TROPICAL NORTH AMERICAN SPECIES OF PANICUM. 
By A. S. Hitchcock and Agnes Cha.se. 
INTRODUCTION. 
In a preceding paper1 our knowledge of the North American species 
of Panicum was presented in a revision of the genus, based upon mate- 
rial in the United States National Herbarium. Since the publication * 
of that paper a large amount of additional material from tropical 
North America has been accumulated. Mr. Hitchcock visited Mexico 
in 1910, Central America and Panama in 1911, and Jamaica and Trini- 
dad in 1912. Mrs. Chase visited Sonora in 1910 and Porto Rico in 
1913. The collections of Mr. H. Pit tier in Panama, of Brother Le6n 
of the Colegio de la Salle, Habana, in Cuba, and of several other col- 
lectors in tropical North America, besides specimens from various 
islands of the West Indies received through Dr. I. Urban, serve to 
augment the amount of material studied. As a result of these further 
studies in the genus the range of nearly all the tropical species 
described in the revision can be presented in much greater detail. 
Several South American species hitherto unknown from North 
America and several new species have been discovered, while a few 
doubtful species and a few referred to synonymy have by field work 
and further study been established as valid. 
While the present paper is supplementary to the revision, it is 
designed to be of use for the region covered without necessarily 
referring to the main work. For this reason keys are given for all 
the tropical species, though descriptions are given of additional species 
only. A study in their native habitat of species before known from 
herbarium specimens alone, has necessitated in some cases a revision 
of the description of habit or of duration, or exceptions have been 
found to characters given as common for groups. Such notes are 
given without other description and are to be understood as additions 
to or corrections of the descriptions given in the revision. Following 
the reference to the original publication of each species is a reference 
1
 Contr. TJ. S. Nat. Herb. 10.1910. 
459 
460 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
to the description in the revision, if the species is there described. 
No synonymy is given except such as is additional to that cited in 
the previous work. 
The distribution here given is based upon all the material at 
present in the National Herbarium, including that already cited. 
As in the earlier paper all specimens cited are in the United States 
National Herbarium, unless otherwise stated. Specimens from other 
herbaria are cited when such specimens add to the known range. 
Through the courtesy of Dr. I. Urban the grasses of the Krug and 
Urban Herbarium in the Berlin Museum were submitted for study. 
Many additional specimens are cited from this herbarium, such 
specimens being indicated by the abbreviation UK. U. Herb." The 
range within the region covered is graphically presented by maps. 
For the sake of comparison these maps indicate also the distribution 
in the southern border of the United States of such, species as extend 
into this region. Trinidad and Tobago, though they are continental 
islands and their flora is essentially South American, are included in 
the West Indies because they are so included by Grisebach1 and by 
Urban.2 In order to connect Panama and Trinidad the detailed 
distribution.includes Venezuela and Colombia. 
KEY TO SPECIES ACT) GROUPS. 
Axis of branchlets produced beyond the base of the uppermost epikelet as a point or 
bristle 1 to 6 mm. long  Subgenus Paurochaetii m, p. 463. 
Axis of branchlets not produced into a bristle. (In P. geminatum and P. pahidivagum 
the somewhat flattened axis pointed but not bristle-form.) 
Basal leaves usually distinctly different from those of the culm, forming a winter 
rosette; culms at first simple, the spikelets of the primary panicle not 
perfecting seed, later usually becoming much branched, the small second- 
ary panicles with cleistogamous, fruitful spikelets. 
Subgenus Dichanthelix'M, p. 512. 
Basal leaves pimilar to culm leaves, not forming a winter rosette; spikelets all 
fertile. 
Plants annual. 
Spikeleta piano-gibbous, the second glume swollen, bristly and burlike 
at maturity; first glume nearly as long as the minute spikelet. 
116. P. hirtum. 
Spikeleta not piano-gibbous nor bur like. 
Panicles consisting of several more or less secund spikelike racemes; 
fruit transversely rugose; glumes and sterile lemma usually 
reticulate-veined  See Fasciculata, p. 467. 
Panicles more or less diffuse (small and narrow in P. vas&janum). 
Blades ovate-lanceolate or elliptical, one-fifth to one-fourth 
as broad as long; plants low; spikeleta minute, not over 
1.5 mm. long. 
Spikelets pyriform, attenuate at base, glabrous. 
71. P. pyrularium. 
Spikelets elliptic, pubescent  67. P. trichoide»r 
1
 Fl. Brit. W. Ind 2
 Symb. Ant 
HITQHOOOK AND CHASE—TROPICAL NORTH AMERICAN PANICXJM. 461 
Blades linear, many times as long as broad; spikelets 1.7 mm. 
or more long. 
First glume not over one-fourth the length of the spikelet, 
truncate or triangular-tipped. 
See Dichotomifloha, p. 473. 
First glume usually as much as half the length of the spike- 
let, acute or acuminate  See Catillaria, p. 476. 
Plants perennial. 
Spikelets short-pediceled along one side of the panicle branches, forming 
more or lew spikelike racemes. 
First glume nearly as long as the obtuse spikelets; plants with long wiry 
stolons with woolly nodes 111. P. obtusum. 
First glume much shorter than the spikelet. 
Spikelets more or less hispid. 
Sterile lemma glandless; second glume inflated-gibbous. 
73, P. ineptum. 
Sterile lemma bearing a pair of crateriform glands. - 
See Stolonifera, p. 500. 
Spikelets glabrous. 
Blades lanceolate or ovate-lanceolate; glumes strongly carinate. 
See Stolonifera, p. 500. 
Blades linear, often elongated; glumes not carinate or but slightly so. 
Fruit transversely rugose  See Geminata, p. 466. 
Fruit not rugose. 
Second glume shorter than the spikelet; fertile lemma subin- 
durate, inrolled only at base, scabrous at the acute apex. 
112. P. stagnatUe. 
Second glume as long as the spikelet; fertile lemma indurate, 
inrolled to the summit See Laxa, p. 492. 
Spikelets in open or sometimes in contracted or congested panicles, but 
not in 1-sided spikelike racemes. 
Fruit transversely rugose See Maxima, p. 433. 
Fruit not transversely rugose (minutely papillose-roughened in P. 
millegrana). 
Culms more or less succulent; plants glabrous or nearly so throughout, 
aquatic or eubaquatic. 
First glume not over one-fifth the length of the spikelet. 
See Dichotomifloba, p. 473. 
First glume one-third to half the length of the spikelet. 
Fruit indurate, the margins of the lemma inrolled; culms erect; 
panicle contracted 48. P. eonden&um. 
Fruit subindurate, the margins of the lemma inrolled only at 
base; culms decumbent at base; panicles open. 
Spikelets 1.8 mm. long; second glume shorter than the fruit; 
secondary panicle branches secund 112. P. stagnatUe. 
Spikelets 2.5 mm. long; second glume exceeding the fruit; 
panicle branches not secund  113. P. grande. 
Culms not succulent. 
First glume hyaline, inconspicuous; spikelets acuminate; lower 
blades with long petiole-like bases  114. P. tuerckheimii. 
First glume evident. 
First glume very small, not over one-fourth the length of the 
small obovate, blunt, glabrous spikelets. 
See Pabviglumia, p. 502. 
462 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM 
First glume usually more than one-third the length of the spikelet, 
if shorter the spikelets not small and blunt. 
Sterile palea enlarged and indurate at maturity, expanding the 
spikelet; blades scarcely wider than their sheaths. 
See Lax a, p. 492. 
Sterile palea if present not enlarged and indurate. 
First glume short, blunt; spikelets pointed; base of culm 
usually decumbent and rooting. 
See Dichotomwlora, p. 473. 
First glume usually more than one-third the length of the 
spikelet. 
Plants forming conspicuous hard creeping scaly roots tocks. 
See Vibgata, p. 486. 
Plants not forming creeping scaly rootstocks. 
Fruit crested at the apex; spikelets 5.5 to 6 mm. long. 
115. P. zizanioides. 
Fruit not crested. 
Panicles narrow and few-flowered; culms erect and 
wiry; blades drying involute.. See Tenera, p. 490. 
Panicles open or contracted, many-flowered. 
Panicles 40 to 60 cm. long, the numerous elongated 
branches in verticils 77. P. megi&ton. 
Panicles mostly much lesa than 40 cm. long; branches 
not verticillate. 
Spikelets ehort-pediceled along the nearly simple 
panicle branches 48. P, condenwm. 
Spikelets long-pediceled; panicle open at maturity. 
Primary panicles open, the secondary reduced, 
narrow, partly inclosed in the sheaths. 
First glume three-fourths as long as the ellipti- 
cal spikelets; fruit apiculate. 
See Cokdovensia, p. 525. 
First glume one-third as long as the pyriform 
spikelets; fruit not apiculate. 
104. P. nodalum. 
Primary and secondary panicles alike, or the 
secondary wanting. 
First glume not pointed, two-thirds the length 
of the spikelet or more; spikelets blunt. 
Panicles not over 6 cm. long; plants some- 
what glaucous, relatively small. 
See Parvifolia, p. 506. 
Panicles 10 to 20 cm. long, very diffuse; 
plants tall, not glaucous. 
Spikelets viscid, 3 mm. long. 
75. P. glutinomm. 
Spikelets not viscid, 2 to 2.3 mm. long. 
74. P. mUUgrana. 
First glume pointed, usually less than two- 
thirds as long as the pointed spikelets. 
Spikelets more or less pubescent. 
Culms slender, straggling; spikelets not 
turgid; glumes and sterile lemma hir- 
sute along the margins. 
72. P. haenkeanum. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 463 
Culms stout, erect or nearly go; spikelets 
turgid, sparsely hirsute. 
76. P. rudgei. 
Spikelets glabrous. 
Culms straggling; spikelets minute. 
68. P. trichanthum. 
Culms erect or stiffly ascending; blades 
linear, usually elongated. 
Sheaths glabrous; culms 1.5 to 2 meters 
high  40. P. icknatrihoidet. 
Sheaths hirsute, or if glabrous the culms 
less than 1 meter high. 
See Diffusa, p. 480. 
ANNOTATED LIST OF THE SPECIES. 
Submenus PAXTROCHAETIUIC Hitchc. & Chase. 
Blades less than 10 cm. long, not narrowed toward the base; 
spikelets about 2.5 mm. long  4. P. ramisetum. 
Blades elongated, usually more than 15 cm. long, narrowed 
toward the base. 
First glume rounded or truncate; second glume about as 
long as fruit  3. P. ckapmani. 
First glume acute; second glume about two-thirds as long as 
fruit. 
Spikelets 1.5 mm. long; blades involute  1. P. distantijlorum,. 
Spikelets 2 mm. long; blades flat  2. P. utouxmaeum. 
1. Panicum distantijlorum A. Rich, 
Panicum distantijlorum, A. Rich, in Sagra, Hist. Cuba 11:304.1850; Contr. U. 8. 
Nat. Herb. 10: 23.1910. 
DISTRIBUTION. 
limestone hills at low altitudes, Bahamas to Cuba and Haiti; Curasao. The type 
specimen from Cuba. 
m 13 I 
i Vv 
K / / V 1 
. T" i fVJ \J 
/ f  ^ v. • 
n \ V C -\L^r\ ] , f , ^ i / [jf i 
- ^ t 
f ^ V n 4 {A 
v .r^ 
Fig. 11.—Distribution of P. iutaniiflorum. 
Santo Domingo: Naranjo, FuerU» 1284. Los Ch&rcos, Fuertes 1427. 
Curasao: BrUton & Shafer 3088. Bonaire, Suringar in 1885. (E. U. Herb,)1 
Bahamas: Inagua, Hitchcock in 
1890, Nash & Taylor 893 (both 
Field Mus. Herb.). 
Cuba: Play a de Cojimar, Hitchcock 
144, Le6n 912. Near Habana, 
Le6n305b,567,2382. Santiago, 
Le6n917. SilladeCayo, Shafer 
2512. Guanoroca, Wright 284. 
Han6bana, Wright 285. Playa 
de Marianao, Le6n in 1909. 
Without locality, Wright 3452. 
1
 Krug & Urban Herbarium 
* 
464 CONTRIBUTIONS FROM THE NATIONAL HEBBARIUM. 
2. Panicum utowanaeum Scribn 
Panicum utowanaeum Scribn. in Millsp. Field Mus. Bot. 2: 25.1900; Contr. TJ. S. 
Nat. Herb. 16: 24.1910. 
DISTRIBUTION. 
Open rocky soil, mostly near the coast, Cuba, Porto Rico, Guadeloupe, and Vene- 
zuela. The type specimen from Porto Rico. 
Cuba: Triscomia, Hitchcock 141, Tracy 
9089. Cayo Guajaba, Shafer 2830. 
Playa Marianao, Wilson 9497. 
Eastern Cuba, Wright 3452. 
Santo Domingo: Azua, Rose, Fitch 
& Russell 3891. 
Porto Rico: Guanica, Mills paugh PI. 
XJtow. 702, Sintenis 3365, 3416, 
3463, Chase 6520. Santa Rita, Chase 
6535, LagunaGuanica, CAose6533. 
Mona, Hess 431, 455, 457. 
FiO. 12.—Distribution of P. utowanaeuni. 
Leeward Islands: Guadeloupe, Dtiss 3177. 
Venezuela: Near Puerto Caballo, Pittier 6434. 
3. Panicum chapmani Vaecv. 
Panicurn chapmani Vaaey, Bull. Tor- 
rey Club 11: 61. 1884; Contr. U. S, 
Nat. Herb. IB: 24.1910. 
distribution. 
Coral sand and shell mounds, south- 
ern Florida and the Bahamas. The 
type specimen from Florida. 
Bahamas: New Providence, Brit- 
ton de Brace 401. Rose la- 
land, Britton & Milhpaugh 
Fig. 13.—Distribution of P. chapmani. 
2137. Great Exuma, Britton & Mills paugh 3076. (All in Field 
Herb.) 
4. Panicum ramisetum Scribn. 
Mus. 
Panicum ramisetum Scribn. TJ. S. Dept. 
Agr. Div. Agrost. Circ. 27: 9.1900; Contr. 
TJ. 8. Nat. Herb. 16: 25. 1910. 
distribution. 
Fit}. 14.—Distribution of P. ramitetttm. 
Sandy plains and prairies, southern Texas 
and northern Mexico. The type specimen 
from Texas. 
Coahuila: Near Dfaz,1 Pringle 8323. 
1
 The "Gu£a Postal de la Reptiblica Mexicans " has been followed in the accent- 
ing of Mexican geographical names. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 465 
TRUE PANICUM. 
GEM IN ATA. 
Nodes bearded  7. P. barbinode. 
Nodes glabrous. 
Spikelets 3 mm. long; glumes and sterile lemma papery 6. P. paludivagum. 
Spikelets not over 2.4 mm. long; glumes and sterile lemma 
not papery  5. P. geminatum. 
5. P&nicum geminatum Forsk. 
Panicum geminatum Forsk. Fl. Aegypt, Arab. 18. 1775; Contr. U. S. Nat. Herb. 
16: 30.1910. 
This species is described by Presl1 under the name Panicum brizoides L. (as well 
as under P. brizaeforme Presl). The Haenke specimen from Mexico is in the herb- 
arium of the National Museum at Prague. 
t 
DISTRIBUTION. 
Moist ground, ditches and swamps, mostly near the coast, tropical regions of both 
hemispheres, in America extending north into southern Florida and Texas. Origi- 
nally described from Kosetta, Egypt. 
Lower California : El Taste, Brandegee in 1893. San Jos6 del Cabo, Brandegee 
14, 36. 
Sonoba : Yaqui River, Palmer 15 in 1869. Guaymas, Palmer 690 in 1887. Sierra 
de Alamos, Hose, Standley & Russell 12996. 
Sinaloa: Mazatlin, Rose, Standley Russell 14109. 
Tamaulifas: Tampico, Hitchcock 5796, Palmer 259 in 1910, 
Colima : Manzanillo, Hitchcock 7032. 
Veracruz; Veracruz, Hitchcock 6586. 
Guerrero: Acapulco, Palmer 289 in 1894. 
YucatAn: Izamal, Gaumer 1027. 
Guatemala: San Jos£, Kelltrman 
6250. 
Salvador: Lake llopango,' Hitch- 
cock 8920. 
Cobta Rica: Bebedero, Jimbiez 
734. 
PANAMA: Corozal, Pittier 2168, 
2170. Pedro Miguel, Hitch- 
cock 7956, Pittier 2509. Em- 
pire, Pittier 3714. Gatun, 
Hitchcock 7980. 
Bahamas: Nassau, Curtiss 175. Fortune Island, Eggers 3992 (K. U. Herb.). 
Cuba: Qabana, Le6n 769,918. CorrientesBay, Britton <fc Cowell9931. Santiago 
de las Vegas, Hitchcock 143. Guanab&coa, Le6n 920. Guant&namo, Britton 
2281. Sancti Spiritus, Sergius 2789. Bataband, Hitchcock 142. Handbana, 
Wright 761. Palma Sola, Wright in 1860. 
Jamaica: Gordon Town, Hart 806. Ferry River, Harris 11352. Appleton, 
Hitchcock 9653. Palisadoes near Kingston, Hitchcock 9752. Black River, 
Hitchcock 9579. Savanna-Ia-Mar, Hitchcock 9877. 
1
 Preal, Rel. Haenk. 1: 302.1830. 
82472°—15 2 
Fio. 15,—Distribution of P. geminatum. 
466 CONTRIBUTIONS FBOM THE NATIONAL HERBARIUM. 
Haiti: La Coupe, Buck 975. Port-au-Prince, Picarda 1430. (Both in K. U. 
Herb.) 
Porto Rico: Guanica, Chase 6527, 6531, Sintenis 3367. Mayaguez, Chase 6288, 
6318. Boqueron, Chase 6511. Coamo Springs, Chase 6552. 
Danish West Indies: St. Croix, Richecher 212. St. Thomas, Eggers in 1882. 
Leeward Islands: Antigua, Wullschlaegel 614. Guadeloupe, Dusa 2690, 3584, 
L 'Herminier. 
Windward Islands: Martinique, Duss 1293. 
Trinidad: Icacos, Hitchcock 10158. 
Curacao: Aruba, Suringar in 1885. 
Venezuela: P&paro, Pittier 6347, 
Colombia : Santa Marta, Smith 151. 
6. Panicum paludivagum Hitchc. & Chase. 
Panicum paludivagum Hitchc. & Chase, Contr. U. S. Nat. Herb. 15: 32. 1910. 
distribution. 
Freeh-water lakes and rivers, the 
base submerged, Florida and Texas, 
Mexico, Central America to Uru- 
guay. The type specimen from Flor- 
ida. 
Jalisco: Guadalajara, Palmer 429 
in 1886, Hitchcock 7294. 
Fig. 15.—Distribution of P. paludivagum. Qrozco, Hitchcock 7374. 
Michoacan: Lake Patzcuaro, Pringle 3336. Zamora Valley, Pringle 9556. 
Morelia, Arsbie in 1909. 
Guatemala: Amatitl&n, Kellerman 6253, 6254, Pittier 101, Tiirckheirri 8790. 
7. Panicum barbinode Tiin. 
Panicum barbinode Trin. M6m. Acad. St, P6tersb. VI. Sci. Nat. 1: 25C. 1S34; Contr. 
U. S. Nat. Herb. 16: 33.1910. 
Commercially known as " Par& grass"; called "malajilla" in Porto Rico. 
distribution. 
Cultivated and waste ground, especially in moist places, tropical America, extend- 
ing into southern Florida and Texas; introduced in the warmer parts of the Old World. 
The type specimen from Bahia, Brazil. 
Lower California : San Jos6 del Cabo, Brandegee 46. La Paz, Palmer 131 in 1890. 
Co LIMA: Paso del Rio, Emrick 8. Manzanilla, Palmer 1078. 
Mexico: Cuantla, Holway 3045. 
Veracruz: Veracruz, Amer. Gr. Nat. Herb. 4.1 
Oaxaca: Tomellfn, Hitchcock 6227. 
YucatAn: M6rida, Collins 27. 
Guatemala: Mazatenango, Maxon & Hay 3476. Cubilquitz, Tiirckheim 7799, 
8617. Escuintla, Hitchcock 9007. 
Salvador: Without locality, Renson 214. 
Nicaragua: Jinotepe, Hitchcock 8701. Chinandega, Baker 2053. 
Costarica: T^rraba, Pittier 412, 4090. Rfo Tufn, Tonduz 11393. Puntarenas, 
Hitchcock 8545. Taboga, Guanacaste, Jiminez 722. Turrialba, Tonduz 8401. 
1
 American Grasses, exsiccatse distributed from the IT. S. National Herbarium, 
Smithsonian Institution. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 467 
PanamA: El Boquete, Hitchcock 8266. Gatun Lake, Pittier 2669, Hitchcock 8031}. 
Pedro Miguel, Hitchcock 7931. Chepo, Pittier 4734. Culebra, Pittier 4807, 
Bahamas: Nassau, Cwrtiss 115. 
* *i 
Cuba: Near Habana, Le6n 568. Camaguey, Shafer 2881. Bomelie, Eggers 4870. 
Cienfuegos, Pringle 26. Almendares River, Le6n 283. Without locality. 
Wright 1545, 
Jamaica: Hope Gardens, Harris 11254, 10930. Mount Hybla, Harris 11565. 
Newcastle, Hitchcock 9339. Black River, Hitchcock 9580. Montego Bay, 
Hitchcock 9693. 
Porto Rico: Bayamon, Hioravi 
360, Millspaugh 324, Heller 
100. Aibonito, GoU 613. 
Playa de Fajardo, Chase 6657. 
Guanica, Chase 6528, Mills- 
paugh 727. Yauco, Heller 
6293. Los Mameyes, Eggers 
1328. Vieques, Chase 6679. 
Mona, Hess 450. 
Danish West Indies: St. Croix, 
Richsecker 300. 
Leeward Islands: Guadeloupe, Duss 2689, L'Herminier. Dominica, Eggers 
1030 (K. U. Herb.). 
Windward Islands: Martinique, Duss 539. Grenada, Buckmire 1423, Broad- 
way in 1904 and in 1905. St. Vincent, Eggers 6834 (K, U. Herb.). Barbar 
dos, Eggers 7079 (K. U. Herb.). 
Trinidad: Port of Spain, Hitchcock 9941, 10044. 
Tobago: Scarborough, Hitchcock 10204, 
Curasao: Santa Cruz, Britton Shafer 3012. 
Colombia: Cartagena, Hitchcock 9901. Santa Marta, Smith 211. Without 
locality, Lehmann 5744, 
FASCICtTLATA. 
Spikelets 5 to 6 mm. long  13. P. texanum. 
Spikelets 2 to 4 mm. long. 
Spikelets strongly reticulate-veined, 2 to 3 mm. long; glabrous. 
Panicle branches long and spreading; blades pubescent or 
glabrous    9. P. fasciculatum. 
Panicle branches short, appressed; blades narrow, pubes- 
cent.  9a. P. fasciculatum 
reticulatum. 
Spikelets scarcely reticulate-veined or only near the apex. 
Spikelets not over 2 mm. long, glabrous  8. P. reptans. 
Spikelets over 3 mm. long, pubescent, 
Rachis scabrous but not bristly  11. P. adspersum. 
Rachis pilose with bristly hairs. 
Plant more or less velvety; sheaths not papillose. 10. P. rmlle. 
Plant not velvety; sheaths papillose.. - -  12. P. arizonicum. 
8. Panicum reptans L. 
Panicum reptans L. Syst. Nat. ed. 10. 2: 870. 1759; Contr. U. S. Nat. Herb. 15: 36. 
1910 
DISTRIBUTION. 
Open ground at low altitudes especially near the coast, frequently a weed in waste 
places and cultivated soil, Gulf Coast of the United States and Atlantic slope of 
Fro. 17.—Distribution of P. barbinode. 
468 CONTRIBUTIONS FBOM THE NATIONAL HEBBABIUM. 
Mexico through the West Indies to northern South America; also introduced in the 
warm regions of the Eastern Hemisphere. The type specimen from Jamaica. 
Tamaulipas: Tampico, Palmer 153 in 1910. 
Veracruz: Banos del Carrizal, Purpus 6213. 
Oaxaca: Tomellln, Hitchcock 6231. Cuicatl&n, Nelson 1622. 
Cuba: Habana, Curtiss 691, Hitchcock 146, Ledn 3474. Habana Yedado, Le6n 
276, 292, 297, 566. Habana Vibora, Le6n 906, 910c. El Cafio, Le6n 1980. 
Baracoa, Shafer 3955, Palmer, Pollard & Palmer 19. Buenaventura, Wilson 
9243, 9399. Rfo Almendares, Wilson 9187. Cojimar, Britton, Earle & 
Gager 6295. Valley of Bacuranao River, Le6n & Wilson 2869. Oienfuegos, 
Pringle 73. Herradura, Tracy 9103. San Antonio, Hitchcock 145. Madruga, 
Cmtiss 536. Arroyo A polo, Le6n 576. La Magdaleua, Earle & Baker 2455. 
Santiago de Cuba, Le6n 910. Guayabal, Le6n 910b. Guantdnamo Bay, 
Britton 1933, 2245. Romelie, Jiggers 5346, Without locality, Wright 762, 
763, 3857. 
Jamaica: Prospect, Harris 11539. Black River, Hitchcock 9631. Montego Bay, 
Hitchcock 9689. Hope, Harris 
6845. Gordon Town, Hart 838. 
Haiti: Without locality, Picarda 1246 
(K. U. Herb.). 
Santo Domingo: Rinc6n, Fuertes 
1377,1417. San Pedro de Ma- 
coris, Rose, Fitch & Russell 4171. 
Porto Rico: Guanica, Chase 6515, 
Millspaugh 726, Sirdenis 3368. 
Bayamon, Chase 6395. Joyuda, 
Chase 6305. Ponce, Heller 497, 
Chase 6326. Cabo Rojo, $intenis 847. Coamo Springs, Goll 662. Fajardo, 
Chase 6664. Vieques, Chase 6700. 
Danish West Indies: St. Croix, Ricksecher 77. St. Thomas, Eggers 293. 
Leeward Islands: Antigua, Wullschlaegel 617. Guadeloupe, Duss 3529. 
St. Martin, Boldingh 3383 (K. U. Herb.). 
Windward Islands: Martinique, Puss 1290. Grenada, Broadway 721. Bequia, 
Smith & Smith 19, 370 (K. U. Herb.). Barbados, Eggers 7081 (K. U. Herb.). 
Bot. Sta. Herb, 538. 
Tobago: Scarborough, Hitchcock 10212. 
Curasao: Santa Cruz, Britton & Shafer 2987, 3002. 
Venezuela: El Valle, Island of Margarita, Miller <& Johnston 171. 
Colombia: Santa Marta, Smith 173, 2753. Cartagena, Hitchcock 9907. Puerto de 
Colombia, Hitchcock 9936. 
9. Panicum fasciculatum Swartz. 
Panicum fasciculatum Swartz, Prodr. Veg. Ind. Occ. 22. 1788; Contr. U. S. Nat. 
Herb. IS: 38. 1910. 
Panicum chartaginense Swartz, Prodr. Veg. Ind. Occ. 22. 1788; Contr. U. S. Nat. 
Ilerb. 15: 40. 1910. "America meridional ia chartagena." A reconsideration of 
Swartz'a type in the light of recent collections from the region whence it came leads 
to the conclusion that it is referable to Panicum fasciculatum rather than to the narrow- 
panicled form confined to the semiarid southwestern United States and Mexico, 
which was described by Torrey as P. reticulatum. 
Fig. 18.—Distribution of 1'. replant. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN FANICUM. 469 
DISTRIBUTION. 
Moist open ground, often a weed in fields and waste places, southern Florida and 
Texas, through Mexico and the West Indies to central South America. The type 
specimen from Jamaica. 
Sonora: Hermosillo, Hitchcock 3598. Imeria to Santa Ana, Griffiths 6857. 
Alamos, Palmer 694 in 1890. Guaymae, Palmer 158 and 207 in 1887. 
Chihuahua: Norogachi, Palmer la in 1885. 
Sinaloa: Rosario, Rose 1834,1884. Topolobampo, Palmer 241 in 1897. Culiac&n, 
Palmer 1557 in 1891. 
San Luis Potosi: Rasc6n, Pvrpua 5418. C&rdenas, Hitchcock 5728. 
Quer^taro: Quer^taro, Hitchcock 5838. 
Couma: Jala, Hitchcock 7011, Alzada, Hitchcock 7097, 7106. Without locality, 
Palmer 19 in 1897. 
Veracruz: Misantla, Purpus 5979. C6rdoba, Find in 1893, Hitchcock 6448. 
Bafios del Carrizal, Purpus 6218. Santa Maria Tlatetla, lAebmann 279. 
Oaxaca: Tomellfn, Rose, Painter <£r Rose 10077, Hitchcock 6214. Totalcingo, 
Liebmann 277. 
Chiapas: Vicinity of Topana, Nelson 2874. Between San Ricardo and Oco- 
zucuantla, Nelson 2958. 
YucatIn: M£rida, Schott 384. __—, , , -—■ ■ ■ . . ^ 
British Honduras: Manatee La- 
goon, Peck 317 (Gray Herb.). 
Guatemala: Nenton, Seler 2704. 
Gual&n, J)eam 6267. Finca 
Trece Aguae, Goll 81. Chic- 
arao, Heyde & Lux 6404. Es- 
cuintla, Smith 2233. 
Honduras: San Pedro Sula, 
Ttneme 195, 5584. Highland Fig. 19.—Distribution of P. fascicuUUum. 
Creek, Wilson 644. 
Salvador: La Unitfn, Hitchcock 8778. 
Nicaragua: Jinotepe, Hitchcock 8670. Masaya, Hitchcock 8663. Without 
locality, Flint in 1868. 
Costa Rica: Puerto Viejo, Biolley 7471. Nueetro Amo, Alajuela, Jimhvez 525. 
Hacienda la Estrella, Tonduz 190. Puntarenas, Hitchcock 8552. Puerto de 
Rio Grande, Pittier 2035. Nicoya, Tonduz 13749. Matina, Pittier 9727. 
Panama: Laguna de Chiriqui, Hart 78. Taboga Island, Pittier 3626, Hitchcock 
8076, 8079. Gatun, Hitchcock 9185. Toro Point, Hitchcock 8057. Culebra, 
Hitchcock 7908, 7932, Pittier 2117. David, Hitchcock 8358. Porto Bello, 
Pittier 2478. Puerto Obaldfa, Pittier 4375. Empire, Pittier 3718. Las 
Case ad as, Pittier 3743. Chepo, Pittier 4695. La Sabana de Panama, Gervais 
165. 
Cuba: Hoyo Colorado, Le6n 573. Herradura, Tracy 9091. Cienfuegos, Pringle 
74, 124. San Antonio, Eggers 4875. Santiago de Cuba, Le6n 813. Banao 
Hills, Le6n 3966. Sancti Spiritus, Le6n 916. Sierra de Anafe, Le6n 1979. 
Santiago de las Vegas, Wilson 593. Without locality, Wright 754. 
Jamaica: Gordon Town, Hart 785, 840, Harris 11345, 11400. Bath, Maxon 2361. 
Hope Gardens, Maxon 1659, Harris 11252, 11255, Hitchcock 9312, Amer. Or. 
Nat, Herb. 5. Flamstead, Harris 11470. Ferry River, Harris 11324, 
Hitchcock 9747. Montego Bay, Hitchcock 9682. Savanna-la-Mar, Hitchcock 
9870. Above Constant Spring, Hitchcock 9259. Ewarton to Linstead, 
Hitchcock 9411. Without locality, Lloyd 1116, Wilson 322. 
470 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
Haiti: Labrante, Buck 48. Without locality, Picarda 409, Christ 2144, (All in 
K. U. Herb.). 
Santo Domingo ; San Pedro de Macoris, Rose, Fitch & Russell 4440, 4174. 
Porto Rico: Aibonito, Underwood <fc Griggs 462. Monte Goyo, Sintenis 1901. 
Guanica, Sintenis 3647, Chase 6523. Rio Piedras, Barrett 63, Heller 135. 
Ponce, Seller 6226, 6302. Caguas, Goll 385. Guayama Road, Goll 588. 
Aguadilia, Heller 4528. Maricao, Chase 6197. Coamo Springe, Chase 6542. 
Bayamon, Chase 6376, Hioram 316. Mayaguez, Chase 6263. Sierra Luquillo, 
Hioram 357. Culebra, Britton & Wheeler 137. Vieques, Chase 6693. With- 
out locality, Fisher 59, Underwood & Griggs 824. 
Danish West Indies: St. Thomas, Eggers in 1880 and in 1882. St. Croix, Rick- 
seeker 317. 
Leeward Islands: Guadeloupe, Duss 2691, L*Herminier. Montserrat, Skafer 
584, St. Kitts, Brit ton <£r Cowell 269 (K. U. Herb.). Dominica, Duss 
(K. U. Herb.). 
Windward Islands: Martinique, Duss 537, 538, Hahn in 1867-70. Grenada, 
Broadway in 1905. St, Vincent, Smith <fc Smith 565 (K. U. Herb.). 
Trinidad: St. Joseph, Hitchcock 10014. Port of Spain, Hitchcock 9976. Without 
locality, Bot. Gard. Herb.1 2283, 3192. 
Tobago: Mount Marie, Broadway 3977. Rockley Vale, Broadway 4630. Scar- 
borough, Hitchcock 10216. 
Curasao: Santa Catherina, Britton & Shafer 3145. Bonaire, Suringar in 1885 
(K. U. Herb.). 
Venezuela: Paparo, Pittier 6336. Island of Margarita, Miller <fc Johnston 180. 
Colombia: Puerto de Colombia, Hitchcock 9934. Rio Frfo, State of Magdalena, 
Pittier 1610. Cartagena, Hitchcock 9911. Santa Marta, Smith 131, 208. 
9a. Panicum fasciculatum reticulatum (Torr.) Beal, 
Panicum reticulatum Torr. in Marcy, Expl, Red Riv. 299.1852. 
Panicum fasciculatum reticulatum Beal, Grasses N. Amer. 2:117.1896. 
For further treatment see Oontr. U. S. Nat. Herb. 15:40.1910 (where this is included 
under P. fasciculatum chartaginense (Swartz) Doell); see also discussion under P. 
fasciculatum. 
distribution. 
Prairies, fields, and waste grounds, Texas to Arizona and southern Mexico. The 
type specimen from Texas. 
Sonora: Guaymas, Hitchcock 3561. 
Chihuahua: Chihuahua, Pringle 379, 380, 
Coahuila: Sabinas, Nelson 6818. 
Nuevo Le6n: Monterrey, Hitchcock 5535, 
5541. 
Tamaulipas: Victoria, Palmer 412 in 1907. 
Tepic: Acaponeta, Rose, Standley de Russell 
14265. 
San Luis PoToaf: San Dieguito, Palmer 152 
in 1904. 
Colima: Manzanillo, Hitchcock 7029. 
Puebla: Tehuacdn, Hitchcock 6050. 
Oaxaca: Ignacia Mejfa, Hitchcock 6115. 
Tomellfn, Hitchcock 6195. 
Fio. 20.—Distribution of P. fasciculatum 
reticulatum. 
'Herbarium of the Botanical Garden, Trinidad. Many specimens are without 
data other than the number. 
HITCHCOCK AND CHASE—TROPICAL NOBTH AMERICAN PANICUM. 471 
1 
10. Panicum molle Swartz. 
Panicum rootle Swartz, Prodr. Veg. Ind. Occ. 22,1788; Contr. U. S. Nat. Herb. 15: 
42.1910. 
DISTRIBUTION. 
Open ground, often a weed in fields, Cuba, Jamaica, Mexico, and Central America 
to Argentina. The type from the West Indies, probably Jamaica. Since at the time 
the revision of Panicum 1 was prepared we had no specimens of the species from the 
West Indies, we doubted the accuracy of the locality given by Swartz. This species 
proves, however, to be not uncommon in the vicinity of Kingston, Jamaica. 
Sinaloa: Lodiego, Palmer 1660 in 1891. 
Colima: Manzanillo, Amer. Gr. Nat. Herb. 7. Alzada, Hitchcock 7107. Colima, 
Palmer 149 in 1897. 
Veracruz: Without locality, Galeotti 5710. 
Guerrero: Balsas, Hitchcock 6813. 
Oaxaca: TomelUn, Hitchcock 6205, 6229. 
Guatemala: Agua Caliente, Beam 6143. 
Salvador: San Salvador, Hitch- 
cock 8860. 
Costa Rica: Puntarenas, Hitch- 
cock 8585. 
Panama: Point Cham6, Amer. Gr. 
Nat. Herb. 8. Taboga Island, 
Hitchcock 8067. 
Cuba: Without locality, Reed. 
Jamaica: Kingston, Alexander in 
1849, Hitchcock in 1890. Gor- 
don Town, Harris 11504, ffltchr 
cock 9323. St. Andrew, Harris 
11483. HealthshireHills, Hitchcock 9758. Hope Gardens, Amer.Gr.Nat.Herb.Q. 
Curasao : Santa Cruz, Britton & Sha/er 2986, 3004. Aruba, Suringar in 1885. 
(K. U. Herb.) 
Colombia: Santa Marta, Smith 209. Cartagena, Hitchcock 9922. Puerto de 
Colombia, Hitchcock 9940. 
11. Panicum adspersum Trio. 
Pcmicum adspersum Trin. Gram. Pan. 146. 1826; Contr. U. S. Nat. Herb. 10: 43. 
1910. 
DISTRIBUTION. 
Moist open ground, Florida and the 
West Indies. The type specimen from 
Santo Domingo. 
BAHAMAS: Nassau, Curtiss 113. 
Anguilla Isles, Wilson 8066. 
Cuba: Habana Vedado, Le6n 291, 
1978. Habana, Le6n 570, Cur- 
tiss 748. Santiago de las Ve- 
gas, Baker de Wilson 512, Tracy 
9109, Hitchcock 147,148. Tris- 
coroia, Hitchcock 159. Cabanas, Palmer & Riley 746, 771. Herraduia, 
Tracy 9102. Sancti Spiiitus, Le&t% 925. Guinea, Le&n 924. Zaza del Sur, 
Sergius 2776. Without locality, Wright 3869. 
Fig. 21.—Distribution of P. mollc. 
Fig. 22.—Distribution of P. adspersum. 
1
 Contr. U. 8. Nat. Herb. 15:42.1910. 
, 472 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM 
Jamaica: Black River, Hitchcock 9630, Hope Gardens, Amer. Or. Nat. Herb. 10, 
11, Harris 11247. Kingston, Hitchcock 9750. Alligator Pond, Hitchcock 9830. 
Gordon Town, Harris 11343,11512. Yallahs Valley, Harris 11498. 
Haiti: Gonaives, Buch 23. 
Santo Domingo: Barahona, Fuertes 43. 
Porto Rico: Between Goamo and Aibonito, Sintenis 1957, Chase 6333. Joyuda, 
Chase 6302. Guanica, Chase 6514. Mayaguez, Chase 6286. Bayamon, 
Hwram 315. Vieques, Chase 6686. 
Danish West Indies: St. Croix, Bggers in 1876, Ricksecler 66, 384. 
Leeward Islands: Antigua, Wullschlaegel 615, 616. Guadeloupe, Duss 3180. 
Saba, Boldingh 1583 (K. U. Herb.). St. Eustatius, Suringar in 1885 (K. U. 
Herb.). St, Bartholomew, Goes (K. U. Herb.). 
Windward Islands: Dim 537b (K. U. Herb.). 
18. Panicum arizonicum Scribn. & Merr. 
Panicum arizonicuin Scribn. & Merr. U. S. Dept. Agr. Div. Agrost. Circ. 32:2.1901; 
Oontr. U. S. Nat. Herb. 16:44.1910. 
distribution. 
Open eandy or stony ground, southwestern United States and northern Mexico. 
The type specimen from Arizona. 
Lower California : Arroyo San Lazaro, Brandegee in 1902. San JW del Oabo, 
Brandegee in 1890. 
Sonora: Nogales, Griffiths 6747, 6759, 
Hitchcock 3637. Lokuka Ranch to Al- 
tar, Griffiths 6891. Guaymas, Palmer 
159 and 208 in 1887. Hitchcock 3562. 
Hermosillo, Hitchcock 3542. 
Chihuahua: Norogachi, Palmer IB in 
1885. Chihuahua, Pringle 487. 
Sinaloa: Topolobampo, Painter 250 in 
1897. 
Durango: Without locality, Rose 2280. 
Guerrero: Rfo Balsas, Orcutt 4195. Bal- 
sas, Hitchcock 6775. 
Oaxaca: Tomellfn, Hitchcock 6192, Amer. Gr. Nat, Herb. 13. 
13. Panicum texanum Buckl. 
Panicum texanum Buckl. Prel. Rep. Geol. 
Agr. Surv. Tex. App. 3.1866; Contr. U. 8. 
Nat. Herb. 15:46.1910. 
distribution. 
Prairies and open ground, often a weed in 
fields, Texas and northern Mexico. The 
type specimen from Texas. 
Nuevo Le6n: Monterrey, Hitchcock 
5540. i'ia. 24.—Distribution of P. texanum. 
Fio. 23,—Distribution of P. arizonicum. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 473 
DIC HOTO M IF LOR A. 
Three species of this group, Panicum aquaticum, P. mcomm, and P. elephantipes, 
are found to be perennial. 
Plants annual. 
Panicles narrow, less than 1 cm. wide  14. P. vaseyanum. 
Panicles open, the branches ascending or spreading. 
Sheaths glabrous  15. P. dichotomijlorum. 
Sheaths papillose-hispid  16. P. bartoivense. 
Plants perennial. 
Fruit not acuminate; panicles rarely over 18 cm. long... 17. P. aquaticum. 
Frtiit acuminate; panicles as much as 40 cm. long, rarely 
less than 20 cm.; culms succulent. 
Spikelets about 3.5 mm. long; fruit abruptly acumi- 
nate  18, P. eucosum. 
Spikelets 4 to 5 mm. long; fruit long-acuminate; 
culms very succulent  19. P. elephantipes. 
14. Panicum vaseyanum Scribn. 
Panicum vaseyanum Scribn.; Beal, Grasses N. Amor. 2:140, 1896; Contr, U. S. 
Nat. Herb. 16: 47.1910. 
DISTRIBUTION. 
Moist open ground, around ponds and de- 
pressions in prairies, northern Mexico. The 
type specimen from Chihuahua. 
Chihuahua: Base of Sierra Madre, 
Pringle 1415. Mifiaca, Hitchcock 
7757. 
Agttascalientes: Aguaecalientes, Bitch- 
cock 7491, Amer. Gr. Not. Herb. 15. 
Jalisco; Guadalajara, Hitchcock 7315. Fig. 25.—Distribution of P. voxtyonum. 
15. Panicum dichotomiflorum Michx. 
Panicum dichotomifiorum Michx. Fl. Bor. Amer. 1:48.1803; Contr. U. S. Nat. Herb. 
15:48.1910. 
The South American Panicum chloroticum Nees, included in the Revision in P. 
dichotomifiorum, is a distinct species, or possibly a subspecies. It is not known from 
north of Brazil. 
DISTRIBUTION*. 
Moist ground along streams and a 
weed in waste places and in cultivated 
soil, Maine to Florida and California 
(rare in the West), and in the West 
Indies and Panama. The type speci- 
men from the Alleghenies. 
Panama: NearChopo, Pittier4614. 
Gatun, Hitchcock 7977. 
Bermuda: Hamilton, MUJspaugh 
126. 
[AMas; Hog Island, Eggers 4405,4512. Watlings, Geogr. Soc. Baltimore 489. 
Nassau, Ourtiss 177. Cat Cay, Brace 3742. 
Fia. 26.—Distribution of P. diehotomifUnwn. 
474 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
Cuba: Herradura, Tracy 9055,9342,Britton Earle 6566. Ariguanabo, Ledn 1975}, 
2774,2775. Santiago de las Vegas, Hitchcock 151. Bataband, Hitchcock 150. 
Leeward Islands: Guadeloupe, Duss 3178. 
16. Panicum bartowense Scribn. & Me nr. 
Panicum bartowense Scribn. & Merr. U. S. Dept. Agr. Div. Agrost. Cire. 35: 3.1901; 
Contr. U. S. Nat. Herb. 15: 52.1910. 
DISTRIBUTION. 
Low ground, often growing in shallow 
water, Florida, Bahamas, Cuba, and 
Jamaica. The type specimen from 
Bartow, Florida. 
Bahamas: Great Bahama, Britton 
& MUlspaugh 2706. North 
Bimini, Brace 3467 (Field Mus. 
Herb.). 
Cuba: Bataban6, Hitchcock 149. 
Without locality, Wright 3860. 
Fig. 27.—Distribution of P. bartowense. 
Jamaica: Black River, Amer. Gr, Nat. Ilerb. 18. 
17. Panicum aquaticum Poir. 
Panicum aquaticum Poir. in Lain. Encycl. Suppl, 4: 281.1816. "Cette plantecrott 
& Porto-Ricco; elle m'a comuniqu^c par Af. Ledru." The type is in the Coeson 
Herbarium. In the Revision1 this species was included in P. dichotomi/forum Michx. 
Subsequent field study has shown it to be a perennial. 
Panicum chloroticum sylvestre Noes; Trin. Gram. Pan. 236. 1826; Oontr. U. S. Nat. 
Herb. 15:48.1910. 
Panicum hygrophilum Salzm.; Steud. Syn. PI. Glum. 1:71.1854; Contr. U. S. Nat, 
Herb. 15:49.1910. 
Panicum proliferumstrictum Griseb. Cat. PI. Cub. 232.1866; Contr. U. S. Nat. Herb. 
15: 49.1910. 
DESCRIPTION. 
Plants perennial, mostly aquatic, glabrous except as noted; culms usually decumbent 
at base, rooting at the nodes, erect branches often arising therefrom, the flowering 
culms erect, simple or sparingly branching, averaging 
lower and more slender than in P. dichotomijlorum; 
sheaths usually overlapping, rarely pilose at the junc- 
tion with the blade; ligule a ring of hairs about 1 mm. 
long; blades commonly elongated, as much as 25 cm. 
long, but sometimes 5 to 10 cm. long and abruptly 
pointed, 5 to 10 mm. wide, linear, acute, occasionally 
sparsely pilose on the upper surface toward the base; 
panicles terminal and axillary, short-exserted or in- 
cluded at base, averaging smaller than those of P. 
dichotomi/lorum, rarely as much as 20 cm, long, less 
freely branching and fewer flowered, the branchlets 
and pedicels smooth or nearly so on the angles; spike- 
lets 3 to 3.4 mm. long, 1 mm. wide, more gradually 
pointed than those of P. dichotomijlorum, the fruit slightly larger and more pointed 
than in that species. 
Herbarium specimens lacking basal parts are difficult to distinguish from P. 
dichotomijlorum. 
Fig. 28.—P. aquaticum. From 
type specimen. 
1
 Contr. U. S. Nat. Herb. 15: 48.1910. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMEBICAN PANICUM, 475 
DISTRIBUTION. 
FiQ. 29.—Distribution of P, aquaiicum. 
Wet places or in shallow water, margins of streams and ponds, mostly at low altitudes, 
Cuba, Porto Rico, and Mexico to Trinidad and Paraguay. 
Couma: Alzada, Hitchcock 7067. 
Guatemala: Puerto BarrioSjiZftcft- 
cock 9147. 
Cuba: Guanabacoa, Le&a 919. 
Ariguanabo, Le6n 1975,1976. 
Habana, Le&n 4155. El Cafio, 
Le&rt 1974. Laguna Jovero, 
Shafer 10912. Without local- 
ity, Wright 3456 in part, 3861. 
Porto Rico: Laguna del Tortu- 
guero,Chase6804. Lake Loisa, 
Chase 6778. Alto de Bandera, Chase 6471. Utuado, Briiton & Cowell 432. 
Leeward Islands: Guadeloupe, Dusa 3178 (K. U. Herb.). 
Trinidad: Pitch Lake, Hitchcock 10099. 
18. Panicum sucosum ep. nov. 
description. 
Plants perennial, glabrous throughout except as noted; culms few to several in a 
tuft, 1 to 1.5 meters long, decumbent at base and rooting at the lower nodes, succulent; 
sheaths rather loose, about as long as the intercedes; ligule a densely ciliate membrane 
about 1 mm. long; blades flat or somewhat involute in drying, 15 to 30 cm. long, 
3 to 9 mm. wide, linear, acuminate, scaberulous on both surfaces, sparsely pilose 
above toward the base; panicle 15 to 30 cm. long, about 
half as wide, the axis and branches somewhat scabrous, 
the branches solitary or in pairs, rather stiffly spreading 
or finally horizontal, as much as 15 cm. long, naked at 
base for 1 or 2 cm., the internodes of the rachis mostly 3 
to 5 cm. long, the branchlets appressed or ascending, 
mostly from the lower side; spikelets short-pediceled, 
somewhat appressed to the rachis, 3.3 to 3.7 mm. long, 
about 1 mm. wide, lanceolate, acuminate, glabrous; first 
glume one-fourth to nearly one-third the length of the 
spikelet, broadly triangular, usually subacute, 3-neryed; 
second glume and sterile lemma subequal, exceeding the 
fruit and pointed beyond it, 7 to 9-nerved; fruit about 
2.5 mm. long, 0.7 mm. wide, rather abruptly acuminate. 
Type in the TJ. S. National Herbarium, no. 693325, col- 
lected in water of a pond at Orozco, near Guadalajara, Jalisco, Mexico, September 29, 
1910, by A. S. Hitchcock (no. 7379). 
This species, which in the Revision 1 was included in P. elephantipes, differs from 
P. dichotomiflorum in being perennial and in having larger spikelets; from P. aquati- 
cum in its larger panicle and more pointed spikelets and fruit; from P. elephardipa 
in having smaller, less succulent culms with narrower blades and in having less 
acuminate fruit. 
FiO.30.—P.swo&um. From 
typo specimen. 
1
 Contr, U. S. Nat. Herb. 15: 54.1910. 
476 CONTRIBUTION'S PROM THE NATIONAL HERBARIUM. 
Fig. 31,—Distribution of P. sucosum. 
DISTRIBUTION* 
In water of ponds at high altitudes, south- 
ern Mexico. 
Jalisco: Qrozco, Hitchcock 7379. 
Mexico; Near Mexico City, Or cut t 
4269, Pringle 6322, 9577, Bourgeau 
529. 
Republic op Mexico : Without locality, 
Schaffner 525 
clcphantiptg. 
16. Panicum elephantipes Neea. 
Panieum ehphantipes Nees, Agrost. Bras. 165. 1829; Oontr. TJ. S. Nat. Herb. 15: 
53. 1910. 
Field study has shown that this species is perennial. 
DISTRIBUTION. 
In ponds and shallow wpter at low altitudes, West Indies and Guatemala, south 
to Argentina. The type specimen from Brazil. 
Guatemala: Near mouth of Bfo Pol.     
ochic, Alta Verapaz, Goll 35A. *'■ ' ^ ^ ^ ^ 
Cuba: San Antonio, Hitchcock 152. 
Almendares River, Le&n 335. 
Jamaica: Middle Quarters, on 
Black River, Amer. Gr. Nat. 
Herb. 19. Savanna-la-Mar, 
Hitchcock 9878. 
Porto Rico: Between Catafio and 
Bayamon, Chase 6407. Dor- 
ado, Chase 6415. Fajardo, Sin- 
tenis 938. 
CAPILLARIA. 
Panicles more or leas drooping  25. P. sonorum. 
Panicles erect. 
Inflorescence elongated, composed of several approximate, 
implicate panicles 27. P. cayennense. 
Inflorescence not composed of approximate nor implicate 
panicles. 
Panicles more than half the length of the entire plant.. 20. P. capfflare. 
Panicles not more than one-third the entire height of 
the plant. 
First glume about one-third the length of the 
spikelets, subacute or blunt 24. P. stramineum. 
First glume usually more than half the length of 
the spikelet, acuminate. 
Spikelets 4.5 to 6 mm. long. 
Spikelets 6 mm. long, scattered   26. P. parcum. 
Spikelets scarcely over 5 mm. long, 
approximate  23. P. decolorans. 
Spikelets not over 4 mm. long. 
First glume more than three-fourths the 
length of the spikelet; spikelets 4 
mm. long 22. P. pampinofum. 
First glume half to two-thirds the length 
of the spikelet; spikelets not over 3.3 
mm. long 21. P. hirticaule. 
HITCHCOCK AND CHASE TROPICAL NORTH AMERICAN PANICUM. 477 
20. Panicum capillare L. 
Panicum capillare L. Sp. PI. 58.1753; 
Contr. U. S. Nat. Herb. 15: 60.1910. 
DISTRIBUTION. 
Open ground, common in eastern 
United States, introduced in Ber- 
muda. The type specimen from Vir- 
ginia. 
Bermuda.: North Shore Road, 
Brovm <fc Britton 21. 
21. Panicum hirticaule Presl. 
Panicum hirticaule Presl, llel. Haenk. 1: 308.1830; Contr. U. S. Nat. Herb. 15: 64, 
1910. 
DISTRIBUTION. 
Rocky or sandy soil, southwestern United States and south through Mexico to 
South America; also in Haiti. The type specimen from Acapulco, Mexico. 
Lower California: San Jos6 del Cabo, Brandegee 42. Furfaima, Brandegee in 
1889. 
Sonora: Nogalee, Hitchcock 3631, Griffiths 6758, 6799. Lokuka Ranch to Altar, 
Griffiths 6892. Llano, Hitchcock 3526. Johnstons Ranch, east of San Pedro 
River, Merton 1705. Guaymaa, Hitchcock 3547, 3553, Palmer 208 and 346 in 
1887. Alamos, Palmer 690, 695, and 750 in 1890. White Water, Mearm 2294. 
Sonora, Hitchcock 3541$. Hermosillo, Hitckcoch 3541, 3573, 3604. 
Chihuahua: Southwestern Chihuahua, Palmer lb in 1885. Between Casas 
Grandes and Sabinal, Nelson 6355. Sierra Madre, Nelson 6297. 
Sinaloa: Culiac&n,Palmer 1544,1545, 
and 1554 in 1891. Topolobampo, 
Palmer 249 and 251 in 1897. 
Tepic: Between Aguacate and Dolo- 
res, Rose 3351. 
Jalisco: Guadalajara, Hitchcock 
7357. 
Guanajuato: Irapuato, Hitchcock 
7424. 
Quer£taro: Quer6taro, Hitchcock 
5846. 
Coljma: Alzada, Hitchcock 7089, 7096, 7104. Without locality, Palmer 14, 143, 
and 145 in 1897. 
Veracruz: Bafioe del Carrizal, Purpus G212. 
Guerrero: Balsas, Hitchcock 6773, 6814, 6815. Rio Balsas, Orcutt 4197. 
Oaxaca: Tomellin, Hitchcock 6248, Amer, Or. Nat. Herb. 28. 
Salvador: La Uni6n, Hitckcock 8794. West side of Late Ilopango, Hitchcock 
8924. 
Nicaragua: Masaya, Hitchcock 8654. Without locality, Flint in 1868. 
Costa Rica: Atenas, Hitchcock 8517. 
Hath: Near the River Artibonite, Picarda 1654. 
Curasao: Boldingh 5379. 
Colombia : Puerto de Colombia, Hitchcock 9927. 
Fig. 33.—Distribution of P. capillare. 
Fio. 34.—Distribution of P. ftirricawte. 
478 CONTRIBUTIONS FBOM THE NATIONAL HERBARIUM. 
22. Panicum painpinosum Hitchc. & Chase. 
Panicum pampinowm Hitchc. & Chase, Contr. U. S. Nat. Herb. 15: 66.1910. 
Culms 15 to 40 cm. high, ascending from a decumbent base or widely spreading. 
DISTRIBUTION. 
Open ground and depressions in mesas, 
southwestern United States to central 
Mexico. The type specimen from Ari- 
zona. 
Chihuahua: Minaca, Hitchcock 7751, 
7756. 
Guanajuato: Ac&mbaro, Hitchcock 
6952. 
Ha. 35.—Distribution of P. pampinotum. 
23. Panicum decolorans H. B. K. 
H 
Panicum decolorans H. B. K. Nov. Gen. & Sp. 1:100.1816; Contr. U. S. Nat. Herb. 
15: 66.328.1910. 
DISTRIBUTION. 
Fields and waste ground, central Mexico. 
The type specimen from Quer^taro. 
San Luis Poroaf: Cardenas, Amer. Gr. 
Nat. Herb. 29. 
Quer£taro: Quer£taro, Basils 51, 52, 
Hitchcock 5822, 5864. 
Pukbla: Tehuac&n, Hitchcock 6057. 
24. Panicum stramineum Hitchc. & Chase. 
Panicum stramineum Hitchc. & Chase, Contr. U. S. Nat. Herb. 15: 67, 1910. 
This species was described as having glabrous (or sometimes ciliate) blades. Later 
collections show that the blades are sometimes sparsely papillose-pilose. 
distribution. 
Rich bottom lands and damp soil, Arizona 
to Michoac&n. The type specimen from 
Guaymas. 
Sonora: Guaymas, Palmer 168a and 
206 in 1887. 
Sinaloa: Culiac&n, Palmer 1538 in 
1891. Between Roeario and Aca- 
poneta, Rose 1878,1883. 
Duranco: Durango, Hitchcock 7658. 
Tepic: Between Concepci6n and Acaponeta, Rose 1889. Acaponeta, Rose 3281. 
MichoacAn: Morelia, ArsZne 60, 2985. 
Guerrero: Lagunillas, Langlasse 263.1 
1
 In the Revision this specimen was doubtfully referred to P.jiUpe* Scribn. (Contr. 
U. S. Nat. Herb. 15: 74.1910). 
FiO. 36.—Distribution of P. decolorojw. 
HITCHCOCK AND CHASE—TROPICAL NOBTH AMERICAN PAN1CUM. 479 
20. Fanicum Bonorum Beal. 
Panieum sonorum Beal, Grasses N. Amer. 3 
15: 67. 1910. 
DISTRIBUTION. 
Rich bottom land, northwestern Mexico. 
The type specimen from Lerdo. 
Sonora: Lerdo, Palmer 947 in 1889. 
Without locality, Palmer in 1885. 
Chihuahua: Southwestern Chihuahua, 
Palmer lc in 1885. 
ctnaloa: Culiacin, Palmer 1539 and 
1554 in 1891, 
Chiapas: Between San Ricardo and 
Ocozucuantla, Nelson 2959. 
130. 1896; Contr. IT. S. Nat, Herb, 
Fia. 38.—Distribution of P. sonorum. 
20. Fanicum pare urn llitchc. & Chase. 
# 
Panieum parewn Hitchc. & Chase, Contr. U. S. Nat. Herb. 15: 68.1910. 
DISTRIBUTION. 
Open rocky soil, Sinaloa to Guerrero. 
The type specimen from Sinaloa. 
Sinaloa: Lodiego, Palmer 1657. Cop- 
radfa, Brandegee in 1904. 
Comma: Alzada, Oreutt 4687, Hitchcock 
7081. 
Guerrero: Balsas, Hitchcock 6808, 
6810, 6811, Amer. Gr. Nat. Herb. 
Fig. 39.—Distribution of P. par cum. 30. 
27. Fanicum cayetmense Lam, 
Panieum cayennense Lain. Tab!. Encycl. 1: 173. 1791; Contr, U. S. Nat. Herb. 15: 
70.1910. 
DISTRIBUTION. 
Open ground and pine woods, Cuba, Costa Rica to Brazil. The type from French 
Guiana. 
CostaRica: Buenos Aires, Tonduz 
3685. 
Panama: Dolega, Hitchcock 8335. 
David, Amer. Gr. Nat. Herb. 
31. 
Cuba: Isle of Pines, Curiiss 267, 
Taylor 34, Palmer <6 Riley 108G. 
Herradura, Tracy 9073, Britton, 
Earle & Gager 6520. Sierra de 
Cabra, Britton, Earle & Gager 
7275. Laguna Jovero, Shafer 10910 
del Rfo, Wright 3865. 
Fio. 40.—Distribution of P, cayennense. 
Sumidero, Le6n & Shafer 13717, Pinar 
480 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM 
DIFFUSA. 
Second glume and sterile lemma elongated, at least three times 
as long as the fruit 28. P, capillarioides. 
Second glume and sterile lemma not elongated. 
Culms as much as 1 cm. thick; blades 2 cm. or more wide.. 33. P. hirsutum. 
Culms slender; blades not over 1 cm. wide. 
Blades 1 to 3 mm. wide; plants spreading or ascending. 29. P. diffusum. 
Blades mostly over 5 mm. wide; plants erect. 
Spikelets 4 to 4.2 mm. long, the midnerves of 
glumes and sterile lemma scabrous toward 
the apex  31. P. lepidulum. 
Spikelets usually less than 3.5 mm. long. 
Blades hirsute on both surfaces (sometimes 
glabrescent), not at all glaucous 32. P. ghiesbreghtii. 
Blades glabrous on both surfaces or with a few 
hairs on either surface, glaucous above.. 30. P. kallii. 
28. Fanicum capillarioides Vasey. 
Panicum capillarioides Vasey in Coulter, 
Contr. U. S. Nat. Herb. 1: 54.1890; Contr. 
U. S. Nat. Herb. 16: 72. 1910. 
DISTRIBUTION. 
Prairies, southern Texas and northern 
Mexico. The type specimen from Texas. 
Nuevo Le6n: Monterrey, Hitchcock 
5547. FlO. 41.—Distribution of P, capillarioidtS- 
29. Fanicum diffusum Swartz. 
Panicum diffusum Swartz, Prodr. Veg. Ind. Occ. 23. 1788; Contr. U. S. Nat. Herb. 
15: 72. 1910. 
Panicum gvadalupense Spreng.; Steud. Nom. Bot. ed. 2. 2: 257. 1841. This is a 
nomen nudum, ascribed to "Spr. hrb. Trin. mpt. Guadal." In the Krug & Urban 
Herbarium is a specimen of Panicum diffusum, collected by "Bertero, S. Domingo," 
marked in Sprengel's hand "Paniqum guadalu-pense" 
DISTRIBUTION • 
Banks, cliffs, and savannas, West Indies. The type specimen from Jamaica or 
Hispaniola. 
Bahamas: New Providence, Britten & Brace 424. 
Cuba: Campo Florido, Le6n 3449. 
Vibora Habana, Le6n 2532. 
Monte la Noria, Le6n 2559. 
13 ah fa Honda, Wilson 9443. 
Holguin, Shafer 1512. La 
Gloria, Shafer 385. Columbia, 
Le6n 305 in part, 923b. Ar- 
royo Apolo, Le6n 190, 923. 
Province of Santa Clara, Le6n 
923c. Santiago dc las Vegas, 
Tracy 9111, Van Hermann 1444 
Fig. 42.—Distribution of P. diffusum. 
Baker & Wilson 511, Wilson 1405." Han&bana, Wright in 1865. Triscornia, 
Tracy 9082. Guanajay, Palmer & Riley 802. Isle of Pines, Curtiss 384, 494. 
Without locality, Wright 1540, 3852, 3860 in part, 3877. 
I 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 481 
Jamaica: Ewarton to Linstead, Hitchcock 9463. 
. Santo Domingo: Without locality, Wright, Parry <tr Brummel 627. Azua, Rose, 
Fitch Russell 4410. 
Porto Rico: Quebradillas, Chase 6567, 6577. Joyuda, Chase 6308. 
Danish West Indies: St. Thomas, Eggers in 1882. 
Leeward Islands: Antigua, Wvllsdilaegel 64. Guadeloupe, Duss 3177, 3181 
{K. U. Herb.). St. Bartholomew, Goes (K, U. Herb.). 
Windward Islands: Martinique, Duss 536, Rahn 1232. 
30. Panicum hallii Vasey. 
Panicum hallii Vasey, Bull. Torrey Club 11: 64.1884; Con.tr. U. S. Nat. Herb. 15: 
74.1910. 
DISTRIBUTION. 
Dry prairie, rocky and gravelly hills and canyons, and in irrigated fields, Texas to 
Arizona and south to central Mexico. The 
type specimen from Texas, 
Chihuahua: Santa Eulalia Mountains, 
Pringle 376. 
Durango: Without locality, Palmer 
525 in 1896. 
Coahuila: Monclova, Palmer 1338 in 
1880. Saltillo, Hitchcock 5605. 
Nuevo Le<5n: Monterrey, Hitchcocl: 
5537. 
Tamaulipas: Victoria, Palmer 554 in 
1907. 
San Luis PoTOsf: Las Canoaa, Hitchcock 5756. Cardenas, Hitchcock 5719$. 
31. Panicum lepidulum Hitchc. & Chase. 
Panicum lepidulum Hitchc. & Chase, Contr, U. S. Nat. Herb. 15: 75. 1910. 
distribution. 
Rocky hills and moist banks, Mexico to Panama. The type specimen from 
Chihuahua. 
Chihuahua: Chihuahua, Pringle 497. 
Durango : Durango, Palmer 525 in 1896, Hitchcock 7591. Tobar, Palmer 533 in 1906. 
Zacatecas: Zacatecas, Hitchcock 
7510. 
San Luis Potosi: Cardenas, Hitch- 
cock 5719. 
Jalisco: San Nicolds, iRfcfcocfe 7199. 
ZapotMn, Hitchcock 7238. 
Quer£taro: San Juan del Rfo, Rose, 
Painter & Rose 9555. 
Mfiuco: Pedregal, Federal District, 
Amer. Gr. Nat Herb. 35. 
Pueblo: Tehuacdn, Hitchcock 6063. 
Fig. 44.—Distribution of P. lepidulum. 
Guatemala: Guatemala City, Hitchcock 9014. 
Panama: Sabano de Juan Corso, Pittier 4526. 
82472°—15 3 
I 
482 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
9 
32. Panicum ghiesbreghtii Fourn. 
Panicum ghiesbreghtii Fourn. Mex. PI. 2: 29.188G; Contr. U. S. Nat. Herb. 15: 76. 
1910. 
DISTRIBUTION. 
Low moist ground, Mexico and West Indies to northern South America. The type 
specimen from Mexico. 
Jalisco: Zapotlan, Hitchcock 7117. 
Veracruz: Cordoba, Hitchcock (5418, 6426. Jalapa, Hitchcock 6546. Orizaba, 
Bourgcait 2751, Hitchcock 6363, Airier. Gr. Nat. Herb. 30. 
Oaxaca : Oaxaea, Hitchcock 6113. 
Yucatan: Izamal, Gaurncr 2477. 
Salvador: Acajutla, Hitchcock 8993. 
Nh aragi a: Corinto, Hitchcock 8755. Han. Juan del Sur, Hitchcock 8601. Jino- 
tepe, Hitchcock 8687. 
|F<~1 : \ V 0 
\\\Tu 
I {*{'Q> 
,  
Fig. 45.—Distribution of QMexbrryhtii. 
dura, Tracy 9068. Santiago de laa Vegaa, Traci/ 9116. La Perla, Le6n 3783, 
El Calvario, Le6n 922, 2674. Guayabal, Lc6n 922b. Without locality, 
Wright 758. Eastern Cuba, Wright. Santiago de Cuba, Le6n 3747. 
Porto Ilico: Coamo Springs, Chase. 0517. Cayey, Chase 6744, Tabucoa, Sintam 
4983. 
Leeward Islands: Guadeloupe, Dims 3184. Antigua, Wullschlaegel 620, 621. 
(K. U. Herb ). 
Colombia: SantaMarta, Smith 165. 
33. Panicum hirsutum Swartz. 
Panicum hirsutum. Swartz, Fl. Ind. Gee. 1: 173. 1797; Contr. U. S. Nat. Herb. 15: 
77, 1910. 
Panicum datum Willd.; Steud. Nom. Bot. cd. 2. 2: 256.1841. In the Willdenow 
I 
Herbarium this unpublished name is written on a Humboldt collection from "America 
merid. *' 
The sheathe, especially the lower, of thia robust species are hirsute with spreading 
stiff hairs that cause mechanical irritation to the skin. The panicles are at first com- 
pact, with ascending branches, but become diffuse at maturity, the branches finally 
widely spreading. 
distribution. 
Open moist mil, West Indies and central Mexico to northern South America. The 
type specimen from Jamaica. 
Colima: Manzanillo, Hitchcock 7031. 
Oaxaca: San Antonio, PringU 5573. 
Costa Rica: Port Limon, Hitchcock 8435. Along Hananitu River, Pittier 3634. 
Zent Farm, Pittier in 1904, Tonduz 266. Puerto Viejo, Biollcy 7467. 
Costa Rica: San Jose, Hitchcock 8452. 
Guanacaate, Jimenez 730, Llano 
Grande de Puriscal, Jimenez 880. 
Nuestro Amo, Jimenez530. Salinas 
Bay, Pittier 2633, Tonduz 2858. 
Panama: South of David, Hitchcock 
8377. Balboa, Hitchcock 7997, 
8014. Taboga Island, Hitchcock 
8065, 8094. 
Bahamas: Crooked Island, Brace4812. 
<' i ba : La Soledad, Eggers 5406. Ilerra- 
+ 
HITCHCOCK AND CHASE TROPICAL NORTH AMERICAN PANICUM. 483 
Fig. 46.—Distribution of P. htrmtum* 
Panama: Between Bohfo and Frijoles, Hitchcock 8397. Near Bohio, Hitchcock 
8392, Tabernilla, Hitchcock 8382. Gorgona, Amer. Gr. Nat. Herb. 37. Pedro 
Miguel, Hitchcock 7957. Puerto 
Obaldfa,Pittier 4374. Culebra, 
Pittier 3441. 
Cuba: Matanzaa, Wright in 1865. 
GuanUnamo, Le&n 3913. 
Leeward Islands: Guadeloupe, 
Dms 3917. 
Windward Islands: Martinique, 
Buss 768 (K. U. Herb.). 
Trinidad: St. Margaritas, Brood- 
way 2629. "Burke's pasture,'' 
Bot. Gard. Herb. 2295. 
Venezuela: Rfo Grande del Tuy, Paparo, Pittier 6332. Boburee, John 353, 355. 
Colombia: Santa Marta, Smith 164. 
MAXIMA. 
Culms with a cormlike base. 
Blades mostly over 5 mm. wide; culms more than 1 meter 
high 36. P. bulbosum. 
Blades less than 5 mm. wide; culms rarely as much as 1 meter 
high 36a. P. bulbosun 
sdaphilum. 
Culms from a creeping rootstock, not cormlike at base. 
Nodes hirsute; ligules 4 to 6 mm. long; fruit strongly rugose 34. P. maximum. 
Nodes glabrous; ligules 2 mm, long; fruit very obscurely 
rugose 35. P. plenum. 
34. Panicum maximum Jacq. 
Panicum maximum Jacq. Coll. Bot. 1:76.1786; Contr. U. S. Nat. Herb. 16:78.1910. 
DISTRIBUTION. 
Open ground, at low altitudes, escaped from cultivation, southern Florida through 
Mexico and West Indies to South America, a native of Africa and now widespread 
in the warmer parts of the Old World. Type locality, Guadeloupe. 
Sinaloa: Mazatldn, Rose, Standley & Russell 14121. 
Co LIMA: Paso del Rfo, Emrick 3. 
Veracruz: C6rdoba, Hitchcock 6410, Finck 8, Kerber 48. Jalapa, Hitchcock 
6621, Huitamalco, Liebmann 425 
T3 1 in part. Veracruz, Hitchcock 6562. 
San Francisco, Smith 1409. Zacu- 
■ apdn, Purpus 3774. 
YucatAk: Izamal, Gaumer 719. 
British Honduras: Manatee Lagoon, 
Peck 195 (Gray Herb.). 
Guatemala: Escuintla, Smith 2705, 
2706, Hitchcock 9001. Morales, Kel- 
lerman 6267. Gualtln, Deam. 6268. 
Finca Sepacuit£, Alta Verapaz, 
Cook & Griggs 579. 
Honduras: Puerto Cortez, Kellerman 4725. 
Salvador: San Salvador, Velasco 2. Izalco, Pittier 1960. Without locality, 
Renson 293. 
Fig. 47.—Distribution of P. maximum. 
484 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
Nicaragua: Corinto, Hitchcock 8702. 
Costa Rica: Zent Farm, Pittier in 1904. Boca Banana, Tonduz 9114. San Joe£, 
Pittier 9050. Nuestro Amo, Pittier 2034, 16662. Alajuelita, Tonduz 2995. 
Panama: Taboga Island, Hitchcock 8080. Cafia, Williams 779. Culebra, Pittier 
2088, 4806, Amer. Gr. Nat. Herb. 38. 
Bermuda: North Shore Road, Brown & Britton 20, Ifarshberger in 1905. 
Bahamas: Nassau, Curtiss 124. Eleuthera, Geogr. Soc. Baltimore 338. 
Cuba: Herradura, Hitchcock 156. Guines, Leon 427, Liebmann 445. Santiago de 
las Vegas, Baker 32, Wilson 438, Hitchcock 157. Santiago de Cuba, Le6n 915, 
Pollard 6c Palmer 283. Guayabal, Le&n 921. Marianao, Le6n 957. Guana- 
jay, Palmer & Riley 816. San Diego de log Bafios, Palmer & Riley 542, 545. 
El Guama, Palmer & Riley 178. Pinar del Rfo, Palmer & Riley 377. Sierra 
de Anafe, Le6n 2871. 
Jamaica: Port Antonio, Fredholm 3319. Kingston, Hitchcock 9256, and in 1890, 
Gordon Town, Hart 797. Hope Gardens, Harris 11249. 
Haiti: Gonai'ves, Buck 468 (K. U. Herb.). 
Santo Domingo: Azua, Rose, Fitch & Russell 3949. San Pedro de Macoris, Rose, 
Fitch & Russell 4172. 
Porto Rico: Guanica, Sintenis3366. Mayaguez, Sintenis 51, Chase 6256. Cayey, 
Sintenis 2468. Aibonito, Chase 6332. Sobrante, Eggers 1226. Martin 
Pefla, Heller 377. Vieques, Chase 6669. Mona, Hess 449. 
Danish West Indies: St. Croix, Ricksecher 200, 413. St. Thomas, Millspaugh 
454, Eggers in 1887, Rose 3184. 
Leeward Islands: Antigua, WuUschlaegel 623b, Rose, Fitch & Russell 3452. 
Dominica, Jones 10. Guadeloupe, Duss 3186. St. Bartholomew, Goes 
(K. U. Herb.). Saba, Boldingh 1580, Suringar in 1885 (both in K. U. Herb.) 
Windward Islands: Martinique, Duss 1288. Barbados, Dash 259. Grenada, 
Broadway in 1905. St. Vincent, Smith & Smith 288 (K. 0. Herb.). 
Trinidad: Four Roads, Hitchcock 10049. 
Tobago: Scarborough, Broadway 4081. Plymouth, Hitchcock 10239. 
Venezuela: Caracas, Pittier 6163. Around Dos Caminos and Los Chorros, 
Pittier 5917. Rio Grande del Tuy, Pittier 6327. Island of Margarita, Miller 
& Johnston 177. 
Colombia: Santa Marta, Smith 2153. Cartagena, Hitchcock 9917. Rfo Frfo, 
Pittier 1617. 
35. Panicum plenum Hitchc. & Chase. 
Panicum plenum Hitchc. & Chase, Contr. U. S. Nat.. Herb. 16: 80. 1910. 
Moist places in rocky hills and canyons, 
southwestern United Statos to central 
Mexico. The type specimen from New 
Mexico. 
Chihuahua: Santa Eulalia Plains, 
So nora: Las Cuevas, Hartman 170. 
Hitchcock 7772. 
Wilkinson in 1885. Chihuahua, 
DISTRIBUTION 
Fig. 48.—Distribution of P. plenum. 
Coahuila: Saltillo, Hitchcock 5609. 
Veracruz: Orizaba, Botteri 160. 
Durango: Durango, Painter 741 in 
1896. 
Jaral, Schumann 1733. 
J 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 485 
36. P&nicum bulbosum H. B. K. 
Panicum bulbosum H. B. K. Nov. Gen. & Sp. 1: 99.1816; Contr. U. S. Nat. Herb. 
15: 81. 1910. 
DISTRIBUTION. 
Moist places in canyons and valleys, southwestern United States to southern Mexico. 
The type specimen from central Mexico. 
Chihuahua: Candelaria, Stearns 2C6. Chihuahua, Pringle 377. Colonia Garcia, 
Nelson 6187, Tovmsend 6c Barber 221. Sierra Madre, Nelson 6301. Mount 
Mohinora, Nelson 4901. Canon de San Diego, Hartman 790. 
Durango: Otfriapa, Palmer 340 in 1906. Durango, Palmer 525a in 1896, With- 
out locality, Palmer 741 in 1896. 
Tepic : Between Aguacate and Dolores, 
Rose 3361. Between Pedro Paulo 
and San Blasito, Rose 1999. 
San Luis PotosI: San Luis Potosf, 
Parry & Palmer 958. Las Canoas, 
Hitchcock 5754. 
Jausco : Road between Mexquitic and 
Monte Escobedo, Rose 2609. Ne- 
vada de Colima, Hitchcock 7150. 
Rfo Blanco, Palmer 207 in 1886. 
MichoacAn: Morelia, Arsbie in 1909. 
Mexico: Federal District, Orcutt 3518, 
Hitchcock 5932, 5941, 7835, Amer. Gr. Nat. Ilerb. 39, Holway 12, Bourgeau 
235, Rose <fc Painter 9249, Pringle 9575,6418. Valley of Toluca, Pringle 5207. 
Puebla: Chinantla, Lkbmann 441, 442, Cerro de Baxtla, Purpus 2908. Espe- 
ranza, Seaton 317. Bords de l'Atoyac, Nicolas in 1909. 
Veracruz: Borrego, Bourgeau 2754. Orizaba, Hitchcock 6384. 
Oaxaca : Oaxaca, Nelson 1374. Cerro de San Felipe, Conzatti & Gonzdlez 243. 
36a. Panicum. bulbosum sciaphilum (Rupr.) Ilitchc. & Chase. 
Panicum bulbosum sciaphilum (Rupr.) Hitchc. & Chase, Contr. U. S. Nat, Herb. 
15: 83.1910. 
DISTRIBUTION. 
Gravelly river banks, southwesternUnited 
States to central Mexico. The type spec- 
imen from central Mexico. 
Sonora: Nogales to Cocospora Ranch, 
Griffiths 6785J. 
Chihuahua: Cusihuiriachic, Pringle 
1406. Siena Madre, Nelson 6298. 
S&nchez, Hitchcock 7666. 
Durango: Tejam&n, Pcdmer46^ in 1906. 
Otfnapa, Palmer 348, 349, and 554 
in 1906. Santiago Papasquiaro, Palmer 467 in 1896. Iron Mountain, Hitch- 
cock 7634. 
Tepic: Between Dolores and Santa Gertrudis, Rose 2053. 
M&uco: Barranca de Rfo Aqueducto, Rose 6c Painter 8634, 
Fig. 49.—Distribution of P. bulbosum. 
FIG, 50.—Distribution of jp. bulbosum 
sciaphiluTB, 
o 
486 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
VIRGATA. 
Perennials, mostly with stout rootstocks, one species, P. ichnanthoides found to be 
without rootstocks. 
Creeping rootstocks wanting  40. P. ichnanthoides. 
Creeping rootstocks present; 
Spikelets not over 2.5 mm. long; first glume less than half the 
length of the spikelet. 
Panicles loosely flowered; first glume truncate, about 
one-fifth the length of the spikelet.  37. P. repens. 
Panicles rather densely flowered; first glume triangular, 
about one-third the length of the spikelet ... 38. P. gomni. 
Spikelets 3 to 7 mm. long (sometimes less than 3 mm. in P. 
virgalum cubense); first glume more than half the length 
of the spikelet. 
Culms decumbent or creeping at base. 
Spikelets 3.2 to 4 mm. long 41. P. altum. 
Spikelets 6 to 8 mm. long  42. P. havardii. 
Culms erect. 
Panicles elongate, strongly contracted; seacoast 
plants 43. P. amarulum. 
Panicles diffuse or only slightly contracted; plants 
sometimes of salt marshes but not littoral. 
Panicles open, loosely-flowered; spikelets 3.5 to 
5 mm. long, beaked; first glume two- 
thirds the length of the spikelet or more, 
acuminate-pointed 39. P. virgalum. 
Panicles somewhat contracted; spikelets not 
over 3.2 mm. long, not beaked; first 
glume about half the length of the spike- 
let, not acuminate  
37. Panicum repens L. 
39a. P. virgatum 
cubense. 
Panievm repens L. Sp. PI. ed. 2. 87. 1702; Contr. U. S. Nat. Herb. 16: 85.1910. 
DISTRIBUTION. 
Sea beaches, warmer regions of both 
hemispheres; in America from Ala- 
bama to Brazil, Originally described 
from the Old World. 
Nicaragua: Without locality, 
Flint 8. 
Cuba: Arroyo Apolo, Le6n 296, 
563. Habana, Le6n in 1909. Fig. SI.—Distribution of repent. 
> 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 487 
38. Panicum gouini Fourn. 
Panicum gouini Fourn. Mex. PI, 2: 28. 
1886; Contr. U. S. Nat. Herb. 15:86.1910. 
DISTRIBUTION. 
Sea beaches along the Gull of Mexico. 
The type specimen from Veracruz. 
Veracruz: Veracruz, Pringle 5569, 
Amer, Gr, Nat. Herb. 40, Miiller 
2177. Antigua, Liebmann 450. 
Coatzacoalcos, Smith 913. 
Fia. 52.—Distribution of P. gouini. 
39. Panicum virgatum L. 
Panicum virgatum L. Sp. PI. 59. 1753; Contr. TJ. S. Nat. Herb. 15: 87.1910. 
distribution. 
Prairies and open woods, eastern and 
central United States, south to north- 
ern South America. The type speci- 
men from Virginia. 
Jausco: Rio Blanco, Palmer 207 
and 510 in 1886. 
Chiapas: Between San Kicardo 
Fig. 63.—Distribution of P. virgatum. an^ Ocozucuantla, Nelson 
2975. 
Honduras: Santa Anna, Thieme 532. 
Venezuela: Alto A pure, John 198. 
Colombia: Santa Marta, Smith lfitt. 
/ 39a. Panicum virg&tum cubense Grieeb. 
Panicum virgatum cubense Grisob. Cat. PI. Cub. 233. 1866; Contr. U. S. Nat. Herb. 
16: 92. 1910. 
" f-j | ] f 
distribution. 
Pine woods, Atlantic Coastal Plain, 
Bermuda and Cuba. The type speci- 
men from Cuba. 
Bermuda: Stone in 1888 (Acad. 
Phil. Herb.). 
Cuba: Bataban6, IlitchcocJc 153, 
Palmer & Riley 1134. Be- 
Ffo. 54.—Distribution of P. virgatum cxtben&t. 
twecn Laguna Jovero and Laguiia Herradura, Sha/er 10927. Without locality, 
Wright 3873. 
488 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
40. Panieum ichnanthoides Fourn. 
Panieum ichnanikoides Fourn. Mex. PI. 2: 30.1886; Contr. U. S. Nat. Herb. 15: 88. 
1910, as synonym under P. virgatum,, The type specimen, Miiller 2002, in the her- 
barium of the Botanical Garden in St. Petersburg, was collected at Orizaba. 
Panieum buchingeri Fourn. Mex. PI. 2: 30. 1886. ''Orizaba (Thomas in herb. 
Buchinger)." Contr. U. S. Nat. Herb. 16: 329: 1910. We have been unable to find 
the type. From the description it ia evident that the species is allied to P. virgatum 
L., while the locality would indicate P. ichnanthoides. 
DESCRIPTION. 
Plants in largo clumps with knotted crowns, without creeping rootstocks; culms 
1.5 to 2 meters high, erect or decumbent at base, smooth, firm, and hard, more or less 
glaucous about the nodes; sheaths shorter than the inter- 
nodes, pubescent on the overlapping margin toward the 
apex, otherwise glabrous; liguleabout3 mm. long, mem- 
branaceous-ciliate; blades elongate, mostly 8 to 15 mm. 
wide, flat except at the narrowed and usually inrolled 
base, scabrous on the margins, villous on the upper 
surface near the base, otherwise glabrous; panicle 20 
to 40 em. long, one-fourth to one-third as wide, the 
long slender branches ascending; spikelets 3.5 to 4.2 
mm. long, about 1.5 mm. wide, similar to those of 
P. virgatum. 
This species is closely allied to P. virgatum L., from 
which it differs in the absence of creeping rootstocks 
and in the narrow panicle. The patch of plants from 
which was collected Hitchcock's no. 6364 is probably 
that from which Botteii, Miiller, and Thomas obtained their specimens. This patch 
lies along a trail leading up a prominent hill at tho western edge of the city of 
Orizaba, a locality likely to be visited by all botanists collecting in that region. 
DISTRIBUTION. 
Open rocky hillsides, southern Mex- 
ico, British Honduras, and Nicaragua. 
Puebla: Tehuac&n, Endlick 1938. 
Veracruz: Orizaba, Botteri 648, 
Hitchcock 6364, Mii ller 2002. 
British Honduras: Manatee La- 
goon, Peck 73 (Gray Herb.). 
Nicaragua: Jinotepe, Hitchcock 
8685. Pig, 56.—Distribution of P. ichiunUhoid(3. 
Fig. 55.—P. ichnanthoides. 
From type specimen. 
41. Panieum altum sp. nov. 
description. 
Plants perennial, in tangled masses, the culms decumbent and straggling at baee 
stout, reedlike, as much as 6 mm. in diameter, ascending to a height of 2 to 4 meters, 
smooth and glaucous, simple or with extra vaginal, mostly erect, sterile branches from 
the swollen nodes; sheaths smooth; ligule a short membrane less than 1 mm. long, the 
ciliate fringe once or twice as long; blades flat, firm, not greatly elongated, mostly 30 
to 45 cm. long, 8 to 15 mm. wide, smooth except the scabrous margins, panicles rather 
more densely flowered than in P. virgatum, 20 to 30 cm. long, narrowly ovoid in out- 
HITCHCOCK AND CHASE—TROPICAL ItOBTH AMERICAN PANICUM. 489 
Fig. 57,—P. altum. From 
type specimen. 
line, open, the slender scabrous branches ascending and spreading, fascicled, some 
naked at base, with short branchlets intermixed; spikelets rather short-pediceled, 3.2 
to 4 mm. long, about 1.2 mm. wide, turgid, glabrous, 
the glumes and sterile lemma strongly nerved, pointed, 
somewhat keeled, usually widely gaping, the first 
glume about two-tliirds the length of the spikelet, 
the second glume and sterile lemma exceeding the 
fruit, subequal; fruit 2 to 2.2 mm. long, 1 mm. wide, 
elliptic. • 
Type in the U. S. National Herbarium, no. 093326, 
collected in moist sandy soil, near sea beach on Point 
Cham6, Province of Panama, September 18,1911, by 
A. S. Hitchcock (no. 8167), 
In the inflorescence this species closely resembles 
P. virgatum, but in habit it differs distinctly. It 
grows in masses like large reeds such as Phragmites but is straggling and much 
tangled at base. 
DISTRIBUTION. 
Sandy marshes or flats near the seacoast, British Honduras to Trinidad. 
British Honduras: Manatee Lagoon, 
Peck 123 (Gray Herb.), 
Costarica: Buenos Aires, Ton<2«z3619. 
Panama: Point Cham£, Bitchcock 8167. 
Trinidad: Near Port of Spain, Bitch- 
cock 10053. Icacos, Hitchcock 10154. 
Three miles south of Cedros, Bitch- 
cock 10159. 
Tobago: Plymouth, Bitchcock 10233. 
Fig. 58.—Distribution of P. aitum. Pigeon Point. Broadway 3068. 
42. Panicum havardii Vasey. 
Panicum havardii Vasey, Bull. Torrey 
Club 14: 95.1887; Contr. U. S. Nat. Herb. 
15: 93.1910. 
DISTRIBUTION. 
Arroyos and sand hills, southwestern 
United States and northern Mexico. The 
type specimen from Texas. 
Chihuahua: Paso del Norte (Ju&rez), 
Pringle 1124. rio. 59—Distribution of P. havardii. 
43. Panicum amarulum Qitchc. & Chase. 
Panicum amarulum Hitchc. & Chase, Contr. U. S. Nat. Herb. 16: 96.1910. 
DISTRIBUTION. 
Sandy seashores and coast dunes, 
*" southeastern United States, Bahamas, 
and Cuba. The type specimen from 
Virginia. 
Bahamas: Great Bahama, Britton 
& Millspaugh 2739. New 
Providence, Britton & Brace 
307,403 (all FieldMus. Herb ). 
Cuba: Camaguey, Shafer 2735. 
Fig, 60,—Distribution of Pr amarulumt 
490 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
TENERA. 
Second glume and sterile lemma exceeding the fruit; spikelets 
pointed 44. P. tenerum. 
Second glume and sterile lemma not exceeding the fruit; epikeleta 
rather blunt. 
Pedicels bearing long stiff erect haira at the summit 46. P. caricoides. 
Pedicels not hairv. 
Spikelets attenuate at base, alxrnt 2 mm. long; leaves more , 
or less pilose 45. P. sienodoides. 
Spikelets not attenuate at base, about 1.5 mm. long; leaves 
glabrous 47. P. stenodes. 
44. Panicum tenerum Beyr. 
Panicum tenerum Beyr. in Trin. M£m. Acad. St. P6tersb. VI. S< i. Nat. 1: 341. 
1834; Contr. U. S. Nat. Ilerb. 15: 97. 
1910. 
DISTRIBUTION. 
Fig. fil.—Distribution of JJ, tcnerurti* 
Margins of sandy swamps and ponds 
and in wet places in sandy woods, 
southeastern United States, Cuba, and 
Porto Rico. The type specimen from 
Georgia. 
Cuba: Herradura, Hitchcoch 154, 
Tracy 9080. Laguna Jovero, 
Shafer 10750. Laguna Los Indios, Shafer 10807. Pinar del Rio, Wright 3870 
in part. 
Porto Rico : ("ampo Alegre, Chase 6617. 
45. Panicum stenodoides Hubbard, 
Panicum stenodoides Ilubbard, Proc. Amer. Acad. 40: 497. 1913. "Type (in the 
Gray Herb.) and only specimen seen, low pine ridge, Ycacos Lagoon, March 5,1907, 
M. E. Peck, no. 681." A portion of the type in Ilubbard's herbarium has been 
examined. 
DESCRIPTION. 
Plants perennial in dense tufts; culms erect, slender and wiry, glabrous or sparsely 
pilose below the glabrous nodes, 20 to 40 cm. high, producing from the upper nodes 
slender leafless panicle-bearing branches; sheaths papillose 
or more or less papillose-pilose, the lowermost bladeless, 
glabrous, in age fibrous; ligule membranaceous, about 0.3 
mm. long; blades erect, flat at base, involute toward the 
apex, 3 to 8 cm. long, 1 to 2 mm. wide, more or less papil- 
lose or papillose-pilose, at least on the upper surface, some- 
times glabrous; panicles short-exserted, usually exceeded 
by the uppermost blade, about 1 cm. long, narrow, bearing 
3 to 7 spikelets on appressed scabrous pedicels, rarely a few 
delicate hairs at the apex of the pedicels; spikelets 2 to 2.1 
mm. long, 0.9 mm. wide, turgid, attenuate at base, glab- 
rous; first glume about half as long as the spikelet; second glume and sterile lemma 
subequal, barely covering the fruit, strongly nerved; fruit 1.7 min. long, 0.8 mm. wide. 
Fig. t>2.—P. stenodoidesr From 
typo specimen. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 401 
This species resembles P, stenodes, but differs in having larger spikelets and papil- 
lose-pilose or at least papillose sheaths. The type specimen and Hitchcock's no. 
8147 are conspicuously pilose on sheaths and blades, but the other specimens of this 
little-known species are papillose only or have a few hairs on the upper surface of the 
blades. 
DISTRIBUTION. 
Open grass land and moist savannas, 
Central America and northern South 
America. 
British Honduras: Ycacos La- 
goon, Peck 681 (Gray Herb.). 
Costa Rica: Buenos Aires, Tonduz 
3583. 
Panama: Chorrera, Hitchcock 8147. 
Near Corozal, Hitchcock 9207. 
Fig. 63.—Distribution of P. stenodoidet. 
Trinidad: Piarco Savanna, south of Arouca, Hitchcock 1C346. 
46. Panicum caricoides Nees. 
Panicum caricoides Nees; Trin. Gram. Pan. 149.1826. Trinius received the type 
specimen from Nees, "V. spp. Brazil. (N. ab Esenb.)" Nees 1 afterwards describes 
the species more fully and gives the original locality as "provincia Paraensi." The 
type specimen is in the Trinius Herbarium at the St. Petersburg Academy of 
Science. It is evidently a portion of the specimen at the Munich Herbarium. This 
specimen, which is the basis of Nees's later description, was collected in Pard by 
Martius. 
Panicum junciforme Steud. Syn. PI. Glum. 1: 82. 1864. Given as a synonym of 
P. caricoides. 
description. 
Perennial, in tufts; culms erect, slender, glabrous, stiff 
and wiry, as much as 90 cm. high; sheaths smooth; ligule 
membranaceous, lacerate, minute; blades appressed, nar- 
rower than the sheath, involute, glabrous, 1 to 5 cm. long, 
the lower rarely longer; panicles terminal and axillary from 
the upper sheaths, short-exBerted, 1 to 1.5 cm. long, narrow, 
nearly simple, the later panicles smaller, in fascicles of 2 or 
3, at first partially inclosed by the somewhat inflated sheaths, the pedicles bearing 
long erect white hairs just below the rather crowded spikelets; spikelets 1.5 to 1.8 
mm. long, about 0.8 mm. wide, turgid, glabrous; first glume about half as long as 
the spikelet, subacute; second glume 
and sterile lemma equal, covering the 
fruit, strongly nerved; fruit 1.4 to 1.5 
mm. long, 0.6 mm. wide. 
distribution. 
Wet savannas, Trinidad to Brazil. 
Trinidad: Aripo Savanna, 
Cumuto Station, Ilitchcork 
10069. J-'iq. 65.—Distribution of P. caricoides. 
Fig. 64.—P. caricoides. 
From type specimen. 
'Agrost. Bras. 108. 1829. 
492 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
47. Panicum stdnodes Griseb. 
Panicum stenodes Griseb. Fl. Brit. \V. Ind. 547. 1864; Contr. U. S. Nat. Herb. 15: 
98.1910. 
DISTRIBUTION. 
* 
Borders of ponds and wet savannas, 
West Indies and Costa Rica to Brazil. 
The type specimen from Jamaica. 
Costa Rica : Buenos Aires, Pittier 
10589, Tonduz 3583. 
Cuba: Herradura, Hitchcock 155. 
Without locality, Wright 3871. 
Jamaica: Bull Head Mountain, 
Amer. Gr. Nat. Herb. 46. 
Santo Domingo: Without locality, Wright, Parry <& Brummel 624. 
Porto Rico: Trujillo Alto, Chase 6769. 
Trinidad: Cumuto Station, Hitchcock 10069. Piarco Savanna, Hitchcock 10347. 
AGROSTOIDIA. 
48. Panicum condensum Nash. 
Panicum condensum Nash in Small, Fl. Southeast. U. S. 93, 1903; Contr. U. S. Nat. 
Herb. 15: 102. 1910. 
DISTRIBUTION. 
Swamps and borders of ponds and streams, Pennsylvania to Florida and Texas 
near the coast, south into Mexico and tlie West Indies. The type specimen from 
Florida. 
Veracruz: Without locality,Goitin 
10. 
Bahamas: Nassau, Cur tins 174. 
Cuba: Banks of Guanabo River, 
Le6n 4141. Along Gibora car- 
retera north of Holguin, Shafcr 
1438. Han&bana, Wright 184 
in 1865. Without locality, 
Wright 3862 in part. 
Jamaica: Savanna-la-Mar, Hitck- 
cock 9884. Black River, Amer. Gr. 2Vat. Herb. 47. 
Porto Rico: Santurce, Chase 6351. Campo Alegre, T agnna del Tortuguero, 
Chase 6802. 
Leeward Islands: Guadeloupe, Duss 3919. 
LAXA. 
All the species are pereitnial. 
Spikelets not expanded at maturity by an enlarged sterile palea, 
pointed. 
Nodes glabrous; spikelets 2.5 mm. long  49. P. fongum. 
Nodes densely pubescent; spikelets not over 1.5 mm. long.. 50. P. polygonatum. 
Fig. 66,—Distribution of P. xlcnodes. 
Fig, (^—Distribution of P. condensum. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICT7M. 493 
Spikelets expanded at maturity by the enlarged sterile palea, 
usually blunt. 
Panicle branches not racemose; blades scarcely wider than 
their sheaths; eterilo palea conspicuously enlarged. 
Spikelets 3 mm. long, congested; panicles dark purple.. 58. P. cupreum. 
Spikelets not over 2A mm. long; panicles green or pale. 
Panicle branches spikelet-bearing or branchlet 
bearing along the upper half or toward the 
ends only 57. P. kians. 
Panicle branches branchlet-bearing throughout their 
length or nearly so.....  !.... 56. P, exiguiflorum. 
Panicle branches subracemose, the epikelets secund; blades 
at least 5 mm. wide; enlarged sterile palea not con- 
spicuous. 
Blades narrowed toward the base  54. P. laxwn. 
Blades cordate or truncate at base. 
Spikelets 2 mm. long; panicle branches erect or 
nearly so 55. P. stevenaianum. 
Spikelets not over 1.6 mm. long, usually less; panicle 
branches spreading or ascending. 
Panicles one-third to half as wide as long; spike- 
lets not conspicuously secund, somewhat 
irregularly disposed; blades cordate- 
clasping   53. P. boliviense. 
Panicles rarely one-fourth as wide as long; spike- 
lets conspicuously secund and regularly 
disposed. 
Culms as much as 2 meters long; panicles 
25 to 30 cm. long  51. P. fnilhjtorum. 
Culms not over 1 meter long; panicles 5 to 
15 cm. long :  52. P. jrilosum. 
49. Panicum longum Hitchc. & Chase. 
Panicum longum Hitchc. & Chase, Contr. 
U. S, Nat. Herb. 15: 111. 1910. 
DISTRIBUTION. 
Swamps, State of Veracruz. The only 
collections known are from near Jalapa, 
Pringle 8195 (the type) and Schiede & Deppe 
674 (Halle Herb.). 
50. Panicum polygonatum Schxad. 
Panicum polygonatum Schrad. in Schult. Mant. 2: 256. 1824; Contr. U. S. Nat. 
Herb. 15:112. 1910. 
DISTRIBUTION. 
Swamps and moist soil, Mexico to Paraguay. The type specimen from Brazil. 
Veracruz: Cdrdoba, Bourgeau 1662 in part. 
Guatemala: SepacuitA, Collins & Goll 08, Cook 6c Griggs 530. Cubilquitz, 
Tiirckheim 7797, 8795. Puerto Barrios, Kellerman 5114. 
Honduras: San Pedro Sula, Thieme 781, 5578, 5587 in part. 
* 
494 CONTRIBUTIONS FBOM THE NATIONAL HERBARIUM. 
Costa Rica: Boruca, Tonduz 4460. Puerto Limon, Hitchcock 8421. Llano 
Grando de Puriscal, Jimenez 881. Hacienda (ie Gu&cimo, Tonduz 27. San 
Josf, Pittier 1183. Echcverrfa, Piliier <fe Tonduz 2479. Carrillo, Biolley 
3107, Pi/t.kr 1182. Talamanca, Tonduz 8557. River Torres, San Francisco, 
Jimenez 126. Tufa, Tonduz 11396. 
Turrialba, Tonduz 4092, 8244. 
Buenos Aires, Tonduz 4042. Rio 
Tub, 'Tonduz 8187. Boca Zhor- 
quln, Tonduz 8557, Quebrada de 
Tocorf, Tonduz 7542, Santo Do- 
mingo de Golfo Dulce, Tonduz 
10099. Piedra del Convento, Pit- 
tier 3650. 
Panama: Boca del Toro, Hart 73. 
Mount Hope, Hitchcock 9172. Be- 
tween Mindi and Co Kin, Hitchcock 
7943. Between Bohlo and Frijoles, Hitchcock 8395. Gorgona, Ainer. Gr. 
Nat. Herb. 58. 
Trinidad: Arima, Hitchcock 10297, 10306. Cumuto Station, Hitchcock 10077. 
Port of Spain, Hitchcock 9953. Tabaquite, Hitchcock 10132. Caparo, Broad- 
way 2837. 
Tobago: Mason Hall, Broadway 4476. 
Colombia: Santa Marta, Smith 206, 2190, 2747. 
51. Panicum milleflorum ep. nov. 
PrcfiCRIITION. 
Plants perennial; culms ascending «ir erect from an elongated creeping base, rooting 
at the nodes, glabrous, 0.8 to 2 meters high, 4 to 5 mm. thick, compressed, the nodes 
pubescent or glabrous, usually with one to several long erect panicle-bearing branches: 
sheaths about as long aa the internodes, keeled toward the summit, ciliate on the 
margin and puberulent at the junction with the blade, otherwise glabroua or sparsely 
papillose-hispid; ligule wanting, the ligular region puberulent; blades flat, 20 to 40 cm. 
long, 1 to 2 cm. wide, rarely wider, long-acuminate, somewhat narrowed to the sub- 
cordate base, glabrous or very sparsely hirsute, the margin scabrous; panicles 20 to 35 
cm. long, about 6 cm. wide, in outline tapering to both ends, consisting 
of numerous ascending or spreading, often curving, slender spikelike 
branches, solitary or in fascicles along an elongated glabrous axis, the 
lower distant, the upper approximate, the scabrous-angled, often 
sparsely pilose rachises bearing throughout their length along their 
lower side evenly disposed short approximate densely flowered branch- 
lets, those of the lower branches sometimes as much as 15 mm. long, the 
others 1 or 2 mm. long; spikelets subsessile, 1.3 mm. long, about 0.5 mm. 
wide, glabrous, the first glume about one-third the length of the spikelet, 
the second glume and sterile lemma equal, the lemma inflated boat- 
shaped with a large membranaceous palca; fruit 1.2 mm. long, 0.4 mm. wide, acute, 
t he lemma boat-shaped. 
Type in the U. S. National Herbarium, no. 093327, collected in water of swamp, 
Frijoles, Canal Zone, Panama, October 12, 1911, by A. S, Hitchcock (no. 8387). 
This species differs from Panicum pilosum in its larger size, its strong decumbent 
rooting base, and its longer blades and panicles. It appears to be the same as Panicum 
distichum tancifolium Griseb.1 which was previously included2 under P. pilosum. 
1
 Fl. Brit. W. Ind. 548. 1864. 3 Contr. U. S. Nat. Herb. 15: 114.1910. 
t iq. 69.—Distribution of P. polyganatuvi. 
Fig. 70. —r, 
miiltfioriim. 
From type 
specimen. 
HITCHCOCK AND CHASE TROPICAL NORTH AMERICAN PANICUM. 495 
Grisebach's variety was based upon Crueger 84 from Trinidad. This has strongly 
appressed-hispid nodes and sparsely pilose panicle branches. Hitchcock's no. 8387, 
the type, has only moderately pubescent nodes and no hairs on the branches of the 
panicle. Hitchcock's no. 7942 has glabrous nodes, evidently, though sparsely, pilose 
blades, and strongly pilose panicle branches. The three collections appear, however, 
to belong to the same species. Frijoles, the type locality, is now covered by the water 
of Gatun Lake. 
DISTRIBUTION. 
Swamps, Panama to Trinidad. 
Panama: Frijolcs, Canal Zone, 
Hitchcock 8387, Between 
Mindf and Col6n, Canal Zone, 
Hitchcock 7942. 
Trinidad: Without locality, Crue- 
ger 84. 
Fio. 71.—Distribution of P. milleflorum. 
52. Panicum pilosum Swartz. 
Panicum pilosum Swartz, Prodr. Vcg. Ind. Occ. 22. 1788; Contr. U. S. Nat. Herb. 
15: 113.1910. 
DISTRIBUTION. 
Moist ground, ditches, and swamps, Mexico and the West Indies to Paraguay. The 
type specimen from Jamaica. 
Veeacruz: C6rdoba, Amcr. Gr. Nat. Herb. CO. Mirador, Liebmann 411. 
Tabasco: San Juan Bautista, Rovirosa 599. 
Chiapas: Ocuilapa, Nelson 3050. 
British Honduras: Manatee Lagoon, Peek 28 (Gray Herb.). 
Guatemala: Finca Trece Aguas, Goll 11. Cubilquitz, Turckheim 8797. Puerto 
Barrios, Hitchcock 9159. Los An- 
des, KeHerman 5119. 
Honduras: San Pedro Sula, Thieme 
5587 in part. Bonacco Island, 
Gaunter in 1887. 
Nicaragua: Wright, U. S. Pacific 
Expl. Exped. 1853^56. 
Costa Rica: IUo Hondo, Cook & 
Doyle 499. Talamanca, Tonduz 
9495. Puerto Viejo, Biolley 7463. 
Fig. 72. Distribution of P. pilot um. Puntaremw, Hitchcock 8564. 
Panama: Chagros, Fcndler 368. Chepo, Pittier 4536. Puerto ObaJdfa, Pittier 
4371. Toro Point, Hitchcock 8044. Taboga Island, Hitchcock 8083. Balboa, 
Hitchcock 8016. Gatun, Hitchcock 8029. Cafia, Williams 776. David, Hitch- 
cock 8347. Culebra, Hitchcock 7897, 9208, Pittier 2082. Between Bohfo and 
Frijoles, Hitchcock 8390. Between Pedro Miguel and Corozal, Hitchcock 7992. 
Cliorrera, Hitchcock 8139. Porto Bello, Pittier 2481. 
Cuba: Dayaniguas, Wright 3451 in part. Sancti Spiritus, Le6n 908. Herradura, 
Van Hermann 763, Tracy 9063. Isle of Pines, Cwrtiss 305, Taylor 36. Sumi- 
dero, Shafer 13505, 13533. Buenaventura, Wilson 9444. Banao Hills, Le&n 
3982. Near Marianao, Le6n 1977. Without locality, Poppig 1834. 
496 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
Jamaica: Troy, Hitchcock 9782. Ramble, Hitchcock 9517. Bull Head Mountain, 
Hitchcock 9548. Castleton Gardens, Hitchcock 9407. Ewarton to Linstead, 
Hitchcock 9423. Ipswich, Hitchcock 9626. Navy Island, Millspaugh 1859. 
Port Morant, Hitchcock in 1890. Portland, Harris 31524. Castleton, Harris 
11280a. Gordon Town, Hart 732. Cedar Hurst, Harris 11547. 
Leeward Islands: St. Kitts, Britton Co well 287 (K. U. Herb.). Guadeloupe, 
Duss 4154. Dominica, Imray 151312, Jones 20. 
Windward Islands: St. Vincent, Eggers 6562, 6633 (K. U. Herb.). Grenada, St. 
Georges, Broadway in 1904. Martinique, Duss 534,4018 (both in K. U. Herb.). 
Barbados, Eggers 7195 (K. U. Herb.). 
Trinidad: Piarco Savanna, ITitchcock 10358. Port of Spain, Hitchcock 9961, 
10045, Hart 3293. Tabaquite, Hitchcock 10121. Cedros, Airier. Gr. Nat. Herb. 
59. Arima, Hitchcock 10309. 
Tobago: Brushfield, Eggers 5534. Scarborough, Hitchcock 10208. Castara, Broad- 
way 4063. Center of island, Hitchcock 10265. Spey Side, Hitchcock 10242. 
Colombia: C6rdoba, Pittier 557. Santa Marta, Smith 203. 
53. Fanicum boliviense Hack. 
Pancium boliviense Hack. Repert. Nov. Sp. Fedde 11: 19, 1912. "Bolivia: Anta- 
huacana, Espirito Santo in alveo arenoso fluminis * * * leg. Dr. O. Buchtien no. 
2501." A portion of the type was sent by Prof. Hackel for deposit in the National 
Herbarium. 
DESCRIPTION. 
Plants perennial with long decumbent or creeping bases, rooting and branching at 
the lower nodes; flowering culms ascending, 0.5 to 1.5 meters high, glabrous, the nodes 
glabrous; sheaths short, glabrous below, usually hirsute toward the summit, especially 
at the junction with the blade, the margins ciliate; ligule a ciliate membrane about 
0.5 mm. long; blades flat, 8 to 15 cm. long. 1 to 2 cm. wide, gradually 
narrowed from the cordate-clasping base to an acuminate apex, ciliate 
at base, otherwise glabrous, obscurely cross-nerved between the veins, 
the margins scabrous; panicle usually short-exserted, 10 to 25 cm. 
long, one-third to half as wide, the solitary or fascicled branches 
increasingly approximate toward the summit of the slender angled 
scabrous axis, usually pilose in the axils, the lowermost branch soli- 
tary and remote, all but the uppermost compound, the branchlets 
somewhat spikelike, rather distant, usually along the lower side of 
the rachis, the axils pilose; spikelets subsessile, clustered, 1.5 to 1.6 
mm. long, about 0,7 mm. wide, turgid, the first glume nearly half the length of spike- 
let, the second glume and sterile lemma equal, the sterile lemma somewhat inflated, 
subtending a membranaceous palea; fruit 1.4 mm. long, 0.6 mm. wide, subacute. 
Several collections of this species were referred in the Revision1 to P. laxurn as 
exceptionally robust specimens with cordate blades and turgid spikelets, 
DISTRIBUTION. 
Ditches, banks of streams, moist open 
or wooded ground, southern Mexico and 
Cuba to Paraguay. 
Veracruz: Veracruz, Hitchcock 
6582. Mirador, Liebmann 419. 
Cdrdoba, Finck 3, Hitchcock 
6435,6457. Zacuapdn, Purpus 
2159, 2160. Misantla, Purpm 
5980. Jalapa, Hitchcock 6666. - no. 74.-Distribution ol P. boliviense. 
1
 Contr. U. S. Nat. Herb. 16:116.1910. 
Fig. 73.—P. boliv~ 
irme. From type 
specimen. 
HITCHCOCK AND CHASE TROPICAL NORTH AMERICAN PANICUM. 497 
Guatemala: Cobdn, Turckheim, 1254. 
Panama: Along ditch to Old Panama, Hitchcock 8400, 
Cuba: Sancti Spiritus, Le&n 909. Isle of Pines, Curtiss 464. Bafios San Vicente, 
Britton & Gager 7452. 
54. Panicum laxum Swartz. 
Pantcum laxum, Swartz, Prodr. Veg. Ind. Occ. 23. 1788; Contr. U, S. Nat. Herb. 
15: 115. 1910. 
DISTRIBUTION. 
Ditches, banks, moist woods and wet savannas, Mexico and West Indies to Paraguay. 
The type specimen from Jamaica. 
Sinaloa: Culiac&n, Palmer 1558 in 1891. 
Veracruz: Coatepec, Hitchcock 6665. Jalapa, Hitchcock 6624. C6rdoba, Hitch- 
cock 6421, 6433. Veracruz, Hitchcock 6583. Sanborn, Orcutt 3238. Mirador, 
Liebmann 412. Orizaba, Botteri 688. 
Tabasco: Mayito, Rovirosa 427, 
British Honduras: Manatee Lagoon, Peck 60 (Gray Herb.). 
Guatemala: Puerto Barrios, Pittier 3d, Hitchcock 9160. Cubilquitz, Turck- 
heim 8803. Santa Rosa, neijde & Lux 3900. Cobdn, Turckheim 3832. 
Honduras : San Pedro Sula, Thieme 5587. 
Salvador: Sonsonate, Hitchcock 8971. 
Nicaragua: Corinto, Hitchcock 8748. 
Costa Rica: Buenos Aires, Tonduz 4864, 4868, 4871, Pittier 3657. Rfo Birrfs, 
Pittier 3117. Turrialba, Ton- 
duz 8267, Pittier 16123. Rfo 
Grande de T6rraba, Tonduz 
3622. Boruca, Tonduz 4473. 
Carrillo, Biolley 3107, Port 
Limon, Hitchcock 8429. San 
Jos6, Hitchcock 8455. Rfo 
Maravilla, Alajuela, Jimenez 
141. Puntarenas, Hitchcock 
8566. Rfo Bebedero, Jime- 
nez 740. Rfo Tilirf, Tonduz 
3071. 
Panama: Gatun, Hitchcock 7968, 8027. Boca del Toro, Hart 86. Porto Bello, 
Pittier 2443. Culebra, Hitchcock 7900. David, Hitchcock 8346,8350. Chepo, 
Pittier 4615. Vicinity of 016, Pittier in 1911. Garaboa, Pittier 4789. Along 
Sambu River, Pittier 5234. Chorrera, Hitchcock 8134. Between Pedro 
Miguel and Corozal, Hitchcock 7993. Dolega, Hitchcock 8334. 
Cuba: Sancti Spiritus, Le&n 907, Sergius 2781, Poppig 6. Han&bana, Wright 
189, 196, 197. Isle of Pines, Taylor 37, Palmer & Riley 1069. Herradura, 
Caldwell <Se Baker 7136, Tracy 9054, 9062, 9072, 9099, Hitchcock 177. Santiago 
de las Vegas, Hitchcock 178, Tracy 9114. Banao Hills, Le&n 3982$. Guana- 
bacoa, Le6n 914. Wajay, Earle Wilson 343. Zaza del Sur, Sergius 2780. 
Camaguey, Sha/er 217. Sumidero, Shafer 13724. Los Palacios, Shafer 
11793. La Gloria, Shafer 174. Without locality, Wright 759, 3451, 3863. 
Jamaica: Bull Head Mountain, Hitchcock 9539, 9545. Castleton Gardens, Hitch- 
cock 9397. Ewarton to Linstead, Hitchcock 9461. Appleton, Hitchcock 9658. 
Gordon Town, Hitchcock 9333. Troy, Hitchcock 9786, Castleton, Harris 
11285. Below Flamstead, Harris 11466. 
Santo Domingo: Jarabacoa, Eggerz 2120. 
82472°—15 4 
Fig. 75.—Distribution oI Pm laxum. 
498 CONTRIBUTIONS FBOM THE NATIONAL HERBARIUM. 
Porto Rico: Between Aiboiiito and Cayey, Chase 6341, Heller 522. Arecibo, 
Chase 6440- Sierra Luquillo, Hioram 368, Chase 6713. Ad juntas Road, 
Heller in 1902. Catafio, Heller 1378. Utuado, Britton & CowellSM. Maya- 
guez, Chase 6155, 6185, Sintenis 360. Along Rio Coamo, Chase 6549. Tru- 
jillo Alto, Chase 6363, 6775. Jajome Alto, Chase 6753. Santurce, Chase 
6354. Mount Morales, Britton & Marble 1068. Fajardo, Sintenis 1254. 
Danish West Indies: St. Thomas, Eggcrs 874 (K. U, Ilerb.). 
Leeward Islands: Antigua, Wullschlaegel 623. Guadeloupe, Duss 3179. 
Dominica, Jones 21, 25. 
Windward Islands: Martinique, Duss 535, 716 (K. TJ. Herb.). Barbados, Dash 
450. St. Lucia, Voyage of the Albatross in 1887-88. Grenada, Broadway 1870, 
3712, 4668. 
Trinidad: Port of Spain, Hitchcock 9984. Piarco Savanna, Hitchcock 10343, 
10351, 10361. Brighton, Hitchcock 10098. Pitch Lake, Hitchcock 10091. 
Cumuto Station, Hitchcock 10068. Cedroe, Hitchcock 10149, Bot. Card. Herb. 
2177, 2289. 
Tobago: Center of island, Hitchcock 10272. Spey Side, Hitchcock 10245. 
Venezuela: Llanos del Alto A pure, Jahn 201. 
Colombia: C6rdoba, Pittier 521, 553. Calf, Pittkr 665. Santa Marta, Smith 
202, 204. 
55. Panicum stevensiaixum Bp. nov. 
description. 
Plants perennial, tufted, glabrous throughout; culms compressed, 30 to 60 cm. long, 
spreading, prostrate, rooting at the nodes, the flowering ends and the branches ascend- 
ing; sheaths loose, much shorter than the interned es, compressed, 
keeled, dilate on the overlapping margin; ligule a delicate mem- 
brane about0.5 mm. long; blades flat, rather firm, more or lees spread- 
ing, 4 to 10 cm. long, 5 to 10 mm. wide, tapering from a little above 
the sparsely ciliate, truncate-cordate base to an acuminate apex; 
panicles terminal on the culm and branches, nhort-exserted, 6 to 
10 cm. long, narrow, consisting of several appressed or ascending 
spikelike branches along an angled axis, the lower branches some- 
what remote, 1.5 to 3 cm. long, usually longer than the interned es, 
the upper approximate, all spiketet-bearing to the base, the short- 
pediroled ppikoleta in crowded clusters, not secund, along a slender 
scabrous rachis; spikelets about 2 mm. long and 0.8 mm. wide, acute, 
somewhat inflated and gaping; first glume about one-fourth the length of the spikelet, 
sub acute, 3-nerved; second glume and sterile lemma equal, pointed, slightly exceed- 
ing the fruit, 3 to 5-nerved, the lemma 
inclosing a large palea and abortive 
perfect flower; fruit 1.9 mm. long, 0.6 
mm. wide, minutely scabrous at the 
acute apex. 
Type in the U. S. National Herbar- 
ium, no. 693323, collected "on wet 
sand around pool, forming a dense car- 
pet, white sand region, Campo Alegre 
near Laguna del Tortuguero," Porto 
Rico, November 25, 1913, by Agnes 
Chase (no. 6616). 
This species appears to be most nearly related to the South Ameiican P. milioides 
Noes, from which it differs in the short erect panicle branches, spikelet-bearing to 
the base. 
Fig. 76.—P. sle- 
vens lanum. 
From typo 
specimen. 
Fiq. 77.—Distribution of P. xtcvemianurn. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 499 
None of the specimens collected is mature. It may be that the large sterile palea 
becomes indurated at maturity as in P. hums and its close allies. The species is 
named in honor of Dr. F, L. Stevens of the University of Illinois, who, during the 
three years he was dean of the College of Agriculture and Mechanic Arts of Porto Rico, 
was an ardent student of the flora of the island and who contributed greatly to the 
success of a botanical trip made by one of the authors to Porto Rico in 1913. 
Since this work went to press a second collection of this species has been re- 
ceived: Wilson 9558, Laguna de Castellano, Cuba. 
Panicum exiguijlorum Griseb. Cat. PI. Cub. 234,1866; Contr. IT. S. Nat. Herb. IB: 
117, 1910. 
Savannas and moist sandy woods, Bahamas and Cuba. The type specimen from 
Cuba. 
Bahamas: Fortune Island, Hitch- 
HQ. Panicum esriguiflorum Griseb. 
DISTRIBUTION. 
Cuba: Han&bana, Wright in 1865. 
Sancti Spiritus, Sergim 2566. 
Obispo Hill, Le6n 902b. Zaza 
del Medio, Le6n 902. Guana- 
bacoa, Le6n 911, 913. Chiri- 
gote, Wright 3450. Herradura, 
Hitchcock 17 Tracy9075. Isle 
of Pines, Taylor 35. Without 
locality, Wrights, 3450,3877. 
cock in 1890. 
Fig. 78.—Distribution of P. exiffuiflorum. 
67. Panicum hians Ell. 
Savannas and moist soil along ponds and 
streams, North Carolina to New Mexico and 
in Veracruz, Mexico. The type specimen 
from South Carolina. 
Panicum Mans E1I. Bot. S. C. & Ga. 1: 
118. 1816; Contr. U. S. Nat. Herb. IB: 118. 
1910. 
DISTRIBUTION. 
Veracruz: Jalapa, Sch&ede. Without 
Fig. 79.—Distribution of i\ Ataiw. locality, MUller 2095. 
Only known from the type collection, in 
"wet hollows in prairies of Flor de Maria* 
State of Mexico," Pringle 3449. 
58. Panicum cupreum Hitchc. & Chase. 
Panicum cupreum Hitchc. & Chase, Contr. 
U. S. Nat. Herb. 15: 120.1910. 
DISTRIBUTION. 
Fie. 80.—Distribution of P. cupreum. 
500 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
STOLONIFERA. 
Spikelets hispid and with 2 crateriform glands on the sterile lem- 
ma; second glume and sterile lemma not boat-shaped. 
Spikelets not over 2 mm. long; blades not over 4 cm. long.. 61. P. pulchelhan. 
Spikelets 3.6 mm. long; blades 4 to 10 cm. long  62. P. biglandulare. 
Spikelets glabrous, glandless; second glumejand sterile lemma 
boat-shaped. 
Blades not over 5 cm., usually 2 or 3 cm., long; second glume 
rather blunt and shorter than the sterile lemma 59. P. stoloniferum. 
Blades 5 to II cm. long; second glume acute, nearly equaling 
the sterile lemma ..   60. P. J'rondescens. 
69. Panicum stoloniferum Foir. 
Panicum stoloniferum Poir. in Lam. Encycl. Suppl. 4: 274.1816; Contr. U. S. Nat. 
Herb. 10: 121. 1910. 
DISTRIBUTION. 
Moist shady places, Guatemala to Brazil. 
Fig. 81.—Distribution of P. stoloniferum. 
The type specimen from Cayenne. 
Guatemala: Puerto Barrios, Pittier 
364. 
Panama : Frijolee, Amer. Gr. Nat. 
Herb. 62. 
Windward Islands: Martinique, 
Dugs 1291 (K. TJ. Herb.). 
Trinidad : St. Anns Valley, Bot. Gard. 
Herb. 2293. Aqua Santa Arima, 
Broadway 2370. Without local- 
ity, Crueger 79. 
60. Paxticum frondescens Meyer. ' 
Panicumfrondescens Meyer, Prim. 11. Esseq. 56. IS 18; Contr. U. S. Nat. Ilerb. 15: 
121.1910. 
Hymenachne frondescens Fourn. Mex. PI. 2 : 36.1886. Based on Panicum frondescens 
Meyer. 
A specimen in the Berlin Herbarium, Sclcr 5024, without locality other than Mexico 
and Central America, is exceptionally lax and is peculiar in having occasional spike- 
lets with glands on the sterile lemma aa in P. pulchelhan and P. biglandulare. In 
most of the spikelets there are a few liairs along the margin of the sterile lemma. Fur- 
ther collections may show this to be a distinct species. In Pittier 6328, cited below, 
a few spikelets bear glands, but all are glabrous. 
distribution. 
Moist woods, Mexico to Brazil. The type specimen from British Guiana. 
Veracruz: Papantla, Liebmann 
405. 
Tabasco: Sebastian, Rovirosa 
497. 
Guatemala: Dept. Petdn, Walker 
1138. 
Costa Rica: Hacienda de Zent, 
Tonduz 259. 
Panama: Trinidad Basin, Pittier 
4027. Fig. 82.—Distribution of P./roudesccus. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 501 
Windward Islands: Smith & Smith 187 (K. U, Herb.). 
Trinidad: Aqua Santa Arima, Broadway 2371. Blanchisseuse, Broadway 2390, 
Without locality, Crueger 85. 
Venezuela: Above Paparo, Pittier 6328. 
Colombia: Santa Marta, Smith 2126, 2571. 
61. Panicum pulchellum Raddi. 
Panicum pulckellum Raddi, Agrost. Bras. 42. 1S23; Contr. TJ. S. Nat. Herb. 15: 
123. 1910. 
Uymenachne leptostachya Fourn. Mex. PI. 3: 3G. 1880. Based on Panicum leptosto 
chyurn Presl. 
DISTRIBUTION. 
Moist shady places, southern Mexico and the Windward Islands to Brazil. The 
type specimen from Rio de Janeiro, 
Veracruz: C6rdoba, Hitchcock 6444, Bourgeau 1455. Minatitl&n, Smith 589. 
British Honduras: Manatee Lagoon, Peck 279 (Gray Herb.). 
Guatemala: Cubilquitz, Tiirckheim 7702, 8794. Secanquim, Cook & Griggs 
284, Maxon & Hay 3153. El 
Palmar, Kellerman 6246. 
Costa Rica: San Francisco, Jime- 
nez 162. San Mateo, Biolley 
7001. Desamparados, Tonduz 
1482, Boruca, Tonduz 4459, 
4460. El General, Pittier 
3362. Piedra del Convento, 
Pittier 3656. Cafias Gordaa, 
Pittier 7360. Buenos Aires, 
Tonduz 4881, 
Panama: Between Panama and 
Fig. 83.—Distribution of P. pulcheUum. 
Corozal, Amer. Gr. Nat. Herb. 63. Culebra, Pittier 2226. Cerro Vaca, 
Pittier 5375. 
Windward Islands: Martinique, Duss 767 (K. U, Herb.). 
Venezuela: C&rdenas, Pittier 5975. 
Colombia: Santa Marta, Smith 2127. 
62. Panicum biglandulare Scribn. & Smith. 
Panicum Mglandulare Scribn. & Smith, U. S. Dept. Agr. Div. Agrost. Bull. 4:13. 
pi, 4. 1897; Contr. U. S. Nat. Herb. 
15:123. 1910. 
distribution. 
Among bushes, southern Mexico and 
Guatemala. The type specimen from 
Chiapas. 
Chiapas: Pinabete, Nelson 3781. 
Guatemala: Cob&n, Tiirckheim 
II. 1342, II. 1956. 
Fig. 84.—Distribution of P. biglanduiare. 
502 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM 
PARVIGLUMIA. 
Fruit glabrous  66. P. parviglume. 
Fruit with scattered appressed silky hairs. 
Culms creeping, with ascending flowering branches, not over 
0.5 meter high; blades falcate 64. P. schmitzii. 
Cuhns clambering, usually 1 to 3 meters long; blades not 
falcate. 
Panicles 2.5 to 7 cm. long; blades n<H over 1 cm. wide, 
symmetrical at base 63. P. virguliorum. 
Panicles 10 to 15 cm. long; blades 1.2 to 2.5 cm. wide, 
unsymmetrical at base  65. P. schiffneri. 
63. Pajiicum virgultonun Hack, 
Panicum virguliorum Hack. Oesterr. Bot. Zeitschr. 51: 369.1901; Contr. U. S. Nat. 
Herb. 15:125,1910. 
DISTRIBUTION. 
Hedge rows, brushy banks, and cultivated fields, southern Mexico to Panama. The 
type specimen from Costa Rica. 
Veracruz: Jalapa, Hitchcock 6630. 
Cdrdoba, Hitchcock 6441. Ori- 
zaba, Amer. Gr. Nat. Herb. 
64. 
Guatemala: Guatemala City, Hitch- 
cock 9074. Chacula, Seler 2708. 
Cob&n, Tiirckhevm 3788. 
Oosta Rica: San Jos6, Hitchcock 
8490. Alajuelita, Tonduz 8818, 
8829. Tres Rfos, Pittier & 
Fig. 85.—Distribution of /'. viryvltorum. 
Tonduz 4326. San Francisco, Rio T6rres, Jimenez 49. San Juan, Jiminez 
919. 
Panama: El Boquete, Hitchcock 8317. 
64. Panicum, schmitzii Hack. 
Panicum schmitzii Hack. Ann. Naturhist. Hofm. Wien 17: 254. 1902; Contr. XI. S, 
Nat. Herb. 15:125.1910. 
DISTRIBUTION. 
Shaded rocky slopes, central and 
southern Mexico. The type specimen 
from Mexico. 
San Luis PoTosf: Laa Canoas, 
Pringle 3817. 
Veracruz: GSrdoba, Hitchcock 
6423, 6446. Orizaba, Amer. 
Gr. Nat. Herb. 65. Fig. 86.—Distribution of P. schmitzii. 
65. Panicum schifEneri Hack. 
Panicum sckiffneri'Eack. Ergeb. Bot. Exped. Akad. Wise. Stidbras. 11.1906; Dent 
Bchr. Kais. Akad. Wiss. Wien 70: 72.1908. The type specimen, in Hackel's herbar- 
ium, was collected in southern Brazil bv Wettstein and Schiffner in 1901. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 603 
Fig. 87. — P. schiffneri. From 
type specimen. 
DESCRIPTION. 
Plants perennial with long branching decumbent base, rooting at the nodes; culms 
straggling, ascend ing, 2 to 3 meters long, slender, more or leas compressed, smooth or 
hirsute or roughened below the nodes, producing long divaricate branches; sheaths 
papillose-hispid especially toward the summit, sometimes glabrate below, densely 
pubescent at the junction with the blade; ligule membranaceous-ciliate, about 0.5 
mm. long; blades flat, rather firm, usually horizontally spreading, 10 to 15 cm. long, 
1.2 to 2.5 cm. wide (rarely larger), narrowed to the usually unsymmetrical base, 
gradually tapering from below the middle to an acumi- 
nate apex, scabrous and sometimes sparsely hispid on 
the upper surface, pubescent above the ligule, scab- 
erulous or puberulent beneath or sometimes sparsely 
hispid, especially along the midnerve, the fine white 
margin undulate, scabrous; panicle 10 to 15 cm. long, 
usually about as wide, the few slender scabrous branches 
remote, pilose in the axils, the lower branches solitary or 
in pairs, widely spreading or reflexed, more than half as 
long as the very scabrous main axis, naked at the base, 
the upper branches much shorter, ascending, the rather densely flowered short 
branchlets appressed along the upper half or two-thirds of the branches, the bract 
at the base of inflorescence usually well developed; spikelets sliort-pediceled, aggre- 
gated, scarcely 2 mm. long, 1 mm. wide, turgid, glabrous; first glume less than one- 
fourth aa long as the spike let; second glume and sterile lemma equal, exceeding the 
fruit, obscurely nerved; fruit about 1.5 mm. long, 0.9 mm. wide, elliptical, sparsely 
covered with long appressed silky hairs. 
This species resembles Panicum parviglume in habit and P. schmitzii and P. virgul- 
torum in spikelet characters. It has longer, more straggling culms than has any other 
species of this group, and larger blades than any except P. parviglume. 
DISTRIBUTION. 
Wet shady banks and slopes, Porto Rico, Windward Islands, and southern Mexico 
to southern Brazil. 
Veracruz: C6rdoba, Amer, Gr. Nat. 
Serb. 66, distributed as P. par- 
viglume. 
Guatemala: Cob&n, Turckheim II. 
1326. 
Costa Rica: San Francisco de 
Guadalupe, Jirn&nez in 1910. 
Panama: El Boquete, Hitchcock 
8278, 8305. 
Porto Rico: Maricao, Chase 6198. 
Indiera Fria, Chase 6247. Vi- 
cinity of Cayey, Chase 6745. 
Alto de Bandera, Chase 6474. 
Windward Islands: Martinique, Hahn 616. St. Vincent, Eggers 6653, Smith 
& Smith 1099 (K. TJ. Herb.). 
66. Panicum parviglume Hack. 
Panicum, parviglume Hack. Oesterr. Bot. Zeitschr. 51: 429. 1901; Contr. U. S. Nat. 
Herb. 16:126.1910. 
Panicum conchatum Fourn. Mex. PI. 2: 25. 1886. This was previously included 
among the doubtful species.1 A few spikelets from Schaffner's no. 204 (the type 
FiG. S3.—Distribution of P, schiffneri. 
1
 Contr. U. S. Nat. Herb. 15:329.1910. 
504 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
collection) in the Berlin Herbarium are in the National Herbarium. A restudy of 
these together with later collections show that they are referable to P. parviglume, 
Fournier's description ia vague and inadequate and appears to have been based on 
more than one species, the vegetative characters as described being unlike those of 
any species in this group. Because of this uncertainty the name P. concha turn is not 
taken up to replace P. parviglume. 
Panicum arundinariae Trin.; Fourn. Mex. PI. 2:25.1886. This name is also included 
among the doubtful species.1 Mtiller's no. 2018, one of the specimens cited by 
Fournier, is P. parviglume, while another, Schaffner 279, also cited, is Panicum vir~ 
gultorum. Aa in the preceding case, the description is inadequate, for which reason 
the name P. arundinariae is not taken up. 
u
—i-/ j \Y 0 
S\v* i \ \ 1 
M crn \ 
Flfl. 8D.—Distribution of'P. parviglume. 
Guadalupe, Tonduz <t Pittier 8448, Jiminez 113. San Jos£, Hitchcock 
8487. 
TRICHOIDIA. 
One of the two species of this group, Panicum trichanthum, ia found to be a perennial. 
Blades more than one-fourth as wide as long; spikelets pubes- 
cent; plants annual  67. P. trichoides. 
Blades less than one-eighth as wide as long; spikelets minutely 
bullate-rugoee; plants perennial  68. P. trichanthum. 
07. Panicum trichoides Swartz. 
Panicum trichoides Swartz, Prodr. Veg. Ind. Occ. 24. 1788; Contr. U. S. Nat. Herb. 
15: 129.1910. 
DISTRIBUTION. 
Damp shady places, often a weed in fields and groves, tropical America. The type 
specimen from Jamaica. 
Sinaloa: I mala, Palmer 1758 in 1801. 
Tepic: Maria Mad re Islands, Nelson 4257. 
Jalisco: Guadalajara, Amer. Gr. Nat. Herb. 70, Pringle 2618, 3828. 
Colima: Manzanillo, Palmer 1083 in 1890. Colima, OrciUt 4515. Alzada, Hitch- 
cock 7110. 
MichoacAn : La Correa, Langlasse 380. Hacienda Ooahuayula, Emrick 53. 
Veracruz: Zacuap&n, Purpu$ 2902. Minatitldn, Smith 601. C6rdoba, Finch in 
1893. Sanborn, Orcutt 3241. 
Guerrero: Acapiilco, Palmer 287 in 1894. 
Oaxaca: Guatulco, Liebmann 317. 
Tabasco: Mayito, Rovirosa 434. San Juan Bautista, Rovirosa 598. 
YucatAn: Izamal, Gaumer 522. San Anselmo, Gaumer 895, 2027. 
1
 Contr. U. S. Nat. Herb. 15: 329.1910. 
distribution. 
Banks and ditches, southern Mexico 
to Costa Rica. The type specimen 
from San Jos6, Costa Rica. 
Veracruz: Orizaba, Bourgeau 
3197, Mutter 2018. Borrego, 
Botteri 150, 152. 
Costa Rica: Alajuelita, Jimenez 
403. San Francisco de 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 505 i 
British Honduras: Manatee Lagoon, Peck 314. Toledo, Peck 637 (both in Gray 
Herb.). 
GUATEMALA: El Palmar, Kellerman 6263. Santa "Rosa, Heyde & Lux42§§. Cubil- 
quitz, Tiirckheim 7801. Morales, Deam 6041. Bine one i to, Heyde & Lux 4299. 
Honduras: Amapala, Hitchcock 8768. Puerto Sierra, Wilson 89, 188. 
Salvador: Sonsonate, Hitchcock 8981. San Salvador, Velasco 10. La Uni6n, 
Hitchcock 8791, 
Nicaragua : Jinotepe, Hitchcock 8693. San Juan del Sur, Hitchcock 8604. Masaya, 
Hitchcock 8631. 
Costa Rica: Hacienda de Zent, Tonduz 363, Puffier in 1904. Atenas, Hitchcock 
8521. Puntarenas, Hitchcock 8555. Alajuola. Jimenez 145. San Jos6, Tonduz 
3133. Buenos Aires, Tonduz 4865, Pittier 3651. Matina, Pittier 9754. 
Nicoya, Cooper 10379, Tonduz 13754. Boruca, Pittier 4458. Hacienda de 
Chirrip6, Pittier 16081, Carrillo, Bioltey 3111. 
Panama: Culebra, Hitchcock 7933, 8022. Alajuola, Pittier 2352, Puerto Obaldfa, 
Pittier 4284. Below Chcpo, 
Pittier 4696. Culebra, Pittier i I   / i [ \/ ET 
2083. Tabernilla, Hitchcock 
8384. Taboga Island, Cdes- 
tine 84. 
Cuba: Habana, Le6n in 1910, 
Curtiss 714. Monte Verde, 
Wright 1538. Sancti Spiritus, 
Ledn 4105. 
Jamaica: Ramble, Hitchcock9475. 
Cedar Hurst, Harris 11566. 
Castleton, Harris 11299,11328. 
Kingston, Hitchcock 9281. Port Antonio, Fredholm 3282. Gordon Town, 
Hitchcock 9332. 
Porto Rico: Maricao, Chase 6191. Vega Baja, Chase 6417. Mayaguez, Chase 
6167, Sintenis 160, Cowell 522, 583. Ponce, Heller 6094. Luquillo Moun- 
tains, Wilson 283. Martin Pefia, Heller 387, Between Aibonito and Cayey, 
Heller 531. Mount Morales, Britton & Covxll 444. Santurce, Heller 157. 
Rio Piedras, Barrett 61. 
Leeward Islands: St. Kitts, Britton & Cowell 295. Guadeloupe, Dues 2681, 
Dominica, Eggers in 1881. Montserrat, Shafer 337, 706, Saba, Suringar in 
1885, Boldingh 2122 (K. U. Herb.). 
Windward Islands: Martinique, Hahn 1047, Duss 1321. Grenada, Broadway in 
1905, Eggers 5987. St. Vincent, Smith de Smith 185 (K. U. Herb.). 
Trinidad: Port of Spain, Hitchcock 9946. 
Venezuela: Above Paparo, Pittier 6494. Tovar, Fendler 2499. 
Colombia: Santa Marta, Smith 167. Sevilla, Pittier 1621. Call, Pittier 971. 
68. Panicuxn trichanthum Neea. 
Panicum trichanthum Nees, Agrost. Bras. 210.1829; Contr. TJ. S. Nat. Herb. 15:131. 
1910. 
The habit of this species as studied in the field indicates that it is a perennial. The 
culms are long and straggling with rootlets at the nodes and form a tangled mass among 
shrubs or vines. 
Fig. 90.—Distribution of P. tTicftoides* 
506 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
DISTRIBUTION, 
Moist thickets and river banks, Mexico, and the West Indies to Paraguay. The type 
specimen from Mexico. 
San Luis PoTosf: San Luis Potosf to Tampico, Palmer 1151 in 1879. 
Co LIMA: Palmer 1257 in 1891, 
Veracruz: Motzorongo, Smith 585. Veracruz, Hitchcock 0584. Jical tepee, IAeb- 
mann 320. Colipa, Liebimnn 432. Without locality, Miiller 2172. 
Campeche: Near Cham pot6n, Collins 37. 
British Honduras: Toledo, Peck 775 (Gray Herb.). 
Guatemala: Giialan, I)eam 424. Puerto Barrios, Hitchcock 9149. Morales, 
Kellerman 0272. Finea Trece 
Aguas, Lewton 377. Cubil- 
quitz, Tiirckheim 7798. 
Honduras: San Pedro Sula, Thieme 
5587. 
Nicaragua: Jinotepc, Hitchcock 
8069, 8682. 
Costa Rica: Port Limon, Hitch- 
cock 8434. Talamanca, Tonduz 
8600, 8070. 
Panama: LagunadeChiriqul, Hart 
87. Culebra, Hitchcock 7895. 
Frijoles, Maxon 4703. Panama, Hitchcock £>212. Tabernilla, Pittier 3824, 
Hitchcock 8383. Gamboa, Pittier 4798, Balboa, Hitchcock 8015. 
Cuba: Trinidad, Wright 753. Vento, Curtiss 598, Leon 557. Romelie, Eggers 
5350. 
Jamaica : Port Antonio, Hitchcock in 1890. 
Porto Rico: Cayey, SirUmu 2471, Chase (5748. Pcmiolas, Chase 6487. Rio Pie- 
dras, Johnston 143. 
Trinidad: Port of Spain, Hitchcock 9943. Cedros, Hitchcock 10146. 
Venezuela: Tovar, Fendler 1443. 
Colombia: Santa Marta, Smith 2151. 
PAR YIFOLIA, 
Perennials, usually glaucous; culms more or less decumbent; panicles small, diffuse, 
the epikelets small, turgid or sub globose, glabrous, the first glume more than half the 
length of the spikelet. 
Culms very slender, decumbent or creeping; blades 1 to 3 cm. 
long 69. P. parmfolium. 
Culms firm, erect or decumbent at base only; blades 3 to 8 cm. 
long  70. P. cyanescens, 
69. Panicum parvifolium Lam. 
Panicum parvifolium Lam. Tabl. Encycl. 1:173.1791; Contr. U. S. Nat. Herb. 15: 
134. 1910.' 
The blades in this species vary in shape from oblong or ovate-lanceolate to narrowly 
lanceolate, in surface from glabrous to densely appressed-hirsute, and in texture from 
thin to firm. One form, in aspect approaching P. cyanescens, has firm pubescent erect 
blades, as much as 5 mm. wide and 4.5 cm. long. This is represented by Broadway 
2372 in part and Ililckcock 10065, both from Trinidad. Panicum brasiliense Spreng. is 
a pubescent form with oblong-lanceolate moderately firm blades. It does not appear 
possible to recognize this form as a distinct species. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 507 
DISTRIBUTION. 
Fig. 92.—Distribution of P. parvifolium. 
Wet savannas and margins of ponds and streams, Costa Rica and the West Indies to 
Paraguay. The type specimen from tropical America. 
Cost a Rica : Buenos Aires, Tonduz 3631, 3659, Pittier 10594. 
Cuba: Los Almacigos, Wright 3458. Herradura, Baker 2078, Tracy 9060, 
9079, Hitchcock 181, Brit ton, 
Earle & Gager 6494. Pinar 
del Rfo, Britton & Gager 
7075. Guane, Shafer 10659. 
Laguna Los Indies, Shafer 
10803. 
Porto Rico: Lake Loisa, Chase 
6786. Campo Alegre, Chase 
6615,6788. Aguada, Sintenis 
5719. Guainabo, Chase 6630. 
Catano, Sintenis 5719. Mar- 
tin Pefia, Chase 6358. Trujillo 
Alto, Chase 6763. VegaBaja, Chase 6796, Heller 1316. 
Trinidad: Cumuto Station, Hitchcock 10065, Amer. Gr, Nat. Herb, 72. Pitch 
Lake, Hitchcock 10100. Arima, Broadway 2372, Without locality, Crueger, 
224. 
70. Panicum cyanescens Nees. 
Panicum, cyanescens Nees, Agrost. Bras. 220.1829, "Habitat in Brasilia meridionali 
(Sellow)." The type is in the Berlin Herbarium. 
Panicum firmifolium Trin,; Nees, loc. cit. A herbarium name giver, as a synonym 
of P. cyanescens. 
DESCRIPTION. 
Plants perennial, bluish or glaucous; culms tufted, erect or with a decumbent base 
rooting at the nodes, slender, smooth, leafy, 30 to 50 cm. high, branching from the mid- 
dle and upper nodes; sheaths striate, glabrous, rarely ciliate on the overlapping margin; 
ligule minute, membranaceous, sparsely cilia te with long hairs or sometimes naked; 
blades flat, rather firm, erect, spreading or refiexed, 3 to 8 cm. long, 4 to 5 mm. wide, 
oblong-lanceolate, slightly narrowed to the base, acute, 
glabrous; panicles short-exserted, terminal and often from 
the upper sheaths, 3 to 6 cm. long, as wide or wider, open, 
the slender flexuous branches rather remote, stiffly ascend- 
ing or spreading, naked below, branching and epikelet- 
bearing toward the ends, the spikelets on slender divaricate 
pedicels; spikelets about 1.5 mm. long, 0.9 mm. wide, tuigid 
or subglobose, obtuse, glabrous; first glume two-thirds to 
three-fourths as long as the spikelet; second glume and sterile 
lemma equal, covering the fruit, or at maturity the glume wrinkled because of the 
turgidily of the fruit, exposing the summit; fruit 1,2 mm. long, 0.8 mm. wide, very 
turgid, subacute, the lemma and palea cellular-roughened as seen under a lens, 
bearing a few very obscure appressed hairs toward the summit. 
This species resembles P. parvifolium, but differs in the less slender, more erect 
culms, longer blades, and stiffly ascending panicle branches, naked below. 
Fio. 93.—P. evanescent. 
From type specimen. 
508 CONTRIBUTIONS FBOM TIIE NATIONAL HERBARIUM. 
Fio. 94.—Distribution of P. cyaiwscens. 
DISTRIBUTION. 
Swampa and wet savannas, British 
Honduras and Trinidad to Brazil. 
British Honduras : Manatee La- 
goon, Peck 271 (Gray Herb.). 
Trinidad: FiarcoSavanna, Hitch- 
cock 10349,10354,10364. Cu- 
mu to Station, Hitchcock 10067. 
Aripo Savanna, Broadway 
2377. 
tTNGROTTPED SPECIES OF TRUE PANICUM. 
71. Panicum pyrularium Bp. nov. 
Plants annual; culms delicate, 10 to 13 cm. high, at first erect and simple, finally 
decumbent and branching, often rooting at the nodes, more or less zigzag, sometimes 
elongating to 25 or 3p cm.; nodes sparsely pilose or glabrous; sheaths shorter than the 
blades, glabrous or sparsely pilose; ligule mcmbranaceous, about 0.5 mm. long; blades 
thin, flat, spreading, 1 to 3 cm. long, 2 to 7 mm. wide (the lower sometimes smaller), 
narrowly elliptical, somewhat clasping at base, obtuse or acutish at the apex, scabrous 
on the margins, glabrous or sparsely pilose on both surfaces, sometimes glaucous; 
panicles terminating the culm and branches, loose and open, 2 to 5 
cm. long, about as wide, ova! or pyramidal in outline, the delicate 
fiexuous branches and branchlets spreading or refiexed, smooth, 
the capillary divaricate pedicels several times longer than the 
spikelets; spikelets 1.5 mm. long, 0.6 mm. wide, turgid, pyriform, 
long-attenuate at base, glabrous; first glume about half as long as 
the spikelet, acute; second glume and sterile lemma 5-nerved, not 
exceeding the fruit, the glume slightly shorter than the sterile 
lemma; fruit 1 mm. long, 0.5 mm. wide, turgid. 
Type in the U. S. National Herbarium, no. 715639, collected 
between Hato del Jobo and Cerro Vaca, eastern Chiriquf, Panama, altitude 700 to 
1,000 meters, December 25 to 28, 1911, by H. Pittier (no. 5416). 
Panicum pyrularium, one of the smallest and most delicate species in the genus, is 
not related to any North American species. 
The South American Panicum polycomum, Trin. is an allied species of more tufted 
habit, with much narrower blades, smaller panicles, and smaller pubescent spikelets 
not attenuate at base. 
None of the label* with the specimens cited below indicates the habitat. 
DISTRIBUTION. 
Panama to Venezuela. 
Panama : Between Hato del Jobo 
and Cerro Vaca, Ohiriquf, 
Pittier 5416. 
Ven ezuela : Tovar, Fendler 2502. 
Colombia: Santa Marta, Smith 
2570. Without locality, JVi- 
ana 281 (2811?). 
FiO. 93.—P. pyru- 
larium. From 
type specimen. 
Fig. yfi.—Distribution ol P. vyrularium. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 509 
72. Panicum haenkeanum Preel. 
Panicum haenkeanum Presl, Rel. Haenk. 1: 304. 1830. "Hab. in Mexico." The 
type specimen is in the herbarium of the National Museum at Prague. The only 
datum on the label is the word "Mexico." 
Panicum costaricense Hack. Oesterr. Bot. Zeitschr. 51: 428. 1901; Contr. IT. S. Nat. 
Herb. 16: 134.1910. 
For discussion of Panicum expansum Fourn,, mentioned in the Revision under P. 
costaricense, see page 525 under P, cordovense. 
Field study of this species shows that it is a perennial with straggling culms as 
much as 2 meters long, often rooting at the lower nodes. 
DISTRIBUTION. 
Moist wooded or grassy banks and slopes, Mexico to Panama. 
Costa Rica: T&raba, Tonduz 3628, 
3636, 3673. Boruca, Pittier 
4626. Cordoncillal, Pittier 
3640. Buenos Aires, Tonduz 
3687, 4860, Pittier 3661. 
Panama: Culebra, Hitchcock 9168. 
Between Corozal and Ancon, 
Pittier 2169,2636. San Felix, 
Pittier 5247. Between Panama 
and Corozal, Hitchcock 9206. 
73. 
Fig. 97.—Distribution of P. hacnkeanuTtt. 
ineptum Bp. no v. 
DESCRIPTION. 
Fig. 98.—P. ineptitvt. From type 
specimen. 
Plants perennial (?); culms slender, apparently ascending, probably 50 em. or 
more long, slender, striate-fluted, glabrous or the upper internodea sparsely pilose, 
producing nearly simple branches as much as 25 
cm. long; nodes retrorsely pubescent; sheaths loose, 
shorter than the intemodes, pilose, the maxgins 
densely ciliate; ligule ciliate, about 0.5 mm. long; 
blades spreading, flat, 3 to 7 cm. long, 3 to 6 mm. 
wide, tapering from the truncate base to an acumi- 
nate apex, softly pilose on both surfaces; panicles 
terminal, those of the branches short-exeerted, 3 to 
5 cm. long, consisting of few to several short spread- 
ing densely flowered branches, remote along the 
slender pilose axis, the branches 4 to 6 mm. 
long, bearing 1 to 8 subsessile spikelets, the rachis pilose; spikelets 2.5 to 2.7 mm. 
long, 1.1 mm. wide, blunt; first glume about half as long as the spikelet, 3-nerved, 
obtuse, pubescent; second glume and 
sterile lemma equal, inflated, much 
larger than the fruit, the glumegibbous 
in the middle, 7-nerved, pubescent, 
the sterile lemma 3-nerved, glabrous, 
inclosing a membranaceous palea; 
fruit 1.6 mm. long, 0.9 mm. wide, 
elliptical, smooth and shining, the 
lemma strongly convex. 
Type in the herbarium of Krug & 
Urban in the Berlin Botanical Mu- 
seum, the ticket reading " 817 Panicum- 
Fig. 99.—Distribution of P. inepttim. 
•S. Domingo, Bertero. Hb. Spr." A slip in 
Sprengel' a hand bears the name'' Panicum nemoroaum Sw.'' [Ichnanthus nemoroms]. 
510 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, 
The type specimen, consists of a culm with four internodes, the base and summit 
wanting, bearing two flowering branches. No other collection of the species is known. 
It appears to bo related to Panicum vene&uelae Hack., from which it differs in the 
smaller panicle and in the smaller and pubescent instead of bristly hispid spikelets. 
74. Panicum xnillegxana Poir. 
Panicum millegrana Poir. in Lam. Encyd. Suppl. 4 : 278. 1816; Contr. TJ. S. Nat. 
Herb. 15 : 135. 1910. 
Panicum, patentmimum Desv.; Poir. in Lam. Encycl. Suppl. 4: 283.1816. "Cette 
pi ante croit & Saint-Dominique & & la Nouvelle-Espagne." The type from the her- 
barium of Desvaux in the Paris Herbarium is a fragmentary specimen consisting of a 
panicle with immature glabrous spikelets together with a few broken leaves. 
DISTRIBUTION. 
Damp woods and shady banks, Mexico and the West Indies to Paraguay. The type 
specimen from tropical America, probably from Cayenne. 
Veracruz: C6rdoba, Arner. Gr. Nat. Herb. 73. Jalapa, Hitchcock 6605, Holway 
3083. Mirador, Liebmann, 275. 
Oaxaca: Without locality, Galeotli 5728. 
Chiapas: Santa Rosa, Heyde <f? 
Lux 3927. 
Guatemala: Cubilquitz, Tiirck- 
heim 8783,8784. Cobdn, Tiirck- 
heim 657. Guatemala City, 
Tfitchcock 9056. 
Costa Rica: El General, Pittier 
10615. 
Panama: El Boquete, Hitchcock 
8299. 
Cuba: TTcrradura, Hitchcock 180, 
Tracy 9098. La Catalina, 
Wright 3455. Pinar del Rfo, Wright 3855. Sumidero, Sha/er 13562. Sierra 
do Cabra, Britton <t Gager 7201. 
Trinidad: Pitch Lake, Hitchcock 10087. Port of Spain, Hitchcock 9979. 
Venezuela: Tovar, Fmdler 1641. 
Colombia: Santa Mart,a, Smith 168, 2146. 
75. Panicum glutinosum Swarlz. 
Panicum glutinosum Swartz, Prodr. Veg. Ind. Ore. 24. 1788; Contr. U. S. Nat. 
Herb. 15: 138. 1910. 
DISTRIBUTION. 
Mountain woods, throughout tropi- 
cal America. The type specimen 
from Jamaica. 
Veracruz : Mirador, Liebmann 
428, Ross 701. Orizaba, 
Bourgcau 3192, Bolter i. 
Jalapa, Amer. Gr. Nat. Herb, 
74. Zacuap&n, Purpus 2156, 
2903, 
Fio. 101.—Distribution of P. glutinosum. 
HITCHCOCK AND CHASE—TROPICAL 2TORTH AMERICAN PANICUM. 511 
Chiapas: Turubula, Nelson 3357. 
Costa Rica: Helechales del General, Pittier 12002. Ca&aa Gordas, PUtier 11017. 
Cabagra, Tonduz 6547. Rio Birrfa, Pittier 3118. 
Cuba: Loma Pelada, Wright 757. La Perla, Ledn 3790. Banao Hills, Le6n 
3998. Gran Piedra, Shafer 9014. Without locality, Linden 2143. 
Jamaica: Gordon Town, Ilart 792, Hitchcock 9321. Troy,Maxon 2816, Hitchcock 
9785. Abbey Green, Hitchcock 9370. Cinchona, Hitchcoch 9708, Harri$ 
11264, 11440, 11508. Ewarton, Hitchcock 9451. Ipswich, Hitchcock 9627. 
Newport, Britton 3193. Road to Salt Hill, Harris 11395. Without locality, 
Wullschlaegel 1163. 
Haiti: Kenskoff, Buch 956. 
Santo Domingo: Picarda 620 (K. U. Herb.). 
Porto Rico: Sierra de Yabucoa, Sintenis 2609, Mayaguez, Sintenis 357. Alto de 
Bander a, Chase 6469. Maricao, Chase 6199, 6250. 
Venezuela: Caracas, Pittier 5910, 
76. Fanicum rudgei Roem. & Schult.1 
Panicum rudgei Roem. & Schult. Syst. Veg. 2: 444. 1817; Contr. U. S. Nat. Herb. 
15:139.1910. 
DISTRIBUTION. 
Savannas, Jamaica and British Hon- 
duras to Brazil. The type specimen 
from British Guiana. 
British Honduras: Monkey 
River, Peck 588 (Gray Herb.). 
Costa Rica: Buenos Aires, Pit- 
tier 10576, Tonduz 3679,4875. 
Los Palm ares, Pittier 10588. 
Helechales del General, Pittier 12064. 
Jamaica: Bunkers Hill Savanna, Harris 11170. 
Trinidad: Pitch Lake, Amer. Gr. Nat. Herb. 75. St. Joseph, Hitchcock 10181. 
Without locality, Crueger. 
77. Fanicum me gist on Schult. 
Panicum megiston Schult. Mant. 2:248.1824; Contr. U. S. Nat. Herb. 15:141.1910. 
DISTRIBUTION. 
Swamps, Mexico and Cuba to Para- 
guay. The type specimen from Brit- 
ish Guiana. 
Tabasco: Ban Juan Bautista, Ro- 
virosa 532. 
Costa Rica: Guanacaste, Jimenez 
727. 
Panama: Empire, Pittier 3726. 
Bohio, Amer. Gr. Nat. Herb. 76. 
Cuba: Santa Cruz de los Pifios, Wright 3872. 
Trinidad: Caroni Savanna, Hart 4197. Without locality, Crueger. 
Venezuela: Santa Catalina, Rushy Squires 355. 
1
 The South American species described as Panicum rotundum Hitchc. & Chase 
(Contr. U. S. Nat. Herb. 15:139.1910) proves to be the same as Panicum campestre 
Nees (Trin. Gram. Pan, 197.1826). 
Pig. 102.—Distribution o[ P. rudgei. 
FiO. MB.—Distribution of P. megiston. 
512 CONTRIBUTIONS PROM THE NATIONAL HERBARIUM. 
Panicum trigonum Retz.,1 an East Indian perennial with low creeping and 
branching culms and spreading pilose blades 2 to 4 cm. long, contracted panicles of 
short-pedicel ed hispidulous spikelets about 1.7 mm. long, the second glume keeled, 
the fruit slightly compressed laterally, has been found "under the shade of a shrub," 
Port of Spain, Trinidad, Broadway 4891. 
Subgenus DICHANTHEUUM Hitchc. & Chase. 
Foliage soft and lax, the flat blades prominently ciliate; 
plants brandling from the base, finally forming ro- 
settes or cushions  See Laxifloua, p. 513. 
Foliage not soft and lax; plants branching from the culm 
nodes. 
First glume nearly as long as the spikelet See Cohijovensia, p. 525. 
First glume much shorter than the spikelet. 
Spikelets glabrous; plants glabrous throughout; 
autumnal form erect, not bushy-branching. See Dichotoma, p. 516. 
Spikelets pubescent. 
Sheaths glabrous or minutely puberulent 
only; ligules minute or obsolete. 
Spikelets attenuate at base, mostly 
prominently pustulose; blades 
narrow, stiff, strongly nerved, ta- 
pering from base to apex  See Angustifolia, p. 514. 
Spikelets not attenuate at base, not pus- 
tulose. 
Spikelets subglobose or pyriform. 
Blades spreading, rarely over 5 
cm. long and 5 mm. 
wide, spikelets pyriform; 
autumnal form bushy- 
branching See Lance a hi a, p. 522. 
Blades erect, 7 to 13 cm. long, 
7 to 14 mm. wide; spike- 
lets spheroid See Spiiaerocaiipa, p. 521. 
Spikelets not subglobose nor pyri- 
form, 
Spikelets 2.8 mm. or moro long. See Commutata, p. 524. 
Spikelets not over 2 mm. long. 
Blades with a cartilaginous 
white margin; spike- 
lets not over 1.5 mm. 
long  101. P. olbomargiTwtum. 
Blades not white-margined; 
spikelets2 mm. long. See Dichotoma, j>. 510. 
Sheaths pubescent. 
Spikelets 4 mm. long, pyriform 104. P. nodutum. 
Spikelets not over 2.6 mm. long. 
Spikelets attenuate at base, mostly 
prominently pustulose; blades 
narrow, stiff, strongly nerved, 
tapering from base to apex; 
autumnal form often bushy- 
branched See Angustifolia, p. 514, 
»Obs. But. 3: 9. 1783. 
t 
HITCHCOCK AND CHASE—TBOPICAL NORTH AMERICAN PANICUM. 5X3 
Spikelets not attenuate at base. 
Culms tall, usually 75 cm. high 
or more; foliage velvety- 
pubescent  See Scop aria, p, 523. 
Culms not over 60 cm. high 
Spikelets not over 1.3 mm. 
long; sheaths and 
culms appressed-pu- 
bescent  See Spreta, p. 518. 
Spikelets 1.7 to 2.4 mm. 
long; plants conspic- 
uously pubescent, at 
least on the sheaths. See Lanuginosa, p. 519. 
LAXIFLORA. 
Spikelets papillose-pilose; sheaths retrorsely pilose 78. P. xalapense. 
Spikelets glabrous; sheaths not retrorsely pilose. 
Blades glabrous on the surface  79. P, polycaulon. 
Blades pilose on the surface  80. P. strigosum. 
78. Panic um xalapense H. B. K. 
Panicum xalapense H. B. K. Nov. Gen. & Sp. 1:103.1816; Contr. U. S. Nat. Herb. 
16: 159.1910. 
DISTRIBUTION. 
Moist banks and rich woods, southeastern United States to Guatemala; also Santo 
Domingo. The type specimen from Jal&pa. 
IIidalgo: Trinidad Iron Works, 
Pringle 13250. 
Puebla: Chinantla, Liebmann 
328. 
Veracruz: Jalapa, Pringle 8083, 
Hitchcock 6687, Amer. Gr. 
Nat. Herb. 84, &mt£A 1752. 
C6rdoba, Bourgeau 2162. 
Boca del Monte, Nelson 201, 
Seler 2160. 
Guatemala : Cobdn, TUrckkeim Fig. 101—Distribution of P. xalapense. 
3831. 
Santo Dominoo: Constanza, Tilrckheim 3059. Rio Yaqui near Jarabacoa, 
Eggera 2129. 
79. Panicum polycaulon Nash. 
Panicum, dichotomum /? glabretcens Griseb. Fl. Brit. W. Ind. 553. 1864. "Pd. 
[Furdie], in mountain-savannahs [Jamaica]." The type specimen is in the Kew 
Herbarium. 
Pameum polycaulon Nash, Bull. Torrey Club 34:200.1897; Contr. XT. S. Nat. Herb. 
16:163. 1910. 
82472°—16 6 
514 CONTRIBUTIONS FROM THE NATIONAL HEKBABIUM. 
DISTRIBUTION. 
Open moist woods and savannas, Florida and the Greater Antilles. The type 
specimen from Florida. 
Cuba: Sierra Nipe near Wood (red, 
Shafer 3083. Herradura, Hitch- 
cock 115. Isle of Pines, Palmer 
& Riley 990. Without locality, 
Wright 3875 in part. 
Jamaica: Hills back of Barican, 
Hitchcock 9561. Bull Head 
Mountain, Hitchcock 9550. 
Porto Rico: Aguada, Sintenti 
5724. Monte Mesa, Chase 6276. 
Trujillo Alto, Chase 6767. Campo Alegre, Chase 6621. 
80. Panicum strigosum Muhl. 
Panicum strigosum Muhl. in Ell. Bot. S. 0. & Ga, 1; 126. 1816; Contr, U. S. Nat. 
Herb. 15: 164.1910. 
distribution . 
Sandy woods and open moist ground, southeastern United States to Colombia and 
in the Greater Antilles. Type specimen from South Carolina or Georgia. 
Veracruz: Minatitl&n, Smith 555. 
Guatemala: Road fromGual&nto 
Cop&n, Pittier 1800. Secan- 
qufm, Pittier 257. 
Costa Rica: Between De Can- 
delaria and San Cristobal, 
Jimenez 459. 
Cuba: El Guama, Palmer & Riley 
213. Without locality, Wright 
3875 in part. 
JAMAICA: Bull Head Mountain, Hitchcock 9551. 
Santo Domingo: Without locality, Eggers 2045 (K. U. Herb.). 
Colombia: Santa Marta, Smith 163. 
ANGCSTIFOLIA. 
Nodes bearded; plants grayish-villous; autumnal blades flat, 
rather soft 82. P. chrysopsidi/olium. 
Nodes not bearded; plants villous only at base, or nearly 
glabrous; autumnal blades involute. 
Spikelets 3 to 3.5 mm. long, pointed 83. P./usi/orme. 
Spikelets less than 3 mm. long, not pointed or obscurely so. 
Plants glabrous or nearly so; autumnal culms erect. 
Spikelets subeecund along the euberect panicle 
branches  86. P. neuranthum. 
Spikelets not subsecund; panicle loose and open. 85. P. ovinum. 
Plants pubescent, at least on the lower half. 
Spikelets about 2.4 mm., rarely only 2.1 mm., 
long; vernal blades 7 to 12 cm. long; 
autumnal blades not falcate 84. P, arenicoloides. 
Spikelets not over 2 mm. long; vernal blades 4 
to 6 cm. long; autumnal blades much 
crowded, falcate 81. P. aciculare. 
Fig. 105.—Distribution of P. polycaulon. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 515 
81. Panicum adculare Deev. 
Fig. 107.—Distribution of P. ocfcufore. 
Panicum adculare Deev.; Poir. in Lam. Encycl, Suppl. 4: 274. 1816; Contr. U. S. 
Nat. Herb. 16: 166,1910. 
In the discussion of the type specimen it is stated that it probably came from the 
southeastern United States. As the species is now known to occur in Porto Rico, it 
becomes probable that this island is the type locality, since Deevaux described many 
species from there. 
DISTRIBUTION. 
Grassy slopes and sandy woods, 
southeastern United States, Cuba, and 
Porto Rico. 
Cuba: Pinar del Rfo, Britton <k 
Gager 7064. Sierra Nipe, 
Shafer 2968. Laguna Jovero, 
Shafer 10733, 10873. With- 
out locality, Wright 3454, 
3461. 
Porto Rico: Monte Mesa, Chase 6273. Campo Alegre, Chase 6624. 
82. Panicum chiysopsidifolium Nash. 
Panicum chrysop&idifoliu m Nash in Small, Fl. Southeast. U. S. 100. 1903; Contr. U. 
S. Nat. Herb. 15: 168.1910. 
DISTRIBUTION. 
Sandy woods and open moist ground, Florida, Louisiana, and the Greater Antilles. 
The type specimen from Florida. 
13 l Cuba: Consolacidn del Bur, Palmer 
& RiUy 481. Isle of Pines, Pal- 
mer & Riley 982. Herradura, 
Hitchcock 116. 
Jamaica: Pedro Morass, Harris 11163. 
Bull Head Mountain, Amer. Gr. 
Nat. Herb. 89. 
Porto Rico: Las Marias, Sintenis 
5985. Trujillo Alto, Chase 6766. 
Fig. 106.—Distribution of P. chrysopsidlfoUum. Santurce, Heller 982. 
Haiti: Without locality, Christ 1744 (K, U, Herb.). 
Santo Dominoo: Without locality, Bertero 818 (K. U. Herb.). 
83. Panicum fusiforme Hitchc. 
Panicum fusiforme Hitchc. Contr. U. S. Nat. Herb. 12: 222. 1909j 15: 172. 1910. 
DISTRIBUTION. ^ '' 1 1  ' f 1 xr ™ 
Sandy pine woods and open moist 
ground, Florida, Cuba, Jamaica, and 
British Honduras. The type speci- 
men from Cuba. 
British Honduras: Manatee La- 
goon, Peck 453a(Gray Herb.). 
Cttba: Herradura, Hitchcock 117, 
Caldwell & Baker 7139, Tracy ri0' l09*-Di3tributioD of p-^0Tme' 
9074. Nueva Gerona, Curtiss 406. Western Cuba, Wright 3453, 3454. 
ajca: Bull Head Mountain. Hitchcock 9552. 
516 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
84. Panicum arenicoloides Ashe. 
Panicum arenicoloides Ashe, Journ. Elisha Mitchell Soc. 16: 89. 1900; Contr. U. S. 
Nat. Herb. 10:173.1910. 
DISTRIBUTION. 
Sandy pine woods, southeastern 
United States; also in Guatemala. 
The type specimen from N orth Caro- 
lina. 
Guatemala: Cuesta de Peinha, 
between Gualin and Cop&n, 
Pittier 1805b. 
Fio. 110.—Distribution of P. arenicoloidts. 
85. Panicum ovinum Scribn. & Smith. 
Panicum ovinum Scribn. & Smith, U. S. 
Dept. Agr. Div. Agrost. Circ. 10: 3.1899, 
Contr. U. S. Nat. Herb. 15:174. 1910. 
DISTRIBUTION. 
Open prairie and meadows, Southern 
States and in Veracruz. The type speci- 
men from Texas. 
Veracruz: Hacienda de la Laguna, 
near Jalapa, Schiede (Berlin Herb.), 
Fia, 111.—Distribution of P. ovinum. 
86. Panicum neuranthum Griseb. 
FlO. 112.—Distribution of P. wuranlhum. 
Panicum neuranthum Griseb. Cat. 
PI. Cub. 232. 1866; Contr. II. S. Nat. 
Herb. 15: 175.1910. 
distribution. 
Moist savannas, Florida and Cuba. 
The type specimen from Cuba. 
Cura: HanfLbana, Wright 3453 
in part. Without locality, 
Wright 3454a. 
DICHOTOMA. 
Nodes, at least the lower, bearded; epikelets pubescent. 
Fruit slightly exposed at maturity; upper sheaths viscid- 
spotted; autumnal form erect or reclining 87. P. nitidum. 
Fruit covered at maturity; sheaths not viscid-epotted; 
autumnal form decumbent, with flabellate-fascicled 
branches 88. P. multirameum. 
Nodes not bearded; epikelets glabrous. 
Spikelets not over 1.6 mm. long; panicles narrow; plants 
glaucous bluish green  91. P. caerulescens. 
HITCHCOCK AND CHASE—TROPICAL NOBTH AMERICAN PAHTCTJM. 517 
Spikelets 2 mm. or more long; panicles open. 
Blades erect, firm; spikelets turgid, strongly nerved; 
plants grayish olive green; spikelets turgid, 
blunt  90, P. Toanohense. 
Blades spreading; spikelets not tuigid, 2.2 mm. or more 
long, pointed; sheaths bearing pale glandular 
spots 89. P. yadkinense. 
87. Panicum nitidum Lam. 
Panicum nitidum Lam. Tabl. Encycl. 1: 172. 1791; Contr. U. S. Nat. Herb. 15: 
183.1910. 
DISTRIBUTION. 
Moist ground and wooded swamps, 
southeastern United States, Bahamas, 
and Cuba. The type specimen from 
[South?] Carolina. 
Bahamas: Great Bahama,Golden 
Grove, Britton & Millspaugh 
2736. 
Cuba: Sierra Nipe, Shafer 2999, 
3001. 
88. Panicum multixameum Scribn. 
Panicum multirameum Scribn. U. S. Dept. Agr. Div. Agrost. Circ. 10: 2. 1900; 
DISTRIBUTION. 
Banks and dry open ground, 
southern Mexico to Guatemala; also 
in Jamaica. The type specimen 
from Jalapa, 
Veracruz: Orizaba, Bourgeon 
2383, Smith 593, Amer. Qr. 
s Nat. Herb. 98. Jalapa, Hitch- 
cock 6610, 6639, Pringle 7882, 
8339a, 9209, 9210. 
JAMAICA: Bull Head Mountain, Hitchcock 9532. 
89. Panicum yadkinextse Asihe. 
Panicum yadhinensc Ashe, Journ. Eliaha 
Mitchell Soc. 16: 85. 1900; Contr. U. S. 
Nat. Herb. 16:195.1910. 
distribution. 
Moist woods and thickets, southeastern 
United States to Mexico. The type speci- 
men from North Carolina. 
Puebla: Without locality, Nicolas in 
1909. Fig. 115.—Distribution of P. yadkinenae. 
Fig. 113.—Distribution of P. nitidum 
Contr. U. S. Nat. Herb. 15:185. 1910. 
Guatemala: CoMn, Turckheim II. 1322. 
518 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
90. Panicum roanokehse Ashe. 
Panicum roanohmse Ashe, Journ. Elisha Mitchell Soc. 15: 44. 1898; Contr. U. S. 
Nat. Herb. 15: 196. 1910. 
DISTRIBUTION. 
Open swampy woods and wet 
meadows, Virginia to Texas; also in 
Jamaica. The type specimen from 
North Carolina. 
Jamaica: Bull Head Mountain, 
Hitchcock 9530. 
Fig. 116.—Distribution of P. roanoUnse. 
01. caerulescena Hack. 
Panicum caerulescena Hack.; Hitchc. Contr. U. S. Nat. Herb. 12: 219. 1909; 15: 
197.1910. 
distribution. 
Marshes and swampy woods, New Jersey to Florida and Mississippi, Bahamas 
Cuba. The type specimen from Florida. 
Bahamas: Great Bahama, Brace 
3524, Britton <fc MUUpaugk 
2506, 2668. Andros, Brace 
7015(allin Field Mus. Herb.). 
New Providence, Britton <t- 
Brace 597, 599, Millspaugh 
2182, Northrup 248, Eggers 
4305 (Hackel Herb.), Eggers 
4312 (K. U. Herb.). 
Cuba: Without locality, Wright 
3463 in part. 
and 
Fig. 117.—Distribution of P. conulHuni. 
SFRETA. 
Spikelets 1.2 to 1.3 mm. long  92. p. leucothrix. 
Spikelets not over 1 mm. long  93. P. wrightianum. 
92. Panicum leucothrix Nash. 
Panicum leucothrix 
16 : 205. 1910. 
Fio. US.—Distribution of P. leucothrix. 
DISTRIBUTION. 
Pine woods and moist open ground, 
New Jersey to Louisiana, Cuba, and 
Porto Rico. The type specimen from 
Florida. 
Cuba: Herradura, Hitchcock 554, 
500. Without locality, Wright 
3460, 3463. 
Porto Rico : Trujillo Alto, Chase 
6770. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 519 
93. Panicum wrightianum Scribn. 
Panicum wrightianum Scribn. U. S. Dept. Agr. Div. Agroet. Bull. 11:44./. 4.1898. 
Contr. U. S. Nat. Herb. 15: 207. 
1910. 
DISTRIBUTION. 
Moist sandy soil, Massachusetts to 
Texas and Cuba. The type specimen 
from Cuba. 
Cuba: Pinar del Rfo, JBritton &■ 
Gager 6952. Without locality, 
Wright 3463 in part. 
LANUGINOSA. 
Spikelets 2.2 mm. or more long. 
Pubescence on culms horizontally spreading; autumnal 
form freely branching  97. P, mllogissimum. 
Pubescence on culms appressed or ascending; autumnal 
form rather sparingly branching  98. P. pseudopubescens, 
Spikelets not over 2 mm. long. 
Plants not velvety; blades usually glabrous on the upper 
surface .  94. P. tennesaeense. 
Plants velvety; blades softly pubescent on both surfaces. 
Plants light or yellow green when dry  95. P. acuminatum. 
Plants dark or olive green when dry 96. P. olivaceum. 
tennesaeense Ashe. 
Panicum tennesseense Ashe, Journ. Elisha 
Mitchell Soc. 15 : 52. 1898; Contr. U. S. 
Nat. Herb. 15: 218. 1910. 
DISTRIBUTION. 
Open ground, Atlantic States to Utah; 
also in southern Mexico, The type speci- 
men from Tennessee. 
Veracruz: Orizaba, Botteri 101. 
Jalapa, Hitchcock 6632. 
acuminatum Swartz. - 
Panicum acuminatum Swartz, Prodr. Veg. Ind. Occ. 23, 1788; Contr. IT. S. Nat* 
Herb. 15: 222. 1910. 
Panicum ornatum Desv.; Hamilt. Prodr. PI. Ind. Occ. 11, 1825. "Herb. Prof. 
Desv. Porto Rico/* 
In the Revision1 this was listed among the doubtful species. The inadequate 
description indicates the common P. acuminatutu. 
1
 Contr. U. 8. Nat. Herb. 16:331.1910. 
Fio. 119.—Distribution of P. WTightianum. 
94. Panicum 
95. Panicum 
520 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, 
DISTRIBUTION. 
Fig. 121.—Distribution of P. acuiainatum. 
mdy pine woods, moist banks and open ground, Greater Antilles; aleo Colombia, 
type specimen from Jamaica. 
Cuba: Isle of Pines, Palmer <fc Riley 989, 1083, Curtiss 307, 328, Taylor 3873. 
Pinar del Rio, Palmer & Riley 447, Herradura, Britton, Earle & Gager 6505, 
Hitchcock 140, Tracy 9078. Rfo Guao, Britton & Cowell 9673. Cuchillas de 
San Sebastian, Le6n & Shafer 13719. Sumidero, Le6n 3472, Shafer 13544. 
Without locality, Wright 3874. 
Jamaica: Clyde River, Harris 11444. Cinchona, Harris 11315, 11371, 11435, 
11494, Hitchcock 9697, 9722, Hills back of Barican. Hitchcock 9564. Bull 
Head Mountain, Hitchcock 9538. 
Gordon Tojyn, Hart 736. Ab- 
bey Green, Amer. Gr. Nat. Herb. 
133, Catherines Peak, Harris 
11554, Hitchcock 9738. Below 
Sir Johns Peak, Harris 11597. 
Upper Clarendon, Harris 11109, 
Haiti: Morne la Sette, Christ 
1850. 
Santo Domingo: Without local- 
ity, Eggers 2021, 2028, 2333 (all 
in K. U. Herb.). 
Poeto Rico: Santnrce, Heller in 1903. Maricao, Sintenis 355. Bayamon, Chase 
6378. Alto de Band era, Chase 6475. Sierra de Luquillo, Chase 6719. Be- 
tween Guainabo and Catano, Chase 6647. Indiera Fria, Chase 6249. Tru- 
jillo Alto, Chase 6765. Jajome Alto, Chase 6752. Lares, Sintenis 5908. 
Fajardo, Sintenis 1224. 
Colombia: La Esmeralda near Jamundi, Pittier 932, 982a. Santa Marta, Smith 
2140. 
06. Paniciun olivaceum Hitchc. & Chase. 
Panieum olivaceum Hitchc. & Chase, Contr. U, S. Nat. Herb. 15: 225. 1910. 
distribution . 
Clay slopes and open ground, Mexico to Venezuela. The type specimen from 
Guatemala. 
Vebaceuz : Orizaba, Bourgeau 
2383 in part, Botteri 99, 100, 
101. Minatitl&n, Smith 571. 
Jalapa, Hitchcock 6631, 6646, 
6651, 6667, Amer. Gr. Nat. 
Herb. 134, Pringle 8339. 
Guatemala: Cob An, Tiirckheim 
428. Without locality, Seler 
3235. 
Costa Rica : San Pedro de la Ca- 
labaza, Tonduz 10745. 
Fig, 122.—Distribution gf P, oHvaceuvL 
Panama: El Boquete, Maxon 5136, Hitchcock 8189, 8252, 8280. 
Venezuela: Tovar, Fendler 1638b. Pico de NaiguatA, Pittier 6252. 
Colombia: La Esmeralda, near Jamundi, Pittier 946. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 521 
97. Panicum viUoaissimum Nash. 
Flo. 123.—Distribution of P. vilkaiisimum. 
98. Panicum pseudopubescen# Nash. 
Panicum pseudopubcsccns Nash, Bull. 
Torrey Club 26: 577. 1899; Contr. U. 8. 
Nat. Herb. 15: 235.1910. 
DISTRIBUTION. 
Sandy open woods eastern United States; 
also in San Luis Potosf. The type speci- 
men from Alabama. 
Panicum viUonstvmum Nash, Bull. 
Torrey Club 33: 149. 1896; Contr. 
U. S. Nat. Herb. 10:233.1910. 
DISTRIBUTION. 
Dry open woods, eastern United 
States; also in Guatemala. The type 
specimen from Georgia. 
Guatemala: Guatemala City, 
Hitchcock 9106. 
San Luis PoTosf: Without locality, 
Schaffner 146. Fig. 124.—Distribution of P. psevdopvbtKtru. 
SPHAEBOCARPA. 
Culms spreading; panicle nearly as broad as long; spikelets 1.6 
to 1.8 mm. long  99. P. sphaerocarpon. 
Culms erect or ascending; panicle never more than two-thirds 
as broad as long; spikelets 1 to 1.2 mm. long 100. P. erectifolium. 
99. Panicum sphaerocarpon Ell. 
Panicum sphaerocarpon Ell. Bot. S. C. <fc Ga. 1: 125. 1816; Contr. U. S. Nat. Herb. 
16: 251. 1910. 
DISTRIBUTION. 
Sandy soil and gravelly banks, eastern United States through Mexico to northern 
South America. The type specimen from Georgia. 
San Luis PoTOsf: Without locality, Schaffner 147. 
MichoacAn : Sierra de San Andres, Ross 381. 
Puebla; Chinantla, Liebmann 327. TeziutlAn, Orrntt 3952. 
Veracruz: Orizaba, Amer. Gr. 
Nat. Herb. 155, Botteri. Ja- 
lapa, Pringle7883,8344, Hitch- 
cock 6607, 6633. 
British Honduras: Manatee La- 
goon, Peck 61 (Gray Herb.). 
Guatemala : Guatemala City, 
Hitchcock 9105. Cob&n, Tiirch- 
heirn 56, 3829. Volcano Pa- 
cay a, Kellerman 6236. 
CostaRica: CoDev. TonAw11866. 
Fig. 135.—Distribution of P. sphaerocarpon. 
San Pedro de la Calabaza, Tonduz 10745. P&ramos del Abejonal, 7Ymduz7878, 
Panama : El Boquete, Hitchcock 8172. Dolega, Hitchcock 8343. 
Venezuela : Without locality, Fendltr 1638. 
♦ 
522 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
100. Panicum ©rectifolium Nash. 
Panicum ereetifolium Nash, Bull. 
Torrey Club 33: 148. 1896; Contr. 
U. S. Nat. Herb. 15: 256.1910. 
DISTRIBUTION. 
Moist pine woods, southeastern 
United States and Cuba. Type speci- 
men from Florida. 
Cuba: Vuelta Abaja, Wright 3462. Laguoa Santa Maria, Britton & Gager 7170. 
EN8IFOLIA. 
101. Panicum albomarg'in.&tum Nash. 
Panicum albomarginatum Nash, Bull. Torrey Club 24: 40. 1897; Contr. U. S. Nat. 
Herb. 16: 260. 1910. 
DISTRIBUTION. 
Moist sandy woods, southeastern 
United States, Cuba and Guatemala, 
Type specimen from Florida. 
Guatemala : Gual&n to CoptLn, 
Pittier 1805a. 
Cuba: Between Sumidero and 
Pinar del "Rio, Le6n 346C. 
Herradura, Hitchcock 555. 
Isle of Pines, Taylor 32. 
LANCE ARIA. 
Spikelets 1.5 to 1.6 mm. long  102. P. portoricen&e. 
Spikelets 2 mm. long—  103, P, lancearium. 
102. Panicum portoricense Dee v. 
Panicum portoricense Desv.; Hamilt. Prodr. PI. Ind. Occ. 11. 1825. "Herb. Prof, 
DeBv. Porto Rico." We have been unable to find the type of this and in the Revision 1 
placed it among doubtful species. Recent study of the grasses of Porto Pico, however, 
leaves no doubt that Desvaux's description was drawn from a specimen in the autumnal 
phase of P. paucicilmtum A she, which is found to be common in the vicinity of ,Saii 
Juan Bay, the region most likely to have been visited by early botanists. 
Panicum pauriciliatum Ashe, Journ. Eliaha Mitchell Soc. 16: 87. 1900; Contr. 
U. S. Nat. Herb. 15: 272. 1910. 
1
 Contr. U. S. Nat. Herb. 15: 331. 1910. 
Fig. 126.—Distribution of P. ereetifolium. 
Fig. 127.—Distribution of P. albomarginatum. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 523 
DISTRIBUTION. 
Moist sandy woods, southeastern United States, Cuba and Porto Rico. 
Cuba: Without locality, Wright 
3876. 
Porto Rico : Martin Pefia, Chase 
6357. Between Catafio and 
Guainabo, Chase 6631. Lake 
Loisa, Chase 6783. Santuree, 
Heller 982b, 6442. VegaBaja, 
Heller 639, Underwood & 
Gfriggs 955. Campo Alegre, 
Chase 6434. Fio. 128.—Distribution of P. portoriceme. 
103. Panicum lancearium Trin. 
Panicum lancearium Trin. Gram. Pan. 223. 1826; Contr. U. S. Nat. Herb. 16: 273. 
1910, 
DISTRIBUTION. 
Sandy pine woods, southeastern 
United States, British Honduras, 
Cuba, and S&nto Domingo. The type 
specimen from North America, the 
exact locality not known. 
British Honduras: Sibune 
River, Peck 425 (Gray Herb.). 
Cuba: San Juan de Buenavista, 
Flo. ^.-Distribution of P. lancearium. 3460 (Gray Herb.). 
Santo Domingo: Constanza, Tiirckheim 3321. 
PEDICELLATA. 
104. Panicum nodatum Hitchc. & Chase. 
Panicum nodatum Hitchc. & Chase, 
Contr. U. S. Nat. Herb. 15: 293. 1910. 
DISTRIBUTION. 
Oak woods and wooded sand dunes, along 
the coast of Texas and adjoining Mexico. 
The type specimen from Texas. 
Mexico : Matamoroe, Berlandier988,2418 
(Gray Herb.). 
Fio. ISO.—Distribution of f, nodatum. 
SCOPARIA. 
Vernal culms erect or ascending; plants velvety throughout; 
spikelets about 2.5 mm. long 105. P. scoparium. 
Vernal culms decumbent at base; upper sheaths mom or less 
glabrate; spikeleta less than 2 mm. long  106. P. viscidellum. 
524 CONTRIBUTIONS FtfOM THE NATIONAL HERBARIUM 
100. Panic urn scoparium Lam. 
Fig. 131.—Distribution of P. scoparium. 
Panicum scoparium Lam. Encycl. 4: 
744.1798; Contr. U. S. Nat. Herb. 15: 
294.1910. 
DISTRIBUTION. 
Wet places, southeastern United 
States and Cuba. The type specimen 
from South Carolina. 
Cuba: Road to Final Mayarl, 
Wright 3467. Sierra Nipe, 
Shafer 2997, 3018. 
Fig. 132.—Distribution oi P. tskcidellum. 
106. Panicum viscidellum Scribii. 
Panicum viscidellum Scribn. U. S. Dept. Agr. Div. Agrost. Circ. Id: 2. 1900; Contr. 
U, S. Nat. Herb. 15: 296.1910. 
DISTRIBUTION. 
Open woods and slopes, Mexico to Colombia; also in Cuba. The type specimen 
from Jalapa. 
Veracruz: Ctfrdoba, Hitchcock 6425. Orizaba, Arner.Gr. Nat. Herb. 179. Jalapa, 
Hitchcock 6606, 6673, Smith 
1617, Pringle 8089. Mirador, 
Liebmann 323. Zacuap&n, 
Purpus 2160, 
Guatemala: Cob&n, Turchheim 
3836,3829 in part. Sierra del 
Mico, Kellerman 6231,6240. 
Nicaragua: TJ. S. Pacific Expl. 
Exped. Wright. 
Costa Rica: Assert, Tonduz 1244. 
El General, Pittier 3358. 
Panama: El Boquete, Hitchcock 8178, Pittier 3043. 
Cuba: Managua, Palmer <£■ Hiley 1065. 
Colombia: La Esmeralda, near JamuncM, Pittier 940. 
COMMUTATA. 
Blades unsymmetrical and falcate; culms decumbent; first 
glume about one-third as long as tlie spikelet  107. P. jodrii. 
Blades symmetrical, nearly linear; culms erect; first glume 
about half aa long as the spikelet 108. P. albomaailatum. 
107. Panicum joorii Vasey. 
Panicum joorii Vasey, TJ. S. Dept. Agr. 
Div. Bot. Bull. 8: 31.1889; Contr. U. S. Nat. 
Herb. 15: 308.1910. 
distribution. 
Damp woods, southeastern United States; 
also in southern Mexico. The type speci- 
men from Louisiana. 
Veracruz: Jalapa, Hitchcock 6650. 
Republic of Mexico : Without locality, 
no. 133.—Distribution of joorii. Pringle 13290. 
* 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 525 
108. Panicum aibomaculatum Scribn. 
Panicum albomaeulatum Scribn. U. S. Dept. Agr. Div. Agrost. Circ. 19: 2. 1900; 
Contr. U. S. Nat. Herb. 15: 311. 1910. 
Autumnal form leaning or ascending, with geniculate nodes, branching from the 
lower and middle nodes, the branches nearly as long as the primary culm, erect, 
simple or sparingly branching, the sheaths overlapping, the blades scarcely reduced, 
the panicles short-exserted; apparently no winter rosette developed. 
One specimen, Hitchcock 6979, has harshly pilose foliage, but in all other respects its 
characters are those of P. aibomaculatum. 
DISTRIBUTION. 
Shady banks and wooded slopes, 
Mexico and Guatemala. The type 
specimen from P&tzcuaro. 
Jalisco : Zapotl&n, Hitchcock 7170, 
Airier. Gr. Nat. Herb. 193. 
Michoac^n : Pitzcuaro, Pringle 
5203, Uru&pan, Hitchcock 
6979. Jorullo, Humboldt 
(Willd. Herb.). 
Guatemala : Antigua, Hitchcock 9139 
Fig. 134.—Distribution of P. albomocviatam. 
CORDOVENSIA. 
Plants perennial, straggling, the culms decumbent at base, rooting at the nodes; 
ligulcs membranaceous, minute; spikelets elliptical, the first glume two-thirds the 
length of the spikelet or more; fruit smooth and shining, apiculate. 
This group, to which are apparently allied the South American Panicum phragmUes 
Nees (P. discolor Trin., not Spreng.) and P. ovulifcrum Trin., appears to be inter- 
mediate between true Panicum and the subgenus Dichanthelium, having the sterile 
primary panicles and secondary panicles with fruitful, probably cleistogamous, 
spikelets, but not forming a winter rosette. 
Plants glabrous or the leaves sparsely pilose  109. P. cordovense. 
Plants velvety pubescent 110. P. chiriquiense. 
109. Panicum cordovenae Fourn. 
Panicum cordovense Fourn. Mex. PI. 2: 26. 1886. "Cordova (Schaffn. n. 293 in 
herb. Franq.)." The type specimen in the herbarium of Drake de Castillo, now a 
part of the herbarium of the Jardin dee Plantee, Paris, has glabrous spikelets (like 
that shown in Fig. 135). This in the Revision was listed among the doubtful species.1 
Panicum expansum Fourn. Mex. PL 2: 26.1886 (following P. cordovense). "Huitar 
malco (Liebm. n. 426)." The type specimen, in the Copenhagen Herbarium, has a 
large immature primary panicle, the spikelets glabrous. In the Revision 3 it was 
suggested that this unknown species was related to P. costaricense. 
1cknanthw apimlatus Scribn. TT. S. Dept. Agr. Div. Agrost. Circ. 30: 1. 1901. 
"Type specimen from near Jalapa, State of Vera Cruz. C. G. Pringle, no. 9208." In 
the type specimen, in the United States National Herbarium, the spikelets are 
glabrous. 
Panicum missionum Ekman, Ark. ffir Bot. II4: 19. pi. S. /. i. 1912. " Bonpland, 
ad rivulum prope praedium 'Almacdn finlandesa,' 26.12. 07. N. 632.", collected by 
E. L. Ekman, in the State of Misiones, Argentina. An immature specimen of the 
type collection is in the U. S. National Herbarium, received from Dr. Ekman. The 
1
 Contr. U. S. Nat. Herb. 15: 329. 1910. 
3
 Contr. V. S. Nat. Herb. 16: 134.1910. 
526 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
Fig. 135.—P. cordovexue. From 
Hitchcock 8328. 
photograph of the type specimen (plate 3, figure I) shows the large open primary 
panicle, lite that of the type of P. expansum, which is characteristic of the early 
phase of the species. 
description . 
Plants perennial in tangled masses; culms slender, subcompressed, a line of pubes- 
cence down one side or glabrous, rarely sparsely pilose, straggling, widely creeping, 
rooting at the lower nodes, as much as 2 meters long, freely branching, the branches 
usually divergent, becoming nearly as long as the 
primary culm; nodes puberulent or glabrate; sheaths 
much shorter than the usually elongate internodes, 
sparsely papillose-pilose or papillose only, densely 
ciliate on the margin, puberulent at the junction with 
the blade; ligule about 0.3 mm. long; blades thin, 
flat, spreading, 5 to 10 cm. (rarely as much as 15 
cm.) long, 5 to 10 mm. wide, narrowly lanceolate, 
acuminate, rounded and usually ciliate at base, ncab- 
ituIous on both surfaces at least on the midnerve, 
usually papillose-hispid above the ligule, sometimes 
sparsely so throughout; panicles of two kinds, the 
primary short-exserted or included at base, loose and 
open, 10 to 15 cm. long, half to two-thirds as wide, the axis and branches slender, sub- 
flexuous, scaberulous, pubescent in the axils, the few branches solitary or in pairs, 
remote, simple or nearly so, finally spreading, few-flowered, the branchlets and rhort- 
pediceled, apparently unfruitful spiltelets more or less appresaed to the rachis, the 
secondary panicles terminal on the branches, 
reduced, narrow, few-flowered, partially 
inclosed in the sheath, the crowded appar- 
ently cleistogamous spikelets fruitful; spike- 
lets 3 to 3.5 mm. long, 1.2 to 1.4 mm. wide, 
those of the primary panicles usually glab- 
rous, those of the secondary panicles usually 
pustulose-villous, turgid; first glume about 
two-thirds as long as the spikelet, obtuse, 
3 to 5-nerved, the second glume and sterile 
lemma equal, covering the fruit, 7-nerved, 
the middls internerves of the sterile lemma 
in the pubescent opikelct* usually glab- 
rous, the midnerve sometimes indistinct; fruit 2.8 to 2.!) mm. long, 1.1 to 1.2 mm. 
wide, apiculate. 
The spikelets shown in figures 135 and 136 are from the terminal and lateral panicles 
respectively of the same individual. 
DISTRIBUTION. 
Shady banks and in deep woods, southern Mexico to Argentina. 
T3  
Veracruz; C6rdwba, Schaffner 293. 
Jalapa, Pringle 9208, Hitchcock 
6611. Huitmalco, IAebmann 
426. 
Costa Rica: Fiedra del Convento, 
Pittier 3655. 
Panama: Chiriquf Volcano, Hitch- 
cock 8196. El Boquete, Sitch- 
cocfc 8271> 8275» 8303, 8328. FiG. 137.—Distribution of P. coriovense. 
Fig. 136.—P. cordovmse. From Hitchcock 8328. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM, 527 
110. Panicum chiriquiense «p. nov. 
DESCRIPTION. 
Plants perennial, olivaceous; culms straggling, creeping and rooting at the lower 
nodes, softly papillose-pilose, freely branching, the leafy fertile branches ascending, 
20 to 30 cm. high; nodes pilose; sheaths nearly as long as the internodes or the upper 
overlapping, softly papillose-pilose; ligiile about 0.5 mm. long; blades flat, somewhat 
spreading, 4 to 7 cm. long, 7 to 10 mm. wide, nar- 
rowly lanceolate, unsymmetrical at base and often 
somewhat falcate, acuminate, softly papillose- 
villous beneath, rather sparsely pilose on the 
upper surface; panicles shart-exsertod or included 
at base, 2.5 to 3.5 cm. long, half to two-thirds as 
wide, the few branches ascending, the axis and 
few nearly simple branches slender, viilous; spike- 
lets short-pediceled, 2.6 to 2.8 mm. long, about 
1.1 mm. wide, elliptical; first glume about three- 
fourths as long as the spikelet, acute, 3-nerved, ^ m _p cWri?t(lejM<. 
viilous; second glume and sterile lemma equal, type spocinifiii. 
covering the fruit, the glume viilous, minutely 
apiculate, the lemma usually subinduratc, smooth and shining in the two middle 
internerves, the midnerve suppressed or evident at the summit only, the lateral 
mternerves viilous; fruit 2.1 mm. long, 1 mm. wide, minutely apiculate. 
Type in the U. S. National Herbarium, no. 725186, collected "in shade along 
trail, hillside jungle, foothills, vicinity of El Boquete, province of Chiriquf, Panama, 
altitude 1,000 to 1,300 meters, October 4, 
1911," by A. S. Hitchcock (no. 8313). 
Known only from the type collec- 
tion. This speciea differs from P. cor- 
doiense in the viilous foliage with 
shorter, unsymmetrical blades, the 
less elongate culms, and the smaller 
spikelets. In the specimens collected 
all the panicles are terminal on the 
branches. Whether or not the plants 
at an earlier season bear large panicles 
on a primary culm is not known. The character of the suppressed midnerve and 
smooth middle internerves, rare in P. eordovense, is usual and emphasised in P. chiri- 
quiense. Only an occasional spikelet in panicles with the usual form has undifFer- 
entiated middle internerves. 
MISCEIiLAltEOTrS SPECIES. 
111. Panicum obtuaum H. B. K. 
Panicum obtusuvn H. B. K. Nov. Gen. & Sp. 1; 98, 1816; Contr. U. S. Nat. Herb. 
15: 321.1910. 
DISTRIBUTION. 
Moist sandy or gravelly soil, southwestern United States to central Mexico. The 
type specimen from Guanajuato. 
Sonora: Nogales to Cocoepora Ranch, Griffiths 6800. 
Chihuahua: Between Casaa Grand ea and Sabinal, Nelson 6352. Chihuahua, 
Pringle 476. Mifiaca, Hitchcock 7734. 
Fia. 139.—Distribution of P. ckiriquieiwe. 
528 CONTRIBUTIONS FBOM THE NATIONAL HERBARIUM 
Durango: Durango, Palmer 175 in 1896, Hitchcock 7618. Torrefin, Hitchcock 7563. 
Coahuila: Saltillo, Palmer 394 and 504 in 1898, Hitchcock 5582. La Ventura, 
Nelson 3908. Jaral, Schumann 1714. 
Nuevo Le6n: Monterrey, Hitchcock 5533. 
Zacatecas: Concepcifin del Oro, Palmer 
266 in 1904. Zacatecas, Hitchcock 7525. 
Aquascalientes: Aguascalientea, Rose <t 
Hay 6231, Hitchcock 7456. 
San Luis PoTosf: San LuisPotosf, Palmer 
590 in 1898, Parry & Palmer 960 in 
1878, Schaffner 148, Hitchcock 5657. 
Guanajuato; Obreg6n, Hitchcock 5801. 
Irapuato, Hitchcock 7407. 
Quer&taro: San Juan del Rfo, Rose, 
Painter <fc Rose 9552,9594. Querdtaro, 
Hitchcock 5813, 5863. 
Hidaloo: Tula, Rose, Painter & Rose 8356. Ixmiquilpan, Rose, Painter & Rose 
9057. 
Puebla: Tehuacin, Hitchcock 6060. 
Republic op Mexico: Without locality, Hartweg 244. 
112. Panicum Btagnatile ep. nov. 
DESCRIPTION. 
Plants perennial, gregarious; culma erect from an ascending or decumbent base 
rooting at the nodes, somewhat lush, 1 to 2 meters high, about 5 mm. thick, glabrous, 
simple or occasionally with sterile branches; nodes glabrous or the lower retroraely 
appresaed-pubescent; sheaths usually about as long as the internodes or the upper 
overlapping, ciliate on the margin, otherwise glabrous or pubescent on the sides at 
the junction with the blade; ligule membranaceous, less than 1 mm. long; blades 
flat, 20 to 35 cm. long, 1.5 to 3 cm. wide, slightly narrowed to the rounded base, taper- 
ing from below the middle to an acuminate apex, sparingly pilose above the ligule, 
otherwise glabrous, the margins scabrous; panicle 20 to 40 cm. long, about half as 
wide, fusiform in outline, the main axis and rachisea strongly 
angled, scabrous, usually sparsely pilose in the lower axils, the 
numerous slender, compound branches rather stiffly ascending 
or spreading, solitary or irregularly fascicled, bearing from near 
the base and usually along the lower side rather stiffly spreading 
slender secondary branchleta, the lower 1 to 2 cm. long; spike- 
lets loosely clustered, short-pediceled, glabrous, 1.8 mm. long, 
about 0.6 mm. wide, acute; first glume about one-third as long 
as the spikelet, acute, scabrous on the keeled midnerve, the *1°' 
,'i , , From type specimen* 
second glume two-thirds as long as the spikelet, somewhat boat- 
shaped, scabrous on the keel, the sterile lemma slightly exceeding the fruit, boat- 
shaped and scabrous on the midnerve at the apex; fruit 1.6 mm, long, 0.4 mm. wide, 
lanceolate, scabrous at the acute apex, the lemma and palea subindurate, the margins 
of the lemma inrolled only at the base. 
Type in the U. 8. National Herbarium, no. 693328, collected in water of swamp, 
Frijoles, Canal Zone, Panama, October 12,1911, by A. S. Hitchcock (no. 8388). 
This species is allied to P. rivulare Trin. but differs in the broader blades, less densely 
flowered panicle branches, and smaller spikelets. The numerous small spikelets 
slender branchleta give the panicle a lacelike appearance. The type locality is now 
covered by the water of Gatun Lake. 
HITCHCOCK ANP CHASE—TROPICAL NORTH AMERICAN PAN1CUM. 529 
DISTRIBUTION. 
Swampe, growing in. the water, south- 
ern Mexico to Panama. 
Tabasco: Between San Juan Ban- 
tista and San Sebastian, 
Rovvrooa 625. 
Guatemala: Puerto Barrios, Hitch- 
cock 9153. 
Panama: Frijoles, Hitchcock 8388. 
Fig. 142.—Distribution of P. staffnatil*. 
113. Panicum gr&nde Bp. nov. 
Plants perennial, gregarious, producing extensively creeping or floating leafy 
stolons about 5 mm. thick; culms 1.5 to 2 meters or more high, erect from a long decum- 
bent base with papery sheaths and tuits of fibrous roots, 1 to 2 cm. thick, simple or 
sparingly branching,, succulent, the nodes densely appressed-hirsute; sheaths over- 
lapping except toward the summit, glabrous, the junction, with the blade in drying 
presenting a darkened triangle on each side; ligule membranaceous, about 2 mm. long; 
blades flat, as much as 1 meter long and 6 cm. wide (the upper and lower smaller), at 
base narrower than the sheath, gradually widening to about the middle, narrowing 
rather abruptly to the acuminate apex, glabrous, striate, somewhat plicate, the 
margins strongly serrulate; panicle as much as 60 cm. long and 40 cm. wide, the axis 
and branches strongly several-angled, scaberulous, the promi- 
nent pulvilli minutely pubescent, the branches stiffly 
spreading, naked at base, the lower in whorls, the short 
ultimate branchlets and the pedicels appressed along the 
rather loose secondary branchlets, the pedicels mostly 1 to 
2 mm. long; spikelets 2.5 mm. long, 0.9 mm. wide, nearly 
terete, pointed, glabrous; first glume slightly over half the 
length of the spikelet, the second glume and sterile lemma 
equal, exceeding the fruit, somewhat beaked beyond it; 
fruit 1.8 mm. long, 0.7 mm. wide, narrowly obovate, smooth 
and shining, the lemma and palea indurate but the lemma 
margins flat. 
Type in U. S. National Herbarium, nos. 693329, 693330, 693331 (parts of the same 
individual), collected in the water of a swamp along the margin of Gatun Lake, Canal 
Zone, Panama, December 15,1911, by A. S. Hitchcock (no. 9178). 
This species grows in large masses in swamps, flowering in December. The broad 
blades give a lily-like aspect to the plants earlier in the season. In Gatun Lake, 
Panama, plants were found growing in 10 feet of water. 
Doell1 describes this species under the name Panicum multifiorum Poir. ,2 but the 
latter name applies to a different species.3 Panicum grande is allied to the South 
American P. grumosum Nees, P. rivulare Trin., and P. prioniiis Nees, from all of which 
it differs in its aquatic habit and open panicle and in the more indurate fertile lemma 
and palea; from P. rivulare and P. prumitis in the equal second glume and fertile 
lemma. 
Fig. 143.—P. grande. From 
type specimen. 
1
 Mart. Fl. Bras. 8a: 215. 1877. 
= Encyci. 4: 282,1816. 
1
 See Contr. U. S. Nat. Herb. 15: 48. 1910. 
82472°—15 « 
530 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM 
Fig. 144—Distribution of P. grande. 
Venezuela: Bobures, John 353. 
DISTRIBUTION. 
Lakes, pond3, and swamps, growing 
in the water, Panama to Pard, 
PANAMA: Gatun, Hitchcock 9178. 
Along the Sambii River, Pit- 
tier 5546. 
Trinidad: Mazanilla, Hitchcock 
10376. Cedros, Hitchcock 
10148. St. Joseph, Hitchcock 
10022. 
114. Panicum tuerckheimii Hack. 
Panicum tuerckheimii Hack. Oesterr. Bot. Zeitschr. 12: CO. 1906. Guatemala, 
Cubilgtiitz [Cubilquitz] im Hochwald, 350 msm. leg H. v.Turckheim(1903 sub nr. II. 
820)." 
DESCRIPTION. 
Plants perennial from a firm knotty base with strong roots; culms erect or ascend- 
ing, simple, 30 to 50 cm. high, compressed, glabrous, the nodes appressed-hirsute, few 
to several long erect leaves borne at the base; sheaths compressed-keel ed, more or less 
hispidulous at the junction with the blade, otherwise glabrous, those of the basal leaves 
loose, overlapping, those of the culm about as long as the in- 
ternodes; ligule membranaceous-ciliate, scarcely 0.5 mm. 
long; blades thin, flat, 10 to 25 cm, long, 2 to 2.5 cm. wide, 
gradually tapering from about the middle to an acuminate 
apex, the uppermost rather abruptly narrowed to a some- 
what rounded ciliate base, the lower and especially the 
basal blades gradually tapering into a narrow, more or less 
folded, petiole-like base, ciliate on the margins, puberu 
lent on the upper surface at the very base; panicle short 
exserted, 15 to 20 cm. long, less than one-fourth as wide, 
the slender angled axis and branches glabrous, the branches 
remote, rather stiffly ascending, with short ascending re- 
mote branchlets, these spikelet-bearing from the base, 
more or less pilose in the axils, the lower branches about 10 cm. long; spikelets 
rather short-pediceled, russet brown, 2.5 to 3 mm. long, about 0.9 mm. wide, acum- 
inate, very sparsely appreased-pilose; first glume minute, hyaline; second glume and 
sterile lemma rather faintly nerved, subequal, or the acuminate tip of the glume 
exceeding the lemma; the sterile palea wanting; fruit 2.4 mm. long, 0.8 mm. wide, 
the lemma and palea not greatly in- 
durated, the lemma minutely pubes- 
cent at the pointed apex, the palea at 
the very tip not inclosed. 
In the original description Professor 
Hackel states that the first glume is 
altogether wanting. This appeared to 
be the case and, failing to note the 
obscure organ, we excluded this species 
from the genus Panicum in the Re- 
vision.1 The first glume is present, 
but is so thin in texture and bo closely appressed that the nerves and the base of 
the sterile lemma are obscurely visible through it. 
This species, known only from the type collection, has no close allies. 
Fig. 145.—P. tuerckheimii. 
From type specimen. 
Pig. 146.—Distribution of P. tuerckheimii. 
1Contr. U. S. Nat. Herb. 15:16. 1910. 
1 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 531 
115, Panicum zizanioides H. B. E. 
Panicum zuanioides H. B. K. Nov. Gen. & Sp. 1:100.1816; Gontr. U. S. Nat. Herb. 
16: 325. 1910. 
DISTRIBUTION. 
Moist usually shaded places, Mexico and the West Indies to Paraguay. The type 
locality, Colombia. 
Veracruz: C6rdoba, Hitchcock 6436. 
Oaxaca: Trapeche de la Concepcitfn, Liebmann 394. 
Tabasco : San Juan Bautista, Rovirosa 624. 
Chiapas: Ocuilapa, Nelson 3023. 
Guatemala: Cubilquitz, TUrckheim 7699, 7700, 8785, 8796. Cob&n, Turckheim 
3828. 
Nicaragua: Jinotepe, Hitchcock 8680. 
Costa Rica: Hacienda de Zent, Tonduz 302. La Florida, Pittier 11276. Matina, 
Pittier 10307, Talamanca, Tonduz 8566. San Rafael, Pittier 2598. 
Tabemilla, Hitchcock 8112, 8385. Gatun, 
j j | I *1 ~~~* 
l^TL ' ' / 
^j f * 
Fiq. 147.—Distribution of P. zizanioides. 
Port Antonio, Mown 2109. Ramble, Hitch- 
cock 9513. Castle ton Gardens, Hitchcock 9398. Above Constant Spring, 
Hitchcock 9278. Buff Bay, Hitchcock 9.779. Ipswich, Hitchcock 9609, 
Ewarton to Linatead, Hitchcock 9419. St. Georges, Portland, Harris 11420. 
Temple Hall, Harris 11385. Troy, Hitchcock 9781. Montego Bay, Hitchcock 
9684. "Without locality, Alexander Prior in 1850, Wullschlaegel 1108. 
Trinidad : Port of Spain, Hitchcock 9958. Mount Pleasant Estate, Bot. Gard, Herb. 
2286. Toco, Broadway 2563. Without locality, Bot. Gard. Herb. 3188. 
Tobago: Great Dog River, Eggers 5810. Roxburgh, Hitchcock 10258. Easter- 
field, Broadway 4556. 
116. Panicum hirtum Lam. 
Panicum hirtum Lain. Encycl. 4: 741. 1798. Lamarck states that "Cette plante 
croit & Cayenne, d'oii elle a £t6 envoyde par Jos. Martin." The type specimen in the 
Lamarck Herbarium is marked "Panicum hirtum Lain. diet. Cayenne. Martin." 
DESCRIPTION. 
Plant# annual, at first simple and erect, later branching and decumbent, rooting 
at the lower nodes; culms slender, puberulent, the fertile branches 20 to 40 cm. high; 
sheaths shorter than the intercedes, loose, short-viUous, densely ciliate on the margin; 
ligule a delicate toothed membrane less than 0.5 mm. long; blades flat, thin, 2 to 6 cm. 
long, 0.8 to 2 cm. wide, ovate to lanceolate, cordate-clasping at base, rather abruptly 
acuminate, scabrous on the upper surface, sparsely hispid on both surfaces, ciliate at 
the base; panicles exserted or included at base, 3 to 7 cm. long, half to two-thirds as 
Panama: David, Hitchcock 8351. 
Hitchcock 7974. Gamboa, Pit- 
tier 4790. Between Gamboa 
and Cruces, Pittier 3781. 
Along the Rfo Sirrf, Pittier 
4026. 
Cuba: Sancti Spiritus, Le6n 903. 
Laguna Caste llano, Baker 
4334, Wilson 9567. Puentes 
Grande?, Le6n 2743. Sumi- 
dero, Le6n 13635. Without 
locality, Wright 3466. 
Jamaica : Gordon Town, Hart 726. 
532 CONTHIBUTIONS FBOM THE NATIONAL HERBARIUM. 
wide, ovoid in outline, the main axis pilose, the numerous delicate but rather stiff 
branches ascending at a uniform angle (in herbarium specimens the branches con- 
spicuously parallel), the numerous but not crowded 
spikelets recurved at right angles on capillary pedi- 
cels; spikeleta 1.2 mm. long, about 0.7 mm. wide, 
strongly plano-convex, turgid; first glume nearly as 
long aa the spikelet, 3-nerved, narrow, covering the 
middle internervea of the sterile lemma, thin in tex- 
ture and bo closely appressed as to be usually invisi- 
ble, sparsely hispid; second glume inflated, gibbous, 
pointed, 5-nerved, papillose-hispid, at maturity sub- 
indurate, the hairs stiffening and as much as 1 mm. 
long; sterile lemma equaling the second glume and 
inclosing a 2-keeled palea, 5-nerved, glabrous, the two 
middle internerves thin; fruit 1 mm. long, 0.6 mm. 
wide, very turgidly plano-convex, at first white, at 
maturity brown, sparsely sprinkled with minute glob- 
ular hairs. 
The spikelets of this unique species at maturity look like tiny burs or, aa Lamarck 
says, like the'' seeds of Daucus." The bristly second glume sometimes falls, leaving 
the turgid fruit, together with the first glume and sterile lemma, attached to the 
pedicel. 
DISTRIBUTION. 
Damp shady places, Trinidad to 
Brazil. 
Trinidad: Arima, Hitchcock. 10310. 
P i a r c o Savanna, Hitchcock 
10363. St. Joseph, Hitchcock 
10177. Port of Spain, Hitch- 
cock 10320. Without locality, 
Bot. Gard. Herb. 3194. Fig. 149.—Distribution ol jP. fcirtutn. 
Fig. 148 — P, hirtum. From type 
specimen. 
EXCLUDED SPECIES. 
The study of genera allied to Panicum and the examination of a 
large number of type specimens has resulted in the identification of 
most of the species of North America included now or in the past by 
some authors in Panicum. Besides the valid species and the names 
accounted for in synonymy within the genus Panicum there are a 
great many names that, according to our present conception of the 
genera of Paniceae, are referable to other genera. While the list is 
not complete it is so nearly complete as to enable us to account for 
almost all the names of North American species that have been re- 
ferred to Panicum. The second name in the column merely indicates 
the genus to which the species belongs. In no case is any name in 
the following list a transfer of a species or a new combination. In 
many cases the name, if transferred to the genus indicated, would 
be untenable. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANIC UM. 
Panicum acutum Raap. Reimarochloa. 
adscendens H. B. E. Synth enema, 
adustum Neee. Syntherisma. 
africanum Trm. Oplismenus. 
agglutinans Kunth. Laaiacis. 
alabamense Trin. Paspalum. 
alopecuroides L. Pennisetum. 
alopecuroideum L. Pennisetum. 
alopecuroideum Walt. Chaetochloa, 
alopecurus Lam. Pennisetum. 
alsinoides Griseb. Ichnanthus. 
americanum L. Pennisetum. 
amphistemon Wright. Scutachne. 
amplexicaule Rudge. Hymenachne. 
amplifolium Steud. Chaetochloa. 
anomalum Walt. Chaetochloa. 
anthaenantia Kuntze. Anthaenantia. 
antillarum Poir, Pennisetum. 
aquaticum Boec. Sacciolepifl. 
aquaticum Muhl. Sacciolepia. 
arborescens Sieber. Laaiacis. 
arbuaculum Sieber. Isachne. 
aristatum Macf. Echinochloa. 
arundinaceum Swartz. Isachne. 
aturense H. B. K. Homolepis. 
aureum Trin. Axonopue. 
auriculatum Willd. Hymenachne. 
autumnale Boec. Leptoloma. 
badium Scribn. & Merr. Syntherisma. 
bambusioidea Deev. Laaiacis. 
barbatum Lam. Chaetochloa. 
berchtholdiae Doell. Chaetium. 
bermudianum Steud. Sacciolepia. 
berteronianum Schult. Chaetochloa. 
bifidum Bertol. Paspalum. 
blepharophorum Preel. Homolepis. 
brachiatum Poir. Chaetochloa. 
brachyphyllum Steud. Syntherisma. 
brevifolium Kunth. Syntherisma. 
buckleyanum Vasey. Error in Index Kewensis for Paspalum. 
californicum Benth. Valota. 
campylostachyum Hack. Thraaya. 
caudatum Lam. Chaetochloa. 
caudatum Salzm. Sacciolepia. 
cenchroidee Rich. Pennisetum. 
cenchroides Ell. Cenchrus. 
cemuum Willd. Chaetochloa. 
chaetium Steud. Chaetium. 
chauvinii Steud. Laaiacis. 
chrysitee Steud. Axonopua 
chrysoblephare Steud. Axonopua. 
chrysodactylon Trin. Axonopua. 
cognatum Schult. Leptoloma. 
534 CONTRIBUTIONS FBOM THE NATIONAL HERBARIUM. 
Panicum colonum L. Echinochloa, 
compactum Swartz. Laaiacis. 
compositum Neee. Chaetochloa. 
compression Balb. Chaetochloa. 
ceresia Kuntze. Paspalum. 
ciliatiflorum Wood. Anthaenantia, 
cirrhosum Scribn. & Men*. Chaetochloa. 
confertum Desv. ? Isachne. 
corrugatum Ell. Chaetochloa. 
crinitum Willd. Pennieetum. 
crueardeae Willd. Chaetochloa. 
cruscorvi L. Echinochloa. 
crusgalli L. Echinochloa. 
aristatum Purah. Echinochloa. 
hispidum Ell. Echinochloa. 
mite Purah. Echinochloa. 
muticum Ell. Echinochloa. 
purpureum Purah. Echinochloa. 
sabulicola Trin. Echinochloa. 
cruspavonis Neee. Echinochloa. 
cubenee Steud, Opliamenus. 
cultratum Trin. Thrasya, 
curvinerve Hack. Syntherisma. 
dactylon L. Capriola. 
decumbens Roem. & Schult. Paspalum. 
denaispica Poir. Pennisetum. 
digitarioides Rasp. Paepalum. 
diapermum Lam. Isachne. 
disaectum L. Paepalum. 
diseitiflorum Steud. Chaetochloa. 
divaricatum Kunth. Laaiacis. 
divaricatum L. Laaiacis. 
glabrum Kuntze. Laaiacis. 
latifolium Fourn. Laaiacis. 
puberulum Griseb. Laaiacis. 
stenoetachyum Griseb. Laaiacis. 
divaricatum Michx. Festuca. 
divergena Muhl. Leptoloma. 
domingense Zucc. Syntherisma. 
drummondii Vasey. Error in Index Kewenais for Paepalum. 
duchaissingix Steud. Valota. 
dumetorum A. Rich. Chaetochloa. 
durum Griaeb. Scutaclme. 
echinatum Willd. Echinolaena. 
echinolaena Nees. Echinolaena. 
eggersli Hack. Valota. 
elliottianum Schult. Sacciolepis. 
erianthum Poir. Anthaenantia. 
falsum Steud. Valota. 
fasciculatum Lam. . Oplismenus. 
fasciculiflorum Steud. Oplismenus. 
filiforme L. Syntherisma. 
HITCHCOCK AND CHASE—TROPICAL NOBTH AMERICAN PAtflCTJM. 535 
Panicum fimbri&tum Preal. Synth eriama. 
setigerum Fourn. Syntherisma. 
firmum Kunth. Oryzopsis, 
flavesceos Sieb. Eriochloa. 
floridanum Trio.. Paspalum. 
fluviatilis Nees. Sacciolepis. 
fragile Kunth. Leptoloma. 
francoi Steud. Oplismenus. 
frumentaceum Roxb. Echinochloa. 
frumentaceum Salisb. Holcus. 
furcellatum S. Moore. Paspalum. 
fuecescens Willd. Chaetochloa. 
fusciflorum Steud. Leptocoryphium. 
fuscum Sieber. LasiaciB. 
gavanianum Steud. Valota. 
geniculatum Lam. Chaetocbloa. 
georgicum Spreng. Eriochloa. 
gibbum Ell. Sacciolepis. 
glaberrimum Ell. Chaetocbloa. 
glabrum Gaud. Syntherisma. 
missisaippiense Gattinger. Syntherisma. 
glandulosum Nees. Echinolaena. 
glauceecena H. B. K. Isachne. 
glaucescens Nees. Syntherisma. 
glaucescens Salzm. Chaetocbloa. 
glaucescens Willd. Isachne. 
glaucum L. Pennisetum. 
glaucum Rupr. Chaetocbloa. 
glaucum Steud. Chaetocbloa. 
flavescens Ell. Chaetocbloa. 
purpurascens Ell. Chaetocbloa. 
glutinosum Lam. Lasiacis. 
gracilentum Poir. Festuca. 
gracillimum Scribn. Syntherisma. 
grisebachii Nash. Lasiacis. 
grossum Salisb. Echinochloa. 
hamiltonii Kunth. Syntherisma. 
heteranthum Link. Echinolaena. 
hirtellum L. Oplismenus. 
hirtellum Walt. Echinochloa. 
hirticalycinum Boec. Anthaenantia. 
hirticalycum Bosc. Echinochloa. 
hirtum Willd. Isachne. 
hispidum Muhl. Echinochloa. 
holciforme Steud. Echinochloa. 
horizontals G. F. W. Meyer. Syntheriama. 
horridum Salzm. Echinochloa. 
humboldtianum Euntze. Paspalum. 
humifusum Kunth. Syntherisma. 
hydrophilum Schult. Sacciolepis. 
hymenachue Desv. Hymenachne. 
ichnodes Griseb. Ichnanthus. 
ignoratum Kunth. Anthaenantia. 
536 CONTRIBUTIONS FBOM THE NATIONAL HERBARIUM. 
Panicum imberbe Poir. Chaetochloa. 
immersum Trin. Axonopus. 
inaequivalve Kuntze. Paspalum. 
incertum Bosc. Eohmochloa. 
inaulare G. F. W. Meyer. Valota. 
iflchaemum Schreb. Syntherisma. 
isocalycinum Meyer. Homolepis. 
italicum L. Chaetochloa. 
jurgensenii Scribn. & Merr. Chaetochloa. 
lachnanthum Torr. Valota. 
laevigatum Lam. Chaetochloa. 
laevigatum Ell. Chaetochloa. 
laevigatum Muhl. Chaetochloa. 
lagaacae Kuntze. Paspalum. 
lagopua Willd. Eriochrysis. 
lagotis Trin. Ichnanthus. 
lanatum Rottb. Valota. 
lanatum Swartz. Lasiacis. 
eorghoideiim Hamilt. Lasiacis. 
langei Fourn. Homolepis. 
laterale Preal. Olyra. 
leandri Trin. Brachiaria. 
leucites Trin. Syntherisma. 
leucocomum Scribn. Syntherisma. 
leucophaeum H. B. K. Valota. 
liebmannianum Fourn. Laaiacis. 
depauperatum Fourn. Lasiacia. 
lindeni Fourn. Ichnanthus. 
lineare L. Syntherisma. 
lineare Krock. Syntherisma. 
mississippiense Gattinger. Syntherisma. 
linkianum Kunth. Syntherisma. 
litorale Kuntze. Paspalum. 
loliaceum Lam. Oplismenus. 
loliiforme Hochst. Mesosetum. 
longiflorum Trin. Homolepis. 
longisetum Torr. Echinochloa. 
longispicula Doell. Homolepis. 
lutescens Weigel. Chaetochloa. 
macros tachya Nees. Chaetochloa. 
macula turn Aubl. Laaiacis. 
piloeum Fourn. Lasiacis. 
malacophyllum Kuntze. Paspalum. 
martinicenae Griseb. Lasiacis. 
mayarense Wright. Ichnanthus. 
melicarium Michx. Panicularia. 
melicarium Muhl. Sporobolus. 
melinie Trin. Melinis. 
mexicanum Scribn. & Merr. Chaetochloa. 
michauxianum Schult. Eriochloa. 
michauxii Poir. Eriochloa. 
minutiflorum Hasp. Melinis. 
molle Michx. Eriochloa. 
HITCHCOCK AND CHASE—TROPICAL NORTH AMERICAN PANICUM 
Panicum mollissimum Kunth. Syntherisma, 
monobotrys Trin, Paapalum. 
monostachyum H. B. K. Paapalum. 
montanum Poit. Ichnanthus. 
multinerve Desv. Isachne. 
muricatum Michx. Echinochloa. 
myoaurus Rich. Sacciolepis. 
myurus Lam. Sacciolepis. 
neesii Kunth. Syntheriama. 
nemorale Schrad. Ichnanthus. 
nemorosum Swartz. Ichnanthus. 
nemoroeum Willd. Echinolaena. 
nudum Walt. Leptoloma. 
nuttallianum Steud. Opliamenua. 
oaxacenae Steud. Lasiacis. 
obtectum Preal. Paapalum. 
occidentale Nieuwl. Chaetochloa, 
onurus Willd. Chaetochloa. 
orinocense Willd. Lasiacis. 
oxyanthum Steud. Reimarochloa. 
pallena Swartz. Ichmmthue, 
palmeri Vasey. Ixophorus. 
palmifolium Poir. Chaetochloa. 
paniculatum Kuntze. Paapalum. 
paniculiferam Steud. Chaetochloa. 
paractaenoides Trin. Chaetochloa. 
parciflorum Steud. Opliamenus. 
patentissimum Roem. & Schult. Feetuca. 
penicillatum Willd. Chaetochloa. 
petrosum Trin. Thrasya. 
phaeocarpum drummondianum Neea. Syntherisma. 
phaeothrix Scribn. Syntherisma. 
phleiforme Preal. Sacciolepis. 
pittieri Hack. Valota. 
plantagineum Link. Brachiaria. 
platyphyllum Munro. Brachiaria. 
plicatulum Kuntze. Paapalum. 
polyrrhizum Preal. Paapalum. 
polystachion Muh.1. Chaetochloa. 
polyBtachyum Preal. Hymenachne. 
praegnans Steud. Lasiacis. 
pringlei Vasey. Ixophorus. 
procerrimum Hack. Lasiacis, 
prorepens Steud. Oplismenus. 
pulchxum Willd. Axonopus. 
pungens Poir. Echinochloa. 
purpurascena H. B. K. Chaetochloa. 
radicosum Preel. Syntheriama. 
rariflorum Lam. Orthoclada. 
rariflorum Preal. Chaetochloa. 
res titu turn Steud. Chaetochloa. 
rhizophorjim Fourn. Lasiacis. 
rigene Swartz. Isachne. 
V 
538 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
Panicum rigidifolium Kunth. leach ne. 
rigidifoliurn Trin. Sacciolepis. 
roscum Steud. Tricholaena. 
rottboellioides H. B. K. Mesosetum. 
rufum Kunth. Anthaenantia. 
nigelii Griseb. Lasiacis. 
ruscilolium H. B. K. Laaiacis. 
sabuiicola Nees. Echinochloa. 
saccharatum Buckl. Valota. 
sacchariflora Steud. Valota. 
Baccharoides A. Rich. Valota. 
saccharoides Kunth, Paspalum. 
Baccharoides Trin. Tricholaena. 
sanctae-marthae Steud. Oplismenus. 
Banguinale L, Syntheriama. 
simpsoni Yaaey. Syntheriama. 
scabrifolium Nees. Chaetochloa. 
scandens Trin. Chaetochloa. 
Bcariosum Trin. Lasiacis. 
schaffneri Kuntze. Paspalum. 
scheelii Steud. Chaetochloa. 
echiedeanum Trin. Paspalum. 
schiedei Spreng. Senitee. 
schlechtendalii Fourn. Ichnanthus. 
monstrosum Fourn. Ichnanthus. 
schultesii Steud. Oplismenus. 
scindens Nees. Echinochloa. 
eemirugoeum Nees. Chaetochloa. 
senescens Trin. Paspalum. 
sericatum Steud. Eriochloa. 
serotinum Trin. Syntheriama. 
setarium Lam. Oplismenus. 
setosum Swartz. Chaetochloa. 
simpsoni Beal. Syntheriama. 
sloanei Griseb. Lasiacis. 
sorghoideum Desv. Lasiacis. 
speciosum Nees. Chaetochloa. 
spectabile Nees. Echinochloa. 
squamatum Fourn. ? Paspalu m. 
stipatum Presl. Syntheriama. 
striatum Lam. Sacciolepis. 
strumosum Presl. Sacciolepis. 
subspicatum Desv. ? Sacciolepis. 
sulcatum Aubl. Chaetochloa. 
swartzianum Hitchc. Lasiacis. 
sylvaticum Lam. Oplismenus. 
ten&x Rich. Chaetochloa. 
thrasya Trin. Thrasya. 
fhiasyoidee Trin. Thrasya. 
trachyspermum Nees. Isachne. 
trinii Kunth. Sacciolepis. 
trinii Mor. Error in Index Kewensis for Echinolaena. 
triquetrum Willd. Chaetochloa. 
triticeum Willd. Cyperaceae. 
HITOHCOOK AND CHASE—TROPICAL NORTH AMERICAN PANICUM. 539 
Panicum tumeecera Trin. Homolepis. 
uncinatum Raddi. Echinolaena. 
unispicatum Scribn. & Merr. Paspalum. 
velutinum Boec. Syntherisma. 
ventenatii Steud. Ghaetocbloa. 
versicolor Nieuwl. Chaetochloa. 
verticillatum L. Chaetochloa. 
vestitunx Eunth. Valota. 
vilvoides Trin. Sacciolepis. 
viride L. Chaetochloa. 
viridiflorum Nees. Homolepis. 
vulpieetum Lam. Fennisetum, 
waiter! Pursh. Echinochloa. 
LIST OF NEW SPECIES. 
Panicum sucosum Hitchc. & Chase  
Panicum altum Hitchc. & Chase  
Panicum milleflorum Hitchc. & Chase  
Panicum stevensi&num Hitchc. & Chase. 
Panicum pyrulaiium Hitchc. & Chase  
Panicum izieptum Hitchc. & Chase  
Panicum chiriquiense Hitchc. & Chase... 
Panicum stagnatile Hitchc. & Chase  
Panicum grande Hitchc. & Chase  
Pftga 
475 
488 
494 
498 
608 
509 
527 
528 
529