ATOLL RESEARCH BULLETIN 
NO. 451 
CRUSTACEA DECAPODA AND STROMATOPODA OF FRENCH POLYNESIA 
BY 
JOSEPH POUPIN 
ISSUED BY 
NATIONAL MUSEUM OF NATURAL HISTORY 
SMITHSONIAN INSTITUTION 
WASHINGTON, D.C., U.S.A. 
SEPTEMBER 1998 
R V  Marara, 43 m long; original drawing by a Polynesian 
This vcsscl has operated in Frcnch Polynesia hetwceo 1973 
and 1996, and realiscd important collections of Cruslacea. 
C O N T E N T S  
SLJMMARY .............. . ............................................................................................. 1 
INTRODUCTION.  
FAMTLY I'ENAEIDA 
FAMII,Y SICYYONIII) 
FA\III .Y ATYTIIA 
.............................................................. ....... FAMILY O I X O P I I O K ~ ) A ~ ~  ... 
FAMII.Y RIIYNCIIOCINF:SII)A 
FAMILY GNIVI'I~OPIIYL.IIUA 
F A A ~ Y  ANCI{ISTIOII) 
FAMILY P A I . A I ~ ~ I O N I D  
FAMIIII HII~IY~ISTII) 
FAMILY PKOCISSII) 
FAMILY ~'ANI~AIII~AI 
FAMII-Y C K A N G  
I I \ T I : K A - O K I ~ ~  T IAI,ASSINI~I~A ................................................................................. 30
FAMIIX AXII1)A 
F~h11l.y CN.I.M: A," I 
FAMILY TIIOMASSINIII)A 
FAM.Y Gom)An-fr.ll),\r 
FAMII.Y ~S~~JI)OSQIJII.I.II> 
FnMr1.u TAK~!II)A 
FAMILY NmOSQUIIAmA 
FAWLY S Q u n  
FAMILY PROTOSQUILI.IIIA 
FAMILY ENOPLOMLTOPIDA 
FAMILY POLYCHXL~ 
FAMILY PALIMIKDEA 
FAMILY PAOURID 
FnM1r.Y I'ARAPAG 
FAMILY GALATBEIUA 
FAMILY CALNTIU 
Fhhav XANTliD 
FAMLY TRAPEZIIDAE.. ... ......................................................................... 39 
FAMILY CRYF'rOCklIRlDAE ............................................................................ 40 
DISCUSSION .......................................................................................................... 41 
................................................................. ...................... LITERATURE CITED .. 44 
...... ........................................................................... ACKNOWLEDGEMEW . . 54 
INWl ...................... .. .................................................................................... 54 
Maria 
LU 
Aeuanuraro . 
. . Nukutcpipi Tureia Anuaouruaga Vanavaoa' 
Terlnram Tmanmga 
Moruroa Vdlangi;". e 
Tcrnatangi * o Mniureivavs Mamtca Sou 
25' AUSTRAL ,, 
lhiers Hank 
Neilson Bank .:, 
- Rapa 
e 
. Marotiri 
Macdonsld Hank : 
155" W 150' 145' 
I I I I I I I I I I I I I I 
CRUSTACEA DECAPODA AND STOMATOPODA 
OF FRENCH POLYNESIA 
(DENDROBRANCHIATA, STENOPODIDEA, CARIDEA, THALASSINIDEA, AND STOMATOPODA, 
WITH ADDITIONS TO ASTACIDEA, PALINURIDEA, ANOMURA, AND BRACHYURA) 
JOSEPH POUPIN 
A B S T R A C T  
French Polynesian Dccapoda, reslrictcd to shrimps, and Stomatopoda are inventoried from literature and 
unpublished fields collcclious. Nuinbcr of species is 205 for the shrimps and 26 for the Stomatopoda. Within 
the shrimps, Caridca dominatc with 170 spccics, mainly Alphciclac (57 spccies), Palaco~onidae (46 species), 
and Pandalidae (29 species). Full literature and distribotiot~ ill French Polynesia are it~dicatcd for each species. 
When available a brief indication is also given on t l~c biotop. New records iiiclude 23 species o i  the genera 
Aristaeonzorpha, F~r~rchal ia ,  Mefapenneol~si,~, Pelogope~loerrs, Sicyonia, Nemnrocarcinns, Jo~~ ice l l a ,  
Srylodactylu.s, A~~clrisras, Ilarpiliopsis, Jocaste, Para~~cl~islrts, P ericlinienes, Philarius, Vir, and nor . ,  For the 
shrimps, atld A h a ,  B~tsquilla, and Go~rndac~yl~ts, for thc Stomatopoda. 
Modificatioi~s to the first docunic~lted checklist of thc French Polynesian lobsters, hermit crabs, end 
crabs (POUI'IN, 1996a) are also inclutled with 28 additional species or the genera Stereonwsris and Panulirus 
for the lobsters, Calcin~rs, Dardarl~rs, Corapagnrus, Nemalol~agariw, Pylopa@irol~si.s, Oncopagurus, Munida, 
and Albrr~tca for the hermit crabs, Calappa, Mio-sia, Alni~~odaeu.~,  I Icraga~ial ia ,  Tetralia, Trapezia, 
Lirhoscaps, Sphenoniain, end Uti~romiella, for the crabs. 
The total numbcr of French Polyocsian Dccapodn and Stomatopda is now 758, mainly Bracllyura (377 
species), Caridea (170 spccies), and Anoinura (126 spccies). Modcrti rcscarches alone, i.e. published after 
1980, have generated 322 new rccords. Despite these efforts, knowledge of the I-cpional krut~a remains 
incomplctc, and it is estimated t l ~ n t  he numbcr ofspecics will easily excccd 800 when rccent collections froin 
the Marquesas are fully studied (MUSORSTOM 9). A lot of genera still need a careful revision and groups 
like the Thahssinidca and Stomatopoda arc obviously under-rcpresei~ted. 
- 
Mus6um national d'Hisloirc naturelie, inboretoire de Zoologie dcs Arlhropodes, Paris. Prcscnl address : 
Laboratoirc d'OcCanograpRic. ~ c o l c  Navalc, 29240 RREST Naval, FRANCE. 
Manuscript receivcd 26 March 1998; reviscd 12 .Jmc 1998 
I N T R O D U C T I O N  
In complement to the first dwumei~ted checklist dealing with the Astacidea, Palinuridea, Anomura, and 
Brachyura (POUPIN, 1996a), the present work conside~s the shrimps (Dendrobranchiata, Stcnopodidae, Caridea, 
arid Thalassinidea) and Stomatopoda of French Polynesia. 
Deep shrimps, i.e. beyond loom, were originally listed separately by POUPIN in: "Recent 
cor~tributions to the deep sea decapod Crustacea of French Polytiesia", submitted to the Proceedings of [he 
International Senckenberg Syniposirinz, Frankfurt, October 1993. However, as these Proceedings were never 
printed, they are again included here. This deep material mainly comes from investigations of the RV Marara 
(POUPIN, 1996b; POUPIN & BENARD, 1996). 
Records from the literatutt: are complemented by unpublisl~ed field collections. These were made by the 
author, during the RV Marara campaigns, or realised during the activities of the French ORSTOM research 
centcr, Tahiti. 
Presentation follows POUI'IN (1996a) and is briefly summarized here. Records are limited to French 
Polynesia, which includes five arcliipclagos: Marquesas, Society, Austral, Tuamotu, and Gainbier Islands (see 
map). A single exception is for Merape~iaeopsis velritina, recorded by CKOSNIER (1991) from Pitcairn Island 
ill the neighbourhood of the Gamhier Islands. The unit of location is the Island, names of hays, rivers, or 
islets on the reef (Polynesian 'Motu') being ignored. It must be noted that distribution patterns of most of the 
specics are well bcyond French Polynesia but are strictly limited to French Polynesia in this report. As far as 
possible, we briefly mentioned the main characteristic of the biotop: freshwater, pelagic, bathypelagic, 
planktonic, sublittoral, deep, burrower, atid association with other organisms (coral, echinoderm, sponge...). 
This information, for exanple depth range or kind of associatioo, is restricted to observations made in French 
Polynesia. Arbitrarily, (see POUPIN, 1996a: 7) "Subliltoral" means between 10-loom, and "Decp" beyond 
IOOtn. When no dcpth range is indicated it means that the species is common in littoral shallow waters. A 
brief history of the collections is related under each Infra-Order section. 
Abbreviations are: MNH, Museum of Natural History, London; BPBM, Bernice P. BISHOP Museum, 
Ho~iolulu; CRIOBE, Centre de  Rechcrche I~isulaire et Obscrvatoire de  I'Environncmcnt Moorca; MNHN, 
M u s h n  national d'Histoire nalurclle, Paris; ORSTOM, Institut Franpis dc Rechcrche Scicntifique pour lc 
DCveloppcmcnt cn CoopQation; RMNH, Nationaal N:ttuurliistorisch Muscum, Lcidcn; RV, Rcscru-cli Vessel; 
SME, Station Marine d'Endoume; USNM, National Museum of Natural History, Washington. 
L I S T  O F  T H E  S P E C I E S  
ORDER DECAPODA 
Dcntlrobranchiata rcportcd liw'c are mostly pelagic, bathypclagic, or deep specics. Oldcst collections 
wcrc madc by the HMS Cliolleri~er (BATE, 1881, 1888). Subsequent species wcre thcn colleclcd by the 
National Geographic Sociely SMITHSONIAN-BISHOP Muscum Marqucsas Expcdition (REHDER, H.A., 1 9 6 7 ~ .  
CROSNIER, 1991). Rcccnt additions to the Dcndroblanchiata mainly come from RV Murara collcctions 
(POUPIN et a / . ,  1990; POUPIN, 1996b) or material from the ORSTOM Center, Tahiti (coll. P. FROUIN, 
E.  JOSSE, and RV Alis Marqucsas Expcdition, August/Scptc~nbcr 1997). This material lias hecn determined 
by A. CROSNIER (MNHN) and includes 7 species reported for the first time: Arisraeomorpha foliacea, 
Fur~chalio raanii~gi, F .  villosa, Metapenaeopsis cornnlensalis, M ,  hilarula, Pelagoperlaeus balboe, and 
Sicyor~ia sp. 
Figure 1 - Arismeon~orplm folincea Risso, 1827 
(Drawing by M.K. RYAN, i n  P~~RI:.LFAR~NTI; & KENSLEY, 1997) 
FAMILY ARISTAEIDAE 
Aristaeorriorpha foliacea Risso, 1827 
DISTRIBUI'ION. -Society (Maupiti) - B n i l i ~ e l a ~ i c  (7181n). 
~~ ~ 
REFI;RENCES. -Aristaeor~~orpha foliacen - NEW MATERIAL. - Coll. RV Mararo, det. A. CRosNlllR (Maupiti MNHN 
Na13433; 718111). 
Aristeus aff. mabaltissae Ramadan ,  1938 
DISTRIBUI'I~N. - Marquesas (Eiao, Hiva Oa) - Bollrypclngic (600-630m). 
REFERENCES. -Aristeus aff. r~tnbahissae - POUPIN el al., 1990: 15 (French Polynesia). - I'OUPIN. 1996b: 4, 
pi. la  (Eiao. Hiva-Oa; 600-630m). 
Plesiopenaeus arrnatus (Rate, 1881) 
DI~'SRIBUI'ION. -Tuamoiu - Barhpclr~gic (4300nr). 
REFERENCES. -Ari~teus  nrmatus - RATE. 1888: 312. pi. 45-46 (Northwest Tuamotu. Chnllenget. st. 276. 
16/09/1875, 13O28'S, 149"30'W). - I'lesiopewaer~s nrmattls - CROSNlER 81 FOREST, 1973: 294, fig. 99c-d 
( S Y ~ . ) .  
l~~- roneous ly  mcntioncd as "HARALD, 1967" in POUI'IN 1996:i 
Benthesicymus crenatus Bate, 1881 
DISTRIBU~ON. -Tuamotu - Barlrypelagic (4300rn). 
REFERENCES. - Benthesicymus crenatus Bate, 1881: 191; 1888: 329, pl. 54-55 (Northwest Tuamotu, 
Challenger st. 276, 16/09/1875, 13'28'S, 149?30'W). 
Benthesicymus investigatoris Alcock & Anderson, 1899 
DISTRIBUTION. - Marquesas (Hiva Oa); Society (Huahine); Tuamotu (Makemo) - Ualhypelagic (630-I025m). 
REFERENCES. - Benthesicymus investigatoris - POUPIN el al., 1990: 15 (French Polynesia). - POIIPIN, 
1996b: 4, pl. l b  (Hiva Oa, Huahine, Makemo; 630-1025n1). 
Benthesicyrnus strabus Burkenroad, 1936 
DISTRIBUTION. - Austral - Ballrypelagic (4346"~). 
REFERENCES. - Benthesicymus brasiliensis Bate 1888: 332, pl. 57-fig. 1 (Southeast of Marotiri, Cltnllenger 
st. 285, 14/10/1875, 32"36'S, 137'43'W, 4346m), Challenger specimens from the South Pacific are not B .  
brasiliensis Bate. 1888 but B .  srrabris aov. in BURKENROAD (1936: 29) - Benthesicymrrs strabus Burkenroad, 
1936: 29, 32, 33, 42, 46, fig. 10, 23, 27, 34, 41, 46, 49 (BATE'S material; Syn.). 
FAMILY PENAEIDAE 
Fenneroperiaeus indicus (H. Milne Edwards, 1837) 
D I S T R I B ~ O N .  -Society (Tahiti) - Aguaculnrre. 
REFERENCES. - Penaeus indicus - AQUACOP & PATROLS, 1990: 67 (Imported for aquaculture; see remark). - 
Fenneropenaeus indicus - P ~ R E Z  FARFANTE & KENSLEY, 1997: 80, fig. 41 (New combination). 
REMARK. - Several other penaeoid sllrimps had been imported for aquaculture purposes in Tahiti: Penneus monodon, 
Litopenuerrs slylirosfris and L. vannamei. 
Funchalia tuaningi Burkenroad, 1940 
DlSTRIBUnON. - Society (Huahine, Maiao, Tahiti) - Pelagic (100-370m). 
REFERENCES. - Funchalia fnaningi -NEW MATERIAL - Coll. ORSTOMrahiti. 1997, fish stomach and pclagic 
trawl. det. A. CROSNIER (Society, off Huahine, Maiao, Tahiti; MNHN Na13426-28; pelagic 100-370m). 
Funchalia villosa (Bouvier, 1905) 
DISTRIBUTION. - Society (Tahiti) - Pelagic ( 5 0 . 7 0 0 ~ ~ ) .  
REFERENCES. - Funchalia villosa - NEW MATERIAL - Coll. ORSTOMflahiti, E. JOssF., fish stomach, det. 3. 
POUPIN & A. CROSNIER (Tahiti; MNHN Na13436). 
Melicertus canaliculatfcs (Olivier, 1811) 
DISTRIBUTION. -Society (Tahiti). 
REFERENCES. - Penaeus cnnaliculntrrs - HBLIBR, 1865: 121 (Tahiti). - Penoerrs (Melicertrrs) 
canaliculntrrs - HOLTlWlS, 1980: 47 (Catalogue). - Melicertus canaliculatus - PARE% FARFANTE & KENSLEY, 
1997: 101 (New combination). 
Metapenaeopsis corumensalis Borradaile, 1898 
DISTRIBUTION. - Society (Tahiti). 
REFERENCES. - Metapenneopsis commensalis - NEW MATERIAL - Coll. ORSTOMrahiti ,  P. FROUIN, det. A. 
CRDSNIER (Tahiti; MNHN Na13430). 
Metapenaeopsis diff ici l is  Crosnier, 1991 
DISTRIBUI'ION. - Marquesas (Tahuata) - St~blirroral (M-79nz). 
REFERENCES. - Metapenaeopsis difficilis Crosnier, 1991: 255, fig. 65-67 (Tahuaia; 64.79111) 
Metapenaeopsis hilarula (de Man, 1911) 
DISTRIBUTION. - Society (Tahiti). 
REFERENCES. -Mefapenaeopsis hilarula - CROSNIER, 1991: 227, fig. 42-44, 45a-d, 46 (Tahiti). 
Metapenaeopsis marquesas Crosnier ,  1991 
D I S T R ~ I ~ I ~ N .  - Marquesas (Hiva Oa, Nuku Hiva, Tahuata, Ua Pou) - S~cblirroral(45-80m). 
REFERENCES. -Mefapenaeopsis marquesas Crosnicr, 1991: 244, fig. 57-59 (Hiva Oa, Nuku Hiva. Tahuata, Ua 
Pou; 45-8Om). 
Metapenaeopsis tarawensis Racek & Dall, 1965 
D I S S R B ~ O N .  -Society (Tahiti); Tuamotu (Marutea). 
REFERENCES. - Melapenaeopsis larawensis - CROSNIER, 1991: 241. fig. 54-56 (Marutea) - NEW MATERIAL - 
Coll. B. THOMASSIN & P. FROUIN, det. A. CROSMER (Tahiti; MNHN Na11431-32). 
Metapenaeopsis velutina (Dana, 1852) 
DISTRIB~TION. - Pitcairn (neighbourhood of Gambier Islands) - Deep (113-124nz). 
REFERENCES. -Melapenaeopsis velulina - CROSNIER, 1991: 247, fig. 60-61 (Pitcairn; 113-l24m). 
Pelagopenaeus balboe (Faron,  1893) 
DISTRIBUTION. -Society (Maiao) -Pelagic. 
REFERENCES. - Pelagopenaeus balboe - NEW MATERIAL - Coll. ORSTOM Tahiti 1997, fish stomach, det. 
A. CROSNIER (Society off Maiao; MNHN Na13425). 
Penaeus monodon Fabricius, 1798 
D l s ~ ~ r n m l o ~ .  - Society (Tahiti). 
REFERENCES. - Penaeus fahitensis Heller, 1862: 528; 1865: 121, pl. 11, fig. 2 (Tahiti). - Penaerrs 
(Penaeas) nwrodon - HOLTHUIS. 1980: 50 (Syn.). 
Hadropenaeus lucasii Bate, 1881 
~ I ~ T R ~ ~ I U I I ~ N .  - Tuamotu (Hao) -Deep (522n1). 
REFERENCES. - Hadropenaeus lrrcasii - NEW MATERIAL - Coll. RV Marara, det. A. CRosNrER (Hao; MNHN 
Na1343.5; 522111). 
Haliporus clrrvirostris Bate, 1881 
DISTRIIIIITION. -Austral - Deep (4346-4365m). 
REFERENCES. -Haliporus curvirosfris - BATE, 1888: 288, pl. 42, fig. 1 (North of Tubuai, CAallenger, st. 281, 
611011875, 2Z021'S, 150?17'W, 4365 m; Southeast off Marotiri, st. 285, 14/10/1875, 32'365, 137"41'W, 43461n). 
- CROSNIER, 1988a: 567, fig. la-b, 2a, f ,  3a, 4, 5, 6a-c, 9a, 10a-b, Ila-c (BATE'S material, st. 285). 
Hytnenopenaects halli Bruce, 1966 
DlsTRruunO~. - Tuamotu (Moruroa) -Deep (71.5~1). 
REFERENCES. -Hymenopenaeus halli - POUPIN, 1996b: 4, pl. 1c (Moruroa; MNHN Na13434; 715m). 
FAMILY SICYONIIDAE 
DISTRIBGSION. -Marquesas (Hatutu, Motu One) -Deep (416-460"~). 
REFERENCES. - Sicyonia sp. - NEW MATERIAL - Coll. RV Alis Marquesas Expedition, det. A. CROSNIER, to be 
included in a rcvision of the family (Hatutu, Motu One; 416-460m). 
FAMILY LUClFERIDAE 
Only two Luciferidae are reported with certainty from French Polynesia, Lucifer chacei Bowan and 
L. Iyi~us H. Milne Edwards. A third species, Luciferpacificus Dana, has been described from the surroundings 
6 
of Rangiroa, Tuamotu (15O20' S, 148' W, see DANA, 1852: 673: 1855, pl. 45,  fig. 2). Th is  matcrial is no  
more  cxtant and with DANA'S description alonc the species is "completely unrecognizable" (cf. HANSEN, 
1919: 49). 
Lucifer  c h n c e i  B o w m a n ,  1967 
DISTRlBU'TioN. - Society (Rora Rora, Moorea. Tahiti); Tuamotu (Moruroa, Nukutipipi, Takapoto, Tikchao) - 
Plonklonic. 
REr;i+KIiNCliS. -Lucifer chacei - ROWMAN, 1967: 266, fig. 1-4 (Bora Bora, Moorea, Tahiti, Tikebau). - RENON, 
1977: 223 (Takapoto); 1985: 391 (List of French I'olynesian zooplankton); 1989: 78 (Syn.). - MEKSCIIARI)T- 
SALVAT. 1991: 80, tab. K (Nukotipipi; larvae in the lagoo). -Lucifer sp. - Mlcm, ,  1969: 122, tab. 5 (Moruroa). 
Lucifer typits H. M i l n e  E d w a r d s ,  1837 
DISTKIIIIJTION. -Gambier (Mangareva); Society (Tahiti) - Plntiklonic. 
I<EI:EKENCES. - Lxcifer reynaadii - R/r'rB, 1888: 466 (Tahiti). - Lucifer bafei Rorradaile, 1915: 228 (BATE'S 
matcrid). -Luci fer  lyprls - NOBIIS, 1907: 352, 01. 1, fig. 1 (Mangareva). - HANSEN, 1919: 49 (Syn.) - SYNONYM 
- Lsc$er bniei Borradaile, 1915 (Lucifer reynrrudii H. Milne Edwards, 1817 is a species incerlne sedis, sce HANSEN, 
1919: 49 "...the name L. Reynnrrdii ought to be droppeX'). 
FAMILY SERGESTIDAE 
S e r g i a  bigenrnrea ( B u r k e n r o a d ,  1940) 
DrsrKll3unoN. - Society ("off Tahiti") - Bnlhypelrrgic. 
KEFI:KENCI;S. - Sergin bigemntea - 1'im.z-FARI~N~E & KENSLEY, 1997: 199 ("off Tahiti"). 
REMARK. - Worid-wide distributed sergestids sliiio~ps are limited here to this single record although undoubtebly 
better reprcscotcd in the area. Sergcsles fo-nterinkii Ratc, 1888 recorded in tile surroundings of Kaevavae (BATE, 
1888: 419, pi. 76, fig. 4. Cknllenger, 24'3, 148"W) is a species lncerlne sedis. HANSEN (1903: 67) whcn revising 
thc Sergesles from the Cl~nllenger Expedition wriles about it: "RATE has examiocd one specimco, captorcd iii the 
'I'acific Ocean. lat. 24'S, long. 148' W,' and tncasuring 5 mm. The specimen is not to be found in the Museum 
collection [MNH]. According to the figure i t  is a very young Mnsiigopns belonging either to S ,  diuponries Rate. 
H.J.H., or to a closely-allied spccics of thc same group." 
Tlic vcry cornlnoi~ Srettoplrs hisl~idirs (Olivier. 181 I)  is the single specics regularly rcportcd from the 
area. SENDLER (1923: 46) also reports an undctcrtnit~cd spccics of thc getllis Slerio[)ascrclus (juvenile from 
Makatca, Tuatnoru: det. DE MAN). Thcrcforc, in addition to the Storopirs sp. nov. listed hereafter, 1111-ce more 
S t e ~ i o p t x l i d x  have bcen recognisctl durilig the recent R V  Alis Marquesas Expedition (J. POUPIN field notes; 
~nnlcrial not studied, deposited it1 MNHN collcctioiis). 
FAMILY STENOI'ODIDAE 
S t e n o p r i s  lrispidrts (Ol iv ie r ,  1811)  
~lsTKli3rIl'lON. - Society (Tahili); Tuamoto (Hao, Makatea, Karaka, Karoin). 
KEI'ERENCES. - S l e n o p x ~  hispidm - DANA,  1852: 607; 1855. pl. 40, fig. 8 (Karaka). - N O I ~ I .  1907: 366 
(Hno). - Slih'l)l.llK, 1923: 45 (Mz~katea). - Holll'nuts. 19538: 60 (Raroiaj. - MOKRISON, 1954: 13 (Karoia). - 
SAI.VAT, 1986a: 23 (Text); 1986b: 71, pliotograph (French Polynesia). - RAGXIS & CIIKIS?.IAN, 1983: 109, 
photograph (Tualnoto). - Sfenoptis - PAKIXIN, 1992: X I ,  photograph (Tahiti; according to the photograpl,, must 
bc S. Irispidrrs). - Slertopris ? - CIlAnO1Ja L. & F., 1954: pl. 44 (Frcacl~ I'olynesia). 
Stenopus sp. nov. (GOY in study) 
DISTRIBUTION. - Tuainotc~ (Fangataufa) -Deep (190111). 
REFERENCES. - Stenopes sp, nov. - POUPIN, 1996b: 4, pl. Id (Fangataufa, 190111; in study with J. Goy). 
INFRA-ORDER CARIDEA 
Caridea have been first reported from French Polynesia by the 19th century voyages: the French 
'Corvette' La Coquille (Gu~RIN-MENEVILLE, 1838), the American US Exploring Eq~edirion (DANA, 1852, 
1855). the American Norrh Pacific Expedirion (STIMPSON, 1860), the Austrian frigate Novara (HELLEK, 
1862, 1865), and the British Challcqer (BATE, 1881, 1888). Through the activities of the J.C. GODEFFROY 
& SON shipping compagny, Polynesian shrimps were also deposited at the Museum GODEFFKOY (1860- 
1886, HAMBURG; sce historical by FRANSEN et al., 1997), and subsequently purchased by the LEIDEN 
Museum (HOLTHUIS, 1950: 175) and the STRASROURG Museum (ORTMANN, 1890,1891). 
Between 1902-1905, G. SEURAT, head of a Zoological laboratory established at Rikitca, Gambic~ 
Islands, sent several collections to Paris Museum. These were studied by NORILI (1906, 1907). A succession 
of expeditions were then conducted in the Polynesian waters, with reports of more freshwater and lagoon 
shrimps: thc Hanscatischen Siidsee-Expedition (SENDLEK, 1923): the Pacific Entomological Survey 
(ADAMSON, 1935); the Yacht Alva world cruise (BOONE, 1935); the Mangarevan Expedition of the Bernice 
P. BISHOP Museum (EDMONSON, 1935, 1944), and the Raroia Atoll Pacific Science Board Expedition 
(HOLTHUIS, 1953a; MORRISSON, 1954). 
The alpheids have mainly been studied by BANNER & BANNER (1967), from a field trip made in 1954 
in the Society Islands. Unfortunately all the material was destroyed in the Hawaii laboratory fire of 1961 
(BANNER & BANNER, 1962). As a result BANNEK & BANNEK (1967) work is based on a tlraft manuscript and 
field notes, without any drawings. It is amazing to realise that type material of Alpheus gracilipes Stimpson, 
1860, described from Tahiti, was destroyed in the Great Chicago fire of 1871 (MANNING, 1993a) and that, 90 
years later, the neotype of that species, selected by BANNER &BANNER (1967) was again destroyed in the 
Hawaii lalmntoly fire. 
In 1965, installation of a French nuclcar base at Moruroa had generated several ecological surveys with 
occasional collections of shrimps. For example, Periclimolaeas tuamorae Bruce, 1969a was collected in 1965 
in Moruroa by B. SALVAT. Later, these collcctinns we[-e pursued during the campaigns of thc RV Marara 
(coll. B. RICHER DE FORGES, J.L. CARSIN, and J. POUPIN) mainly with deep pandalid shrimps (CROSNIER, 
1986a,b, 1988b; CHAN & CROSNIER, 1991, 1997) but also with littoral component of the fauna, such as 
Ciiletorhynchus her~dersorli (Kemp, 1925) (OKUNO, 1997). Part of this material (13 species) is hereafter 
reported for the first time: Neniarocarcirrus gracilis Bate, 1888; Ja~~icella spinicauda (A. Millie Edwards, 1883), 
Slylodactylus aff. libratus Filhol, 1885, At~chisr~ts deniarii Kemp, 1922, Harpiliopsis spiriigera (Ortmann, 
1890), .locasce japoriica (Ortmann, 1890), Para~lcl~isri~s sereriei Bruce, 1983, Pet-iclime~lcs elegam (Paulson, 
18751, Periclinienes grandis (Stimpson, 1860), Periclinlenes rrlahei Bruce, 1969a. Philarius gerlachei (Nobili, 
1905), Vir sp. nov. Bruce (in preparation), and Thor anzboii~errsis (De Man, 1888). Though ORSTOM Tahiti 
center activity (coll. E. JOSSE and P. FKOUIN), four other shrimps are also added to the French Polyoesian 
fauna: Acar~rhephyra snzirhi Kemp, 1939, Oplophorus gracilirosfris A. Milnc Edwxds. 1881, Salmone~rs ? 
serraridiginrs (Coutikre, 1896), Nikoirles aff. darlae Paulson, 1875, and Processa aff. nwana Yaldwyn, 1971. 
Since 1971, several ecological studies have been initiated by  the French ~ c o l e  Pratique des  Hautes 
~ t u d c s  research center, uow dcnominatcd CRIOBE and located at Moorea, Society Islands. These studics 
mainly report coral or algae associalcd shrimps (NAIM, 1980, identifications by HOLTHUIS; ODINETZ, 1983, 
identifications by BRUCE for the Palaemonidae, and BANNER for thc Alpheidae; and KROI'I' & BIKKELAND, 
1981) atid freshwater shrimps (MARQUET, 1988, 1991, 1993, identifications by  HOLTHUIS). T h e  freshwater 
species are still curfcutly under study (see KEITH & VIGNEUX, 1997). 
FAMILY NEMATOCARCMIDAE 
Nernntocarcinrrs grac i l i s  Bate,  1888 
DISTRIB~KION, -Tuarnotu (Fangataufa. Marutca South) - Deep (840-IOOOm). 
RE1:liRENcEs. - Nenratocarcinris grncilis - NEW MATERIAL - Coll. RV Marnrn, dct. T. KOMAI (Fangataufa. Marutca 
South; 840-10001~1). 
Nernatocarcinr~s nndula t ipes  Bate, 1888 
D I S T R I R V ~ N .  - MRI~IICSRS (Fatu Hiva); Society (Raiatea) - Deep (570-850m). 
RIII:J:KENCES. - Nenlatocnrcinli~ rindelatipes - POUPIN, 1996h: 4, pl. l e  (Fatu Hiva, Raiatea; 570-8501~1). 
FAMILY ATYIDAE 
Atyoida  pilipes ( N e w p o r t ,  1847) 
DlsTRmUrIoN. - Austral (Ruruto, Tubuai); Gambicr (Mangarcva); Marquesas (Eiao, Fatu ' h a ,  Hiva Oa, Nuku Hiva, 
Ua Nuka, Ua I'ou); Society (Moorea, Tahiti) - Freshwnler. 
REFERENCES. - Afyoida fahitermis Stimpson, 1860: 28 1971 (Tahiti). - Afya brevirostris - SI:IJKAT. 1934: 
50 (Tahiti). RJCARD. 1986: 88 (French Polynesia). - Atyoida serrafa - ADAMSON. 1935: 16 (Eiao. Fatu Hiva, 
Hiva Oa, Nuku Hiva, Ua Pou). - EDMONSON, 1935: 14 (Mangarcva, Rapa, Tahiti; Maquesas) - Both not Alyoida 
serrata (Bate, 1888). - Orfrrrorrrrin allsaudi - ADAMSON, 1935: 16 (Eiao, Fatu Hiva). - Vanderbiltin 
ro sa~no~ idae  Boonc, 1935: 160. pl. 41-42 (Tahiti). - HOLTI~UIS,  1953b: 113, fig. 1 (Tahiti; redescription of 
R o o ~ ~ i ' s  material as a juvcnile stage of Alyu serruln; lint A. serrnm Rate, 1888, in CIIACE, 1983); 1980: 70 (Syn.). - 
Atyoida pilipes - CrmcE, 1983: 10, fig. 3-8 (Marquesas, Tahiti; Syn.). - MARQUET, 1988: 87, fig. 48, tab. 23; 
1991: 129, tab. 1 & 2; 1993: tab. 1 & 3 (Hiva On, Mangarcva, Moorca, Nuku Hiva, Rurutu, Tahiti, Tubuai, Ua Hoka, 
Ua Pou). -- MOSSERON, 1994: 4, fig. 321 (Nuku Hiva). - M o s s s n o ~  & Fosscri,  1994: 16, fig. Ra (Nuku I-Iiva). - 
P o u r w ,  1994: 5, fig. 1, pl. l a  (Tahiti). - K r x r ~  & V I G N ~ U X ,  1997: 22, tab. 2 (Same Islands than MARQUW) - 
SYNONYMS - Alya brevi~mlris De Man, 1892; Orunnnnin nllr~nrrdi Roovicr, 1905. 
Afyopsis spinipes ( N e w p o r t ,  1847) (doubtful)  
D i s r ~ u ~ u n o ~ .  - Gambier (Mangareva) - Freslrwnier. 
REFERENCES. -Atyo spinipes - NODILI, 1907: 353 ("Gatavakc" = Mangareva). - SeuRhr, 1934: 51 ("Gatwake" 
= Mangarcva). - Atyopsis spinipes - CIMCF.. 1983: 35, figs 20-22 (Syn.; cf. remark). 
REMARK. - CRACE (1983: 40) writes "Verification is also nceded of the Magarcva record in N o n m  (1907) and 
SEURAT (1934) in order to eliminate the possibility of confusion hetwecn Alyopsis spinipes and Aryoida pilipes, 
which seems to be established on the Gambier Islands". To confirm C l I ~ c a ' s  hesitations we rcmark that mccnt 
prospections in the Galnbier rivers (electric fishing by MARQUE?: 1988. 1991, 1993) lhave brought only Alyoida 
pilipes and never Alyopsis ,spinipes. 
Caridirra rapaensis  E d m o n s o n ,  1935 
Dr5TRlBmoN. - Austral (Rapa, Rurutu) - Preslrwnler. 
REFERENCES. - Cnridina rnpaensis Edmonson, 1935: 12 (Kapa). - ELI~REDFB, 1967: 12 (Rapa; catalog). - 
MARQUPT, 1988: 87, fig. 48, tab. 23 ; 1991: 130, tab. 1 & 2 ; 1993: tab. 1 & 3 (Rurutu). - K E n r  & V I G N ~ I X ,  1997: 
24, tab. 2 (MARQUET'S data). 
Caridirra serrat irostr is  De M a n ,  1892 
DISTKIIXI~ION. - Austral (Tubuai); Socicty (Moorea) - Pres/twnrer. 
REFERENCES. - Caridina serralirostris - MARQuEr, 1988: 87, fig. 48, tab. 23 ; 1991: 130, tab. 1 & 2 ; 1993: 
tab. 1 & 3 (Moorea. Tubuai). - K e r n  & VIGNEUX, 1997: 23, tab. 2 (Same Islands than MARQUET). 
Carid ina  weberi  De Man, 1892 
DISTRIBUTION. -Austral (Rurutu, Tubuai); Marquesas (Fatu Hiva, Hiva Oa, Nuku Hiva, Ua Huka, Ua Pou); Society 
(Moorea, Tahiti) - Freshwater. 
REFERENCES. - Caridina weberi - ADAMSON, 1935: 16 Vatu Hiva, Nuku Hiva, Ua Pou). - EDMONSON, 1935: 10 
(Tahiti, Marquesas). - MARQUET, 1988: 87, fig. 48, tab. 23: 1991: 129, tab. 1 & 2; 1993: tab. 1 & 3 (Hiva Oa, 
Moorea, Nuku Hiva. Rurutu, Tahiti, Tubuai, Ua Huka, Ua Pou). - MOSSERON, 1994: 4, fig. 3a (Nuku Hiva). - 
MOSSERON & F O S S A ~ ,  1994: 16, fig. 8b-c (Nuku Hiva). - K E m  & V I G N F . ~ ~ ,  1997: 22, tab. 2 (Same Islands than 
MARQUIT). 
FAMILY OPLOPHORIDAE 
Acan tkephyra  ex imia  Smith, 1884 
DlsrmBmoN. - Austral (Tubuai): Tuamotu (Fanrataufa. Hao. Moruroa) - Bnrhvuelanic (720-I050ml 
. . . " 2. " . 
REFERENCES. - Acatrt/rephyra eximia - POUPIN et al., 1990: 15 (French Polynesia). - POUPIN, 1996b: 4, pl. l e  
(Fangataufa, Hao, Moruroa; 720-1050m) - NEW MTERIAL - Coll. Y. PLESSIS, 14/05/79, det. A. CROSNIER (Paris, 
MNHN Na7259; Tubuai; 800m). 
Acanthephyra  srnithi Kenlp, 1939 
DISTR~BUTION. -French Polynesia - Barltypelagic (Up lo 3450m). 
RLFERENCES. - Acanthephyra snrilhi - NEW MATERIAL - Coil. E. JOSE 1995, fish stomach, det. J. POUPIN, 
material deposited at ORSTOMRabiti (French Polynesia, probably between Society and Marquesas). 
Acan thephyra  stylorostrat is  (Bate, 1888) 
DlsrRIBlJTIoN. - Austral - Barhypelagic (4346m). 
REFERENCES. - Re~lheocaris  exuens Bate, 1888: 724, pl. 123-fig. 3 (Southeast off Marotiri, Challenger, 
st. 285, 14/10/1875, 32"36'S, 137"43'W, 4346111). - Acanthephyra slylorostrata - CHACE, 1940: 144, 
fig. 22 (Syn.; see remark). 
R E M A R K .  - CEIACE writes "It is very probable that BATE'S Benlheocnris exuens is the same species [than 
A .  srylorosfratis] but, inasmuch as CALMAN states that the type is in such condition that comparison is almost 
impossible and as BATE'S figure is inadequate its seems best to retain the much more suitable name, srylorosfruta, for 
this form." 
Heterogenys  microphthairna (Smith, 1885) 
DI~TRIBLI~ON. -Auseal - Bntlzyl,elagic (4346m). 
REFERENCES. - Acatrthephyra longidens Bate, 1888: 735, pl. 124-fig. 4 (Southeast off Marotiri, Cllalienger, 
st. 285, 14/10/1875, 32'363, 137'43'W, 4346111). - Heteragenys nricrophthalnra - CHACE, 1986: 38. 
fig. 20 (Syn.). 
Janicel la spinicnuda ( A .  Milne Edwards, 1883) 
DiSTRIBUnON. - Probably Tuamotu (Moruroa) - Barlrypelagic (SOOm?). 
REFERENCES. - Janicella spinicauda - NEW MATERlAL - Colt. Y. PLESSIS & B. RICHER DE FORGES,  det. 
A. CROSNIER (Paris. MNHN Na7255, no location but probably Moruroa according to number given by the 
collector). 
Oplophorus  graci l irostr is  A.  Milne Edwards, 1881 
DISTRIBUTION. - Society (Maiao); Tuamotu (Mataiva) - Barlzypelagic (100-370n1). 
REFERENCES. - Oplophorus gracilirostris - NEW MATERIAL - Coll. ORSTOM 1997, fish stomach and pelagic 
trawl, det. A. CKOSNIER (Off Mataiva and Maiao; MNHN Na13429; bathypelagic 100-370111). 
Oplophorus  sp inosus  (Brullk, 1839) 
D1sT~lBvrIoN. - Tuaniom (Fangataufa) - Bnrlrypclagic (30-loom). 
R E I ~ I I N C E S .  -Oplophorus spirzosus - POUPIN, 1996b: 4, pl. lg (Fangataufa; in plankton nct 30-100m). 
Oplophorus  typus Milne Edwards, 1837 
D I S T R I B ~ O N .  - Marquesas (Nuku Hiva. Tahuata) - Barl~ypelagic (541-580n1). 
REFERENCES. - Oplophorus typus - POUPIN er (11.. 1990: 15 (French Polynesia). - POUPIN, 1996b: 4, pi. Ih 
(Noku Hiva, Tahuata; 541-580m). 
Systellaspis debil is  (A. Milne Edwards, 1881) 
DISTRIBUTION. -Tuamotu (Fangataufa) - Bathypelagic (50-100n1). 
REFERENCES. - Systellaspis debilis - POUPIN, 1996b: 6 ,  pl. 2a (Fangataufa; in plankton net 50-100111). 
Systellaspis pel lucida  Filhol, 1885 
DISTRIBUTION. -Tuatnotu (Hao) - Bull~ypelagic (1020m). 
REFERENCES. -Systellaspis pellucidu - POUPIN, 1996b: 6, pl. 2h (Hao; 1020m). 
FAMILY STYKIDA~YLIDAE 
S ty lodac ty lus  aff. l ibra t l t s  Chace, 1983 
DISTRIBUTION. -Tuamotu (Moruroa) -Deep (512n1). 
REFERENCES. - Stylodactylus aff. libratus - NEW MATERIAL - Call. J. POUPIN, det. R. CLEVA (MNHN Na 13400; 
Moruroa; 5121n). 
FAMILY KHYNCHOCINETIDAE 
Cinetorhynchlts  heridersorti (Kemp, 1925) 
DrssRmunoN. -Austral (Rapa). 
REFERENCES. -Cinetorhynchus herrdersoni - OKUNO, 1997: 46. f ~ g .  9. 12d-f, pl. If (Comh. nov.; Kapa). 
Cinetorhyricltus h ia t t i  (Holthuis & Hayashi, 1967) 
DISTRIBUTION. -Marquesas (Nuku Hiva); Tuamotu (Raroia). 
KEI:ERENCES. - Rhynchocinctes hiatti n. sp. - HoLTfiUls. 1953a: 54 (Raroia; nonlina nuda). - MORRISON, 
1954: 18 (Same material). - Rhynchocinetes hiatti Holthuis & Hayashi, 1967: 162, fig. 1-2 (Raroia; 
holotype from Formosa). - FRANSEN el al., 1997: 14 (Paratypcs RMNH D8939; Raroia). - Cirretorhynchus 
hiatti - OKONO, 1997: 40, fig. 5, pl. l c  (Comb. nov.; Nuku Hiva). 
Cinetorhynchus  re t icula tus  Okuno, 1997 
DISTRIBUTION. -Marquesas (Nuku Hiva). 
REFERENCES. -Cinetorhynchus reticulatus Okuno, 1997: 49, fig. 10.11, 12a-c, pl. Ig, h (Nuku Hiva) 
FAMILY GNATHOI'HYLLIDAE 
Gnatkophyl lum arnericanurn Guerin, 1856 
DISTRIBuTlON. - Gambier (Mangareva); Society (Tahiti); Tuamotu (Raroia) - Free-living or coral associale (Gen. 
Pocillopora). 
REFERENCES. - Gnuthophyllunz pallidunt Ortmann, 1890: 537 (Tahiti). - Gnathophyllum triderrs 
Nobili, 1906: 259; 1907: 365, pl. 1, fig. 4 ("Rikitea" = Mangareva). - Gnathophyllrrn~ arnericnnunz - BOONE, 
1935: 197, pl. 54 (Tahiti; Syn.). - HOLTHUIS, 1949: 210 (NOBILI'S material; Syn.); 1953a: 60 (Raroia). - O ~ N E T Z ,  
19R3: 207 (Tahiti; in coral PociI/oj~orn, nor1 obligate associate). 
FAMILY ANCHJSTIOWWAE 
Anchis t io ides  cornpressus P a u l s o n ,  1875 
DISTRIBUTION. - Gambier (Mangareva) - ? Sponge associale. 
REFERENCES. - Antphipalaemon seurafi Nobili, 1906: 259; 1907: 364, pl. 1. fig. 3 ("Tearia" = Mangareva; 
22m). - SEURAT, 1934: 51, 60 ("Gatavake" = Mangareva; in Freshwater. see remark). - Anchistioides 
compressus - BRUCE, 1967: 570 (NonrLl's material). 
REMARK. -Anchisloides species are common in a variety of sponge and, on occasion, in tunicate; freshwater record 
of SEURAT is certainly erroneous (A.J. BRUCE, personal communication). 
FAMILY HYMENOCERIDAE 
Hyrnenocera picta D a n a ,  1852 
DlsrRmunON. - Tuamotu (Raraka) - ?Echinoderm ussociare. 
RE1;ERENCES. - Hymenocern picfn Dana, 1852: 593; 1855, pl. 39, fig. 3a-c (Raraka; "Coral reef" without 
mention of echinoderm host).- BRUCE, 1982: 209 (Possibly an echinoderm associate). - C m c e  & BRUCE, 1993: 
137 (Syn.). 
FAMILY PALAEMONiDAE 
SUB-FAMILY PALAEMOMNAE 
Bracitycarpus biunguici i latus ( L u c a s  1846) 
DISTNBUTION. - Tuamotu (Raroia). 
REFERENCES. - Rrachycarpus biunguic~ilatus - HOI.THUIS, 1953.1: 55 (Raroia). - MORRISON. 1954: 18 
(Raroia). 
Macrobrachii irr~ aernulum (Nobil i ,  1906) 
DISTRIBUTION. - Awral  (RUIU~I.  Tubuai); Ga~nbier (Mangareva); Society (Moorea, Tahiti) - Frcslrwnler. 
REFERENCES. - Palaemon (Parapalaemon) aem~l lrs  Nobili, 1906: 258; 1907: 362, pl. 1, fig. 5 ("Gatavake" 
= Mangareva; elevation 180m). - SEIJRAT, 1934: 51  ("Gatavake" = Mangarcva). - Macrobrachiunt aemulrtnt - 
HOLTHUIS, 1953a: 54 (Tahiti). - MARQUET, 1988: 90, fig. 48, tab. 23; 1991: 135, tab. 1 & 2; 1993: tab. 1 & 3 
(Mangareva, Moorea, Rurutu, Tahiti, Tubuai). - KErnl& VIGNEUX, 1997: 22, tab. 2 (Same Islands than M A R Q ~ T ) .  
Macrobrachiurn aiistrale ( G u k r i n - M h e v i l l e ,  1838) 
DlsTRlBUTlON. - Austral (Ruruto, Tubuai); Gambier (Mangareva); Marquesas (Fatu Hiva, Hiva Oa. Nuku Hiva, Ua 
Huka, Ua Pou); Society (Moorea. Tahiti) - Freslrnmler. 
REFERENCES. - Palaemon ausfrolis Gu6rin-M6ncviIle, 1838: 37 (Tahiti). - Palaemon dispar - ADAMSON, 
1935: 17 (Fatu Hiva, Hiva Oa, Ua I'ou). - IJalaenton (Eupalaenron) dispar - NOBILI, 1907: 361 (Tahiti). - 
SEURAT, 1934: 50  (Tahiti). - Macrobrachium arrstrale - HOLTHUIS, 1950: 124, fig. 27-30 (Tahiti); 1953a: 55 
(Tahiti); 1980: 87 (Syn.). - GRAND. 1986: 84, fig. 2-3 (French Polynesia). - RICARD, 1986: 88 (French 
Polynesia). - MARQUES, 1988: 87, fig. 48, tab. 23; 1991: 133, tab. 1 & 2; 1993: tab. 1 & 3 (Hiva Oa, Mangareva, 
Moorea, Nuku Hiva, Kurutu. Tahiti, Tubuai, Ua Huka, Ua I'ou). - DANIGO, 1991: 5, fig. 8, pl. l a  (Nuku Hiva). - 
FOUILLAND, 1993: 10, fig. 6 (Nuku Hiva). - KEsslr & VlaNeux, 1997: 22, tab. 2 (Same Islands than MARQUET) - 
SYNONYM - Palaenlon dispar von Martens, 1868. 
Macrobrachiurn lar  ( F a b r i c i u s ,  1798) 
DISTRIBUTION. - Austral (Rurutu, Tubuai): Gambier (Mangareva); Marqnesas (Eiao, Fatu Hiva, Hiva Oa. Nuku Hiva. 
Tabuata, Ua Huka, Ua Pou); Society (Moorea, Tahiti) - Freshwafer. 
REFERENCES. - Palaemo~r specfabilis Heller, 1862: 527; 1865: 113, pl. 10, fig. 8 (Tahiti). - Palaemon 
vagus Heller, 1865: 113 (Tahiti). - Hithynis lor - BATE, 1888: 789, pl. 124-fig. I (Tahiti). - Palaemon lar 
- ADAMSON. 1935: 17 (Eaio, Fatu Hiva, H i m  Oa, Nuku Hiva, Tahuata, Ua Huka, Ua Pou). - Palaenton 
(Eupalaemon) lar - DE MAN, 1904: 291, pl. 18, fig. 1 (Tahiti). - NOBILI, 1907: 361 (Tahiti). - SEURAT, 1934: 
50 (Tahiti). - BOONE, 1935: 150, pl. 38 (Tahiti). - Macrobrachium lar - HOVI'HUIS, 1950: 176, fig. 37; 1980: 
96 (Syn.). -GKANIX 1986: 84, fig. 1 (French Polynesia). - RICAKD, 1986: 88, photograph (French Polynesia). - 
1 L 
MARQUET, 1988: 87. fig. 48, tab. 23; 1991: 134, tab. 1 & 2; 1993: tab. 1 & 3 (Hiva Oa, Mangareva, Moorea, Nuku 
Hiva, Rurutu, Tahiti, Tubuni, Ua Huka, Ua Pou). -DhNIGO, 1991: 5, fig. 8, pl. l b  (Nuku Hiva). - FouII.l.ANIir,, 1993: 
10, fig. 6 (Nuku Hiva). - CrtAci: & BRUCE: 1993: 20 (Key). - KEIT11 & VIGNEUX, 1997: 22, tab. 2 ( S a m  Islands 
than M A R Q ~ T ) .  
M a c r o b r a c i t i u r n  latirrtanrts (von Martens, 1868) 
DISTRIBUTION. -Austral (Rmutu, Tubuai); Gambier (Mangareva); Marquesas (Fatu Hiva, Hiva Oa, Motanc. Nuku 
Hiva. Ua Huka, Ua Pou); Society (Moorca, Tahiti) - Freslnerrrer. 
R1~l:ERENcEs. - I'alaemon latinlanns - ORTMANN, 1890: 737, pl. 47, fig. 11, l l z  (Tahiti). - ADAMSON, 1935: 
17 (Fatu Hiva, Hiva Oa. "Mohotani" = Motane, Ua Pou). - Macrobranchiun~ lofirnanus - HOLTHUIS, 1980: 97 
(Syn). - GRAND, 1986: 84-85, fig. 4 (French Polynesia). - KICARD. 1986: 88 (Frcnch Polynesia). - MARQUET, 
1988: 90, fig. 48, tab. 23; 1991: 134, tab. 1 & 2; 1993: tab. 1 & 3 (Hiva Oa, Mangareva, Moorea, Nuku Hiva, 
Kurulu, Tahiti, Tubuai, UR Huka, Ua Pou). - DANIGO, 1991: 5, fig. 6, 8, pl. 1c (Nuku Hiva). - C w c e  & BRUCE 
1991: 20 (Key). - FOUlI.l.AND, 1993: 10, fig. 6 (Nuku Hiva). - K m s  & V I G N I : ~ ~ ,  1997: 22, tab. 2 (Same Islands 
than MARQUET) - RELEVANT MATERIAI, - Macrobrnchiunz sp. nov. - MNHN Na 1251, Na 1379, coll. PBre Simon 
DELMAS, 4/7/1926, det. J. SHORT; MNHN Na 14404, coll. M.  de TRevlERE 1937, del. J. SHORT (Moorca) - 
Macrobrackium sp. 1 & 2 - KEITH & VIGNEUX, 1997: 11, 22, tab. 2 (Macrobracliirrn~ sp. 1, Hiva Oa, Moorea, 
Nuku Hiva. Tahiti. Ua Huka, Ua Pou; Macrobrachirmi sp. 2. Moorea, Tahiti). 
REMARK. - In Paris collections, J. SHORT has recognised a new species, related to Mao-ohrachi~m~ Iatin~anus. This 
species, denominated Onann in Polynesian language, is currently under study. 
Arbitrarily we place here the Macrobrachi~yunl sp. 1 and sp. 2. mentioned by KElnI & VIGNEUX (1997). According to 
tl~ese authors (personal communication), HOLTHUIS think that they could belong to two ncw species. SHORT study 
should bring some clarification on that point. 
M a c r o b r a c h i r t m  lepidactylotdes (De Man, 1892) 
DISTRIUCTON. - Marquesas (Hiva Oa, Nuku Hiva, Ua Huka. Ua Pou); Society (Moorca) - Freshwarer, 
KE~:EKI~NCES. -Macrobrachirrm sp. - MAKQUET, 1988: 90, fig. 48, tab. 23; 1991: 135, tab. 1 & 2; 1993: tab. 1 
& 3 (Hiva Oa, Moorca, Nuku Hiva, Ua Huka, Ua Pou). - DANIGO. 1991: 5, fig. 8 (Nuku Hiva). - Macrobracltinnr 
lepidactyloiiies - CIIACE & BRUCE: 1993: 20 (Key). - SIIORT. in prcpartion (Syn., pcrsnnal communication). 
M a c r o b r a c h i u r r t  rose i tbe rg i i  (de Man, 1879) 
DlsTRlllrmoN. - Socicty (Tahiti) - Freskwalm., aqryuacrdiere. 
KlililiRENCES. - Macrobrachiun~ rosenbergii - GRAND. 1986: 85, photograp11 (Frcnch I'olynesia). -- I'ARDON, 
1992: 116, pllotograpll (Tahiti). - Ctihcli & BKIJCE: 1993: 20, 36, fig. 15 (Syn. & key). - AQUACOP & I'A'CROIS, 
1990: 67 (Aquacolturc). 
Palaerrtort cor tc inr tus  Dana, 1852 
DlsTRlBtiSION. - Austral (Rurutu, Tubuai); Society (Moorea, Tahiti); Tuamotu (Makatea) - Brnckish water. 
KEI:ERENCI:S. - L e a ~ d e r  cortcinnus - SF.NDLF.R, 1923: 46 (Makatea). - Palnentorr corrcinrzus - Hor : r~~uls ,  
1950: 61, fig. 12 (Tahiti); 1953a: 54 (Tahiti); 1980 (Syn.). - MARQUET, 1988: 90, fig. 48, tab. 23; 1991: 136, 
tab. 1 & 2; 1993: tab. 1 & 3 (Moorea. Rurutu. Tahiti. Tubuai; "estuaries and shallow stagnant waters"). - C l m w  & 
RKIJCE: 1993: 40 (Syn. & kcy). - KEITII & VIaNEux, 1997: 22, tab. 2 (Same Islands than MARQUE?'). 
Palaearon deb i l i s  Dana, 1852 
DlsSRlnUTroN. - Austral (Tubuai); Gaolbicr (Mangarcva); Tuamoto (Hao, Rangiroa. Raroia) - Brncki.sli ~:uler .  
RE~ERIINCBS. -Learrder debilis - Nonlr.1, 1907: 363 (Hao, "Kikitea" = Mangareva). - Periclirrzenes Leorrdcr 
debi l is  - SEURAT,  1934: 60 (Hao, "Rikitea" = Mar~garcva). - Pnlnernon debilis - Hot;riiu~S, 1953a: 54 
(Karoia). - MORRISON. 1954: 5 (Karoia). - MAKQUET, 1988: 90, fig. 48, tab. 23; 1991: 136, tab. 1 & 2; 1993: 
tab. 1 & 3 (Maagarcva, Rangiroa, Tubuai; "estuaries and shallow stagnant waters"). - CNACE & RKIJCE: 1993: 40 
(Syn. & key). - K m r  & VlGNl;Ux, 1997: 22. tab. 2 (Same Islands than M ~ K Q ~ T ) .  
SUBFAMILY PDNTONIMAE 
A n c l r i o p o n t o n i a  h r t r i i  (Holthuis, 1981) 
DISTRIIIIJTION. -Tuamotu (Raroia) - Bioahv nssociote (Gen. Spondylus). 
Kl3'1inENcEs. - I'onfonia hrrrii 11. sp. - HOI.TIIUIS, 19%: 57 (Karoia; rromina nirdn). - MORRISON, 1954: 6 (San~c 
material; c o n ~ r n e ~ ~ s a l  in the spiny oyslcr Spondylrrs vnriss). - Pontorria huri i  Holtl~ois, 1981: 796, fig. 4 
(Paratype from Raroia). - Anchiupontonin huri i  - B R U C ~ ~ ,  1992a: 1276, fig. 1-4 (Comb. oov.). - Fransen e i  
a/., 1997: 15 (Paratypes RMNH D9248; Raroia). 
Figure 2 - Anchioponronia ittrrii (Holthuis, 1981) 
(From Buuce, 1992a) 
A n c h i s t u s  derr tani  Kemp, 1922 
DISTRIBUTION. - Tuatuotu (Moruroa) - Ohmlve associare (Gen. Tridacna). 
KEFEReNcEs. -Anchistus denratti - NEW MATERIAL - Coll. J. 1'OUPIN April-May 1997, det. A.J. BRUCE (Moruroa 
MNHN Na13416; in bivalve Tridacna maxima). 
A n c h i s t u s  rrtiersi (1)e Man, 1888) 
D1srRlnrrsroN. - Gambier (Mangareva) - Bivalve associde (Gen. Pinctada). 
REPEKENCES. -Anchi.stus n t i e r ~ i  - No1111.1, 1907: 359 ("Mangarcva-Tearia" = Mangareva). - SEUKAT, 1934: 60 
(Gambier; in pcarl oystcr Pinclada r~zargaritifern). - CIiAcri & BRUCE, 1993: 71, 72 (Syn. & key). - FKANSEN, 
1994: 89, pl. Id (I'hotograpb). 
C o n c l t o d y t e s  r r te ieagrir tae  Peters, 1852 
DIsrRlulmoN. - Gambicr (Mangareva); Tuamotu (Amaoo, Raioia) - Bivalve associare (Gcn. Pinctada). 
RI~FEKENCES. -Conchodytes nreleagrinae - Norr11.1. 1907: 359 (Mangareva). - Sr:rrnhr. 1934: 60 (Gambier. 
Tuamotu; in pcarl oystcr Pinctada nmrgarit+ra). - HoI.Tl~uis, 1 9 5 3 :  59 (Amanu. Raroia). - BRUCE. 1989a: 1R4 
(Key). - CHACE & BKUCII, 1993: 74 (Syn.). - FRASSEN, 1994: 96, fig. 21, pl. l e  (Photograph). 
Conc t tody tes  ? t r i d a c n a e  Peters, 1852 
DTSSRIBC~ON.  - Turmoru (Fangalaufa) - O i d v e  msociarr (Gm. I'inctada). 
Ra?auENcr.s. - Conchodyles tridncrrae - BRUCE. 1989a: 184 (Kcy) - NEW MATLRIA~. - Coll. J. I'O~JPIN April-May 
1997, det. A.J. BRUCE, with a ?  (Fangataula MNHN Na13415; in pcarl oyster Pinctodn margnririfo-a). 
C o r a l l i o c a r i s  g r a r n i n e a  (Dana, 1852) 
DrsrRlnlJlloN. - Socicty (Mooren, Tahiti); Tuamotu (Takapoto) - Coral associare (Gen. Pocillopora). 
K E I ~ ? E N C I : S .  - C o r a l l i n c a r i ~  grarninea - OI~INI:T%, 1983: 205 (Moorca, Tahiti, Takapoto; in coral 
PocilIo/)oro). - ODINKIL-COI.I.AK.T & KlC1li.K I)E FORGI~S, 1985: 201 (Same tnialeriai). - CIlAcE & BRIJCE, 1993: 77 
(Syn. & key). - BKIJSI~, in preparalion (Host gcncra Acro/mrrr, rarely Pocilloporn. Seriaioporu, Slylophora). 
Coralliocaris n u d i r o s t r i s  (Heller, 1861) 
~isrRInLir.loN. - Socicty (Tahiti) - Coral associate ((;en. Acropora). 
Rr.?Eni:Ncr:s. - Coralliocaris tnhiloei Boonc, 1935: 180, pl. 49 (Tahiti without mention of association). - 
Cora l l iocar i .  t tudirorlris - C i r ~ c E  & BRUCL'., 1993: 47 (Syn.). - BRUCE, in picparation (Host genus 
A c r o p r a ) .  
Coralliocaris superba (Dana,. 1852) 
DISTR~m'ION. -Society (Raiatea. Tahiti); Tuamotu (Fangataura, Moruroa) - C o r d  associae (Gen. Acropora). 
RS.FEUaNCES. - Coralliocaris superba - STIMPSON, 1860: 40 [I071 (Tahiti; "in coralliis"). - BOONE, 1935: 
170, pl. 46 (Raiatea, Tahiti; "in coral"). - CHACE & BRUCE, 1993: 76, 77 (Syn. & key) - NEW MATERIAL - Coll. 
J. POUPIN AprilIMay 1996 & April-May, 1997, det. A.J. BRUCE (Fangataufa MNHN Na13409, Moruroa; in coral 
Acropora). 
Coralliocaris viridis Bruce, 1974 
DISTK~I~UTION. - Society (Tahiti); Tuamoti~ (Fangataufa) - Free-livbzg or coral nssociate (Gen. Acropora, Pocillopora). 
REFERENCES. - Cornlliocaris viridis - ODlNWZ. 1983: 207 (Tahiti; in coral Poc i l loporn ,  non obligate 
associate). - NEW MATERIAL. -Coil. 1. I'OUPIN April-May 1997, det. A.J. BRUCE (Fangataufa MNHN Na13412; in 
coral Acropora). 
REMARK. - In  OUINETZ (1983) this species is reported as "Gnarhocaris ? viridis Bruce", but it is coricctly recorded 
under Coralliocaris siridis in Paris MNHN (Na6625). 
Exoclimenella denticulata (Nobili, 1906) 
DlSCR~UI'tON. - Gambier (Mangareva); Tuamotu (Raroia) - Free-living or  coral associare (Gen. Acropora). 
R E r ; e n e ~ c e s .  - Periclimenes pefithouarsi var. denficulnta  Nobili. 1906: 257; 1907: 358 ("Gatavake" = 
Mangareva). - Polaenrorrelln denticulafa - HOLTI1UIS, t953a: 55 (Raroia; in coral Acropora). - MORRISON. 
1954: 5 (Raroia). - Periclinrenea denticulotus - BR!JCEi1992b: 59. fig. 12-14 (Syn.; "32~1, trap, on sand 
wit11 Hal imeda") .  - Ilxoclinrenelln denticnlnta - DORIS & BRIJCE. 1995: 637, fig. 9-11 ("Gatavake" = 
Mangareva, Raroia; new combination and this remark "free-living browser, and only incidentaly associated wit11 
coral"). - BRUCE, in preparation (Host genus: Acropora). 
Fennera ckncei Holthuis, 1951 
DISTRIUUI'ION. -Soclety (Moorea); Tuamoto (Takapolo) - Coral associare (Gen. Pocilloporn). 
R E ~ ; E R E N C E ~ .  - F e n n e r a  chacei - KKOPP & BIRKELAND, 1981: 629, tab. 5 (Moorea, Takapoto; in coral 
Pocillopora). - B w c a ,  in prcparation (Host gcnera: Pocilloporn, Porires. Stylophora). 
Harpiliopsis beaupresii (Audouin, 1825) 
DISTRIBUTION. - Society (Moorea, Tahiti); Tuamotu (Fangataufa, Moruroa, Takapoto) - Coral associate (Gen. 
Pocillopora). 
REFERENCES. - Hnrpiliopsis benapressi - KROPI' & BIKKELAND, 1981: 629, tab. 5 (Moorea, Takapoto; in coral 
Pocillopora). - ODINETZ, 1983: 205 (Moorea, Tahiti, Takapoto; in coral Pocilloporn). - ODINETZ-COILART & 
RICHER DE FORGES,  1985: 201 (Samc material). - ClIhCE & BRUCE. 1993: 82 (Syn. & key). - BRUCE, in 
preparatioll (Host gcncra: Pocil/opora. Seriaropliom, Stylopl~orn, rarely Acroyora) - NEW MA'SEUIAI. - Coll. 1. PoUPlN 
AprilIMay 1996 and April-May 1997 det. A.J. BRUCE (Fangatauia MNHN Na13402-13413, Moruroa MNHN 
Na13403; in coral Pocilloporn). 
Harpiliopsis depressa (Stimpson, 1860) 
DISTRIBUTION. -Marqucsas (Fatu Hiva, Hiva Oa); Society (Moorea, Tahiti); Tuamoto (Moruroa. Takapoto) - Coral  
ossocinte (Gew. I'ocillopora). 
REFERENCES. - Hnrpiliopsis depressus - KROPP & BIRKELAND. 1981: 629, tab. 5 (Moorea, Takapoto; in coral 
Poci l lopora) .  - ODINETZ, 1983: 205 (Moorea, Tahiti, Takapolo; in coral Poci l lopora) .  - Hnrpi l iops i s  
depressa - CIIACE & BRUCE. 1993: 82 (Syn. & key). - BRUCE, in preparation (Host genera: Poci l loporn,  
Serinroporn, St.ylophom, rarely Acropora, Porires) - NEW MATERIAL - Call. J. POUPIN, February 1996 & April-May, 
1997, det. A.J. BRIJCI: (Fato Hiva. Hiva Oa, Moruroa MNHN Na13406; in coral Pocilloporn). 
Hnrpiliapsis spiriigera (Ortmanri, 1890) 
~ISTR~I~UTION. - Marquesas (Hiva Oa); Tuamotu (Fangataufa, Moruroa) - Coral nssocinre (Gen. l'ocillopora). 
RtiPERENcES. - Horpiliopsis spinigera - CI<ACE & BRUCE, 1993: 82 (Syn. & key). - FKANSEN, 1994: 108, fig. 
60, 62, pl. 2b (Syn.). - BRUCE, in preparation (Host genera: Pocilloporn, Stylophorrr) - NEW MATEKIAI. - Coll. 
1. POUPIN October 1995, February 1996. and April-May, 1997. dct. A.J. BRIJCE (Fangataufa MNHN Na13401, Hiva 
Oa, Moruroa; in coral Pocillopora). 
Jocaste jnponicn (Ortmann, 1890) 
DtsTKlnlrrION. - Tuamotu (Fangataufa) - C o r d  associofe (Gen. Acropora). 
15 
REFERENCES. - Jocaste japotrica - BRUCE, 19G9b: 300, fig. 1 (Host genera: Acropora, rarely Pocilloporn); in 
preparatioll (Host genus: Ao'oporn). - NEW MATERIAL - Coll. J. POIJNN April-May 1997, det. A.J. BRUCE 
(Fangataufa MNHN Na13411; in coral Acropora). 
Jocas te  laci~ia (Nobili ,  1901)  
DISTRIBLTION. - Society (Tahiti); Tuatnotu (Fangataufa, Moruroa) - Coral associale (Gen. Acropora). 
R ~ F E R R N C E S .  - Coral l iocar is  lamellirosfris - BOONF.. 1935: 174, pl. 47 (Tahiti). - Jocaste lrrcina - 
CI-IACE & BRUCE, 1993: 49, 8 4  (Syn. & key; BOONE'S reference wit11 a ?). - BRUCE, in preparation (Host gencra: 
Ao.opora, rarely Pocillopora, Styloplrora) - NEW MATERIAL - Coll. J. POUPIN April-May 1996 & April-May 1997, det. 
A.J. B n u c a  (Fangataufa MNHN Na13410, Moruroa) - SYNONYM - Coralliocaris ianrellirostris Stimpson, 
1860.  
O f i y c o c a r i s  q u a d r a t o p h t h a l r n a  (Balss,  1921)  
D I S T R I I ~ ~ I ~ N .  - Society (Tahiti) - Free-living or coral associare (Gen. I'ocillopora). 
Ker:maNcss.  - Onycocaris qrradratophthalnza - CAsrRo, 1971: 396 (Host Toxadocia violuren, Porilcra, 
Demospongia). - OOINETZ 1983: 207 (Tahiti; in coral Pocilloportz. non obligate associate). 
P a l a e m o n e l l a  lata K e m p ,  1 9 2 2  
DISTRIBUTION. -Society (Tahiti) - Free-living or coral associate (Gen. Pocillopora). 
R a r a n e N c e s .  - I 'alnen~onella lala - BRUCE, 1970: 274 (Key). - O D I N I ? ~ ,  1983: 207 (Tahiti; in coral 
Pocillopora, non obligatc associate). -NEW MATERUE. - Coll. P. FROUIN 109311994, dct. A.J. BRUCE (FROIIIN st. 
n01030, Tahiti). 
P a l a e f n o t i e l l a  rotrcrnana ( B o r r a d a i l e ,  1898)  
Drsr~lurrrrON. -Society (Moorea) - Free-living, coral,  or seo-weed nssociale. 
Kr:FERl?NceS. - 1'alaenronella rofuntana - Bnucs. 1970: 274 (Key); 1976b: table 8 ("Society & Marqucsas"; 
BRUCE perso~ial ~ o n u i i ~ n i c a t i ~ t i :  locatio~i must be a table entry error). - NAIM. 1980, aiincx 1-2 (Moorca; in 
filamentous sea-weed Gffordia milclrellae). - CHACE & BRUCE, 1993: 87, 89 (Syn. & key). - FRANSIIN, 1994: 117 
(Syn.; "Free-living and occuring among corals"). 
P a l a e t n o n e l l a  te~iuipes D a n a ,  1852 
D1STRlBUTloN. - Society (Moorea, Tahiti); Tuamotu (Takapoto) - Free-living or coral associate (Gen. I'ocillopora). 
REFERENCES. -Palaenronella tenuipes -BRUCE. 1970: 274 (Kcy). - NAIM, 1980, annex I ,  tab. 3 (Moorca). - 
ODINETZ. 1983: 207 (Moorea, Tahiti, Takapoto; in coral Poc;!lopor.a, non obligate associate). - OIXNE~Z-COI.I.ART 
& RICIIER I>E FORGILT, 1985: 201 (Samc material). - C m c e  & BRUCE: 1993: 87, 89 (Syn. & key). 
P a r a n e h i s t r c s  seret te i  Brnce ,  1983 
D t s r ~ r o m m o ~ .  -Tualiiotu (Fangataufa) - 8ivulve associate (Gen. Pinctada). 
REFEReNCss. - Paranchis tus  serenei - NEW MATERIAL - Coll. J .  I'OIIPIN, February 1996, dcl. A.J. BRUCF. 
(Fangataufa, in pearl oyster Pinctnda nrargarilifera, 10m; MNI-IN Na13352) - KtLevAwr NEW M A T ~ I A I .  - ? 
Parnrrchistrrs sp. - Coll. J. POUPIN April-May 1997, dct. A.J. BRUCE (Fangataufa; in pearl oyster P inc tado  
margaritifern). 
P e r i c l i t n e n a e r t s  cf. triderltatrts (Miers ,  1884) 
Drs ' s~~sunoN.  -Society (Moorea) - Seu-weed associate (Gen. Halinieda). 
KEFaRENC1:s. -Periclirtrer~aeus fridentatus - NAIM, 1980, annex 1-2 (Moorca; i n  sea-weed liali~izedn op~mrin). 
- CBACE & BRIJCE: 1993: 92. 93 (Syn. & key). - Periclinrennerrs cf. fridetrtatus - BRUCE personnl 
communication (cf, remark). 
REMARK. - According to BRUCE, personal communication: "I did get hold of the NAIM Periclinienaees rridenralus 
sprns from Leiden and in my opinion they werc definitely 1101 that species. They could be ncw. I made some drawings 
and then unfortunately lost the best spccitneti! ... What I would like is some more specimens." 
P e r i c l i m e t z a e v s  tuat t iotae B r u c e ,  1 9 6 9 a  
DlSrRtnlJIION. - Tuamotu (Moruroa). 
KSreRENCEs. - Periclin~enaeus frrnnrotae Brucc, 1969a: 170 (Moruroa). - C l r ~ c e  & BRUCE, 1993: 54 (list).  
P e r i c l i f n e n e l l a  spitrifera (de M a n ,  1902)  
DISTRlBUTlON. - Society (Moorea, Tahiti); Tuatnotu (Takaporo) - Free-living or coral associuie (Gen. I'ocillopora). 
REFERENCES. - Anchistin inaeqrrirnona - HELLER, 1865: 109 (Tahiti). - Periclintenes (Harpi l ias )  
spinifertrs - HOLTHUIS, 1952: 76, fig. 30 (Syn.). - Periclimenes spifiiferus - ODINETZ, 1983: 207 (Moorea, 
Tahiti. Takapoto; non obligate coral Pocillopora associate). - ODINETZ-COLLART & RICHER DE FORGES, 1985: 201 
(Moorca, Tahiti. Takapoto; in coral Pocillopora). - Periclintenella spinifera - DURIS & BRUCE, 1995: 656, 
fig. 19-20 (New combination; free-living, and associated with corals of the genera Acropora, Millcporn, Pnvona, 
Pocillopora, Porites and Seriatopora, or with a hydroid Millepora tenera). - BRUCE, in preparation (Host genera: 
various) - SYNONYM - ? Anchistia inaeqsinmna Heller. 
Periclittienes elegatis  (Paulson, 1875) 
DISSRIBUTION. - Society (Tahiti); Tuamotu (Moruroa) - Free-living or coral associate (Gen. Pocillopora). 
REFERENCES. - Periclinzene~ elegans - BRUCE, 1983: 884. 898 (Free-living shrimp). - NEW MATERIAL - Coll. 
0. ODINETZ, 1982, det. A.J. BRUCE 10/12/82, (Tahiti MNHN Na6618; in coral Pocillopora damicornis and 
P.  elegans) -RELEVANT MATERIAL - Coll. 3. POUPIN April-May 1997, det. A.J. BRUCE with a ? (Moruroa MNHN 
Na13408; in Pocillopora). 
Pericl i tnenes ensifrons (Dana, 1852) 
DISl'RIBu~TlON, - Tuan~otu (Amanu. Fakahina) - Free-living. 
REFERENCES. -Periclin~elres ensifrons - NOBILI, 1907: 359 (Amaou, Fakahina). - CMACE & BRUCE, 1993: 95, 
111 (Syn. & key). 
Periclirnenes grandis  (Stimpson, 1860) 
DlsTRlBmoN. - Tuamotu (Fangataufa, Moruroa) - Cord associnte (Gen. Pocillopora). 
REFERENCES. - Periclimenes grandis - New MATERIAI. - Coll. J. POUPIN April-May 1997, det. A.J. BRUCE 
(Fangataofa MNHN Na13414, Moruroa MNHN Na13407; in coral Pocillopora). - FRANSBN, 1994: 151 ("Free- 
living") - RELEVANT MATERIAL - ? Anchistia danae Stimpson, 1860: 41 [I081 (Tahiti; cf. remark). 
. . - - 
ever be resolved. It probably is a Periclinrenes, at least. It is probably a well known species under some other name, 
possibly one of thc P. grandis group (KEMP, 1922: 138). It lacked both second pereiopods and the ambulatory 
pereiopods, so is not really worth wasting further time on. The sooner it is completely Forgotten the better ..." 
Periclitnettes lutescens (Dana, 1852) (doubtful) 
DlsTRmIJTION. - Marquesas (Nuku Hiva); Society (Tahiti) - Coral associate (Gen. Acropora). 
REFERENCES. - Hnrpilius latescens - BOON& 1935: 167, pl. 45 (Nnku Hiva, Tahiti). - Periclimenes 
latescens - BRUCE, 19768: 6 ("Tlm species has been confused with P, consobrim<s De Man, and some records need 
to be checked"); 1976b: 98 ("The records from Samoa. Tahiti and the Marquesas islands are in particular need of 
confirmation"); in preparation (Host genera: Acropora, rarely Pocillopora, Seriatopora). - CUACE & BRUCE, 1993: 
94, 117 (Syn. & key). 
Periclitnenes tnadreporae Bruce, 1969a 
DISTRIBUTION. - Society (Moorea, Tahiti); Tuamotu (Takapoto) - Coral associnte (Gen. Pocillopora). 
REFERENCES.  - Periclinrenes ntadreporae - ODINETZ, 1983: 205 (Moorea, Tahiti, Takapoto; in coral 
Pocillopora). - BRUCE, in preparation (Host genera: Acropora, Pocillopora, Serintopora, Stylophora, Twbirmria). 
Periclitnenes niahei Bruce, 1969a 
DlSTRlBWSlON. - Tuamotu (Fangataufa) - Coral associule (Gcn. Acropora). 
REFERENCES. - Periclintenes nrahei - BRUCE, in preparation (Host gellera: Acropora, Pocillopora, Seriatopora) - 
NEW MATERIAL -Coil. J. POUPIN, February 1996, det. A.J. BRUCE (Fangataufa; in coral Acropora, lorn). 
Periclitrienes poupini  Bruce, 1989 
DISTRIBUTION. - Austral (Maria, Rapa, Rurutu. Tobuai); Gambier; Society (Bora Bora); Tuamotu (Fangataufa, 
Moruroa) - Ascidian associate; deep (409-600m). 
REFERENCES. - Periclintenes porrpini Bruce. 1989b: 852, fig. 1-5, 6a (Bora Bora, Fangataufa, Gambier, Maria, 
Momroa, Kapa, Rurutu, Tubuai; 409-600m). - POUPIN, 1996b: 6, pl. 2c (Same material). - Periclinrenes sp. 
nov. - POUPIN el al., 1990: 16 (Frcncll Polynesia). - POUPIN & RICHER DE FORGES. 1991: 211 (French Polynesia). 
REMARK. - Lives on a the sea-anemone carried on the shell occupicd by a parapagurid. Sympogrrr~rs dofleini (Balss, 
1912). 
17 
Periclirnettes soror Nobili, 1904 
DISTRIBUTION. -Society (Tahiti); Tuamotu (Moruroa) - Ecltinodern~ nssocinle (Genus Culcita). 
REFERENCES. -Peridin~enes soror - BRUCE, 1978: 299. fig. 1-6 (Moruroa, Tahiti); 1982: 200 (Asteroids hosts 
genera: Ckorinsler, Culcila, Penrocerasler, Proloreaster, Acunlhasler, Mifltrodia. Echinnsrer). - CIIACE & BRUCE, 
1993: 95, 122 (Syn. & key). - FRANSEN, 1994: 129, pl. 3f (Photograph) - NEW MATERIAL - Colt. J. I'OUPlN, det. 
A.J. BRUCE (Moruroa; on a sea star, Culcila ? novacguineae). 
Ph i l a r ius  ger lachei  (Nobili, 1905) 
DISTRIBUI'ION. -Tt~amotu (Moruroa) - Cord ussociote (Gen. Acropora). 
REFERENCES. -I'hilarius gerluchei - NEW MATERIAL - Coll. J. POUPIN AprilJMay 1996, det. A.1. BRUCE (Morura: 
in coral Acropora -10m; specimens with A.J. BRUCE). 
Pontonides unciger  Caiman, 1939 
DIS~RIBWON. -Marquesas (Tahuata) - Coral associale (Anthipataria). 
REFERENCES. - Ponfonides unciger - MONOD, 1979: 10, fig. 9-35 (Tahuata; associated with a coral 
Anthipataria). - FRANSEN. 1994: 136, pl. 4d (Photograph; Anthipataria associate). 
S tegopontonia  c o r n r ~ ~ e n s a l i s  Nobili, 1906 
D~STRIBUTION. - Tuamotu (Hao, Raroia) - Ecltinodern~ associn~e (Gen. Echinotbrix). 
KEFERENCES. -Stegopontonia comniensalis Nobili, 1906: 258; 1907: 360, pl. 1, fig. 2 (Hao). - SEURAT, 
1934: 60 (Commensal of sea urchin Echirrorris rrrrcarum). - Hol:rllurs, 1953a: 58 (Raroia; co~nrnensal of 
Echinotlrrix diadernn). - MORRISON, 1954: 6 (Raroia). - BRtJCa. 1982: 204 (Echinoid hosts gcnera: Diadenra and 
Eclrinorhrix). 
Vir sp. nov. Bruce (in preparation) 
DISTRlBunON. - Tuamotu (Momroa) - Coral ossociale (Gen. Pocillopora). 
REFERENCES. -Vir sp. nov. - BRUCE in preparation (Momroa. coll. 1. POUPIN, February 1996, AprilIMay 1997; 
in coral Pocilloporn, 10m). 
FAMILY AU'HEIDAE 
Alpheopsis dinbolus Banner, 1956 
DlSrRlBUT1oN. - Society (Bora Bora, Tahiti) - Free-livbrg or coral associnle (Gen. Pocillopora). 
REFERENCES. -Alpheopsis diabolus - BANNER & BANNER, 1967: 262 ('BD8, BD22 = Rora Bora). - ODINETZ, 
1983: 208 (Tahiti; in coral Pocillopora, non obligate associate). - Cllhca. 1988: 5 (Syn.). 
Alpkeopsis equal is  Couti&re, 1896 
DlSTRmWrlON. - Society (Bora Rora, Tahiti). 
REI:ERENCES. - Alphenpsis epuali.~ - B A N ~ R  & BANNER. 1967: 262 ("BD3-4, BD6, BD30" = Tahiti, "BD9, 
BD13-14, BD18" = Bora Bora); 1983: 11 ("collected from reef flat to 150m"). 
Alpheus  aculeipes var. t r y p k o p n s  Nobili, 1906 
DISI'RIB~~~ION. - Tuamotu (Marokau). 
REFERENCES. -Alpheus aculeipes var. tryphoprts Nobili, 1906: 257; 1907: 355 (Marokau). 
Alpltens nmirantei  sizo~c Banner & Banner, 1967 
DISTRIB~ION.  - Society (Bora Bora, Tahiti). 
REFERENCES. -Alpherts arriirarrlei siznu Banner & Banner, 1967: 265 (Bora Bora, Tahiti; outer reel platfolm) 
- Mwn. 1984: 91 (Collected in living or dead coral). 
Alpiceus bradypus Contiere, 1905a 
DlSrRlBUT1oN. -Society (Bora Bora). 
REr+3RENcas. -Alpheus bradypns -BANNER & BANNLR, 1967: 275 ("BDX = Bora Bora, inner rcct). 
18 
Alpkeus bidens (Olivier, 1811) 
DISTRIBUTION. -Tuamotu (Takapoto) - Free-living or coral associare (Gen. Pocillopora). 
REFERENCES -Alpheus bidens - ODINETZ, 1983: 207 (Takapoto; in coral Pocillopora, non obligate associate). 
Alpheus brevipes Stimpson, 1860 
DISTRIBUTION. -Society (Moorea, Tahiti). 
REFERENCES. -Alpheus brevipes -BANNER & BANNER. 1967: 274 ("BD2" = Tahiti. from various coral heads. 
"BD33" = Moorea, probably on the fringing reeo. 
Alpkeus bucepltulus Coutiere, 1905a 
DISTRIBUTION. - Socicty (Moorea); Tuamotu (Takapoto) - Non obligare coral associate (Gen. Pocillopora). 
REFERENCES. - Alpheus bucephuius - KROPP & BIRKELAND, 1981: 630, tab. 5 (Moorea, Takapoto; in coral 
Pocillopora). - CHACE, 1988: 17 ("Intertidal to 80m"). 
Alpheus collumianus Stimpson, 1860 
D~STRIBUTION. - Society (Bora Bora, Moorea, Tahiti); Tuamotu (Makatea, Marutea South, ?Takapto) - Free-living 
or coral associate (Gen. Pocillopora). 
REFERENCES. -Alpheus seurafi Coutihre 1905% 881, pl. 75, fig. 20, 20a-e; 1905b: 22 (Marutea). - SEURAT, 
1934: 60 (Marutea South). - Alpheris coNumianus - NoBILI, 1907: 354 (Marutea South). - SENDLER, 1923: 46 
(Makatea). - CHACE, 1988: 19 (Syn.). - MIYA, 1974: 109, pl. 19, fig. a-d (See remark); 1984: 92  (Under coral 
rock and among branches and in holes in bases of living Acropora). - ODINETL, 1983: 207 (Tahiti, but recorded as 
"Takapoto" in Paris collections; in coral Pocillopora, non obligate associate). - B A N ~ R  & BANNER, 1983: 27 
(Specimens found in association with a sea star, Opliiorhrix keysroniaj - RELEVANT MATERIAL - Alpheus 
collrrnrianus nredius - BANNER & BANNER, 1967: 264 ("BD3, BD30" = Tahiti, "BD8, BD22" = Bora Bora, 
"BD33" = Moorea). - MIYA. 1974: 112, pl. 16, fig. e-g (See remark). - Alpheus C O ~ ~ U ~ Z ~ U N U S  inermis - 
BANNER & BANNER, 1967: 264 ("BD2-3. BD5-6, B D 3 0  = Tahiti, " B D 1 6  = Bora Bora. "BD33" = Moorea; see 
remark). - Aipheus probabilis - ODINETZ, 1983: 207 (Moorea, Tahiti; in coral Pocillopora, non obligate 
associate; recorded under A. col1umianu.s probabilis Banner. 1956, det. BANNER, in MNHN collections). 
RBMARK. - From inaterial collected in the Marianas. BANNER (1956) has established three collamianu subspecies: 
Alplzetis collrin~ianrcs probablis, A. c. medim, and A. c. inermis. He supposed that Alpheus collrimianus probabilis 
would probably be identical with STIMPSON'S typical species but because of the distance from type locality (Bonin 
Islands) and the Marianas, and the lack of specimens from the Bollins be considered that "it was inadvisable to 
aswme this ( A .  c .  probabilis) to be exactly the same form as would be found there". This question remains still open 
(see MIYA, 1974) and the exact identity o i  these three subspecics must be clarified. For example, BANNER & BANNER 
(1967: 265) write: "According to field notes, a pair of cherry-red specilnens apparently cohabiting were collected off 
southern Bora Bora and were bottled together as BD16. By the criteria estahlisl~ed to separate the two subspecies, 
one appears to be A. c .  nledius and one A .  c .  inermis. Obviously this collection throws considerable doubt on the 
validity of the subspccies, but more specimens should be examined before the subspecies, distinct anatomically, are 
discarded". 
Afplteus crockeri (Armstrong, 1941) 
DIsrRlom.roN. - Society (Bora Bora. Moorea. Tahiti). 
REI:ERENCES. -Alpheus crockeri - BANNER & BANNER. 1966a: 159 (Syn.); 1967: 264 ("BD3" = Tahiti, from 
coral heads, "BD8, inner reef, BD12, mud and sandflats most specimens under rocks" = Bora Bora; Moorca). - 
MIYA. 1974: 124, pi. 21 (Syn.; living in dead coral head and on corallin algae). 
Alplteus cytkereus Banner & Banner, 1966b 
D l ~ ~ R l l 3 1 ~ 7 0 ~ .  - Society (Tahiti). 
REFERENCES. -Alphells cythereus Banner 8: Banner, 1966b: 125, fig. 45 (Tahiti; from head of dead coral); 
1967: 282 (Tahiti). 
Alpheus diaderrza Dana, 1852 
D l s r n l n u n o ~ .  - Garnbier (Mangareva); Society (Bora Bora, Moorea, Tahiti) - Free-living, coral (Gen. Pocillopora), 
or algae (Gen. Actinotrichia. Halimeda, Hydroclathrus, Sytnploca, Turbinaria) associate. 
REFERENCES. -Alpheus insignis - NOBILI. 1907: 355 (Mangareva). - Alpheus diodema - BANNER, 1956: 
357 (Syn.). - BANNI'.R & BANNER, 1967: 267 ("BD3, coral heads, BD37, inner reef" = Tahiti, "BD8, inner reef, 
BD13, barrier rcef, BD21, sandy area from scattered Ileads o f  dcads corals, BD26. BD28, from dead coral heads, 
behind growing rcef" = Bora Bora). - MIYA, 1974: 152, pl. 29 (Syn.; collected in bases of coral Pocillopora, 
fleliopora, and Poritcs, or in coralline algal massesj. - NAIM. 1980, annex 1-2 (Moorea; in algae Hydroclathrlrs 
19 
claclrrdm, ilalinreda oprnria, Syrtrplocn Irypnoidcs, Twbinurin ornam, Acrinorricl~ia rigidu). - ODINETZ. 1983: 207 
(Moorca. Tahiti; in coral Pocilloporo, non obligate associate). - ODINETZ-COI.I.ART & RICIIt5R DE FORGES, 1985: 
201 (Moorea, Tahiti; in coral Pocillopora) - SYNONYM - Alplieus insignis Heller. 1861 
Alpheics dolerus Banner, 1956 
DlsTRrlIuilON. - Society (Bora Bora, Moorea, Tahiti). 
REFERENCES. -Alpheris dolerus - BANNER & BANNER, 1967: 283 ("BD4, reef flat" = Tahiti, "BD14. BD18, 
BD20, from massivc dead heads of Porires" = Bora Bora,"BM3" = Moorea). - BANNER & RANW,R, 1983: 126 (Sand 
with shells and corals, hard blocks). 
Alphem edamensis I k  Man, 1888 
Dts'rRlBullON. -Society (Tahiti). 
REFERENCES. - Alphem acanthonlerus Ortmano, 1890: 474, pl. 36, fig. 12 (Tahiti). - Alpheus edanrensis 
- B A ~ W E R  & BANNER, 1966b: 157, fig. 61 (At low tide undcr rocks; "Society"). - MIYA, 1984: 92 (Under dead coral 
rocks in fine coral sands). - CHACE, 1988: 24 (Syn.; intertidal to 50111). 
Alpheus edwardsii (Audouin, 1826) 
D1sTRInmlON. - Society (Tahiti). 
RcFsRENCIis. -Alpheus edwardsii - ORTMANN, 1890: 470 (Tahiti). - CllAce, 1988: 25 (Syn.; intertidal under 
rocks to 251n) - RI;LEVANT MATSKIAL - A l p h e i ~ s  cf. edwardsii - NAIM. 1980, annex 1-2 (Moorea; in algae 
Z'wbinaria ornala, Giffordia nriickellae, Acrinorrickia rigidu). 
Alplzeirs frontalis H .  Milne Edwards, 1837 
D l s ~ ~ l l I U ' f l o ~ .  - Society (Moorea, Tahiti); Tuamotu (Raroia) - Under rocks in a l g d  tube, or c o r d  associate (Gen. 
Pacilloporaj. 
R r ; ~ o ~ a ~ c a s .  - Alpheus fro~ifal is  - HELLER, 1865: 107 (Tahiti). - Alpheus frnnfalis - BANNER. 1959: 141 
("Hanial~omo" = Kairoa; in algal tube, undcr rocks). - MIYA, 1974: 135, pl. 23, fig. d-g ("Could be found living io 
pairs in tubes which they made from filamentous blue-green or dark brown algac (Gen. Lyngbya). These tubes were 
found branching and penelratitig between the fronds of dead corals ..." ). - BANNER & BANNER, 1983: 30 (Living in 
felted tubes of fila~oentous blue-green algae, genera Plecronema, Lyngbyn, Microcolerrs). - ODINETZ, 1983: 207 
(Moorca - locality not indicated clearly in the work but mentioned in MNHN cataloguc; in coral Pocilloporo, no11 
obligare associate). - CHACE. 1988: 28 (Syn.; "Alburross specimens were found swimming at the surface undcr an 
elcctric light"). 
Alpheus grncilipes Stimpson, 1860 
DISTRIIIUTION. -Marquesas; Society (Bora Bora, Moorea, Tahiti) - In  algae (Gen. Halirneda, Symplocaj or  coral 
(Gen. Pocilloporaj. 
KEFBHENCRS. -Alpheus gracillipes Stinipson, 1860: 32 [I001 (Tahiti "intcr corallia"). - HEI.l.aR, 1865: 108 
(Tahiti). - OWMANN, 1890: 488 (Marquises). - BANNER & BANNER, 1967: 276 rRD3,  coral hcads, BD30, BD36, 
inncr rcer' = Tahiti. "BD7, shallow fringing rccf, BD13, sandy hottorn horn various living and dead corals, RDI4, 
BD19, from Pocillopora, BD21-22. BD25-26, BD28-29" = Bora Bora; Moorea; redescription). - MIYA, 1974: 154, 
pl. 30 (In dcad and overgrown corals of Pocillopora spp., Slylophora n~ordux and others). - NAIM, 1980, annex 1-2 
(Moorca; in algac Hnlimedn opicnlirr and Synploca Itypnoides). - KKOPP & BIRKEI.AND, 1981: 630. tab, 5 (Moorea; 
in coral Pocillopora). - ODINETZ, 1983: 207 (Moorea, Tahiti; in coral Pocilloporn, non obligate associate). 
Alpheus gracilis Heller, 1861 
DIssRlerrrroN. - Society (Bora Bora, Tahiti). 
REFERENCES. - Alpheus grncilis sinrplex - BANNER & BANNER, 1967: 268 ("BD18, fro111 consolidated rccf flat 
made of massive hcads of Porires sp." = Rora Bora, "RD30, from various heads of coral" = Tahiti). - Alplieus 
gmcil1.r - MIYA. 1984: 94 (With dcad corals). - C~IACE.  1988: 30 (Syn.). 
Alpheus Itoplites Nobili, 1906 
Dls~RlnU'noN. -Tuamotu (Marutea South). 
K E ~ ; E R I ~ N C E ~ .  -Alphens haplites Nobiii, 1906: 257; 1907: 357, 111. 1, fig. 7 (Tuamotu). 
REMARK. -Since its description, Alplreus hopliler secnis to h a w  disappeared from the literature. Type specinien is 
liowever still in MNHN collections (Na 2508, 1 sp. in bad condition; coll. G. SEURAT Marutca South 1905, det. 
Nortar). 
Alpheus  idiocheles Coutikre, 1905a 
DISTRIIIUTION. - Society (Rora Rora, Moorea, Tahiti) - Brcrrower. 
REI:ERENCES. -Alpheris idiocheles - BANER & BANNF.R. 1967: 271 ("BD2, various coral head" = Tahiti, "BD9, 
outer ridge of coralline alga" = Bora Rora, "BD32, wave-washed ridge of calcareous alga" = Moorea; "specimens of 
A. idiocheles ... were found living in galleries"). 
Alplieus lobidens De Haan 1850 
D~~TRIBWION. -Gambier (Mangareva); Society (Bora Bora, Maupiti. Moorea, Tahiti). 
REFERENCES. -Alpheus lobidens - N O I ~ I ,  1907: 356 (Matigareva). - BANNER & BANNER, 1983: 36 (Syn.). - 
Alpheris crassinranrr.s - BANNER & BANNER, 1967: 283 ("BD1, under algal mat with small musscls, BD35, under 
coral hcads" = Tahiti, "BD12, mud and sand flat, under rocks, BD24, frotn coral boulders, BD25, on a sand mud spit" 
= Bora Rora. "RD23" = Maupiti. "RD33" = Moorca) - SYNONYM -Alpl?erts crtrssinzanrrs Heller, 1865. 
Alpheus  lot t ini  Gutkin-Meneville, 1829 
DISTRIBUTION. - Society (Moorca, Raiatca, Tahiti); Tuamotu (Moruroa, Takapoto) - Coral associnte ( G e n .  
Pocillopora). 
REFERENCES. -Alpheris lnevis - HELLER, 1865: 107 (Tahiti). - Alpheris obesonranus - BooNE, 1935: 135 
(Raiatea, Tahiti; in coral) "partim" in BANNER & BANNER. 1964, not A .  obesornanrrs Dana. 1852. - Alpheris 
lottini - BANNER & BANNER, 1964: 88 (Syn.); 1967: 267 ("BD2, various coral head" = Tahiti, "BD15, from 
Pocilloporn, BD26, living and dead coral" = Bora Bora; Moorea). - KROPP & BIRKEr.ANl), 1981: 629, tab. 5 
(Moorea, Takapoto; in coral Pocilloporn). - ODlNwz, 1983: 205 (Moorea, Tahiti, Takapoto; in coral Pocilloporn). 
- KIM & ABEI.R, 1988: 33, fig. 11 (Syn.; Exclusively i n  coral I'ocilloporidae). - FRANS'N el a / . ,  1997: 154 ( B y  
collections RMNH D5544; Tahiti) - FIELD Ol3SERVA'SlON - J. POUPIN. 1997 (Moruroa; in coral Pocilloporn) - SYNONYM 
- Alpherrs lneeis Randall, 1839. 
Alpheus  mal leodig i tus  (Date, 1888) 
DISSRIBVSION. -Society (Bora Rora, Moorca, Tahiti) - Free-living or coral ussocinre (Gm.  I'ocillopora). 
REFERENCES. -Alpheus nrulleodigifris - BANER & BANNER, 1966a: 174 (Syn.); 1967: 270 ("BD3, from coral 
head, RD4, frarn reef flat, RDh, RD30, various heads of corals" =Tahiti, "BD8, BD13, From various living and dead 
coral, BD14, BD18, BD20, from massive dead hcads of  Poriles, BD28-29, frotn dead coral heads" = Bora Bora, 
"BD33-34" = Moorea). - MIYA, 1974: 130. pl. 22 fig. f-g (Syn.; in holes of the basal part of living, overgrown and 
dead corals). - ODINETZ, 1983: 207 (Tahiti; i n  coral Pocillopora. non obligate associate). 
Alpheus  rnicroscaphis (Banner, 1959) 
DlsrRIBuTrON. - Society (Moorea). 
REFERtNCES. - Alpheris nzicroscaphis - BANNER & B A N N E R ,  1967: 273 ("BD32, wave-washed ridge of 
calcareous alga" = Moorea). 
A l p l w i s  rrzitis Dana, 1852 Ilwerlae sedis 
DIsTRn%uIlON. - Tuaniotu (Fakabina, Makatca). 
REFI~RENCES. -Alpheus nritis - Noslr.1, 1907: 355 (Fakaliiiia). - SENDLER, 1923: 46 (Makatca). - BANNER & 
BANNER, 1983: 55 (cf remark). 
REMARK. -BANNER & BANNER (1983: 55-57) write "There is a serious question in our minds about the separation of 
A .  pnrncritiwns and A .  ntiris Dana ... Ui~fortunakly we cannot reexamine the type specimens for that of DANA is 
missing ... Thercforc the question of the scparation of the two species must at prescnt be left unanswered ..." 
Alpheus rrobili Banner & Banner, 1966a 
DISTRIBUTION. -Society (Tahiti). 
RetmENCrs. -Alpheus nohili - BANNER & BANNER, 1967: 269 ("BD30, from various heads of coral" = Tahiti). 
Alpheus oahuertsis (Banner, 1953) 
DISTRIRUTION. -Tuamotu (Takapoto) - Free-living or coral nssocinle (Gen. Pocillopraj. 
REFERENCES. - Alpherrs oulraensis - Ol)lNl;l'%, 1983: 207 (Takapoto; i n  coral Pocilloporn, non obligate 
associate). 
Alpheus  obesomnnus Dana, 1852 
D I ~ S R I ~ ~ ~ I ' I O N .  - Society (Bora Bora, Moorca, Raiaiea, Tahiti) - Free-living or coral ossociae (Gen. Pocillopora) 
REFERENCES. -Alpheus obesonranrrs - BOONE, 1935: 135, pl. 35, in coral; "partim" in BANNER, & BANNER 
(1964: 88), (Raiatea, Tahiti). - BANNER & BANNER. 1966a: 174 (Syn.); 1967: 269 ("BD2-3. from coral heads, BD4, 
from reef flat, BD5, outer ridge encrusted with smooth coralline alga, BD30, various heads of coral, BD37" = Tahiti, 
"BD8-9, BD13, frotn various living and dead corals, BD14, BD19, from dead and overgrown heads of Pocilloporn 
spp., BD20, from massive dead heads of Porites, BD21, from scattered heads of dead coral. BD26, Corn living and 
dead coral" = Bora Bora, " B D 3 2  = Moorea). - MIYA, 1974: 127, pl. 22, fig. a-e (Syn.). - OIXNETZ, 1983: 207 
(Tahiti, not indicated in work but mentioned in MNHN collections; in coral Pocilloyora, non obligate associate). 
Figure 3 - Alpheus obesomanirs Dana, 1852 
(From MRA, 1974) 
Alpheus  ovnliceps Coutiere, 190% 
DIsTRIBUnoN. - Society (Bora Bora. Tahiti). 
REFERENCES. -Alpheus ovaliceps - BANNER & BANNER, 1967: 275 ("BD3, from coral lieads" = Tahiti. "BD8, 
frotn inner reef' = Bora Bora). - CHACE, 1988: 44 (Syo.). 
DIsTRIBUrloN. - Society (Bora Bora, Moorea, Tahiti) - Free-lising or coral associate (Gen. Pocillopora). 
RBI:ERENCBS. -Alpheus pachychirus - DB MAN, 1890: 116, pl. 6, fig. 14 (Tahiti). - ORTMANN, 1890: 489, pl. 
26, fig. 17a-k (Taliiti). - BANNER & BANNER, 1967: 275 ("BD7, BD13, frotn various living and dead corals, BD14, 
BD15, from Pocillopora spp.. RD20, from massive dead heads of Porites, BD26, from living and dead coral" = Bora 
Bora; Moorea). - MIYA, 1974: 133, pl. 23, fig. a-c (Syn.). - ODINB~Z, 1983: 207 (Moorea, Tahiti, Takapoto; in 
coral Pocillopora, non obligate associate). 
Alpheus pacif icus Dana, 1852 
D1sTRlBUT1oN. - Gambier (Mangareva); Society (Bora Rora, Moorca, Tahiti); Tuamotu (Taiaro). 
REFERENCES. -Alpheris pacificus - N ~ B I I J ,  1907: 356 (Mangareva). -BANNER & BANNER, 1967: 283 ("BD8" = 
Bora Bora; Moorea). - KIM & ABE1.E. 1988: 100, fig. 42 (Syn.). - POUPIN, 1994: 6, fig. 2 (Mangareva, Taliiti, 
Taiaro; at low tide, sandy area, under rocks). 
Alpheus  paracritiniis Miers, 1881 
DlsTRmuTloN. - Society (Bora Bora, Moorea. Tahiti); Tuatnotu (Takapoto) - Free-living or coral associate (Gen. 
Pocillopora). 
REFERENCES. -Alpheus paracrifinos - B A N N ~  & BANNER. 1967: 278 ("BD4, BD30, various lheads of corals. 
BD36-37" = Tahiti. "BD11, from dcad hcads of Porites, BD13, various living and dcad corals, BD21, scattered l~eads 
of dead corals, RD25, BD28, dead coral hcads" = Rora Bora, "RD34" = Moorca). - MIYA, 1974: 157, pl. 31 (Syn.). 
- ODINI:TZ. 1981: 207 (Moozea. Tahiti. Takapoto; in coral Pocillopora, non obligate associatcj. - OnlNlrrz- 
COIIAR'r & RICHER 1)E FORGES. 1985: 201 (Takapoto). 
Alptieus p a r a d e ~ f i p e s  C o o t i h ? ,  1905a 
DT.STR~BW~N.  -Society (Tahiti); Tuamotu (Takapoto) - Fr~divinj i  or coral associme (Gen. Pocillopora) 
REI:BRENCES. - Alpheus pnrndentipes - ODINETZ. 1983: 207 (Tabiti, Takapoto; in coral Pocillopora, non 
obligate associate). 
Alpheus  parvirostr is  Dana, 1852 
DISTRIBU~ON.  -Gambier (Mangareva); Society (Bora Bora, Moorea, Tahiti); Tuamoto (Marutea South, Takapoto) - 
Free-living, coral associare (Gen. Pocillopora), or bivalve associnte (Gen. Pinctada). 
REFBRIINCES. -Alplreus pnrvirodris - C O U T I ~ R E  1905b: 23 (Marutea). - Nonrl,r, 1907: 356 (Mangareva). - 
SEURAT, 1934: 60 (In pearl oyster Pincrada mrrrgarirfera). - BANNER & BANNER, 1967: 284 ("BD4, BD6, BD36-37" 
= Tahiti, "BD7-8, BDI1. 8013-14, BD17, BD19-22, BD25-26, BD28-29" = Rora Bora. "BD33-34" = Moorea). - 
KROPP & BIRKELANU, 1981: 630, tab. 5 (Moorea, Takapoto). - ODINETZ, 1983: 207 (Moorea, Tahiti, Takapoto; in 
coral Pocillopora, non obligate associate). - CHACE. 1988: 47 (Syn.) - Alpheus euchiroides Nabili, 1906: 
257; 1907: 356, pl. 1. fig. 6 (Marutea). 
Alpheus  perplexus Banner, 1956 
DISTR~BUTXON. -Societv (Tahiti). , , 
REFERENCES. -Alpheus perplexus -BANNER & BANNER, 1967: 270 ("BD30, from various heads o f  coral" = 
Tahiti). 
Alpheus  platyurtguiculatus (Banner, 1953) 
DISSRIB~I'ION. - Society (Bora Bora, Moorea, Tahiti) - Free-lh'ing or coral ~ssociare (G.  Acropora). 
Rer;anr.wcr:s. -Alpheus platyurrgrriculotrrs - BANNER & BANNER, 1967: 281 ("BD17, from partially living 
Aoopora and coralline alga" = Bora Bora. "BD30, from various heads of coral" =Tahiti, "BD33" = Moorca). 
Alplteus ros t ra t ipes  Pocock, 1890 
DISTKIIIUTION. -Society (Bora Dora. Moorca, Tahiti). 
KEFERENCES. -Alpheus rosfrntipes - BANNER & BANNER. 1967: 268 ("BD2, BD30, BD38" = Tahiti, "BD8-9, 
BD14. BD17-18, BD22" = Bora Bora, "RD32" = Moorea). 
Alpheus  strertuus Dana, 1852 
DrsTRlBuTIoN. - Society (Bora Bora. Moorea, Tahiti); Tua~notu (Hao. Makatea. Takapoto) - Free-living or coral 
nsrssociare (Gen. I'ocillopora). 
REI'I~RENCES. - Alpheus strenuus - ORTMANN. 1890: 475 (Tahiti) - N O I ~ L .  1907: 356 (Hao). - SENDIXR, 1923: 
46 (Makatca). - BANNER & BANNER, 19668: 181 (Syn.); 1967: 284 ("BD12, mud and sand flats, most specimens 
under rocks" = Bora Rora; Moorea). - OUINETZ, 1983: 207 (Takapoto; in coral Pocillopora, non obligate associate). 
- C m c e ,  1988 (Syn.). 
Alpheus  sublucanus  (PorskBI, 1775) 
D I S T R I I I ~ ~ O N .  - Gambier (Mangareva); Marqucsas (Nuku Hiva); Society (Moorca. Raiatea, Tahiti); Tuamotu 
(Makatca). 
RE1:ERENCtis. -Alpheus vetrtrosus - NOBll.1, 1907: 355 (Kikitea). - SENDI.ER, 1923: 4 6  (Makatea). - BOONII, 
1935: 126, pl. 33 ("Anaho bay" = Nuku Hiva, "l'oint Venus" = Tahiti. "Tcviatoa" = Raiatea). - Alpheus 
sriblucnnes - HOLTHUIS, 1980: 122 (Syn.). - NAIM, 1980, annex 1, tab. 3 (Moorca) - SYNONYM - AlpAcr~s 
rwnlrosss H. Milne Edwards. 1837. 
Alpheus  sulca tus  Kingsley, 1878 
D l s ~ u l u m ~ o ~ .  - Marquesas (Nuku Hiva); Society (Bora Rora, Moorca, Tahiti). 
REFERENCES. -Alpheus trracrochirris - ORTMANN,  1890: 485 (Tahiti). - BANNER & BANNER,  1967: 268 
("RD14, from ridgc dclnarking outer margin of inner reef flat" = Bora Bora; Moorea; Nuku Hiva).- Alpheus 
sulcnfus - BANNER & BANNER, 1985: 65 (Syn.) -SYNONYM - Alpheus rnacrochirrrs Richtcrs, 1880. 
Athanas  areteforttzis Coutihre, 1903 
~ l S T R l l l ~ ~ l O h '  -Society. 
KI~FEUENC~IS. -Athntms nretefornris - B A N N I ~ R  & B A N N ~ .  1960: 138 (Syn.); 1967: 261 (Socicty). - C r r ~ c ~ ,  
1988: 61 (Syn.). 
Athanas  d j iboutens is  Coutikre, 1897 
DISTRIBUTION. - Gatnbicr (Mangareva); Marquesas (Nuku Hiva); Society (Moorca, Tahiti); Toamoto (Hikueru, 
Takapoto) - Free-living or coral ossocinre (Gen I'ocillopora). 
23 
REFERENCES. -Athanas djiboufensis - NOBILI, 1907: 353 (Hikueru. "Rikitea" = Mangareva). - BOONE, 1935: 
119, pl. 31. ("Anabo bay" = Nuku Hiva). - BANNER & BANNER, 1960: 140 (Tahiti); 1967: 261 (Society). - NAIM. 
1980, annex 1, tab. 3 (Moorea). - ODINETZ, 1983: 208 (Moorea, Tahiti, Takapoto; in coral Pocillopora, non 
obligate associate). - ODINETZ-COLLART & RICHER DE FORGES. 1985: 201 (Moorea and/or Tahiti, Takapoto). - 
CHACE, 1988: 62  (Syn.) - RELEVANT MATERIAL - Athanas cf. djiboutensis - KROPP & BIRKEI.AND. 1981: 630. 
tab. 5 (Moorea). 
Athanas dorsalis (Stimpson, 1860) 
DISTRIBUTION. - Gambier (Mangareva); Tuamotu (Fakarava, Hao, Marutea South) - Echinoid associate (Gen. 
Heterocentrotus). 
REFERENCES. - Arete dorsal*. var. Pacificus - COUTIBRE, 1904: 58 (Marurea South). - Arete nzarrrteerrsis 
Coutihre, 1905a: 864; 190%: 18, fig. 1-6 (Marutea South: in sea urchin Heferocentrofus nzantillafr~s). - NOBILI. 
1907: 353 (Mangareva, Hao, Fakarava, Marutea). - SEURAT, 1934: 60 (Marutea South). -Athnnas dorsalis - 
BANNER & BANNER. 1960: 151, fig. 5-6; 1967: 261 (Syn. and Tuamotu in distribution). - BRUCE, 1982: 207 
(Echinoid hosts genera: Centrosleplranus, EchinotArix. Stomoprreustes, Tr ipneus fe s ,  Heliocidnris ,  
Centrostephanus); 1990: 578 (Redescription with illustrations). - CHACE, 1988: 62 (Syn.). 
Athanas gracilis Boone, 1935 
DISTRIBUTION. -Society (Raiatea). 
REFERENCES. -Allranas gracilis Boone, 1935: 122, pl. 32 (Raiatea). - BANNER & BANNER, 1960: 132 (List). 
Athanas indicus (Coutiire, 1903) 
DlsTRlBmoN. - Society (Bora Bora) - Echbtoid associare (Get,. Echinometra). 
REFERENCES. - Athanas indicus - BANNER & BANNER, 1960: 149 (Bora Bora); 1967: 260-261 (Society; 
"Athanas indicus was found living in exacavations made by the sea urchin Ecitinometra"). - BRUCE, 1982: 207 
(Echinoid hosts genera: Diadeaa. Echinontetra, Anfhocidaris). 
Athanas rhotltionastes Banner & Banner, 1960 
DISTRIBUTION. - Society (Tahiti). 
REFERENCES. -Athanos rhothionasles Banner & Banner, 1960: 142, fig. 3 (Tahiti); 1967: 261 (Society). 
Automate gardineri Coutiire, 1902 
DISTRIBUTION. -Society (Bora Bora, Moorea) - Coral associate (Gen. Acropora). 
REFERENCES. -A~rtontate gardineri - BANNER & BANNER, 1966a: 150 (Syn.); 1967: 261 ("BD17, from partially 
living Acropora and coralline alga" = Bora Bora, "BD33" = Moorca). 
Metalpheus paragracilis (Coutikre, 1897) 
DISTRIBUTION. - Societv (Bora Bora. Tahiti): Tuamotu (Marutea South) - Free living or coral associate (Gcn. 
. . . . " 
Acropora). 
REFERENCES. - Alphees paragracilis - COUTIGRE 1905a: 883, pl. 76. fig. 22, 22a-g (Tahiti); 1905b: 23 
(Marutea South). - SEURAT, 1934: 60 (Marutea South). - BANNXR & BANNER. 1967: 268 ("BD6, barrier reef. BD30. 
from various coral heads, BD35, under coral heads on a gravel and sand expanse" = Tahiti. "BDR, inner reef, BD14, 
BD17, from partially living Acropora and coralline alga, BD18, from consolidated reef flat made mostly of massive 
heads of Porites, BD20, from massive dead heads of Porites" = Bora Bora). - Metalpheus paragracilis - 
BRUCE, 1976a: 44 (Syn.). 
Saltnoneus ? serratidigitus (Coutiire, 1896) 
DISTRIBUTION. - Society (Tahiti). 
REFERENCES. - Salnroneus ? serratidigitus - NEW MATERIAL - Coll. P. FROUIN, det. A.J. BRUCE "to be checked 
by Y. MIYA" (st. FRoUlN 1225, Tahiti; material with BRUCE). 
Salmonens sibogae (de Man, 1910) (presence uncertain) 
DaTRlBUTIoN. - Society (?). 
REFERENCES. - Salntoneus sibogae - BANNER & BANNER, 1964: 86 (Syn.); 1967: 261 ("Locality notes lost"; 
could be Society and/or Cook Islands). 
Salrnoneus tricristatns Banner, 1959 
DnTRlBU'r1ON. - Society (Tahiti). 
REFERENCES. -Salmo~~eus  fricristnta - BANNER & BANNER, 1967: 262 ('BD30, from various heads of coral" = 
Tahiti). - Saln~oneus tricrislatus - MIYA, 1972: 43, pl. 5 (Syn.). 
Syna lpheus  brachyceros Nobiii, 1906 
DrsTRrevnON. - Tuamotu (Makatea). 
REFERENCES. -Synalpheus hracbyceros Nobili, 1906: 256; 1907: 354, pl. 1, fig. 8 (Makatea). - dc MAN, 
1911: 190. 199 (List & kev). 
. , 
REMARK. - Species never rchieved since N0Ull.l'~ description. 
Synalplteus cha ron  (Heller, 1861) 
DISTRIBUTION. - Society (Bora Bora, Moorea, Tahiti); Tuatnotu (Takapoto) - Free living or cornl associate (Gen. 
Pocillopora). 
REFERENCES. - Sytlalpheus charon - BANNER & BANNER, 1967: 262 ("BD2. various coral heads" = Tahiti, 
"BD20, from massive dcad heads of Porites" = Bora Bora). - MIYA, 1972: 51, pl. 8 (Syn.). - BRUCE, 1976a: 48 
(Coral host, genus Stylophorn). - KROPP & BIRKELAND, 1981: 629, tab. 5 (Moorea. Takapoto; in coral 
Pocillopora). - ODINETZ, 1983: 205 (Moorea, Tahiti, Takapoto; in coral Pocillopora). - Crrhca, 1988: 77 (Syo.). 
Synalpheus  heroni  Coutiere, 1909 
D1sTRlnrrnO~. - Society (Bora Bora. Moorea, Tahiti) - Free living or cornl associare (Gen. Acropora). 
REFERENCES. - Synalphem herani - BANNER & BANNER, 1967: 264 ("BD2, various coral heads" = Tahiti, 
"BD17, from partially living Acropora and co:orallinc alga" = Bora Bora. "BD32. from ridge of calcareous alga" = 
Moorea). 
Synalpheus  nilandensis Cuutikre, 1905a 
D ~ S T R ~ B I ~ O N .  - Gamhicr (Mangareva) -Bivalve associale (Gen. I'inctada). 
RBFEKENCES. -Syrmlpheus nilandetrsis - Nonrl.1. 1907: 353 (Mangareva). - SEURAT, 1934: 60 (Mangareva; 
in pearl oyster Pincmdri margaririfera). -BANNER & BANNER, 1983: 101 (Syn.). - Cmcr-. 1988: 82 (Syn.) 
Synalplrerrs paraneorneris  Coutiere, 190Sa 
DlsT~rI3rrrlo~. - Society (Bora Bora, Moorea, Tahiti); Tuamotu (Marutea Sooth. Takapoto) - Free-living or corrrl 
associac (Gen. Acropora, l'ocillopora). 
R~~PEKENCES.  - ynalpheus pamneomeris - COUTI~RE.  1905b: 22 (Marutea). - SEURAT, 1934: 60 (Maiutca 
South). -- BANNER & BANNER, 1967: 262 ("BD2. various o r a l  heads, BD6, barrier reef, BD30, various coral heads" 
= Tahiti, "BD17, frotn partially living Acropora and coralline alga" = Bora Bora. "BD32, frotn ridge of calcareous 
alga, RD33" = Moorca). - MrYn, 1972: 54, pl. 9 (Syn.). - ODINESZ, 1983: 208 (Tahiti; in coral Pocillopora, non 
obligate associate; Takapoto also indicated in Paris collections). - CrmcE, 1988: 83 (Syn.). 
Synalpheus  streptodactylus streptodactylus Coutikre, 1905a 
DrSTRlnunON. - Society (Moorca, Tahiti); Tuamotu (Takapolo) - Free-living or cornl associate (Gm. Pocillopora). 
REI:ERENCES. - Synalpheus strepfodactylus streptodactylus - BANNER & BANNER, 1966a: 157 (Syn.); 
1967: 264 ("BD34, behind the rcei" = Moorea). - MIYA, 1972: 69, pl. 14 (Syn.) - S y n a l p h e u s  
slrepfodaclylus - ODINGTZ. 1983: 208 (Tahiti; in coral Pocillopora, non obligate associate; Takapoto also 
indicated in Paris collections). 
FAMILY HLPPOLYTDAE 
Hippolyte ventricosa H. Milne ISdwards, 1837 
D I S T R I B ~ O N .  -Society (Moorea) - In algae (Gen. Turbinaria). 
REFERENCES. -Hippolyte ventricosa - HOLTIIUIS, 1947: 55, fig. 7-9 (Syn.). - NAIM, 1980, anltexe 1-2 
(Moorea; in algae Turbinaria ornam). 
REMARK. -According to C. <UI>EKEM n'Acoz (personal cotntnunication) most of tile Xndo-Pacific references to 
I f .  venrricosu need a revision. 
L igur  ensiferns (Risso, 1816) 
DlSSRlnunON. -Austral (Raevavae, Rapa. Rurutu); Ganlbier; Tuarnotu (Maria) - Deep (290-380n1). 
REFI~RENCES. -Ligur ensiferas - POUPIN el d.. 1990: 15 (French Polynesia). - POUIXN, 1996h: 6, pl. 2d 
(Gambier. Maria. Racvavae, Rapa, Rurutu; 290-380m). 
Lyysnlata arnboinensis (de Man, 1888) 
DISTNBI~ON. - Marquesas (Ua Pou) - Lirroral to deep (120n1). 
REFERENCES. - Lysmata arnboinensis - POUPIN. 1996b: 6, pl. 2e (Ua Pou; 120n1). 
Merhippolyte kauiaensis RatLbun, 1906 
DlsTRIBlmON. - Marquesas (Tahuata) -Deep (530m). 
REFERENCES. - Merhippolyte kauiaensis - POUPIN el a/ . ,  1990: 15 ("French Polynesia"; exact location, not 
indicated in text, is Tahuata, 530m; MNHN Na13299). 
Parhippolyte nveae Borradaile, 1900 
DlsTRmmION. - Society (Maupiti, Raiatea) - Deep (110-13Ont). 
REFERENCES. - Parkippolyte rrveae - POUPIN, 1996b: 6, pl. 2f (Maupiti. Raiatea; 110-l30m; usually this 
species is littoral). 
Saron rnarrnoratus (Olivier, 1811) 
DIsTRlBullON. - Gambier (Mangareva); Marquesas (Nuku Hiva); Society (Tahiti); Tuamotu (Amanu, ?Mororoa) - 
Cord  associnre (Gen. Pocillopora). 
REFERENCES. - Hippolyte gibbosus - HELLER, 1865: 120 (Tahiti). - Hippolyte gibberosn - ORTMANN, 
1890: 497 (HELLER'S reference). - Snrorr gibberosas - NoBn.1, 1907: 358 ("Rikitea" = Mangareva. "Omanu" = 
Amanu). - SENDLEK, 1923: 46 (Tahiti). - Saron nmrmoratas - BOONE, 1935: 187, pl. 51 (Nuku Hiva). - 
HOI;~HUIS, 1947: 6 (Syn.) - RELEVANT MATERIAL - Snron sp. a f f .  nmrntorotrrs - Coll. J .  POUPIN, det. A.J. BRUCE 
(Moruroa; in coral Pocillopora) - SYNONYM - liippolyle gibberosus H. Milne Edwards, 1837. 
Figure 4 - TAorina ~imldivonis (Rorradailc, 1915) 
(From BRUCE, 1997) 
Sarort neglectus De Man 1902 
DISTRIBUTION. - Society (Moorca. Tahiti); Tuainotu (Takapoto) - In algae (Gen. Giffordia, Ilalimcda, Liagora, 
Turbinaria) or in coral (Gen. I'ocilloporaJ. 
REFERENCES. - Saron rreglectus - NAIM, 1980, annex 1-2 (Moorea; in algae Giffordia nrifchellae, ilalinredn 
olmrin,  Liagora hawaiiarza, Turbinaria oranam). - KROPP & BIRKELAND. 1981: 630, tab. 5 (Moorea; in coral 
Pocillopora, non obligate associate). - ODINETZ, 1983: 207 (Moorea. Tahiti. Takapoto; in coral Pocillopora, non 
obligate associate) - ODINETZ-COILART & RlCllER DL FORGES, 1985: 201 (Moorea. Tahiti. TakaPoto; in coral 
Pocilloporn). 
T h o r  amboinens is  (De Man, 1888) 
DIS'SRlJ3IinON. - Tuamoto (Fangataufa) - Coral associrile (Gen. Acropora). 
~EIXRENCES. - Thor nnrboinensis - NEW MATERIAL - Coll. J. POUPIN, February 1996, dct. A.J. BRIJCE 
(Fangataufa; 111 coral Acropora, 10m; MNHN Na13303). 
T h o r  paschalis Heller, 1862 
DIS~KIIIWION. -Society (Moorca) - In algae (Gen Codium). 
REFERI:NCI.:S. - TIzor paschalis - HOLTHUIS, 1947: 49 (Syn.). - NAIM. 1980, annex 1-2 (Moorea; in algae 
Codirm geppii). 
Thor ina  rrialdiv~nsis  (Borradaile, 1915) 
DISTRIBU'SION. - Society (Moorca, Tahiti); Tuamotu (Takapolo) - Free-living or coral nssociale (Gen. Pocillopora). 
RE1:ERENCEs. - Thnr maldivensis - ODINBE, 1983: 207 (Moorea, Tahiti, Takapoto; in coral Pocillopora, non 
obligate associatcj. - Tltorina maldivertsis - BRUCE, 1997: 14, fig. 1-6 (Tahiti). 
FAMlLY PRDCESSDAE 
Nikoides aff. danae Paulson, 1875 
DlSrRlnmroN. - Society (Tahiti). 
REI:ERI:NCES. - Nikoides aff. dnane - NEW MATJiRlAI. - Coll. P. FROUIN, preliminary det. P. Nor:]., MNHN 
Na13053-54 (Tah~ti). 
Processa aff. moana Yaldwyn, 1971 
DISTRIIIUTLON. -Society (Tahili). 
R111:BRl:NCliS. - l'roce~sa a f f .  nloann - NEW MATERIAL. - Coll. 1'. FROUIN, preliminary dct. 1'. Noar.. MNHN 
Na13055 (Tahiti). 
Chlorocur t i s  jartarrs (Nobili, 1904) 
~lsTR1nu.slON. - Society (Moorca, Tahiti) - Free-living, algae (Gen. Turbinaria) or coral associare (Cen. I'ocillopora) 
Kr:.l;ERENcr;s. - Chforocurtis jnctons - NAIM, 1980, annex f -2 (Moorea; in algae Turbinariu orntna). - ODINETZ, 
1983: 207 (Moorea, Tahiti; in coral Pocillopora, IIOII obligate associate). 
Heterocarplis aff. ensifer A .  Milne Edwartls, 1881 
D I S S R T I ~ ~ ~ I O N .  - Marqucsas (Eiao, Fatu Hiva, Hiva O d .  Tahuata, Ua Pou) -Deep (217.730111, n~osrly 300-400m). 
RT~I%KI:.NCES. - Heferncnr~~us aff. erisifer - POIJI'IN el 01. .  1990: 16, 50, fig. 5-7. 11, pl. Id (French Polynesia). 
- I 'O~PIN.  1996h: 6, pl. 2g (Eiao, Fatu Hiva. Hiva Oa, Tahuata, Ua Pou; 217.730111). - Heterocnrpus errsifer - 
POUlW & RI(:IER I X  FORGES, 1991: 211, fig. 1 (Frcnch Polynesia). - Not Heterocar-pis ensifer - MANAC'R & 
CARSIN, 1985: 473. - CROSNIER, 19868: 362. - POUPIN, 1988: 27, fig. 2, 5 - TIlcse three refcrences co~~ccrned  
11. rm,ocrrla Crosnicr and/or /I. pnrvispinn de Man. 
Heterocarplis arrtacrila Crosnier, 1988b 
DJS~.KIIIU.I.ION. - Austral (Kapa, Tubuai); Gambia; Society (Tahiti); Tuamotu (Fangataufa, Mor~~ioa)  - Deep (250- 
ROOm, mosrly 4.70-650111). 
KEI:BKENcI:S. - Heferocarpus ertsifer - MANAC'II & CARSIN, 1985: 473 (Moruroa andior Fangataufa). 
CROSNIER, 1986a: 162  (Moruroa, Talliti, Tubuai). - POUPIN, 1988: 27, fig. 2, 5 (Moruroa) - All not /I. cnsifer 
A. Milne Edwards = I f ,  nnrt~culn sp. nov. in CROSNIER (1988b); some sp. arc N .  parvispina de Man). - 
Heferocnrpus nrnacriln Crasnicr, 1988b: 72, fig. 4a, 111. lb, 211 (Tuboai, Moruroa). - I'OUPIN el al., 1990: 15, 
56, fig. 5. 6, 7, 12, pi. I c  (Frcnch Polynesia). - Pou1)lN & RlCllEKDE FORGES. 1991: 211, fig. 1 (Frc~~ch  Polynesia). 
- I'O~JPIN, 1996h: 6. pl. 211 (Fangataufa, Gambier. Moruroa. Rapa, Tahiti, Tubuai; 250.800111). 
Nelerocarpris dorsal is  Bate, 1888 
DISTRlI3UflON. - Marqucsns (Tahuatn); Society (Tahiti); Tuamotu (Makano) - Deep (680-1025nz). 
REFERENCES. - Heterocarpus dorsalis - CROSNIER, 1986a: 362; 1988b: 62, fig. 2-3 (Tahiti). - POUPIN EI a / . ,  
1990: 16 (French Polynesia). - POUPIN, 1996b: 8, pl. 3a (Makemo, Tahiti, Tahuata; 680-l025m). 
Heterocarpus laevigatus Rate, 1888 
DISTRIBUTION. - Austral (Rapa, Tubuai); Gambier; Marquesas; Society (Bellingshausen, Huahine, Tahiti); Tuamotu 
(Fangataufa, Moruroa) - Deep (410-1050m; ntos~ly 500-800nl). 
REFERENCES. - Heterocarpu~ laevigatus - CROSNIER, 1986a: 362 (Tahiti, Tubuai); 1988b: 74 (Tubuai). - 
POUPIN el nl., 1990: 16, 37, fig. 5, 6, 7, 9, pl. l e  (French Polynesia). - POIJPIN, 1996b: 8, pl. 3b (Bellingshausen = 
Motu One, Fangataufa, Gambier, Huahine, Marquesas, Moruroa, Kapa. Tubuai; 410-1050m). 
Heterocarpus  longirostris  McGilchrist, 1905 
Heterocarpus parvispina de Man, 1917 
DISTRIBUTION. - Austral (Tubuai); Society; Tuamotu (Fangataufa, Moruroa) -Deep (230-730m; nzoslly 350-500n1). 
REFERENCES. - Heterocnrpes parvispina - CKOSNIER. 1988b: 70. 72, fig. 5a, pl. la ,  2g (Moruroa, Tubuai). - 
PDUPIN el a/., 1990: 16, 60, fig. 5-7, 13, pl. l b  (French Polynesia).- POUPIN & RICIIER DE FORGES, 1991: 211, 
fig. 1 (French Polynesia). - POUPIN, 1996b: 8, pl. 3d (Fangataufa. Moruroa; Austral, Society; 230-730m). - 
Heterocarpns ensifer. - MANAC'II & CARSIN, 1985: 473. - CROSNIER, 1986a: 362. - POIJPIN, 1988: 27, 
fig. 2, 5, 6 - AlLpro pnrlc not 11. ensfer A. Milne Edwanls, 1881 but a mixture of If, parvispinn and 11. rmraciilu. 
Heterocarpus sibogae de Man, 1917 
DisrRmrrnoN. - Austral (Tubuai); Society (Tahiti); Toamotu (Moruroa) - Deep (190-950nr). 
REPERBNCES. - Heterocarpus sibogae - CROSNIER, 19868: 362 (Tahiti); 1988b: 79, fig. Sc, pl. Ic, 3a-b (Tahiti, 
Tubuai; Tuamotu). - POUPIN, 1988: 27, fig. 2 (Moruroa); 1996b: 8, pl. 3e (Moruroa, Tahiti, Tubuai; 190-950m). - 
POUPIN el 01.. 1990: 16, 45, fig. 5-7, 10, pl. I f  (French Polynesia). - POUPIN & KICKER I>E FORGES, 1991: 211 
(French l'olynesia). 
Plesionika carsini  Crosnier, 1986a 
DISTRIBUTION. - Austral (Maria, Raevavae, Tubuai); Gambicr; Marquesas (Ua 1%"); Tuamotu (Makcmo, Maria, 
Moruroa, Tureia) - Deep (410-600m). 
REFERENCES. - I'lesionika carsini Crosnier, 1986a: 369, fig. 3a-i (Moruroa). - POUI'IN el a/ . ,  1990: 16, pi. 2b 
(French Polynesia). - PDUPIN & RICHER DB FORGES, 1991: 211 (French Polynesia). - POUPIN, 1996b: 8, pl. 3f 
(Gambier, Makemo, Maria, Moruroa, Kaevavae, Tubuai, Tureia, Ua Pou; 430-600m; in these locations s o m e  
specimens most belong to Plesionika pictn Chan & Crosnier). - CNAN & CROSNIER. 1997: 190, fig. 2a. 2b, fig. 21 
(Fangataufa, Gambier, Makemo, Maria, Moruroa, Tureia; 410-510m). 
Plesionika curvata Chan & Crosnier, 1991 
D I S T R I I ~ U ~ O N .  - Austral (Rapa, Rimatara, Tubuai); Gambicr; Tuamotu (Fangataufa) - Deep (150-400nt). 
REFERENCES. - Plesionikn sp. nov. 2 - POUPIN el a/ . .  1990: 16, pl. 31 (French Polynesia). - Plesionika 
crrrvata Chan & Crosnier, 1991: 440, fig. lob, Ild-f, 33 (Fangataufa, Gambier, Kapa. Tubuai). - POUPIN, 1996b: 
8, pl. 3.5 (Fangataufa, Gambier, R a p ,  Rimatara, Tubuai; 150-400m). 
Plesionika edwardsii (Urandt, 1851) 
DrsTRmu'rloN. - Austral (Kaevavae, Rapa. Tubuai); Gambier; Marquesas; Society (Tahiti); Tuamotu (Fangataufa, 
Moruroa) - Deep (200-500nt: nlosrly 250-350n1). 
REFERENCES. -- I'lesionika edwardsii - MANAC% & CARSIN, 1985: 473 (Fangataufa and/or Moruroa). - 
CROSNIER,  1986s: 362 (Moruroa, Tahiti, Tubuai). - POUPIN, 1988: 28, fig. 2 (Momroa); 1996b: 8. pi. 311 
(Fangataufa, Gambier, Marqucsas, Moruroa, Tahiti, Tubuai; 220-500m). - POUPIN el a / . ,  1990: 16, 68, fig. 5, 6, 7, 
15, pl. Ig (French Polynesia). -CHAW & CROSNIER, 1997: 193, fig. 23 (Moruroa, Raevavae, Rapa, Tahiti, Tubuai; 
200-450m). 
Plesionika erythrocyclus Chan & Crosnier 1997 
D I S T R I ~ I ~ I O N .  - Austral (Rurutu); Tuamoto (Moruroa) - Deep (200-260nr). 
R F ~ R B N C E S .  - Plesionika sp. nov. 1 - POIJPIN,  19961,: 10. pi. 4a (Mororoa, Kurotu; 200-260m). - 
Plesionika erythrocyclus Chan & Crosnier, 1997: 214, fig. 16. 34-35 (Moruroa, Rurutu; 200-260m; Syn.] 
Plesiortika fenneri  Crosnier, 198613 
DI~TKII~UTION.  - Austral (Kapa, Rurutu, Tubuai); Gambicr; Marqucsas; Society; Tuarnotu (Fangataufa, Maria, 
Moruroa) - Deep (260-820n1; nzoslly 500-700n1). 
REFERI.:NCES. - Plerionikn chncei Crosoier. 1986a: 363, fig. la-b, 2a-h (Moruroa, Tubuai). - Plssionikn 
fenneri Crosnier, 1986h: 691 (new name for P, cliocei). - POUI~IN,  1988: 28, fig. 2. 4-6 (Moruroa); 1996b: 10, 
pl. 4b (Fangataufa, Moruroa, Rapa, Tubuai; Marquesas & Society; 260-820m). - POUPIN el nl., 1990: 16, 29, fig. 
5-8, pl. l a  (French I'olyncsia). - POUPIN & R I ~ I B R  DBFORGES, 1991: 211 (French Polynesia). - CIIAN &CROSNIER, 
1997: 196, fig. 4, 26-27 (Gambicr. Maria. Kumtu; 470-660m). - Plesioniko aff. trispinus (sic) - POUPIN ci 
nl., 1990: 16 ("French Polynesia"; cxact location, not indicated, is Maria, 680111) - Not P, rrispinostrs Squires & 
Rarraguan, 1976 = P. fenneri juvcnilefide CCBAN & CROSNIER (1997: 196). 
Plesiorrika f lavicauda Chan & Crosnier, 1991 
DISTRIBIJ~ON. - Austral (Rurutu, Tubuai); Society (Tahiti); Tuamotu (Fangataufa. Moruroa, Takapoto) - Dcep (100- 
260mJ. 
KEI:ERENCES. - I'lesionikn sp.  nov. 1 - POUPIN el nl.. 1990: 16, pl. 3e (French Polynesia). - Plesinnikn 
flovicnuda Chan & Crosoier, 1991: 438, fig. IOa, l la-c, 29-32 (Fangataufa, Moruroa, Rurutu, Tahiti, Takapoto). 
- POUPIN, 1996b: 10. pl. 4c (same islands; 100-260m). - ? Plesionika serrntifrow - CROSNIER, 1986a: 362 
(Tahiti, Tubuai) not P. serrarift.or!s (Borradaile. 1900); matcrial not retrieved in MNHN but most probably belonging 
to P. flavicarrdn. 
Plesiorcika rrracropoda Chace, 1939 
Dlsr~mU'SloN. - Austral (Maria); Gambier; Marquesas (Ua Pou); Society (Huahine); Tuamotu (Maria. Moruroa, 
Tureia) - Deep (300-600n1j. 
RIIFERENCI~S. -Plesionikn aff. willionrsi - POUPIN el a/ . .  1990: 16 (French I'olyncsia), in part not P, williamsi 
Forest, 1964. - Plesinnika nracropodn - POUPIN. 1996b: 10, pl. 4d (Gambier. Huahine. Maria, Moruroa, 
Tureia, Ua Pou; 300-600m). - CRAN & CROSNIER, 1997: 205, fig. 11-13, 30-31 (Gambier, Huahine, Maria [Austral 
& Tunmotu], Moruroa. Ua Pou; 310-500m; Syn.). 
Plesiorrika nrzorfia. ( A .  Milne Edwards, 1883) 
DlsTulBUTloN. - Mnrqocsas (Nuku Hiva, Ua Pou) - Dcep (590-liS0nl). 
RE~rua,NcIs. -- I'le~ionikn ntnrtia - CHAN & CROsNlEn, 1997: 213 (Ua Pou, Nuku Hiva; 590-650m). 
Plesionika nurval (Pabricius, 1787) 
DISTRIBII?.ION. - Society (Tahiti) - S~~blirlorrrl (40-50ni). 
REFERENCES. - I1le,sioniko ~rnrvnl - CIUN & CKOSNICR, 1991: 443, fig. 12a-c, 13a, 14a-c, 15a-e, 34-36 (Tahiti; 
40-50m). 
Plesionika nes i s i  (Durukovsky, 1986) 
DISTRIBU'I'ION. - Marquesas (Hiva Oa); Society (Huahine); Tuamotu (Faogataufa, Mskerno, Moruroa) - Deep (550- 
1080ni; n~osrly 800-I000m). 
REFERENCES. - f'lesionikn aff. laevis - POUPIN el ul., 1990: 16, 72, fig. 16, pl. 2a (French I'olyncsia). - 
I'lesioniko nesiri - POIJPIN, 1996h: 10, pl. 4e (Fangataufa, Hiva Oa, Huahine, Makemo, Moruroa; 550-1080m). 
- CNAN & CKOS~?BR, 1997: 197, fig. 5-10, 28 (Faogataufa, Hiva Oa. Makemo, Moruroa; 600-l050m). 
Plesionika payeni Chan & Crosnier, 1997 
DrsTKlBvrION. - Tuamotu (Moruroa) -Deep (200-250mj. 
RRrEKENCES. - Plesionikn sp.  nov. 5 - POIIPIN, 1996b: 12, pl. 5g (Moruroa; 250m). - Plesionikn pnyeni 
Chan & Crosnier, 1997: 221: fig. 19, 39  (Moruroa, 200.250111). 
Plesiorcikn picta Chan & Crosnier, 1997 
DIsTRJI3u?lON. - Ausual (Maria, Rurutu); Tuamoto (Fangataufa, Moruroa) - D e q  (520-710nr). 
Re~Enl iNces .  - Plesionika sp.  nov. 2 - Poul'lN, 1996b: 10, pi. 4f (Fangataufa, Maria, Moruroa. Rurutu; 650- 
710m). - I'lssiorrika picfn Chan & Crosnier. 1997: 223, fig. 20, 40 (Fnagataufa, Gambicr. Maria. Kurolu, Ua 
Pou; 520-7 lorn). 
Plesionika porcpirci Chan & Crostiier, 1997 
~~~~~~~~ITON. - Ausiral (Raevavae, Rimatara, Rurutu); Gainhicr; Tuan~otu (Fangataufa. Makmno. Maria. Mororoa, 
Tureia) - Dee,) (260-6.70n1j. 
29 
REFERENCES. - Plesiortika carsini - CKOSNIER. 1986% 369. - POUPIN & RICER DE FORGES, 1991: 211 - Both 
references, p r o  pnrle, not P. cnrsini Crosnier 1986 = P. porrpini Chan & Crosnier, 1997. - Plesionika aff. 
carsini - POUPIN pf ol., 1990: 16, pl. 3h (French Polyncsia). - Plesionika sp. nov. 3 - POIJPIN, 1996b: 10, 
pl. 4g (Fangataufa, Gambicr, Makemo, Maria, Moruroa. Raevavae. Kimatara, Rurutu, Tureia; 280.6501~). - 
Plesionika poupini Chan & Crosnier, 1997: 189, Cig. 1-2, 22 (Fangataofa. Makerno, Maria, Moruroa, 
Racvavae, Ri~natara. Rurutu; 260-600m). 
Plesionika pro taf i  Chan & Crosnier, 1997 
DISTRIHUI'ION. - Marquesas (Fatu Hiva. Nuku Hiva, Tahuata) -Deep (190-230m). 
REFERENCES. - Plesionika sp. nov. 4 - POUPIN, 1996b: 4. pl. 411 (Fatu Hiva, Tahuata; 190-210m). - 
Plesionika profati Chan & Crosnier, 1997: 219. fig. 18. 38 (Falu Hiva. Nuku Hiva, Tahuata; 190-230m). 
Plesionika reflexa Chace, 1985 
DlsTRlolJirON. - Awlral (Kimarara, Tubuai); tiambier; Marquesas (Fatu Hiva); Society; Tuamotu (Moruroa, Tureia) - 
Deep (360-910nz; mostly 550-700m). 
REFERENCES. - Plesiorrika reflexa - CROSNIER, 1986a: 362 (Tubuai). - POUPIN, 19961): 12, pi. 5a (Gamhicr, 
Fatu Hiva. Mororoa, Rimatma, Tubuai, Turcia; Socicty; 360-910m). - CHAN & CROSNIER, 1997: 194, fig. 3, 24 
(Fato Hiva, Maria, h4omroa. Rimatsra, Tubuai. Tureia; 430-800m). - Plesionika easis - I'OUPIN el (11.. 1990: 16. 
64, fig. 5-7, 14, pl. lh  (French Polyncsia). - I'lesiorrika aff. ensis. - POIIFIN, 1988: 28, fig. 2 (Moruroa) - 
Both, not P. ensis A. Milnc Edwards. 1881 = P. reflexa Chace, 1985. 
Plesiortika rubr ior  Chan & Crosnier, 1991 
DtSrRtliUTlON. - Austral (Kimatara, Kurutu); Society (Maiao. Tahiti); Tuamotu (Moruroa, Nihuru. Takapolo, 
Tenaronga) - Dee11 (120-350nt). 
R E F I ~ : N C E S .  - ? Plesiorrika pacificn - MANACII & CARSIN.  1985: 473 (Moruroa andlor Fangataufa). - 
CKOSNIBR, 1986a: 362 - Both, ? not P. paci/icn Edmonson, 1952 = P.  rubrior (cf. remark). - Plesio~~ika sp.  
nov. 3 - POUPIN el 01.. 1990: 16 (French Polynesia). - I'lesionikn rubrior Chan & Crosnicr. 1991: 433. 
fig. 7a-f, 25-28 (Syn.; Maiao, Moruroa. Nihoru, Kimatara. Kurutu, Tahiti, Takapato, Tenarunga; 120-350m). - 
Pout'tN, 1996h: 12, pl. 5h (Same material). 
KEMARK. - Plesioniko pocificn is mentioned from a lost specimen, caught around Moruroa. Although not verifiable, 
the exact identity of this inaterial would rathcr be P, rirhr.ior. 
Ples ionika  serrlilaevis bate, 1888 
DlSl'RIIiLTtON. - Austral (Tuh~ai)  - Deep (800n1). 
REFERENCES. -Plesionika sentilnevis - C I ~ A N  & CROSNIER. 1997: 213 (Tubuai; 800111). 
Plesionikn sindoi  (Rathbun, 1906) 
DISTNII~UTION. - Austral (Maria, Raevavac, Rurutu); Gamhicr; Marquesas (Fato Uiva, Talulata); Society (Rora Rora); 
Tuamotu (Fangatanfa, Makeom. Moruroa, Tuanake) - Deep (150-800n1, most/)' 300-600111). 
REFERENCI:~ .  - Pldsiortikn aff .  ocelirrr - ? POIII'IN ef a/., 1990: 16 (French Polyncsia). - Plesiorrika 
sindoi - I'OUlWi, 1996b: 12, pl. 5c (Rora Bora, Fangataula, Fatu Hiva, tiantbier, Makcmo, Maria [Austral & 
Tuamotol, Moruroa, Kacvavae, Ruruto, Tahuata; 150-800m). - CIIAN & CROSNIEK, 1997: 216, fig. 17. 36-37 (Bola 
Rora. Fangataula, Fatu Hiva, Gambicr, Makemo. Maria [Austral & Tuamotu], Moruroa, Raevavac, Rurutu, Tunakc; 
3 8 0 - 7 8 0 ~  ). 
Ple s io r~ ika  spinidorsal is  (Ratlibun, 1906) 
DrSsR1nrrlroN. - Austral (Tubuai); C;ambicr, Socicty (Rora Rora, Huahine); Tuamotu (Moruroa) - Deep (300-750mJ. 
K~:rr:uv.~ctis. - Plesionikn spinidorsolis - POUPIN cl a/., 1990: 16 (French Polyncsia). - POTIPIN. 1996b: 12. 
pl. 5 c  ( R o n  Rora, Gainhicr. Huahinc, Moruroa, Tubmi; 300-750in). - CIIAN & CROSNIER, 1997: (Rora Rora, 
Gambier, IIoahine, Moruroa. Tubuai; 540-720m). 
Plesiorrika sp in iyes  Iiate, 1888 
~ISTRIIiISrION. -Society (Maiao, Tahiti) - I h e p  (320-60011~). 
RErERkNCllS. - Plesionikn spiaipes - CROSNII :~ .  19863: 362 (Tahiti). - CIlnN & CROSNII:~, 1991: 422, fig. 
Ib-c, 3c, 21 (Maiao. "Taravao" =Tahiti: 320-6001n). - POIIFIN, 1996h: 12, pl. 5d (Samc material). 
Plesiortika willicrrrrsi Forest, 1964 
DlSl'~ltiIISt0N.;. Austral (Rorotu); Gamhier; Marquesas (Eiao, Tahuata, Ua 1'0"); Society (Huahine, Rclliogshausen. 
Tahiti); Tualnoto (FangaiauCa, Makcmo, Mariltea Sooth, Moruroa) - Deep (3I0-60Ont). 
REFERENCES. -Plesionika aff. williarrrsi - CROSNIEK, 1986a: 373, fig. 4a-g (Tahiti). - PoUPlN el nl., 1990: 
16 (French Polynesia), in part, see under P. mncropodu. - Plesiorrika williamsi - POUPIN. 1996b: 12, pl. 5f 
(Rellinghausen. Eiao, Fangataufa. Gatnbier, Makcaio, Marutca South, Moruroa, Rurutu, Tahiti. Tahuata, Ua Pou; 
310-470m). - CRAN & CROSNIEK, 1997: 209, fig. 15-15, 32-33 (Syn.; Rellingshausen, Eiao, Fangataufa, Makerno, 
Marutea South, Moruroa. Tahiti, Tahuata, Tubuai. Ua Pou; 350.600). 
Figurc 5 - Plesionikn spinidorsalis (Kathbun, 1906) 
(From Ci l~ce ,  1985) 
FAMILY CRANGONlDAE 
Prescnce of the deep crangonids in French Polynesia is attcsted by a single small specimen in bad 
condition (tclson is inissing) in the MNHN collections (Coll. RV Marara, B. RICHEK DE FOKGES, May 
1979, Tubuai, 300m; material not recorded). It is attributed to Philocheras sp. (dct. A. CKOSNIER). This first 
report is confirmcd by collection of sevcral other deep cra~igoiiids around the Marquesas during Lhe recent RV 
Alis Maquesas Expedition, August/Scptcmber 1997 (1. POUPIN field notes). 
Tltalassocaris aff .  crinita (Dana, 1852) 
DIsTRIBuT~~N. - Society? - Bn~i~~~pelagic (IOOn!). 
RI~FERENCES. - Thala,mocaris aff.  crinila - NEW MATERIAL - Coll. E. JossE, dct. J. I'OUPIN (Frencll Polynesia, 
?Society; 100m; material rcturned to ORSTOMiTahiti). 
Thalassinidca are often hiddcn in deep burtnws and are very difficult to collcct. As a rcsult, our 
knowledge of this group is superficial and the species listed below certainly repmerit only a small part of the 
rcal French Polynesian rauna. An example of an unknown specics is give11 by a beautiful axiid photographed 
in Tah i t i a~~  watcrs and illustmtcd i n  ;I local iicwsp;iper ("LC homarct aux pinccs d'or", La Dc!pc^rhe, Tahiti, 
August 31th, 1994, p. 19, photograph D. PAKDON). 
First reports of Thalassinidea from French Polynesia were made by  SENDLER (1923) and EDMONSON 
(1944). Subsequent collectiotis made  by, H.G. MOLLER, researchers sojourning at the C R I O B E  center (B. 
THOMASSIN, J.  tic VAUGELAS), or  prescot author, are r e p r l e d  by THOMASSIN el a[. (1982). DE VAUGELAS 
(1983, 1985), POOKE & SUCI-IANEK (1988). SAKAI (1992). POUPIN (1994), NGOC-HO (1995, 1998), and  
POORE (1997). In AusWal, Gambier, or Marquesas Islands, we still have no record o f  Thalassinidea. 
FAMILY AXllDAE 
E r t t r i c h o c l t e l e s  t u n r n o t u  Ngoc-Wo, 1 9 9 8  
DISTRIBIII'ION. - Tuamotu (Momma) - Deep (140h-1). 
II~1:t:Kt:NCI:S. Ilufrichocheles tuzlmolu Ngoc-Ho, 1998: in press (Moruroa; 140111). 
RI:MAKK. -According to NGOC-MO (personnal conm~~nication) this species would better be placed in the genus 
Pornxiolxis (sce KemI .n ,  1996). This point will be discussed in her fuubre publications. 
P n r a s c y t o l e p t t t s  t r i d e n s  ( R a t h b u n ,  1906)  
DrsrmltlrrrtoN. -- Socicly (Rora Bora. Moorea); Tuamotu (Makalca). 
RI:I;I:R~;NCES. - Pnr.axi,rs triderrs - S E N D I . ~ ,  1923: 44 (Maketea). - I'arascytoleplus fridens - SAKAI & D E  
SAINTLAIJRIWS, 1989: 40 (New combination). - SAKAI. 1992: 212 ("Vaitapu" = Bora Bora, "Maharepa & Afareaitu" 
= Moorca; coral and dead coral, 0.5-2m). 
P a ~ a x i o p s i s  b r o c k i  (De Man, 1888)  
DlsrRit3unON. - Society (Rora Rora); Toamoti~ (Fangataufa). 
Rt:l~liRliN~~is. -Ilutrichncheles hrocki - SAKAI. 1992: 216 ("Vaitapc" = Rora Rora; in coral 0.5-lm). - NFOC- 
1-10, 1998: in press (Faiigataufa; outcr reef). --- I'araxinpsi,~ brocki - KENSLEY, 1996 (New combination). 
FAMILY CALLIANASSrDAE 
C/rerar~zrts a r t i c u l a t u s  ( R a t h b u n ,  1906)  - N e w  combina t ion  
DlssKIn~IsIoN. - Tunmotu (Mataiva) - Dl~rrouw. 
Ra1:ERENcEs. - Callickirus nrliculafus - VAIJGEL.AS, 1985: 316 (Mataiva). 
REMARK. -MANNING & FI:I.DI:K (1986) do not includc K~sn~rrirJ 's species in their definition of the genus Callichirrrs. 
in MNHN collections, material from thc Tunmoti~ is rcprescnted by one male (c.1. 11.5 mm, MNHN Th1232). Its 
examination stlows that: a) Maxillipedal 3 dactylus is digitifarm; b) Plcopods 3-5 have digitiform appendices 
intcrnac; c) Maxillipcd 3 is without exopod. In I'OORil (1994: 101), this key out to the genus Chernn~rrs. We therefore 
propose here a new combination for this spfcies. 
Figure 6 - Parascyroleptirs fridens (Katlibun, 1906) 
(From SAKAI & IIr; SAINS  LAUREN^. 1989) 
Coral l ianassa  collaroy (Poore & Griffin, 1979) 
DISTRIB~ION. - Society (Moorea, Tahiti). 
REFERENCES. -Corallianassa collaroy - SAKAI, 1992: 212. fig. la-d (Moorea, Tahiti). 
Glypturus a rma tus  (A. Milne Edwards,  1870) 
D I S T ~ U T I O N .  - Society (Moorea, Tahiti); Tuamotu (Mataiva, Takapoto, Tikehau) - Burrower. 
REFERENCES. -Callichirus arnrafus - VAUCELAS. 1983: 23 (Mataiva. Moorea. Tahiti); 1985: 314 (Mataiva). - 
DELESALE, 1985: 288, 295, 299 (Mataiva). - HARMELIN-VIVIEN, 1985: 251 (Tikehau). - SALVAT & RICHARD, 
1985: 353 (Takapoto). - VAUGELAS el 01.. 1986: 204, fig. 1-3 (Mataiva, Moorea, Tahiti). - Collianassn 
armnfa - MANNING, 1987: 390 (Synonymize this species with Glypturrrs acanlhochirus). - Glypturus nrnmtrrs - 
POORE & SUCIIANEK, 1988: 197, 201 (Mataiva, VAUGELAS' material, with this remark "We do not agree with 
MANNING (1987) who believed that G. arntalus is a junior synonym of G. aconlhochirss"). 
Neocall ichirus taiaro Ngoc-Ho, 1995 
D I S T R ~ ~ I O N .  - Tuan~otu (Taiaro) - Barrower. 
REFERENCES. - NeocnNichirus taiaro Ngoc-Ho, 1995: 21 1, fig. 1-2 (Taiaro) 
FAMILY CALLIANIDEIDAE 
Callinnidea typa H. Milne Edwards,  1837 
DIS~RIBWION. -Tuamotu (Moruroa, Taiaro) - Birrrower. 
REFERENCES. -Callio~rideu typo - EDMONSON, 1944: 38, fig. 2 (Tahiti). - POUPIN, 1994: 7, fig. 3, pl. l b  
(Taiaro). - POORE, 1997: 349, fig. 1-3, 4a-b (Taiaro) - New MATERIAI. - Coll. & del. J. I'OUI)IN, Octobcr 1995 
(Moruroa; sandy area, under rocks). 
Thornassinia inoorea Poore,  1997 
DISTRIBUTION. - Society (Moorea). 
REFERENCES. - Tkonrassinia sp. - TROMASSIN el al., 1982: 394 (Moorea). - Thonrassirria nrnorca Poore, 
1997: 416, fig. 38 (Moorea). 
RF.MARK. - Two others Thomassiniidae, attributed by POORE (1997: 408) to "? Mictaxius sp.", were also collccted 
with this new species. The spccimcns are in poor condition and POORE writes "While the species is represented by 
such incolnplete speci~nens, its generic placelnenl is uncertain". 
ORDER STOMATOPODA 
Collections of Stoinatopoda roughly follow those sccordcd in the Caridea section with the addition ol 
the following Expeditions: the U.S.S. Wachrrsetr Expedition, Marquesas 1884, coll. Dr. William H. JONES, 
US .  Navy (BIGELOW, 1931), the 1899-1900Albatross Voyage (MANNING, 1978c), the 1901-1902 American 
Alvin Scale cluise (EDMONSON, 1921). and the 1957 SMITHSONIAN BREDIN Expcdition (MANNING, 1978~).  
Material from these cruises is deposited at USNM, Washington, or BPBM, Honolulu. 
Surprisingly, two species are deposited at the India11 Museum, Calcutta, Pscudosquillisn~a oculala 
(Brull6, 1837) and Gorzodactylus platysoma Wood Mason, 1895 (typc specics) (KEMP, 1913; GOSH & 
MANNING, 1988). This material was donated to the Indian Museum by the OTAGO Museum, before 1895 
(MANNING pcrsonal communication). OTAGO Muscum is the Muscum of Olago province, New Zealand, and 
was founded in 1868 (HOLTHUIS, pcrsollal communication). 
Individual collectors of Stomatopoda include, J.M. CLEMENTS (1927, Society), priest S i m h  
DELMAS (1927, Marqucsas), G. PINCHOT (1929, Tuamotu), G .  RANSON (1952, Society), J. RANDALL 
(1957. Marquesas), R. WATKINS (1962, Socicty), and A. MICHEL (1969, Marquesas). People sojour~ting at 
Moorea research ccotcrs, CRIOBE and/or BERKELEY station, have also collected stomatopods: 0. NAIM, 
M. MONTEFORTE, B. THOMASSIN, M. PEYROT-CLAUSADE and R. CALDWELL (cf. MANNING, 1978a). 
This material is deposited at the BPBM, Honolulu, USNM, Washington, or SME, Endoume. 
Deep collcctiot~s by the RV Mar-ara include the big indosquilla rnanihiriei Inglc & Merrctt, 1971 
(illushated it1 POUPIN, 1996b: 42, pl. 20 f-g) and two littoral to sublittoral spccies, hcrc reported for tile first 
tirnc: Odor~roclactylus brevirostris (Micrs, 1884) arid Orato.squilla cal~rnlr~ia (Townsley, 1953). From 
ORSTOM Tahiti rcscarch centcr (coll. P. FROUIN) and unpublished material in MNHN and USNM (dct. R.B. 
MANNING), thrce othcr species are also added to the rcgional fauna: A h a  alha (Bigclow, 1893), Busqrtilla 
quadraticauda (Fukuda, 191 I), and Gonodac/y/u.r child Manning, 1971. 
Matcrial collected by H.G. MOLI.ER in 1988 at Bora Bora and Moorea, Socicty Islands, is properly 
registcrcd in MNHN (Sto 1658-1674) bul was not rctrievcd. It could he from thesc field collections that 
Gorrodactylus platysorna Wood-Mason, 1895 and Kaou1,serertea o r n a ~ a  (Micrs, 1880) are listctl from Moorea, 
to our knowledge for the first time, in MOLLEK (1994)'s world bibliography of Stomatopoda. 
FAMILY INDOSQUILLDAE 
Indosqrtilla ttrarrihirrei Ingle & Merrett ,  1971 
D ~ s r ~ r s l m o ~ .  - Tuamotu (Faogataufa) - Deep (55Onij. 
KaPERENcEs. - 1,rdosqeilla nmnihinei - MANNING el nl., 1990: 314 (Biogeography). - MOLLBR, 1994: R 
(Syn.). - MANNING, 1995: 18. 28 (List; new family). - I'OUPIN, 1996: 42, pl. 20 f - g  (Fangatnufa; 550rn). 
FAMILY GONODACIYLIDAE 
Gotrodactylaceus fa lca tus  (I'orskAl, 1775) 
DrsTRlnrrnoN. - Society (Moorea). 
REFERENCES.  -Gonoda~fy les  falcalus - NAIM, 1980: 51, with a ? - Gonodacfylncerrs fa/catris - 
MANNING, 1995: 19, 42-43 (List & key; new combination). 
Gorrodactylaceus randall i  (Manning,  1978) 
DISTRIIIUTION. - Society (Moorea). 
KI;I:ERENCES. - Gomdacfyl~s randalli Manning. 1978b: 9, fig. 6, 12 (Holotype from Moorea). - MOLLEK, 
1994: 41 (Syn.). - Conodnctylacerrs rarrdalli - M A N N I N G ,  1995: 19, 42, 43, 47 (List & key; new 
combination). 
Gonodaclylellris espittosus (Borratlaile, 1898) 
DrsrRterirroN. - Marqucsas (Fatu Hiva); Society (Bora Bora, Moorca, Tahiti); Tunmotu (Fakaravn. Mataiva). 
R~FERENCI~S .  - Gortodacty1rr.s espirtosrrs - MANNING, 19678: 21, fig. 8 (Bora Bora, Fakarava). - MONTEFORTI:, 
1984: 173, aluicn 1, tab. a (Mataiva, Moorea. Tahiti); 1987: 8 (Moorea). - MOr.1.m 1994: 30 (Syo.). -- 
Corrodactylellrrs espinosris - MANKING, 1995: 19, 56, 57 (List & key; new co~nbination) - NEW ~ i \ ? . t i R t ~ L  - 
February 1996, Coil. J. POIJI)IK, det. K.B. MANNING (Falu Hiva MNHN Sto 1960; i n  basc o l  a coral Pocillol>orn). 
Gonodactylellus incipiens ( L a n c h e s t e r ,  1903) 
DISTRIBUTION. - Society (Moorea. Tahiti); Tuamotu (Fakarava). 
REFERENCES. - Gonodacty l~~s  incipiens - MANNING, 1967a: 18, fig. 7 (Fakarava). - MOLLER. 1994: 34 
(Syn.). - Gonodactylellus incipiens - MANNING,  1995: 19, 56, 57. 63, fig. 24. 25b (List & key; new 
combination) - NEW MATERIAL - MNHN Sto 1165-1166, det. R.B. MANNING (Moorea. Tahiti). 
Gonodactylinus viridis (Serene,  1954) 
DISTRIBLWtON. - Society (Moorea, Tahiti); Tuamotu (Mataiva). 
REFERENCES. - Gonodactylus viridis - MONTEFORTE. 1984: 173, annex 1, tab. a (Mataiva, Moorea, Tahiti); 
1987: 8 (Moorea). - DELESALLE, 1985: 305 (Mataiva). - MOLLER, 1994: 44 (Syn.). - Gonodactylinus 
viridis - MANNING, 1995: 19. 66, pl. 4, fig. 8c,d, 9c, 10e. l l c ,  25a (List: new combination). 
Gonodactylus childi M a n n i n g ,  1971 
DISTRIBUTION. - Society (Moorea). 
REFERENCES. - Gonodactylus childi - NEW MATERIAL - MNHN Sto 1170, det. R.B. MANNING (Moorea). 
REMARK. - According to MANNING (in preparation), the original account of childi included two species in two 
different genera. The holotype is a dwarf species. Gonodacrylus child senso stricto, and tho paratypes are a new 
Gonodacrylellus. French polynesian specimens belong to G. childi sensu stricto. 
Gonodactylus chiragra ( F a b r i c i u s ,  1781) 
DlSTRmWrtON. - Gambier (Mangareva); Marquesas (Nuku Hiva); Society (Moorea. Tahiti); Tuan~otu (Hao. Kaukura, 
Fakahina, Makatea). 
REFERENCES. - Got todac t y l~~  chiragra - HELLER, 1865: 126 (Tahiti). - NOBILI. 1907: 410 ("Ohura-Hao" = 
Hao, Kaukura, Fakahina, "motu Marakuraku"=?). - SENDLER, 1923: 47 (Makatea). - EDMONSON, 1921: 300 
(Marquesas). - BOONS. 1934: 11, pl. 2, fig. a, part (Nuku Hiva, Tahiti; not Bora Bora and Raiatea, see MANNING, 
1967c: 103). - SEuRAT, 1934: 61 ("Rikitea" = Mangareva. Hao, Fakahina, Kaukura). - MANNING, 1967c: 102 
(Marquesas and Society islands; revision of BOONE'S material but without precision of i s l a ~ ~ d s  concerned and: "these 
specimens were not all examined"); 1995: 20, 69, 71, pl. 5-8, fig. 8e.f, 9a.b. 10a, l l a ,  27a, 28-30 (List & key). - 
NAIM, 1980: 51 (Moorea; in algae Nalinteda oprtria). - MOl.I.l:R, 1994: 27 (Syn.) - NEW MATERIAL - Coll. C. HILY, 
dct. K. MOOSA (Tahiti MNHN Sto 1959). 
REMARK.  - Exccpt for BOONE'S material, revised by MANNING (1967c), most of the old collections sl~ould be 
checked. "A suite of species, including virtually all members of Gonodacrylus, has been confused with G .  chiragra at 
one time or another" (MANNING, 1995: 74). 
Gonodactylus platysotna W o o d - M a s o n ,  1895 
DISTRIBUITON. - Marqoesas; Society (Bora Bora. Moorea, Raiatea); Tuamotu (Raroia). 
REFERENCES. - Gonodactylus platysoma Wood-Mason, 1895: 11, pl. 3, fig. 1-9 (Society Islands). - 
Gonodactylus chiragra var. acufrrs Lanchester - EDMONSON, 1921: 300 (Marqoesas). - Gonodactylns 
chiragra var. tunlidus - BOONE, 1934: 11, pl. 1, 2b, part (Bora Bora. Raiatea; see MANNING.  1 9 6 7 ~ ) .  - 
Gonodactylus chirngra var. platysonra - KEMP, 1913: 162 (Society Islands; WOOI>-MASON'S material). - 
BIGELOW: 1931: 119 (Bora Bora). - HOLTHUIS, 1953a: 62 (Raroia). - MORRISON, 1954: 13 (Raroia). - 
Gorrodactylrrs plafysonta - MANNING, 1967c: 103 (Bora Bora, Raiatea; BOONE'S material); 1995: 20, 69, 75, 
pl.9. 10, fig. 9d, lob, l l b ,  27b, 31 (List & key; "type locality Socicty Islands, 17?00'S, 150?00'W"). - GHOSH & 
MANNING. 1988: 654 (Sclectio~l of a lectotype from Society Islands in the collections of Zoological Surey of India, 
Calcutta). - MULLER, 1994: 40 (Syn.; with "Moorea" in the Distribution). 
FAMILY ODONTODACMIDAE 
Odontodactylus brevirostris (Miers ,  1884) 
DlsrRlnunON. - Marqucsas (Fatu Hiva) - Sublirroral(4911z). 
RI:FERENCES. - Odor~fodactylus brevirostris - MANNING, 1967b: 22, fig. 6-8 (Revision of the gcnus); 1995: 
20, 82 (List & key). - MOI.I.ER, 1994: 50 (Sytl.) - NEW MATERIAI. - Coll. RV Mnrara, dct. R.B. MANNING (Fatu Hiva 
MNHN Sto 1962; 491n). 
Pseudosquilla c i l i a t a  (Fahricius, 1787) 
DlsrRlBunON. - Gainbier (Mangareva); Marquesas; Society (Tahiti). 
REFERENCES. - 1?seudosq~ilka ciliata - NOBILI, 1907: 410 (Mangareva "Rikitea"). - SEURAT,  1934: 61 
(Mangareva). - EDMONSON. 1921: 288 (Marquesas). - CHABOUIS L. & F., 1954: 92, unurnbered fig. (French 
Polynesia). - MOI.I.ER, 1994: 72 (Syn.). - MANNING, 1995: 20, 111, pl. 20, 21, fig. 59a, 60a,b,e, 61-63 (List & 
syn.) - Naw MA'I'ERIAI. - Coll. P. FROUIN, det. R.B. MANNING ("Mahaeila" =Tahiti MNHN Sto 1958). 
Pse r tdosqu i l l i s rna  o c u l a t a  (BrulI6, 1837) 
DISTRIBUTION. -Society. 
RSFERENCES. - Pse~~dosqui l ln  oculafa - KEMP, 1913: 102 (Society Islands). - MIILLER, 1994: 76 (Syn.). - 
Psendosquillisnza oculnta - CAPPOIA & MANNING, 1995: 285 (New combination). - MANNING, 1995: 21 (List) 
- Not Psendosqidla ocrdara - HELLEn, 1865: 124 (see undcr Raoulsermea ornara). 
R a o u l s e r e n e a  h ie rog lyp l i i ca  (Manning, 1972) 
DISTRIBUTION. -Society (Tahiti). 
Raor t l se re r iea  o r n a t a  (Miers, 1880) 
DIsTRm~moN. - Society (Moorca. ?Tahiti) - Liltoral to srcblirroral(31m). 
REFERENCES. - Pseudosquilla oculafa - ? HELLER, 1865: 124 (Tahiti) ? non P.  O C U I ~ ~ O  (BrullC, 1837). - 
Pseudosquilla o rna ta  - Mlsins, 1880: 33, pl. 3, fig. 5, 6 (Syn.; ?Tahiti see remark). - SEURAT, 1934: 61 
(HELLER'S inatcrial). - MOLLER, 1994: 77 (Syn.; Moorea in Distribulion; up to 31m). - Ronsl.rerensa nrnafo - 
MANNING, 1995: 21. 118, pl. 22, fig. 59b, hOc.d.f, 64 (List & key; new combination), 
REMARK. - MIERS (1880: 34) writes undcr Psercdosqriilla orwla:  "I believe that the specimens from Tahiti, referred 
by HELLER (1.c.) to P. ocrrlnm, belong to this species, as the rostrum is described as 'vorn abgerundet;' and of the 
terminal post-abdominal segment he observes, 'gewahrt man suf der Oberflache mchrere L#ngsleistchen, und zwar 
eine lange mittlerc und zwei seitliclm."' 
R a o u l s e r e n e a  nsp.  Manning & Caldwell in preparation 
DISTRIBWION. - Society (Moorea); Tuamoto (Mataiva). 
KEFE.RENCES. - Raoeberenea nsp. -MANNING & CALDWEIJ., in preparation: coll. R. C ~ m w e r s . ,  Moorea; coll. M. 
MONTEFORTE, 23/05/1982, Mataiva, outer reef, intertidal, dead wral fragments, MNHN Sto 1964. 
FAMILY TAKUIDAE 
M e s a c t u r u s  f u r c i c a u d a t u s  (Miers, 1880) 
DIsTRrr~rmN. - Tuarnotu (Makatea, Mataiva). 
REFBRENCSS. - Conodncfylrrs f ~ ~ r c i c a n d a f s s  - N o n l u  1907: 410 (Makatca). - SEURfl. 1934: 61 (Makatca). 
- Mesaclnrus  furcicaudafus - MONTEFORTE, 1984: 173, annex 1, tab. a (Mataiva). - MOl.l.en, 1994: 50  
(Syn.). -MANNING. 1995: 21, 119 (List & new family). 
FAMILY CORONIDIDAE 
P a r v i s q u i l l a  r n u l t i t u b e r c u l a t a  (Borradaile, 1898) 
DIsTRlBmroN. - Society (Moorea). 
ReFIIRENcES. - Parvisqeilla n~rilfitrrberculatn - MANNING, 1978a: 16, fig. 8a-f (Moorea); 1995: 22 (List). - 
MOI.LER, 1994: 84 (Syn.). 
Lys iosqui l l ina  sulcata (Manning, 1978) 
D I S T R I B ~ I O N .  -Society (Moorea). 
REFERENCES. - Lysiosquilla sulcatd - STEGER & BENIS-STEGER, 1988: 94 (Moorea). - MOLLER, 1994: 89 
(Syn.). - Lysiosquillina sulcata - MANNING, 1995: 22, 133. 134 (List & key; new combination). 
Lys iosqui l l ina  rnaculata (Fabricius, 1793) 
DISTRIBUTION. -Marquesas (Nuku Hiva); Society (Bora Bora, Moorea, Raiatea. Tahiti); Tuaruotu (Mataiva. Toau) - 
Lirrornl to sublittoral (28nt). 
REFERENCES. - Lysiosquilla maculata - HELLER, 1865: 260, 265 (Tahiti; distribution table only). - BIGELOW, 
1894: 508 ("Tawhae" = Taiohae, Nuku Hiva). - EDMONSON, 1921: 292 (Tahiti). - BOONE, 1934: 21-28, pl. 5-6 
(Tahiti). - MANNING, 1967c: 103 (Tahiti; BOONE'S material). -MANNING, 1978c: 3, fig. 1-3, 9 (Bora Bora, Nuku 
Hiva, Raiatea, Tahiti, Toau). - MONTEFORTE, 1984: 173, annexe I, tab, a (Mataiva). - DELESALLI.:, 1985: 288, 293 
(Mataiva). - MOLLER, 1994: 91 (Syn.). - Lysiosquillina niaculata - MANNING,  1995: 22, 134, fig. 68c, 
70a.b. 71a.b. 72a.b. 74-77. 78a, 80a (List & key: new combination) - NEW MATERIAL - Coll. & det. 3. POUPIN, May 
1997 (Moorea). 
Figure 7 - Lysiosquillina mncrlrrla (Fabricius, 1793) 
(Carapace, anterior appendages, and raptorial claws, from MANNING, 1995) 
Pul losqui l la  l i toral is  (Michel & Manning, 1971) 
DlSTRl8UTlON. - Marquesas (Nuku Hiva); Society (Moorea). 
REFERENCES. - Austrosquilla litoralis Michel & Manning, 1971: 237. fig. 1 (Nuku Hiva). 
litoralis - MANNING, 1978a: 19 (Moorea); 1995: 23 (List). -MOLLER, 1994: 120 (Syn.). 
Pul losqui l la  thornassirri Manning, 1978 
DISTRIBU~ION. -Society (Moorea). 
. . 
REFERENCES. - I'ullosq~rilla thonrassini Manning, 1978a: 20, fig. 9a-e (Moorea); 1995: 23 (List). - M~I.I.ER, 
1994: 121 (Syn.). 
FAMILY SQULLLIDAE 
Aliina alba (Bigelow, 1893) 
DlsTRlBUrlo~. - Society (Moorea). 
REFERENCES. -Alin~a alba - SCH~TTE & MANNING, 1993: 577 (Syn.). - MANNING, 1995: 23 (List) - NEW 
MATERIAL - MANNING pessonal communication, in USNM collections, R. CALDWELL leg. 1980 (Moorea; in rubble, 
1111). 
Busquilla quadraticauda (Fukuda,  1911) 
DISTRIBUTION. -Society (Tahiti). 
REFERENCES. -Busqlliiia quadraticnuda - MULLER, 1994: 135 (Syn.). - MANNING, 1995: 23, 170 (List & key) 
- NEW MATERIAL -Coil. P. FROUIN, det. R.B. MANNING (Tahiti MNHN Sto 1961; a juvenile). 
Miyakea nepa (Latreille, 1828) 
DISTRIBUTION. -Society (Tahiti). 
REFERENC'S. - Squilla nepa - HELLER, 1865: 124 (Tahiti). - SEURAT, 1934: 61 (HELLER'S reference). - 
Oralosquilla nepa - MUI.LER, 1994: 167 (Syn.). - Miyakea nepa -MANNING, 1995: 25, 213, 216, pl. 37, fig. 
130a.b. 131d,e, 132-134 (List & key; new combination). 
REMARK. - MANNING (Personal communication) has examined HELLER'S material from Tahiti in the Vienna Museum. 
It is labeled as taken by the Nosara and is correctly identified. 
Oratosquilla calumnia (Townsley, 1953) 
DISTRIBUI.ION. -Marquesas (Eiao) - Slrblitloral(55n1). 
REFERENCES. - Oratosquilla colunrnia - MULLER, 1994: 161 (Syn.). - MANNING, 1995: 25, 220 (List & key) - 
NEW MATERIAL - Coll. KV Marara, 19/01/91, det. R.B. MANNING (Eiao MNHN Sto 1963; 55m). 
FAMILY PROTOSQUILLIDAE 
Echinosquilla guerinii (White,  1861) 
DlsTRmvrloN. - Marquesas (Nuku Hiva). 
REFERENCES. -Gonodactylus guerinii - GRAVIER, 1928: 337, fig. 1-3 (Nuku Hiva). - BIGELOW, 1931: 139 
(Syn.). - Echinosquilla grrerirzii - M~LI.ER. 1994: 61 (Syn.). - MANNING. 1995: 21 (List). 
A D D I T I O N S  
Additions or modifications to POUPIN (199Ga) checklist of Astacidea, Palinuridea, Anomura and 
Brachyura are given here. They mostly come from revised or unstudied RV Marara collections. One 
Polychelidae m d  two Calappidae are identified from material recently collected in the Marquesas (RV Alis, 
AugustJSeptember, 1997), and three crabs Cryptochiridae are reported from re-examination of MNHN 
collections (det. R.K. KROPP). 
FAMILY ENOPLOMETOPIDAE 
The Enoplomclopus sp. nov. from the Marquesas (POUPIN el al., 1990: 16, pl. 3c; POUPIN, 1996a: 8: 
199Gb: 12, pl. 5h) has been recognised in Taiwanese waters and submitted for description by CHAN & YU 
(1998). 
38 
FAMILY POLYCHELIDAE 
Srereon~asfis phosphorus (Alcock, 1894), a common Indo-Pacific spccies, has been collected during the 
RV Alis Marquesas Expedition (Nuku Hiva, 405m). 
Figure 8 - RV Alis. 28.4 m long. lateral vicw 
Crustacea have becn recently collected by this vessel around the Marquesas, between 20. 
lOOOm (MUSORSTOM 9 Marquesas Expedition, August 18111 - September llth, 1997) 
FAMILY PALINURIDEA 
Palinuridae could now includc, Pariulim albif7ageNum Chan & Chu. 1996. CHAN & CHU (1996: 381) 
mentioned that Pariulirus lorlgipes fenlorisrriga (A. Milne Edwards, 1868), reported in French Polynesia by 
GEORGE (1972). is "without cross-bands on the antcnnular flagella" and could be Pa~iulirus albiflagelli~nz". 
FAMILY DIOGENIDAE 
Six ilcw records of Cnlciftrts arc givcn by POUPIN (1997): Calcirrcis gouti Poupin, 1997: C, haigae 
Wooster, 1984; C. imperialis Whiteleggc, 1901; C. isahellae Ponpio, 1997; C. orchidue Poupin, 1997: and 
C. vachor~i Forest, 1958. Additionally, threc more ncw Cnlcirr~rs have becn submitted for publication 
(POUI'IN & MCLAUGHLIN, in press). 
A large spcciincn of Dardarzus sa~igrrirma,pus Dcgener, in Edmonson 1925 has been collcclcd at Eiao, 
during the RV Alis Marquesas Expcdition. Coloration of this specimeo agrecs very well with the illustration 
givcn in HOOVER (1997; dct. P. MCLAUGHLIN). 
From unstudicd RV Marara material, three species has been identified by P. MCLAUGHIN: Carapagur.us 
ensifer Hcndcrson, 1893 (Eiao; 541n), Nemaropagarus cl. spifiulosensoris McLaughtin & Brock, 1974 
(illustrated in POUI'IN, 1996b: 18, pi. 8f), and Pylopaguropsis lewir~soh?ri McLaughlin & Haig 1988 
(Fangataufa, Moruroa; 110-190111). Morcover, Solirnriopagurus sp. nov. (POUPIN, 1996a: 96) is now 
dcscribcd under Solilario/~agitr~rs r iproholcrs Poupin & McLaughlin, 1996 (illustrated in POUPIN, 19961): 18, 
PI. 88). 
FAMILY PARAPAGURIDAE 
From RV Marara collections (Moruroa, 200m) Oncopagurus o imos  sp. nov. is described by 
LEMAITRE (1998). 
FAMILY GALATHEIDAE 
Two galatheids, Munida distiza Macphcrson, 1994 (Moruroa; 290m) and Munida elegarilissima de 
Man, 1902 (Eiao, 55m), have been recognised during the last RV Marara campaigns (illustrated in POUPIN, 
1996b, pl. lob, c, pl. 10d). 
FAMILY ALBUNEIDAE 
From RV Marara collections Alburlea sp. cf. synznysla (Linnt, 1758) (Eiao, Fatu Hiva; 100m) is 
reported and illustrated in POUPIN (1996b: 26, pl. 12c). 
FAMlLY CALAPPIDAE 
Calappa sp. nov. in POUPIN (1996a: 97; 1996b: 30, pl. 14d) is now described under Calappa sebastie~~i 
Galil (1997a). Thercfore, from RV Alis Marquesas collections, two other Calappidae are tentatively attributed 
here to Calal~pa aff. gallus (Herbst, 1803) (Hiva Oa, 98-100m; st. DW1210) and Mursia aff. s p i ~ i i m a f ~ u s  
Rathbun, 1906(Hatutu, 163-245m; st. DW1287). 
FAMILY PARTHENOPUIAE 
Attention is drawn to Parthertope cof~lrarius (Herbst, 1796) illustrated in POUPIN (1996b: 32, pl. 15d), 
and again collected during the RV Alis Marquesas Expedition. This material seems to be similar to Lanrbrus 
(Rhirtolambrus) l o f t g i s ~ ~ i r ~ i s  M ers, reported and illustrated from Hawaii by TINKER (1965: 88, pl. 32). 
Revision of that material is liecessary for a correct determination. 
FAMILY GONEPLACIDAE 
The lriresius sp. nov. illustrated in POUPIN el a / .  (1990: 19 pl. 3a) and POUPIN (1996b: 34, pi. 16e) is 
proposed lor publication by DAVlE (1998). 
FAMILY XANTHIDAE 
Alainodaeus rluklr Davic, 1997 has been described from RV Marara unstudied collections (Eiao, Nuku 
Hiva, Raevavae; 100-400rn). 
FAMILY TRAPEZUDAE 
MNHN material, collected by P. LABOUTE in the Tuamotu (Tikehau, 50/60m, in Slylaster coral) has 
been described under Hexagonalia laboutei Galil, 1997b. Other uapeziids collected during RV Marara 
campaigns or deposited in MNHN collections have been studied by CASTRO (1997a, b) with the following 
species added to the fauna: Terralia fulva Serkoe, 1984; Tetralia rub,ldactyla Garth 1971; Tetralia vanriirtii 
Galil & Clark, 1988; Trapezia globosa Castro 1997b; and Trapezia lutea Castro, 1997a. Moreover, CASTRO 
has removed three trapeziids from the regional fauna: Tetralia glaberrifna (Herbst, 1790) whose records are 
attributed either to T. cirtcri/~es Paulson, T. f i~ lva  Serhe, or T vanninii Galil & Clark; Trapezia fornwsa 
Smith, 1869 re-identified to Trapezia globosa Castro 1997b; and Trapezia septara Dana, 1852, re-identified to 
Trapezia areolara Dana, 1852. 
FAMILY CKI 
MNHN material from Marutea South, attributed by NOBILI (1907) to Cr)plochirus coralliodyles 
Hellcr, 1861, has been re-examined by KROI'I' (Personal con~muoication). I t  is re-idcnlificd to Li lhoscaps  
paradoxus A. Milnc Edwards (1862) and Syhenomaiapyrifnrnm (Edmonson, 1933). These observations throw 
a doubt on  thc cffcctive prescncc of Cr)plochirfts coralliodyfes Hellcr, 1861 in French Polynesia (cf. rccords 
from Mavokau by NORILI, 1907. and Makatea by SENDLEK, 1923). From RV Marar-a collections (Fatu Hiva, 
Marquesas) KKOPI' has also idetitificd U~i,lo?niella dimo,pha (Henderson, 1906). 
Table 1 - French Polynesian shrimps 
number of spccies by Itifra-Order and Family 
IWKA-ORDER FAME.Y B w m  
D e n d r o b r a n c h i a f a  2 5 
Aristaeidae 3 
Renthesicymidae 3 
Penaeidae 1 2  
Solenoceridae 3 
Sicyoniidae 1 
Luciferidae 2 
Scrgestidac I 
S t e n o p o d i d e a  2 
Stenopodidae 2 
C a r i d e a  1 7 0  
Nematocarcinidac 2 
Atyidac 5 
Oploplioridae 1 0  
Slylodactylidac 1 
Rhyncliocinetidae 3 
Gnathophyllidae 1 
Ancliistioididae 1 
Hymenoceridae 1 
I'alaernonidae 46 
Alplieidae 57  
Hippolytidac 1 0  
I'roccssidae 2 
Pandalidae 29 
Tbalassocarididae 1 
Crangonidae 1 
T h a l a s s i n i d e a  9 
Axiidae 3 
Callianassidae 4 
Callianidcidae 1 
Thomassiniidac 1 
T o t a l  2 0 5  
From this compilation 205 shrimps and 26 Stomatopoda are reported from French Polynesia (Tab. 1 & 
2). Within the shrimps Caridea alone are represented by 170 species (83 % of the shrimps) mainly Alpheidae 
(57 species), Palaemonidae (46 species), and Pandalidae (29 species). 
Table 2 - French Polynesiaa Stomatopoda 
number of species by Family 
Stomatopoda 2 6 
Indosquillidae 1 
Gonodactylidae 8 
Odontodactylidae 1 
I'seudosquillidae 5 
Takuidae 1 
Coronididae 1 
Lysiosquillidae 2 
Nannosquillidae 2 
Squillidae 4 
Protosquillidae 1 
If the previous list of French Polynesian Astacidea, Palinuridea, Anomura, and Brachyura is considered 
(POUPIN, 1996a; with above additions), the total number of Decapoda and Stomatopoda comes to 757 species. 
The number oi species by Infra-Order is summarized in Table 3 where it clearly appears that Brachyura alone 
rcpresents almost 50 % of the species. 
Table 3 - French Polynesian Decapoda and Stomatopoda 
~iumber of species by Infra-Order 
INFRA-ORDER NUMBER OFSPECIES P E K C ~ A G E  (%) 
Dendrobranchiata 25 3 2  - 
Stenopodidea 2 0.3 
Caridea 170 22,5 
Thalassinidea 9 1 . 2  
Astacidea 4 0.5 
Paliouridea 19 2.5 
Anomura 126 16,6 
Brachvura 377 49 ,8  
Stomatopoda 26 3 ,4  
T o t a l  7 5 8  100 % 
Although drawn from more than a century of rcsearch, this report is only a first step in the 
elahoratation of a comprehensive fauna of French Polynesia. Iticrcascd effort in collection is nwded as wcll as 
careful revision of many genera. 
Few groups have been relalivcly wcll studied. They include: Pandalidac (CIIOSNIEK, 1986a,b, 1988b; 
CHAN & CROSNIEK, 1991, 1997); Palinuridea and Cocnobitidac (sce POUPIN, 1996a). Diogcnidae of thc 
genus Calcirrus (POUPIN, 1997; POUI'IN & MCLAUGFILIN, in press): Parapaguridea (L.EMAITKE, 1994, 
1998), deep Galatheidae (MACPHEKSON & DE SAINT LAURENT, 1991), Homolidae (GUINOT & RICHEK DE 
FORGES, 1995); Geryonidae (MANNING, 1992, 1993b); Etisinac and Chlorodiinae (sce POUPIN, 1996a); 
Trapeziidae (ODINETZ, 1981,1984; CASTRO, 1997a, b; GALIL, 1997b); and Ocypodidac (cf. POUPIN, 1996a, 
1997b). 
A lot of groups need further researches. Frcshwatcr Dccapoda havc been intensively collcctcd 
(MAKQUET, 1988, 1991,1993; KEITH & VIGNEUX, 1997) but revision of the Macrobrachiuni is necessary to 
establish the status of one to three new species (cf. under Macrobrachirmz Iarirnanus). 
Alpheidae, rcprcsenting thc hulk of the shrimps, arc almost exclusively known from the collections 
made in 1954 in the Society Islands by A.H BANNEK (BANNEK & BANNEK, 1967). It is rcgrcuable that all 
this material has been destroyed in tlie Hawaii laboratory fire of 1961 (sce Caridea section), thus exclutlit~g any 
revision of the collection and limiting BANNEK & BANNEK (1967) to a documcntcd list written from a draft 
manuscript and field notcs, without illustsations or key. 
Pontoniinac, although reprcsetitcd by scores of spccics, are certainly under-cstimatcd. Without 
particular effort, ficld collcclions realiscd by the present autlior and dctcrmined by A.J. BKUCE havc added 10 
more species to this list, and several world-wide distributcd spccies, such as (;riarho~~lrylloide,s rriineri Schtnitt, 
1933 (Iudo-Pacific and Atlantic), are still not recorded in French Polynesia although it is inconceivable that 
they we  not occuring there. 
The Infra-Order Thalassinidca lias only 9 spccics. This low figure is undouhtctlly the rcsult of the 
burrowing mode of life of these specics, and the resulting difriculty to collect them. Revision of tlic Indo- 
Pacific Upogebiidac (SAKAI, 1982) docs not mention a single Frcnch Polynesian mcmbcr. Axiiclae arc 
represclitcd by only 3 spccics, whcrcas SAKAl & DE SAINTLAUKENT (1989) report 9 species around Hawaii. 
Ncvcrthcless, we know that the number of specics is usually greater in French Polynesia than in Hawaii (cf. 
POUPIN, 1996a: 78). 
A large RV Marara collection of Portunidae is still undcr study by MOOSA & CKOSNIEK and will 
include sevcml news sccords (Brrtsiriia, Porlurircs, arid Thainnrila spp.). 
T o  conclude this brief overview of groups that nced further i@scarcli, Stomatopotla appcar obvioilsly 
utidcr-reprcsentcd. In New Caledonia, itivcsligations in the lagoon conducted by means of a dredge (RICKEK DE 
FORGES, 1991) have boosted the number of Slomatopodn Sroin I0 lo 69 (MOOSA. 1991). French Polyncsian 
lagoons are usually inadeqontc for navigation. However, if tlrcdgcs wcrc opcralcd by nie:iiis of small boats, we 
could expect a significant increase of numbcr of  stoma pod:^, as well as for many Dccapoda (Majitlac, 
Portunidae, Xanthidae ... ). 
A special mention must be givcn here to the rccent MUSORSTOM 9 Marquesas Expcdition, August 
18th - Septembrc I ltli, 1997, RV Alis. During that cruise, dictlgc and 1x:rm-trawl were opcmtctl hctwecn 20- 
1000m, and a large collection of Crustacca was realised. This material, dcpositcd at the MNHN, is still 
unstudied. Howcver, from ficld notes taken by the prescnt author, it appcars that this collection will bring a 
minimum of 60 new records. These include bathypclagic shrimps (Aristeidac, Bcnthcsicytnidnc, Solenoccritlac. 
Sergestidae), Stenopodidca (at least three species), lobsters (Scyllaritlca, Axiidac), C1mgonidca. Dioycnidae (a 
new Ciliopagurus), Parapaguridea, Galatheidea (Manida spp.), Chirostylidae (first record of an Uroplychus 
sp.), Dromiidae, Dorsipidae (first report of the family), Calappidae, Leucosiidae (at least five additional 
species), Majidae, Portunidae, Xanthidae, and Palicidae (first report of the family). This estimation is reported 
on figure 9 where evolulioli of number of  specics within the years is illustrated. Number of Polynesian 
Decapoda and Stomatopoda will thus exceed 800 species. Moreover, it appears that researches conducted after 
1980 (CRIOBE Moolea studies and RV Marara campaigns) have generated 322 new records, i.e. an increase of 
74 O/o ! If this effort was continued in the future, it is to be expected that regional fauna will reach the level of 
a thousand spccies. 
Number 
ofsnecies G 
Figure 9 - Historical of collections of the French Polynesian Decapoda and Stomatopoda 
Number of species, and cumulated oumbcr of specics, rccordcd each 10 years, with 
indication of main episodes in collections 
In the present state of knowledge, biogeographical consideration are rather hazardous for most of the 
groups. The main trend, as generally admitted, would be towards a decreasing biodiversity from Indonesia and 
Australia to French Polynesia. For example, BRUCE (1990: 13) mentioned 168 Australian pontoniins for 
only 36 in French Polynesia. Howcvcr, this difference is probably amplified by imperfection of our data, and 
could be reduced when pontoniids are better studied in French Polynesia. In fact, for genera that have k e n  well 
studied, biodiversity appears only slighly lower, or even gl-eater, than in western Pacific. For examplc 
Plesionika count 21 species in French Polynesia 1)s 29 in western Pacific (CHACE, 1985) or 26 around 
Taiwati (CHAN, personal communication). Hermit crabs of the genus Calcinus were cited as at1 example of 
low diversity in French Polynesia by POUPIN (1996a), with only 10 species vs 23 in Indonesia. However, 
after more collections and revision of the genus (see above) the balance is now 18 vs 23 and biodiversity of 
Polynesian Calcitrus appears even greater than in Australia (17 species in MORGAN, 1991). The same 
observation is verified after more collections and re-examinalioo of the genus Trapezia by CASTRO (1997a, b) 
who records 14 Polynesian species vs only 11 in New Caledonia and CASTRO concludes (1997b: 131): "The 
decrease in species diversity that is expcted as one moves eastward from axas  of high divcrsity in the western 
Pacific is not observed in trapeziids" 
L I T E R A T U R E  C I T E D  
AUAMSON, A.M. 1935. - Non-Marine invertebrate fauna of the Marquesas (exclusive of insects). Occasioiial 
papers of rhe Bernice P. BISHOP Miiscum, Hawaii, 11 (10): 1-39. 
AQUACOP & J .  PATKOIS, 1990. - Aquaculture of edible species in French Polynesia. Advarms in  
Tropical Aquaculrure, Workshop at Tahiti, Freiich Polyiiesia, February 20 - March 4,  1989, 
AQUACOP, IFREMER, Acres du colloqlie 9: 67-71. 
BAGNIS, R. & E. CHRISTIAN, 1983. - Guide sous-nzarin de ihhiti.  Les Cditions du Pacifique: 1-152, 
illuslratetl. 
BANNER, A.H., 19.56. - Conhibutions to the knowledge of the Alpheid shrimp of the Pacific occan. Part 1. 
Collections from the Mariana archipelago. Pacific Science, 10 (3): 318-375, fig. 1-23. 
BANNEK, A.H., 1959. - Ibid. P x t  IV. Various small collections from the cenlral Pacific area, including 
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SAKAI, K. & M. de SAINT LAURENT, 1989. -A check list of Axiidae (Decapoda, Crustacca, Thalassinidea, 
Anomura), with remarks and in addition descriptions of one ncw sublamily, clevcn new genera and 
two ncw spccies. Narriralists, 3: 1-104, fig. 1-25. 
SALVAT, B., l986a. -LC lirroral corulliet~. 111: C. GLEIZAL (ed.) EncyclopCdie tic la PolynCsie, Tome 3, Lc 
monde marin. C. GLEIZALIMultipress: 6-24, illustrC. 
SALVAT, B., 1986b. - Crabes, langoustes, ti'an'e et varo. 111: C. GLEIZAL (ed.) EncyclopCdic de la 
PolynCsie, Tome 3, Le mondc marin. C. GLEI%AL/Multipress: 70-72, illustld. 
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& B. SALVAT (eds). Proceedings of the Fifrh Inter~rarional C o m l  Reef Corigr.ess, Tahiti, 27 May - 
1 June 1985, 1: 323-362, fig. 1-34. 
SCHOTTE, M. & R.B. MANNING, 1993. - Stomatopod crustacea frotn Tobago, Wcst Indies. Proceedings of 
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54 
THOMASSIN, B.A., JOUIN, C., RENAUD-MORNANT, J., RICHARD, G. & B. SALVAT, 1982. - Macrofauna 
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A C K N O W L E D G E M E N T  
I am most graleful to the following people for their help and support during this work. 
Deterniinations published here for the first time have beeii made by A. CROSNIEK (Aristaeidae, 
Penaeidae, Solenoceridae, Sicyoniidae, and Oplophoridae), T. KOMAI (Neinatocarcitiidae), A.J. BRUCE 
(Pontoniinae, and Hippolytidae), P. NOEL (Processidae), P.A. MCLAUGHLIN (Paguridae), R.K. KROI'P 
(Cryptochiridae), and R.B. MANNING (Stomatopoda). F.A. CHACE, T. KIKUCHI, and B. KENSLEY solved 
problems encountered with old records of the Oplophoridae, Sergestidae, and Lucifcridae. Dr. L.B. HOLTHUiS 
kindly gave information about two old institutions, GODEFFROY and OTAGO Museums, a i d  P. F'ROUIN sent 
several interesting shrimps and stotnatopoda collected during his thesis at ORSTOM Tahiti research center. 
I N D E X  
aenrrrlr~nt, Macrobraclrircnr ............................ I 1  
oentrrlrrs 
Pulne,mrr ............................ . ........ 11 
Alpheus 
acanrhonzerus ..................................... 19 
acrtleipes ......... ..... .................. 17 
amirante i  ............................................ 17 
crassin~anus ........................................ 20 
ntitis ................................................. 20 
. . pacrftcus ................... . .................. 21 
parrrcr i t inus ...... . .......................... 20; 21 
paradentipes ....... .. .......................... 21 
paragraci l is  ............. .. ...................... 23 
parv i ros l r i s  ......................................... 22 
anrboinerrsis 
25 
26 
........................ 10 
amirantei. Alpheus .................................... 17 
Amphipalaemon seura ' II 
Ancl~ioponlonia hur 12 
Anchisria 
Ancltistioides corn~ressus ........................... 11 
arereformis. Arhanas ...............  ............. 22 
............................... Arislaeomorphafoliacea 3 
Aristeus 
arnmtus ..... 
arn~atus 
3 
32 
Clypturss ........................................... 32 
Plesiopenaeus ......... ... ...................... 3 
articulalr~s 
Atya 
8 
8 
. . prlrpes ................. ........................... 8 
assrrule. Mucrobrachirm ........................... I I 
australis. Palaenton ........................... .... . . .  I I  
Auslrosqlrilla l iroralis .......................... ....... 36 
Auromare gardineri .................................... 23 
balboe. Pelagopenaeus ................................. 5 
Benrltesicyntlrs 
4 
4 
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . .  4 
4 
bidens. Alplieus ...................................... I8 
bigentnteu. Sergiu ............................ .... . . .  6 
biunguiculatus. Braclrycarprrs ...................... I1 
Brackycarptrs biungrricrrlarus ........................ I1 
brachyceros. Synalpheus ................. . ..... 24 
bradypus. Alpheus .............. . ...... . ........ I7 
brasiliensis. Benrhesicymus .......................... 4 
brevipes. Alpheus .................................... 1 8  
br ocki 
Erctricl~oclreles 
Calliannssa arn~frtn .............. .................. 32 
Call innidea rypa ..................................... 32 
....... .......... calrinrnia. Omrosquillo ......... . . 37 
conaliculul~rs 
............................................ Melicertrrs 4 
.......................................... 8 
............................... 
cl!acei 
Chm-risnrris rirticrrlut 
................................ childi. Gonodoctylus 34 
chiragra. Gonodactylrrs ............................. 34 
Cklorocrir l is joctans ................................... 26 
ci l intn, Pserrdosqrrilla .................................. 35 
relicrrIa11is ....................................... 10 
collaroy, Corallianass~r .............................. 32 
collrinlianrrs, Alplrerrs ................................. I8 
conznrensalis 
Melopcnaeopsis ..................................... 4 
Stegopontonirr .................................... I7 
con~pressus, Anckislioides .......................... 11 
Conchodj'les 
nrcleagri~ae .......................... ........... 13 
tridncna 13 
concinnrrs 
Lemder . . . . . . . . . . . . .  ......... 
.................... ..... ..... 39 
Cortrlliocuris 
nrrdirostris ........................................ 13 
. . .  
vrrrdrs ........................... .... . . .  I 4  
coralliodytes . Crvploclrirrrs ......................... 40 
crassinranus. Alpheris .............................. 20 
crenatiis. Benlhesicynzrrs ............................ 4 
crinita. Tlrulassocaris ...... . .... . ............. 30 
crockeri. Alplieus .............. .. ................... I8  
Cryptoclrirus cornlliodytes .......................... 40 
carvala. Plesionika ................................. 27 
crirvirostris . Hnliporris ................ .. ............ 5 
cyhereus. Alpllerrs .................................... I 8  
Kclrinosqrcilla geerinii ................................ 37 
edanrensis. Alplieus ................................... I 9  
edwardsii 
Alpheus .......................................... ... 19 
eleganlissima. Munida .............. . ............ 39 
Enoplo~~~etopus sp . nov .............................. 37 
ensifer 
......... .......................... Calapagurus .. 38 
Heterocarprrs .................................. 26; 27 
ensiferrrs. Ligur ........................ .. ........... 24 
ensifrons. Periclimenes ......................  .... 16 
emis. Plesionika .................................... 29 
equalis. Alplzeopsis ................................. 17 
erythrocycl~rs. Plesionikn .......................... 27 
espinosus 
Goriodactylellus .... ...... ................ 33 
Gonodacrylus ...................................... 33 
euchiroides. Alpheus .................................. 22 
Eupalaenion dispar ................................... I1 
Eutrichocl~eles 
31 
31 
eximia. Acanttrephym .................................. 9 
Exoclin~enella denticulata ............................ 14 
exrrens. Benfheocnris .................... ............ 9 
falcalus 
Gonoductylaceus .................................. 33 
Gonodactylus .................................... 33 
Jenloristriga. Panrrlirus .............................. 38 
Fennera chacei .................  ................. I4 
fenneri. Plesionika .................................... 28 
Fenneropenaerrs indicirs ................................ 4 
fermerinkii. Sergesres ................... . ....... 6 
flavicar~da. Plesionika ................................ 28 
foliacea. Aristaeonzorpha .............................. 3 
Jorniosa . Trapezia 39 
frontalis. Alpheus 19 
f u h n .  Z'errnli 39 
Funcl~ulia 
taaningi ............................................... 4 
uillost 4 
firrcicmrrloius 
Gonodacrylris .................................... 35 
Mesocturrrs 35 
.................. ............. gallris. Calappn ... 39 
gardineri, Autonxrle ................................... 23 
gerlrrclrei. Philnrius ................................. 17 
gibberosa. /lippolt.ye ...................... .. ....... 25 
gibbcrosris, Saron ...................... . ....... .... 25 
gibbosr~s, Nippolyte .................................. 25 
glribcrrinlu. Tetrnlin ................................... 39
globosa, Trapezin ................. .. ............. 39 
Glypterus 
Gnrnkophylloidcs mineri ............................ 42 
Gnarhop/tyllr~m 
americonunl ..................................... 10 
10 
10 
33 
33 
33 
34 
rr . sp ........................ .. .................. 34 
Gonodncrylinus viridis ................... .... .. 34 
Gonodacrylus 
acrrtus .............................................. 34 
. . 
chrldi ................................................ 34
fitrcicnudar~rs ................................ 35 
.. .. 34 ........  .......... ................ 
tsmidus ................... .... . . . . . . . . . . . . . . .  34 
gouti, Cnl ................................. ...... 
gracilipes . AIp11e1rs ............... .. .............. 19 
gracilirostris, Oplophorus ........................... 9 
grocilis 
Alpheus 19 
Arlrirnos ............................................. 23
Nenzorocarcinus ... ...... . ....................... 8
gruminea, Coralliocoris .............................. I3 
grundis, Periclinzenes ................................ 16 
guerinii 
...................................... Ecltinosqrrilla 37 
...................................... Gonodacryles 37 
tladropcnrrerrs lucosii .................................... 5 
hnlli. Ilynrenopermeus 
llorpiliopsis 
benuprcsii ............................. 
.................... keroni. Synnlpireus .......... . 24 
llelerocnrprrs 
anracrrl ............. 
dorsrr 
ensife 
Iaevigatus ............  ........................... 27 
longiroslris ........................................ 27
. . pmw.~pino ..................................... 26; 27 
sibogae .................. ........................ 27 
.......................... Hererogenys microphthalma 9 
idioc1rele.s. Alplretr. s ................................... 20
inrperinlis . Calcin, rs ............ ... ........ ...... 38 
inneqeinznna. Anchislia ............................ 16 
incipierrs 
............... Gonodaclylcllrrs ... . . . . .  34 
...................................... Gonodaclylus 34 
indicrrs 
Arlmnas ............................................. 23 
Fewnoopenoerrs ..................................... 4 
Penoeris ............... ... . . . . . . . . . . . . . . . . . .  4 
.................... Indosyr~i l ln  nranihinci ........... 33 
inern~is. Alplreri 18 
insignis. Alphcir 18 
........... . ................. Intesirrs sp nov .... 39 
....................... investignloris. Bcnrhesicynws 4 
........ isnbellne. Cnlcinu.~ ..................... . 38 
.... 
jaclons, Clr lnrocsrt is ..... . ....... . ........ . .  26 
Jrlnicel l~ spinicnrida .. .. ................................. 9 
joponico, Jocaste ...................................... 14 
. Ioctlsre 
japo 14 
Iircin 15 
Inn~elliro.sfris. Cornlliocar is ..... .. ........... 15 
lar 
Macrobrachirrnr .................................... I 1  
Pnlnenro II 
Irzln. Palnenronello .................... .. ... . . . . . . .  15 
lalinrnniis 
.................................... Macrobraclchiirnr 12 
lewirzsolmi . Pylopogrrropsis ........................ 38 
l ihmlus. Stylodaclylus ............... .. ........... 10 
.......................................... Ligur ensiferirs 24 
Lithoscaprrs paradorrrs .......... . ................. 40 
Aa.slrosqrril1~1 ....... .. ......................... 36 
Pril losqrril la ....................................... 36 
lohidens. Alplreris ..................................... 20 
longidens. Acnnrhephyra .............................. 9
.................................. longipes. Pansl i r~rs 38 
longiroslris. fIererocarprcs ........................... 27 
long ispinis 
......................... ............. Lamhrus  39 
Rlrinolambrvs ........ ... ................... 39 
loil ini. Alphess ....................... ... . . . . . .  20 
................................... Iricasii. Iladropenrrerrs 5 
Lrrcfer 
6 
6 
5 
6 
6 
Irrcinn. ocas e 15 
39 
lrirescens 
16 
I 6  
Lysiosqirilln 
niacrclnra .......................................... 36 
srilcarn 36 
Lysiasqrril 
nrncirlora ............ . ............................ 36 
nzuhalrissae. Ari.slerr.s ................... . .......... 3 
Macrobrtlclrircm 
aemi~ l r rn~  ............ .. ............................. I 1  
rosenbergii ......... .. ......................... 12 
sp., . s p  nov ........................................ 12 
moo-ochirrrs. Alplreas ................ .. ............ 22 
Ly.siosqrrillina ..................................... 36 
................................. nzadreporne. Periclinrenes ... 16 oahirensis. Alplzerrs 20 ........... ......... 
............................ nzahei. Periclintencs I6 obesomnnes. Alpkerrs 20 ................................... 
............ ................... n~nldiwrrsis ocellss. Plr~sionikn . 29 
ocrrlnln 
......................... ....... Pserrdosquilln .. 35 
n1u1leodigilrr.s. Alpheirs .. 20 Pserrdosyrrillisnr 35 ............... ........... 
nmnilririei. Irrdosquilia ............................ 33 Odonrodacrylrrs brel 34 
. . . . . . . . . . .  ntarntoraru.~. Saron .. 25 oimos. Oncopngirras 39 ....................  .......... 
................................... niarqsesas. Meiapenaeopsis 5 O~tcopagrrrris oinlos 39 ........................... 
.................... marlin Plesionikn 28 Onycocoris quodrnlopirriial,,lo 15 . .....................................
marrrlcensis. Arere 23 Oplopl~orrrs .....................................
......................................... n~edirrs. Alplreus ....................................... 18 gmciliroslris 9 
meleogrinne. Conckodgres ........................... 13 
Melicerl, rr c~molicrrlaircs ................................ 4 
Merliippolyre knrriaensis .............  ............ 25 
Mesacrrrrrrs Jrtrcicarrdairrs ............................ 35 
Merdphcus prirrrgroci1i.s .... . .. .. ............... 23 
Merupenaeopsis 
con~n~en.snli.s 4 orn(rr(1 ......................................... 
..................... ........... 4 Pseirdosqrrilln .. 35 
......... .................... 4 Rnorrlserenea ... 35 
.............. . . . . . . . . . . . . .  5 Orlninnnirr allrrairdi  8 
.................................... 5 ovalicep. Alpheirs 21 
............................................... wlu l ina  5 
microphllcalma. Nererogenys ......................... 9 
nricroscnphis. Alpheus ............................... 20 
mirrsi. Anchislrrs ....................... ........ . .  13
miner;. Gnarlrophylloides ........................... 42 
nziiis. Alplieu 20 
Miyukea nepa 37 
rnoann. Processa ....................................... 26 
monodon. Perrrrerrs ................ .- ............... 4; 5 
moorea. Titonmssinia ........ . ....... .. . . . . . . .  32 
nznltirribercrrlaia. Par~~isyrril la ....................... 35 
nnrvol. Plesionika ..................................... 28 
Nemnlocnrcinrrs 
Neocnlliclrirrrs lnimo .................................. 32 
nepa 
................ ....................... Miyakcn . 37 
Orrrfosqrcilla ........................... . ....... 37 
Nikoides dam 
............................. 24 
nadirosrris . Co,-n1liocnri.s ............................ 13 
nekrr. A1ainodaeir.s ..................................... 39 
11~ic1zyckira.s. Alplreris ................. ... 2I 
paci/rcn. Plesionika ....................... ...... 29 
pacificus 
Prms1ire.s 
rilbiJlngellrrnr ........ ................. . . . . . .  38 
feniorislriga ........................................ 38
pnrndenlipcs. Alpheris ................................ 21
parodosrr.~. Lirhoscapics ............................... 40 
23 
Meln1plrerr.s ......................................... 23 
Pornrrchis1rr.s 
Pm-osc)'~oleples tridens ............................... 31 
Prrr(r.vioi,si.s brocki .................................... 31 
Parnriirs o'idcns ........................................ 31 
Parllrcnope conlrarius ................ . ........... 39 
paroirostris. Alp1rerr.s ................................. 22 
paruispinn. Ilelerocarpss ....... .. ............. 26; 27 
Pnr~isgvi l ln nie~i~r~berc~rlo~lm ......................... 35 
pnsclralis, n o r  ..................................... 26 
poyeni, Plesioniko ................................... 28 
Pelagopcnnerrs haiboe .................................. 5 
p e l l ~ ~ c i d o ,  Sgsrellnspis ....... . ................. 10 
den1icr11nrrr.s ....................................... 14 
peliIlrorrorsi, Prriclin~enes ........ .............. 14 
Plrilarirrs gerlnchei .............. . ................ 17 
Plii loclierns sp ....................... ... ......... 30 
plrospliorvs . Slo.eonras1i.s ........................... 38 
picra 
Il jnrrnoceru ........................................ II 
Plesionikn ..................... ............. 27; 28 
pl(ilgrrngsicula,irs ...... .. ....... . . . . . . . . . . . .  22 
Plcsioniko 
narvol  ........................... ... ........... 28 
. . 
I ~ I S ~ ~ O T I L Y  ........................................ 28 
iv i l l inmsi ............ . .......................... 29 
Plesiopennerrs mrnnlss .................................. 3 
Ponlonin Irrrrii .............. ..... . . . . . . . . . . . . .  I2 
Ponfonidcs rrnciger .................................... I7 
42 
Periclinienes ....................................... 16 
28 
probubiiis. Alpheus ................... .. ......... 18 
Processo nlounu ........................................ 26 
prolnli. Plerioniko .................................... 29 
Pscrrdosqirillo 
. . 
cr l io lo .............................................. 35 
Iiierogiyplricn ...... ... .................... 35 
oculul 35 
ornrrlo ................. .. ........ .... .... . . . . . . . . . . .  35 
Pseirdosqrriliisn, n ocslirm ..............  ........... 35 
P~rllosqeilla 
l i iorii l is . . . .  36 
rlionrossini 36 
Pylopug rrropsis ......................... 38 
p)s@orn~n. Sphenonroiu .............................. 40 
......... rsbridaciyl<~. 7'errnlia ...................  39 
rirbrior. Plesionikn .................................... 29 
Snlnroneus 
.serrrrlidigirrrs .... ...................... 23 
sihogne .................... ...... .... . . . . . . . . .  23 
sebrrstieni. Cnlnppn ................................... 39 
senril(revis. Plesionikn ........................... 29 
. sepmrtr. Trapezia ....................................... 39 
sermei. Pornncliislus ...... ..- ......... 15 
Sergin higemmen . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .  6 
sermtif,.ons. Plesionikn .................... . .... 28 
. . 
swrorirosrris. Cnrrdrrrn .......................... . 8 
soumi 
18 
I 1  
sibognc 
/lereroca,p rrs ..................................... 27 
sindoi. Plesionikn .......... . ................... 29 
sizort. Alpl,  eu 17 
.............. ................ sniiri! i. Acnnrheplryro .. 9 
Solirnriol~oglgsrr~s 1,- iproholss ........................ 38 
. soror. Pcriclimcncs ...... .. ............... . 17 
................................ ~ ' e c l i ~ b i l i s .  Palnen~on I 1  
Spl~cnonmin p\irfmnm ............................... 40 
................. ............... spirricosdn. Jnnicclln .. 9 
spinidorso1i.s. Plesiotrika ............................ 29 
spinifern. Per iclinienelln ... .. .................... I5  
,sp;triferrr.s 
lfnrpii i j 16 
Periclin~enes .................................... 16 
........... . spinigern. I l rrr l , i l iol~si.  s .......... 14 
.spino.srrs . Oplopl~orris .................................. P 
.s/>i,irriosen.sori.s . Nen,aiopngera.r ................... 38 
Syirilln rrepa ............................................. 37 
Sic,-conzn.s~i.s p1iosl~l1ore.s . . . . . . . . . . . . . . . . . . . . . . . . . . .  38 
srrnbas. Den1kesicyn11i.s ................................ 4 
s1renait.s. Alpiter<.s. ...... . 22 . ........ . ........... 
slreplodudylrrs. Synnlpl~eus ...................... 24 
s1yliro.slris. Lilopenaerrs ............. .. ............ 4 
Stylodncrgl~rs librulrrs .............................. 10 
.......................... .slylorosrrurn. Acanlhephyru 9 
...................... slyloro.slrnlis. Aconlirephyyra .... 9 
sublrrcnnns. Alpheus ................................ 22 
srilcnlo 
Lysiosyrrill(r ........... . ......................... 36 
Lys io  sqaillina ..................................... 36 
sirlct,rr~s. Alplreris ...................................... 22 
. srrperbir. Coralliocnris ................................ I 4  
symnysra. Albrrneu .................................... 39 
S g n ~ p a g u ~ ~  dofleini .................................. 16 
Synalpl~errs 
brackyceros ........................................ 24 
running;. Frrncltnlin .............. .. ................ 4 
rnirirensis 
Alyoidu ...........  .... ... ..................... 8 
larawensis. Me1rrpenneo)~sis ........................... 5 
lenu.pes. Pnla~n~onella ............... .... . . . . . .  I 5  
Telrulia 
7lrtrlrrssocm-is crirrirtr ...........  ..... . . . . . . . . . . . .  30 
Ihomossirri. Pdlosyrrilln ............................. 36 
l'hon~nssinin 
32 
32 
onrboinensis ....................................... 26 
nrnldivensis ........................................ 26 
pnsclinlis ................. . ...... .. . . . . . . . . . . . .  26 
77rorinn nmldiaensis .................................. 26 
Trapmio 
seplnrn ............... ............................. 39 
widens 
Gnalltophylluni . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .  10 
................................. Parascylolep~us 31 
Paraxius ............. . .......................... 31 
rridenlnlirs. Periclin~enaeirs .......................... 15 
lriprobolrrs. Solilnriol~ngrrrrrs ....................... 38 
Irispinosus, Plesionika ............................. 28 
................................... Iryphoprrs, Alpheas 17 
trianrorae, Periclinlenaeus ............................ 15 
lrcanrollr 
Pnrnsiopsis ......................................... 31 
Eslricl~ocl~eles ........... .. .................... 31 
............................ lamidus, Gonodaclyl~cs 34 
........... ................... typn, Callinnidea ... 32 
typi1s 
Lucfe 6 
Oplophorus ......................................... 10 
............ .............. uncigcr. Ponlonidcs . 17 
undrrlalipes. Nemnlocorcines .......................... 8 
................................. Uroplyclrus ........ . 43 
Ulinon~ielln diniorphn ................................ 40 
vnclioni . Crrlcinris ................................... 38 
wglglds. Palaenlon ...................................... I I 
Vmderbillia rosw~ondnc ............................... 8 
\~nnnnnrei. Lilopenneics ........... . ................. 4 
vanninii, Telrnli 39 
velulinn, Moo 5 
ventricosn . Hippolyle .............................. 24 
venlrosss. Alpi~eus ..............  ................ 22 
~illosa, Fenchnlio ....................................... 4 
Vir sp. nov ..................... . ................. I7 
viridis 
.................................... Gonodnclylinl~s 34 
Gonodnclylrrs . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .  34
........... ............... viridis. Cornlliocnris .. I4 
........................ 
. . ............ 
. weberi Cmrrdrna . 9
n~illinmsi. Plesionika ................................. 29