PROCEEDINGS OF THE UNITED STATES NATIONAL MUSEUM
issued
SMITHSONIAN INSTITUTIONU. S. NATIONAL MUSEUMVol. 87 Washington : 1939 No. 3075A TAXONOMIC STUDY OF NEOTROPICAL BEETLES OFTHE FAMILY MORDELLIDAE, WITH DESCRIPTIONSOF NEW SPECIES By Eugene Ray
The Morclellidae are an obscure family of small or minute beetlesthat have never commanded much attention, owing to the relativelyinnocuous habits of the adults and larvae. Very little has ever beendone with this group except in a taxonomic way, and not a singlecomplete life history is known.Unfortunately, many of the forms collected in obscure places areknown only from unique specimens. It seems unlikely that taxono-mists will ever arrive at a point where unlimited material will beavailable for all species, and, no doubt, many uniques will remainalone in collections for years to come. However, some of thesesingle individuals are sufficiently distinct from their closest alliesin the matter of actual description and structural characters to per-mit their proposal as new names. It has been the policy of thepresent writer to describe such uniques only when they can be sepa-rated on the basis of characters that are known not to vary in othermembers of the family. Many more, which seem distinct but areclosely allied to known forms, have been set aside without names,because the characters needed for their separation are known to bevariable in certain cases. No description has been made on thebasis of color and pubescent markings alone, since these featuresare often considerably aberrant, but rather, when one or more struc-tural points of differentiation can be found, the color and pattern165175?39 1 271
272 PROCEEDINGS OF THE NATIONAL MUSEUM vou 87
of pubescence are distinct enough to make immediate, superficialseparation possible. In all cases a combination of characters, ratherthan a single one, has been used for specific and generic diagnosis.A peculiar characteristic has been observed in the present study.At the ends of the tarsi are the tarsal claws, which are distinctstructures bearing one or more teeth on their mesal surfaces. Theseteeth apparently vary in number, position, and shape according tothe species being considered. At present it seems premature to statedefinitely that these will form a new character of specific value,but there is apparently a diversity of detail that seems to providemuch the same diagnostic character as that provided by the geni-talia in certain other groups.Closely appressed to the tarsal claws, and scarcely distinguishablewithout being mounted on a slide, is an additional pair of tarsal
"claws," somewhat less heavily chitinized than the other pair. These
"claws" are attached to the pretarsus at the same point as the otherpair. Their specific value is probably somewhat less than that ofthe external pair, owing to their lesser chitinization, which permitsa bending into various shapes before the integument hardens uponemergence of the adult insect" This character has been observedonly in the subfamily Mordellinae, the Anaspinae apparently beingwithout it. In addition, all the Anaspinae that the writer has ob-served have but the single terminal tooth on the tarsal claw.One new genus and 29 new species are herein recorded. Thirtyothers are found to be much more widely distributed than has hith-erto been supposed. Mention of described species is limited to thoseforms known only from restricted localities. One other species istaken out of synonymy.This study has been based primarily on material received fromthe United States National Museum and the Mexican Departmentof Agriculture, together with a small series of specimens from theUniversity of Puerto Rico and the Deutsches Entomologisches In-stitut. The types of all new species are deposited in the NationalMuseum.^ Paratypes, when available, remain in the collection ofthe writer.Grateful acknowledgment is made to officials of the NationalMuseum ; Dr. A. Dampf , chief entomologist of the Mexican Depart-ment of Agriculture; Prof. S. A. Danforth, of the University ofPuerto Rico; and Dr. Walther Horn, director of the DeutschesEntomologisches Institut. My thanks are due to Miss Kathryn Sum-merman and Julius Bosen, graduate students in the Department ofEntomology at the University of Illinois, for their kind assistance
? The types of Mordella militaris, new species, received from the Deutsches Entomo-logisches Institut, are returned to that institution.
TAXONOMIC STUDY OF MORDELLIDAE RAY 273in the preparation of the figures. Finally, I wish to express myindebtedness to Prof. C. L. Metcalf, head of the Department ofEntomology at the University of Illinois, under whose direction thisstudy w^as made. Genus GLIPA LeConteOUpa LeConte, Coleoptera of Kansas and eastern New Mexico, p. 17, 1857.GLIPA HIEROGLYPHICA SchwarzFigure 17, kOUpa hieroglypliica Schwarz, Proc. Amer. Philos. Soc, vol. 17, p. 372, 1878.Tomoxia hilaris Smith, Trans. Amer. Ent. Soc., vol. 10, p. 80, pi. 1, figs. 30, 32,1882.Six specimens: Two from the Dominican Republic, June-July1913 (H. W, Foote, Yale Expedition) ; one from Alhajuelo, Panama,April 12, 1911 (A. Busck) ; one from Porto Bello, Panama, March 4,1911 (A. Busck) ; one from Taboga, Panama, February 23, 1912(A. Busck) ; one from Trinidad Rio, Panama, March 8, 1912 (A.Busck)
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This species, originally described from Florida, has, I believe,been wrongly suppressed as a synonym of G. hilaris (Say) (1835,p. 190) since the publication of the J. B. Smith paper in 1882.Additional specimens, from the Dominican Republic and Panama,as recorded above, tally more closely with the Florida types in theUnited States National Museum than with true specimens of hilaris{vide Barber, on pin label of specimen, det. 1935). Structurally,there can be no doubt that the two species are distinct. Comparisonof specimens of hilaris from Illinois with the above-mentioned exam-ples of hieroglyphica shows distinct differences in the maxillary palpi,antennae, and tarsal claws (fig. 17, k^ I). In addition, the pattern ofthe dermal color and pubescence of the dorsal surface is distinctlydifferent. In my opinion there can be no doubt that hieroglyphicamerits valid specific standing.Genus TOMOXIA CostaTomoxia Costa, Fauna Regni Napoli, Mordellidae, p. 8, 1854.TOMOXIA SPINIFER ChampionTomoxia spinifer Champion, Biologia Centrali-Americana, Coleoptera, vol. 4,pt. 2, pp. 261, 462, pi. 9, figs. 8, 8a, 1S91.Three specimens: One from Turrialba, Costa Rica (Schild andBurgdorf) ; one from Livingston, Guatemala, April 5 (Barber andSchwarz) ; one from Peru, October 3, 1935, found dead in dried woodat Washington, D. C. (H. Y. Gouldman).
274 PROCEEDINGS OF THE NATIONAL MUSEUM vol.87TOMOXIA FULVICEPS ChampionFiGUBE 17, fTonioxia fulviceps Champion, Biologia Centrali-Americana, Coleoptera, vol. 4,pt. 2, p. 262, pi. 11, fig. 10, 1891.Three specimens: One from Cabima, Panama, May 25, 1911 (A.Busck) ; one from Porto Bello, Panama, March 14, 1911 (A. Busck) ;one from San Carlos, Costa Kica (Schild and Burgdorf).TOMOXIA FIEBRIGI, new speciesFigure 17, dThis species is closely allied to auratonotata Eay (1936a, p. 145)but may easily be separated from the latter member of the genus bythe broader scutellum, shorter intermediate tibiae, and the equalityin length between abdominal segments 3 and 4.Length: 3.95 mm.; including anal style, 4.65 mm. Form short,quite robust, subcuneate, broadest at middle of thorax, sides narrow-ing posteriorly. Derm entirely black, except for labial palpi, whichare orange-red. Body densely ,covered with fine, recumbent, silverypubescence, rufobrunneous on elytra.Antennae 1.25 mm. long, reaching middle of lateral pronotal mar-gin; segments 3 and 4 equal; 5-10 each as long as 4, considerablybroadened apically; 11 distinctly longer than 10, oval, sides rounded,apex slightly rounded. Distal segment of maxillary palpi enlarged,form of a scalene triangle, basal angle a right angle, outer marginbroadly rounded, inner margin almost straight, corners rounded, basearcuate, midbasal lobe short, sides angular, base truncate. Scutellumtwice as broad as long, sides slightly angidar, posterior emarginationwanting. Elytra four-tenths longer than broad (2.65 by 1.85 mm.),sides attenuate for their entire length, apices broadly, individuallyrounded. Intermediate tarsi distinctly longer than their tibiae;penultimate segments of anterior and intermediate tarsi simple'.Tarsal claws with four distinct, broad, blunt teeth, distal one longest,and a small subacute, premedian tooth (fig. IT, d). "Median claws"twice as thick at base as medially, apical half of mesal margin withfour thin, very acute teeth, outer edge rounded. Outer spine ofposterior tibiae twice as long as inner one. Anal style twice as longas apical ventral segment (0.7 by 0.35 mm.), very stout, short, evenlyattenuate, broadly truncate at apex.Type locality.?SdiTi Bernardino, Paraguay.Type.~M^\Q, U.S.N.M. No. 52926, a unique specimen collected byK. Fiebriff. ^
TAXONOMIC STUDY OF MORDELLIDAE RAY 275TOMOXIA BREVIPENNIS, new speciesFigures 17, a; 19, cThis species is most closely allied to hinotata, new species, but maybe easily separated from that member of the genus by the greaternumber of elytral spots, the longer, narrower body, the shorter,broader, anterior and intermediate tibiae, the shorter distal segmentof the maxillary palpi, and the narrower scutellum. From latenotataPic (1924, p. 23) it may be separated by the much shorter anal style,the narrower scutellum, the shorter anterior and intermediate tarsi,and the different position of the elytral spots.Length : 6.6 mm. ; including anal style, 8.5 mm. Form moderatelyrobust, elongate, sides narrowing posteriori}^, broadest near middle ofpronotum. Derm piceous, eyes, antennae, mouth parts, anteriorfemora and tarsi, and spurs of posterior tibiae castaneous, remainderof legs and ventral abdominal segments fuscocastaneous. Bodydensely covered with fine, recumbent pubescence, reddish brown ex-cept for the following whitish areas : mixed on head ; covering mar-gins of pronotum broadly, with a curved band extending from an-terior angles into disk and meeting posterior marginal line on eitherside of middle; scutelkim entirely; an indeterminate humeral spotshortly curving into disk, an oblong scutellar spot extending alongsuture, a large, rounded, postmedian, subsutural spot, and a pair ofirregular postmedian spots on middle of disk curving posteriorly,reaching neither suture nor margins ; intermediate and posterior legs,parts of mesosternum and metasternum, and bases of abdominal seg-ments, basal one and anal style broadly, whitish.Antennae 2.1 mm. long, reaching posterior angles of pronotum;segment 4 twice as long as and much broader than 3; 5-10 each aslong as 4 but successively narrower distad ; 11 distinctly longer than10, broadest beyond middle, sides and apex rounded. Distal segmentof maxillary palpi enlarged, with form of a broad scalene triangle,sides curved, apex straight. Pronotum much broader than long (2.8by 2.1 mm.), sides rounded, broadest medially, base arcuate, midbasallobe short, but rounded. Scutellum of a broad, modified triangularshape, twice as broad as long, sides angularly cutting into elytra,apex broadly rounded, subtruncate. Elytra three-fourths longer thanbroad (4.5 by 2.6 mm.), sides attenuate posteriorly, apices individu-ally rounded. Anterior tarsi much shorter than their tibiae; inter-mediate pair equal in length ; three intermediate segments of anteriorand intermediate tarsi dilated, flattened. Tarsal claws bearing threeteeth, distal one longest (fig. 17, a). Outer spur of posterior tibiaeone-half longer than apical ventral segment (1.9 by 1.2 mm.), stout.
276 PROCEEDINGS OF THE NATIONAL MUSEUM VOL, 87evenly attenuate, truncate at apex; last ventral segment twice aslong as penultimate one.Type locality.?Barro Colorado Island, Canal Zone, Panama.ry;?e.?Female, U.S.N.M. No. 52923, collected April 26, 1926, byC. T. Greene. TOMOXIA BINOTATA, new speciesFiGUEE 17, CThis species is most closely allied to innotata Pic (1924, p. 23) andmay be separated from the latter member of the genus by the bino-tate elytra, the peculiar character of the antennae, the broad distalsegment of the maxillary palpi, and the much larger size.Length : 7 mm. ; including anal style, 8.5 mm. Form short, mod-erately robust, subcuneate, sides narrowing posteriorly, broadest nearbase of prothorax. Derm fuscopiceous, femora of anterior and inter-mediate legs, mesal half of distal segment of maxillary palpi, andantennae, fuscocastaneous. Body densely covered with fine recum-bent brownish pubescence, which assumes a violaceous tinge in cer-tain lights; a pair of postmedian, elytral spots as far from suture aswidth of spot and almost toucliing lateral margins, and basal mar-gins of ventral abdominal segments, white.x\ntennae 1.8 mm. long, reaching middle of lateral pronotal mar-gin ; segment 3 slightly longer than 4 ; 5-10 each succeedingly shorterthan preceding, 10 two-thirds as long as 3 ; 11 as long as 4, broadestsubapically, subtriangular in profile, rounded at corners. Distal seg-ment of maxillary palpi enlarged, form of an isosceles triangle, mesaledge straight, outer edge curved, corners rounded. Pronotumbroader than long (3 by 2 mm.), sides rounded, base arcuate, mid-basal lobe short, truncate. Scutellum slightly broader than long,sides angular, apex truncate, not emarginate, apical angles rounded.Elytra two-thirds longer than broad (4 by 2.3 mm.), sides attenuateposteriorly, apices broadly, individually rounded. Intermediate tarsilonger than their tibiae ; penultimate segment of anterior and inter-mediate tarsi simple. Tarsal claws with three distinct, broad teeth,distal one longest, and a large, blunt, premedian tooth (fig. 17, c)."Median claws" twice as thick at base as medially, apical three-fourthsof mesal margin and entire lateral margin broadly rounded. Outerspur of posterior tibiae one-half longer than inner one, densely cov-ered with coarse pubescence, broadest subapically, thence rounded toapex. Anal style two and one-half times as long as apical ventralsegment (1.3 by 0.5 mm.), stout, evenly attenuate, truncate at apex.Type locality.?Barro Colorado Island, Canal Zone, Panama.Type.?K unique male, U.S.N.M. No. 62925, collected July 17,1923, by R. C. Shannon.
TAXONOMIC STUDY OF MORDELLIDAE?RAY 277TOMOXIA SERRICORNIS, new speciesFiGUBE 16, gThis species is most closely allied to Komi Ray (1936b, p. 147)but may adequately be separated from the latter member of thegenus by the more parallel-sided form, difference in color and patternof pubescence, the greater length of the intermediate tarsi and ofthe outer tibial spur, and the shorter elytra.Length : 3.4 mm. ; including anal style, 4.1 mm. Robust, sides sub-parallel; body fuscobrunneous, maxillary palpi and anterior legsflavocastaneous. Surface densely covered with fine, recumbent, whit-ish pubescence, uniformly covering derm, except on disk of elytra,where it is irregularly disposed and not forming definite spots, exceptfor a postmedian, transverse arcuate band.Antennae 1.1 mm. long, reaching base of pronotum, segments 3 and4 large, equal; 5 to 10 subequal, each much broader and longer than4; 11 distinctly longer than 10, apex and sides rounded. Distal seg-ment of maxillary palpi enlarged, ovate, broadest at middle, sides andapex rounded, outer side greatly so (fig. 16, g). Pronotum distinctlybroader than long (1.4 by 1 mm.), apex and sides rounded, basalangles obtuse, base arcuate, midbasal lobe short, truncate. Scutellumbroader than long, sides angular, apical angles obtuse. Elytra one andseven-tenths times as long as broad (2.4 by 1.4 mm.), sides parallel towithin one-fourth of apex, broadly curved posteriorly, apices individ-ually rounded. Intermediate tarsi longer than their tibiae ; penulti-mate segment of anterior and intermediate tarsi simple. Tarsal clawswith two distinct teeth, and a large, thick, blunt, premedian eleva-tion. Outer spur of posterior tibiae three and one-half times as longas inner one. Anal style one-half longer than apical ventral segment(0.7 by 0.48 mm.), stout, apex broadly truncate.Type locality.?Roxborough, Tobago.Type.?K single female, U.S.N.M. No. 52924, taken at the ArchibaldEstate, Roxborough, Tobago, November 6, 1918 (H. Morrison).MORDELLOIDES, new genusThis genus is allied to both Zeamordella Broun (1886, p. 847) andMordella Linnaeus (1758, p. 420). From the former it may be sepa-rated by the angular insertion of the distal segment of the maxillarypalpi, the much longer length of the posterior legs, the thin, filiformtarsi, and the much longer, acute anal style. From Mordella it maybe separated by the semicircular scutellum, the lengthy intermediatetibiae, the angulate anterior angles of the pronotum, the single flat-tened spur of the posterior tibiae, and the truncate apex of the lastventral segment. From Reynoldsiella Ray (1930, p. 184), to which
278 PROCEEDINGS OF THE NATIONAL MUSEUM vol.87genus it is also allied, it may be separated by the presence of a sub-apical ridge on the posterior tibiae, the much narrower shape of thebody, the flat tibial spur, and characters as mentioned in comparisonwith MordelJa above. Mordelloides, Zeamordella^ and Reynoldsiellamay be linked together because of the semicircular scutellum.The single broad, flattened spur of the posterior tibiae is uniqueamong known Mordellidae and will readily separate this form fromany other member of the family.Form suboval, moderately robust, convex. Distal segment of maxil-lary palpi broadly, equilaterally triangular, inserted at the corner.Antennae subserrate, distal segment distinctly longer than penulti-mate segment. Pronotum much broader than long, anterior anglesobtuse, not rounded. Scutellum fairly large, semicircular. Mesoster-num produced posteriorly in the form of a rounded lobe. Posteriorcoxae elongate, contiguous, meeting in a depressed sutural line ; ante-rior margin of latter part closely connected with rounded lobe ofmesosternum. Anal style long, more than four times length of lastventral segment, broadest at its basal fifth, then suddenly narrowedand attenuate to apex, curved ventrad. Intermediate tibiae longerthan their tarsi, the latter filiforta ; posterior tibiae with but a single,broad, flattened, tibial spur, concave on its anterior surface, convexon its posterior surface.The genotype is Mordelloides acuticauda^ new species, describedbelow. MORDELLOIDES ACUTICAUDA. new speciesFigures 17, i; 19, hThe single, flattened spur of the posterior tibiae, the semicircularscutellum, the peculiar antennae, and other generic characters willserve adequately to separate this species from any of its allies thatmight superficially resemble it.Length : 2.3 mm. ; including anal style, 5.8 mm. Form moderatelyrobust, convex, suboval, sides attenuate, narrowing posteriorly, broad-est near base of pronotum. Head castaneous, darkening evenlycaudad, piceous at occipital margin, mouth parts and antennae flavo-castaneous, distal segment of latter somewhat fuscous in certainlights; pronotum fuscous, sides broadly castaneous; elytra fusco-castaneous, a broad, curved, humeral stria extending toward but notreaching the suture, castaneous ; anterior and middle legs, posteriortarsi, and middle of ventral abdominal segments, flavocastaneous
;
posterior tibiae and anal style brunneous. Surface densely coveredwith fine, recumbent pubescence, yellowish on head, pronotum, andelytra, condensed on latter part to form the following fasciae : a curved
TAXONOMIC STUDY OF MORDELLIDAE RAY 279humeral line covering underlying castaneous derm, a transverse, post-median fascia, extending to and along lateral margins but not reach-ing humeri, extending posteriorly along suture and coalescing with apreapical curved fascia that fails to reach lateral margins; a mixedyellowish white on ventral surface and legs.Antennae 0.9 mm. long, scarcely reaching base of lateral prothoraciomargin; segments 3 and 4 equal; 5-10 subserrate, each broadestmedially and each twice as long as 4; 11 broadly rounded on sidesand apex, distinctly broader and twice as long as 10, broadest pre-medially, apparently truncate at apex. Distal segment of maxillarypalpi enlarged, form of an equilateral triangle, sides straight, anglesrounded. Pronotum almost twice as broad as long (1.05 by 0.6 mm.),apical angles obtuse, pointed, sides rounded, basal angles obtuse, basearcuate, midbasal pronotal lobe short, declivious, truncate. Elytraalmost twice as long as broad (1.9 by 1 mm.), sides visibly attenuatefrom base to apex, apices individually rounded. Intermediate tibiaedistinctly longer than their tarsi; penultimate segment of anteriorand intermediate tarsi filiform. Tarsal claws with four distinctteeth (fig. 17, i). Posterior tibiae with a single, broad, flattenedtibial spur, attenuate to apex, two-thirds length of basitarsus, con-cave on anterior surface, convex on posterior surface. Anal stylefour and one-half times as long as apical ventral segment (1.4 by3 mm.), narrow, thin, curved ventrad at its middle, broad at basalfifth, thence narrowing abruptly distad and attenuate to apex; lastventral segment truncate at apex, but one-half longer than penulti-mate one.Type locality.?Tierra Blanca, Veracruz, Mexico.Type.?A unique female, U.S.N.M. No. 52922, collected on Sep-tember 17, 1924, from weeds of the coastal plain, near railway of theIsthmian line (A. Dampf )
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Remarks.?The dermal color of the elytra is changeable accordingto the angle from which it is viewed, a more noticeable castaneoustinge being visible from certain diagonal directions. The descriptionof the elytra was made from a direct dorsal view.Genus MORDELLA LinnaeusMordella Linnaeus, Systema naturae, ed. 10, p. 420, 1758.MORDELLA DIVERGENS, new speciesFiGUKE 19, aThis species is most closely allied to the North American M.lunulata Helmuth (1865, p. 96), but it may be separated from the165173?39 2
280 PROCEEDINGS OF THE NATIONAL MUSEUM vou 87latter member of the genus by the divergent line of pubescence onthe elytra, the rounded mesal edge of the terminal segment of themaxillary palpi, and the larger size.Length : 5.4 mm. ; including anal stjde, 6.8 mm. Moderately ro-bust, subcuneate; dermal color black; surface densely covered withfine, recumbent pubescence, partaking of ground color, except forthe folloAving whitish areas: head and pronotum completely, con-densed on latter part to form three short, longitudinal, basal bands;scutellum completely; elytra with humeri narrowly and two diver-gent stripes, anterior extending submarginally to within one-sixth ofapex and separated from margin by its own width, interior one ex-tending diagonally across disk and joining whitish suture subapically,sides of mesosternum and metasternum densely, bases of abdominalsegments, and legs somewhat less distinctly.Antennae 1.4 mm. long, scarcely reaching posterior coxae; seg-ments 3 and 4 equal in length ; 5-10 each one-half longer than 4 andconsiderably broadened; 11 rounded, one-half longer than 10. Distalsegment of maxillary palpi enlarged, subtriangular, rounded onmesal surface and apex. Pronotum one-third broader than long(2.1 by 1.6 mm.), convex, sides finely margined, basal angles obtuse,midbasal lobe short, broad, subtruncate. Scutellum twice as broadas long, rounded-triangular. Elytra broadest subbasally, almosttwice as long as broad (3.8 by 2.1 mm.), sides attenuate to apex,latter parts individually rounded, distinctly obtuse. Intermediatetibiae and tarsi equal in length. Tarsal claws with four teeth, allsomewhat blunt at ends, proximal one rudimentary. Anal style 3times length of apical ventral segment, narrowly truncate at apex.Type locality.?Mount Puilsboreau, Haiti.Type.?K unique male, U.S.N.M. No. 52930, collected on May 291927, by G. N. Wolcott.MORDELLA LATITHORAX, new specie*This species is somewhat allied to Uformis Champion (1891, p. 295)but differs from the latter member of the genus in numerous particu-lars, the most striking of which are the decidedly transverse prono-tum, the equilateral condition of the distal segment of the maxillarypalpi, the much narrower and more elongate antennae, and the dif-ferent color.Length : 3.5 mm. ; including anal style, 4.8 mm. Form moderatelybroad, sides parallel posteriorly to middle of elytra, thence broadlyrounded to apex; body broadest near base of pronotum. Head cas-taneous, with a piceous rounded spot on vertex reaching occiput andextending laterally in a narrow line to eyes, anteriorly not extending
TAXONOMIC STUDY OF MORDELLIDAE?RAY 281beyond middle of eyes ; pronotum piceous, with a pair of castaneousspots on either side of middle, separated by a piceous line, extendingposteriorly along lateral margins, but not reaching base; elytrapiceous, with apex narrowly castaneous; mesosternum and metaster-num, posterior femora and tibiae, and distal half of anal style fus-copiceous; antennae, mouth parts, anterior and intermediate legs,posterior tarsi, abdomen, and base of anal style castaneous. Surfacedensely covered with fine, recumbent pubescence, partaking of groundcolor, except on dorsal surface, wdiere it has been abraded and onlya few yellowish hairs remain.Antennae 1.1 mm. long, slender, reaching base of lateral pronotalmargin ; segment 4 one-half longer than 3 ; 5-10 each one-third longerthan 4 and but slightly broader; 11 missing. Distal segment of max-illary palpi enlarged, with form of an equilateral triangle. Prono-tum six-tenths broader than long (1.6 by 1 mm.), apex and sidesbroadly rounded, basal angles obtuse, base arcuate, midbasal lobemoderately produced, rounded. Scutellum large, broadly triangular,sides but slightly curved, apical angle rounded. Elytra but seven-tenths longer than broad (2.5 by 1.45 mm.), sides parallel on basalhalf, thence strongly curved to apex, apices individually rounded.Intermediate tibiae and tarsi approximately equal in length, secondand third segments of latter part short, somewhat dilated, penulti-mate segment considerably broadened, fifth segment being insertednear base of latter and only claws visible beyond distal margin.Outer spur of posterior tibiae two and one-half times as long as innerone. Anal style two and one-half times as long as apical ventralsegment (1.3 by 0.5 mm.), moderately thick at base, attenuate toapex; last ventral segment eight times length of penultimate one.Type locality.?Limon Plantation, Chagres Eiver, Panama.Type.?Male, U.S.N.M. No. 52927, a unique individual taken onJuly 14, 1918, from sweepings around a cornfield (H. F. Dietz andJ. Zetek).Remarks.?The three distal segments of the posterior tarsi aremissing in the type and no mention can therefore be made of thetarsal claws. MORDELLA SCUTELLARIS FabriciusMordella scutellaris Fabricius, Systema eleutheratorum, vol. 2, p. 123. 1801.Five specimens : One from Arecibo, Puerto Rico, November 1, 1932f,on flower of Gurte amor; one from Ponce, Puerto Rico, August 21,1933, on flower of Senegaliu (R. G. Oakley) ; one from Bayamon,Puerto Rico, June 2, 1934, on milkweed (Lesesne et al.) ; two fromArecibo, Puerto Rico, March 13, 1934, and March 21, 1935, on Mar-garita (Anderson).
282 PROCEEDINGS OF THE NATIONAL MUSEUM vol.87MOEDELLA HAITIENSIS, new speciesFigure 16, oIn addition to its difference in color pattern this species may beseparated from its closest ally, dimidiata Champion (1891, p. 234),by the greater length of the fourth antennal segment, the emarginateapex of the distal segment of the maxillary palpi, and the very longlast ventral abdominal segment.Length : 2.75 mm. ; including anal style, 4 mm. Moderately robust,sides attenuate, narrowing posteriorly, broadest at middle of pro-notum. Head piceous, antennae and mouth parts flavous; prothoraxflavobrunneous ; elytra piceous, with a diagonal, quadrangular, flavousspot on disk near base, separated from lateral margin by less thanone-half its width ; ventral surface black ; anterior legs flavous, inter-mediate pair fuscobrunneous. Surface densely covered with fine, re-cumbent pubescence, yellowish white, except on elytra, where it formsgolden areas, as follows: covering subbasal flavous spot and entireapical half, whence it extends in a broad line along suture to base.Antennae 0.37 mm. long, reaching base of pronotum, segment 4 dis-tinctly longer than 3; 5-10 serfate, each one-half longer than 4 andmuch broader at apex, slightly decreasing in length distally; 11 dis-tinctly longer 10, broadest at apical third, sides and apex broadlyrounded. Distal segment of maxillary palpi enlarged, form of a sca-lene triangle, outer side longest, apical angle emarginate (fig. 16, a).Pronotum distinctly broader than long (1 by 0.78 mm.), apex andsides rounded, basal angles obtuse, base arcuate, midbasal lobe veryshort, truncate. Scutellum small, triangular. Elytra twice as long asbroad (2 by 1 mm.), sides visibly attenuate on their apical two-thirds,broadly curved subapically, apices individually rounded. Interme-diate tarsi distinctly longer than their tibiae ; penultimate segment ofanterior and intermediate tarsi simple. Tarsal claws with three dis-tinct teeth. Outer spur of posterior tibiae twice as long as innerone. Anal style one-half longer than apical ventral segment (1.2 by0.82 mm.), narrow, attenuate to apex; last ventral segment seven timesas long as penultimate one.Type locality.?Mariani, Haiti.ry/je.?Male, U.S.N.M. No. 52929, a unique individual taken onSeptember 17, 1918, by W. A. Hoffman.MORDELLA VIDUA SolierMordcVa vidua Solier, in Gay, Historia fisica y politica de Chile, vol. 5, p. 270,1851.Three specimens : Two from Angol, Chile, January 14, 1928 (D. S.Bullock) ; one from Chile (E. C. Reed).
I
TAXONOMIC STUDY OF INIORDELLIDAE RAY 283MORDELLA MILITARIS, new speciesThis species is most closely allied to ogloUini Pic (1930, p. 29)and may be separated from the latter member of the genus by theabsence of distinct maculations on the elytra, the more slender, atten-uate anal style, and the smaller size. In the synopsis of the Argen-tinian species of Mordella (Pic, 1936, pp. 114, 115) jnilitaiis keys outto oglohlini.Length: (male) 5.85 and (female) 7.35 mm.; including anal style,(male) 8.1 and (female) 9.35 mm. Elongate, sides narrowing pos-teriorly to apex of elytra, body subcuneate in shape; dermal colorpiceous, except for basal three-fourths of elytra, which are golden-brown. Surface densely clothed with fine, recumbent pubescence, par-taking of gTound color, except for the following whitish areas : head
;
pronotum, densest on lateral and basal margins ; elytra, condensed onsuture and apex of golden-brown area; mesosternum and meta-sternum; basal margins of ventral abdominal segments, densest atlateral edge ; base of anal style ; legs.Antennae (male) 2.2 and (female) 2.4 mm. long, reaching posteriorcoxae ; segments 3 and 4 equal in length ; 5-10 distinctly serrate, eachslightly longer and twice as broad as 4; 11 rounded, slightly longerthan 10. Distal segment of maxillary palpi enlarged, with form of ascalene triangle, outer margin distinctly curved, cephalic surface witha median, elongate depression. Prothorax distinctly broader thanlong (1.6 by 1.5 mm.), sides rounded, base arcuate, basal lobe short,subtruncate. Scutellum small, subtriangular, sides and apical anglerounded. Elytra twice as long as broad (3.6 by 1.8 mm.), sidesstraight, attenuate to apex, apices individually rounded. Tarsalclaws with seven teeth, the distal one longest, successively shorteningproximad, the proximal one scarcely more than rudimentary. Analstyle slightly more than twice as long as apical ventral segment,slender from a dorsal view, attenuate to apex.Type locality.?Argentina.Type.?Male, in the collection of the Deutsches EntomologischesInstitut.Remarks.?A male and female example (type and allotype), withthe data merely as "Argentina?Coll. Kraatz," contains all availableinformation. There is no visible external difference between the sexes.The middle tarsi are missing in both specimens.MORDELLA SUMMERMANAE, new speciesThis species cannot be closely allied with any of the known NorthAmerican forms. The form of the body, with its subovate form, theextremely short anal style, the short elytra, and the narrow distal
284 PROCEEDIISrGS OF THE NATIONAL MUSEUM VOL. 87segment of the maxillary palpi separate it at once from any speciesrecorded in the literature. For systematic position it may be placednear seriata Champion (1891, p. 298).Length : 2.2 mm. ; including anal style, 2.7 mm. Form short, broad,subovate, broadest near base of pronotum ; dermal color black, anten-nae and legs fuscous, spurs of posterior tibiae and genitalia flavous.Surface densely covered with fine, recumbent, yellowish-brown pubes-cence, except on ventral abdominal segments, where it is whitish.Antennae 0.5 mm. long, scarcely reaching base of pronotum; seg-ments 3 and 4 equal in length; 5-10 each as broad as long, stronglyserrate, broadest apically; 11 distinctly longer than 10, rounded onsides and apex, broadest medially; distal segment of maxillary palpienlarged, with form of an elongate isosceles triangle, sides slightlycurved, corners rounded. Pronotum distinctly broader than long(0.85 by 0.6 mm.), broadest subbasally, apex and sides rounded, basalangles almost right angles, base arcuate, midbasal lobe short, butrounded. Scutellum small, broadly triangular, sides visibly curved,apical angle rounded. Elytra less than twice as long as broad (1.6 by0.85 mm.), sides distinctly curved from base to apex, apices indi-vidually rounded. Anterior and intermediate tibiae longer than theirtarsi, second and third segments of latter part short, but not dilated.Tarsal claws with three teeth, all blunt, basal one obtuse, very muchflattened. Anal style short (0.5 by 0.4 mm.), but one-fourth longerthan apical ventral segment, very broad at base, but one-third longerthan broad from a dorsal view, sides straight, apex rounded.Type locality.?Constanza, Dominican Republic.Type.?A^ unique female, U.S.N.M. No. 52928, taken on May 22,1927, by A. Wetmore.MORDELLA FLAVOFASCIATA ChampionMordella flavofasciata Champion, Biologia Centrali-Americana, Coleoptera, vol. 4,pt. 2, p. 287, pi. 11, flg. 18, 1891.One specimen: Potrero, Veracruz, Mexico, October 28, 1924, onweeds, 600 meters elevation (A. Dampf).MORDELLA SERIATA ChampionMordella seriata Champion, Biologia Centrali-Americana, vol. 4, pt. 2, p. 298, pi.13, fig.s. 9, 9a, 1891.One specimen : Finca Vergel, Huixtla Valley, Chiapas, Mexico, May15, 1935, 700 meters elevation (A. Dampf).Genus CALYCE ChampionCalyce Champion, Biologia Centrali-Americana, vol. 4, pt. 2, p. 307. 1891.
TAXONOMIC STUDY OF MORDELLIDAE?RAY 285CALYCE MACULATA PicCalyce maculata Pic, L'lSchange, vol. 27, p. 191, 1911.One specimen: Trinidad (A. Busck).CALYCE FULVA ChampionCalyce fulva Champion, Biologia Centrali-Ainericana, Coleoptera, vol. 4, pt. 2,p. 307, pi. 13, figs. 21, 21a, 1891.One specimen: San Carlos, Costa Kica (Schild and Burgdorf).Genus GLIPODES LeConteGlipodes LeConte, Proc. Acad. Nat. Sci. Philadelphia, vol. 14, p. 48, 1862.GLIPODES SERICANS (Melsheimer)Mordella sericans Melsheimer, Proc. Acad. Nat. Sci. Philadelphia, vol. 2, p. 312,1846.Olipodes sericans LeConte, Proc. Acad. Nat. Sci. Philadelphia, vol. 14, p. 48, 1862.Two specimens : One from Cacao, Trece Aguas, Alta V. Paz, Guate-mala, March 27 (Schwarz and Barber) ; one from Paraiso, CanalZone, Panama, April 29, 1911 (E. A. Schwarz).GLIPODES HELVA LeConteOlipodes helva LeConte, Proc. Acad. Nat. Sci. Philadelphia, vol. 14, p. 48, 1862.Three specimens: One from Xolitla, Guererro, Mexico, August 3,1930, at light (J. Parra) ; one from Trinidad Rio, Panama, June 9,1912 (A. Busck) ; one from Cano Saddle, Gatun Lake, Panama, May3, 1923 (R.C. Shannon).Genus CONALIA Mulsant and ReyConalia Mulsant and Rey, Ann. Soc. Agr. Lyon, ser. 3, vol. 2, p. 313, ]858.CONALIA EBENINA ChampionConalia elenina Champion, Biologia Centrali-Americana, Coleoptera, vol. 4, pt. 2,p. 306, pi. 13, figs. 20, 20a, b, c, 1891.Conalia fulvoplagiata Champion, Trans. Ent. Soc. London, 1896, p. 50.
?
Ray,Coleopt. Contr., vol. 1, pt. 3, p. 163, 1930.Twenty-nine specimens: One from San Rafael, Jicaltepec, Vera-cruz, Mexico (Townsend) ; one from Montemorelo, Nuevo Leon, Mex-ico, June 3, 1931, grass-citrus, 432 meters elevation (A. Dampf ) ; fivefrom Santa Anita, Jalisco, Mexico, August 7, 1927, on weeds, 2,000meters elevation (A. Dampf) ; one from Plancha Piedra, Peten De-})artment, on the border between British Honduras and Guatemala,October 27, 1925, swept from weeds (A. Dampf) ; one from La Ceiba,Honduras, August 27, 1916 (F. J. Dyar) ; one from Cacao, TreceAguas, Alta Vera Paz, Guatemala, February 25 (Schwarz and Bar-ber) ; one from Gamboa, Canal Zone, Panama, July 17, 1918, sweep-
286 PROCEEDINGS OF THE ISTATIONAL MUSEUM vol.87ings (H. F. Dietz and J, Zetek) ; two from Pueblo Buevo, PapayaPlantation, 4 miles from Panama City, Panama, August 1926 (H.Morrison) ; three from Limon Plantation, Cliagres River, Pan-ama, July 14, 1918, sweepings around cornfield (H. F. Dietzand J. Zetek) ; one from Montserrat, Trinidad, June 29 (A.Busck) ; two from DAbadie, Trinidad, October 13, 1918 (H. Mor-rison) ; one from Department of Agriculture grounds, Port-of-Spain, Trinidad, November 23, 1918 (H. Morrison) ; three fromTobago Island, Panama, September 21, 22, 1918 (H. F. Dietz) ; onefrom La Romana Centrale, Dominican Republic, July 15, 1917 (H.Morrison) ; one from San Pedro de Maoris, Dominican Republic, July15, 1917 (H. Morrison) ; one from Tucuman, Argentina (G. F.Mozuetta) ; and three from San Bernardino, Paraguay.Genus MORDELLISTENA CostaMordelUstena Costa, Fauna Regni Napoli, Mordellidae, pp. 16, 31, 1854.MORDELLISTENA CRUX ChampionMordellistena crux Champion, Biologia Centrali-Americana, Coleoptera, vol. 4, pt.2, p. 346, pi. 15, figs. 21, 21a, 1891.Two specimens: One from Tampico, Mexico, December 8 (E. A.Schwarz) ; one from Cacao, Trece Aguas, Alta Vera Paz, Guatemala,March 26 (Schwarz and Barber).MORDELLISTENA BICINCTELLA LeConteMordellistena Mcinctella LeConte, Proc. Acad. Nat. Sci. Philadelphia, vol. 14,p. 48, 1862.Five specimens : All from Trece Aguas, Alta Vera Paz, Guatemala(Barber and Schwarz), on the following dates: December 12 (1),March 26 (1), April 5 (2), no date (1).MORDELLISTENA RHENOLEPIS, new speciesFiGXiREs 16, c; 18, aThis species is allied to ozarJcensis Ray (1936a, p. 125) and may beseparated from the latter member of the genus by the additionalridge on the posterior tibiae, the completely flavocastaneous conditionof the head and pronotum, the more parallel form, and the longer,hammerlike distal segment of the maxillary palpi. Among the Cen-tral American forms rhenolepis is most closely allied to leata Cham-pion (1891, p. 347) and may easily be separated by the unicoloredpronotum, the difference in the number of comblike ridges on theposterior tibiae and tarsi, the different position of the basal cinereousvitta on the elytra, and the greater length of the fourth antennalsegment.
TAXONOMIC STUDY OF MOEDELLIDAE?RAY 287Length: 2.3 mm.; including anal style, 3.2 nun. Form narrow,elongate, sides subparallel to within one-fourth of apex of elytra,broadest near base of pronotum. Head, pronotum, antennae, mouthparts, legs, last two ventral abdominal segments and anal style, middleof mesosternum and metasternum, and a rounded, oblique, subbasalspot on elytra reaching lateral margins but not suture, flavocastane-ous; elytra, sides of mesosternum and metasternum, basal abdominalsegments, and eyes, piceous-black. Body densely covered w^ith fine,j'exiumbent pubescence, partaking of ground color, except on elytra,where it forms the following whitish vittae: covering subbasalrounded spot mentioned above, and broad, transverse, postmedianvitta, reaching both lateral margins and suture, and prolonged an-teriorly near suture to form a blunt, rounded area.Antennae 0.7 mm. long, filiform, reaching metacoxae; segments 3and 4 equal in length, latter somewhat broader; 5-10 each one-halflonger and nuicli broader than 4; 11 one-third longer than 10, sidesand apex rounded, broadest postmedially. Distal segment of maxil-lary palpi enlarged, subrectangnlar, slightly narrower subapically,lateral margin slightly convex, mesal margin slightly concave,apical margin subangulate, corners at both apex and base rounded,two and two-thirds times as long as broad (fig. 16, c). Pronotum one-third broader than long (0.8 by 0.6 mm.), sides and apex rounded,basal angles obtuse, base arcuate, midbasal lobe short, but rounded.Scutellum moderately large, rounded-triangular, broader than long,sides and apical angle rounded. Elytra slightly more than twice aslong as broad (1.7 by 0.8 nmi.), sides broadest at base, subparallel towithin a fourth of apex, thence strongly rounded, apices individuallyrounded. Intermediate tibiae and tarsi approximately equal inlength ; penultimate segment of anterior and intermediate tarsi short,filiform, simple. Posterior tibiae with two short, oblique ridges,neither extending more than one-third across outer surface ; basitarsiwith two short ridges extending one-third across outer surface.Tarsal claws with three long, acute, postmedian teeth (fig. 18, a).Anal style two and one-fourth times length of apical ventral segment(0.9 by 0.4 mm.), narrow, evenly attenuate to apex; last ventral seg-ment about twice length of penultimate one.Type locality.?Guatemala.ry/>6.?Male, U.S.N.M. No. 52938.R&nuvrks.?The type and two male paratypes, all with protrudinggenitalia, were taken at Washington, D. C, October 17, 1916, uponemergence from the rhizome of Rhenolepis sp. received fi"om Guate-mala (H. L. Sanford)
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288 PROCEEDINGS OF THE NATIONAL MUSEUM vol, 87MORDELLISTENA HONDURENSIS, new speciesFiGtJKE 16, I; 18, 6This species is most closely allied to sehioarzi^ new species, describedbelow, and may be separated from the latter member of the genus bythe difference in the maculation of the elytra, the difference in thebreadth of antennal segments 3 and 4, the peculiar shape of the distalantennal segment, the additional tooth on the tarsal claws, and thepeculiar condition of the penultimate segment of the anterior andintermediate tarsi. From xanthopyga Champion (1891, p. 344),which it resembles in color, it may be distinguished by the shorterlength of the basal ridge of the posterior tibiae, the flavobrunneouscondition of the pronotum, and the difference in the shape of themaxillary palpus.Length: 0.9-1.15 mm.; including anal style, 1.15-1.55 mm. Formelongate, narrow, sides subparallel, narrowing only one-fourth fromapex of elytra, broadest near middle of pronotum. Head (includingantennae and mouth parts), pronotum, apex and middle ofmetasternum, last two ventral abdominal segments, and anal style,flavobrunneous; elytra piceous, with following flavous areas: a sub-triangular, rounded, subbasal area reaching sides but not suture anda broad, transverse, postmedian fascia extending entirely across diskfrom lateral margins to suture. Surface densely covered with fine,recumbent pubescence, partaking of ground color, a mixture offlavous and piceous hairs on head and pronotum, giving these partsa darker tinge, except along margins.Antennae 0.28-0.4 mm. long, reaching metacoxae, somewhatthickened but not serrate; segments 3 and 4 equal in length; 5-10each almost as long as 3 and 4 together, and distinctly broader; 11one-half longer than 10, broadest medially, outer edge straight, mesalmargin rounded from middle to apex, the latter subacute. Distalsegment of maxillary palpi enlarged, subquadrate, mesal marginstraight, lateral and apical margins broadly rounded (fig. 16, I).Pronotum but slightly broader than long (0.28-0.35 by 0.25-0.3 mm.),sides rounded, basal angles obtuse, base arcuate, midbasal lobe short,rounded. Scutellum small, broadly triangular. Elytra slightly lessthan two and two-thirds times as long as broad (0.65-0.8 by 0.28-0.35mm.), sides parallel to within one-fourth of apex, thence attenuateand broadly rounded to apex, apices individually rounded. Inter-mediate tibiae and tarsi equal in length; penultimate segment ofanterior and intermediate tarsi filiform, excavate beneath, bearingfifth segment on middle of ventral surface. Posterior tibiae withtwo oblique ridges, basal one extending three-fourths, other one-half,across outer surface; basitarsi with two, second segment with one
TAXONOMIC STUDY OF MORDELLIDAE?RAY 2^9
oblique ridge, all extending halfway across outer surface. Tarsalclaws with four distinct teeth, basal one shorter than others andobtuse (fig. 18, 5). A single spur on posterior tibiae, thick, two-thirds as long as basitarsus. Anal style almost three times as longas apical ventral segment (0.28-0.4 by 0.1-0.15 mm.), fairly stout,attenuate to apex; last ventral segment but twice as long aspenultimate one.Type locality.?La Ceiba, Honduras.rype.?Male, U.S.N.M. No. 52913.Remarks.?The type was collected at La Ceiba, Honduras, August25, 1916 (F. J. Dyer). A male paratype was taken at Peto, Yucatan,Mexico, August 9, 1925, at about 30 meters elevation, in a dense bushforest (A. Dampf ) . This latter specimen has the pronotum fuscopice-ous, except for a small flavous area at middle of lateral margin, theseven apical segments are slightly darker than in the type, and thebasal elytral spot is smaller, so that it does not reach the lateralmargin. Structurally the paratype, even including the nature, dis-position, and length of the teeth on the tarsal claws, is identicalwith the type. MORDELHSTENA SCHWARZI, new speciesFigures 16, & ( 5 ), d ( 9 ) ; 18, dThis species may be separated from both diluta Champion (1891,p. 338) and emarginata., new species, described below, by the complexmaculation of the elytra, the very peculiar shape of the distal seg-ment of the maxillary palpi in the male, the much greater lengthof the second over the first antennal segment, the equal length butgreater width of segment 4 over segment 3, the longer com-parative length of the apical ventral segment, and a distinct differ-ence in the form of the tarsal claws.Length: 1.07 mm.; including anal style, 1.5 mm. Elongate, nar-row, sides subparallel, narrowing only one-fourth from apex ofelytra, broadest at middle of pronotum. Head (including mouthparts and antennae), pronotum, and elytra, flavocastaneous, latterpart with the following piceous areas : a triangular humeral patch,a Y-shaped sutural area at base narrowly connected to humeralpatch, a broad marginal spot at middle extending halfway to suture,and an ai-ea covering apical sixth; ventral surface flavocastaneous,abdomen slightly darker (base and sides of mesosternum and sidesof metasternum up to a point parallel with posterior coxae, piceocas-taneous in female) ; ridges of posterior tibiae and tarsi piceous. Sur-face densely covered with fine, recumbent pubescence, partaking ofground color.
290 PROCEEDINGS OF THE NATIONAL MUSEUM vol.87Antennae 0.35 mm. long, reaching middle of lateral prothoracicmargin ; segment 2 one-half longer than 1 ; 4 equal in length to 3 butbroader; 5-10 filiform, each one-half longer than 4 and distinctlybroader; 11 longer than 10, sides subparallel, rounded from apicalthird to apex. Distal segment of maxillary palpi enlarged, showinga sexually dimorphic form, viz: in male, two anteapical segmentsthickened, distal one oblong-ovate, sides subparallel, angles allrounded (fig. 16, h) \ in female, anteapical segments normal, thin,distal one with the form of an elongate scalene triangle, apicalangle acute, mesal angle rounded, broadest at apical third (fig. 16,d). Eyes large, coarsely faceted, entire, without emargination.Pronotum slightly broader than long (0.32 by 0.26 mm.), apex andsides rounded, basal angles obtuse, base arcuate, midbasal lobe short,but rounded. Scutellum small, broadly triangular. Elytra abouttwo and two-thirds times as long as broad (0.8 by 0.32 mm.),sides parallel to within one-fourth of apex, thence attenuate andbroadly rounded to apex, apices individually rounded. Interme-diate tibiae and tarsi long, equal in length ; penultimate segment ofanterior and intermediate tarsi ?imple. Posterior tibiae with twooblique ridges extending halfway across outer surface ; basitarsi withtwo, second segment with one oblique ridge, all extending halfwayacross outer surface. Tarsal claws with three distinct teeth (fig.18, d). A single spur visible on posterior tibiae, thick, three-fourthsas long as basitarsus. Anal style three times as long as apical ven-tral segment (0.45 by 0.15 mm.), narrow, attenuate to apex; lastventral segment two and one-half times as long as penultimate one.Typ^ locality.?Paraiso, Canal Zone, Panama.7'y;?e.?Male, U.S.N.M. No. 52914.Remarks.?Both the type and allotype were taken at Paraiso,Canal Zone, Panama, January 25 and 30, 1911 (E. A. Schwarz). Asnoted above there exists a distinct sexual dimorphism in the differ-ence of the color of the ventral surface and in the shape of the distalsegment of the maxillary palpi.MORDELLISTENA SPARSA ChampionMordelUstena sparsa Champion, Biologia Centrali-Americana, Coleoptera, vol. 4,pt. 2, p. 335, pi. 15, figs. 4, 4a, 1891.Five specimens: District Federale, Mexico (L. Conradt).
TAXONOMIC STUDY OF MORDELLIDAE?RAY 291
Figure 16.?Maxillary palpi of Mordella, Mordellistena, Glipodes, and Tomoxia.
a, Mordella haitiensis, new species; h, Mordellistena schwarzi, new species, male; c, M.rhenolepia, new species ; d, M. schwarzi, new species, female ; e, M. funereaChampion; f, Glipodes sericans (Melslieimer), male; (/, Tomoxia serricornis, newspecies ; h, Mordellistena laticomis, new species ; /, M. ioseni, new species ; /, M.dampfi, new species ; k, M. candelabra, new species ; I, M. hondurensis, new species.
292 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 87
FIGURE 17.?Tarsal claws Tomoxia, Calyck, Mokdeli.oiues. Morublla, and Glipa.a, Tomowia brevipennis, new species; b, T. bidentata (Say) ; c, T. binotata, new species;d, T. fiebrigi, new species; e, T. spinifer Champion; /, T. fulviceps Champion;g, Oalyce maculata Pic; ft, O. fulva Champion; i, Mordelloides acuticauda, newgenus and species ; ;, Mordella 4-signata Chevrolat ; k, Glipa hieroglyphica Schwarz
;
I, O. hilaris (Say).
I TAXONOMIC STUDY OF MORDELLIDAE?RAY 293
mFiGDEB 18.?Tarsal claws of Moedkllistbna.
a, Mordellistena rhenolepis, new species ; 6, M. hondurensis, new species ; c, M. emarffi-ndta, new species ; <?., M. sciiwarzi. new species; e, M. damvfi. new species; f. M.sonorensis, new species ; a, M. candelabra, new species ; h, M. decora, new species
;
i, M. ornatipennis, new species ; /, M. latipalpalis, new species ; k, M. laticortvis, newspecies ; I, M. l)o.<teni, new species ; m, M. flavoooUaris, new species ; n, M. chiapenHs,new species.
294 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 87
PiOTJHB 19.?Dorsal views of Mordella, Mordelloides, and Tomoxia, and tarsal claws ofMORDELLA and MORDELLISTENA.Dorsal view of: a, Mordella divergena, new species; 6, Mordelloides acuticauda, newgenus and species; c, Tomoxia hrevipennis, new species. Tarsal claws of: d.Mordella clavicornia Kirby ; e, MordelUstena sepia, new species.
TAXONOMIC STUDY OF MORDELLIDAE?BAY 295MORDELLISTENA EMARGINATA, new speciesFiGtniE 18, VThis species is closely allied to dilufa Champion (1891, p. 338) butmay easily be separated from the latter member of the genus by themuch smaller size, the equality in length between antennal segments3 and 4, the filiform condition of segments 5-10, and the presence ofan emargination at the frontal edge of the eye.Length: 1.1 mm.; including anal style, 1.4 mm. Elongate, narrow,sides subparallel, narrowing only at apical fourth of elytra, broadestat middle of pronotum. Head (including antennae and mouth parts)and pronotum flavotestaceous, latter with a narrow, dark brown linealong base and posterior half of lateral margin; elytra flavobrun-neous, somewhat darker along suture and sides; ventral surface fla-vopiceous, three apical ventral segments flavobrunneous ; anterior andintermediate legs flavotestaceous, posterior pair flavobrunneous;ridges of posterior tibiae and tarsi piceous. Surface densely cov-ered with fine, recumbent pubescence, partaking of ground colorexcept on dark areas of ventral surface, where it is flavous.Antennae 0.35 mm. long, almost reaching base of abdomen; seg-ments 3 and 4 small, equal; 5-10 filiform, each almost as long as 3and 4 together, but each slightly broader than 4; 11 distinctly longerthan 10, broadest at middle, rounded to apex. Distal segment ofmaxillary palpi enlarged, fairly robust, elongate-ovate, broadest atmiddle. Eyes large, coarsely gi^anulated, with a distinct emargina-tion on frontal edge near antennal declivity. Pronotum slightlybroader than long (0.35 by 0.28 mm.), apex and sides rounded, basalangles obtuse, base arcuate, midbasal lobe moderately long, rounded.Scutellum small, broadly triangular. Elytra about two and one-halftimes as long as broad (0.8 by 0.28 mm.), sides parallel to within one-fourth of apex, thence attenuate and broadly rounded to apex, apicesindividually rounded. Intermediate tibiae and tarsi equal in length
;
penultimate segment of anterior and intermediate tarsi simple. Pos-terior tibiae with two oblique ridges extending halfway across outersurface; basitarsi with two, second segment with one oblique ridge,all extending halfway across outer surface. Tarsal claws with threedistinct teeth and a premedian, blunt, very small elevation (fig. 18,c). Outer spur of posterior tibiae three times as long as inner one.Anal style three times as long as apical ventral segment (0.3 by 0.1mm.), narrow, attenuate to apex; last ventral segment but slightlylonger than penultimate one.Type locality?La Ceiba, Honduras.Type?A unique male, U.S.N.M. No. 52915, taken on October 18,1916, by F. J. Dyer,
296 PROCEEDINGS OF THE NATIONAL MUSEUM vou 8tMORDELLISTENA TESSELLATA ChampionMordellistena tessellata Champion, Biologia Centrali-Americana, Coleoptera,vol. 4, pt. 2, p. 336, pi. 15, figs. 5, 5a, 1891.Five specimens : One from Antigua, Guatemala, May 2, 1926 (J. M.Aldrich) ; one from Monterey, Nuevo Leon, Mexico, November 25,1909 (F. C. Bishopp) ; two from Coapa, District Federale, Mexico,June 12, 1922 (E. G. Smyth) ; one from Mexico City, Mexico (O. W.Barrett)
.
MORDELLISTENA DAMPFI, new speciesFiGUBES 16, j; 18, eThis species is allied to diluta Champion (1891, p. 338) and may beseparated from the latter member of the genus by the black head, theequality in length between antenna! segments 3 and 4, the elongate-triangular shape of the distal segment of the maxillary palpi, themuch smaller size, and the peculiar condition of the tarsal claws.Length: 0,8 mm.; including anal "style, 1.18 mm. Form elongate,narrow, sides subparallel, narrowing approximately one-fourth fromapex of elytra, broadest near middle of pronotum. Head piceousblack, thorax, elj'tra, antennae, mouth parts, and legs, flavocastaneous,sides of mesosternum and metasternum and abdomen (except analstyle and apical ventral segment) piceocastaneous. Body denselycovered with fine, recumbent, flavous pubescence, thickest on analstyle, where it gives this structure a coarse appearance.Antennae 0.25 mm. long, slender, filiform; segments 3 and 4 equalin length, latter distinctly broader of two; 5-10 each as long as 3and 4 together, each twice as thick as 3, with straight sides; 11 dis-tinctly longer than 10, apically rounded. Distal segment of maxil-lary palpi enlarged, with form of an elongate scalene triangle, sidesstraight, corners rounded (fig. 16, j). Pronotum distinctly broaderthan long (0.25 by 0.2 mm.), sides rounded, basal angles obtuse, basearcuate, midbasal lobe short, rounded. Scutellum small, broadlytriangular. Elytra two and one-third times as long as broad (0.6 by0.26 mm.), sides parallel to within a fourth of apex, thence attenuateand broadly rounded to apex, apices individually rounded. Inter-mediate tibiae and tarsi equal in length; penultimate segment ofanterior and intermediate tarsi dilated, excavate beneath, bearingfifth segment on middle of its ventral surface. Posterior tibiae withtwo oblique ridges, basal one extending slightly more than half dis-tance across outer surface ; basitarsi with three (basal one rudimen-tary), second segment with one oblique ridge. Tarsal claws with buttwo teeth, distal one long, acute, terminal, other one broad, short,median, the two sides forming a right angle (fig. 18, e). Outer spur
TAXONOMIC STUDY OF MORDELLIDAE?RAY 297of posterior tibiae twice as long as inner one. Anal style almostthree times length of apical ventral segment (0.28 by 0.1 mm.), stout,dorsal surface rounded to apex, latter acute, bearing several short,black bristles ; ventral surface straight ; last ventral segment but twiceas long as penultimate one.Type locality.?Paraiso, Canal Zone, Panama.Type.?A unique male, U.S.N.M. No. 52919, collected on April 22,1911 (E. A. Schwarz).Reiivirks.?This species is named for Dr. A, Dampf, chief en-tomologist of the Mexican Department of Agriculture, who so largelycontributed to the possibility of this work by sending an assortmentof 802 specimens collected over a period of years.MORDELLISTENA AZTECA ChampionMordellistena azteca Champion, Biologia Centrali-Americaua, Coleoptera, vol. 4,pt. 2, p. 334, pi. 15, figs. 3, 3a, 1891.Seventeen specimens: San Rafael, Jicaltepec, Veracruz, Mexico(Townsend). MORDELLISTENA LUTEIFRONS ChampionMordellistena luteifrons Champion, Biologia Centrali-Americana, Coleoptera,vol. 4, pt. 2, p. 330, pi. 14, fig. 24a, 1891.Five specimens : All from District Federale, Mexico, July 11, 1924,on Arracia Tnultiflda (A. Dampf).MORDELLISTENA CANDELABRA, new speciesFiGXiRES 16, k; 18, gThis species is most closely allied to xanthopyga Champion (1891,p. 344) and may be separated from the latter member of the genusby the difference in color pattern, the peculiar shape of the distalsegment of the maxillary palpi, the longer length of the thirdantennal segment, and the presence of but a single spur at the apexof the posterior tibiae.Length : 1.82 mm. ; including anal style, 2.38 mm. Elongate, nar-row, sides subparallel; body flavocastaneous, eyes and ridges onposterior tibiae and tarsi fuscocastaneous, elytra and ventral ab-dominal segments (except distal segment) piceocastaneous. Sur-face covered with fine, recumbent, uniform pubescence, partaking ofground color, with contrasting areas on elytra, as follows: a broadhumeral patch extending obliquely into disk and reaching suture;a broad, transverse, postmedian band; and a broad sutural line con-necting the preceding two.Antennae 0.48 mm. long, reaching base of lateral pronotal margin
;
segments 1 and 2 large, equal ; 3 distinctly longer but narrower than
298 PROCEEDINGS OE THE NATIONAL MUSEUM voh.811 ; 5-10 each successively shorter and narrower than preceding ; 11 aslong as 1, spindle-shaped, broadest subbasaliy, attenuate to apex.Distal segment of maxillary palpi enlarged, elongate, with form of ascalene triangle, sides and apex but slightly rounded (fig, 16, k).Pronotum but slightly broader than long (0.56 by 0.52 mm.), apexand sides rounded, basal angles obtuse, base arcuate, midbasal lobeshort, rounded. Scutellum small, triangular. Elytra two and four-tenths times as long as broad (1.26 by 0.52 mm.), sides graduallynarrowing to within one-fourth of apex, where they become broadlyrounded, apices individually rounded. Penultimate segment of an-terior and intermediate tarsi filiform. Posterior tibiae with a shortsubapical ridge, a short median one, and a long oblique one extendingentirely across outer surface near basal articulations; basitarsi withtwo short, oblique ridges, each very oblique ; second segment with oneshort, oblique ridge. Tarsal claws with two distinct, broad teeth,distal one longest, and a small, blunt, median elevation on concavesurface (fig. 18, g). Outer spur of posterior tibiae more than halfiis long as basitarsus; inner spur ladling. Anal style tAvo and two-thirds times as long as apical ventral segment (0.55 by 0.2 mm.),narrow, evenly attenuate to apex.Type locality.?Panama City, Panama.ry;>f.?Male, U.S.N.M. No. 52939.Remarks.?Seven specimens: The type and a male paratype weretaken at Panama City, Panama, April 12 (A. H. Jennings) ; the allo-type and two female paratypes were taken at Paraiso, Canal Zone,Panama, on January 17, 1911, and April 18, 1911, respectively (E. A.Schwarz) ; a female paratype was taken at Pedro Miguel, Canal Zone.Panama, April 17, 1911 (E. A. Schwarz).I have been unable to find the shorter spur of the posterior tibiae inany of the seven specimens at hand ; in the absence of any damage tothe series this peculiarity is unaccountable. If this is the normal casefor this and several otlier species, it is the first occurrence of such anomissioi\ in the family.The males of candelahra are darker tlian the females, with ventralsurface of abdomen a uniform piceocaslaneous (except distal segment).MORDELLISTENA XANTHOPYGA ChampionMordellistena wanthopyga Champjon. Biologia Centrali-Americana, Coleoptera.vol. 4. pt. 2, p. .344, 462, pi. 15, figs. 18, 18a, 1891.One specimen : Paraiso, Canal Zone, Panama, April 26, 1911 (E. A.Schwarz). MORDELLISTENA TIARA RayMordellistena tiara Ray, Cau. Ent., vol. 68, p. 127, pi. 9, figs. 2, 5, 8, 1936.One specimen; Tampico, Mexico, December 14 (E. A. Schwarz).
TAXONOMIC STUDY OF MORDELLIDAE?RAY 299MORDELLISTENA SONORENSIS, new speciesFiGUBE 18, fThis species is most closely allied to funerea Champion (1891, p.343) and may be separated from the latter member of the genus bythe bicolored antennae, the much shorter, distal segment of the maxil-lary palpi, the longer antennae, the larger size, the oval form of thebody, and the emarginate anterior margin of the compound eye.Length: 2.4 mm.; including anal style, 3.2 mm. Form elongate,narrow^, sides rounded, subovate, elytra broadest at base, distinctlynarrowing and rounded caudad. Derm piceous black, five distal seg-ments of antennae and anterior and intermediate legs fuscous, basalsix segments of antennae and mouth parts flavocastaneous. Bodydensely covered with fine, recumbent, yellowish-white pubescence,except on castaneous parts, where it partakes of ground color.Antennae 1.5 mm. long, reaching to posterior coxae ; segments 3 and4 very short, equal ; 5-10 each as long as' 3 and 4 together and dis-tinctly broader; 11 distinctly longer than 10, sides and apex rounded,broadest medially. Distal segment of maxillary palpi enlarged, withform of a scalene triangle, lateral margin almost straight, corners, andmesal and apical margins distinctly rounded, broadest premedially.Eyes coarsely faceted, distinctly emarginate on anterior margins.Prothorax four-tenths broader than long (0.7 by 0.5 mm.), sides andapex rounded, basal angles obtuse, base arcuate, midbasal lobe short,but rounded. Scutellum short, subtriangular, sides and apex rounded,broader than long. Elytra tw^o and two-thirds times as long as broad(1.9 by 0.7 mm.), broadest at base, sides distinctly rounded to apex,curvature greatest on apical fourth, apices individually rounded.Intermediate tarsi distinctly longer than their tibiae ; penultimate seg-ment of anterior and intermediate tarsi simple, filiform. Posteriortibiae with two oblique ridges, basal one extending entirely acrossouter face, the other halfway; basitarsi with three, second segmentwith two oblique ridges, each extending halfway across outer surface.Tarsal claws with four teeth, distal one long, acute, two intermediateones short, their sides almost forming right angles, the proximal, sub-basal one rudimentary (fig. 18, /). Only a single spur visible at apexof posterior tibiae, one-half length of basitarsus. Anal style short,fairly stout, but one-half longer than apical ventral segment (0.75 by0.5 mm.), evenly attenuate to apex; apical ventral segment almosttwice length of penultimate one.Type locality.?Nainari, Sonora, Mexico.Type.?A unique male, U.S.N.M. No. 52940, collected on August 19.1927, on grass in^ricefield (A. Dampf).
300 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 87MORDELLISTENA DECORA, new speciesFiGUBE 18, hThis species is closely allied to flUcomis Champion (1891, p. 342),but may easily be separated from the latter member of the genus bythe greater length of the fourth antennal segment, the shorter, bicol-ored antennae, and the broader, much shorter distal segment of themaxillary palpi.Length : 3.7 mm. ; including anal style, 5 mm. Form narrow, elon-gate, sides subparallel, but slightly narrowing caudad until apicalfourth of elytra, broadest near base of pronotum. Head, pronotum,anterior and intermediate legs, four basal segments of antennae andspurs of posterior tibiae, castaneous ; elytra, eyes, and ventral surfacepiceous, seven distal segments of antennae and posterior legs piceo-castaneous. Body densely covered with fine, recumbent pubescence,partaking of ground color on castaneous parts, yellowish white else-where, short and indistinct on eyes.Antennae 1.2 mm. long, almost^, reaching base of lateral pronotalmargin ; segment 4 slightly longer and broader than 3 ; 5-10 each one-half longer and distinctly thicker than 4, not serrate, however, broad-est subapically; 11 large, one-half longer than 10, sides and apexrounded, broadest postmedially. Distal segment of maxillary palpienlarged, form of a broad, scalene triangle, lateral margin straight,corners and other two sides rounded. Pronotum but slightly broaderthan long (1.1 by 1 mm.), sides rounded, basal angles obtuse, basearcuate, midbasal lobe short, subtruncate. Scutellum small, broadlytriangular, sides straight, apical angle rounded. Elytra almost twoand one-half times as long as broad (2.7 by 1.1 mm.), sides gentlynarrowing on basal three-fourths, thence broadly rounded to apex,apices individually rounded. Intermediate tarsi distinctly longer thantheir tibiae; penultimate segment of anterior and intermediate tarsishort, filiform. Posterior tibiae with two oblique ridges, basal oneextending entirely across outer surface, the other halfway; basitarsiwith three, second segment with two oblique ridges, each extendinghalfway across outer surface. Tarsal claws with four teeth, threeacute and long, fourth short, blunt, premedian (fig. 18, h). Outerspur of posterior tibiae two and one-half times as long as inner one.Anal style two and six-tenth times length of apical ventral segment(1.3 by 0.5 mm.), narrow, attenuate to apex; last ventral segmentslightly more than twice length of penultimate one.Type JocaUfy.?Centinela. Colima, Mexico.Type.~K unique male, U.S.N.M. No. 52941, collected on Jauuarv28, 1930, on cotton (A. Dampf).
TAXONOMIC STUDY OF MORDELLIDAE?RAY 301MORDELLISTENA FLAVICORNIS ChampionMordellistena flavicornis Champion, Biologia Centrali-Americana, Coleoptera,vol. 4, pt. 2, p. 342, pi. 15, fig. 15, 1891.One specimen: Juan Mina Plantation, Canal Zone, Panama, July13, 1928, swej)t from grass and cowpeas (H. J. Dietz and J. Zetek).MORDELLISTENA MUTABILIS ChampionMordellistena mutabilis Champion, Biologia Centrali-Americana, Coleoptera, vol.4, pt. 2, p. 339, pi. 15, figs. 9-11, 1891.Eight specimens: Six from Cordoba, Veracruz, Mexico, (3) April30, (1) May 5, and (2) May 17, 1908 (A. Fenyes) ; two from Cacao,Trece Aguas, Alta Vera Paz, Guatemala, March 30 and April 29(Schwarz and Barber).MORDELLISTENA ORNATIPENNIS, new speciesFigure 18, iThis species is most closely allied to hasimacula Champion ( 1891, p.341) and may be separated from the latter member of the genus bythe shorter, broader, terminal segment of the maxillary palpi and thetiDO long ridges of the posterior tibiae.Length: (male) 2.25 mm., (female) 2.75 mm.; including anal style,(male) 3.25, (female) 3.75 mm. Elongate, narrow, sides subparallel;head fuscocastaneous ; pronotum fuscocastaneous with a large,rounded, castaneous spot on each side at middle ; elytra fuscopiceous,with an oblique, narrow^, castaneous, humeral spot extending obliquelytoward suture and separated from latter by less than its own width;front and middle legs and anal style castaneous, ventral surface andposterior legs fuscopiceous. Surface densely covered with fine,recumbent, flavous pubescence.Antennae 0.7 mm. long, reaching posterior coxae ; segments 3 and 4equal; 5 to 10 each almost as long as 3 and 4 together; 11 rounded,one-third longer than 10. Distal segment of maxillary palpi en-larged, subtriangular, outer margin straight, corners rounded, apicaland inner margins curved. Pronotum as long as broad, apex andsides rounded, basal angles obtuse, base arcuate, midbasal lobe short,rounded. Scutellum small, triangular, apical angle rounded. Elytrathree times as long as broad (1.75 by 0.58 mm.), sides parallel towithin one-fourth of apex, apices individually rounded. Penultimatesegment of anterior and intermediate tarsi dilated. Posterior tibiaewith a short subapical ridge and two long, oblique ridges, each ex-tending entirely across outer surface; basitarsi with three obliqueridges, each extending two-thirds across outer surface; second seg-ment with two ridges, each almost completely spanning articulation.Tarsal claws with four rather thick, blunt teeth, distal one longest,
302 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 87proximal, median one broad, the sides forming a right angle (fig.18, ^). Anal style more than three times as long as apical ventralsegment, narrow, attenuate to apex.Type locality.?Paraiso, Canal Zone, Panama.Tyj)e.?M.2\^ : U.S.N.M. No. 52916.Remarks.?The type was taken at Paraiso, Canal Zone, Panama,January 17, 1911 (E. A. Schwarz), the allotype at Pedro Miguel,Canal Zone, Panama, April 17, 1911 (A. H. Jennings) ; and a maleparatype at Palo Seco, Trinidad, October 20, 1918 (H. Morrison).The two long ridges of the posterior tibiae in this species is thefirst record of such an occurrence in any member of the family andthis character alone will separate ornatiperwiis from any species re-corded in the "Biologia." The castaneous humeral spot is variablein size and in the paratype extends the length of the elytra, leavingonly a narrow, piceous, sutural and marginal line.MORDELLISTENA GRACILIFORMIS, new speciesThis species is mostly closely alKed to ^-fa^sciata Champion (1891,p. 337) , a Mexican form, but may easily be separated from the lattermember of the genus by the completely black dermal color, thesmaller, more irregular pubescent patches on the pronotum and ely-tra, and by the shorter distal segment of the maxillary palpi.Length: 2.75 mm.; including anal style, 3.65 mm. Elongate, sub-parallel ; dermal color entirely black. Surface densely covered withfine, yellowish-white pubescence, dense on pronotum and elytra, whereit forms a number of irregular, broken, transverse fasciae.Antennae 1 mm. long, reaching basal abdominal segment ; segment4 equal in length to 3, but slightly broader; 5-10 each one-halflonger than 4 and considerably broadened; 11 rounded, but littlelonger than 10. Distal segment of m.axillary palpi enlarged, broadlytriangular, rounded on mesal edge and apex. Prothorax slightlybroader than long (0.8 by 0.75 mm.), sides rounded, base arcuate,midbasal lobe short, subtruncate. Scutellum small, triangular, apicalangle rounded. Elytra twice as long as broad (2 by 1 mm.), sidesgradually narrowing from broadest posthumeral position to apex,apices individually rounded. Tarsal claws with three teeth, all short,blunt, proximal one situated medially. Posterior tibiae with twooblique ridges extending halfway across outer surface ; basitarsi withthree, second segment witli two oblique ridges, all extending halfwayacross outer surface. Anal style three times as long as apical ventralsegment (0.9 by 0.3 mm.), narrow, attenuate to apex, sides straight;last ventral segment but twice length of penultimate one.Tyj)e locality.?Coliseo, Cuba.Ti/pe.?Male, U.S.N.M. No. 52917.
TAXONOMIC STUDY OF MORDELLIDAE RAY 303Remarks.?The type and a male paratype were collected at Coliseo,Province of Matanzas, Cuba, June 15, 1934 (S. T. Danfortli). Theallotype and a female paratype were collected at Central Jaronu,Cuba, June 2, 1930, on weeds and grasses (L. C. Scaramuzza) andCentral Jagueyal, Cuba, June 30, 1927, on grasses (L. C. Scara-muzza). MORDELLISTENA LATIPALPALIS, new speciesFigure 18, iThis species is rather closely allied to aequinoctialis Champion(1891, p. 311) but may be separated from the latter member of thegenus without difficulty by means of the much smaller size, the dif-ferent character of the antennal segments, the broader terminal seg-ment of the maxillary palpi of the male, and the general differencein color.Length : 2.95 mm. ; including anal style, 4.15 mm. Form elongate,narrow, sides subparallel, attenuate, and gradually rounded caudadfrom middle of elytra. Dermal color dark fuscocastaneous, legs,except posterior femora, maxillary palpi, antennae, two basal seg-ments lighter, apex of front, and clypeus, flavocastaneous. Surfacerather densely covered with fine, recumbent, yellowish-white pubes-cence, densest on elytra, abdomen, and anal style ; eyes sparsely cov-ered with short, whitish hairs.Antennae 1 mm. long, slender, reaching intermediate coxae; seg-ments 1 and 2 subequal; 3 and 4 equal in length, narrow; 5-10 eachone-third longer and broader than 4, lateral margins straight, mesalmargins curved, broadest at apex; 11 one-half longer than 10, sidesand apex rounded, broadest medially. Distal segment of maxillarypalpi enlarged, somewhat pear-shaped, broad, mesal margin straight,lateral margin and apex broadly rounded, greatest width on apicalthird. Pronotum distinctly broader than long (0.95 by 0.85 mm.),apex and sides rounded, basal angles subacute, forming a rightangle, base arcuate, midbasal lobe moderately produced, subtruncate.Scutellum broadly subtriangular, broader than long, apex abruptlytransversely truncate, angles obtuse. Elytra distinctly more thantwice as long as broad (2.1 by 0.95 mm.), sides subparallel on basalhalf, narrowly attenuate caudad, broadly rounded on caudal fourth,apices individually rounded. Hind tibiae with two oblique ridgesextending halfway across outer surface ; basitarsi with three, secondsegment with two oblique ridges, all extending halfway across outersurface. Tarsal claws with four teeth, the distal three long, acute,postmedian, proximal one short, blunt, and median in position (fig.18, j). Anal style elongate, narrow, attenuate to apex, three timeslength of apical ventral segment (1.2 by 0.4 mm.) ; last ventral seg-ment three times as long as penultimate one.
304 PROCEEDINGS OF THE NATIONAL MUSEUM vol.87Type locality.?Plancha Piedra, between British Honduras andGuatemala.ry;^^.?Male, U.S.N.M. No. 52942.Remarks.?The single male at hand was collected at PlanchaPiedra, on the border between British Honduras and Guatemahi, inthe Peten Department, October 27, 1925, swept from weeds (A.Dampf). MORDELLISTENA LATICORNIS, new speciesFigure 16, U; 18, fcThis species is most closely allied to hexastigma Champion (1891,p. 321) and may be separated from the latter member of the genusby the unicolored antennae, the different nature and position of thespots on the elytra, the much greater length and breadth of thefourth antennal segment, the different shape of the distal segmentof the maxillary palpi, the small number of combiike ridges on theposterior tibiae and tarsi, and the smaller size.Length: 2.5 mm.; including anal style, 3.45 mm. Form moder-ately robust, subcuneate, elytra broadest at base, thence narrowingto apex, body broadest near base of pronotum. Head, pronotum,antennae, mouth parts, spurs of posterior tibiae, and legs, exceptposterior femora, fiavocastaneous; ventral surface, except last ventralsegment and base of anal style, and elytra, piceous black, the elytraw'ith a castaneous, semilunate, humeral fascia, curving posteriorlyand mesad, not reaching suture and extending one-fourth into disk;posterior femora, last ventral segment, and base of anal style fusco-castaneous. Body densely covered with fine, recumbent pubescence,partaking of ground color on castaneous areas, whitish yellow onventral surface, black on elytra, except for following whitish areas:a semilunate fascia overlying castaneous dermal spot mentionedabove, a broad, transverse, postmedian fascia, interrupted at sutureand not quite reaching lateral margins, and a short area coveringthe extreme apex.Antennae 1.1 nun. long, reaching basal abdominal segment; segment4 one-half longer and much broader than 3; 5-10 each one third longerthan 4, broad, flattened, but not serrate, broadest subapically; 11 dis-tinctly longer than 10, sides and apex rounded, broadest medially.Distal segment of maxillary palpi enlarged, form of an elongatescalene triangle, outer margin straight, mesal and apical edges andcorners rounded, broadest at apical third (fig. 16, A). Pronotum dis-tinctly broader than long (0.85 by 0.7 mm.), sides and apex rounded,basal angles obtuse, base arcuate, midbasal lobe short, but rounded.Scutellum moderately large, triangular, sides straight, apical anglerounded. Elytra slightly more than twice as long as broad (1.8 by
TAXONOMIC STUDY OF MORDELLIDAE?RAY 3050.85 mm.'), sides broadest at base, evenly attenuate to apex, apicesindividually rounded. Intermediate tibiae and tarsi approximatelyequal in length; penultimate segment of anterior and intermediatetarsi dilated, emarginate at apex, bearing distal segment on its ventralsurface. Posterior tibiae with two and a rudimentary third obliqueridge, each extending approximately halfway across outer surface;basitarsi with three, second segment with two oblique ridges, eachextending halfwaj' across outer surface. Tarsal claws with fourteeth, two distal ones long, acute, two proximal ones short, blunt(fig. 18, k). Outer spur of posterior tibiae almost twice length ofinner one. Anal style slightly more than tw^ice length of apical seg-ment (0.95 by 0.45 mm.), rather stout at base, evenly attenuate toapex; last ventral segment almost three times length of penultimateone.Type locality.?Tegucigalpa, Honduras.Type.?K unique male, U.S.N.M. No. 52912, taken on March 31,1917, by F. J. Dyer. MORDELLISTENA SEPIA, new speciesFiQUKE 19, eThis species is most closely allied to tahascana Champion (1891, p.330) and may be separated from the latter member of the genus by adistinct difference in color and pubescent maculation, the shorter formand the isosceles condition of the distal segment of the maxillarypalpi, the greater length of the fourth antennal segment, and the oddlyshaped scutellum.Length: 1.5 mm.; including anal style, 2 mm. Form moderatelyelongate, robust, sides attenuate beyond basal third of elytra, broadestnear base of pronotum. Derm warm sepiaceous in color, meso- andmetasternum and anal style piceocastaneous. Body densely coveredwith fine, recumbent, flavobrunneous pubescence, partaking of groundcolor in the following areas: a somewhat rounded spot on middle ofdisk of pronotum, a subbasal, subsutural spot extending to middle ofelytra, produced posteriorly and recurved anterolaterad, but not reach-ing lateral margins; and a broad, transverse, arcuate band reachingneither suture nor lateral margins and produced anteriorly, almosttouching previously mentioned subsutural line.Antennae 0.7 mm. long, reaching base of abdomen, filiform, slender,segment 4 twice as long and much broader than 3 ; 5-10 but slightlylonger than 4 and subequal in length and width; 11 slightly longerthan 10, broadest at apical third, thence rounded to apex. Distalsegment of maxillary palpi enlarged, with form of an elongatescalene triangle, apical margin and angles rounded, mesal and lateralmargins straight. Pronotum slightly broader than long (0.47 by
306 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 870.37 mm.), sides rounded, base arcuate, midbasal lobe short, rounded.Scutellum subangulate, base straight, sides convex, very obtuselyangulate, apical angle acute. Elytra twice as long as broad (1.02 by0.53 mm.), sides attenuate and narrowing from basal third to apex,apices broadly, individually rounded. Intermediate tibiae and tarsisubequal in length; antepenultimate segment of anterior and inter-mediate tarsi short, slightly broadened, penultimate segment dilated,excavate beneath. Posterior tibiae with three subequal, obliqueridges extending halfway across outer surface; basitarsi with three,second segment with two oblique ridges, all extending halfway acrossouter surface. Tarsal claws with three distinct, acute teeth. Outerspur of posterior tibiae twice as long as inner one. Anal style threetimes as long as apical ventral segment (0.5 by 0.16 mm.), moderatelyrobust, attenuate to apex.Type locality.?La Ceiba, Honduras.Type.?A unique male, U.S.N.M. No. 52920, collected on August23, 1916, by F. J. Dyer.MORDELLISTENA BOSENI, new speciesFiGUBEs 16, i; 18, IThis species bears close resemblance to both in-fima LeConte (1863a,p. 49) and tiai^a Ray (1936a, p. 127), but may easily be separatedfrom either of these members of the genus by the black, enclosed areaof pubescence on the elytra, the bicolored antennae and legs, thelarger size, the larger number of comblike ridges on the posteriortibiae and tarsi, the larger size, and the greater length of the fourthantennal segment.Length: 3.3 mm.; including anal style, 4.55 mm. Form narrow,elongate, sides subparallel, narrowing from apical fourth of elytra,broadest near base of pronotum. Derm piceous black, apical mar-gins of abdominal segments a shining, castaneous color, basal foursegments of antennae, mouth parts, anterior legs, and spurs of pos-terior tibiae, flavocastaneous. Body densely covered with fine, re-cumbent pubescence, a distinct yellowish-greenish white, condensedon elytra to form the following fasciae: a broad humeral area curv-ing posteromesad and reaching suture; a narrow line along base,embracing scutellum and continuing along entire length of suture;an irregular, transverse, postmedian band broadening laterally andextending along lateral margins to reach both humeral area and anapical fascia ; apical sixth of elytra, longest basally near suture, pubes-cence between these areas consisting chiefly of dark, fuscopiceoushairs.Antennae 1.2 mm. long, reaching posterior coxae; segment 4 one-half longer than 3; 5-10 each one-third longer and considerably
TAXOirOMIC STUDY OF MORDELLIDAE RAY 307broader than 4, broadest subapically; 11 one-half longer than 10mesal edge, apex and corners rounded, broadest postmedially. Distalsegment of maxillary palpi enlarged, with form of a scalene triangle,broad, lateral margin straight, other edge and corners rounded (fig.16, i). Pronotum four-tenths broader than long (1.1 by 0.8 mm.),sides and apex rounded, basal angles obtuse, base arcuate, midbasallobe short, distinctly truncate. Scutellum moderately large, trian-gular, sides straight, apical angle rounded. Elyira two and three-tenths times as long as broad (2.5 by 1.1 mm.), sides subparallel onbasal three-fourths, thence broadly rounded to apex, apices individu-ally rounded. Intermediate tarsi longer than their tibiae; penulti-mate segment of anterior and intermediate tarsi dilated, emarginateat apex, bearing the distal segment on its ventral surface. Posteriortibiae with three short, oblique ridges, each extending less than half-way across outer surface; basitarsi with four, second segment withtwo oblique ridges, each extending less than halfway across outersurface. Tarsal claws with four teeth, all long, acute (fig. 18, I).Outer spur of posterior tibiae twice length of inner one. Anal styletwo and one-half times length of apical ventral segment (1.25 by 0.5mm.), slender, attenuate to apex; last ventral segment three timeslength of penultimate one.Type locality.?San Cristobal, Chiapas, Mexico.Type.?Mule, U.S.N.M. No. 52943.BeTnarks.?The type was taken at San Cristobal, Chiapas, Mexico,July T, 1926, by sweeping, at 2,200 meters elevation (A, Dampf )
;
a male paratype was taken at Cerro Ecatepec, Chiapas, Mexico, June24, 1926, on shrubs, at 2,400 meters elevation (A. Dampf).MORDELLISTENA FLAVOCOLLARIS, new speciesFiGtJBE 18, mThis species is most closely allied to sanguinicollis Champion (1891,p. 320) and may be separated from the latter member of the genus bythe difference in the pattern of the elytral pubescence, the equalityin length between antennal segments 3 and 4, the broader distal seg-ments of the same, the difference in the color of the ventral surface,legs, and antennae, and the smaller number of comblike ridges onthe posterior tibiae.Length : 3.6 mm. ; including anal style, 4.7 mm. Form moderatelyrobust, sides attenuate, narrowing slightly but visibly from base ofelytra posteriorly, broadly rounded from apical fourth of elytra toapex, apices individually rounded, broadest near base of pronotum.Head, except clypeus, and elytra, piceous black ; pronotum castaneous,with an indefinite dorsal cloud on posterior half of disk; antennae,mouth parts, anterior legs, caudal margins of metasternum and
308 PROCEEDINGS OF THE NATIONAL MUSEUM vol.87abdominal segments, last one entirely, flavocastaneoiis ; intermediateand posterior legs piceocastaneoiis. Body covered with fine, re-cumbent pubescence, whitish on all parts of body except pronotum,where it is distinctly yellowish and on apical half of anal style,where it is piceous ; condensed on elytra to form the following whitishspots : an indefinite area along base and embracing humeri, with twoblack areas enclosed, a sutural line continuing for two-thirds length,a postmedian band extending from suture almost to lateral marginand broadened anteriorly and posteriorly for a short distance atlatter point, and a small, irregular, apical area.Antennae 1.5 mm. long, filiform, distal segments fairly stout ; seg-ments 3 and 4 equal in both length and width; 5-10 each one-halflonger and much broader than 4, distinctly longer than broad, broad-est at distal end; 11 one-half longer than 10, rounded on sides andapex, broadest medially. Distal segment of maxillary palpi en-larged, with form of a broad isosceles triangle, thick, lateral andmesal surfaces noticeably convex, angles rounded. Pronotum dis-tinctly broader than long (1.3 by 0.9 mm.), sides and apex rounded,basal angles obtuse, midbasal lobe short, rounded. Scutellum small,subtriangular, corners and apex broadly rounded, so as to indicate asubtruncate appearance. Elytra slightly more than twice as long asbroad (2.7 by 1.3 mm.), sides attenuate, broadest at base, gentlynarrowing from latter point to three-fourths length, where it becomesbroadly rounded to apex, apices individually rounded. Interme-diate tibiae and tarsi equal in length ; penultimate segment of anteriorand intermediate tarsi dilated, excavate beneath, bearing fifth segmenton middle of its ventral surface, posterior tibiae with three obliqueridges, all extending less than half distance across outer surface;basitarsi with four, second segment with two oblique ridges, eachextending less than halfway across outer surface. Tarsal claws withfour teeth, distal one blunt, others acute (fig. 18, m). Outer spurof posterior tibiae stout, but twice as long as inner one. Anal stylebut twice length of apical ventral segment (1.1 by 0.55 mm.), stout,suddenly constricted at middle, thence attenuate to apex, latter nar-rowly truncate ; last ventral segment but twice as long as penultimateone.Ty^e locality.?Jalapa, Mexico.Type.?A unique male, U.S.N.M. No. 52918, collected by J. T.Mason. MORDELLISTENA TEXANA SmithMordellistena texana Smith, Trans. Amer. Ent. Soc, vol. 10, pp. 89, 98, pi. 3,fig. 28, 1882.One specimen: Colima, Colima, Mexico (L. Conradt).
TAXONOMIC STUDY OF MORDELLIDAE?RAY 309MORDELLISTENA CHIAPENSIS, new speciesFiGUBE 18, nThis species is most closely allied to gracilicornis Champion (1891,p. 318) and may be separated from the latter member of the genusby the distinct difference in color pattern of the dorsal surface, theshorter, broader antennae, the broader, more rounded distal segmentof the maxillary palpi, and the longer anal style.Length: (male) 3.9 and (female) 4.9 mm.; including anal style,(male) 4.9 and (female) 6 mm. Form elongate, narrow, sides sub-parallel, distinctly narrowing at apical third of elytra, broadest atmiddle of pronotum. Body brunneous, legs a little lighter, eyes, tipsof mandibles, and ridges of posterior tibiae and tarsi black. Surfacedensely covered with fine, recumbent pubescence, distinctly yellowishon entire body, except on posterior tibiae and tarsi and anal style,where it partakes of ground color.Antennae (male) 1.4 o.nd (female) 1.7 mm. long, reaching base ofabdomen ; segment 4 distinctly longer but very little broader than 3
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5-10 somewhat flattened and a little dilated at apex, each as long asand noticeably broader than 4; 11 one-third longer than 10, sides andapex rounded, broadest at distal third. Distal segment of maxillarypalpi enlarged, strongly robust, with the form of an isosceles triangle,mesal and apical sides equal in length, ail sides and corners distinctlyrounded, broadest medially. Eyes large, coarsely granulated, an-terior margin emarginate (male) or truncate (female). Pronotumbut slightly broader than long (1.2 by 1 mm. in male, 1.5 by 1.3 mm.in female), sides subparallel to within a third of apex, thence attenu-ate and broadly rounded to apex, apices individually rounded. In-termediate tibiae and tarsi equal in length; penultimate segment ofanterior and intermediate tarsi dilated, emarginate at apex, receivingdistal segment on its ventral surface. Posterior tibiae with foursubequal, oblique ridges extending halfway across outer surface ; basi-tarsi with four, second segment with two oblique ridges, all extendinghalfway across outer surface. Tarsal claws with five distinct teeth,distal four long, acute, proximal one premedian, short, blunt, obtuse(fig. 18, n). Outer spur of posterior tibiae less than twice as long asinner one. Anal style distinctly more than twice as long as apicalventral segment (1 by 0.4 mm. in male, broken in female), narrow,attenuate to apex; last ventral segment twice length of penultimateone.Type locality.?Finca Belen, Chiapas, Mexico.Type.?Male, U.S.N.M. No. 52944.Remarks.?Two specimens, type and allotype, were taken at FincaBelen, Chiapas, Mexico, April 22, 1932, at light, 800 meters elevation(A. Dampf).
310 PROCEEDINGS OF THE NATIONAL MUSEUM vol.87As noted in the description above, there exists a distinct sexual dif-ference in the condition of the anterior margin of the compound eye.The antennae, maxillary palpi, and the ridges of the posterior tibiaeand tarsi do not indicate a further possibility of sexual separation.MORDELLISTENA HEXASTIGMA ChampionMordelUMena hexastigma Champion, Biologia Centrali-Americana, Coleoptera,vol. 4, pt. 2, p. 321, pi. 14, figs. 12, 12a, 1891.One specimen: Porto Bello, Panama, February 18, 1911 (E. A.Schwarz). MORDELLISTENA RUFONOTATA ChampionMordellistcva riifonotata Champion, Biologia Centrali-Americana, Coleoptera,vol. 4, pt. 2, p. 318, pi. 14, fig. 7, 1891.Two specimens : Both from San Cristobal las Casas, Chiapas, Mex-ico, June 4, 1926 (A. Dampf).MORDELLISTENA OALLENS ChampionMordellii-tena callens Champion, Biologia Centrali-Americana, Coleoptera, vol. 4,pt. 2, p. 315, 1891.Three specimens : One from Finca Vergel, Chiapas, Mexico, May 19,1935, on wing, TOO meters elevation (A. Dampf) ; one from La Ceiba,Honduras, September 23, 1916 (F. J, Dyer) ; one from Paraiso, CanalZone, Panama, April 29, 1911 (E. A. Schwarz).MORDELLISTENA VAFER ChampionMordellistena vafcr Champion, Biologia Centrali-Americana, Coleoptera, vol. 4,pt. 2, p. 314, 1891.One specimen: Panama City, April 13, 1911 (A. H. Jennings).MORDELLISTENA CASTANEICOLOR ChampionMordellisiena castaveirolor Champion, Biologia Centrali-Americana, Coleoptera,vol. 4, pt. 2, p. 313, pi. 14, figs. 1, la, 1891.Five specimens : Three from Port Bello, Panama, February 15, 20^and March 1, 1911 (E. A. Schwarz) ; two from Finca Vergel, HuixtlaValley, Chiapas, Mexico, May 19, 1935, at light, 1,200 meters eleva-tion (A. Dampf). Genus DICLIDIA LeConteDiclidia LeConte, Proe. Acad. Nat. Sci. Philadelphia, vol. 14, p. 43, 1862.DICLIDIA UNDATA ChampionDiclidia tnidata Champion, Biologia Centrali-Americana, Coleoptera, vol. 4, pt. 2,p. 250. pi. 11, figs. 1, la, 1891.Two specimens : Both from Zacatepec, Pueblo, Mexico, eTune 28, 1932,at light, 2,000 meters elevation (A. Dampf). <
TAXONOMIC STUDY OF MORDELLIDAE RAY 311DICLIDIA MEXICANA, new speciesThis species may easily be separate from the only other CentralAmerican member of the genus, undata Champion (1891, p. 250), bythe difference in color, the broader distal segments of the antennae,the broader, more depressed form, and the shorter antennae. Frominyoensis Liljeblad (1921, p. 181) it may be distinguished by the dif-ference in color, the much larger size, the peculiar character of theantennae, and the greater compression and elevation of the meso-sternum.Length: 3.5 mm. Form oblong-ovate, rounded anteriorly and po-teriorly, broadest near middle of elytra. Head fuscocastaneous, pro-notum castaneous, elytra piceous; antennae flavous at base, succes-sively darker distally, last five segments piceous ; ventral surface pice-ous, anterior legs castaneous, intermediate and posterior pairfuscocastaneous. Body densely covered with fine, whitish-yellowpubescence ; surface of pronotum and elytra covered with fine, whitish-yellow f)ubescence ; surface of pronotum and elytra covered with fine,transverse strigae, generally obscured by the pubescence.Antennae 1.1 mm. long, reaching base of lateral pronotal margin;segments 1 and 2 equal; 4 more than one-half longer than 3; 5 equalto 4 ; 6 equal to 2 ; 7-10 short, dilated, forming a loose, elongate club,as broad as long; 11 almost twice as long as 10 and equally broad,broadest medially, sides almost straight, apex rounded. Maxillarypalpi missing. Pronotum one-half broader than long (1.2 by 0.8mm.), apex and sides broadly rounded in a continuous semicircle,broadest at base, basal angles acute, base but slightly arcuate, no mid-basal lobe being distinguishable. Scutellum large, triangular. Elytraslightly more than twice as long as broad at base (2.5 by 1.2 mm.),sides visibly curved from base to apex, apices individually rounded.Anterior and intermediate tibiae slender, as long as their tarsi, secondand third segments of latter part short, penultimate segment dilatedwith emarginate apex, bearing distal segment on its ventral surface.Tarsal claws with but the single distal prolongation.Type locality.?Santa Anita, Jalisco, Mexico.Type.?Female, U.S.N.M., No. 52931, a unique specimen taken Au-gust 7, 1927, on weeds, 2,000 meters elevation (A. Dampf).Genus NAUCLES ChampionNancies Champion, Biologia Centrali-Americana, Coleoptera, vol. 4, pt. 2, p. 257,1891.
312 PROCEEDINGS OF THE NATIONAL MUSEUM VOU 87NAUCLES BASALIS ChampionNancies basalis Champion, Biologia Centrali-Americana, Coleoptera, vol. 4, pt. 2,p. 258, 1891.Six specimens : Four from Tampico, Mexico, February 16 (3) andDecember (E. A. Schwarz) ; two from Progreso, Yucatan, Mexico,June 7 (F. Knab)
.
NAUCLES FASCIATA, new speciesSuperficially this species resembles Anfhohatula quadriguttataChampion (1891, p. 255), but the much smaller size, the entirely dif-ferent character of the maxillary palpi and antennae, and the fivevisible ventral abdominal segments adequately distinguish fasciatafrom quadriguttata. This species is unquestionably a species of thegenus Naucles and may easily be separated from the four previouslyknown members of the genus by the difference in the length of thevarious antennal segments, the maculation of the elytra, and thelarger size.Length : 1.7 mm. Form oblong-bvate, rounded anteriorly and pos-teriorly, broadest near middle of elytra. Derm flavous, eyes black,nine distal segments of antennae, mesosternum and metasternum andventral abdominal segments fuscocastaneous ; elytra with base, suture,except apical sixth, lateral margin, except broadly rounded apicalportion, and a broad, postmedian, transverse band, fuscous. Bodydensely covered with fine, recumbent, flavous pubescence.Antennae 0.5 mm. long, reaching intermediate coxae; segment 2twice as long as 1; 3 slightly longer than 4; 4-6 equal 7-10 broad-ened, forming a loose, elongate club, each segment distinctly broaderthan long; 11 as long as 10, but narrower, broadest subbasally, sidesand apex rounded. Distal segment of maxillary palpi enlarged,elongate, with form of a scalene triangle, apical margin veryobliquely truncate, corners and sides rounded; terminal segment oflabial palpi slender, apical half prolonged, acuminate. Pronotumdistinctly broader than long (0.65 by 0.45 mm.), apex and sidesbroadly rounded in a continuous semicircle, broadest at base, basalangles acute, base arcuate, midbasal lobe short, but rounded; entiresurface covered with fine, transverse strigae. Scutellum moderatelylarge, semicircular. Elytra almost two-thirds longer than broad(1.25 by 0.8 mm.), lateral margin distinctly curved, broadest sub-medially, apical fourth broadly rounded, apices individually curved
;
basal fourth of surface faintly covered with transverse strigae. Pe-nultimate and antepenultimate segments of anterior and intermediatelegs much shortened, as broad as long. Tarsal claws with but thesingle distal prolongation.Type locality.?Ponce, Puerto Rico.ry;?e.?Female, U.S.N.M. No. 52921.
TAXONOMIC STUDY OF MORDELLIDAE?RAY 313
'Remarks.?The type and seven female paratypes were collected atPonce, Puerto Rico, August 7, 1933, on flowers of Randia mitis(R. G. Oakley). Another female paratype, from the same locality,was taken on August 8, 1935 (R. G. Oakley). Since six of the speci-mens have protruding genitalia and all are alike in other respects,I am obliged to consider them as apparently all females.The markings on the elytra vary considerably in both width offasciae and depth of color. In one specimen they are visible onlywhen turned at an angle to the disk.Genus ANTHOBATULA StrandAnthobatula Strand, Acta Univ. Latviensis, vol. 20, p. 23, 1929.Anthohates LeConte, nee Gistel, in Agassiz and Cabot, Lake Superior, p. 231,1850.Pentaria Mulsant, Ann. Soc. Linn. Lyon, ser. 2, vol. 3, p. 391, 439, 1856.ANTHOBATULA MULTIPILIS, new speciesIn Champion's key to the Central American species of Pemtaria(1891, p. 252) this species runs to fiavipes Champion (1891, p. 253).It may be separated from that member of the genus, however, bythe unicolorous antennae, which do not have the distal segmentstransverse, in the color of the ventral surface and legs, and in thebroader form of the body. A. multipilis is not closely allied to anyof the North American members of the genus.Length : 2.1 mm. Form oblong-ovate, rounded anteriorly and pos-teriorly, broadest near middle of elytra. Derm piceous, dull, apexof front, clypeus, metasternum, abdomen, legs, and antennae casta-neous, maxillary palpi and genitalia flavous. Body densely coveredwith fine, short, recumbent pubescence, silvery on dorsal surface, par-taking of ground color ventrally.Antennae 0.7 mm. long, reaching intermediate coxae; segments3-6 equal; 7-10 each twice as long as 5 but slightly broader; 11slightly longer and narrower than 10, sides straight, apex rounded.Distal segment of maxillary palpi enlarged, with form of an elon-gate, scalene triangle, basal and apical angles acute, mesal angleobtuse, all rounded, sides straight, except apical fifth of lateralmargin, which is distinctly rounded. Pronotum one-half broaderthan long (0.9 by 0.6 mm.), sides and apex broadly rounded in asingle continuous curve, basal angles obtuse, base arcuate, midbasallobe very short, but rounded. Scutellum subtriangular, sides andapical angle rounded. Elytra but one-half longer than broad (1.5by 1 mm.), sides visibly rounded, broadest postmedially, apicesindividually rounded; surface completely covered with numerous,fine, transverse strigae. Intermediate and posterior tibiae approxi-
314 PROCEEDIlSrGS OF THE NATIONAL MUSEUM vol. 87mately equal in length to their tarsi, third and fourth segments oflatter structure short, fourth segment dilated, receiving distal seg-ment in a cavity near basal margin. Tarsal claws with but a singletooth, the distal prolongation.Type locality.?Cayamas, Cuba.Type.?M^\Q., U.S.N.M. No. 52932.Reriiarlcs.?The type was taken on January 17, the allotype onJanuary 16, seven paratypes (2 males, 5 females) were taken Jan-uary 22, February 3, January 20, February 10 (2), February 24,and June 10; all were taken at Cayamas, Cuba (E. A. Schwarz).There is some variation in the color of the ventral surface and inthe extent of the castaneous area on the front and clypeus. It isentirely possible that some specimens in a larger series would beentirely piceous. LITERATURE CITEDBroun, T.18S6. Manual of New Zealand Coleoptera, pt. 4.Champion, G. C. *1891. Mordellidae. In Biologia Centrali-Americana, Coleoptera, vol. 4, pt.2, pp. 250-350, 5 pis.Helmuth, 0. A.1S65. New species of Mordellidae collected In Illinois. Proc. Acad. Nat.Sci. Philadelphia, vol. 17, p. 96.LeContB, John L.1862. Synopsis of the Mordellidae of the United States. Proc. Acad. NatSci. Philadelphia, vol. 14, pp. 43-51.LiT.JEBLAD, E.1921. Notes on the Mordellidae of the United States, with descriptions ofnew species. Can. Ent., vol. 53, pp. 181-186.Linnaeus, Carolus.1758. Systema naturae, ed. 10.Pic, M.1924. Nouveautes diverses. Mel. Exot.-Ent., fasc. 42, 32 pp.1930. Mordella oglohlini. Rev. Soc. Ent. Argentina, vol. 3, p. 30.1936. Contribution h I'etude du genre Heteromfere. Mordella L. Rev. Soc.Ent. Argentina, vol. 9, pp. 111-115.Ray, Eugene.1930. A study of South American Mordellidae. Coleopt. Contrib., vol. 1,pt. 3, pp. 161-171, 1 pi.1936a. Studies on North American Mordellidae, I. Can. Ent., vol. 68, pp.124-129, 1 pi.1936b. The Neotropical Mordellidae of the Deutsches Entomologisches In-stitut, with descriptions of new species. Arb. Morph. Taxon. Ent.Berlin-Dahlem, vol. 3, Nos. 2, 3, pp. 142-150, 215-221, 8 figs.Say, Thomas.1835. Descriptions of new North American Coleoptera. Boston Journ. Nat.Hist., vol. 1, pp. 151-203.Smith, J. B.1882. A synopsis of the Mordellidae of the United States. Trans. Amer. Ent.Soc, vol. 10, pp. 73-100, 3 pis. U. S. GOVERNMENT PRINTING OFFICE: 1939