NORTH AMERICAN TWO-WINGED FLIES OF THE GENUSCYLINDROMYIA MEIGEN (OCYPTERAOF AUTHORS) By J. M. AldrichAssociate Curator, Division of Insects, United States National Museum In the preparation of this paper the Avriter has studied the mate-rial contained in the United States National Museum, including hisown earlier collectino-; the Canadian National Collection; the Uni-versity of Kansas; the California Academy of Sciences; and theprivate collections of C. W. Johnson, James S- Hine; A. L. Melander,H. J. Reinhard, and H. W. Allen, together with a few specimensfrom various other sources. The total number of specimens ex-amined is nearly 800. Special acknowledgment is gratefully madeto C. Howard Curran, who when he learned of my work turned overall the material of the Canadian National Collection with a partlyprepared paper and numerous drawings. R. E. Snodgrass kindlyprepared drawings of the remarkable holding apparatus of thefemales of CylindroTnyia nana. Several European species deter-mined by Prof. M. Bezzi are in the United States National Museum,and have been very useful.The genus Cylimlroniyia was described by Meigen in his famous1803 paper, the real beginning of dipterous taxomony. Two yearslater Latreille described the same genus, with the identical genotype,as Ocyjjtera. Meigen, solely out of deference to his distinguishedcolleague, as far as can be perceived now, waived his own priorityand adopted Ocyptera in his next treatment of the genus in 1824.This disposition of the names was accepted for many years. OstenSacken cited this case in his paper on Priority or Continuity? in1882, as an illustration of the undesirable effect of too strict enforce-ment of priority. Professor Bezzi later argued that Ocyptera wasincluded in Latreille's 1802 volume, and hence really was prior; thatthis was an error was not discovered until after he had published histhird volume of the Palaearctic Catalogue. Coquillett, in his TypeSpecies paper of 1910, accepted Cylindromyia^ and was followed byTownsend in 1912.No. 2624.?Proceedings U. S. National Museum, Vol. 68, Art. 23.70648?26 1 1 2 PROCEEDINGS OF THE NATIONAL MUSEUM Vol. 68In the present case we have the consolation, in abandoning a well-known and long-used name, that we are going back to a classic au-thor and a historic work, and choosing a name based on an unques-tioned genotype. Even with this much in favor of Cylindromyia^ Iconfess I debated the matter a long time in my own mind, and finallyyielded Avith reluctance to what seems the logical conclusion.Genus CYLINDROMYIA MeigenMeigen, Illig. Mag., vol. 2, 1803, p. 279. Sole species, Musca brassicaria Fab-ricius.Latreille, Hist. nat. Crust, et Ins., vol. 14, 1805, p. 378 {Ocyptera, with 3species, of which Musca brassicaria was designated as type by Curtis, Brit.Ent, 1837, p. 629).Fallen, Rhizomyzides, 1820, p. 5 (Ocyptcra).Meigen, Syst. Beschr., vol. 4, 1824, p. 209 ( Ocypteru ) . Robineau-Desvoidy, Myodaires, 1830. pp. 229, 231 {Ocyptera and Partheniu,the latter with 3 species, of which Coquillett designated boscii as type in hiaType-Species, 1910, p. 585).Macquart, Hist. Nat. Dipt., vol. 2, 1835, p. 180; Dipt. Nord Fr.. 1834, p. 215;Dipt. Exot., vol. 2, pt. 3, 1843, p. 232 (all Ocyptcra).Zetterstedt, Dipt. Scandinav., vol. 3, 1845, p. 1219 {Ocyptera).Rondani, Dipt. Italicae. Prod., vol. 1, 1856, p. 78; vol. 4, 1861, p. 122 (bothOcyptera).ScHiNER, Fauna Austriaca, vol. 1, 18G2, p. 412 (Ocyptcra).Osten Sacken, Wien. Ent. Ztg., vol. 1, 1882, p. 191, says Cylindromyia is prior,but prefers to retain Ocyptera.Brauer and Bergenstamm, Zweifl. Kais. Mus.. vol. 4, 1889. p. 138; vol. 5, 1891,pp. 387, 407; vol. 6, 1893, p. 143 (all Ocyptera).Pandelle, Revue Entom., vol. 13, 1894, pp. 32, 57 (Ocyptera).CoQuiLLETT, Revis. Tachin., 1897, p. 86, key to North Amei'ican species (Ocyp-tera) ; Type-Species, 1910, 529, adopts Cylindromyia (Proc. U. S. Nat. Mus.,vol. 37).ViLLENEUVE, Wien. Ent. Ztg., vol. 22, 1903, p. 37 (Ocyptera, European species.with notes on Meigen's types).Van uer Wulp, Biologia, Dipt., vol. 2, 1903, p. 449, key to North Americanspecies (Ocyptera).Bezzi, Wien. Ent. Ztg., vol. 26, 1907, p. 55 ; Kat. Pal. Dipt., vol. 3, 1908, p. 429(both Ocyptcra) . TowNSEND, Muscoid Flies, 1908 (Smiths. Misc. Colls., vol. 51, No. 1803), p. 81,notes on species (Ocyptera) ; Proc. Ent. Soc. Wash., vol. 14, 1912, p. 48,adopts Cylindromyia, notes on habits ; Insecutor Ins. Menst., vol. 3, 1915, p.94 (Apinocyptera, type nignakt, new) ; Journ. New York Ent. Soc, vol. 23,1915, p. 233 (Odontocyptcra, type nana, new) ; Insecutor Ins. Menst., vol. 4,1916, p. 32 (Ncocypiera, type dosiadcs Walker) ; Proc. U. S. Nat. Mus.. vol.49, 1916, p. 631 (Ocyptcrodes, type cuchcnor Walker).Apstein, Sitzungsber. Naturforsch. Freunde Berlin, 1915, No. 5, p. 166, listsOcyptera, type brassicaria Fabricius, in his proposed nomina conservanda.Stein, Archiv. f. Naturgesch., vol. 96, 1924, p. 173, key to European species,with brief descriptions (Ocyptera).The species of Cylindromyia' b,vq, remarkably uniform in most oftheir characters; even those which show specializations in the female Art. 23 GENUS CYLINDROMYIA MEIGEN ALDRICH 6 ovipositing apparatus are in the male sex, as far as Imown, quitewithin the range of the rest of the genus. Several species show inthe females some specialization tending toward a single holdinghook instead of a symmetrical pair, and this is more or less associ-ated with the development of special groups of spines on the venter.Attention is especially called to the remarkable condition in 7ianaTownsend, in which one of the hooks is weak at base and regularlybreaks off at an early stage in the activity of the adult female tly.As a detailed description of all the species would consist mostly ofunprofitable repetition, it is thought best to begin with the genericcharacters (liberally interpreted) of the type species hrassieaiia ofEurope, taken from specimens identified by Professor Bezzi. Afterthe ke}^, each North American species is described only as far as thecharacters seem important for separating the species from hrasslcanaor known North American species. By this method it is hoped tosave a good deal of space and yet omit nothing of significance.GeneHc characters of Cylindromym (from the type species)?head in profile almost as long as high, from front a little wider thanhigh (length 39, height 42, width 51 units of micrometer) : the loweredge long and straight, the back bulging, so that the bare eyes arein profile about as far from the back edge as from the front; frontmoderately wide, .30 of head width in both sexes; frontal stripebroad, of uniform width to antennae, which are inserted above mid-dle of head, moderately long, the arista bare, with distinct but notelongated penultimate joint. Face not receding, slightly keeledabove, the epistoma jutting out between the vibrissae, facial ridgesbare. Parafacials bare. Palpi generally absent, occasionally presentin a very rudimentary condition (in European material?I have seenbut one case in the American; strangely enough this variabilityseems not to be even of specific value) ; proboscis slender but notmuch elongated. A^erticals one large pair; ocellars one smallishpair directed forward and outward, behind the ocellar triangle an-other pair directed outward, and still another small pair behind theseon the occiput directed upward; two pairs orbitals in female, nonein male; frontals about eight, none of the upper distinctly largerand roclinate, the lowest just at insertion of antennae, the row notdiverging toward the eye; bucca with only a single row of bristlesalong lower edge, disappearing behind.Thorax rather narrow, barely as wide as the head, with promi-nent humeri, the suture behind the middle. Postscutellum distinct.Chaetotaxy: acrostichai 0, 0; dorsocentral 3, 3; humeral 2, andnumerous outstanding stout hairs; posthumeral 1; presutural 1;notopleural 2; supraalar 1, stout; intraalar 2, only a little above thepreceding; postalar 2; scutellum with two stout lateral pairs and asmall decussate apical, no discal; sternopleural 2; pteropleural 0; 4 PROCEEDINGS OF THE NATIONAL MUSEUM Vol.. 68hypopleural 3 to 4; mesopleural 5 or 6 behind, one at lower frontangle; prothoracic 2 (just above front coxae); prosternum bare;postalar declivity bare; postnotum at side below calypter destituteof minute hairs.Abdomen narrow and elongate, the basal segment which in mostTachinidae is not visible from above is quite distinct though short(following custom I have ignored this in counting the segments) ; this short segment is without bristles; the first principal segmenthas a pair of median bristles not near the margin, and on eachlateral margin three, one behind the other ; second segment with onemedian marginal pair and one lateral near hind margin; third seg-ment with marginal row of four, no laterals ; fourth segment with arow of about 8, which are discal in the middle, marginal at the sides(the American are mostly marginal in the middle) ; fifth or firstgenital segment in male with one pair of small bristles projectingbackward. Basal sternite of abdomen overlapping the tergites, thefollowing sternites however concealed; second tergite with a fewbristles at hind angles below near median line.Male (fig. 1) with the usual posterior (or inner) and anterior (orouter) forceps, the former pale and much less chitinized than thelatter. Female with last genital segment forming a theca or cup,open apically, where a small sting protrudes; on each side at tip thechitin is produced into a curved horn or hook, not jointed, no doubtused in holding the host during the insertion of the sting. Thestructure is almost identical with that of inter^nedia (fig. 25).Legs stout, claws and pulvilli elongated in male, not in female.Wings narrow; fourth vein with stump at bend, the last sectionsinuous, joining the third considerably before its apex, petiole ofthird vein curving forward to join the costa far before apex. Veinsbare except a few hairs at base of third. Hind crossvein sinuous,joining fourth vein about last fifth of its penultimate section.Our species, like those of Europe, separate into three groups onthe bristles of the scutellum ; Pandelle used these characters in 1894,and Coquillett in 1897. Our dosiades has a single pair of scutellars(the posterior laterals of Pandelle), while intermedia and nigrahave in addition a small crossed apical pair. Compressa, knownfrom a single female, has in the type only one of the apicals, hencemay belong to either of the two groups. All the rest of our specieshave in addition to the two pairs mentioned a lateral pair near base(anterior lateral of Pandelle).Females of our species separate into three groups on the presenceor absence of groups of short ventral setules on the abdomen, on aswelling of the segment. G. dosiades and cmnjrressa have such aswelling with spinules on the first principal segment; noma and Akt. 23 GENUS CYLINDROMYIA MEIGEN ALDEICH O signata have it on the second ; while the other species do not have itat all.We have but one species, interviedia.^ with the hind tibiae villouson the flexor side in the male; and only one species, dosiades, withdiscal bristles (both sexes).The principal male genital organs for taxonomy are the forceps.The posterior (or inner) originate at the sides of the anus, and areusually of soft consistency. The anterior (or outer) are well chiti-nized, blade-like. The penis is quite simple and seems to present notangible characters. The fifth sternite is here a delicate plate, hardlymore than a membrane, and usually invisible until the genitalia havebeen pulled out of their cavity; it then shows important charactersin the width of the median incision, etc.The female genitalia are adapted for holding the host insectduring the insertion of the sting, which is much like that of a wasp.The last segment turns up against the preceding portion of theabdomen, and in most species has two lateral terminal unjointedhooks; the presence of these and their shape are of importance inseparating the species.In the wing the sinuosity of the hind cross vein is of some im-portance, but most of the supposed characters are variable and Ihave made little use of them.Systematic position.?Fallen was the first to attempt subdivisionscorresponding to what we would now call families in the muscoidgroup. He separated Ocyptera, Gymnosomu, and Alophora (whichhe called Thereva) from his Muscides in a family Rhizomyzides(Faun. Suec, 1820), Robineau-Desvoidy (Myodaires, 1830, p. 222)proposed a group Ocypteratae, which apparently would rank be-tween a subfamily and a tribe in the terms now used. Macquart(Dipt, Nord France, 1834, pp, 142, 212) adopted the same name;but as he greatly reduced the number of groups he in effect madethis of family rank, Schiner adopted Macquart's major groups withbut a slight change, having a subfamily Ocypterinae of equal rankwith his Tachininae and Dexiinae, Brauer and Bergenstamm atfirst ^ recognized a family Ocypteridae ; but later they receded frommost of their family names and ^ then designated a " Gruppe Ocyp-tera," which they placed between " Gruppe Pseudominthoidae " and " Gruppe Micropalpus." In their third paper ^ they have a slightlj'^extended " Sectio Ocyptera," Girschner * seems to have made noplace for the genus, as it does not run to any of his divisions. Pan-delle ' has a subfamily Ocypterinae in his family Tachinidae. ? Zvveifl. Kais. Mus., vol. 4, 1889, p. l.'^S.2 Vol. 5, 1891, p. 407. 'Vol. 6, 1893, p. 143.* must. Wochenschrift f. Eiit., vol. 1, 1890, p. Ill and proccdinfr."Kcv. Ent.. vol. 13, 1894, p. L'8. 6 PROCEEDINGS OF THE NATIONAL MUSEUM Vol. 68Bezzi ''' places Ocjjptern in Dexiinae. Baer ' does the same. Stein'sposthumous work ^ treats all muscoids with hypopleiiral bristles asthe single family Tachinidae, without introdiicin<>- subfamilies.In the United States, Coquillett ^ included Ocyptera in his Tachi-nidae. Townsend ^" in 1912 placed Cylindromyia in a tribe dylin-dromyiirii of the subfamily Phaniinae, family Exoristidae; he addedthe unique suggestion that the tribe Qonipsilurini would go in thesame subfamily. In a manuscript outline of his classification, sub-mitted quite recently, he puts the tribe Cylindronvyiini in the familyPhasiidae.In the present paper it is proposed to consider Cylindromyia thechief member of the small subfamily Cylindromyiinae of the familyTachinidae. No attempt will here be made to analyze the othercomponents of the group.Biology.?About a century ago, Dufour observed the parasitismof the European Cylindromyia hicolor Olivier on the PentatomidRhaphigaster gnsea Fabricius;'^ he figured adult and larva.Nielsen has given a report of some studies of his own, of Cylindromyiahrassicaria Fabricius parasitic on another Pentatomid, Dolycorishaccarum}'^ He found larvae in the first and third stages at thesame time in the last of July, but only one in each host. He notedthat the larva hibernates in its host in the second stage, and thatit does not kill the host, even at the time of exit. It pupates in theground. He thinks the fly is ovoviviparous (Pantel's group 9).Caffrey and Barber, in Bulletin 779 of the United States Depart-ment of Agriculture, have noted on page ;^1 that they reared eightspecimens of Cylindroinyia armata (as Ocypterodes euchenorWalker) from adults of the Pentatomid Chlorochroa sayi Stal atMaxwell, New Mexico. I correct the species from one of the speci-mens. Townsend reported the rearing of euchenor by Forbes inIllinois from Acrididae and from Leucania u)iipu?icfa; ^^ but theserecords may well be doubted, as all the others are from Pentatomidae. KliY TO NORTH AMERICAN .SPKCIES OF CYLI XDUO-MY lA1. Scutelluin with only one pair of bristles (fig. 17) ; abdomen with discnl niac-rochaetiie (widespread; dosiadcs Walker) dosiades Walker.Scutellum with two pairs of bristles?a small decussate apical iiair andanother much larger not far from them (fig. 16) 2Scutellum with three pairs of bristles, the additional pair being near thebase at the sides (fig. 15) 4 "Kat. Pal. Dipt., vol. 3, 1908, p. 42!)."Die Tachincii, 1921, p. 157.SArch. f. XaturKescli., p. 96, 1924.? Revls. Tacliin., 1897, p. 86.?? Prop. Ent. Soc. Wash., vol. 14, p. 48."Annales Sci. Nat., vol. 10, 1827, pp. 246-260, pi. 11." Entom. Medd., ser. 2, vol. 4, 1909, pp. 76-81. flss." Psycho, 1893, p. 400. Art. 23 GENUS CYLINDEOMYIA MEIGEN ALDRICH 7 2. Abdomen without red on sides, wliolly black ; front basitarsus shorter thanthe four following joints (eastern; argentea of Coquillett 1897, not ofTownsend) nigra, new species.Abdomen broadly red on sides 33. Female with distinct, symmetrical hooks on last genital segment, which areplaced quite far apart ; male with the front basitarsus as long as the fol-lowing four joints and concave on outer side, also with hind tibia villouson inner side (widespread west and north ; European) __ intermedia Meigen.Female with last genital segment compressed, pointed, without hooks ; fourthabdominal segment with a pair of stout bristles near base above; male pre-sumably without the characters just mentioned (Alberta).compressa, new species.4. Abdomen wholly shining, the incisures not white poUinose (Colorado).alticola, new species.Abdomen with very evident white pruinosity 55. Tibiae red ; veins around discal and first posterior cells with sharply blackishborders (California, Mexico) limbata, new species.Tibiae black; the dark color of the wings more diffused 66. A conspicuous white poUinose spot on the sclerite just above the posteriorthoracic spiracle (Cuba) bakeri, new species.No such spot, the sclerite shining black 77. Abdomen without any red; large species (Porto Rico) atra Koeder.Abdomen broadly red on sides or apically 88. Males J-- 9Females 149. (MALES)?The posterior forceps short and great y swollen, soft and cush-ion-like, but suddenly contracted into a small apical beak 10The posterior forceps long and soft, basal half swollen, suddenly reduced atmiddle, the apical portion reaching almost to tips of anterior forceps (eastern, widespread) argentea Townsend.Posterior forceps not as described 1110. The fifth sternite with a wide, shallow emargination in the middle (east-ern) euchenor Walker.The fifth sternite with two narrow prolongations close together, forming adeep narrow notch at the middle line (Mexico City; Dominica).uniformis, new species.11. Both pairs of forceps yellow, shining, alike in consistency, the posteriorehitinized, both pairs small and somewhat concealed by the thin yellowexpanded margin of the last genital segment (widespread).nana Townsend.The posteiior pair of forceps much less ehitinized than the anterior pair,which are hard and shining 1212. Anterior forceps very broad apically, broader than at base, subtruncate(widespread) armata, new species.Anterior forceps tapering toward tip, not so wide apically 1313. Posterior forceps short and in profile rather thick, but not strikingly .swol-len, about half as long as anterior ones, which are grooved behind at thebend ; back of head with some black hair on each side of the occipitalregion above (wulcspread) decora, new si>ecies.Posterior forceps more than two-thirds as long as anterior, which are notgrooved behind at the bend ; back of head with only pale hair inside theorbital row (widespread; euchenor of Townsend 1916).vulgaris, new species. 8 PROCEEDINGS OF THE NATIONAL MUSEUM Vol. 6i 14. (FEMALES)?Second principal segment underneatli with a large crescent-shaped area of short stubby spines on a prominence, behind which is aconcavity ; terminal hooks of last genital segment unsymmetrieal or onlyone present 15Second abdominal segment destitute of such spiny, swollen area ; hooksof last genital segment symmetrical 1615. Large tropical species with only one genital hook, which is strikingly large ; fifth and sixth abdominal segments red (Guatemala) ._ signata Townsend.Small, more northern species, the left genital hook deeply grooved at base ifpresent, usually broken off; tip of abdomen shining black (widespread).nana Townsend.16. The hooks on last genital segment attached below a square upper apicalshoulder of the segment 17Last genital segment without such shoulder, sloping down to the hook 1917. With only 2 sternopleurals ; genital hooks as in armata (widespread).decora, new species.With 3 sternopleurals 1818. The genital hooks pointing decidedly forward, gradually turned upward(widespread) armata, new species.The genital hooks turned upward from base, standing close to shoulder, thetips divaricate (widespread) euchenor Walker.19. Usually with only 4, sometimes with 6, bristles on tergite of fourth abdomi-nal segment; large, common species (eastern) argentea Townsend.With more than 6 bristles on the tergite of fourth abdominal segment (wide-spread) vulgaris, new species.CYLINDROMYIA DOSIADES Walker(Figs. 7, 22)Ocyptera dosiades Walker, List. Dipt. Ins., vol. 4, 1849, p. 695. ? Van debWxn:p, Tijdschr. Ent., vol. 26, 1883, p. 15, occurs in Quebec. ? Giglio-Tos,Ditt. del Messico, vol. 3, 1893, p. 3. ? Coquillett, Revis. Tachin., 1897, p.86. ? Johnson, List Ins. New Jersey, 1899 and 1910; List Dipt, of Florida,1895 and 1913.?Hine, Canad. Ent., vol. 36. 1904, p. 91.?Van der Wxjlp,Biologia, Dipt., vol. 2, 1903.?Chagnon, Eutom. Student, vol. 2, 1901, p. 15,occurs in Montreal. ? Gibson, Entomol. Record, 1911, 1914, 1915, occurs inCanada. ? Van Duzee, Canad. Ent., vol. 43, 1911, p. 237, occurs in Kearny,Ontario. Winn and Beaulieu, Ins. Prov. Quebec, 1915, p. 141. Britton,Check-List Ins. Connecticut, 1920. ? Cole and Lovett, List Dipt. Oregon, 1921,p. 301. ? Brimley, Ent. News. vol. 33, 1922, p. 232, occurs in North Carolina.Ocyptera euchenor Townsend, Journ. New York Ent. Soc, vol. 5. 1897, p.176 (makes dosiades a synonym) ; Psyche, 1897, p. 149.Ocyptera euchenor, var. dosiades Townsend, Psyche, 1898, p. 212.Neocyptcra dosiades Townsend, Insecutor Ins. Menst., vol. 4, 1916, p. 32.The species has the usual red or yellow areas on the sides of theabdomen, leaving a rather wide median black stripe, and the wholeposterior end of the abdomen shining black.The male has the posterior forceps entirely united, brown andenlarged at base, where they bear a tuft of black hair; apically theyrun out in a long yellowish point, slender in profde, grooved behind, Art. 23 GENUS CYLINDROMYIA MEIGEN ALDRICH 9 nearly as long as the anterior forceps; the latter are shininc; black,moderately slender, gently curved to a rounded apex. The sides ofthe last genital segment project in a thin yellow process along theside of the cavity containing the genital organs.In the female the last abdominal segment is in the form of a deepspoon with a deep median notch in its apex, from which the stingusually projects; there are no hooklike processes on the segment.The type of dosiades from Nova Scotia is in the British INIuseum.The species is quite recognizable from the description, and I amassured by Maj. E. E. Austen that we have it correctly identified(as dosiades, he is not responsible for the s3monymy). Sixty-eightspecimens before me of both sexes show the following distribution : In National Museum.?Vicinity of Washington, District of Colum-bia, including adjacent Maryland and Virginia, 14 specimens col-lected b}'' McAtee, Walton, Knab, Roliwer, Barber, Greene, Myers,Jackson, and Aldrich; others from Bar Harbor, Maine (C. W. John-son) ; White Mountains, New Hampshire (Morrison) ; base of MountWashington, New Hampshire (Townsend) ; Wilmington Notch,Adirondacks, New York (Aldrich) ; Oswego, New York; Harrisburg,Pennsylvania (Walton) ; La Fayette, Indiana (Aldrich) ; SouthGeorgia (Morrison) ; Minot, North Dakota, and Moscow, Idaho(Aldrich) ; Lindsa}^, California (McGregor) ; Rampart, Alaska, *' through B. Preston Clark." The last locality is almost at theArctic Circle, on the Yukon.In the Canadian National Collection.?Oliver, British Columbia(Garrett) ; Agassiz, British Columbia (Glendenning) ; Banff,Alberta (Garrett) ; Aylmer, Quebec (Miss G. Beaulieu) ; Ottawa,Ontario.In Professor Melander's collection (all collected by him except asnoted): Woods Hole, Massachusetts; Lyndon, Vermont; Chicago,Illinois; Moscow and Chatcolet, Idaho; Perma, Montana; Pullman,Seattle, and Lake Cushman, Washington; Stanford University,California (Mann); Pullman, Washington (Mann).In Professor Hine's collection.?Pelee Island, Ontario (Gaige) ; Kaslo, British Columbia (Osburn) ; Winona Lake, Indiana.In H. J. Reinhard's collection.?Ohio (Reinhard).Two specimens in Professor Melander's collection (Waubamic,Ontario and Almota, Washington) have no discals on the second andthird segments; one specimen from Laggan, Alberta (Osburn, inHine's collection), and one from Colorado, have the regular pair ofsmall decussate apical scutellars occurring on other species of thisgenus; and one specimen from Bottineau, North Dakota (Ainslie),has the head unusually long on its lower edge. With the availablematerial I do not consider these aberrations specific.79G48?26 2 10 PROCEEDINGS OF THE NATIONAL MUSEUM Vol. 68CYLINDROMYIA COMPRESSA. new species(Fig. 23)A small species of ordinary appearance, known only in the female,and having very unusual egg-laying apparatus. Parafacials andparafrontals silvery, the former a little wider than the third an-tennal joint. Antennae black, rather small, the second joint con-siderably more than half the third. Sternopleural two. Scutellumwith one large pair of bristles near the apex, between which in thesingle specimen there is an apical bristle on one side but not on theother. First and second abdominal segments red except the base offirst and a very little of the apex of the second above. The firsthas only two lateral bristles, one near the base, the other near thehind edge. No discals except a pair near the base on fourth seg-ment. On the median line below the ends of the first principaltergite seem to fuse together behind, and in this region bear about20 distinct small spiny bristles on a slight swelling. In profile theventer of the second segment is a little concave. The third andfollowing segments of abdomen are shining black. The third seg-ment below with a median groove for the reception of the tip ofthe last genital segment; this groove continues on the fourth seg-ment where it is mostly filled up by a globose shining black mediansclerite; fifth segment contracted and tapering; sixth segment ratherlong, compressed, curved, shining black, drawn out into a sharpupturned point as viewed in profile; the point is composed of twoblade-like portions divided below for some distance and betweenthem the sharp sting projects. Front tarsi with a slight widening ofthe last three joints. Wings with rather uniform smoky coloration,more yellow in the costal cell and at base.Length, 7.G mm.Described from a single female specimen taken at Banff, Alberta,by C. B. D. Garrett.Type in the Canadian National Collection.This species is most nearly related to dosiades, in which occa-sionally the discals are partly absent. Whether compressa normallyhas one pair of scutellars or two can not be decided from the singlespecimen, in which the apical pair is represented only by a bristleon one side. Both of these species have in the female a swollen,spiny area on the venter of the first abdominal segment, not thesecond as in signata and nana', they also have no hooks on the lastgenital segment. There is no doubt of the distinctness of compressahowever, as the last genital segment is much more acute and com-pressed than in dosmdes.The Eui'opean interrupta Meigen is very much like this species, asI note from a female in the Canadian collection kindly sent me forstudy. It has discal abdominal bristles however. Art. 23 GENUS CYLINDROMYIA MEIGEN ALDEICH 11CYLINDROMYIA NIGRA, new species(Figs. 10. 32)Ocyptera argentea Townsend, Coquillett, Revision Tachin., 1897, p. 86 (misideri-tification). ? Johnson, Cat. Ins.. New Jer.sey, 1899 and 1909. ? Banks, Ent.News, vol. 23, 1912, p. 110.?Brimley, Ent. News, vol. 33, 1922, p. 24(argentata, a slip of the pen).A small species easily distinguished by the characters in the key.The vertex is shining, and the parafacials and parafrontals havesmooth pale yellow pollen: the parafacials are rather narrow, andthe frontal stripe is narrower than one parafrontal in its middle.Antennae black, but the tip of second joint and base of third red-dish; third joint one and one-half times the second. In profile theeye is nearer the front than back of the head. Bucca about one-eighth of the eye.Thorax black, the silvery pollen of the sides extending upon thedorsum to include the presutural bristle.Abdomen wholly black, the white pruinose bands arcuate, so asto include a little of the back edge of the first and second segmentsat the middle, and also extending upon the venter. First principalsegment Avith only two bristles on the lateral margin ; fourth segmentwithout bristles except one on each side.Wings brown, fourth vein generally not with stump at bend, theapical cell wide distally; hind crossvein straight, almost erect.Male.?Second principal abdominal segment with a cluster of fourspines at the median line below, on the inflexed tergite before itsposterior edge. Posterior forceps united to tip, of soft texture,basally brownish, apically whitish and swollen, as long as anteriorones, which are shining brown and end in a sharp point on theanterior edge.Fem, Proc U. S. Nat. Mus.. vol. 49, 1916. p. 631.Townsend designated euchenor Walker as type of his genusOcypterodes; the specimen so labeled by him belongs to the si3eciesherein described as new under the name vulgaris. For euchenor'as reared from a Pentatomid by Caffrey and Barber, see ai^iata.The species here selected as the true euchenor is the one agreeingw^ith the type, as made out by Major Austen from my key and somenumbered specimens that I sent him (his letter October 14, 1923).It is strikingly similar to argentea in almost every detail, buteasily recognizable by the male genitalia and the females can alsobe separated without any difficulty. In the male the posterior for-ceps are .soft and swollen, subglobose, almost white, with a minutebeak on the anterior side at the tip, between the anterior forceps.The soft cushionlike swelling of the posterior forceps is very re-markable. 'Jliere seems to be no separation between the forceps 16 PROCEEDINGS OF THE NATIONAL MUSEUM Vol.68although there is a deep groove. The anterior forceps are shiningbrown, large, considerably curved forward toward the apex and theyhave on the posterior and inner edge a groove beginning at the tipof the posterior forceps and extending nearly to their own apex.The thin margins of this groove curve slightly up or down, con-siderably changing the shape of the organ. The fifth sternite has awide excision, with a cluster of hairs each side, and outside of thisanother excision beyond which are about three bristles. This ispractically the same in argentea. The female can be distinguishedby the character given in the table?the terminal hooks of the lastgenital segment being attached below a square shoulder with whichthey are nearly jjarallel in profile. There are no special groups ofspines on the venter.Length, 9 to 11.5 mm.Redescribed from 35 specimens, of both sexes. Of these 26 arein the National Museum, including 16 from the District of Columbiaand adjacent parts of Maryland and Virginia, collected by Town-send, Currie, Quinter, Shannon, Crawford, and Walton. The restare from Franconia, New Hampshire (Townsend)^ Ohio (Rein-hard) ; Union Springs, New York (Anderson) ; Clementon, NewJersey (Johnson) ; White Springs, Florida (Townsend) ; CollegeStation, Texas (Reinhard) ; Galveston, Texas (Trotter) ; Boulder,Colorado (Cockerell).In C. W. Johnson's collection are five specimens, from Province-town, Massachusetts (Morse) ; Durham, New Hampshire (Brid-well) ; Manomet, Massachusetts (Johnson) ; Monmouth, Maine(Frost), and Wellesley, Massachusetts (Morse).In Professor Hine's collection are three specimens, from Hinckley,Ohio (Hine) ; Bluffton, Indiana (Williamson), and Golden, Colo-rado (Hine).In the California Academy of Sciences is one s[)ecimen from SanMarcial, Socorro County, New Mexico (Duncan).In the TTniversity of Kansas collection are 9 specimens, fromConnecticut (Williston) ; Illinois (Forbes) ; South Illinois; Ithaca,New York; Ames, Iowa; Pennsylvania; Minnesota, and Galveston,Texas (Snow). CYLINDROMYIA ALTICOLA. new species(Figs. 13, 24)The only North American species at present known in which theabdomen is entirely shining without any pollinose markings. Frontalstripe decidedly red, at the middle about three times as wide aseither parafrontal, the latter and the parafacial silvery; the para-facial at narrowest is hardly wider than the third antennal joint. Art. 23 GENUS CYLINDROM YIA MEIGEN ALDEICH 17Antennae rather small, blackish, the arista short. Thorax black,with a distinct pollinose spot on the humerus extendin<]^ almost tothe suture, and on the sides with silvery pollen on all of the coxaeand extending vaguely above the first and second ones. Sterno-pleural 2, with an additional small one on one side in 1 male. Abdo-men broadly dark red on the sides. Wings rather narrow, evenlyinfuscated from front to back; the fourth vein with a long branchat its bend, its last segment far from the apex of the wing.Male.?Claws and pulvilli all elongated, second principal segmentof the abdomen a little swollen along the median line below, thethird segment with a flattened V-shaped opening toward the geni-talia. Genital segment blackish, the hind forceps united, of rathersoft texture, deeply grooved behind, straight almost to the apex,where they bend sharply forward and are not very sharp. Anteriorforceps concealed by a long swollen yellow lobelike expansion ofthe side of the last genital segment, so that only the tips, whichare strongly bent forward and rounded, not sharp, are visible. Fronttarsi plain.Female.?Front tarsi elongated and distinctly flattened beginningwith the second joint. The second principal segment of the abdomena very little swollen on the under side, somewhat less than in themales. The genital segments black, the last one ending w^ith a rathersquare shoulder which bears an almost imperce^^tible tooth, therebeing no hooklike structure developed.Length. G to 7 mm.Described from two males and two females collected at MarshallPass, Colorado, by the writer on July 28, 1908, at an altitude of10,856 feet.One of the females possesses very minute but unmistakable palpi.7V/^e.?Male, Cat. No. 28280, U.S.N.M.CYLINDROMYIA NANA Townsend(Figs. 14, 19, 20)Odontocyptera nana Townsend, ,Jouni. New York Ent. Soc, vol. 23, 1915,p. 233, female.The parafrontals and parafacials silvery, the brown frontal stripeabout as wide in the middle as the width on both sides combined.Parafacial almost twice as wide as the third antennal joint; theposition of the eye a very little more vertical than usual. Thoraxwith considerable silvery pollen, viewed from behind there is abroad black stripe outside the dorsocentrals before the suture and aslender stripe on the inner side of the same bristles which does notreach the suture. The abdomen varies a good deal in the amount ofpollen that it shows and the red portion varies in extent but tends 18 PROCEEDINGS OF THE NATIONAL MUSEUM Vol. 6Sto cover most of the second and third principal segments. Some-times the median black stripe is absent.Male.?Last genital segment varying from red to black, the pos-terior forceps yellow, shining, apparently united, ending in a sharppoint; outer forceps of the same color, rounded at tip, straight.Margin of last genital segment thin and shining yellow, projectingabout as far as the tips of the posterior forceps in a broad lobe;penis longer than usual, slender in the middle, enlarged apically;fifth sternite so retracted as to be practically invisible.Female.?The first principal segment of the abdomen bears fourbristles on the hind edge where the corners come together at themiddle of the venter, the second segment on the median ventralportion has a transverse protuberance somewhat arcuated, bearingnumerous, very short, stiff spines. The third segment has an ovalbasin-shaped concavity on the middle ventral line into which thehook folds; the fifth apparent segment of the abdomen is elongatedand conical, covered with short, retrorse hairs; the terminal genitalsegment is small, composed chiefly of the elongated hooks, which areunsymmetrical in shape, the left one being strongly constricted atthe base, bulging beyond this region; the right one is stout at thebase and the cavity which ordinarily lies in a symmetrical positionbetween the two hooks is in this case thrown up along the side ofthe larger hook. Townsend's type specimen and one from SouthernPines, North Carolina, are the only females we possess that havetwo hooks; 15 other females differ only in having the left hookbroken off at the base, the fractured portion is exceedingly slenderand lies along the edge of the median cavity, the stinglike ovipositorso characteristic of this group lies in a groove along the side of theremaining, or right-hand hook.Length, 5.5 to 8 mm.Redescribed from 46 specimens (29 males, 17 females). H. J.Reinhard collected 20 males and 12 females at College Station,Texas, and this lot enabled me to feel certain that the two sexesbelong together. In Professor Melander's collection are 2 malesfrom Wawawai, Washington, collected by him.In the Kansas collection are 3 males from Santa Rita ]Mountains,Arizona (Snow).The remainder are in the National Museum as follows: 1 femalewith both hooks, the type, collected by Townsend at Head of RioPiedras Verdes, Sierra Madre, Chihuahua, INIexico; 1 female, withboth hooks. Southern Pines, North Carolina (Manee) ; 2 females,Claremont, California (Baker) ; 1 male, 1 female, Moscow, Idaho(Aldrich) ; 1 male, Lewiston, Idaho (Aldrich) ; 1 male, DevilVRiver, Texas (Bishopp) ; 1 male, Kerrville, Texas (Pratt). Art. 23 GENUS CYLINDROMYTA MEIGEN ALDEICH 19CYLINDROMYIA ATRA Roeder(Figs. 11. 29)Oci/ptera atra Roder, Stett. Ent. Zeit., 1885, p. 344.?Van der Wulp, BioloRia,Dipt., vol. 2, 1903, p. 450.?GiGLio-Tos, Ditt. del Me.ssico, vol. 3. p. 3, 1894(Mem. R. Accad. Sci. Torino, ser. 2, vol. 44.)Ocyptera minor Roder, Stett, Ent. Zeit., 1885, p. 344.A large shining black species Avithoiit red on the abdomen.Three sternopleiirals. Abdomen with two silvery, pruinose cross-bands, the posterior one including most of the third segment; fourthsegment shining black with eight bristles.Male.?Fifth sternite with a narrow excision in the center, on eachside of which is a slight prolongation followed by an emargination,beyond which at the side are a couple of flattened spines; secondprincipal tergite of the abdomen with two or three pairs of long de-pressed bristles at the hind corners below next to the median line.Genital segments dark brown or black. Posterior forceps united,soft, deeply grooved behind in profile, of uniform width almost totheir tip, then drawn out in a point on the anterior edge. Anteriorforceps shining brown, rather broadly and evenly curved forwardwith a rather sharp apex.Female.?Similar to male, the abdomen in profile rather deeper,not provided with a special group of short spines. Genital segmentsshining black, the last one piceous on the upper side, sloping rathergradually toward the hooks which are large and long, moderatelydivergent and curve gently forward. Front tarsi not perceptiblyflattened.Length, 9 mm.Redescribed from 1 male from Porto Rico (W. V. Tower) in theUnited States National Museum, and 1 female from the AmericanMuseum of Natural History, collected at Barros, Porto Rico.CYLINDROMYIA BAKERI, new species(Fig. 5)Male.?Very similar to atra., but readily distinguished by thecharacter given in the table. In the single specimen the abdominalbands are narrower and more sharply defined, especially the posteriorone. The genital segments are brown, the posterior forceps of thesame color are chitinized, very short and rather bluntly pointed.The anterior forceps are shining brown and broader on the basal partthan in any other known species. They are bent but little forwardand have a broadly rounded apex ; the fifth sternite is rather broadlyemarginate in the middle and deeply excavated outside of this.The female is imknown, but I have ventured to assume that itThis is a medium-sized species with brightly m^irked abdomen,very common through the western part of the United States, morerarely occurring in the region of Washington, District of Columbia.The red markings of the abdomen are very distinct, the median blackstripe often lacking. The distinctive characters are given in thekey and those of the male are most easily made out. The forcepsare almost exactly as figured for atra. The posterior forceps areshining brown or black on the sides, widening at the apex. Asusual, they are united and deeply grooved behind. In profile theyare of about uniform width to the tip, where they end in a si;ialltoothlike projection closely applied to the anterior forceps. Thelatter are shining brown or black, the hind edge broadly curvedforward, not grooved at the bend, the width nearly uniform formost of the length, the front edge curving forward beyond the mid-dle to a rather sharp tip. The projecting edge of the last genitalsegment is not very prominent ; the fifth sternite has a narrow notchin the center with a considerable lobe each side, beyond whichthere is a deep, rounded excavation, and at the lateral margin threellattened bristles of increasing size. The second principal segmentof the abdomen has on the under side at the posterior corners two orthree pairs of depressed long bristles. The fourth segment hasabout eight marginal bristles. In profile the eye is as far from thefront edge of the head as from the hind edge. There are threesternopleurals in the female, the abdomen is without any charac-teristic cluster of ventral spines, and the depressed bristles of thehind edge of the second segment below are small and inconspicuous.The hooks of the last genital segment are large and strongly curved,but there is no square shoulder above their origin.Length, 7 to 9 mm.Described from 143 specimens of both sexes.In the National Museum are 47 specimens, of which 15 are fromthe District of Columbia and the adjacent parts of Maryland andVirginia, collected by Greene, Currie, McAtee, Walton, Townsend,and Rohwer. The remainder are from Chesapeake Beach, Maryland(Shannon) ; Colonial Beach, Virginia (Silver) ; Michigan City,Indiana (Aldrich) ; Cadet, Missoiu'i (Riley coll.) ; East Point, Art. 23 GENUS CYLINDROMYIA MEIGEN ALDEIGH 21Louisiana (Bishopp) ; Brookings, South Dakota, and Minot, NorthDakota (Aldrich) ; Colorado; Denver, Colorado (Jackson); NorthCheyenne Canyon, Colorado (Champlain) ; Binford, North Dakota(Mabbott) ; Quinlan, Texas (Bishopp) ; Austin, Texas (Melander) ;Socorro, New Mexico (Williston) ; Tempe, Arizona (Caffrey) ;Ormsby County, Nevada (Baker) ; Moscow, Idaho (Aldrich) ; Pull-man, Spokane, and Fishtrap Lake, Washington (Aldrich) ; We-natchee, Washington (Melander) ; Bead Lake, Newport, Washing-ton(Lane) ; Vernon, British Columbia (Treherne) ; Yosemite, Califor-nia (Westcott) ; Visalia and Los Angeles, California.In the Kansas collection are 32 specimens, of which 15 are fromDouglas, Decatur, Pratt, Marion, Johnson, Anderson, Clark, Mc-Pherson, Smith, and Linn Counties in Kansas; collected by Wil-liams, Snow, Beamer, Martin, and Hoffman; the other are fromRock Creek, Kansas (Moodie) ; Illinois (Forbes) ; southern Illinois(^Robertson) ; Santa Rita Mountains, Arizona (Snow) ; Bailey, Colo-rado; Montana; California and Washington Territory.In H. J. Reinhard's collection are 23 specimens from CollegeStation, Texas, and 3 from Dallas, Texas, all collected by himself.In the California Academy of Sciences are 25 specimens, of which21 are from localities in California?Yosemite Valley, StrawberryValley, Vivian Park, Clayton, ISIineral King, Davis Creek, BuckCreek, Mendocino County, Stockton, Panoche Hills, and San Diego,collected by Van Duzee, Van Dyke, Fox, Leach, and Blaisdell. Theother four are from Garfield, Logan, and Salt Lake City, all in Utah,and collected by Van Duzee.In C. W. Johnson's collection are two specimens; 1 from Stam-ford, Connecticut (Morse), the other from Cranbrook, British Co-lumbia (Garrett).In the Canadian collection are 8 specimens, all from British Co-lumbia, as follows: Victoria, William Head, and Saanich District(all by Downes) ; Vernon (Cutler) ; Oliver (Vroom, Buckell) ;Penticton (Treherne); Lillooet (Anderson).Type.?M^\Q, Cat. No. 28282, U.S.N.M., from Falls Church,Virginia (Greene) ; allotype female from Great Falls, Virginia(Currie). CYLINDROMYIA DECORA, new species(Fiss. 12, 30)This species is strikingly similar to vulgaris^ but seems to occurmore eastward and can be readily separated, at least in the male,by the characters given in the table. The second abdominal seg-ment has rather smaller bristles at the lower hind corners. The fifthsternite is the same. The posterior forceps are more white in color,noticeably shorter and in profile thicker than in vulgay^ls. The 22 PEOCEEDINGS OF THE NATIONAL MUSEUM Vol.68 anterior forceps are more stronglj^ curved forward, beginning totaper at about the middle and have on the hind edge, somewhatbefore the tip, a distinct groove on the inner side.The female, if I have associated it correctly, has on the last genitalsegment a square shoulder on each side at the lowest edge of whicha strong upcurved hook originates which is directed more forwardthan in eucKenor. The presence of dark hairs on the occiput oneach side of the center above seems to distinguish the species fromvulgaris.Length, 7 to 9 mm.Described from 40 specimens of both sexes.In the National Museum are 11 specimens, from Franconia, NewHampshire (Townsend) ; Ithaca, New York (Anderson) ; Fishkill,New York (Kalmbach) ; Ann Arbor, Michigan (Anderson) ; Penn-sylvania (Walton); Missoula, Montana; Pollock, Idaho (Aldrich) ; Chevy Chase Lake, Maryland; and Mojave, California.In C. W. Johnson's collection are 5 specimens, from Colebrook,Connecticut (Wheeler); Rutland, Vermont; Niagara Falls, NewYork ; Delaware County, Pennsylvania ; and Point Antonio, Jamaica(last 4 by Johnson).In J. S. Hine's collection, 1, "G. W. Lake" (Osburn).In the California Academy of Sciences is 1, from Pleyto, Men-docino County, California (Van Duzee).In Professor Melander's collection are 3, from Waubamick, On-tario (Parish) ; Chicago, and Hayti.In the Canadian collection are 6; 3 from Jordan, Ontario; theothers from Ottawa, Ontario (Beaulne) ; Jordan, Ontario (Ross) ;and Osoyoos, British Columbia (Anderson).In the Kansas collection are 13 specimens, from Barton, Ness, andClark Counties, Kansas (Williams) ; Champaign Coimty, Illinois(Snow) ; Illinois (Forbes) ; Ithaca, New York; near Lander, Wyo-ming (Moodie) ; and California; "Brooks."In the American Museum of Natural History are 2, from SantaRosa Island, California (Wheeler).Typ^.?Male, Cat. No. 28283, U.S.N.M., from Franconia, NewHampshire ; the allotype female is from Pollock, Idaho.The females are very close to amiata^ and I fear may not all sepa-rate on the sternopleural character.CYLINDROMYIA ARMATA, new species(Figs. 9, 26)Ocyptcra caroUnae Gkeene, Proc. U. S. Nat. Mus., vol. 60. 1922, art. 10, p. 11,fis. 20, puparium.Parafrontals and parafacials silvery, the latter much wider thanthe third antennal joint, the former very narrow to the vertex. Art. 2.T GENUS CYLIXDROMYTA MEIGEN ALDRTCH 23Sternopleurals three; in the male the posterior forceps are soft andrather short, somewhat thick in profile, united and deeply groovedbehind, varying in color from almost white to brownish. The ante-rior forceps are the most characteristic feature of this species, beingrather broad at base and still more so toward the apex, where theyare almost obliquely truncate, with a rounded point on the frontedge; the fifth sternite has a notch in the middle separating twosmall lobes and beyond these at the sides two or three somewhatflattened small bristles. On the venter close to the middle line thereare two or three pairs of bristles on the hind corners of the tergite ofthe second segment. In the female the hooks at the tip of the lastgenital segment are directed more straight forward than in otherspecies and there is no shoulder at their base. The abdomen isdestitute of groups of special short spines on the venter.Length, 7 to mm.Described from 26 specimens of both sexes.In the National Museum are 13 specimens, from Bethayres, Penn-sylvania (Harbeck) ; Rock Creek, District of Columbia (Town-send) ; East Falls Church, Virginia (Rohwer) ; Chesapeake Beach,Maryland (Shannon) ; Beaver Creek, Montana (Hunter) ; Brock-ton, Montana (Sperry) ; Boulder, Colorado (Cockerell) ; Pueblo,Colorado (Wetmore) ; Northern New Mexico (Weese) ; Socorro,New Mexico (Williston) ; Maxwell, New Mexico (Barber) ; Mouthof Bear River, Utah (Wetmore), and Rocky Ford, Colorado, pu-parium found in beet field (McMillan).In C W. Johnson's collection are 5 specimens, from Cohasset,Massachusetts (O. Bryant) ; Anglesea, New Jersey (Johnson) ; andColorado (Baker).In the California Academy of Sciences are 3 specimens, from Gar-field and Saltair, Utah, and Panoche Hills, Western Merced County,California, all collected by E. P. Van Duzee.In the Kansas collection are 5 specimens, from Douglas County,Kansas (Snow) ; Beaver Creek, Montana (Hunter) ; Creede, Col-orado (Hunter) ; and New Jersey (Johnson).7'ype.?Male, Cat. No. 28284, U.S.N.M., from Chesapeake Beach,Maryland. The allotype female is the one from Rock Creek, Dis-trict of Columbia.The puparium of the Rocky Ford specimen, a female, was figuredby Greene under the name Ocyptera carolinae. I correct the identi-fication from the adult.CYLINDROMYIA SIGNATA Townsend(Fig. IS)Apinocyptcra signata Townse?.d, Insecutor Ins. Meust., vol. 3. 1915, j). 94.This is a large dark, tropical species, known only in the female,which has very peculiar genitalia. Parafacials silvery, the para- 24 PROCEEDINGS OF THE NATIONAL MUSEUM Vol. 08frontals slightly yellow, a double pollinose spot between the verticalbristles behind the ocellar. The front and lower part of the headappears to be yellow in ground color, the parafacial is considerablywider than the third antennal joint. First and second joints ofantennae and base of third yellowish, Sternople'ural two. Firstprincipal segment of the abdomen entirely black, except the verynarrow hind edge above; second segment broadly yellow on eachside, its ventral part strongly projecting, with numerous very shortstubby spines on the swollen portion, which forms a flattened areaextending back upon the following segment; third and fourth seg-ments reddish, nearly covered with a very thin whitish pruinosity.The spines on the fourth segment become irregularly discal in themiddle; the fifth segment bent forward and under the abdomen,long and conical, reddish-brown in color, covered with short, stiff,retrorse hairs; the last genital segment small on the basal portion,being mostly developed into a single, very large up-turned, curvedholding organ, which is on the right side of the apical opening andhas on its mesal side a very distinct groove for the reception of thestinglike ovipositor. This apical opening occupies a sloping posi-tion upon the base of the prolongation or hook, and I can see noindication of a broken off left hook, such as would occur in nana.It is possible, however, that such a hook may have been presentwhen the specimen emerged. Front tarsi normal; claws andpulvilli small. Wings more yellow than brown except around thefirst posterior cell, from the costa to the hind crossvein.Length, 10.6 mm.Described from a single female specimen taken at Gualan, Guata-mala, by W. P. Cockerell (Mrs. T. D. A. Cockerell).ry/?e.?Female, Cat. No. i9960, U.S.N.M.The species nana connects this form with the ordinary typicalmembers of the genus Cylindroniyia- so well that at least until themale is loiown I think it not expedient to recognize the genusApinocyptera. CYLINDROMYIA UNIFORMIS, new species(Fig. 4)This species is almost like euchenor, but is somewhat smaller anddiffers quite decidedly in the shape of the fifth sternite, which cer-tainly must be a specific character. In uniforniis the median exci-sion of the fifth sternite is narrow, only a little rounded at itsbase, and on each side of it there is a rather elongated, narrowlobe bearing a few hairs. This lobe forms a rather acute angle withthe margin of the sternite laterally, and this margin bears at a littledistance three erect bristles of which the hindmost is the largest. ART. 23 GENUS CYLINDROMYIA MEIGEN ALDRICH 25The posterior forceps are smaller and a little less swollen than ineuchenor but of the same general pattern. The outer forceps arealso smaller and a little less robust. In both species the hind edgeof the forceps is grooved at the base.Length, 6.5 to 9 mm.Described from 2 males; 1 in the National Museum collection isfrom San Jacinto, Mexico, Federal District, collected by E. G.Smyth ; the other in C. W. Johnson's collection from San Dominica,AVest Indies.Type.?UfilQ, Cat. No. 28285, U.S.N.M.Paratype in C. W. Johnson's collection.CYLINDROMYIA LIMBATA. new species(Figs. 6, 21)Male.?Parafacials and parafrontals slightly yellowish, the formeras wide as the third antennal joint, the latter narrow above; antennaereddish-brown almost to arista. Mesonotum and scutellum with adecided brassy and metallic reflection. Sternopleurals three. Ab-domen rather brownish in color throughout, the second and thirdsegments only a little reddish on the sides ; the suture between themnarrowly and sharply bordered with white pollen ; the third segmentwith the pollen much more extended; fourth segment with eightbristles; fifth segment with one strong pair directed backwards.The last genital segment mostly yellow, the genitalia greatly resem-bling those of nana, the hind forceps minute, yellow, shining,slender, about two-thirds as long as the anterior, which are alsoyellow and shining, straight with broadly rounded tip. The marginof the last genital segment is also shining yellow, and projects enoughto partially conceal the forceps. Fifth sternite with wide andextremely shallow emargination along the center, at the sides, how-ever, with the usual row of three bristles increasing in size on aslight prominence. Wings brown along the veins and on the anteriorpart. The discal and first posterior cells much lighter in the middle,the brown color following the veins with quite uniform width.Tibia rather dark red (the only species known with this character) ;claws and pulvilli elongated.Female.?The specimen is teneral so that the dark border on thewing veins is only suggested. The second principal segment of theabdomen has on the median line below, just before the middle, aslight swelling with numerous small and somewhat stubby spines,much less distinct than in nana. The fifth segment is reddish, elon-gated, conical, covered with rather dense retrorse hair. The lastsegment consists almost entirely of the hooks, which are shiningblack, the right one stout at the base, slightly notched in profile 26 PROCEEDINGS OF THE NATIONAL MUSEUM Vol.68beyond the middle; the left one deeply grooved at the base as in nanaand swollen on the outer side beyond the groove.Length, 7 to 8 mm.Described from 2 males and 1 female, the males from the CaliforniaAcademy of Sciences, were collected at Mill Creek Canyon, SanBernardino County, California, by E. P. Van Diizee. One of themis retained for the National Museum collection. The female wascollected at Atencingo, Mexico, by E. G. Smyth.Type.?M2i\Q, Cat. No. 28286, U.S.N.M.; paratype, male in Cali-fornia Academy of Sciences.LIST OF DESCRIBED SPECIES OF CYLINDROMYIA FROM NORTHAMERICA, NOT HEREIN IDENTIFIED(All heretofore mentioned under Ocyptera except as noted. The types willhave to be reexamined before the species can be identified ; in the case ofcarolinae this is impossible, as no types are in existence.)hinotata Bigot, Annates Soc. Ent. Fr., 1878, p. 44, female.?Baltimore.carolinae Robineau-Desvoidy, Myodaires, 1830, p. 232 (Parthenia).?Carolina.CoQuiixETT, Revis. Tachin., 1897, p. 86.?Massachusetts to California, etc.A mixture of several si>ecies ; the following references are based onCoquillett's key and equally uncertain.Aldutch, Annals Ent. Soc. Amer., vol. 8, 191.5, p. 82.^?^Widespread.Cbevecobur, Kans. Acad. Sci., vol. 20, 1900, p. 93.?Kansas.Also reported from New Jersey, Florida, and Jamaica (Johnson) ;Ontario (Gibson) ; Quebec (Fyles, and Winn and Beaulieu) ; BritishColumbia (Iline) ; Minnesota (Washburn) ; California and Nevada(Baker) ; Kansas (Snow) ; Montreal (Chajiuon) ; Illinois (Hart) ; NorthCarolina (Brimley) ; Connecticut (Britton).dotadas Walker. List, vol. 4, 1849, p. 694.?Jamaica.epijtus Walker, List, vol. 4, 1849, p. 694.?Georgia.fumipennis Bigot, Annates, 1878, p. 43, male.?Colorado.Bbaueb, Sitzungsber. Kais. Akad. Wiss. Wien., vol. 107, 1898, p. 1, note ontype.signatipenniH Van der Wulp, Tijdsch. v. Ent., vol. 35, p. 187 ; Biologia, Dipt.,vol. 2, 1903, p. 450. pi. 13, fig. 11.?Guerrero, Mexico.simplex Bigot, Annales, 1878, p. 47, female.?Mexico.soror Bigot, Annales, 1878, p. 46.^?Mexico. (The name is preoccupied.)BBAUEaj, Sitzungsber. Kais. Akad. Wiss. Wien, vol. 107, 1898, p. 1, note ontype. EXPLANATION OF PLATE < Figures 2, 3. 7, 8, 12, 22, 31, by C. Howard Curran ; 19, 20, by R. E.grass. The remainder by the autlior) Snod- generally only the posterior andVariously magnified)Plate 1(Figs. 1-14 are male genitalia, side view,anterior forceps being drawn.Fig. 1. Cylindi'omyia brassicaria Fabricius.2. " intermedia Meigen.3. " argentea Townseud.4. " uniformis, new species.5. *' bakeri, new species.6. '* linibata, new species (imperfectly drawn out).7. " dosiades Walker.8. " euchenor Walker.9. " armata, new species.10. nigra, new species.11. " atra Roeder.12. " decora, new species. *13. " alticola, new species.14. " naria Townsend.1"). Scutellum with three pairs of bristles.16. Scutellum with two pairs of bri.stles.17. Scutellum with one pair of bristles.(Female genitalia from the side (19 and 20 diagonally from below),)Fig. is. Cylindromyia signata Townsend.19. U. S. NATIONAL MUSEUM PROCEEDINGS. VOL. 68, ART. 23 PL. I Details of Cylindromyia MeigenFor explanation of plate see paqe 27