A Check List of the /V^phibiaijs and Reptiles of EcuadorWITHA Bibliography of EcuadoriaisI Herpetology
KENNETH MIYATA
Museum of Comparative ZoologyHarvard University(present address)lational Museum of Natural HistorySmithsonian Institution
smithsonianherpetological informationSERVICENO. 541982 ^^6RaRIES

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INTRODUCTION
There has been a renewed interest in the herpetology of Ecuadorover the past twenty years, initiated largely through the efforts ofthe late Dr. James A. Peters. During this period large collections ofEcuadorian material have been made by a number of workers and thismaterial is now deposited in North American collections, notably theUniversity of Kansas Museum of Natural History, the United StatesNational Museum of Natural History, and the Museum of Comparative Zoo-logy. These collections have shown that the Ecuadorian herpetofauna isexceptionally rich, and until now a complete checklist has not beenaval lable
.
As is the case with any checklist, this is very much a work-in-progress. Of the 682 taxa recorded herein from Ecuador, 136 have beendescribed since 1970, and I am aware of many more species awaitingdescription. There is no shortage of basic exploratory work yet to bedone. Although most of the major habitats and regions have been sam-pled, there still remain large blank areas on collecting maps that willno doubt turn up many surprises. I have made efforts to check outsuspicious literature records, but in some cases species may beincluded on this list due to either misidenti f icat ions or incorrectlocality data.I chose not to include information on distribution in this listfor two reasons. The distributions of many species are poorly known,and many species appear to have extremely restricted ranges. Asimplified notation of distribution would be misleading because itwould imply that distributions are well known even when they are not.Also, many of the older records, both in the literature and in col-lections, are suspect, and if these taxa are not well represented inrecent collections it is not possible to make realistic estimations ofdistribution. I have made every effort to eliminate these, but some nodoubt remain. Field work in Ecuador has characteristically beenexploratory, and only a few localities have been sampled adequately.More faunal work concentrating on specific areas is needed, and thiswork is probably best done by resident naturalists.The bibliography is a current (to March 1982) and fairly compre-hensive guide to the systematic literature on the Ecuadorian herpeto-fauna. It contains all references to original descriptions ofEcuadorian taxa, as well as ecological and systematic papers thatmention Ecuadorian specimens. This latter coverage is incomplete, but Ibelieve that the bibliography includes most references likely to be use-ful to any student of the Ecuadorian herpetofauna. I relied heavily onVanzolini's bibliography of South American reptiles (1978) for locatingreferences on reptiles, and his comments on many of the older papersshould be consulted before any great effort is made to locate obscurepapers
.
I hope that publication of this list and bibliography will stimu-late interest in the enormously diverse Ecuadorian herpetofauna. Inmany respects Ecuador is a microcosm of tropical South America, andstudents of amphibian and reptile biology are sure to find much ofinterest there.
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ACKNOWLEDGEMENTS
Drafts of this checklist were read by Ronald I. Crombie, WilliamE. Duellman, W. Ronald Heyer, John D. Lynch, Roy W. McDiarmid, andErnest E. Williams. I am grateful for their comments, which helpedbring many names up to date and added several additional species to thelist. I also thank the many sources of funding for the field work inEcuador that was the impetus for creating this list: Earthwatch and theCenter for Field Studies in Belmont, Massachusetts, the Barbour Fund ofthe Museum of Comparative Zoology, Harvard University, the Richmond andAnderson Funds of the Department of Biology, Harvard University, andthe Museu de Zoologia of the Universidade de SIo Paulo. GabrielleDundon and the Friends Public Programs of the MCZ allowed me the use oftheir printer for typing the manuscript.
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AMPHIBIA
ANURA
BUFONIDAE
Atelopus arthuri Peters 1973A. balios Peters 1973A. bomolochos Peters 1973A. boulengeri Peracca 1904A. coynei Miyata 1980aA. elegans (Boulenger 1882a)A. halihelos Peters 1973A. ignescens (Cornalia 1849)A. longirostris Cope 1868A. lynchi Cannatella 1981A. mindoensis Peters 1973A. nepiozoraus Peters 1973A. pachydermus (0. Schmidt 1857)A. palraatus Andersson 1945A. planispinus Jimenez de la Espada 1875A. pulcher pulcher (Boulenger 1882a)Bufo blombergi Myers and Funkhouser 1951B. caeruleocellatusFowler 1913B. caeruleostictus Gunther 1859dB. ceratophrys Boulenger 1882aB, chanchanensis Fowler 1913B. coniferus Cope 1862aB. dapsilis Myers and Carvalho 1945B. glaberrimus Gunther 1868B. haematiticus Cope 1862aB. marinus (Linnaeus 1758)B. spinulosus Wiegraann 1835B. typhonius (Linnaeus 1758)Dendrophryniscus minutus Melin 1941Osornophryne bufoniformis (Peracca 1904)Rhamphophryne festae (Peracca 1904)
CENTROLENIDAECentrolene geckoideum Jimenez de la Espada 1872Centrolenella anoraala Lynch and Duellman 1973C. audax Lynch and Duellman 1973C. balionota Duellman 1981C. buckleyi (Boulenger 1882a)C. cochranae (Goin 1961a)C. flavopunctata Lynch and Duellman 1973C. fleischmanni (Boettger 1893)C. grand isonae Cochran and Goin 1970C griffithsi (Goin 1961a)C. heloderma Duellman 1981
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Centrolenella lynchi Duellman 1980C. medemi Cochran and Coin 1970C. megacheira Lynch and Duellman 1973C. raidas Lynch and Duellman 1973C. munozorum Lynch and Duellman 1973C. ocellifera (Boulenger 1899c)C. pellucida Lynch and Duellman 1973C. peristicta Lynch and Duellman 1973C. pipilata Lynch and Duellman 1973C. prosoblepon (Boettger 1892)C. resplendens Lynch and Duellman 1973C. siren Lynch and Duellman 1973C. spiculata Duellman 1976C. spinosa Taylor 1949C. valerioi (Dunn 1931)
DENDROBATIDAE
Colostethus anthracinus Edwards 1971C. bocagei (Jimenez de la Espada 1871)C. brunneus (Cope 1887)C. elachyhistus Edwards 1971C. fuliginosus (Jimenez de la Espada 1871)C. infraguttatus (Boulenger 1898a)C. intermedins (Andersson 1945)C. kingsburyi (Boulenger 1918)C. latinasus (Cope 1863)C. marchesianus (Melin 1941)C. pratti (Boulenger 1899a)C. pulchellus (Jimenez de la Espada 1871)C. sauli Edwards 1974C. taeniatus (Andersson 1945)C. vertebralis (Boulenger 1899b)C. whymperi (Boulenger 1882a)Dendrobates abditus Myers and Daly 1976aD. anthonyi (Noble 1921)D. boulengeri (Barbour 1909)D. erythromos Vigle and Miyata 1980D. espinosai (Funkhouser 1956)D. femoralis (Boulenger 1883a)D. histrionicus (Berthold 1846a)D. parvulus Boulenger 1882aD. pictus (Bibron in Tschudi 1838)D. quinquevittatus Steindachner 1864D. tricolor (Boulenger 1899b)D. trivittatus (Spix 1824a)D. zaparo (Silverstone 1976)
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HYLIDAE
Agalychnis calcarifer Boulenger 1902bA. craspedopus (Funkhouser 1957)A. litodryas (Duellman and Trueb 1967)A. spurrelli (Boulenger 1912)Amphignathodon guentheri Boulenger 1882aGastrotheca angustifrons (Boulenger 1898a)G. cavia Duellman 1974aG. cornuta (Boulenger 1898a)G. humbertoi Lutz 1977G. lojana Parker 1932G. longipes (Boulenger 1882a)G. monticola Barbour and Noble 1920G. orophylax Duellman and Pyles 1980G. plumbea (Boulenger 1882a)G. psychrophila Duellman 1974aG. riobambae (Fowler 1913)G. testudinea (Jimenez de la Espada 1871)G. weinlandii (Steindachner 1892)Hemiphractus bubalus (Jimenez de la Espada 1871)H. fasciatus Peters 1862bH. johnsoni (Noble 1917)H. proboscideus (Jimenez de la Espada 1871)H. scutatus (Spix 1824a)Hyla alboguttata Boulenger 1882aH. albopunctulata Boulenger 1882aH. alytolylax Duellman 1972bH. bifurca Andersson 1945H. boans (Linnaeus 1758)H. bokermanni Goin 1960H. brevifrons Duellman and Crump 1974H. calcarata Troschel 1848H. carnifex Duellman 1969bH. Columbiana Boettger 1892H. crepitans Wied-Neuwied 1824H. fasciata Gunther 1859aH. favosa Cope 1885H. geographica Spix 1824aH. granosa Boulenger 1882aH. gryllata Duellman 1973aH. lanciformis lanciformis (Cope 1871a)H. larinopygion Duellman 1973aH. leucophyllata (Beireis 1783)H. lindae Duellman and Altig 1978H. marmorata (Laurenti 1768)H. rainuta Peters 1872aH. parviceps Boulenger 1882aH. pellucens Werner 1901H. phyllognatha Melin 1941H. picturata Boulenger 1882a
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Hyla punctata Schneider 1799H. rhodopepla Gunther 1859aH. riveroi Cochran and Coin 1970H. rosenbergi Boulenger 1898aH. rossalleni Coin 1957H. sarayacuensis Shreve 1935H. torrenticola Duellman and Altig 1978H. triangulura Gunther 1869H. tuberculosa Boulenger 1882aNyctimantis rugiceps Boulenger 1882aOlolygon cruentonmia (Duellman 1972a)0. funerea (Cope 1874)0. garbei (Miranda-Ribeiro 1924)0. quinquefasciata (Fowler 1913)0. rubra (Daudin 1802)0. sugillata (Duellman 1973a)Osteocephalus buckleyi (Boulenger 1882a)0. leprieurii (Dum^ril and Bibron 1841)0. taurinus Steindachner 18620. verrucigerus (Werner 1901)Phrynohyas coriacea (Peters 1867)P. venulosa (Laurenti 1768)Phyllomedusa buckleyi (Boulenger 1882a)P. palliata Peters 1872bP. perinesos Duellman 1973aP. tarsius (Cope 1868)P. toraopterna (Cope 1868)P. vaillanti Boulenger 1882aSrailisca phaeota (Cope 1862a)Sphaenorhynchus carneus (Cope 1868)S. dorisae (Coin 1957)S. eurhostus Rivero 1969Trachycephalus jordani (Stejneger and Test 1891)
LEPTODACTYLIDAE
Adenoraera andreae (Muller 1923)A. hylaedactyla (Cope 1868)Barycholos pulcher (Boulenger 1898a)Ceratophrys cornuta (Linnaeus 1758)C. stolzmanni scaphiopeza Peters 1967aEdalorhina perezi Jimenez de la Espada 1871Eleutherodactylus achatinus (Boulenger 1898a)E. acerus Lynch and Duellman 1980E. actites Lynch 1979bE. acuminatus Shreve 1935E. altamzonicus Barbour and Dunn 1921E. anomalus (Boulenger 1898a)E. append iculatus (Werner 1894b)E. atratus Lynch 1979aE. balionotus Lynch 1979a
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Eleutherodactylus baryecuus Lynch 1979aE. buckleyi (Boulenger 1882a)E. cajamarcensis Barbour and Noble 1920E. calcarulatus Lynch 1976aE. caprifer Lynch 1977E. carvalhoL Lutz _in Lutz and Kloss 1952E. celator Lynch 1976aE. cerastes Lynch 1975cE. chalceus (Peters 1873)E. chloronotus Lynch 1969aE. colodactylus Lynch 1979aE. condor Lynch and Duellman 1980E. conspicillatus (Gunther 1858)E. cornutus (Jimenez de la Espada 1871)E. cremnobates Lynch and Duellman 1980E. crenunguis Lynch 1976aE. croceoinguinus Lynch 1968aE. crucifer (Boulenger 1899b)E. cruentus (Peters 1873)E. cryophilus Lynch 1979aE. cryptoraelas Lynch 1979aE. curtipes (Boulenger 1882a)E. devillei (Boulenger 1880)E. diadematus (Jimenez de la Espada 1875)E. diastema (Cope 1876a)E. dolops Lynch and Duellman 1980E. duellmani Lynch 1980cE. elassodiscus Lynch 1973bE. eriphus Lynch and Duellman 1980E. galdi (Jimenez de la Espada 1871)E. gladiator Lynch 1976cE. glandulosus (Boulenger 1880)E. gularis (Boulenger 1898a)E. helonotus (Lynch 1975c)E. ignicolor Lynch and Duellman 1980E. incanus Lynch and Duellman 1980E. incomptus Lynch and Duellman 1980E. inusitatus Lynch and Duellman 1980E. lacrimosus (Jimdnez de la Espada 1871)E. lanthanites Lynch 1975aE. latidiscus (Boulenger 1898a)E. leoni Lynch 1976cE. leucopus Lynch 1976dE. lividus Lynch and Duellman 1980E. longirostris (Boulenger 1898a)E. loustes Lynch 1979cE. luteolateralis Lynch 1976cE. lymani Barbour and Noble 1920E. malkini Lynch 1980bE. martiae Lynch 1974cE. modipeplus Lynch 1981E. rauricatus Lynch and Miyata 1980
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Eleutherodactylus necerus Lynch 1975cE. nigrogriseus (Andersson 1945)E. nigrovittatus Andersson 1945E. nyctophylax Lynch 1976aE. ockendeni (Boulenger 1912a)E. ocreatus Lynch 1981E. orcesi Lynch 1972E. orestes Lynch 1979aE. ornatissimus (Despax 1911c)E. orphnolairaus Lynch 1970E. parvillus Lynch 1976aE. pastazensis Andersson 1945E. paululus Lynch 1974cE. percultus Lynch 1979aE. peruvianas (Melin 1941)E. peters i Lynch and Duellman 1980E. phoxocephalus Lynch 1979aE. prolatus Lynch and Duellman 1980E. proserpens Lynch 1979aE. pseudoacurainatus Shreve 1935E. pugnax Lynch 1973bE. pycnodermis Lynch 1979aE. pyrrhomerus Lynch 1979aE. quaquaversus Lynch 1974cE. quinquagesimus Lynch and Trueb 1980E. riveti (Despax 1911c)E. roseus (Boulenger 1902b)E. rubicundus (Jimenez de la Espada 1875)E. ruidus Lynch 1979aE. sobetes Lynch 1980cE. spinosus Lynch 1979aE. subsigillatus (Boulenger 1902b)E. sulcatus (Cope 1874)E. supernatis Lynch 1980aE. surdus (Boulenger 1882a)E. taeniatus (Boulenger 1912a)E. tenebrionis Lynch and Miyata 1980E. thymalopsoides Lynch 1976aE. thymelensis Lynch 1972E. trachyblepharis (Boulenger 1918)E. trepidotus Lynch 1968bE. unistrigatus (Giinther I859d)E. variabilis Lynch 1968aE. ventrimarmoratus (Boulenger 1912a)E. versicolor Lynch 1979aE. vertebralis (Boulenger 1886)E. vidua Lynch 1979aE. walkeri Lynch 1974bE. w-nigrum (Boettger 1892)Ischnocneraa quixensis (Jimenez de la Espada 1872)I. simmonsi Lynch 1974aLeptodactylus amazonicus Heyer 1978
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Leptodactylus knudseni Heyer 1972L. labrosus Jimenez de la Espada 1875L. melanonotus (Hallowell 1861)L. pentadactylus Laurenti 1768L. rhodomystax Boulenger 1883L. stenodema Jimenez de la Espada 1875L. ventrimaculatus Boulenger 1882aL. wagneri (Peters 1862c)Lithodytes lineatus (Schneider 1799)Phrynopus brunneus Lynch 1975bP. flavomaculatus (Parker 1938)P. peraccai Lynch 1975bPhyllonastes lochites (Lynch 1976b)Physalaemus petersi (Jimenez de la Espada 1872)P. pustulatus (Shreve 1941)Telmatobius cirrhacelis Trueb 1979T. niger Barbour and Noble 1920T. vellardi Munsterraann and Leviton 1959Vanzolinius discodactylus (Boulenger 1883)
MICROHYLIDAEChiasmocleis anatipes Walker and Duellman 1974C. bassleri Dunn 1949C. ventrimaculata (Andersson 1945)Ctenophryne geayi Mocquard 1904Glossostoraa aequatoriale (Peracca 1904)G. aterrimura Gunther 1901Hamptophryne boliviana (Parker 1927)Synapturanus rabus Pyburn 1976Syncope antenori Walker 1973
PIPIDAE
Pipa pipa (Linnaeus 1758)
RANIDAE
Rana palmipes Spix 1824a
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CAUDATA
PLETHODONTIDAE
Bolitoglossa altaraazonica (Cope 1874)B. chica Brame and Wake 1963B. ecuatoriana Brame and WakeB. palraata (Werner 1897)B. sima (VaiUant 1911) 1972
APODACAECILIIDAECaecilia abitaguae Dunn 1942C. albiventris Daudin 1802C. attenuata Taylor 1968C. bokermanni Taylor 1968C. crassisquama Taylor 1968C. disossea Taylor 1968C. dunni Hershkovitz 1938C. nigricans Boulenger 1902bC. orientalis Taylor 1968C. pachynema Glinther 1859dC. subterminalis Taylor 1968C. tentaculata Linnaeus 1758C. tenuissima Taylor 1973Microcaecilia albiceps (Boulenger 1882b)Oscaecilia bassleri (Dunn 1942)0. equatorialis Taylor 1973Siphonops annulatus (Mikan 1820)
ICHTHYOPHIIDAE
Epicrionops bicolor Boulenger 1883bE. marraoratus Taylor 1968E. petersi petersi Taylor 1968
TYPHLONECTIDAE
Potomotyphlus kaupii (Berthold 1859)
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REPTILES
CHELONIA
CHELIDAEChelus firabriatus (Schneider 1783)Phrynops geoffroanus tuberculosa (Peters 1870)P. gibbus (Schweigger 1812)P. nasutus (Schweigger 1812)Platerays platycephala (Schneider 1792)
CHELYDRIDAEChelydra serpentina acutirostris Peters 1862d
EMYDIDAE
Rhinoclemmys annulata (Gray 1860)R. nasuta (Boulenger 1902b)R. punctularia melanosterna (Gray 1861)
KINOSTERNIDAE
Kinosternon scorpioides scorpioides (Linnaeus 1758)K. spurrelli Boulenger 1913b
PELOMEDUSIDAE
Podocnemis expansa (Schweigger 1812)P. unifilis Troschel 1848
TESTUDINIDAEGeochelone denticulata (Linnaeus 1758)
CROCODYLIA
ALLIGATOR I DAECaiman crocodilus crocodilus (Linnaeus 1758)Melanosuchus niger (Spix 1825)Paleosuchus palpebrosus (Cuvier 1807)
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Paleosuchus trigonatus (Schneider 1801)
CROCODYLIDAE
Crocodylus acutus Cuvier 1807
SAURIA
ANGUIDAE
Diploglossus monotropis (Kuhl 1820)
GEKKONIDAE
Gonatodes caudiscutatus (Gunther 1859d)G. concinnatus (0' Shaughnessy 1881)G. humeralis (Guichenot 1855)Heraidactylus mabouia (Moreau de Jonnes 1818)Lepidodactylus lugubris (Duradril and Bibron 1836)Lepidoblephar is buchwaldi Werner 1910L. festae festae Peracca 1897L. intermedius Boulenger 1914L. ruthveni Parker 1926Phyllodactylus inaequalis Cope 1876bP. purailus Dixon and Huey 1970P. reissii Peters 1862dPseudogonatodes guianensis Parker 1935Sphaerodactylus scapularis Boulenger 1902bThecadactylus rapicauda (Houttuyn 1782)
IGUANIDAE
Anolis aequator ialis Werner 1894aA. binotatus Peters 1863aA. biporcatus parvauritus Williams 1966A. bitectus Cope 1864A. bombiceps Cope 1876bA. chloris Boulenger 1898aA. chocorum Williams and Duellman 1967A. chrysolepis scypheus Cope 1864A. fasciatus Boulenger 1885A. festae Peracca 1904A. fraseri Gunther 1859dA. fuscoauratus fuscoauratus D'Orbigny 1837A. gemmosus 0' Shaughnessy 1875A. gracilipes Boulenger 1898aA. granuliceps Boulenger 1898a
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Anolis maculiventris Boulenger 1898aA. nigrolineatus Williams 1965A. ortonii Cope 1868A. parilis Williams 1975A. peraccae Boulenger 1898aA. princeps Boulenger 1898aA. proboscis Peters and Orces-V. 1956A. punctatus boulengeri 0' Shaughnessy 1881A. trachyderma Cope 1876bA. transversalis Dum^ril in Dum^ril and Dum^ril 1851Basiliscus basiliscus basiliscus (Linnaeus 1758)B. galeritus Dum^ril iii Dum^ril and Dum^ril 1851Enyalioides cofanorum Duellman 1973bE. heterolepis (Bocourt 1874)E. laticeps laticeps (Guichenot 1855)E. 1. festae Peracca 1897E. microlepis (0' Shaughnessy 1881)E. oshaughnessyi (Boulenger 1881
)
E. praestabilis (0' Shaughnessy 1881)Iguana iguana iguana (Linnaeus 1758)Morunasaurus annularis (0' Shaughnessy 1881)Ophryoessoides aculeatus angulifer Werner 19010. erythrogaster Hallowell 18570. iridescens (Gunther 1859d)Phenacosaurus orcesi Lazell 1969Plica plica (Linnaeus 1758)P. umbra ochrocollaris (Spix 1859c)Polychrus femoralis Werner 1910P. gutturosus gutturosus Berthold 1846P. g. spurrellii Boulenger 1914P. marmoratus (Linnaeus 1758)Stenocercus carrioni Parker 1934aS. festae (Peracca 1897)S. guentheri (Boulenger 1885)S. haenschi (Werner 1901)S. humeralis (Gunther 1859b)S. ornatus (Gray 1845)S. rhodomelas (Boulenger 1899b)S. simonsii Boulenger 1899bS. varius Boulenger 1885Tropidurus occipitalis occipitalis Peters 1871bT. peruvianus peruvianas (Lesson 1830)Uracentron flaviceps (Guichenot 1855)
TEIIDAE
Alopoglossus atriventris Duellman 1973bA. buckleyi (0* Shaughnessy 1881)A. carinicaudatus (Cope 1876b)A. copii Boulenger 1885
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Alopoglossus festae Peracca 1904Ameiva ameiva petersii Cope 1868A. bridgesii (Cope 1869)A. edrecantha Bocourt 1874A. orcesi Peters 1964bA. septemlineata Dum^ril and Dum^ril 1851Anadia petersi Oftedal 1974A. rhorabifera (Glinther 1859d)Arthrosaura reticulata (0' Shaughnessy 1881)Bachia trisanale trisanale (Cope 1868)Callopistes f lavipunctatus (Dum^ril and Bibron 1839)Dicrodon guttulatum Dum^ril and Bibron 1839Dracaena guianensis Daudin 1802Echinosaura horrida horrida Boulenger 1890Euspondylus guentheri (O' Shaughnessy 1881)E. maculatus Tschudi 1845E. ocellifer (Werner 1901)Iphisa elegans elegans Gray 1851Kentropyx altaraazonicus Cope 1876bK. calcaratus Spix 1825K. pelviceps Cope 1876bLeposoma parietale (Cope 1885)Neusticurus cochranae Burt and Burt 1931N. ecpleopus Cope 1876bN. strangulatus strangulatus (Cope 1868)Pholidobolus affinis (Peters 1862a)P. annectens (Parker 1930)p. raacbrydei Montanucci 1973p. montium (Peters 1862a)p. prefrontalis Montancucci 1973Prionodactylus argulus (Peters 1862a)p. dicrus Uzzell 1973p. manicatus manicatus (0' Shaughnessy 1881)p. vertebralis (0' Shaughnessy 1879)Proctoporus columbianus Andersson 1914p. hypostictus Boulenger 1912b
^p. meleagris Boulenger 1885p. oculatus (0' Shaughnessy 1879)p. simoterus (0' Shaughnessy 1879)p. unicolor (Gray 1858)Ptychoglossus brevifrontalis Boulenger 1912bp. picticeps (Cope 1885)Teuchocercus keyi Fritts and Smith 1969aTupinarabis teguixin (Linnaeus 1758)
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AMPHISBAENIA
AMPHISBAENIDAE
Amphisbaena fuliginosa bassleri Vanzolini 1951A. f. varia Laurenti 1768
SERPENTES
ANOMALEPIDIDAE
Anoraalepis flavapices Peters 1957Liotyphlops petersi (Boulenger 1889)
ANILIIDAE
Anilius scytale scytale (Linnaeus 1758)
BOIDAE
Boa constrictor constrictor Linnaeus 1758B. c. imperator Daudin 1803aCorallus annulatus blombergi (Rendahl and Vestergren 1941)C. a. colombianus (Rendahl and Vestergren 1941)C. caninus (Linnaeus 1758)C. enydris enydris (Linnaeus 1758)Epicrates cenchria cenchria (Linnaeus 1758)Eunectes murinus raurinus (Linnaeus 1758)
COLUBRIDAE
Atractus badius (Boie 1827)A. bocourti Boulenger 1894aA. carrioni Parker 1930bA. collaris Peracca 1897A. dunni Savage 1955A. ecuadorensis Savage 1955A. elaps (GUnther 1858)A. gaigei Savage 1955A. lehmanni (Boettger 1898)A. major Boulenger 1894aA. microrhynchus (Cope 1868)A. modes tus Boulenger 1894aA. multicinctus (Jan in Jan and Sordelli 1865)A. occidentalis Savage 1955A. occipitoalbus (Jan 1862)A. paucidens Despax 1910
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Atractus resplendens Werner 1901A. roulei Despax 1910Chironius carinatus (Linnaeus 1758)C. flavopictus (Werner 1909)C. fuscus (Linnaeus 1758)C. grand is quamis (Peters 1868)C. multiventris Schmidt and Walker 1943C. schleuteri (Werner 1899)C. scurrulus (Wagler jji Spix 1824b)C. vicinus (Boulenger 1915)Clelia clelia clelia (Daudin 1803b)C. equatoriana (Amaral 1924)Coniophanes dromiciformis (Peters 1863b)C. fissidens fissidens (Gunther 1858)Dendrophidion bivittatus (Dumdril, Bibron, and Dum^ril 1854)D. brunneus (Gunther 1858)D. dendrophis (Schlegel 1837)Diaphoralepis wagneri Jan 1863Dipsas catesbyi (Sentzen 1796)D. elegans (Boulenger 1896a)D. gracilis (Boulenger 1902b)D. indica indica Laurenti 1768D. i. ecuadorensis Peters 1960aD. latifasciata (Boulenger 1913a)D. latifrontalis (Boulenger 1905)D. oreas (Cope 1868)D. pavonina Schegel 1837D. temporalis (Werner 1901)D. variegata variegata (Dum^ril, Bibron, and Dum^ril 1854)D. V. nicholsi (Dunn 1933)D. vermiculata Peters 1960aDrepanoides anomalus (Jan 1863)Drymarchon corais corais (Boie 1827)D. c. melanurus (Dum^ril, Bibron, and Dum^ril 1854)Drymobius rhombifer (Gunther 1860)Dryraoluber dichrous (Peters 1863b)Emmochliophis fugleri Fritts and Smith 1969bErythrolamprus aesculapii aesculapii (Linnaeus 1758)E. guentheri Garraan 1883E. rairaus raimus (Cope 1868)E. m. raicrurus Dunn and Bailey 1939Helicops angulatus (Linnaeus 1758)H. pastazae Shreve 1934H. petersi Rossman 1976Imantodes cenchoa cenchoa (Linnaeus 1758)I. inornatus (Boulenger 1896a)I. lentiferus (Cope 1894)Lampropeltis triangulum micropholis (Cope 1860a)L. bimaculatus lamonae Dunn 1944L. epinephalus epinephalus (Cope 1862b)L. e. ecaudorensis Laurent 1949Leptodeira annulata annulata (Linnaeus 1758)
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Leptodeira septentr ionalis ornata (Bocourt 1884)L. s. larcorura Schmidt and Walker 1943Leptophis ahaetulla bocourti Boulenger 1898aL. a. nigromarginatus (Glinther 1866)L. a. occidentalis (Glinther I859d)L. cupreus (Cope 1868)L. depressirostris (Cope 1861)L. riveti Despax 1910Liophis albiventris Jan 1863bL. bimaculatus laraonae (Dunn 1944)L. breviceps Cope 1860aL. cobella (Linnaeus 1758)L. epinephalus epinephalus Cope 1862bL. e. ecuadorensis (Laurent 1949)L. festae (Perraca 1897)L. fraseri Boulenger 1894aL. poecilogyrus (Wied-Neuwied 1825)L. purpurans (Dum^ril, Bibron, and Dum^ril 1854)L. reginae (Linnaeus 1758)L. taeniurus Tschudi 1845L. typhlus (Linnaeus 1758)L. undulatus (Wied-Neuwied 1824)Mastigodryas boddaerti boddaerti (Sentzen 1796)M. pulchriceps (Cope 1868)Ninia atrata (Hallowell 1845)N. hudsoni Parker 1940Nothopsis rugosus Cope 1871bOxybelis aeneus (Wagler in Spix 1824b)0. argenteus (Daudin 1803b)0. brevirostris (Cope 1861)0. fulgidus (Daudin 1803b)Oxyrhopus fitzingeri frizzelli Schmidt and Walker 19430. formosus (Wied-Neuwied 1820)0. leucomelas (Werner 1916)0. melanogenys (Tschudi 1845)0. petola digitalis (Reuss 1834)0. p. sebae Dum^ril, Bibron, and Dum^ril 1854Philodryas elegans rufidorsatus (Glinther 1858)P. simonsii Boulenger 1900P. viridissimus (Linnaeus 1758)Pliocercus euryzonus euryzonus Cope 1862bPseudoboa coronata Schneider 1801Pseustes poecilonotus polylepis (Peters 1867)P. shropshirei (Barbour and Amaral 1924)P. sulphureus sulphureus (Wagler in Spix 1824b)Rhadinaea brevirostris (Peters 1863b)R. decorata (Glinther 1858)R. fulviceps Cope 1886R. lateristriga Berthold 1859Rhinobothryura bovallii Andersson 1916Saphenophis atahuallpae (Steindachner 1901)S. boursieri (Jan in Jan and Sordelli 1867)
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Sibon dunni Peters 1957bS. nebulata nebulata (Linnaeus 1758)S. n. leucomelas (Boulenger 1896b)Siphlophis cervinus (Laurenti 1768)Spilotes pullatus pullatus (Linnaeus 1758)Stenorhina degenhardtii degenhardtii (Berthold 1846)Synophis bicolor Peracca 1896S. lasallei (Niceforo-Maria 1950)S. miops Boulenger 1898aTantilla andinista Wilson and Mena 1980T. equatoriana Wilson and Mena 1980T. insulimontana Wilson and Mena 1980T. melanocephala (Linnaeus 1758)T. petersi Wilson 1979T. supracincta Peters 1863bThamnodynastes strigilis (Thunberg 1787)Tretanorhinus taeniatus Boulenger 1903aTripanurgos compressus (Daudin 1803b)Umbrivaga pygmaeus (Cope 1868)Xenodon rabdocephalus rabdocephalus (Wied-Neuwied 1824)X. severus (Linnaeus 1758)Xenopholis scalaris (Wucherer 1862)
ELAPIDAE
Leptomicrurus narduccii (Jan 1863b)Micrurus ancoralis ancoralis (Jan jja Jan and Sordelli 1872)M. annelatus annelatus (Peters 1871a)M. bocourti bocourti (Jan rn Jan and Sordelli 1872)M. dumerilii transandinus Schmidt 1936M. filiforrais filiformis (Glinther 1859b)M. hemprichi ortoni Schmidt 1953aM. langsdorffi langsdorffi Wagler ui Spix 1824bM. 1. ornatissimus (Jan 1858)M. leraniscatus helleri Schmidt and Schmidt 1925M. mertensi Schmidt 1936M. mipartitus decussatus (Dum^ril, Bibron, and Dura^ril 1854)M. spixii obscurus (Jan jji Jan and Sordelli 1872)M. steindachneri steindachner i (Werner 1901)M. s. orcesi Roze 1967M. s. petersi Roze 1967M. surinamensis surinamensis (Cuvier 1817)M. tschudii olssoni Schmidt and Schmidt 1925
LEPTOTYPHLOPIDAE
Leptotyphlops amazonicus Ore jas-Miranda 1969L. anthracinus Bailey 1946L. guayaquilensis Ore jas-Miranda and Peters 1970
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Leptotyphlops subcrotilla Klauber 1939L. tenella Klauber 1939
TROPIDOPHIDAE
Trachyboa boulengeri Peracca 1904T. gularis Peters 1861aTropidophis battersbyi Laurent 1949T. taczanowskyi (Steindachner 1860)
TYPHLOPIDAE
Typhlops reticulatus (Linnaeus 1758)
VIPERIDAE
Bothrops albocarinatus Shreve 1934B. alticolus Parker 1934aB. atrox (Linnaeus 1758)B. bilineatus smaragdinus Hoge 1966B. castelnaudi Dumdril, Bibron, and Dum^ril 1854B. hyoprorus Amaral 1935B. lojanus Parker 1930aB. microphthalmus microphthalmus Cope 1876bB. nasutus Bocourt 1868B. pulcher (Peters 1862e)B. punctatus (Garcia 1896)B. schlegelii (Berthold 1846)B. xanthogrammus (Cope 1868)Lachesis muta rauta (Linnaeus 1758)
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1927. The brevicipitid frogs allied to the genus Gastrophryne .Occ . Pap. Mus . Zool. Univ. Michigan, (187): 1-6.
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1932. Some new or rare reptiles and amphibians from southernEcuador. Ann. Mag. Nat. Hist., (10)9:21-26.
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1861b. Zwei neue Schlangen aus dem Gattung Mizodon und Bothriop-sis. Monats. Akad. Wiss. Berlin, 1861:358-359.
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1862c. Eine neue Gattung von Laubfrbschen , Plectromantis , ausEcuador vor , Monats . Akad . Wiss. Berlin, 1862:232-233.
1862d. Mittheilung liber einige neuen Phyllodactylus aus Guaya-quil. Monats. Akad. Wiss. Berlin, 1862:626-627.
1862e. Einen Vortrag liber die craniologischen Verschiedenheitender Grubbenottern ( Trigonocephali ) und liber eine neueArt der Gattung Bothrioechis . Monats. Akad. Wiss. Berlin,1862:670-674.
1863a. Eine Mittheilungen liber einige neue Arten der Saurier-Gattung Anolis . Monats. Akad. Wiss. Berlin, 1863:135-149.
1863b. Uber einige neue oder weniger bekannte Schlangenarten deszoologisches Museums zu Berlin. Monats. Akad. Wiss. Berlin,1863:272-289.
1867. Uber Flederthiere und Amphibien. Monats. Akad. Wiss.Berlin, 1867:703-712.
1868. Uber neue oder wenige bekannte Amphibien. Monats. Akad.Wiss. Berlin, 1868:449-453.
1870. Uber Platemys tliberosa , eine neue Art von Schild-krbten aus British-Guiana. Monats. Akad. Wiss. Berlin,1870:641-653, 2 pis.
1871a. Mittheilungen uber eine von Hrn . Dr. Robert Abendroth indem Hochlande von Peru gemachte Sammlung von Amphibien,welche derselbe dem Kbnigl. zoologischen Museum geschenkthat. Monats. Akad. Wiss. Berlin, 1871:397-404.
1871b. Uber einige Arten der herpetologische Sammlung des Berlin-er zoologischen Museums. Monats. Akad. Wiss. Berlin,1871:644-652.
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1872b. Uber eine, zwei neue Gattungen enthaltende, Sammlung vonBatrachiern des Hrn. Dr. 0. Wucherer aus Bahia, so wieliber einige neue oder weniger bekannte Saurier. Monats.Akad. Wiss. Berlin, 1872:767-776.
1873. Uber eine neue Schildkrbtenart , Cinosternon effeld-tii und einige neuer oder weniger bekannte Amphibien.Monats. Akad. Wiss.
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1881. Vorkommen schildformiger Verbreiterungen der Dornfortatzebei Schlangen und uber neue oder weniger bekannt Artendieser Abtheilung der Reptilien. Sitz. Ber . Naturf. Freunde,1881:49-52.
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ADDENDA
Subsequent to printing the final draft of the manuscript, I receivedsome helpful comments from Paulo E. Vanzolini. He suggested that thefollowing Amazonian reptiles might be part of the Ecuadorian herpeto-fauna
:
TESTUDINIDAEGeochelone carbonariaSCINCIDAEMabuya bistriataM. ficta
AMPHISBAENIDAEAmphisbaena alba
COLUBRIDAEHelicops leopardinusHydrodynastes bicinctusRhinobothryvmt lentiginosumPseudoboa neuwiedii