above
mouth, Cobb County, Georgia.
RANGE.?Chattahoochee drainage system from Lumpkin
and Hall counties, Georgia, to Lee County, Alabama.
HABITAT.?Streams, usually in riffle areas.
Cambarus (Cambarus) ortmanni Williamson
FIGURE 34
Bartonius ortmanni Williamson, 1907:749 [Bartonius intended as a subgeneric
name].
Cambarus (Bartonius) ortmanni Williamson, 1907:754, pi. 35: figs. 1-3.
Cambarus (Cambarus) ortmanni.?Fowkr, 1912:341 [by implication].
Hobbs, 1969b: 109. figs. 5,19o; 1974b:l 1, fig. 28.
Cambarus ortmanni.?Faxon, 1914:394.
TYPES.-^Syntypes, CM 74.826 (male I, female), MCZ 7587
(female).
TYPE LOCALITY.?Six-Mile Creek and Craven Ditch, tribu-
tary to Wabash River, above Bluffton, Wells County, Indiana.
RANGE.?Southern Indiana and Ohio, and adjacent northern
Kentucky.
HABITAT.?Lentic situations and burrows (secondary bur-
rower).
Cambarus (Cambarus) sciotensis Rhoades
FIGURE 33
Cambarus bartoni sciotensis Rhoades, 1944b:96.
Cambarus sciotensis?Holt, 1954:170.?Hobbs, Holt, and Walton, 1967:27,
fig. 5.
Cambarus bartonii sciotensis.?Hobbs, 1955b:330.
Cambarus (Cambarus) sciotensis.?Hobbs, 1969b: 109, figs. 5,19p; 1974b:12,
fig. 27.?Jezerinac and Thoma, 1984:120, figs. 6,7.
TYPES.?Holotype, allotype, and "morphotype," USNM
81533 (male I, female, male II); paratypes, MCZ, OSM.
TYPE LOCALITY.?Limestone cliffs below O'Shaughnessy
Dam, Scioto River, Sec. 3, Liberty Township, Delaware
County, Ohio.
RANGE.?Scioto and Little Scioto river drainages in Ohio,
and Kanawha drainage in southwestern Virginia and West
Virginia Jezerinac and Thoma (1984:123) stated that this
crayfish seems to be restricted to the Scioto and Little Scioto
rivers and to tributaries of the Ohio River in Scioto and
Lawrence counties, Ohio. They did not mention the popula-
tions in the Kanawha Basin of Virginia and West Virginia that
formerly have been assigned to this species.
HABITAT.?Riffle areas in small to large streams.
Subgenus Depressicambarus Hobbs
Astacus.?LeConte, 1856:402.
Cambarus.?Hagen, 1870:83.
Subgenus Bartonius Ortmann, 1905c:97 [in part].
Subgenus Cambarus.?Fowler, 1912:341 [in part].
Subgenus Depressicambarus Hobbs, 1969b: 102. [Type species, by original
designation, Astacus latimanus LeConte, 1856:402. Gender masculine.]
Cambarus (Depressicambarus) catagius Hobbs and Perkins
FIGURE 42
Cambarus catagius Hobbs and Perkins, 1967:141, figs. 1-11.
Cambarus (Depressicambarus) catagius.?Hobbs, 1969b: 104, figs. 8, 18c;
1974b:12,fig. 36.?Bouchard, 1978a:30,figs. Id, 2a.
TYPES.?Holotype, allotype, and morphotype, USNM
117779, 117780, 117781 (male I, female, male II); paratypes,
USNM, MCZ, BMNH.
TYPE LOCALITY.?Lawn at East Whittington Street in
southeastern section of Greensboro, Guilford County, North
Carolina.
RANGE.?Known only from the type locality.
HABITAT.?Burrows (primary burrower).
Cambarus (Depressicambarus) cymatilis Hobbs
FIGURE 44
Cambarus (Depressicambarus) cymatilis Hobbs, 1970b:251, figs. 3, 4;
NUMBER 480 15
1974b: 12, fig. 37; 1981:100, figs. 23h, 37e, 39a, 43, 44, 198.?Bouchard,
1978a:30,figs. lcj,m,2b.
Cambarus cymatilis.?Hobbs, 1970b:241.
TYPES.?Holotype, allotype, and morphotype, USNM
129860, 129861,129862 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Western city limits of Chatsworth,
Murray County, Georgia, in lawn and rose garden of Mr.
Charles S. Dunn, off Chestnut Street.
RANGE.?Vicinity of Chatsworth, Murray County, Georgia,
and along Mill Creek, upstream from State Route 74, Bradley
County, Tennessee.
HABITAT.?Burrows (primary burrower).
Cambarus (Depressicambarus) deweesae Bouchard and
Etnier
FIGURE 46
Cambarus (Depressicambarus) deweesae Bouchard and Etnier, 1979:589, fig.
1.
Cambarus deweesae.?Bouchard and Etnier, 1979:596.
TYPES.?Holotype and allotype, USNM 148363, 148364
(male I, female), and morphotype, CM 74.1092; paratypes,
USNM.
TYPE LOCALITY.?Seepage area near east bank of Poplar
Creek at State Highway 61 (south side of highway), Anderson
County, Tennessee.
RANGE.?Known from only four localities in the Clinch
(Anderson County) and Emory (Roane County) river basins in
Tennessee.
HABITAT.?Burrows (primary burrower).
Cambarus (Depressicambarus) englishi Hobbs and Hall
FIGURE 36
Cambarus (Depressicambarus) englishi Hobbs and Hall, 1972:151, fig.
la-n.?Hobbs, 1974b:12. fig. 30; 1981:92, figs. 23j, 37a, 39b, 40, 41,
199.?Bouchard, 1978a:31, figs. lb.h,3c.
Cambarus englishi.?Wharton, 1978:46.
TYPES.?Holotype, allotype, and morphotype, USNM
131700, 131701, 132519 (male I, female, male II); paratypes,
USNM.BMNH.
TYPE LOCALITY.?Tallapoosa River, in riffle area 1 mile
(1.6 km) north of Tallapoosa, Haralson County, Georgia, a few
hundred yards east of bridge on State Route 100.
RANGE.?Endemic in the Tallapoosa Basin, where known
from two subjacent localities in Georgia and from Clay,
Clcburne, and Tallapoosa counties, Alabama.
HABITAT.?Streams.
Cambarus (Depressicambarus) graysoni Faxon
FIGURE 49
Cambarus graysoni Faxon. 1914:393.?Hobbs, 1974b:14.
Cambarus (Depressicambarus) striatus.?Hobbs, 1969b, fig. 8; 1974b, fig. 38.
Cambarus (Depressicambarus) graysoni.?Bouchard, 1976b:589; 1978a:31,
figs. lk,2c,3d,e,4a-L
TYPES.?Syntypes, MCZ 3593 (male 11,3 female).
TYPE LOCALITY.?Bear Creek, a tributary of Green River,
Grayson Springs, Grayson County, Kentucky.
RANGE.?Nashville Basin, Highland Rim, and western edge
of the Cumberland Plateau in Kentucky, Tennessee, and
Alabama.
HABITAT.?Streams.
Cambarus (Depressicambarus) halli Hobbs
FIGURE 35
Cambarus halli Hobbs. 1968a:269,figs. 12-22.
Cambarus (Depressicambarus) halli.?Hobbs, 1969b:104, figs. 8, 18e;
1974b:13, fig. 29; 1981:96, figs. 23i, 37b, 39c, 42, 43, 200.?Bouchard,
1978a:33,figs. la,e/,3f.
TYPES.?Holotype, allotype, and morphotype, USNM
129288, 129289, 129290 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Tributary of the Tallapoosa River, 1.3
miles (2.1 km) south of the river on U.S. Highway 27, Haralson
County, Georgia.
RANGE.?Tallapoosa drainage system in Alabama and
Georgia.
HABITAT.?Streams.
Cambarus (Depressicambarus) hard Hobbs
FIGURE 47
Cambarus (Depressicambarus) harti Hobbs, 1981:104. figs. 23e, 38b, 39d, 41.
45,201.
TYPES.?Holotype, allotype, and morphotype, USNM
148348, 148349, 148350 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Seepage and wooded area adjacent to the
National Fish Hatchery at Warm Springs, Meriwether County,
Georgia.
RANGE.?Known from only two localities in the piedmont
section of the Chattahoochee and Flint river basins in
Meriwether County, Georgia.
HABITAT.?Burrows (primary burrower).
Cambarus (Depressicambarus) latimanus (Le Conte)
FIGURE 38
Astacus latimanus LeConte, 1856:402.
Cambarus latimanus.?Hagen, 1870:83, pi. 1: figs. 43-46; pi. 3: fig.
162.?Faxon, 1885a:69,pl. 2: fig. 3.?Hobbs. 1942b:158,figs. 186-190.
Cambarus obesus var. latimanus.?Packard, 1880:222.
Cambarus Jordani Faxon, 1884:119; 1885a:83,pl. 3: fig. 3. [Type: MCZ 3561
(male II). Type locality: Etowah River, near Rome, Floyd County, Georgia.]
Cambarus jordani.?Hay, 1899b:959.
Cambarus latimus.?Steele, 1902:7 [erroneous spelling].
16 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambarus (Bartonius) latimanus.?Ortmaim, 1905c:120.
Cambarus (Bartoniusjjordani.?Ortmann, 1905c: 120 [by implication].
Cambarus (Cambarus) latimanus.?Fowler, 1912:341 [by implication].
Cambarus (Cambarus) jordani.?Fowler, 1912:341 [by implication].
Cambarus (Depressicambarus) latimanus.?Hobbs, 1969b: 104, figs. If, 8,
13g, 14g, 18f; 1974b:13, fig. 33; 1981:109, figs. 23f. 37c,d, 39e-g. 46-51.
202.?Bouchard 1978a:34.figs. 11,2a,b. 5-8.
Cambarus (Depressicambarus) jordani.?Hobbs, 1969b:104 [by implication].
TYPES.?Syntypes, MCZ 3378 (male I, dry), MCZ 236
(listed by Faxon, 1914:425, as paratypes), ANSP 329 (female).
TYPE LOCALITY.?Athens, Clarke County, Georgia (by
subsequent designation, Faxon 1914:395).
RANGE.?Piedmont and coastal plain from the Tar and Cape
Fear basins in North Carolina southward to the Altamaha and
Apalachicola basins in Georgia and Florida, westward to the
Coosa Basin in Alabama.
HABITAT.?Small to moderately large streams and burrows
(secondary burrower).
Cambarus (Depressicambarus) obstipus Hall
FIOURE37
Cambarus obstipus Hall, 1959:221, figs. 1-9.
Cambarus (Depressicambarus) obstipus.?Hobbs, 1969b: 104, figs. 8, 18g;
1974b:13,fig. 31.?Bouchard, 1978a:37,figs. li.2f,3g.
TYPES.?Holotype and allotype, USNM 102123, 102124
(male I, female); paratypes, USNM.
TYPE LOCALITY.?Black Warrior River at Underwood's
Ferry, 8 miles (12.8 km) downstream from Cordova, Walker
County, Alabama.
RANGE.?Known only from the Black Warrior River system
in Alabama.
HABITAT.?Streams.
Cambarus (Depressicambarus) pyronotus Bouchard
FIGURE 40
Cambarus. latimanus.?Hobbs, 1942b: 159 [in part].
Cambarus (Depressicambarus) pyronotus Bouchard, 1978a:37, fig. 9.
Cambarus pyronotus. ?Bouchard, 1978a:27.
TYPES.?Holotype, allotype, and morphotype, USNM
146761,146762,146763 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Burrows along tributary of the Apalachi-
cola River south of Indian Ridge, Torreya State Park, Liberty
County, Florida.
RANGE.?Known only from the vicinity of the type locality.
HABITAT.?Burrows (primary burrower).
Cambarus (Depressicambarus) reduncus Hobbs
FIGURE 50
Cambarus reduncus Hobbs, 1956b:61, figs. 1-11.
Cambarus (Depressicambarus) reduncus.?Hobbs, 1969b:104, figs. 8, 18h;
1974b:13,fig. 39.
TYPES.?Holotype, allotype, and morphotype, USNM
99183,99184,99185 (male I, female, male II); paratypes, TU,
USNM.
TYPE LOCALITY.?Floodplain pools of Little River, 10.3
miles (16.3 km) west of Winnsboro on State Route 22,Fairfield
County, South Carolina.
RANGE.?Piedmont Province from Orange County, North
Carolina to Richland County, South Carolina.
HABITAT.?Streams, ponds, and burrows (primary or secon-
dary burrower).
Cambarus (Depressicambarus) reflexus Hobbs
FIGURE 39
Cambarus (Depressicambarus) reflexus Hobbs, 1981:120. figt. 23c, 38a, 39h,
52.53,203.
TYPES.?Holotype, allotype, and morphotype, USNM
148116, 148117, 148118 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Savannah River floodplain at U.S. High-
way 301, Allendalc County, South Carolina.
RANGE.?Coastal plain of the Savannah through the Santcc
basins in Georgia and South Carolina.
HABITAT.?Burrows (primary burrower).
Cambarus (Depressicambarus) sphenoides Hobbs
FIOURE41
Cambarus bartonii bartonii.?Hobbs and Shoup, 1942:636, figs. 5,8.
Cambarus sphenoides Hobbs, 1968a:262, figs. 1-11.
Cambarus (Depessicambarus) sphenoides.?Hobbs, 1969b: 104. figs. 8, 18i;
1974b: 13, fig. 35.?Bouchard. 1978a:40, figs. lg,3j.
TYPES.?Holotype, allotype, and morphotype, USNM
129325, 129326, 129327 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Tributary to Clear Creek (Emory River
drainage), 11.2 miles (17.9 km) north of Crossville, Cumber-
land County, Tennessee, on U.S. Highway 127.
RANGE.?Stream tributaries of the Cumberland, Emory,
Sequatchie, and Tennessee river systems on the Cumberland
Plateau in Kentucky and Tennessee.
HABITAT.?Streams.
Cambarus (Depressicambarus) striatus Hay
FIGURE 48
Camburus sp. [Ashland City, Tennessee].?Faxon. 1885c:358 [lapsus for
Cambarus].
Cambarus latimanus striatus Hay, 19O2b:437.
Cambarus (Bartonius) latimanus var. striatus.?Ortmann, 1905c:119 [by
implication].
Cambarus (Cambarus) latimanus striatus.?Fowler, 1912:341 [by implica-
tion].
Cambarus (Cambarus) bartoni striatus.?Ortmann, 1931:140.
Cambarus (Cambarus) bartonii striatus.?Fleming, 1938:303.
NUMBER 480 17
Cambarus latimanus.?Fleming, 1939:311 [erroneous spelling].
Cambarus lalimus.?Fleming, 1939:319 [erroneous spelling].
Cambarus (Cambarus) floridanus Hobbs, 1941b:l 14, figs. 1,4,5, 8,9,16,19,
22, 25. 31, 32. [Holotype and allotype: USNM 79341. "Morphotype":
USNM 79344. Paratypes: USNM. MCZ, FSBC. Type locality: 12 miles (19.2
km) west of Tallahassee on State Route 19, Leon County, Florida.]
Cambarus bartonii striatus.?Hobbs, 1942a:354 [by implication].
Cambarus floridanus.?Hobbs, 1942a:356; 1942b:161,pl. 11: figs. 191-195.
Cambarus barloni striatus.?Rhoades, 1944a: 114.
Cambarus striatus.?Hobbs, 1956b:61.
Cambarus (Depressicambarus) striatus.?Hobbs, 1969b: 102, fig. 18j;
1974b:14 [not fig. 38, see C. (D.) graysoni herein]; 1981:127, figs. 23g,
38c,d,39i. 53-55.204.?Bouchard, 1978a:40, figs. 2d,e, 10-14.
Cambarus (Depressicambarus) floridanus.?Hobbs, 1969b: 104, figs. 8,18d.
TYPES.?Syntypes, USNM 20519 (male I, 3 male II, 4
female), MCZ 7348 (male I, male II, 3 female).
TYPE LOCALITY.?Nashville, Davidson County, Tennessee.
RANGE.?From the Savannah and Ochlockonee basins in
South Carolina, Georgia, and Florida westward into Missis-
sippi and northward through Tennessee into the Green
watershed in Kentucky.
HABITAT.?Streams and burrows (primary burrower in the
southern part of the range).
Cambarus (Depressicambarus) strigosus Hobbs
F1OURE45
Cambarus sp.?Wharton, 1978:46.
Cambarus (Depressicambarus) strigosus Hobbs, 1981:134, figs. 23d, 38e,
39jJ.41.56.2O5.
TYPES.?Holotype, allotype, and morphotype, USNM
148284,148574, 148575 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Roadside ditch within 30 meters of Susan
Smith Branch (tributary to Long Creek and Broad River) west
of State Route 17 on unnumbered county road, Wilkes County,
Georgia.
RANGE.?Known from only five localities in the Broad and
Little river basins in Elbert, Oglethorpe, and Wilkes counties,
Georgia.
HABITAT.?Burrows (primary burrower).
Cambarus (Depressicambarus) truncatus Hobbs
FIGURE 43
Cambarus (Depressicambarus) truncatus Hobbs, 1981:140, figs. 23b, 38f,
39k,m,41.57,206.
TYPES.?Holotype, allotype, and morphotype, 116966,
146649,146650; paratypes, USNM.
TYPE LOCALITY.?Roadside ditch 15.4 miles (24.6 km) east
of Irwinton, Wilkinson County, Georgia, on State Route 57.
RANGE.?Known from only four localities on the Fall Line
Hills District in the Oconee Basin of Laurens and Wilkinson
counties, Georgia.
HABITAT.?Burrows (primary burrower).
Submenus Erebicambarus Hobbs
Cambarus.?Hay, 1902a:232.
Subgenus Bartonius Ortmann, 1905c:120.
Subgenus Cambarus.?Fowler, 1912:341.
Subgenus Bartontius.?Rioja, 1941:193 [erroneous spelling].
Subgenus Erebicambarus Hobbs, 1969b:99 [Type species, by original
designation, Cambarus barloni tenebrosus Hay, 19O2a:232. Gender
masculine.]
Subgenus Erebicambaruss.?Hobbs III, 1975:291 [erroneous spelling].
Subgenus Eribkambarus.?Hobbs III, 1980:148 [erroneous spelling].
Cambarus (Erebicambarus) hubbsi Creaser
FIGURE 51
Cambarus (Bartonius) hubbsi Creaser, 193la:4,figs. 7-12.
Cambarus hubbsi.?Hobbs, 1942a:356.?Williams, 1954:904. figs. 233.
235-245.?Pflieger. 1987a:23; 1987b:7.
Cambarus (Erebicambarus) hubbsi.?Hobbs, 1969b: 100, fig. 17d; 1974b:14,
fig. 40.?Gardner, 1986:19.
TYPES.?Holotype and allotype, USNM 98332,98333 (male
I, female); paratypes, USNM.
TYPE LOCALITY.?Little Creek, tributary to St. Francis
River, 1 mile (1.6 km) northeast of Chloride, Iron County,
Missouri.
RANGE.?Southeastern Missouri and northeastern Arkansas.
HABITAT.?Streams, burrows, and caves.
Cambarus (Erebicambarus) hubrichti Hobbs
FIGURE 53
Cambarus hubrichti Hobbs, 1952c:689. figs. 1-8.?Hobbs and BUT, 1960:23,
figs. 21-30.?Pflieger, 1987a:24; 1987b:8.
Cambarus hubricht.?Hobbs, 1952c:693 [erroneous spelling].
Cambarus Hubrichti.?Cooper and Poulson, 1968:130, fig. 8.
Cambarus (Erebicambarus) hubrichti.?Hobbs, 1969b:100, fig. 17e;
1974b:14,fig. 42.?Gardner, 1986:19.20 [fig.].
TYPES.?Holotype and allotype, USNM 92295,92296 (male
II, female); paratypes, USNM.
TYPE LOCALITY.?Stream in Lewis Cave, 15 miles (24 km)
northwest of Doniphan, Ripley County, Missouri.
RANGE.?Current, Eleven Point, and Meramec river basins
in Carter, Oregon, Phelps, Pulaski, Ripley, Shannon, and Texas
counties, Missouri.
HABITAT.?Subterranean waters and springs.
Cambarus (Erebicambarus) laevis Faxon
FIGURE 56
?Cambarus barloni brevis Rietz, 1912 [nomen nudum; see Page, 1985b:335].
Cambarus bartonii laevis Faxon, 1914:391.
Cambarus (Cambarus) bartoni laevis.?Ortmann, 1931:143.
Cambarus bartoni laevis.?Fleming, 1939:311.
Cambarus (Erebicambarus) laevis.?Hobbs, 1969b:100, figs. 6, 17f;
1974b: 15, fig. 46.
Cambarus laevis.?Hobbs III, 1971:140.
Cambarus (erebicambarus) laevis.?Hobbs III, 1973:58 [lapsus].
18
Cambarus laevi.?Holt, 1973:250 [erroneous spelling].
Cambarus (Erebicambarus) Laevis.?Hdbbs III, 1975:280.
Cambarus (Erebicambaruss) laevis.?Hobbs III, 1975:291 [erroneous spell-
ing].
Cambarus (Eribicambarus) laevis.?Hobbs III, 1980:148 [erroneous spelling].
TYPES.?Holotype and paratype.MCZ 3812 (holotypic male
II, female); paratypes, MCZ, USNM.
TYPE LOCALITY.?Bloomington, Monroe County, Indiana.
RANGE.?Southern parts of Illinois(?) and Indiana, and
northern Kentucky(?).
HABITAT.?Streams, springs, and caves.
REMARKS.?Page (1985b:428) tentatively referred the Illi-
nois populations possibly assignable to this taxon to Cambarus
(E.) tenebrosus, and as pointed out by him, it is probable that
when a careful study of the subgenus Erebicambarus is made,
C. (E.) laevis will be declared a synonym of C. (E.) tenebrosus.
Until such a study is conducted, I deem it advisable to maintain
specific status for it. Jezerinac and Thoma (1984:121) omitted
this crayfish in their key to the genera Cambarus and
Fallicambarus in Ohio because all of the material that they
examined from the state that had been assigned to C. (E.) laevis
was "assignable to either C. (C.) ortmanni or C. (C.) bartonii
cavatus."
Cambarus (Erebicambarus) ornatus Rhoades
FIGURE 55
Cambarus bartoni ornatus Rhoades, 1944a: 144, fig. lOa-f.
Cambarus bartonii ornatus.?Hobbs and Shoup, 1947:145.
Cambarus ornatus.?Hobbs and Barr, 1960:15.
Cambarus (Erebicambarus) ornatus.?Hobbs, 1969b:100, figs. 6, 17g;
1974b:15,fig. 45.
TYPES.?Holotype, allotype, and "morphotype," USNM
81330, 81331, 81332 (male I, female, male II); paratypes,
MCZ, USNM, OSM.
TYPE LOCALITY.?Ellis Branch, 4 miles (6.4 km) west of
Carrollton, Carroll County, Kentucky.
RANGE.?Ohio Basin from the lower Kentucky River system
to the Salt River, Kentucky.
HABITAT.?Intermittent streams.
Cambarus (Erebicambarus) rusticiformis Rhoades
FIGURE 52
Cambarus rusticiformis Rhoades, 1944a:133, fig. 8a-f.?Page, 1985b:432,
figs. 164-166.
Cambarus (Erebicambarus) rusticiformis.?Hobbs, 1969b: 100, figs. 2a, 6,
13h,17h;1974b:15.fig. 41.
TYPES.?Holotype, allotype, and "morphotype," USNM
81324, 81325, 81326 (male I, female, male II); paratypes,
MCZ, OSM.
TYPE LOCALITY.?Little River, 0.5 mile (0.8 km) west of
Cadiz, Trigg County, Kentucky.
RANGE.?Cumberland and Green drainage systems in
Kentucky and Tennessee, and Ohio River in southern Illinois.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
HABITAT.?Swift streams.
Cambarus (Erebicambarus) tenebrosus Hay
FIGURE 54
Cambarus bartoni tenebrosus Hay, 1902a:232.
Cambarus (Bartonius) bartoni.?Ortmann, 19O5c:12O [in part].
Cambarus (Cambarus) bartoni tenebrosus.?Fowler, 1912:341 [by implica-
tion].
Cambarus bartonii tenebrosus.?Faxon, 1914:424.
Cambarus tenebrosus.?Garman, 1920:42.?Hobbs, 1959:898.?Page,
1985b:428,figs. 160-163.
Cambarus Bartoni tenebrosus.?Chappuis, 1927:92.
Cambarus (Bartonius) Bartoni tenebrosus.?Fage, 1931:373.
Cambarus (Cambarus) bartoni tenebrosus.?Ortmann, 1931:145.
Cambarus (Bartontius) Bartoni tenebrosus.?Rioja, 1941:193 [erroneous
spelling].
Cambarus (Cambarus) cahni Rhoades, 1941b:146, fig. 36a-d. [Holotype and
allotype: USNM 80031,80032 (male Il.female). Paratypes: MCZ, Alabama
Mus. Nat. Hist., ANSP, Leslie Hubricht, OSM. Type locality: Belgreen
Cave,NWV4.SWy,4,Sec. 12.T. 7S.R. 13W,Franklin County, Alabama.]
Cambarus (Cambarus) bartonii tenebrosus.?Hobbs, 1941b: 114.
Cambarus cahni.?Hobbs, 1942a:354.?Hobbs and Barr, 1960:25. figs.
31-37.
Cambarus (Bartonius) bartoni tenebrosus.?Hart, 1962:121.
Cambarus (Erebicambarus) tenebrosus.?Hobbs, 1969b: 100, figs. 6,14h, 17i;
1974b: 15, fig. 43.
Cambarus (Erebicambarus) cahni.?Hobbs, 1969b: 100, figs. 6,17c.
Cambarus tenebrorus.?H rod sky, 1981:49 [erroneous spelling].
TYPES.?Syntypes, USNM 22346 (male 11,2 female).
TYPE LOCALITY.?Echo River, Mammoth Cave, Edmonson
County, Kentucky.
RANGE.?Mammoth Cave southward to northern Alabama.
Illinois records based on assumption of C. (E) laevis being a
synonym.
HABITAT.?Streams and subterranean waters (troglophilic).
REMARKS.?See "Remarks" under Cambarus (E.) laevis.
Subgenus Exilicambarus Bouchard and Hobbs
Subgenus Exilicambarus Bouchard and Hobbs, 1976:2. [Type species, by
original designation, Cambarus (Exilicambarus) cracens Bouchard and
Hobbs, 1976:2. Gender masculine.]
Cambarus (Exilicambarus) cracens Bouchard and Hobbs
FIGURE 57
Cambarus (Exilicambarus) cracens Bouchard and Hobbs, 1976:2, figs. 1,2.
TYPES.?Holotype, allotype, and morphotype, USNM
146082, 146083,146084 (male I, female, male II); paratypes,
USNM,BMNH.
TYPE LOCALITY.?Short Creek at State Route 75,1.1 mile
(1.8 km) southwest of the junction with State Route 68 (Sec.
36, T. 8S, R. 4E), Marshall County, Alabama.
RANGE.?Southeastern tributaries of Guntersville Lake
(impounded Tennessee River) in DeKalb and Marshall coun-
ties, Alabama.
HABITAT.?Streams.
NUMBER 480 19
Subgenus Hiaticambarus Hobbs
Cambarus.?Girard, 1852:90.
Subgenus Bartonius Ortmann, 1905c:97 [in part].
Subgenus Cambarus.?Fowler, 1912:341 [in part].
Subgenus Hiaticambarus Hobbs, 1969b: 105; 1981:145. [Type species, by
original designation, Cambarus longulus Girard, 1852:90. Gender: mascu-
line.]
Cambaroides.?Unestam, 1969:204 [lapsus].
Cambarus (Hiaticambarus) chasmodactylus James
FIGURE 64
Cambarus longulus chasmodactylus James, 1966:14, fig. 1, pi. lb,d,e,q,u-z,zz.
Cambarus chasmodactylus.?Hobbs, 1968b:K17.
Cambarus (Hiaticambarus) chasmodactylus.?Hobbs, 1969b: 106, figs. 10,
18k; 1974b: 15, fig. 50.
TYPES.?Holotype, allotype, and morphotype, USNM
115513,115514, 115516 (male I, female, male II); paratypes,
MCZ, TU, IBM, USNM, Hugo L. James.
TYPE LOCALITY?East Fork of the Greenbrier River, 9.7
miles (15.5 km) west of Virginia state line on U.S. Highway
250, Pocahontas County, West Virginia.
RANGE.?New River drainage system in northwestern North
Carolina and Virginia, and the Blue Stone and Greenbrier
basins in eastern West Virginia.
HABITAT.?Rocky streams.
Cambarus (Hiaticambarus) coosawattae Hobbs
FIGURE 58
Cambarus (Hiaticambarus) coosawattae Hobbs, 1981:150, figs. 22b, 58a, 59,
60,207.
TYPES.?Holotype, allotype, and morphotype, USNM
148112,148113,148114 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Cartecay River, 6 miles (9.6 km) east
southeast of Ellijay, just off Route S1010, near Flint Hill
Church, Gilmer County, Georgia.
RANGE.?Known only from the Coosawattee watershed
(Coosa River basin) in Gilmer County, Georgia.
HABITAT.?Streams (primarily riffle areas).
Cambarus (Hiaticambarus) fasciatus Hobbs
FIGURE 59
Cambarus (Hiaticambarus) fasciatus Hobbs, 1981:156, figs. 22e, 58b, 60,61,
208.
TYPES.?Holotype, allotype, and morphotype, USNM
147917, 147918, 147919 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Etowah River, 0.2 mile (0.32 km) west of
State Route 52 on unpaved road near Davis Chapel, Lumpkin
County, Georgia.
RANGE.?Etowah River basin in northwestern Georgia.
HABITAT.?Streams.
Cambarus (Hiaticambarus) girardianus Faxon
FIGURE 60
Cambarus Girardianus Faxon 1884:117; 1885a:78,pL 4: fig. Lpl. 9: fig. 2a,
2a'.
Cambarus girardianus.?Faxon 1898:650.
Cambarus extraneus girardianus.?Faxon 1898:650.
Cambarus (Cambarus) extraneus girardianus.?Fowler, 1912:341 [by impli-
cation].
Cambarus (Bartonius) girardianus.?Ortmann, 1931:97.
Cambarus extraneus Girardanius.?Fleming, 1938:299 [erroneous spelling].
Cambarus (Hiaticambarus) girardianus.?Hobbs, 1969b:106, figs. 10, 181;
1974b:16,fig. 47; 1981:162,figs. 22f.58c,60,62,63,209.
TYPES.?Syntypes, MCZ 3560 (male II, female), USNM
4882 (male II, 2 female).
TYPE LOCALITY.?Cypress Creek, Lauderdale County, Ala-
bama.
RANGE.?Tennessee River drainage system from northeast-
ern Mississippi to Georgia and northward to the vicinity of
Knoxville, Tennessee. Possibly the dominant Hiaticambarus in
the upper Coosa Basin in Alabama is referable to this species.
HABITAT.?Streams.
Cambarus (Hiaticambarus) longirostris Faxon
FIGURE 61
Cambarus Bartonii, var. longirostris Faxon, 1885a:64.
Cambarus bartonii longirostris.?Faxon, 1890:623.
Cambarus bartonii spinirostris.?Faxon 1890:623 [lapsus forC. b. longiros-
tris].
Cambarus longulus longirostris.?Hay, 1899b:959.?James, 1966:9, fig. 2a,b,
pi. lc,f,g,n,o,r,s.
Cambarus (Bartonius) bar lord longirostris.?Ortmann, 1905c: 135 [by impli-
cation].
Cambarus (Cambarus) bartonii longirostris.?Fowler, 1912:341 [by implica-
tion].
Cambarus (Cambarus) longulus longirostris.?Ortmann, 1931:121.
Cambarus longerosilis.?Brimley, 1938:503 [erroneous spelling].
Cambarus longulus longerostris.?Schurr and Stamper, 1962:474 [erroneous
spelling].
Cambarus bartonii var. longirostris.?James, 1966:2.
Cambarus spinirostris.?James, 1966:2.
Cambarus longirostris.?James, 1966:2.
Cambarus longulus longirosilis.?James, 1966:8 [erroneous spelling].
Cambarus (Hiaticambarus) longirostris.?Hobbs, 1969b:106, figs. 10, 18m;
1974b:16,fig. 48.
Cambarus (Hiaticambarus) girardianus.?Bouchard, 1976a:572 [in part];
1976b:588 [in part].
TYPE.?Holotype, MCZ 3629 (male II).
TYPE LOCALITY.?"Eastern Tennessee and West Virginia"
(Faxon, 1885a). Restricted to Doe River, Elizabethton, Carter
County, Tennessee, designated by Ortmann, 1931:121.
RANGE.?Tennessee River Basin upstream from below the
mouth of the Sequatchie Riven in Georgia, Tennessee, North
Carolina, and Virginia. Introduced into headwaters of the
Savannah River in South Carolina.
HABITAT.?Rocky segments of streams.
20 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambarus (Hiaticambarus) longulus Girard
FIGURE 62
Cambarus longulus Girard, 1852:90.
Cambarus Bartonii longulus.?Faxon, 1885a:66 [by implication].
Cambarus longulus longulus.?Hay 1899b:959 [by implication].?James,
1966:4, fig. 2c, pi. la, h-m,p,giun.
Cambarus (Bartonius) longulus.?Orimann, 1905c: 120.
Cambarus (Cambarus) longulus longulus.?Fowler, 1912:341 [by implica-
tion].?Ortmann, 1931:106.
Cambarus bartonii longulus.?Faxon, 1914:387.
Cambarus bartoni longulus.?Ortmann, 1931:118.
[Cambarus longulus] longulous.?James, 1966:22 [erroneous spelling].
Cambaroides longulus.?Unestam, 1969:204 [lapsus for Cambarus].
Cambarus (Hiaticambarus) longulus.?Hobbs, 1969b:106, figs, la, 10, 13c,
14c,18n;1974b:16,fig. 49.
TYPES.?Probably destroyed in the Chicago fire, 1871
(Faxon, 1914:124).
TYPE LOCALITY.?"Middle States" (Girard, 1852). South
River, Waynesboro, Augusta County, Virginia, designated by
Ortmann, 1931. Emended by Hobbs (1967a:129) to Rockfish
River, south of Afton, Nelson County, Virginia.
RANGE.?Atlantic watershed from the James drainage in
Virginia and West Virginia south to the Yadkin Basin in North
Carolina, in mountains and piedmont
HABITAT.?Rocky areas of streams.
Cambarus (Hiaticambarus) manningi Hobbs
FIGURE 63
Untitled color photograph.?Bouchard in Anonymous, 1978:175.
Cambarus (Hiaticambarus) manningi Hobbs, 1981:175, figs. 22c, 58e, 65, 67,
68,211.
TYPES.?Holotype, allotype, and morphotype, USNM
147911,147912,147913 (male I, female, male II); paratypes,
BMNH.USNM.
TYPE LOCALITY.?Little Cedar Creek (Coosa River basin)
near school for deaf on outskirts of Cave Spring, Floyd County,
Georgia.
RANGE.?Coosa River basin in northwestern Georgia,
southeastern Tennessee, and northeastern Alabama.
HABITAT.?Rocky areas of streams.
Cambarus (Hiaticambarus) speciosus Hobbs
FIGURE 65
Cambarus (Hiaticambarus) species H?Hobbs, 1969b: 106,141,fig. 10.
Cambarus (Hiaticambarus) speciosus Hobbs, 1981:181, figs. 22g, 58f, 65,69,
212.
TYPES.?Holotype, allotype, and morphotype, USNM
146023,146024,146025 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Talking Rock Creek at State Route 5,
Pickens County, Georgia.
RANGE.?Coosawattee River basin in Pickens, Gilmer, and
Murray counties, Georgia.
HABITAT.?Moderately to swiftly flowing streams.
Subgenus Jugicambarus Hobbs
Astacus.?Erichson, 1846:87.
Subgenus Cambarus Erichson, 1846:96.?Fowler, 1912:341 [in part].
Cambarus.?Girard, 1852:88 [in part].
Subgenus Bartonius Ortmann, 1905c:97 [in part].
Subgenus Jugicambarus Hobbs, 1969b: 106. [Type species, by original
designation, Cambarus bartonii asperimanus Faxon, 1914:391. Gender:
masculine.]
Subgenus Depressicambarus ?Hobbs and Hall, 1969:287.
Subgenus Veticambarus.?Hobbs, 1970b:243.
Subgenus Jugocambarus.?Hart and Hart, 1974:74 [erroneous spelling].
Cambarus (Jugicambarus) asperimanus Faxon
FIOURE72
Cambarus bartonii asperimanus Faxon, 1914:391.
Cambarus (Cambarus) bartoni asperimanus.?Ortmann, 1931:137.
Cambarus asperimanus.?Brimley, 1938:503.
Cambarus (Jugicambarus) asperimanus.?Hobbs, 1969b: 107, figs. Id, 9,13d,
14d, 18o; 1974b:16.fig. 57; 1981:189, figs. 24b, 70a, 71.72.213.
TYPES.?Syntypes, USNM 47375 (2 male I).
TYPE LOCALITY.?Flat Creek, Montreat, Buncombe County,
North Carolina.
RANGE.?Mountains of North Carolina, South Carolina, and
Georgia, in the headwaters of the French Broad, Little
Tennessee, Catawba, Broad, Saluda, and Savannah rivers and
in the Watauga Basin in Tennessee.
HABITAT.?Seepage areas and mountain streams.
Cambarus (Jugicambarus) batchi Schuster
FIGURE 80
Cambarus (Jugicambarus) batchi Schuster, 1973:24,fig. 3; 1976:227,fig. 1.
TYPES.?Holotype, allotype, and morphotype, USNM
144390, 144391,144392 (male I, female, male II); paratypes,
USNM, MMZ, UTM, Donald Batch.
TYPE LOCALITY.?Burrows around ponds in the Central
Kentucky Wildlife Management Area, off U.S. Highway 421
approximately 2.5 miles (4.0 km) southeast of Kingston,
Madison County, Kentucky.
RANGE.?Known only from the type locality.
HABITAT.?Burrows (primary burrower).
Cambarus (Jugicambarus) bouchardi Hobbs
FIGURE 66
Cambarus (Veticambarus) bouchardi Hobbs, l970b:245, figs. 1,2.
Cambarus bouchardi.?Hobbs, 1970b:241.
Cambarus (Jugicambarus) bouchardi.?Bouchard, 1973a: 105.?Hobbs,
1974b:17,fig. 51.
TYPES.?Holotype, allotype, and morphotype, USNM
NUMBER 480 21
130295, 130296, 130297 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Perkins Creek, 6.9 miles (11 km) north of
Oneida, Scott County, Tennessee, on U.S. Highway 27.
RANGE.?Known only from the Big South Fork of the
Cumberland basin in Scott County, Tennessee.
HABITAT.?Streams.
Cambarus (Jugicambarus) brachydactylus Hobbs
FIGURE 74
Cambarus brachydactylus Hobbs, 1953c:20, pi. 1: figs. 1-10.
Cambarus (Jugicambarus) brachydactylus.?Hobbs, 1969b: 107, figs. 9, 18p;
1974b: 17, fig. 58.
TYPES.?Holotype and allotype, USNM 93155 (male I,
female); morphotype, USNM 93156 (male II); paratypes,
MCZ.USNM.TU.
TYPE LOCALITY.?Louise Creek, 13.9 miles (22.2 km) south
of Clarksville, Montgomery County, Tennessee.
RANGE.?Tributaries of the Cumberland River on the
western Highland Rim in Montgomery and Dickson counties,
Tennessee.
HABITAT.?Riffle areas of streams.
Cambarus (Jugicambarus) carotinus (Erichson)
FIGURE 77
Astacus Carolina* Erichson, 1846:87.
Astacus (Cambarus) Carolinus Erichson, 1846:96.
Astacus (Cambarus) carolinus.?Dana, 1852:522 [by implication].
Cambarus carolinus.?Girard, 1852:88.?Dewees, 1972:1, figs. 1,2,5.
Cambarus Carolinus.?Hagen, 1870:8.
Cambarus (Bartonius) carolinus.?Ortmann, 1905c: 120.
Cambarus carolintnsis.?Adams, 1907:899 [erroneous spelling].
Cambarus (Cambarus) carolinus.?Fowler, 1912:341 [by implication].
Cambarus carolinus carolinus.?Faxon, 1914:339.
Cambarus (Jugicambarus) carolinus.?Hobbs, 1969b: 107 [in part], figs. 9,
19a; 1974b:17,fig. 62.?Hobbs and Bouchard, 1973:51, figs. 3,4.
TYPES.?Not extant. Topotypes, USNM 144164 (2 male I,
female, female with young), and others.
TYPE LOCALITY.?Greenville, Greenville County, South
Carolina. Restricted by Hobbs and Bouchard (1973:55) to
"along Wildcat Creek, 8.8 miles (14.1 km) north of the junction
of U.S. Highway 29 and State Route 101 on latter
RANGE.?From the upper Broad River basin in the northern
part of South Carolina westward into North Carolina and
Tennessee (south of the French Broad River) almost to the
Tennessee River (see Dewees, 1972).
HABITAT.?Burrows (primary burrower).
Cambarus (Jugicambarus) causeyi Reimer
FIGURE 83
Cambarus causeyi Reimer, 1966:9, figs. 1 -8.
Cambarus (Jugicambarus) causeyi.?Hobbs, 1969b:107, fig. 19b; 1974b:17,
fig. 65.
TYPES.?Holotype, allotype, USNM 116678,116679 (male
I, female); paratype (juvenile female), USNM.
TYPE LOCALITY.?Spring and natural pond, 4 miles (6.4 km)
west of Sandgap, Pope County, Arkansas, on State Route 124.
RANGE.?Known only from the Arkansas River basin in
Pope County, Arkansas.
HABITAT.?Streams and complex burrows.
Cambarus (Jugicambarus) conasaugaensis Hobbs and
Hobbs
FIGURE 71
Cambarus conasaugaensis Hobbs and Hobbs, 1962:41, figs. 1-10.
Cambarus (Jugicambarus) conasaugaensis.?Hobbs, 1969b: 107, figs. 9, 19c;
1974b:17,fig. 56; 1981:192,figs. 24c,70b,72,73-76,214.
Cambarus conasougaensis.?Schuster, 1973:9 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
107156, 107157, 107158 (male I, female, male II); paratypes,
BMNH.USNM.
TYPE LOCALITY.?Small cascading, but sluggish, tributary
of Holly Creek (to Conasauga River), 2 miles (3.2 km) east of
Chatsworth, Murray County, Georgia, on U.S. Highway 76.
RANGE.?Headwater tributaries of the Conasauga, Coo-
sawattee, and Etowah rivers in Dawson, Fannin, Gilmer,
Murray, Lumpkin, and Pickens counties, Georgia, and in the
Hiwassee Basin in Bledsoe and Polk counties, Tennessee.
HABITAT.?Streams.
Cambarus (Jugicambarus) crinipes Bouchard
FIGURE 73
Cambarus bartonii bartonii.?Hobbs and Shoup, 1942:636, fig. 6,7.
Cambarus crinipes Bouchard, 1973a: 106, fig. 1.
Cambarus (Jugicambarus) crinipes Bouchard, 1973a: 109.
TYPES.?Holotype, allotype, and morphotype, USNM
132342, 132343,132344 (male I, female, male II); paratypes,
USNM.UTM.
TYPE LOCALITY.?White Oak Creek, a tributary of Clear
Fork Creek (Big South Fork of the Cumberland River system)
at U.S. Highway 27 in Sunbright, Morgan County, Tennessee.
RANGE.?Headwaters of the East Fork of the Obey River,
Clear Creek (Emory River system) and Clear Fork Creek (Big
South Fork of the Cumberland River in Cumberland, Fentress,
Morgan, Overton, Pickett, Putnam, and Scott counties, Tennes-
see, and Rockcastle County, Kentucky.
HABITAT.?Streams, mostly small.
Cambarus (Jugicambarus) cryptodytes Hobbs
FIGURE 82
Cambarus (Cambarus) cryptodytes Hobbs, 1941b:110, figs. 2,3,7,11,13,15,
18, 21, 24, 28, 29; 1942b:162, pi. 11: figs. 196-200; 1974b:18, fig. 64;
1981:197, figs. 24e, 77, 87, 215.?Hobbs and Barr, 1960:31, figs.
48-57.?Hobbs, Hobbs, and Daniel, 1977:83, fig. 40.
22 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambarus cryptodytes Hobbs, 1941b:113.?Caine, 1974:12, 43, 62.?Franz
and Lee, 1982:56,69.
Cambarus (Jugicambarus) cryptodytes.?-Hobbs. 1969b: 107, figs. 9, 19d;
1981:197, figs. 24e, 77,87,215.
cambarus cryptodytes.?Padgett, 1970:19 [lapsus calami].
Cambarus (Jugicambarus) crytodytes.?Hobbs, 1974b, fig. 64 [erroneous
spelling].
TYPES.?Holotype, allotype, and "morphotype," USNM
79339, 79340, 79343 (male I, female, male II); paratypes,
MCZ.USNM.
TYPE LOCALITY.?R.W. Williams farm, 2 miles (3.2 km)
south of Graceville, Jackson County, Florida (NW lM, SW 74,
NE 74, Sec. 15, T. 6W, R. 13W), 0.2 mi (320 m) W of St. Rte.
77 on St. Rte. 277.
RANGE.?Known from several caves in Jackson and Wash-
ington counties, Florida, and Climax Cave, 3 miles (4.8 km)
north of Climax, Decatur County, Georgia.
HABITAT.?Subterranean waters.
Cambarus (Jugicambarus) distans Rhoades
FIGURE 68
Cambarus distans Rhoades, 1944a:136, fig. 9a-f.?Bouchard, 1976a:573;
1976b:585.?Hobbs, 1981:200, fig. 78.
Cambarus diistans.?Rhoades, 1944a: 139 [erroneous spelling].
Cambarus (Jugicambarus) distans.?Hobbs, 1969b:107, figs. 9, 19e;
1974b: 18, fig. 53.
TYPES.?Holotype, allotype, and "morphotype," USNM
81327,81329,81328 (male I, female, male II); paratypes, CM,
USNM,OSM.
TYPE LOCALITY.?"Cumberland River and small tributary,
just above Cumberland Falls, McCreary County, Kentucky"
(Rhoades, 1944a: 139).
RANGE.?Kentucky, Cumberland, and Tennessee river
drainages in Kentucky and Tennessee southward to Sand
Mountain in Alabama and Georgia.
HABITAT.?Streams.
Cambarus (Jugicambarus) dubius Faxon
FIGURE 78
Cambarus dubius Faxon, 1884:114; 1885a:70, pi. 4: fig. 3; pi. 8: figs. 7,
7'.?Dewees, 1972:1,figs, lb-1,2c-h, 3,4,5b-h,6-16.
Cambarus carolmus dubius.?Faxon, 1914:396,425.
Cambarus (Jugicambarus) dubius.?Hobbs and Bouchard, 1973:62.?Hobbs,
1974b: 18. fig. 60.
Cambraus dubius.?Peters, 1975:iii [erroneous spelling].
TYPES.?Holotype, MCZ 3631 (male I).
TYPE LOCALITY.?Terra Alta (Cranberry Summit), Preston
County, West Virginia.
RANGE.?Poorly understood, but extending southward
through the Allegheny Mountains from southwestern Pennsyl-
vania and Kentucky into southwestern Virginia, northwestern
North Carolina and as far south in Tennessee as the junction of
the Clinch and Holston rivers, and westward on the Cumber-
land Plateau to headwaters of the South Fork of the
Cumberland and Caney Fork rivers in Fentress and Cumber-
land counties, Tennessee. (See Dewees, 1972.)
HABITAT.?Burrows (primary burrower).
Cambarus (Jugicambarus) friaufi Hobbs
FIGURE 75
Cambarus friaufi Hobbs, 1953c:24, figs. 11-18.
Cambarus (Jugicambarus) friaufi.?Hobbs, 1969b: 107, figs. 9,19f; 1974b: 18,
fig. 59.
TYPES.?Holotype and allotype, USNM 93157 (male I,
female); paratypes, MCZ, USNM.
TYPE LOCALITY.?Small stream tributary of Cumberland
River at Elmwood, Smith County, Tennessee.
RANGE.?Highland Rim in Smith, De Kalb, Cannon,
Humphreys, and Stewart counties, Tennessee.
HABITAT.?Swift, pebbly reaches of small streams.
Cambarus (Jugicambarus) gentryi Hobbs
FIGURE 81
Cambarus (Jugicambarus) gentryi Hobbs, 1970a: 163, fig. 1; 1972b: 125, fig.
109b; 1974b:18,fig. 63.
Cambarus gentryi.?Hobbs and Bouchard, 1973:54.
TYPES.?Holotype, allotype, and morphotype, USNM
130283, 130284,130285 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Boggy area below seepage along small
tributary to Turnbull Creek (Harpeth-Cumberland drainage
system), 1 mile (1.6 km) west of Kingston Springs, Cheatham
County, Tennessee.
RANGE.?Portions of the Cumberland (Stones River, Har-
peth River, and Yellow Creek) and Duck river basins in the
Nashville Basin and Western Highland Rim of Tennessee.
HABITAT.?Burrows (primary burrower).
Cambarus (Jugicambarus) monongalensis Ortmann
FIGURE 79
Cambarus monongalensis Ortmann, 1905a:395.
Cambarus (Bartonius) monongalensis.?Ortmann, 1905c: 120;
1906b:398,pL B: fig. 4,pL 39: figs. 4,10, pL 40: fig. 5,pL 41: figs. 2-4.
Cambarus (Cambarus) monongalensis.?Fowler, 1912:341 [by implication].
Cambarus carolinus monongalensis.?Faxon 1914:3%.
Cambarus carolinus monongalenses.?Hand, 1954:437 [erroneous spelling].
Cambarus mongalensis.?Meredith and Schwartz, 1959:1 [erroneous spelling].
Cambarus (Jugicambarus) monongalensis.?Hobbs, 1969b: 109, figs. 9, 19g;
1974b: 18, fig. 61.
TYPES.?Syntypes, CM 74.316, USNM 30613 (male I, 2
female), MCZ 6953 (male I, female), ZIAS 1/44968 (4
specimens), ANSP 1601 (male 1,3 females).
TYPE LOCALITY.?Edgewood Park, Allegheny County,
Pennsylvania.
NUMBER 480 23
RANGE.?Southwestern Pennsylvania and adjacent northern
West Virginia southward to Pocahontas County, West Virginia,
and Highland County, Virginia.
HABITAT.?Burrows (primary burrower).
Cambarus (Jugicambarus) nodosus Bouchard and Hobbs
FIGURE 76
Cambarus (Jugicambarus) nodosus Bouchard and Hobbs, 1976:8, fig. 3.
TYPES.?Holotype, allotype, and morphotype, USNM
146756, 146757, 146758 (male I, female, male II); paratypes,
USNM.BMNH.
TYPE LOCALITY.?Small unnamed tributary of North Potato
Creek (Hiwassee River system via Ocoee River) between 0.7
and 0.9 miles (1.1 and 1.4 km) west of Tennessee-North
Carolina border on U.S. Highway 64, Polk County, Tennessee.
RANGE.?Headwater areas of the Hiwassee (Ocoee, Nottely,
and Toccoa river basins), Savannah (Chattooga, Tugaloo, and
Koewee river basins), and Chattahoochee river systems in
Tennessee, North Carolina, South Carolina, and Georgia.
HABITAT.?Burrows and occasionally in streams.
Cambarus (Jugicambarus) obeyensis Hobbs and Shoup
FIGURE 67
Cambarus obeyensis Hobbs and Shoup, 1947:138, figs. 1-11.
Cambarus (Jugicambarus) obeyensis.?Hobbs, 1969b:107, figs. 9, 19h;
1974b: 19, fig. 52.?Bouchard, 1973a: 105.
Cambarus (Velicambarus) obeyensis.?Hobbs, 1970b:243.
TYPES.?Holotype, allotype, and "morphotype," USNM
82260 (male I, female, male II); paratypes, MCZ, USNM, TU.
TYPE LOCALITY.?Big Hurricane Creek on Monterey-
Clarkrange road, Putnam-Cumberland county line, Tennessee.
RANGE.?Headwaters of East Fork of Obey River in
Cumberland, Fentress, Overton, and Putnam counties, Tennes-
see.
HABITAT.?Rocky streams.
Cambarus (Jugicambarus) parvoculus Hobbs and Shoup
FIOURE70
Cambarus parvoculus Hobbs and Shoup, 1947:142, figs. 12-22.
Cambarus (Jugicambarus) parvoculus.?Hobbs, 1969b:107, figs. 9, 19i;
1974b:19,fig. 55; 1981:208,figs. 24g.70e,79,82,218.
TYPES.?Holotype, allotype, and "morphotype," USNM
82259 (male I, female, male II); paratypes, USNM, MCZ, TU.
TYPE LOCALITY.?Tributary to Big Hurricane Creek, Fen-
tress County (southwestern part), Tennessee.
RANGE.?Largely restricted to the Cumberland Plateau
section of the Appalachian Plateau Province, ranging through
the Cumberland River basin from Fentress, Overton, and
Putnam counties, Tennessee, and Bell County, Kentucky, to
headwaters of the Kentucky River in Letcher County, Ken-
tucky; and in the Tennessee Basin in Lee County, Virginia,
southward along the plateau to the northwestern part of Dade
County, Georgia.
HABITAT.?Rocky streams.
Cambarus (Jugicambarus) setosus Faxon
FIGURE 84
Cambarus setosus Faxon, 1889:237, pL 1: figs. 1-3,7, pi. 2: fig. 1.?Hobbs
and Barr, 1960:26, figs. 38-47.?Pfiieger,1987a:24; 1987b:8.
Cambarus ayersii Steele, 1902:18, pi. 5: fig. A, pi. 6: fig. 14. [Types: USNM
62316 (male II). Type locality: Fisher's Cave (= Sequiota Cave in Galloway),
near Springfield, Green County, Missouri (Sec. 9, T. 28N, R. 21W).]
Cambarus (Bartonius) setosus.?Ortmann, 1905c: 120.
Cambarus (Cambarus) setosus.?Fowler, 1912:341 [by implication].
Cambarus ayersi.?Ortmann, 1918:838.
Cambarus (Jugicambarus) setosus.?Hobbs, 1969b:107, fig. 19j; 1974b:19,
fig. 66.?Marquart, 1979:17.?Gardner, 1986:20.
TYPES.?Syntypes: MCZ 4200 (4 male II, 1 female),4201 (3
male 1,1 male 11,4 female, 4 juv. male, 2 juv. female), 4202 (2
male II, 3 female, 3 juv. male, 4 juv. female); USNM 25828
(male II, female).
TYPE LOCALITY.?Wilson's Cave (= Whisner Cave),2 miles
(3.2 km) northwest of Sarcoxie, near Springfield, Jasper
County, Missouri (Sec. 6,T. 27N,R. 29W).
RANGE.?Southwestern Missouri (Christian, Dade, Greene,
Jasper, Lawrence, Newton, and Stone counties) and ^north-
eastern Oklahoma (Delaware and Mayes counties).
HABITAT.?Subterranean streams.
Cambarus (Jugicambarus) tartarus Hobbs and Cooper
FIGURE 86
Cambarus (Jugicambarus) tartarus Hobbs and Cooper, 1972:51, figs.
1-12.?Hobbs, 1974b:19,fig. 68.
Cambarus tartarus.?Looney, 1975:13, fig. 2.
TYPES.?Holotype, allotype, and morphotype, USNM
131951,131411,132754 (male I, female, male II).
TYPE LOCALITY.?Stansberry-January Cave System, 4 miles
(6.4 km) north of Colcord, Delaware County, Oklahoma (Sec.
l l .T. 21N,R. 22E).
RANGE.?Known only from the type locality.
HABITAT.?Subterranean stream.
Cambarus (Jugicambarus) unestami Hobbs and Hall
FIGURE 69
Cambarus (Depressicambarus) unestami Hobbs and Hall, 1969:287, figs.
13-24.?Hobbs, 1974b: 19, fig. 54.?1981:210, figs. 24f,70f,81,83,219.
Cambarus unestami.?Bouchard, 1972b:92.?1976b:585.
Cambarus (Jugicambarus) unestami.?Bouchard, 1973a: 105.
TYPES.?Holotype, allotype, and morphotype, USNM
129863, 129864,129865 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Daniel Creek, 2.5 miles (4.0 km) west of
24 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
the Walker County line on State Route 143, Dade County,
Georgia.
RANGE.?Tributaries of Chattanooga, Cole City, Lookout,
and Long Island creeks (Tennessee River basin) in Walker and
Dade counties, Georgia, and Jackson County, Alabama. This
crayfish has also been collected from tributaries of the Little
River (Chattooga-Coosa Basin) in the northwestern part of
Chattooga County, Georgia. All localities are on Sand or
Lookout mountains.
HABITAT.?Rocky streams.
Cambarus (Jugicambarus) zophonastes Hobbs and
Bedinger
FIOURE85
Cambarus zophonastes Hobbs and Bedinger, 1964:11, figs. 1,2,4-11.?K.L
Smith, 1984:1.
Cambarus (Jugicambarus) zophonastes.?Hobbs, 1969b:107, fig. 19k;
1974b:20, fig. 67.?Hobbs, Hobbs, and Daniel, 1977:88, fig. 43.
Cabmarus (Jugicambarus) zophonastes.?Thome, 1987:421 [erroneous spell-
ing].
TYPES.?Holotype and allotype, USNM 108356, 108357
(male I, female); paratypes, USNM.
TYPE LOCALITY.?Hell Creek Cave, Stone County, Arkan-
sas (NE V4.NE 74, Sec. 30, T. 15N.R. 10W).
RANGE.?Known only from the type locality.
HABITAT.?Subterranean stream.
Subgenus Lacunicambarus Hobbs
Astacus.?Rafinesque, 1817:42.
Cambarus.?Girard, 1852:88 [in part].
Subgenus Bartonius Ortmann, 1905c:97 [in part].
Subgenus Cambarus.?Fowler, 1912:341 [in part].
Subgenus Lacunicambarus Hobbs, 1969b: 110. [Type species, by original
designation, Cambarus diogenes Girard, 1852:88. Gender masculine.]
Cambarus (Lacunicambarus) acanthura Hobbs
FIGURE 87
Cambarus (Lacunicambarus) acanthura Hobbs, 1981:215, figs. 25g, 84a,
85-87,220.
Cambarus acanthura.?Bouchard, 1982:90.
TYPES.?Holotype, allotype, and morphotype, USNM
129758,146601, 146602 (male I, female, male II); paratypes,
BMNH.USNM.
TYPE LOCALITY.?Field on west side of Holly Creek (farm
of Homer Robinson) about one mile northeast of Chatsworth,
Murray County, Georgia.
RANGE.?Tennessee Basin in Alabama, southwestern Ten-
nessee, and northwestern Georgia; Coosa Basin from its
headwaters in Georgia to Chilton County, Alabama, and in
headwaters of the Black Warrior River in Blount and
Tuscaloosa counties, Alabama; Chattahoochee Basin from
Walton and De Kalb counties, Georgia, to Russell County,
Alabama; and in the upper Ocmulgee Basin in Georgia.
HABITAT.?Burrows and streams.
Cambarus (Lacunicambarus) diogenes Girard
FIGURE 88
Astacus fossor Rafinesque, 1817:42. [Name suppressed by International
Commission, Opinion 522. (See Hemming, 1958.)].
Cambarus diogenes Girard, 1852:88.?Huner, 1977:11.?Page, 1985b:434,
figs. 167-170.?Pflieger. 1987a:23; 1987b:7.
Cambarus nebrascensis Girard, 1852:91. [Location of types (if extant)
unknown. Type locality: Fort Pierre, Nebraska (now in Stanley County,
South Dakota).]
Cambarus Nebrascensis.?Hagen, 1870:8.
Cambarus Diogenes.?Hagen, 1870:6.
Cambarus fossor.?Hagen, 1870:8.
Cambarus obesus Hagen, 1870:81, pi. 1: figs. 39-42, pi. 3: fig. 163, pi. 9
[Syntypes: MCZ 195 (2 male L 1 female), 165 (female), 3361 (male I), 229
(male I), 3363 (male); MHNP (2 dry); St. Petersburg Mus. (= ZIAS),
(female) not extant Type locality: Lawn Ridge, Illinois.]
Cambarus Diogenenes.?Abbott, 1873:83 [erroneous spelling].
Cambarus Diogenes Diogenes.?Faxon, 1884:144 [by implication].
Cambarus diogenes diogenes.?Hay, 1899b:959 [by implication].?Marlow,
1960:229.
S[ambarus] diogenes.?Steele, 1902:11 [erroneous spelling].
Cambarus (Bartonius) diogenes.?Ortmann, 1905c: 120; 1906b:402,pl. A: fig.
3, pi. 39: fig. l l .pl . 40: figs. 6,7.
Bartonius diogenes.?Williamson, 1907:749 [Bartonius intended as a subgen-
eric name].
Cambarus (Cambarus) diogenes.?Fowler, 1912:341.
Cambarus (Lacunicambarus) diogenes diogenes.?Hobbs, 1969b: 110, figs. 2c,
11, I S , 14i,20a; 1974b:20,fig. 69.
Cambarus (Lacunacambarus) diogenes diogenes.?Bouchard, 1972b: 103
[erroneous spelling].
Cambarus (Lacunicambarus) diogenes Diogenes.?Hobbs III, Thorp, and
Anderson, 1976:5 [lapsus calami].
Cambarus Diogenes diogenes.?Phillips, 1981:17 [lapsus calami].
Cambarus (Depressicambarus) diogenes.?Hobbs, 1986:16 [lapsus calami].
TYPES.?Not known to be extant except for a questionable
paratype (female), ANSP.
TYPE LOCALITY.?Vicinity of Washington, D.C.
RANGE.?Very widespread east of the Rockies and south of
the Great Lakes, except peninsular Florida and the Alleghenies;
not reported northeast of New Jersey in the East and east of
western Pennsylvania in the Mississippi drainage system.
HABITAT.?Burrows (primary burrower); often found in
ponds or streams in the spring season.
REMARKS.?This is a species complex and needs consider-
able attention.
Cambarus (Lacunicambarus) ludovicianus Faxon, new
status
FIGURE 89
Cambarus Diogenes var. Ludoviciana Faxon, 1884:144.
Cambarus diogenes ludovicianus.?Hay, 1899b:959.?Marlow, 1960:229.
Cambarus diogenes, ludoviciana.?Steele, 1902:21.
Cambarus diogenes ludoviciana.?Penn, 1941:8.
Cambarus ludovicianus.?Roberts, 1944:363.
NUMBER 480 25
Cambarus (Lacunicambarus) diogenes ludovicianus.?Hobbs, 1969b: 110, fig.
20b; 1974b:20,fig. 70.
Cambarus I. ludovicianus.?Hart and Hart, 1974:107 [lapsus calami].
TYPES.?Syntypes, USNM 5504 (male I, male 11,4 female),
USNM 5625 (male II, female), MCZ 3617 (male I).
TYPE LOCALITY.?New Orleans, Orleans Parish, Louisiana.
RANGE.?Lower Mississippi drainage in Arkansas, Tennes-
see, Louisiana, Mississippi, and in eastern Texas (probably
more extensive).
HABITAT.?Burrows (primary burrower).
REMARKS.?With a range that overlaps that of C. (L.)
diogenes, recognition of it as a subspecies of that crayfish is
rejected, and specific rank is accorded it herein.
Cambarus (Lacunicambarus) miltus Fitzpatrick
FIGURE 90
Cambarus (Lacunicambarus) miltus Fitzpatrick, 1978a:749, figs. 1-13.
Cambarus miltus.?Bouchard, 1982:90.
TYPES.?Holotype, allotype, and morphotype, USNM
148556, 148557, 148558 (male I, male II, female); paratypes,
USNM.
TYPE LOCALITY.?"In burrows along d'Olide Creek under
old highway 98 bridge, 0.1 mi (161 m) S of the junction of U.S.
highways 90, 98, and 1-10 at Spanish Fort, Baldwin County,
Alabama" (Fitzpatrick, 1978a:752).
RANGE.?Known only from the type locality.
HABITAT.?Burrows (primary burrower).
Subgenus Puncticambarus Hobbs
Cambarus.?Girard, 1852:90 [in part].
Subgenus Barlonius Ortmann, 1905c:97 [in part].
Subgenus Cambarus Fowler, 1912:341 [in part].
Subgenus Puncticambarus Hobbs, 1969b:101. [Type species, by original
designation, Cambarus extraneus Hagen, 1870:73. Gender: masculine.]
Cambarus (Puncticambarus) acuminatus Faxon
FIGURE 99
Cambarus acuminatus Faxon, 1884:113; 1885a:67,pl. 3: fig. 5, pi. 8: figs. 6a,
6a'.?Hobbs and Walton, 1966:136.
Cambarus (Bartonius) acuminatus.?Ortmann, 1905c: 120.
Cambarus (Cambarus) acuminatus.?Fowler, 1912:341 [by implication].
Cambarus bartonii acuminatus.?Faxon, 1914:424.
Cambarus montanus acuminatus.?Ortmann, 1931:106.
Cambarus bartoni acuminatus.?Ortmann, 1931:108.
Cambarus (Cambarus) montanus acuminatus.?Ortmann, 1931:108.
Cambarus (Puncticambarus) acuminatus.?Hobbs, 1969b:101, figs. 7, 14a,
171; 1974b:20,fig. 74.
P[rocambarus] acuminatus.?Hart and Hart, 1974:28 [lapsus calami].
TYPES.?Syntypes, MCZ 3624 (female). "Cotypes" for-
merly said to have been at Butler University (Faxon, 1914:424)
are no longer there.
TYPE LOCALITY.?Saluda River, at Fair's Mills, west of
Greenville, Greenville County, South Carolina.
RANGE.?Patapsco Basin in Maryland southward to the
Saluda drainage system in South Carolina.
HABITAT.?Streams.
REMARKS.?This highly variable species is also in need of
attention. Even in the Santee Cooper Basin (of which the
Saluda River is a tributary), South Carolina, more than one
rather distinct "variant" is recognizable.
Cambarus (Puncticambarus) buntingi Bouchard
FIGURE 98
Cambarus (Puncticambarus) sp. D.? Hobbs, 1969b:102,fig. 7.
Cambarus (Puncticambarus) ms. sp.? Bouchard, 1972b: 103.
Cambarus (Puncticambarus) buntingi Bouchard, 1973b:407, fig. 1.
Cambarus buntingi.?Bouchard, 1976b:585.
TYPES.?Holotype, allotype, morphotype, USNM 133057,
133058, 133059 (male I, female, male II); paratypes, UTM,
USNM.
TYPE LOCALITY.?"Elk Creek, a tributary of Clear Fork
Creek (Cumberland River system) at Co. Rd. 2345, approxi-
mately 1 mile [1.6 km] S of Newcomb, Campbell Co.,
Tennessee" (Bouchard, 1973b:410).
RANGE.?Cumberland and Tennessee drainage systems in
Anderson, Campbell, Claiborne, and Scott counties, Tennessee
and in Bell County, Kentucky.
HABITAT.?Small to large streams.
Cambarus (Puncticambarus) chaugaensis Prins and Hobbs
FIGURE 101
Cambarus (Puncticambarus) sp. E.?Hobbs. 1969b:102,134,135.
Cambarus (Puncticambarus) chaugaensis Prins and Hobbs, 1972:413, fig.
1.?Hobbs, 1974b:21,fig. 75; 1981:245, figs. 21h, 87,89c, 94,222.
TYPES.?Holotype, allotype, and morphotype, USNM
131926, 131927,131928 (male I, female, male II); paratypes,
USNM.BMNH.
TYPE LOCALITY.?Chauga River at Cassidy Bridge (off
County Road 290), Oconee County, South Carolina.
RANGE.?Tributaries of the Savannah River in Oconee
County, South Carolina, and Rabun County, Georgia.
HABITAT.?Streams.
Cambarus (Puncticambarus) coosae Hobbs
FIGURE 91
Cambarus (Cambarus) extraneus.?Ortmann, 1931:99 [in part].
Cambarus (Puncticambarus) species B.?Hobbs, 1969b:102,fig. 7.
Cambarus sp. nov. A.?Anonymous, 1969:C-23; 1970:162.
Cambarus species A.?Hobbs and Hall, 1974:204.
Cambarus (Puncticambarus) sp.?Hart and Hart, 1974:58.
Cambarus nov. sp. A.?Wharton, 1978:220.
Cambarus (Puncticambarus) coosae Hobbs, 1981:230, ftgs. 21c, 89a, 90,91,
223.
Cambarus (Hiaticambarus) coosae.?Hobbs and Peters, 1982:312 [lapsus
calami].
26 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
TYPES.?Holotype, allotype, and morphotype, USNM
145603,145604,145605 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Spring Creek, 8 miles (12.8 km) east-
southeast of Rome, Floyd County, Georgia, on U.S. Highway
411.
RANGE.?The Coosa River basin from Polk and Bradley
counties, Tennessee, Murray, Pickens, and Paulding counties,
Georgia, to Bibb, Calhoun, Cherokee, Chilton, Clay, Coosa,
Shelby, and Talladcga counties, Alabama.
HABITAT.?Streams.
Cambarus (Puncticambarus) cumberlandensis Hobbs and
Bouchard
FIGURE 95
Cambarus (Puncticambarus) species A.?Hobbs, 1969b: 102, fig. 7.
Cambarus (Puncticambarus) sp.?Bouchard, 1972b:91.
Cambarus (Puncticambarus) sp. nov.?Bouchard, 1973b:411.
Cambarus (Puncticambarus) cumberlandensis Hobbs and Bouchard, 1973:41,
figs. 1,2.
Cambarus cumberlandensis.?Hobbs and Bouchard, 1973:41.
TYPES.?Holotype, allotype, and morphotype, USNM
132989,132990,132991 (male I, female, male II); p?ratypes,
USNM.
TYPE LocALrrY.-^oplar Cove Creek (a tributary to the East
Fork of the Obey River), 5.8 miles (9.3 km) west of Jamestown,
Fentress County, Tennessee, off State Route 52, just east of
Helena.
RANGE.?Cumberland River system from Jellico Creek
(Scott County, Tennessee) downstream to and including
Roaring River in Jackson County, Tennessee; and Green River
system in Adair and Metcalf counties, Kentucky. Introduced
into Caney Fork River system in DcKalb County, Tennessee.
HABITAT.?Streams.
Cambarus (Puncticambarus) extraneus Hagen
FIGURE 92
Cambarus extraneus Hagen, 1870:73, pi. 1: figs. 88,89, pL 3: fig. 156.
Cambarus extraneous.?Adams, 1901:849 [erroneous spelling].
Cambarus extranus.?Steele, 1902:7 [erroneous spelling].
Cambarus (Bartonius) extraneus.?Ortmarm, 1905c: 120.
Cambarus (Cambarus) extraneus.?Fowler, 1912:341 [by implication].
Cambarus extranius.?Unestam, 1969:203 [erroneous spelling].
Cambarus (Puncticambarus) extraneus.?Hobbs, 1969b: 101, figs, lb, 7,13b,
14b, 17m; 1974b:21,fig. 71; 1981:247, figs. 21f,89g,93,95.224.
TYPES.?Syntypes, MCZ 175 (3 male II, 1 female), USNM
4957 (female). Hagen (1870:74) indicated that among the 6
specimens available there were "Male Form I, and II. Fern."
The first form male is no longer among the types and was
missing from them when I examined the series in the early
1940s.
TYPE LOCALITY.?"Tennessee River, (near the borders of?)
Georgia" (Hagen, 1870:74). The emendation offered by
Ortmann (1931:97) must be rejected because Cambarus (P.)
extraneus does not occur in the Coosa (Etowah) Basin.
Ortmann confused C. (P.) coosae with Hagen's species.
RANGE.?Known only from the South Chickamauga Creek
basin (Tennessee River drainage) in Catoosa, Walker, and
Whitfield counties, Georgia, and in the same basin in Hamilton
County, Tennessee (Bouchard, 1972b:58).
HABITAT.?Streams.
Cambarus (Puncticambarus) georgiae Hobbs
FIGURE 93
Cambarus (Puncticambarus) sp.?Anonymous, 1973b:2.
Cambarus (Puncticambarus) georgiae Hobbs, 1981:251, figs. 21g, 87, 89d,96,
225.
TYPES.?Holotype, allotype, and morphotype, USNM
118944, 144954, 144955 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Little Tennessee River at U.S. Highway
441, south of Dillard, Rabun County, Georgia.
RANGE.?Known only from the Little Tennessee River in
Rabun County, Georgia, and Macon County, North Carolina.
HABITAT.?Streams.
Cambarus (Puncticambarus) hiwasseensis Hobbs
FIGURE 102
Cambarus (Puncticambarus) species C?Hobbs, 1969b: 102,135, fig. 7.
Cambarus hiwasseensis Anonymous, 1973a:66 [nomen nudum].
Cambarus (Puncticambarus) sp.?Bouchard and Hobbs, 1976:13 [in part].
Cambarus (Puncticambarus) species D.?Hobbs and Peters, 1977:6.
Cambarus hiawasseensis Wharton, 1978:220 [nomen nudum].
Cambarus (Puncticambarus) hiwasseensis Hobbs, 1981:256, figs. 21 e, 89f,97,
98,226, frontispiece.
Cambarus Hiwasseenis.?Hamey, 1983:B4 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
129366,129365,129364 (male I, female, male II); paratypes,
BMNH.USNM.
TYPE LOCALITY.?Tributary to Peachtree Creek, 0.8 mile
(1.3 km) north of Peachtree School on U.S. Alternate Highway
64, Cherokee County, North Carolina.
RANGE.?The upper Hiwassee River basin in Georgia and
North Carolina.
HABITAT.?Streams.
Cambarus (Puncticambarus) nerterius Hobbs
FIGURE 97
Cambarus nerterius Hobbs, 1964:189, figs. 1-10.
Cambarus (Puncticambarus) nerterius.?Hobbs, 1969b:101, figs. 1, 17n;
1974b:21,fig. 73.?flobbs, Hobbs, and Daniel, 1977:91, fig. 44.
TYPES.?Holotype, allotype, and morphotype, USNM
111295, 111296, 111297 (male I, female, male II); paratypes,
USNM.
NUMBER 480 27
TYPE LOCALITY?Matt's Black Cave, 2 miles (3.2 km)
south of Renick, Greenbrier County, West Virginia.
RANGE.?Greenbrier River drainage system in Greenbrier
and Pocahontas counties, West Virginia.
HABITAT.?Subterranean waters.
Cambarus (Puncticambarus) parrishi Hobbs
FIGURE 96
Cambarus (Puncticambarus) sp.?Bouchard and Hobbs, 1976:13 [in part].
Cambarus (Puncticambarus) parrishi Hobbs, 1981:263, figs. 2Id, 89e, 98,99,
227.
TYPES.?Holotype, allotype, and morphotype, USNM
144957, 144958, 144959 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Hiwassee River, 2 miles (3.2 km) north
of the junction of Georgia routes 17 and 66 on Route 17,
approximately 7 miles (11.2 km) southeast of Hiawassee,
Towns County, Georgia.
RANGE.?Headwaters of the Hiwassee River in Towns
County, Georgia, and Clay County, North Carolina.
HABITAT.?Streams.
Cambarus (Puncticambarus) reburrus Prins
FIGURE 103
Cambarus reburrus Prins, 1968:458, figs. 1-11.
Cambarus (Puncticambarus) reburrus.?Prins and Hobbs, 1972:412.?Hobbs
1974b:21,fig. 76.
TYPES.?Holotype, allotype, and morphotype, USNM
129501,129502, 129503 (male I, female, male II); paratypes,
USNM, MCZ, Rudolph Prins.
TYPE LOCALITY.?Small tributary to Horsepasture River
from Sapphire Lake off U.S. Highway 64,5.5 miles (8.8 km)
east of Cashiers, Jackson County, North Carolina.
RANGE.?Type locality and tributaries of the French Broad
River in Buncombe, Henderson, Jackson, Madison, and
Transylvania counties, North Carolina.
HABITAT.?Streams.
Cambarus (Puncticambarus) robustus Girard
FIGURE 104
Cambarus robustus Girard, 1852:90.?Hagen, 1870:80, pL 3: fig. 167.?
Crocker and Barr, 1968:118, figs. 30, 39, 48, 55, 65, 67, 81.?Page,
1985b:439-440,figs. 171-173.
Cambarus Bartonii robustus.?Faxon, 1885a:9.
Cambarus Bartonii var. robusta.?Faxon, 1885a:61.
Cambarus bartonii robustus.?Faxon 1890:622.
Cambarus bartonii robusta.?Shufeldt, 1897:55.
Cambarus (Bartonius) bartoni var. robustus.?Ortmann, 1905c: 122 [by
implication].
Cambarus bartoni robustus.?Ortmann, 19O5c:135.
Cambarus (Cambarus) bartonii robustus.?Fowler, 1912:341 [by implication].
Cambarus (Cambarus) bartoni robustus.?Ortmann, 1931:126.
Cambarus (Bartonius) robustus.?Creaser, 1931b:260.
Cambarus (Puncticambarus) robustus.?Hobbs, 1969b:101,figs. lc, 13a, 17o;
1974b:21,fig. 77.
Cambarus robustis.?Espinet and Golding, 1976:B18 [erroneous spelling].
TYPES.?Syntype, ANSP 328 (male II); others probably
destroyed in the Chicago fire, 1871 (Faxon, 1914:423).
TYPE LOCALITY.?Humber River, near Toronto, Canada.
RANGE.?Ontario and New York, to Illinois, and southward
to Tennessee, North Carolina, and Virginia. Its presence in
New England east of the Hudson River basin is believed to
have resulted from introductions (Douglas G. Smith, in litt.).
HABITAT.?Streams.
Cambarus (Puncticambarus) scotti Hobbs
FIGURE 100
Cambarus aspinosus.?Anonymous, 1970:211 [nomen nudurn].
Cambarus (Puncticambarus) sp.?Hart and Hart, 1974:58.
Cambarus (Puncticambarus) scotti Hobbs, 1981:238, figs. 21b, 89b, 92, 93,
228.
TYPES.?Holotype, allotype, and morphotype, USNM
146479, 146480, 146481 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Clarks Creek, 1 mile (1.6 km) north of
Holland, Chattooga County, Georgia, on State Route 100.
RANGE.?Chattooga River basin in Chattooga and Walker
counties, Georgia, and Calhoun, Cherokee, and St. Clair
counties, Alabama.
Cambarus (Puncticambarus) spicatus Hobbs
FIGURE 94
Cambarus spicatus Hobbs, 1956c: 116, figs. 1-11.
Cambarus (Puncticambarus) spicatus.?Hobbs, 1969b:102, figs. 7, 18a;
1974b:22.fig. 72.
TYPES.?Holotype, allotype, and morphotype, USNM
99323,99324,99325 (male I, female, male II); paratypes, TU,
USNM.
TYPE LOCALITY.?Little River, 10.3 miles (16.5 km) west of
Winnsboro, Fairfield County, South Carolina.
RANGE.?Little River in Fairfield and Richland counties,
South Carolina.
HABITAT.?Streams.
Cambarus (Puncticambarus) veteranus Faxon
FIGURE 105
Cambarus bartonii veteranus Faxon 1914:389, pi. 13: fig. 2.
Cambarus bartoni veteranus.?Ortmann, 1931:112.
Cambarus (Cambarus) montanus veteranus.?Ortmann, 1931:112 [in part].
Cambarus veteranus.?Hobbs, 1955b:330.
Cambarus (Puncticambarus) veteranus.?Hobbs, 1969b: 102, figs. 7, 18b;
1974b:22,fig. 78.
TYPES.?Holotype, USNM 44712 (male I); paratypes,
28 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
USNM, MCZ.
TYPE LOCALITY.?Indian Creek, Baileysville, Wyoming
County, West Virginia.
RANGE.?Guyandot and Big Sandy rivers drainage in
Virginia, West Virginia, and Kentucky. Limits of range very
indefinite and poorly known.
HABITAT.?Streams.
Subgenus Veticambarus Hobbs
Subgenus Veticambarus Hobbs, 1969b:96. [Type species, by original designa-
tion, Cambarus pristinus Hobbs, 1965:268. Gender: masculine.]
Cambarus (Veticambarus) pristinus Hobbs
FIGURE 106
Cambarus pristinus Hobbs, 1965:268, figs. 1-12.
Cambarus (Veticambarus) pristinus.?Hobbs, 1969b:98, figs, le, 5, 131,14k,
17a;1974b:22.fig. 79.
TYPES.?Holotype, allotype, and morphotype, USNM
115528, 115529, 115530 (male I, female, male II); paratypes,
MCZ, USNM.
TYPE LOCALITY.?White Oak Creek, a tributary to the
Caney Fork of the Cumberland River, 3.9 miles (6.2 \m) east
of the White-Cumberland county line and 0.1 mile (160 m)
south of U.S. Highway 70S, Cumberland County, Tennessee.
RANGE.?Upper Caney Fork drainage in Cumberland and
Bledsoe counties, Tennessee.
HABITAT.?Streams, under rocks.
Genus Distocambarus Hobbs
Procambarus.?Hobbs, 1981:12,301 [in part].
Subgenus Distocambarus Hobbs, 1981:301. [Type species, by original
designation, Procambarus (Distocambarus) devexus Hobbs, 1981:302.
Gender masculine.]
Distocambarus.?Hobbs and Carlson, 1983:420.?Hobbs, 1984:18, fig. 7.
Subgenus Fitzcambarus Hobbs, 1983:430. [Type species, by original designa-
tion, Distocambarus (Fitzcambarus) carlsoni Hobbs, 1983:430. Gender
masculine.]
Subgenus Distocambarus Hobbs
Procambarus.?Hobbs, 1981:301.
Subgenus Distocambarus Hobbs, 1981:301; 1983:429. [Type species, by
original designation, Procambarus (Distocambarus) devexus Hobbs,
1981:302. Gender masculine.]
Distocambarus.?Hobbs and Carlson, 1983:420.
Distocambarus (Distocambarus) crockeri Hobbs and
Carlson
FIGURE 107
Distocambarus crockeri Hobbs and Carlson, 1983:421. fig. 1.
Distocambarus (Distocambarus) crockeri.?Hobbs, 1983:430.
TYPES.?Holotype, allotype, and morphotype, USNM
178582, 178583, 178584 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Roadside ditch 0.7 mile (1.1 km) south of
Parksville, McCormick County, South Carolina, on U.S.
Highway 221.
RANGE.?Savannah and Saluda river basins in Abbeville,
Edgefield, Greenwood, McCormick, and Saluda counties,
South Carolina.
HABITAT.?Burrows and occasionally flooded roadside
ditches and pools (primary burrower).
Distocambarus (Distocambarus) devexus (Hobbs)
FIGURE 108
Procambarus (Distocambarus) devexus Hobbs, 1981:302, figs, l l j , 105,
115-117,234.
Distocambarus devexus.?Hobbs and Carlson, 1983:420.
Distocambarus (Distocambarus) devexus.?Hobbs, 1983:430.
TYPES.?Holotype, allotype, and morphotype, USNM
148569, 148570, 148571 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Marshy area under bridge within 200 m
south of and in the flood plain of the Broad River on State
Route 17, Wilkes County, Georgia.
RANGE.?Savannah River Basin in Elbert, Oglethorpe, and
Wilkes counties, Georgia.
HABITAT.?Burrows, juveniles occasionally in flooded
roadside ditches (primary burrower).
Subgenus Fitzcambarus Hobbs
Procambarus.?Hobbs, 1983:430.
Subgenus Fitzcambarus Hobbs, 1983:430. [Type species, by original designa-
tion, Distocambarus (Fitzcambarus) carlsoni Hobbs, 1983:430. Gender:
masculine.]
Distocambarus (Fitzcambarus) carlsoni Hobbs
FIGURE 109
Distocambarus (Fitzcambarus) carlsoni Hobbs, 1983:430, fig. 1.
TYPES.?Holotype, allotype, and morphotype, USNM
178599, 178600, 178601 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Swampy area bordering unnamed tribu-
tary of Saluda River about 1 mile (1.6 km) north of State Route
81 on Route 106, Anderson County, South Carolina, about 4.5
airmiles (7.2 km) east-northeast of Belton.
RANGE.?Saluda and Savannah river basins from Greenville
to Abbeville and Saluda counties, South Carolina.
HABITAT.?Burrows (primary burrower).
Distocambarus (Fitzcambarus) youngineri Hobbs and
Carlson
FIGURE 110
Distocambarus (Fitzcambarus) youngineri Hobbs and Carlson, 1985:82, fig. 1.
NUMBER 480 29
TYPES.?Holotype, allotype, and morphotype, USNM
208413, 208414, 208415 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?About 50 m southwest of State Route 58
on Route 22 (11 airmiles, 17.6 km) west of Newberry,
Newberry County, South Carolina (34?18'N,81O48'W).
RANGE.?Known from the type locality and another nine
airmiles (14.4 km) away on State Route 121, 0.15 mile (0.24
km) northeast of Route 34.
HABITAT.?Burrows (primary burrower).
Genus Fallicambarus Hobbs
Astacus.?Cottle, 1863:217.
Cambarus.?Faxon, 1884:115 [in part].
Subgenus Bartonius Ortmann, 1905c:120 [in part].
Bartonius.?Williamson, 1907:749 [intended as subgeneric name].
Subgenus Cambarus.?Fowler, 1912:341 [in part].
Fallicambarus Hobbs, 1969b:111. [Type species, by original designation,
Cambarus strawni Reimer, 1966:11. Gender: masculine.]
Subgenus Creaserinus Hobbs, 1973a:463. [Type species, by original designa-
tion, Astacus fodiens Cottle, 1863:217. Gender: masculine.]
Subgenus Creaserinus Hobbs
Astacus.?Cottle, 1863:217.
Cambarus.?Faxon, 1884:115.
Subgenus Bartonius Ortmann, 1905c:120 [in part].
Bartonius.?Williamson, 1907:749 [intended as subgeneric name].
Subgenus Cambarus.? Fowler, 1912:341 [in part].
Fallicambarus.?Hobbs, 1969b:l 11 [in part].
Subgenus Creaserinus Hobbs, 1973a:463. [Type species, by original designa-
tion, Astacus fodiens Cottle, 1863:217. Gender: masculine.]
Pfrocambarus].?Hart and Hart, 1974:88 [lapsus; in part].
Subgenus Creserineus.?lezerinac, 1983:5 [erroneous spelling].
Fallicambarus (Creaserinus) byersi (Hobbs)
FIGURE 123
Cambarus byersi Hobbs, 1941b:118, figs. 6, 10, 14, 17, 20, 23, 26, 27, 30;
1942b: 167, pi. 12: figs. 206-210.
Procambarus byersi.?Hobbs, 1942b: 14 [lapsus calami].
Fallicambarus byersi.?Hobbs, 1969b:l 11, fig. 20c; 1972b: 101,figs. 81e, 83e;
1974b:22,fig. 87.
Fallicambarus (Creaserinus) byersi.?Hobbs, 1973a:463.
TYPES.?Holotype, allotype, USNM 79342 (male I, female),
and "morphotype," USNM 87969 (male II); paratypes, USNM,
MCZ.FSBC.
TYPE LOCALITY.?Burrows along Phifer Creek, 5.5 miles
(8.8 km) northwest of Pensacola, near U.S. Highway 90,
Escambia County, Florida.
RANGE.?Okaloosa County, Florida, west to Bay St. Louis,
Hancock County, Mississippi.
HABITAT.?Burrows (primary burrower).
Fallicambarus caesius.?Hobbs, 1975:28.
TYPES.?Holotype, allotype, and morphotype, USNM
144921, 144922, 144923 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Roadside ditch at Hot Spring-Saline
county line, Arkansas, on State Route 67.
RANGE.?Basins of Ouachita River and Dorcheat Bayou in
southern Arkansas.
HABITAT.?Burrows.
Fallicambarus (Creaserinus) danielae Hobbs
FIGURE 121
Cambarus lobdelli Lyle, 1938:76 [nomen nudum].
Fallicambarus (Creaserinus) danielae Hobbs, 1975:28, fig. 8.
TYPES.?Holotype, allotype, USNM 145997, 145998 (male
I, female); paratypes, USNM.
TYPE LOCALITY.?Burrows in a roadside ditch 9.0 miles
(14.4 km) east of Ocean Springs, Jackson County, Mississippi,
on U.S. Highway 90.
RANGE.?Known only from Jackson County, Mississippi,
and Mobile County, Alabama.
HABITAT.?Burrows.
Fallicambarus (Creaserinus) fodiens (Cottle)
FIGURE 116
Astacus fodiens Cottle, 1863:217.
Cambarus argillicola Faxon, 1884:115. [Types: MCZ 3459 (male 1,4 female,
3 juvenile). Type locality: Detroit, Michigan.]
Cambarus (Bartonius) argillicola.?Ortmann, 1905c: 120.
Bartonius argillicola.?Williamson, 1907:749 [Bartonius intended as subgen-
eric name].
Cambarus (Cambarus) fodiens.?Fowler, 1912:341 [by implication].
Cambarus fodiens.?Huntsman, 1915:158, figs. 8f,9d,10e, l l a , 12c?Crocker
and Barr, 1968:129-135,figs. 28,37,46,57,62, 85.
Cambarus (Bartonius) fodiens.?Creaser, 1931b:260,fig. 37.
Fallicambarus fodiens.?Hobbs, 1969b: 111, fig. 20e; 1972b: 102, figs. 83c,
84b, 85b; 1974b:23, fig. 82.?Page, 1985b:422,figs. 155-158.
Fallicambarus (Creaserinus) fodiens.?Hobbs, 1973a:463.
P[rocambarus] Fallicambarus fodiens.?Hart and Hart, 1974:88 flaPsus
calami].
Fallicambarus (Creaserineus) fodiens.?Jezerinac, 1983:5 [erroneous spell-
ing].
TYPES.?If extant, location unknown.
TYPE LOCALITY.?"Upper Canada," probably Ontario.
RANGE.?Southern Ontario, Michigan, Ohio, Indiana, and
Illinois southward to Arkansas, Mississippi, and Alabama.
HABITAT.?Lentic and lotic habitats and burrows.
REMARKS.?The range of this crayfish and that of the closely
allied, if not conspecific, F. (C.) hedgpethi abut or overlap in
Arkansas, Mississippi, and Tennessee.
Fallicambarus (Creaserinus) caesius Hobbs
FIGURE 118
Fallicambarus (Creaserinus) caesius Hobbs, 1975:24, fig. 7.
Fallicambarus (Creaserinus) hedgpethi (Hobbs)
FIGURE 117
Cambarus hedgpethi Hobbs, 1948d:224, fig. 17a-f,h-j,L?Perm, 1959:14,
30 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
figs. 9,27.46.64.86?Reimer. 1969:53.figs. 2,39.
Cambarus hedgepelhi.?Wallsand Black. 1967:60 [erroneous spelling].
Fallicambarus hedgpethi?Hobbt, 1969b:111.fig. 20f; 1972b: 102.figs. 82c,
83d;1974b:23.fig.83.
Fallicambarus (Creasermus) hedgpethi.?Hotbt, 1973* 463
Fallicambarus hedgepelhi.?Huner, 1977:11 [erroneous spelling].
TYPES.?Holotype, morphotype, USNM 85146 (male I.
male II). and allotype, USNM 85147 (female); paratypes.
USNM.
TYPE LOCALITY.?Lower middle part of the Aransas
National Wildlife Refuge, Aransas County, Texas.
RANGE.?Texas and Oklahoma eastward to western Tonnes
see, (?)panhandle of Florida, and (?)southwestern Georgia.
HABITAT.?Temporary bodies of water and burrows.
REMARKS.?See "Remarks" under F. (C.)fodiens.
Fallicambarus (Creaserinus) hortoni Hobbs and
Fitzpatrick
I u;rHI 120
Fallicambarus hortoni Hobbs and Fitzpatrick. 1970:829. Tigs. 1 -12?Hobbs .
1972b: 100. fig. 8Id; I974b:23. fig. 85.
Fallicambarus (Creaserinus) Hortoni?Wdbbt, I973a:463.
TYPES.?Holotype, allotype, and morphotype, USNM
129895. 129897, 1298% (male I. female, male II); paratypes.
USNM.
TYPE LOCALITY.?Low area along roadside ditch leading to
tributary of Cypress Creek, 7.5 miles (12 km) east of the
Hardeman County line on State Route 57 (Hatchie River
drainage), McNairy County, Tennessee.
RANGE.?Known from only two localities in McNairy
County, Tennessee.
HABITAT.?Burrows (primary burrower).
Fallicambarus (Creaserinus) oryktes (Penn and
Marlow)
FIGURE 122
Cambarus oryktes Perm and Marlow, 1959:197. figs. 1-14.
Fallicambarus orykies.?Hobbs. 1969b: 111. fig. 20h; 1972b:101. figs. 81f,
83a; 1974b:24. fig. 86.
Fallicambarus (Creaserinus) oryktes.?Hobbs. 1973a:463.
TYPES.?Holotype, allotype, and morphotype, USNM
101735,101736,101737 (male I, female, male II); paratypes,
TU.USNM.
TYPE LOCALITY.?Roadside ditch at Alton, St. Tammany
Parish, Louisiana.
RANGE.?St. Tammany Parish, Louisiana, and Forest,
Hancock, Harrison, Jackson, and Pearl River counties in
southern Mississippi eastward to Baldwin County, Alabama.
HABITAT.?Burrows (primary burrower).
Fallicambarus (Creaserinus) uhleri (Faxon)
Puuu 119
Cambarus UkUri Faxon. 1884 116; 1885a 77. pi 8: fig. 8. t \ 8a. la'.
Cambarus uhleri?Hiy, 1899b:959?Meredith and Schwartz. 1960:28.rigs.
11.16.
Cambarus (Bariomius) uhleri?Ortmarm. 1905c: 120.
Cambarus (Cambarus) uhlen.?Fcmla. 1912:341 [by implication).
Fallicambarusuhleri.?Hobbs. 1969b:111.fig, 2Oj; 1972b: 102.figs. 82a. 13b.
84a. 85a; 1974b-.24.Ag. 84.
Fallicambarus (Creaserinus) uhleri?Hobbs. 1973a 463.
TYPES.?Holotype, MCZ 3634 (male I); paratypes. MCZ.
TYPE LOCALITY.?"Swamp on Eastern Road near Fclsbury,
Somerset County. Maryland" (restricted by Faxon. 1914:426).
RANGE.?Atlantic coastal plain, from Maryland to South
Carolina.
HABITAT.?Louc and lcntic habitats and burrows.
Subgrnus Fallicambarus Hobbs
Cambarut.? Perm. 1933:73.
FaUicambanu He***. 1969b:III |.n pwi|
Subgenus Fallicambarus.?iUMm. 1973*462. nVp* * ? ? ? * . by anginal
designation. Cambarus ttrawnt Rennet, 1966.11. Gander masailine.)
Fallicambarus (Fallicambarus) dissltus (Penn)
PIOUBIIIS
Cambarus Jusitus Penn. I955:73.fig?. 1-13.
Fallicambarus AMI/M?Hobbs. 1969b 111. fig 20d. 1972b"99. figs l i e .
82d; 1974b:23.fig 88.
Fallxambarus (FalUambarus) dusuut ?Hobbt I973a:463
TYPES.?Holotype, allotype, and morphotype, USNM
98125. 98126. 98127 (male I. female, male II); paratypes.
ANSP, AMNH. CM. TU. USNM.
TYPE LOCALITY.?Three miles (4.8 km) east of Choudrant,
Lincoln Parish, Louisiana.
RANGE.?Red and Ouachita watersheds in Caldweli, Grant,
and Lincoln parishes, Louisiana, and Columbia and Union
counties, Arkansas.
HABITAT.?Burrows (primary burrower).
Fallicambarus (Fallicambarus) harpi Hobbs and
Robison
PK1UBB112
Fallicambarus (Fallicambarus) harpi Hobbs and Robison. 1985:1035. fig. 1.
TYPES.?Holotype, allotype, and morphotype, USNM
217946, 217947, 217948 (male I. female, male II); paratypes.
MHNP.RNHL.USNM.
TYPE LOCALITY.?Seepage area 0.2 mile (0.32 km) east of
Glenwood, Pike County, Arkansas (Sec. l.T. 5S.R. 24W).
RANGE.?Known from only two localities, both in the
Ouachita River basin in Pike County, Arkansas.
HABITAT.?Burrows (primary burrower).
NUMBER 480 31
Falticambarus (Fallicambarus) jeanae Hobbs
FIGURE 113
FaUicambana (FaUicambana) jeanae Hobbs. 1973*463. fig. 1.
Fallicambarus (FaUicambana) spectrum Hobbs. 1973a:469, fig. 2. [Typei:
holotype and allotype. USNM 144674.14467S (male I. female); pantypes.
USNM TVpe locality: two miles (3.2 km) east of Daisy, Pike County.
Arkansas, on VS. Hwy. 70.]
TYPES.?Holotype and allotype. USNM 144672, 144673
(male I, female); paratypes, USNM.
TYPE LOCALITY.?Seepage area 1.8 miles (2.9 km) east of
the Clark County line, Hot Spring County, Arkansas, on St.
Rte. 84.
RANGE.?Ouachita River basin in Clark, Hot Spring,
Montgomery, and Pike counties, Arkansas.
HABITAT.?Burrows (primary burrower).
REMARKS.?Fallicambarus (F.) spectrum appears to repre-
sent only a distinctive color morph of F. (F.) jeanae and is
therefore placed herein in the synonomy of that species.
Fallicambarus (FaUlcambarus) macneesei (Black)
FWURBlll
Cambarus macneesei Black. 1967:173. fig*. 1-12.
Fallicambarus macneesei.?Hobbs. 1969b:III, fig. 20g; 1972b:99. figs. 4c.
8L; 1974b:23.fi?. 80.
FaUicambana (FaUicambana) macneesei.?Hot**. 1973a:463.
TYPES.?Holotype, allotype, and morphotype, USNM
117681,117683,117682 (male I. female, male II); paratypes,
USNM, TU. MCZ, Joe B. Black.
TYPE LOCALITY.?Roadside ditches along East McNeese
Road, 1.8 miles (2.9 km) west of intersection with State Route
14, Lake Charles, Calcasieu Parish, Louisiana.
RANGE.?Known only from Calcasieu, and Lafayette par-
ishes, Louisiana, and Brazoria County. Texas.
HABITAT.?Burrows (primary burrower).
Fallicambarus (Falticambarus) strawni (Reimer)
FkHJKBlM
Cambana strawmReimer. 1966:11. figs. 9-18.
FaUicambana strawnL?ttdbU, 1969b:l 11. figs. 2f. 13j. 20i; 1972b.-99. figs.
4a. 81b; 1974b:24,fig. 81.
FaUicambana (FaUicambana) Jtrmwu.?Hobbs, 1973a:463.
TYPES.?Holotype, allotype, and morphotype, USNM
116675,116676,116677 (male I, female, male II); paratypes,
Rollin D. Reimer.
TYPE LOCALITY.?Small marshy area in the Saline River
drainage, 2.7 miles (4.3 km) north of Dierks, Howard County,
Arkansas.
RANGE.?Saline River basin in Sevier, Howard, and Pike
counties, Arkansas.
HABITAT.?Burrows, bom simple and complex (primary
burrower).
Genus FaxoneUa Creaser
Cambana?Hay, 1899a:122 [in part].
Subgenus FaxoneUa Creaser, 1933b:21. [Type species, by monotypy,
Cambana clypeatus Hay, 1899a:122. Gender feminine.]
Faxonius.?Creaser. 1933b:19 [inpart).
Orconectes.?Hobbs. 1942a:352 [in part].
Procambana.?Hott, 1944:349 [lapsus].
FaxoneUa.?Fitzpatrick. 1963:61.
FaxoneUa beyeri (Penn)
FIGURE 127
Orconectes (FaxoneUa) beyeri Penn. 19S0a:166, figs. 1-9.
FaxoneUa beyeri?Creaser. 1962:3 [by implication].?Fitzpatrick, 1963:61.?
Hobbs. 1972b:29.fig. 19b; 1974b:24.fig. 91.
TYPES.?Holotype, allotype, and morphotype, USNM
90361,90362.90363 (male I. female, male II); paratypes, TU.
USNM.
TYPE LOCALITY.?Roadside ditch on U.S. Highway 84, 2
miles (3.2 km) northeast of Naborton, De Soto Parish,
Louisiana.
RANGE.?Known only from De Soto and Natchitoches
parishes, Louisiana, and Angelina, Jasper, and San Augustine
counties, Texas.
HABITAT.?Lcntic habitats.
FaxoneUa blairi Hayes and Reimer
FIGURE 125
FaxoneUa blairi Hayes and Reimer, 1977:1, figs. 1-7.
TYPES.?Holotype and allotype, USNM 145705,145706.
TYPE LOCALITY.?Woodland swamp approximately 100
yards (35 m) west of U.S. Highway 259 on north side of Little
River, 7 miles (11.2 km) south of Broken Bow [not "Box"],
McCurtain County, Oklahoma.
RANGE.?Southeastern Oklahoma and southwestern Arkan-
sas in the Little and Red river basins. (See "Remarks" under
FaxoneUa clypeata.)
HABITAT.?Lentic situations.
FaxoneUa clypeata (Hay)
FIGURE 124
Cambana clypeatus Hay, 1899a: 122, fig. 2.
Faxonius clypeatus.?Creaser, 1933b: 19. pi. 1: figs. 7.8. pl.2: figs. 1.2.
Faxonius (FaxoneUa) clypeatus.?Creaser, 1933b:21 [by implication].
Orconectes clypeatus.?Hobbs, 1942a:352 [by implication].
Orconectes c/jjwola.?Hobbs. 1942b: 14.
Orconectes (FaxoneUa) clypeata.?Hobbs, 1942b: 154, pL 11: figs. 181-185.
Procambarus clypeata.?Hoff, 1944:349 [lapsus calami].
32 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Orconectes (Faxonella) clypeatus.?Penn, 1952a:746.
Faxonella clypeata,?Creaser, 1962:3 [by implication].?Fitzpatrick,
1963:61.?Hobbs, 1972b:29, fig. 19c; 1974b:24, fig. 89.?Huner,
1977:10.?Pflieger,1987a:29; 1987b:13.
Faxonella clypeta.?Unestam, 1969:203 [erroneous spelling].
Faxonella clyptea.?Spitzy, 1976:445 [erroneous spelling].
Faxonella cylpeata.?Hobbs III, Thorp, and Anderson, 1976:13 [erroneous
spelling].
Faxonella clypeatus.?Rao and Fingeiman, 1983:518.
TYPES.?Holotype.USNM 17277 (female).
TYPE LOCALITY.?Bay St. Louis, Hancock County, Missis-
sippi.
RANGE.?Le Flore County, Oklahoma(?), and Marion
County, Texas, east to Gadsden County, Florida, and Richland
County, South Carolina.
HABITAT.?Sluggish streams, lentic situations, and burrows
(tertiary burrower).
REMARKS.?The ranges of F. clypeata, F. creaseri, and F.
blairi have not been clearly defined. A study is needed to
ascertain whether or not current records for Missouri, Arkansas,
and Oklahoma are based on correct identifications.
Faxonella creaseri Walls
FIGURE 126
Faxonella creaseri Walls, 1968:413, figs. 1-9.?Hobbs, 1972b:29, figs. 5w,
19a;1974b:25,fig. 90.
TYPES.?Holotype, allotype, and morphotype, USNM
129215, 129216, 129214 (male I, female, male II); paratypes,
TU, MCZ, IBM, AMS, Joe B. Black, and Jerry G. Walls.
TYPE LOCALITY.?Roadside ditch 3.0 miles (4.8 km) north
of Grayson on U.S. Highway 165, Caldwell Parish, Louisiana.
RANGE.?Known with certainty from only several localities
in Caldwell Parish, Louisiana. Tentatively reported here from
Arkansas and Missouri.
HABITAT.?Temporary pools in roadside ditches.
REMARKS.?Specimens of the genus from eastern Arkansas
and southeastern Missouri may be assignable to F. creaseri or
perhaps this taxon represents only a local variant of F. clypeata.
(See "Range" and "Remarks" under F. clypeata) Studies of
populations in eastern Arkansas and southern Missouri are
badly needed.
Genus Hobbseus Fitzpatrick and Payne
Cambarus.?Hobbs, 1955t:95 [in part].
Hobbseus Fitzpatrick and Payne, 1968:15. [Type species, by original
designation, Cambarus cristatus Hobbs, 1955a:95. Gender: masculine.]
Hobbseus attenuates Black
FIGURE 132
Hobbseus attenuatus Black, 1969:193, figs. 1-12.?Hobbs, 1972b:75, fig.
58c;1974b:25,fig. 95.
TYPES.?Holotype, allotype, and morphotype, USNM
129535, 129536, 129537 (male I, female, male II); paratypes,
USNM, MCZ, TU, Joe B. Black.
TYPE LOCALITY.?"Roadside ditch adjacent to Noxapater
Creek, on gravel road two miles [3.2 km] north of State Route
395, six miles [9.6 km] west of Noxapater, Winston County,
Mississippi (R. 11E.T. 14N)."
RANGE.?Pearl River drainage in Winston and Neshoba
counties, Mississippi.
HABITAT.?Lentic situations.
Hobbseus cristatus (Hobbs)
FIGURE 129
Cambarus cristatus Hobbs, 1955a:95, figs. 1-11.
Hobbseus cristatus.?Fitzpatrick and Payne, 1968:15.?Hobbs, 1972b:75, figs.
5o,58b;1974b:25,fig. 93.
TYPES.?Holotype, allotype, and morphotype, USNM
96985, 96986, 96987 (male I, female, male II); paratypes,
MCZ, TU, USNM, BMNH.
TYPE LOCALITY.?Roadside ditch, 11.3 miles (18.1 km)
south of Macon on U.S. Highway 45, Noxubee County,
Mississippi.
RANGE.?Tombigbee River basin in Kempcr, Lauderdale,
Lowndes.and Noxubee counties, Mississippi.
HABITAT.?Lentic and lotic habitats and burrows (probably
secondary burrower).
Hobbseus orconectoides Fitzpatrick and Payne
FIGURE 128
Hobbseus orconectoides Fitzpatrick and Payne, 1968:17, figs. 1-12.?Hobbs,
1972b:75,fig. 58a; 1974b:25, fig. 92.
TYPES.?Holotype, allotype, and morphotype, USNM
129191, 129193, 129194 (male I, female, male II); paratypes,
MCZ, TU.OSM, USNM.
TYPE LOCALITY.?Woodland pond near Rock Hill commu-
nity, about 4.25 miles (6.8 km) north of Starkville, Oktibbeha
County, Mississippi (Sec. 1,T. 19N,R. 14E).
RANGE.?Known only from Lowndes, Oktibbeha, and
Webster counties, Mississippi.
HABITAT.?Lentic situations and burrows (probably secon-
dary burrower).
Hobbseus petilus Fitzpatrick
FIGURE 133
Hobbseus petilus Fitzpatrick, 1977:367, fig. 1.
TYPES.?Holotype, allotype, and morphotype, USNM
147673, 147674, 147675 (male I, female, male II); paratypes,
USNM, Joe B. Black.
TYPE LOCALITY.?Tributary of Patch Creek (Tombigbee
River drainage), 7.4 miles (11.8 km) E of U.S. Highway 45 in
Saltillo, on State Route 363, Lee County, Mississippi.
NUMBER 480 33
RANGE.?Itawamba, Lee, and Monroe counties, Mississippi.
HABITAT.?Lentic and sluggish lotic situations.
Hobbseus prominens (Hobbs)
FIGURE 130
Cambarus prominens Hobbs, 1966a:110,figs. 1-10.
Hobbseus prominens.?Fitzpatrick and Payne, 1968:15.?Hobbs, 1972b:76,
figs. 58e, 59b; 1974b:25, fig. 94.
TYPES.?Holotype, allotype, and morphotype, USNM
115603, 115604, 115605 (male I, female, male II); paratypes,
USNM, MCZ.
TYPE LOCALITY.?Roadside ditch, 3 miles (4.8 km) west of
Demopolis, on U.S. Highway 80, Marengo (not Sumter)
County, Alabama.
RANGE.?Mobile River drainage in Choctaw, Clarke, Dal-
las, Hale, Marengo, Perry, Sumter, and Webster counties,
Alabama. Also present in northern Mississippi.
HABITAT.?Lentic and lotic situations and burrows (prob-
ably secondary burrower).
Hobbseus valleculus (Fitzpatrick)
FIGURE 131
Cambarus valleculus Fitzpatrick, 1967b: 163, figs. 1-12.
Hobbseus valleculus.?Fitzpatrick and Payne, 1968:15.?Hobbs, 1972b:76,
figs. 58d, 59a; 1974b:25, fig. 96.
TYPES.?Holotype, allotype, and morphotype, USNM
117963,117964, 117965 (male I, female, male II); paratypes,
MCZ,TU.
TYPE LOCALITY.?Small creek, 6.8 miles (10.9 km) south of
junction of State Routes 15 and 12 on State Route 15, Choctaw
County, Mississippi.
RANGE.?Known only from a few localities in Attala,
Choctaw, and Kemper counties, Mississippi.
HABITAT.?Streams.
Genus Orconectes Cope
Astacus.?Rafinesque, 1817:42.
Subgenus Cambarus Erichson, 1846:95 [in part].
Cambarus.?Girard, 1852: 87 [in part].
Orconectes Cope, 1872:419. [Type species, by subsequent designation
(Fowler, 1912:339), Orconectes inermis Cope, 1872:419. Gender mascu-
line.].?Hobbs. 1942a:35Z
Camtarus.?Packard, 1888:156 [erroneous spelling].
Oreonectes.?Lonnberg, 1894:126 [erroneous spelling].
Camborus.?Williamson, 1899:47 [erroneous spelling].
Cambrus.?Price, 1900:155 [erroneous spelling].
Orconectis.? Harris, 1903:113 [erroneous spelling].
Subgenus Faxonius Ortmarm, 19O5c:97. [Type species, by original designation,
Astacus limosus Rafinesque, 1817:42. Gender masculine. Proposed as
subgenus of Cambarus.].?Fitzpatrick, 1987a:51.
G[ambarus].?Ortmann, 19O5c:112 [erroneous spelling].
Fraxonius.?Turner, 1926:43 [erroneous spelling].
Faxonius.?Creaser, 1933a: 1.
Orconetes.?Wolf, 1934-1938:104.?Avault, 1973:242. [Erroneous spelling.]
Faxonicus.?Fleming, 1938:302 [erroneous spelling].
Cambarrus.?Fleming, 1939:305 [erroneous spelling].
Cityphobius.?Joleaud, 1939:14 [error for C. typhlobius].
Fexonius.?Rioja, 1941:193 [erroneous spelling].
Orconecctes.?Williams, 1952:330 [erroneous spelling].
Orconectas.?Villalobos, 1953:351 [erroneous spelling].
Orconects.?Lewis, 1955:146 [erroneous spelling].
Gambarus.?Croizat, 1958:908 [erroneous spelling].
Orconnectes.?Threinen, 1958:1 [erroneous spelling].
Camburus.?Bacescu, 1967:218 [erroneous spelling].
Oronectes.?Bacescu, 1967:218 [erroneous spelling].
Orchonectes.?Dimond et al., 1968:760 [erroneous spelling].
Orconectus.?Ray and Stevens, 1970:58 [erroneous spelling].
Orconcectes.?Fielder, 1972:133 [erroneous spelling].
Orconeotes.?Unestam, 1973:4 [erroneous spelling].
Oroncestes.?Wielgus, 1973:119 [erroneous spelling].
Orcenectes.?Hart and Hart, 1974:45 [erroneous spelling].
Oroconectes.?Hobbs III, 1975:273 [erroneous spelling].
Orcnoectes.?O'Brien, 1976:84 [erroneous spelling].
Oreconectes.?Yassini, 1977:201 [erroneous spelling].
Onconectes.?Price and Payne, 1977:79 [erroneous spelling].
Procambarus.?Huner, 1978:4 [lapsus for Orconectes].
Orconelctes.?Brodsky, 1981:122 [erroneous spelling].
Orconcetes.?Huner and Naqvi, 1984:405 [erroneous spelling].
Subgenus Billecambarus Fitzpatrick, 1987a:47. [Type species, by original
designation, Cambarus Harrisonii Faxon, 1884:130. Gender masculine.]
Subgenus Buannulifictus Fitzpatrick, 1987a:49. [Type species, by original
designation, Cambarus Palmeri Faxon, 1884:124. Gender: masculine.]
Subgenus Crockerinus Fitzpatrick, 1987a:50. [Type species, by original
designation, Cambarus Sanbornii Faxon, 1884:128. Gender masculine.]
Subgenus Gremicambarus Fitzpatrick, 1987a:53. [Type species, by original
designation, Cambarus virilis Hagen, 1870:63. Gender: masculine.]
Subgenus Hespericambarus Fitzpatrick, 1987a:54. [Type species, by original
designation, Cambarus difficilis Faxon, 1898:656. Gender masculine.]
Subgenus Procericambarus Fitzpatrick, 1987a:57. [Type species, by original
designation, Cambarus forceps Faxon, 1884:133. Gender: masculine.]
Subgenus Rhoadesius Fitzpatrick, 1987a:58. [Type species, by original
designation, Cambarus sloanii Bundy, 1876:24. Gender masculine.]
Subgenus Tragulicambarus Fitzpatrick, 1987a:60. [Type species, by original
designation, Cambarus lancifer Hagen, 1870:59. Gender masculine.]
REMARKS.?Hobbs (1942a:350) erroneously indicated that
Orconectes inermis Cope was the type-species by monotypy.
Cope (1872:419) stated: "The present genus embraces two
species, 0. inermis of the Wyandotte and the 0. pellucidus of
the Mammoth Cave."
Subgenus Billecambarus Fitzpatrick
Cambarus.?Faxon, 1884:130 [in part].
Subgenus Faxonius Ortmann, 1905c: 111 [in part].
Faxonius.?Creaser, 1933a:3 [in part].
Orconectes.?Hobbs, 1942a:352 [in part].
Subgenus Billecambarus Fitzpatrick, 1987a:47. [Type species, by original
designation, Cambarus Harrisonii Faxon, 1884:130. Gender masculine.]
Orconectes (Billecambarus) harrisonii (Faxon)
FIGURE 149
Cambarus Harrisonii Faxon, 1884:130; 1885a:94,pl. 3: fig. l.pl. 9: figs. 9,9'.
Cambarus harrisonii?Hay, 1899b:960.
Cambarus harrisoni.?Ortmann, 1902:278.
34 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambarus (Faxonius) harrisoni.?Ortmann, 1905c: 111.
Faxonius harrisonii.?CKXKT, 1933a:3 [by implication]; 1934a: 1; 1962:2 [by
implication].
Faxonius (Faxonius) harrisonii.?Creaser, 1933b:21 [by implication].
Orconectes harrisonii.?Hobbs, 1942a:352 [by implication].
Orconectes (Orconectes) harrisonii.?Hobbs, 1942b: 154 [by implication].
Orconectes harrisoni.?Rhozdes, 1944a:117.?Pflieger,1987a:28; 1987:12.
Orconectes harrisonii.?Williams, 1954:839, figs. 35-41.?Fitzpatrick,
1963:61 [by implication].?Hobtx, 1974b:29,fig. 111.
Orconectes (Billecambarus) harrisonii.?Fitzpatrick, 1987a:49, fig. 1.
TYPES.?Syntypes, MCZ 3586 (11 male 1,5 female),USNM
25826 (male I, female).
TYPE LOCALITY.?Irondale, Washington County, Missouri.
RANGE.?St. Francis, St. Genevieve, and Washington
counties, Missouri.
HABITAT.?Streams.
Subgenus Buannulifictus Fitzpatrick
Cambarus.?Faxon, 1884:124 [in part].
Subgenus Faxonius Ortmann, 1905c: 103 [in part].
G[ambarus].?Ortmann, 1905c:l 12 [erroneous spelling].
Faxonius.?Creaser, 1933a:3 [in part].
Orconectes.?Hobbs, 1942a:352 [in part].
Subgenus Orconectes.?Hobbs, 1942b: 154 [in part].
Subgenus Buannulifictus Fitzpatrick, 1987a:49. [Type species, by original
designation, Cambarus Palmeri Faxon, 1884:124. Gender masculine.]
Orconectes (Buannulifictus) hobbsi Penn
FIGURE 197
Orconectes hobbsi Penn, 195Oc:381, figs. 1-10; 1959:18, figs. 32. 52, 68,
82.?Fitzpatrick, 1963:61 [by implication].?Hobbs, 1974b:29, fig. 160.
Orconectes (Orconectes) hobbsi.?Hobbs, 1959:895.
Faxonius hobbsi.?Creaser, 1962:2 [by implication].
Orconectes (Buannulifictus) hobbsi.?Fitzpatrick, 1987a:49.
TYPES.?Holotype, allotype, and morphotype, USNM
90952, 90953, 90954 (male I, female, male II); paratypes,
MCZ, ANSP, USNM, TU, AMNH.
TYPE LOCALITY.?Headwater creek of Bayou Lacombe at
SL Tammany, 6 miles (9.6 km) north of Lacombe on State
Route 187, St. Tammany Parish, Louisiana.
RANGE.?Lake Pontchartrain watershed in Louisiana and
Mississippi.
HABITAT.?Rapidly flowing streams.
Orconectes (Buannulifictus) meeki brevis Williams
FIGURE 193
Orconectes meeki brevis Williams, 1952:348, figs. 41-48; 1954:881,figs. 166,
175-182.?Fitzpatrick, 1963:61 [by implication].?Hobbs, 1974b:34, fig.
156.
Orconectes (Orconectes) meeki brevis.?Hobbs, 1959:895 [by implication].
Faxonius meeki brevis.?Creaser, 1962:2 [by implication].
Orconectes (Buannulifictus) meeki brevis.?Fitzpatrick, 1987a:50.
TYPES.?Holotype, allotype, and "morphotype," USNM
118848 (male I, female, male II); paratypes, USNM, MCZ.
TYPE LOCALITY.?Streams, approximately 5 miles (8 km)
north of Stilwell on U.S. Highway 59, Adair County,
Oklahoma.
RANGE.?Tributaries of the Red River in eastern Oklahoma
and extreme northwestern Arkansas.
HABITAT.?Rocky streams.
Orconectes (Buannulifictus) meeki meeki (Faxon)
FIGURE 192
Cambarus meeki Faxon, 1898:657, pi. 65: figs. 5-9.
Cambarus meekii.?Hay, 1899b:960 [erroneous spelling].
Cambarus (Faxonius) meeki.?Ortmann, 1905c:l 12.
G[ambarus] (Faxonius) meeki.?Ortmann, 1905c: 112 [erroneous spelling].
Faxonius meeki.?Creaser, 1933a:3 [by implication].
Faxonius (Faxonius) meeki.?Creaser, 1933b:21 [by implication].
Orconectes meeki.?Hobbs, 1942a:352 [by implication].?Pflieger, 1987a:28;
1987b:12.
Orconectes (Orconectes) meeki.?Hobbs, 1942b: 154 [by implication].
Orconectes meeki meeki.?Williams, 1952:350; 1954:878, figs. 166-174.?
Fitzpatrick, 1963:61 [by implication].?Hobbs, 1974b:34, fig. 155.
Orconectes (Orconectes) meeki meeki.?Hobbs, 1959:895 [by implication].
Faxonius meeki meeki.?Creaser, 1962:2 [by implication].
Orconectes (Buannulifictus) meeki meeki.?Fitzpatrick, 1987a:50.
TYPES.?Syntypes, MCZ 4363 (2 male II, 3 female),USNM
19680 (10 male II, 13 female), Mus. Zool. Torino 433.
TYPE LOCALITY.?Walnut Fork of Big Piney Creek, Swain,
Newton County, Arkansas.
RANGE.?Arkansas and White river systems north of the
35th parallel in Arkansas.
HABITAT.?S treams.
Orconectes (Buannulifictus) palmeri creolanus (Creaser)
FIGURE 1%
Faxonius (Faxonius) creolanus Creaser, 1933b: 16,pi. 1: figs. 1,2 [subgenus by
implication].
Cambarus creolanus.?Penn, 1941:8.
Faxonius creolanus.?Hobbs, 1942a:352.
Orconectes creolanus.?Hobbs, 1942a:352.
Orconectes (Orconectes) creolanus.?Hobbs, 1942b: 154 [by implication].
Orconectes (Orconectes) palmeri creolanus.?Penn, 1952a:744.
Orconectes palmeri creolanus.?Penn, 1953e:6; 1957:231, figs. 24, 27,
30.?Fitzpatrick, 1963:61 [by implication].?Hobbs, 1974b:37, fig. 159.
Faxonius palmeri creolanus.?Creaser, 1962:2 [by implication].
Orconectes palmeri croelanus.?Godley, McDiarmid, and Rojas, 1984:83
[erroneous spelling].
Orconectes (Buannulifictus) palmeri creolanus.?Fitzpatrick, 1987a:49.
TYPES.?Holotype and allotype, USNM 101972 and 98341
(male I, female); paratypes, MCZ, USNM.
TYPE LOCALITY.?Tributary to Amite River, 0.5 mile (0.8
km) north of Ethel, East Feliciana Parish, Louisiana.
RANGE.?Mississippi and Louisiana in "the Lake Pontchar-
train drainage area plus the Pearl and Pascagoula rivers" (Penn,
1957:258).
HABITAT.?Rapid and sluggish streams.
NUMBER 480 35
Orconectes (Buannulifictus) palmeri longimanus (Faxon)
FIGURE 195
Cambarus palmeri longimanus Faxon, 1898:655, pi. 64: figs. 1-6.
Cambarus longimanus.?Crcaser and Ortenburger, 1933:16,38, figs. 4,16,26.
Faxonius longimanus.?Creaser, 1933a:3 [by implication].
Faxonius (Faxonius) longimanus.?Crcaser, 1933b: 19 [by implication].
Cambarus longimonus.?Goodnight, 1940:41 [erroneous spelling].
Orconectes palmeri longimanus.?Hobbs, 1942a:352 [by implication];
1974b:37, fig. 158.?Williams, 1954:894, figs. 200, 209-216.?Penn,
1957:231, figs. 25,28.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) palmeri longimanus.?Hobbs, 1942b:154 [by impli-
cation].
Orconectes longimanus.?Hoff, 1943:282.
Faxonius palmeri longimanus.?Crcaser, 1962:2 [by implication].
Orconectes (Buannulifictus) palmeri longimanus.?Fitzpatrick, 1987a:50.
TYPES.?Holotype, MCZ 7390 (male I); paratypes, MCZ,
USNM, Mus. Zool. Torino.
TYPE LOCALITY.?Red River, Arthur City, Lamar County,
Texas.
RANGE.?Texas, Oklahoma, Kansas, Arkansas, and Louisi-
ana, "all western tributaries of the Mississippi River from the
Arkansas River to the Gulf, and Gulf drainage streams from the
Mississippi River to the Guadalupe River in Texas" (Penn,
1957:258).
HABITAT.?Streams.
Orconectes (Buannulifictus) palmeri palmeri (Faxon)
FIGURE 194
Cambarus Palmeri Faxon, 1884:124; 1885a:103, pL 3: fig. 6,pL 10: figs. 5a,
5a'.
Cambarus palmeri.?Underwood, 1886:371.
Cambarus palmeri palmeri.?Faxon, 1898:655 [by implication].
Cambarus (Faxonius) palmeri.?Ortmann, 1905c: 113.
Faxonius palmeri.?Creaser, 1933a:3 [by implication].
Faxonius (Faxonius) palmeri.?Creaser, 1933b: 19 [by implication].
Cambarus (Faxonius) creaseri Lyle, 1938:76 [nomen nudum, fide J.F.
Fitzpatrick, in lilt.].
Orconectes palmeri palmeri.?Hobbs, 1942a:352 [by implication]; 1974b:37,
fig. 157.?Hobbs and Marchard, 1943:28, figs. 7, 13, 19, 26.?Penn,
1957:231, figs. 26,29.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes)palmeri.?Hobbs, 1942b:l54 [by implication].
Faxonius palmeri palmeri.?Creaser, 1962:2 [by implication].
Orconectes palmeri.-?Huner, 1977:11.?Pflieger, 1987a:3O; 1987b:14.
Orconectes (Buannulifictus) palmeri palmeri.?Fitzpatrick, 1987a:49, fig. 2.
TYPES.?Syntypes, USNM 4872 (8 male 11,7 female), MCZ
3564 (2 male II, 2 female).
TYPE LOCALITY.?Creek at eastern side of Reelfoot Lake,
Obion County, Tennessee.
RANGE.?Lower Mississippi Valley, occurring in western
Kentucky, Tennessee, Louisiana, and Mississippi, and in
eastern Missouri and Arkansas.
HABITAT.?Streams.
Subgenus Crockerinus Fitzpatrick
Cambarus.?Ginrd, 1852:88 [in part].
Astacus.?Hagen, 1870:61 [lapsus].
Camborus.?Williamson, 1899:47 [erroneous spelling].
Subgenus Faxonius Ortmann, 1905c:l 12 [in part].
Faxonius.?Williamson, 1907:754 [intended as subgeneric name].?Creaser,
1933a:3 [in part].
Camarus.?Fleming, 1939:304 [erroneous spelling].
Cambarrus.?Fleming, 1939:305 [erroneous spelling].
Orconectes.?Hobbs, 1942a:352 [in part].
Subgenus Orconectes.?Hobbs, 1942b:154 [in part].
Orconecctes.?Williams, 1952:330 [erroneous spelling].
Orconnectes.?Thrcinen, 1958:1 [erroneous spelling].
Orconcectes.?Fielder, 1972:132 [erroneous spelling].
Subgenus Crockerinus Fitzpatrick, 1987a:50. [Type species, by orgininal
designation, Cambarus Sanbornii Faxon, 1884:128. Gender masculine.]
Orconectes (Crockerinus) bisectus Rhoades
FIGURE 164
Orconectes bisectus Rhoades, 1944a:129, fig. 7a-d?Fitzpatrick, 1963:61 [by
implication].?Prins and Fitzpatrick, 1965:141, figs. 1-12.?Hobbs,
1974b:27,fig. 125.
Orconectes (Orconectes) bisectus.?Hobbs, 1959:890 [by implication].
Faxonius bisectus.?Creaser, 1962:2 [by implication].
Orconectes (Crockerinus) bisectus.?Fitzpatrick, 1987a:51.
TYPES.?Holotype and allotype.USNM 81322,81323 (male
II, female); paratypes, MCZ, OSM.
TYPE LOCALITY.?Brushy Fork, 1 mile (1.6 km) west of
Repton, Crittenden County, Kentucky.
RANGE.?Ohio River drainage in eastern part of Crittenden
County, Kentucky.
HABITAT.?Medium-sized streams.
Orconectes (Crockerinus) erichsonianus (Faxon)
FIGURE 162
Cambarus erichsonianus Faxon, 1898:659,pl. 64: figs. 7-12.
Cambarus erichsoni.?Ortmann, 1905c:109 [erroneous spelling].
Cambarus (Faxonius) erichsonianus.?Ortmann, 19O5c:l 12.
Cambarus spinosus gulielmi Faxon, 1914:375. [Syntypes, USNM 26379 (6
male II, 6 female, 4 juvenile male), MCZ 7448 (male II, female). Type
locality, streams from John Ross Spring, near Rossville, Walker County,
Georgia.]
Cambarus gulielmi.?Ortmann, 1931:68.
Faxonius erichsonianus.?Creaser, 1933a:3 [by implication]; 1962:2 [by
implication].
Faxonius (Faxonius) erichsonianus.?Creaser, 1933b:7 [by implication].
Cambarus erichsonionus.?Fleming, 1938:299 [erroneous spelling].
Orconectes erichsonianus.?Hobbs, 1942a:352 [by implication]; 1974b:29,
fig. 123; 1981:284, figs. 19d, 108a,b, 109-111,231.?Fitzpatrick, 1963:61
[by implication]; 1967a: 147, figs. 2-18,20.
Orconectes (Orconectes) erichsonianus.?Hobbs, 1942b: 154 [by implication].
Orconectes (Crockerinus) erichsonianus.?Fitzpatrick, 1987a:51.
TYPES.?Syntypes, USNM 20787 (3 male 1,4 female), MCZ
4347 (male I, female, male II).
TYPE LOCALITY.?Rip Roaring Fork, 5 miles (8.0 km)
northwest of Greeneville, Greene County, Tennessee.
RANGE.?Tennessee, Elk, and Coosa river systems in
southwestern Virginia, eastern Tennessee, northwestern Geor-
gia, and northern Alabama.
HABITAT.?Streams.
36 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Orconectes (Crockerinus) eupunctus Williams
FIGURE 154
Orconecctes eupunctus Williams, 1952:330 [erroneous spelling].
Orconectes eupunctus Williams, 1952:334, pL 1: figs. 1-8; 1954:840, figs.
41-49.?Fitzpatrick, 1963:61 [by implication].?Hobbs, 1974b: 19, fig.
116.?Pflieger, 1987a:24; 1987b:8.
Orconectes (Orconectes) eupunctus.?Hobbs, 1959:894.
Faxonius eupunctus.?Creaser, 1962:2 [by implication].
Orconectes (Crockerinus) eupunctus.?Fitzpatrick, 1987a:51.
TYPES.?Holotype, allotype, and "morphotype" USNM
129200 (male I, female, male II); paratypes, USNM, MCZ.
TYPE LOCALITY.?Eleven Point River at Riverton, Oregon
County, Missouri.
RANGE.?Spring River and Eleven Point drainage systems
in Arkansas and Missouri.
HABITAT.?Clear, cold, rapid streams.
Orconectes (Crockerinus) iUinoiensis Brown
FIGURE 166
Orconectes illinoiensis Brown, 1956:163, figs. 1-9.?Hobbs, 1974b:30, fig.
128.?Page, 1985b:386, figs. 116-119, frontispiece.
Faxonius illinoiensis.? Creaser, 1962:2 [by implication].
Orconectes Utinoisensis.?Hobbs and Fitzpatrick, 1962:207.?Fitzpatrick,
1967a: 149, figs. 2-18.21. [Erroneous spelling.]
Orconectes (Crockerinus) illinoiensis.?Fitzpatrick, 1987a:51.
TYPES.?Holotype, allotype, and morphotype, USNM
97997,97998,97999 (male I, female, male II); paratypes, Max
R. Matteson, Troy C. Dorris, USNM.
TYPE LOCALITY.?Cypress Creek, 3.2 miles (5.1 km) south
of Mount Pleasant, Union County, Illinois.
RANGE.?Ohio and Big Muddy basins in Alexander, Hardin,
Jackson, Johnson, Massac, Pope, Pulaski, Union, and William-
son counties, Illinois (see Page, 1985b:386).
HABITAT.?Streams.
Orconectes (Crockerinus) jeffersoni Rhoades
FIGURE 157
Orconectes propinquus jeffersoni Rhoades, 1944a: 123, fig. 5a-f.?Fitzpatrick,
1963:61 [by implication].
Orconectes (Orconectes) propinquus jeffersoni.?Hobbs, 1959:894 [by impli-
cation].
Faxonius propinquus jeffersoni?Creaser, 1962:2 [by implication].
Orconectes jeffersoni.?Fitzpatrick, 1967a:152, figs. 2-18, 22.?Hobbs,
1974b:31,fig. 119.
Orconectes (Crockerinus) jeffersoni.?Fitzpatrick, 1987a:51.
TYPES.?Holotype, allotype, and "morphotype" (designated
by Rhoades as allotype), USNM 81316,81318,81317 (male I,
female, male II); paratypes, MCZ, CM, USNM, OSM.
TYPE LOCALITY.?Tributary of Muddy Fork of Beargrass
Creek, 2 miles (3.2 km) east of Louisville corporation line on
U.S. Highway 42, Jefferson County, Kentucky.
RANGE.?Tributaries to Beargrass Creek, Ohio River,
Jefferson County, Kentucky.
HABITAT.?Streams.
Orconectes (Crockerinus) marchandi Hobbs
FIGURE 153
Orconectes marchandi Hobbs, 1948b: 140, figs. 1-4,10-14,17; 1974b:34,fig.
115.?Williams, 1954:843, figs. 1, 50-57.?Fitzpatrick, 1963:61 [by
implication].?Pflieger, 1987a:24; 1987b:8.
Orconectes (Orconectes) marchandi.?Hobbs, 1959:895.
Faxonius marchandi.?Creaser, 1962:2 [by implication].
Orconectes (Crockerinus) marchandi Fitzpatrick, 1987a:51.
TYPES.?Holotype, allotype, and "morphotype," USNM
82072, 82073, 82074 (male I, female, male II); paratypes,
MCZ, USNM, TU.
TYPE LOCALITY.?Stream 3.2 miles (5.1 km) southeast of
Hardy, Sharp County, Arkansas, on U.S. Highway 63.
RANGE.?Spring River drainage system in Lawrence and
Sharp counties, Arkansas, and Oregon County, Missouri.
HABITAT.?Clear, rocky streams.
Orconectes (Crockerinus) obscurus (Hagen)
FIGURE 155
Cambarus obscurus Hagen, 1870:69, pi. 1: figs. 72-75.pl. 3: fig. 154.
Astacus obscurus.?Hagen, 1870:5 [lapsus].
Cambarus propinquus var. obscura.?Faxon, 1885a:92.
Cambarus propinquus obscurus.?Hay, 1899b:960.
Cambarus (Faxonius) obscurus.?Ortmann, 1905c: 112; 19O6b:369,pl. A: figs.
1.2,pi. 39: fig. 7,pi. 40: fig. l.pl. 42: figs. 2,3.
Cambarus propinquus obscura.?Fowler, 1912:564.
Faxonius obscurus.?Creaser, 1933a:5; 1962:2 [by implication].
Faxonius (Faxonius) obscurus.?Creaser, 1933a:5 [by implication].
Orconectes obscurus.? Hobbs, 1942a:352 [by implication]; 1974b:36, fig.
117.?Crocker, 1957:36,53, 75, pi. 3: figs. 5,6, pi. 5: fig. 2.?Fitzpatrick,
1963:61 [by implication]; 1967a: 160,figs. 3,11-15.25.
Orconectes (Orconectes) obscurus.?Hobbs, 1942b: 154 [by implication].
Cambarus obscura.?Filzpatrick, 1967a: 161 [erroneous spelling].
Orconcedes obscurus.?Fielder, 1972:133 [erroneous spelling].
Orconectes (Crockerinus) obscurus.?Filzpatrick, 1987a:50.
TYPES.?Lectotype, MCZ 181 (male I); paralectotypes,
MCZ 181 (9 male 1,6 female), MCZ 3353 (7 male I, female),
MCZ 3354 (3 male II), USNM 4971 (male I), MHNP (male,
female), Wiirzburg Mus.; "cotypes" previously reported to be
in the Australian Museum are not there. The lectotype (male I)
was selected by Fitzpatrick (1967a:162) from MCZ 181, and all
other "types" were designated paralectotypes by him.
TYPE LOCALITY.?Genesee River, Rochester, Monroe
County, New York.
RANGE.?Southeastern Ontario, New York, Pennsylvania,
eastern Ohio, West Virginia, northern Virginia, and western
Maryland.
HABITAT.?Rocky streams.
Orconectes (Crockerinus) propinquus (Girard)
FIGURE 156
Cambarus propinquus Girard, 1852:88.
Astacus propinquus.?Hagen, 1870:61.
Cambarus propinquus propinquus.?Faxon, 1885a:91 [by implication].
NUMBER 480 37
Camborus propinquus.?Williamson, 1899:47 [erroneous spelling].
Cambarus propinquus typicus.?Ortmann, 19O5c:133.
Cambarus (Faxonius) propinquus.?Ortmann, 1905c: 112; 1906b:358, pi. 39:
fig. 6a,b.
Faxonius propinquus.?Williamson, 1907:754.?Creaser, 1933a: 1.
Cambarus (Fraxonius) propinquus.?Turner, 1926:43 [erroneous spelling].
Faxonius (Faxonius) propinquus propinquus.?Creaser, 1933a:3 [by implica-
tion].
Camarus propinguus.?Fleming, 1939:304 [erroneous spelling].
Cambarus propinkuus.?Fleming, 1939:305 [in part; erroneous spelling].
Cambarrus propinquus.?Fleming, 1939:305 [erroneous spelling].
Cambarus Propinquus.?Goodnight, 1940:40.
Orconectes propinquus propinquus.?Hobbs, 1942a:352 [by implication].?
Crocker, 1957:35, 48, 75, frontispiece, pL 3: figs. 1, 2, pi. 5: fig.
1.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) propinquus propinquus.?Hobbs, 1942b: 154 [by
implication].
Orconectes propinquus.?Hoff, 1943:281.?Fitzpatrick, 1967a: 145, figs. 2 -
19.?Hobbs, 1974b:39, fig. 118.?Page, 1985a:569; 1985b:406, figs.
141-144.
Orconnectes propinquus.?Threinen, 1958:1 [erroneous spelling].
Faxonius propinquus propinquus.?Creaser, 1962:2 [by implication].
Orconectes sp. A.?Fitzpatrick, 1967a:150-151,167-169,figs.2-18,27.
Orconectes iowaensis Fitzpatrick, 1968a:507, figs. 1-10 [Holotype, allotype,
and morphotype, USNM 117963,117964,117965 (male I, female, male II);
paratypes, USNM, Coe College, MCZ. Type locality: Turkey River, 2.5
miles (4 km) south of Cresco, Howard County, Iowa].?Hobbs, 1974b:31,
fig. 127.?Phillips, 1980:87, figs. 2c. 6c; 1982:27.
Orconectes propoinquus.?Hart and Hart, 1974:120 [erroneous spelling.]
Orconectes propinguis.?Nolfi, 1978:186 [erroneous spelling).
Orconectes propinques.?Buikema and Benfield, 1979:1711 [erroneous spell-
ing].
Orconectes iowaenis.?Phillips and Reiss, 1979:19 [erroneous spelling].
Orconectes kinderhookensis Fitzpatrick and Pickett, 1980:375, figs. 1-11.
[Holotype, allotype, and morphotype. USNM 148883, 148884, 148885;
paratypes, USNM, Joseph F. Pickett, Sr. Type locality. Kinderhook Creek
0.4 mi (0.6 km) south of the Rensselaer County line on St. Rte. 66, Columbia
County, New York].?D.G. Smith, 1981b:389.
Orconectes (Crockerinus) propinquus.?Fitzpatrick, 1987a:51.
TYPES.?Probably destroyed in the Chicago fire in 1871
(Faxon, 1914:417).
TYPE LOCALITY.?Three localities were originally given by
Girard. From these, Faxon (1914:417) designated Garrison
Creek, Sackett's Harbor, Jefferson County, New York, as the
type locality. Ortmann (1906b:363) cited Oswego, Oswego
County, New York. Crocker (1957:35) and Fitzpatrick
(1967a: 146) accepted Ortmann's designation.
RANGE.?Ontario, Quebec, and New York westward to Iowa
and southeastern Minnesota, and southward to Illinois, Ohio,
Pennsylvania, and Massachusetts.
HABITAT.?Streams and rocky shores of lakes.
Orconectes (Crockerinus) rafinesquei Rhoades
FIGURE 163
Orconectes rafinesquei Rhoades, 1944a: 116, fig. la-f.?Fitzpatrick, 1963:61
[by implication].?Hobbs, 1974b:40, fig. 124.
Orconectes (Orconectes) rafinesquei?Hobbs, 1959:893.
Faxonius rafinesquei.?Creaser, 1962:2 [by implication].
Orconectes (Crockerinus) rafinesquei.?Fitzpatrick, 1987a:51.
TYPES.?Holotype, allotype, and morphotype, USNM
81304, 81306, 81305 (male I, female, male IT); paratypes,
ANSP, MCZ, OSM, BCPM.
TYPE LOCALITY.?Rough River, at Falls-of-Rough, Gray-
son-Breckenridge counties, Kentucky.
RANGE.?Green River basin of western Kentucky.
HABITAT.?Streams.
Orconectes (Crockerinus) sanbornii erismophorous
Hobbs and Fitzpatrick
FIGURE 160
Orconectes propinquus erismophorous Hobbs and Fitzpatrick. 1962:208, figs.
1-15.
Orconectes sanborni erismophorus.?Fitzpatrick, 1967a:131, figs. 2-18, 24
[erroneous spelling].
Orconectes sanborni erismophorous.?Fitzpatrick, 1967a: 160, fig. 24.
Orconectes sanbornii erismophorous.?Hobbs, 1974b:40, fig. 122.
Orconectes (Crockerinus) sanbornii erismophorous.?Fitzpatrick, 1987a:50.
TYPES.?Holotype, allotype, and morphotype, USNM
107597, 107598, 107599 (male I, female, male II); paratypes,
TU.USNM.
TYPE LOCALITY.?Crane Nest Creek at Pee Wee, Wirt
County, West Virginia.
RANGE.?Little Kanawha River system, West Virginia.
HABITAT.?Streams.
Orconectes (Crockerinus) sanbornii sanbornii (Faxon)
FIGURE 159
Cambarus Sanbornii Faxon, 1884:128.
Cambarus propinquus Sanbornii.?Faxon, 1885a:91, pi. 5: fig. 3, pi. 9: figs.
10,10*. 10a, 10a'.
Cambarus propinquus var. sanborni.?Underwood, 1886:372.
Cambarus propinquus var. sanbornii.?Osborn and Williamson,1898:21.
Cambarus propinquus sanbornii.?Faxon, 1898:660.
Cambarus propinquus sanborni.?Ortmann, 1905c: 128.
Cambarus (Faxonius) propinquus sanborni.?Ortmann, 1906b: 365.
Cambarus obscurus sanborni.?Ortmann, 1906b:437.
Faxonius sanborni.?Creaser. 1933a:3 [by implication].
Faxonius (Faxonius) sanborni.?Creaser, 1933b:21 [by implication].
Orconectes propinquus sanborni.?Hobbs, 1942a:352 [by implication].?
Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) propinquus sanborni.?Hobbs, 1942b:154 [by impli-
cation].
Faxonius sanborni sanborni.?Creaser, 1962:2 [by implication].
Orconectes sanborni sanborni.?Fitzpatrick, 1967a: 157, figs. 2-18,23.
Orconectes sanborni.?Stevenson, 1967:208.
Orconectes sanbornii sanbornii.?Hobbs, 1974b:40, fig. 121.
Orconectes senborni.?Campos, 1982:18 [erroneous spelling].
Orconectes (Crockerinus) sanbornii sanbornii.?Fitzpatrick, 1987a:50,fig. 3.
TYPES.?Lectotype, MCZ 3692 (male I); paralectotypes,
MCZ 3587 (lost), MCZ 3692 (male II, female). (See
Fitzpatrick, 1967a: 157.)
TYPE LOCALITY.?Oberlin.Lorain County, Ohio. Restricted
by Faxon, 1914:418.
38 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
RANGE.?Southern Ohio and adjacent West Virginia and
Kentucky.
HABITAT.?Streams.
Orconectes (Crockerinus) shoupi Hobbs
FIGURE 167
Cambarus propinkuus.?Fleming, 1939:305 [in part; erroneous spelling].
Cambarus propinquus sanborni.?Fleming, 1939:305 [inpart],pl. 14.
Orconectes shoupi Hobbs, 1948a: 14,figs. 1-5,9,15,16,19,20; 1974b:41,fig.
129.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) shoupi.?Hobbs, 1959:893.
Faxonius shoupi.?Creaser, 1962:2 [by implication].
Cambarus shoupi.?Bouchard, 1972b:18 [lapsus calami].
Orconectes (Crockerinus) shoupi.?Fitzpatrick, 1987a:51.
TYPES.?Holotype, allotype, and morphotype, USNM
84072 (male I, female, male II); paratypes, MCZ, USNM.
TYPE LOCALITY.?Mill Creek, tributary of Cumberland
River, east of Oglesby near Antioch Pike, 10 miles (16 km)
south of Nashville, Davidson County, Tennessee.
RANGE.?Known only from the vicinity of Nashville,
Tennessee.
HABITAT.?Streams.
Orconectes (Crockerinus) stannardi Page
FIGURE 161
Orconectes stannardi Page, 1985a:564, fig. 1.
Orconectes (Crockerinus) stannardi.?Fitzpatrick, 1987a:50.
TYPES.?Holotype, allotype, and morphotype, INHS (male
I, female, male II); paratypes, INHS, USNM.
TYPE LOCALITY.?Little Wabash River at secondary road
719,3.75 miles (6 km) NNW of Louisville (Sec. 33SW, T. 5N,
R. 6E),Clay County, Illinois.
RANGE.?Little Wabash River system in southeastern
Illinois.
HABITAT.?Streams, principally riffle areas.
Orconectes (Crockerinus) tricuspis Rhoades
FIGURE 165
Orconectes tricuspis Rhoades, 1944a:117, fig. 2a-f.?Hobbs, 1948a:20;
1974b:41,fig. 126.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) tricuspis.?Hobbs, 1959:894.
Faxonius tricuspis.?Creaser, 19622 [by implication].
Orconectes (Crockerinus) tricuspis.?Fitzpatrick, 1987a:51.
TYPES.?Holotype, allotype, and "morphotype" (latter des-
ignated as allotype by Rhoades), USNM 81307,81309,81308
(male I, female, male II); paratypes, ANSP, MCZ, OSM.
TYPE LOCALITY.?Pete Lights' spring, 3 miles (4.8 km) east
of Canton, Trigg County, Kentucky.
RANGE.?Lower Green, Tradewater, and Cumberland drain-
ages in western Kentucky.
HABITAT.?Small streams.
Orconectes (Crockerinus) virginiensis Hobbs
FIGURE 158
Orconectes virginiensis Hobbs, 1951a:122, figs. 1-10; 1974b:42, fig.
120.?Fitzpatrick, 1963:61 [by implication]; 1967a: 164, figs. 2-18,26.
Orconectes (Orconectes) virginiensis.?Hobbs, 1959:893.
Faxonius virginiensis.?Creaser, 1962:2 [by implication].
Orconectes virginianus.?Fitzpatrick, 1967a:133 [erroneous spelling].
Orconectes (Crockerinus) virginiensis.?Fitzpatrick, 1987a:50.
TYPES.?Holotype, allotype, and morphotype, USNM
91659, 91660, 91661 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Rowanty Creek, tributary of the Not-
toway River, 3.3 miles (5.3 km) south of Reams Station on U.S.
Highway 301, Dinwiddie County, Virginia.
RANGE.?Chowan drainage system in North Carolina and
Virginia.
HABITAT.?Streams.
Subgenus Faxonius Ortmann
Astacus.?Rafinesque, 1817:42 [in part].
Cambarus.?Girard, 1852:87 [in part].
Subgenus Astacus.?Herklots, 1861:144 [in part].
Subgenus Faxonius Ortmann, 1905c:97 [in part]. [Type species, by original
designation, Astacus limosus Rafinesque, 1817:42. Gender masculine.].?
Fitzpatrick, 1987a:51.
Faxonius.?Creaser, 1933a:3 [in part].
Orconectes?Hobbs, 1942a:352 [inpart].
Subgenus Orconectes.?Hobbs, 1942b:154 [in part].
Orconetes.?A vault, 1973:242 [erroneous spelling].
Orconeotes.?Unestam, 1973:4 [erroneous spelling].
Orconelctes.?Brodsky, 1981:122 [erroneous spelling].
Orconectes (Faxonius) indianensis (Hay)
FIGURE 142
Cambarus indianensis Hay, 1896:494, fig. 9.
Cambarus (Faxonius) indianensis.?Ortmann, 1905c: 112.
Faxonius indianensis.?Creaser, 1933a:3 [by implication]; 1962:2 [by implica-
tion].
Faxonius (Faxonius) indianensis.?Creaser, 1933b:21 [by implication].
Orconectes indianensis.?Hobbs, 1942a:352 [by implication]; 1974b:31, fig.
105.^itzpatrick, 1963:61 [by implication).?Page, 1985b:394, figs. 124-
126.
Orconecters (Orconectes) indianensis.?Hobbs, 1942b: 154 [by implication].
Orconectes (Faxonius) indianensis.?Fitzpatrick, 1987a:53.
TYPES.?^Syntypes, USNM 14624 (2 male 1,2 female), MCZ
3859 (2 male 1,2 female).
TYPE LOCALITY.?Patoka River, Patoka, Gibson County,
Indiana.
RANGE.?Wabash Basin in southern Indiana and Sabine
River and Honey Creek watersheds in southern Illinois. (See
Page, 1985b:394.)
Orconectes (Faxonius) limosus (Rafinesque)
FIGURE 140
Astacus limosus Rafinesque, 1817:42.
Astacus affinis Say, 1817:168. [Types, not extant. Type locality, Delaware
NUMBER 480 39
River.]
Astacus (Cambarus) qffinis.?Erichson, 1846:96.
Cambarus Pealti Girard, 1852:87. [Syntypes, USNM 2081 (2 male I, 2
female). Type locality, Potomac River at Washington, D.C.]
Cambarus affinis.?Girard, 1852:87.
Astacus (Astacus) affinis.?HeikloU, 1861:144.
Cambarus (Faxonius) limosus.?Ortmann, 1905c: 107.?Fowler, 1912:353,
ph. 104,105.
Cambarus limosus.?Ortmann, 1905c:107.
Cambarus pealei.?Ortmann, 1906b:352.
Faxonius limosus.?Creaier, 1933a:3 [by implication]; 1962:2 [by implica-
tion].
Faxonius (Faxonius) limosus.?Creaser, 1933b:21 [by implication].
Orconectes limosus.?Hobbs 1942a:352 [by implication]; 1948a:20, fig. 22;
1974b:33,fig. 103.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) limosus.?Hobbs, 1942b:154 [by implication].
Faxonius qffinis.?Hobbs, 1942c:55.
Orconects limosus.?Lewis, 1955:146 [erroneous spelling].
Orconectes affinis.?Holthuis, 1956:228.
Orconectes limosa.?Mobberly, 1963:80 [erroneous spelling].
Camburus affinis.?Bicescu, 1967:218 [erroneous spelling].
Orconetes limosus.?Avault, 1973:242 [erroneous spelling].
Orconeotes limosus.?Unestam, 1973:4 [erroneous spelling].
Oroncestes limosus.?Wielgus, 1973:119 [erroneous spelling].
Orconectes limnosus.?Ingle and Thomas, 1974:525 [erroneous spelling].
Orconectes limusus.?Reynolds, 1979:57 [erroneous spelling].
Orconelctes limosus.?Brodsky, 1981:122 [erroneous spelling].
Orconectes Limosus.?D.G. Smith, 1981a:169 [lapsus].
Orconectes (Faxonius) limosus.?Fitzpatrick, 1987a:53, fig. 4.
TYPES.?Not known to exist
TYPE LOCALITY.?Delaware River near Philadelphia, Penn-
sylvania.
RANGE.?The Atlantic watershed from Maine to the lower
James River in Virginia. Introduced into western Europe.
HABITAT.?Streams.
Orconectes (Faxonius) wrighti Hobbs
FIGURE 141
Orconectes wrighti Hobbs, 1948c:85, pi. 3: figs. 1-9; 1974b:43, fig.
104.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) wrighti.?Hobbs, 1959:893.
Faxonius wrighti?Creaser, 1962:2 [by implication].
Orconectes (Faxonius) wrighti.?Fitzpatrick, 1987a:53.
TYPES.?Holotype, USNM 85144 (male I); allotype and
morphotype, USNM 85145 (female, male II); paratypes,
USNM.
TYPE LOCALITY.?Robinson Creek on State Route 57,
Hardin County, Tennessee.
RANGE.?Known only from Hardin County, Tennessee.
HABITAT.?Streams.
Subgenus Gremicambarus Fitzpatrick
Cambarus.?Hagen, 1870:63 [in part].
Subgenus Faxonius Ortmann, 1905c: 113 [in part].
Faxonius.?Williamson, 1907:749 [in part; intended as subgeneric name].?
Creaser, 1933a:3 [in part].
Orconectes.?Hobbs, 1942a:352 [in part].
Subgenus Orconectes.?Hobbs, 1942b: 154 [in part].
Orconnectes.?Thrcinen, 1958:1 [erroneous spelling].
Orconectis.?Dudel, 1965:324 [erroneous spelling].
Camburus.?Bicescu, 1967:218 [erroneous spelling].
Orchonectes.?Dimond et al., 1968:760 [erroneous spelling].
Orconectus.?Ray and Stevens, 1970:58 [erroneous spelling].
Oronectes.?Durliat and Vranckx, 1976:474 [erroneous spelling].
Orcnelctes.?Brodsky, 1981:122 [erroneous spelling].
Orconetes.?Huner, 1981:47 [erroneous spelling].
Orconcetes.?Huner and Naqui, 1984:405 [erroneous spelling].
Subgenus Gremicambarus Fitzpatrick, 1987a:53. [Type species, by original
designation, Cambarus virilis Hagen, 1870:63. Gender masculine.]
Orconectes (Gremicambarus) alabamensis (Faxon)
FIGURE 206
Cambarus Alabamensis Faxon 1884:125; 1885a:104,pl. 4: fig. 4, pi. 10: figs.
3,3', 3a, 3a'.
Cambarus alabamensis.?Hay, 1899b:960.
Cambarus (Faxonius) alabamensis.?Ortmann, 1905c: 112.
Faxonius alabamensis.?Creaser, 1933a:3 [by implication]; 1962:2 [by
implication].
Faxonius (Faxonius) alabamensis.?Creaser, 1933b:16 [by implication].
Orconectes alabamensis.?Hobbs, 1942a:352 [by implication]; 1974b:26, fig.
166.?Fitzpatrick, 1963:61 [by implication].?Cooper and Hobbs, 1980:2,
figs. 1.2a.
Orconectes (Orconectes) alabamensis.?Hobbs, 1942b: 154 [by implication].
Orconectes (Gremicambarus) alabamensis.?Fitzpatrick, 1987a:54.
TYPES.?Syntypes, USNM 4876 (14 male I, 16 female, 4
male II), MCZ 3565 (2 male 1,2 female, 2 male II.
TYPE LOCALITY.?Second Creek, Waterloo, Lauderdale
County, Alabama.
RANGE.?Tennessee River basin from Shoal Creek in
Lawrence County, Tennessee, and Lauderdale County, Ala-
bama, to Hardin County, Tennessee.
Orconectes (Gremicambarus) causeyi Jester
FIGURE 200
Orconectes causeyi Jester, 1967:518, figs. 1-12,?Reiner, 1969:52, figs. 4,
10,24.37.?Hobbs, 1974b:17, fig. 163.
Orconectes acuseyi.?Loring and Hill, 1976:219 [erroneous spelling].
Orconectes nais form causeyi.?Unger, 1978:14.
Orconectes (Gremicambarus) causeyi.?Fitzpatrick, 1978a:54.
TYPES.?Holotype, allotype, and morphotype, USNM
115698,115699,1156700 (male I, female, male II); paratypes,
USNM.TU.
TYPE LOCALITY.?Conchas Reservoir, 32 miles (51.2 km)
northwest of Tucumcari on State Route 104, San Miguel
County, New Mexico.
RANGE.?"New Mexico, Texas, Arkansas, Oklahoma, Mis-
souri, and Kansas" (Reimer, 1969:55). To this may be added
southwestern Colorado.
HABITAT.?Streams and impoundments.
REMARKS.?As stated in my earlier checklist (Hobbs,
1974b) and elsewhere, this crayfish appears to me to be
indistinguishable from Orconectes (G.) virilis (Hagen). Until
the relationships of these two species and O. (G.) nais (Faxon)
40 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
become more adequately understood, the conservative recogni-
tion of the three seems advisable. Particularly does this seem
prudent in light of Unger's (1978) having presented evidence
that the three taxa are perhaps distinct and that 0. (G.) causeyi
is more similar to O. (G.) nais than to 0. (G.) virilis.
Unfortunately, his samples were quite limited both in quantity
and in representing the range of the complex.
Orconectes (Gremicambarus) chickasawae Cooper and
Hobbs
FIGURE 209
Orconectes chickasawae Cooper and Hobbs, 1980:29, fig. 10.
Orconectes (Gremicambarus) chickasawae.?Fitzpatrick, 1987a:54.
TYPES.?Holotype, allotype, and morphotype, USNM
144924,144925,144926); paratypes,USNM,NCSM.
TYPE LOCALITY.?Town Creek, 3.5 miles (5.6 km) south-
east of Muldon, 3 miles (4.8 km) east of State Route 45W
(CS:T. 16S, R. 7E, Sec. 7 and 18), Monroe County,
Mississippi.
RANGE.?"Known from western tributaries of the Tom-
bigbee River in Chickasaw, Clay, Lowndes, and Monroe
counties, Mississippi" (Cooper and Hobbs, 1980:34).
HABITAT.?Lentic and sluggish lotic situations.
Orconectes (Gremicambarus) compressus (Faxon)
FIGURE 210
Cambarus compressus Faxon, 1884:127; 1885a:105,pL 5: fig. 6, pi. 10: figs. 2,
2', 2a, 2a'.
Cambarus (Faxonius) compressus.?Ortmann, 1905c: 112.
Faxonius compressus.?Creaser, 1933a:3 [by implication]; 1962:2 [by implica-
tion].
Faxonius (Faxonius) compressus.?Creaser, 1933b:16 [by implication].
Orconectes compressus.?Hobbs, 1942a:352 [by implication]; 1949:17;
1974b:27.fig. 169.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) compressus.?Hobbs, 1942b: 154 [by implication].
Orconectes (Gremicambarus) compressus.?Fitzpatrick, 1987a:54.
TYPES.?Syntypes, USNM 4878 (11 male I, 13 female),
USNM 4879 (5 male 1,3 female, 1 male II),MCZ 3583 (3 male
1,2 female, 1 male II).
TYPE LOCALITY.?Second Creek, Waterloo, Lauderdale
County, Alabama. Restricted by Faxon (1914:419).
RANGE.?Tennessee River basin, below Walden Ridge, in
Alabama, Mississippi, and Tennessee; Cumberland River basin
in Tennessee and Kentucky; and the Barren River watershed in
Tennessee and Kentucky.
HABITAT.?Clear, rock- or gravel-bedded streams.
Orconectes (Gremicambarus) cooperi Cooper and Hobbs
FIOURE204
Orconectes cooperi Cooper and Hobbs, 1980:17, fig. 7.
Orconectes (Gremicambarus) cooperi.?Fitzpatrick, 1987a:54.
TYPES.?Holotype, allotype, and morphotype, USNM
147722,147723,147724; paratypes, USNM, NCSM.
TYPE LOCALITY.?Brier Fork of the Flint River at U.S.
Highway 231-431, about 2 miles (3.2 km) north of Meridian-
ville (Sec. 7, T. 2S, R. IE), Madison County, Alabama.
RANGE.?Known only from the Flint River watershed in
Madison County, Alabama, and Lincoln County, Tennessee.
HABITAT.?Streams.
Orconectes (Gremicambarus) etnieri Bouchard and
Bouchard
FIGURE 205
Orconectes etnieri Bouchard and Bouchard, 1976b:459, fig. 1.
Orconectes (Gremicambarus) etnieri.?Fitzpatrick, 1987a:54.
TYPES.?Holotype, allotype, and morphotype, USNM
146583, 146584,146585 (male I, female, male II); paratypes,
USNM,TU.
TYPE LOCALITY.?Robinson Creek at Tennessee State
Highway 57, Hardin County, Tennessee.
RANGE.?Western tributaries of the Tennessee River from
Robinson Creek northward to Snake Creek in Hardin County,
and in the Forked Deer, Hatchie, and Loosahatchie drainage
systems in Tennessee and Mississippi.
HABITAT.?Streams.
Orconectes (Gremicambarus) holti Cooper and Hobbs
FIGURE 202
Orconectes holti Cooper and Hobbs, 1980:23, figs. 8,9.
Orconectes (Gremicambarus) holti.?Fitzpatrick, 1987a:54.
TYPES.?Holotype, allotype, and morphotype, USNM
147149,147150,147151 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Bogue Chitto Creek, 2.3 miles (3.7 km)
west of Marion on State Route 14 (Sec. 11, T. 19N, R. 7E),
Perry County, Alabama.
RANGE.?Alabama River basin in Dallas, Lowndes,
Montgomery, Perry, and Wilcox counties, Alabama.
HABITAT.?Streams.
Orconectes (Gremicambarus) immunis (Hagen)
FIGURE 201
Cambarus immunis Hagen, 1870:71, pi. lrfigs. 101,102, pi. 3: fig. 160; pi. 8:
fig. b.
Cambarus signifer Herrick, 1882:253, fig. 7. [Types, MCZ 3515 (2 male L 1
female). Type locality, Grass Lake, Richfield, Hennepin County, Minne-
sota.]
Cambarus immunis spinirostris Faxon, 1884:146. [Types, USNM 4655 (6 male
II, 8 female), MCZ 3562 (2 male II, 2 female). Type locality, Reelfoot Lake.
Obion County, Tennessee.]
Cambarus immunis immunis.?Faxon, 1884:146 [by implication].
Cambarus immunis var. spinostris.?Harris, 1901:115 [erroneous spelling].
Cambarus rinmunis.?Steele, 1902:3 [erroneous spelling].
NUMBER 480 41
Cambarus (Faxonius) immunis.?Ortmann, 1905c:l 13.
Faxonius immunis.?Williamson, 1907:749 [Faxonius intended as subgeneric
name].?Creaser, 1933a:4; 1962:2.
Faxonius (Faxonius) immunis immunis.?Creaser, 1933b:21 [by implication].
Faxonius (Faxonius) immunis pedianus Creaser, 1933b: 14 [Subgenus by
implication. Holotype and allotype, USNM 98334,98335 (male I, female);
paratypes, USNM. Type locality, reservoir at Englewood, Denver, Colo-
rado].
Cambarus immunis Hagenianus.?Fleming, 1938:303 [(?) lapsus for C.
immunis Hagen].
Orconectes immunis immunis.?Hobbs, 1942a:352 [by implication].?
Williams, 1954:890, figs. 200-208.?Fitzpatrick, 1963:61 [by implication].
Orconectes immunis pedianus.?Hobbs, 1942a:352.
Orconectes (Orconectes) immunis immunis.?Hobbs, 1942b: 154 [by implica-
tion].
Orconectes immunis.?Penn, 1950d:645.?Hobbs, 1974b:30,fig. 164.?Page,
1985b:388,figs. 120-123.?Pflieger, 19 87a :20; 1987b:6.
Orconnectes immunis.?Threinen, 1958:2 [erroneous spelling].
Orconectes (Orconectes) immunis.?Hobbs, 1959:894.
Orchonectes immunis.?Dimond et al., 1968:760 [erroneous spelling],
Orconectes imunis.?Ono and Nakajima, 1979:1681 [erroneous spelling].
Orconcetes immunis.?Huner and Naqvi, 1984:405 [erroneous spelling].
Orconectes (Gremicambarus) immunis.?Fitzpatrick, 1987a:54.
TYPES.?Syntypes, MCZ 188 (2 male 1,2 ovigerous female.)
The "paratypes" of Faxon (1914), MCZ 3355 (1 male I), are
dubious "types" because this number was not listed by Hagen
(1870:73). Douglas G. Smith has examined MCZ lots 3355,
listed as paratypes by Faxon (1914:421), and 1841, listed as
such by Hagen 1870:73, and found the former to contain a first
form male of 0. immunis from Belleville, Illinois, the latter, a
marine crustacean from Society Island. Probably Hagen erred
in recording the specimen from Belleville as having been
catalogued under number 1841. The "paratype" said to be in
MHNP could not be located.
TYPE LOCALITY.?Lawn Ridge, Marshall County, Illinois.
RANGE.?Maine and Connecticut to eastern Colorado and
Wyoming, and from Alabama to southern Manitoba, Ontario,
and Quebec. Its presence in New England probably resulted
from introductions (Douglas G. Smith, in litt).
HABITAT.?Ponds, roadside ditches, and sluggish streams.
Orconectes (Gremicambarus) mississippiensis (Faxon)
FIGURE 198
Cambarus Mississippiensis Faxon, 1884:123; 1885a:101,pl. 3: fig. 2, pi. 10:
figs. 4,4', 4a, 4a'.
Cambarus mississippiensis.?Hay, 1899b:960.
Cambarus (Faxonius) mississippiensis.?Ortmann, 1905c: 113.
Faxonius mississippiensis.?Creaser, 1933a:3 [by implication]; 1962:2 [by
implication].
Faxonius (Faxonius) mississippiensis.?Creaser, 1933b:19 [by implication].
Orconectes mississippiensis.?Hobbs, 1942a:352 [by implication]; 1974b:35,
fig. 161.?Fitzpatrick, 1963:61 [by implication].?Cooper and Hobbs,
1980:35, figs. 11-13.
Orconectes (Orconectes) mississippiensis.?Hobbs, 1942b:154 [by implica-
tion].
Orconectes (Gremicambarus) mississippiensis.?Fitzpatrick, 1987a:54.
TYPES.?Syntypes, USNM 19760 (male I, female), USNM
19759 (male II, female), MCZ 3563 (female).
TYPE LOCALITY.?Eastern Mississippi. Restricted to Macon,
Noxubee County, Mississippi, by Faxon (1914:422).
RANGE.?"Limited to the Prairie Section of the Noxubee
River basin and in other tributary systems of the Tombigbee
River northward to the Catalpa Creek basin in Kemper,
Noxubee, Lowndes, and Oktibbeha counties, Mississippi"
(Cooper and Hobbs, 1980:40).
Orconectes (Gremicambarus) nais (Faxon)
FIGURE 203
Cambarus Nais Faxon, 1885b:140.
Cambarus nais.?Underwood, 1886:371.
Cambarus nais.?Faxon, 1890:631.
Cambarus pilosus Hay, 1899a:121, fig. 1. [Types, USNM 19761 (6 male II),
MCZ 7389 (male II). Type locality, Beloit, Mitchell County, Kansas.]
Cambarus pelosus.?Harris, 1901:115 [erroneous spelling].
Cambarus (Faxonius) nais.?Ortmann, 1905c:l 13.?Creaser and Ortenberger,
1933:37, figs. 10,14,24.
Cambarus (Faxonius) pilosus.?Ortmann, 1905c: 113.
Faxonius nais.?Creaser, 1933a:3; 1962:2 [by implication].
Faxonius (Faxonius) nais.?Creaser, 1933b:5.
Orconectes nais.?Hobbs, 1942a:352 [by implication]; 1974b:35, fig. 165.?
Williams, 1954:886, figs. 191-199.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) nais.?Hobbs, 1942b:154 [by implication].
Orconectus nais.?Ray and Stevens, 1970:58 [erroneous spelling].
Orconectes mais.?Craelius, 1976:406 [erroneous spelling].
Orconectes (Gremicambarus) nais.?Fitzpatrick, 1987a:54.
TYPES.?Syntypes, MCZ 3755 (2 male I, male II, female),
USNM 129213 (3 male 1,3 male II); Mus. Washburn Coll. (no
longer there).
TYPE LOCALITY.?Labette County, Kansas.
RANGE.?Great Plains: Oklahoma and Texas; the limits not
known. (See Bouchard and Robison, 1981:28.)
HABITAT.?Streams.
REMARKS.?The syntypes in the National Museum of
Natural History are specimens collected by W.S. Newlon from
the type locality and were received from the Kansas Museum of
Natural History; presumably they represent a part of the lot that
was said by Faxon (1914:420) to have been at Washburn
College. See "Remarks" under Orconectes (G.) causeyi.
Orconectes (Gremicambarus) rhoadesi Hobbs
FIGURE 207
Cambarus validus Faxonianus.?Fleming, 1938:303 [error for "C. validus
Faxon"].
Orconectes rhoadesi Hobbs, 1949:19, pi. 3: figs. 1-10; 1974b:40, fig.
167.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) rhoadesi?flobbs, 1959:895.
Faxonius rhoadesi.?Creaser, 1962:2 [by implication].
Orconectes (Gremicambarus) rhoadesi.?Fitzpatrick, 1987a:54.
TYPES.?Holotype, morphotype, USNM 87953 (male I,
male II), and allotype, USNM 87954 (female); paratypes,
USNM.
42 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
TYPE LOCALITY.?Otter Creek, between Granny White Pike
and Hillsboco Pike, about 7 miles (11.2 km) south of Nashville,
Davidson County, Tennessee.
RANGE.?Cumberland, Duck, and Tennessee basins in
central Tennessee.
HABITAT.?Streams.
Orconectes (Gremicambarus) validus (Faxon)
FIGURE 208
Cambarus validus Faxon, 1914:382, pi. 7: figs. 3 ,4 ,8, pi. 13: fig. 1.
Faxonius validus.?Crcaser, 1933a:3 [by implication]; 1962:2 [by implica-
tion].
Faxonius (Faxonius) validus.?Crcascr, 1933b:21 [by implication].
Orconectes validus.?Hobbs, 1942a:352 [by implication]; 1974b:42, fig.
168.?Fitzpatrick, 1963:61 [by implication].?Cooper and Hobbs, 1980:8,
figs. 2b, 3-6.
Orconectes (Orconectes) validus.?Hobbs, 1942b: 154 [by implication].
Cambarus (Faxonius) validus.?Bouchard, 1972b: 103.
Orconectes (Gremicambarus) validus.?Fitzpatrick, 1987a:54.
TYPE.?Holotype, MCZ 301 (male I).
TYPE LOCALITY.?Huntsville, Madison County, Alabama.
RANGE.?Tennessee and Black Warrior river systems in
northern Alabama and southern Tennessee.
HABITAT.?Streams.
Orconectes (Gremicambarus) virilis Hagen
FIGURE 199
Cambarus virilis Hagen, 1870:63, pi. 1: figs. 23-28, pi. 2: figs. 128-132, pi.
3: fig. 155.pl.8.
Cambarus wisconsinensis Bundy, 1876:4. [Type, MCZ 3448 (male II). Type
locality, Racine, Racine County, Wisconsin.]
Cambarus debilis Bundy, 1876:24. (Type, MCZ 3449 (male II). Type locality,
Bamboo River, Ironton, Sauk County, Wisconsin.]
Cambarus couesi Streets. 1877:803. [Syntypes, USNM 3154 (1 male L 3
juvenile male, 5 juvenile female), MCZ 3545 (male I, female, male II). Type
locality. Red River of the North, near Pembina, Pembina County, North
Dakota.]
Cambarus viriles.?Bundy, 1882:180 [erroneous spelling].
Cambarus cousii.?Bundy, 1883:402 [erroneous spelling].
Cambarus Couesi.?Faxon, 1885a.-97.
Cambarus wisconsiensis.?Harris, 1900271 [erroneous spelling].
Cambarus cousei.?Harris, 1903:134 [erroneous spelling].
Cambarus viridis.?Moenkhaus, 1904:111 [erroneous spelling].
Cambarus (Faxonius) virilis.?Oitmaim, 1905c:113.?Creaser, 1932:326, figs.
1,2,8.
cambarus virilis.?Harmon, 1910: fig. 1 [lapsus].
Cambarus virile.?Leger, 1924:1206 [erroneous spelling].
Faxonius virilis.?Creaser, 1933a:3; 1962:2 [by implication].
Faxonius (Faxonius) virilis.?-Creaser, 1933b:21 [by implication].
Orconectes virilis.?Hot**, 1942a:352 [by implication]; 1972b:91,figs. 72h,
73e; 1974b:42, fig 162.?Fitzpatrick, 1963:61 [by implication].?Page,
1985b:417.figs. 151-154.?Pflieger, 1987a:22; 1987b:6.
Orconectes (Orconectes) virilis.?Hobbs, 1942b:154 [by implication].
Orconnectes virilis.?Threinen, 1958:1 [erroneous spelling].
Cambarus virilut.?Hama, 1961:329 [erroneous spelling].
Orconectis virilis.?Dudel, 1965:324 [erroneous spelling).
Camburus viridis.?Bicescu, 1967218 [erroneous spelling].
Orconectes (Cambarus) virilis.?Mason, 1970b:975.
Oronectes virilis.?Durliat and Vranckx, 1976:474 [erroneous spelling].
Orconelcus virilis.?Brodsky, 1981:122 [erroneous spelling].
Orconetes virilis.?Huner, 1981:47 [erroneous spelling].
Orconectes (Gremicambarus) virilis.?Fitzpatrick, 1987a:54,fig. 5.
TYPES.?Syntypes, MCZ 194 (male I, male II, 4 female, 2
ovigerous female), MCZ 196 (7 male 1,3 male II, 3 female, juv.
male, juv. female), MCZ 203 (male II, 3 female), MCZ 1151
(male I), MCZ 3342 (male 1,5 male II, female, 2 juv. male, juv.
female), MCZ 3343 (4 male I, female), MCZ 3344 (male II, 2
female, juv. male, 3 juv. female), MHNP, Wurzburg Mus.,
AMS.
TYPE LOCALITY.?"Lake Superior," restricted by Faxon,
(1914:420).
RANGE.?From Saskatchewan to Ontario, Canada, and from
Montana and Utah to Arkansas, New York, and Maine.
Introduced into California (Riegel, 1959:29, 37); Maryland
(Meredith and Schwartz, 1960:28); parts of New England
(Crocker, 1979:247; D.G. Smith, 1979:388); Alabama, Tennes-
see, Mississippi, and West Viginia (Bouchard, 1976a:583);
Utah (Johnson, 1986:630); New York (Crocker, 1979:247);
Chihuahua, Mexico (Campos and Contreras, 1985:218); and
Pennsylvania (specimen in National Museum of Natural
History from Adams County). See "Remarks" under 0. (G.)
causeyi.
HABITAT.?Lentic and lotic situations.
Subgenus Hespericambarus Fitzpatrick
Cambarus.?Faxon, 1898:656 [in part].
Subgenus Faxonius Ortmann, 1905c:l 13 [in part].
Faxonius.?Creaser, 1933a:3 [in part].
Orconectes.?Hobbs, 1942a:352 [by implication, in part].
Subgenus Orconectes.?Hobbs, 1942b: 154 [in part].
Subgenus Hespericambarus Fitzpatrick, 1987a:54. [Type species by original
designation, Cambarus difficilis Faxon, 1898:656. Gender masculine.]
Orconectes (Hespericambarus) blacki Walls
FIGURE 144
Orconectes difficilis blacki Walls, 1972:454, figs. 3a-e, 4a.?Hobbs,
1974b:28,fig. 107.
Orconectes (Hespericambarus) blacki.?Fitzpatrick, 1987a:55.
TYPES.?Holotype, USNM 131255 (male I); paratypes,
USNM, MCZ, TU, Joe B. Black, and Jerry G. Walls.
TYPE LOCALITY.?Bearhead Creek, 4 miles (6.4 km)
southwest of Juanita on State Route 109, Beauregard Parish,
Louisiana.
RANGE.?Known from only six localities in Beauregard and
Calcasieu parishes, Louisiana, and, according to Walls,
1972:455, intergrading with 0. d. hathawayi "in Calcasieu
River drainage of Allen, Beauregard, Calcasieu, and Vernon
Parishes, Louisiana." In his recent revision of the genus
Orconectes, however, Fitzpatrick (1987a), as noted above,
recognized no geographic races of Orconectes (H.) difficilis,
according Walls' subspecies specific rank.
HABITAT.?Streams.
NUMBER 480 43
Orconectes (Hespericambarus) deanae Reimer and
Jester, new combination
FIGURE 148
Orconectes deanae Reimer and Jester, 1975:17, fig. 1-7.?Bouchard,
1978c:415.
Orconectes denae.?Bouchard, 1978b:451 [erroneous spelling].
Orconectes (Buannuliftctus) denae.?Fitzpatrick, 1987a:49 [erroneous spell-
ing].
TYPES.?Holotype and allotype, USNM 133759, 133760;
paratypes, USNM, Rollin D. Reimer.
TYPE LOCALITY.?Conchas Lake, located at junction of
Conchas and South Canadian rivers in San Miguel County, on
State Road 104, approximately 32 miles (51.2 km) northwest of
Tucumcari, New Mexico.
RANGE.?Known only from Canadian River basin in San
Miguel and Quay counties, New Mexico.
HABITAT.?Lentic and lotic habitats.
Orconectes (Hespericambarus) difficilis (Faxon)
FIGURE 143
Cambarus difficdis Faxon, 1898:656,pi. 65: figs. 1-4.
Cambarus (Faxonius) diffic'dis.?Ortmaim, 1905c: 113.
Faxonius difficilis.?Creaser, 1933a:3 [by implication]; 1962:2 [by implica-
tion].
Faxonius (Faxonius) difficilis.?Creaser, 1933b: 16 [by implication].
Cambarus (Faxonius) difficult.?Fleming, 1938:300 [erroneous spelling].
Orconectes difficilis.?Hobbs, 1942a:352 [by implication].?Williams,
1954:898, figs. 217-224,233.^itzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) difficilis.?Hobbs, 1942b: 154 [by implication];
1959:892.
Orconectes difficilis difficUis.?WaOa, 1972:456.?Hobbs, 1974b:28, fig. 106.
Orconectes (Hespericambarus) difficilis.?Fitzpatrick, 1987a:55, fig. 6.
TYPES.?Syntypes, USNM 19687 (9 male I, 10 female, 5
male II), USNM 19688 (male I), MCZ 4359 (male I, female,
male II), Mus. Zool. Torino.
TYPE LOCALITY.?McAlester, Pittsburg County, Oklahoma.
RANGE.?Pittsburg and Latimer counties, Oklahoma, Up-
shur County, Texas, to Washington County, Arkansas, south-
eastward into Louisiana (see Walls, 1972).
HABITAT.?Rocky streams.
REMARKS.?See "Range" under O. (H.) blacki.
MCZ, CM, ANSP, USNM, AMNH, TU.
TYPE LOCALITY.?Spring Creek, 5.1 miles (8.2 km) east of
Hineston on State Route 85,Rapides Parish, Louisiana.
RANGE.?Jackson and Rapides parishes south to Vermilion
Parish, Louisiana.
HABITAT.?Streams.
REMARKS.?See "Range" under 0. (H.) blacki.
Orconectes (Hespericambarus) maletae Walls
FIGURE 145
Orconectes difficilis maletae Walls, 1972:456, figs. 2a-e, 4b.?Hobbs,
1974b:28.fig. 108.
Orconectes (Hespericambarus) maletae.?Fitzpatrick, 1987a:55.
TYPES.?Holotype, USNM 131254 (male I); paratypes,
USNM, MCZ, TU, Joe B. Black, and Jerry G. Walls.
TYPE LOCALITY.?Bayou Santabarb "(third branch north of
Kisatchie)" on State Route 117, Natchitoches Parish, Louisi-
ana.
RANGE.?Known from only seven localities in Natchitoches
and Sabine parishes, Louisiana, and Upshur County, Texas.
Specimens from Coal County, Oklahoma, were interpreted by
Walls (1972) as intergrades between this crayfish and 0. (H.)
difficilis.
HABITAT.?Streams.
REMARKS.?See "Range" under 0. (H.) blacki.
Orconectes (Hespericambarus) perfectus Walls
FIGURE 147
Orconectes perfectus Walls, 1972:451, figs. la-?, 4c.?Hobbs, 1974b:38, fig.
110.
Orconectes (Hespericambarus) perfectus.?Fitzpatrick, 1987a:55.
TYPES.?Holotype, USNM 131256 (male I); paratypes,
USNM.
TYPE LOCALITY.?Satilpa Creek, 8.7 miles (13.9) km
northwest (not "east") of Grove Hill on Chilton Road, Clarke
County, Alabama.
RANGE.?Widespread in the Tombigbee and lower Alabama
river basins in Alabama and Mississippi.
HABITAT.?Streams.
Orconectes (Hespericambarus) hathawayi Penn
FIGURE 146
Orconectes (Orconectes) hathawayi Penn, 1952b:l, figs. 1-14.
Orconectes hathawayi.?Penn, 1953d:7.?Fitzpatrick, 1963:61 [by implica-
tion].
Faxonius hathawayi.?Creaser, 1962:2 [by implication].
Orconectes difficdis hathawayi.?Wills, 1972:453.?Hobbs, 1974b:28, fig.
109.
Orconectes (Hespericambarus) hathawayi.?Fitzpatrick, 1987a:55.
TYPES.?Holotype, allotype, and morphotype, USNM
92998, 92999, 93000 (male I, female, male II); paratypes,
Subgenus Orconectes Cope
Astacus.?Tellkampf, 1844a:684 [in part].
astacus.?Tellkampf, 1845:93 [lapsus].
Subgenus Cambarus Erichson, 1846:95 [in part].
Cambarus.?Girard, 1852:87 [in part].
Orconectes Cope, 1872:419 [in part]. [Type species.by subsequent designation
(Fowler, 1912:339), Orconectes inermis Cope, 1872:419. Gender mascu-
line.
Subgenus Orconectes.?Hovey, 1882:222 [in part].?Fitzpatrick, 1987a:55.
Camtarus.?Packard, 1888:156 [erroneous spelling].
Oreonectes.?Ldnnberg, 1894:126 [erroneous spelling].
Cambrus.?Price, 1900:155 [erroneous spelling].
44 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Orconectis.?Harris, 1903:113 [erroneous spelling].
Cityphobius.?Joleaud, 1939:14 [lapsus for C. typhlobius].
Subgenus Faxonius.?Rhoades, 1941b:142 [in part].
G[ambarus}.?Ctoizal, 1958:908 [erroneous spelling].
Oreonectes.?Hobbs III and Welch, 1972:66 [erroneous spelling].
Oronectes.?Hobbs III and Wells, 1972:125 [erroneous spelling].
Orconectes (Orconectes) australis australis (Rhoades)
FIGURE 134
Cambarus (Faxonius) pellucidus australis Rhoades, 1941b: 142, fig. 35a-f.
Orconectes pellucidus australis.?Hobbs, 1942a:353; 1948a:20, fig. 6,
13.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) pellucidus australis.?Hobbs, 1942b: 154 [by impli-
cation].
Cambarus pellucidus australis.?Jeannel and Henrot, 1949:84.
Orconectes australis australis.?Barr, 1967:161.?Hobbs and Barr, 1972:3,
11, figs. 2,3,5o-v,6,7,8i-cc.?Hobbs, 1974b:26, fig. 97.
Orconectes Pellucidus.?Cooper and Poulson, 1968:130, fig. 8.
Orconectes australis.?Barr and Hoi singer, 1971:115 [in part].
Orconectes (Orconectes) australis australis.?Fitzpatrick, 1987a:57.
TYPES.?Holotype, allotype, and "morphotype," USNM
79363, 79364, 79365 (male I, female, male II); paratypes,
ANSP, OSM, Alabama Mus. Nat. Hist., Leslie HubrichL
TYPE LOCALITY.?Shelta Cavern (SE 74, NE 'A, Sec. 27, T.
3S, R. 1W), north of Huntsville, Madison County, Alafama.
RANGE.?"Jackson and Madison counties, Alabar i, north-
ward on the western edge of the Cumberland Plateau to
Fentress County, Tennessee, and Wayne County, Kentucky,
where it intergrades with Orconectes australis packardi"
(Hobbs, Hobbs, and Daniel, 1977:95).
HABITAT.?Subterranean streams and pools.
Orconectes (Orconectes) australis packardi Rhoades
FIGURE 135
Orconectes pellucidus packardi Rhoades, 1944a:121, fig. 3a-f.?Hobbs,
1948a:20,figs. 8,11.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) pellucidus.?Hobbs, 1959:890 [in part].
Orconectes pellucidus packardii.?Nicholas, 1960:133.
Orconectes australis packardi.?Barr, 1967:161.?Hobbs and Barr, 1972:4,
22, figs. 2,4,5a-n, 8a-h.?Hobbs, 1974b:27, fig. 98.
Orconectes (Orconectes) australis packardi.?Fitzpatrick, 1987a:57.
TYPES.?Holotype, allotype, and "morphotype," USNM
81310, 81312, 81311 (male I, female, male II); paratypes,
MCZ, USNM, OSM.
TYPE LOCALITY.?Cumberland Crystal Cave (Sloans Valley
Cave), Alpine, Pulaski County, Kentucky.
RANGE.?Subterranean waters of the Cumberland River
basin in southeastern Kentucky, intergrading with the nominate
subspecies in Wayne County, Kentucky, and Fentress County,
Tennessee.
HABITAT.?Subterranean streams and pools.
Orconectes (Orconectes) incomptus Hobbs and Barr
FIGURE 136
Orconectes incomptus Hobbs and Barr, 1972:32,fig. 9.?Hobbs, 1974b:31,fig.
99.?Hobbs, Hobbs, and Daniel, 1977:98, fig. 47.
Orconectes (Orconectes) incomptus.?Fitzpatrick, 1987a:57.
TYPES.?Holotype, allotype, and morphotype, USNM
130299, 130300, 130301 (male I, female, male II); paratypes,
USNM,H.H. Hobbs III.
TYPE LOCALITY.?Cherry Cave, 36?28' 09"N, 85?36'28"W,
Jackson County, Tennessee.
RANGE.?Known from only three localities, all in Jackson
County, Tennessee.
HABITAT.?Subterranean waters.
Orconectes (Orconectes) inermis inermis Cope
FIGURE 137
Orconectes inermis Cope, 1872:419.?Hobbs, 1948a:20, figs. 10,14.
Cambarus pellucidus inermis.?Faxon, 1885a:83.?Packard, 1888:41.
Cambarus pellucidus variety inermis.?Hay, 1893:284.
Oreonectes inermis.?Lonnberg, 1894:126 [erroneous spelling].
Cambarus inermis.?Faxon, 1898:647.
Orconectis inermis.?Harris, 1903:113 [erroneous spelling].
Orconectes (Orconectes) inermis.?Hobbs, 1942b: 154 [by implication].
Orconectes pellucidus inermis.?Rhoades, 1959:401 [in part].
Orconectes pellacious.?Anonymous, 1965:7360.
Orconectes inermis inermis.?Barr, 1967:161.?Hobbs and Barr, 1972:4,36,
figs. 1,10,12c-z, 13.?Hobbs, 1974b:31,fig. 100.
Oreonectes inermis inermis.?Hobbs III and Welch, 1972:66 [erroneous
spelling].
Oronectes inermis inermis.?Hobbs III and Wells, 1972:125 [erroneous
spelling].
Orconectes i. inernis.?Hobbs III and Wells, 1972:125 [erroneous spelling].
Oroconectes inermis inermis.?Hobbs III, 1975:278 [erroneous spelling].
Orconectes (Orconectes) inermis inermis.?Fitzpatrick, 1987a:57, fig. 7,8.
TYPES.?Not extant. Topotypes, USNM 131405 (male I,
female, male II).
TYPE LOCALITY.?Wyandotte Cave, Indiana. Actually Si-
bert's Well Cave, just below Wyandotte Cave, Crawford
County, Indiana, according to Hobbs (1942a:335).
RANGE.?From Hart County, Kentucky, northwestward to
Crawford and Harrison counties, Indiana, where it intergrades
with 0. (0.) inermis testii.
HABITAT.?Subterranean waters.
Orconectes (Orconectes) inermis testii (Hay)
FIGURE 138
Cambarus pellucidus var. testii Hay, 1891:148.
Cambarus pellucidus testii.?Hay, 1893:285.
Cambarus pellucidus testi.?Hay, 19O2a:233.
Cambarus (Faxonius) pellucidus testi.?Ortmann, 1931:64 [by implication].
Orconectes pellucidus testii.?Hobbs, 1942a:352 [by implication].
Orconectes (Orconectes) pellucidus testii.?Hobbs, 1942b: 154 [by implica-
tion].
Orconectes pellucidus inermis.?Rhoades, 1959:401 [in part].
Orconectes inermis testii.?Hobbs and Barr, 1972:2, 49, figs. 1, 11,
12a,b.?Hobbs, 1972b:79,148,fig. 60f,61e; 1974b:31,fig. 101.
Orconectes (Orconectes) inermis testii.?Fitzpatrick, 1987a:57.
TYPES.?Syntypes, USNM 17702 (2 male II, female), MCZ
7431 (male II, female).
NUMBER 480 45
TYPE LOCALITY.?Mayfield's Cave, SW 7-*, NE ?/4, SW 74,
Sec. 26, T. 9N, R. 2W, Monroe County, Indiana.
RANGE.?Monroe County, Indiana, intergrading immedi-
ately to the south with the typical subspecies.
HABITAT.?Subterranean waters.
Orconectes (Orconectes) pellucidus (Tellkampf)
FIGURE 139
Astacus pellucidus Tellkampf, 1844a:684; 1844b:383; 1845:85.
astacus pellucidus.?Tellkampf, 1845:93.
Astacus (Cambarus) pellucidus.?Erichson, 1846:95.
Cambarus pellucidus.?Girard, 1852:87.?Hagen, 1870:55. pi. 1: figs. 68-71,
pi. 3: fig. 148, pi. 6.
Cambarus ptlulcidus.?Hagen, 1870:106 [erroneous spelling].
Orconectes pellucidus.?Cope, 1872:409.?Hobbs and Barr, 1972:58, figs. 1,
14-16.?Hobbs, 1974b:37, fig. 102.?Hobbs, Hobbs, and Daniel, 1977:108,
fig. 50.
Cambarus typhlobius Joseph, 1880:202 [see Holthuis, 1964].
Cambarus coecus Joseph, 1881:237 [nomen nudum; see Holthuis, 1964].
Cambarus stygius Joseph, 1881:241 [nomen nudum]; 1882:12 [validation of
nomen nudum of 1881, becoming homonym of Cambarus stygius Bundy (=
Procambarus acutus acutus (Girard)); see Holthuis, 1964].
Cambarus (Orconectes) pellucidus.?Hovey, 1882:222 [in part].
Cambarus Stygius.?-Faxon, 1884:139.
Cambarus pellucidus pellucidus.?Packard, 1888:41 [by implication].
Camtarus (Orconectes) pellucidus Jorm inermis.?Packard, 1888:156 [errone-
ous spelling].
Astacus Cambarus Stigius.?Rate, 1890:9 [erroneous spelling].
Astacus Cambarus pellucidus.?Rate, 1890:9.
Cambrus pellucidus.?Price, 1900:155 [erroneous spelling].
Cambarus pelucidus.?Eigenmann, 1900:228 [erroneous spelling].
Cambarus (Faxonius) pellucidus.?Ortmann, 1905c:97.
Cityphlobius.?Joleaud, 1939:14 [lapsus forC. typhlobius].
Cambarus caecus.?Bouvier, 1940:68 [erroneous spelling].
Cambarus pellucidus pellucidus.? Rhoades, 1941b: 144.
Cambarus (Fexonius) pellucidus.?Rioja, 1941:194 [erroneous spelling].
Cambarus pelludicus.?Dearolf, 1942:50 [erroneous spelling].
Cambarus Pellucidus.?Jackson, 1942:4.
Orconectes pellucidus pellucidus Hobbs, 1942a:352 [by implication].?
Filzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) pellucidus pellucidus.?Hobbs, 1942b: 154 [by
implication].
Cambarus (Cambarus) pellucidus.?Balss, 1944:402.
Orconectes pellucidus pelluicdus.?Hobbs, 1948a:16 [erroneous spelling].
Orconectes pellucidus pellucidus pellucidus.?Hobbs, 1948a:19 [lapsus ca-
lami].
Cambarus Coecus.?Boa, 1950:25.
Gambarus typhlobius.?Croizat, 1958:908 [erroneous spelling].
Orconectes (Orconectes) pellucidus.?Hobbs, 1959:890.?Fitzpatrick,
1987a:57.
Orconectes pallucidus.?Elliott, 1986:5 [erroneous spelling].
TYPES.?Holotype, ZBM 1562 (male I).
TYPE LOCALITY.?Mammoth Cave, Edmonson County,
Kentucky.
RANGE.?Area extending "southwestward from Hart County
to Trigg County, Kentucky, and Montgomery County, Tennes-
see" (Hobbs and Barr, 1972:69).
Subgenus Procericambarus Fitzpatrick
Cambarus.?Hagen, 1870:65 [in part].
Subgenus Faxonius Ortmann, 1931:76 [in part].
Faxonius.?Creaser, 1933a:3 [in part].
Orconectes.?Hobbs, 1942a:352 [in part].
Subgenus Orconectes.?Hobbs, 1942b:154 [in part].
Subgenus Procericambarus Fitzpatrick, 1987a:57 [Type species, by original
designation, Cambarus forceps Faxon, 1884:133. Gender: masculine.]
Orconectes (Procericambarus) acares Fitzpatrick
FIGURE 170
Orconectes leptogonopodus acares Fitzpatrick, 1965:87, figs. A-J.
Orconectes acares.?Hobbs, 1972b:94,fig. 74f; 1974b:26. fig. 131
Orconectes (Procericambarus) acares.?Fitzpatrick, 1987a:58.
TYPES.?Holotype, allotype, and morphotype, USNM
115517, 115518, 115519 (male I, female, male II); paratypes,
USNM,MCZ,TU.
TYPE LOCALITY.?Stream tributary to Ouachita River, 6
miles (9.6 km) northwest of Mount Ida, Montgomery County,
Arkansas.
RANGE.?Tributaries of the Ouachita River in Garland, Hot
Spring, Montgomery, Perry, Pike, Polk, and Saline counties,
Arkansas.
HABITAT.?Streams.
Orconectes (Procericambarus) barrenensis Rhoades
FIGURE 178
Orconectes rusticus barrenensis Rhoades, 1944a: 125, fig. 6a-f.?Fitzpatrick,
1963:61 [by implication].
Orconectes (Orconectes) rusticus.?Hobbs, 1959:893 [in part].
Faxonius rusticus barrenensis.?Creaser, 1962:2 [by implication].
Orconectes rusticus barrensis.?Fitzpatrick, 1967a: 143 [erroneous spelling].
Orconectes barrenensis.?Hobbs, 1972b:97, figs. 74h,76b; 1974b:27, fig. 139.
Orconectes (Procericambarus) barrenensis.?Fitzpatrick, 1987a:58.
TYPES.?Holotype, allotype, and "morphotype" (designated
as allotype by Rhoades), USNM 81319,81321,81320 (male I,
female, male II); paratypes, CM, MCZ, USNM, OSM.
TYPE LOCALITY.?Barren River, Beech Bend, 2 miles (3.2
km) north of Bowling Green, Warren County, Kentucky.
RANGE.?Barren River drainage in Kentucky and Tennes-
see, and Green River system in Kentucky.
HABITAT.?Swift streams.
Orconectes (Procericambarus) forceps (Faxon)
FIGURE 176
Cambarus forceps Faxon, 1884:133; 1885a:119,pL 5: fig. 4, pi. 9: figs. 5,5',
5a, 5a'.
Cambarus (Faxonius) forceps.?Ortmann, 1905c: 112.
Cambarus rusticus forceps.?Ortmann, 1931:66.
Cambarus (Faxonius) rusticus forceps.?Ortmann, 1931:72.
Faxonius forceps.?Creaser, 1933a:3 [by implication]; 1962:2 [by implica-
tion].
Faxonius (Faxonius) forceps.?Creaser, 1933b: 10 [by implication].
Orconectes rusticus forceps.?Hobbs, 1942a:352 [by implication].?
Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) rusticus forceps.?Hobbs, 1942b: 154 [by implica-
tion].
46 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Orconectes forceps.?Fitzpatrick, 1967t:143.?Hobbs, 1972b:98, figs. 77i.
80d; 1974b:29,fig. 137; 1981:289, figs. 19c, 107b, 108c, 112,113,232.
Orconectes (Procericambarus) forceps.?Fitzpatrick, 1987a:58, fig. 9.
TYPES.?Syntypes, USNM 4880 (3 male 1,4 female), MCZ
3582 (male I, female).
TYPE LOCALITY.?Cypress Creek, Lauderdale County, Ala-
bama.
RANGE.?Tennessee River drainage, chiefly above Walden
Ridge, from southwestern Virginia to Wayne County, Tennes-
see.
HABITAT.?Streams.
Orconectes (Procericambarus) hylas (Faxon)
FlOURE 172
Cambarus hylas Faxon, 1890:632.
Cambarus (Faxonius) kylas.?Oitmaim, 1905c:l 12.
Faxonius hylas.?Creaser, 1933a:3 [by implication]; 1962:2 [by implication].
Faxonius (Faxonius) hylas.?Creaser, 1933b:21 [by implication].
Orconectes hylas.?Hobbs, 1942a:352 [by implication]; 1974b:30, fig.
134?Williams, 1954:847, figs. 74-82.?Fitzpatrick, 1963:61 [by implica-
tion].?1987a:25.
Orconectes (Orconectes) hylas.?Hobbs, 1942b: 154 [by implication].
Orconectes (Procericambarus) hylas.?Fitzpatrick, 1987a:58.
TYPES.?Syntypes, MCZ 3858 (3 male II, 1 female), USNM
25827 (male II, female).
TYPE LOCALITY.?West Fork of Black River, Reynolds
County, Missouri.
RANGE.?Tributaries of the Big and Black rivers in
southeastern Missouri.
HABITAT.?Streams.
Orconectes (Procericambarus) leptogonopodus Hobbs
FIGURE 169
Orconectes leptogonopodus Hobbs, 1948b:146, figs. 24-32; 1972b:87, figs.
68a,e, 69b; 1974b:32, fig. 131.?Williams, 1954:854, figs. 18, 99-106.?
Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) leptogonopodus.?Hobbs, 1959:892.
Faxonius leptogonopodus.?Creaser, 1962:2 [by implication].
Orconectes leptogonopodus leptogonopodus.?Fitzpatrick, 1965:87.?Reimer,
1969:55. fig. 27.
Orconectes leptogonapodus.?May den, 1985:198 [erroneous spelling].
Orconectes (Procericambarus) leptogonopodus.?Fitzpatrick, 1987a:58.
TYPES.?Holotype, allotype, and "morphotype," USNM
82262 (male I, female, male II); paratypes, USNM, MCZ, TU.
TYPE LOCALITY.?McKinney's Creek, 4.7 miles (7.5 km)
northeast of Hatfield.Polk County, Arkansas,on U.S. Highway
71.
RANGE.?Red River drainage in Arkansas and eastern
Oklahoma, Ouachita Mountain Province.
HABITAT.?Small, clear, rocky streams.
Orconectes (Procericambarus) longidigitus (Faxon)
FIGURE 181
Cambarus longidigitus Faxon, 1898:653, pL 62: figs. 6-9.
Cambarus whitman! Steele, 1902:24, pi 3: figs, cl.c2.pt. 5: fig. b. [Location
of types unknown. Type locality, James River, Missouri.]
Cambarus (Faxonius) longidigitus.?Ortmann. 19O5c:l 13.
Faxonius longidigitus.?Creaser. 1933a:3 [by implication]; 1962:2 [by
implication].
Faxonius (Faxonius) longidigitus.?Creaser, 1933b 21 [by implication].
Orconectes longidigitus.?Hobbs. 1942a:352 [by implication]; 1974b:33,fig.
142.?Williams, 1954:884. figs. 149. 183-190?Pflieger. 1987a26;
1987b: 10.
Orconectes (Orconectes) longidigitus.?Hobbt, 1942b: 154 [by implication).
Orconectes (Procericambarus) longidigUus.?VwipUndt, 1987a:58.
TYPES.?Syntypes, MCZ 4364 (2 male II, female, 2 juv.
female), USNM 19683 (male II, 3 female, juv. male, 2 juv.
female).
TYPE LOCALITY.?Oxford Bend, White River (Izard
County?), Arkansas.
RANGE.?Tributaries of the White and Little Red rivers in
Missouri and Arkansas.
HABITAT.?Streams.
Orconectes (Procericambarus) luteus (Creaser)
nounitt
Faxonius (Faxonius) luteus Creaser, 1933b:7, pi. 1: figs. 3, 4, (subgenus by
implication].
Faxonius luteus?Creaser, 1934a: I; 1962:2 (by implication!.
Orconectes luteus?HobN. 1942a:353; 1974b:33. fig. 150.?Williams,
1954:872,figt. 149-157.?Rlzpatnck, 196361 [by implication].?Pflieger.
1987a:27; 1987b: 11.
Orconectes (Orconectes) luteus.?Hobba, 1942b: 154 (by implication].
Orconectes luelus.?Fitzpatrick, I967a:143 [erroneous spelling).
Orconectes (Procericambarus) luteus.?Fitzpalrick, I987a:58.
TYPES.?Holotype and allotvpc. USNM 98344,98345 (male
I, female); paratypes, USNM.
TYPE LOCALITY.?Niangua River at mouth of Greasy Creek,
5 miles (8 km) southeast of Buffalo, Dallas County, Missouri.
RANGE.?From the Marais des Cygnes River, Kansas,
eastward through southern Missouri and into Lawrence
County, Arkansas.
HABITAT.?Clear, swiftly flowing streams.
Orconectes (Procericambarus) macrus Williams
FIGURE 184
Orconectes nana macrus Williams. 1952:337. figs. 17-24; 1954:851. figs. 2,
91-98.?Fitzpatrick, 1963:61 [by implication].?Hayes. 1973:111.
Orconectes (Orconectes) nana macrus.?Hobbs, 1959:892 [by implication].
Faxonius nana macrus.?Creaser, 1962:2 [by implication].
Orconectes macrus.?Hobbs, 1972b.-93. fig. 74e; 1974b:33. fig. 145.?
Pflieger, 1987a:28; 1987b:12.
Orconectes (Procericambarus) macrus.?Fitzpatrick, 1987a:58.
TYPES.?Holotype, allotype, and "morphotype," USNM
118847 (male I, female, male II); paratypes, USNM, MCZ.
TYPE LOCALITY.?Spring River, 2 miles (3.2 km) southwest
of Mount Vernon, Lawrence County, Missouri.
RANGE.?Upper Arkansas River system in southwestern
NX'MBER 480 47
Missouri and northwestern Arkansas.
HABITAT.?Streams and shallow burrows along them.
REMARKS.?Hayes (1973:111) reported what he interpreted
to be intergrade populations between 0. (P.) nana and O. (P.)
macrus in Delaware and Ottawa counties, Oklahoma, but in his
recent revision of the genus, Fitzpatrick (1987a: 58) maintained
specific status for these crayfishes.
Orconectes (Procericambarus) medius (Faxon)
FIGURE 187
Cambarus medius Faxon, 1884:121; 1885a:107,pl. 3: fig. 4,pl. 9: figs. 4,4'.
Cambarus (Faxonius) medius.?Ortmann, 19O5c:l 12.
Faxonius medius.?Creaser, 1933a:3 [by implication]; 1962:2 [by implication].
Faxonius (Faxonius) medius.?Crcaser, 1933b:21 [by implication].
Orconectes medius.?Hobbs, 1942a:352 [by implication]; 1974b:34, fig.
149? Williams, 1954:876, figs. 158-166?Fitzpatrick, 1963:61 [by impli-
cation].? Pflicger, 1987a:27; 1987b:ll.
Orconectes (Orconectes) medius.?Hobbs, 1942b:154 [by implication].
Orconectes (Procericambarus) medius.?Filzpalrick, 1987a:58.
TYPKS.?Syntypcs, MCZ 3585 (male I, female).
TYPE LOCALITY.?Irondalc, Washington County, Missouri.
RANGE;.?Headwaters of the Big and Meramec rivers in
eastern Missouri.
HABITAT.?Streams.
Orconectes (Procericambarus) menae (Creaser)
FIGURE 189
Faxonius (Faxonius) menae Crcaser, 1933b:5, pi. 1: figs. 9,10 [subgenus by
implication].
Orconectes menae.? Hobbs, 1942a:353; Hobbs, 1974b:24, fig. 151.?
Williams, 1954:863, figs. 123-131.?Filzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) menae.?Hobbs, 1942b:154 [by implication].
Faxonius menae.?Creaser, 1962:2 [by implication].
Orconectes (Procericambarus) menae.?Fitzpatrick, 1987a:58.
TYPES.?Holotype and allotype, USNM 98330,98331 (male
I, female); paratypes, USNM, MCZ, FSBC.
TYPE LOCALITY.?Stream tributary to Irons Fork of the
Ouachita River, Mena, Polk County, Arkansas.
RANGE.?Tributaries of the Ouachita River in Polk and
Montgomery counties, Arkansas, and of the Red River in
LeFlore and McCurtain counties, Oklahoma.
HABITAT.?Rocky creeks.
Orconectes (Procericambarus) mirus (Ortmann)
FIGURE 182
Cambarus (Faxonius) rustic us mirus Ortmann, 1931:81.
Faxonius (Faxonius) mirus.?Crcaser, 1933b: 10 [by implication].
Orconectes rusticus mirus.?Hobbs, 1942a:352 [by implication].?Filzpatrick,
1963:61 [by implication].
Orconectes (Orconecles) rusticus mirus.?Hobbs, 1942b: 154 [by implication].
Faxonius mirus.?Creaser, 1962:2 [by implication].
Orconectes mirus.?Hobbs, 1972b:97,figs. 74i,76c,e; 1974b:35, fig. 143.
Orconectes (Procericambarus) mirus.?Fitzpatrick, 1987a:58.
TYPES.?Syntypes, CM 74.1378 (male 1,3 male II, female).
TYPE LOCALITY.?Hurricane Creek, Cumberland Springs,
Moore County, Tennessee.
RANGE.?Northern tributaries of the westward flowing
segment of the Tennessee River in Alabama and Tennessee,
and the Duck and Harpeth rivers in Tennessee.
HABITAT.?Streams.
Orconectes (Procericambarus) nana Williams
FIGURE 183
Orconectes nana nana Williams, 1952:333, figs. 9-16; 1954:849, figs.
82-90.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) nana nana.?Hobbs, 1959:892 [by implication].
Faxonius nana nana.?Creaser, 1962:2 [by implication].
Orconectes nana.?Hobbs, 1972b:96, figs. 74g,76a; 1974b:35, fig. 144.
Orconectes (Procericambarus) nana.?Fitzpatrick, 1987a:58.
TYPES.?Holotype, allotype, and "morphotype," USNM
129824, 129825, 129826 (male I, female, male II); paratypes,
USNM, MCZ.
TYPE LOCALITY.?Flint Creek, 5 miles (8 km) east of
Kansas, Delaware County, Oklahoma.
RANGE.?Neosho River basin in Arkansas and eastern
Oklahoma.
HABITAT.?Rocky streams.
REMARKS.?See "Remarks" under O. (P.) macrus.
Orconectes (Procericambarus) neglectus chaenodactylus
Williams
FIGURE 179
Orconectes neglectus chaenodactylus Williams, 1952:344, figs. 33-40;
1954:869, figs. 140-148.?Fitzpatrick, 1963:61 [by implication].?Hobbs,
1974b:36, fig. 140.
Orconectes (Orconectes) neglectus chaenodactylus.?Hobbs, 1959:893 [by
implication].
Faxonius neglectus chaenodactylus.?Creaser, 1962:2 [by implication].
Onconectes neglectus chaenodactylus.?Price and Payne, 1977:79 [erroneous
spelling].
Orconectes neglectus chaenodaytylus.?Morales, Bozada, and Casanova,
1987:175 [erroneous spelling].
Orconectes (Procericambarus) neglectus chaenodactylus.?Fitzpatrick,
1987a:58.
TYPES.?Holotype, allotype, and "morphotype," USNM
118840 (male I, female, male II); paratypes, USNM, MCZ.
TYPE LOCALITY.?Whites Creek, 8.6 miles (13.7 km)
southeast of Ava, Douglas County, Missouri.
RANGE.?North Fork of the White River in Arkansas and
Missouri, and as far south as Sylamore Creek in Stone County,
Arkansas; intergrading in the headwaters with the nominate
subspecies.
HABITAT.?Streams.
Orconectes (Procericambarus) neglectus neglectus (Faxon)
FIGURE 180
Cambarus neglectus Faxon, 1885b: 142.
Cambarus (Faxonius) neglectus.?Ortmann, 19O5c:112.?Creaser and
48 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Ortenburger, 1933:17, figs. 9,15,23.
Faxonius neglectus.?Cntser, 1933?:3 [by implication].
Faxonius (Faxonius) neglectus.?Creaser, 1933b:9 [by implication].
Cambarus negfccla.?Parks, Smith, and Garrett, 1939:2 [erroneous spelling].
Orconectes neglectus.?Hobbs, 1942a:352 [by implication].?Pflieger,
1987a:25; 1987b:9.
Orconectes (Orconectes) neglectus.?Hobbs, 1942b: 154 [by implication].
Orconectes neglectus neglectus.?Williams, 1952:344 [by implication];
1954:866, figs. 132-139, 148.?Fitzpatrick. 1963:61 [by implication].?
Hobbs, 1974b:36,fig. 141.?Unger, 1978:7, figs. 9-13,29.
Faxonius neglectus neglectus.?Creaser, 1962:2 [by implication].
Orconectes transfuga Fitzpatrick, 1966b: 178, figs. A-T. [Types: holotype and
allotype, USNM 114763 (male I, female); morphotype, USNM 114762
(male II); paratypes, MCZ, USNM. Type locality: Rogue River, 6 miles (9.6
km) upstream from Grants Pass, Jackson County, Oregon. (See Bouchard,
1977a:38 and 1978c:412,or 1980a:412).]
Orconectes transfugus.?Fitzpatrick, 1967a:143 [erroneous spelling].
Orconectes (Procericambarus) neglectus neglectus.?Fitzpatrick, 1987a:58.
Orconectes (Procericambarus) transfuga.?Fitzpatrick, 1987a:58.
TYPES.?Syntypes, MCZ 3757 (male I, female), MCZ 3590
(male II, questionable type), Mus. Washburn Coll., no longer
there.
TYPE LOCALITY.?Mill Creek, Wabaunsee County, Kansas.
RANGE.?Streams in the White (except North Fork) and
Arkansas river systems in Missouri, Oklahoma, and Arkansas,
and in tributaries of the Kansas River in Colorado, Nebraska,
and east central Kansas. Introduced into the Rogue River basin
in Oregon (Bouchard, 1978c:412).
HABITAT.?Rocky streams.
Orconectes (Procericambarus) ozarkae Williams
FlOURE 190
Orconectes ozarkae Williams, 1952:339, figs. 25-32; 1954:860, figs.
115-123.?Fitzpatrick, 1963:61 [by implication].?Hobbs, 1974b:36, fig.
153.?Pflieger, 1987a:25; 1987b:9.
Orconectes (Orconectes) ozarkae.?Hobbs, 1959:892.
Faxonius ozarkae.?Creaser, 1962:2 [by implication].
Orconectes (Procericambarus) ozarkae.?Fitzpatrick, 1987a:58.
TYPES.?Holotype, allotype, USNM 118846 (male I, fe-
male) and "morphotype," USNM 118844 (male II); paratypes,
USNM, MCZ.
TYPE LOCALITY.?Hackney Creek, 4.6 miles (7.3 km)
west-southwest of Stuart, Sharp County, Arkansas.
RANGE.?White and Black river systems in southern
Missouri and northern Arkansas.
HABITAT.?Streams.
Orconectes (Procericambarus) peruncus (Creaser)
FIGURE 173
Cambarus (Faxonius) peruncus Creaser, 1931a:7, figs. 13-17.
Faxonius (Faxonius) peruncus.?Crcaser, 1933b: 12 [by implication].
Faxonius peruncus.?Crcaser. 1934a:l; 1962:2 [by implication].
Orconectes peruncus.?Hobbs, 1942a:353; 1974b:38. fig. 135.?Williams,
1954:845, figs. 18, 66-73.?Fitzpatrick, 1963:61 [by implication].?
Pflieger, 1987a:26; 1987b:10.
Orconectes (Orconectes) peruncus.?Hobbs, 1942b:154 [by implication].
Orconectes (Procericambarus) peruncus.?Fitzpatrick, 1987a:58.
TYPES.?Holotype and allotype, USNM 98339,98340 (male
I, female); paratypes, USNM.
TYPE LOCALITY.?Little Creek, tributary to St. Francis
River, 1 mile (1.6 km) northeast of Chloride, Iron County,
Missouri.
RANGE.?Headwaters of the St. Francis River in central
Missouri.
HABITAT.?Rocky streams.
Orconectes (Procericambarus) placidus (Hagen)
FIGURE 177
Cambarus placidus Hagen, 1870:65,pl. 1: figs. 76-79.pl. 3: fig. 158.
Cambarus rusticus, var. placida.?Faxon, 1885a:65.
Cambarus rusticus, var. placidus.?Faxon, 1885b: 142.
Cambarus rusticus placidus.?Faxon, 1890:630.
Cambarus (Faxonius) rusticus placidus.?Ortmann, 1931:76.
Faxonius placidus.?Creaser, 1933a:3, [by implication]; 1962:2 [by implica-
tion].
Faxonius (Faxonius) placidus.?Creaser, 1933b: 10 [by implication].
Cambarus rustics placidus.?Fleming, 1938:302 [erroneous spelling].
Orconectes rusticus placidus.?Hobbs, 1942a:352 [by implication].?
Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) rusticus placidus.?Hobbs, 1942b: 154 [by implica-
tion].
Orconectes placidus.?Hobbs, 1966b:69; 1974b:38, fig. 138.?Page,
1985b:4O4,figs. 136-140.
Orconectes (Procericambarus) placidus.?Fitzpatrick, 1987a:58.
TYPES.?Syntypes, MCZ 170 (3 male 1,1 male II), MCZ 289
(4 male II, 3 female),MCZ 296 (male I),USNM 4966 (male II),
MHNP (male II, 3 female), Wurzburg Museum (?, see Faxon,
1914:418), AMS.
TYPE LOCALITY.?Lebanon, Wilson County, Tennessee.
RANGE.?Lower Ohio, Cumberland, Duck, and Tennessee
drainage systems in southern Illinois, Kentucky, Tennessee,
and northern Alabama.
HABITAT.?Streams.
Orconectes (Procericambarus) punctimanus (Creaser)
FIGURE 171
Faxonius (Faxonius) punctimanus Creaser, 1933b: 1, pi. 1: figs. 5,6 [subgenus
by implication].
Faxonius punctimanus.?Creaser, 1934a:l; 1962:2 [by implication].
Orconectes punctimanus.?Hobbs, 1942a:353; 1974b:39,fig. 133.?Williams,
1954:856, figs. 107-114, 200.?Fitzpatrick, 1963:61 [by implication].?
Pflieger, 1987a:27; 1987b:ll.
Orconectes (Orconectes) punctimanus.?Hobbs, 1942b: 154 [by implication].
Orconectes (Procericambarus) punctimanus.?Fitzpatrick, 1987a:58.
TYPES.?Holotype and allotype, USNM 98342,98343 (male
I, female); paratypes, USNM, MCZ.
TYPE LOCALITY.?Rubidoux Creek at Waynesville, Pulaski
County, Missouri.
RANGE.?Between the White, St. Francis, and Missouri
rivers in Arkansas and Missouri.
HABITAT.?Streams.
NUMBER 480 49
Orconectes (Procericambarus) putnami (Faxon)
FIGURE 186
Cambarus Putnami Faxon, 1884:131; 1885a: 118,pi. 5: fig. 5, pi. 9: figs. 6,6',
6a, 6a'.
Cambarus putnami.?Underwood, 1886:372.
Cambarus (Faxonius) putnami.?Ortmann, 19O5c:l 12.
Orconectes putnami.?-Rhoades, 1944a:129.?Hobbs, 1972b:86, figs. 68d,
69a;1974b:39,fig. 148.
Orconectes (Procericambarus) putnami.?Fitzpatrick, 1987a:58.
TYPES.?Syntypes, MCZ 3568 (4 male I, 7 female, 8 juv.
male, 5 juv. female), MCZ 3569 (13 male I), MCZ 3570 (3
male I), USNM 10130 (male I), ZIAS 1/44971 (male, 2
female).
TYPE LOCALITY.?Bear Creek, Grayson Springs, Grayson
County, Kentucky.
RANGE.?Ohio drainage system in western Kentucky,
southern Indiana(?),and Tennessee(?).
HABITAT.?Streams.
REMARKS.?Following Ortmann (1931), this species has
been largely relegated to synonymy with O. juvenilis (Hagen)
(= O. (P.) rusticus). Even Rhoades, in using the combination
Orconectes putnami for the first time, considered it a synonym
of 0. juvenilis. Not until 1972 was it proposed that it again be
accorded specific status, but its range and that of the closely
allied O. (P.) spinosus have not been adequately investigated.
Orconectes (Procericambarus) quadruncus (Creaser)
FIGURE 168
Faxonius (Faxonius) quadruncus Creaser, 1933b: 10, pi. 1: figs. 11, 12
[subgenus by implication].
Faxonius quadruncus.?Creaser, 1934a:l; 1962:2 [by implication].
Orconectes quadruncus.?Hobbs, 1942a:353; 1974b:39, fig. 130.?Williams,
1954:844, figs. 1,58-65.?Fitzpartick, 1963:61 [by implication].?Pflieger,
1987a:26; 1987b: 10.
Orconectes (Orconectes) quadruncus.?Hobbs, 1942b: 154 [by implication).
Orconectes (Procericambarus) quadruncus.?Fit2patrick, 1987a:58.
TYPES.?Holotype and allotype, USNM 98346,98347 (male
I, female); paratypes.USNM, MCZ.
TYPE LOCALITY.?Stout's Creek, tributary to St. Francis
River, between Ironton and Arcadia, Iron County, Missouri.
RANGE.?Restricted to the headwaters of the St. Francis
River system in Iron, Genevieve, and Madison counties,
Missouri.
HABITAT.?Rapidly flowing, rocky streams.
Orconectes (Procericambarus) rusticus (Girard)
FIGURE 174
Cambarus rusticus Girard, 1852:88.?Faxon, 1885a:108, pi. 9: figs. 8, 8', 8a,
8a'.
Cambarus juvenilis Hagen, 1870:66. pL 1: figs. 29-33, pL 3: fig. 157.
[Syntypes, MCZ 3347 (26 male II, female, 8 juv. male), USNM 4967 (male
I), USNM 25830 (male I, female), MHNP (2 male, 2 female). Other types
cited by Faxon (1914:418) apparently no longer extant. Type locality,
Kentucky River, Little Hickman, Jessamine County, Kentucky.]
Cambarus juvenalis.?Harris, 1900:271 [erroneous spelling].
Cambarus (Faxonius) rusticus.?Ortmann, 19O5c:l 12.
Faxonius rusticus.?Williamson, 1907:753 [Faxonius intended as a subgeneric
name].?Creaser, 1933a:5.
Cambarus (Faxonius) rusticus rusticus.?Ortmann, 1931:82.
Cambarus (Faxonius) juvenilis.?Ortmann, 1931:84 [in part].
Faxonius juvenilis.?Creaser, 1933a:3 [by implication]; 1962:2 [by implica-
tion].
Faxonius (Faxonius) rusticus rusticus.?Creaser, 1933b:21 [by implication].
Orconectes rusticus rusticus.?Hobbs, 1942a:352 [by implication].?
Fitzpatrick, 1963:61 [by implication].
Orconectes juvenilis.?Hobbs, 1942a:352 [by implication]; 1974b:32, fig.
147.?Fitzpatrick, 1963:61 [by implication].
Orconectes (Orconectes) rusticus rusticus.?Hobbs, 1942b:154 [by implica-
tion].
Orconectes (Orconectes) juvenilis.?Hobbs, 1942b: 154 [by implication].
Orconectes rusticus.?Pennak, 1953:465.?Hobbs, 1972b:92, figs. 74c,75b,d;
1974b:4O,fig. 136.?Page, 1985b:412, figs. 145-147.
Orconnectes rusticus.?Threinen, 1958:3 [erroneous spelling].
Faxonius rusticus rusticus.?Creaser, 1962:2 [by implication].
Orconectes Juvenils.?Hobbs and Walton, 1966:136 [erroneous spelling in
title].
Orconectes juvenalis.?Merkle, 1969:228 [erroneous spelling].
Orcenectes juvenilis.?Hart and Hart, 1974:45 [erroneous spelling].
Cambarus vusticus.?Lake and Newcombe, 1975:212 [erroneous spelling].
Orconectus rusticus.?DuBois and Sharma, 1977:27 [erroneous spelling].
Procambarus rusticus.?Huner, 1978:4 [lapsus calami].
Cambarus rustrineus rustrineus.?Brodsky, 1981:49 [erroneous spelling].
Orconectes (Procericambarus) rusticus.?Fitzpatrick, 1987a:58.
TYPES.?"Types probably destroyed in the Chicago fire in
1871" (Faxon, 1914:418). Questionable types: USNM 4968
(male II), in very poor condition; ANSP 195 (dry female).
TYPE LOCALITY.?Ohio River at Cincinnati, Hamilton
County, Ohio.
RANGE.?Southern Ontario and Michigan to Kentucky and
Tennessee; presumably introduced in one or more of the
following: New England, Illinois, Iowa, Minnesota, New
Mexico, West Virginia, and Wisconsin. (See Page, 1985b:412;
Lawton, 1979:90.)
HABITAT.?Streams, ponds, and lakes.
Orconectes (Procericambarus) saxatilis Bouchard and
Bouchard
FIGURE 175
Orconectes saxatilis Bouchard and Bouchard, 1976a:439, fig. 1.
Orconectes (Procericambarus) saxatilis.?Fitzpatrick, 1987a:58.
TYPES.?Holotype, allotype, and morphotype, USNM
146577, 146578, 146579 (male I, female, male II); paratypes,
USNM, Raymond W. Bouchard.
TYPE LOCALITY.?Pigeon Creek at Oklahoma State High-
way 63, LaFlore County, Oklahoma.
RANGE.?Known only from the type locality, Red River
Basin.
HABITAT.?Stream.
50 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Orconectes (Procericambarus) spinosus (Bundy)
FIGURE 185
Cambarusspinosus Bundy, 1877:173.?Faxon, 1885a:115,pl. 9: figs. 7,7',7a,
7a'.
Cambarus (Faxonius) spinosus.?Ortmann, 1905c: 112.
Cambarus (Faxonius) juvenilis.?Ortmann, 1931:84 [in part].
Orconectes spinosus.?Hobbs, 1944:340; 1972b:87, figs. 68b.g, 69d;
1974b:41,fig. 146; 1981:293, figs. 19b, 107a, 108d, 113.114,233.
Orconectes spinous.?Fitzpatrick, 1983b: 190 [erroneous spelling].
Orconectes (Procericambarus) spinosus.?Fitzpatrick, 1987a:58.
TYPES.?Syntypes, MCZ 3540 (2 male II, 2 female), MCZ
3541, (male II, female), USNM 19779 (3 male II, 2 female).
TYPE LOCALITY.?"Etowah, Oostanaula, and Coosa Rivers,
in the vicinity of Rome, Georgia" (Bundy, 1877:174).
RANGE.?Coosa, Tennessee, and Kanawha(?) river systems
in Alabama, Georgia, eastern Tennessee, Virginia(?), and West
Virginia(?). The limits of the range of O. (P.) spinosus and the
closely related O. (P.) putnami have not been determined.
HABITAT.?Streams.
REMARKS.?Following Ortmann (1931), this species has
been largely relegated to synonomy with O. juvenilis (Hagen)
(= O. (P.) rusticus (Girard)). Not until 1972 was it again
accorded specific status.
Orconectes (Procericambarus) williamsi Fitzpatrick
FIGURE 191
Orconectes williamsi Fitzpatrick, 1966a:145, figs. 1-10.?Hobbs, 1974b:43,
fig. 154.
Orconectes (Procericambarus) williamsi.?Fitzpatrick, 1987a:58.
TYPES.?Holotype, allotype, and morphotype, USNM
115520,115521, 115522 (male I, female, male II); paratypes,
MCZ.OSM.
TYPE LOCALITY.?White River, 2.8 miles (4.5 km) east of
Pettigrew, Madison County, Arkansas.
RANGE.?Headwaters of the White River from Madison and
Washington counties, Arkansas, to Barry County, Missouri.
HABITAT.?Pools in small streams.
Subgenus Rhoadesius Fitzpatrick
Cambarus.?Bundy, 1876:24 [in part].
Subgenus Faxonius Ortmann, 1905c: 107 [in part].
Faxonius.?Crcaser, 1933a:3 [by implication; in part].
Orconectes.?Hobbs, 1942a:352 [in part].
Subgenus Orconectes.?Hobbs, 1942b: 154 [by implication; in part].
Subgenus Rhoadesius Fitzpatrick, 1987a:58. [Type species, by original
designation, Cambarus sloanii Bundy, 1876:24. Gender masculine.]
Orconectes (Rhoadesius) kentuckiensis Rhoades
FIGURE 151
Orconectes kentuckiensis Rhoades, 1944a: 122, fig. 4a-f.?Fitzpatrick,
1963:61 [by implication].?Hobbs, 1974b:32, fig. 113.?Page 1985b 396
figs. 128-131.
Orconectes (Orconectes) kentuckiensis.?Hobbs, 1959:894.
Faxonius kentuckiensis.?Creaser, 1962:2 [by implication].
Orconectes kentuckiens.?Magnuson et al., 1975:71 [erroneous spelling].
Orconectes (Rhoadesius) kentuckiensis.?Fitzpatrick, 1987a:60.
TYPES.?Holotype, allotype, and "morphotype" (designated
as allotype by Rhoades), USNM 81313,81315,81314 (male I,
female, male II); paratypes, MCZ, ANSP, OSM.
TYPE LOCALITY.?Piney Creek, 3 miles (4.8 km) west of
Shady Grove, Crittenden County, Kentucky.
RANGE.?Caldwell, Crittenden, Livingston, and Union
counties, Kentucky, and Hardin County, Illinois.
HABITAT.?Mud-bottomed streams.
Orconectes (Rhoadesius) sloanii (Bundy)
FIGURE 150
Cambarus sloanii Bundy, 1876:24.
Cambarus Sloanii.?Faxon, 1885a:89,pl. 4: fig. 5, pi. 10: figs. 1,1', la, la'.
Cambarus sloanei.?Ortmann, 1902:278 [erroneous spelling].
Cambarus (Faxonius) sloanei.?Ortmann, 1905c: 107 [erroneous spelling].
Faxonius sloanii.?Creaser, 1933a:3 [by implication]; 1962:2 [by implication].
Faxonius (Faxonius) sloanii.?Creaser, 1933b:21 [by implication].
Cambarus sloani.?Rhoades, 1941a:93.
Orconectes sloanii.?Hobbs, 1942a:352 [by implication]; 1974b:41, fig.
112.?Eberly, 1955:281.
Orconectes (Orconectes) sloanii.?Hobbs, 1942b:154 [by implication].
Orconectes sloani.?Hobbs, 1948a:20.
Orconectes sloanei.?Eberly, 1958:2 [erroneous spelling].
Orconectes (Orconectes) sloani.?Hobbs, 1959:894.
Orconectes sloanni.?Hobbs III, 1975:284 [erroneous spelling].
Orconectes (Rhoadesius) sloanii.?Fitzpatrick, 1987a:60, fig. 10.
TYPES.?Syntypes, MCZ 3806 (male I, female).
TYPE LOCALITY.?New Albany, Floyd County, Indiana.
RANGE.?Southern Indiana and southwestern Ohio.
HABITAT.?Streams.
Subgenus Tragulicambarus Fitzpatrick
Cambarus.?Hagen, 1870:59.
Subgenus Faxonius Ortmann, 19O5c:l 13 [in part].
Faxonius.?Perm, 1939:215.
Orconectes.?Hobbs, 1942a:352 [by implication; in part].
Subgenus Orconectes.?Hobbs, 1942b: 154 [by implication; in part].
Subgenus Tragulicambarus Fitzpatrick, 1987a:60. [Type species, by original
designation, Cambarus lancifer Hagen, 1870:59. Gender: masculine].
Orconectes (Tragulicambarus) lancifer (Hagen)
FIGURE 152
Cambarus lancifer Hagen, 1870:59, pi. 1: figs. 86,87, pi. 3: fig. 159.
Cambarus faxonii Meek, 1894:1042, figs. 1-4. [Syntypes, MCZ 4220 (4 male
II, 3 female), USNM 19331 (40 male II and 25 female), USNM 22425 (male
II, female). Type locality, St. Francis River at Greenway and Big Bay, Clay
County, Arkansas.]
Cambarus (Faxonius) lancifer.?Ortmann, 19O5c:l 13.
Faxonius lancifer.?Penn, 1939:215.
Orconectes lancifer.?Hobbs, 1942a:352 [by implication]; 1974b:32, fig.
114.?Fitzpatrick, 1963:61 [by implication].?Page, 1985b:4O0, figs. 131-
135.?Pflieger, 1987a:29; 1987b:13.
NUMBER 480 51
Orconectes (Orconectes) lancifer.?Hobbs, 1942b:154 [by implication].
Orcnoectes lancifer.?O'Brien, 1976:84 [erroneous spelling].
Orconectes (Tragulicambarus) lancifer.?Fitzpatrick, 1987a:60.
TYPE.?Holotype, MCZ 306 (male I).
TYPE LOCALITY.?"Root Pond, Miss." Rocky Ford near
Etta, Union County, Mississippi, fide Penn (1939:215).
RANGE.?Known from the coastal plain and Mississippi
River floodplain of eastern Texas, Louisiana, Arkansas,
Mississippi, Missouri, Tennessee, Kentucky, and extreme
southern Illinois.
HABITAT.?Sluggish streams and lentic situations.
Genus Procambarus Ortmann
Astacus.?Harlan, 1830:464 [not Fabricius, 1775].
Subgenus Cambarus Erichson, 1846:95 [in part]. [Type species,by subsequent
designation (Faxon, 1898:644), Astacus Bartonii Fabricius, 1798:407.
Gender: masculine.]
Cambarus.?Cinid, 1852:91 [in part].
Camborus.?Williamson, 1899:47 [erroneous spelling].
Subgenus Cambarus Ortmann, 1905c:96 [in part]. [Type species, by invalid
designation, Astacus Blandingii Harlan, 1830:464. Gender: masculine.]
Subgenus Procambarus Ortmann, 1905d:435, 437. [Type species, by subse-
quent designation (Fowler, 1912:340), Cambarus Digueti Bouvier,
1897:225. Gender masculine.]
Subgenus Paracambarus Ortmann, 1906a: 1. [Type species, by monotypy,
Cambarus (Paracambarus) paradoxus Ortmann, 1906a:3. Gender: mascu-
line. Proposed as subgenus of Cambarus, elevated to generic rank by Hobbs,
1942a:344, and reduced to subgenus of Procambarus by Hobbs, 1972a: 10.]
Subgenus Ortmannicus Fowler, 1912:341. [Type species, by original designa-
tion, Abacus Blandingii Harlan, 1830:464. Gender: masculine. Replacement
name for Ortmann's subgenus Cambarus. Utilized as generic name by
Rhoades, 1944a: 114, and as a subgenus of Procambarus by Hobbs,
1972a:9.]
Cambarellus.?Creaser, 1933b:21 [lapsus calami].
Subgenus Girardiella Lyle, 1938:76. [Type species, by monotypy, Cambarus
Hagenianus Faxon, 1884:141. Gender: feminine.]
Cambaru.?Goodnight, 1941:72 [erroneous spelling].
Procambarus.?Hobbs, 1942a:341 [Ortmann's (19O5d) subgenus elevated to
generic rank]; 1984: fig. 1.
Ortmanmanicus.?Hobbs, 1942a:342 [erroneous spelling].
Cambaus.?Okada, 1948:133 [erroneous spelling].
Pracambarus.?Villalobos, 1953:352 [erroneous spelling].
Paracamburus.?Villalobos, 1953:354 [erroneous spelling].
Procambrus.?Suko, 1961:37 [erroneous spelling].
procambarus.?Padgett, 1970:19 [lapsus calami].
Subgenus Acucauda Hobbs, 1972a:5. [Type species, by original designation,
Procambarus fitzpatricki Hobbs, 1971a:461. Gender feminine.]
Subgenus Austrocambarus Hobbs, 1972a:5. [Type species, by original
designation, Procambarus vazquezae Villalobos, 1954b:328. Gender: mas-
culine.]
Subgenus Capillicambarus Hobbs, 1972a:6. [Type species, by original
designation, Cambarus (Cambarus) hinei Ortmann, 1905b:401. Gender:
masculine.]
Subgenus Hagenides Hobbs, 1972a:7. [Type species, by original designation,
Astacus advena LeConte, 1856:402. Gender masculine.]
Subgenus Leconticambarus Hobbs, 1972a:7. [Type species, by original
designation, Cambarus barbatus Faxon, 1890:621. Gender: masculine.]
Subgenus Lonnbergius Hobbs, 1972a:8. [Type species, by original designation,
Cambarus acherontis Lonnberg, 1894:125. Gender masculine.]
Subgenus Mexicambarus Hobbs, 1972a:8. [Type species, by original designa-
tion, Cambarus (Cambarus) bouvieri Ortmann, 1909:159. Gender: mascu-
line].
Subgenus Pennides Hobbs, 1972a:10. [Type species, by original designation,
Procambarus natchitochae Penn, 1953d:5. Gender: masculine].
Subgenus Remoticambarus Hobbs, 1972a: 11. [Type species, by original
designation,Procambarus pecki Hobbs, 1967b:2. Gender: masculine].
Subgenus Scapulicambarus Hobbs, 1972a: 11. [Type species, by original
designation, Cambarus clarkii paeninsulanus Faxon, 1914:369. Gender:
masculine].
Subgenus Tenuicambarus Hobbs, 1972a: 12. [Type species, by original
designation, Procambarus tenuis Hobbs, 1950b:194. Gender: masculine.]
Subgenus Villalobosus Hobbs, 1972a: 12. [Type species, by original designa-
tion, Paracambarus riojae [sic] Villalobos, 1944a:161. Gender: masculine.].
Procamborus.?Franzini-Armstrong, 1976:218 [erroneous spelling].
Procambararus.?Hobbs III, Thorp, and Anderson, 1976:12 [erroneous
spelling].
Procambasrus.?Huner, 1977:10 [erroneous spelling].
Procambaris.?Wharton, 1978:46 [erroneous spelling].
Procamburus.?Brown and Bowler, 1978:34 [erroneous spelling].
Combarus.?Shinozaki and Ishida, 1981:50 [erroneous spelling].
Procambarius.?Mauro, Thompson, and Melacha, 1983:938 [erroneous spell-
ing].
Procambambarus.?Appelberg, 1983:67 [erroneous spelling].
Procamberus.?Welcome, 1985:91 [erroneous spelling].
Subgenus Acucauda Hobbs
Cambarus.?Lyle, 1938:76 [in part].
Subgenus Paracambarus.?Lyle, 1938:76.
Procambarus.?Hobbs, 1972a:5 [in part].
Subgenus Acucauda Hobbs, 1972a:5 [Type species, by original designation,
Procambarus fitzpatricki Hobbs, 1971a:461. Gender: feminine.]; 1984, fig.
7.
Procambarus (Acucauda) fitzpatricki Hobbs
FIGURE 211
Cambarus (Paracambarus) harnedi Lyle, 1938:76 [nomen nudum].
Procambarus fitzpatricki Hobbs, 1971a:461,fig. 1.
Procambarus (Acucauda) fitzpatricki.?Hobbs, 1972a:5, figs. 11, 2a, 4a-g;
1974b:43,fig. 170.
TYPES.?Holotype, allotype, and morphotype, USNM
131205, 131206, 131207 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Roadside ditch at junction of State Route
67 and Woolmarket Road, just north of D'lberville, Harrison
County, Mississippi.
RANGE.?Southern Mississippi between the Wolf and
Pascagoula rivers.
HABITAT.?Burrows (primary or secondary burrower).
Subgenus Austrocambarus Hobbs
Astacus.?Erichson, 1846:87 [in part].
Cambarus.?Girard, 1852:87 [in part].
Subgenus Cambarus Ortmann, 1905c: 101 [in part].
Subgenus Procambarus Ortmann, 19O5d:438 [in part].
Procambarus.?Hobbs, 1942a:342 [in part].
Subgenus Austrocambarus Hobbs, 1972a:5 [Type species, by original
designation, Procambarus vazquezae Villalobos, 1954b:328. Gender mas-
culine.]; 1984, fig. 6.
52 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Procambarus (Austrocambarus) acanthophorus Villalobos
FIGURE 223
Procambarus acanthophorus Villalobos, 1948:175,0. 1: figs. 1-10,pi. 2: figs.
1-9; 1955:218. ph. 49,50; 1983:209, pis. 49,50.
Procambarus {Austrocambarus) acanthophorus.?Hobbs, 1972a:6; 1974b:43,
fig. 180.
Procambarus acantophorus.?Spitzy, 1976:443 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, IBM
90847(035) (male I, female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?Four kilometers (2.5 mi) east of Tuxte-
pec, Oaxaca, Mexico.
RANGE.?Atlantic drainage in southern Veracruz, Chiapas,
and Oaxaca, Mexico.
HABITAT.?Streams and canals.
Procambarus (Austrocambarus) atkinsoni (Ortmann)
FIGURE 226
Cambarus (Procambarus) atkinsoni Ortmann, 1913:414.
Procambarus atkinsoni.?Hobbs, 1942a:342 [by implication].?Hobbs and
Villalobos, 1964:346, figs. 40-50.
Procambarus Atkinsoni.?Creaser, 1962:7.
Cambarus atkinsoni.?Hobbs and Villalobos, 1964:323.
Procambarus (Austrocambarus) atkinsoni.?Hobbs, 1972a:6; 1974b:44, fig.
183.
TYPES.?Lectoholotype, lectoallotype, and lectomor-
photype, CM 74.924 (male I, female, male II); paralectotypes,
CM 74.924, 74.925. Designated by Hobbs and Villalobos,
1964:347,349,350.
TYPE LOCALITY.?Tributaries of Rio de los Indios, Isla de
Pinos, Cuba.
RANGE.?Isla de Pinos, Cuba.
HABITAT.?Streams (?).
Procambarus (Austrocambarus) cubensis cubensis
(Erichson)
FIGURE 228
Astacus Cubensis Erichson, 1846:87.
Astacus (Cambarus) Cubensis Erichson, 1846:100.
Cambarus Cubensis.?Guard, 1852:87 [by implication].?Faxon, 1885a:51,
pL 2: fig. L pL 8: figs. 5.5', 5a, 5a'.
Cambarus consobrinus Saussure, 1857a: 101. [Syntypes, GM (uncatalogued,
dry. 2 male), specimens formerly in MHNP no longer there. BMNH 1163 (2
female), USNM 20684 (male II). Type locality, ponds in the central part of
Cuba.]
Cambaus cubensis.?Huxley, 188Oa:3O5.
Cambarus (Cambarus) cubensis.?Ortmann, 1905c: 101.
Cambarus (Procambarus) cubensis.?Ortmann, 19O5d:438.
Astacus Cubensis.?VtXAis Ragues, 1909:180.
Cambarus cubensis cubensis.?Faxon, 1912:458 [by implication].
Cambarus cubensis consobrinus.?Faxon, 1912:458.
Cambarellus cubensis.?Creaser, 1933b:21 [lapsus calami].
Procambarus cubensis cubensis.?Hobbs, 1942a:342 [by implication].?
Hobbs and Villalobos, 1964:319, figs. 1-13.
Procambarus cubensis consobrinus.?Hobbs, 1942a:342 [by implication].
Procambarus consobrinus.?Villalobos, 1954b:303.
Procambarus C. cubensis.?Straskraba, 1969:18.
Astacus cubensis.?Hobbs, 1972a:l.
Procambarus (Austrocambarus) cubensis cubensis.?Hobbs, 1972a:6;
1974b:44,fig. 185.
TYPES.?BMNH 4140.
TYPE LOCALITY.?Cuba.
RANGE.?Scattered localities throughout Cuba except in the
southwestern mountains.
HABITAT.?Lentic and lotic situations.
Procambarus (Austrocambarus) cubensis rivalis (Faxon)
FIGURE 227
Cambarus cubensis rivalis Faxon, 1912:459.
Procambarus cubensis rivalis.?Hobbs, 1942a:342 [by implication].?Hobbs
and Villalobos, 1964:335, figs. 14-24.
Procambarus (Austrocambarus) cubensis rivalis.?Hobbs, 1972a:6; 1974b:44,
fig. 184.
TYPES.?Syntypes, MCZ 7406 (2 male II, 3 female).
TYPE LOCALITY.?San Diego de los Bafios, Province of
Pinar del Rio, Cuba.
RANGE.?Known only from a few localities in the province
of Pinar del Rio, Cuba.
HABITAT.?Mountain streams.
Procambarus (Austrocambarus) llamasi Villalobos
FIGURE 224
Procambarus llamasi Villalobos, 1954b:364,pls. 16,17; 1955:214,pls. 51,52;
1983:213,pls. 51,52.
Procambarus (Austrocambarus) llamasi.?Webbs, 1972a:6; 1974b:44, fig.
181.?Valenzuela, 1986:1, figs. 1-18.
Procambarus llamesi.?Covich, 1976:5 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, IBM (male I,
female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?Santa Rita, 47 kilometers (29.5 mi) east
of Escaicega, Campeche, Mexico.
RANGE.?Southern Veracruz to Yucatan, Mexico, and
northern Guatemala.
HABITAT.?Small streams and drainage ditches.
Procambarus (Austrocambarus) mexicanus (Erichson)
FIGURE 214
Astacus (Cambarus) mexicanus Erichson, 1846:99.
Astacus Mexicanus.?Erichson, 1846:87.
Astacus mexicanus.?Dana, 1852:522.
Cambarus aztecus Saussure, 1857b:503, fig. 23. [Type locality: "Pris a
Tomatlan, dans les Terres-Chaudes," Veracruz, Mexico. Syntypes: GM, 13
dry specimens; USNM 20682 (male I, dry).].?Hobbs, 1987:206, fig. 2.
Cambarus Mexicanus.?Saussure, 1858:460.
Cambarus mexicanus.?Saussure, 1858:461.
Cambarus Aztecus.?Hagen, 1870:11.
Astacus Aztecus.?von Martens, 1872:131.
Cambarus (Cambarus) mexicanus.?Ortmann, 1905c: 101.
Cambarus (Procambarus) mexicanus.?Ortmann, 1905d:437.
NUMBER 480 53
Astacus Astecus.?Vald6s Ragues, 1909:180 [erroneous spelling].
Procambarus aztecus.?Hobbs, 1942a:342 [by implication].
Procambarus mexicanus.?Hobbs, 1942a:342 [by implication].?Villalobos,
1954b:307,pls. 1,2; 1955:162,pis. 34,35; 1983:156,pis. 34,35.
Procambarus mexiacnus.?Villalobos, 1954b:3O4 [erroneous spelling].
Procambarus Mexicanus.?Creaser, 1962:7.
Procambarus (Austrocambarus) mexicanus.?Hobbs, 1972a:6; 1974b:45, fig.
173; 1987:200, fig. 1.
Procambarus (Austrocambarus) aztecus.?Hobbs, 1972a:6; 1972b [1976]:43,
figs. 30e,34c,36b; 1974b:44,fig. 172; 1987:201.
TYPES.?Lost, formerly in ZBM. Neoholotype, ANSP 421
(male I); neoallotype and neoparatypes, IBM (Villalobos,
1954b:312).
TYPE LOCALITY.?Mexico. Restricted by Villalobos
(1954b:312) to "El Mirador de Zacuapan," 8 kilometers (5 mi)
northeast of Huatusco, Veracruz.
RANGE.?Veracruz, Mexico, south of the Cordillera Vol-
canica Transversal.
HABITAT.?Lentic and lotic situations.
REMARKS.?A review of what is known about this crayfish
is reported by Hobbs (1987), but its range and limits of
variation are still poorly understood.
Procambarus (Austrocambarus) mirandai Villalobos
FIGURE 219
Procambarus mirandai Villalobos, 1954b:355, pis. 13-15; 1955:210, pis.
46-48; 1983:200, pis. 46-48.
Procambarus mirandae.?Hobbs and Villalobos, 1964:313 [erroneous spell-
ing].
Procambarus (Austrocambarus) mirandai.?Hobbs, 1972a:6; 1974b:45, fig.
176;1977b:198,fig. 1.
TYPES.?Holotype, allotype, and morphotype, IBM (male I,
female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?Cerro Hueco, 4 kilometers (2.5 mi)
southeast of Tuxtla Gutierrez, Chiapas, Mexico.
RANGE.?Known from only four localities in Chiapas.
HABITAT.?"Los ejemplares fueron colectados dentro de una
grata, que es la salida de un rio subterraneo" (Villalobos,
1954b:361).
Procambarus (Austrocambarus) niveus Hobbs and
Villalobos
FIOURE229
Procambarus niveus Hobbs and Villalobos, 1964:342, figs. 26-39.
Procambarus (Austrocambarus) niveus.?Hobbs, 1972a:6; 1974b:45, fig.
186.?Hobbs, Hobbs, and Daniel, 1977:114, fig. 51.
TYPES.?Holotype and allotype, USNM 109076, 109077
(male I, female).
TYPE LOCALITY.?"Cuevas de Santo Tomas, Sierra de los
Organos, cerca de Ponce, Pinar del Rio, Cuba" (Hobbs and
Villalobos, 1964:346).
RANGE.?Known only from the type locality.
HABITAT.?Subterranean waters.
Procambarus (Austrocambarus) oaxacae oaxacae Hobbs
FIGURE 216
Procambarus (Austrocambarus) oaxacae oaxacae Hobbs, 1973b:29, figs.
3-5.?Hobbs, Hobbs, and DanieL 1977:116, fig. 52.?Reddell, 1981:111,
fig. 16.
TYPES.?Holotype, allotype, and morphotype, USNM
144341, 144342, and 144343 (male I, female, male II);
paratypes, USNM, IBM, TTM.
TYPE LOCALITY.?Cueva del Guano, 10 kilometers (6.3 mi)
northeast of Valle Nacional, Oaxaca, Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Subterranean waters.
Procambarus (Austrocambarus) oaxacae reddelli Hobbs
FIGURE 217
Procambarus (Austrocambarus) oaxacae reddelli Hobbs, 1973b:33, figs.
6-8.?Hobbs, Hobbs, and Daniel, 1977:118, fig. 53.?Reddell, 1981:112,
fig. 16.
Procambarus oaxacae reddelli.?Hobbs, 1973c:73.
TYPES.?Holotype, allotype, and morphotype, USNM
144346, 144347, 144348 (male I, female, male II); paratypes,
USNM, IBM, TTM.
TYPE LOCALITY.?Cueva del Nacimiento del Rio San
Antonio, 10 kilometers (6.3 mi) south-southwest of Acatlan,
Oaxaca, Mexico.
RANGE.?Known from five caves in Oaxaca and Veracruz,
Mexico.
HABITAT.?Subterranean waters.
Procambarus (Austrocambarus) pilosimanus (Ortmann)
FIGURE 225
Cambarus (Procambarus) pilosimanus Ortmann, 1906a:6,fig. 2.
Cambarus pilosimanus.?Pearse, 1911:110.
Procambarus pilosimanus.?Hobbs, 1942a:342 [by implication].?Villalobos,
1955:231,pis. 53,54; 1983:220,pls. 53,54.
Procambarus pelosimanus.?Creaser, 1962:7 [erroneous spelling].
Procambarus (Austrocambarus) pilosimanus.?Hobbs, 1972a:6; 1974b:45,
fig. 182.
TYPES.?Syntypes, MHNP (8 male, 6 female), CM 74.678
(2 specimens).
TYPE LOCALITY.?"Coche, pres de la rivere de Coban
[Province of Alta Verapaz], Guatemala" (Ortmann, 1906a:9).
Probably Rio Cahabdn at Coban.
RANGE.?Guatemala, Belize, and Chiapas and Quintana
Roo, Mexico.
HABITAT.?Lentic, lotic(?), and subterranean habitats.
Procambarus (Austrocambarus) primaevus (Packard)
Cambarus primaevus Packard, 1880:222; 1881a:832; 1881b:391.
Cambarus primoevus.?Villalobos, 1955:14 [erroneous spelling].
Procambarus (Austrocambarus) primaevus.?Feldmann et al., 1981:794, pis.
2,3.
54 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
TYPES.?Neotype, AMNH 42675 (male).
TYPE LOCALITY.?Green River beds (Eocene) of western
Wyoming in "the fish beds of the Bear River Valley, situated
from 75 to 100 miles [120 to 160 km] a little west of north of
Evanston, Wyo., and near the Utah line" (Packard, 1881b:391).
RANGE.?Known only from the Green River beds (Eocene)
of Wyoming.
HABITAT.?Fossil lake.
Procambarus (Austrocambarus) rodriguezi Hobbs
FIGURE 215
Procambarus rodriguezi Hobbs, 1943b:203,pl. 2.
Paracambarus rodriguezi.?Rioja, 1953:293 [lapsus calami].
Procambarus (Austrocambarus) rodiguezi.?Hobbs, 1972a:6 [erroneous spell-
ing].
Procambarus (Austrocambarus) rodriguezi.?Hobbs, 1974b:45, fig. 174.?
Hobbs, Hobbs, and Daniel, 1977:118,fig. 54.-*eddell, 1981:112, fig. 16.
TYPES.?Holotype, allotype, and "morphotype," USNM
81302 (male I, female, male II); paratypes, USNM, MCZ.
TYPE LOCALITY.?Cueva de Ojo de Agua, west-northwest
of Hacienda Potrero Viejo, Paraje Nuevo, C6rdoba, Veracruz,
Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Subterranean stream.
REMARKS.?Reddell (1981:112) noted that
the type-locality of this species has been stated to be "Cueva del Agua" and
"Cueva del Ojo de Agua Grande," but the brief description of the type-locality
as given by Hobbs (1943[bJ) does not fit Cueva del Ojo de Agua Grande.
Furthermore, Cueva del Ojo de Agua Grande has no habitat suitable for
crayfish. It contains only a single passage and this contains a rapid rock-floored
stream. A single juvenile crayfish was seen trapped in a tiny drip pool in the
latter cave where flood water apparently isolated it. It must be assumed that the
type-locality has not been rediscovered.
Procambarus (Austrocambarus) ruthveni Pearse
FIGURE 220
Cambarus ru/Mvm'Pearse, 1911:110,fig. 3.
Procambarus ruthveni.?Hobbs, 1942a:342 [by implication]; 1954b:l, fig. 1.
Procambarus ruthveni ruthveni.?Villalobos, 1954b:336,pls. 9,10; 1955:191,
pis. 42,43; 1983:183, pis. 42,43 [by implication].
Procambarus (Austrocambarus) ruthveni ruthveni.?Hobbs, 1972a:6;
1974b:46,fig. 177.
TYPES.?Holotype and allotype, USNM 98348,98349 (male
I, female).
TYPE LOCALITY.?Cuatotolapan, CanuSn de Acayucan,
Veracruz, Mexico.
RANGE.?Known from only one additional locality: San
Juan Diaz Covarrubias, 10 kilometers (6.3 mi) southwest of
Cuatotolapan, Veracruz (Villalobos, 1954b:344).
HABITAT.?Temporary ponds, pools, and burrows (secon-
dary burrower).
REMARKS.?See "Remarks" under P. (A.) zapoapensis.
Procambarus (Austrocambarus) sbordonii Hobbs
FIGURE 213
Procambarus (Austrocambarus) sbordonii Hobbs, 1977b:201, fig. 2.
TYPES.?Holotype, USNM 146588 (male I).
TYPE LOCALITY.?Cueva del Nacimiento de Rio S. Do-
mingo, near the village of Bochil, Chiapas, Mexico, at an
altitude of 1250 m.
RANGE.?Known only from the type locality.
HABITAT.?Subterranean waters.
Procambarus (Austrocambarus) vazquezae Villalobos
FIGURE 212
Procambarus vazquezae Villalobos, 1954b:328, pis. 7, 8; 1955:184, pis. 40,41;
1983:176, pis. 40,41.
Procambarus vasquezae.?Hobbs, 1966b:71 [erroneous spelling].
Procambarus (Austrocambarus) vazquezae.?Hobbs, 1972a:6, figs. 2b, 5a-g;
1974b:46,fig. 171.
TYPES.?Holotype, allotype, and morphotype, IBM (male I,
female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?"Playa Norte dc la Laguna dc Catemaco"
(Villalobos, 1954b:355), Veracruz, Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Under rocks along the shore of Laguna de
Catemaco.
Procambarus (Austrocambarus) veracruzanus Villalobos
FIGURE 218
Procambarus veracruzanus Villalobos, 1954b:323,pls. 5,6; 1955:178,pis. 38,
39; 1983:171, pis. 38,39.
Procambarus (Austrocambarus) veracruzanus.?Hobbs, 1972a:6; 1974b:46,
fig. 175.
TYPES.?Holotype, allotype, and morphotype, IBM (male I,
female, male II); paratypes, IBM.
TYPE LOCALITY.?Presidio, 30 kilometers (18.8 mi) south-
east of C6rdoba, Veracruz, Mexico. "Cuenca del rio Papa-
loapan" (Villalobos, 1954b:328).
RANGE.?Known only from the vicinity of C6rdoba,
Veracruz.
HABITAT.?Streams.
Procambarus (Austrocambarus) williamsoni (Ortmann)
FIGURE 222
Cambarus (Procambarus) williamsoni Ortmann, 1905d.-439, figs. 1-3.
Procambarus williamsoni.?Hobbs, 1942a:342 [by implication].
Procambarus (Austrocambarus) williamsoni.?Hobbs, 1972a:6; 1974b:46, fig.
179.
TYPES.^Syntypes, CM 74.560 (male I, female, male II),
CM 74.561 (male I, female, male II).
TYPE LOCALITY.?LOS Amates, Province of Izabal, 6 to 8
kilometers (4 to 5 mi) south of Rio Motagua, Guatemala. (Not
NUMBER 480 55
"Malagua" as recorded by Ortmann, 1905d:440.)
RANGE.?Known only from the vicinity of the type locality
and from Lago Yojoa, Honduras.
HABITAT.?Lentic situations.
Procambarus (Austrocambarus) zapoapensis Villalobos, new
status
FIGURE 221
Procambarus ruthveni zapoapensis Villalobos, 1954b:347, pis. 11, 12;
1955:202,pls. 44,45; 1983:192,pls. 44,45.
Procambarus ruthveni zapaopensis Villalobos, 1955:203 [erroneous spelling].
Procambarus (Austrocambarus) ruthveni zapoapensis.?Hobbs, 1972a:6;
1974b:46,fig. 178.
TYPES.?Holotype, allotype, and morphotype, IBM (male I,
female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?Zapoapan de Cabafla, 11 kilometers (6.9
mi) southeast of Catemaco, Veracruz, Mexico.
RANGE.?Known only from the vicinity of the type locality.
HABITAT.?Small, rocky streams.
REMARKS.?In the absence of evidence of intergradation
between this crayfish and P. (A.) ruthveni, it is accorded
specific rank herein.
Subgenus Capillicambarus Hobbs
Cambarus.?Ortmann, 1905b:401 [in part].
Subgenus Cambarus Ortmann, 1905b: 401 [in part].
Subgenus Ortmannicus Fowler, 1912:341 [in part, by implication].
Procambarus.?Hobbs, 1942a:342 [in part].
Subgenus Capillicambarus Hobbs, 1972a:6 [Type species, by original
designation, Cambarus (Cambarus) hinei Ortmann, 1905b:401. Gender:
masculine.]; 1984, fig. 7.
Procambarus (Capillicambarus) brazoriensis Albaugh
FIGURE 231
Procambarus (Capillicambarus) brazoriensis Albaugh, 1975:1, figs. 1,2.
Procambarus (Capillicambarus) brazorensis.?Fitzpatrick, 1983b:203 [erro-
neous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
133916, 133917, 133918 (male I, female, male II); paratypes,
USNM, D.W. Albaugh.
TYPE LOCALITY.?Ditch along County Road 400,0.1 mile
(160 m) southeast of Missouri Pacific railroad and 0.25 mile
(0.4 km) southwest of Brazos River at south edge of Brazoria,
Brazoria County, Texas.
RANGE.?Brazoria County, Texas between the San Bernard
River and Oyster Creek, extending inland from the Gulf of
Mexico only about 20 miles (32 km).
HABITAT.?Lentic and sluggish lotic situations.
Procambarus (Capillicambarus) hinei (Ortmann)
FIGURE 230
Cambarus (Cambarus) hinei Ortmann, 1905b:401, fig. 1.
Cambarus heini.?Cary and Spaulding, 1909:11 [erroneous spelling].
Cambarus (Ortmannicus) hinei.?Fowler, 1912:341 [by implication].
Cambarus hinei.?Faxon, 1914:370.
Procambarus hinei.?Hobbs, 1942a:342 [by implication].?Penn, 1953b:63,
figs. 1-15.
Procambarus (Capillicambarus) hinei.?Hobbs, 1972a:6, figs. In, 2c, 6a-g;
1974b:46,fig. 187.?Albaugh, 1973:102, figs. 1,23-28.
Procambarus hineyi.?Huner, Meyers, and Avault, 1976:150 [erroneous
spelling].
TYPES.?Syntypes, OSM 443.1 (male I), OSM 443.2
(female), OSM 444.1 (female), OSM 444.2 (female), USNM
98355 (male I, female).
TYPE LOCALITY.?One-fourth mile (0.3 km) from Gulf
Beach, near Cameron, Cameron Parish, Louisiana.
RANGE.?Southern Louisiana and east of the Brazos River in
southeastern Texas.
HABITAT.?Pools and roadside ditches.
Procambarus (Capillicambarus) incilis Penn
FIGURE 232
Procambarus incilis Penn, 1962:222, figs. 1-11.
Procambarus (Capillicambarus) incilis.?Hobbs, 1972a:6; 1974b:47,fig. 188.
Procambarus incillis.?Albaugh, 1975:7 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
106794, 106795, 106796 (male I, female, male II); paratypes,
USNM,TU.
TYPE LOCALITY.?7.5 miles (12 km) east of Edna, Jackson
County, Texas.
RANGE.?Southeastern Texas in Jackson, Matagorda, and
Wharton counties.
HABITAT.?Roadside ditches.
Subgenus Girardiella Lyle
Cambarus.?Bundy, 1876:5 [in part].
Subgenus Cambarus.?Ortmann, 1905c:101 [in part].
Subgenus Ortmannicus.?Fowler, 1912:341 [by implication, in part].
Subgenus Girardiella Lyle, 1938:76 [Type species, by monotypy, Cambarus
Hagenianus Faxon, 1884:141. Gender feminine.].?Hobbs, 1972a:6; 1984,
fig. 6.?Fitzpatrick, 1978b:57. figs. 1-121.
Procambarus.?Hobbs, 1942a:342 [by implication, in part].
Procambarus (Girardiella) barbiger Fitzpatrick
FIGURE 246
Cambarus (Girardiella) hagenianus forestae Lyle, 1938:76 [nomen nudum].
Procambarus hagenianus.?Fitzpatrick, 1968b:37 [in part].
Procambarus (Girardiella) hagenianus.?Hobbs, 1972b:47 [in part].
Procambarus (Girardiella) sp. B.?Fitzpatrick, 1976a:385, figs, lb, 2b [by
implication].
Procambarus (Girardiella) barbiger Fitzpatrick, 1978b:69, figs. 37-52,
109-121.
Cambarus (Girardiella) hagenianus forrestae.?Fitzpatrick, 1978b:69 [errone-
ous spelling].
Pfrocambarus] h[agenianus] forrestae.?Fitzpatrick, 1978b:73 [erroneous
spelling].
P[rocambarus] barbiger.?Stewart, 1986:15, unnumbered fig.
56 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
TYPES.?Holotype and allotype, USNM 146258, 146259
(male I, female); paratypes, USNM.
TYPE LOCALITY.?Forest, Scott County, Mississippi.
RANGE.?Known only from the Jackson Prairie area, chiefly
in Newton and Scott counties, Mississippi.
HABITAT.?Burrows (primary burrower).
Procambarus (Girardiella) cometes Fitzpatriek
FIGURE 244
Procambarus (Girardiella) hagenianus.?Hobbs, 1972b:47 [in part].
Procambarus (Girardiella) sp. E.?Fitzpatriek, 1976a:385, fig. le, 2e [by
implication].
Procambarus (Girardiella) cometes Fitzpatriek, 1978b:74, figs. 53-71,
109-121.
Pfrocambarus] cometes.?Stewart, 1986:15.
TYPES.?Holotype, allotype, and morphotype, USNM
130227, 146260, 131280 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Field behind Luxury Mobile Homes (SW
V4 Sec. 3, T. 18N, R. 14E), Starkville, Oktibbeha County,
Mississippi.
RANGE.?Lowndes and Oktibbeha counties, Mississippi.
HABITAT.?Burrows (primary burrower).
Procambarus (Girardiella) connus Fitzpatriek
FIGURE 243
Procambarus hagenianus.?Fitzpatriek, 1968b:37 [in part].
Procambarus (Girardiella) hagenianus.?Hobbs, 1972b:47 [in part].
Procambarus (Girardiella) sp. C.?Fitzpatriek, 1976a:385, figs, lc, 2c [by
implication].
Procambarus (Girardiella) connus Fitzpatriek, 1978b:76, figs. 72-90, 109-
121.
Procambarus (Girardiella) connos.?Fitzpatriek, 1978b:57 [erroneous spell-
ing].
P[rocambarus] connus.?Stewart, 1986:15.
TYPES.?Holotype, allotype, and morphotype, USNM
146261,146262,146263; paratypes, USNM.
TYPE LOCALITY.?Carrollton, Carroll County, Mississippi.
RANGE.?Vicinity of the type locality.
HABITAT.?Burrows (primary burrower).
Procambarus (Girardiella) curdi Reimer
FIOURE 235
Procambarus A.?Reimer and Clark, 1974:172, figs. 20-23.
Procambarus (Girardiella) curdi Reimer, 1975:22, figs. 1-9.?Hobbs and
Robison, 1988:405, figs. 10,16.
TYPES.?Holotype and allotype, USNM 144250, 144251
(male I, female); paratypes, Rollin D. Reimer.
TYPE LOCALITY.?Navasota River, northeast of Bryan on
U.S. Highway 190, Brazos County, Texas.
RANGE.?Navasota and Trinity river watersheds in Texas,
and the Red River basin in southwestern Arkansas and
southeastern Oklahoma.
HABITAT.?Lentic and sluggish lotic situations.
Procambarus (Girardiella) gracilis (Bundy)
FIGURE 239
Cambarus gracilis Bundy, 1876:5.
Cambarus gracillis.?Bundy, 1882:182 [erroneous spelling].
Cambarus (Cambarus) gracilis.?Ortmann, 1905c: 101.
Cambarus (Ortmannicus) gracilis.?Fowler, 1912:341 [by implication].
Cambarus (Girardiella) gracilis.?Lyle, 1938:76 [by implication].
Procambarus gracilis.?Hobbs, 1942a:342 [by implication].?Williams,
1954:826, figs. 10-18.?Page, 1985b:370, figs. 98-102.?Pflieger,
1987a:22; 1987b:6.
Procambarus (Girardiella) gracilis.?Hobbs, 1972a:7, fig. Ik; 1974b:47, fig.
193.?Hobbs III and Rewolinski, 1985:26, figs. 1,2.
TYPES.?Syntypes, INHS (no longer there), MCZ 3794
(male I, female), MCZ 3454 (male I).
TYPE LOCALITY.?Normal, McLean County, Illinois. (Re-
stricted by Hobbs, 1974b:47).
RANGE.?Wisconsin, Iowa, Illinois, Indiana, Missouri, and
perhaps Kansas, Oklahoma, and Texas.
HABITAT.?Burrows (primary burrower).
REMARKS.?As is true of most of the wide-ranging primary
burrowers, it is possible, if not probable, that many of the
reported localities (particularly those from which first form
males are not known) are based on misidentifications.
Procambarus (Girardiella) hagenianus hagenianus (Faxon)
FIGURE 242
Cambarus advena.?Hagen, 1870:86-87 [in part: figs. 90-92, 164. The
description is of hagenianus. The Georgia specimens mentioned are
members of P. (H.) pygmaeus and P. (H.) advena.]
Cambarus Carciinus.?Hagen, 1870:31.
Cambarus carolinus.?Brocchi, 1875:27.
Cambarus Hagenianus Faxon, 1884:141.
Cambarus hagenianus.?Hay, 19O2c:38.?Faxon, 1914:366, pi. 1, pi. 7: figs.
la-c,7.
Cambarus (Cambarus) hagenianus.?Ortmann, 1905c: 101.
Cambarus (Ortmannicus) hagenianus.?Fowler, 1912:341 [by implication].
Cambarus (Girardiella) hagenianus hagenianus.?Lyle, 1938:76.
Procambarus hagenianus.?Hobbs, 1942a:342 [by implication].
Procambarus (Girardiella) hagenianus.?Hobbs, 1972a:7, figs. 2d, 7a-g;
1974b:47,fig. 192.?Fitzpatriek, 1976a:385, figs. lh,2h [by implication].
Procambarus (Girardiella) hagenianus hagenianus.?Fitzpatriek, 1978b:60,
figs. 1-18,109-121.
Procambarus hagenianus hagenianus.?Fitzpatriek, 1983b:198, fig. 193.
TYPES.?Holotype, MCZ 232 (male I).
TYPE LOCALITY.?Given as Charleston, South Carolina, but
definitely based on mislabeled specimen.
RANGE.?Cited by Fitzpatriek (1978b:61) as "south and east
of Tibbie Creek in the Tombigbee drainage of Lowndes,
Noxubee and Oktibbeha counties, Mississippi, and Marengo,
Pickens and Sumter counties Alabama "
HABITAT.?Burrows (primary burrower).
NUMBER 480 57
Procambarus (Girardiella) hagenianus vesticeps Fitzpatrick
FIGURE 241
Cambarus (Girardiella) hagenianus evansi Lyle, 1938:76 [nomen nudum].
Procambarus hagenianus.?Fitzpatrick, 1968b:37 [in part].
Procambarus (Girardiella) hagenianus.?Hobbs, 1972b:47 [in part].
Procambarus (Girardiella) subsp. A.?Fitzpatrick, 1976a:385,figs. la, 2a [by
implication].
Procambarus (Girardiella) hagenianus vesticeps Fitzpatrick, 1977:372 [nomen
nudum]; 1978b:64, figs. 19-36,109-121.
C[ambarus] h[agenianus] evansi.?Fitzpatrick, 1978b:69.
TYPES.?Holotype, allotype, and morphotype, USNM
146265, 146266,146267 (male I, female, male II); paratypes,
USNM, BMNH.
TYPE LOCALITY.?Egypt, Chickasaw County, Mississippi.
RANGE.?Fitzpatrick (1978b:67) recorded the range as
"north of Tibbie Creek in Tombigbee drainage of Chickasaw,
Clay, Monroe and Pontotoc counties [Mississippi]" inter-
grading with the nominate subspecies in Monroe County.
HABITAT.?Burrows (primary burrower).
Procambarus (Girardiella) liberorum Fitzpatrick
FIGURE 240
Procambarus (Girardiella) liberorum Fitzpatrick, 1978c:533, figs. 1-14.?
Hobbs andRobison, 1988:407, figs. 13,15.
Procambarus liberorum.?Bouchard and Robison, 1981:28.
TYPES.?Holotype and allotype, USNM 148353, 148354
(male I, female); paratypes, USNM.
TYPE LOCALITY.?Yard at 206 S.W. Seventh Street,
Bentonville, Benton County, Arkansas.
RANGE.?Known only from the Arkansas River basin from
Benton and Washington to Pope and Yell counties, Arkansas.
HABITAT.?Burrows (primary burrower).
Procambarus (Girardiella) parasimulans Hobbs and
Robison
FIGURE 237
Procambarus (Girardiella) parasimulans Hobbs and Robison, 1982:545, fig.
1; 1988:407, figs. 8,16.
TYPES.?Holotype, allotype, and morphotype, USNM
177698, 177699,177700 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Unnamed tributary to Prairie Bayou, 10.2
miles (16.3 km) east of Bismarck on State Route 84 (Sec. 35, T.
4S, R. 19W), Hot Spring County, Arkansas.
RANGE.?Arkansas, Ouachita, and Red river basins in
southwestern Arkansas.
HABITAT.?Lentic and sluggish lotic situations and burrows.
Procambarus (Girardiella) pogum Fitzpatrick
FIGURE 245
Cambarus (Girardiella) hagenianus carri Lylc, 1938:76 [nomen nudum].
Procambarus hagenianus.?Hobbs, 1968b:K25,fig. 19c.
Procambarus (Girardiella) hagenianus.?Hobbs, 1972b:47 [in part].
Procambarus (Girardiella) sp. D.?Fitzpatrick, 1976a:385, figs. Id, 2d [by
implication].
Procambarus (Girardiella) pogum Fitzpatrick, 1978b:83,figs. 91-121.
Procambarus poqum.?Stewart, 1986:15 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
146270, 146271, 146272 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Houston, Chickasaw County, Missis-
sippi.
RANGE.?Known only from the Houlka-Tibbie Creek basin
in Chickasaw County, Mississippi.
HABITAT.?Burrows (primary burrower).
Procambarus (Girardiella) regiomontanus Villalobos
FIGURE 238
Procambarus simulans regiomontanus Villalobos, 1954a:289, pis. 1, 2;
1955:47, pis. 9, 10; 1983:51, pis. 9, 10.?Campos, 1982:18, photographs
1-18.
Procambarus simulans regimontanus.?Hobbs, 1966b:70 [erroneous spelling].
Procambarus (Girardiella) simulans regiomontanus.?Hobbs, 1972a:7;
1974b:47,fig. 191.
Procambarus (Girardiella) regiomontanus.?Hobbs and Robison, 1988:398,
fig. 6.
TYPES.?Holotype, allotype, and morphotype, IBM (male I,
female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?Five kilometers (3.1 mi) north of Monter-
rey, Nuevo Le6n, Mexico.
RANGE.?Known only from Nuevo Le6n and Tamaulipas
provinces, Mexico.
HABITAT.?Lentic and lotic situations and burrows.
REMARKS.?This taxon was elevated by Hobbs and Robison
(1988) to specific rank for, on the basis of present data, there is
no evidence that it intergrades with P. (G.) simulans.
Procambarus (Girardiella) reimeri Hobbs
FIGURE 236
Procambarus (Girardiella) reimeri Hobbs, 1979:804. fig. 1.?Hobbs and
Robison, 1988:409, figs. 11,16.
Procambarus reimeri.?Bouchard and Robison, 1981:26.
TYPES.?Holotype, allotype, and morphotype, USNM
14880, 14881, 14882 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Roadside ditch about 5 miles (8 km)
northeast of Mena on unnumbered road to Irons Fork River,
Polk County, Arkansas.
RANGE.?Known only from the Ouachita River basin in
Polk county, Arkansas.
HABITAT.?Burrows and temporary pools.
Procambarus (Girardiella) simulans (Faxon)
FIGURE 234
Cambarus simulans Faxon, 1884:112; 1885a:48.pl.I: fig. 2, pi. viii: figs. 3,3',
3a, 3a'.
58 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambarus similans.?Ortmann, 1892:12 [erroneous spelling].
Cambarus gallinus Cockerell and Porter, 1900:434, fig. 1. [Syntypes, MCZ
7342 (3 male ID. USNM 23915 (male II, 2 female), USNM 23916 (male I.
female),USNM 129197 (2malell,2female),ANSP 1455 (2female),ANSP
1456 (male I). Type locality, Gallinas River at Las Vegas, New Mexico.]
Cambarus gallinas.?Harris, 1900:264 [erroneous spelling].
Cambarus baumgartneri Harris, 1901:115 [nomen nudum].
Cambarus (Cambarus) simulans.?Ortmann, 1905c:101.
Cambarus (Ortmannicus) simulans.?Fowler, 1912:341 [by implication].
Procambarus simulans.?Hobbs, 1942a:342 [by implication].
Cambarus simulana.?Villalobos, 1953:368 [erroneous spelling].
Procambarus simulans simulans.?Villalobos, 1954a:296.
Procambarus (Girardiella) simulans simulans.?Hobbs, 1972a:7; 1974b:47,
fig. 190.
Procambarus stimulans.?Airaksinen, Calkama, and Lindqvist, 1978:354
[erroneous spelling].
Procambarus (Girardiella) simulans.?Hobbs and Robison, 1988:409, figs. 9,
16.
TYPES.?Syntypes, MCZ 3646 (male I, 4 female), MCZ
3647 (23 male 11,30 female, 4 juv. male, 5 juv. female), USNM
4150 (male 1,3 male II, 5 juv. male), ZIAS 1/44975 (2 male,
female). Three second form male syntypes (MCZ 7342) could
not be located.
TYPE LOCALITY.?Dallas, Texas.
RANGE.?New Mexico, Texas, Louisiana, Oklahoma, Kan-
sas, Arkansas(?), and Colorado.
HABITAT.?Lentic and lotic situations, and burrow (secon-
dary burrower).
REMARKS.?The range of this crayfish is by no means
clearly defined. Further study, particularly of first form males,
throughout the recorded range is badly needed.
Procambarus (Girardiella) tulanei Penn
FIGURE 233
Procambarus tulanei Penn, 1953a: 163, figs. 1-12.
Procambarus (Girardiella) tulanei?Hobbs, 1972a:7; 1974b:48. fig. 189.?
Hobbs and Robison, 1988:410, figs. 7,15.
TYPES.?Holotype, allotype, and morphotype, USNM
93655, 93656, 79928 (male I, female, male if); paratypes,
USNM, TU, La. Polytechnic Inst.
TYPE LOCALITY.?Tributary of Bayou D'Arbonne, 4 miles
(6.4 km) west of Dubach on State Route 288, Lincoln Parish,
Louisiana.
RANGE.?Drainage systems of Arkansas, Ouachita, and Red
rivers in Louisiana and Arkansas, and the lower Arkansas River
basin in Arkansas.
HABITAT.?Lentic and lotic situations, and burrows (secon-
dary burrower).
Subgenus Hagenides Hobbs
Astacus.?LeConte, 1856:402 [in part].
Cambarus.?Hagen, 1870:87 [in part].
Subgenus Cambarus Ortmann, 1905c:97 [in part].
Subgenus Ortmannicus Fowler, 1912:341 [in part].
Procambarus.?Hobbs, 1942a:342 [by implication].
Subgenus Hagenides Hobbs, 1972a:7 [Type species, by original designation,
Astacus advena LeConte, 1856:402. Gender: masculine.]; 1984, fig. 8.
Procambarus (Hagenides) advena (LeConte)
FIGURE 247
Astacus advena LeConte, 1856:402.
Cambarus Carolinus.?Hagen, 1870:31, 87-88 [latter in part], pi. 1: fig,.
51-54,pl. 3: fig. 165.
Cambarus advena.?Hagen, 1870:87, pi. 3: fig. 164, pi. 7.?Faxon, 1884:140.
Cambarus (Cambarus) advena.?Ortmann, 1905c:101.
Cambarus (Ortmannicus) advena.?Fowler, 1912:341 [by implication].
Procambarus advena.?Hobbs, 1942a:342 [in part].
Procambarus (Hagenides) advena.?Hobbs, 1972a:7 [in part]; 1974b:48 [in
part],fig. 194; 1981:311,figs. 17f, 118a, 119-121,235.
TYPES.?Syntypes, MCZ 3379 (female), ANSP 321 (fe-
male).
TYPE LOCALITY.?"Georgia inferiore." Restricted to 2.5
miles (4 km) west of Riceboro, Liberty County, Georgia, by
Hobbs, 1974b:48.
RANGE.?Lower coastal plain of Georgia, between Savan-
nah and Altamaha rivers.
HABITAT.?Burrows (primary burrower).
REMARKS.?As pointed out by Faxon (1884:140), the
descriptions of P. advena and C. carolinus were transposed in
Hagen's monograph, and pi. 1: figs. 51-54 and pi 3: fig. 165
are actually of P. advena. (See Hobbs, 1981:311-312 for a
detailed bibliography and explanation).
Procambarus (Hagenides) caritus Hobbs
FIGURE 249
Procambarus advena.?Hobbs, 1942b:77 [in part].
Procambarus (Hagenides) caritus Hobbs, 1981:319, figs. 118b, 122,123.
TYPES.?Holotype, allotype, and morphotype, USNM
117598,133678, and 116952; paratypes, USNM.
TYPE LOCALITY.?Seepage area, 3.7 miles (5.9 km) west of
Glenwood, Wheeler County, Georgia, on U.S. Highway 280.
RANGE.?Altamaha and Satilla watersheds in the coastal
plain of Georgia.
HABITAT.?Burrows (primary burrower).
Procambarus (Hagenides) geodytes Hobbs
FIGURE 250
Cambarus advena geodytes Hobbs, 1937:154 [nomen nudum].
Procambarus geodytes Hobbs, 1942b:80,figs. 61-65,276-285.
Procambarus (Hagenides) geodytes.?Hobbs, 1972a:7; 1974b:48, figs. 195,
196b [latter labeled P. (H.) pygmaeus].
Procambarus (Hagenides) pygmaeus.?Hobbs, 1974b, fig. 196b.
TYPES.?Holotype and allotype, USNM 81279 (male I,
female); "morphotype," USNM 81280 (male II); paratypes,
MCZ, USNM, UF (now in USNM), FSBC.
TYPE LOCALITY.?Orange Springs in the northeastern part
NUMBER 480 59
of Marion County, Florida (Sec. 2, T. 12S, R. 23E); found in
burrows near the springs.
RANGE.?Along the St. Johns River and its tributaries from
Putnam County south to Seminole County, Florida.
HABITAT.?Burrows in flatwoods, seepage areas, and in
vicinities of springs (primary burrower).
Procambarus (Hagenides) pygmaeus Hobbs
FIGURE 251
Procambarus pygmaeus Hobbs, 1942b:83,figs. 66-70,296-304.
Procambarus (Hagenides) pygmaeus.?Hobbs, 1972a:7; 1974b:48, fig.
196a,c-g [not b, which is P. (H.) geodytes]; 1981:324, figs. 17c, 102, 118c,
124,237.
TYPES.?Holotype, allotype, and "morphotype," USNM
81285 (male I, female, male II); paratypes, MCZ, USNM,
FSBC.
TYPE LOCALITY.?Roadside ditch 15.7 miles (25.1 km)
north of Fargo on State Route 89, Clinch County, Georgia.
RANGE.?From the Savannah River southward to Putnam
County, Florida, and in the panhandle of the latter in Gulf,
Liberty, Leon, and Wakulla counties.
HABITAT.?Lentic and lotic situations and burrows (secon-
dary burrower).
Procambarus (Hagenides) rogersi campestris Hobbs
FIGURE 253
Procambarus rogersi campestris Hobbs, 1942b:90, figs. 81-85; 1945a:255,
figs. 3,4,7,16,18,21,24.
Procambarus (Hagenides) rogersi campestris.?Hobbs, 1972a:7; 1974b:49,
fig. 198.
TYPES.?Holotype, allotype, and "morphotype," USNM
79384 (male I, female, male II); paratypes, MCZ, USNM.
TYPE LOCALITY.?About 12 miles (19.2 km) south of
Tallahassee, Leon County, Florida (Sec. 16-17, T. 2S, R. 2W).
RANGE.?Leon and Wakulla counties, Florida.
HABITAT.?Burrows in flatwoods and seepage areas (pri-
mary burrower).
Procambarus (Hagenides) rogersi expletus Hobbs and
Hart
FIGURE 254
Procambarus rogersi expletus Hobbs and Hart, 1959:179, figs. 27-39.
Procambarus (Hagenides) rogersi expletus.?Hobbs, 1972a:7; 1974b:49, fig.
199.
TYPES.?Holotype, allotype, and morphotype, USNM
102125, 102127, 102126 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Seepage slope, 0.9 mile (1.4 km) north of
Clarksville, Calhoun County, Florida, on State Route 73 (Sec.
25, T. 1N,R. 10W).
RANGE.?Known only from the type locality.
HABITAT.?Burrows in flatwoods and seepage areas (pri-
mary burrower).
Procambarus (Hagenides) rogersi ochlocknensis Hobbs
FIGURE 252
Procambarus rogersi ochlocknensis Hobbs, 1942b:89, figs. 76-80; 1945a:252,
figs. 1,10.11,13,15,19,20,26.
Procambarus rogersi ochlockensis.?Hobbs III, 1969:25 [erroneous spelling].
Procambarus (Hagenides) rogersi ochlocknensis.?Hobbs, 1972a:7;
1974b:49,197.
Procambarus (Hagenides) rogersi ocklocknensis.?Hobbs, 1974b, fig. 197
[erroneous spelling].
TYPES.?Holotype, allotype, and "morphotype," USNM
79383 (male I, female, male II); paratypes, MCZ, USNM.
TYPE LOCALITY.?7.3 miles (11.7 km) west of Quincy, on
U.S. Highway 90, Gadsden County, Florida (Sec. 1, T. 2N, R.
5W).
RANGE.?Ochlockonee River basin in Gadsden and Liberty
counties, Florida.
HABITAT.?Burrows in seepage areas (primary burrower).
Procambarus (Hagenides) rogersi rogersi (Hobbs)
FIGURE 255
Cambarus rogersi Hobbs, 1938a:62, figs. 1-11.
Procambarus rogersi.?Hobbs, 1942a:344.
Procambarus rogersi rogersi.?Hobbs, 1942b:89, figs. 71-75.
Procambarus (Hagenides) rogersi rogersi.?Hobbs, 1972a:7; 1974b:49, fig.
200.
TYPES.?Holotype, allotype, and "morphotype," USNM
75120, 75121, 75589 (male I, female, male II); paratypes,
MCZ, USNM.
TYPE LOCALITY.?Flatwoods, 4 miles (6.4 km) north of
Blountstown on State Route 76 (71, old number), Calhoun
County, Florida (Sec. 14, T. 1N.R. 9W).
RANGE.?Eastern Calhoun County, Florida.
HABITAT.?Burrows in flatwoods and seepage areas (pri-
mary burrower).
Procambarus (Hagenides) talpoides Hobbs
FIGURE 248
Cambarus advena.?Hagen, 1870:87 [in part].
Procambarus advena.?Hobbs, 1942b:73-80 [in part], figs. 56-60.
Procambarus (Hagenides) advena.?Hobbs, 1972b:50, 150, 154 [in part];
1974b:48 [in part].
Procambarus (Hagenides) talpoides Hobbs, 1981:329. figs. 17e. 118d,e, 120,
125-127.238.
TYPES.?Holotype, allotype, and morphotype, USNM
115002,115018, and 133347; paratypes, USNM.
TYPE LOCALITY.?Burrows along Hogtown Creek just
southwest of the junction of 13th (U.S. Highway 441) and
Boundary streets in Gainesville, Alachua County, Florida.
60 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
RANGE.?Coastal plain from the Satilla River system
westward to Tift County, Georgia, and southward to Thomas
County, Georgia, and Alachua and Putnam counties, Florida;
also present on Cumberland Island, Camden County, Georgia.
HABITAT.?Burrows (primary burrower).
Procambarus (Hagenides) truculentus Hobbs
FIGURE 256
Procambarus truculentus Hobbs, 1954a: 111, figs. 1-13.
Procambarus Truculentus.?Hobbs, 1954a:l 11.
Procambarus (Hagenides) truculentus.?Hobbs, 1972a:7; 1974b:49, fig. 201;
1981:337, figs. 17b, 118f, 120,128,129.239.
TYPES.?Holotype, allotype, and morphotype, USNM
95670, 95671, 95672 (male I, female, male II); paratypes,
MCZ,TU,USNM.
TYPE LOCALITY.?Seepage area 11 miles (17.6 km) north of
Lyons on U.S. Highway 1, Emanuel County, Georgia.
RANGE.?Upper Coastal Plain between the Oconee and
Ogeechee rivers in Georgia.
HABITAT.?Burrows in flatwoods and seepage areas (pri-
mary burrower).
Subgenus Leconticambarus Hobbs
Astacus.?LcCatic, 1856:401 [in part].
Cambarus.?Hagen, 1870:16 [in part].
Subgenus Cambarus.?Ortmann, 19O5c:97 [in part].
Subgenus Ortmannicus Fowler, 1912:341 [in part].
Genus Procambarus.?Hobbs, 1942a:342.
Subgenus Ortmanmanicus.?Hobbs, 1942a:342 [in part, erroneous spelling].
Subgenus Leconticambarus Hobbs, 1972a:7 [Type species, by original
designation, Cambarus barbatus Faxon, 1890:621. Gender masculine.];
1984, fig. 6.
Procambarus (Leconticambarus) alleni (Faxon)
FIGURE 257
Cambarus Alleni Faxon, 1884:110; 1885a:35,pL 1: fig. l.pL 8: figs. 2,2'.
Cambarus alleni.?Faxon, 1890:619.
Cambarus (Cambarus) alleni.?Ortmann, 1905c: 102.
Cambarus (Ortmannicus) alleni?fowler, 1912:341 [by implication].?
Creaser, 1934b:4 [by implication].
Procambarus alleni?Hobbs, 1942a:342 [by implication]; 1942b:69 figs
51-55.
Procambarus allenti .?Bovbjerg, 1956:612 [erroneous spelling].
Procambarus allenii?Gifford, 1962:209.
Procambarus (Leconticambarus) alleni.?Hobbs, 1972a:8; 1974b:49, fig. 202.
TYPE.?Holotype, MCZ 3531 (male I).
TYPE LOCALITY.?SL Johns River, Hawkinsville, 1 mile
(1.6 km) upstream from bridge at Crow's Bluff, between Eustis
and Deland), Lake County, Florida.
RANGE.?East of St. Johns River and all of peninsular
Florida in and south of Levy and Marion counties; also present
on some of the Keys.
HABITAT.?Lentic and lotic situations and burrows.
Procambarus (Leconticambarus) apalachicolae Hobbs
FIGURE 261
Procambarus apalachicolae Hobbs, 1942b:55,figs. 26-30,256-265.
Procambarus (Leconticambarus) apalachicolae.?Hobbs, 1972a:8; 1974b:50,
fig. 206.
TYPES.?Holotype and allotype, USNM 81272 (male I,
female); "morphotype," USNM 81273 (male II); paratypes,
MCZ, USNM.
TYPE LOCALITY.?Roadside ditch, 11.1 miles (17.8 km)
west of Beacon Hill on U.S. Highway 98, Bay County, Florida
(Sec. 27, T. 5S,R. 13W).
RANGE.?Coastal flatwoods in Bay and Gulf counties,
Florida.
HABITAT.?Lentic situations and burrows (secondary bur-
rower).
Procambarus (Leconticambarus) barbatus (Faxon)
FIGURE 259
Astacus penicillatus LeConte, 1856:401 [not Aslacus penic.Hiatus Olivier,
1791].
Cambarus penicillatus.?Hagen, 1870:16,53, figs. 93,94,149 [not 95 and 96,
which are P. (0.) ancylus].
Cambarus barbatus Faxon, 1890:621 [substitute name; in part].
Cambarus (Cambarus) barbatus.?Ortmann, 1905c: 102.
Cambarus (Ortmannicus) barbatus.?Fowler, 1912:341 [by implication].
Procambarus barbatus.?Hobbs, 1942a:340; 1942b:39.figs. 1-5.
Procambarus (Leconticambarus) barbatus.?Hobbs, 1972a:8, figs, lm, 2f,
9a-g; 1974b:50,fig. 204; 1981:342, figs. 18b, 130a,b, 131,132,240.
TYPE.?MCZ 279 (male I), designated neotype of A
penicillatus LeConte by Hobbs, 1974b:50. Specimens from
Flomaton, Alabama, cited by Faxon (1890) and listed as
paratypes by him (1914:414) are paratypes of Procambarus
(L.) escambiensis Hobbs.
TYPE LOCALITY.?"Habitat in fossis Georgiae inferioris"
(LeConte, 1856:401). Restricted by Hobbs (1974b:50) to 2.5
miles (4 km) west of Riceboro, Liberty County, Georgia, in
pine flatwoods.
RANGE.?In the coastal plain from the Altamaha River in
Georgia to the Edisto River in South Carolina.
HABITAT.?Lentic situations and burrows (secondary bur-
rower).
REMARKS.?The type of LeConte's Astacus penicillatus is
no longer extant. Inasmuch as Hagen (1870) and Faxon (1884,
1885a, 1890, and 1914) confused other species with it, and, in
the latter work, Faxon included members of two species in his
type series of C. barbatus (= A. penicillatus), his "Type" was
designated the neotype of LeConte's A. penicillatus to establish
the identity of the latter's species.
Procambarus (Leconticambarus) capillatus Hobbs
FIGURE 267
Procambarus capillatus Hobbs, 1971b:83, fig. 1.
Procambarus (Leconticambarus) capillatus.?Hobbs, 1972a:8; 1974b:50, fig-
211.
NUMBER 480 61
TYPES.?Holotype, allotype, and morphotype, USNM
131454, 131455,131456 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Drainage ditch adjacent to Burnt Corn
Creek (Escambia River drainage) on State Route 41 northwest
of Brewton, Escambia County, Alabama.
RANGE.?Escambia River basin in Conecuh, Escambia, and
Monroe counties, Alabama, and Escambia County, Florida.
HABITAT.?Temporary ponds, pools, and burrows (secon-
dary burro wer).
Procambarus (Leconticambarus) econfinae Hobbs
FIGURE 262
Procambarus econfinae Hobbs, 1942b:49,figs. 16-20,236-245.
Procambarus (Leconticambarus) econfinae.?Hobbs, 1972a:8; 1974b:50, fig.
207.
TYPES.?Holotype, allotype, and "morphotype," USNM
81276 (male I, female, male II); paratypes, MCZ, USNM,
BMNH.FSBC.
TYPE LOCALITY.?Flatwoods in the northern part of Panama
City, Bay County, Florida, between railroad and U.S. Highway
231 (Sec. 33, T. 3S.R. 14W).
RANGE.?Environs of Panama City, Bay County, Florida.
HABITAT.?Flatwoods, in ditches and temporary ponds
(secondary burro wer).
Procambarus (Leconticambarus) escambiensis Hobbs
FIGURE 263
Cambarus barbatus Faxon, 1890:621 [in part].?Faxon, 1914:414 [in part].
Procambarus escambiensis Hobbs, 1942b:46, figs. 11-15,226-235.
Procambarus (Leconticambarus) escambiensis.?Hobbs, 1972a:8; 1974b:51,
fig. 208.
TYPES.?Holotype, allotype, USNM 81278 (male I, female),
and "morphotype," USNM 81277 (male II); paratypes, MCZ,
USNM,FSBC.
TYPE LOCALITY.?About 100 yards (90 m) east of Perdido
River on U.S. Highway 90, Escambia County, Florida (Sec. 10,
T. 1S.R. 32W).
RANGE.?Known from the type locality and from Flomaton,
Escambia County, Alabama. Reported from the latter as
Cambarus barbatus by Faxon (1890); specimens listed as
paratypes of that species by him (1914), designated paratypes
of P. (L.) escambiensis by Hobbs (1942:48).
HABITAT.?Temporary bodies of water in flatwoods and
floodplains (secondary burrower).
Procambarus (Leconticambarus) hubbeltt (Hobbs)
FIGURE 269
Cambarus hubbelli Hobbs, 1938b:90 [nomen nudum; validated by Hobbs,
194Oa:4O6,fig. 19].
Procambarus hubbelli.?Hobbs, 1942a:342; 1942b:67,figs. 46-50.
Procambarus (Leconticambarus) hubbelli.?Hobbs, 1972a:8; 1974b:51, fig.
213.
TYPES.?Holotype and allotype, USNM 76593 (male I,
female); "morphotype," USNM 79351 (male II); paratypes,
MCZ, USNM, Chas. Mus.
TYPE LOCALITY.?One mile (1.6 km) east of Bonifay on
U.S. Highway 90, Holmes County, Florida (Sec. 5, T. 3N, R.
14W).
RANGE.?Choctawhatchee and Yellow river drainages in
Alabama and Florida.
HABITAT.?Temporary ponds, pools, and burrows (secon-
dary burrower).
Procambarus (Leconticambarus) kilbyi (Hobbs)
FIGURE 268
Cambarus kilbyi Hobbs, 1940a:410, fig. 20.
Procambarus kUbyi.?Edbbs, 1942a:343; 1942b:64, figs. 41-45.?Caine,
1974:10, figs. ld,4d,5a.
Procambarus (Leconticambarus) kilbyi.?Hobbs, 1972a:8; 1974b:51,fig. 212.
TYPES.?Holotype, allotype, and "morphotype," USNM
76594 (male I, female, male II); paratypes, MCZ, USNM,
FSBC.
TYPE LOCALITY.?Seven miles (11.2 km) northwest of
Blountstown on State Route 71 (6, old number), Calhoun
County, Florida (Sec. 3,T. 1N,R. 9W).
RANGE.?Coastal flatwoods from Calhoun and Gulf coun-
ties east and south to Levy County, Florida.
HABITAT.?Temporary ponds, pools, and burrows (secon-
dary burrower).
Procambarus (Leconticambarus) latipleurum Hobbs
FIGURE 260
Procambarus latipleurum Hobbs, 1942b:52,figs. 21-25.246-255.
Procambarus (Leconticambarus) latipleurum.?Hobbs, 1972a:8; 1974b:51,
fig. 205.
TYPES.?Holotype, allotype, and "morphotype," USNM
81281 (male I, female, male II); paratypes, MCZ, USNM, UF
(now in USNM), FSBC.
TYPE LOCALITY.?5.8 miles (9.3 km) west of Wewahitchka
on State Route 22 (52, old number), Gulf County, Florida (Sec.
18, T. 4S,R. 10W).
RANGE.?Gulf County, Florida, north of the Wetappo Canal.
HABITAT.?Temporary ponds, pools, and burrows (secon-
dary burrower).
Procambarus (Leconticambarus) milleri Hobbs
FIGURE 258
Procambarus milleri Hobbs, 1971c:115,figs. 1-11,16.
Procambarus (Leconticambarus) milleri.?Hobbs, 1972a:8; 1974b:51, fig.
203.?Hobbs, Hobbs, and Daniel, 1977:121, fig. 5.
TYPES.?Holotype and morphotype, USNM 131257,
62 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
131258 (male I, male II); paratypes, USNM.
TYPE LOCALITY.?Well at Little Bird Nursery and Garden
Store at 8427 Bird Road, Miami, Dade County, Florida (Sec. 15,
T.54S.R. 40E).
RANGE.?Known only from the type locality.
HABITAT.?Subterranean waters.
Procambarus (Leconticambarus) pubischelae deficiens
Hobbs
FIGURE 265
Procambarus pubischelae Hobbs, 1942b:43 [in part].
Procambarus (Leconticambarus) pubischelae.?Hobbs, 1974b:82 [in part].
Procambarus (Leconticambarus) pubischelae deficiens Hobbs, 1981:356, figs.
18d, 132,135,241.
TYPES.?Holotype, allotype, and morphotype, USNM
133315, 133316,133317 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Roadside ditch, 3 miles (4.8 km) south of
Baxley, Appling County, Georgia.
RANGE.?Coastal plain of Georgia between the Ocmulgee-
Altamaha and Satilla rivers (one locality situated immediately
north of the Altamaha River in Montgomery County);
intergrading with the nominate subspecies in the extreme upper
and lower Satilla and in the upper Suwannee basins.
HABITAT.?Lentic and sluggish lotic situations and burrows
(secondary burrower).
Procambarus (Leconticambarus) pubischelae pubischelae
Hobbs
FIGURE 264
Procambarus pubischelae Hobbs, 1942b:41, figs. 6-10,216-225 [in part].
Procambarus pubischaelae.?Villalobos, 1955:38 [erroneous spelling].
Procambarus (Leconticambarus) pubischelae.?Hobbs, 1972a:8 [in part];
1974b:52 [in part],fig. 209.
Procambarus (Leconticambarus) pubischelae pubischelae.?Hobbs,
1981:350, figs. 18c, 130c,d, 132-134,242.
TYPES.?Holotype, allotype, and "morphotype," USNM
81284 (male I, female, male II); paratypes, MCZ, ANSP,
USNM.
TYPE LOCALITY.?Cypress ponds and roadside ditch 9.4
miles (15 km) north of Lake City on U.S. Highway 441,
Columbia County, Florida (Sec. 17, T. 2S,R. 17E).
RANGE.?From the Satilla River in Georgia south to
Alachua and Flagler counties, Florida; intergrading with
Procambarus (L.) pubischelae deficiens in the extreme upper
and lower Satilla and in the upper Suwannee basins.
HABITAT.?Lentic and sluggish lotic situations and burrows
(secondary burrower).
Procambarus (Leconticambarus) rathbunae (Hobbs)
FIGURE 266
Cambarus rathbunae Hobbs, 1940a:414,fig. 21.
Procambarus rathbunae.?Hobbs, 1942a:344; 1942c:59.figs. 31-35.
Procambarus (Leconticambarus) rathbunae.?Hobbs, 1972a:8; 1974b:52, fig.
210.
TYPES.?Holotype, allotype, and "morphotype," USNM
76595 (male I, female, male II); paratypes, MCZ, USNM,
FSBC, BMNH.
TYPE LOCALITY.?Near the Yellow River at Milligan,
Okaloosa County, Florida, at the intersection of U.S. Highway
90 and State Route 4 (Sec. 22, T. 3N, R. 24W).
RANGE.?Okaloosa and Holmes counties, Florida.
HABITAT.?Temporary bodies of water and burrows (secon-
dary burrower).
Procambarus (Leconticambarus) shermani Hobbs
FIGURE 270
Procambarus shermani Hobbs, 1942b:61. figs. 36-40,266-275.
Procambarus (Leconticambarus) shermani.?Hobbs, 1972a:8; 1974b:52,214.
TYPES.?Holotype and "morphotype," USNM 81287 (male
I, male II); allotype, USNM 81288 (female); paratypes, MCZ,
USNM, FSBC, BMNH.
TYPE LOCALITY.?About 12 miles (19 km) southwest of Jay,
Santa Rosa County, Florida, in the Escambia River swamp
along McCaskill's Mill Creek (Sec. 28,31,T. 5N,R. 30W).
RANGE.?From the Escambia River drainage in Florida
westward to the Pearl River basin in Mississippi and Louisiana.
HABITAT.?Streams, sloughs, and burrows.
Subgenus Lonnbergius Hobbs
Cambarus.?Lonnberg, 1894:125 [in part].
Subgenus Cambarus.?Ortmarm, 1905c:97 [in part].
Subgenus Ortmannicus Fowler, 1912:341 [in part].
Procambarus.?Hobbs, 1942a:342 [in part].
Subgenus Lonnbergius Hobbs, 1972a:8 [Type species, by original designation,
Cambarus acherontis Lonnberg, 1894:125. Gender masculine.]; 1984, fig.
7.
Procambarus (Lonnbergius) acherontis (Ldnnberg)
FIGURE 271
Cambarus acherontis Lonnberg, 1894:125; 1895:6, figs. l-5b.?Hobbs,
194Oa:39O,fig. 15a-k.
Cambarus archerontis.?Hay, 1902b:437 [erroneous spelling].
Cambarus (Cambarus) acherontis.?Ortmann, 1905c: 102.
Cambarus (Ortmannicus) acherontis.?Fowler, 1912:341 [by implication].
Procambarus acherontis.?Hobbs, 1942a:342 [by implication]; 1942b:91,figs.
86-90.
Procambarus (Lonnbergius) acherontis.?Hobbs, 1972a:8, figs. 2g, lOa-g;
1974b:52, fig. 215.?Hobbs, Hobbs, and Daniel, 1977:122, fig. 56.?Franz
and Lee, 1982:58.
TYPE.?Holotype, ZIAS 1/4412(1 specimen).
TYPE LOCALITY.?Subterranean rivulet about 42 feet (12.6
m) from surface, near Lake Brantley, Seminole County, Florida
(probably Sec. 3, T. 21S, R. 29E); these co-ordinates were
furnished by R. Franz who searched the tax records of the
property of Mr. A.E. Sjoblom on which the well mentioned by
NUMBER 480 63
Ltinnberg was situated).
RANGE.?Known from only one locality in Orange County,
Florida, and three in Seminole County, Florida (see Franz and
Lee, 1982).
HABITAT.?Springs and underground streams.
REMARKS.?Hobbs (1940a:393), endeavoring to clarify the
confusion engendered by Faxon as to the identity of the species
(see P. (0.) lucifugus lucifugus) and believing that both of
LOnnberg's types were lost, designated neotypes from the Palm
Springs locality. Dr. Y. Starobogatov kindly has informed me
that one of LOnnberg's specimens is still extant (see above);
thus, the neotype designation should be rejected.
Subgenus Mexicambarus Hobbs
Cambarus.?Ortmann, 1909:159.
Subgenus Cambarus.?Ortmann, 1909:159.
Subgenus Ortmannicus Fowler, 1912:341 [in part, by implication].
Procambarus.?Hobbs, 1942a:342 [in part, by implication].
Subgenus Mexicambarus Hobbs, 1972a:8 [Type species, by original designa-
tion, Cambarus (Cambarus) bouvieri Ortmann, 1909:159. Gender: mascu-
line.]; 1984, fig. 8.
Procambarus (Mexicambarus) bouvieri (Ortmann)
FIGURE 272
Cambarus (Cambarus) bouvieri Ortmann, 1909:159, figs. 1,2.
Cambarus (Ortmannicus) bouvieri.?Fowler, 1912:341 [by implication].
Cambarus bouvieri.?Van Straelen, 1942:5.
Procambarus bouvieri.?Hobbs, 1942a:342 [by implication].?Villalobos,
1946:224,pl. 2: figs. 1-12; 1955:61,pi. 12; 1983:66,pi. 12.
Procambarus (Mexicambarus) bouvieri.?Hobbs, 1972a:9, figs. 2h, l la-g;
1974b:53,fig. 216.
procambarus bouvieri.?Barrera-Mera, 1973:242 [lapsus].
TYPES.?Syntypes, MHNP (2 male I, female), CM 74.864
(male I).
TYPE LOCALITY.?Uruapan, Michoacdn, Mexico.
RANGE.?Several localities in the vicinity of Uruapan,
Michoac?n, Mexico.
HABITAT.?Streams.
REMARKS.?There are two lots of this crayfish in the
Smithsonian collection which appear to me to bear erroneous
data: one from Valles, San Luis Potosi, Mexico, and the other
from Tuxtla Gutierrez, Chiapas, Mexico.
Subgenus Ortmannicus Fowler
Astacus.?Harlan, 1830:464.
Subgenus Cambarus Erichson, 1846:95 [in part].?Ortmann, 19O5c:97 [in
part].
Cambarus.?Girard, 1852:91 [in part].
Subgenus Ortmannicus Fowler, 1912:341 [in part. Type species, by original
designation, Astacus Blandingii Harlan, 1830:464. Gender: masculine.
Ortmannicus is a replacement name for Ortmann's subgenus Cambarus.].?
Hobbs, 1972a:9; 1984: fig. 3.
Procambarus.?Hobbs, 1942a :341 [in part].
Subgenus Ortmanmanicus.?Hobbs, 1942a:342 [in part; erroneous spelling].
Subgenus Orimanicus.?Hobbs III, Thorp, and Anderson, 1976:58 [erroneous
spelling].
Procambasrus.?Huner, 1977:10 [erroneous spelling].
Procambaris.?Wharton, 1978:46 [erroneous spelling].
Procambarus (Ortmannicus) acutissimus (Girard)
FIGURE 308
Cambarus acutissimus Girard, 1852:91.
Cambarus acutus.?Most authors prior to 1942 [in part].
Procambarus blandingii acutus.?Most authors from 1942 to 1955 [in part].
Procambarus acutissimus.?Hobbs, 1955a:98; 1962a:286, fig. 51.
Procambarus (Ortmannicus) acutissimus.?Hobbs, 1972a:9; 1974b:53, fig.
246; 1981:366, figs. 15c, 136b, 138a, 139-142,243.
Procambarus acutissumus.?Hart and Hart, 1974:31 [erroneous spelling].
TYPES.?ANSP 309 (2 male II). These specimens could not
be located in 1980 or 1985.
TYPE LOCALITY.?Affluent (= tributary to Tombigbee
River) of Mobile River, Kemper County, Mississippi.
RANGE.?Tombigbee River basin in Mississippi eastward to
the middle Chattahoochee River basin in Georgia.
HABITAT.?Sluggish streams, ponds, and roadside ditches.
Procambarus (Ortmannicus) acutus acutus (Girard)
FIGURE 301
Cambarus acutus Girard, 1852:91.
Astacus Blandingii.?LeConte, 1856:400 [in part].
Cambarus acutus var. A.?Hagen, 1870:36.
Cambarus acutus var. B.?Hagen, 1870:36.
Cambarus blandingii.?Many authors from 1870 to 1962 [in part].
Cambarus stygius Bundy, 1876:3. [Type not extant Type locality. Lake
Michigan at Racine, Racine County, Wisconsin].
Cambarus Blandingii, var. acuta.?Faxon, 1884:113; 1885a:20, pi. 7: figs. 2,
2'. 2", 2a, 2a'.
Cambarus Stygius.?Underwood, 1886:373.
Cambarus blandingii acutus.?Faxon, 1890:619.
Cambarus blandingii blandingii.?Many authors from 1890 to 1962 [in part].
Cambarus blandingii var. acuta.?Hay, 1891:147.
Camborus blandingii acutus.?Williamson, 1899:47 [erroneous spelling].
Cambarus blandingi acutus.?Ortmann, 1905c:105.
Cambarus (Cambarus) blandingi acutus.?Ortmann, 19O5c:126 [by implica-
tion].?Fleming, 1938:301.
Cambarus blandingiacutus.?Harmon, 1910, fig. 2 [lapsus].
Cambarus (Ortmannicus) blandingii acutus.?Fowler, 1912:341 [by implica-
tion].
Cambarus (Cambarus) blandingii acutus.?Fleming, 1938:301.
Cambarus (Ortmannicus) evictus Lyle, 1938:76 [nomen nudum].
Procambarus blandingii acutus.?Hobbs, 1942a:343 [by implication].
Procambarus blandingii blandingii.?Many authors from 1942 to 1962 [in
part].?Villalobos. 1959:326 [in part].
Ortmannicus blandingi acutus.?Rhoades, 1942:1.
Cambarus blandingi acutis.?Roberts, 1944:364 [erroneous spelling].
Procambarus acutus acutus.?Hobbs, 1967a:130; 1968b:K-8,figs. 21o,23d.
Procambarus acutus.?Walls and Black, 1967:60.?Pflieger, 1987a:30;
1987b: 14.
Procambarus (Ortmannicus) acutus acutus.?Hobbs, 1972a:9; 1974b:53, fig.
240; 1981:372,figs. 15b, 136a, 138b, 140,143-145,244.
Procambarus (Ortmannicus) act us actus.?Payne and Riley, 1974:127
[erroneous spelling].
64 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Procambarus act us.?Comeaux, 1976:614- [erroneous spelling].
Procambarus (Ortmanicus) acutus acutus.?Hobbs III, Thorp, and Anderson,
1976:58 [erroneous spelling].
Procambasrus acutus acutus.?Huner, 1977:10 [erroneous spelling].
Procambarus acutus acustus.?Chea, Avault, and Graves, 1979:349 [errone-
ous spelling].
TYPES.?Lost in Chicago fire, 1871 (Faxon, 1914:413).
TYPE LOCALITY.?Affluent (= tributary to Tombigbee
River) of Mobile River in Kemper County, Mississippi.
RANGE.?Coastal plain and piedmont from Maine to
Georgia, from Florida panhandle to Texas, and from Minnesota
to Ohio; intergrading with P. (O.) a. cuevachicae in southwest-
ern Texas and northern Mexico.
HABITAT.?Sluggish to moderately flowing streams and
most lentic situations.
REMARKS.?With little doubt, the populations currently
assigned to this subspecies constitute a species complex.
Procambarus (Ortmannicus) acutus cuevachicae (Hobbs)
FIGURE 302
Cambarus blandingii cuevachicae Hobbs, 1941a:l, fig. 1.
Procambarus blandingii cuevachicae.?Hobbs, 1942a:342.?Villalobos,
1955:70, pis. 13,14; 1983:72, pis. 13,14.
Procambarus acutus cuevachicae.?Hobbs, 1966b:70.
Procambarus (Ortmannicus) acutus cuevachicae.?Hobbs, 1972a:9; 1974b:53,
fig. 241.
TYPES.?Holotype, allotype, and morphotype, USNM
80030 (male I, female, male II); paratypes, USNM, PM.
TYPE LOCALITY.?La Cueva Chica, a limestone cave about
1.6 kilometers (1.0 mile) northeast of Pujal, San Luis Potosi,
Mexico.
RANGE.?Known from the states of Hidalgo, Puebla, San
Luis Potosf, Tamaulipas, and Veracruz, Mexico.
HABITAT.?Subterranean and lentic situations.
Brocambarus (Ortmannicus) ancylus Hobbs
FIGURE 293
Cambarus penicillatus.?Hagen, 1870:54 [in part], figs. 95,96.
Procambarus ancylus Hobbs, 1958a:164, figs. 13-24.
Procambarus (Ortmannicus) ancylus.?Hobbs, 1972a:9; 1974b:54,fig. 233.
TYPES.?Holotype, allotype, and morphotype, USNM
115050 (male I, female, male II); paratypes, USNM.
TYPE LOCALITY.?Summerville, Dorchester County, South
Carolina.
RANGE.?In the coastal plain from the Cape Fear River basin
in North Carolina southward to the Edisto and Ashepoo basins
in Colleton County, South Carolina.
HABITAT.?Lentic and lotic situations and burrows.
Procambarus (Ortmannicus) angustatus (LeConte)
FIGURE 282
Astacus angustatus LeConle, 1856:401.
Cambarus angustatus.?Hagen, 1870:50, figs. 65-67,146.
Cambarus (Cambarus) angustatus.?Ortmann, 1905c: 102.
Cambarus (Ortmannicus) angustatus.?Fowler, 1912:341 [by implication].
Procambarus angustatus.?Hobbs, 1942a:342 [by implication].
Procambarus (Ortmannicus) angustatus.?Hobbs, 1972a:9; 1974b:54, fig.
226; 1981:384,figs. 13c, 148,149,245.
TYPE.?Holotype, ANSP 444 (male I); could not be located
in 1981.
TYPE LOCALITY.?"Georgia inferiore, in aquae purae rivulos
qui inter colliculos arenosos (sand-hills) currunt" (LeConte,
1856:402).
RANGE.?Known from the type specimen only. Repeated
searches for this species in Georgia have been unsuccessful.
HABITAT.?Streams.
Procambarus (Ortmannicus) bivittatus Hobbs
FIGURE 319
Procambarus bivittatus Hobbs, 1942b:96,figs. 96-100,305-314.
Procambarus (Ortmannicus) bivittatus.?Hobbs, 1972a:10; 1974b:54,fig. 255.
TYPES.?Holotype, USNM 81274 (male I); allotype and
"morphotype," USNM 81275 (female, male II); paratypes,
MCZ,USNM,FSBC.
TYPE LOCALITY.?Sloughs along Escambia River on State
Route 4 (old number 62), Escambia County, Florida (Sec. 9, T.
6N,R. 30W).
RANGE.?From the Escambia River drainage in Escambia
and Santa Rosa counties, Florida, to the Pearl River drainage in
Pearl River County, Mississippi, and SL Tammany and
Washington parishes, Louisiana.
HABITAT.?Streams and sloughs.
Procambarus (Ortmannicus) blandingii (Harlan)
FIGURE 300
Astacus Blandingii Harlan, 1830:464.
Astacus blandingii.?DeKay, 1844:23.
Astacus (Cambarus) Blandingii.?Erichson, 1846:98.
Cambarus Blandingii.?Ginrd, 1852:91.
Cambarus Blandingii var. Blandingii.?Faxon, 1884:136 [by implication].
Cambarus blandingii blandingii.?Faxon 1890:619 [by implication].
Cambarus (Cambarus) blandingi.?Ortmann, 1905c:102.
Cambarus blandingi-typicus.?Ortmann, 1905c: 105.
Cambarus (Cambarus) blandingi blandingi.?Ortmann, 1905c:126 [by impli-
cation].
Cambarus (Ortmannicus) blandingii.?Fowler, 1912:341.
Procambarus blandingii blandingii.?Hobbs, 1942a:342 [by implication].
Procambarus blandingii.?Hobbs, 1967a: 130.
Procambarus (Ortmannicus) blandingii.?Hobbs, 1972a:9, figs. 2i, 12a-g;
1974b:54,fig. 239.
Procambarus a. blandingii.?Page, 1985b:376 [lapsus calami].
TYPE.?Holotype, ANSP 4177 (male I).
TYPE LOCALITY.?Camden, Kershaw County, South Caro-
lina.
RANGE.?In the coastal plain from the Pee Dee River system
in South Carolina and southern North Carolina southward to
the Santee River basin in South Carolina.
HABITAT.?Lentic and lotic situations and burrows.
NUMBER 480 65
Procambarus (Ortmannicus) caballeroi VHlalobos
FIOURE 321
Procambarus caballeroi ViUalobos, 1944b:175, pis. 1,2; 1955:87, pis. 15,16;
1959:314, figs. 47-50; 1983:86, pis. 15,16.
Procambarus (Ortmannicus) caballeroi?Hobbs, 1972a:9; 1974b:54,fig. 257.
TYPES.?Holotype, allotype, and morphotype, IBM
50040(004) (male I, female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?"Riachuelos del Sur de Villa Judrez,"
Puebla, Mexico.
RANGE.?Known only from the vicinity of the type locality.
HABITAT.?S treams.
Procambarus (Ortmannicus) chacei Hobbs
FIGURE 275
Procambarus chacei Hobbs, 1958b:5, pi. 2: figs. 14-26.
Procambarus sp. A.?Hobbs, 1958c:72,76,77,79,83-86,90.
Procambarus (Ortmannicus) chacei.?Hobbs, 1972a:9; 1974b:55, fig. 219.
IProcambarus (Ortmannicus) enoplosternum.?Hobbs, 1981:388 [in part].
TYPES.?Holotype, allotype, and morphotype, USNM
101289,101290, 101291 (male I, female, male II); paratypes,
MCZ, ANSP, IBM, TU, USNM.
TYPE LOCALITY.?Cedar Creek, 3 miles (4.8 km) east of
Lykes, Richland County, South Carolina, on U.S. Highway 76.
RANGE.?"From the Wateree River in South Carolina south
to the Canoochee River in Georgia" (Hobbs, 1974b:55). The
range is in need of further investigation; see "Remarks" below.
HABITAT.?Streams.
REMARKS.?Procambarus (O.) chacei was synonymized
with P. (O.) enoplosternum Hobbs by Hobbs (1981:388); the
reasons cited were presented on page 398. On the basis of
additional materials from South Carolina, there is reason to
question this action; therefore,/*. (0.) chacei is retained herein
tentatively.
Procambarus (Ortmannicus) delicatus Hobbs and Franz
FIGURE 291
Procambarus (Ortmannicus) delicatus Hobbs and Franz, 1986:509, figs. l-3d.
TYPES.?Holotype, allotype, USNM 218528, 144848 (male
II, female); paratype, USNM.
TYPE LOCALITY.?Alexander Springs, about 5.5 miles (9
km) south of Astor Park, Lake County, Florida (Levy Grant 39,
T. 16S.R. 27E; 19o04/50wN,29?04'3(rW).
RANGE.?Known only from the type locality.
HABITAT.?Subterranean waters and spring.
Procambarus (Ortmannicus) enoplosternum Hobbs
FIGURE 274
Procambarus enoplosternum Hobbs, 1947a:5, figs. 2, 5, 10, 12, 14, 18, 20,
23-25,32.
?Procambarus sp. A.?Hobbs, 1958c:72,76,77,79,83-86,90.
Procambarus enoplosterum.?Hobbs III, 1969:42 [erroneous spelling].
Procambarus (Ortmannicus) enoplosternum.?Hobbs, 1972a:9, 1974b:55, fig.
218; 1981:388,figs.l3e, 137b-d, 138c. 150-154,246.
Procambaris enoplosternum.?Wharton, 1978:46 [erroneous spelling].
TYPES.?Holotype, allotype, and "morphotype," USNM
82263 (male I, female, male II); paratypes, MCZ, USNM, TU.
TYPE LOCALITY.?Rocky Creek, 6 miles (9.6 km) south of
Lyons on U.S. Highway l.Toombs County,Georgia.
RANGE.?Lower Oconee and Ohoopee drainages in Georgia
eastward to the Santee River basin in Richland County, South
Carolina.
HABITAT.?Streams.
Procambarus (Ortmannicus) epicyrtus Hobbs
FIGURE 276
Procambarus epicyrtus Hobbs, 1958b:l.pL l:figs. 1-13.
Procambarus sp. C?Hobbs, 1958c:72,fig. 17.
Procambarus (Ortmannicus) epicyrtus.?Hobbs, 1972a:9; 1974b:55, fig. 220;
1981:398, figs. 13f, 137a, 138d, 155-157,247.
TYPES.?Holotype, allotype, and morphotype, USNM
101286, 101287, 101288 (male I, female, male II); paratypes,
MCZ, ANSP, IBM, TU, USNM.
TYPE LOCALITY.?South Ogeechee Creek, 6.8 miles (10.9
km) south of Sylvania on U.S. Highway 301 (not 306 as cited
in Hobbs, 1974b: 55), Screven County, Georgia.
RANGE.?Tributaries of the Ogeechee and lower Savannah
rivers in southeastern Georgia.
HABITAT.?Streams.
Procambarus (Ortmannicus) erythrops Relyea and
Sutton
FIGURE 286
Procambarus (Ortmannicus) erythrops Relyea and Sutton, 1975b:8, figs. 1,
2.?Hobbs, Hobbs, and Daniel, 1977:124. fig. 57.?Franz and Lee, 1982:59,
figs. 2,6.
Procambarus erythrops.?Relyea and Sutton, 1975b: 13.
TYPES.?Holotype, allotype, and morphotype, USNM
133471, 133472, 133473 (male I, female, male II); paratypes,
USNM.RNHL.BMNH.
TYPE LOCALITY.?Sim's Sink, 1 mile (1.6 km) west of the
junction of U.S. Highway 27 and 129, and 0.1 mile (160 m)
south of Highway 27, Suwannee County, Florida (Sec. 24, T.
6S.R. 14E).
RANGE.?Known from only four localities, all situated north
of the Santa Fe River, east of the Suwannee River, and west of
Ichetucknee Springs, Suwannee County, Florida: the type
locality and three other sinks.
HABITAT.?Subterranean waters.
66 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Procambarus (Ortmannicus) evermanni (Faxon)
FIGURE 297
Cambarus evermanni Faxon, 1890:620.
Cambarus (Cambarus) evermanni?Ortmann, 1905c:102.
Cambarus (Ortmannicus) evermanni.?Fowler, 1912:341 [by implication].
Procambarus evermanni.?Hobbs, 1942a:342 [by implication]; 1942b:107,
figs. 111-115.
Procambarus evermani.?Hobbs III, 1969:37 [erroneous spelling].
Procambarus (Ortmannicus) evermanni.?Hobbs, 1972a: 10; 1974b:55, fig.
237.
TYPE.?Holotype, MCZ 3834 (male I).
TYPE LOCALITY.?Escambia River at Flomaton, Escambia
County, Alabama.
RANGE.?In addition to the type locality, known from the
Yellow River basin in Santa Rosa County, Florida, to the Pearl
River basin in Jackson County, Mississippi.
HABITAT.?Sluggish streams. Specimen from Mississippi
dug from simple burrows in drying creek bed.
Procambarus (Ortmannicus) fallax (Hagen)
FIGURE 295
Cambarus fallax Hagen, 1870:45. figs. 103-105.
Cambarus Fallax.?Hagen, 1870:45.
Cambarus (Cambarus) fallax.?Ortmann, 19O5c:102.
Cambarus (Ortmannicus) fallax.?Fowler, 1912:341 [by implication].?
Creaser, 1934b:4 [by implication].
Procambarus fallax.?Hobbs, 1942a:342 [by implication]; 1942b:lll, figs.
116-120.
Procambarus (Ortmannicus) fallax.?Hobbs, 1972a:9; 1974b:55, fig. 235;
1981:416, fig. 14e. 136e, 138e, 156,164,248.
Procambaris fallax.?Wharton, 1978:46 [erroneous spelling].
TYPES.?Syntypes, MCZ 3526 (male I, male II); USNM
63294 (male II, female), formerly in the Boston Society of
Natural History.
TYPE LOCALITY.?Florida. Restricted by Hobbs (1974b:55)
to St. Johns River at Welaka, Putnam County, Florida.
RANGE.?Tributaries of the Satilla River, Georgia, south-
ward through peninsular Florida.
HABITAT.?Lentic and lotic situations and burrows.
Procambarus (Ortmannicus) franzi Hobbs and Lee
FIGURE 287
Procambarus (Ortmannicus) franzi Hobbs and Lee, 1976:384, fig. 1.?Franz
and Lee, 1982:59, figs. 2,6.
TYPES.?Holotype, allotype, and morphotype, USNM
146992,146993,146994; paratypes, USNM.
TYPE LOCALITY.?Orange Lake Cave, 0.4 mile (0.6 km)
south of junction of U.S. Highway 441 and State Route 318 off
Highway 411 (Sec. 33/34, T. 12S, R. 21E), Marion County,
Florida.
RANGE.?Known only from the type locality and Hell Hole
(Sec. 6,T. 12S.R. 21E),also in Marion County, Florida.
HABITAT.?Subterranean waters.
Procambarus (Ortmannicus) geminus Hobbs
FIGURE 305
Procambarus (Ortmannicus) geminus Hobbs, 1975:1, fig. 1.
Procambarus geminus.?Bouchard and Robison, 1981:26.
TYPES.?Holotype, allotype, and morphotype, USNM
145756, 145757, 145758 (male I, female, male II); paratypes,
BMNH,USNM,H.W. Robison.
TYPE LOCALITY.?"Roadside pool (perhaps a streambed.but
no flow was detected), 1.7 miles [2.7 km] south of Taylor,
Columbia County, Arkansas, on State Route 132" (Hobbs,
1975:4-5).
RANGE.?Red River Basin in Columbia, Lafayette, and
Miller counties, Arkansas, and Webster and Sabinc parishes,
Louisiana.
HABITAT.?Lentic and sluggish lotic situations.
Procambarus (Ortmannicus) gonopodocristatus Villalobos
FIGURE 324
Procambarus gonopodocristatus Villalobos, 1958:279, pis. 1-3; 1959:314,
figs. 38-45.
Procambarus (Ortmannicus) gonopodocristatus.?Hobbs, 1972a:10;
1974b:56, fig. 260.
TYPES.?Holotype, allotype, and morphotype, IBM (male I,
female, male II); paratypes, USNM, AMNH.
TYPE LOCALITY.?Paso Largo, 16 kilometers (10 mi)
northeast of Martinez de la Torre, Veracruz, Mexico.
RANGE.?Only one other locality known: "Maria de la
Torre, km. 406 de la carretera Martinez de la Torre, Nautla,"
Veracruz, Mexico (Villalobos, 1958:287).
HABITAT.?Lentic situations.
Procambarus (Ortmannicus) hayi (Faxon)
FIGURE 306
Cambarus Hayi Faxon, 1884:108.
Cambarus hayi.?Usy, 1899b:959.
Cambarus (Cambarus) hayi.?Ortmann, 1905c: 102.
Cambarus (Ortmannicus) hayi.?Fowler, 1912:341 [by implication].?
Creaser, 1934b:4 [by implication].
Procambarus hayi.?Hobbs, 1942a:342 [by implication].
Procambarus (Ortmannicus) hayi.?Hobbs, 1972a:9; 1974b:56,fig. 244.
Procambarus layi.?Hazlett, 1983:187 [erroneous spelling].
TYPES.-^Syntypes, MCZ 3533 (male I, 2 male II, female,
juv. male), USNM 19751 (male II), USNM 19752 (male II,
female), USNM 21850 (2 male I, female).
TYPE LOCALITY.?Macon, Noxubee County, Mississippi.
RANGE.?Tombigbee and Tallahatchie river basins in Ala-
bama and Mississippi, and the Hatchie drainage in Tennessee.
HABITAT.?Sluggish streams and lentic situations.
NUMBER 480 67
Procambarus (Ortmannicus) hirsutus Hobbs
FIGURE 280
Procambarus hirsutus Hobbs, 1958a: 160, figs. 1-12
Procambarus sp. B.?Hobbs, 1958c:72,76,77,79,84,86,90.
Procambarus (Ortmannicus) hirsutus.?Hobbs, 1972a:9; 1974b:56, fig. 224;
Hobbs III.Thorp, and Anderson, 1976:35, figs. 15,22.
Procambararus (Ortmannicus) hirsutus.?Hobbs III, Thorp, and Anderson,
1976:12 [erroneous spelling].
Procambarus (Ortmanicus) hirsutus.?Hobbs III, Thorp, and Anderson,
1976:59 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
101148,101150,1001149 (male I, female, male II); paratypes,
TU.Chas. Mus.,USNM.
TYPE LOCALITY.?Salkehatchie River, 1.9 miles (3 km)
south of Barnwell, Barnwell County, South Carolina, on State
Route 3.
RANGE.?Santee, Edisto, Salkehatchie, and Savannah drain-
age systems in South Carolina.
HABITAT.?Streams.
Procambarus (Ortmannicus) horsti Hobbs and Means
FIGURE 289
Procambarus horsti Hobbs and Means, 1972:401, fig. 2.?Hobbs, 1974b:56,
fig. 230.?Caine, 1974:11, figs. 2a, 4c, 5b.?Franz and Lee, 1982:61, fig. 4.
Procambarus (Ortmannicus) horsti?Holt, 1973:246.?Hobbs, 1974b:56, fig.
230.?Hobbs, Hobbs, and Daniel. 1977:126, fig. 58.
TYPES.?Holotype, allotype, and morphotype, USNM
132043,132044, 132045 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Big Blue Springs (tributary to the
Wacissa River); 2.2 miles (3.5 km) south of the crossroads in
the town of Wacissa, Jefferson County, Florida (SE 'A, NW V*,
Sec. 12, T. 2S.R. 3E).
RANGE.?In addition to the type locality, known from
Shepard Spring, St. Marks Wildlife Refuge, Wakulla County,
and from one locality in Leon County, Florida.
HABITAT.?Subterranean waters.
Procambarus (Ortmannicus) jaculus Hobbs and Walton
FIGURE 310
Cambarus (Ortmannicus) cookae Lyle, 1938:76 [nomen nudum].
Procambarus jaculus Hobbs and Walton, 1957:48, figs. 24-34.?Walls and
Black, 1967:60.
Procambarus (Ortmannicus) jaculus.?Hobbs, 1972a:10; 1974b:56,fig. 248.
TYPES.?Holotype, allotype, and morphotype, USNM
99584, 99585, 99586 (male I, female, male II); paratype,
USNM.
TYPE LOCALITY.?Roadside ditch 1 mile (1.6 km) west of
Scott-Rankin county line on U.S. Highway 80,Rankin County,
Mississippi.
RANGE.?Black and Pearl river basins in Attala, Carroll,
Hinds, Montgomery, Rankin, and Scott counties, Mississippi,
and Avoyelles Parish, Louisiana.
HABITAT.?Temporary lentic situations and burrows (secon-
dary burrower).
Procambarus (Ortmannicus) lecontei (Hagen)
FIGURE 304
Cambarus LeContei Hagen, 1870:47,figs. 15-18,145.
Cambarus Lecontei.?Faxon, 1884:110.
Cambarus lecontei.?flay, 1899b:959.
Cambarus (Cambarus) lecontei.?Ortmann, 1905c:102.
Cambarus (Ortmannicus) lecontei.?Fowler, 1912:341 [by implication].?
Creaser, 1934b:4 [by implication].
Procambarus lecontei.?Hobbs, 1942a:342 [by implication]; 1952a:210, fig.
81.
Procambarus (Ortmannicus) lecontei.?Hobbs, 1972a:9; 1974b:57,fig. 243.
TYPES.?Syntypes, MCZ 217 (male 1,7 female, 2 male IT),
USNM 4958 (male I), AMS (lost), MHNP (male I, female),
Wiirzburg Mus., ZIAS 1/44966 (2 specimens).
TYPE LOCALITY.?Mobile County, Alabama.
RANGE.?Pascagoula and Mobile river basins in Mobile and
Washington counties, Alabama, and George, Jackson, and
Stone counties, Mississippi.
HABITAT.?Streams.
Procambarus (Ortmannicus) hybus Hobbs and Walton
FIGURE 311
Procambarus hybus Hobbs and Walton, 1957:39, figs. 1-12.
Procambarus (Ortmannicus) hybus.?Hobbs, 1972a:10; 1974b:56, fig. 249.
TYPES.?Holotype, allotype, and morphotype, USNM
99581,99582,99583 (male I, female, male II); paratypes, TU,
USNM.BMNH.
TYPE LOCALITY.?Roadside ditch 1.7 miles (2.7 km) north
of Boligee, Greene County, Alabama.
RANGE.?Lower Alabama and Tombigbee drainages in
Alabama and Mississippi, and in the Yalobusha River basin in
Calhoun and Lee counties, Mississippi.
HABITAT.?Temporary ponds, pools, and burrows (secon-
dary burrower).
Procambarus (Ortmannicus) leitheuseri Franz and
Hobbs
FIGURE 283
Procambarus (Ortmannicus) leitheuseri Franz and Hobbs, 1983:323, fig. 1.
TYPES.?Holotype, allotype, and morphotype, USNM
178361, 178362, 178585 (male I, female, male II); paratypes,
USNM.FSM.
TYPE LOCALITY.?"Eagle's Nest" (= Lost Sink or Eagle
Hole), 4.0 miles (5.4 km) northwest of the junction of U.S.
Highway 19 and State Road 50 (NE V4,NE 74, SW V4,Sec. 21,
T. 22S, R. 17E), Hernando County, Florida.
RANGE.?Known from only six localities in Hernando and
Pasco counties, Florida.
HABITAT.?Subterranean waters.
68 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Procambarus (Ortmannicus) leonensis Hobbs
FIGURE 296
Procambarus leonensis Hobbs, 1942b: 114, figs. 121-125; 1943a:49,figs. 1,6,
7,10,12-14,16.17,22,26,29,31.?Caine. 1974:9, figs, lc, 4c, 5?.
Procambarus (Ortmannicus) leonensis.?Hobbs, 1972a:9; 1974b:57, fig. 236.
TYPES.?Holotype, allotype, and "morphotype," USNM
81091 (male I, female, male II); paratypes, MCZ, USNM,
ANSP.
TYPE LOCALITY.?Sinkhole pond, 3 miles (4.8 km) south-
west of Tallahassee on State Route 20, Leon County, Florida
(now beneath runway at airport).
RANGE.?Between the Apalachicola and Suwannee rivers,
Florida.
HABITAT.?Lentic and lotic situations, and burrows.
Procambarus (Ortmannicus) lepidodactylus Hobbs
FIGURE 278
Procambarus lepidodactylus Hobbs, 1947b:25, figs. 1-12.
Procambarus (Ortmannicus) lepidodactylus.?Hobbs, 1972a:9; 1974b:57, fig.
222.
TYPES.?Holotype, and allotype, USNM 84198 (male I,
female); morphotype, USNM 84199 (male II); paratypes,
USNM.
TYPE LOCALITY.?Juniper Creek, 1 mile (1.6 km) southwest
of Patrick on U.S. Highway 1, Chesterfield County, South
Carolina.
RANGE.?Santee and Pee Dee drainage systems in eastern
South Carolina and Columbus County, North Carolina.
HABITAT.?Streams.
Procambarus (Ortmannicus) lewisi Hobbs and Walton
FIGURE 318
Procambarus lewisi Hobbs and Walton, 1959:39, figs. 1-12.
Procambarus (Ortmannicus) lewisi?Hobbs, 1972a:10; 1974b:57, fig. 254.
TYPES.?Holotype, allotype, and morphotype, USNM
102467, 102468, 102469 (male I, female, male II); paratypes,
TU.USNM.
TYPE LOCALITY.?Roadside ditch, 18.8 miles (30 km) east
of Montgomery on U.S. Highway 80, Macon County,
Alabama.
RANGE.?Barbour, Bulloch, Lowndes, Macon, Montgom-
ery, and Russell counties, Alabama.
HABITAT.?Sluggish streams and roadside ditches.
Procambarus (Ortmannicus) titosternum Hobbs
FIGURE 277
Procambarus litosternum Hobbs, 1947a:9, figs. 3 .4 ,9 ,11,13 [not 15], 16,19,
21,26,29,30.
Procambarus (Ortmannicus) litosternum.?Hobbs, 1972a:9; 1974b:57, fig.
221; 1981:403, figs. 13d, 137e, 138g, 158-160,250.
TYPES.?Holotype, allotype, and "morphotype," USNM
82261 (male I, female, male II); paratypes, USNM.
TYPE LOCALITY.?Stream, 5 miles (8 km) northeast of
Swainsboro on U.S. Highway 25, Emanuel County, Georgia.
RANGE.?Canoochee, Ogeechee, Newport, and Savannah
river basins in southeastern Georgia.
HABITAT.?Streams.
Procambarus (Ortmannicus) lophotus Hobbs and Walton
FIGURE 307
Procambarus lophotus Hobbs and Walton, 1960:123, figs. 1-11.
Procambarus (Ortmannicus) lophotus.?Hobbs, 1972a:9; 1974b:58, fig. 245;
1981:379, figs. 2,15d, 136c, 138h, 146,147,251.
TYPES.?Holotype, allotype, and morphotype, USNM
104404, 104405, 104406 (male I, female, male II); paratypes,
TU.USNM.
TYPE LOCALITY.?Roadside ditch, 3.4 miles (5.4 km)
northeast of Haynesville on State Route 111, Lowndes County,
Alabama.
RANGE.?Alabama drainage system from Clarke County,
Alabama, to Chattooga and Whitficld counties, Georgia, and
the Tennessee river basin in Walker and Catoosa counties,
Georgia, and Polk County, Tennessee.
HABITAT.?Lentic and lotic situations and burrows (tertiary
burrower).
Procambarus (Ortmannicus) lucifugus alachua (Hobbs)
FIGURE 284
Cambarus lucifugus alachua Hobbs, 1940a:402,fig. 18a-l.
Procambarus lucifugus alachua.?Hobbs, 1942a:343.?Franz and Lee,
1982:60, figs. 1,6.
Procambarus (Ortmannicus) lucifugus alachua.?Hobbs, 1972a:10; 1974b:58,
fig. 227.?Hobbs, Hobbs, and Daniel, 1977:130, figs. 4,60.
TYPES.?Holotype, allotype, and "morphotype," USNM
76592 (male I, female, male II); paratypes, USNM, FSBC.
TYPE LOCALITY.?Small cave, Hog Sink, about 10 miles (16
km) west of Gainesville, Alachua County, Florida (Sec. 24, T.
10S.R. 18E).
RANGE.?Southwestern Alachua, western Gilchrist, and
northern and eastern Levy counties, Florida; intergrading with
P. (0.) lucifugus lucifugus in Marion County.
HABITAT.?Subterranean waters.
Procambarus (Ortmannicus) lucifugus lucifugus (Hobbs)
FIGURE 285
Cambarus acherontis.? Faxon, 1898:645, pi. 62: figs. 1-5.
Cambarus lucifugus lucifugus Hobbs, 194Oa:398,figs. 17,18m,n.
Procambarus lucifugus lucifugus.?Hobbs, 1942a:343.?Franz and Lee,
1982:60, figs. 1.6.
Procambarus (Ortmannicus) lucifugus lucifugus.?Hobbs, 1972a:9.?Hobbs,
Hobbs, and Daniel, 1977:128, fig. 59.
TYPES.?Holotype, allotype, and "morphotype," USNM
NUMBER 480 69
77916, 77917, 77918 (male I, female, male II); paratypes,
MCZ, USNM, FSBC, OSM.
TYPE LOCALITY.?Gum Cave, about 7 miles (11.2 km)
southwest of Floral City, Citrus County, Florida (NW 1/A, NE
74, Sec. 36.X 20S,R. 19E).
RANGE.?Citrus and Hernando counties northward to Ma-
rion County Florida, where it intergrades with P. (O.) lucifugus
alachua.
HABITAT.?Subterranean waters.
Procambarus (Ortmannicus) lunzi (Hobbs)
FIGURE 294
Cambarus lunzi Hobbs, 194Ob:3,figs. 1-10.
Procambarus lunzi.?Hobbs, 1942a:343.
Procambarus (Ortmannicus) lunzi.?Hobbs, 1972a:9; 1974b:58, fig. 234;
1981:424, figs. 14c, 136d, 138i, 166-168,252.
TYPES.?Holotype and allotype, USNM 79301 (male I,
female).
TYPE LOCALITY.?1.4 miles (2.2 km) southeast (not south-
west) of Early Branch on State Route 68, Hampton County,
South Carolina.
RANGE.?Lower coastal plain from Aiken, Bamberg, and
Colleton counties, South Carolina, southward to Mclntosh
County, Georgia (between the Edisto and Satilla rivers).
HABITAT.?Sluggish lotic and lentic situations and burrows
(secondary or tertiary burrower).
Procambarus (Ortmannicus) mancus Hobbs and Walton
FIOURE313
Procambarus mancus Hobbs and Walton, 1957:44, figs. 13-23.
Procambarus (Ortmannicus) mancus.?Hobbs, 1972a:10; 1974b:58,fig. 251.
TYPES.?Holotype, allotype, and morphotype, USNM
99578,99579,99580 (male I, female, male II); paratypes, TU,
USNM.
TYPE LOCALITY.?Roadside ditch, 5 miles (8 km) south of
Meridian, Lauderdale County, Mississippi.
RANGE.?Pascagoula and Tombigbee river basins of Missis-
sippi between the 31st and 34th parallels.
HABITAT.?Burrows (secondary burrower).
Procambarus (Ortmannicus) marthae Hobbs
FIGURE 317
Procambarus (Ortmannicus) marthae Hobbs, 1975:6, fig. 2.
TYPES.?Holotype, allotype, and morphotype, USNM
145994, 145995, 145996 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Small, very sluggish stream, 2.3 miles
(3.7 km) west of the Alabama River on U.S. Highway 84,
Monroe County, Alabama.
RANGE.?Alabama River basin in Dallas, Hale, Monroe, and
Perry counties, Alabama.
HABITAT.?Lentic and sluggish lotic situations.
Procambarus (Ortmannicus) medialis Hobbs
FIGURE 315
Procambarus (Ortmannicus) medialis Hobbs, 1975:10, figs. 3,4.
TYPES.?Holotype, allotype, and morphotype, USNM
144942, 144943, 144944 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Pool in roadside ditch, 0.6 mile (1 km)
south of Scotland Neck, Halifax County, North Carolina, on
U.S. Highway 258.
RANGE.?Neuse and Pamlico river basins in the coastal plain
of North Carolina.
HABITAT.?Lentic and sluggish lotic situations.
Procambarus (Ortmannicus) orcinus Hobbs and Means
FIGURE 290
Procambarus orcinus Hobbs and Means, 1972:394, fig. 1.?Caine, 1974:14,
figs. 2b, 4f, 5b.?Franz and Lee, 1982:61, fig. 4.
Procambarus (Ortmannicus) orcinus.?Hobbs, 1974b:58, fig. 231.?Hobbs,
Hobbs, and Daniel, 1977:132, fig. 61.
TYPES.?Holotype, allotype, and morphotype, USNM
132031, 132032, 132033 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Gopher Sink, 3.1 miles (5 km) southwest
of State Route 61 and 0.2 mile (0.3 km) east of State Route 369
(SW 74, NW 74, NE 74, Sec. 16, T.2S, R. 1W), Leon County,
Florida.
RANGE.?Known from four localities in Leon and Wakulla
counties, Florida.
HABITAT.?Subterranean waters.
Procambarus (Ortmannicus) pallidus (Hobbs)
FIGURE 288
Cambarus acherontis.?Hobbs, 1937:154.
Cambarus acherontis pallidus Hobbs, 1938b:90 [nomen nudum].
Cambarus pallidus Hobbs, 1940a:394, fig. 16.
Procambarus pallidus.?Hobbs, 1942a:343.
procambarus pallidus.?Padgett, 1970:19 [lapsus calami].
Procambarus (Ortmannicus) pallidus.?Hobbs, 1972a:10; 1974b:59, fig.
229.?Hobbs, Hobbs, and Daniel, 1977:132, figs. 4, 62.?Franz and Lee,
1982:61, fig. 4.
Procambrus pallidus.?Relyea and Sutton, 1975a:174, fig. 1 [erroneous
spelling].
TYPES.?Holotype and allotype, USNM 76591 (male I,
female); "morphotype," USNM 79350 (male II); paratypes,
MCZ, USNM, FSBC.
TYPE LOCALITY.?Warren's Cave, 11 miles (17.6 km)
northwest of Gainesville, Alachua County, Florida (Sec. 13, T.
9S.R. 18E).
70 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
RANGE.?Subterranean waters from the northwestern part of
Alachua and northeastern Levy counties to the Suwannee River
basin in southern Suwannee County, Florida.
HABITAT.?Subterranean waters.
Procambarus (Ortmannicus) pearsei (Creaser)
FIGURE 316
Cambarus (Ortmannicus) pearsei Creaser, 1934b: 1, figs. 1-3.?Hobbs,
1975:1 [by implication].
Procambarus pearsei.?Hobbs, 1942a:343.
Procambarus pearsei pearsei.?Hobbs and Walton, 1958:7.
Procambarus (Ortmannicus) pearsei pearsei.?Hobbs, 1972a:10; 1974b:59,
fig. 253.
Procambarus (Ortmannicus) pearsei.?Hobbs, 1975:15.
TYPES.?Holotype and allotype, USNM 98336,98337 (male
I, female); paratypes, USNM.
TYPE LOCALITY.?Pond and ditch on State Route 22, south
of Fayetteville, Cumberland County, North Carolina.
RANGE.?Johnston and Samson counties, North Carolina,
south to Horry and Marion counties, South Carolina.
HABITAT.?Lentic situations and burrows (secondary bur-
rower).
Procambarus (Ortmannicus) pictus (Hobbs)
FIGURE 273
Cambarus pictus Hobbs, 1940a:419,fig. 22.
Procambarus pictus.?Hobbs, 1942a:344.
Procambarus (Ortmannicus) pictus.?Hobbs, 1972a:9; 1974b:59, fig. 217.?
Franz and Franz, 1979:13, fig. 1.
TYPES.?Holotype, allotype, and "morphotype," USNM
76596 (male I, female, male II); paratypes, MCZ, USNM.
TYPE LOCALITY.?Small, swift swamp stream, about 2 miles
(3.2 km) southwest of Green Cove Springs, Clay County,
Florida. This is probably Peters Creek at State Route 315, Sec.
30,T.6S,R. 26E. (See Franz and Franz, 1979:14.)
RANGE.?Known from several small tributaries of Black
Creek in Clay County, and tributary to the St. Johns River in
Duval County, Florida.
HABITAT.?Streams.
Procambarus (Ortmannicus) planirostris Penn
FIGURE 312
Procambarus planirostris Penn, 1953c:71,figs. 1-12.
Procambarus (Ortmannicus) planirostris.?Hobbs, 1972a: 10; 1974b:59, fig.
250.
Procambarus plantiuoslrusg.?Huner, Meyers, and Avault, 1976:152 [errone-
ous spelling].
Procambarus planirostrus.?Huner, Meyers, and Avault, 1976:156 [erroneous
spelling].
Procambarus plantirostrus.?Huner and Barr, 1980:146 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
95674, 95675, 95676 (male I, female, male II); paratypes,
ANSP,TU,USNM.
TYPE LOCALITY.?Flatwoods, 1 mile (1.6 km) south of
Walker on State Route 336, Livingston Parish, Louisiana.
RANGE.?"Florida" parishes of southeastern Louisiana, and
Harrison, Jones, Jackson, Perry, Pearl River, and Winston
counties, Mississippi.
HABITAT.?Temporary ponds, pools, and burrows (secon-
dary burrower).
Procambarus (Ortmannicus) plumimanus Hobbs and
Walton
FIGURE 314
Procambarus pearsei plumimanus Hobbs and Walton, 1958:8, figs. 1-13.
Procambarus (Ortmannicus) pearsei plumimanus.?Hobbs, 1972a:10;
1974b:59.fig. 252.
Procambarus plumimanus.?Hobbs, 1975:1.
Procambarus (Ortmannicus) plumimanus.?Hobbs and Peters, 1977:8.
TYPES.?Holotype, allotype, and morphotype, USNM
100940, 100941, 100942 (male I, female, male II); paratypes,
TU, USNM.
TYPE LOCALITY.?Roadside ditch 2.2 miles (3.5 km)
southeast of Havelock on U.S. Highway 70, Craven County,
North Carolina.
RANGE.?Carteret, Craven, Duplin, and Jones counties,
North Carolina.
HABITAT.?Temporary ponds, pools, and burrows (secon-
dary burrower).
Procambarus (Ortmannicus) pubescens (Faxon)
FIGURE 279
Cambarus pubescens Faxon, 1884:109.
Cambarus (Cambarus) pubescens.?Ortmann, 1905c:101.
Cambarus (Ortmannicus) pubescens.?Fowler, 1912:341 [by implication].
Procambarus pubescens.?Hobbs, 1942a:342 [by implication]; 1947a: 1, figs.
1,6-8,15 [not 14], 17,22,27,28.31.
Procambarus (Ortmannicus) pubescens.?Hobbs, 1972a:9; 1974b:60, fig. 223;
1981:408, figs. 13b, 137f, 138j, 161-163,253.
Procambarus (Ortmanicus) pubescens.?Hobbs III, Thorp, and Anderson,
1976:59 [erroneous spelling].
TYPES.?Syntypes, USNM 3181 (male II, female), MCZ
3551 (2 female).
TYPE LOCALITY.?McBean Creek, south of Augusta, Burke-
Richmond County line, Georgia.
RANGE.?The Oconee, Ogeechee, and Savannah river
systems in eastern Georgia and South Carolina.
HABITAT.?Streams.
Procambarus (Ortmannicus) pyenogonopodus Hobbs
FIGURE 299
Procambarus pyenogonopodus Hobbs, 1942b: 117, figs. 126-130; 1943a:53,
figs. 3,4,8,9,11,15,19-21,24,25,27,28,30,32.
Procambarus (Ortmannicus) pyenogonopodus.?Hobbs, 1972a:9; 1974b:60,
fig. 238.
NUMBER 480 71
TYPES.?Holotype, allotype, and "morphotype," USNM
81092 (male I, female, male II); paratypes, MCZ, ANSP,
USNM.
TYPE LOCALITY.?Roadside excavation and adjoining inter-
mittent stream in flatwoods, 5.8 miles (9.3 km) west of
Wewahitchka on State Route 22, Gulf County, Florida.
RANGE.?From the Apalachicola River westward to the
Choctawhatchee Bay drainage basin in Walton County,
Florida.
HABITAT.?Lentic and lotic situations, and burrows (tertiary
burro wer).
Procambarus (Ortmannicus) seminolae Hobbs
FIGURE 292
Procambarus seminolae Hobbs, 1942b: 142, figs. 166-170,335-346.
Procambarus (Ortmannicus) seminolae.?Hobbs, 1972a:9; 1974b:60, fig. 232;
1981:430, figs. 14b, 136f, 138k, 169-172.254.
Procambaris seminolae.?Wharton, 1978:46 [erroneous spelling].
TYPES.?Holotype, allotype, and "morphotype," USNM
81286 (male I, female, male II); paratypes, MCZ, ANSP,
USNM, UF (now in USNM, FSBC).
TYPE LOCALITY.?Roadside excavation about 9 miles (14.4
km) northeast of Gainesville on State Route 24 (old number
13), Alachua County, Florida (Sec. 32, T. 8S, R. 21E).
RANGE.?From the Altamaha River drainage in Georgia to
Marion County, Florida. A few localities are known in Bryan,
Montgomery, and Toombs counties, Georgia (see Hobbs,
1981:433).
HABITAT.?Lentic and lotic situations, and burrows (secon-
dary or tertiary burrower).
Procambarus (Ortmannicus) texanus Hobbs
FIGURE 303
Procambarus texanus Hobbs, 1971b:87, fig. 2.
Procambarus (Ortmannicus) texanus.?Hobbs, 1972a:9; 1974b:60, fig. 242.
TYPES.?Holotype, allotype, and morphotype, USNM
131457, 131458, 131459 (male I, female, male II); paratypes,
USNM, BMNH.
TYPE LOCALITY.?Fish hatchery near Smithville, Bastrop
County, Texas.
RANGE.?Known only from the type locality.
HABITAT.?Ponds.
Procambarus (Ortmannicus) toltecae Hobbs
FIGURE 323
Procambarus toltecae Hobbs, 1943b: 198, figs. 1-14.?Villalobos, 1959:315,
figs. 53-57.
Procambarus toltecas.?Villalobos, 1953:356 [erroneous spelling].
Procambarus (Ortmannicus) toltecae.?Hobbs, 1972a:10; 1974b:60,fig. 259.
TYPES.?Holotype and allotype, USNM 81301 (male I,
female); paratypes, USNM.
TYPE LOCALITY.?Puente de Xilitla (not Palitla as originally
recorded), small stream 8 km (5 mi) north of Tomazunchale,
San Luis Potosf, Mexico.
RANGE.?Southeastern San Luis Potosf, Hidalgo, and
Tamaulipas.
HABITAT.?Epigean and hypogean streams.
Procambarus (Ortmannicus) verrucosus Hobbs
FIGURE 309
Procambarus verrucosus Hobbs, 1952a:212, fig. 82.
Procambarus (Ortmannicus) verrucosus.?Hobbs, 1972a:9; 1974b:61, fig.
247.
TYPES.?Holotype, allotype, and morphotype, USNM
90743 (male I, female, male II); paratypes, USNM.
TYPE LOCALITY.?Tributary of Calebea Creek, 3.9 miles
(4.6 km) south of Tuskegee, Macon County, Alabama, on U.S.
Highway 29.
RANGE.?Southeastern Alabama from Lee and Macon
counties southward between the Alabama and Chattahoochee
rivers.
HABITAT.?Streams.
Procambarus (Ortmannicus) viaeviridis (Faxon)
FIGURE 320
Cambarus viae-viridis Faxon, 1914:370, pi. 5.
Cambarus (Ortmannicus) viae viridis.?Creaser, 1934b:4 [by implication].
Procambarus viae-viridis.?Hobbs, 1942a:342 [by implication].
Procambarus viaeviridis.?Penn, 1959:5.?Pflieger, 1987a:31; 1987b:15.
Procambarus (Ortmannicus) viaeviridis.?Hobbs, 1972a:9.
TYPES.?Syntypes, MCZ 7336 (2 male I, 2 male II, 6
female).
TYPE LOCALITY.?St. Francis River, Greenway, Clay
County, Arkansas.
RANGE.?From southern Illinois, eastern Arkansas, and
northeastern Louisiana eastward to the northward flowing
segment of the Tennessee River in Kentucky and Tennessee,
and through Mississippi to Tuscaloosa County, Alabama.
HABITAT.?Sluggish streams and lentic situations.
Procambarus (Ortmannicus) villalobosi Hobbs
F I G U R E 322
Procambarus villalobosi Hobbs, 1969a:41,pl. 1: figs. 1-11.
Procambarus (Ortmannicus) villalobosi.?Hobbs, 1972a:10; 1974b:61, fig.
258.
TYPES.?Holotype and allotype, USNM 118644 and 118645
(male I, female).
TYPE LOCALITY.?Cueva del Agua, 75 kilometers (46.9 mi)
east of Valles, San Luis Potosf, Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Streams.
72 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Procambarus (Ortmannicus) xilitlae Hobbs and Grubbs
FIGURE 298
Procambarus (Scapulicambarus) xilitlae Hobbs and Grabbs, 1982:45, fig. 1.
Procambarus (Ortmannicus) xilitlae.?Hobbs and Grabbs, 1986:737, fig. 1.
Procambarus (Ortmannicus) Xilitlae.?Anonymous, 1987b:22 [lapsus].
Procambarus (Ortmannicus) xilitae.?Anonymous, 1987b:22 [erroneous spell-
ing].
TYPES.?Holotype, allotype, and paratype, USNM 177140,
177141,177142 (male II, female, female).
TYPE LOCALITY.?Hoya de las Guaguas, 10 kilometers (6.2
mi) south-southwest of Aquismon, and very near the town of
Xilitla, San Luis Potosf, Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Subterranean waters.
Procambarus (Ortmannicus) youngi Hobbs
FIGURE 281
Procambarus youngi Hobbs, 1942b:131,figs. 146-150,325-334.
Procambarus (Ortmannicus) youngi.?Hobbs, 1972a:9; 1974b:61,fig. 225.
TYPES.?Holotype and allotype, USNM 81289 (male I,
female); "morphotype," USNM 81290 (male II); paratypes,
MCZ,USNM,FSBC.
TYPE LOCALITY.?Guard House Branch, about 1 mile (1.6
km) west of Wewahitchka on State Route 22, Gulf County,
Florida (Sec. 23, T. 4S,R. 10W).
RANGE.?Known from the type locality; Wetappo Creek, 6.4
miles (10 km) east of Bay County line on State Route 22; and
St. Marks River in Leon and Wakulla counties, Florida.
HABITAT.?Streams.
Subgenus Paracambarus Ortmann
Cambarus.?Ortmann, 1906a:3.
Subgenus Paracambarus Ortmann, 1906a:l [Type species, by monotypy,
Cambarus (Paracambarus) paradoxus Ortmann, 1906a:3. Gender: mascu-
line. Proposed as subgenus of Cambarus, elevated to generic rank by Hobbs,
1942a:344 and reduced to subgeneric status in the genus Procambarus by
Hobbs, 1972a:3].?Hobbs, 1984: fig.2.
Paracambarus.?Hobbs, 1942a:344.
Paracamburus.?Villalobos, 1953:354 [erroneous spelling].
Procambarus.?Hobbs, 1972a:3 [in part].
Procambarus (Paracambarus) ortmannii (Villalobos)
FIGURE 325
Paracambarus ortmannii Villalobos, 1949:331, pi. 1: figs. 1-7, pi. 2: figs. 1-7.
Paracambarus ortmanni.?Villalobos, 1953:346.
Procambarus (Paracambarus) ortmanni.?Hobbs, 1972a: 10.
Procambarus (Paracambarus) ortmannii.?Hobbs, 1974b:61, fig. 261.
Procambarus (Paracambarus) ortmanii.?Bouchard, 1978b:452 [erroneous
spelling].
TYPES.?Holotype, allotype, and morphotype, IBM 2-1149-
la (OF6) (male I, female, male II); paratype, IBM.
TYPE LOCALITY.?"Los Estajos, 6 kilometers (3.7 mi) NE
[sic] de Xihuateutla, Pue. en un pequefio arroyito que corre por
el lado derecho del camino que va al rancho El Mirador"
(Villalobos, 1949:338), Mexico.
RANGE.?Known only from the vicinity of the type locality.
HABITAT.?Epigean and hypogean streams.
Procambarus (Paracambarus) paradoxus (Ortmann)
FIGURE 326
Cambarus (Paracambarus) paradoxus Ortmann, 1906a:3, fig. 1.
Paracambarus paradoxus.?Andrews, 1908a:123 [lapsus].?Hobbs,
1942a:344.?Villalobos, 1947a:233, pi. l.pl. 3: figs. 1-4.
Cambarus paradoxus.?Andrews, 1908b: 167.
Cambarus (Paracamburus) paradoxus.?Villalobos, 1953:354 [erroneous
spelling].
Procambarus Paradoxus.?Creaser, 1962:7 [lapsus calami].
Procambarus (Paracambarus) paradoxus.?Hobbs, 1972a: 10, figs. 2j, 13a-g;
1974b:61.fig. 262.
TYPES.?Syntypes, AMS P.5385 (3 specimens), ANSP 5358
(3 specimens), MHNP (25 plus specimens), CM 74.677 (5
specimens), MCZ 6955 (male I, male II, female).
TYPE LOCALITY.?Tetela de Ocampo y La Canada, 35
kilometers (21.9 mi) northeast of Zacapoaxtla, Estado de
Puebla, Mexico.
RANGE.?Known only from the area of the type locality.
HABITAT.?Streams.
REMARKS.?Ortmann (1906a:6) noted that there were
"numerous specimens in the Mus. Paris." Some of these were
subsequently sent to the other museums listed (to the
Australian Museum in September 1921).
Subgenus Pennides Hobbs
Artacus.?LeConte, 1856:401 [in part].
Cambarus.?Hagen, 1870:48 [in part].
Subgenus Cambarus.?Ortmann, 1905c:101 [in part].
Subgenus Ortmannicus.?Fowler, 1912:341 [in part].
Cambaru.?Goodnight, 1941:72 [erroneous spelling].
Procambarus.?Hobbs, 1942a:342 [in part].
Subgenus Pennides Hobbs, 1972a: 10 [Type species, by original designation,
Procambarus natchitochae Penn, 1953d:5. Gender masculine.]; 1984,fig. 2.
Procambarus (Pennides) ablusus Penn
FIGURE 329
Procambarus ablusus Penn, 1963:121, figs. 1-10.
Procambarus (Pennides) ablusus.?Hobbs, 1972a: 10; 1974b:62,fig. 265.
TYPES.?Holotype, allotype, and morphotype, USNM
108200, 108202, 108201 (male I, female, male II); paratypes,
TU.
TYPE LOCALITY.?Hatchie River, 12.1 miles (19.3 km) east
of Ripley on State Route 4, Tippah County, Mississippi.
RANGE.?Hatchie River system in Hardeman, Hardin, and
McNairy counties, Tennessee, and Alcorn, Tippah, and
Tishomingo counties, Mississippi.
HABITAT.?Streams.
NUMBER 480 73
Procambarus (Pennides) clemmeri Hobbs
FIGURE 335
Procambarus (Pennides) clemmeri Hobbs, 1975:19, fig. 6.
Procambarus clemmeri.?Hobbs, 1975:24.
TYPES.?Holotype, allotype, morphotype, USNM 145607,
145608,145609 (male I, female, male II); paratypes,USNM.
TYPE LOCALITY.?Tributary to the Jourdan River, 8.7 miles
(13.9 km) southeast of the Pearl River-Hancock County line on
State Route 43,4.3 miles (6.9 km) northwest of the junction of
State routes 43 and 603 north of Kiln, Hancock County,
Mississippi.
RANGE.?From the Pearl River to the Escatawpa River basin
in southern Mississippi and Alabama.
HABITAT.?Streams.
Procambarus (Pennides) dupratzi Penn
FIGURE 336
Procambarus dupratzi Penn, 1953d:l,figs. 1-7,10-14.
Procambarus (Pennides) dupratzi.?Hobbs, 1972a:10; 1974b:62, fig. 270.
TYPES.?Holotype, allotype, and morphotype, USNM
93652, 93653, 93654 (male I, female, male II); paratypes,
USNM, AMNH, ANSP, MCZ, CM, TU.
TYPE LOCALITY.?Attoyac Bayou, 5.6 miles (8.9 km)
southwest of Timpson on U.S. Highway 59, Shelby County,
Texas.
RANGE.?Drainage systems of the Trinity, Red, Neches,
Sabine, and Calcasieu rivers in Texas, Arkansas, and Louisi-
ana.
HABITAT.?Streams.
Procambarus (Pennides) echinatus Hobbs
FIGURE 334
Procambarus echinatus Hobbs, 1956a: 117, figs. 1-17.
Procambarus (Pennides) echinatus.?Hobbs, 1972a:10; 1974b:62,fig. 269.
TYPES.?Holotype, allotype, and morphotype, USNM
99180,99181,99182 (male I, female, male II); paratypes, TU,
USNM.
TYPE LOCALITY.?Salkehatchie River, 1.9 miles (3 km)
south of Barn well on State Route 3, Barnwell County, South
Carolina.
RANGE.?Salkehatchie and Edisto drainage systems in
South Carolina.
HABITAT.?Streams.
Procambarus (Pennides) elegans Hobbs
FIGURE 331
Procambarus elegans Hobbs, 1969c:329, figs. 1-11,24,25.
Procambarus (Pennides) elegans.?Hobbs, 1972a: 10; 1974b:62,fig. 266.
TYPES.?Holotype, allotype, and morphotype, USNM
129892, 129893, 115838 (male I, female, male II); paratype,
USNM.
TYPE LOCALITY.?Spillway from Corney Lake, an im-
pounded tributary of the Ouachita River, Claiborne Parish,
Louisiana.
RANGE.?Ouachita drainage system in southern Arkansas
and Claiborne and Union parishes, Louisiana, and in the Pearl
River basin in Montgomery and Simpson counties, Mississippi.
HABITAT.?Streams.
Procambarus (Pennides) gibbus Hobbs
FIGURE 342
Procambarus gibbus Hobbs, 1969c:337, figs. 12-23.
Procambarus (Pennides) gibbus.?Hobbs, 1972a:10; 1974b:62, fig. 275;
1981:440, figs. 12e, 173f, 174,175,183a, 255.
TYPES.?Holotype, allotype, and morphotype, USNM
129804, 129803, 129805 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Tributary to Muckalee Creek (Flint River
drainage), 3.2 miles (5.1 km) north of Americus on U.S.
Highway 19, Sumter County, Georgia.
RANGE.?Flint River drainage system in Baker, Crawford,
Lee, Marion, Schley, and Sumter counties, Georgia.
HABITAT.?Streams.
Procambarus (Pennides) lagniappe Black
FIGURE 339
Procambarus lagniappe Black, 1968:5, figs. 1-12.
Procambarus (Pennides) lagniappe.?Hobbs, 1972a:10; 1974b:62,fig. 273.
TYPES.?Holotype, allotype, and morphotype, USNM
119088, 119089, 119090 (male I, female, male II); paratypes,
TU, Joe B. Black.
TYPE LOCALITY.?Pawticfaw Creek (tributary to Sucar-
noochee Creek, tributary to Tombigbee River), 6.0 miles (9.6
km) south of De Kalb, Kemper County, Mississippi, on State
Route 39.
RANGE.?Tributaries of the Tombigbee River in Kemper
County, Mississippi.
HABITAT.?Streams.
Procambarus (Pennides) lylei Fitzpatrick and Hobbs
FIGURE 333
Procambarus lylei Fitzpatrick and Hobbs, 1971 ^5, figs. 1-14.
Procambarus (Pennides) lylei.?Hobbs, 1972a: 10; 1974b:63,fig. 268.
TYPES.?Holotype, allotype, and morphotype, USNM
131533, 131534,131535 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Shutispear Creek, 2.7 miles (4.3 km)
south of State Route 8 on State Route 9, Calhoun County,
Mississippi.
74 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
RANGE.?Known from only a few localities in the Yalo-
busha River drainage in Calhoun County, Mississippi.
HABITAT.?Streams.
Procambarus (Pennides) natchitochae Penn
FIGURE 328
Procambarus natchitochae Penn, 1953d:5,figs. 8,9,15-19.?Huner, 1977:11.
Procambarus (Pennides) natchitochae.?Hobbs, 1972a: 10, figs. 2k, 14a-g;
1974b:63,fig. 264.
TYPES.?Holotype, allotype, and morphotype, USNM
93649, 93650, 93651 (male I, female, male II); paratypes,
USNM, AMNH, ANSP, TU.
TYPE LOCALITY.?Tributary of Spring Creek at Melder, on
State Route 85,Rapides Parish, Louisiana.
RANGE.?Tributaries of Bayou Teche, Red, and Calcasieu
rivers in Louisiana, Arkansas, and Texas.
HABITAT.?Streams.
Procambarus (Pennides) ouachitae Penn
FIGURE 343
Procambarus vioscae.?Williams, 1954:819 [erroneous spelling for Procam-
barus vioscai].
Procambarus ouachitae Penn, 1956a: 109, figs. 1-17.
Procambarus quachitae.?Penn, 1956a:113 [erroneous spelling].
Procambarus ouichitae.?Hobbs, 1968b:K26 [erroneous spelling].
Procambarus (Pennides) ouachitae.?Hobbs, 1972a:10; 1974b:63, fig. 276.
TYPES.?Holotype, allotype, and morphotype, USNM
99222, 99223, 99224 (male I, female, male II); paratypes,
USNM, ANSP, AMNH, CM.
TYPE LOCALITY.?Tributary of the South Fork of the Saline
River, 2.4 miles (3.8 km) north of U.S. Highway 70 on a local
road 6.5 miles (10.4 km) northeast of Hot Springs (7.7 miles
(12.3 km) south-southwest of Owensville), Garland County,
Arkansas.
RANGE.?Ouachita and Arkansas river systems in south-
western Arkansas, and the Yazoo Basin in Mississippi.
HABITAT.?Streams.
Procambarus (Pennides) penni Hobbs
FIGURE 338
Procambarus penniHobbs, 1951 b:273, figs. 1-11.
Procambarus (Pennides) penni.?Hobbs, 1972a:10; 1974b:63,fig. 272.
TYPES.?Holotype, allotype, and morphotype, USNM
91662,91663,91664 (male I, female, male II); paratypes, TU,
USNM.
TYPE LOCALITY.?Talisheek Creek, at Talisheek, St. Tam-
many Parish, Louisiana.
RANGE.?Pearl and Pascagoula drainage systems in eastern
Louisiana and south-central Mississippi.
HABITAT.?Streams.
Procambarus (Pennides) petersi Hobbs
FIGURE 340
Procambarus spiculifer.?Hart and Hart, 1974:21.
Procambarus (Pennides) petersi Hobbs, 1981:442, figs. 193d, 176.
Procambarus petersi.?Hobbs, 1981:449.
TYPES.?Holotype, allotype, and morphotype, USNM
144960, 144961, and 144962 (male I, female, male II);
paratypes, BMNH, RNHL, USNM.
TYPE LOCALITY.?Rocky Comfort Creek, 1.7 miles (2.7 km)
north of Gibson, Glascock County, Georgia, on Route S2126,
and 0.4 mile (0.64 km) east on Chalker Smith Road.
RANGE.?Ogeechee River basin in Georgia.
HABITAT.?Streams.
Procambarus (Pennides) raneyi Hobbs
FIGURE 344
Procambarus raneyi Hobbs, 1953b:412,figs. 1-13.
Procambarus rayeni.?Anonymous, 1967a, table 3 in Appendix [erroneous
spelling].
Procambarus (Pennides) raneyi.?Hobbs, 1972a:10; 1974b:63, fig. 277;
1981:449, figs. 12c, 173c, 175,178,183c, 257.
Procambarus ranei.?K. Brown, 1981:225 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, USNM
95124, 95125, 95126 (male I, female, male II); paratypes,
MCZ,TU,USNM.
TYPE LOCALITY.?South fork of the Broad River, 1 mile (1.6
km) south of Carlton on the Oglethorpe-Madison county line,
Georgia.
RANGE.?Savannah River drainage in Georgia and South
Carolina and headwaters of the Ocmulgee River in DeKalb,
Gwinnett, Newton, and Walton counties, Georgia.
HABITAT.?Streams.
Procambarus (Pennides) roberti Villalobos and Hobbs
FIGURE 330
Procambarus (Pennides) roberti Villalobos and Hobbs, 1974:8, fig. 2.
Procambarus roberti.?Villalobos and Hobbs, 1974:2.
TYPES.?Holotype, allotype, and morphotype, USNM
132345, 132346, 133826 (male I, female, male II); paratypes,
IBM.
TYPE LOCALITY.?"Ditch from La Media Luna, 4.8 miles
(7.7 km) south of Rio Verde (on highway to Pedro Montoya)
and 2.5 miles (4 km) west on dirt road to Mina El Refugio, San
Luis Potosi, Mexico" (Villalobos and Hobbs, 1974:6).
RANGE.?Known only from the vicinity of the type locality.
HABITAT.?Streams.
Procambarus (Pennides) spiculifer (LeConte)
FIGURE 341
Astacus spiculifer LeConte, 1856:401.
Cambarus spiculifer.?Hagen, 1870:48, figs. 59-62,147.
NUMBER 480 75
Cambarus (Cambarus) spiculifer.?Ortmaim, 1905c: 101.
Cambarus (Ortmannicus) spiculifer.?Fowler, 1912:341 [by implication].?
Creaser, 1934b:4 [by implication].
Cambaru spiculifer.?Goodnight, 1941:72 [erroneous spelling].
Procambarus spiculifer.?Hobbs, 1942a:342 [by implication]; 1942b: 119,
figs. 131-135.
Procambarus (Pennides) spiculifer.?Hobbs, 1972a:10; 1974b:64, fig. 274;
1981:453, figs. 12d, 173a,b, 177,179,180,183d, 258.
Procambarus spiculifier.?Bouchard, 1978b:445 [erroneous spelling].
TYPES.?Syntypes, ANSP 316 (male II), MCZ 172 (2 male
1,4 female, 2 juv. male, 3 juv. female), MCZ 3376 (male I),
USNM 4962 (specimens decayed), MHNP (male I, female).
TYPE LOCALITY.?Athens, Clarke County, Georgia.
RANGE.?From the upper Savannah Basin southward to the
SL Marys and Suwannee river systems in Florida and westward
to the Alabama-Mobile River basin.
HABITAT.?Streams.
Procambarus (Pennides) suttkusi Hobbs
FIGURE 327
Procambarus suttkusi Hobbs, 1953a:173,pl. 9.
Procambarus suttkuzi.?Hobbs, 1968b:Kll [erroneous spelling].
Procambarus (Pennides) suttkusi.?Hobbs, 1972a:10; 1974b:64,fig. 263.
TYPES.?Holotype, allotype, and morphotype, USNM
93730, 93731, 93732 (male I, female, male II); paratypes,
USNM.TU.BMNH.
TYPE LOCALITY.?Tributary of Claybank Creek, 2 miles
(3.2 km) west of Ozark, Dale County, Alabama.
RANGE.?Choctawhatchee drainage system in Alabama and
northern Florida.
HABITAT.?Streams.
Procambarus (Pennides) versutus (Hagen)
FIGURE 332
Cambarus versutus Hagen, 1870:51, figs. 55-58,150.
Cambarus (Cambarus) versutus.?Ortmann, 1905c: 101.
Cambarus (Ortmannicus) versutus.?Fowler, 1912:341 [by implication].?
Creaser, 1934b:4 [by implication].
Procambarus versutus.?Hobbs, 1942a:342 [by implication]; 1942b: 126, figs.
136-140; 1981:463,figs. 12f, 173e, 175,181,183e,259.
Procambarus (Pennides) versutus.?Hobbs, 1972a:10; 1974b:64, fig. 267.
TYPES.?Syntypes, MCZ 190 (3 male II, female), USNM
4963 (male I), MHNP (2 specimens), AMS (no longer there).
TYPE LOCALITY.?Spring Hill, Mobile County, Alabama
(see Faxon, 1914:412).
RANGE.?"From the Mobile River drainage (as far north as
Choctaw, Tuscaloosa, Blount, and Tallapoosa counties) in
Alabama eastward to the Chattahoochee-Apalachicola drainage
in Alabama, Florida, and Georgia" (Hobbs, 1981:465).
HABITAT.?Streams.
Procambarus (Pennides) vioscai Penn
FIGURE 337
Procambarus vioscai Penn, 1946:27, fig. 1.
Procambarus vioscae.?Williams, 1954:819 [erroneous spelling].
Procambarus viosci.?Hobbs, 1968b:K26 [erroneous spelling].
Procambarus (Pennides) vioscai.?Hobbs, 1972a: 10; 1974b:64,fig- 271.
TYPES.?Holotype and allotype, USNM 79925,79931 (male
I, female); paratypes, USNM, Alabama Mus. NaL Hist., TU.
TYPE LOCALITY.?Big Creek at Fishville, about 3 miles (4.8
km) east of Pollock, Grant Parish, Louisiana.
RANGE.?Tributaries of the Red River in Arkansas and
Louisiana eastward to the Pascagoula River in Mississippi and
lower Tombigbee River in Alabama.
HABITAT.?Streams.
Subgenus Procambarus Ortmann
Cambarus.?Bouvier, 1897:225 [in part].
Subgenus Cambarus Ortmann, 1905c:98,101 [in part].
Subgenus Procambarus Ortmann, 1905d:437 [Type species, by subsequent
designation (Fowler, 1912:340), Cambarus Digueti Bouvier, 1897:225.
Gender: masculine.].?Hobbs, 1972a: 11; 1984, fig. 7.
Procambarus.?Hobbs, 1942a:342.
REMARKS.?Hobbs (1942a:341; 1972a:ll) erred in citing
himself as having designated the type species of Procambarus.
See above.
Procambarus (Procambarus) digueti (Bouvier)
FIGURE 345
Cambarus Digueti Bouvier, 1897:225.
Cambarus carinatus Faxon, 1898:648, pi. 63. [Type, USNM 17699 (male I).
Type locality, Guadalajara, Mexico.]
Cambarus digueti?Hay, 1899b:960.
Cambarus (Cambarus) digueti.?Ortmann, 1905c: 101.
Cambarus (Procambarus) digueti.?Ortmann, 1905d:438.
Cambarus digneti.?Harrison, 1926:380 [erroneous spelling].
Cambarus digniti.?Metcalf, 1929:11 [erroneous spelling].
Procambarus digueti.?Hobbs, 1942a:342 [by implication].?Villalobos,
1946:220, pi. 1.
Procambarus Digueti.?Villalobos, 1953:352.
Procambarus (Procambarus) digueti.?Hobbs, 1972a:ll, figs. 21, 3a, 15a-g;
1974b:64,fig. 278.
TYPES.?Syntypes, USNM 30579 (2 male I, female); MHNP
(8 male, 6 female), CM 74.483 (2 specimens).
TYPE LOCALITY.?Affluents of Rio Santiago, Jalisco,
Mexico.
RANGE.?Pacific slope in states of Jalisco and Michoadin,
Mexico.
HABITAT.?Streams.
Subgenus Remoticambarus Hobbs
Procambarus.?Hobbs, 1972a:l 1.
Subgenus Remoticambarus Hobbs, 1972a: 11 [Type species, by original
designation, Procambarus pecki Hobbs, 1967b:2. Gender: masculine.];
1984, fig. 6.
Procambarus (Remoticambarus) pecki Hobbs
FIGURE 346
Procambarus pecki Hobbs, 1967b:2, figs. 1-12.
Procambarus (Remoticambarus) pecki.?Hobbs, 1972a:ll, figs. 2m, 16a-g;
76 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
1974b:65,fig. 279.?Hobbs. Hobbs, and Daniel, 1977:136, fig. 63.
TYPES.?Holotype, allotype, and morphotype, USNM
117684, 117685, 117686 (male I, female, male II); paratypes,
MCZ.TU.USNM.
TYPE LOCALITY.?McKinney Pit Cave, about 2.5 miles (4
km) west of Tuscumbia, Colbert County, Alabama (Sec. 10, T.
4S,R. 12W).
RANGE.?Tennessee drainage system in Colbert, Lauder-
dale, and Morgan counties, Alabama.
HABITAT.?Subterranean streams and pools.
Subgenus Scapulicambarus Hobbs
Cambarus.?Girard, 1852:91 [in part].
Subgenus Cambarus.?Ortmann, 1905b:401 [in part].
Subgenus Ortmannicus.?Fowler, 1912:341 [in part].
Procambarus.?Hobbs, 1942a:342 [in part].
Cambaus.?Okada, 1948:133 [erroneous spelling].
Procambrus.?Suko, 1961:37 [erroneous spelling].
Subgenus Scapulicambarus Hobbs, 1972a:ll [Type species, by original
designation, Cambarus clarkii paeninsulanus Faxon, 1914:369. Gender
masculine.]; 1984, fig. 5.
Procamborus.?Franzini-Armstrong, 1976:218 [erroneous spelling].
Procambaris.?Wharton, 1978:46 [erroneous spelling].
Combarus.?Shinozaki and Ishida, 1981:50 [erroneous spelling].
Procamburus.?Shinozaki and Ishida, 1981:50 [erroneous spelling].
Procambarius.?Mauro, Thompson, and Melacha, 1983:938 [error ous spell-
ing]-
Procambambarus.?Appelberg, 1983:67 [erroneous spelling].
Procambarus (Scapulicambarus) clarkii (Girard)
FIGURE 351
Cambarus Clarkii Girard, 1852:91.?Hagen, 1870:39, figs. 7-10.99,100,133,
134,142, pi. 4.
Cambarus clarkii.?Faxon, 1898:645.
Cambarus clarki.?Hay, 1902b:437.
Cambarus (Cambarus) clarki.?Ortmann, 1905b:401.
Cambarus (Ortmannicus) clarkii.?Fowler, 1912:341 [by implication].
Cambarus clarkii clarkii.?Faxon, 1914:369 [by implication].
Procambarus clarkii clarkii.?Hobbs, 1942a:342 [by implication].
Procambarus clarkii.?Hobbs, 1942b:103; 1962b:273, figs. 1-9.?Huner,
1977:10.?Pflieger, 1987a:30; 1987b:14.
Cambaus clarkii.?Okada, 1948:133 [erroneous spelling].
Procambarus clarki.?Fingermarm and Lago, 1957:383.?Penn and Hobbs,
1958:466, figs. 2,21,36,49,65.
Procambrus clarkii.?Suko, 1961:37 [erroneous spelling].
Procambarus (Cambarus) clarkii.?Niiyama, 1962:232.
Procambarus clarcii.?Unestam, 1969:203 [erroneous spelling].
Procambarus (Scapulicambarus) clarkii.?Hobbs, 1972a:12; 1974b:65, fig.
283.
Pfrocambarus] (OfrtmannicusJ) clarki.?Bouchard, 1972b:102 [lapsus].
Procamborus clarkii.?Franzini-Armstrong, 1976:218 [erroneous spelling].
Procamburus clarkii.?Brown and Bowler, 1978:34 [erroneous spelling].
Combarus clarkii.?Shinozaki and Ishida, 1981:50 [erroneous spelling].
Procambarius clarki.?Mauro, Thompson, and Melacha, 1983:938 [erroneous
spelling].
Procambambarus clarkii.?Appelberg, 1983:67 [erroneous spelling].
Cambarus calrkii.?Morales, Bozada, and Casanova, 1987:188 [erroneous
spelling].
TYPES.?Destroyed in Chicago fire, 1871 (Faxon,
1914:414).
TYPE LOCALITY.?Between San Antonio and El Paso del
Norte, Texas.
RANGE.?Northern Mexico to Escambia County, Florida,
and north to southern Illinois and Ohio. Widely introduced in
the United States and elsewhere; Huner (1986) recorded the
following introductions: Arizona, California, Georgia, Hawaii,
Idaho, Indiana, Maryland, Nevada, New Mexico, North
Carolina, Ohio, Oregon, South Carolina, Belize, Brazil, Costa
Rica, Dominican Republic, Island of Cyprus, Japan, Kenya,
People's Republic of China, Portugal, Spain, Taiwan, and
Uganda. It has also been introduced into Oklahoma and into
Utah (Johnson, 1986:630). Limited cultivation is in progress in
Colombia, France, and Zambia, and introductions are planned
or have been made in Ecuador, El Salvador, Guatemala,
Honduras, Nicaragua, Venezuela, and Zimbabwe.
HABITAT.?Lentic and lotic situations and burrows (tertiary
burrower).
Procambarus (Scapulicambarus) howellae Hobbs
FIGURE 349
Procambarus howellae Hobbs, 1952b:167, figs. 1,4,5, 8-14.
Procambarus (Scapulicambarus) howellae.?Hobbs, 1972a:12; 1974b:65, fig.
281; 1981:469, figs. 16b, 182b, 183f, 184-188,260.
TYPES.?Holotype and allotype, USNM 93158 (male I,
female); morphotype, USNM 93159 (male II); paratypes,
MCZ,USNM.
TYPE LOCALITY.?Drainage ditch on campus of Wesleyan
College, Rivoli, Bibb County, Georgia.
RANGE.?Lower piedmont and upper coastal plain of
Georgia in the middle Flint and Altamaha river basins.
HABITAT.?Lotic and lentic situations and burrows (tertiary
burrower).
Procambarus (Scapulicambarus) okaloosae Hobbs
FIGURE 350
Procambarus okaloosae Hobbs, 1942b:100,figs. 101-105,315-324.
Procambarus (Scapulicambarus) okaloosae.?Hobbs, 1972a:12; 1974b:65,
fig. 282.
TYPES.?Holotype, USNM 81282 (male I); allotype and
"morphotype," USNM 81283 (female, male II); paratypes,
MCZ.USNM.FSBC.
TYPE LOCALITY.?Roadside ditch at intersection of U.S.
Highway 90 and State Route 4, Milligan, Okaloosa County,
Florida (Sec. 22, T. 3N,R. 24W).
RANGE.?Perdido, Escambia, and Yellow drainage systems
in Alabama and Florida.
HABITAT.?Lentic and lotic situations.
Procambarus (Scapulicambarus) paeninsulanus (Faxon)
FIGURE 348
Cambarus clarkii paeninsulanus Faxon, 1914:369.
Cambarus clarkipaenensulanus.?Hobbs, 1937:154 [erroneous spelling].
NUMBER 480 77
Cambarus clarki paeninsulanus.?Goodnight, 1941:72.
Procambarus clarkii paeninsulanus.?Hobbs, 1942*:342 [by implication].
Procambarus paeninsulanus.?Hobbs, 1942b:104, figs. 106-110.?Caine,
1974:7,14,36.47, figs, lb, 4b, 5a.
Procambarus paeninsulanis.?Anonymous, 1967b, table 3 in unnumbered
Appendix [erroneous spelling].
Cambarus paeninsulanus.?Hobbs, 1972a: 12 [lapsus].
Procambarus (Scapulicambarus)paeninsulanus.?Hobbs, 1972a:12,figs. le,f,
2n, 3b, 17a-g; 1974b:65, fig. 280; 1981:475, figs. 16c, 182c, 183g, 185,
189-191,261.
Procambarus paeninsulans.?Hart and Hart, 1974:128 [erroneous spelling].
Procambaris paeninsulanus.?Wharton, 1978:46 [erroneous spelling.]
TYPES.?Holotype, MCZ 3530 (male II); paratypes, MCZ,
USNM.
TYPE LOCALITY.?Three miles (4.8 km) below Horse
Landing, St. Johns River, Putnam County, Florida.
RANGE.?Southern Georgia, and Honda from the Choc-
tawhatchee Basin east and south to Flagler, Marion, and
Hillsboro counties.
HABITAT.?Lentic and lotic situations, and burrows (tertiary
burrower); rarely in subterranean waters.
Procambarus (Scapulicambarus) strenthi Hobbs
FIGURE 347
Procambarus (Scapulicambarus) strenthi Hobbs, 1977a:412,2 figs.
TYPES.?Holotype, allotype, and morphotype, USNM
147725,147726,147727; paratypes, USNM.
TYPE LOCALITY.?Small stream 15 miles (24 km) west of
Ciudad Valles, San Luis Potosi, Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Stream.
Procambarus (Scapulicambarus) troglodytes (LeConte)
FIGURE 352
Aslacus troglodytes LeConte, 1856:400.
Astacus fossarum LeConte, 1856:401. [Syntypes, MCZ 3377 (male I, dry),
ANSP 314 (female). Type locality, ditches,lower Georgia.]
Astacus maniculatus LeConte, 1856:401. [Types not extant. Type locality,
ditches, lower Georgia.]
Cambarus troglodytes.?Hagen, 1870:41, figs. 11-14,141.
Cambarus troglodytus.?Hagen, 1870:33 [erroneous spelling].
Cambarus maniculatus.?Hagen, 1870:34.
Cambarus fossarum.?Hagen, 1870:53.
Cambarus troglodytes.?Herrick, 1882:254 [erroneous spelling].
Cambarus (Cambarus) troglodytes.?Ortxnann, 1905c: 102.
Cambarus (Ortmannicus) troglodytes.?Fowler, 1912:341 [by implication].?
Creaser, 1934b:4 [by implication].
Procambarus troglodytes.?Hobbs, 1942a:342 [by implication].
Procambarus (Scapulicambarus) troglodytes.?Hobbs, 1972a:12; 1974b:66,
fig. 284; 1981:484,figs. 16d, 182a, 183h, 185,192-194,262.
TYPES.?Syntypes, MCZ 3375 (male I, dry), ANSP 4175
(fragments).
TYPE LOCALITY.?Rice fields in Georgia.
RANGE.?Between the Altamaha and Pee Dee rivers, in
Georgia and South Carolina.
HABITAT.?Lentic and lotic situations, and burrows (tertiary
burrower).
Subgenus Tenuicambarus Hobbs
Procambarus.?Hobbs, 1950b: 194 [in part].
Subgenus Tenuicambarus Hobbs, 1972a: 12 [Type species, by original
designation, Procambarus tenuis Hobbs, 1950b:194. Gender masculine.];
1984, fig. 8.
Procambarus (Tenuicambarus) tenuis Hobbs
FIGURE 353
Procambaruus tenuis Hobbs, 1950b:194, figs. 1-12.
Procambarus (Tenuicambarus) tenuis Hobbs, 1972a:12, figs. 2o, 3d, I8a-g;
1974b:66,fig. 285.
TYPES.?Holotype, allotype, and morphotype, USNM
90390 (male I, female, male II); paratypes, USNM.
TYPE LOCALITY.?Six miles (9.6 km) east of Page and just
west of Oklahoma state line, Le Flore County, Oklahoma.
RANGE.?Arkansas, Ouachita, and Red river basins in
eastern Oklahoma and western Arkansas.
HABITAT.?In, and burrowing adjacent to, springs and clear,
cool streams.
Subgenus Villalobosus Hobbs
Cambarus.?Creaser, 193la: 1 [in part].
Subgenus Cambarus.?Creaser, 1931a:l [in part].
Procambarus.?Hobbs, 1942a:342 [in part].
Paracambarus.?Villalobos, 1944a: 161 [in part].
Subgenus Procambarus.?Villalobos, 1953:354 [lapsus].
Subgenus Villalobosus Hobbs, 1972a:12 [Type species, by original designa-
tion, Paracambarus riojae [sic] Villalobos, 1944a:161. Gender masculine.];
1984, fig. 7.
Procambarus (Villalobosus) contrerasi (Creaser)
FIGURE 358
Cambarus (Cambarus) contrerasi Creaser, 1931a:l, figs. 1-6.
Cambarus contrerasi.?Creaser, 193 la: 1.
Procambarus contrerasi.?Hobbs, 1942a:342.?Villalobos, 1950:373, pis. 2,
3; 1955:142,pis. 29.30; 1983:137,pls. 29,30.
Cambarus (Procambarus) contrerasi.?Villalobos, 1953:354 [lapsus calami].
Procambarus (Villalobosus) contrerasi?Hobbs, 1972a:12; 1974b:66. fig.
289.
TYPES.?Holotype, USNM 98353 (male I); paratype,
USNM.
TYPE LOCALITY.?Tributary of the Rio Cazones near Agua
Fria, 19.2 kilometers (12 mi) south of Miahuapan, State of
Puebla, Mexico.
RANGE.?Tributaries of the Rio Cazones and the Rio
Tecolutla in the vicinity of Zihuateutla and Agua Fria, Puebla,
Mexico.
HABITAT.?Streams.
Procambarus (Villalobosus) cuetzalanae Hobbs
FIGURE 355
Procambarus (Villalobosus) cuetzalanae Hobbs, 1982:39, fig. 1.
TYPES.?Holotype, allotype, and morphotype, USNM
78 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
177202, 177203, and 177204 (male I, female, male II);
paratypes, BM, RNHL, TMM, USNM.
TYPE LOCALITY.?Cueva de Tasalolpan, 5 kilometers (3.1
mi) southwest of Cuetzalan, Puebla, in the Rio Tecuantepcc
(tributary to Rio Tecolutla) watershed, Mexico.
RANGE.?Known from only five localities in the vicinity of
Cuetzalan, Puebla, Mexico.
HABITAT.?Subterranean waters and springs.
Procambarus (Villalobosus) erichsoni Villalobos
FIGURE 357
Procambarus erichsoni Villalobos, 1950:384, ph. 5-7; 1955:131, pis. 26, 27;
1983:127,pls. 26,27.
Procambarus (Villalobosus) erichsoni?Hobbs, 1972a: 12; 1974b:66, fig. 288.
Procambarus ercichsoni.?Bouchard, 1978b:445 [erroneous spelling].
TYPES.?Holotype, allotype, and morphotype, IBM
52048(048) (male I, female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?"Arroyos Puendo, Mamay y Bojoy,
Tenango de Doria, Hidalgo [Mexico]" (Villalobos, 1950:390).
RANGE.?Tributaries of the Rio Pantepec (to Rio Tuxpan) in
the vicinity of Tenango de Doria, Hidalgo, Mexico.
HABITAT.?Streams.
Procambarus (Villalobosus) hoffmanni (Villalobos)
FIGURE 356
Paracambarus hoffmanni Villalobos, 1944a: 169, pi. 2.
Procambarus hoffmanni.?Villalobos, 1950:411; 1955:102, pis. 18, 19;
1983:100, pis. 18,19.
Procambarus Hoffmanni.?Villalobos, 1955:285.
Procambarus Hoffmani.?Hobbs, 1962a:279 [erroneous spelling].
Procambarus (Villalobosus) hoffmanni?Hobbs, 1972a:12; 1974b:67, fig.
287.
TYPES.?Holotype, allotype, and "morphotype," IBM
100043(013) (male I, female, male II); paratypes, IBM.USNM.
TYPE LOCALITY.?"Vertedor de Demasia de la Presa de
Necaxa, Necaxa, [Puebla, Mexico]" (Villalobos, 1955:107).
RANGE.?Tributaries of the Rio Tecolutla and Rio Nautla in
Puebla and Veracruz, Mexico.
HABITAT.?Streams.
Procambarus (Villalobosus) hortonhobbsi Villalobos
FIGURE 359
Procambarus hortonhobbsi Villalobos, 1950:402, pis. 10, 11; 1955:107, pis.
19,20; 1983:105, pis. 19,20.
Procambarus hortonhobsi.?Villalobos, 1955:288 [erroneous spelling].
Procambarus (Villalobosus) hortonhobbsi?Hobbs, 1972a:12; 1974b:67, fig.
290.
TYPES.?Holotype, allotype, and morphotype, IBM
21249(085) (male I, female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?El Coyular, 7 kilometers (4.4 mi)
northeast of La Uni6n, Municipio de Zihuateutla, Puebla,
Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Streams.
Procambarus (Villalobosus) riojai (Villalobos)
FIGURE 354
Paracambarus riojae Villalobos, 1944a:161,pl. 1.
Procambarus riojae.?Villalobos, 1950:401; 1955:96,pl. 17; 1983.95,pl. 17.
Procambarus riojai.?Hobbs, 1971d:3.
Procambarus (Villalobosus) riojai.?Hobbs, 1972a: 12, figs. 2p, 3c, 19a-g;
1974b:67, fig. 286.
TYPES.?Holotype, allotype, and "morphotype," IBM
50045(027) (male I, female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?"Huauchinango, Estado de Puebla, ria-
chuelos tributarios del no Necaxa, [Mexico]" (Villalobos,
1944a:169).
RANGE.?Headwater tributaries of the Rio Cazones and Rio
Tecolutla in Hidalgo and Puebla, Mexico.
HABITAT.?Streams.
Procambarus (Villalobosus) teziutlanensis (Villalobos)
FlGURK 362
Paracambarus teziutlanensis Villalobos, 1947?:240. pi. 2, pi. 3: figs. 5-9.
Procambarus teziutlanensis.?Villalobos. 1950:372; 1955:115. pis. 21, 22;
1983:112, pis. 21,22.
Paracambarus teziutlanensi.?Villalobos, 1953:357 [erroneous spelling).
Procambarus tziutlanensis.?Hobbs, 1971d:12 [erroneous spelling).
Procambarus (Villalobosus) teziutlanensis.?Hobbs, 1972a: 12; 1974b:67, fig.
292.
TYPES.?Holotype, allotype, and morphotype, IBM
121245(029) (male I, female,male II); paratypes,IBM,USNM.
TYPE LOCALITY.?"Chignautla, 5 kilometers [3.1 mi] al
Oriente de Teziutlan, Estado de Puebla, [Mexico]" (Villalobos,
1947a:247).
RANGE.?Headwater tributaries of the Rio Nautla in Puebla
and Veracruz, Mexico.
HABITAT.?Streams.
Procambarus (Villalobosus) tlapacoyanensis (Villalobos)
FIGURE 363
Paracambarus tlapacoyanensis Villalobos, 1947b:537, pis. 1,2.
Procambarus tlapacoyanensis.?Villalobos, 1950:372; 1955:123, pis. 23, 24;
1983:119, pis. 23,24.
Procambarus (Villalobosus) tlapacoyanensis.?Hobbs, 1972a:12; 1974b:67,
fig. 293.
TYPES.?Holotype, allotype, and morphotype, IBM
51947(031) (male I, female, male II); paratypes, IBM, USNM.
TYPE LOCALITY.?Canada de Tomata, Tlapacoyan, Ver-
acruz, Mexico.
RANGE.?Tributaries of the Rio Nautla in the vicinity of
Tlapacoyan, Veracruz, Mexico.
HABITAT.?Streams.
NUMBER 480 79
Procambarus (VUIalobosus) xochitlanae Hobbs
FIGURE 361
Procambarus (VUIalobosus) xochitlanae Hobbs, 1975:16, fig. 5.
TYPES.?Holotype, allotype, morphotype, USNM 145610,
145611, 145612 (male I, female, male II); paratypes USNM,
IBM.TTM.
TYPE LOCALITY.?Cueva de Los Camarones (Rio Tecolutla
Basin), 3 kilometers (1.9 mi) northwest of Xochitl?n, Puebla,
Mexico.
RANGE.?Known only from the type locality.
HABITAT.?Subterranean stream.
Procambarus (VUIalobosus) zihuateutlensis Villalobos
FIGURE 360
Procambarus zihuateutlensis Villalobos, 1950:394, pis. 8,9; 1955:149,pls. 31,
32; 1983:144,pls. 31,32.
Procambarus (VUIalobosus) zihuateullensis.?Hobbs, 1972a: 12; 1974b:67,
fig. 291.
Procambarus (VUIalobosus) zihuatentlensis.?Villalobos, 1982:220 [errone-
ous spelling].
TYPES.?Holotype, allotype, and morphotype, IBM
21149(080) (male I, female, male II); paratypes, IBM.
TYPE LOCALITY.?Arroyo de Tlatentiloyan, Los Estajos,
Municipio de Zihuateutla, Puebla, Mexico.
RANGE.?Headwater tributaries of the Rio Tecolutla in the
vicinity of Zihuateutla, Puebla, Mexico.
HABITAT.?Streams.
Genus Troglocambarus Hobbs
Troglocambarus Hobbs, 1942a:345. [Type species, by original designation,
Troglocambarus maclanei Hobbs, 1942a:345. Gender masculine.]
Trolocambarus.?Bouchard, 1978b:428 [erroneous spelling].
Troglocambarus maclanei Hobbs
FIGURE 364
Troglocambarus maclanei Hobbs, 1942a:345, pis. 1,2; 1971c:122,figs. 17a-g,
19a; 1974b:68, fig. 294.?Hobbs, Hobbs, and DanieL 1977:138, figs. 4,64,
65.?Hobbs and Franz, 1986:509, fig. 3e/ .
Trolocambarus maclanei.?Bouchard, 1978b:428; 1980b:428 [erroneous
spelling].
TYPES.?Holotype, allotype, and "morphotype," USNM
79385, 79386, 79387 (male I, female, male II); paratypes,
MCZ.USNM.
TYPE LOCALITY.?Squirrel Chimney, a small cave about 11
miles (17.6 km) northwest of Gainesville, Alachua County,
Florida (Sec. 21, T. 9S,R. 18E).
RANGE.?Florida: Hernando and Marion counties to
Alachua and Suwannee counties.
HABITAT.?Subterranean waters.
Family PARASTACIDAE Huxley, 1879
Genus Parastacus Huxley
Astacus.?Poeppig, 1835:314.
Parastacus Huxley, 1879:771. [Type species, by subsequent designation
(Faxon, 1898:683), Astacus pilimanus von Martens, 1869:15. Gender:
masculine].
Parastacus brasiliensis (von Martens)
FIGURE 365
Astacus Brasiliensis von Martens, 1869:16, pi. 2: fig. 2.
Astacus brasiliensis.?Huxley, 1879:771.
Parastacus brasUiensis.?Hwdey, 1879:771 [by implication]; 188Oa:25O, fig.
64.?Riek, 1971:133.?Buckup and Rossi, 1980:667, figs. 5-7,8,21.
Parastacus brazUiensis.?Moreira, 1901:80 [erroneous spelling].
Astacus brazUiensis.?Moreira, 1901:80 [erroneous spelling].
Parastacus brasUensis.?Brodsky, 1981:120 [erroneous spelling].
TYPES.?Syntypes,ZBM 3322,3348.
TYPE LOCALITY.?Porto Alegre, Rio Grande do Sul, Brazil.
RANGE.?Known only from the east-central part of the state
of Rio Grande do Sul, Brazil.
HABITAT.?Lotic habitats (see Buckup and Rossi, 1980:
668-669).
REMARKS.?This taxon was considered to be a junior
synonym of Parastacus pilimanus (von Martens) by Riek,
1971, but was recognized as being distinct by Buckup and
Rossi, 1980.
Parastacus defossus Faxon
FIGURE 370
Parastacus defossus Faxon, 1898:686, pi. 67: figs. 3, 4.?Riek, 1971:134.?
Buckup and Rossi, 1980:677, figs. 15-17,21.
TYPES.?Syntypes.USNM 19647 (3 specimens); MCZ 4776
(1 specimen).
TYPE LOCALITY.?Montevideo, Uruguay.
RANGE.?From Montevideo, Uruguay, to east central Rio
Grande do Sul, Brazil. (See Buckup and Rossi, 1980:677.)
HABITAT.?Burrows.
Parastacus laevigatus Buckup and Rossi
FIGURE 369
Parastacus laevigatus Buckup and Rossi, 1980:677, figs. 18-21.
TYPES.?Holotype, MN.UFRJ (male); paratypes,
MN.UFRJ.
TYPE LOCALITY.?"Joinville, (Estrada da Cidra, Chdcara
dos Ipes), Santa Catarina, Brasil" (Buckup and Rossi,
1980:680).
RANGE.?Known from only two localities, in the extreme
northeastern part of Santa Catarina, Brazil.
HABITAT.?Not reported.
80 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Parastacus nicoleti (Philippi)
FIGURE 372
Astacus chUensis.?Nicdct, 1849:211 [not H. Milne Edwards, 1837].
Astacus Nicoleti Philippi, 1882:624.
Astacus Ricoieli.?Philippi, 1882:626 [erroneous spelling].
Parastacus nicoletii Faxon, 1898:689 [as a new species].
Parastacus nicoleti.?Ortmann, 1902:293.?Porter, 1917:98, fig. 61.
Bahamonde, 1958:183, figs. 1-17.?Riek, 1971:133.
Parastacus Nicoleti.?Porter, 1904:255.
Astacus nicoleti.?Faxon, 1914:354.
Parastacus nicoletti.?Bahamonde, 1951:91 [erroneous spelling].
TYPE.?Not designated.
TYPE LOCALITY.?Valdivia, Chile.
RANGE.?Valdivia region of Chile.
HABITAT.?Burrows.
Parastacus pilimanus (von Martens)
FIGURE 366
Astacus pilimanus von Martens, 1869:15, pi. 2: fig. 1.
Parastacus pilimanus.?Huxley, 1879:771 [by implication].?Riek,
1971:133.?Buckup and Rossi, 1980:665, figs. 2 -4 , 8, 21.?Amestoy,
1983:2, fig. 3.
Astacus polmanus.?Brodsky, 1981:120 [erroneous spelling].
TYPES.?Syntypes: ZBM 3323,3447.
TYPE LOCALITY.?Porto Alegre.Rio Grande do Sul, Brazil.
RANGE.?Eastern Uruguay and southern Rio Grande do Sul,
Brazil.
HABITAT.?Burrows; in open water seasonally (associated
with the liberation of young). (See Buckup and Rossi,
1980:665, and Amestoy and Pin, 1984:2).
Parastacus pugnax (Poeppig)
FIGURE 371
Astacus pugnax Poeppig, 1835:314.
Astacus chUensis H. Milne Edwards, 1837:333 [Type: MHNP. Type locality:
"cotes du Chile"].
Astacus chUiensis.?Gray, 1845:411 [erroneous spelling].
Astacus (Cambarus) chUensis.?Erichson, 1846:100.
Astacus (Astacus) Chilensis.?HeMots, 1861:144.
Parastacus ChUensis.?Von Diering, 1893:46.
Parastacus hassleri Faxon, 1898:687,pi. 70: figs. 1-3. [Types: holotype.MCZ
3401; paratypes, USNM 19689. Type locality: Talcahuano, Chile.]
Parastacus Hassleri.?Lonnberg, 1898:349, figs. 1-3.
Parastacus chUensis.?Rathbun, 1910:602.?Holthuis, 1952:81.
Astacus chUensis aua.?Bahamonde, 1951:92.
Parastacus pugnax.?Holthuis, 1952:84.?Riek, 1971:133, fig. lb.
TYPES.?J^ot extant Neotype, here designated: MCZ 3401,
holotype of Parastacus hassleri Faxon, 1898.
TYPE LOCALITY.?Talcahuano, Chile.
RANGE.?Assuming the identity of H. Milne Edwards' type
to be conspecific with Poeppig's species (as believed by Riek,
1971), this Chilean crayfish ranges from Valparaiso southward
to the Taitao Peninsula.
HABITAT.?Burrows.
REMARKS.?The synonomy and taxonomic problems in-
volved with the identities of Astacus chUensis Poeppig and
Astacus chUensis H. Milne Edwards are discussed by Holthuis
(1952).
Parastacus saffordi Faxon
FIGURE 368
Parastacus saffordi Faxon, 1898:683, pi. 68.?Riek, 1971:134.?Buckup and
Rossi, 1980:673, figs. 9-11.21.
TYPES.?Syntypes, USNM 12581 (2 specimens), MCZ 4775
(1 specimen).
TYPE LOCALITY.?Montevideo, Uruguay.
RANGE.?Until the report of the occurrence of this crayfish
in two widely separated localities in the eastern part of the state
of Santa Catarina by Buckup and Rossi (1980:675), this
crayfish was known only from the vicinity of Montevideo,
Uruguay. It is surprising that it has not been found in the state
of Rio Grande do Sul.
HABITAT.?Faxon reported its occurrence in burrows, and
Buckup and Rossi found it swimming in a "cacimba" (= small,
perhaps temporary, pool).
Parastacus varicosus Faxon
FIGURE 367
Parastacus varicosus Faxon, 1898:685,pl. 69.?Riek, 1971:134,fig. la.d.e.?
Buckup and Rossi, 1980:675, figs. 12-14,21.?Amestoy, 1983:2, fig. 2.
TYPE.?Holotype, USNM 4133.
TYPE LOCALITY.?Colima, Mexico. As pointed out by
others, these are obviously erroneous locality data.
RANGE.?Eastern parts of Rio Grande do Sul, Brazil, and
departments of Maldonado and Rocha, Uruguay.
HABITAT.?Lotic situations.
Genus Samastacus Riek
Astacus.?Philippi, 1882:627.
Parastacus.?Faxon, 1898:690.
Samastacus Riek, 1971:135. [Type species, by original designation, Astacus
spinifrons Philippi, 1882:627. Gender masculine.]
Samastacus araucanius (Faxon)
FIGURE 374
Parastacus araucanius Faxon, 1914:353, plate 4.
Samastacus araucanius.?Riek, 1971:135.
TYPE.?Holotype, MCZ 7355 (male).
TYPE LOCALITY.?Corral, Chile.
RANGE.?Known only from the type locality.
HABITAT.?Cascading stream, but according to Faxon, "it
has a strongly compressed cephalothorax, indicating a burrow-
dwelling species."
NUMBER 480 81
Samastacus spinifrons (Philippi)
FIGURE 373
Astacus spinifrons Philippi, 1882:627, fig. B.
Actacus bimacidatus Philippi, 1894:378 [Types: unknown. Type locality:
presumably Chile, but no locality cited. Erroneous spelling of generic name.]
Parastacusagassizii Faxon, 1898:690,pi. 70: figs. 4,5. [Syntypes: MCZ 3400,
USNM 12045. Type locality: Talcahuano, Chile.].?Doflein, 1900:132.
Parastacus agassizi.?Ortmann, 1902:293.
Parastacus Agassizi.?Porter, 1904:258, pi. 9.
Astacus bimacidatus.?Faxon, 1914:354.
Parastacus spinifrons.?Faxon, 1914:354,pl. 9: fig. 1.?Bahamonde, 1951:85,
6 figs.?Holthuis, 1952:81.
Parastacus bimacidatus.?Faxon, 1914:356.
?Parastacus nicoleti.?Porter, 1917:98, fig. 61.
Parastacus agassizzi.?Bahamonde, 1951:89 [erroneous spelling].
Parastacus efinifrons.?Castro, 1966:11 [erroneous spelling].
Samastacus spinifrons.?Riek, 1971:135, fig. lc/.g.?Hobbs, 1974a:26, fig.
26.
TYPES.?Unknown.
TYPE LOCALITY.?"Llico i de Valdivia," Chile (Philippi,
1882:627).
RANGE.?Region of Talcahuano southward to the Taitao
Peninsula and the Island of Chiloe\ and questionably in the
Pacific Basin of neighboring Argentina.
HABITAT.?Lotic (and possibly lentic?) situations.
Appendix 1
Alphabetical List of American Crayfishes with Their Distribution
Crayfishes in Astacidae, Cambaridae, and Parastacidae are listed along with the
countries, states, and/or provinces in which they live. Parentheses enclosing political units
denote known or suspected introductions; if the latter, a question mark follows the closing
parenthesis. The number for each species/subspecies is referenced in Appendix 2 (it does
not correspond to the figure number for that taxon). A dagger indicates the species is
extinct.
Family ASTACIDAE
Pacifastacus
(Hobbsastacus)
f l . chenoderma.?USA: Idaho, Oregon (Miocene and
Pliocene deposits; extinct)
2. connectens.?USA: Idaho, Oregon
3. fortis.?USA: California
4. gambelii.?USA: (California)?, Idaho, Montana,
Nevada, Oregon, Utah, Washington, Wyoming
5. nigrescent.?USA: California (probably extinct)
(Pacifastacus)
6. leniusculus klamathensis.?CANADA: British Co-
lumbia.?USA: California, Idaho, Oregon, Wash-
ington
7. leniusculus leniusculus.?CANADA: British Co-
lumbia.?(JAPAN).?(SWEDEN).?USA: (Cali-
fornia)?, Idaho, (Nevada), Oregon, (Utah)?, Wash-
ington
8. leniusculus trowbridgii.?CANADA: British Co-
lumbia.?(JAPAN).?USA: (California)?, Idaho,
(Nevada), Oregon, Washington
Family CAMBARIDAE
Subfamily CAMBARELUNAE
Cambarellus
(Cambarellus)
9. alvarezi.?MEXICO: Nuevo Le<5n
10. areolatus.?MEXICO: Coahuila
11. chapalanus.?MEXICO: Jalisco, Michoacan
12. chihuahuae.?MEXICO: Chihuahua
13. montezumae.?MEXICO: Distrito Federal, Gua-
najuato, Hidalgo, Jalisco, Mexico, Michoacdn
14. occidentalis.?MEXICO: Michoacan, Sinaloa
15. patzcuarensis.?MEXICO: Michoacin
16. prolixus.?MEXICO: Jalisco
17. zempoalensis.?MEXICO: Morelos
(Dirigicambarus)
18. shufeldtii.?USA: Alabama, Arkansas, (Georgia?),
Illinois, Kentucky, Louisiana, Mississippi, Mis-
souri, Tennessee, Texas
(Pandicambarus)
19. blacki.?USA: Florida
20. diminutus.?USA: Alabama, Mississippi
21. lesliei.?USA: Alabama, Mississippi
22. ninae ?USA: Texas
23. puer.?USA: Arkansas, Illinois, Kentucky, Lou-
isiana, Mississipppi, Tennessee, Texas
24. schmitti.?USA: Florida
25. texanus ?USA: Texas
Subfamily CAMBARINAE
Barbicambarus
26. cornutus.?US A: Kentucky, Tennessee
Bouchardina
27. robisoni.?USA: Arkansas
Cambarus
(Aviticambarus)
28. hamulatus.?USA: Alabama, Tennessee
29. jonesi.?USA: Alabama
(Cambarus)
30. bartonii bartonii.?CANADA: New Brunswick,
Ontario, Quebec.?USA: Alabama, Connecticut,
Delaware, District of Columbia, Georgia, Ken-
tucky, Maryland, Massachusetts, New Jersey, New
York, North Carolina, Pennsylvania, Rhode Island,
South Carolina, Tennessee, Vermont, Virginia,
West Virginia.
31. bartonii carinirostris.?USA: Ohio, West Virginia
32. bartonii cavatus.?USA: Alabama, Georgia, Ken-
tucky, North Carolina, Ohio, Tennessee, Virginia,
West Virginia
82
NUMBER 480 83
33. howardi.?USA: Alabama, Georgia
34. ortmanni.?USA: Indiana, Kentucky, Ohio
35. sciotensis.?USA: Ohio, Virginia, West Virginia
(Depressicambarus)
36. catagius.?USA: North Carolina
37. cymatilis.?USA: Georgia, Tennessee
38. deweesae.?USA: Tennessee
39. englishi.?USA: Alabama, Georgia
40. graysoni.?USA: Alabama, Kentucky, Tennessee
41. halli.?US A: Alabama, Georgia
42. hard.?USA: Georgia
43. latimanus.?USA: Alabama, Florida, Georgia,
North Carolina, South Carolina, Tennessee
44. obstipus.?USA: Alabama
45. pyronotus.?USA: Florida
46. reduncus.?USA: North Carolina, South Carolina
47. reflexus.?USA: Georgia, South Carolina
48. sphenoides.?USA: Kentucky, Tennessee
49. striatus.?USA: Alabama, Florida, Georgia, Ken-
tucky, Mississippi, South Carolina, Tennessee
50. strigosus.?USA: Georgia
51. truncatus.?USA: Georgia
(Erebicambarus)
52. hubbsi.?USA: Arkansas, Missouri
53. hubrichti.?USA: Missouri
54. laevis.?USA: Illinois?, Indiana, Kentucky?
55. ornatus.?USA: Kentucky
56. rusticiformis.?USA: Illinois, Kentucky, Ten-
nessee
57. tenebrosus.?USA: Alabama, Illinois?, Kentucky,
Tennessee
(Exilicambarus)
58. cracens.?USA: Alabama
(Hiaticambarus)
59. chasmodactylus.?USA: North Carolina, Virginia,
West Virginia
60. coosawattae.?USA: Georgia
61. fasciatus.?USA: Georgia
62. girardianus.?USA: Alabama, Georgia, Missis-
sippi?, Tennessee
63. longirostris.?USA: Georgia, North Carolina,
(South Carolina), Tennessee, Virginia
64. longulus.?USA: North Carolina, Virginia, West
Virginia
65. manningi.?USA: Alabama, Georgia, Tennessee
66. speciosus.?USA: Georgia
(Jugicambarus)
67. asperimanus.?USA: Georgia, North Carolina,
South Carolina, Tennessee
68. batchi.?USA: Kentucky
69. bouchardi.?USA: Tennessee
70. brachydactylus.?USA: Tennessee
71. carolinus.?USA: North Carolina, South Carolina,
Tennessee
72. causeyi.?USA: Arkansas
73. conasaugaensis.?USA: Georgia, Tennessee
74. crinipes.?USA: Kentucky, Tennessee
75. cryptodytes.?USA: Florida, Georgia
76. distans.?USA: Alabama, Georgia, Kentucky,
Tennessee
77. dubius.?USA: Kentucky, Maryland, North Caro-
lina, Pennsylvania, Tennessee, Virginia, West
Virginia
78. friaufi.?USA: Tennessee
79. gentryi.?USA: Tennessee
80. monongalensis.?USA: Pennsylvania, Virginia,
West Virginia
81. nodosus.?USA: Georgia, North Carolina, South
Carolina, Tennessee
82. obeyensis.?USA: Tennessee
83. parvoculus.?USA: Georgia, Kentucky, Tennes-
see, Virginia
84. setosus.?USA: Missouri, Oklahoma?
85. tartarus.?USA: Oklahoma
86. unestami.?USA: Alabama, Georgia
87. zophonastes.?USA: Arkansas
(Lacunicambarus)
88. acanthura.?USA: Alabama, Georgia, Tennessee
89. diogenes ?CANADA: Ontario.?USA: Alabama,
Arkansas, Colorado, Delaware, District of Colum-
bia, Florida, Georgia, Illinois, Indiana, Iowa,
Kansas, Kentucky, Louisiana, Maryland, Michi-
gan, Minnesota, Mississippi, Missouri, Nebraska,
New Jersey, North Carolina, North Dakota, Ohio,
Oklahoma, Pennsylvania, South Carolina, South
Dakota, Tennessee, Texas, Virginia, West Vir-
ginia, Wisconsin
90. ludovicianus.?USA: Arkansas, Louisiana, Mis-
sissippi, Tennessee, Texas
91. miltus.?US A: Alabama
(Puncticambarus)
92. acuminatus.?USA: Maryland, North Carolina,
South Carolina, Virginia
93. buntingi.?USA: Kentucky, Tennessee
94. chaugaensis.?USA: Georgia, South Carolina
95. coosae.?USA: Alabama, Georgia, Tennessee
96. cumberlandensis.?USA: Kentucky, Tennessee
97. extraneus.?US A: Georgia, Tennessee
98. georgiae.?USA: Georgia, North Carolina
99. hiwasseensis.?USA: Georgia, North Carolina
100. nerterius.?USA: West Virginia
101. parrishi.?USA: Georgia, North Carolina
102. reburrus?USA: North Carolina
103. robustus.?CANADA: Ontario.?USA: Con-
necticut, Illinois, Indiana, Kentucky, Michigan,
New York, North Carolina, Ohio, Pennsylvania,
Tennessee, Virginia, West Virginia
104. scotti.?USA: Alabama, Georgia
84 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
105. spicatus.?USA: South Carolina
106. veteranus.?USA: Kentucky, Virginia, West Vir-
ginia
(Veticambarus)
107. pristinus.?USA: Tennessee
Distocambarus
(Distocambarus)
108. crockeri.?USA: South Carolina
109. devexus.?USA: Georgia
(Fitzcambarus)
110. carlsoni.?USA: South Carolina
111. youngineri.?USA: South Carolina
Fallicambarus
(Creaserinus)
112. byersi.?USA: Alabama, Florida, Mississippi
113. caesius.?US A: Arkansas
114. danielae.?USA: Alabama, Mississippi
115. fodiens.?CANADA: Ontario.?USA: Alabama,
Arkansas, Illinois, Indiana, Kentucky, Michigan,
Mississippi?, Missouri, Ohio, Tennessee, Wiscon-
sin?
116. hedgpethi ?USA: Alabama?, Arkansas, Florida?,
Georgia?, Louisiana, Mississippi, Oklahoma, Ten-
nessee?, Texas
117. hortoni.?USA: Tennessee
118. oryktes.?USA: Alabama, Louisiana, Mississippi
119. uhleri.?USA: Maryland, North Carolina, South
Carolina, Virginia
(Fallicambarus)
120. dissitus.?USA: Arkansas, Louisiana
121. harpi.?USA: Arkansas
122. jeanae.?USA: Arkansas
123. macneesei.?USA: Louisiana, Texas
124. strawni.?USA: Arkansas
Faxonella
125. beyeri.?USA: Louisiana, Texas
126. blairi.?US A: Arkansas, Oklahoma
127. clypeata.?USA: Alabama, Arkansas?, Florida,
Georgia, Louisiana, Mississippi, Oklahoma?,
South Carolina, Texas
128. creaseri.?USA: Arkansas? Louisiana, Missouri?
Hobbseus
129. attenuatus.?USA: Mississippi
130. cristatus.?USA: Mississippi
131. orconectoides.?USA: Mississippi
132. petilus.?USA: Mississippi
133. prominens.?USA: Alabama, Mississippi
134. valleculus.?USA: Mississippi
Orconectes
(Billecambarus)
135. harrisoni.?USA: Missouri
(Buannulifictus)
136. hobbsi.?USA: Louisiana, Mississippi
137. meeki brevis.?US A: Arkansas, Oklahoma
138. meeki meeki.?USA: Arkansas
139. palmeri creolanus.?USA: Louisiana, Mississippi
140. palmeri longimanus.?USA: Arkansas, Kansas,
Louisiana, Oklahoma, Texas
141. palmeri palmeri.?USA: Arkansas, Kentucky,
Louisiana, Mississippi, Missouri, Tennessee
(Crockerinus)
142. bisectus.?USA: Kentucky
143. erichsonianus.?USA: Alabama, Georgia, Ten-
nessee, Virginia
144. eupunctus.?USA: Arkansas, Missouri
145. illinoiensis.?USA: Illinois
146. jejfersoni.?USA: Kentucky
147. marchandi.?USA: Arkansas, Missouri
148. obscurus.?CANADA: Ontario.?USA: Mary-
land, New York, Ohio, Pennsylvania, Virginia,
West Virginia
149. propinquus.?CANADA: Ontario, Quebec.?
USA: Illinois, Indiana, Iowa, Massachusetts,
Michigan, Minnesota, New York, Ohio, Pennsyl-
vania, Vermont, Wisconsin
150. rafinesquei.?USA: Kentucky
151. sanbornii erismophorous.?USA: West Virginia
152. sanbornii sanbornii.?USA: Kentucky, Ohio,
West Virginia
153. shoupi.?USA: Tennessee
154. stannardi.?USA: Illinois
155. tricuspis.?USA: Kentucky
156. virginiensis.?USA: North Carolina, Virginia
(Faxonius)
157. indianensis.?USA: Illinois, Indiana
158. limosus.?USA: Connecticut, Delaware, District of
Columbia, Maine, Maryland, Massachusetts, New
Hampshire, New Jersey, New York, Pennsylvania,
Rhode Island, Vermont, Virginia, West Virginia.?
(EUROPE, widespread in western part)
159. wrighti.?USA: Tennessee
(Gremicambarus)
160. alabamensis.?USA: Alabama, Mississippi, Ten-
nessee
160a. causeyi.?USA: Arkansas? Colorado, Kansas,
Missouri? (New Mexico), Oklahoma, Texas
161. chickasawae?USA: Mississippi
162. compressus.?USA: Alabama, Kentucky, Mis-
sissippi, Tennessee
163. cooperi.?USA: Alabama, Tennessee
164. etnieri.?USA: Mississippi, Tennessee
165. holti.?USA: Alabama
166. immunis.?CANADA: Manitoba, Ontario,
Quebec.?USA: Alabama, Arkansas, Colorado,
(Connecticut)?, Illinois, Indiana, Iowa, Kansas,
Kentucky, (Maine)?, (Massachusetts)?, Michigan,
Minnesota, Mississippi, Missouri, Montana, Ne-
braska, (New Hampshire)?, New York, North
NUMBER 480 85
Dakota, Ohio, (Rhode Island)?, South Dakota,
Tennessee, (Vermont)?, Wisconsin, Wyoming
167. mississippiensis.?USA: Mississippi
168. nais.?US A: Kansas, Oklahoma, Texas?
169. rhoadesi.?USA: Tennessee
170. validus.?USA: Alabama, Tennessee
171. virilis.?CANADA: Alberta, Manitoba, Ontario,
Quebec, Saskatchewan.?(MEXICO: Baja Califor-
nia Norte, Chihuahua).?USA: (Alabama), (Ari-
zona), Arkansas, (California), Colorado, (Con-
necticut)?, (District of Columbia), Illinois, Indiana,
Iowa, Kansas, Kentucky, (Maine)?, (Maryland),
(Massachusetts)?, Michigan, Minnesota, (Missis-
sippi), Missouri, Montana, Nebraska, (New Hamp-
shire)?, (New Jersey), (New Mexico)?, New York,
North Dakota, Ohio, Oklahoma, (Pennsylvania),
(Rhode Island)?, South Dakota, (Tennessee),
Texas?, Utah, (Vermont)?, (Virginia), (West Vir-
ginia), Wisconsin, Wyoming
(Hespericambarus)
172. blacki.?USA: Louisiana
173. deanae.?USA: New Mexico
174. difficilis.?USA: Arkansas, Louisiana, Oklahoma,
Texas
175. hathawayi.?USA: Louisiana
176. maletae.?USA: Louisiana, Texas
177. perfectus.?USA: Alabama, Mississippi
(Orconectes)
178. australis australis.?USA: Alabama, Tennessee
179. australis packardi.?USA: Kentucky
180. incomptus.?USA: Tennessee
181. inermis inermis.?USA: Indiana, Kentucky
182. inermis testii.?US A: Indiana
183. pellucidus.?USA: Kentucky, Tennessee
(Procericambarus)
184. acares.?USA: Arkansas
185. barrenensis.?USA: Kentucky, Tennessee
186. forceps.?USA: Alabama, Georgia, Tennessee,
Virginia
187. hylas.?USA: Missouri
188. leptogonopodus.?USA: Arkansas, Oklahoma
189. longidigitus.?USA: Arkansas, Missouri
190. luteus.?USA: Arkansas, Kansas, Missouri
191. macrus.?US A: Arkansas, Missouri
192. medius?USA: Missouri
193. menae.?US A: Arkansas, Oklahoma
194. mirus.?US A: Alabama, Tennessee
195. nana.?USA: Arkansas, Oklahoma
1%. neglect us chaenodactylus.?USA: Arkansas, Mis-
souri
197. neglectus neglectus.?USA: Arkansas, Colorado,
Kansas, Missouri, Nebraska, Oklahoma, (Oregon)
198. ozarkae.?USA: Arkansas, Missouri
199. peruncus.?USA: Missouri
200. placidus.?USA: Alabama, Illinois, Kentucky,
Tennessee
201. punctimanus.?USA: Arkansas, Missouri
202. putnami.?USA: Indiana, Kentucky, Tennessee
203. quadruncus.?USA: Missouri
204. rusticus.?CANADA: Ontario.?USA: (Con-
necticut)?, Illinois, Indiana, Iowa, Kentucky,
(Maine)?, (Massachusetts)?, Michigan, Minnesota,
(New Hampshire)?, (New Jersey), (New Mexi-
co), (New York), Ohio, (Pennsylvania)?, (Tennes-
see)?, (Vermont)?, (West Virginia)?, Wisconsin
205. saxatilis.?USA: Oklahoma
206. spinosus.?USA: Alabama, Georgia, Tennessee,
Virginia?, West Virginia?
207. williamsi.?USA: Arkansas, Missouri
(Rhoadesius)
208. kentuckiensis.?USA: Kentucky, Illinois
209. sloanii.?USA: Indiana, Ohio
(Tragulicambarus)
210. lancifer.?USA: Arkansas, Illinois, Kentucky,
Louisiana, Mississippi, Missouri, Tennessee,
Texas
Procambarus
(Acucauda)
111. fitzpatricki.?USA: Mississippi
(Austrocambarus)
212. acanthophorus.?MEXICO: Chiapas, Oaxaca,
Veracruz
213. atldnsoni.?CUBA: Isla de Pinos
214. cubensis cubensis.?CUBA: Camagiiey, Isla de
Pinos, La Habana, Las Villas, Matanzas, Oriente,
Pinar del Rio
215. cubensis rivalis.?CUBA: Pinar del Rio
216. llamasi.?GUATEMALA.?MEXICO: Cam-
peche, Chiapas, Tabasco, Veracruz, Yucatan
217. mexicanus ?MEXICO: Veracruz
218. mirandai.?MEXICO: Chiapas
219. niveus.?CUBA: Pinar del Rio
220. oaxacae oaxacae.?MEXICO: Oaxaca
221. oaxacae reddelli.?MEXICO: Oaxaca, Veracruz
222. pilosimanus.?BELIZE.?GUATEMALA.?
MEXICO: Chiapas, Quintana Roo
|223. primaevus.?USA: Wyoming (Eocene fossils?
extinct)
224. rodriguezi.?MEXICO: Veracruz
225. ruthveni?MEXICO: Veracruz
226. sbordonii.?MEXICO: Chiapas
227. vazquezae.?MEXICO: Veracruz
228. veracruzanus.?MEXICO: Veracruz
229. williamsoni.?GUATEMALA.?HONDURAS
230. zapoapensis.?MEXICO: Veracruz
(Capillicambarus)
231. brazoriensis.?USA: Texas
232. hinei.?USA: Louisiana, Texas
233. incilis.?USA: Texas
(Girardiella)
234. barbiger.?USA: Mississippi
86 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
235. cometes.?USA: Mississippi
236. connus.?USA: Mississippi
237. curdi.?USA: Arkansas, Oklahoma, Texas
238. gracilis.?USA: Illinois, Indiana, Iowa, Kansas?,
Missouri, Oklahoma?, Texas?, Wisconsin 276.
239. hagenianus hagenianus.?USA: Alabama, Mis-
sissippi 277.
240. hagenianus vesticeps.?USA: Mississippi 278.
241. liberorum.?USA: Arkansas 279.
242. parasimulans.?USA: Arkansas
243. pogum.?USA: Mississippi 280.
244. regiomontanus.?MEXICO: Nuevo Leon, Ta- 281.
maulipas
245. reimeri.?USA: Arkansas 282.
246. simulans.?USA: Arkansas, Colorado, Kansas?, 283.
Louisiana, New Mexico, Oklahoma, Texas
247. tulanei.?USA: Arkansas, Louisiana 284.
(Hagenides) 285.
248. advena.?USA: Georgia 286.
249. caritus.?USA: Georgia 287.
250. geodytes.?USA: Florida 288.
251. pygmaeus ?USA: Florida,Georgia 289.
252. rogersi campestris ?USA: Florida 290.
253. rogersi expletus.?USA: Florida 291.
254. rogersi ochlocknensis.?USA: Florida 292.
255. rogersi rogersi.?-USA: Florida 293.
256. talpoides.?US A: Florida, Georgia 294.
257. truculentus ?USA: Georgia 295.
(Leconticambarus) 296.
258. alleni.?USA: Florida 297.
259. apalachicolae.?USA: Florida 298.
260. barbatus.?USA: Georgia, South Carolina 299.
261. capillatus.?USA: Alabama, Florida
262. econfinae.?USA: Florida 300.
263. escambiensis.?US A: Alabama, Florida 301.
264. hubbelli.?USA: Alabama, Florida 302.
265. Hlbyi.?USA: Florida 303.
266. latipleurum.?USA: Florida 304.
267. milleri.?USA: Florida 305.
268. pubischelae deflciens.?USA: Georgia 306.
269. pubischelae pubischelae.?USA: Florida, Georgia 307.
270. raihbunae.?USA: Florida 308.
271. shermani.?USA: Alabama, Florida, Louisiana, 309.
Mississippi 310.
(Lonnbergius) 311.
272. acherontis.?USA: Florida 312.
(Mexicambarus) 313.
273. bouvieri.?MEXICO: MichoacSn 314.
(Ortmannicus) 315.
274. acutissimus.?USA: Alabama, Georgia, Missis- 316.
sippi 317.
275. acutus acutus.?USA: Alabama, Arkansas, (Cali- 318.
fornia), (Connecticut), Delaware, Florida, Georgia, 319.
Illinois, Indiana, Iowa, Kansas, Kentucky, Louisi-
ana, (Maine), Maryland, (Massachusetts), Michi- 320.
gan, Minnesota, Mississippi, Missouri, New Jer-
sey, New York, North Carolina, Ohio, Oklahoma,
Pennsylvania, (Rhode Island), South Carolina,
Tennessee, Texas, Virginia, Wisconsin.
acutus cuevachicae.?MEXICO: Hidalgo, Puebla,
San Luis Potosi, Tamaulipas, Veracruz
ancylus.?USA: North Carolina, South Carolina
angustatus.?USA: Georgia (probably extinct)
bivittatus.?USA: Alabama?, Florida, Louisiana,
Mississippi
blandingii.?USA: North Carolina, South Carolina
caballeroi.?MEXICO: Puebla
chacei.?See enoplosternum
delicatus.?USA: Florida
enoplosternum (including P.(O.) chacei).?USA:
Georgia, South Carolina
epicyrtus.?USA: Georgia
erythrops.?USA: Florida
evermanni.?USA: Alabama, Florida, Mississippi
fallax?USA: Florida, Georgia
franzi.?USA: Florida
geminus.?USA: Arkansas, Louisiana
gonopodocristatus.?MEXICO: Veracruz
hayi.?USA: Alabama, Mississippi, Tennessee
hirsutus.?USA: South Carolina
horsti?USA: Florida
hybus.?USA: Alabama, Mississippi
jaculus.?USA: Louisiana, Mississippi
lecontei.?USA: Alabama, Mississippi
leitheuseri.?USA: Florida
leonensis.?USA: Florida
lepidodactylus.?USA: North Carolina, South
Carolina
lewisi.?USA: Alabama
litosternum.?USA: Georgia
lophotus.?USA: Alabama, Georgia, Tennessee
lucifugus alachua.?US A: Florida
lucifugus lucifugus.?USA: Florida
lunzi.?USA: Georgia, South Carolina
mancus.?USA: Mississippi
marthae.?USA: Alabama
medialis.?USA: North Carolina
orcinus.?USA: Florida
pallidus.?USA: Florida
pearsei.?USA: North Carolina, South Carolina
pictus.?USA: Florida
planirostris.?USA: Louisiana, Mississippi
plumimanus.?USA: North Carolina
pubescens.?USA: Georgia, South Carolina
pycnogonopodus.?USA: Florida
seminolae.?US A: Florida, Georgia
texanus.?USA: Texas
toltecae.?MEXICO: Hidalgo, San Luis Potosi,
Tamaulipas
verrucosus.?USA: Alabama
NUMBER 480 87
321. viaeviridis.?USA: Alabama, Arkansas, Illinois,
Kentucky, Louisiana, Mississippi, Missouri, Ten-
nessee
322. villalobosi.?MEXICO: San Luis Potosi
323. xilitlae.?MEXICO: San Luis Potosi
324. youngi.?USA: Florida
(Paracambarus)
325. ortmanni.?MEXICO: Puebla
326. paradoxus.?MEXICO: Puebla
(Pennides)
327. ablusus.?USA: Mississippi, Tennessee
328. clemmeri.?USA: Alabama, Mississippi
329. dupratzi.?USA: Arkansas, Louisiana, Texas
330. echinatus.?USA: South Carolina
331. elegans.?USA: Arkansas, Louisiana, Mississippi
332. gibbus ? USA: Georgia
333. lagniappe.?USA: Mississippi
334. lylei.?USA: Mississippi
335. natchitochae.?USA: Arkansas, Louisiana, Texas
336. ouachitae.?USA: Arkansas, Mississippi
337. penni.?USA: Louisiana, Mississippi
338. petersi.?USA: Georgia
339. raneyi.?USA: Georgia, South Carolina
340. roberti.?MEXICO: San Luis Potosi
341. spiculifer.?USA: Alabama, Florida, Georgia,
Mississippi, South Carolina
342. suttkusi.?USA: Alabama, Florida
343. versutus.?USA: Alabama, Florida, Georgia
344. vioscai.?USA: Alabama, Arkansas, Louisiana,
Mississippi
(Procambarus)
345. digued.?MEXICO: Jalisco, Michoacan
(Remoticambarus)
346. pecki.?USA: Alabama
(Scapulicambarus)
347. clarkii.?(BELIZE).? (BRAZIL).?(COSTA
RICA).?(DOMINICAN REPUBLIC).?
(ECUADOR).?(EL SALVADOR).?{GUATE-
MALA).?(HONDURAS).?(JAPAN).?MEX-
ICO: (Baja California Norte), (Chihuahua), (Coa-
huila), Nuevo Leon, Sonora.?(NICARAGUA).?
USA: Alabama, (Arizona)?. Arkansas, (Califor-
nia), Florida, (Georgia), (Hawaii), (Idaho), Illinois,
Indiana, Kentucky, Louisiana, (Maryland), Missis-
sippi, Missouri, (Nevada), (New Mexico), (North
Carolina), (Ohio), Oklahoma, (Oregon), (South
Carolina), Tennessee, Texas, (Utah), (Virginia).?
(VENEZUELA). (See also P. (S.) clarkii in
checklist for introductions on other continents and
islands.)
348. howellae.?USA: Georgia
349. okaloosae.?USA: Alabama, Florida
350. paeninsulanus.?USA: Florida, Georgia
351. strenthi ?MEXICO: San Luis Potosi
352. troglodytes.?USA: Georgia, South Carolina
(Tenuicambarus)
353. tenuis.?Arkansas, Oklahoma
(Villalobosus)
354. contrerasi.?MEXICO: Puebla
355. cuetzalanae.?MEXICO: Puebla
356. erichsoni.?MEXICO: Hidalgo
357. hoffmanni.?MEXICO: Puebla, Veracruz
358. hortonhobbsi.?MEXICO: Puebla
359. riojai.?MEXICO: Hidalgo, Puebla
360. teziutlanensis.?MEXICO: Puebla, Veracruz
361. tlapacoyanensis.?MEXICO: Veracruz
362. xochitlanae.?MEXICO: Puebla
363. zihuateutlensis ?MEXICO: Puebla
Troglocambarus
364. maclanei.?USA: Florida
Family PARASTACIDAE
Parastacus
365. brasiliensis.?BRAZIL: Rio Grande do Sul
366. defossus ? BRAZIL: Rio Grande do Sul.?
URUGUAY: Montevideo
367. laevigatus.?BRAZIL: Santa Catarina
368. nicoleti.?CHILE: Aysen, Valdivia
369. pilimanus.?BRAZIL: Rio Grande do Sul.?
URUGUAY: Canelones
370. pugnax.?CHILE: Aysen, Conception, Malleco,
Nuble
371. saffordi.?URUGUAY: Montevideo
372. varicosus ? BRAZIL: Rio Grande do Sul.?
URUGUAY: Maldonada, Rocha
Samastacus
373. araucanius.?CHILE: Valdivia
374. spinifrons.?CHILE: Arauco, Concepcidn, Llan-
quihue, Nuble, Valparaiso.?ARGENTINA?: Neu-
quen
Additional Species Described in 1987 and 1988
375. Cambarus (Jugicambarus) aculabrum.?USA: Ar-
kansas
376. Fallicambarus (Creaserinus) burrisi.?USA: Ala-
bama, Mississippi
377. Fallicambarus (Creaserinus) gordoni.?USA:
Mississippi
378. Fallicambarus (Fallicambarus) devastator.?
USA: Texas
379. Procambarus (Austrocambarus) olmecorum.?
MEXICO: Veracruz
380. Cambarus (Erebicambarus) maculatus.?USA:
Missouri
381. Procambarus (Girardiella) ferrugineus.?USA:
Arkansas
382. Procambarus (Girardiella) regalis.?USA: Ar-
kansas
Appendix 2
Alphabetical List of American Countries and States, or Provinces, and Their Crayfish Faunas
Only those political units in which crayfishes are known to occur are included. The
bibliographic citations are to regional summary works listed in the "Literature Cited" that
should be helpful in studying the crayfishes of the various areas. The numbers refer to the
list of species in Appendix 1 (not to those identifying the illustrations). A question mark
immediately following a number signifies that the record is believed to be in need of
confirmation. Areas occupied as a result of known, virtually certain, or suspected (if
followed by a question mark) introductions are noted by parentheses. The total number of
species reported or known to occur in each political unit is noted in brackets following the
species list.
ARGENTINA (Riek, 1971).?374? [1]
BELIZE (Villalobos, 1955,1983).?222, (347) [2]
BRAZIL (Buckup and Rossi, 1980; Riek, 1971)
Rio Grande do Sul.?(347), 365,366,369,372 [5]
Santa Catarina.?367 [1]
CANADA (Crocker and Barr, 1968; Riegel, 1959)
Alberta.?171 [1]
British Columbia.?6,7,8 [3]
Manitoba.?166,171 [2]
New Brunswick.?30, [1]
Ontario.?30,89,103,115,148,149,166,171,204 [9]
Quebec.?30,149,166,171 [4]
Saskatchewan.?171 [1]
CHILE (Bahamonde, 1951; Holthuis, 1952; Riek, 1971).?
368,370,373,374 [4]
COSTA RICA.?<347) [1]
CUBA (Hobbs and Villalobos, 1964)
Camaguey.?214 [1]
Isla de Pinos.?213,214 [2]
La Habana.?214 [1]
Las Villas.?214 [1]
Matanzas.?214 [1]
Oriente.?214 [1]
Pinar del Rio.?214,215,219 [3]
DOMINICAN REPUBLIC?(347) [1]
ECUADOR.?(347) [1]
EL SALVADOR.?<347) [1]
GUATEMALA (Villalobos, 1955, 1983).?216, 222, 229,
(347) [4]
HONDURAS (Villalobos, 1955,1983).?229, (347) [2]
MEXICO (Villalobos, 1955,1983)
Baja California Norte.?(171), (347) [2]
Campeche.?216 [1]
Chiapas.?212,216,218,222,226 [5]
Chihuahua.?12, (171), (347) [3]
Coahuila.?10, (347) [2]
Distrito Federal.?13 [1]
Guanojuato.?13 [1]
Hidalgo.?13,276,319,356,359 [5]
Jalisco.?11,13,16,345 [4]
Mexico.?13 [1]
Michoacan.?11,13,14,15,273,345 [6]
Morelos.?17 [1]
Nuevo Lo5n.?9,244,347 [3]
Oaxaca.?212,220,221 [3]
Puebla.?276, 281, 325, 326, 354, 355, 357, 358, 359, 360,
362,363 [12]
Quintana Roo.?222 [1]
San Luis Potosi.?276,319,322,323,340,351 [6]
Sinaloa.?14 [1]
Sonora.?(347) [1]
Tabasco.?216 [1]
Tamaulipas.?244,276,319 [3]
Veracruz.?212, 216, 217, 221, 224, 225, 227, 228, 230,
276,290,357,360,361,379 [15]
Yucatan.?216 [1]
NICARAGUA.?(347) [1]
VENEZUELA.?(347) [1]
UNITED STATES
Alabama (Hobbs, 1942b, 1968b, 1969b, 1981).?18,20,21,
28, 29, 30, 32, 33, 39,40, 41, 43,44, 49, 57, 58, 62, 65,
76, 86, 88,89,91,95,104,112,114,115,116?, 118,127,
133, 143, 160, 162, 163, 165, 166, 170, (171), 177, 178,
186, 194, 200, 206, 239, 261, 263, 264, 271, 274, 275,
279?, 286, 291, 294, 296, 300, 302, 307, 320, 321, 328,
341,342,343,344,346,347,349,376 [72]
Arizona (Penn and Hobbs, 1958; Reimer, 1969).?(160a),
(171?), (347) [3]
NUMBER 480 89
Arkansas (Penn, 1959, Reimer, 1969; Williams, 1954;
Bouchard and Robison, 1981).?18, 23, 27, 52, 72, 87,
89, 90, 113, 115, 116, 120, 121, 122, 124, 126, 127?,
128?, 137, 138, 140, 141, 144, 147, (160a?), 166, 171,
174, 184, 188, 189, 190, 191, 193, 195, 196, 197, 198,
201, 207, 210, 237, 241, 242, 245, 246, 247, 275, 289,
321, 329, 331, 335, 336, 344, 347, 353, 375, 381, 382
[60]
California (Bouchard, 1980a; Eng and Daniels, 1982; Riegel,
1959).?3,4?,5,6, (7),(8)?,(171),(275), (347) [9]
Colorado (Engle, 1926; Unger, 1978).?89, 160a, 166, 171,
197,246 [6]
Connecticut (Crocker, 1979; D.G. Smith, 1979).?30, 103,
158, (166)?, (171)?, (204)?, (275) [7]
Delaware (Bouchard, 1982; Francois, 1959; Meredith and
Schwartz, I960).?30,89,158,275 [4]
District of Columbia (Meredith and Schwartz, I960).?30,
89,158, (171)[4]
Florida (Hobbs, 1942b, 1981).?19, 24, 43, 45, 49, 75, 89,
112, 116?, 127, 250, 251, 252, 253, 254, 255, 256, 258,
259, 261, 262, 263, 264, 265, 266, 267, 269, 270, 271,
272, 275, 279, 282, 285, 286, 287, 288, 293, 297, 298,
303, 304, 309, 310, 312, 316, 317, 324, 341, 342, 343,
347,349,350,364 [55]
Georgia (Hobbs, 1981).?(18)?, 30, 32, 33, 37, 39, 41, 42,
43, 47, 49, 50, 51, 60, 61, 62, 63, 65, 66, 67, 73, 75, 76,
81, 83, 86,88,89,94,95,97,98,99,101,104,109,116?,
127, 143, 186, 206, 248, 249, 251, 256, 257, 260, 268,
269, 274, 275, 278, 283, 284, 287, 301, 302, 305, 315,
317,332,338,339,341,343, (347), 348,350,352 [69]
Hawaii.?(347), [1]
Idaho (Riegel, 1959).?1,2,3,4,6,7,8, (347) [8]
Illinois (Page, 1985b).?18, 23, 54?, 56, 57?, 89, 103, 115,
145, 149, 154, 157, 166, 171, 200, 204, 208, 210, 238,
275,321,347 [22]
Indiana (Page, 1985b; Hobbs III and Jass, 1988).?34, 54,
89, 103, 115, 149, 157, 166, 171, 181, 182, 202, 204,
209,238,275,347 [17]
Iowa (Phillips, 1980, 1981).?89, 149, 166, 171, 204, 238,
275 [7]
Kansas (Williams and Leonard, 1952).?89,140,160a, 166,
168,171,190,197,238?, 246?, 275 [11]
Kentucky (Burr and Hobbs, 1984; Ortmann, 1931; Rhoades,
1944a)._18, 23, 26, 30, 32, 34,40, 48, 49, 54?, 55, 56,
57,68,74, 76,77,83,89,93,96,103,106,115,141, 142,
146, 150, 152, 155, 162, 166, 171, 179, 181, 183, 185,
200,202,204,208,210,275,321,347 [45]
Louisiana (Penn, 1952a, 1956b, 1959; Penn and Marlow,
1959).?18, 23, 89, 90, 116, 118, 120, 123, 125, 127,
128, 136, 139, 140, 141, 172, 174, 175, 176, 210, 232,
246, 247, 271, 275, 279, 289, 295, 313, 321, 329, 331,
335,337,344,347 [36]
Maine (Crocker, 1979; D.G. Smith, 1979).?158, (166)?,
(171)?, (204)?, (275) [5]
Maryland (Meredith and Schwartz, I960).?30, 77, 89, 92,
119,148,158, (171), 275, (347) [10]
Massachusetts (Crocker, 1979; D.G. Smith, 1979, 1988).?
30,149,158, (166)?, (171)?, (204)?, (275) [7]
Michigan (Creaser, 1931b).?89, 103, 115, 149, 166, 171,
204,275 [8]
Minnesota (Creaser, 1932, Page, 1985b; Hobbs III and Jass,
1988).?89,149,166,171,204,275 [6]
Mississippi (Hobbs, 1942b, 1981; see also references for
Louisiana).?18, 20, 21, 23, 49, 62?, 89, 90, 112, 114,
115?, 116, 118, 127, 129, 130, 131, 132, 133, 134, 136,
139, 141, 160, 161, 162, 164, 166, 167, (171), 177, 210,
211, 234, 235, 236, 239, 240, 243, 271, 274, 275, 279,
286, 291, 294, 295, 296, 306, 313, 321, 327, 328, 331,
333,334,336,337,341,344,347,376,377 [63]
Missouri (Gardner, 1986; Page, 1985b; Pflieger, 1987a, b;
Williams, 1954).?18, 52, 53, 84, 89, 115, 128?, 135,
141, 144, 147, 160a?, 166, 171, 187, 189, 190, 191, 192,
196, 197, 198, 199, 201, 203, 207, 210, 238, 275, 321,
347,380 [32]
Montana (Riegel, 1959).?4,166,171 [3]
Nebraska (Reimer, 1969; Unger, 1978; Williams, 1954).?
89,166,171,197 [4]
Nevada (Engle, 1926; Reimer, 1969; Riegel, 1959).?4, (7),
(8), (347) [4]
New Hampshire (Aiken, 1965; Crocker, 1979; D.G. Smith,
1979).?158,(166)?, (171)?, (204)? [4]
New Jersey (Bouchard, 1982; Francois, 1959).?30,89,158,
(171), (204), 275 [6]
New Mexico (Penn and Hobbs, 1958; Reimer, 1969).?
(160a), (171?), 173, (204), 246, (347) [6]
New York (Crocker, 1957).?30, 103, 148, 149, 158, 166,
171, (204), 275 [9]
North Carolina (Bouchard, 1976a, 1976b; Hobbs, 1968b,
1969b, 1981; Ortmann, 1931).?30, 32, 36, 43, 46, 59,
63, 64, 67, 71, 77, 81, 89, 92, 98,99, 101,102,103, 119,
156,275,277,280,299,308,311,314, (347) [29]
North Dakota (Phillips, 1980; Hobbs III and Jass, 1988).?
89,166,171 [3]
Ohio (Jezerinac and Thoma, 1984; Rhoades, 1944a, 1944b;
Turner, 1926).?31, 32, 34, 35, 89, 103, 115, 148, 149,
152,166,171,204,209,275, (347) [16]
Oklahoma (Creaser and Ortenberger, 1933; Dunlap, 1951;
Reimer, 1969).?84?, 85, 89, 116, 126, 127?, 137, 140,
160a, 168, 171, 174, 188, 193, 195, 197, 205, 237, 238?,
246,275,347,353 [23]
Oregon (G.C. Miller, 1960; Riegel, 1959).?1, 2,4, 6, 7, 8,
(197), (347) [8]
Pennsylvania (Ortmann, 1906b).?30, 77, 80, 89, 103, 148,
149,158, (171), (204)?, 275 [11]
Rhode Island (Crocker, 1979; D.G. Smith, 1979).?30,158,
(166)?, (171)?, (275) [5]
South Carolina (Hobbs, 1968b, 1969b, 1981; Ortmann,
1931).?30, 43, 46, 47, 49, (63), 67, 71, 81, 89, 92, 94,
90 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
105, 108, 110, 111, 119, 127, 260, 275, 277, 280, 283, Virginia (Hobbs, 1968b, 1969b; Meredith and Schwartz,
292,299,305,311,315,330,339,341, (347), 352 [33] 1960; Ortmann, 1931).?30, 32, 35, 59, 63, 64, 77, 80,
South Dakota (Phillips, 1980; Page, 1985b).?89, 166, 171 83, 89,92, 103,106,119, 143,148,156,158, (171), 186,
[3] 2067,275,(347) [23]
Tennessee (Bouchard, 1972b, 1976a, 1976b, 1978a; Hobbs, Washington (G.C. Miller, 1960; Riegel, 1959).?4, 6, 7, 8,
1969b, 1981; Ortmann, 1931).?18, 23, 26, 28, 30, 32, (152) [5]
37, 38, 40, 43, 48, 49, 56, 57, 62, 63, 65, 67, 69, 70, 71, West Virginia (Lawton, 1979; Ortmann, 1931).?30,31,32,
73, 74, 76, 77, 78, 79, 81, 82, 83, 88, 89, 90, 93, 95, 96, 35, 59, 64, 77, 88, 80, 89, 100, 103, 106, 115, 148, 151,
97, 103, 107, 115, 116?, 117, 141, 143, 153, 159, 160, 152,158, (171), (204)?, 206? [20]
162, 163, 164, 166, 169, 170, (171), 178,180,183,185, Wisconsin (Hobbs III and Jass, 1988; Creaser, 1932).?89,
186,194,200,202, (204)?, 206,210,275,291, 302, 321, 115?, 149,166,171,204,238,275 [8]
327,347 [71] Wyoming (Creaser, 1932; Feldmann et al., 1981; Packard,
Texas (Penn and Hobbs, 1958).?18,22,23,25,89,90,116, 1880; Riegel, 1959).?4,166,171,223 [4]
123,125,127,140, 160a, 168?, 171?, 174,176,197,210, URUGUAY (Amestoy, 1983; Buckup and Rossi, 1980; Riek,
231, 232, 233, 237, 238?, 246, 275, 318, 329, 335, 347, 1971)
378 [30] Canelones.?369 [1]
Utah (Riegel, 1959; Johnson, 1986).?4, (7)?, (171), (347) Maldonada.?372 [1]
[4] Vermont (Crocker, 1979; D.G. Smith, 1979).?30, Montevideo.?366 371 [2]
149,158, (166)?, (171)?, (204)? [6]
Appendix 3
Alphabetical List of American Crayfishes Described during 1987 and 1988
Descriptions of the species listed here appeared after the manuscript had been
completed. Moreover, arrangements of the illustrations of the previously described
crayfishes had already been accomplished, and new ones could not be properly inserted
among them.
Cambarus (Jugicambarus) aculabrum Hobbs and Brown
FIGURE 375
Cambarus (Jugicambarus) aculabrum Hobbs and Brown, 1987:1040, figs. 1,
2a-c,ej,n.
TYPES.?Holotype, allotype, and morphotype, USNM
219149,219150,219151 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Logan Cave, about 11 km east of Siloam
Springs, Benton County, Arkansas (Gallatin Quadrangle, T.
18N, R. 32W, Sec. 33; 36?1 l'50"N, 94?22'5(TW).
RANGE.?Known from only two localities in Benton
County, Arkansas.
HABITAT.?Subterranean waters.
Fallicambarus (Creaserinus) burrisi Fitzpatrick
FIGURE 376
Fallicambarus (Creaserinus) burrisi Fitzpalrick, 1987b:433,figs. 1,2a-d.
TYPES.?Holotype, allotype, and morphotype, USNM
206881, 206882,206883 (male I, female, male II); paratypes,
USNM, MMNH.
TYPE LOCALITY.?Hillside Sarracenia bog, Greene County,
Mississippi (E l/i, Sec. 21, T. 4N.R. 5W).
RANGE.?Chickasawhay and Escatawpa river basins in
Alabama and Mississippi.
HABITAT.?Burrows in Sarracenia bogs (primary bur-
rower).
Fallicambarus (Creaserinus) gordoni Fitzpatrick
FIGURE 377
Fallicambarus (Creaserinus) gordoni Fitzpatrick, 1987b:439, figs. 2e-i,3.
TYPES.?Holotype, allotype, USNM 206877, 206878 (male
I, female); paratypes, USNM, MMNH.
TYPE LOCALITY.?DeSoto National Forest, Camp Shelby
Military Reservation, Perry County, Mississippi (SE 'A.NE 74,
SE 74, Sec. 5,T. 2N,R. 10W).
RANGE.?Known only from the Pascagoula River basin in
the DeSoto National Forest, Perry County, Mississippi.
HABITAT.?Burrows (primary burrower).
Fallicambarus (Fallicambarus) devastator Hobbs and
Whiteman
FIGURE 378
Fallicambarus (Fallicambarus) devastator Hobbs and Whiteman, 1987:403,
figs. 1,2.
TYPES.?Holotype, allotype, and morphotype, USNM
218546, 218547, 218548 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?Angelina County Airport in Burke
Community, about 5 miles (8 km) south of Lufkin, Angelina
County, Texas.
RANGE.?Neches River basin in eastern Texas.
HABITAT.?Burrows (primary burrower).
Procambarus (Austrocambarus) olmecorum Hobbs
FIGURE 379
Procambarus (Austrocambarus) olmecorum Hobbs, 1987:208, fig. 3.
TYPES.?Holotype, allotype, morphotype, USNM 217626,
217627,217628 (male I, female, male II); paratypes, USNM.
TYPE LOCALITY.?Arroyo to Rio Metlac near "edge" of
Fortin de Las Flores, Veracruz, Mexico.
RANGE.?Basins of the Atoyac and Jamapa rivers at
elevations of 800 to 1600 meters, Veracruz, Mexico.
HABITAT.?Lentic and lotic situations.
Cambarus (Erebicambarus) maculatus Hobbs and Pflieger
Cambarus (Erebicambarus) maculatus Hobbs and Pflieger, 1988:644, figs.
1-3.
TYPES.?Holotype, allotype, morphotype, USNM 219292,
219293,219280 (male I, female, male II); paratypes, Ronald D.
Oesch.USNM.
91
92
TYPE LOCALITY.?Hazel Creek at Route C, 8 miles (12.8
km) northeast of Courtois, Washington County, Missouri (T.
36N.R. 1W,SE V4,Sec. 24).
RANGE.?Meramec River basin in eastern Missouri.
HABITAT.?Streams.
Procambarus (Girardiella) ferrugineus Hobbs and Robison
Procambarus (Girardiella) ferrugineus Hobbs and Robison, 1988:391, figs. 1,
2,12,15.
TYPES.?Holotype and allotype, USNM 218841, 218842
(male I, female); paratypes, USNM.
TYPE LOCALITY.?Roadside ditch 10 miles (16 km) south of
Lonoke on State Route 31, Lonoke County, Arkansas (T. IS, R.
8W, Sec. 6).
RANGE.?Known from only two localities in Lonoke
County, Arkansas.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
HABITAT.?Has been taken only from simple burrows.
Procambarus (Girardiella) regalis Hobbs and Robison
Procambarus (Girardiella) regalis Hobbs and Robison, 1988:398, figs. 3, 4,
14,15.
TYPES.?Holotype, allotype, and morphotype, USNM
219244, 219245, 219246 (male I, female, male II); paratypes,
USNM.
TYPE LOCALITY.?DeAnn Cemetery, about 1 mile (1.6 km)
west of the junction of State Routes 19 and 24, on latter, at
western city limit of Prescott, Nevada County, Arkansas (T.
11S.R. 22W,Sec. 8).
RANGE.?Ouachita and Red River basins of Nevada,
Howard, and Sevier counties, Arkansas.
HABITAT.?Simple burrows and temporary pools.
Literature Cited
Abbott, Charles C.
1873. Notes on the Habits of Certain Crawfish. American Naturalist,
7:80-84.
Adams, Charles C.
1901. Baseleveling and Its Faunal Significance, with Illustrations from
Southeastern United States. American Naturalist, 35:839-852,
figures 1-5.
1907. The Crawfishes of the State of Pennsylvania. [Review.] Science,
25(649):897-901.
Aiken, David E.
1965. Distribution and Ecology of Three Species of Crayfish from New
Hampshire. American Midland Naturalist, 73(1): 240-244.
Airaksinen, Mauno, Heva-Liisa Valkama, and Ossi V. Lindqvist
1978. Distribution of DDT in the Crayfish Astacus astacus L. in Acute
Test. In Ossi V. Lindqvist, editor. Freshwater Crayfish 3, pages
349-356,1 figure. Kuopio, Finland: University of Kuopio.
Albaugh, Douglas W.
1973. Life Histories of the Crayfishes Procambarus acutus and Procam-
barus hinei in Texas, xiii + 1 3 5 pages, figures 1-28. Doctoral
dissertation, Texas A&M University, College Station.
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33(1,2):273-276,9 figures.
A New Cave-Dwelling Crayfish from the Greenbrier Drainage
System, West Virginia (Decapoda, Astacidae). Proceedings of the
Biological Society of Washington, 77(21):189-194,figures 1-10.
A New Crayfish of the Genus Cambarus from Tennessee with an
Emended Definition of the Genus (Decapoda, AsUcidae). Proceed-
ings of the Biological Society of Washington, 78(33):265-273,
figures 1-12.
A New Crayfish from Alabama with Observations on the Crislatus
Section of the Genus Cambarus (Decapoda, Astacidae). Proceedings
of the Biological Society of Washington, 79(15):109-l 16, figures
1-10.
An Illustrated Key to the Species of the Genus Ankylocythere with a
Description of a New Species from Louisiana (Ostracoda, Entocyth-
eridae). Proceedings of the Louisiana Academy of Sciences,
29:67-75, figures 1-18.
The Current Status of the Crayfishes Listed by Girard (1852) in His
"A Revision of the North American Astaci . . . . " Cruslaceana,
12(2): 124-132.
A New Crayfish from Alabama Caves with Notes on the Origin of
the Genera Orconectes and Cambarus (Decapoda: Astacidae).
Proceedings of the Untied States National Museum, 123(3621): 1-
17, figures 1-21.
Two New Crayfishes of the Genus Cambarus from Georgia,
Kentucky, and Tennessee (Decapoda, AsUcidae). Proceedings of the
Biological Society of Washington, 81(31):261-274,figures 1-22.
Crusucea: Malacostraca. in Fred K. Parrish, Keys to Water Quality
Indicative Organisms (Southeastern Untied States), pages K1-K36,
32 figures. Federal Water Pollution Control Administration, Depart-
ment of the Interior.
Procambarus villalobosi, un Nuevo Cambarina de San Luis Potosf,
Mexico (Decapoda, AsUcidae). Anales del Instituto de Biologia,
Universidad Nacional Autonoma de Mexico, Serie Ciencias del Mar
yLimnologui,38(l)Al -46, figures 1-11.
On the Distribution and Phylogeny of the Crayfish Genus
Cambarus. In Perry C. Holt, Richard L Hoffman, and C. Willard
Hart, Jr., The Distributional History of the BioU of the Southern
Appalachians, Part I: Invertebrates. Virginia Polytechnic Institute,
Research Division Monograph, 1:93-178, figures 1-20.
Two New Species of the Crayfish Genus Procambarus (Decapoda,
AsUcidae) with Keys to the Members of the Spiculifer Group.
Proceedings of the Biological Society of Washington, 83(24):329-
348, figures 1-38.
A New Crayfish from the Nashville Basin, Tennessee. Proceedings
of the Biological Society of Washington, 83(14):161-169, figure 1.
New Crayfishes of the Genus Cambarus from Tennessee and
Georgia (Decapoda, AsUcidae). Proceedings of the Biological
Society of Washington, 83(23):241-259, figures 1-4.
NUMBER 480 101
1971a. A New Crayfish of the Genus Procambarus from Mississippi
(Decapods: Astacidac). Proceedings of the Biological Society of
Washington, 83(4O):459-468,figure 1.
1971b. New Crayfishes of the Genus Procambarus from Alabama and
Texas (Decapoda, Astacidae). Proceedings of the Biological Society
of Washington, 84(11): 81-94, figures 1,2.
1971c. A New Troglobitic Crayfish from Florida. Quarterly Journal of the
Florida Academy of Sciences, 34(2): 114-124, figures 1-19.
197Id. The Entocytherid Ostracods of Mexico and Cuba. Smithsonian
Contributions to Zoology, 81: 55 pages, figures 1-31.
1972a. The Subgenera of the Crayfish Genus Procambarus (Decapoda:
Astacidae). Smithsonian Contributions to Zoology, 117: 22 pages,
figures 1-20.
1972b. Crayfishes (Astacidae) of North and Middle America. In Biota of
Freshwater Ecosystems, Identification Manual, 9: x + 173 pages,
115 figures. United States Environmental Protection Agency.,
[Water Pollution Research Control Series; reprinted 1976 with same
pagination.]
1973a. New Species and Relationships of the Members of the Genus
Fallicambarus. Proceedings of the Biological Society of Washing-
ton, 86(40):461-481, figures 1-4.
1973b. Three New Troglobitic Decapod Crustaceans from Oaxaca, Mexico.
Association for Mexican Cave Studies, Bulletin, 5:25-38, figures
1-8.
1973c. Two New Troglobitic Shrimps (Decapoda: Alpheidae and Palaemo-
nidae) from Oaxaca, Mexico. Association for Mexican Cave Studies,
Bulletin, 5:73-80,figures 1-3.
1974a. Synopsis of the Families and Genera of Crayfishes (Crustacea,
Decapoda). Smithsonian Contributions to Zoology, 164: 32 pages,
figures 1-27.
1974b. A Checklist of the North and Middle American Crayfishes
(Decapoda: Astacidae and Cambaridae). Smithsonian Contributions
to Zoology, 166: iii+ 161 pages, figures 1-294.
1975. New Crayfishes (Decapoda: Cambaridae) from the Southern United
States and Mexico. Smithsonian Contributions to Zoology, 201:1-
34, figures 1-8.
1977a. A New Crayfish (Decapoda: Cambaridae) from San Luis Potosi,
Mexico. Proceedings of the Biological Society of Washington,
90(2):412-419,figuresl,2.
1977b. Cave-inhabiting Crayfishes of Chiapas, Mexico (Decapoda: Cam-
baridae). Accademia Nazionale dei Lincei, 171:197-206, figures
1-2
1977c. The Crayfish Bouchardina robisoni, a New Genus and Species
(Decapoda, Cambaridae) from Southern Arkansas. Proceedings of
the Biological Society of Washington, 89(62):733-742, figure 1.
1979. A New Crayfish from the Ouachita River Basin in Arkansas
(Decapoda: Cambaridae). Proceedings of the Biological Society of
Washington, 92(4):804-811, figure 1.
1980. New Dwarf Crayfishes (Decapoda: Cambaridae) from Mexico and
Florida. Proceedings of the Biological Society of Washington,
93(l):194-207,figuresl,2.
1981. The Crayfishes of Georgia. Smithsonian Contributions to Zoology,
318: viii + 549 pages, figures 1-262.
1982. A New Crayfish (Decapoda: Cambaridae) from the State of Puebla,
Mexico, with New Locality Records for Procambarus (Villalobosus)
xochitlanae and Entocytherid Ostracod Symbionts. Mexican Cave
Studies, Bulletin, 8:39-44, figure 1.
1983. Distocambarus (Fitzcambarus) carlsoni, a New Subgenus and
Species of Crayfish (Decapoda: Cambaridae) from South Carolina.
Proceedings of the Biological Society of Washington, 96(3):429-
439, figure 1.
1984. On the Distribution of the Crayfish Genus Procambarus (Decapoda:
Cambaridae). Journal of Crustacean Biology, 4(1): 12-24, figures
1-11.
1986. Highlights of a Half Century of Crayfishing. In Per Brinck, editor,
Freshwater Crayfish 6, pages 12-23,3 figures. Lund, Sweden: Btj
Datafilm.
1987. On the Identity of Astacus (Cambarus) mexicanus Erichson (1846)
and Cambarus aztecus Saussure (1857) (Decapoda: Cambaridae)
with the Description of Procambarus olmecorum. New Species from
Veracruz, Mexico. Proceedings of the Biological Society of
Washington, 100(1): 198-215,3 figures.
Hobbs, Horton H., Jr., and Thomas C. Barr, Jr.
1960. The Origins and Affinities of the Troglobitic Crayfishes of North
America (Decapoda, Astacidae), I: The Genus Cambarus. American
Midland Naturalist, 64(l):12-33, figures 1-57.
1972. Origins and Affinities of the Troglobitic Crayfishes of North
America (Decapoda: Astacidae), II: Genus Orconectes. Smithsonian
Contributions toZoology, 105: 84 pages,figures 1-16.
Hobbs, Horton H., Jr., and M.S. Bedinger
1964. A New Troglobitic Crayfish of the Genus Cambarus (Decapoda,
Astacidae) from Arkansas with a Note on the Range of Cambarus
cryptodytes Hobbs. Proceedings of the Biological Society of
Washington. 77(3) :9-15, figures 1-11.
Hobbs, Horton H., Jr., and Raymond W. Bouchard
1973. A New Crayfish from the Cumberland River System with Notes on
Cambarus carolinus (Erichson). Proceedings of the Biological
Society of Washington, 86(5):41-68, figures 1-4.
Hobbs, Horton H., Jr., and Arthur V. Brown
1987. A New Troglobitic Crayfish from Northwestern Arkansas (Deca-
poda: Cambaridae). Proceedings of the Biological Society of
Washington, 100(4): 1040-1048, figures 1,2.
Hobbs, Horton H., Jr., and Paul H. Carlson
1983. Distocambarus (Decapoda: Cambaridae) Elevated to Generic Rank,
with an Account of D. crockeri, New Species, from South Carolina.
Proceedings of the Biological Society of Washington, 96(3):420-
428, figure 1.
1985. A New Member of the Genus Distocambarus (Decapoda: Cambari-
dae) from the Saluda Basin, South Carolina. Proceedings of the
Biological Society of Washington, 98(1 ):81-89, figure 1.
Hobbs, Horton H., Jr., and Martha R. Cooper
1972. A New Troglobitic Crayfish from Oklahoma (Decapoda: Astacidae).
Proceedings of the Biological Society of Washington, 85(3):49-56,
figure 1-12.
Hobbs, Horton H., Jr., and J.F. Fitzpatrick, Jr.
1962. A New Crayfish of the Propinquus Group of the Genus Orconectes
from the Ohio Drainage System in West Virginia (Decapoda,
Astacidae). Proceedings of the Biological Society of Washington,
75(30):207-214, figures 1-15.
1970. A New Crayfish of the Genus Fallicambarus from Tennessee
(Decapoda, Astacidae). Proceedings of the Biological Society of
Washington, 82(64):829-836, figures 1-12.
Hobbs, Horton H., Jr., and Richard Franz
1986. New Troglobitic Crayfish with Comments on Its Relationship to
Epigean and Other Hypogean Crayfishes of Florida. Journal of
Crustacean Biology, 6(3):509-519, figures 1-3.
Hobbs, Horton H., Jr., and Andrew G. Grubbs
1982. Description of a New Troglobitic Crayfish from Mexico and a list
of Mexican Crayfishes Reported Since the Publication of the
Villalobos Monograph (1955) (Decapoda, Cambaridae). Association
for Mexican Cave Studies, Bulletin, 8:45-50, figure 1.
1986. Notes on the Crayfish Procambarus (Ortmannicus) xilitlae (Deca-
poda: Cambaridae). Proceedings of the Biological Society of
Washington, 99(4):735-738, figure 1.
Hobbs, Horton H., Jr., and Edward T. Hall, Jr.
1969. New Crayfishes from Georgia (Decapoda, Astacidae). Proceedings
of the Biological Society of Washington, 82(21 ):281-294, figures
1-24.
102 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
1972. A New Crayfish from the Tallapoosa River in Georgia (Decapoda:
Astacidae). Proceedings of the Biological Society of Washington,
85(12):151-161,figure 1.
1974. Crayfishes (Decapoda: Astacidae). In C.W. Hart, Jr., and S.L.H.
Fuller, editors. Pollution Ecology of Freshwater Invertebrates, pages
195-214. New York: Academic Press.
Hobbs, Horton H., Jr., and C.W. Hart, Jr.
1959. The Freshwater Decapod Crustaceans of the Apalachicola Drainage
System in Florida, Southern Alabama, and Georgia. Bulletin of the
Florida State Museum, 4(5): 145-191. figures 1-39.
Hobbs, Horton H., Jr., and H.H. Hobbs III
1962. A New Crayfish of the Genus Cambarus from Georgia (Decapoda,
Astacidae). Proceedings of the Biological Society of Washington,
75(5):41-45, figures 1-10.
Hobbs, Horton H., Jr., H.H. Hobbs III, and Margaret A. Daniel
1977. A Review of the Troglobitic Crustaceans of the Americas.
Smithsonian Contributions to Zoology, 244: v + 183 pages, figures
1-70.
Hobbs, Horton H., Jr., Perry C. Holt, and Margaret Walton
1967. The Crayfishes and Their Epizootic Ostracod and Branchiobdellid
Associates of the Mountain Lake, Virginia, Region. Proceedings of
the United States National Museum, 123(3602): 1-84, figures 1-22.
Hobbs, Horton H., Jr., and David S. Lee
1976. A New Troglobitic Crayfish (Decapoda, Cambaridae) from Peninsu-
lar Florida. Proceedings of the Biological Society of Washington,
89(32):383-391,figurel.
Hobbs, Horton H., Jr., and Lewis J. Marchand
1943. A Contribution toward a Knowledge of the Crayfishes of the
Reelfoot Lake Area. Journal of the Tennessee Academy of Science,
18(l):6-35, figures 1-27.
Hobbs, Horton H., Jr., and D. Bruce Means
1972. Two New Troglobitic Crayfishes (Decapoda, Astacidae) from
Florida. Proceedings of the Biological Society of Washington,
84(46):393-4O9, figures 1.2.
Hobbs, Horton H., Jr., and Frank O. Perkins
1967. A New Burrowing Crayfish from North Carolina (Decapoda,
Astacidae). Proceedings of the Biological Society of Washington,
80(21):141-146,figures 1-11.
Hobbs, Horton H., Jr., and Daniel J. Peters
1977. The Entocytherid Ostracods of North Carolina. Smithsonian Contri-
butions to Zoology, 247: iv + 73 pages, figures 1 -33.
1982. The Entocytherid Ostracod Fauna of Northern Georgia. Proceedings
of the Biological Society of Washington, 95(2):297-318, figures
1-9.
Hobbs, Horton H., Jr., and William L. Pflieger
1988. Cambarus (Erebicambarus) maculatus, a New Crayfish (Decapoda:
Cambaridae) from the Meramec River Basin of Missouri. Proceed-
ings of the Biological Society of Washington, 101(3):644-652,
figures 1-3.
Hobbs, Horton H., Jr., and Henry W. Robison
1982. A New Crayfish of the Genus Procambarus from Southwestern
Arkansas. Proceedings of the Biological Society of Washington,
95(3):545-553. figure 1.
1985. A New Burrowing Crayfish (Decapoda: Cambaridae) from South-
western Arkansas. Proceedings of the Biological Society of
Washington, 98(4):1035-1041, figure 1.
1988. The Crayfish Subgenus Girardiella (Decapoda: Cambaridae) in
Arkansas, with the Descriptions of Two New Species and a Key to
the Members of the gracilia Group of the Genus Procambarus.
Proceedings of the Biological Society of Washington, 1O1(2):391-
413, figures 1-16.
Hobbs, Horton H., Jr., and Charles S. Shoup
1942. On the Crayfishes Collected from the Big South Fork of the
Cumberland River in Tennessee during the Summer of 1938.
American Midland Naturalist, 28(3):634-644, figures 1-8.
1947. Two New Crayfishes (Decapoda, Astacidae) from the Obey River
Drainage in Tennessee. Journal of the Tennessee Academy of
Science, 22(2):138-145,figures 1-22.
Hobbs, Horton H., Jr., and Alejandro Villalobos
1964. Los Cambarinos de Cuba. Anales del Instituto de Biologia,
Universidad Nacional Autonoma de Mexico, 34(l,2):307-366,
figures 1-50.
Hobbs, Horton H., Jr., and Margaret Walton
1957. Three New Crayfishes from Alabama and Mississippi (Decapoda:
Astacidae). Tulane Studies in Zoology, 5(3):39-52, figures 1-34.
1958. Procambarus pearsei plumimanus, a New Crayfish from North
Carolina (Decapoda, Astacidae). Journal of the Elisha Mitchell
Scientific Society, 74(1):7-12, figures 1-13.
1959. A New Crayfish of the Genus Procambarus from Alabama
(Decapoda, Astacidae). Proceedings of the Biological Society of
Washington, 72(10):39-44, figures 1-12.
1960. A New Crayfish of the Genus Procambarus from Southern Alabama
(Decapoda, Astacidae). Proceedings of the Biological Society of
Washington, 73(20):123-129,figures 1-11.
1966. Orconectes Juvenils (Hagen) in Mountain Lake, Virginia: an
Unplanned Experiment in Interspecific Competition (Decapoda,
Astacidae). Virginia Journal of Science, 17(3):136-14O.
Hobbs, Horton H., and Mike Whiteman
1987. A New, Economically Important Crayfish (Decapoda: Cambaridae)
from the Neches River Basin, Texas, with a Key to the Subgenus
Fallicambarus. Proceedings of the Biological Society of Washing-
ton, 100(2):403-411, figures 1,2.
Hobbs III. H.H.
1969. Studies in Ecological and Host Specificity in Entocytherid Ostracods
(Ostracoda: Entocytheridae). vii + 93 pages, 8 figures. Master's
thesis, Mississippi State University, Starkville.
1971. New Entocytherid Ostracods of the Genera Ankylocythere and
Dactylocythere. Proceedings of the Biological Society of Washing-
ton, 84(17):137-146, figures 1,2.
1973. The Population Dynamics of Cave Crayfishes and Their Commensal
Ostracods from Southern Indiana. [Abstract.] Fifteenth Annual
Report of the Cave Research Foundation, pages 58-59.
1975. Distribution of Indiana Cavemicolous Crayfishes and Their Ecto-
Commensal Ostracods. International Journal of Speleology, 7:273-
302, figures 1-6.
1980. Population Studies of Indiana Cavemicolous Ostracods (Ostracoda:
Entocytheridae). Proceedings of the Indiana Academy of Science,
89:147-148.
Hobbs III.H.H., and Joan P. Jass
1988. The Wisconsin Decapod Crustaceans (Cambaridae, Palaemoniaae).
viii + 177 pages, 78 figures. Milwaukee Public Museum.
Hobbs III, H.H., James H. Thorp, and Gilbert E. Anderson
1976. The Freshwater Decapod Crustaceans (Palaemoniaae, Cambari-
dae) of the Savannah River Plant, South Carolina. 63 pages, figures
1-24. Savannah River Plant, National Environment Research Park
Program.
Hobbs III, H.H., and Norbert Welch
1972. Will Lost River be Lost? NSS News. 30(3):66-67.
Hobbs III, H.H., and Steve G. Wells
1972. The Lost River Karst?Problems in Conservation and Land
Management. JVSSJVewj, 30(8): 123-128, figure 1.
Hobbs III, H.H., and Stanley A. Rewolinski
1985. Notes on the Burrowing Crayfish Procambarus (Girardiella)
gracilis (Bundy) (Decapoda, Cambaridae) from Southeastern Wis-
consin, U.S.A. Crustaceana, 48(1 ):26-33, figures 1-3.
Hoff, C. Clayton
1943. Two New Ostracods of the Genus Entocythere and Records of
Previously Described Species. Journal of the Washington Academy
NUMBER 480 103
of Sciences, 33(9):276-286, figures 1,2.
1944. New American Species of the Ostracod Genus Entocythere.
American Midland Naturalist, 32(2):327-357, figures 1-33.
Holt, Perry C.
1954. A New Branchiobdellid of the Genus Cambarincola (Oligochaeta,
Branchiobdellidae) from Virginia. Virginia Journal of Science,
5(3): 168-172, figures 1-5.
1960. The Genus Ceratodrilus Hall (Branchiobdellidae, Oligochaeta), with
the Description of a New Species. Virginia Journal of Science,
ll(2):53-77,figuresl-16.
1973. Branchiobdellids (Annelida: Clitellata) from Some Eastern North
American Caves, with Descriptions of New Species of the Genus
Cambarincola. International Journal of Speleology, 5:219-256,
figures 1-7.
Holthuis, L.B.
1952. The Crustacea Decapoda Macrura of Chile. Lunds Universitets
Arsskrift, new series 2,47(10):l-109,figures 1-19.
1956. Proposed Suppression under the Plenary Powers (a) Certain Names
Given by C.S. Rafinesque to Genera and Species of the Orders
Decapoda and Stomatopoda (Class Crustacea) and (b) of Certain
Specific Names Currently Regarded as Senior Subjective Synonyms
of the Names of the Type Species of the Genera "Homola" and
"Lissa," Both of Leach, 1815, Belonging to the Foregoing Class.
Bulletin of Zoological Nomenclature, 12(9):227-239.
1964. On the Status of the Two Allegedly European Crayfishes, Cambarus
typhlobius Joseph, 1880, and Austropotamobius pallipes bispinosus
Karaman, 1961 (Decapoda, Astacidae). Cruslaceana, 7(l):42-48,
plate 3.
Hopkins, C.L.
1967. Breeding in the Freshwater Crayfish Paranephrops planifrons
White. Journal of Marine and Freshwater Research, l(l):51-58,
figures 1-7.
Hovey, Horace C.
1882. Celebrated American Caverns, Especially Mammoth, Wyandot, and
Luray, together with Historical Scientific and Descriptive Notices of
Caves and Grottoes in other Lands; with Maps and Illustrations, xii
+ 288 pages, illustrated. Cincinnati: Robert Clarke and Company.
Huner, Jay V.
1977. Ditch Bugs, Good Bait . . . Good to Eat! Bayou State Sportsman,
2(2):10-13, illustrated.
1978. Exploitation of Freshwater Crayfishes in North America. Fisheries,
3(6):2-5, illustrated.
1981. Information about the Biology and Culture of the Red Crawfish,
Procambarus clarkii (Girard, 1852) (Decapoda, Cambaridae) for
Fisheries Managers in Latin America. Anales del Instituto Ciencia
del Mar y Limnologia, Universidad Nacional Autdnoma de Mexico,
8(l):43-50.
1986. Crawfish Introductions Affect Louisiana Industry. Crawfish Tales,
5(3):16-18.
Huner, J.V., and J.E. Barr
1980. Red Swamp Crawfish: Biology and Exploitation, xi + 148 pages, 61
figures. Baton Rouge: Louisiana State University Center for
Wetland Resources. [Sea Grant Publication Number LSU-T-80-
001.]
Huner, J.V., Samuel P. Meyers, and James W. Avault, Jr.
1976. Response and Growth of Freshwater Crawfish to an Extruded,
Water-Stable Diet In James W. Avault, Jr., editor. Freshwater
Crayfish, pages 149-157. Baton Rouge, Louisiana: Division of
Continuing Education, Louisiana State University.
Huner, J.V., and Syed Naqvi
1984. Invertebrate Faunas and Crawfish Food Habits in Louisiana
Crawfish Ponds. Proceedings of the Annual Conference of the
Southeastern Association of Fish and Wildlife Agencies, 38:395-
406.
Huntsman, A.G.
1915. The Fresh-Water Malacostraca of Ontario. Contributions to Cana-
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1879. On the Classification and the Distribution of the Crayfishes.
Proceedings of the Zoological Society of London, 1878:751-788,
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1880a. The Crayfish: An Introduction to the Study of Zoology. Interna-
tional Scientific Series, 28: xiv + 371, 81 figures and frontispiece.
1880b. L'ecrevisse: Introduction a l'etude de la zoologie. Bibliotheque
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1974. Mating and Spawning of the Crayfish Austropotamobius pallipes
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Jackson, George F.
1942. Animal Life of Caverns. Bulletin of the National Speleological
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James, Hugo A.
1966. Range and Variations of Subspecies of Cambarus longulus
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Jeannel, R., and H. Henrot
1949. Les coleopteres cavemicoles de la region des Appalaches. Notes
Biospeoleogiques, Museum National d'Histoire Naturelle, 4(12):7-
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Jester, Douglas B.
1967. A New Crawfish of the Genus Orconectes from New Mexico
(Decapoda, Astacidae). American Midland Naturalist, 77(2):518-
524, figures 1-12.
Jezerinac, Raymond F.
1983. Possible Correlations of Present Distributions of Ohio Crayfishes
(Decapoda: Cambaridae) with Teays-Age Drainages. In Anony-
mous, Teays-Age Drainage Effects on Present Distributional
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[Abstracts and supplements.] Ohio Biological Survey Informative
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Jezerinac, Raymond F., and Roger F. Thoma
1984. An Illustrated Key to the Ohio Cambarus and Fallicambarus
(Decapoda: Cambaridae) with Comments and a New Subspecies
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1986. Inventory of Utah Crayfish with Notes on Current Distribution.
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1939. Atlas de Paleobiogeographie. 39 pages, 99 plates. Paris: Paul
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Jordan, Stephen D., and D.W. Dunham
1981. Two New Color Morphs of the Crayfish Orconectes propinquus
(Girard) from Lake Simcoe, Ontario. American Midland Naturalist,
105(2):384-385, figure 1.
Joseph, Gustav
1880. Demonstration eines neuen zur Familie der Astaciden gehorenden
blinden Grottenkrebses aus Krain. Schlesische Gesellschaft fur
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104 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
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1970. Studies on the Fresh-Water Shrimps, Prawns and Crawfishes of
Japan. 213 pages. Matsue.
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(Dobutsugaku Zasshi), 83(2):123-126,figurcs 1-4.
Kingsley, J.S.
1899. Synopses of North-American Invertebrates, IV: Astacoid and
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1955. Two New Species of Entocythere (Ostracoda: Cytheridae) Commen-
sal on Pacifastacus Gambelii (Girard). American Midland Natural-
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LaCaze, Cecil
1970. Crawfish Farming. 27 pages. Louisiana Wildlife and Fisheries
Commission.
Laguarda, Alfredo
1961. Contribucidn al estudio comparative de la formula branquial en la
Familia Astacidae (Crustacea: Decapoda). 74 pages, 22 plates.
Facultad de Ciencias, Departamento de Biologia, Universidad
Nacional Autonoma de Mexico.
1962. Estudio comparado de las pleurobranquias vestigiales del genero
Astacus. Anales del Instituto de Biologia, Universidad Nacional
Autonoma de Mexico, 33(1,2):227-285,4 plates.
Lake, P.S., and KJ. Newcombe
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1952. Six New Crayfishes of the Genus Orconectes (Decapoda: Astacidae)
no SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
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Figures 1-379
Explanation of Illustrations
All of the illustrations except Figure I4d, which was redrawn from Villalobos
(1952:527), and Figures 365 and 369, which were modified from Buckup and Rossi
(1980), were made from camera lucida drawings sketched during the preparation of this
manuscript or its predecessor (Hobbs, 1974b). The lettering used in Figure 1 applies to
Figures 2 through 7; that in Figure 8, to Figures 9 through 123,128 through 210, and 375
through 378; that in Figure 124, to Figures 125 through 127; that in Figure 211, to Figures
212 through 364, and Figure 379; and that in Figure 365, to Figures 366-374.
FIGURES 1 -7.?a, dorsal view of carapace of male; b, mesial view of first pleopod of male;
c, lateral view of first pleopod of male; d, dorsal view of chela of male.
FIGURES 8-123, 128-210, 375-378.?a, dorsal view of carapace of male, form I; b,
mesial view of first pleopod of male, form I; c, lateral view of first pleopod of male, form
I; d, lateral view of first pleopod of male, form II; e, annulus ventralis and postannular
sclerite;/, dorsal view of right chela of male, form I. (Figure \4d after Villalobos, 1952.)
FIGURES 124-127.?a, dorsal view of carapace of male, form I; b, mesial view of first
pleopod of male, form I; c, caudal view of first pleopod of male, form I; d, caudal view of
first pleopod of male, form II; e, annulis ventralis and postannular sclerites;/, dorsal view
of right chela of male, form I.
FIGURES 211-364,379.?a, dorsal view of carapace of male, form I; b, lateral view of first
pleopod of male, form I; c, lateral view of distal part of first pleopod of male, form I; d,
mesial view of distal part of first pleopod of male, form I; e, lateral view of first pleopod
of male, form 11;/, annulus ventralis and postannular sclerite; g, dorsal view of right chela
of male, form I.
FIGURES 365-374.?a, dorsal view of carapace; b, epistome and basal podomere of
antennae; c, caudal view of basal podomeres of fifth pereiopods; d, dorsal view of carpus
of right cheliped; e, lateral view of carpus of right cheliped; /, dorsal view of chela.
(Figures 365 and 369 after Buckup and Rossi, 1980.)
I l l
112 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
I. Pacifastacus (Hobbsastacus) connectens
2. Pacifastacus IHobbsastacus) gambelii
4 Pacifastacus I Hobbsastacus) nigrescens
3. Pacifastacus (Hobbsastacus) fortis
5. Pacifastacus (Pacifastacus) leniusculus leniusculus
ft Pacifastacus {Pacifastacus) leniusculus klamathensts 7. Pacifastacus (Pacifastacus} leniusculus trowbridgii
NUMBER 480 113
a b
8. Cambarellus (Cambarellus) occidentalis
9. Cambarellus (Cambarellus) alvarezi
10. Cambarellus (Cambarellus) chapalanus
,
11. Cambarellus (Cambarellus) chihuahuae
114 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
12. Cumbarellus (Cambarellus) areolatui
13. Cumbarellus fCamburellus) prolixus
14 Cambarellus (Cambarellus) montezumae
15. Cambarellus (Cambarellus) patzcuarensis
NUMBER 480 115
16. Cambarellus (Cambarellus) zempoalensis
17. Cambarellus (Dirigicambarus) shufeldtii
18. Cambarellus (Pandicambarus) blacki
19. Cambarellus (Pandicambarus) schmitti
116 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
20. Cambarellus (Pandicambarus) lesliei
21. Cambarellus (Pandicambarus) puer
22. Cambarellus (Pandicambarus) texanus
v
23. Cambarellus (Pandicambarus) ninae
NUMBER 480 117
24. Cambarellus (Pandicambarus) diminutus
25. Barbicambarus cornutus
26. Bouchardina robisoni
27. Cambarus (Aviticambarus) hamulatus
118 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
28. Cambarus (Aviticambarus) jonesi
29. Cambarus (Cambarus) bartonii hartnnii
30. Cambarus (Cambarus) bartonii carinirostris
31. Cambarus (Cambarus) bartonii cavatus
NUMBER 480 119
32. Cambarus (Cambarus) howardi
33. Cambarus (Cambarus) sciotensis
34. Cambarus {Cambarus) ortmanni
35. Cambarus (Depressicambarus) halli
120 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
36. Cambarus (Depressicambarus) englishi
37. Cambarus (Depressicambarus) obstipus
38. Cambarus (Depressicambarus) latimanus
39. Cambarus (Depressicambarus) reflexus
NUMBER 480 121
40. Cambarus (Depressicambarus) pyronotus
41. Cambarus (Depressicambarus) sphenoides
42. Cambarus (Depressicambarus) catagius
43. Cambarus (Depressicambarus) truncatus
122 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
44. Cambarus (Depressicambarus) cymatilis
45. Cambarus (Depressicambarus) strigosus
46. Cambarus (Depressicambarus)
unknown
47. Cambarus (Depressicambarus) harti
NUMBER 480 123
48. Cambarus (Depressicambarus) striatus
49. Cambarus (Depressicambarus) graysoni
50. Cambarus (Depressicambarus) reduncus
51. Cambarus (Erebicambarus) hubbsi
124 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
52. Cambarus (Erebicambarus) rusticiformis
53. Cambarus (Erebicambarus) hubrichti
54. Cambarus (Erebicambarus) tenebrosus
55. Cambarus (Erebicambarus) ornatus
NUMBER 480 125
56. Cambarus (Erebicambarus) laevis
57. Cambarus (Exilicambarus) cracens
58. Cambarus (Hiaticambarus) coosawattae
59. Cambarus (Hiaticambarus) fasciatus
126 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cnmhnrus IHintirnmhnru<!\ oirnrilinnuR
61. Cambarus (Hiaticambarus) longirostris
62. Cambarus (Hiaticambarus) loneulus
63. Cambarus {Hiaticambarus) manningi
NUMBER 480 127
64. Cambarus (Hiaticambarus) chasmodactylus
65. Cambarus (Hiaticambarus) speciosus
66. Cambarus (Jugicambarus) bouchardi
67. Cambarus (Jugicambarus) obeyensis
128 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
69. Cambarus (Jueicambarus) unestami
70. Cambarus (JuQicamhnru<:\ nnrvnrulm
71. Cambarus (Jugicambarus) conasaugaensis
NUMBER 480 129
72. Cambarus (Jugicambarus) asperimanus
73. Cambarus (Jugicambarus) crinipes
74. Cambarus (Jugicambarus) brachydactylus
75. Cambarus (Jugicambarus) friaufi
130 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
76. Cambarus (Jugicambarus) nodosus
77. Cambarus (Jugicambarus) carolinus
78. Cambarus (Jugicambarus) dubius
79. Cambarus (Jugicambarus) monongalensis
NUMBER 480 131
80. Cambarus (Jugicambarus) batchi
81. Cambarus (Jugicambarus) gentryi
82. Cambarus (Jugicambarus) cryptodytes
83. Cambarus (Jugicambarus) causeyi
132 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
84. Cambarus (Jugicambarus) setosus
85. Cambarus (Jugicambarus) zophonastes
86. Cambarus (Jugicambarus) tartarus
87. Cambarus (Lacunicambarus) acanthura
NUMBER 480 133
88. Cambarus (Lacunicambarus) diogenes
89. Cambarus (Lacunicambarus) ludovicianus
90. Cambarus (Lacunicambarus) miltus
91. Cambarus (Puncticambarus) coosae
134 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
92. Cambarus (Puncticambarus) extraneus
94. Cambarus (Punrtirnmhnr,M*\ or,;?...-
95. Cambarus (Puncticambarus) cumberlandensis
NUMBER 480 135
96. Cambarus (Puncticambarus) parrishi
97. Cambarus (Puncticambarus) nerterius
98. Cambarus (Puncticambarus) buntingi
4 1
?
99. Cambarus (Puncticambarus) acuminatus
136
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
100. Cambarus (Puncticambarus) scotti
101. Cambarus (Puncticambarus) chaueaensis
102. Cambarus (Puncticambarus) hiwasseensis
MM it*tf
103. Cambarus (Puncticambarus) reburrus
NUMBER 480 137
104. Cambarus (Puncticambarus) robustus
105. Cambarus (Puncticambarus) veteranus
106. Cambarus fVeticambarus) pristinus
107. Distocambarus (Distocambarus) crockeri
138 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
108. Distocambarus (Distocambarus) devexus
HIV. Distocambarus (Fitzcambarus) carlsoni
110. Distocambarus {Fitzcambarus) xoungineri
III. Fallicambarus (Fallicambarus) macneesei
NUMBER 480 139
112. Fallicambarus (Fallicambarus) harpi
113. Fallicambarus (Fallicambarus) jeanae
114. Fallicambarus (Fallicambarus) strawni
115. Fallicambarus (Fallicambarus) dissitus
140 SMfraSONIAN CONTRIBUTIONS TO ZOOLOGY
116. Fallicambarus (Creaserinus) fodiens
117. Fallicambarus (Creaserinus) hedgpethi
118. Fallicambarus {Creaserinus) caesius
119. Fallicambarus (Creaserinus) uhleri
NUMBER 480 141
120. Fallicambarus (Creaserinus) hortoni
121. Fallicambarus (Creaserinus) danielae
122. Fallicambarus (Creaserinus) oryktes
123. Fallicambarus (Creaserinus) byersi
142 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
a b
124. Faxonella clypeata
125. Faxonella blairi
126. Faxonella creaseri
127. Faxonella beveri
NUMBER 480 143
128. Hobbseus orconectoides
129. Hobbseus cristatus
130. Hobbseus prominens
131. Hobbseus valleculus
144
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
132. Hobbseus attenuatus
133. Hobbseus petilus
134. Orconectes (Orconectes) australis australis
135. Orconectes (Orconectes) australis packardi
NUMBER 480 145
136. Orconectes (Orconectes) incomptus
137. Orconectes (Orconectes) inermis inermis
138. Orconectes (Orconectes) inermis testii
\
i 1}'
7 v ?' .
139. Orconectes (Orconectes) pellucidus
146 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
140. Orconectes (Faxonius) limosus
141. Orconectes (Faxonius) wrighti
142. Orconectes (Faxonius) indianensis
143. Orconectes (Hespericambarus) difficilis
NUMBER 480 147
144. Orconectes (Hespericambarus) blacki
145. Qrcnnpctps (H4><tnerirnmhnrus\ mnlptnp
146. Orconectes (Hespericambarus) hathawavi
147. Orconectes (Hespericambarus) perfectus
148 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
148. Orconectes (Hespericambarus) deanae
149. Orconectes (Billecambarus) harrisonii
150. Orconectes (Rhoadesius) sloanii
151. Orconectes (Rhoadesius) kentuckiensis
NUMBER 480 149
152. Orconectes (Tragulicambarus) lancifer
153. Orconectes (Crockerinus) marchandi
154. Orconectes {Crockerinus) eupunctus
155. Orconectes (Crockerinus) obscurus
150 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
156. Orconectes (Crockerinus) propinquus
157. Orconectes (Crockerinus) jeffersoni
158. Orconectes (Crockerinus) virginiensis
159. Orconectes (Crockerinus) sanbornii sanbornii
NUMBER 480 151
760. Orconectes (Crockerinus) sanbornii erismophorous
161. Orconectes (Crockerinus) stannardi
162. Orconectes (Crockerinus) erichsonianus
163. Orconectes (Crockerinus) rafinesquei
152 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
164. Orconectes (Crockerinus) bisectus
165. Orconectes {Crockerinus) tricuspis
166. Orconectes (Crockerinus) illinoiensis
167. Orconectes (Crockerinus) shoupi
NUMBER 480 153
168. Orconectes (Procericambarus) quadruncus
169. Orconectes (Procericambarus) leptogonopodus
170. Orconectes (Procericambarus) acares
171. Orconectes (Procericambarus) punctimanus
154 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
172. Orconectes (Procericambarus) hylas
173. Orconectes {Procericambarus) peruncus
174. Orconectes {Procericambarus) rusticus
175. Orconectes {Procericambarus) saxatilis
NUMBER 480 155
776. Orconectes (Procericambarus) forceps
177. Orconectes (Procericambarus) placidus
178. Orconectes (Procericambarus) barrenensi
179. Orconectes (Procericambarus) neglectus chaenodactyh
156 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
[
?
180. Orconectes (Procericambarus) neglectus neglectus
181. Orconectes (Procericambarus) longidigitus
182. Orconectes (Procericambarus) mirus
183. Orconectes (Procericambarus)
NUMBER 480 157
184. Orconectes (Procericambarus) macrus
185. Orconectes (Procericambarus) spinosus
186. Orconectes (Procericambarus) putnami
187. Orconectes (Procericambarus) medius
158 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
188. Orconectes (Procericambarus) luteus
189. Orconectes (Procericambarus) menae
190. Orconectes (Procericambarus) ozarkae
191. Orconectes (Procericambarus) Williams
NUMBER 480 159
792. Orconectes (Buannulifictus) meeki meeki
193. Orconectes (Buannulifictus) meeki brevis
194. Orconectes (Buannulifictus) palmeri palmeri
195. Orconectes (Buannulifictus) palmeri longimanus
160 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
196. Orconectes (Buannulifictus) palmeri creolanus
197. Orconectes (Buannulifictus) hobbsi
198. Orconectes (Gremicambarus) mississippiensis
'
F
/99. Orconectes (Gremicambarus) virilis
NUMBER 480 161
200. Orconectes (Gremicambarus) causeyi
201. Orconectes (Gremicambarus) immunis
202. Orconectes (Gremicambarus) holti
203. Orconectes (Gremicambarus) nais
162 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
204. Orconectes (Gremicambarus) cooperi
205. Orconectes (Gremicambarus) etniet
206. Orconectes (Gremicambarus) alabamensis
, J K,
207. Orconectes (Gremicambarus) rhoade
NUMBER 480 163
205. Orconectes (Gremicambarus) validus
209. Orconectes (Gremicambarus) chickasawae
210. Orconectes {Gremicambarus) compressus
a b
211. Procambarus (Acucauda) fitzpatricki
164 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
212. Procambarus (Austrocambarus) vazquezae
213. Procambarus (Austrocambarus) sbordonii
214. Procambarus (Austrocambarus) mexicanus
u n k n o w n u n k n o w n
yj .
215. Procambarus (Austrocambarus) rodriguezi
NUMBER 480 165
? ?
216. Procambarus (Austrocambarus) oaxacae oaxacai
217. Procambarus (Austrocambarus) oaxacae reddelli
218. Procambarus (Austrocambarus) veracruzanus
i-J
v .1
r
Vv;;.: ?".'.?-.':'!
219. Procambarus (Austrocambarus) miranda
166 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
220. Procambarus (Austrocambarus) ruthveni
221. Procambarus (Austrocambarus) zapoapensis
222. Procambarus {Austrocambarus) williamsoni
223. Procambarus (Austrocambarus) acanthophorous
NUMBER 480 167
224. Procambarus (Austrocambarus) llamasi
225. Procambarus (Austrocambarus) pilosimanus
\ ,11
226. Procambarus (Austrocambarus) atkinsoni
227. Procambarus (Austrocambarus) cubensis rivalis
168 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
228. Procambarus (Austrocambarus) cubensis cubensis
229. Procambarus (Austrocambarus) niveus
230. Procambarus (Capillicambarus) hinei
u n k n o w n
H
'}::?'.':?'-':'?:{>'
r.".'..-.?;.--.?
r
231. Procambarus (Capillicambarus) brazoriensis
NUMBER 480 169
232. Procambarus (Capillicambarus) incilis
233. Procambarus (Girardiella) tulanei
234. Procambarus (Girardiella) simulans
1/
235. Procambarus (Girardiella) curdi
170 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
236. Procambarus (Girardiella) reimeri
237. Procambarus (Girardiella) parasimulam
238. Procambarus (Girardiella) regiomontanus
239. Procambarus (Girardiella) gracilis
NUMBER 480 171
u n k n o w n
240. Procambarus (Girardiella) liberorum
241. Procambarus (Girardiella) hagenianus vesticeps
242 Procambarus (Girardiella) hagenianus hagenianus
243. Procambarus (Girardiella) connus
172 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
244. Procambarus (Girardiella) cometes
245. Procambarus (Girardiella) pogum
246. Procambarus (Girardiella) barbiger
247. Procambarus (Hagenides) advena
NUMBER 480 173
248. Procambarus (Hagenides) talpoides
249. Procambarus (Hagenides) carilus
250. Procambarus (Hagenides) geodytes
251. Procambarus (Hagenides) pygmaeus
174 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
252. Procambarus (Hagenides) rogersi ochlocknensis
253. Procambarus (Hagenides) rogersi campestris
254. Procambarus (Hagenides) rogersi expletus
255. Procambarus (Hagenides) rogersi rogersi
NUMBER 480 175
256. Procambarus (Hagenides) truculent us
257. Procambarus (Leconticambarus) alleni
258. Procambarus (Leconticambarus) milleri
unknown
259. Procambarus (Leconticambarus) barbatus
176 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
260. Procambarus (Leconticambarus) latipleurum
261. Procambarus (Leconticambarus) apalachicolae
262. Procambarus {Leconticambarus) econfinae
263. Procambarus (Leconticambarus) escambiensis
NUMBER 480 177
264. Procambarus (Leconticambarus) pubischelae pubischelae
265. Procambarus (Leconticambarus) pubischelae deficiens
266. Procambarus (Leconticambarus) rathbunae
267. Procambarus (Leconticambarus) capillatus
178 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
268. Procambarus (Leconticambarus) kilbyi
269. Procambarus (Leconticambarus) hubbelli
270. Procambarus {Leconticambarus) shermani
r >^
? . - - ? - !
271. Procambarus (Lonnbergius) acherontis
NlflSffiER 480 179
a
272. Procambarus (Mexicambarus) bouvieri
273. Procambarus (Ortmannicus) pictus
274. Procambarus (Ortmannicus) enoplosternum
275. Procambarus (Ortmannicus) chacei
180 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
276. Procambarus (Ortmannicus) epicyrtus
277. Procambarus (Ortmannicus) litosternum
278. Procambarus (Ortmannicus) lepidodactylus
279. Procambarus (Ortmannicus) pubescens
NUMBER 480 181
280. Procambarus (Ortmannicus) hirsutus
281. Procambarus (Ortmannicus) youngi
282. Procambarus (Ortmannicus) angustatus
unknown unknown
283. Procambarus (Ortmannicus) leitheuseri
182 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
284. Procambarus (Ortmannicus) lucifugus alachua
285. Procambarus (Ortmannicus) lucifugus lucifugus
286. Procambarus (Ortmannicus) erythrops
v.
f
r *,,.nininii HJ i \sr imunmcus) jranzi
NUMBER 480 183
288. Procambarus (Ortmannicus) pallidus
289. Procambarus (Ortmannicus) horsti
290. Procambarus (Ortmannicus) orcinus
unknown
291. Procambarus (Ortmannicus) delicatus
184 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
r
292. Procambarus (Ortmannicus) seminolae
293. Procambarus (Ortmannicus) ancylus
294. Procambarus (Ortmannicus) lunzi
295. Procambarus (Ortmannicus) fallax
NUMBER 480 185
296. Procambarus (Ortmannicus) leonensis
297. Procambarus (Ortmannicus) evermanni
298. Procambarus (Ortmannicus) xilitlae
299. Procambarus (Ortmannicus) pycnogonopodus
186 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
300. Procambarus (Ortmannicus) blandingii
301. Procambarus (Ortmannicus) acutus acutus
302. Procambarus {Ortmannicus) acutus cuevachicae
303. Procambarus (Ortmannicus) texanus
NUMBER 480 187
304. Procambarus (Ortmannicus) lecontei
305. Procambarus (Ortmannicus) geminus
r.
306. Procambarus (Ortmannicus) hayi
307. Procambarus (Ortmannicus) lophotus
188 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
508. Procambarus (Ortmannicus) acutissimus
309. Procambarus (Ortmannicus) verrucosus
310. Procambarus (Ortmannicus) jaculus
311. Procambarus (Ortmannicus) hybus
NUMBER 480 189
312. Procambarus (Ortmannicus) planirostris
313. Procambarus (Ortmannicus) mancus
314. Procambarus (Ortmannicus) plumimanus
315. Procambarus (Ortmannicus) medialis
190 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
316. Procambarus (Ortmannicus) pearsei
317. Procambarus (Ortmannicus) marthae
318. Procambarus (Ortmannicus) lewisi
319. Procambarus (Ortmannicus) bivittatus
NUMBER 480 191
320. Procambarus (Ortmannicus) viaeviridis
321. Procambarus (Ortmannicus) caballeroi
t
322. Procambarus (Ortmannicus) villalobosi
u n k n o w n
323. Procambarus (Ortmannicus) toltecae
192 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
324. Procambarus (Ortmannicus) gonopodocristatus
u n k n o w n
325. Procambarus (Paracambarus) ortmannii
326. Procambarus (Paracambarus) paradoxus
327. Procambarus (Pennides) suttkusi
NUMBER 480 193
328. Procambarus (Pennides) natchitochae
329. Procambarus (Pennides) ablusus
330. Procambarus (Pennides) roberti
331. Procambarus (Pennides) elegans
194 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
r
332. Procambarus (Pennides) versutus
7
i i i . Procambarus (Pennides) lylei
334. Procambarus {Pennides) echinatus
335. Procambarus (Pennides) clemmeri
NUMBER 480 195
336. Procambarus (Pennides) dupratzi
337. Procambarus (Pennides) vioscai
338. Procambarus (Pennides) penni
339. Procambarus (Pennides) lagniappe
1% SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
340. Procambarus (Pennides) petersi
341. Procambarus (Pennides) spiculifer
342. Procambarus (Pennides) gibbus
343. Procambarus (Pennides) ouachitae
NUMBER 480 197
344. Procambarus (Pennides) raneyi
a
345. Procambarus (Procambarus) digueti
346. Procambarus (Remoticambarus) pecki
347. Procambarus (Scapulicambarus) strenthi
198 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
348. Procambarus (Scapulicambarus) paeninsulanus
349. Procambarus (Scapulicambarus) howellae
350. Procambarus (Scapulicambarus) okaloosae
351 Procambarus (Scapulicambarus) clarkii
NUMBER 480 199
352. Procambarus (Scapulicambarus) troglodytes
353. Procambarus (Tenuicambarus) tenuis
354. Procambarus (Villalobosus) riojai
J55. Procambarus (Villalobosus) cuetzalanae
200 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
356. Procambarus (Villalobosus) hoffmanni
357. Procambarus (Villalobosus) erichsoni
358. Procambarus (Villalobosus) contrerasi
359. Procambarus (Villalobosus) hortonhobbsi
NUMBER 480 201
360. Procambarus (Villalobosus) zihuateutlensis
361. Procambarus (Villalobosus) xochitlanae
362. Procambarus (Villalobosus) teziutlanensis
363. Procambarus (Villalobosus) tlapacoyanensis
202 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
364. Troglocambarus maclanei
no spec imens a v a i l a b l e
365. Parastacus brasiliensis
366. Parastacus pilimanus
367. Parastacus varicosus
NUMBER 480 203
371. Parastacus pugnax
204 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
372. Parastacus nicoleti
374. Stimo\tui us araucanius
375. Cambarus (Junicambarus) aculabrum
NUMBER 480 205
376. Fallicambarus (Creuserinus) burrisi
377. Fallicambarus (Creaserinus) gordoni
378. Fallicambarus (Fallicambarus) devastator
/ '
?>;;
379. Procambarus (Austrocambarus) olmecorum
Index
(Each name is followed by page numbers and, where applicable, a figure number in
parentheses.)
ablusus, Procambarus, 72
ablusus, Procambarus (Permides), 72,87 (329)
acanihophorus, Procambarus, 52
acanthophorus, Procambarus (Austrocambarus),52,85 (223)
acanthura, Cambarus, 24
acanthura, Cambarus (Lacunicambarus), 24, 83 (87)
acantophorus, Procambarus, 52
acares, Orconectes, 45
scares, Orconectes (Procericambarus),45,85 (170)
acheronlis, Cambarus, 51,62,68,69
acherontis, Cambarus (Cambarus), 62
acherontis, Cambarus (Ortmannicus), 62
acherontis, Procambarus, 62
acherontis, Procambarus (Lonnbergius), 62,86, (271)
acherontis pallidus, Cambarus, 69
Actacus bimaculatus, 81
actus, Procambarus, 64
actus actus, Procambarus (Ortmannicus), 63
Acucauda, subgenus, 4,51
aculabrum, Cambarus (Jugicambarus), 87,91 (375)
acuminatus, Cambarus, 25
acuminatus, Cambarus (Bartonius), 25
acuminatus, Cambarus (Cambarus), 25
acuminatus, Cambarus (Puncticambarus), 25,83 (99)
acuminatus, Procambarus, 25
acuseyi, Orconectes, 39
acutissimus, Cambarus, 63
acutissimus, Procambarus, 63
acutissimus, Procambarus (Ortmannicus), 63,86 (308)
acutissumus, Procambarus, 63
acutus, Cambarus, 63
acutus, Procambarus, 63
acutus acustus, Procambarus, 64
acutus acutus, Procambarus,63
acutus acutus, Procambarus (Ortmanicus),63
acutus acutus, Procambarus (Ortmannicus), 63,86 (301)
acutus acutus, Procambasrus, 64
acutus blandingii. Procambarus,64
acutus cuevachicae, Procambarus, 64
acutus cuevachicae, Procambarus (Ortmannicus), 64,86 (302)
acutus var. A, Cambarus, 63
acutus var. B, Cambarus, 63
ad vena. A JUcus. 51,58
advena, Cambarus, 56,58,59
advena, Cambarus (Cambarus), 58
advena, Cambarus (Ortmannicus), 58
advena, Procambarus, 58,59
advena, Procambarus (Hagenides). 58,59,86 (247)
advena geodytes, Cambarus, 58
af finis, Asucus, 38
?ffinis. Asucus (Asucus), 39
af fin i j , Asucus (Cambarus), 39
af finis, Cambarus, 39
af finis, Camburus, 39
af finis. Faxonius, 39
affinis, Orconectes, 39
Agassizi, Parastacus, 81
agassizi, Parastacus, 81
agassizii, Parastacus, 81
agassizzi, Parastacus, 81
Alabamensis, Cambarus, 39
alabamensis, Cambarus, 39
alabamensis, Cambarus (Faxonius), 39
alabamensis, Faxonius, 39
alabamensis, Faxonius (Faxonius), 39
alabamensis, Orconectes, 39
alabamensis, Orconectes (Gremicambarus), 39,84 (206)
alabamensis, Orconectes (Orconectes), 39
Alleni, Cambarus, 60
alleni, Cambarus, 60
alleni, Cambarus (Cambarus), 60
alleni, Cambarus (Ortmannicus), 60
alleni, Procambarus, 60
alleni, Procambarus (Leconticambarus),60,86 (257)
allenii, Procambarus, 60
allenti, Procambarus, 60
alvarezi, Camharellus, 8
alvarezi, Cambarellus (Cambarellus), 8,82 (9)
americanus, Cambaras, 7
americunus, Cambals, 7
ancylus, Procambarus, 64
ancylus, Procambarus (Ortmannicus), 64, 86 (293)
angustatus, Astacus, 64
angustatus, Cambarus, 64
angustatus, Cambarus (Cambarus), 64
angustatus, Cambarus (Ortmannicus), 64
angustatus, Procambarus, 64
angustatus, Procambarus (Ortmannicus), 64, 86 (282)
apalachicolae, Procambarus, 60
apalachioolae, Procambarus (Leconticambarus) 60,86 (261)
araucanius, Parastacus, 80
araucanius, Samasucus, 3,80,87 (374)
archerontis, Cambarus, 62
areolatus, Cambarellus, 9
areolatus, Cambarellus (Cambarellus), 8,9,82 (12)
argillicola, Bartonius, 29
argillicola, Cambarus, 29
argillicola, Cambarus (Bartonius), 29
asperimanus, Cambarus, 20
asperimanus, Cambarus (Jugicambarus),20,83 (72)
aspinosus, Cambarus, 27
Astacidae, 3 ,4 ,6
Astacidea.3,4
Astacoidea, 3,4
Asucus, 3 ,6 ,7 ,12,13,14,20,24,29, 33,35,38, 43, 51, 58, 60, 63,72,79,80
Asucus, subgenus, 6,7,12,13,38
astacus, 43
Asucus advena, 51,58
Asucus affinis, 38
Asucus angustatus, 64
206
NUMBER 480 207
Astacus Astecus, 53
Astacus Aztecus, 52
Astacus Bartoni, 13
Astacus bartoni, 13
Astacus Bartonii, 3,12,13,51
Astacus bartonii, 13
Astacus bimaculatus, 81
Astacus Blandingii,3,51,63,64
Astacus Handingii,64
Astacus Brasiliensis,79
Astacus brasiliensis, 79
Astacus braziliensis,79
Astacus breviforceps, 6
Astacus Cambarus pellucidus, 45
Astacus Cambaras Stigius, 45
Astacus Carolinus, 21
Astacus chenoderma, 6
Astacus chilensis, 4,80
Astacus chilensis auct., 80
Astacus chiliensis, 80
Astacus chinoderma, 6
Astacus ciliaris, 13
Astacus Cubensis, 52
Astacus cubensis, 52
Astacus fodiens, 29
Astacus fossarum, 77
Astacus fossor, 5,24
Asucus Gambeli, 6
Astacus gambeli, 7
Astacus Gambelii, 6
Astacus gambelii, 7
Asucus gambelii connectens, 6
Asucus gambelii gambelii, 7
Asucus gambelii, 6
Asucus gambelii gambelii, 7
Astacus Gambellii, 7
AsUcus Klamathensis.6,7
Asucus klamathensis,7
Asucus klaymathensis,7
AsUcus laeviusculus, 7
Asucus latimanus, 12,14,15
Asucus leenisculus, 7
Asucus leniusculus,7
Asucus limosus, 33,38
Asucus maniculatus, 77
AsUcus Mexicanus, 52
Asucus mexicanus, 52
Asucus Nicoleti, 80
Asucus nicoleti, 80
Asucus nigrescens,7
Asucus nigrescens fortis,6
Astacus nigrescens nigrescens, 7
Asucus obscurus, 36
AsUcus oreganus, 5
Asucus pellucidus, 45
asucus pellucidus, 45
Asucus penicillatus,60
Asucus pilimanus, 79,80
AsUcus polmanus, 80
Asucus propinquus, 36
Asucus pugnax, 3,80
Asucus pusillus, 13
Asucus Ricoleti, 80
Asucus spiculifer,74
AsUcus spinifrons, 80,81
AsUcus subgrundialis, 6
AsUcus troglodytes, 77
AsUcus trowbridgei, 8
AsUcus trowbridgi, 8
AsUcus Trowbridgii, 8
AsUcus trowbridgii, 8
AsUcus Trownbridgi, 8
Asucus (AsUcus) affinis, 39
Asucus (AsUcus) Bartonii, 13
Astacus (Asucus) Chilensis, 80
Asucus (Asucus) Gambelii, 6
AsUcus (Asucus) gambelii, 7
Asucus (AsUcus) Klamathensis, 7
AsUcus (AsUcus) klamathensis, 7
AsUcus (AsUcus) leniusculus, 7
AsUcus (AsUcus) nigrescens, 7
AsUcus (AsUcus) Trowbridgii, 8
Astacus (AsUcus) trowbridgii, 8
AsUcus (C.) Chilensis, 3
AsUcus (Cambarus) affinis, 39
AsUcus (Cambarus) Bartonii, 13
AsUcus (Cambarus) Blandingii, 64
AsUcus (Cambarus) Carolinus, 21
AsUcus (Cambarus) carolinus, 21
Asucus (Cambarus) chilensis, 80
AsUcus (Cambarus) Cubensis, 52
AsUcus (Cambarus) mexicanus, 52
AsUcus (Cambarus) pellucidus, 45
AsUcus (Cambarus) Wiegmanni, 5
AsUcus (PacifasUcus) leniusculus, 7
Astecus, Asucus, 53
atkinsoni, Cambarus, 52
atkinsoni, Cambarus (Procambarus), 52
Atkinsoni, Procambarus, 52
atkinsoni, Procambarus, 52
atkinsoni, Procambarus (Austrocambarus), 52, 85 (226)
attenuatus, Hobbseus, 32,84
australis, Orconectes, 44
australis australis, Orconectes, 44
australis australis, Orconectes (Orconectes), 44,85 (134)
australis packardi, Orconectes 44
australis packardi, Orconectes (Orconectes), 44,85 (135)
Austrocambarus, subgenus, 4,12,51
Aviticambarus, subgenus, 4,12
Aviticambarus jonesi, 13
ayersi, Cambarus, 23
ayersii, Cambarus, 23
Aztecus, Asucus, 52
Aztecus, Cambarus, 52
aztecus, Cambarus, 52
aztecus, Procambarus, 53
aztecus, Procambarus (Austrocambarus), 53
barbatus, Cambarus, 51,60
barbatus, Cambarus (Cambarus), 60
barbatus, Cambarus (Ortmannicus),60
barbatus, Procambarus, 60,61
barbatus, Procambarus (Leconticambarus),60, 86 (259)
Barbicambarus,3,4,11
Barbicambarus, subgenus, 3
Barbicambarus comutus, 12,82 (25)
barbiger, Procambarus, 55
barbiger, Procambarus (Girardiella), 55,85 (242)
bamenensis, Orconectes, 45
barrenensis, Orconectes (Procericambarus),45,85 (178)
Bartoni, AsUcus, 13
208 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
bartoni, Astacus, 13
bartoni, Cambaras, 13
Bartoni, Cambarus, 13
bartoni, Cambarus, 12,13
bartoni, Cambarus (Bartonius), 13,14,18
bartoni, Canbarus, 13
bartoni, Procambarus, 13
bartoni acuminates, Cambarus, 25
bartoni asperimanus, Cambarus (Cambarus), 20
bartoni bartoni, Cambarus, 13
bartoni bartoni, Cambarus (Cambarus), 13
bartoni bartonii, Cambarus (Cambarus), 13
bartoni brevis, Cambarus, 17
bartoni carinirostris, Cambarus (Cambarus), 13
bartoni cavatus, Cambarus, 14
bartoni cavatus, Cambarus (Cambarus), 14
bartoni laevis, Cambarus, 17
bartoni laevis, Cambarus (Cambarus), 17
bartoni longirostris, Cambarus (Bartonius), 19
bartoni longulus, Cambarus, 20
bartoni montanus, Cambarus, 13
bartoni ornatus, Cambarus, 18
bartoni robustus, Cambarus, 27
bartoni robustus, Cambarus (Cambarus), 27
bartoni scioiensis, Cambarus, 14
bartoni striatus, Cambarus, 16,17
bartoni striatus, Cambarus (Cambarus), 16
Bartoni tenebrosus, Cambarus, 18
bartoni tenebrosus, Cambarus, 12,17,18
Bartoni tenebrosus, Cambarus (Bartonius), 18
bartoni tenebrosus, Cambarus (Bartonius), 18
Bartoni tenebrosus, Cambarus (Bartontius), 18
bartoni tenebrosus, Cambarus (Cambarus), 18
bartoni typicus, Cambarus, 13
bartoni var. robustus, Cambarus (Bartonius), 27
bartoni veteranus, Cambarus, 27
Bartonii, Astacus, 12,13, SI
bartonii, Astacus, 13
Bartonii, Astacus (Astacus), 13
Bartonii, Astacus (Cambarus), 13
Bartonii, Cambarus, 13
bartonii, Cambarus, 13
bartonii, Cambarus (Cambarus), 13
bartonii, Camparus, 14
bartonii acuminatus, Cambarus, 25
bartonii asperimanus, Cambarus, 12,20,82
Bartonii Bartonii, Cambarus, 13
bartonii bartonii, Cambarus, 13,16,21
bartonii bartonii, Cambarus (Cambarus), 13 (29)
bartonii bartonii, Cambarus (cambarus), 13
bartonii carinirostris, Cambarus, 13
bartonii carinirostris, Cambarus (Cambarus), 13,82 (30)
bartonii cavatus, Cambarus, 14
bartonii cavatus, Cambarus (Cambarus), 14,82 (31)
bartonii laevis, Cambarus, 17
bartonii longirostris, Cambarus, 19
bartonii longirostris, Cambarus (Cambarus), 19
Bartonii longulus, Cambarus, 20
bartonii longulus, Cambarus, 20
bartonii montanus, Cambarus, 13
bartonii omatus, Cambarus, 18
bartonii robusta, Cambarus, 27
Bartonii robustus, Cambarus, 27
bartonii robustus, Cambarus, 27
bartonii robustus, Cambarus (Cambarus), 27
bartonii sciotensis, Cambarus, 14
bartonii spinirostris, Cambarus, 19
bartonii striatus, Cambarus, 17
bartonii striatus, Cambarus (Cambarus), 16
bartonii tenebrosus, Cambarus, 18
bartonii tenebrosus, Cambarus (Cambarus), 18
Bartonii var. longirostris, Cambarus, 19
bartonii var. longirostris, Cambarus, 19
Bartonii var. robusta, Cambarus, 27
bartonii veteranus, Cambarus, 27
Bartonius, 3,13,29
Bartonius, subgenus, 11,12,13,14,17,19,20,24,25,29
bartonius, Cambarus, 13
Bartonius argillicola, 29
bartonius bartoni, Cambarus, 13
Bartonius diogenes, 24
Bartonius ortmanni, 14
bartonni, Cambarus, 13
Bartontius, 12
Bartonii us, subgenus, 17
batchi, Cambarus (Jugicambarus),20,83 (80)
baumgartneri, Cambarus, 58
beyeri, Faxonella, 31.84 (127)
beyeri, Orconectes (Faxonella), 31
Billecambarus, subgenus, 3,4,33
bimaculatus, Astacus, 81
bimaculatus, Parastacus, 81
bisectus, Faxonius, 35
bisectus, Orconectes, 35
bisectus, Orconectes (Crockerinus), 35, 84 (164)
bisectus, Orconectes (Orconectes), 35
bivittatus, Procambarus, 64
bivittatus, Procambarus (Ortmannicus), 64, 86 (319)
black i, Cam barellus, 10
blacki.Cambarellus (Pandicambarus), 10, 82, (18)
blacki, Orconectes (Hespericambarus), 42,43,85 (144)
blairi, Faxonella, 31,84
blandingi, Cambarus (Cambarus), 64
blandingi acutis, Cambarus, 63
blandingi acutus, Cambarus,63
blandingi acutus, Cambarus (Cambarus), 63
blandingi acutus, Ortmannicus, 63
blandingi blandingi, Cambarus (Cambarus), 64
blandingiacutus, Cambarus, 63
blandingi-typicus, Cambarus, 64
Blandingii, Astacus, 3,51,63,64
blandingii, Astacus, 64
Blandingii, Astacus (Cambarus), 64
Blandingii, Cambarus, 63,64
blandingii, Cambarus, 63
blandingii, Cambarus (Ortmannicus), 64
blandingii, Procambarus, 64
blandingii, Procambarus (Ortmannicus), 64, 86 (300)
Blandingii, var. acuta, Cambarus,63
blandingii acutus, Cambarus, 63
blandingii acutus, Cambarus (Cambarus), 63
blandingii acutus, Cambarus (Ortmannicus), 63
blandingii acutus, Camborus, 63
blandingii acutus, Procambarus, 63
blandingii blandingii, Cambarus, 63,64
blandingii blandingii, Procambarus, 63,64
blandingii cuevachicae, Cambarus, 64
blandingii cuevachicae, Procambarus,64
blandingii var. acuta, Cambarus, 63
Blandingii var. Blandingii, Cambarus, 64
NUMBER 480 209
bouchardi, Cambarus, 20
bouchardi, Cambarus (Jugicambarus), 20, 83 (66)
bouchardi, Cambarus (Veticambarus),20
Bouchardina, 3, 4, 12
Bouchardina robisoni, 12, 82 (26)
Bourchardina, 12
Bourchardina robinsoni, 12
boutoni, Cambarus, 13
bouvieri, Cambarus, 63
bouvieri, Cambarus (Cambarus), 51,63
bouvieri, Cambarus (Ortmannicus), 63
bouvieri, Procambarus, 63
bouvieri, procambarus, 63
bouvieri, Procambarus (Mexicambarus), 63,86 (272)
brachydactylus, Cambarus, 20
brachydactylus, Cambarus (Jugicambarus), 21,83 (74)
brasilensis, Parastacus, 79
Brasiliensis, Astacus, 7
brasiliensis, Astacus, 79
brasiliensis, Parastacus, 79, 87 (365)
braziliensis, Astacus, 79
braziliensis, Parastacus, 79
brazorensis, Procambarus (Capillicambarus), 55
brazoriensis, Procambarus (Capillicambarus), 55, 85 (231)
breviforceps, Astacus, 6
Buannuliflctus, subgenus, 3,4,33,34
buntingi, Cambarus, 25
buntingi, Cambarus (Puncticambarus), 25,83 (98)
burrisi.Fallicainbarus (Creaserinus), 87, 91 (376)
byersi, Cambarus, 29
byersi, Fallicambarus, 29
byersi, Fallicambarus (Creaserinus), 29,84 (123)
byersi, Procambarus, 29
Caambarus, 8
Caambarus montezumae areolatus, 9
caballeroi, Pocambarus, 65
caballeroi, Procambarus (Ortmannicus), 65,86 (321)
cabarus, 12
Cabmarus, 12
Cabmarus (Jugicambarus) zophonastes, 24 (85)
caecus, Cambarus, 45
caesius, Fallicambarus, 29
caesius, Fallicambarus (Creaserinus), 29,84 (118)
cahni, Cambarus, 18
cahni, Cambarus (Cambarus), 18
cahni, Cambarus (Erebicambarus), 18
calrkii, Cambarus, 76
Camabarus, 12
Camarus, 35
Camarus hamuetus, 12
Camarus propinquus, 37
Cambals.6,7
Cambals americunus, 7
Cambaras,6,7,12,13
Cambaras americanus, 7
Cambaras bartoni, 13
Cambaraus, 12
CambareUa.8,10
Cambarella shufeldti, 10
Cambarellinae, 3,4,8
Cambarellus, 1,3,4,8,10,51
Cambarcllus, subgenus, 4,8,10
Cambarellus alvarezi, 8
Cambarellus areolatus, 9
Cambarellus blacki, 10
Cambarellus campalanus,9
Cambarellus chapalanus, 9
Cambarellus chihauhuae, 9
Cambarellus chihuahuae, 9
Cambarellus cubensis, 52
Cambarellus diminutus, 10
Cambarellus leslei, 11
Cambarellus lesliei, 11
Cambarellus montazumae, 9
Cambarellus Montezumae, 9
Cambarellus montezumae, 9
Cambarellus montezumae areolatus, 8
Cambarellus montezumae chapalanus, 9
Cambarellus montezumae dugesi, 9
Cambarellus montezumae dugesii, 9
Cambarellus montezumae forma lermensis,9
Cambarellus montezumae forma zempoalensis, 10
Cambarellus montezumae lermensis, 9
Cambarellus montezumae montezumae, 9
Cambarellus montezumae occidentalism
Cambarellus montezumae patzcuarensis,9
Cambarellus montezumae var. areolata, 9
Cambarellus montezumae var. tridens, 9
Cambarellus montezumae zempoalensis, 10
Cambarellus ninae, 11
Cambarellus occidentalis, 9
Cambarellus patzacuarensis, 9
Cambarellus patzcuarensis,9
Cambarellus prolix us, 10
Cambarellus puer, 11
Cambarellus schmioi, 8,10,11
Cambarellus schufeldi, 10
Cambarellus shufeldti, 10
Cambarellus shufeldtii, 10
Cambarellus shufeldtii, 10
Cambarellus shufeltdii, 10
Cambarellus shulfeldtii, 10
Cambarellus texanus, 11
Cambarellus zempoalensis, 10
Cambarellus (Cambarellus) alvarezi, 8,82 (9)
Cambarellus (Cambarellus) areolatus, 9,82 (12)
Cambarellus (Cambarellus) chapalanus, 9,82 (10)
Cambarellus (Cambarellus) chihuahuae, 9,82 (11)
Cambarellus (Cambarellus) montezumae, 9,82 (14)
Cambarellus (Cambarellus) occidentalis, 9,82 (8)
Cambarellus (Cambarellus) patzcuarensis, 9,10, 82 (15)
Cambarellus (Cambarellus) patzquarensis, 9
Cambarellus (Cambarellus) prolixus, 10,82 (13)
Cambarellus (Cambarellus) zempoalensis, 10,82 (16)
Cambarellus (Dirigicambarus) shufeldtii, 10,82 (17)
Cambarellus (Pandicambarus) blacki, 10,82 (18)
Cambarellus (Pandicambarus) diminutus, 10,82 (24)
Cambarellus (Pandicambarus) lesliei, 11,82 (20)
Cambarellus (Pandicambarus) ninae, 11, 82 (23)
Cambarellus (Pandicambarus) puer, 11,82 (21)
Cambarellus (Pandicambarus) schmitti, 11, 82 (19)
Cambarellus (Pandicambarus) texanus, 11,82 (22)
Cambaridae.4,8
Cambarinae, 3,4,11
Cambarinus, 12
Cambaroides, 12,19
Cambaroides longulus, 20
Cambaroidinae, 3
Cambarrus,33,35
Cambarrus propinquus, 37
210 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambani.51,72
Cambani spiculifer, 75
Cambarug, 12
Cambarus. 3. 4, 6. 8, 10,11. 12, 14, 17, 19, 20,24, 25, 29, 30, 31, 32, 33, 34,
35,38,39,42,43,45,50,51,55,58,60,62,63,72,75,76,77
cambarus, 12
Cambarus. subgenus, 3 ,4 ,11,12,13.14,19,20,24, 25,29,33,43, 51,55, 58,
60,62,63,72,75,76,77
Cambarus acanthura, 24
Cambarus acheroniis, 51,62,68,69
Cambarus acherontis pallidus, 69
Cambarus acuminatus, 25
Cambarus acutissimus, 63
Cambarus acutus, 63
Cambarus acutus var. A, 63
Cambarus acutus var. B, 63
Cambarus advena, 56,58,59
Cambarus advena geodytes, 58
Cambarus affinis, 39
Cambarus Alabamensis, 39
Cambarus alabamensis, 39
Cambarus Alleni, 60
Cambarus alleni, 60
Cambarus angustatus, 64
Cambarus archeronlis, 62
Cambarus argillicola, 29
Cambarus asperimanus, 20
Cambarus aspinosus, 27
Cambarus atkinsoni, 52
Cambarus ayersi,23
Cambarus ayersii, 23
Cambarus Aztecus, 52
Cambarus aztecus, 52
Cambarus barbatus, 51,60,61
Cambarus Bartoni, 13
Cambarus bartoni, 12,13
Cambarus bartoni acuminatus, 25
Cambarus bartoni bartoni, 13
Cambarus bartoni brevis, 17
Cambarus bartoni cavatus, 14
Cambarus bartoni laevis, 17
Cambarus bartoni longulus, 20
Cambarus bartoni montanus, 13
Cambarus bartoni ornatus, 18
Cambarus bartoni robustus, 27
Cambarus bartoni sciotensis, 14
Cambarus bartoni striatus, 17
Cambarus Bartoni tenebrosus, 18
Cambarus bartoni tenebrosus, 12,17
Cambarus bartoni typicus, 13
Cambarus bartoni veteranus, 27
Cambarus Bartonii, 13
Cambarus bartonii, 13
Cambarus bartonii acuminatus, 25
Cambarus bartonii asperimanus, 12,20
Cambarus Bartonii Bartonii, 13
Cambarus bartonii bartonii, 13,16,21
Cambarus bartonii carinirostris, 13
Cambarus bartonii cavatus, 14
Cambarus bartonii laevis, 17
Cambarus bartonii longirostris, 19
Cambarus Bartonii longulus, 20
Cambarus bartonii longulus, 20
Cambarus bartonii montanus, 13
Cambarus bartonii ornatus, 18
Cambarus bartonii robusta, 27
Cambarus Bartonii robustus, 27
Cambarus bartonii robustus, 27
Cambarus bartonii sciotensis, 14
Cambarus bartonii spinirostris, 19
Cambarus bartonii striatus, 17
Cambarus bartonii tenebrosus, 18
Cambarus Bartonii var. longirostris, 19
Cambarus bartonii var. longirostris, 19
Cambarus Bartonii var. robusta, 27
Cambarus bartonii veteranus, 27
Cambarus bartonius, 13
Cambarus bartonius bartoni, 13
Cambarus bartonni, 13
Cambarus baumgartneri,58
Cambarus blandingi acutis,63
Cambarus blandingi acutus, 63
Cambarus blandingiacutus,63
Cambarus blandingi-typicus,64
Cambarus Blandingi i, 64
Cambarus blandingii, 63
Cambarus blandingii acutus, 63
Cambarus blandingii blandingii, 63,64
Cambarus blandingii cuevachicae, 64
Cambarus Blandingii, var. acuu, 63
Cambarus blandingii var. acuu, 63
Cambarus Blandingii var. Blandingii, 64
Cambarus bouchardi,20
Cambarus boutoni, 13
Cambarus bouvieri, 63
Cambarus brachydactylus, 21
Cambarus buntingi, 25
Cambarus byersi, 29
Cambarus caecus,45
Cambarus cahni, 18
Cambarus calrkii, 76
Cambarus carinatus, 75
Cambarus carolinensis, 21
Cambarus Carolinus, 21,56,58
Cambarus carolinus, 21,56
Cambarus carolinus carolinus, 21
Cambarus carolinus dubius, 22
Cambarus carolinus monongalenses, 22
Cambarus carolinus monongalensis, 22
Cambarus catagius, 14
Cambarus causeyi, 21
Cambarus chapalamus, 9
Cambarus chapalanus, 9
Cambarus chaplanus, 9
Cambarus chasmodactylus 19
Cambarus clarki,76
Cambarus clarki paeninsulanus, 77
Cambarus Clarkii, 76
Cambarus clarkii, 76
Cambarus clarkii clarkii, 76
Cambarus clarkii paeninsulanus, 51,77
Cambarus clarkipaenensulanus,76
Cambarus clypeatus, 3,31
Cambarus Coecus, 45
Cambarus coecus, 45
Cambarus compressus, 40
Cambarus conasaugaensis,21
Cambarus conasougaensis, 21
Cambarus consobrinus, 52
Cambarus contrerasi,77
NUMBER 480 211
Cambarus cornutus, 12
Cambarus Couesi, 42
Cambarus couesi, 42
Cambarus cousei, 42
Cambarus cousii, 42
Cambarus creolanus, 34
Cambarus crinipes, 21
Cambarus crisUtus, 32
Cambarus cryptodytes, 22
cambarus cryptodytes, 22
Cambarus Cubcnsis, 52
Cambarus cubensis consobrinus, 52
Cambarus cubensis cubensis, 52
Cambarus cubensis rivalis,52
Cambarus cumberlandensis, 26
Cambarus cymatilis, 15
Cambarus debilis, 42
Cambarus deweesae, 15
Cambarus difficilis, 33,42,43
Cambarus digneti, 75
Cambarus digniti, 75
Cambarus Digueti, 51,75
Cambarus digueti, 75
Cambarus diistans, 22
Cambarus Diogenenes, 24
Cambarus Diogenes, 24
Cambarus diogenes, 12,24
Cambarus diogenes, ludoviciana, 24
Cambarus Diogenes Diogenes, 24
Cambarus Diogenes diogenes, 24
Cambarus diogenes diogenes, 24
Cambarus diogenes ludoviciana, 24
Cambarus diogenes ludovicianus, 24
Cambarus Diogenes var. Ludoviciana, 24
Cambarus dissitus, 30
Cambarus distans, 22
Cambarus dubius, 22
Cambarus englishi, 15
Cambarus erichsoni, 35
Cambarus erichsonianus, 35
Cambarus erichsonionus, 35
Cambarus evermanni, 66
Cambarus extraneous, 26
Cambarus extraneus, 12,25,26
Cambarus extraneus Girardanius, 19
Cambarus extraneus girardianus, 19
Cambarus extranius, 26
Cambarus extranus, 26
Cambarus Fallax, 66
Cambarus fallax, 66
Cambarus faxonii, 50
Cambarus floridanus, 17
Cambarus fodiens, 29
Cambarus forceps, 33,45
Cambarus fossarum,77
Cambarus fossor, 24
Cambarus friaufi, 22
Cambarus gallinas,58
Cambarus gallinus, 58
Cambarus Gambeli, 7
Cambarus gambelli, 7
Cambarus gentryi, 22
Cambarus Girardianus, 19
Cambarus girardianus, 19
Cambarus gracilis, 56
Cambarus gracillis, 56
Cambarus graysoni, 15
Cambarus gulielmi, 35
Cambarus Hagenianus, 51,55,56
Cambarus hagenianus, 56
Cambarus hagenianus evansi, 57
Cambarus halli, 15
Cambarus hamueatus, 12
Cambarus hamuetus, 12
Cambarus hamulatus, 12
Cambarus harrisoni, 33
Cambarus Harrisonii, 33
Cambarus harrisonii, 33
Cambarus Hayi, 66
Cambarus hayi, 66
Cambarus hedgpethi, 29
Cambarus hedgepethi, 30
Cambarus heini, 55
Cambarus hiawasseensis, 26
Cambarus hinei, 55
Cambarus Hiwasseenis, 26
Cambarus hiwasseensis, 26
Cambarus howardi, 14
Cambarus hubbelli, 61
Cambarus hubbsi, 17
Cambarus hubricht, 17
Cambarus Hubrichti, 17
Cambarus hubrichti, 17
Cambarus hylas, 46
Cambarus Iatimanus, 17
Cambarus immunis, 40,41
Cambarus immunis Hagenianus 41
Cambarus immunis immunis, 40
Cambarus immunis spinirostris, 40
Cambarus immunis var. spinostris, 40
Cambarus indianensis, 38
Cambarus inermis, 44
Cambarus jonesi, 13
Cambarus Jordani, 15
Cambarus jordani, 15
Cambarus juvenalis, 49
Cambarus juvenilis, 49
Cambarus kilbyi, 61
Cambarus Klamathensis, 7
Cambarus laevi, 18
Cambarus laevis, 17
Cambarus lancifer, 33,50
Cambarus Iatimanus, 15,16,17
Cambarus Iatimanus striatus, 16
Cambarus latimus, 15,17
Cambarus LeContei,67
Cambarus Leconlei, 67
Cambarus lecontei, 67
Cambarus limosus, 39
Cambarus lobdelli, 29
Cambarus longerosilis, 19
Cambarus longidigitus, 46
Cambarus lungimanus, 35
Cambarus longimonus, 35
Cambarus longirostm, 19
Cambarus longulus, 12,19,20
Cambarus longulus chasmodactylus, 19
Cambarus longulus longerostris, 19
Cambarus longulus longirosilis, 19
Cambarus longulus longirostns, 19
212 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambanis longulus longulous, 20
Cambarus longulus longulus, 20
Cambanis lucifugus, S
Cambarus lucifugus alachua, 68
Cambarus lucifugus lucifugus, 68
Cambarus ludovicianus, 24
Cambarus ludovicianus ludovicianus, 25
Cambanis lunzi, 69
Cambarus macneese, 31
Cambarus maniculatus,77
Cambanis medius, 47
Cambanis meeki, 34
Cambanis meekii, 34
Cambanis Mexicanus, 52
Cambanis mexicanus, 52
Cambanis miltus, 25
Cambanis Mississippiensis, 41
Cambanis mississippiensis, 41
Cambanis mongalensis, 22
Cambanis rnonongalensis, 22
Cambanis montanus, 13
Cambanis montanus acuminatus, 25
Cambanis montanus montanus, 13
Cambanis Montezumae, 8,9
Cambanis montezumae, 9
Cambanis montezumae areolatus, 8
Cambanis montezumae chapalanus, 9
Cambanis montezumae dugesi, 9
Cambanis montezumae dugesii, 9
Cambanis Montezumae Montezumae, 9
Cambanis montezumae montezumae, 9
Cambanis montezumae occidenialis, 9
Cambanis Montezumae var. areolata, 8
Cambanis Montezumae var. tridens, 9
Cambanis montezumae var. tridens, 9
Cambanis nais, 41
Cambanis Nais, 41
Cambanis nais, 41
Cambanis Nebrascensis, 24
Cambanis nebrascensis, 24
Cambanis neglecta, 48
Cambanis neglectus, 47
Cambanis nerterius,26
Cambarus obesus, 24
Cambarus obesus var. latimanus, 15
Cambarus obeyensis, 23
Cambanis obscura, 36
Cambanis obscunis, 36
Cambanis obscunis sanbomi, 37
Cambanis obstipus, 16
Cambanis omatus, 18
Cambanis ortmanni, 14
Cambanis oryktes, 30
Cambanis paeninsulanus, 77
Cambarus pallidus, 69
Cambanis Palmeri, 33,34,35
Cambanis palmeri, 35
Cambanis palmeri longimanus, 35
Cambarus palmeri palmeri, 35
Cambarus paradoxus, 72
Cambarus parvoculus, 23
Cambarus Pealei, 39
Cambarus pealei, 39
Cambarus Pellucidus, 45
Cambanis pellucidus, 45
Cambarus pellucidus australis,44
Cambarus pellucidus inermis, 44
Cambarus pellucidus pellucidus, 45
Cambarus pellucidus testi, 44
Cambarus pellucidus testii, 44
Cambarus pellucidus var. testii, 44
Cambarus pellucidus variety incrmis, 44
Cambarus pelludicus, 45
Cambarus pelosus, 41
Cambarus pelucidus, 45
Cambarus pelulcidus, 45
Cambarus penicillatus, 60,64
Cambarus pictus, 70
Cambarus pilosimanus, 53
Cambarus pilosus, 41
Cambarus placidus, 48
Cambanis primaevus, 5,53
Cambanis primoevus, 53
Cambarus pristinus, 12,28
Cambanis prominens, 33
Cambarus propinguus, 39
Cambarus propinkuus, 37,38
Cambarus Propinquus, 37
Cambarus propinquus, 36
Cambanis propinquus obscura, 36
Cambarus propinquus obscurus, 36
Cambarus propinquus propinquus, 36
Cambarus propinquus sanbomi, 37,38
Cambarus propinquus Sanbornii, 37
Cambarus propinquus sanbornii, 37
Cambarus propinquus typicus, 37
Cambarus propinquus var. obscura, 36
Cambarus propinquus var. sanbomi, 37
Cambarus propinquus var. sanbornii, 37
Cambarus pubescens, 70
Cambanis pusillus, 13
Cambarus Putnami, 49
Cambarus putnami, 49
Cambanis pyronotus, 16
Cambarus rathbunae, 62
Cambarus reburms, 27
Cambanis reduncus, 16
Cambarus rinmunis, 40
Cambarus robustis, 27
Cambarus robustus, 27
Cambarus rogersi, 59
Cambarus rusticiformis, 18
Cambarus rustics placidus, 48
Cambarus rusticus, 49
Cambanis rusticus, var. placida, 48
Cambarus rusticus, var. placidus, 48
Cambanis rusticus forceps, 45
Cambanis rusticus placidus, 48
Cambarus rustrineus rustrineus,49
Cambarus ruthveni, 54
Cambarus Sanbornii, 33,35,37
Cambarus schmittii,5,11
Cambarus sciotensis, 14
Cambarus setosus, 23
Cambarus shoupi, 38
Cambarus shufeldti, 10
Cambarus Shufeldtii, 8,10
Cambarus shufeldtii, 10
Cambarus signifer, 40
Cambarus similans, 58
NUMBER 480 213
Cambarus simulana, 58
Cambarus simulans,57
Cambarus sloanei, 50
Cambarus sloani, 50
Cambarus Sloanii, 50
Cambarus sloanii, 33,50
Cambarus sphenoides, 16
Cambarus spicatus, 27
Cambarus spiculifer, 74
Cambarus spinirostris, 19
Cambarus spinosus, 50
Cambarus spinosus gulielmi, 35
Cambarus strawni, 29,30,31
Cambarus striatus, 17
Cambarus Stygius, 45,63
Cambarus stygius, 45,63
Cambarus tartarus, 23
Cambarus tenebrorus, 18
Cambarus tenebrosus, 18
Cambarus troglodyles, 77
Cambarus troglodytes, 77
Cambarus troglodytus, 77
Cambarus typhlobius. 44,45
Cambarus Uhleri, 30
Cambarus uhleri, 30
Cambarus unesiami, 23
Cambarus validus, 42
Cambarus validus Faxonius, 41
Cambarus valleculus, 33
Cambarus versutus,75
Cambarus veteranus, 27
Cambarus viae-viridis,71
Cambarus viridis,42
Cambarus virile, 42
Cambarus viriles, 42
Cambarus virilis, 33,39,42
cambarus virilis, 42
Cambarus virilus, 42
Cambarus vusticus, 49
Cambarus whitmani, 46
Cambarus Wiegmanni, 5
Cambarus wiegmanni, 5
Cambarus wisconsiensis, 42
Cambarus wisconsinensis, 42
Cambarus zophonastes, 24
Cambarus (Aviticambarus) hamulatus, 12,82 (27)
Cambarus (Aviticambarus) jonesi, 13,82 (28)
Cambarus (Barbicambarus) comutus, 10
Cambarus (Bartonius) acuminatus, 25
Cambarus (Bartonius) argillicola, 29
Cambarus (Bartonius) barton i, 13,14,18
Cambarus (Bartonius) bartoni longirostris, 19
Cambarus (Bartonius) Bartoni tenebrosus, 18
Cambarus (Bartonius) bartoni tenebrosus, 18
Cambarus (Bartonius) bartoni var. robustus, 27
Cambarus (Bartonius) carolinus, 21
Cambarus (Bartonius) comutus, 12
Cambarus (Bartonius) diogenes, 24
Cambarus (Bartonius) extraneus, 26
Cambarus (Bartonius) fodiens, 29
Cambarus (Bartonius) girardianus, 19
Cambarus (Bartonius) hamulatus, 12
Cambarus (Bartonius) hubbsi, 17
Cambarus (Bartonius) jordani, 16
Cambarus (Bartonius) latimanus, 16
Cambarus (Bartonius) latimanus var. striatus, 16
Cambarus (Bartonius) longulus, 20
Cambarus (Bartonius) monongalensis, 22
Cambarus (Bartonius) ortmanni, 14
Cambarus (Bartonius) robustus, 27
Cambarus (Bartonius) setosus, 23
Cambarus (Bartonius) uhleri, 30
Cambarus (Bartontius) Bartoni tenebrosus, 18
Cambarus (Cambarellus) chapalanus, 9
Cambarus (Cambarellus) Montezumae, 9
Cambarus (Cambarellus) montezumae, 9
Cambarus (Cambarellus) montezumae areolatus, 8
Cambarus (Cambarellus) montezumae dugesi, 9
Cambarus (Cambarellus) montezumae occidentals, 9
Cambarus (Cambarellus) shufeldti, 10
Cambarus (Cambarus) acherontis, 62
Cambarus (Cambarus) acuminatus, 25
Cambarus (Cambarus) advena, 58
Cambarus (Cambarus) alleni, 60
Cambarus (Cambarus) angustatus, 64
Cambarus (Cambarus) barbatus, 60
Cambarus (Cambarus) bartoni asperimanus, 20
Cambarus (Cambarus) bartoni bartoni, 13
Cambarus (Cambarus) bartoni bartonii, 13
Cambarus (Cambarus) bartoni carinirostris, 13
Cambarus (Cambarus) bartoni cavatus, 13
Cambarus (Cambarus) bartoni laevis, 17
Cambarus (Cambarus) bartoni robustus, 27
Cambarus (Cambarus) bartoni striatus, 16
Cambarus (Cambarus) bartoni tenebrosus, 18
Cambarus (Cambarus) bartonii, 13
Cambarus (Cambarus) bartonii bartonii, 13,82 (29)
Cambarus (cambarus) bartonii bartonii, 13
Cambarus (Cambarus) bartonii carinirostris, 13,82 (30)
Cambarus (Cambarus) bartonii cavatus, 14,82 (31)
Cambarus (Cambarus) bartonii longirostris, 19
Cambarus (Cambarus) bartonii robustus, 27
Cambarus (Cambarus) bartonii striatus, 16
Cambarus (Cambarus) bartonii tenebrosus, 18
Cambarus (Cambarus) blandingi, 64
Cambarus (Cambarus) blandingi acutus, 63
Cambarus (Cambarus) blandingi blandingi, 64
Cambarus (Cambarus) blandingii acutus, 63
Cambarus (Cambarus) bouvieri, 51,63
Cambarus (Cambarus) cahni, 18
Cambarus (Cambarus) carolinus, 21
Cambarus (Cambarus) clarki, 76
Cambarus (Cambarus) contrerasi, 77
Cambarus (Cambarus) comutus, 12
Cambarus (Cambarus) cryplodytes, 21
Cambarus (Cambarus) cubensis, 52
Cambarus (Cambarus) digueti, 75
Cambarus (Cambarus) diogenes, 24
Cambarus (Cambarus) evermanni, 66
Cambarus (Cambarus) extraneus, 25,26
Cambarus (Cambarus) extraneus girardianus, 19
Cambarus (Cambarus) fallax, 66
Cambarus (Cambarus) floridanus, 17
Cambarus (Cambarus) fodiens, 29
Cambarus (Cambarus) gracilis, 56
Cambarus (Cambarus) hagenianus, 56
Cambarus (Cambarus) hamulatus, 12
Cambarus (Cambarus) hayi, 66
Cambarus (Cambarus) hinei, 51,55
Cambarus (Cambarus) howardi, 14,83
214 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Cambarus (Cambarus) jordani, 16
Cambarus (Cambarus) latimanus, 16
Cambarus (Cambarus) latimanus striatus, 16
Cambarus (Cambarus) lecontei, 67
Cambarus (Cambarus) longulus longirostris, 19
Cambarus (Cambarus) longulus longulus, 20
Cambarus (Cambarus) mexicanus, 52
Cambarus (Cambarus) monongalensis, 22
Cambarus (Cambarus) montanus acuminatus,25
Cambarus (Cambarus) montanus montanus, 13
Cambarus (Cambarus) montanus veteranus, 27
Cambarus (Cambarus) ortmanni, 14, 83 (34)
Cambarus (Cambarus) peUucidus, 45
Cambarus (Cambarus) pubescens, 70
Cambarus (Cambarus) scioiensis, 14, 83 (33)
Cambarus (Cambarus) setosus, 23
Cambarus (Cambarus) simulans, 58
Cambarus (Cambarus) spiculifer, 75
Cambarus (Cambarus) troglodytes, 77
Cambarus (Cambarus) uhleri, 30
Cambarus (Cambarus) versutus, 75
Cambarus (Depressicambarus) catagius, 14,83 (42)
Cambarus (Depressicambarus) cymatilis, 14,83 (44)
Cambarus (Depressicambarus) deweesae, 15, 83 (46)
Cambarus (Depressicambarus) diogenes, 24
Cambarus (Depressicambarus) englishi, 15,83 (36)
Cambarus (Depressicambarus) floridanus, 17
Cambarus (Depressicambarus) graysoni, 15,17,83 (49)
Cambarus (Depressicambarus) halli, 15, 83 (35)
Cambarus (Depressicambarus) harti, 15,83 (47)
Cambarus (Depressicambarus) jordani, 16
Cambarus (Depressicambarus) latimanus, 15,16,83 (38)
Cambarus (Depressicambarus) obstipus, 16,83 (37)
Cambarus (Depressicambarus) pyronotus, 16, 83 (40)
Cambarus (Depressicambarus) reduncus, 16,83 (50)
Cambarus (Depressicambarus) reflexus, 16,83 (39)
Cambarus (Depressicambarus) sphenoides, 16,83 (41)
Cambarus (Depressicambarus) striatus, 15,16,17,83 (48)
Cambarus (Depressicambarus) strigosus, 17,83 (45)
Cambarus (Depressicambarus) truncatus, 17,83 (43)
Cambarus (Depressicambarus) unestami, 23
Cambarus (Erebicambarus) cahni, 18
Cambarus (Erebicambarus) hubbsi, 17, 83 (51)
Cambarus (Erebicambarus) hubrichti, 17,83 (53)
Cambarus (Erebicambarus) Laevis, 18
Cambarus (Erebicambarus) laevis, 17.18, 83 (56)
Cambarus (erebicambarus) laevis, 17
Cambarus (Erebicambarus) maculatus, 87,91
Cambarus (Erebicambarus) omatus, 18,83 (55)
Cambarus (Erebicambarus) rusticiformis, 18,83 (52)
Cambarus (Erebicambarus) tenebrosus, 18,83 (54)
Cambarus (Erebicambaruss) laevis, 18
Cambarus (Eribicambarus) laevis, 18
Cambarus (Exilicambanis) cracens, 12.18,83 (57)
Cambarus (Faxonius) alabamensis, 39
Cambarus (Faxonius) compressus, 40
Cambarus (Faxonius) creaseri.35
Cambarus (Faxonius) difficilis, 43
Cambarus (Faxonius) difficulis, 43
Cambarus (Faxonius) erichsonianus, 35
Cambarus (Faxonius) forceps. 45
Cambarus (Faxonius) harrisoni. 34
Cambarus (Faxonius) hylas, 46
Cambarus (Faxonius) immunis, 41
Cambarus (Faxonius) indianemis, 38
Cambarus (Faxonius) juvenilis, 49,50
Cambarus (Faxonius) lancifer, 50
Cambarus (Faxonius) limosus, 39
Cambarus (Faxonius) longidigitus, 46
Cambarus (Faxonius) medius, 47
Cambarus (Faxonius) meeki, 34
Cambarus (Faxonius) mississippiensis, 41
Cambarus (Faxonius) nais, 41
Cambarus (Faxonius) neglectus, 47
Cambarus (Faxonius) obscurus, 36
Cambarus (Faxonius) palmeri, 35
Cambarus (Faxonius) pellucidus, 45
Cambarus (Faxonius) pellucidus australis,44
Cambarus (Faxonius) pellucidus testi, 44
Cambarus (Faxonius) peruncus, 48
Cambarus (Faxonius) pilosus, 41
Cambarus (Faxonius) propinquus, 37
Cambarus (Faxonius) propinquus sanborni, 37
Cambarus (Faxonius) putnami, 49
Cambarus (Faxonius) rusticus, 49
Cambarus (Faxonius) rusticus forceps, 45
Cambarus (Faxonius) rusticus minis, 47
Cambarus (Faxonius) rusticus placidus, 48
Cambarus (Faxonius) rusticus rusticus, 49
Cambarus (Faxonius) sloanei, 50
Cambarus (Faxonius) spinosus, 50
Cambarus (Faxonius) validus, 42
Cambarus (Faxonius) virilis, 42
Cambarus (Fexonius) pellucidus, 45
Cambarus (Fraxonius) propinquus, 36
Cambarus (Girardiella) gracilis, 56
Cambarus (Girardiella) hagenianus carri, 57
Cambarus (Girardiella) hagenianus evansi, 57
Cambarus (Girardiella) hagenianus forestae, 55
Cambarus (Girardiella) hagenianus forrestae, 55
Cambarus (Girardiella) hagenianus hagenianus, 56
Cambarus (Hiaticambarus) chasmodactylus, 19, 83 (64)
Cambarus (Hiaticambarus) coosae, 25
Cambarus (Hiaticambarus) coosawattae, 19, 83 (58)
Cambarus (Hiaticambarus) fasciatus, 19, 83 (59)
Cambarus (Hiaticambarus) girardianus, 19,83 (60)
Cambarus (Hiaticambarus) longirostris, 19, 83 (61)
Cambarus (Hiaticambarus) longulus, 20,83 (62)
Cambarus (Hiaticambarus) manningi, 20, 83 (63)
Cambarus (Hiaticambarus) speciosus,20,83 (65)
Cambarus (Jugicambarus) aculabrum, 87,91 (375)
Cambarus (Jugicambarus) asperimanus, 20, 83 (72)
Cambarus (Jugicambarus) batchi, 20, 83 (80)
Cambarus (Jugicambarus) bouchardi, 20, 83 (66)
Cambarus (Jugicambarus) brachydactylus, 21,83 (74)
Cambarus (Jugicambarus) carolinus, 21, 83 (77)
Cambarus (Jugicambarus) causeyi, 21,83 (83)
Cambarus (Jugicambarus) conasaugaensis, 21,83 (71)
Cambarus (Jugicambarus) crinipes, 21,83 (73)
Cambarus (Jugicambarus) cryptodytes, 21,22,83 (82)
Cambarus (Jugicambarus) crytodytes, 22
Cambarus (Jugicambarus) distans, 22, 83 (68)
Cambarus (Jugicambarus) dubius, 22, 83 (78)
Cambarus (Jugicambarus) friaufi, 22,83 (75)
Cambarus (Jugicambarus) gentryi,22,83 (81)
Cambarus (Jugicambarus) monongalensis, 22,83 (79)
Cambarus (Jugicambarus) nodosus, 23,83 (76)
Cambarus (Jugicambarus) obeyensis, 23,83 (67)
Cambarus (Jugicambarus) parvoculus, 23. 83 (70)
Cambarus (Jugicambarus) setosus, 23,83 (84)
NUMBER 480 215
Cambarus (Jugicambarus)lartarus,23, 83 (86)
Cambarus (Jugicambarus) unestami, 23, 83 (69)
Cambarus (Jugicambarus) zophonastes,24,83 (85)
Cambarus (Lacunacambarus) diogenes diogenes, 24
Cambarus (Lacunicambarus) acanthura, 24,83 (87)
Cambarus (Lacunicambarus) diogenes, 24,83 (88)
Cambarus (Lacunicambarus) diogenes Diogenes, 24
Cambarus (Lacunicambarus) diogenes diogenes, 24
Cambarus (Lacunicambarus) diogenes ludovicianus, 25
Cambarus (Lacunicambarus) ludovicianus, 24,83 (89)
Cambarus (Lacunicambarus) miltus, 25,83 (90)
Cambarus (Orconectes) pellucidus, 45
Cambarus (Ortmannicus) acherontis, 62
Cambarus (Ortmannicus) advena, 58
Cambarus (Ortmannicus) alleni, 60
Cambarus (Ortmannicus) anguslatus, 64
Cambarus (Ortmannicus) barbatus, 60
Cambarus (Ortmannicus) blandingii, 64
Cambarus (Ortmannicus) blandingii acutus, 63
Cambarus (Ortmannicus) bouvieri, 63
Cambarus (Ortmannicus) clarkii, 76
Cambarus (Ortmannicus) cookae, 67
Cambarus (Ortmannicus) evermanni, 66
Cambarus (Ortmannicus) evictus, 63
Cambarus (Ortmannicus) fallax, 66
Cambarus (Ortmannicus) gracilis, 56
Cambarus (Ortmannicus) hagenianus, 56
Cambarus (Ortmannicus) hayi, 66
Cambarus (Ortmannicus) hinei, 55
Cambarus (Ortmannicus) lecontei, 67
Cambarus (Ortmannicus) pearsei, 70
Cambarus (Ortmannicus) pubescens, 70
Cambarus (Ortmannicus) simulans, 58
Cambarus (Ortmannicus) spiculifer,75
Cambarus (Ortmannicus) troglodytes, 77
Cambarus (Ortmannicus) versutus, 75
Cambarus (Ortmannicus) viae viridis, 71
Cambarus (Paracambarus) harnedi, 51
Cambarus (Paracambarus) paradoxus, 51,72
Cambarus (Paracamburus) paradoxus, 72
Cambarus (Procambarus) atkinsoni, 52
Cambarus (Procambarus) contrerasi, 77
Cambarus (Procambarus) cubensis, 52
Cambarus (Procambarus) digueti, 75
Cambarus (Procambarus) mexicanus, 52
Cambarus (Procambarus) pilosimanus, 53
Cambarus (Procambarus) williamsoni, 54
Cambarus (Puncticambarus) acuminatus, 25,83 (99)
Cambarus (Puncticambarus) buntingi, 25,83 (98)
Cambarus (Puncticambarus) chaugaensis, 25,83 (101)
Cambarus (Puncticambarus) coosae, 25,26,83 (91)
Cambarus (Puncticambarus) cumbeiiandensis,26,83 (95)
Cambarus (Puncticambarus) extraneus, 26,83 (92)
Cambarus (Puncticambarus) georgiae, 26,83 (93)
Cambarus (Puncticambarus) hiwasseensis, 26, 83 (102)
Cambarus (Puncticambarus) nertcrius, 26,83 (97)
Cambarus (Puncticambarus) parrishi, 27,83 (96)
Cambarus (Puncticambarus) rebumis, 27,83 (103)
Cambarus (Puncticambarus) robustus, 27,83 (104)
Cambarus (Puncticambanis) scotti, 27,83 (100)
Cambarus (Puncticambarus) spicatus, 27,84 (94)
Cambarus (Puncticambarus) veteranus, 27,84 (105)
Cambarus (Veticambanis) bouchardi, 20
Cambarus (Veticambanis) obeyensis, 23
Cambarus (Veticambanis) pristinus, 28,84 (106)
Cambaurs, 12
Cambaus, 12,51,76
Cambaus clarkii, 76
Cambaus cubensis, 52
Camberellus, 8
Camberus, 12
Camborus,33,35,51
Camborus blandingii acutus, 63
Camborus propinquus, 37
Cambrarus, 12
Cambraus, 12
Cambraus dubius, 22
Cambrus.6,33,43
Cambrus Gambelii, 6
Cambrus pellucidus, 45
Camburus, 12,16.33,39
Camburus affinis, 39
Camburus viridis, 42
campalanus, Cambarellus, 9
Camparu s, 12
Camparus bartonii, 14
Camurus, 33,43
Camtarus (Orconectes) pellucidus, form incrmis, 45
Canbams, 12
Canbarus bartoni, 13
capillatus, Procambarus, 60
capillatus, Procambarus (Leconlicambarus), 60,86 (267)
Capillicambarus, subgenus, 4,51,55
Carbarinae, 11
Carbarus, 12
Carbarus jonesi, 13
carinatus, Cambarus, 75
caritus, Procambarus (Hagenides), 58,86 (249)
carlsoni, Distocambarus (Fitzcambarus),28,84 (109)
carolinensis, Cambarus, 21
Carolinus, Astacus, 21
Carolinus, Astacus (Cambarus), 21
carolinus, Astacus (Cambarus), 21
carolinus, Cambarus, 21,56
Carolinus, Cambarus, 21,56
carolinus, Cambarus (Bartonius), 21
carolinus, Cambarus (Cambarus), 21
carolinus, Cambarus (Jugicambarus), 21,83 (77)
Carolinus Cambarus, 21,58
carolinus carolinus, Cambarus, 21
carolinus dubius, Cambarus, 22
carolinus monongalenses, Cambarus, 22
carolinus monongalensis, Cambarus, 22
catagius, Cambarus, 14
caugius, Cambarus (Depressicambarus), 14, 83 (42)
causey i, Cambarus, 21
causeyi, Cambarus (Jugicambarus), 21, 83 (83)
causeyi, Orconectes, 39
causeyi, Orconectes (Gremicambarus), 39,40,84 (200)
chacci, Procambarus, 65
chacei, Procambarus (Ortmannicus),65,86 (275)
chapalamus, Cambarus, 9
chapalanus, Cambarellus, 9
chapalanus, Cambarellus (CambareUus). 9, 82 (10)
chapalanus, Cambarus, 9
chapalanus, Cambarus (Cambarellus), 9
chaplanus, Cambarus, 9
chasmodactylus, Cambarus, 19
chasmodactylus, Cambarus (Hiaticambarus), 19,83 (64)
chaugaensis, Cambarus (Puncticambarus), 25,83 (101)
216 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
chenoderma, Asiacus, 6
chenoderma, Pacifastacus, 6
chenoderma, Hacifastacus (Hobbsastacus), 6, 82
ch ickasawae, Orconectes, 40
chickasawae, Orconectes (Gremicambarus),40,84 (209)
chihauhuae, Cambarellus, 9
chihuahuae, Cambarellus, 9
chihuahuae, Cambarellus (Cambarellus),9, 82 (11)
chilensis, Asiacus, 4, 80
Chilensis, Asiacus (Asiacus), 80
Chilensis, Astacus (C), 3
chilensis, Astacus (Cambarus), 80
Chilensis, Parastacus, 80
chilensis, Parastacus, 80
chilensis auct., Asiacus, 80
chiliensis, Astacus, 80
chinoderma, Astacus, 6
ciliaris, Astacus, 13
Cityphlobius, 45
Cityphobius, 33,44
darci i, Procambarus, 76
clarki, Cambarus, 76
clarki, Cambarus (Cambarus), 76
clarki, Procambarius, 76
clarki, Procambarus, 76
clarki, Procambarus (Ortmannicus), 76
clarki paeninsulanus, Cambarus, 77
Clarku, Cambarus, 76
clarkii, Cambarus, 76
clarkii, Cambarus (Ortmannicus), 76
clarkii, Cambaus, 76
clarkii, Comharus, 76
clarkii, Procambambarus, 76
clarkii, Procambarus, 76
clarkii, Procambarus (Cambarus), 76
clarkii, Procambarus (Scapulicambarus),76,87 (351)
clarkii, Procambonis, 76
clarkii, Procambrus, 76
clarkii, Procamburus, 76
clarkii clarkii, Cambarus, 76
clarkii clarkii, Procambarus, 76
clarkii paeninsulanus, Cambarus, 51,76
clarkii paeninsulanus, Procambarus, 77
darkipaenensulanus, Cambarus, 76
demmeri, Procambarus, 73
demmeri. Procambarus (Permides), 73,87 (335)
dypeau, Faxonella. 32,84 (124)
dypeau, Orconectes, 31
dypeau, Orconectes (Faxonella), 31
dypeau, Procambarus, 31
dypeatus, Cambarus, 3,31
dypeatus, Faxonella. 32
dypeatus, Faxonius, 31
dypeatus, Faxonius (Faxonella), 31
dypeatus, Orconectes, 31
dypeatus, Orconectes (Faxonella), 32
dypeu, Faxonella, 32
dyptea, Faxonella, 32
Coecus, Cambarus, 45
coecus, Cambarus, 45
Combarus.51,76
Combarus darkii,76
cometes, Procambaru, 56
cometes. Procambarus (Girardiella),56, 86 (244)
compressus, Cambarus, 40
compressus, Cambarus (Faxonius), 40
compressus, Faxonius, 40
compressus, Faxonius (Faxonius), 40
compressus, Orconectes, 40
compressus, Orconectes (Gremicambarus), 40,84 (210)
compressus, Orconectes (Orconectes), 40
conasaugaensis, Cambarus, 21
conasaugaensis, Cambarus (Jugicambarus),21, 83 (71)
conasougaensis, Cambarus, 21
conectens, Pacifastacus, 6
connectens, Pacifastacus, 6
connectens, Pacifastacus (Hobbsastacus), 6, 82 (1)
connos, Procambarus (Girardiella), 56
connus, Procambarus, 56
connus, Procambarus (Girardiella), 56, 86 (243)
consobrinus, Cambarus, 52
consobrinus, Procambarus, 52
contrerasi, Cambarus, 77
contrerasi, Cambarus (Cambarus),77
contrerasi, Cambarus (Procambarus), 77
contrerasi, Procambarus, 77
contrerasi, Procambarus (Villalobosus), 77,87 (358)
cookae, Cambarus (Ortmannicus), 67
cooperi, Orconectes, 40
cooperi, Orconectes (Gremicambarus), 40, 84 (204)
coosae, Cambarus (Hiaticambarus), 25
coosae, Cambarus (Puncticambarus), 25,26, 83 (91)
coosawattae, Cambarus (Hiaticambarus), 19, 83 (58)
cornutus, Barbicambarus, 12,82 (Fig. 25)
cornutus, Cambarus, 11,12
cornutus, Cambarus (Barbicambarus), 12
cornutus, Cambarus (Bartonius), 12
cornutus, Cambarus (Cambarus), 12
Couesi, Cambarus, 42
couesi, Cambarus, 42
cousei, Cambarus, 42
cousii, Cambarus, 42
cracens, Cambarus (Exilicambarus), 12,18, 83 (57)
Crangonyx lucifugus, 5
creaseri, Cambarus (Faxonius), 35
creaseri, Faxonella, 32, 84 (126)
Creasehnus, subgenus, 29
creolanus, Cambarus, 34
creolanus, Faxonius, 34
creolanus, Faxonius (Faxonius), 34
creolanus, Orconectes, 34
creolanus, Orconectes (Orconectes), 34
Creserineus, subgenus, 29
crinipes, Cambarus, 21
crinipes, Cambarus (Jugicambarus),21, 83 (73)
en status, Cambarus, 32
crisutus, Hobbseus, 32,84
crockeri, Distocambarus, 28
CTOckeri.EHstocambarus (Distocambarus), 28,84 (107)
Crockerinus, subgenus, 3,4,33,35
cryptodytes, Cambarus, 22
cryptodytes, cambarus, 22
cryptodytes, Cambarus (Cambarus), 21
cryptodytes, Cambarus (Jugicambarus),21,22, 83 (82)
crytodytes, Cambarus (Jugicambarus), 22
Cubensis, Astacus, 52
cubensis, AsUcus, 52
Cubensis, AsUcus (Cambarus), 52
cubensis, Cambarellus, 52
Cubensis, Cambarus, 52
NUMBER 480 217
cubensis, Cambarus (Cambarus), 52
cubensis, Cambarus (Procambarus), 52
cubensis, Cambaus, 52
cubensis, Procambarus C , 52
cubensis consobrinus, Cambarus, 52
cubensis consobrinus, Procambarus, 52
cubensis cubensis, Cambarus, 52
cubensis cubensis, Procambarus, 52
cubensis cubensis, Procambarus (Auslrocambarus), 52, 85 (228)
cubensis rivalis, Cambarus, 52
cubensis rivalis, Procambarus, 52
cubensis rivalis, Procambarus (Austrocambarus), 52,85 (227)
cuetzalanae, Procambarus (Villalobosus), 77, 87 (355)
cumberlandensis, Cambarus, 26
cumberlandensis, Cambarus (Puncticambarus), 26,83 (95)
curdi, Procambarus (Girardiella), 56, 86 (235)
cylpeata, Faxonella, 32
cymatilis, Cambarus, 15
cymatilis, Cambarus (Depressicambarus), 14, 83 (44)
danielae, Fallicambarus (Creaserinus),29,84 (121)
deanae, Orconectes, 43
deanae, Orconectes (Hespericambarus),43, 85 (148)
debilis, Cambarus, 42
defossus, Parastacus,79,87
delicatus, Procambarus (Ortmannicus), 65,86 (291)
denae, Orconectes, 43
denae, Orconectes (Buannulifictus), 43
Depressicambarus, subgenus, 4,12,14,20
devastator, Fallicambarus (Fallicambarus), 87,91 (378)
devexus, Distocambarus, 28
devexus, Distocambarus (Distocambarus), 28,84 (108)
devexus, Procambarus (Distocambarus), 28
deweesae, Cambarus, 15
deweesae, Cambarus (Depressicambarus), 15, 83 (46)
difficilis, Cambarus, 33,42,43
difficilis, Cambarus (Faxonius), 43
difficilis, Faxonius 43
difficilis, Faxonius (Faxonius), 43
difficilis, Orconectes, 43
difficilis,Orconectes (Hespericambarus),42,43, 85 (143)
difficilis, Orconectes (Orconectes) 43
difficilis blacki, Orconectes, 42
difficilis difficilis, Orconectes, 43
difficilis hathawayi, Orconectes, 42,43
difficilis maleiac, Orconectes, 43
difficulis, Cambarus (Faxonius), 43
digneti, Cambarus, 75
digniti, Cambarus, 75
Digueti, Cambarus, 51,75
digueti, Cambarus, 75
digueti, Cambarus, (Cambarus) 75
digueti, Cambarus, (Procambarus) 75
Digueti, Procambarus, 75
digueti, Procambarus, 75
digueti, Procambarus (Procambarus), 75,87 (345)
diistans, Cambarus, 22
diminutus, Cambarellus, 10
diminutus, Cambarellus (Pandicambarus), 10, 82 (24).
Diogenenes, Cambarus, 24
diogenes, Bartonius, 24
Diogenes, Cambarus, 24
diogenes, Cambarus, 12.24
diogenes, Cambarus (Bartonius), 24
diogenes, Cambarus (Cambarus), 24
diogenes, Cambarus (Depressicambarus), 24
diogenes, Cambarus (Lacunicambarus),24, 83 (88)
diogenes, Sambarus, 24
Diogenes Diogenes, Cambarus, 24
Diogenes diogenes, Cambarus, 24
diogenes diogenes, Cambarus, 24
diogenes Diogenes, Cambarus (Lacunacambarus), 24
diogenes diogenes, Cambarus (Lacunicambarus), 24
diogenes ludoviciana, Cambarus, 24
diogenes ludovicianus, Cambarus, 24
diogenes ludovicianus, Cambarus (Lacunicambarus), 25
Diogenes var. Ludoviciana, Cambarus, 24
Dirigicambarus, subgenus, 3,4,8,10
dissitus, Cambarus, 30
dissitus, Fallicambarus, 30
dissitus, Fallicambarus (Fallicambarus), 30,84 (115)
distans, Cambarus, 22
distans, Cambarus (Jugicambarus),22,83 (68)
Distocambarus, 4,28
Distocambarus, subgenus, 3,4,28
Distocambarus (Distocambarus) crockeri, 28,84 (107)
Distocambarus (Distocambarus) devexus, 28,84 (108)
Distocambarus (Fitzcambarus) carkoni, 2S, 84 (109)
Distocambarus (Fitzcambarus) youngineri, 28,84 (110)
Distocambarus crockeri, 28
Distocambarus devexus, 28
dubius, Cambarus, 22
dubius, Cambarus (Jugicambarus), 22,83 (78)
dubius, Cambraus, 22
dupratzi, Procambarus, 73
dupratzi, Procambarus (Pennides), 73, 87 (336)
echinatus, Procambarus, 73
echinatus, Procambarus (Pennides), 73, 87 (334)
econf inae, Procambarus, 61
econfinae, Procambarus (Leconticambarus),61,86 (262)
efinifrons, Parastacus, 81
elegans, Procambarus, 73
elegans, Procambaus (Pennides), 73, 87 (331)
englishi, Cambarus, 15
englishi, Cambarus (Depressicambarus), 15,83 (36)
enoplosternum, Procambaris, 65
enoplosternum, Procambarus, 65
enoplosternum, Procambarus (Ortmannicus),65,86 (274)
enoploslerum, Procambarus, 65
epicyrtus, Procambarus, 65
epicyrtus, Procambarus (Ortmannicus), 65,86 (276)
ercichsoni, Procambarus, 78
Erebicambarus, subgenus, 4,12,17
Erebicambaruss, 17
tin hi cambarus, 17
erichsoni, Cambarus, 35
erichsoni, Procambarus, 78
erichsoni, Procambarus (VUlalobosus),78,87 (357)
erichsonianus, Cambarus, 35
erichsonianus, Cambarus (Faxonius), 35
erichsonianus, Faxonius, 35
erichsonianus, Faxonius (Faxonius), 35
erichsonianus, Orconectes, 35
erichsonianus, Orconectes (Crockerinus),35,84 (162)
erichsonianus, Orconectes (Orconectes), 35
erichsonionus, Cambarus, 35
cncichsoni. Procambarui,78
erythrops, Procambarus, 65
erythrops, Procambarus (Ortmannicus), 65, 86 (286)
escambiensis, Procambarus, 61
escambiensis, Procambarus (Leconticambarus),60,61,86 (263)
218 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
elnieri, Orconectes, 40
einieri, Orconectes (Grcmicambarus), 40,84, (205)
eupunctus, Faxonius, 36
eupunctus, Orconecctes, 36
eupunctus, Orconectes, 36
eupunctus, Orconectes (Crockerinus), 36,84 (154)
eupunctus, Orconectes (Orconectes), 36
evansi, Cambarus hagenianus, 57
evermani, Procambarus, 66
evermanni, Cambarus, 66
evermanni, Cambarus (Cambarus), 66
evermanni, Cambarus (Ortmannicus),66
evermanni, Procambarus, 66
evermanni, Procambarus (Ortmannicus),66, 86 (297)
evictus, Cambarus (Ortmannicus), 63
Exilicambarus, subgenus, 3,4,12,18
extraneous, Cambarus, 26
extraneus, Cambarus, 12,25,26
extraneus, Cambarus (Bartonius), 26
extraneus, Cambarus (Cambarus), 25,26
extraneus, Cambarus (Puncticambarus), 26,83 (92)
extraneus Girardanius, Cambarus, 19
extraneus girardianus, Cambarus (Cambarus), 19
extranius, Cambarus, 26
extranus, Cambarus, 26
Fallax, Cambarus, 66
fallax, Cambarus, 66
fallax, Cambarus (Cambarus), 66
fallax, Cambarus (Ortmannicus),66
fallax, Procambaris, 66
fallax, Procambarus, 66
fallax, Procambarus (Ortmannicus), 66,86 (295)
Fallicambanis,3.4.29,30
Fall icambarus, subgenus, 4,30
Fallicambarus byersi. 29
Fall icambarus caesius, 29
Fall icambarus dissitus, 30
[?'allicambarus fodiens, 29
Faliicambarus hedgepethi, 30
Fall icambarus hedgpethi, 30
Faliicambarus hortoni, 30
Faliicambarus macneesei, 31
Faliicambarus oryktes, 30
Faliicambarus strawni, 31
Faliicambarus uhleri, 30
Fallicambanis (Creaserineus), fodiens 29
Faliicambarus (Creaserinus) burrisi, 87.91 (376)
Faliicambarus (Creaserinus) byersi, 29. 84 (123)
Faliicambarus (Creaserinus) caesius, 29, 84 (118)
Faliicambarus (Creaserinus) danielae, 29.84 (121)
Faliicambarus (Creaserinus) fodiens, 29,84 (116)
Faliicambarus (Creaserinus) gordoni, 87,91 (377)
Fallicambanis (Creaserinus) hedgpethi. 29,30,84 (117)
Faliicambarus (Creaserinus) hortoni, 30, 84 (120)
Faliicambarus (Creaserinus) oryktes, 30, 84 (122)
FaUicambarus (Creaserinus) uhleri, 30.84 (119)
Faliicambarus (Faliicambarus) devastator, 87,91 (378)
Fallicambanis (Faliicambarus) dissitus, 30,84 (115)
Faliicambarus (Faliicambarus) harpi. 30,84 (112)
Faliicambarus (Fallicambanis) jeanae, 31,84 (113)
Faliicambarus (Fallicambarus) macneesei, 31.84 (111)
Faliicambarus (Fallicambarus) spectrum, 31
Faliicambarus (Fallicambarus) strawni, 31, 84 (114)
fasaatus. Cambarus (Hiaucambarus), 19,83 (59)
Faxonella.3.4.31
Faxonella, subgenus, 3,31
Faxonella beyeri,31,84 (127)
Faxonella blairi,31,84 (125)
Faxonella dypeata, 31,32,84 (124)
Faxonella clypeatus, 32
Faxonella dypeta, 32
Faxonella clyptea, 32
Faxonella creaseri, 32,84 (126)
Faxonella cylpeata, 32
Faxonicus, 33
faxonii, Cambarus, 50
Faxonius, 3.4,31,33,34,35,38,39,50
Faxonius, subgenus, 3,35,38,39,42,44,45,50
Faxonius affinis, 39
Faxonius alabamensis,39
Faxonius bisectus, 35
Faxonius clypeatus, 31
Faxonius compressus, 40
Faxonius creolanus, 34
Faxonius difficilis, 43
Faxonius erichsonianus, 35
Faxonius eupunctus, 35
Faxonius forceps, 45
Faxonius harrisonii, 34
Faxonius hathawayi, 43
Faxonius hobbsi, 34
Faxonius hylas, 46
Faxonius illinoiensis, 36
Faxonius immunis, 41
Faxonius tndianensis, 38
Faxonius juvenilis, 49
Faxonius kentuckiensis, 50
Faxonius lancifer,50
Faxonius leptogonopodus, 46
Faxonius limosus, 39
Faxonius longidigitus, 46
Faxonius longimanus, 35
Faxonius luteus, 46
Faxonius marchandi, 36
Faxonius medius, 47
Faxonius meeki, 34
Faxonius meeki brevis, 34
Faxonius meeki meeki, 34
Faxonius menae, 47
Faxonius minis, 47
Faxonius mississippiensis, 41
Faxonius nais, 41
Faxonius nana macros, 46
Faxonius nana nana, 47
Faxonius neglectus, 48
Faxonius neglectus chaenodactylus, 47
Faxonius neglectus neglectus, 48
Faxonius obscurus, 36
Faxonius ozarkae, 48
Faxonius palmeri, 35
Faxonius palmeri creolanus, 34
Faxonius palmeri longimanus, 35
Faxonius palmeri palmeri, 35
Faxonius peruncus,48
Faxonius placidus, 48
Faxonius propinquus, 37
Faxonius propinquus jeffersoni, 36
Faxonius propinquus propinquus, 37
Faxonius punctimanus, 48
Faxonius quadruncus, 49
NUMBER 480 219
Faxonius rafinesquei, 37
Faxonius rhoadesi,41
Faxonius rusticus, 49
Faxonius rusticus barrenensis, 45
Faxonius rusticus rusticus, 49
Faxonius sanborni, 37
Faxonius sanborni sanborni, 37
Faxonius shoupi, 38
Faxonius sloanii.SO
Faxonius tricuspis,38
Faxonius validus, 42
Faxonius virginiensis, 38
Faxonius virilis, 42
Faxonius wrighti, 39
Faxonius (Faxonella) clypeatus, 31
Faxonius (Faxonius) alabamensis, 39
Faxonius (Faxonius) compressus, 40
Faxonius (Faxonius) creolanus, 34
Faxonius (Faxonius) difficilis, 43
Faxonius (Faxonius) erichsonianus, 35
Faxonius (Faxonius) forceps, 45
Faxonius (Faxonius) harrisonii, 34
Faxonius (Faxonius) hylas, 46
Faxonius (Faxonius) immunis immunis, 41
Faxonius (Faxonius) immunis pedianus,41
Faxonius (Faxonius) indianensis,38
Faxonius (Faxonius) limosus, 39
Faxonius (Faxonius) longidigitus, 46
Faxonius (Faxonius) longimanus, 35
Faxonius (Faxonius) luteus, 46
Faxonius (Faxonius) medius, 47
Faxonius (Faxonius) meeki, 34
Faxonius (Faxonius) menae, 47
Faxonius (Faxonius) minis, 47
Faxonius (Faxonius) mississippiensis, 41
Faxonius (Faxonius) nais,41
Faxonius (Faxonius) neglectus, 48
Faxonius (Faxonius) obscums, 36
Faxonius (Faxonius) palmeri,35
Faxonius (Faxonius) peruncus, 48
Faxonius (Faxonius) placidus, 48
Faxonius (Faxonius) propinquus propinquus, 37
Faxonius (Faxonius) punctimanus, 48
Faxonius (Faxonius) quadruncus, 49
Faxonius (Faxonius) rusticus rusticus, 49
Faxonius (Faxonius) sanbomi, 37
Faxonius (Faxonius) sloanii,50
Faxonius (Faxonius) validus, 42
Faxonius (Faxonius) virilis, 42
ferrugineus, Procambarus (Girardiclla), 87,91
Fexonius, 33
Fitzcambarus, subgenus, 4,28
fitzpatricki, Procambarus, 51
fitzpalricki, Procambarus (Acucauda), 51,85 (211)
floridanus, Cambarus, 17
floridanus, Cambarus (Cambarus), 17
floridanus, Cambarus (Deprcssicambarus), 17
fodiens, Astacus, 29
fodiens, Cambarus, 29
fodiens, Cambarus (Bartonius),29
fodiens, Cambarus (Cambarus), 29
fodiens, Fallicambarus, 29
fodiens, Fallicambarus (Creaserineus),29,84 (116)
fodiens, Procambarus Fallicambarus, 29
forceps, Cambarus, 33,45
forceps, Cambarus (Faxonius), 45
forceps, Faxonius, 45
forceps, Faxonius (Faxonius), 45
forceps, Orconectes, 46
forceps, Orconectes (Procericambarus),45,46, 85 (176)
fortis, Pacifastacus, 6
fortis, Pacifastacus (I Iobbsasiacus), 6, 82 (3)
fossarum, Astacus, 77
fossarum, Cambarus, 77
fossor, Astacus, 5,24
fossor, Cambarus, 24
franzi, Procambarus (Ortmannicus), 66, 86 (287)
Fraxonius, 33
friauf i, Cambarus, 22
friaufi, Cambarus (Jugicambarus), 22,83 (75)
gallinas, Cambarus, 58
gallinus, Cambarus, 58
Gambarus, 12,33,44
Gambarus typhlobius, 45
Gambarus (Faxonius) meeki, 12,33,44
Gambeli, Astacus, 7
gambeli, Astacus, 7
Gambeli, Cambarus, 7
gambeli, Pacifastacus, 7
gambeli, Potamobius (Potamobius), 7
gambeli connectens, Pacifastacus, 6
Gambelii, Astacus, 6, 7
gambelli, Astacus, 7
Gambelii, Astacus (Astacus), 7
gambelii, Astacus (Astacus), 7
Gambelii, Cambrus, 6
gambelii, gambelii, Pacifastacus, 7
gambelii, Pacifastacus, 6,7
gambelii, Pacifastacus (Hobbsastacus), 6,7,82 (2)
gambelii connectans, Pacifastacus, 6
gambelii connectens, Astacus, 6
gambelii connectens, Pacifastacus, 6
gambelli gambelii, Astacus, 7
gambelii, Astacus, 7
gambelli, Cambarus, 7
gambelli gambelli, Astacus, 7
gambelli gambelli, Pacifasucus, 7
Gambelli i, Astacus, 7
Gam belly, 6
Gam belly nigrescens, 7
geminus, Procambarus, 66
geminus, Procambarus (Ortmannicus), 66,86 (305)
gentryi, Cambarus, 22
gentryi, Cambarus (Jugicambarus), 22, 83 (81)
geodytes, Procambarus, 58
geodytes, Procambarus (Hagenides), 58, 86 (250)
georgiae, Cambarus (Puncticambirus), 26, 83 (93)
gibbus, Procambarus, 73
gibbus, Procambarus (Permides),73. 87 (342)
Girardianus, Cambarus, 19
girardianus, Cambarus, 19
girardianus, Cambarus (Bartonius), 19
girardianus, Cambarus (Hiaticambarus), 19, 83 (60)
girardianus, Cambarus extraneus, 19
Girardiella, subgenus, 3,4,51,55
gonopodocristatus, Procambams, 66
gonopodocrisutus, Procambarus (Ortmannicus),66, 86 (324)
gordoni, Fallicambarus (Creaserinus),87,91 (377)
gracilis, Cambarus, 56
gracilis, Cambarus (Cambarus), 56
220 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
gncflis,Cambuas (Girmrdidl*),56
gracflu.Carnbanu (Ortmannicus), 56
grac0is,Procambuus,S6
gracto, Procambaras (Gir?rdiella),56,86 (241)
gracillis, Cambaras, 56
gnysoni, Cambuns, IS
graysoni, Cambuut (Dqpresticambaros), IS, 83 (49)
Gremicambaras, subgenus, 3,4,33,39
gulielmi, Cambaras, 35
Hagenianus, Cambuut, 55,56
hagenianns, Cambuus, 56
hageniamu, Cambaras (Cambaras), 56
tugenianus, Cambaras (Ortmaimicu$),56
hagenianus, Procambarat, 55,56,57
hagenianui, Procambaras (Giiaidiella), 55,56,57
hagenianus cam, Cambaras (GirardieUa), 57
hagenianus evansi, Cambanu (GinudieUa), 57
hagenianus forestae, Cambaras (Girardiella),55
hagenianus fonestae, Cambaras (Ginrdiella),55
hagenianus fonestae, Procambaius, 55
hagenianus hagenianus, Cambaras (Girardiella),56
hagenianus hagenianus, Procambaras, 56
hagenianus hagenianus, Procambaras (Giiaidiella), 56,86 (239)
hagenianus vesticeps, Procambaras (GinudieUa), 57,86 (240)
Hagenides, subgenus, 4,51,58
halll, Cambanu, 15
hafli. Cambaras (Deprcsskambaras), 15,83 (35)
hamueatus, Cambaras, 12
hamoetos, Cambaros, 12
hamulatus, Cambaras, 12
haroulatus, Cambaras (Avittcambaras), 12,82 (27)
hamulatus, Cambaras (Bartonius), 12
hamulatus, Cambaras (Cambaras), 12
hamulatus, Oiconectes, 12
hanmlatus, Oreonectes, 12
haroedi, Cambaras (Paracambaras),51
harpi, Famcambaras (FaDicambaras, 30,84 (112)
harrisoni, Cambaras, 33
hanisoni, Cambaras (Faxonius), 34
harrisoni, Oreoncctes. 34
Harrisonii, Cambaras, 33
hanisonii, Cambaros, 33
harrisonii, Faxonius, 34
harrisonii, Faxonius (Faxonius), 34
harrisonii, Oreonectes, 34
hanisonii, Oreonectes (Bilkcambara?),34.84 (149)
harrisonii, Oreonectes (Oreonectes), 34
harti, Cambaras (Depressicambaras), 15,83 (47)
Haasleri, Parastacus, 80
hasskri, Parastacus, 80
hathawayi, Faxonius, 43
hathawayi, Oreonectes, 43
hathawayi. Oreonectes (Hespericambaras), 43.85 (146)
hathawayi. Oreonectes (Oiconectes), 43
Hayi. Cambaras, 66
hayi, Cambaras, 66
hayi, Cambaras (Cambanu), 66
hayi. Cambaras (Orunannicns),66
hayi. Procambaras, 66
hayi, Procambaras (Ortmanniau),66,86 (306)
hedgepethi,FaUkambaras,30
bedgpeuu, Cambaros, 30
hcdgpethi,Fa0icamoanis.30
hedgpedu. Famcambaras (CfeaserimuX29.30.84 (117)
hemi. Cambaras, 55
Hespericambaras, subgenus, 3 ,4,33,42
Hiaticambarus, subgenus, 4,12,19
hiawasseensis, Cambarus, 26
hinei, Cambaras, 55
hinei, Cambaras (Cambaras), 51,55
hinei, Cambaras (Ortmannicus),55
hinei, Procambaras, 55
hinei, Procambaras (Capillicambarus),55,85 (230)
hineyi, Procambaras, 55
hirsutus, Procambararus (Ortmannicus), 67
hirsutus, Procambaras, 67
hirsutus, Procambaras (Ortmanicus),67,86 (280)
hirsutus, Procambaras (Ortmarmicus),67
Hiwasseenis, Cambaras, 26
hiwasseensis, Cambarus (Puncticambarus),26,83 (102)
Hobbsastacus, subgenus, 3 ,4 ,6
Hobbseus.3,4,32
Hobbseus attenuatus,32,84 (132)
Hobbseus crisutus, 32,84 (129)
Hobbseus orconectoides, 32,84 (128)
Hobbseus petilus, 32,84 (133)
Hobbseus prominens, 33,84 (130)
Hobbseus vaUeculus, 33,84 (131)
hobbsi, Faxonius, 34
hobbsi, Oreonectes, 34
hobbsi, Oreonectes (Buannulinctus),34,84 (197)
hobbsi, Oreonectes (Oreonectes), 34
Hoffmani, Procambarus, 78
hoffmanni, Paracambarus, 78
Hoffmanni, Procambaras, 78
hoffmanni, Procambarus (Vulalobosus),78,87 (356)
holii, Oreonectes, 40
holti, Oreonectes (Gremicambarus),40,84 (202)
horsti, Procambaras, 67
horsti, Procambaras (Ortmannicus), 67,86 (289)
hortonhobbsi, Procambarus, 78
hortonhobbsi. Procambarus (ViUalobosus),78,87 (359)
hortonhobsi, Procambaras, 78
hortoni, Fallicambaras, 30
hortoni, Fallicambaras (Creaserinus), 30,84 (120)
howardi, Cambaros, 14
howardi, Cambaras (Cambarus), 14,83 (32)
howardi, Procambarus, 14
howcUac, Procambaras, 76
howeUae, Procambaras (Scapulicambarus),76,87 (349)
hubbeUi, Cambarus, 61
hubbeUi, Procambaras, 61
hubbeUi, Procambaras (Leconticambarus),61,86 (269)
hubbsi, Cambarus, 17
hubbsi, Cambaros (Bartonius), 17
hubbsi, Cambaras (Erebkambarus), 17,83 (51)
hubricht, Cambaras, 17
Hubrichti, Cambaras, 17
hubrichli, Cambarus, 17
hubrichti, Cambaras (Erebicambaros), 17,83 (53)
hybus, Procambaras, 67
hybus, Procambaras (Ortmannicus), 67,86 (311)
hylas, Cambaras, 46
hylas, Cambaras (Faxonius), 46
hylas, Faxonius, 46
hylas, Faxonius (Faxonius), 46
hylas, Oreonectes, 46
hylas, Oreonectes (Oiconectes), 46
hylas, Oreonectes (Procericarnbarus), 46,85 (172)
L memis, Oreonectes, 44
NUMBER 480 221
latimanus.Cambarus, 17
Ieniusculus klamathcnsis, Pacifastacus,7
illinoiensis, Faxonius, 36
illinoiensis, Orconectes, 36
illinoiensis, Orconectes (Crockcrinus),36,84 (166)
illinoisensis, Orconectes, 36
immunis, Cambarus, 40,41
immunis, Cambarus (Faxonius), 41
immunis, Faxonius, 41
immunis, Orchonectes, 41
immunis, Orconcetes, 41
immunis, Orconectes, 41
immunis, Orconectes (Gremicambanis), 41, 84 (201)
immunis, Orconectes (Orconectes), 41
immunis, Orconnecte s, 41
immunis Hagenianus, Cambarus, 41
immunis immunis, Cambarus, 40
immunis immunis, Faxonius (Faxonius), 41
immunis immunis, Orconectes, 41
immunis immunis, Orconectes (Orconectes), 41
immunis pcdianus, Faxonius (Faxonius), 41
immunis pedianus, Orconectes,41
immunis spinirosuis, Cambarus, 40
immunis var. spinostris, Cambarus, 40
imunis, Orconectes, 41
incilis, Procambarus, 55
incilis, Procambarus (CapiUicambarus), 55, 85 (232)
inci His, Procambarus, 55
incomptus, Orconectes, 44
incomptus, Orconectes (Orconectes), 44,85 (136)
indianensis, Cambarus, 38
indianensis, Cambarus (Faxonius), 38
indianensis, Faxonius, 38
indianensis, Faxonius (Faxonius), 38
indianensis, Orconectes, 38
indianensis, Orconectes (Faxonius), 38,84 (142)
indianensis, Orconectes (Orconectes), 38
inermis, Cambarus, 44
inermis, Orconectes, 33,44
inermis, Orconectes (Orconectes), 44
inermis, Orconectis, 44
inermis, Oreonectes, 44
inermis inermis, Orconectes,44
inermis inermis, Orconectes (Orconectes), 44,85 (137)
inermis inermis, Orconectes, 44
inermis inermis, Oroconectes, 44
inermis inermis, Oronectes, 44
inermis tcsiii, Orconectes, 44
inermis tesui. Orconectes (Orconectes), 44,85 (138)
inemis inemis, Orconectes, 44
iowaen is, Orconectes, 37
iowaensis, Orconectes, 37
jaculus, Procambarus, 67
jaculus, Procambarus (Ortmannicus),67, 86 (310)
jeanae, Fallicambarus (Fallicambarus),31, 84 (113)
jeffersoni, Orconectes, 36
jcffersoni, Orconectes (Crockerinus),36.84 (157)
joncsi, Aviticambarus, 13
jonesi, Cambarus, 13
jonesi, Cambarus (Aviticambarus), 13,82 (28)
jonesi. Carbarns, 13
Jordani, Cambarus, 15
Jordan i, Cambarus, 15
jordani, Cambarus (BartonIUS), 16
jordani, Cambarus (Cambarus), 16
jordani, Cambarus (Depressicambarus), 16
Jugicambarus, subgenus, 4,12,20
Jugocambarus, 20
juvenal is, Cambarus, 49
juvenalis, Orconectes, 49
juvenilis, Cambarus, 49
juvenilis, Cambarus (Faxonius), 50
juvenilis, Faxonius, 49
juvenilis, Orcenectes, 49
juvenilis, Orconectes, 49
juvenilis, Orconectes (Orconectes), 49
Juvenils, Orconectes, 49
kentuck iens, Orconectes, 50
kentuckiensis, Faxonius, 50
keniuck iensis, Orconectes, 50
kentuckiensis, Orconectes (Orconectes), 50
kentuckiensis, Orconectes (Rhoadesius), 50,85 (151)
kilbyi, Cambarus, 61
kilbyi, Procambarus, 61
kilbyi, Procambarus (Leconticambarus),61,86 (268)
kinderhookensis, Orconectes, 37
klamathenis, Pacifasucus, 7
Klamathensis, Asiacus, 6,7
klamaihensis, Aslacus, 7
Klamathensis, Astacus (AsUcus),7
klamathensis, Astacus (Astacus), 7
Klamathensis, Cambarus, 7
klamathensis, Pacifasucus, 7
klamathensis, Pacifasucus Ieniusculus, 7
klamathensis, Poumobius (Poumobius),7
klaymathensis, Asucus, 7
Lacunicambarus, subgenus, 4,12,24
laevi, Cambarus, 18
laevigaius, ParasUcus, 4,79,87 (369)
laevis, Cambarus, 17
laevis, Cambarus (Krcbicambarus), 17,18, 83 (56)
Laevis, Cambarus (Erebicambarus), 18
laevis, Cambarus (erebicambarus), 17
laevis, Cambarus (Hrcbicambaruss), 18
laevis, Cambarus (Eribicambarus), 18
laeviusculus, Astacus, 7
lagniappe, Procambarus, 73
lagniappe, Procambarus (Pcrmides),73,87 (339)
lancifer, Cambarus, 33,50
lancifer, Cambarus (Faxonius), 50
lancifer, Faxonius, 50
lancifer, Orcnoectes, 51
lancifer, Orconectes, 50
lancifer, Orconectes (Orconectes), 51
lancifer, Orconectes (Tragulicambarus), 50,51,85
latimanus, Asucus, 12,14,15
latimanus.Cambarus, 15,17
latimanus, Cambarus (Bartonius), 16
latimanus.Cambarus (Cambarus), 16
latimanus, Cambarus (Depressicambarus), 15,16,83 (38)
latimanus striatus, Cambarus, 16
latimanus striatus, Cambarus (Cambarus), 16
latimanus var. striatus, Cambarus (Bartonius), 16
latimus, Cambarus, 15,17
latiplcurum, Procambarus, 61
latipleurum, Procambarus (Leconlicambarus), 61,86 (260)
layi, Procambarus, 66
LeContei, Cambarus, 67
I jecontei, Cambarus, 67
lecontei, Cambarus, 67
222 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
lecontei, Cambarus (Cambarus), 67
lecon tei, Cambarus (Ortmannicus),67
lecontei. Procambanis, 67
leconlei, Procambarus (Ortmannicus), 67,86 (304)
Lecont icambarus, subgenu s, 4,51,60
leenisculus, Astacus, 7
leiiheusen,Procambarus (Ortmannicus),67,86 (283)
lenisculus, Pacifastacus, 7
lenisculus klamathensis, Pacifastacus, 7
lemsculus lenisculus, Pacifastacus, 8
leniuscilus klamathensis, Pacifastacus, 7
lenisculus trowbridgii, Pacifastacus, 8
lcni suculus, PacifasUcus, 8
leniuaculus leniuaculus, Pacifastacus, 8
leniusculas, Pacifastacus, 8
leniusculus, Astacus, 7
leniusculus, Asucus (Astacus), 7
leniusculus, Astacus (Pacifastacus), 7
leniusculus, PacifasUcus, 7
leniusculus, Pacifasticus, 8
leniusculus, Pasifasticus, 7
leniusculus, Potamobius, 7
leniusculus, Potamobius (Potamobius), 7
leniusculus klamathensis, PacifasUcus, 7,8
leniusculus klamathensis, PacifasUcus (PacifasUcus), 7,82 (6)
leniusculus leniusculus, PacifasUcus, 7
leniusculus leniusculus, PacifasUcus (PacifasUcus), 7,8,82 (5)
leniusculus trowbridgii, PacifasUcus, 8
leniusculus trowbridgii, PacifasUcus (PacifasUcus), 8,82 (7)
leniusculus trowbridgii, Pacifasticus, 8
leniusculus trowbridgii, Pacifiasucus, 8
kniuscus, Pacifasucuhis,7
leniuskulus, PacifasUcus, 8
leniuslus trowbridgii, PacifasUcus, 8
lcnu isculu J . PacifasUcus, 7,8
lcnusculus knuscuhis, PacifasUcus,7
leonensis, Piocambaras, 68
feonensis, Procambanis (Ortmannicus), 68,86 (296)
tepidodactyhis, Procambarus, 68
lepidodactyhis, Procambarus (Ortmannicus), 68,86 (278)
leptogonapodus, Orconectes, 46
leptogonopodus, Faxonius, 46
leptogonopodus, Orconectes, 46
leptogonopodus, Orconectes (Orconectes), 46
leptogonopodus. Orconectes (Procericambanis),46,85 (169)
leptogonopodus acares, Orconectes, 45
leptogonopodus leptogonopodus, Orconectes, 46
leslei, Cambafdhis, 11
lesliei, Camoarellus. 11
lesliei, CambareOus (Pandicambanis). 11,82 (20)
lewisi, Procambanu, 68
lewui, Procambanu (Ortmannicus), 68,86 (318)
liberoran, Procambanis, 57
uberorum, Procambanis (Girardiella). 57,86 (246)
1 imnosu?. Orconectes, 39
limou, Orconectes, 39
umocus. Asucus, 33,38
linracus, Cambaras, 39
limosus, Cambaras (Faxonius), 39
tinxwus, Faxonius, 39
limosus, Faxonius (Faxonius), 39
I jimnu i. Orconectes, 39
lanows. Orconectes, 39
limoras, Orconectes (Faxonius), 38,84 (140)
limosus. Orconectes (Orconectes), 39
limosus, Orconects, 39
limosus, Orconelctes, 39
limosus, Orconeotes, 39
limosus, Orconetes, 39
limosus, Oroncestes, 39
limusus, Orconectes, 39
liniusculus, PacifasUcus, 8
litosternum, Procambanis, 68
litostemum, Procambanis (Ortmannicus), 68, 86 (277)
Ilamasi, Procambarus, 52
llamasi, Procambarus (Austrocambaras),52,85 (224)
llamesi, Procambanis, 52
lobdelli, Cambarus, 29
longerosilis, Cambarus, 19
longerostris, Cambarus longulus, 19
longidigitus, Cambarus, 46
longidigitus, Cambarus (Faxonius), 46
longidigitus, Faxonius, 46
longidigitus, Faxonius (Faxonius), 46
longidigitus, Orconectes, 46
longidigitus, Orconectes (Orconectes), 46
longidigitus, Orconectes (Procericambarus),46,85 (181)
longimanus, Cambarus, 35
longimanus, Faxonius, 35
longimanus, Faxonius (Faxonius), 35
longimanus, Orconectes, 35
longimonus, Cambarus, 35
Ion girostris, Cambarus, 19
longirostris, Cambarus (Hiaticambarus), 19,83 (61)
longulus, Cambaroides, 20
longulus, Cambarus, 12,19,20
longulus, Cambarus (Bartonius), 20
longulus, Cambarus (Hiaticambarus), 20,83 (62)
longulus chasmodactylus, Cambarus, 19
longulus longirosilis, Cambarus, 19
longulus longirostris, Cambarus, 19
longulus longirostris, Cambarus (Cambarus), 19
longulus longulous, Cambaras, 20
longulus longulus, Cambaras, 20
longulus longulus, Cambaras (Cambaras), 20
Lonnbergius, subgenus, 4,51,62
lophotus, Procambarus, 68
lophotus, Procambarus (Ortmannicus), 68,86 (307)
lucifugus, Cambarus, S
lucifugus, Crangonyx, 5
lucifugus alachua, Cambaras, 68
lucifugus alachua, Procambarus, 68
lucifugus alachua, Procambarus (Ortmannicus), 68,86 (284)
lucifugus lucifugus, Cambaras, 68
lucifugus lucifugus, Procambarus, 68
lucifugus lucifugus, Procambanis (Ortmannicus), 68,86 (285)
ludoviciana, Cambarus diogenes, 24
ludovicianus, Cambarus, 24
ludovicianus, Cambarus (Lacunicambams),24,83 (89)
ludovicianus ludovicianus, Cambarus, 25
luetus, Orconectes, 46
lunzi, Cambaras. 69
lunzi, Procambarus, 69
lunzi. Procambarus (Ortmannicus), 69, 86 (294)
luteus, Faxonius, 46
luteus, Faxonius (Faxonius), 46
luteus, Orconectes, 46
luteus, Orconectes (Orconectes), 46
luteus, Orconectes (Procericambarus),46,85 (188)
lylei, Procambarus, 73
NUMBER 480 223
lylei, Procambarus (Pennidcs), 73,87 (333)
maclanei, Troglocambarus, 79, 87 (364)
maclanei, Trolocambarus, 79
macneesei, Cambarus, 31
macnecsei, Fallicambaru s, 31
macneesei, Fallicambams (Fallicambarus), 31, 84 (111)
macrus Orconectes, 46
macrus, Orconectes (Procericambarus), 46, 85 (184)
maculatus, Cambarus (Erebicambarus), 87,91
mais, Orconectes, 41
maletae, Orconectes (Hespericambarus),43, 85 (145)
mancus, Procambarus, 69
mancus, Procambarus (Ortmannicus),69,86 (313)
rnaniculatus, Astacus, 77
maniculatus, Cambarus, 77
manningi, Cambarus (Hiaticambarus), 20, 83 (63)
marchandi, Faxonius, 36
marchandi, Orconectes, 36
marchandi, Orconectes (Crockerinus), 36, 84 (153)
marchandi, Orconectes (Orconectes), 36
marthae, Procambarus (Ortmannicus), 69, 86 (317)
medialis, Procambarus (Ortmannicus), 69,86 (315)
medius, Cambarus, 47
medius, Cambarus (Faxonius), 47
medius, Faxonius, 47
medius, Faxonius (Faxonius), 47
medius, Orconectes, 47
medius, Orconectes (Orconectes), 47
medius, Orconectes (Procericambarus), 47,85 (187)
meeki, Cambarus, 34
meeki, Cambarus (Faxonius), 34
meeki, Faxonius, 34
meeki, Faxonius (Faxonius), 34
meeki, Gambarus (Faxonius), 34
meeki, Orconectes, 34
meeki, Orconectes (Orconectes), 34
meeki brevis, Faxonius, 34
meeki brevis, Orconectes, 34
meeki brevis, Orconectes (Buannul if ictus), 34,84 (193)
meeki brevis, Orconectes (Orconecles),34
meeki meeki, Faxonius, 34
meeki meeki, Orconectes, 34
meeki meeki,Orconectes (Buannulif ictus), 34,84 (192)
meeki meeki, Orconectes (Orconectes), 34
meekii, Cambarus, 34
menae, Faxonius, 47
menae, Faxonius (Faxonius), 47
menae, Orconectes, 47
menae, Orconectes (Orconectes), 47
menae,Orconectes (Procericambarus), 47,85 (189)
mexiacnus, Procambarus, 53
Mexicambarus, subgenus, 4,51,63
Mexicanus, Astacus, 52
mexicanus, Astacus, 52
mexicanus, Astacus, (Cambarus) 52
Mexicanus, Cambarus, 52
mexicanus, Cambarus, 52
mexicanus, Cambarus (Cambarus), 52
mexicanus,Cambarus (Procambarus), 52
mexicanus, Procambarus, 53
Mexicanus, Procambarus, 53
mexicanus, Procambarus (Austrocambarus),52,53,85 (214)
milled, Procambarus, 61
milled, Procambarus (Leconticambarus),61, 86 (258)
miltus, Cambarus, 25
miltus, Cambarus (Lacunicambarus), 25,83 (90)
mirandae, Procambarus, 53
mirandai, Procamharus, 53
mirandai, Procambarus (Austrocambarus),53,85 (219)
minis, Faxonius, 47
minis, Faxonius (Faxonius), 47
minis, Orconectes, 47
minis, Orconectes (Procericambarus), 47,85 (182)
Mississippiensis, Cambarus, 41
mississippiensis, Cambarus, 41
mississippiensis, Cambarus (Faxonius), 41
mississippiensis, Faxonius, 41
mississippiensis, Faxonius (Faxonius),41
mississippiensis, Orconectes, 41
mississippiensis, Orconectes (Grcmicambarus),41, 85 (198)
mississippiensis, Orconectes (Orconectes), 41
mongalensis, Cambarus, 22
monongalensis, Cambarus, 22
monongalensis, Cambarus (Bartonius), 23
monongalensis, Cambarus (Cambarus), 22
monongalensis, Cambarus (Jugicambarus),22,83 (79)
montanus, Cambarus, 13
montanus acuminatus, Cambarus, 25
montanus acuminatus, Cambarus (Cambarus), 25
montanus montanus, Cambarus, 13
montanus montanus, Cambarus (Cambarus), 13
montanus veteranus, Cambarus (Cambarus), 27
montazumae, Cambarcllus, 9
Montezumae, Cambarellus, 9
montezumae, Cambarellus, 9
montezumae, Cambarellus (Cambarellus), 9,82 (14)
Montezumae, Cambarus, 8,9
montezumae, Cambarus, 9
Montezumae, Cambarus (Cambarellus),9
montezumae, Cambarus (Cambarellus), 9
montezumae areolatus, Caambarus, 9
montezumae areolatus, Cambarellus, 8
montezumae areolatus, Cambarus, 8
montezumae areolatus, Cambarus (Cambarellus), 8
montezumae chapalanus, Cambarellus, 9
montezumae chapalanus, Cambarus, 9
montezumae dugesi, Cambarellus,9
montezumae dugesi, Cambarus, 9
montezumae dugesi, Cambarus (Cambarcllus), 9
montezumae dugesii, Cambarellus, 9
montezumae dugesii, Cambarus, 9
montezumae forma lermensis, Cambarellus, 9
montezumae forma zempoalensis, Cambarellus, 10
montezumae lermensis, Cambarellus, 9
montezumae montezumae, Cambarellus, 9
Montezumae Montezumae, Cambarus, 9
montezumae montezumae, Cambams, 9
montezumae occidenulis, Cambarellus, 9
montezumae occidenulis, Cambarus, 9
montezumae occidenulis, Cambarus (Cambarellus), 9
montezumae patzcuarensis, Cambarellus, 9
Montezumae var. areolau, Cambarus, 8
montezumae var. areolau,Cambarcllus. 9
montezumae var. tridens, Cambarellus, 9
Montezumae var. tridens, Cambarus, 9
montezumae var. tridens, Cambarus, 9
montezumae zempoalensis, Cambarellus, 10
nali, Cambarus, 41
Nais, Cambarus, 41
nais, Cambarus, 41
224 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
nais, Cambanis (Faxonius), 41
nais, Faxonius, 41
nais, Faxonius (Faxonius), 41
nais, Orconectes, 41
nais, Orconectes (Gremicambams), 39,40,41,85 (203)
nais, Orconectes (Orconectes), 41
nais, Orconectus, 41
nais form causeyi, Orconectes, 39
nana, Orconectes, 47
nana, Orconectes (Procericambams),47,85 (183)
nana macros, Faxonius, 46
nana macros, Orconectes, 46
nana macros, Orconectes (Orconectes), 46
nana nana, Faxonius, 47
nana nana, Orconectes, 47
nana nana, Orconectes (Orconectes), 47
natchitochae, Procambarus, 72,74
natchitochae, Procambarus (Pennides), 74,87 (328)
Nebra scens i s, Cambanis, 24
nebrascensis, Cambanis, 24
neglecu, Cambarus, 48
neglectus, Cambarus, 47
neglecuis, Cambarus (Faxonius),47
neglectus, Faxonius, 48
neglectus, Faxonius (Faxonius), 48
neglectus, Orconectes, 48
neglectus, Orconectes (Orconectes), 48
neglectus chaenodactylus, Faxonius, 47
neglectus chaenodactylus, Onconectes, 47
neglectus chaenodactylus, Orconectes, 47
neglectus chaenodactylus, Orconectes (Orconectes), 47
neglectus chaenodactylus, Orconectes (Procericambarus),47,85 (179)
neglectus chaenodaytylus, Orconectes, 47
neglectus neglectus, Faxonius, 48
neglectus neglectus, Orconectes, 48
neglectus neglectus, Orconectes (Procericambarus), 48,85 (180)
Nephropoidea, 3,4
nerterius, Cambarus. 26
nerterius, Cambarus (Puncticambarus),26,83 (97)
Nicoleti, Astacus, 80
nicoleii, Asucus. 80
Nicoleti, Parastacus, 80
nicoleti. Parastacus, 80,81,87 (372)
nicoietii, Parastacus, 80
mcolcit i, Parastacus, 80
nigrescens, Astacus, 7
nigrescens, Astacus (Astacus), 7
nigrescens, Gambelly, 7
nigrescens, Pacifastacus, 6,7
nigrescens, Pacifastacus (Hobbsasucus), 7,82 (4)
nigrescens, Polamobius (Potamobius), 7
nigrescens fortis, Astacus, 6
nigrescens fortis, Pacifastacus, 6
nigrescens nigrescens, Astacus, 7
nigrescens nigrescens, Pacifastacus, 7
nigresens, Pacifastacus, 7
ninae, CambareOus 11
ninae, Cambarellus (Pandicambarus). 11,82 (23)
niveus, Procambarus, 53
niveus, Procambarus (Austrocambarus), 53,85 (229)
nodosus, Cambaras (Jugicambaros),23,83 (76)
ouacae oaxacae, ProcBmbarus (Austrocambarus), 53,85 (216)
oaxacae reddeOi, Procambarus (Austrocambarus), 53,85 (217)
obesus. Cambarus, 24
obesus var. Utimanus, Cambarus, 15
obeyensis, Cambarus, 23
obeyensis, Cambarus (Jugicambarus),23, 83 (67)
obeyensis, Cambarus (Veticambarus),23
obscura, Cambarus, 36
obscurus, Astacus, 36
obscurus, Cambarus, 36
obscurus, Cambarus (Faxonius), 36
obscurus, Faxonius, 36
obscurus, Faxonius (Faxonius), 36
obscurus, Orconcectes, 36
obscurus, Orconectes, 36
obscurus, Orconectes (Crockerinus), 36,84 (155)
obscurus sanbomi, Cambarus, 37
obstipus, Cambarus, 16
obstipus, Cambanis (Depressicambarus), 16,83 (37)
occidentalis, Cambarellus, 9
occidentalis.Cambarellus (Cambarellus), 9,82 (8)
okaloosae, Procambarus, 76
okaloosae, Procambarus (Scapulicambarus),76,87 (350)
olmecorum, Procambarus (Austrocambarus), 87,91 ( 379)
Onconectes, 33
Onconectes neglectus chaenodactylus, 47
Orcenectes,33
Orcenectes juvenilis, 49
Orchonectes, 33,39
Orchonectes immunis,41
orcinus, Procambarus, 69
orcinus, Procambarus (Ortmannicus), 69,86 (290)
Orcnoectes,33
Orcnoectes lancifer, 51
Orconcectes, 33,35
Orconcectes obscurus, 36
Orconcetes.33,39
Orconcetes immunis, 41
Orconecctes, 33,35
Orconecctes eupunctus, 36
Orconecus,33
Orconectes, 1,3,4,12,31,33,34,35,38,39.42,43,45,50
Orconectes, subgenus, 3,4,34,35,38,39,42,43,50
Orconectes acares, 45
Orconectes acuseyi, 39
Orconectes affinis, 39
Orconectes alabamensis, 39
Orconectes australis,44
Orconectes australis australis,44
Orconectes australis packardi, 44
Orconectes barrenensis, 45
Orconectes bisectus, 35
Orconectes causey i, 39
Orconectes chickasawae, 40
Orconectes clypeata, 31
Orconectes clypeatus, 31
Orconectes compressus, 40
Orconectes cooperi, 40
Orconectes creolanus, 34
Orconectes deanae, 43
Orconectes denae, 43
Orconectes difficilis, 43
Orconectes difficilis blacki, 42
Orconectes difficilis difficilis, 43
Orconectes difficilis hathawayi, 42,43
Orconectes difficilis maletae, 43
Orconectes erichsonianus, 35
Orconectes etnieri, 40
Orconectes eupunctus, 36
NUMBER 480 225
Orconectes forceps, 46
Orconectes hamulatus, 12
Orconectes haninlatus, 12
Orconectes harrisoni, 34
Orconectes harrisonii, 34
Orconectes hathawayi, 43
Orconectes hobbsi, 34
Orconectes holti, 40
Orconectes hylas 46
Orconectes illinoiensis, 36
Orconectes illinoisensis, 36
Orconectes immunis, 41
Orconectes immunis immunis, 41
Orconectes immunis pedianus, 41
Orconectes imunis, 41
Orconectes incomptus, 44
Orconectes indianensis, 38
Orconectes inermis, 33,44
Orconectes inermis inermis, 44
Orconectes inermis testii, 44
Orconectes inemis inemis 44
Orconectes iowaenis, 37
Orconectes iowaensis, 37
Orconectes jeffersoni, 36
Orconectes juvenalis, 49
Orconectes juvenilis, 49
Orconectes Juvenils, 49
Orconectes kentuckiens, SO
Orconectes kentuckiensis, SO
Orconectes kinderhookensis, 37
Orconectes lane ifer, SO
Orconectes leptogonapodus, 46
Orconectes leptogonopodus, 46
Orconectes leptogonopodus acares,45
Orconectes leptogonopodus leptogonopodus, 46
Orconectes limnosus, 39
Orconectes limosa, 39
Orconectes Limosus, 39
Orconectes limosus, 39
Orconectes limusus, 39
Orconectes longidigitus, 46
Orconectes longimanus, 35
Orconectes luetus, 46
Orconectes luteus, 46
Orconectes macrus, 46
Orconectes mais, 41
Orconectes marchandi, 36
Orconectes medius, 47
Orconectes meeki, 34
Orconectes meeki brevis, 34
Orconectes meeki meeki, 34
Orconectes menae, 47
Orconectes minis, 47
Orconectes mississippiensis,41
Orconectes nais, 41
Orconectes nais form causeyi, 39
Orconectes nana, 47
Orconectes nana macrus, 46
Orconectes nana nana, 47
Orconectes neglectus, 48
Orconectes neglectus chaenodactylus, 47
Orconectes neglectus chaenodaytylus, 47
Orconectes neglectus neglectus, 48
Orconectes obscurus, 36
Orconectes ozarkae,48
Orconectes pallucidus, 45
Orconeaes palmeri, 35
Orconeaes palmeri creolanus, 34
Orconectes palmeri croelanus, 34
Orconectes palmeri longimanus, 35
Orconectes palmeri palmeri, 35
Orconectes pellacious, 44
Orconectes Pel lucid us, 44
Orconectes pellucidus, 33,45
Orconectes pellucidus australis, 44
Orconectes pellucidus inermis, 44
Orconectes pellucidus packardi, 44
Orconectes pellucidus packardii, 44
Orconectes pellucidus pellucidus, 45
Orconectes pellucidus pellucidus pellucidus, 45
Orconectes pellucidus pelluicdus, 45
Orconectes pellucidus testii, 44
Orconectes perfectus, 43
Orconectes peruncus, 48
Orconectes placidus, 48
Orconectes propinguis, 37
Orconectes propinques, 37
Orconectes propinquus, 37
Orconectes propinquus erismophorous, 37
Orconectes propinquus jeffersoni, 36
Orconectes propinquus propinquus, 37
Orconectes propinquus sanbomi, 37
Orconectes propoinquus, 37
Orconectes punctimanus, 48
Orconeaes putnami, 49
Orconectes quadruncus, 49
Orconeaes rafinesquei,37
Orconeaes rhoadesi, 41
Orconeaes rusticus, 49
Orconeaes rusticus barrenensis, 45
Orconeaes rusticus barrensis, 45
Orconeaes rusticus forceps, 45
Orconeaes rusticus minis, 47
Orconeaes rusticus placidus, 48
Orconeaes rusticus rusticus, 49
Orconeaes sanbomi, 37
Orconeaes sanbomi erismophorous, 37
Orconeaes sanbomi erismophorus, 37
Orconeaes sanbomi sanbomi, 37
Orconectes sanbomii erismophorous, 37
Orconectes sanbomii sanbomii, 37
Orconeaes saxatilis, 49
Orconeaes senbomi, 37
Orconeaes shoupi, 38
Orconeaes sloanei, SO
Orconectes sloani, SO
Orconeaes sloanii, SO
Orconectes sloanni, SO
Orconeaes spinosus, SO
Orconeaes spinous, SO
Orconeaes stannardi, 38
Orconeaes transfuga, 48
Orconeaes transfugus, 48
Orconeaes tricuspis, 38
Orconectes validus, 42
Orconectes virginianus, 38
Orconectes virginiensis, 38
Orconectes virilis, 42
Orconectes williamsi, SO
Orconectes wrighti, 39
226 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Orconecles (Billecambarus) harrisonii, 34,84 (149)
Orconectes (Buannulificius) dense, 43
Orconecles (Buannulifictus) hobbsi, 34,84 (197)
OTConecles (Buannulifictus) meeki brevis, 34,84 (193)
Orconectes (Buannulifictus) meeki meeki, 34,84 (192)
Orconectes (Buannulifictus) palmeri creolanus, 34,84 (196)
Orconectes (Buannulifictus) palmeri longimanus, 35,84, (195)
Orconecles (Buannulifictus) palmeri palmeri, 45,84 (194)
Orconectes (Cambarus) virilis, 42
Orconectes (Crockerinus) bisectus,35, 84 (164)
Orconectes (Crockerinus) erichsonianus, 35,84 (162)
Orconectes (Crockerinus) eupunctus, 36,84 (154)
Orconectes (Crockerinus) illinoiensis, 36,84 (166)
Orconectes (Crockerinus) jeffersoni, 36,84 (157)
Orconectes (Crockerinus) march and i, 36,84 (153)
Orconectes (Crockerinus) obscurus, 36,84 (155)
Orconectes (Crockerinus) propinquus, 36,84 (156)
Orconectes (Crockerinus) rafinesquei,37,84 (164)
Orconectes (Crockerinus) sanbornii erismophorous, 37,84 (160)
Orconectes (Crockerinus) sanbornii sanbornii, 37,84 (159)
Orconectes (Crockerinus) shoupi, 38,84 (167)
Orconectes (Crockerinus) stannardi, 38,84 (161)
Orconectes (Crockerinus) tricuspis, 38,84 (165)
Orconectes (Crockerinus) virginiensis, 38,84 (158)
Orconectes (Faxonella) beyeri, 31
Orconectes (Faxonella) clypeata, 31
Orconectes (Faxonella) clypeaius, 32
Orconectes (Faxonius) indianensis, 38,84 (142)
Orconectes (Faxonius) limosus, 38,84 (140)
Orconectes (Faxonius) wrighti, 39,84 (141)
Orconectes (Gremicambarus) alabamensis,39,84 (206)
Orconectes (Grcmicambarus) causeyi, 39,40, (200)
Orconectes (Gremicambaius) chickasawae, 40,84 (209)
Orconectes (Gremicambarus) compressus, 40,84 (210)
Orconectes (Gremicambarus) cooperi, 40,84 (204)
Orconectes (Gremicambarus) etnieri, 40,84 (f05)
Orconectes (Gremicambarus) holti, 40.84 (202)
Orconectes (Gremicambarus) immunis, 40,84 (201)
Orconectes (Gremicambarus) mississippiensis, 41,85 (198)
Orconectes (Gremicambarus) nais, 39,40,85 (203)
Orconectes (Gremicambarus) moadesi, 41,85 (207)
Orconectes (Gremicambarus) validus, 42,85 (f i8)
Orconectes (Gremicambarus) virilis, 5,39,40,42,85 (199)
Orconectes (Hespericambarus) blacki, 42,43,85 (144)
Orconectes (Hespericambarus) deanae, 43,85 (148)
Orconectes (Hespericambarus) difficilis, 42,43.85 (143)
Orconectes (Hespericambarus) hathawayi, 43,85 (146)
Orconectes (Hespericambarus)maletae. 43,85 (145)
Orconectes (Hespericambarus) perfectus, 43,85 (147)
Orconectes (Orconectes) alabamensis, 39
Orconectes (Orconectes) australis australis, 44,85, (134)
Orconectes (Orconectes) australis packardi, 44,85 (135)
Orconectes (Orconectes) bisectus, 35
Orconectes (Orconectes) compressus, 40
Orconectes (Orconectes) creolanus, 34
Orconectes (Orconectes) difficilis, 43
Orconectes (Orconectes) erichsonianus. 35
Orconectes (Orconectes) eupunctus, 36
Orconectes (Orconectes) harrisonii, 34
Orconectes (Orconectes) hathawayi, 43
Orconectes (Orconectes) hobbsi, 34
Orconectes (Orconectes) hylas, 46
Orconectes (Orconectes) immunis, 41
Orconectes (Orconectes) immunis immunis, 41
Orconectes (Orconectes) incomptus. 44,85 (136)
Orconectes (Orconectes) indianensis, 38
Orconectes (Orconectes) inermis,44
Orconectes (Orconectes) inermis inermis,44,85 (137)
Orconectes (Orconectes) inermis testii,44, 85 (138)
Orconectes (Orconectes) juvenilis, 49
Orconectes (Orconectes) kentuckiensis, 50
Orconectes (Orconectes) lancifer,51
Orconectes (Orconectes) leptogonopodus, 46
Orconectes (Orconectes) limosus, 39
Orconectes (Orconectes) longidigitus, 46
Orconectes (Orconectes) luteus, 46
Orconectes (Orconectes) marchandi, 36
Orconectes (Orconectes) medius, 47
Orconectes (Orconectes) meeki, 34
Orconectes (Orconectes) meeki brevis, 34
Orconectes (Orconectes) meeki meeki, 34
Orconectes (Orconectes) menae, 47
Orconectes (Orconectes) mississippiensis, 41
Orconectes (Orconectes) nais, 41
Orconectes (Orconectes) nana macrus, 46
Orconectes (Orconectes) nana nana, 47
Orconectes (Orconectes) neglectus, 48
Orconectes (Orconectes) neglectus chaenodactylus, 47
Orconectes (Orconectes) obscurus, 36
Orconectes (Orconectes) ozarkae, 48
Orconectes (Orconectes) palmeri, 35
Orconectes (Orconectes) palmeri creolanus, 34
Orconectes (Orconectes) palmeri longimanus, 35
Orconectes (Orconectes) pellucidus, 44,45,85 (139)
Orconectes (Orconectes) pellucidus australis, 44
Orconectes (Orconectes) pellucidus pellucidus, 45
Orconectes (Orconectes) pellucidus testii, 44
Orconectes (Orconectes) peruncus, 48
Orconectes (Orconectes) propinquus jeffersoni, 36
Orconectes (Orconectes) propinquus propinquus, 37
Orconectes (Orconectes) propinquus sanbomi, 37
Orconectes (Orconectes) punctimanus, 48
Orconectes (Orconectes) quadruncus, 49
Orconectes (Orconectes) rafinesquei, 37
Orconectes (Orconectes) rhoadesi, 41
Orconectes (Orconectes) rusticus, 45
Orconectes (Orconectes) rusticus forceps, 45
Orconectes (Orconectes) rusticus minis, 47
Orconectes (Orconectes) rusticus placidus, 48
Orconectes (Orconectes) rusticus rusticus, 49
Orconectes (Orconectes) shoupi, 38
Orconectes (Orconectes) sloani, 50
Orconectes (Orconectes) sloanii, 50
Orconectes (Orconectes) tricuspis, 38
Orconectes (Orconectes) validus, 42
Orconectes (Orconectes) virginiensis, 38
Orconectes (Orconectes) virilis, 42
Orconectes (Orconectes) wrighti, 39
Orconectes (Procericarnbarus)acares,45,85 (170)
Orconectes (Procericambarus) barrcnensis, 45,85 (178)
Orconectes (Procericambarus) forceps, 46,85 (176)
Orconectes (Procericambarus) hylas, 46,85 (172)
Orconectes (Procericambarus) leptogonopodus, 46,85 (169)
Orconectes (Procericambarus) longidigitus, 46,85 (181)
Orconectes (Procericambarus) luteus, 46,85 (188)
Orconectes (Procericambarus) macrus, 46,85 (184)
Orconectes (Procericambarus) medius, 47,85 (187)
Orconectes (Procericambarus) menae, 47,85 (189)
Orconectes (Procericambarus) minis, 47,85 (182)
Orconectes (Procericambarus) nana, 47,85, (183)
NUMBER 480 227
Orconectes (Procericambarus) neglectus chaenodactylus, 47,85 (179)
Orconectes (Procericambarus) negleaus neglectus, 47,48,85 (180)
Orconectes (Procericambarus) ozarkae, 48,85 (190)
Orconectes (Procericambarus) peruncus, 48, 85 (173)
Orconectes (Procericambarus) placidus, 48,85 (177)
Orconectes (Procericambarus) punctimanus, 48,85 (171)
Orconectes (Procericambarus) putnami, 49,85 (186)
Orconectes (Procericambarus) quadruncus, 49,85, (168)
Orconectes (Procericambarus) rusticus, 49,85 (174)
Orconectes (Procericambarus) saxatilis, 49,85 (175)
Orconectes (Procericambarus) spinosus, 50,85 (185)
Orconectes (Procericambarus) transfuga, 48
Orconectes (Procericambarus) williamsi, 50,85 (191)
Orconectes (Rhoadesius) kentuckiensis, 50,85 (151)
Orconectes (Rhoadesius) sloanii, 50,85 (150)
Orconectes (Tragulicambarus) lancifer, 50,51,85 (152)
Orconectis,33,39,44
Orconectis inermis,44
Orconectis virilis, 42
orconectoides.Hobbseus, 32,84 (128)
Orconects,33
Orconects limosus, 39
Orconectus.33,39
Orconectus nais, 41
Orconectus rusticus, 49
Orconelctes,33,38
Orconelctes limosus, 39
Orconelctes virilis, 42
Orconeotes,33,38
Orconeotes limosus, 39
Orconetes.33,38,39
Orconetes limosus, 39
Orconetes virilis 42
Orconnectes, 33,35,39
Orconnectes immunis, 41
Orconnectes propinquus, 37
Orconnectes rusticus, 49
Orconnectes virilis, 42
Oreconectes,33
oreganus, Astacus, 5
Oreonectes.33,43,44
Orconectes incrmis, 44
Orconectes inermis inermis,44
omatus, Cambarus, 18
omatus, Cambarus (Erebicambarus), 18,83 (55)
Oroncestes.33
Oroncestes limosus, 39
Oroconectes,33
Oroconectes inermis inermis,44
Oronectes.33,39.44
Oronectes inermis inermis, 44
Oronectes virilis, 42
Ortmanicus, subgenus, 3,63
ortmanii, Procambarus (Paracambarus),72,87 (325)
Ortmanmanicus, 51 ,60,63
ortmanni. Bartonius, 14
ortmanni, Cambarus, 14
ortmanni, Cambarus (Bartonius), 14
ortmanni, Cambarus (Cambarus), 14,83 (34)
ortmanni, Paracambarus,72,87
ortmanni, Procambaras (Paracambarus), 72
Ortmann icus, 63
Ortmannicus, subgenus, 3,4,51.55,58,60,62,63,72,76
Ortmarmicus blandingi acutus, 63
ortmann ii, Paracambarus, 72
ortmannii, Procambarus (Paracambarus), 72,87 (325)
oryktes, Cambarus, 30
oryktes, Fallicambarus, 30
oryktes, Fallicambarus (Creaserinus),30,84 (122)
ouachitae, Procambarus, 74
ouachitae, Procambarus (Pennides),74,87 (343)
ouichitae, Procambarus, 74
ozarkae, Faxoniut, 48
ozarkae, Orconectes, 48
ozarkae, Orconectes (orconectes), 48
ozarkae, Orconectes (Procericambarus), 48,85 (190)
Pacifastaculus, 6,7
Pacifastaculus leniuscus,7
Pacifastaculus trowbridgii, 8
Pacif astacus, 3 ,4 ,6 ,7
Pacifastacus, subgenus, 4,7
Pacif astacus chenoderma,6
Pacifastacus conectens, 6
Pacifastacus connectens, 6
Pacifastacus fortis, 6
Pacifastacus gambeli, 7
Pacifastacus gambeli connectens, 6
Pacifastacus gambelii, 6,7
Pacifastacus gambelii connecians,6
Pacifastacus gambelii connectens, 6
Pacifastacus gambelii gambelii, 7
Pacifastacus gambelii gambelii, 7
Pacifastacus Ieniusculus klamathensis,7
Pacifastacus klamathenis, 7
Pacifastacus klamathensis, 7
Pacifasucus lenisculus, 7
Pacifastacus lenisculus klamathensis, 7,8
Pacifastacus lenisculus lenisculus, 8
Pacifasucus lenisculus trowbridgii, 8
Pacifastacus lenisuculus, 8
Pacifasucus leniuaculus leniuaculus,8
Pacifasucus lcniuscilus klamathensis,7
Pacifasucus leniusculas, 8
Pacifasucus Ieniusculus, 7
Pacifasucus Ieniusculus klamathensis, 7,8
Pacifasucus Ieniusculus Ieniusculus, 8
Pacifasucus Ieniusculus trowbridgii, 8
Pacifasucus leniuskulus, 8
Pacifasucus leniuslus trowbridgii, 8
Pacifasucus lenuisculus,7
Pacifasucus lenusculus lenutculus, 7
Pacifasucus liniusculus, 8
Pacifasucus nigrescens, 6,7
Pacifasucus nigrescens fortis, 6
Pacifasucus nigrescens nigrescens, 7
Pacifasucus nigresens, 7
Pacifasucus trawbridgii, 8
Pacifasucus trowbridgei, 8
Pacifasucus trowbridgeii, 7,8
Pacifasucus trowbridgi, 8
Pacifasucus Trowbridgii, 8
Pacifasucus trowbridgii, 8
Pacifasucus (Hobbsastacus) chenoderma, 6,82
Pacifasucus (Hobbsasucus) connectens, 6,82 (1)
Pacifasucus (Hobbsasucus) fortis, 6,82 (3)
Pacifasucus (Hobbsasucus) gambelii, 6,82 (2)
Pacifasucus (Hobbsasucus) nigrescens, 7,82 (4)
Pacifasucus (Pacifasucus) Ieniusculus klamathensis, 7,82 (6)
Pacifasucus (Pacifasucus) Ieniusculus Ieniusculus, 7,82 (5)
Pacifasucus (Pacifastacus) Ieniusculus trowbridgii, 8,82
228 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Pacifasticus.6,7,8
Pacifaslicus leniusculus, 7,8
Pacifasticus leniusculus irowbridgii, 8
Pacifiastacus,6, 7
Pacifiasiacus leniusculus trowbridgii, 8
Pacificastacus, 6
paeninsulanis, Procambarus, 77
paeninsulans, Procambarus,77
paen insulanus, Cambanis, 77
paeninsulanus, Procambaris,77
paeninsulanus, Procambarus, 77
paeninsulanus, Procambarus (Scapulicambarus),77, 87 (348)
pall idus, Cambarus, 69
pallidus, Procambarus, 69
pall idus, procambarus, 69
pall idus, Procambarus (Onmannicus), 69, 86 (288)
pall idus, Procambrus, 69
paJlucidus, Orconectes, 45
Palmeri, Cambarus, 33,34,35
palmen, Cambarus, 35
palmeri, Cambanis (Faxonius), 35
palmeri, Faxonius, 35
palmeri, Faxonius (Faxonius), 35
palmeri, Orconectes, 35
palmeri, Orconectes (Orconectes), 35
palmen creolanus, Faxonius, 34
palmeri creolanus,Orconectes (Buannulifictus),34,38 (196)
palmeri creolanus, Orconectes, 34
palmeri creolanus, Orconectes (Orconectes), 34
palmeri croelanus, Orconectes, 34
palmeri longimanus, Cambanis, 35
palmeri longimanus, Faxonius, 35
palmen longimanus, Orconectes,35
palmeri longimanus, Orconectes (Buannulifictus), 35,84 (195)
palmeri longimanus, Orconectes (Orconectes), 35
palmeri palmeri, Cambanis, 35
pafaneri pafaneri, Faxonius, 35
palmeri palmen, Orconectes, 35
palmeri palmeri, Orconectes (Buannulifictus), 35,84 (194)
Pandicambarus, 3 ,4 ,8 ,10
Paracambarus, 3,72,77
Paracambarus, subgenus, 3,4,51,72
Paracambarus hof fmanni. 78
Paracambarus ortmanni, 72
Paracambarus ortmarmii, 72
Paracambarus paradoxus, 72
Paracambarus riojae, 51,77,78
Paracambarus rodriguezi, 54
Paracambarus tezhitlanensi,78
Paracambarus tcziutlanemt J. 78
Paracambarus tlapacoyanensis, 78
Paracamburus, 51,72
paradoxus, Cambarus, 72
paradoxus, Cambarus (Paracambanis),51.72,87 (326)
paradoxus, Cambanis (Paracambarus), 72
paradoxus, Paracambarus, 72
Paradoxus, Procambarus, 72
paradoxus, Procambarus (Paracambarus), 72
parasimulans, Procambarus (Girardiella), 57,86
Parastacidae. 4,79.87
Paraitacoidea.3.4
Parastacus,3.4,79.8O
Parastacus Agassizi, 81
Panutacus agassizi, 81
Parastacus agassizii, 81
Parastacus agassizzi,81
Parastacus araucanius, 80
Parastacus bimaculatus, 81
Parastacus brasilensis, 79
Parastacus brasiliensis,79,87 (365)
Parastacus braziliensis,79
Parastacus Chilensis, 80
Parastacus chilensis, 80
Parastacus defossus, 79,87 (370)
Parastacus efinifrons, 81
Parastacus Hassleri, 80
Parastacus hassleri, 80
Parastacus laevigatus, 4,79,87 (369)
Parastacus Nicoleti, 80
Parastacus nicoleti, 80,81,87
Parastacus nicoletti, 80
Parastacus pilimanus, 80,87
Parastacus pugnax, 80, 87
Parastacus saffordi, 80,87
Parastacus spinifrons, 81
Parastacus varicosus, 80,87
parrishi, Cambanis (Puncticambarus), 27, 83 (%)
parvoculus, Cambarus, 23
parvoculus, Cambarus (Jugicambarus), 23, 83 (70)
Pasifastacus, 6
Pasifastacus trowbridgeii, 8
Pasifasticus.6
Pasifasticus leniusculus, 7
patzacuarensis, Cambarellus, 9
patzcuarensis, Cambarellus, 9
patzcuarensis,Cambarellus (Cambarellus),9, 82 (15)
patzquarensis, Cambarellus (Cambarellus), 9
Peak i, Cambanis, 39
pealei, Cambarus, 39
pearsei, Cambarus (Ortmannicus),70
pearsei, Procambarus, 70
pearsei, Procambarus (Ortmannicus),70,86 (316)
pearsei pearsei, Procambarus, 70
pearsei pearsei, Procambarus (Onmannicus), 70
pearsei plumimanus, Procambarus, 70
pearsei plumimanus, Procambarus (Ortmannicus), 70
pecki, Procambarus, 75
pecki, Procambarus (Remoticambarus), 75, 87
pellacious, Orconectes, 44
pellucidus, Astacus, 45
pellucidus, astacus, 45
pellucidus, Astacus Cambarus, 45
pellucidus, Astacus (Cambarus), 45
Pellucidus, Cambarus, 45
pellucidus, Cambarus, 45
pellucidus, Cambarus (Cambarus), 45
pellucidus, Cambanis (Faxonius), 45
pellucidus, Cambanis (Fexonius), 45
pellucidus, Cambanis (Orconectes), 45
pellucidus, Cambrus, 45
pellucidus, form inermis, Camtarus (Orconectes), 45
Pellucidus, Orconectes, 44
pellucidus, Orconectes, 33,45
pellucidus, Orconectes (Orconectes), 44,45,85 (139)
pellucidus australis, Cambarus, 44
pellucidus australis, Cambanis (Faxonius), 44
pellucidus australis, Orconectes, 44
pellucidus australis, Orconectes (Orconectes), 44
pellucidus inermis, Cambarus, 44
pellucidus inermis, Orconectes, 44
NUMBER 480 229
pellucidus packardi, Orconectes, 44
pellucidus packardii, Orconectes, 44
pellucidus pellucidus, Cambarus, 45
pellucidus pellucidus, Orconectes, 45
pellucidus pellucidus, Orconectes (Orconectes), 45
pellucidus pellucidus pellucidus, Orconectes, 45
pellucidus pelluicdus, Orconectes, 45
pellucidus testi, Cambarus, 44
pellucidus testi, Cambarus (Faxonius), 44
pellucidus testii, Cambarus, 44
pellucidus testii, Orconectes, 44
pellucidus testii, Orconectes (Orconectes), 44
pellucidus var. testii, Cambarus, 44
pellucidus variety inermis, Cambarus, 44
pelludicus australis, Orconectes (Orconectes),44
pelludicus, Cambarus, 45
pelosimanus, Procambarus, 53
pelosus, Cambarus, 41
pelucidus, Cambarus, 45
pelulcidus, Cambarus, 45
penicillatus, Astacus, 60
penicillatus, Cambarus, 60,64
penni, Procambarus, 74
penni, Procambarus (Pennides), 74, 87
Pennides, subgenus, 4,51,72
perfectus, Orconectes, 43
perfectus, Orconectes (Hespericambarus),43,85 (147)
peruncus, Cambarus (Faxonius), 48
peruncus, Faxonius, 48
peruncus, Faxonius (Faxonius), 48
peruncus, Orconectes, 48
peruncus, Orconectes (Orconectes), 48
peruncus, Orconectes (Procericambarus), 48,85 (173)
petersi, Procambarus (Pennides), 87
petilus, Hobbseus. 32,84 (133)
pictus, Cambarus, 70
picius, Procambarus, 70
pictus, Procambarus (Ortmannicus),70, 86 (273)
pilimanus, Astacus, 79,80
pilimanus, Parastacus, 80,87 (366)
pilosimanus, Cambarus, 53
pilosimanus, Cambarus (Procambarus), 53
pilosimanus, Procambarus, 53
pilosimanus, Procambarus (Austrocambarus), 53,85 (225)
pilosus, Cambarus, 41
pilosus, Cambarus (Faxonius), 41
placidus, Cambarus, 48
placidus, Faxonius, 48
placidus, Faxonius (Faxonius), 48
placidus, Orconectes, 48
placidus, Orconectes (Procericambarus), 48,85 (177)
planirosiris, Procambarus, 70
planirostris, Procambarus (Onmannicus),70,86 (312)
planirostrus, Procambarus, 70
planti rostrus, Procambanis, 70
plantiuostrusg, Procambarus, 70
pi um imanus, Procambarus, 70
plumimanus, Procambarus (Ortmannicus),70,86 (314)
pogum, Procambarus (Girardiella), 57,86 (245)
polmanus, Asucus, 80
poqum, Procambarus, 57
Poumobius, 6
Potamobius, subgenus, 6
Potamobius leniusculus,7
Potamobius (Potamobius) gambeli, 7
Potamobius (Potamobius) klamathensis, 7
Potamobius (Potamobius) leniusculus,7
Potamobius (Poumobius) nigrescens, 7
Poumobius (Poumobius) trowbridgei, 8
Pracambarus,51
primaevus, Cambarus, 5,53
primaevus, Procambarus (Austrocambarus), 5,53,85
primoevus, Cambarus, 53
pristinus, Cambarus, 12,28
pnstinus, Cambarus (Veticambarus), 28, 84 (106)
Procambambarus, 51,76
Procambambarus clarkii, 76
Procambarerus, 51
Procambararus (Ortmannicus) hirsutus, 67
Procambaris, 51,63,76
Procambaris enoplosternum,65
Procambaris fallax, 66
Procambaris paeninsulanus,77
Procambaris seminolae, 71
Procambarius, 51,76
Procambarius clarki, 76
Procambarus. 3, 4, 5, 12, 13,28, 29, 31, 33, 51, 55. 58, 60, 62,63,72,75,76.
77
procambarus, 51
Procambarus, subgenus, 3,4,51,72,77
Procambarus a. blandingii, 64
Procambarus ablusus, 72
Procambanis acanthophorus, 52
Procambarus acantophorus, 52
Procambarus acherontis,62
Procambarus actus, 63
Procambarus acuminatus, 25
Procambarus acutissimus,63
Procambarus acutissumus, 63
Procambarus acutus, 63
Procambarus acutus acustus, 64
Procambarus acutus acutus, 45,63
Procambarus acutus cuevachicae, 64
Procambarus advena, 58,59
Procambarus allcni, 60
Procambarus allenii, 60
Procambarus allenti, 60
Procambarus ancylus, 64
Procambarus angustatus, 64
Procambarus apalachicolae,60
Procambarus Atkinsoni, 52
Procambarus atkinsoni, 52
Procambarus aztecus, 53
Procambarus barbatus,60
Procambarus barbiger, 55
Procambarus bartoni, 13
Procambarus bivitutus, 64
Procambarus blandingii, 64
Procambarus blandingii acutus, 63
Procambarus blandingii blandingii, 63,64
Procambarus blandingii cuevachicae, 64
Procambarus bouvieri,63
procambarus bouvieri, 63
Procambarus byersi, 29
Procambarus C. cubensis, 52
Procambarus caballeroi, 65
Procambarus capillatus, 60
Procambarus chacci,65
Procambarus clarcii, 76
Procambarus clarki. 76
230 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Procambarus clarkii, 76
Procambarus clarkii clarkii, 76
Procambarus clarkii paeninsulanus, 76
Procambarus clemmeri,73
Procambarus clypeata, 31
Procambarus cometes, 56
Procambarus connus, 56
Procambarus consobrinus, 52
Procambarus contrerasi, 77
Procambarus cubensis consobrinus, 52
Procambarus cubensis cubensis, 52
Procambarus cubensis rivalis, 52
Procambarus Digueii,75
Procambarus digueii, 75
Procambarus dupratzi, 73
Procambarus echinatus,73
Procambarus econfinae, 61
Procambarus elegans, 73
Procambarus enoplostemum, 65
Procambarus enoplosterum, 65
Procambarus epicyrtus, 65
Procambarus ercichsoni, 78
Procambarus erichsoni, 78
Procambarus erythrops, 65
Procambarus escambiensis, 61
Procambarus everrnani, 66
Procambarus e vermann i, 66
Procambarus fallax, 66
Procambarus Fallicambarus fodiens, 29
Procambarus fitzpatricki, 51
Procambarus geminus, 66
Procambarus geodytes, 58
Procambarus gibbus, 73
Procambarus gonopodocrisutus, 66
Procambarus gracilis, 56
Procambarus hagenianus, 55,56,57
Procambarus hagenianus forrestae.55
Procambarus hagenianus hagenianus, 56
Procambarus hayi, 66
Procambarus hinei, 55
Procambarus hineyi, 55
Procambarus hinutus, 67
Procambarus Hcffrnani, 78
Procambarus Hoffmanni,78
Procambanis hoffmanni, 78
Procambarus honti, 67
Procambarus hortonhobbsi,78
Procambanis hottonhobsi, 78
Procambanis howardi, 14
Procambarus howellae, 76
Procambarus bubbelli, 61
Procambarus hybus, 67
Procambanis inalis, 55
Procambanis inciUn, 55
Procambanis jaculus, 67
Procambarus kilbyi, 61
Procambarus lagniappe, 73
Procambanis latipfeurum, 61
Procambanis layi, 66
Procambarus lecontei, 67
Procambanis lecnensts, 68
Procambanis Iepidodactylus,68
Procambarus lewis i, 68
Procambarus liberorum, 57
Procambarus utocternum,68
Procambarus llamasi, 52
Procambarus llamesi, 52
Procambarus lophotus, 68
Procambanis lucifugus alachua, 68
Procambarus lucifugus lucifugus, 68
Procambarus lunzi, 69
Procambarus lylei, 73
Procambarus mancus, 69
Procambarus mexiacnus, 52
Procambarus Mexicanus, 53
Procambarus mexicanus, 52
Procambarus milleri, 61
Procambarus mirandae, 53
Procambarus mirandai, 53
Procambarus natchitochae, 51,72,74
Procambarus niveus,53
Procambarus oaxacae reddelli, 53
Procambarus okaloosae, 76
Procambarus orcinus, 69
Procambanis ouachitae, 74
Procambarus ouichitae, 74
Procambarus paeninsulanis, 77
Procambarus paeninsulans,77
Procambarus paeninsulanus, 77
Procambarus pallidus, 69
procambarus pallidus, 69
Procambarus Paradoxus, 72
Procambarus pearsei,70
Procambarus pearsei pearse t, 70
Procambarus pearsei plumimanus, 70
Procambarus pecki, 51,75
Procambarus pelosimanus, 53
Procambarus penni, 74
Procambarus petersi, 74
Procambarus pictus, 70
Procambanis pilosimanus, 53
Procambarus planirostris, 70
Procambarus planirostms, 70
Procambarus plantirostms,70
Procambarus plantiuostrusg, 70
Procambarus plumimanus, 70
Procambarus poqum, 57
Procambanis pubescens, 70
Procambarus pubischaelae, 62
Procambarus pubischelae, 62
Procambanis pycnogonopodus, 70
Procambarus pygmaeus, 59
Procambarus quachitae, 74
Procambarus ranei,74
Procambarus raneyi, 74
Procambanis raihbunae, 62
Procambanis rayeni, 74
Procambarus reimeri,57
Procambarus riojae, 78
Procambarus riojai, 78
Procambarus roberti, 74
Procambarus rodriguezi,54
Procambanis rogersi, 59
Procambarus rogersi campestris, 59
Procambarus rogersi expletus, 59
Procambarus rogersi ochlockensis, 59
Procambarus rogersi ochlocknensis, 59
Procambarus rogersi rogersi, 59
Procambarus rusticus, 49
Procambarus ruthveni, 54
NUMBER 480 231
Procambanis nithveni ruthveni, 54
Procambarus nithveni zapaopcnsis, 55
Procambarus nithveni zapoapensis, 55
Procambanis seminolae, 71
Procambanis shermani, 62
Procambanis simulans, 58
Procambanis simulans regimontanus, 57
Procambanis simulans regicmontanus, 57
Procambanis simulans simulans, 58
Procambanis spiculifer.74,75
Procambanis spiculifter, 75
Procambanis stimulans, 58
Procambanis sutikusi, 75
Procambanis suttkuzi, 75
Procambanis tenuis, 51,77
Procambanis texanus, 71
Procambanis teziutlanensis,78
Procambarus tlapacoyanensis.78
Procambanis toliecae, 71
Procambanis toltecas,71
Procambanis troglodytes, 77
Procambanis Truculentus, 60
Procambarus truculentus, 60
Procambarus tulanei, 58
Procambanis tziutlanensis,78
Procambanis vasquezae, 54
Procambanis vazquezae, 51,54
Procambarus veracnizanus, 54
Procambanis verrucosus, 71
Procambarus versutus,75
Procambarus viae-viridis,71
Procambanis viaeviridis, 71
Procambanis vUlalobosi,71
Procambanis vioscae, 74,75
Procambarus vioscai, 75
Procambarus viosci, 75
Procambarus Weigmanni, 5
Procambanis weigmanni, 5
Procambarus wiegmanni, 5
Procambanis williamsooi, 54
Procambarus youngi, 72
Procambanis zihuateutlensis,79
Procambarus (Acucauda) rnzpatricki, 51,85 (211)
Procambanis (Austrocambarus) acanthophorus, 52,85 (223)
Procambanis (Austrocambarus) alkinsoni, 52,85 (226)
Procambanis (Austrocambarus) aztecus, 53
Procambanis (Austrocambarus) cubensis cubensis, 52, 85, (228)
Procambanis (Austrocambarus) cubensis rivalis, 52,85 (227)
Procambarus (Austrocambarus) llamasi, 52, 85 (224)
Procambarus (Austrocambarus) mexicanus, 52,53,85 (214)
Procambanis (Austrocambarus) mirandai, 53, 85 (219)
Procambanis (Austrocambarus) niveus, 53,85 (229)
Procambanis (Austrocambarus) oaxacac oaxacae, 53,85 (216)
Procambanis (Austrocambarus) oaxacae reddclli, 53,85 (217)
Procambanis (Austrocambarus) olmecorum, 87,91 (379)
Procambanis (Austrocambarus) pilosimanus, 53,85 (225)
Procambanis (Austrocambarus) primaevus, 5,53,85
Procambarus (Austrocambarus) rodiguezi, 54
Procambanis (Austrocambarus) rodriquezi, 54,85 (215)
Procambarus (Austrocambarus) ruthveni, 54,85 (220)
Procambanis (Austrocambarus) ruthveni ruthveni, 54
Procambanis (Austrocambarus) sbordonu, 54,85 (213)
Procambanis (Austrocambarus) vazquezae, 54,85 (212)
Procambanis (Austrocambarus) veracnizanus, 54,85 (218)
Procambarus (Austrocambarus) williamsoni, 54,85 (222)
Procambarus (Austrocambarus) zapoapensis, 55,85 (221)
Procambanis (Cambarus) daikii.76
Procambanis (Capillicambarus) brazorensis, 55
Procambanis (Capillicambarus) brazoriensis,55,85 (231)
Procambarus (Capillicambarus) hinci, 55,85 (230)
Procambanis (Capillicambarus) incilis, 55,85 (232)
Procambanis (Distocambarus) devexus, 28
Procambanis (Girardiella) barbiger. 55,85 (246)
Procambanis (Girardiella) cometes, 55,86 (244)
Procambarus (Girardiella) connos, 56
Procambanis (Girardiella) cormus, 56,86 (243)
Procambarus (Girardiella) curdi, 56,86 (235)
Procambarus (Girardiella) ferrugineus, 87,92
Procambanis (Girardiella) gracilis, 56,86 (239)
Procambarus (Girardiella) hagenianus, 55,56,57
Procambanis (Girardiella) hagenianus hagenianus, 56,57,86 (242)
Procambanis (Girardiella) hagenianus vesticeps, 57,86 (241)
Procambarus (Girardiella) libcrorum, 57,86 (240)
Procambarus (Girardiella) parasimulans, 57,86 (237)
Procambarus (Girardiella) pogum, 57,86 (245)
Procambarus (Girardiella) regalis, 87,92
Procambarus (Girardiella) regiomontanus, 57,86 (238)
Procambarus (Girardiella) reimeri, 57,86 (236)
Procambarus (Girardiella) simulans, 57,58,86 (234)
Procambarus (Girardiella) simulans regiomontanus, 57
Procambarus (Girardiella) simulans simulans, 58
Procambarus (Girardiella) tulanei, 58,86 (233)
Procambanis (Hagenides) advena, 58,59,86 (247)
Procambarus (Hagenides) caritus, 58.86 (249)
Procambarus (Hagenides) geodytes, 58,86 (250)
Procambanis (Hagenides) pygmaeus, 58,59,86 (251)
Procambarus (Hagenides) rogersi campestris, 59,86 (253)
Procambanis (Hagenides) rogersi exptetus, 59,86 (254)
Procambarus (Hagenides) rogersi ochlocknensis, 59,86 (252)
Procambarus (Hagenides) rogersi ocklocknensis, 59
Procambarus (Hagenides) rogersi rogersi, 59,86 (255)
Procambarus (Hagenides) talpoides, 59.86 (248)
Procambanis (Hagenides) truculentus, 60,86 (256)
Procambanis (Leconticambanis) alleni, 60,86 (257)
Procambarus (Leconticambanis) apalachicolae, 60,86 (261)
Procambanis (Leconticambarus) barbatus, 60,86 (259)
Procambarus (Leconticambanis) capiUmtui, 60,86 (267)
Procambanis (Leconucarnbarus) econfinae, 61,86 (262)
Procambanis (Leconticambanis) escambiensis, 60,61,86 (263)
Procambanis (Leconticambanis) hubbelli, 61,86 (269)
Procambarus (Leconticambarus) kilbyi, 61,86 (268)
Procambarus (Leconticambarus) latipleurum, 61,86 (260)
Procambanis (Leconticambarus) milleri, 61,86 (258)
Procambanis (Leconticambanis) pubischelae, 62
Procambanis (Leconticambanis) pubischelae deficiens, 62,86 (265)
Procambarus (Leconticambanis) pubischelae pubischelae, 62,86 (264)
Procambarus (Leconticambanis) rathbunae, 62,86 (266)
Procambanis (Leconticambarus) shermani, 62,86 (f70)
Procambanis (Lonnbergius) acherontis, 62,86 (271)
Procambarus (Mexicambarus) bouvieh, 62,86 (272)
Procambarus (Ortmanicus) acutus acutus, 64
Procambanis (Ortmanicus) hirsutus, 67
Procambarus (Ortmanicus) pubescens, 70
Procambanis (Orunannicus) actus actus, 63
Procambarus (Ortmamicus) ?cutisiimus, 63,86 (308)
Procambanis (Orunannicus) acutus acutus, 63,86 (301)
Procambarus (Ortmamicus) acutus cuevachicae. 64,86 (302)
Procambanis (Oronannicus) ancylus, 64,86 (293)
Procambarus (Ortmamicus) angusutus, 64,86 (282)
Procambanu (Ortmamicus) hi vitutui, 64.86 (319)
232 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Procambarus (Ortmannicus) blandingii, 64,86 (300)
Procambarus (Ortmannicus) caballeroi, 65,86 (321)
Procambarus (Ortmannicus) chacei, 65,86 (275)
Procambarus (Ortmannicus) clarki, 76
Procambarus (Ortmannicus) delicatus, 65,86 (291)
Procambarus (Ortmannicus) enoplostemum, 65,86 (274)
Procambarus (Ortmannicus) epicyrtus, 65. 86 (276)
Procambarus (Ortmannicus) erythrops, 65,86 (286)
Procambarus (Ortmannicus) evermanni, 66,86 (297)
Procambarus (Ortmannicus) fallax, 66,86 (295)
Procambarus (Ortmannicus) franzi, 66,86 (287)
Procambarus (Ortmannicus) geminus, 66,86 (305)
Procambarus (Ortmannicus) gonopodocristatus, 66,86 (324)
Procambarus (Ortmannicus) hayi, 66,86 (306)
Procambarus (Ortmannicus) hirsutus, 67,86 (280)
Procambarus (Ortmannicus) horsti, 67,86 (289)
Procambarus (Ortmannicus) hybus, 67,86 (311)
Procambarus (Ortmannicus) jaculus. 67,86 (310)
Procambarus (Ortmannicus) lecontei, 67,86 (304)
Procambarus (Ortmannicus) leitheuseri, 67,86 (283)
Procambarus (Ortmannicus) leonensis, 68,86 (296)
Procambarus (Ortmannicus) lepidodactylus, 68,86 (278)
Procambarus (Ortmannicus) lewisi, 68,86 (318)
Procambarus (Ortmannicus) 1 itostemum, 68,86 (277)
Procambarus (Ortmannicus) lophotus, 68,86 (307)
Procambarus (Ortmannicus) lucifugus alachua, 68,86 (284)
Procambarus (Ortmannicus) lucifugus lucifugus, 63,68,86 (285)
Procambarus (Ortmannicus) lunzt, 69,86 (294)
Procambarus (Ortmannicus) mancus, 69,86 (313)
Procambarus (Ortmannicus) marthae, 69,86 (317)
Procambarus (Ortmannicus) medialis, 69. 86 (315)
Procambarus (Ortmannicus) orcinus, 69. 86 (290)
Procambarus (Ortmannicus) pallidus, 69. 86 (288)
Procambarus (Ortmannicus) pearsei,70,86 (316)
Procambarus (Ortmannicus) pearsei pearsei, 70
Procambarus (Ortmannicus) pearsei plumimanus, 70
Procambarus (Ortmannicus) pictus, 70,86 (273)
Procambarus (Ortmannicus) planirostris, 70,86 (312)
Procambarus (Ortmannicus) plumimanus, 70,86 (314)
Procambarus (Ortmannicus) pubcscens, 70,86 (279)
Procambarus (Ortmannicus) pycnogonopodus, 70,86 (299)
Procambarus (Ortmannicus) seminoUe, 71,86 (292)
Procambarus (Ortmannicus) simulans, 58
Procambaius (Ortnunnicus) texanu s, 71,86 (303)
Procambarus (Ortmannicus) toltccae, 71,86 (323)
Procambarus (Ortmannicus) vemicosus, 71,86 (309)
Procambarus (Ortmannicus) viaeviridis, 71,87 (320)
Procambarus (Ortmannicus) viOalobosi, 71,87 (322
Procambarus (Ortmannicus) xiliuc, 72
Procambarus (Ortmannicus) Xilitlae, 72
Procambarus (Ortmannicus) xilitlae, 72,87 (298)
Procambarus (Ortmannicus) youngi, 72,87 (281)
Procambarus (Pancambarus) ortmanii, 72
Procambarus (Paracambtrus) ortmarmi, 72
Procambarus (Pancambarus) ortmamii, 72,87 (325)
Procambarus (Pancambarus) paradoxus, 72,87 (326)
Procambarus (Permides) ablusus.72,87 (329)
Procambarus (Pennides) dcmmeri,73, 87 (335)
Procambarus (Pennides) dupntzi, 73,87 (336)
Procambarus (Pennides) echinatus, 73,87 (334)
Procambarus (Pennides) elegans, 73,87 (331)
Procambarus (Pennides) gibbus, 73.87 (342)
Procambarus (Pennides) l?gniappe,73,87 (339)
Procambarus (Pennides) lyki,73,87 (333)
Procambarus (Pennides) natchitochae. 74,87 (328)
Procambarus (Pennides) ouachitae,74,87 (343)
Procambarus (Pennides) penni, 74, 87 (338)
Procambarus (Pennides) petersi, 74,87 (340)
Procambarus (Pennides) raneyi,74,87 (344)
Procambarus (Pennides) roberti,74,87 (330)
Procambarus (Pennides) spiculifer, 74,75,87 (341)
Procambarus (Pennides) suttkusi,75,87 (327)
Procambarus (Pennides) versutus, 75,87 (332)
Procambarus (Pennides) vioscai, 75, 87 (337)
Procambarus (Procambarus) digueti,75,87 (345)
Procambarus (Remoticambarus) pecki, 75,87 (346)
Procambarus (Scapulicambarus) clarkii,76,87 (351)
Procambarus (Scapulicambarus) howellae, 76,87 (349)
Procambarus (Scapulicambarus) okaloosae, 76,87 (350)
Procambarus (Scapulicambarus) paeninsulanus, 76,77,87 (348)
Procambarus (Scapulicambarus) strenthi, 77,87 (347)
Procambarus (Scapulicambarus) troglodytes, 77,87 (352)
Procambarus (Scapulicambarus) xilitlae, 72
Procambarus (Tenuicambarus)tenuis,77,87 (353)
Procambarus (Villalobosus) contrerasi, 77,87 (358)
Procambarus (Viflalobosus) cuetzalanae,77,87 (355)
Procambarus (Villalobosus) erichsoni,78,87 (357)
Procambarus (Villalobosus) hoffmanni, 78, 87 (356)
Procambarus (Villalobosus) hortonhobbsi, 78, 87 (359)
Procambarus (Villalobosus) riojai,78,87 (354)
Procambarus (Villalobosus) teziutlanensis,78,87 (362)
Procambarus (Villalobosus) tlapacoyanensis,78,87 (363)
Procambarus (Villalobosus) xochitlanae,79,87 (361)
Procambarus (Villalobosus) zihuatentlensis, 79
Procambarus (Villalobosus) zihuateutlensis,79,87 (360)
Procambasrus, 51,63
Procambasrus acutus acutus, 64
Procamberus,51
Procamborus, 51,76
Procamborus clarkii, 76
Procambrus, 51,76
Procambrus darkii, 76
Procambrus pallidus, 69
Procamburus, 51,76
Procamburus clarki i, 76
Procericambarus, subgenus, 3,4,33,45
prolixus, Cambarellus, 10
prolixus.Cambarellus (Cambarellus), 10,82 (13)
prominens, Cambarus, 33,84
prominens, Hobbseus, 33
propinguis, Orconectes, 37
propinkuus, Cambarus, 37,38
propinques, Orconectes, 37
propinquus, Astacus, 36
propinquus, Camarus, 37
propinquus, Cambanrus, 37
Propinquus, Cambarus, 37
propinquus, Cambarus, 36
propinquus, Cambarus (Faxonius), 37
propinquus, Cambarus (Fraxonius), 37
propinquus, Camborus, 37
propinquus, Faxonius, 37
propinquus, Orconectes, 37
propinquus, Orconectes (Crockerinus), 37,84 (156)
propinquus, Orconnectes, 37
propinquus erismophorous, Orconectes, 37
propinquus jef f ersotii, Faxonius, 36
propinquus jeff ersoni, Orconectes, 36
propinquus jeff ersoni, Orconectes (Orconectes), 36
propinquus obscura, Cambarus, 36
NUMBER 480 233
propinquus obscurus, Cambarus, 36
propinquus propinquus, Cambarus 36
propinquus propinquus, Faxonius, 37
propinquus propinquus, Faxonius (Faxonius), 37
propinquus propinquus, Orconectes, 37
propinquus propinquus, Orconectes (Orconectes), 37
propinquus sanbomi, Cambarus, 37,38
propinquus sanbomi, Cambarus (Faxonius), 37
propinquus sanbomi, Orconectes, 37
propinquus sanbomi, Orconectes (Orconectes), 37
propinquus Sanbomii, Cambarus, 37
propinquus sanbomii, Cambarus, 37
propinquus typicus, Cambarus, 37
propinquus var. obscura, Cambarus, 36
propinquus var. sanbomi, Cambarus, 37
propinquus var. sanbomii, Cambarus, 37
propoinquus, Orconectes, 37
pubescens, Cambarus, 70
pubescens, Cambarus (Cambarus),70
pubescens, Cambarus (Ortmannicus),70
pubescens, Procambarus, 70
pubescens, Procambarus (Ortmanicus),70
pubescens, Procambarus (Orunannicus), 70,86 (279)
pubischaelae, Procambarus, 62
pubischelae, Procambarus, 62
pubischelae, Procambarus (Leconticambarus), 62
pubischelae deficiens, Procambarus (Leconticambarus), 62,86 (265)
pubischelae pubischelae, Procambarus (Leconticambarus), 62, 86 (264)
puer, Cambarellus, 11
puer, Cambarellus (Pandicambarus), 11,82 (21)
pugnax, Astacus, 3,80
pugnax, Parastacus, 80, 87 (371)
Puncticambarus, subgenus, 4,25
punctimanus, Faxonius, 48
punctimanus, Faxonius (Faxonius), 48
punctimanus, Orconectes, 48
punctimanus, Orconectes (Orconectes), 48
punaimanus, Orconectes (Procericambarus), 48, 85 (171)
pusillus, Astacus, 13
pusillus, Cambarus, 13
Putnami, Cambarus, 49
putnami, Cambarus, 49
putnami, Cambarus (Faxonius), 49
putnami, Orconectes, 49
putnami, Orconectes (Procericambarus), 49,85 (186)
pycnogonopodus, Procambarus, 70
pycnogonopodus, Procambarus (Orunannicus), 70,86 (299)
pygmaeus, Procambarus, 59
pygmaeus, Procambarus (Hagenides), 58,59,86 (251)
pyronotus, Cambarus, 16
pyronotus, Cambarus (Depressicambarus), 16,83 (40)
quachitae, Procambarus, 74
quadruncus, Faxonius, 49
quadruncus, Faxonius (Faxonius), 49
quadruncus, Orconectes, 49
quadruncus, Orconectes (Orconectes), 49
quadruncus, Orconectes (Procericambarus), 49,85 (168)
rafinesquei, Faxonius, 37
rafinesquei, Orconectes, 37
rafinesquei. Orconectes (Crockerinus), 37,84 (163)
rafinesquei, Orconectes (Orconectes), 37
ranei, Procambarus, 74
rancy i, Procambarus, 74
raneyi, Procambarus (Permides), 74,87 (344)
rathbunae, Cambarus, 62
rathbunae, Procambarus, 62
rathbunae, Procambarus (Leconticambarus), 62,86 (266)
rayeni, Procambarus, 74
reburrus, Cambarus, 27
reburrus, Cambarus (Puncticambarus), 27,83 (103)
reduncus, Cambarus, 16
reduncus, Cambarus (Depressicambarus), 16,83 (50)
reflexus, Cambarus (Depressicambarus), 16,83 (39)
regal is, Procambarus (Girardiella), 87,91
regiomontanus, Procambarus (Girardiella), 57,86 (238)
reimeri, Procambarus, 57
reimeri.Procambarus (Girardiella), 57,86 (236)
Remoti cambarus, subgenus, 4,51,75
rhoadesi, Faxonius, 41
rhoadesi, Orconectes, 41
rhoadesi, Orconectes (Gremicambarus),41,85 (207)
rhoadesi, Orconectes (Orconectes), 41
Rhoadesius, subgenus, 3,4,33,50
Ricoleti, Astacus, 80
rinmunis, Cambarus, 40
riojae, Paracambarus, 51,77,78
riojae, Procambarus, 78
riojai, Procambarus, 78
riojai, Procambarus (Villalobosus),78,87 (354)
roberti, Procambarus, 74
roberti, Procambarus (Pennides), 74,87 (330)
robin son i, Bourchardina, 12
robisoni, Bouchardina, 12,82 (26)
robustis, Cambarus, 27
robustus, Cambarus, 27
robustus, Cambarus (Bartonius), 27
robustus, Cambarus (Puncticambarus), 27,83 (104)
rodiguezi, Procambarus (Austrocambarus), 54
rodriguezi, Paracambarus, 54
rodriguezi, Procambarus, 54
rodriguezi, Procambarus (Austrocambarus), 54,85 (215)
rogersi, Cambarus, 59
rogersi, Procambarus, 59
rogersi campestris, Procambarus, 59
rogersi campestris, Procambarus (Hagenides), 59,86 (253)
rogersi expletus, Procambarus, 59
rogersi expletus, Procambarus (Hagenides), 59,86 (254)
rogersi ochlockensis, Procambarus, 59
rogersi ochlocknensis, Procambarus, 59
rogersi ochlocknensis, Procambarus (Hagenides), 59,86 (252)
rogersi ocklocknensis, Procambarus (Hagenides), 59
rogersi rogersi, Procambarus, 59
rogersi rogersi, Procambarus (Hagenides), 59,86 (255)
rusticiformis, Cambarus, 18
rusticiformis, Cambarus (Erebicambarus), 18,83 (52)
rustics placidus, Cambarus, 48
rusticus, Cambarus 49
rusticus, Cambarus (Faxonius), 49
rusticus, Faxonius, 49
rusticus, Orconectes, 49
rusticus, Orconectes (Orconectes), 45
rusticus, Orconectes (Procericambanis),49,85 (174)
rusticus, Orconectus, 49
rusticus, Orconnectes, 49
rusticus, Procambarus, 49
rusticus, var. placida, Cambarus, 48
rusticus, var. placidus, Cambarus, 48
rusticus barrenensis, Faxonius, 45
rusticus barrenensis, Orconectes, 45
rusticus barrensis, Orconectes, 45
234 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
rusticus forceps, Cambarus, 45
msticus forceps, Cambarus (Faxonius), 45
rusticus forceps, Orconectes, 45
rusticus forceps, Orconectes (Orconectes), 45
rusticus minis, Cambarus (Faxonius), 47
rusticus minis, Orconectes, 47
rusticus minis, Orconectes (Orconectes),47
rusticus placidus,Cambarus, 48
rusticus placidus, Cambarus (Faxonius), 48
rusticus placidus, Orconectes, 48
rusticus placidus, Orconectes (Orconectes), 48
rusticus rusticus, Cambarus (Faxonius), 49
rusticus rusticus, Faxonius, 49
rusticus rusticus, Faxonius (Faxonius), 49
rusticus rusticus, Orconectes, 49
rusticus rusticus, Orconectes (Orconectes), 49
rustrineus rustrineus, Cambarus, 49
ruthveni, Cambarus, 54
ruthveni, Procambarus, 54
ruthveni, Procambarus (Austrocambarus), 55,85 (220)
ruthveni ruthveni, Procambarus, 54
ruthveni ruthveni, Procambarus (Austrocambarus), 55
ruthveni zapaopensis, Procambarus, 55
ruthveni zapoapensis, Procambarus, 55
ruthveni zapoapensis, Procambarus (Austrocambarus), 55
saffordi, Parastacus, 80,87 (368)
Samasucus.3,4,80
Samastacus araucanius, 3,80,87 (374)
Samastacus spinifrons, 3,81,87 (373)
Sambarus, 12
Sambarus diogenes, 24
sanbomi, Faxonius, 37
sanbomi, Faxonius (Faxonius), 37
sanborni, Orconectes, 37
sanbomi erismophorous, Orconectes, 37
sanbomi erismophorus, Orconectes, 37
sanbomi sanbomi, Faxonius, 37
sanbomi sanbomi, Orconectes, 37
Sanbomii, Cambarus, 33,35,37
sanbomii erismophorous, Orconectes, 37
sanbomii erismophorous, Orconectes (Crockerinus), 37,84 (160)
sanbomii sanbomii, Orconectes, 37
sanbomii sanbomii, Orconectes (Crockerinus), 37,84 (159)
saxaulis, Orconectes, 49
saxaulis, Orconectes (Procericambarus), 49,85 (175)
sbordonii, Procambarus (Austrocambarus), 54,85 (213)
Scapulicambarus, subgenus, 4,51,76
jchmitii, Cambarellus, 8,10,11
schmitti, Cambarellus (Pandicambarus), 11,82 (19)
schmittii, Cambarus, 5,11
schufeldi, Cambarellus, 10
sciotensis, Cambarus, 14
sciotaisis, Cambarus (Cambarus), 14,83 (33)
scotii, Cambarus (Puncucambarus), 27, 83 (100)
seminolae, Procambaris, 71
seminolae, Procambarus, 71
seminolae, Procambarus (Ortmannicus),71,86 (292)
tenborni, Orconectes, 37
setocus, Cambarus, 23
setosus, Cambarus (Bartonius), 23
setosus, Cambarus (Cambarus) 23
setosus, Cambarus (Jugicambarus), 23,83 (84)
shermani, Procambarus, 62
shermani, Procambarus (Lecontkambarus),62,86 (270)
shoupi, Cambarus, 38
shoupi, Faxonius, 38
shoupi, Orconectes, 38
shoupi, Orconectes (Crockerinus), 38,84 (167)
shoupi, Orconectes (Orconectes), 38
shufeldti, Cambarella, 10
shufeldti, Cambarellus, 10
shufeldti, Cambarus, 10
shufeldti, Cambarus (Cambarellus), 10
shufeldtii, Cambarellus, 10
shufeldtii, Cambarellus (Dirigicambarus), 10,82 (17)
Shufeldtii, Cambarus, 8,10
shufeldtii, Cambarus, 10
shufeldtii, Cambarellus, 10
shufeltdii, Cambarellus, 10
shulfeldtii, Cambarellus, 10
signifer, Cambarus, 40
similans, Cambarus, 58
simulana, Cambarus, 58
simulans, Cambarus, 57
simulans, Cambarus (Cambarus),58
simulans, Cambarus (Ortmannicus), 58
simulans, Procambarus, 58
simulans, Procambarus (Girardiella), 57,58,86 (234)
simulans regiomontanus, Procambarus, 57
simulans regiomontanus, Procambarus (Girardiella), 57
simulans simulans, Procambarus, 58
simulans simulans, Procambarus (Girardiella), 58
sloanei, Cambarus, 50
sloanei, Cambarus (Faxonius), 50
sloanei, Orconectes, 50
sloani, Cambarus, 50
sloani, Orconectes, 50
sloani, Orconectes (Orconectes), 50
Sloanii, Cambarus, 50
sloanii, Cambarus, 33,50
sloanii, Faxonius, 50
sloanii, Faxonius, (Faxonius) 50
sloanii, Orconectes, 50
sloanii, Orconectes (Orconectes), 50
sloanii, Orconectes (Rhoadesius), 50,85 (150)
sloanni, Orconectes, 50
speciosus, Cambarus (Hiaticambarus), 20,83 (65)
spectrum, Fallicambarus (Fallicambarus), 31
sphenoides, Cambarus, 16
sphenoides, Cambarus (Depressicambarus), 16,83 (41)
spicatus, Cambarus, 27
spicatus, Cambarus (Puncticambarus), 27,84 (94)
spiculifer, Astacus, 74
spiculifer, Cambaru, 75
spiculifer, Cambarus, 74
spiculifer, Cambarus (Cambarus), 75
spiculifer, Cambarus (Ortmannicus), 75
spiculifer, Procambarus, 75
spiculifer, Procambarus (Permides),75,87 (341)
spiculifter, Procambarus, 75
spinifrons, Astacus, 80,81
spinifrons, Parastacus, 81
spinifrons, Samastacus, 3, 81,87 (373)
spinirostris, Cambarus, 19
spinosus, Cambarus, 50
spinosus, Cambarus (Faxonius), 50
spinosus, Orconectes, 50
spinosus, Orconectes (Procericambarus), 50,85 (185)
spinosus gulielmi, Cambarus, 35
spinous, Orconectes, 50
NUMBER 480 235
stannardi, Orconectes, 38
stannardi, Orconectes (Crockerinus), 38,84 (161)
Stigius, Astacus Cambams, 45
stimulans, Procambarus, 58
strawni, Cambarus, 29,30,31
slrawni, Fallicambarus, 31
strawni, Fallicambarus (Fallicambarus), 31,84 (114)
strenthi, Procambanis (Scapulicambarus),77,87 (347)
striatus, Cambarus, 17
striatus, Cambarus (Depressicambarus), 15,16,17,83 (48)
strigosus, Cambarus (Depressicambarus), 17,83 (45)
Stygius, Cambarus, 45,63
stygius, Cambarus, 45,63
subgrundialis, Astacus, 6
suttkusi, Procambarus, 75
suttkusi, Procambarus (Permides),75,87
suttkuzi, Procambarus, 75
talpoides, Procambarus (Hagenides), 59,86 (248)
tartarus, Cambarus, 23
tartarus, Cambarus (Jugicambarus), 23,83 (86)
lenebrorus, Cambarus, 18
tenebrosus, Cambarus, 18
tenebrosus, Cambarus (Erebicambarus), 18,83 (54)
Tenuicambarus, subgenus, 4,51,77
tenuis, Procambarus, 51,77
tenuis, Procambarus (Tenuicambarus),77,87 (353)
texanus, Cambarellus, 11
texanus, Cambarellus (Pandicambarus), 11,82 (22)
texanus, Procambarus, 71
texanus, Procambarus (Ortmannicus),71,86 (303)
teziutlanensi, Paracambarus, 78
teziutlanensis, Paracambarus, 78
teziutlanensis, Procambarus, 78
teziutlanensis, Procambarus (Villalobosus), 78,87 (362)
tlapacoyanensis, Paracambarus, 78
tlapacoyanensis, Procambarus, 78
tlapacoyanensis, Procambarus (Villalobosus), 78,87 (363)
toltecae, Procambanis, 71
toltecae, Procambarus (Ortmannicus), 71,86 (323)
toltecas, Procambarus, 71
Tragulicambarus, subgenus, 3,4,33,50
transfuga, Orconectes, 48
transfuga, Orconectes (Procericambarus),48
transfugus, Orconectes, 48
trawbridgii, Pacifastacus, 8
tricuspis, Faxonius, 38
tricuspis, Orconectes, 38
tricuspis, Orconectes (Crockerinus), 38,84 (165)
tricuspis, Orconectes (Orconectes), 38
Troglocambarus, 3,4,79
Troglocambarus madanei, 79,87 (364)
troglodyles, Cambarus, 77
troglodytes, Astacus, 77
troglodytes, Cambarus, 77
troglodytes, Cambarus (Cambarus), 77
troglodytes, Cambarus (Ortmannicus), 77
troglodytes, Procambarus, 77
troglodytes, Procambarus, (Scapulicambarus) 77,87 (352)
troglodytus, Cambarus, 77
Trolocambarus, 79
Trolocambarus madanei, 79
trowbridgei, Astacus, 8
trowbridgei, PacifasUcus, 8
trowbridgei, Potamobius (Potamobius), 8
trowbridgeii, PacifasUcus, 7,8
trowbridgeii, Pasifastacus, 8
trowbridgi, Astacus, 8
trowbridgi, Pacifastacus, 8
Trowbridgii, Astacus, 8
trowbridgii, Astacus, 8
trowbridgii, Astacus (Astacus), 8
Trowbridgii, Astaus (Astacus), 8
trowbridgii, Pacifastaculus, 8
Trowbridgii, Pacifastacus, 8
trowbridgii, Pacifastacus, 8
Trownbridgi, Astacus, 8
Truculentus, Procambarus, 60
truculentus, Procambarus, 60
truculentus, Procambarus (Hagenides), 60,86 (256)
truncatus, Cambarus (Depressicambarus), 17,83 (43)
tulanei, Procambarus, 58
tulanei, Procambarus (Girardiella),58, 86 (233)
typhlobius, Cambarus, 45
typhlobius, Gambarus, 45
tziutlanensis, Procambarus, 78
Uhleri, Cambarus, 30
uhleri, Cambarus, 30
uhleri, Cambarus (Bartonius), 30
uhleri, Cambarus (Cambarus), 30
uhleri, Fallicambarus, 30
uhleri, Fallicambarus (Creaserinus),30,84 (119)
uneslami, Cambarus, 23
unestami, Cambarus (Depressicambarus), 23,83 (69)
unestami, Cambarus (Jugicambarus), 23
validus, Cambarus, 41,42
validus, Cambarus (Faxonius), 42
validus, Faxonius, 42
validus, Faxonius (Faxonius), 42
validus, Orconectes, 42
validus, Orconectes (Gremicambarus),42,85 (208)
validus, Orconectes (Orconectes), 42
validus Faxonianus, Cambarus, 41
valleculus, Cambarus, 33
valleculus, Hobbseus, 33,84 (131)
varicosus, Parasucus, 80,87 (367)
vasquczae, Procambarus, 54
vazquezae, Procambarus, 50,54
vazquezae, Procambarus (Austrocambarus), 54,85 (212)
veracruzanus, Procambarus, 54
veracruzanus, Procambarus (Austrocambarus), 54,85 (218)
verrucosus, Procambarus, 71
verrucosus, Procambarus (Ortmannicus), 71,86 (309)
versutus, Cambarus, 75
versutus, Cambarus (Cambarus), 75
versutus, Cambarus (Ortmannicus), 75
versutus, Procambarus, 75
versutus, Procambarus (Pennides), 75,87 (332)
veteranus, Cambarus, 27
veteranus, Cambarus (Puncticambarus), 27,84 (105)
Veticambarus, subgenus, 4,12,20,28
viae viridis, Cambarus (Ortmannicus), 71
viae-viridis, Cambarus, 71
viae-viridis, Procambarus, 71
viaeviridis, Procambarus, 71
viaeviridis, Procambarus (Ortmannicus), 71,87 (320)
villalobosi, Procambarus, 71
villalobosi, Procambarus (Ortmannicus), 71,87 (322)
Villalobosus, subgenus, 4,51,77
vioscae, Procambarus, 74,75
vioscai, Procambarus, 75
236 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
vioscai, Procambarus (Pennides), 75,87 (337)
viosci, Procambarus, 75
virginianus, Orconecles, 38
virginiensis, Faxonius, 38
virginiensis, Orconectes, 38
virginiensis, Orconectes (Crockerinus), 38,84 (158)
virginiensis, Orconectes (Otconectes), 38
viridis, Cambarus, 42
viridis, Camburus, 42
virile, Cambarus, 42
viriles, Cambarus, 42
virilis, Cambarus, 33,39,42
virilis, cambarus, 42
virilis, Cambarus (Faxonius), 42
virilis, Faxonius, 42
virilis, Faxonius (Faxonius), 42
virilis, Orconectes, 42
virilis, Orconectes (Cambarus), 42
virilis, Orconectes (Gremicambarus), 5,39,40,42,85 (199)
virilis, Orconectes (Orconectes), 42
virilis, Orconectis, 42
virilis, Orconelctes, 42
virilis, Orconetes, 42
virilis, Orconnectes, 42
virilis, Oronectes, 42
virilus, Cambarus, 42
vusu'cus, Cambarus, 49
Weigmanni, Procambarus, 5
weigmanni, Procambarus, 5
whitmani, Cambarus, 46
Wiegmanni, Astacus (Cambarus), 5
Wiegmanni, Cambarus, 5
wiegmanni, Cambarus, 5
wiegmanni, Procambarus, 5
williamsi, Orconectes, 50
williamsi, Orconectes (Procericambarus), 50,85 (191)
williamsoni, Cambarus (Procambarus), 54
williamsoni, Procambarus, 54
williamsoni, Procambarus (Austrocambarus), 54,85 (222)
wisconsinensis, Cambarus, 42
wrighti, Faxonius, 39
wrighti, Orconectes, 39
wrighti, Orconectes (Faxonius), 39,84 (141)
wrighti, Orconectes (Orconectes), 39
Xilitlae, Procambarus (Ortmannicus),72
xilitae, Procambarus (Ortmannicus),72
xilillae, Procambarus (Ortmannicus), 72,87 (298)
xilitlae, Procambarus (Scapulicambarus),72
xochitlanae, Procambarus (Villalobosus), 79,87
youngi, Procambarus, 72
youngi, Procambarus (Ortmannicus), 72,87 (281)
youngineri, Distocambarus (Fitzcamharus),28, 84 (110)
zapoapensis, Procambarus (Austrocambarus), 54,55,85 (221)
zempoalensis, Cambarellus, 10
zempoalensis, Cambarellus (Cambarellus), 10,82 (16)
zihuatentlensis, Procambarus (Villalobosus), 79
zihuateutlensis, Procambarus, 79
zihuateutlensis, Procambarus (Villalobosus), 79, 87 (360)
zophonastes, Cabmams (Jugicambarus), 24
zophonastes, Cambarus, 24
zophonastes, Cambarus (Jugicambarus), 24,83 (85)
An Illustrated Checklist
of the American Crayfishes
(Decapoda: Astacidae, Cambaridae,
and Parastacidae)
HORTON H. HOBBS, JR.
m
i
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY ? NUMBER 480
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Use the parentheses/colon system for volume (number):
pagination: "10(2):5-9." For alignment and arrangement of ele-
ments, follow the format of recent publications in the series for
which the manuscript is intended. Guidelines for preparing bibli-
ography may be secured from Series Section, SI Press.
Legends for illustrations must be submitted at the end of the
manuscript, with as many legends typed, double-spaced, to a
page as convenient.
Illustrations must be submitted as original art (not copies)
accompanying, but separate from, the manuscript. Guidelines for
preparing art may be secured from Series Section, SI Press. All
types of illustrations (photographs, line drawings, maps, etc.) may
be intermixed throughout the printed text. They should be termed
Figures and should be numbered consecutively as they will
appear in the monograph. If several illustrations are treated as
components of a single composite figure, they should be desig-
nated by lowercase italic letters on the illustration; also, in the
legend and in text references the italic letters (underlined in copy)
should be used: "Figure 9b." Illustrations that are intended to
follow the printed text may be termed Plates, and any components
should be similarly lettered and referenced: "Plate 9b." Keys to
any symbols within an illustration should appear on the art rather
than in the legend.
Some points of style: Do not use periods after such abbrevi-
ations as "mm, ft. USNM, NNE." Spell out numbers "one' through
nine" in expository text, but use digits in all other cases if possible.
Use of the metric system of measurement is preferable; where
use of the English system is unavoidable, supply metric equiva-
lents in parentheses. Use the decimal system for precise meas-
urements and relationships, common fractions for approximations.
Use day/month/year sequence for dates: "9 April 1976." For
months in tabular listings or data sections, use three-letter abbre-
viations with no periods: "Jan, Mar, Jun," etc. Omit space between
initials of a personal name: "J.B Jones."
Arrange and paginate sequentially every sheet of manu-
script in the following order: (1) title page. (2) abstract, (3) con-
tents, (4) foreword and/or preface, (5) text, (6) appendixes, (7)
notes section, (8) glossary, (9) bibliography (10) legends, (11)
tables. Index copy may be submitted at page proof stage, but
plans for an index should be indicated when manuscript is sub-
mitted.
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