Neotropical Microlepidoptera, XXI: New Genera and Species of Oecophoridae from Chile J. F. GATES CLARKE SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY ? NUMBER 273 SERIES PUBLICATIONS OF THE SMITHSONIAN INSTITUTION Emphasis upon publication as a means of "diffusing knowledge" was expressed by the first Secretary of the Smithsonian. In his formal plan for the Institution, Joseph Henry outlined a program that included the following statement: "It is proposed to publish a series of reports, giving an account of the new discoveries in science, and of the changes made from year to year in all branches of knowledge." This theme of basic research has been adhered to through the years by thousands of titles issued in series publications under the Smithsonian imprint, commencing with Smithsonian Contributions to Knowledge in 1848 and continuing with the following active series: Smithsonian Contributions to Anthropology Smithsonian Contributions to Astrophysics Smithsonian Contributions to Botany Smithsonian Contributions to the Earth Sciences Smithsonian Contributions to the Marine Sciences Smithsonian Contributions to Paleobiotogy Smithsonian Contributions to Zoology Smithsonian Studies in Air and Space Smithsonian Studies in History and Technology In these series, the Institution publishes small papers and full-scale monographs that report the research and collections of its various museums and bureaux or of professional colleagues in the world cf science and scholarship. The publications are distributed by mailing lists to libraries, universities, and similar institutions throughout the world. Papers or monographs submitted for series publication are received by the Smithsonian Institution Press, subject to its own review for format and style, only through departments of the various Smithsonian museums or bureaux, where the manuscripts are given substantive review. Press requirements for manuscript and art preparation are outlined on the inside back cover. S. Dillon Ripley Secretary Smithsonian Institution S M I T H S O N I A N C O N T R I B U T I O N S T O Z O O L O G Y ? N U M B E R 2 7 3 Neotropical Microlepidoptera, XXI: New Genera and Species of Oecophoridae from Chile J. F. Gates Clarke SMITHSONIAN INSTITUTION PRESS City of Washington 1978 A B S T R A C T Clarke, J. F. Gates. Neotropical Microlepidoptera, XXI: New Genera and Species of Oecophoridae from Chile. Smithsonian Contributions to Zoology, number 273, 80 pages, 54 figures, 6 plates, 1978.?The following genera are described as new and their type-species are indicated: Doina (D. lagneia, n. sp.); Perzelia (P. arda, n. sp.); Teresita (T. isaura, n. sp.); Aliciana (A. geminata, n. sp.); Lelita (L. acmaea, n. sp.); Dita (D. phococara, n. sp.); Atha (A. trimacula, n. sp.); Afdera (Cryptolechia orphnaea Meyrick); Corita (C. amphichroma, n. sp.); Utilia (U. florinda, n. sp.); Alynda (A. sarissa, n. sp.); Deia (D. lineola, n. sp.); Doshia (D. miltopeza, n. sp.); Altiura (A. maculata, n. sp.); Muna (M. zostera, n. sp.); Lucyna (Cryptolechia fenestella Zeller); Talitna (anomala, n. sp.); Irenia (I. leucoxantha, n. sp.); Revonda (R. eschara, n. sp.); Despina (Borkhausenia rhodosema Meyrick); Zymrina (Borkhausenia xanthosema Meyrick; Eraina (E. thamnocephala, n. sp.); Retha (R. rustica, n. sp.); Nedenia (N. rhodochra, n. sp.); Aniuta (A. ochroleuca, n. sp.); Osmarina (O. argilla, n. sp.). Thirty-nine (39) species, including generic types, are described as new. Previously described species are placed in this present system as far as possible. The subfamily Metachandinae, new status, is noted. OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is recorded in the Institution's annual report, Smithsonian Year. SERIES COVER DESICN: The coral Montastrea cavernosa (Linnaeus). Library of Congress Cataloging in Publication Data Clarke, John Frederick Gates, 1905- Neotropical Microlepidoptera, XXI. (Smithsonian contributions to zoology ; no. 273) Bibliography: p. Supt. of Docs, no: SI 1.27:273 1. Oecophoridae. 2. Insects?Classification. 3. Insects?Chile. I. Title. II. Series: Smithsonian Institution. Smithsonian contributions to zoology ; no. 273. QL1.S54 no. 273 [QL561.043] 591'.08s [595.7'81] 77-608260 Contents Page Introduction 1 Key to the Chilean Genera of Oecophoridae 2 Eraina, new genus 3 Eraina thamnocephala, new species 3 Aniuta, new genus 5 Aniuta ochroleuca, new species 5 Aniuta melanoma, new species 7 Mattea Duckworth 7 Mattea phoenissa (Butler) 7 Pisinidea Butler 7 Pisinidea viridis Butler 7 Talitha, new genus 9 Talitha anomala, new species 10 Deia, new genus 10 Deia lineola, new species 12 Macarocosma Meyrick 12 Macarocosma philochrysa Meyrick 12 Perzelia, new genus 12 Perzelia arda, new species 14 Corita, new genus 15 Corita amphichroma, new species 15 Muna, new genus 17 Muna zostera, new species 17 Altiura, new genus 18 Altiura maculata, new species 19 Doina, new genus 19 Doina paralagneia, new species 21 Doina lagneia, new species 21 Doina flinti, new species 24 Doina inconspicua, new species 24 Doina asperula, new species 26 Doina truncata, new species 27 Doina subicula, new species 28 Doina annulata, new species 29 Doina eremnogramma, new species 30 Doina phaeobregma, new species 30 Doina trachycantha, new species 33 Doina glebula, new species 33 Doina increta (Butler), new combination 34 Doina edmondsii (Butler), new combination 34 Doina scariphista (Meyrick), new combination 35 Doshia, new genus 35 Doshia miltopeza, new species 36 iii j v SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY Page Ajdera, new genus 37 Afdera orphnaea (Meyrick), new combination 38 Melaneuha Butler 38 Melaneulia hectate Butler 38 Depressariod.es Turati 39 Depressariodes lusciosa (Meyrick), new combination 39 Depressariodes relegata (Meyrick), new combination 39 Gonionota Zeller 40 Gonionota mimulina (Butler), new combination 40 Nedenia, new genus 40 Nedenia rhodochra, new species 40 Revonda, new genus 42 Revonda eschara, new species 42 Despina, new genus 43 Despina rhodosema (Meyrick), new combination 44 lrenia, new genus 44 Irenia curvula, new species 44 lrenia leucoxantha, new species 45 Lucyna, new genus 45 Lucyna fenestella (Zeller), new combination 45 Dita, new genus 48 Dita phococara, new species 48 Dita fasciatipedella (Zeller), new combination 48 Utilia, new genus 49 Utilia luridella (Zeller), new combination 50 Utilia ochracea (Zeller), new combination 51 Utilia falcata, new species 51 Utilia florinda, new species 53 Alynda, new genus 53 Alynda striata, new species 54 Alynda sarissa, new species 55 Alynda cinnamomea, new species 56 Atha, new genus : 57 Atha trimacula, new species 57 Retha, new genus 58 Retha rustica, new species 58 Zymrina, new genus 59 Zymrina xanthosema (Meyrick), new combination 60 Callistenoma Butler 60 Callistenoma ustimacula (Zeller) 60 Lelita, new genus 60 Lelita acmaea, new species 60 Teresita, new genus 64 Teresita isaura, new species 64 Aliciana, new genus 64 Aliciana geminata, new species 65 Aliciana albella (Blanchard), new combination 66 Osmarina, new genus 66 Osmarina argilla, new species 67 NUMBER 273 Page Arctopoda Butler 67 Arctopoda maculosa Butler 69 Philomusea Meyrick 69 Philomusea craterias Meyrick 71 Philomusea brachyxista Meyrick 71 Philomusea meniscogramma, new species 71 Literature Cited 73 Plates 74 Neotropical Microlepidoptera Series 80 Neotropical Microlepidoptera, XXI: New Genera and Species of Oecophoridae from Chile J. F. Gates Clarke Introduction The purpose of this paper is to make known cer- tain genera and species that are representative of what appears to be a very large oecophorid fauna and to present these with the hope that my col- leagues in the Southern Hemisphere will attempt to relate the Chilean fauna to those, particularly, of Australia and New Zealand. I hope, also, that lepi- dopterists in Chile will attempt to expand our knowledge of that fauna. To date, only a comparatively few taxa in the Oecophoridae have been described from Chile. Blanchard (1852) appears to have been the first to describe an oecophorid from Chile when he pub- lished Epigraphia albella. Zeller (1874) seems to have been next when he described Cryptolechia ochracea, C. fasciatipedella, C. luridella, C. fene- stella, and C. ustimacula. Next, Butler added (1883) Arctopoda maculosa (=Polypseustis cuprea Dognin, 1908), which he described in the Tortricidae; Melaneulia hectate was also described in the Tor- tricidae. The following species were placed by Butler in the Gelechiidae: Orthotelia increta, De- pressaria edmondsii, Oecophora minneta, Hyper- skeles choreutidea, Callistenoma ustimacula zelleri, Cryptolechia phoenissa, Pisinidea viridis, and Ag- riocoma mimulina. In 1893 Bartlett-Calvert de- scribed Antithesia montana, without family designa- tion, but which has been placed in the Oecophoridae by Meyrick, and others, and which in fact belongs /. F. Gates Clarke, Department of Entomology, National Museum of Natural History, Smithsonian Institution, Wash- ington, D. C. 20560. in the Tortricidae. In 1915 Meyrick described Depressaria lusciosa, subsequently recognized from Chile, and added Depressaria relegata (1922a). It was not until 1931 that more Chilean Oecophoridae were described by Meyrick. He added Borkhausenia rhodosema, B. xanthosema, B. praesul (question- ably from Chile), Harpella semnodoxa, Macarocosma philochrysa, Philomusea craterias (described from Argentina but present in Chile), P. brachyxista, Cryptolechia scariphista, and C. orphnaea. Of the described species I have placed albella in Alicinia, new genus, ochracea and luridella in Utilia, new genus; fasciatipedella in Dita, new genus; fenetstella in Lucyna, new genus; increta, edmondsii, and scariphista in Doina, new genus; Cryptolechia phoenissa was transferred to Mattea (Duckworth, 1966); I place rhodosema in Despina, new genus; xanthosema in Zymrina, new genus; praesul of ques- tionable occurrence in Chile and not treated; orphanea in Afdera, new genus; mimulina in Goni- onota Zeller; lusciosa was transferred to Martyrhilda Clarke by Clarke (1963) and relegata to the same genus by Clarke (1965), but herein both are placed in the genus Depressariodes Turati. Of the described species, I am unable to place in this classification the following: Oecophora min- netta, Hyperskeles choreutidea, and Harpella sem- nodoxa. For the most part the species key to Australian, African, or Indian genera but those genera are described as having ocelli; all the Chilian species lack ocelli. The majority of the Chilean genera, therefore, appear to be endemic. The only tropical American genus that intrudes on the Chilean scene SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY is Gonionola Zeller. The genus Deprcssariodes Turati is widespread in the holarctic region and extends down the Andes and to the Juan Fernan- dez Islands. Specimens that are deposited in the National Museum of Natural History (NMNH), Smithsonian Institution, Washington, D.C., are listed under the catalog number of the former United States Na- tional Museum (USNM). ACKNOWLEDGMENTS.?Several people have helped in the preparation of this paper. I appreciate the help I have received from my old friend and col- league Dr. John D. Bradley, Commonweath Insti- tute of Entomology, London, who has repeatedly answered questions concerning certain types in the British Museum (Natural History); my thanks are due my friend and colleague Dr. Klaus Sattler, British Museum (Natural History), for arranging the loan of several types and other pertinent mate- rial housed in the collections of that institution. I wish to thank Jack F. Marquardt and Mrs. Carolyn Hahn, Smithsonian Library, for much help with library materials. My thanks are due several illus- trators who have contributed drawings: Andre1 del Campo Pizzini (deceased), formerly on the Smith- sonian staff, George L. Venable, Elaine Hodges, Lawrence M. Druckenbrod, and Ambrose Nelson, presently staff illustrators. My thanks are also due Victor E. Kranz for the photographs. Key to the Chilean Genera of Oecophoridae 1. Forewing with lc absent 2 Forewing with lc present 3 2. Abdominal terga setose Eraina, new genus Abdominal terga not setose Aniuta, new genus 3. Abdominal terga not setose 4 Abdominal terga setose 20 4. Veins 7 and 8 of forewing coincident to termen Matte a Duckworth Veins 7 and 8 of forewing otherwise 5 5. Veins 7 and 8 of forewing separate Pisinidea Butler Veins 7 and 8 of forewing otherwise 6 6. Antennal pecten present 7 Antennal pecten absent 8 7. Veins 2 and 3 of forewing stalked Depressariodes Turati Veins 2 and 3 of forewing separate Talitha, new genus 8. Vein 7 of forewing to costa 9 Vein 7 of forewing otherwise 15 9. Third segment of labial palpus as long as second 10 Third segment of labial palpus shorter than second 12 10. Forewing with vein lb simple Deia, new genus Forewing with vein lb furcate 11 11. Veins 3 and 4 of hind wing stalked Macarocosma Meyrick Veins 3 and 4 of hind wing connate Philomusea Meyrick 12. Hind tibia smooth 14 Hind tibia roughened with hairlike scales 13 IS. Vein 3 of forewing from well before angle of cell Perzelia, new genus Vein 3 of forewing from angle of cell Corita, new genus 14. Labial palpus upturned, exceeding vertex Mima, new genus Labial palpus porrect, or nearly so Altiura, new genus 15. Tongue vestigial; labial palpus greatly reduced Arctopoda Butler Tongue and labial palpus otherwise 16 16. Forewing tortriciform 17 Forewing otherwise Doitia, new genus 17. Vein 7 of forewing to termen 18 Vein 7 of forewing to apex; termen convex Doshia, new genus 18. Veins 4 and 5 of forewing connate Melaneulia Butler Veins 4 and 5 of forewing separate 19 19. Third segment of labial palpus less than half the length of second Ganionota Zeller Third segment of labial palpus more than half length of second Nedenia, new genus NUMBER 273 20. Vein 7 and 8 of forewing coincident 21 Vein 7 and 8 of forewing otherwise 22 21. Antennal pecten present; veins 4 and 5 of forewing connate Revonda, new genus Antennal pecten absent; veins 4 and 5 of forewing separate Despina, new genus 22. Vein 7 of forewing to apex 23 Vein 7 of forewing to tcrmen or costa 24 23. Scape with well-developed pecten Irenia, new genus Scape without pecten Lucyna, new genus 24. Apex of forewing falcate 25 Apex of forewing not falcate 27 25. Veins 3 and 4 of hind wing connate 26 Veins 3 and 4 of hind wing stalked Dita, new genus 26. Upper internal vein of forewing well developed Utilia, new genus Upper internal vein of forewing obsolete Alynda, new genus 27. Vein 7 of forewing to costa 28 Vein 7 of forewing to termen 32 28. Hind tibia roughened with hairlike scales 29 Hind tibia smooth Atha, new genus 29. Veins 3 and 4 of hind wing shortstalked Retha, new genus Veins 3 and 4 of hind wing connate 30 30. Third segment of labial palpus as long as second Zymrina, new genus Third segment of labial palpus noticeably shorter than second 31 31. Second segment of labial palpus broad, compressed Callistenoma Butler Second segment of labial palpus otherwise Ajdera, new genus 32. Hind tibia smooth 33 Hind tibia roughened with hairlike scales 34 33. Veins 3 and 4 of hindwing stalked Lelita, new genus Veins 3 and 4 of hindwing connate Teresita, new genus 34. Antennal pecten present Aliciana, new genus Antennal pecten absent Osmarina, new genus Eraina, new genus TYPE-SPECIES.?Eraina thamnocephala, new spe- cies (by monotypy and present designation). The gender of this generic name is feminine. Labial palpus upturned, exceeding vertex, second segment with some projecting scales at apex; third segment very slender, acute, as long as second. Maxillary palpus slender, free. Head roughened with loosely appressed scales; ocellus absent. Antenna simple; scape without pecten. Forewing costa slightly arched, termen oblique, 12 veins; 1b furcate; lc absent; 2 remote from 3, 4 nearer to 5 than to 3; 7 and 8 stalked, 7 to costa; 10 much closer to 9 than to 11; 11 from before middle of cell. Hind wing with 8 veins; 2 remote from 3; 3 and 4 con- nate; 5 nearer to 6 than to 4; 6 and 7 subparallel. Hind leg tibia smooth. Abdomen setose. Male genitalia with gnathos and uncus well developed. Female genitalia unknown. Eriana has many characters in common with Aniuta but differs from it by the much longer palpus, the loose scaling at apex of second segment, the furcation of lb of forewing, and the setose abdomen. Eraina thamnocephala, new species FIGURE I; PLATE la Alar expanse 17-21 mm. Labial palpus light cream color; third segment sparsely irrorate with fuscous. Antenna light cream color with small fuscous spot on each segment; scape with fine fuscous longitudinal line dorsally. Head and thorax light cream color. Forewing ground color pale cream color; extreme edge of basal fifth of costa fuscous; fuscous discal spots as follows: one in cell at basal third, one slightly be- yond in fold, and a transverse pair at end of cell; surface with or without ill-defined clay color patches and sparse fuscous irrorations; around termen a series of 4 or 5 fuscous spots, sometimes confluent; cilia grayish mixed with cream color. Hind wing ocherous white; surface, especially toward margins, SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 1.?Eraina thamnocephala, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus; d, ventral view of female genitalia. NUMBER 27S sprinkled with grayish-fuscous scales; outer edge narrow grayish fuscous; cilia mixed grayish-tipped cream and ocherous white. Foreleg cream color; tibia and tarsal segments heavily overlaid fuscous; tarsal segments cream color distally; midleg similar; hind leg suffused grayish fuscous. Abdomen cream color strongly suffused grayish fuscous; setose. Male genitalia slides USNM 24137, 24241. Harpe narrow; cucullus small, fingerlike; costa extended as a long, fleshy process, swollen distally; sacculus produced as a strongly sclerotized, slender, curved process. Gnathos a slender hook. Uncus broad basally, then narrowed and slightly dilated distally. Vinculum narrowly U-shaped, saccus slightly de- veloped. Tegumen slightly concave laterally. Anellus strongly sclerotized, narrow basally, broader at mid- dle, produced posteriorly as two sharply pointed processes. Aedeagus moderately slender; vesica armed with 4 or 5 weak, bluntly pointed cornuti. Female genitalia slide USMN 24242. Ostium with ventral lip deeply incised. Antrum granular. Incep- tion of ductus seminalis posterolateral. Ductus bursae partly membranous, anterior two-thirds granular. Bursa copjulatrix very finely granular. Signum an oval dentate plate. HOLOTYPE?USNM 73690. TYPE-LOCALITY.?Chile, Maule, Pelluhue, 600 m. Described from the male holotype (2 Dec 1953, L. E. Pefia), 13 male and one female paratypes all from the type-locality with same data as holotype. At first glance E. thamnocephala reminds one of Doina lagneia, new species, but averages paler in color. The two are, of course, generically distinct and can be separated easily on characters of the palpus and genitalia. Aniuta, new genus TYPE-SPECIES.?Aniuta ochroleuca, new species (by present designation). The gender of this generic name is feminine. Labial palpus upturned not reaching vertex; second segment slightly roughened beneath; third segment slender, acute, about as long as second. Maxillary palpus minute. Head roughened with loosely appressed scales; ocellus absent. Antenna of male serrulate, female simple; scape without pecten. Forewing costa nearly straight, termen oblique; 12 veins; lb furcation obsolescent; lc absent; 2 remote from 3; 3, 4, and 5 equidistant; 7 and 8 stalked, 7 to costa; 10 much closer to 9 than to 11; 11 from before middle of cell. Hind wing with 8 veins; 2 remote from 3; 3 and 4 connate (or very short stalked); 5 well removed from 4; 6 and 7 parallel. Hind leg tibia smooth. Abdominal terga not setose. Male genitalia with gnathos absent. Female genitalia with signum present. Aniuta is near Borkhausenia but differs from it by the absence of an ocellus, absence of pecten on the scape, and lack of cilia on the flagellum. More- over, the hind tibia of Aniuta is smooth, that of Borkhausenia is clothed with long, fine, hairlike scales. In the forewing lb is furcate in Borkhausenia but in Aniuta the furcation is obsolescent, one branch being scarcely indicated. Aniuta ochroleuca, new species FIGURE 2; PLATE lc Alar expanse 12-13 mm. Labial palpus pale ocherous white; second seg- ment, except apex, fuscous on outer side; third segment fuscous apically. Antenna buffy brown; scape fuscous dorsally. Head white; scales around base of scape fuscous. Thorax ocherous white; tegula white except base fuscous. Forewing ground color ocherous white to white; base of costa fuscous; at apical third of costa a fuscous spot; in middle of cell an oblique fuscous dash and a spot of same color at end of cell; at middle, dorsally, a fuscous suffusion and a shade of similar color around ter- men; cilia ocherous white with a suffused grayish subbasal line. Hind wing white, pale buff distally; cilia pale buff. Foreleg ocherous white with fuscous suffusion on outer side; midleg similar; hind leg ocherous white with a suggestion of a few obscure grayish spots. Abdomen ocherous white, slightly darker posteriorly; not setose. Male genitalia slides USMN 24121, 24238. Harpe ample, dorsal and ventral edges nearly parallel; cucullus rounded; sacculus produced distally as a sharp spine. Uncus broad basally, narrowed dis- tally. Vinculum with saccus produced, V-shaped. Tegumen short, broad. Anellus a small rectangular plate with a broad median anterior arm; posterior corners of central plate articulated with elements of transtilla. Aedeagus rather slender except broad- ened proximally; cornuti absent. SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 2.?Aniuta ochroleuca, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus; d, ventral view of female genitalia. NUMBER 273 Female genitalia slide USMN 24239. Ostium pear shaped. Antrum a sclerotized band. Inception of ductus seminalis near posterior end of ductus bursae. Ductus bursae very slender and sclerotized most of its length. Bursa copulatrix membranous. Signum a large, transverse, triangular, sclerotized plate with serrate anterior edge. HOLOTYPE.?USNM 73691. TYPE-LOCALITY.?Llanquihue, Peulla. Described from the male holotype (9 Mar 1959, J. F. G. Clarke), 2 male and 2 female paratypes as follows: male, Peulla (7 Mar 1959); male, 2 females, Petrohue (10-12 Mar 1959), all from Llanquihue Province, collected by J. F. G. Clarke. Aniuta melanoma, new species FIGURE 3; PLATE \b Alar expanse 12-14 mm. Labial palpus light buff; second segment with grayish-fuscous shade on outer side. Antenna pale ochraceous buff with grayish-fuscous annulations; scape fuscous dorsally, light ochraceous buff ven- trally. Head light buff, lightly infuscated. Thorax light buff, suffused grayish fuscous; tegula grayish fuscous basally. Forewing ground color grayish fuscous with scattered darker scales; in cell, about middle, and at end of cell, fuscous spots with a fuscous bar between and slightly dorsad; at apical third of costa a fuscous streak and inside that a broken fuscous shade; around apex and along termen a series of indistinct fuscous spots; cilia pale grayish mixed with grayish fuscous. Hind wing pale grayish buff, slightly darker toward mar- gins; cilia pale grayish buff, becoming darker around apex. Foreleg light buff, femur and tibia heavily overlaid fuscous on outer side; tarsal segments banded fuscous; midleg similar but dark markings not so extensive; hind leg light buff with scattered fuscous irrorations. Abdomen grayish fuscous dor- sally; ventrally buff with grayish fuscous irroration laterally. Male genitalia slide USNM 24240. Harpe of about equal width except narrowed before cucullus; cucullus truncate; sacculus extended as a sharply pointed process exceeding cucullus. Gnathos absent. Uncus very weak, short, triangular. Vinculum rounded. Tegumen constricted posteriorly. Anellus tubular, with basal plate and lateral flaps. Aedeagus stout proximally, pointed distally; vesica unarmed. Female genitalia slide USNM 24218. Ostium small, round, slightly protruding. Antrum not dif- ferentiated. Inception of ductus seminalis slightly before ostium. Ductus bursae slender, membranous with slight granulation before bursa copulatrix. Bursa copulatrix finely granular. Signum anterior, a folded plate, the folded edge forming a serrated keel, each outer edge with a median flange. HOLOTYPE.?USNM 73692. TYPE-LOCALITY.?Llanquihue, Peulla. Described from the male holotype (7 Mar 1959, J. F. G. Clarke) and one female paratype with same data except date (8 Mar 1959). This is similar to A. ochroleuca but with less contrasting markings of forewing. Moreover, the extension of sacculus exceeds cucullus in A. mela- noma but in A. ochroleuca the extension of sacculus is curved and does not exceed cucullus. Mattea Duckworth Mattea Duckworth, 1966:2, fig. 1, pi. 1: fig. a. [Type-species: Ciyptolechia phoenissa Butler, 1883:81, pi. 11: figs. 12, 12a; by monotypy and original designation.] Mattea phoenissa (Butler) Cryptolechia phoenissa Butler, 1883:81, pi. 11: figs. 12, 12a.? Bartlett-Calvert, 1886:347. Mattea phoenissa (Butler).?Duckworth, 1966:2, fig. 1, pi. 1: fig. a. TYPE.?British Museum (Natural History). TYPE-LOCALITY.?Corral. The single species M. phoenissa, belonging to this genus, exhibits the extraordinary character of hav- ing veins 7 and 8 of forewing coincident to termen. I know of no other American oecophorid genus with this character. Previously figured in Duck- worth (1966). Pisinidea Butler Pisinidea Butler, 1883:83. [Type-species: Pisinidea viridis Butler, 1883:83, pi. 11: fig. 10; by monotypy.] Pisinidea viridis Butler FIGURE 4 Pisinidea viridis Butler, 1883:83, pi. 11; fig. 10.?Bartlett- Calvert, 1886:347. Male genitalia slide USNM 24243. Harpe of nearly equal width throughout; cucullus broadly SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 3.?Aniuta melanoma, new species: a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus; c, ventral view of female genitalia. NUMBER 273 FIGURE 4.?Pisinidea viridis Butler: a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus; c, venation of right wings. rounded; sacculus broadly sclerotized; costa scle- rotized, terminating distally in a short point; clasper arising in middle of basal third of harpe, directed toward saccular margin. Gnathos absent. Transtilla U-shaped, arms of U pointed distally and profusely clothed with setae basally. Uncus slender, curved, pointed. Vinculum U-shaped; lateral arms broadest at articulation with tegumen. Tegumen hood shaped, broader than long. Anellus a large, divided, sclerotized plate. Aedeagus simple, moder- ately long; vesica apparently unarmed. Female genitalia unknown. HOLOTYPE.?British Museum (Natural History). TYPE-LOCALITY.?"Chiloe ? . . . " Unfortunately, the abdomen of the single speci- men available to me was destroyed when the geni- talia slide was prepared in 1929. Dr. John D. Brad- ley of the Commonwealth Institute of Entomology has examined a specimen in the British Museum and has informed me that the abdomen is not spined. The species is keyed accordingly. The generic type is the only species I place here. Talitha, new genus TYPE-SPECIES.?Talitha anomala, new species (by monotypy and present designation). The gender of this generic name is feminine. 10 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY Labial palpus slender, ascending; second segment slightly roughened anteriorly; third segment shorter than third. Maxillary palpus prominent, filiform, free. Head smooth, sidetufts spreading; ocellus ab- sent. Antenna simple (at least in female); scape with pecten. Thorax smooth. Forewing smooth, costa nearly straight, 12 veins; lb simple; lc present; 2, 3, and 4 nearly equidistant; 4 much nearer to 5 than to 3; 7 and 8 stalked, 7 to apex; 6, 7 + 8 and 9 equidistant, 10 much nearer to 9 than to 11; 11 from well before middle of cell; upper internal vein well developed. Hind wing with 8 veins; 2, 3, and 4 well separated; 5 approximate to 4; 5, 6, and 7 about equidistant, subparallel. Hind tibia clothed with hairlike scales. Abdominal terga setose. Female genitalia with signum present. This is an anomalous genus combining oecophorid and blastobasid characters. In the forewing veins 2 to 10 are bunched toward the outer end of cell but veins 2 and 3 arise from the cell as in the Oecophoridae. The signum is likewise oecophorid. The setae of the terga are missing in Talitha and are not present as a band of setae on the posterior edges of the terga as in the blastobasids. The genus differs from Depressariod.es by the separate veins 2 and 3 of forewing and by the bunching of veins 2 to 10. Talitha anomala, new species FIGURE 5; PLATE Id Alar expanse 23 mm. Labial palpus blackish fuscous; second segment white basally and speckled with white on inner side; inner side of third segment white. Antenna blackish fuscous; pecten white. Head blackish fuscous streaked with elongate white scales. Thorax blackish fuscous mixed with gray and white scales; tegula white tipped. Forewing ground color black- ish fuscous heavily overlaid white; at basal fifth, in cell, a blackish-fuscous spot followed at basal third of cell by a pair of confluent blackish-fuscous spots, these in turn followed by a pair of blackish- fuscous spots on upper vein of cell; at end of cell a transverse broken blackish-fuscous bar extends to tornus; from apex, along termen a short blackish- fuscous bar; cilia grayish mixed with white and blackish-fuscous scales. Hind wing grayish fuscous except cell that is gray devoid of grayish-fuscous scales; cilia ocherous white. Foreleg sordid white, heavily overlaid blackish fuscous on outer side; midleg similar but with less dark marking; hind leg ocherous white sparsely irrorate with grayish fuscous. Abdomen grayish fuscous dorsally, sordid white ventrally suffused grayish; abdominal terga not setose. Female genitalia slide USNM 24244. Ostium broadly funnel shaped. Antrum not differentiated. Inception of ductus seminalis well before ostium. Ductus bursae membranous in posterior two-fifths with most of this part with granular inner surface; anterior three-fifths spiraled, membranous. Bursa copulatrix membranous. Signum a triangular den- tate plate. HOLOTYPE.?USNM 73693. TYPE-LOCALITY.?Santiago, Guatacan. Described from the female holotype (Oct 1952, 1100 m, L. E. Pefia) and one female paratype with identical data. The paratype is slightly larger than the holotype and is not as contrastingly marked. In general facies T. anomala reminds one of the North American Valentinia glandulella (Riley) (Blastobasidae) but T. dnomala lacks the transverse submedian fascia of that species. Deia, new genus TYPE-SPECIES.?Deia lineola, new species (by monotypy and present designation). The gender of this generic name is feminine. Labial palpus slender, recurved but second seg- ment not reaching vertex, very slightly roughened anteriorly; third segment as long as second, slender, acute. Tongue well developed; maxillary palpus slender, appressed to base of tongue. Head rough, sidetufts spreading; ocellus absent. Antenna (female) slightly serrate and finely pubescent; scape without pecten. Thorax smooth. Forewing smooth, costa nearly straight, termen slightly sinuate, oblique, 12 veins; lb simple; lc present; 2 remote from 3; 3, 4, and 5 approximate; 7 and 8 stalked from well below angle, both to costa; 9 from angle; 10 much closer to 9 than to 11; 11 from beyond middle of cell; upper internal vein absent. Hind wing with 8 veins; 2 distant from 3; 3 and 4 connate; 5 ap- proximate to 4; 6 and 7 subparallel. Hind tibia roughened with a few loose, long scales. Abdominal terga not setose. NUMBER 273 11 FIGURE 5.?Talitha anomala, new species: a, venation of right wings; b, ventral view of female genitalia; c, signum enlarged. 12 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY Male genitalia unknown. Female genitalia without signum. Deia is similar to Borkhausenia but differs from it by the absence of pecten, the absence of an ocellus, having a smoother hind tibia, lb simple, and vein 2 of forewing remote from 3. Deia lineola, new species FIGURE 6; PLATE le Alar expanse 11-12 mm. Labial palpus white; second segment blackish fuscous in apical half; third segment with blackish scales scattered along anterior edge. Antenna black- ish fuscous spotted with white ventrally; scape longitudinally striped white and blackish fuscous. Head olivaceous gray. Thorax blackish fuscous, streaked with olivaceous gray. Forewing ground color blackish fuscous; from middle of costa to dorsum a broad white fascia with scattered blackish- fuscous scales; from apical fourth of costa to tornus a straight, white, transverse line; beyond this line a patch of olivaceous-gray scales; termen narrowly white; cilia gray mixed with olivaceous gray and blackish-fuscous scales basally. Hind wing light gray, somewhat darker toward margins. Foreleg ocherous white; femur and tibia suffused fuscous on outer side; tarsal segments spotted fuscous; mid- leg similar; hind leg ocherous white with slight grayish suffusion on outer side of tibia distally; tarsal segments spotted fuscous on outer side. Ab- domen grayish fuscous dorsally; ventrally sordid white. Female genitalia USNM 24207. Ostium small, funnel shaped. Antrum a sclerotized ring. Inception of ductus seminalis dorsal, at posterior edge of an- trum. Ductus bursae membranous except for a sclerotized area at about posterior third. Bursa copulatrix membranous. Signum absent. HOLOTYPE.?USNM 73694. TYPE-LOCALITY.?Llanquihue, Petrohue. Described from the female holotype (10 Mar 1959, J. F. G. Clarke) and three female paratypes with identical data, except one, 12 Mar 1959. Aside from certain species of Chionodes (Gele- chiidae) I know of no species with which D. lineola can be favorably compared, unless one refers to Borkhausenia praesul Meyrick from Argentina. In the case of B. praesul, it is a larger moth and the dark markings of D. lineola are more extensive than those of B. praesul. Moreover, B. praesul lacks the white transverse line at apical fourth of forewing, present in D. lineola. Macarocosma Meyrick Macarocosma Meyrick, 1931:393. [Type-species: Macarocosma philochrysa Meyrick, 1931; by monotypy.] Macarocosma philochrysa Meyrick FIGURE 7 Macarocosma philochrysa Meyrick, 1931:393.?Clarke, 1963, 4:314, pi. 154; figs. 1-lc. Female genitalia slide USNM 24215. Ostium pro- truding; ventral surface granular. Antrum cylin- drical, sclerotized. Inception of ductus seminalis ventral, at about anterior two-thirds of ductus bursae. Dustus bursae very slender, membranous. Bursa copulatrix with finely granular inner sur- face; signum a long, curved sclerotized rod with serrate edges. The male genitalia, wings, and wing venation have been figured in Clarke (1963) but the female genitalia are here illustrated for the first time. So far, only the single species of this genus has been discovered. Perzetia, new genus TYPE-SPECIES.?Perzelia arda, new species (by monotypy and present designation). The gender of the generic name is feminine. Labial palpus recurved, second segment exceed- ing vertex, very slightly roughened anteriorly; third segment slender, acute, slightly shorter than second. Tongue well developed; maxillary palpus, minute, free. Head smooth; side tufts spreading; ocellus absent. Antenna simple; scape without pecten. Thorax smooth. Forewing smooth, costa gently arched, termen nearly straight, oblique, 12 veins; lb furcate; lc well preserved; 2 remote from 3; 3 arising well before angle of cell; 4 and 5 connate, from angle of cell; 6 nearer to 7 than to 5; 7 and 8 stalked, 7 to costa; 9 nearer to stalk of 7 and 8 than to 10; 10 nearer to 9 than to 11; 11 from middle; upper internal vein absent. Hind wing with 8 veins; 2 remote from 3; 3 and 4 connate; 5 nearer NUMBER 273 13 FIGURE 6.?Deia lineola, new species: a, venation of right wings; b, ventral view of female genitalia. 14 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 7.?Macarocosma philochrysa Meyrick, ventral view of female genitalia. to 4 than to 6; 6 and 7 subparallel. Hind tibia roughened with hairlike scales. Abdominal terga not setose. Male genitalia with well-developed uncus and gnathos. Female genitalia unknown. In genitalia this genus is close to Doshia, new genus, but the position of vein 3 of forewing well before angle in Perzelia at once distinguishes it from Doshia. Perzelia arda, new species FIGURE 8; PLATE 1/ Alar expanse 15-18 mm. Labial palpus sordid white; second segment with light grayish suffusion anteriorly and on outer side; third segment grayish fuscous anteriorly. Antenna very pale grayish, annulated grayish fuscous; scape sordid white, infuscated dorsally. Head sordid white; face infuscated. Thorax sordid white. Fore- wing ground color sordid white; basally some pale ocherous suffusion; extreme edge of costa, at base, fuscous; underside of wing strongly overlaid fus- cous; some specimens show a few widely scattered grayish-fuscous scales on upper surface of forewing; cilia white. Hind wing pale grayish, darker toward margins; cilia mixed sordid white and grayish. Foreleg sordid white; tibia and tarsal segments gray- ish fuscous on outer sides; midleg similar; hind leg sordid white. Abdomen sordid white heavily over- laid grayish fuscous dorsally; first segment of male with median eversible scale tuft. Male genitalia slides USNM 24245, 24246. Harpe broad, tapering rather abruptly to the truncated cucullus; inside costa, near middle of harpe, a cluster of stout, truncated setae; before cucullus, on sacculus, a series of strong, short setae. Gnathos a spined, heart-shaped knob. Uncus large, hood shaped. Vinculum U-shaped. Tegumen subrectangu- lar, slightly longer than uncus. Anellus a subrec- tangular, sclerotized plate with median posterior projection and a slender digitate process from base on each side. Aedeagus long, curved, pointed; vesica armed with one long, strong cornutus. HOLOTYPE.?USNM 73695. TYPE-LOCALITY.?Maule, Pelluhue, 600 m. Described from the male holotype (2 Dec 1953, L. E. Pena) and 8 male paratypes, all with the same data. NUMBER 273 15 FIGURE 8.?Perzelia arda, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus. Perzelia arda is similar to T. isaura, new species, but has a much darker hind wing. Corita, new genus TYPE-SPECIES.?Corita amphichroma, new species (by monotypy and present designation). The gender of this generic name is feminine. Labial palpus recurved; second segment exceed- ing base of antenna, slightly roughened anteriorly; third segment slender, acute, shorter than second. Tongue well developed; maxillary palpus reduced, filiform, free. Head roughened with loosely ap- pressed scales, side tufts spreading; ocellus absent. Antenna of male ciliated, female antenna serrate; scape without pecten. Thorax smooth. Forewing smooth, costa gently arched, apex slightly produced, termen below apex slightly concave, 12 veins, lb furcate; lc strongly preserved; 2 distant from 3; 3 from angle of cell; 3, 4, and 5 nearly equidistant, approximate; 7 and 8 stalked, both to costa; 9 much nearer to stalk of 7 and 8 than to 10; 10 nearer to 9 than to 11; 11 from well before middle of cell. Upper internal vein absent. Hind wing with 8 veins; 2 remote from 3; 3 and 4 stalked; 5 about equidistant from 4 and 6; 6 and 7 subparallel. Hind tibia roughened with long hairlike scales. Ab- dominal terga not setose. Male genitalia with well-developed uncus and gnathos. Female with signum present. This genus appears to be very closely related to the New Zealand Gymnobathra Meyrick; in fact the only characters by which Corita differs from Gymnobathra are the absence of an ocellus and the stalking of veins 3 and 4 in the hind wing. In-as- much as Meyrick attributed an ocellus to many genera where there is none, we can discount that character and have left only the difference in the condition of veins 3 and 4 in the hind wing. Corita amphichroma, new species FIGURE 9; PLATE Ig Alar expanse 16-19 mm. Labial palpus second segment clay color; third segment buff. Antenna pale clay color with ill- defined fuscous annulations. Head pale clay color 16 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 9.?Corita amphichroma, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus; d, ventral view of female genitalia. NUMBER 273 17 laterally and anteriorly; buff dorsally. Thorax clay color. Forewing ground color clay color; three black discal spots, one on fold, one in cell at two-fifths and one at end of cell; costa with two buff spots, one, the smaller of the two slightly before middle, the larger one slightly beyond middle, the latter followed by an ill-defined fuscous blotch; from the outer angle of this blotch an outwardly curved row of small fuscous spots ending at tornus; cilia clay color except a fuscous apical group, the latter pre- ceded and followed by a buff patch. Hind wing ocherous white; veins and outer margin infuscated; cilia pale grayish fuscous. Foreleg buff; outer side clay color; midleg and hind leg light ochraceous buff. Abdomen ochraceous buff ventrally, pale gray- ish fuscous dorsally. Male genitalia slide USNM 24247. Harpe simple, costa strongly sclerotized; cucullus rounded. Gnathos an elongate, triangular spined plate. Uncus narrow, curved, pointed. Vinculum narrowly rounded. Teg- umen rather narrow, half as long as harpe. Anellus semitubular, deeply incised posteroventrally. Aedea- gus short, stout, distally pointed; vesica armed with a single strong cornutus. Female genitalia slides USNM 24248, 24249. Ostium cupped, transverse. Antrum very strongly sclerotized, broadly dilated anteriorly and with a dorsal spine at about two thirds. Inception of ductus seminalis dorsal about middle of antrum. Ductus bursae membranous, rather wide with two coils before bursa copulatrix. Bursa copulatrix very finely granular; appendix bursae membranous, small. Signum an oval, dentate plate. HOLOTYPE.?USNM 73696. TYPE-LOCALITY.?Centro-Austral. Described from the female holotype (Jan-Mar 1898, V. Izquerdo), three female and two male para- types with same data as holotype. The males have a much darker hind wing than the females but other characters agree. This species is similar superficially to some species of the genus Gonionota, particularly G. oligarcha (Meyrick). On the costa of the forewing of C. amphichroma there are two pale spots, on the costa of G. oligarcha only one. Moreover, G. oligarcha lacks the curved row of fuscous spots before termen present on the forewing of C. amphichroma. The labial palpus and genitalia immediately separate the two. Muna, new genus TYPE-SPECIES.?Muna zostera, new species (by monotypy and present designation). The gender of the generic name is feminine. Labial palpus recurved, exceeding vertex; second segment slightly roughened anteriorly; third seg- ment shorter than second, slender, acute. Tongue well developed; maxillary palpus slender, free. Head smooth, side tufts spreading; ocellus absent. An- tenna slightly thickened, pubescent (in male); scape without pecten. Thorax smooth. Forewing smooth, costa arched, termen straight, 12 veins; 1b furcate; lc very strongly preserved; 2 distant from 3; 3, 4, and 5 equidistant; 7 and 8 stalked, both to costa; 9, 10, and 11 about equidistant; 11 from before middle of cell; upper internal vein preserved. Hind wing with 8 veins; 2 remote from 3; 3 and 4 con- nate; 4, 5, and 6 about equidistant, subparallel. Hind tibia smooth. Abdominal terga not setose. Male genitalia with oval spined gnathos and hood-shaped uncus. Female genitalia not available. In Meyrick's key (1922b) this genus keys to the Indian and African genus Porthmologa. Muna differs from Porthmologa, however, by the absence of an ocellus, the smooth third segment of labial palpus, in vein 2 of forewing being remote from 3, and the smooth hind tibia. Moreover, the forewing of Porthmologa, is narrow with a very oblique termen while that of Muna is tortriciform with a straight termen. Muna zostera, new species FIGURE 10; PLATE \h Alar expanse 21 mm. Labial palpus straw color; second segment with ochraceous-tawny blotches on outer side; third seg- ment ochraceous tawny on outer side and anteriorly. Antenna grayish fuscous above, buff ventrally. Head grayish buff. Thorax light cinnamon brown; tegula concolorous. Forewing ground color light cinnamon brown with violaceous tinge; at basal third two ill-defined fuscous spots, one in cell, the other on fold; from three-fifths of costa a broad straw-yellow transverse fascia extends outwardly to middle of wing, then curves inwardly to middle of dorsum; both inner and outer edges of this fascia irregularly marked with cinnamon-buff scales; between outer 18 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 10.?Muna zostera, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus. edge of fascia and termen, a series of five ill-defined, grayish-fuscous spots; cilia cinnamon brown mixed with a few straw color scales. Hind wing ocherous white shading to very pale cinnamon brown at apex. Legs ocherous white; foreleg tibia and tarsal segments suffused grayish fuscous on outer side. Abdomen sordid whitish with violaceous tinge. Male genitalia slide USNM 24250. Harpe long and narrow, with a cluster of strong setae at base near costa; cucullus bluntly pointed; sacculus moderately sclerotized and with a very small point at its outer extremity. Gnathos an elongate, spined plate. Uncus hood shaped with median posterior point; posterior half of surface clothed with very fine, short setae. Vinculum rounded. Tegumen subrectangular, about as long as uncus. Anellus U-shaped. Transtilla a very narrow sclerotized band with a cluster of strong setae on each side. Aedeagus moderately slender, slightly curved; vesica unarmed. HOLOTYPE.?USNM 73697. TYPE-LOCALITY.?Chiloe Island, Dalcahue. Described from the unique male holotype (10-12 Feb 1954, L. E. Pena). The only species with which M. zostera can be compared is Cqptotelia cyathopa (Meyrick) but M. zostera has a complete pale transverse band of forewing and the remaining ground color of M. zostera is darker than in C. cyathopa. Altiura, new genus TYPE-SPECIES.?Altiura maculata, new species (by monotypy and present designation). The gender of this generic name is feminine. Labial palpus slightly ascending, nearly porrect; second segment slightly tufted anteriorly and pos- teriorly; third segment slender, acute, less than half the length of second. Tongue well developed; maxillary palpus filiform, free. Head very rough, scales on crown forming a tuft; ocellus absent. Antennae ciliated in male, simple in female; scape without pecten. Thorax smooth. Forewing smooth, costa arched, termen oblique, convex, 12 veins; lb furcate; lc strongly preserved; 2 well before 3; 3, 4, and 5 closely approximate; 7 and 8 stalked, 7 to costa; 9 much nearer to stalk of 7 and 8 than NUMBER 273 19 to 10; 11 from before middle; upper internal vein absent. Hind wing with 8 veins; 2 remote from 3; 3 and 4 connate or very short stalked; 4, 5, 6, and 7 about equidistant, subparallel. Hind tibia smooth. Abdominal terga not setose. Male genitalia with uncus and gnathos. Female genitalia with signum. Altiura keys to the New Zealand Izatha Walker but differs from it by the absence of tufts of scales on the forewing, the slightly ascending labial pal- pus, and the smooth posterior tibia. In female genitalia there appears to be a close relationship to the South American genus Gonionota Zeller. Altiura maculata, new species FIGURE 11; PLATE 2a,b Alar expanses 21-22 mm. Labial palpus light ochraceous buff; second seg- ment tawny on outer side and on inner side dis- tally; third segment pale ochraceous buff with a faint tawny streak at basal third on inner side. An- tenna tawny dorsally, pale ochraceous buff ven- trally. Head tawny mixed with ochraceous buff. Thorax ochraceous buff; tegula tawny. Forewing ground color light orange yellow; basal fifth of costa broadly tawny with a violaceous tinge; on middle of costa a similarly colored spot (in male a similar spot at end of cell and a smaller one in cell at one third); cilia orange yellow. Hind wing ocherous white, cilia concolorous. Legs light ochra- ceous buff. Abdomen light ochraceous buff; ab- dominal terga not setose. Male genitalia slide USNM 24251. Harpe of almost even width throughout; cucullus broadly rounded; sacculus an undulating sclerotized ridge in outer half; at base, inside costa, a small raised ridge. Gnathos broad, spoon shaped, broadly rounded distally. Uncus a half cylinder with a deep U-shaped excavation on posteroventral edge. Vin- culum rounded. Tegumen very broad basally, nar- rowed posteriorly. Anellus a broad, sclerotized, curved plate with a large winglike structure on each side, also from each basal angle a fleshy papilla. Aedeagus stout, curved, relatively short, pointed distally; vesica armed with a single, stout, curved cornutus. Female genitalia slide USNM 24206. Ostium very small, slitlike, transverse. Antrum a sclerotized ring. Inception of ductus seminalis lateral, slightly an- terior to antrum. Ductus bursae lightly rugose; anterior section before bursa copulatrix sclerotized. Bursa copulatrix very finely granular. Signum a broad dentate plate. HOLOTYPE.?USNM 73698. TYPE-LOCALITY.?Santiago Province, Cajon de Maypo, Cordillera, El Canelo. Described from the female holotype (12-20 Jan 1948, Tito Ramirez) and one male paratype (Peumo, 15 Mar 1951, J. Herrera). This has nearly the ground color of Gonionota mimulina (Butler) but the dark markings of A. maculata are much reduced. Doina, new genus TYPE-SPECIES.?Doina lagneia, new species (by present designation). The gender of this generic name is feminine. Labial palpus recurved, second segment reaching or exceeding vertex, smooth; third segment slender, acute, shorter than second. Tongue well developed; maxillary palpus very small, filiform, appressed to base of tongue. Head smooth; side tufts spreading; ocellus absent. Antenna pubescent in male, simple in female, scape without pecten. Thorax smooth. Forewing smooth, costa gently arched, termen nearly straight, slightly oblique, 12 veins; lb fur- cate; 1c strongly preserved, 2, 3, 4 approximate; 4 and 5 connate or very closely approximate; 5, 6, 7 nearly parallel; 7 and 8 stalked, 7 to apex, or to termen very slightly below apex; 9 nearer to stalk of 7 and 8 than to 10; 10 nearer 9 than to 11; 11 from middle. Hind wing with 8 veins; 2 distant from 3; 3 and 4 connate or very short stalked; 5 nearer 6 than to 4; 5-8 nearly parallel. Hind tibia smooth or with a few short, loosely appressed scales at base. Abdominal terga not setose. Male genitalia with well-developed uncus and gnathos. Female genitalia with signum (except D. phaeo- bregma). This "genus" is closely related to Psilocorsis but differs from it by the absence of the longitudinally striped labial palpus found consistently in Psilo- corsis, vein 11 of forewing from middle of cell and vein 5 of hind wing nearer to vein 6 than to 4. The species grouped under Doina show some variation. Of the species in which males are known, 20 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 11.?Altitira macttlata, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and acdeagus removed; c, aedeagus with part of anellus attached; d, ventral view of female genitalia; e, signum, enlarged. NUMBER 273 21 some have a median ventral eversible scale tuft from the first abdominal segment, some do not. Of the known males, six have a cluster of strong setae at the base of harpe, five do not, although in other features they are consistent, except D. glebula has a divided uncus and probably requires a different genus, but keys to this group. The known females (except D. scariphista and D. phaeobregma) have a multibranched signum. Doina paralagneia, new species FIGURE 12; PLATE 2c Alar expanse 18-20 mm. The description of the following, D. lagneia, will suffice for D. paralagneia except spot at middle of costa of forewing D. paralagneia more conspicu- ous than in D. lagneia and forewing with more of an orange tinge. Foretibia brownish or fuscous. Abdomen with ventromedian pocket containing scale tuft on first and second segments of male. Male genitalia slides USNM 24122, 24124, 24125. Harpe broad, stubby, simple, scarcely longer than tegumen; cucullus rounded. Gnathos a transverse, oval, spined knob. Uncus as long as tegumen, some- what truncated distally. Vinculum U-shaped. Tegu- men about as broad as long. Anellus a strongly sclerotized subrectangular plate; from each corner, basally, a strongly sclerotized lobe, dilated distally; opposite corners, each with a papilla directed an- teriorly. Aedeagus C-shaped; vesica armed with cluster of slender cornuti. Female genitalia silde USNM 24123. Ostium very small, protruding lamella antivaginalis sclerotized, cupped. Antrum not differentiated. Inception of ductus seminalis from slightly before ostium. Ductus bursae long, looped, and reversed on itself. Bursa copulatrix oval, membranous; appendix bursae present, membranous. Signum a long, branched, sclerotized plate extending full length of bursa copulatrix. HOLOTVPE.?USNM 73700. TYPE-LOCALITY.?Centro-Austral. Described from the male holotype, 2 male para- types, Centro-Austral (Jan-Mar 1898, V. Izquerdo); male paratype, Arauco, Caramavida (1-6 Jan 1954, L. E. Pena): and one female paratype, Province Cautin, near Pucon (4 Jan 1966, O. Flint and Cekalovic). This and the following fly together and are, for all intents and purposes, indistinguishable in the field. Even when one has the two side by side the only superficial character that can be used to separate them is the color of the foretibia, which is carmine or carmine-tinged on outer side in D. lagneia and brownish or fuscous in D. paralag- neia. Moreover, the male of D. paralagneia has a ventromedial pocket, containing a scale tuft, on the first and second abdominal segments that is lacking in D. lagneia. Unfortunately, this character can only be seen in dissected specimens. The male genitalia present excellent characters for separation as can be seen from the figures. The differences between the aedeagi are so striking that there is no difficulty in distinguishing between them. Unfortu- nately, there is only one female specimen that I have placed with D. paralagneia on the color of the foretibia, so no comparison between the two species, based on females, can be made. Doina lagneia, new species FIGURE 13; PLATE 2d Alar expanse 20-22 mm. Labial palpus light ochraceous buff; second seg- ment overlaid clay color. Antenna brown dorsally, clay color ventrally; scape light ochraceous buff ventrally. Head light buff to light ochraceous buff. Thorax buff to light ochraceous buff, sometimes overlaid and irrorate clay color; tegula with extreme base fuscous. Forewing ground color light ochrace- ous buff; extreme base of costa fuscous; from mid- costa around apex and termen a series of 11 fuscous spots, the last, on tornus, more of a longitudinal streak; at basal third, in cell, a fuscous spot; a similar, but larger, spot on fold and one at end of cell (in some specimens these spots are obscure or absent); surface of wing sparsely irrorate with clay color and fuscous scales; cilia various, clay color, grayish or cinnamon buff, sometimes mixed. Hind wing ocherous white, darker toward margins; cilia light buff. Foreleg light buff; tibia carmine on outer side; tarsal segments annulated fuscous; midleg light buff; outerside of femur, distally, infuscated; tibia shaded with fuscous basally and irrorate with fuscous on outer side; hind leg light buff; terminal four basal segments clouded grayish fuscous. Ab- domen light buff to light ochraceous buff; ventro- laterally a few ill-defined grayish-fuscous spots on 22 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 12.?Doina paralagneia, new species: a, ventral view of male genitalia with left harpe and actleagus removed; b, aedeagus; c, ventral view of female genitalia; d, detail of signum, enlarged. NUMBER 275 23 FIGURE. 13.?Doina lagneia, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus; d, lateral outline of male genitalia with aedeagus in situ; e, ventral view of female genitalia. 24 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY posterior segments; dorsally centers of segments grayish fuscous; abdominal terga not setose. Male genitalia slides USNM 24126, 24127, 24128, 24136, 24252, 25253. Harpe simple, longer than tegumen and uncus combined; cucullus narrowly rounded. Gnathos a pear-shaped, spined knob. Uncus as long as tegumen, broad basally and bluntly pointed distally. Vinculum U-shaped. Tegu- men subrectangular, nearly as broad as long. Anel- lus a lighty sclerotized plate with a fleshy, finger- like lobe from base on each side. Aedeagus about as long as uncus and tegumen combined, curved, pointed distally; vesica armed with a cluster of long slender cornuti. Female genitalia slides USNM 24134, 24136. Ostium very small, conical. Antrum not differenti- ated. Inception of ductus seminalis well before os- tium, ventral. Ductus bursae long, membranous, reversed on itself. Bursa copulatrix membranous. Signum posterior, a short multibranched plate. La- mella antevaginalis a narrow sclerotized band. HOLOTYPE.?USNM 73699. TYPE-LOCALITY.?Lautara. Described from the male holotype (Nov 1895, V. Izquerdo), and 11 male and two female paratypes as follows: 9 males, Centro-Austral (Jan-Mar, V. Izquerdo); male, Malleco, Rio Blanco (21-24 Nov 1954, L. E. Pena); male, 2 females, Llanquihue, Petrohue (10 Mar 1959, J.F.G. Clarke). This is a variable species, one male exhibiting considerable grayish clouding in the basal half of forewing, and a paler, yellowish ground color than is found in most specimens. One female has a pale yellowish ground color to the forewing and con- spicuous discal spots. Under D. paralagneia, I have discussed the dif- ferences between the two. Doina flinti, new species Fict'RF. 14; PLATE 2e Alar expanse 22 mm. Labial palpus light buff; second segment light ochraceous buff on outer side; third segment light ochraceous buff laterally. Thorax light ochraceous buff; tegula paler posteriorly. Forewing ground color light buff; extreme costal edge white; in- wardly, narrowly clay color; from middle of costa, around outer end of cell to fold, a clay color blotch; from basal third of costa an ill-defined clay color fascia extends to dorsum; in middle of cell a fus- cous speck; at end of cell a well-defined fuscous spot; cilia light buff suffused basally with light ochraceous buff. Hind wing whitish with slight buff shading terminally; cilia concolorous. Foreleg light buff shaded light orchraceous buff on outer side; midleg similar; hind leg light buff. Abdomen buff; first two segments shaded light ochraceous buff dor- sally. Female genitalia slide USNM 24209. Ostium funicular, rather broad. Antrum not differentiated. Inception of ductus seminalis lateral, slightly before ostium. Ductus bursae finely granular posteriorly, membranous anteriorly, very long, twisted. Bursa copulatrix very finely granular; appendix bursae small, membranous. Signum a branched plated with 8 to 9 branches on a side. HOLOTYPE.?USNM 73701. TYPE-LOCALITY.?Province Cautin, near Pucon. Described from the unique female holotype (4 Jan 1966, O. Flint and Cekalovic). Structurally, D. flinti is probably nearest D. subi- cula, new species, but has none of the dark mark- ings of that taxon. I am pleased to dedicate this species to Dr. Oliver S. Flint, one of the collectors. Doina inconspicua, new species FIGURE 15; PLATE 2/ Alar expanse 26-30 mm. Labial palpus sordid white; second segment heav- ily overlaid with drab and gray on outer side and anteriorly. Antenna very pale cinereous; scape drab. Head drab. Thorax drab. Forewing ground color light drab; surface of wing irrorate with scattered grayish-fuscous scales; at two-fifths, in cell, an ill- defined, small fuscous discal spot; at end of cell a similar but larger spot; a third similar spot on fold (in some specimens these spots are missing); between end of cell and termen a series of three or four small, ill-defined fuscous spots; along termen, to tornus, a series of seven ill-defined fuscous spots; cilia a mixture of drab and cinereous scales. Hind wing sordid white, the surface toward margins irro- rate with fuscous scales; cilia sordid white with narrow drab subbasal and subterminal lines. Fore- leg and midleg sordid white, strongly overlaid drab NUMBER 273 25 FIGURE 14.?Doina flinti, new species, ventral view of female genitalia. 26 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 15.?Doina inconspicua, new species: a, ventral view of male genitalia with left harpe and aedeagus removed, b, aedeagus. on outer sides; hind leg sordid white mottled drab. Abdomen sordid white heavily overlaid drab and irrorate fuscous; abdomen of male with eversible median scale tuft from first segment. Male genitalia slide USNM 24254. Harpe elon- gate, simple, about twice as long as tegumen; cucul- lus broadly rounded. Gnathos a pear-shaped, spined knob. Uncus very board, posterior edge excavated. Vinculum U-shaped. Tegumen, one-and-a-half times length of uncus. Anellus broadly oval with a pair of digitate processes from base. Aedeagus stout, curved, pointed distally, about as long as harpe; vesica armed with a strong cornutus arising from a rectangular plate. HOLOTYPE.?USNM 73702. TYPE-LOCALITY.?Arauco, Caramavida. Described from the male holotype (25-31 Dec 1953, L. E. Pefia), one male paratype with same data (except 1-6 Jan 1954), and one male paratype, Santiago, Guayacan (Oct 1952, 1100 m, L. E. Pefia). This species averages a little large than D. erem- nogramma and lacks any of the longitudinal dark markings of that species. Moreover, D. inconspicua lacks the cluster of stout setae at the base of harpe as in D. eremnogramma. Doina asperula, new species FIGURE 16; PLATE 2g Alar expanse 23-24 mm. Labial palpus sordid white; second segment speckled with grayish-fuscous scales. Antenna gray- ish with very narrow fuscous annulations; scape fus- cous. Head sordid white, laterally edged fuscous. Thorax sordid white infuscated anteriorly. Fore- wing ground color light cinnamon buff; base of wing a narrow white patch edged outwardly by fuscous; from slightly beyond middle of costa a broad, trans- verse gray shade extends to dorsum and curves in- wardly to basal white patch; outer edge of gray shade marked by an outwardly curved line of black spots; costa, beyond beginning of gray shade and termen, grayish fuscous; cilia a mixture of cinereous and grayish fuscous scales. Hind wing very pale grayish basally, lightly infuscated toward margin; margin narrowly grayish fuscous; cilia sordid white, slightly infuscated. Foreleg ocherous white; tibia and tarsal segments suffused fuscous on outer sides; midleg similar; hind leg ocherous white. Abdomen grayish fuscous dorsally, ocherous white ventrally. Female genitalia slide 24255. Ostium small, fun- nel shaped. Antrum not differentiated. Inception of ductus seminalis lateral, slightly anterior to ostium. Ductus bursae membranous. Bursa copulatrix mem- branous. Signum an elongate, many branched sclerotized plate. HOLOTYPE.?USNM 73703. TYPE-LOCALITY.?Maule, Constitution. Described from the female holotype. (26 Nov 1953, L. E. Pefia) and one female paratype with identical data. Like D. subicula, D. asperula has a small white patch at base of forewing. In addition, D. asperula has a cinnamon buff shade inside basal half of costa, entirely lacking in D. subicula. NUMBER 273 27 Doina truncata, new species FIGURE 17; PLATE 2h Alar expanse 16-22 mm. Labial palpus pale buff suffused and irrorate with fuscous on outer surface of second segment; an- terior and posterior edges of second segment very pale pink; third segment very sparsely dusted fus- cous. Antenna buff, suffused fuscous. Head drab mixed fuscous and cinereous. Thorax light ochrace- ous buff. Forewing ground color light ochraceous buff shading to ochraceous tawny before a broad light buff terminal area; outer half of wing very sparsely irrorate with fuscous; on costa, just before apex, an ill-defined, tiny fuscous dot; along termen a series of 5 fuscous spots; lying on tornal edge a blackish streak; cilia drab. Hind wing pale grayish FIGURE 16.?Doina asperula, new species: a, ventral view of female genitialia; b, signum, enlarged. FIGURE 17.?Doina truncata, new species: a, ventral view of male genitalia with left harpe and aedcagus removed; b, aedcagus. 28 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY fuscous, paler basally; on termen three or four ill- defined fuscous spots and apical portion of wing speckled with fuscous. Foreleg pale buff; tibia suf- fused and irrorate fuscous; 1st tarsal segment with longitudinal orange-ocherous dash; midleg pale buff; femur and tibia suffused and speckled grayish fuscous; hind leg pale buff with a few scattered fuscous scales. Abdomen pale buff, suffused light ocherous posteriorly. Male genitalia slide USNM 24256. Harpe ample, gently tapered to the rounded cucullus; base of in- ner surface clothed with a cluster of stout setae. Gnathos a spined knob. Uncus triangular; broad basally, tapering to a point posteriorly. Vinculum narrow U-shaped. Tegumen narrow, about as long as vinculum, finely scobinate laterally. Anellus a small sclerotized plate with large lateral lobes. Aedeagus short, stout, curved, bulbous basally; vesica armed with a few slender cornuti. HOLOTYPE.?USNM 73704. TYPE-LOCAUTY.?Centro-Austral. Described from the male holotype (Jan-Mar 1898, V. Izquerdo), and one female paratype (with- out abdomen), Lautaro (Nov 1895, V. Izquerdo). The female is more ochraceous orange than the male before the buff terminal area. In genitalia D. truncata is probably nearest to D. subicula but the pointed uncus of D. truncata im- mediately distinguishes it from D. subicula. Doina subicula, new species FIGURE 18; PLATE 3a Alar expanse 24-25 mm. Labial palpus fuscous; second segment with longi- tudinal area and apex buff; third segment apex buff. Antenna fuscous; scape with buff spot dorsally at apex. Head drab, side tufts and frons infuscated. Thorax drab suffused fuscous, tegula fuscous with light ochraceous-buff apex. Forewing ground color fuscous; basal angle light ochraceous buff; on costa, to apex, a series of ill-defined light ochraceous-buff dots; in cell a blackish dash joined at end of cell by a transverse bar of the same color; between cell and termen an ill-defined, outwardly curved series of light ochraceous-buff spots; termen narrowly edged blackish fuscous; cilia drab and fuscous, tipped with sordid buff; underside of costa buff. Hind wing light grayish olive with darker line around outer edge; cilia light grayish olive. Foreleg FIGURE 18.?Doina subicula, new species: a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus. buff overlaid fuscous on outer side; tarsal segments annulated buff; midleg similar but tibia with two buff annuli; hind leg buff with some infuscation on tibia and tarsal segments. Abdomen buff with median fuscous line dorsally and series of four in- distinct fuscous spots laterally. Male genitalia slide USNM 24257. Harpe ample, tapering slightly to rounded cucullus; base of in- ner surface clothed with cluster of stout setae; sac- culus narrowly sclerotized. Gnathos a broad, trans- verse, subtriangular spined knob. Uncus as long as tegumen, hood shaped, narrowed posteriorly. Vin- culum U-shaped. Tegumen nearly as broad as long, lightly scobinate laterally. Anellus a small, sclero- NUMBER 273 29 tized plate with large flattened lateral lobes. Aedea- gus stout, pointed, strongly curved, and bulbous basally; vesica with elongate roughened area. HOLOTYPE.?USNM 73705. TYPE-LOCALITY.?Chile, Llanquihue, Peulla. Described from the male holotype (7 Mar 1959, J.F.G. Clarke) and one male paratype with identical data. The nearest relative of this is D. phaeobregma, from which it is distinguished by the darker ground color and the presence of the light ochraceous-buff basal angle absent in D. phaeobregma. The male genitalia are similar but the gnathos of D. subicula is short and broad, that of D. phaeobregma long and narrow. Doina annulata, new species FIGURE 19; PLATE 3b Alar expanse 24 mm. Labial palpus fuscous, narrowly buff on the in- side posteriorly. Antenna light gray color, the light clay color almost obscured by fuscous annulations; scape fuscous except for buff apex posteriorly. Head grayish clay color. Thorax mixed clay color and fuscous. Forewing ground color light clay color; from extreme base of costa to fold a short black, transverse dash; from fold to costa basal two-thirds of wing marked with fuscous and black scales, espe- cially along fold and between end of cell and costa; in middle of cell a black dash followed at end of cell by a similarly colored L-shaped mark; before termen three ill-defined fuscous blotches; around apex, along termen to tornus, a series of fuscous dashes forming almost a continuous narrow line; cilia clay color mixed with fuscous. Hind wing olive buff with narrow grayish-fuscous terminal line; cilia similarly colored with a grayish-fuscous sub- basal band. Foreleg buff heavily overlaid fuscous; midleg similar; hind leg buff with slight infuscation on outer side. Abdomen pale grayish with slight yellowish tinge; first segment almost wholly infus- cated dorsally, other segments with only slight in- fuscation; ventrally buff, somewhat infuscated pos- teriorly. Male genitalia slide USNM 24258. Harpe broad- est at base, tapering gradually to a narrowly rounded cucullus; near base, in middle, a cluster of stout setae. Gnathos a broadly triangular spined knob. Uncus broadly hood shaped, slightly longer FIGURE 19.?Doina annulata, new species: a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus. than tegumen. Vinculum narrowly U-shaped. Tegu- men short, less than, half the length of harpe, broader posteriorly than anteriorly. Anellus a scle- rotized plate with a fleshly lobe on each side at base. Aedeagus robust, sharply bent, pointed dis- tally; vesica armed with a cluster of long strong setae. HOLOTYPE.?USNM 73706. TYPE-LOCALITY.?Gen tro-Austral. Described from the male holotype (Jan-Mar 1898, V. Izquerdo). The markings of D. annulata are similar to those of Cryptolechia veniflua Meyrick, but are more sharply defined; and D. annulata is a much smaller insect. In genitalia D. annulata and D. subicula, new species, are very closely related but the vesica of D. annulata is armed with a cluster of strong cornuti, absent in D. subicula. 30 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY Doina eremnogramma, new species FIGURE 20; PLATE 3C Alar expanse 28-31 mm. Labial palpus sordid white speckled with grayish fuscous. Antenna grayish; scape fuscous dorsally. Head cinereous mixed with a few fuscous scales. Thorax cinereous with fuscous lateral line and gray- ish suffusion centrally; tegula fuscous. Forewing ground color cinereous; from base of wing, between veins 11 and 12, a fuscous line extends to almost middle of costa; between veins 9 and 10 and 10 and 11, short fuscous streaks; similar but ill-defined lines between other veins; from apical third of costa, around apex and along termen to tornus, a series of 11 fuscous streaks; in cell a fuscous longitudinal streak and a grayish-fuscous shade; on fold, at about two-fifths, a short fuscous streak; apical two-fifths of wing lightly infuscated; cilia mixed cinereous and grayish fuscous. Hind wing sordid white, lightly infuscated and sparsely irrorate with small grayish- fuscous scales. Foreleg sordid white; femur, tibia, and tarsal segments heavily overlaid fuscous on outer sides; midleg sordid white; femur fuscous apically; tibia and tarsal segments lightly infus- cated; hind leg ocherous white, irrorate grayish fuscous; tarsal segments lightly suffused grayish fuscous. Abdomen sordid white dorsally with slight grayish suffusion; ventrally irrorate with grayish fus- cous. Male genitalia slide USNM 24259. Harpe simple, with rounded cucullus; at base a cluster of short, stout setae. Gnathos spined, reniform. Uncus as long as tegumen, hood shaped, bluntly pointed dis- tally. Vinculum U-shaped. Tegumen subrectangular. Anellus a rectangular plate, with a strong, curved blade extending beyond base of harpe on each side. Aedeagus stout, strongly curved, pointed; vesica armed with a single, long cornutus. Female genitalia slide USNM 24260. Ostium small, funnel shaped. Antrum undifferentiated. In- ception of ductus seminalis slightly before ostium. Ductus bursae broad, slightly sclerotized and looped posteriorly, remainder membranous. Bursa copula- trix membranous. Signum a long, multibranched, sclerotized plate. HOLOTYPE.?USNM 73707. TYPE-LOCALITY.?Arauco, Caramavida. Described from the male holotype (1-6 Jan 1954, L. E. Pena) and one female paratype, Maule, Con- stituti6n (16 Nov 1953, L. E. Pena). In male genitalia D. eremnogramma appears to be nearest D. trachycantha but the gnathos of D. eremnogramma is reniform, that of D. trachycantha is split into two separate elements. The two blade- like structures on each side of the anellus immedi- ately separate D. eremnogramma from any near relatives. Doina phaeobregma, new species FIGURE 21; PLATE 3e Alar expanse 20-23 mm. Labial palpus brussels brown; inner surface of second segment with narrow buff longitudinal streak; third segment buff with anterior edge fus- cous for entire length. Antenna buff with bronze cast ventrally, semiannulated grayish fuscous dor- sally; scape fuscous. Head drab, vertex violaceous. Thorax light brussels brown. Forewing ground color light brussels brown; extreme edge of costa rufous; from apical half of costa, around termen to tornus, twelve tiny salmon-buff spots; in middle of cell a fuscous spot; on fold a similarly colored, larger spot; at end of cell a short fuscous transverse dash; subterminally, from vein 2 to vein 7 a series of five fuscous spots, each preceded inwardly by buff scales; termen narrowly edged fuscous; underside of costa and longitudinal streaks between the veins, buff; cilia light brussels brown. Hind wing very pale grayish fuscous; cilia buff with some very pale gray- ish fuscous at apex. Foreleg buff; outer sides of tibia and tarsal segments brussels brown; midleg similar; hind leg buff with slight grayish suffusion on outer side. Abdomen grayish fuscous dorsally, buff ven- trally; laterally, a series of four blackish spots in posterior half; with paired ventral hair pencil from first sternum. Male genitalia slides USNM 24261, 24262. Harpe long, slender; cucullus rounded; sacculus very nar- rowly sclerotized; at base, on inner surface, a patch of strong setae. Gnathos a spined knob. Uncus al- most as long as tegumen, sharply pointed. Vincu- lum subtriangular. Tegumen narrow, about two- thirds the length of the harpe; scobinate laterally. Anellus a narrow transverse plate with large, flat, lateral lobes. Aedeagus stout, slightly curved; apex sharply pointed; base bulbous; cornuti absent. NUMBER 273 31 FIGURE 20.?Doina eremnogramma, new species: a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus; c, ventral view of female genitalia. 32 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 21.?Doina phaeobregma, new species: a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus; c, ventral view of female genitalia. NUMBER 273 33 Female genitalia slide USNM 24263. Ostium fun- nel shaped; lamella postvaginalis a broadly sclero- tized transverse band; lamella antevaginalis a very narrow sclerotized band. Antrum not differentiated. Inception of ductus seminalis slightly anterior to ostium. Ductus bursae membranous. Bursa copu- latrix membranous. Signum absent. HOLOTYPE.?USNM 73708. TYPE-LOCALITY.?Centro-Austral. Described from the male holotype (Jan?Mar 1898, V. Izquerdo), 11 male and 1 female paratypes with same data as holotype; two males (Chile, V. Izquerdo). In male genitalia D. phaeobregma is very near D. truncata but with a more sharply pointed uncus. Moreover, D. phaeobregma lacks the buff terminal area found in D. truncata. The absence of a signum at once separates D. phaeobregma from all other species of this group. Doina trachycantha, new species FIGURE 22; PLATE 3d Alar expanse 26 mm. Labial palpus buff; second segment with tawny suffusion on outer side; third segment pale tawny. Antenna buff, strongly overlaid tawny. Head buff, mixed tawny. Thorax buff mixed with tawny; tegula tawny proximally. Forewing ground color buff; tawny suffusion along costa; surface of wing sprin- kled with tawny irrorations; in middle of cell a slender fuscous streak; at end of cell a small fuscous spot followed by tawny; from just before apex, on costa, around tornus, a series of small fuscous spots between the veins; cilia mixed buff and tawny. Hind wing very pale cinereous with slight grayish suffu- sion; cilia sordid white. Foreleg buff; femur and tibia suffused tawny on outerside, tibia with a streak of carmine; tarsi almost wholly tawny; midleg simi- lar; hind leg buff. Abdomen pale brassy yellow strongly overlaid anterodorsally with fuscous, less so posterodorsally; ventrally ocherous white; on each side ventrolaterally a row of two or three fuscous spots. Male genitalia slide USNM 24264. Harpe about twice length of tegumen, of nearly equal width throughout; cucullus rounded; at base, just inside sacculus, a cluster of sharply pointed stout setae. Gnathos a pair of spined knobs. Uncus as long as tegumen, rather narrow, bluntly pointed dis- tally. Vinculum U-shaped. Tegumen about as long as broad; outer surface granular. Anellus a broadly crescentic plate with a fleshy lobe on each side at base. Aedeagus stout, curved, pointed; vesica armed with a pair of pear-shaped cornuti. HOLOTYPE.?USNM 73709. TYPE-LOCALITY.?Centro-Austral. Described from the unique male holotype (Jan- Mar 1898, V. Izquerdo). This species is nearest D. eremnogramma, new species, but the setal cluster of harpe of D. trachy- cantha is much nearer sacculus than in D. eremno- gramma and the latter has a single long cornutus, not two as in D. trachycantha. The genitalia, with double gnathos, is atypical but on all other char- acters the species belongs here. Doina glebula, new species FIGURE 23; PLATE 3/ Alar expanse 20 mm. Labial palpus sordid white, shaded and dusted with fuscous on outer side. Antenna grayish fuscous with very narrow paler annulations; scape fuscous. Head a mixture of grayish and white-tipped fuscous scales. Thorax a mixture of fuscous and grayish scales. Forewing ground color drab; costa marked with a series of short fuscous dashes and spots; at apical third two of the costal spots are longer than the remainder and are conspicuous; at basal third, in cell, a transverse fuscous dash; at end of cell an oblique transverse fuscous dash followed by a patch of brownish scales and between this and costa a grayish-fuscous suffusion forming an indistinct blotch; surface of wing with scattered fuscous irro- rations; cilia composed of grayish white-tipped scales. Hind wing pale grayish fuscous; cilia sordid white with a darker subbasal line. Foreleg sordid white heavily overlaid fuscous; midleg similar; hind leg sordid white; tibia with slight grayish suffusion. Abdomen pale cinereous, dorsally overlaid with fuscous except posterior edges of segments; ventrally irrorate with fuscous. Male genitalia slide USNM 24265. Harpe broad- est at middle and heavily clothed with fine setae; cucullus rounded. Gnathos a broad, finely spined knob. Uncus divided. Vinculum rather narrowed distally and rounded. Tegumen about half as long 34 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 22.?Doina trachycantha, new species: a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus. as harpe; rectangular. Anellus a round plate with fleshy lobe basolaterally. Aedeagus rather stout, curved, pointed distally; vesica armed with one strong cornutus. HOLOTYPE.?USNM 73710. TYPE-LOCALITY.?Santiago, Rio Colorado. Described from the unique male holotype (10 Apr 1953, L. E. Pena). Although atypical by reason of its divided uncus, D. glebula nevertheless keys to this group. A differ- ent generic designation is indicated here but in the absence of more material I leave D. glebula in this genus. Superficially, D. glebula resembles closely darker examples of the North American Agonopterix ar- gillacea (Walsingham), but is immediately distin- guishable structurally. Doina increta (Butler), new combination FIGURE 24; PLATE 66 Orthotelia increta Butler, 1883:75.?Bartlett-Calvert, 1886: 346. Cryptolechia increta Butler.?Meyrick, 1922b: 197. Male genitalia slide BM 12693. Harpe ample, of nearly equal width throughout; cucullus broadly rounded; sacculus lightly sclerotized. Gnathos a spined curved plate. Uncus hood shaped, as long as tegumen. Vinculum a U-shaped band. Tegumen about as long as width of harpe. Anellus an oval sclerotized plate with elongate basolateral lobes. Aedeagus stout, strongly curved; vesica armed with a cluster of three or four cornuti. HOLOTYPE.?British Museum (Natural History). TYPE-LOCALITY.?"Corral, Valdivia." Butler described D. increta in the Gelechiidae but Meyrick (1922b: 197) placed it correctly in the Oceo- phoridae. So far, the species is known only from Butler's type. Doina edmondsii (Butler), new combination Depressaria edmondsii Butler, 1883:76. Depressaria Edmondsii Butler.?Bartlett-Calvert, 1886:346.? Meyrick, 1922b: 177.?Gaede, 1939:322. TYPE.?British Museum (Natural History). TYPE-LOCALITY.?"Mountains of the hacienda of Cauquenes." NUMBER 273 35 FIGURE 23.?Doina glebula, new species: a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus. According to my notes D. edmondsii has genitalia with a spined gnathos, and harpe and other features like other species of Doina. This species is certainly not a Depressaria. Doina scariphista (Meyrick), new combination FIGURE 25; PLATE 6a Cryptolechia scariphista Meyrick, 1931:396.?Gaede, 1939: 393.?Clarke, 1963:170, pi. 82: fig. 1-lr. Female genitalia slide JFGC 4732 [in British Mu- seum (Natural History)]. Ostium transverse, oval. Antrum narrowly sclerotized. Inception of ductus seminalis dorsal, slightly before ostium. Ductus bur- sae rather long, slender, membranous. Bursa copu- latrix membranous, finely spiculate. Signum a small, elongate, sclerotized, branched plate. HOLOTYPE.?British Museum (Natural History). TYPE-LOCALITY.?Castro, Chiloe Island. FIGURE 24.?Doina increta (Butler): a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus. This species is known only from the unique type in the British Museum. The female genitalia are somewhat atypical but on other characters D. scarphista clearly belongs with other members of this genus. In the figure in Clarke (1963) the signum is obscured by long cor- nuti, of which there are at least five. Doshia, new genus TYPE-SPECIES.?Doshia miltopeza, new species (by monotypy and present designation). The gender of this generic name is feminine. Labial palpus compressed, recurved, second seg- ment reaching base of antenna, roughened slightly anteriorly; third segment shorter than second. Tongue well developed; maxillary palpus small, free. Head rough, side tufts spreading; ocellus ab- 36 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY sent. Antenna slightly thickened, simple; scape with- out pecten. Thorax smooth. Forewing smooth, costa sinuate, termen convex, oblique, 12 veins; lb fur- cate; lc strongly preserved; 2, 3, and 4 approximate, about equidistant; 5 nearer to 6 than to 4; 7 and 8 stalked, 7 to apex; 10 nearer to 9 than to 11; upper internal vein obsolete. Hind wing with 8 veins; 2 remote from 3; 3 and 4 very short stalked or con- nate; 5 nearer to 6 than to 4; 7 and 8 subparallel. Hind tibia smooth. Abdominal terga not setose. Male genitalia with well-developed gnathos and uncus. Female genitalia unknown. Doshia miltopeza, new species FIGURE 26; PLATE 3g Alar expanse 20 mm. Labial palpus ochraceous tawny; first segment white ventrally, reddish tawny dorsally; second seg- ment cinnamon buff on inner side. Antenna ochraceous tawny; scape buff ventrally. Head cin- namon buff. Thorax cinnamon buff, tegula slightly darker. Forewing ground color cinnamon buff; ex- treme costal edge white, bordered inwardly by a russet-vinaceous line; at costal two-thirds the slender white edge of costa broadens to form a white spot, and beyond this two small russet- vinaceous spots invade the white of costa; basal third of wing marked with numerous, very small, fuscous dots and outer third of wing similarly marked, the spots following the veins; apical half of cilia white, the cilia preceded by a narrow fuscous shade at base; remainder of cilia slightly paler than ground color; underside of costa broadly white, marked with russet-vinaceous spots. Hind wing grayish fus- cous; cilia sordid white; underside of hind wing whitish, spotted with fuscous. Foreleg white; femur shaded with russet on outer side; tibia shaded with carmine on outer side; midleg similar; hind leg white. Abdomen brownish buff dorsally, creamy white ventrally. Male genitalia slide USNM 24266. Harpe broad- est at base, tapering to a bluntly pointed cucullus; at base, inside costa a group of short, stout setae and beyond this group a less dense cluster of setae. FIGURE 25.?Doina scariphista (Meyrick): ventral view of female genitalia. NUMBER 273 37 FIGURE 26.?Doshia miltopeza, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus. Gnathos a spined disc. Uncus very broad, hood shaped, with posteromedian excision. Vinculum rounded. Tegumen broad as long. Anellus broadly semicydindrical with a short papilla on each side basally. Aedeagus nearly straight, stout; vesica armed with two small, sclerotized plates with ser- rate edges. HOLOTYPE.?USNM 73711. TYPE-LOCALITY.?San Ignacio. Described from the unique male holotype (Nov 1892, V. Izquerdo). This species is similar in coloring to a few species of Gonionota, but particularly to the darker speci- mens of G. insulana Clarke. The white costal spot of D. miltopeza is nearer the midde of costa than that of G. insulana, and the veins of the hind wing of D. miltopeza show ill-defined spots above, con- spicuous beneath, absent in the hind wing of G. insulana. Afdera, new genus TYPE-SPECIES.?Cryptolechia orphanaea Meyrick, 1931:397 (by monotypy and present designation). The gender of this generic name is feminine. Labial palpus recurved, second segment reaching, or slightly exceeding, base of antenna, very slightly roughened anteriorly; third segment shorter than second, slender, acute. Tongue well developed; maxillary palpus filiform, free. Head rather smooth With closely appressed scales; side tufts spreading; ocellus absent. Antenna of male finely and short ciliated; antenna of female finely serrate; scape without pecten. Thorax smooth. Forewing smooth, costa gently arched, termen slightly convex, oblique, 12 veins; lb furcate; lc weakly preserved at margin; 2 remote from 3; 3, 4, and 5 equidistant and ap- proximate; 7 and 8 stalked, both to costa; 9 much nearer the stalk of 7 and 8 than to 10; 10 nearer 9 than to 11; 11 from before middle of cell. Hind wing with 8 veins; 2 remote from 3; 3 and 4 con- nate; 5 nearer 6 than to 4; 6 and 7 subparallel. Hind leg roughened with hairlike scales. Abdomi- nal terga not setose. Male genitalia with uncus and gnathos well de- veloped. Female genitalia with signum present, but mi- nute. Admittedly close to Cryptolechia Zeller, Afdera differs primarily by a well-developed, recurved ex- tension of the sacculus and an unarmed aedeagus. Meyrick's diagnosis of Cryptolechia is vague and contains a large number of unrelated forms belong- ing to several well-known and distinct genera. Afdera differs from Corita primarily by the condi- 38 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY tion of veins 3 and 4 of hind wing, which are con- nate in Afdera and stalked in Corita. Ajdera orphnaea (Meyrick), new combination FIGURE 27 Cryptolechia orphnaea Meyrick, 1931:397.?Clarke, 1963:166, pi. 80: figs. 1-la. HOLOTYPE.?British Museum (Natural History). TYPE-LOCALITY.?Llanquihue, Casa Pangue. The species is apparently common. Arauco, Cara- mavida (January) Arauco, Cordillera de Nahuel- buta, Pichinahuel (January) collected by L. E. Pena and "Centro-Austral" (Jan-Mar) V. Izquerdo. The figure of the male genitalia in Clarke, cited above, was made from a damaged specimen; the genitalia of the male are refigured here (slide USNM 24267). Melaneulia Butler Melaneulia Butler, 1883: 70. [Type-species: Melaneulia hecate Butler, 1883; by monotypy.] Melaneulia hecate Butler FIGURE 28 Melaneulia hecate Butler, 1883: 70.?Bartlett-Calvert, 1886: 344. Cryptolechia hecate (Butler).?Meyrick, 1922b: 197. Male genitalia slides USNM 24269, 24270. Harpe ample, broadest at base; cucullus narrowly rounded; sacculus narrowly sclerotized and continued termi- nally across harpe as a broad, flattened clasper. Gnathos a broad spined knob with small median point. Uncus broad, hoodlike. Vinculum rounded. Tegumen quadrate, about half as long as harpe. Anellus a pear-shaped plate with a spiny ridge on each side from base. Aedeagus slightly curved, bulbous proximally, pointed distally; vesica un- armed. HOLOTYPE.?British Museum (Natural History). TYPE-LOCALITY.?Valdivia. The genus remains monobasic. In addition to the type in the British Museum (Natural History) there FIGURE 27.?Afdera orphnaea (Meyrick): a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus. NUMBER 273 39 FIGURE 28.?Melaneulia hecate Butler: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus. are two males in the National Museum of Natural History. Butler described M. hecate as a tortricid but it is indeed oecophorid. Depressariodes Turati Martyrhilda Clarke, 1941:125, pi. 2: fig. 17, pi. 6: fig. 44, pi. 10: figs. 67, 67a, pi. 16: fig. 100. [Type-species: Depressaria canella Busck, 1904: 764; by original designation.] Depressariodes Turati, 1924:175. [Type-species: Depressariodes marmaricellus Turati, 1924, by monotypy; D. marmaricellus Turati is a synonym of Depressaria fuscicostella Christoph, 1887.] Depressariodes Turati, Hannemann, 1976:209 [synonymy]. This is a widespread genus occurring in the palaearctic region, in the nearctic region from Alaska to Mexico, and down the length of the Andes with one species in the Juan Fernandez Islands. There are two described species of this genus from Chile: Depressariodes lusciosa (Meyrick), new combination, and Depressariodes relegata (Mey- rick), new combination. Depressariodes lusciosa (Meyrick), new combination Depressaria lusciosa Meyrick, 1915:211; 1922b:177. Martyrhilda lusciosa (Meyrick).?Clarke, 1963:326, pi. 160: figs. 2-2b. This species was described from Peru but ranges into Chile. Depressariodes relegata (Meyrick), new combination Depressaria relegata Meyrick, 1922a:268. Martyrhilda relegata (Meyrick).?Clarke, 1965:78, figs. 78, 79. The range of M. relegata is apparently confined to Masatierra in the Juan Fernandez Islands. 40 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY Gonionota Zeller Hypercallia (Gonionota) Zeller, 1877:381. [Type-species: Hy- percallia (Gonionota) notodontella Zeller, 1877; subsequent designation by Meyrick, 1922b, p. 161.] Gonionota mimulina (Butler), new combination FIGURE 29 Agriocoma mimulina Butler, 1883:83, pi. 11: fig. 9. Hypercallia mimulina (Butler.)?Bartlett-Calvert, 1886:347.? Meyrick, 1922b: 163 (No. 57).?Gaede, 1939:261. Agriocoma mimulina araucana Bartlett-Calvert, 1893:831. HOLOTYPE.?British Museum (Natural History) (A. mimulina); (A. m. araucana). TYPE-LOCALITIES.?Valparaiso (A. mimulina)', Araucania (A. m. araucana). In the National Museum of Natural History there are specimens from Malleco and Maule provinces. So far this is the only species known in the Chilean fauna clearly referable to this genus. Male genitalia figured from slide USNM 24271. =*? FIGURE 29.?Gonionota mimulina (Butler): a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus. Nedenia, new genus TYPE-SPECIES.?Nedenia rhodochra, new species (by monotypy and present designation). The gender of this generic name is feminine. Labial palpus slender, recurved, exceeding ver- tex; second segment smooth, third segment acute, shorter than second. Tongue well developed; maxil- lary palpus filiform, well developed, free. Head smooth centrally, with loose scales laterally and posteriorly; ocellus absent. Antenna finely pubes- cent and slightly thickened in male, simple in female; scape without pecten. Thorax with small, posteromedian tuft. Forewing smooth, costa arched, termen straight; 12 veins; lc very strongly pre- served; 2 remote from 3; 3, 4, and 5 about equi- distant; 7 and 8 stalked, 7 to termen; 6, stalk of 7 and 8, and vein 9 about equidistant; 10 slightly nearer 9 than 11; 11 from slightly before middle of cell. Hind wing with 8 veins; 2 remote from 3; 3 and 4 connate; 5 well removed from 4. Hind leg tibia slightly roughened above and beneath with short, hairlike scales. Abdominal terga not setose. Male genitalia with uncus and gnathos present; socius absent. Female genitalia with two signa. In Meyrick's key, this genus runs to the Aus- tralian genus Machetis Meyrick but differs from it by the absense of an ocellus, lacks the second segment of labial palpus thickened with dense ap- pressed scales, and posterior tibia not clothed with very long hairs. Moreover, Nedenia is a broader winged genus with much shorter cilia on hind wing. Nedenia rhodochra, new species FIGURE 30; PLATE 3/I Alar expanse 18-22 mm. Labial palpus buff; second segment argillaceous anteriorly and coral red posteriorly on outer side; on inner side a slight tinge of coral red anteriorly; third segment apical third brown. Antenna light ochraceous buff basally, shading to brown distally; scape narrowly brown apically. Head buff. Thorax buff; thoracic tuft mixed coral red and yellow scales. Forewing ground color buff; extreme edge of costa narrowly coral red and yellow; at extreme base of costa a small, outwardly oblique blackish dash; at basal third scattered blackish scales form an ill-defined blackish dash; at apical two-thirds a conspicuous triangular vinaceous-brown spot with NUMBER 273 41 FIGURE SO.?Nedenia rhodochra, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus; d, ventral view of female genitalia. 42 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY its base on costa; in cell and on fold some scattered blackish scales; on dorsum a vinaceous brown suf- fusion; in female extreme dorsal edge blackish fuscous; cilia vinaceous brown except yellowish around tornus. Hind wing whitish to pale buff; cilia concolorous except around apex and termen yellowish. Foreleg buff; femur and tibia coral red on outer side; basal segments suffused fuscous; midleg pale buff; tibia with blackish dash proxi- mally; tarsal segments with coral red scattered scales; hind leg pale buff; tarsal segments slightly darker. Abdomen buff; terga not setose. Male genitalia slide USNM 24237, 24275, 24188. Harpe simple, cucullus rounded; basally a cluster of very strong, short setae. Gnathos a broad spinu- late lobe. Uncus broad, hood shaped, with strong setae laterally on ventral surface. Vinculum U- shaped, narrow. Tegmen short, rectangular, about two-thirds length of uncus. Anellus rectangular, deeply incised laterally. Aedeagus short, curved, bulbous basally; cornuti at most very small, spinu- late. Female genitalia slide USNM 24274, 24189. Os- tium very small, elongate. Antrum not differenti- ated. Inception of ductus seminalis slightly anterior to ostium. Ductus bursae long, membranous, spi- raled. Bursa copulatrix membranous; signa two elongate, scobinate patches. HOLOTYPE.?USNM 73712. TYPE-LOCALITY.?Llanquihue, Petrohue. Described from the male holotype (12 Mar 1959, J. F. G. Clarke) 11 male and 3 female paratypes as follows: 4 males, Llanquihue, Petrohue (11-12 Mar 1959); male Peulla (8 Mar 1959) J. F. G. Clarke; 3 males, 3 females Malleco, Rio Blanco, 1050-1300 m, (21/24 Nov 1954) L. E. Pefia; 4 males, Centro-Austral (Jan-Mar 1898) V. Izquerdo. This species reminds one somewhat of the Euro- pean Carcina quercana (Fabricius) but is imme- diately distinguished by the shorter antennae. Revonda, new genus TYPE-SPECIES.?Revonda eschara, new species (by monotypy and present designation). The gender of this generic name is feminine. Labial palpus slender, recurved, slightly exceed- ing vertex; second and third segments of about equal length; second segment slightly roughened anteriorly. Maxillary palpus minute. Head smooth, side tufts spreading; ocellus absent. Antenna cili- ated in male; scape with well-developed pecten. Thorax smooth. Forewing smooth, costa nearly straight, stigma present, 11 veins; lc strongly pre- served; 2 remote from 3; 3 near 4; 4 and 5 connate; 7 and 8 coincident to costa; 9 nearer to 7 4- 8 than to 10; 11 from two-fifths. Hind wing with 8 veins; 2 remote from 3; 3 and 4 stalked; 5 connate with stalk of 3 and 4. Hind tibia smooth. Abdominal terga setose. Male genitalia with uncus and gnathos present; socius indicated by a few weak setae. Female unknown. Revonda is near Despina but the antennal pecten is present in RevOnda, absent in Despina and veins 4 and 5 of forewing are connate in Revonda, separate in Despina. Revonda eschara, new species FIGURE 31; PLATE 4a Alar expanse 17-21 mm. Labial palpus creamy white; second segment fuscous on outer side and anteriorly in basal half; apex of third segment fuscous. Antenna grayish fuscous; scape with pecten. Head ocherous white. Thorax ocherous white; base of tegula blackish fuscous. Forewing ground color ocherous white; on costa three blackish fuscous spots, one at base, one at middle, and one slightly before apex; dorsal two-thirds of wing lightly shaded grayish; from end of cell a longitudinal shade, somewhat darker than the grayish dorsal shade; on fold, at two-fifths, a loose aggregarion of fuscous scales forming an ill- defined longitudinal dash; at end of cell, a fuscous spot; scattered over surface irregularly placed fus- cous scales; cilia pale grayish buff. Hind wing gray; cilia grayish buff. Foreleg ocherous white; tibia and tarsal segments strongly overlaid blackish fuscous; midleg similar; hind leg ocherous white shaded with gray. Abdomen ocherous white ventrally, gray- ish dorsally and laterally; terga setose. Male genitalia slides USNM 24194, 24276. Harpe simple, broadest about middle, tapering to a nar- rowly rounded cucullus. Gnathos long, slender, hooked at distal end. Uncus triangular, pointed distally. Vinculum U-shaped with slender median projection. Tegumen as broad as long. Anellus a small sclerotized plate with two digitate basal processes, Aedeagus moderately stout, slightly NUMBER 273 43 FIGURE 31.?Revonda eschara, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus. curved, sharply pointed; vesica armed with a single strong cornutus. HOLOTYPE.?USNM 73713. TYPE-LOCALITY.?Arauco, Cordillera de Nahuel- buta, Pichinahuel, 11-1400 m. Described from the male holotype (23-31 Jan 1954, L. E. Pefia) and one male paratype with identical data. Strikingly similar to Oecophora minnetta Butler (Figure 54) but the harpe of R. eschara is simple, that of O. minnetta with a strong clasper. The aedeagus of O. mifinetta is slender and armed with a longitudinal series of teeth; the aedeagus of R. eschara is short and unarmed and the vesica has a single cornutus. The anellus of R. eschara lacks the series of strong setae found in O. minnetta. The females of both species are unknown. Despina, new genus TYPE-SPECIES.?Borkhausenia rhodosema Meyrick, 1931:390 (by monotypy and present designation). The gender of this generic name is feminine. Labial palpus slender, recurved; second segment smooth; third segment slightly shorter than second, acute. Maxillary palpus minute. Head slightly 44 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY roughened with closely appressed scales; ocellus absent. Antenna of male very short ciliated; an- tennal pecten absent. Thorax smooth. Forewing smooth, pointed, costa nearly straight, 11 veins; lc strongly preserved; 2 remote from 3; 3, 4, and 5 nearly equidistant; 7 and 8 coincident to costa; 9 nearer to 7 + 8 than to 10; 11 from middle of cell. Hind wing with 8 veins; 2 remote from 3; 3 and 4 very short stalked; 5 nearer to 6 than 4. Hind tibia smooth. Abdominal terga setose. Male genitalia with simple harpe. Gnathos and uncus well developed. Socius absent. Female unknown. Despina is nearest Revonda but differs from it by the absence of antennal pecten, absence of the stigma of forewing, veins 4 and 5 of forewing well separated whereas in Revonda they are connate, and vein 5 of hind wing is well separated from 4, not connate. The male genitalia have been adequately figured in Clarke (1963, pi. 62: figs. 1-lb). Despina rhodosema (Meyrick), new combination Borkhausenia rhodosema Meyrick, 1931:390.?Clarke, 1965, 4:130, pi. 62: figs. l i b . HOLOTYPE.?British Museum (Natural History). TYPE-LOCALITY.?Llanquihue Province, Ense- nada. This is the only species referable to this genus, and it has been figured previously by Clarke (1963). Irenia, new genus TYPE-SPECIES.?Irenia leucoxantha, new species (by present designation). The gender of this generic name is feminine. Labial palpus slender, recurved, exceeding ver- tex; third segment shorter than second. Maxillary palpus well developed, free. Head smooth; side tufts spreading; ocellus absent. Antenna ciliated in male, simple in female; scape with well-developed pecten. Thorax smooth. Forewing smooth, costa arched, 12 veins; lb furcate; 2 distant from 3; 3 farther from 4 than 4 is from 5; 7 and 8 stalked, 7 to apex; 9 nearer to stalk of 7 and 8 than to 10; 11 from before middle of cell. Hind wing with 8 veins; 2 remote from 3; 3 and 4 connate. Hind tibia clothed with long, hairlike scales. Abdominal terga setose. Male genitalia with uncus and gnathos present; socius indicated by a few weak setae. Cornuti ab- sent. Female genitalia with signum present. Irenia is near Revdfrda but veins 7 and 8 of fore- wing are stalked, not coincident as in Revonda. Irenia curvula, new species FIGURE 32; PLATE AC Alar expanse 21 mm. Labial palpus fuscous. Antenna pale ochraceous buff. Head light ochraceous tawny. Thorax ochra- ceous tawny. Forewing ground color ochraceous buff; costa, especially toward apex, sayal brown; cilia light clay color. Hind wing straw color with slight brownish suffusion toward apex; cilia concolorous with ground color. Foreleg straw color heavily over- laid fuscous; midleg similar; hind leg straw color; tibia with slight fuscous suffusion ventrally toward apex; tarsal segments heavily infuscated. Abdomen sparsely scaled dorsally, ochraceous buff; ventrally light brown with coppery luster. Male genitalia slide USNM 24277. Harpe very broad basally, slightly longer than tegumen; cucul- lus divided into a costal lobe and ventral lobe, the latter terminating as a short, pointed, recurved hook. Gnathos spatulate, broader distally than proximally. Uncus broad basally, tapering to mid- FIGURE 32.?Irenia curvula, new species: a, ventral view of male genitalia with left harpe and aedeagus removed; b, aedeagus. NUMBER 273 45 die, terminating as a long pointed median process. Vinculum U-shaped. Tegumen broad basally, nar- rower at base of uncus. Anellus a broad, rectangular plate, fused with base of harpe, posterior edge deeply concave and with a fleshy, digitate, setaceous lobe on each side. Aedeagus slender, slightly curved, sharply pointed distally; vesica unarmed. HOLOTYPE.?USNM 73715. TYPE-LOCALITY.?Arauco, Caramavida. Described from the unique male holotype (1/6 Jan 1954, L. E. Pefia). In general appearance similar to Aliciana gemi- nata, new species, but with a distinctly brown costa, a character not found in A. geminata. Irenia leucoxantha, new species FIGURE 33; PLATE 4ft Alar expanse 18-20 mm. Labial palpus brown; second segment with apex and scattered scales white; base of third segment white. Antenna brown; apex of scape white. Head clay color; face white. Thorax clay color anteriorly, shading to ochraecous buff posteriorly. Forewing ground color orange yellow; extreme edge of costa white; surface of wing with a few widely scattered black scales; cilia orange yellow except suffused brownish in tornal area. Hind wing pearl white, cilia concolorous. Foreleg brown; midleg ochraceous buff; hind leg ochraceous buff; spurs brown. Male genitalia slides USNM 24139, 24278. Harpe broad basally, and thick; cucullus narrow, rounded; sacculus strongly sclerotized, produced, terminating in a very long, slender, recurved sclerotized process. Gnathos spoon shaped. Transtilla fleshy, with a weak lobe on each side. Uncus broad basally, tri- angular, strongly sclerotized distally. Vinculum U-shaped with median process. Tegumen broad basally, narrowed distally. Anellus narrow, V-shaped to U-shaped basally, fused with base of harpe and transtilla. Aedeagus slender, produced apically as a long, strongly sclerotized point. Female genitalia USNM 24142. Ostium slitlike, transverse, ventral lip slightly convex. Antrum not differentiated. Inception of ductus seminalis from near opening of ostium. Ductus bursae membranous. Bursa copulatrix half membranous, half weakly granular. Signum a minute dentate plate. HOLOTYPE.?USNM 73714. TYPE-LOCALITY.?M aule, Consti tucion. Described from the female holotype (26 Nov 1953, L. E. Pena), and two male paratypes as follows: male, Arauco, Pillem-Pilli (15 Jan 1954, L. E. Pefia); male, Arauco, Pichinahuel, Cordillera de Nahuel- buta, 11-1400 m. (23-31 Jan 1954, L. E. Pena). Although similar to /. curvula, new species, /. leucoxantha is a smaller and much paler species. The long, curved process from the sacculus of /. leucoxantha immediately distinguishes it from /. curvula. Lucyna, new genus TYPE-SPECIES.?Cryptolechia fenestella Zeller, 1875:439 (by monotypy and present designation). The gender of the generic name is feminine. Labial palpus slightly ascending but not strongly recurved; second segment nearly twice as long as third, smooth; third segment slender with small tuft posteriorly at middle. Tongue well developed; maxillary palpus reduced, filiform, free. Head smooth, side tufts spreading; ocellus absent. An- tenna simple (at least in female); scape without pecten. Thorax smooth. Forewing with slightly raised tufts of scales at end of cell and on fold, opposite; costa gently arched, termen strongly oblique, 12 veins; lb furcate, lc strongly preserved; 2 remote from 3; 3, 4, and 5 approximate and equi- distant; 7 and 8 stalked, 7 to apex; 6 and 9 approxi- mate to the stalk of 7 and 8; 10 nearer to 9 than to 11; 11 from middle of cell; upper internal vein absent. Hind wing with 8 veins; 2 remote from 3; 3 and 4 stalked; 5, 6, and 7 equidistant, subparallel. Hind tibia clothed with long scales dorsally. Ab- dominal terga setose. Male genitalia not available. Female genitalia with signum. Lucyna shows affinity to Gonionota Zeller, but differs from it by the setose abdominal terga, and vein 7 to apex, not to termen as in Gonionota. Moreover, the termen in Lucyna is much more oblique than that of Gonionota and forewing is narrower. Lucyna fenestella (Zeller), new combination FIGURE 34 Cryptolechia fenestella Zeller, 1874:439, pi. 12: fig. 9. Machimia fenestella {Zeller).?Zeller, 1877:259. Hypercallia fenestella (Zeller).?Bartett-Calvert, 1886:346. 46 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 53.?Irenia leucoxantha, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus; d, ventral view of female genitalia. NUMBER 273 47 FIGURE 34.?Lucyna fenestella (Zeller): a, venation of right wings; b, ventral view of female genitalia. 48 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY Meyrick, 1922b: 163 [as synonym of thyridopa Meyrick]. Coptotelia thyridopa Meyrick, 1912:701. Hypercallia thyridopa (Meyrick).?Meyrick, 1922b: 163. HOLOTYPE.?British Museum (Natural History). TYPE-LOCALITY.?Valparaiso. Lucyna fenestella resembles species of the genus Coptotelia but is a very much narrower-winged spe- cies with a very oblique termen. Dita, new genus TYPE-SPECIES.?Dita phococara, new species [by monotypy and present designation]. The gender of this generic name is feminine. Labial palpus upturned; second segment slightly roughened ventrally toward apex; third segment shorter than second, acute. Tongue well developed; maxillary palpus filiform, free. Head roughened with loosely appressed scales; ocellus absent. An- tenna ciliated (in male); scape with pecten. Thorax smooth. Forewing smooth, costa slightly arched, apex falcate, termen nearly straight below apex, oblique, 12 veins; 1b furcate; lc well preserved; 2 rather remote from 3; 3, 4, and 5 approximate, nearly equidistant; 7 and 8 stalked, 7 to termen; 9 nearer to stalk of 7 and 8 than to 10; 10 nearer to 9 than to 11; 11 from middle. Hind wing with 8 veins; 2 remote from 3; veins 3 and 4 short stalked; 4 to 7 equidistant. Hind tibia roughened by hairlike scales. Abdominal terga setose. Male genitalia with uncus and gnathos well developed. Female genitalia unknown. This genus is apparently very closely related to the Australian and African Tanyzancla Meyrick, from which it differs mainly by the upturned labial palpus. Dita phococara, new species FIGURE 35; PLATE Ad Alar expanse 20 mm. Labial palpus sordid white irrorate and overlaid grayish fuscous on outer side. Antenna grayish fuscous; scape sordid white ventrally. Head gray, mixed with sordid white. Thorax cinnamon brown. Forewing ground color buckthorn brown; costa, from base to apical fifth, cinnamon brown; from apical fifth of costa a transverse cinnamon brown shade extends to dorsum, where it is broadest; in center of cell a small cinnamon brown spot and at end of cell a sordid white dot; cilia mixed buck- thorn brown and cinnamon brown. Hind wing gray- ish fuscous; cilia considerably lighter. Foreleg sordid white; femur cinnamon brown; tibia and tarsal seg- ments grayish fuscous, the latter faintly annulated sordid white; midleg similar; hind leg sordid white, suffused and blotched grayish. Abdomen fuscous dorsally, somewhat paler ventrally. Male genitalia USNM 24201. Harpe rectangular, cucullus forming a point on costal edge and fleshy portion fused to an extension of the sacculus; ex- tension of sacculus heavily sclerotized, terminating in a clawlike appendage. Gnathos very large, as broad as long. Uncus stout, triangular, pointed dis- tally. Vinculum U-shaped. Tegumen as broad as long. Anellus a small sclerotized plate with a long, curved, pointed dorsoposterior process. Aedeagus moderately slender, pointed; vesica armed with a series of graduated cornuti. HOLOTYPE.?USNM 73716. TYPE-LOCALITY.?Llanquihue, Rio Maullin. Described from the unique male holotype (6 Jan 1966, O. Flint and Cekalovic). This species is similar to C. fasciatipedella but is darker, has a conspicuous white spot at end of cell, and a postmedian dark shade absent in C. fas- ciatipedella. The figures illustrate adequately the genitalic differences. Dita fasciatipedella (Zeller), new combination FIGURE 36; PLATE 6C Cryptolechia fasciatipedella Zeller, 1874:437, pi. 12: fig. 7.? Butler, 1883:80.?Bartlett-Calvert, 1886:347. Machimia fasciatipedella (Zeller).?Zeller, 1877:259. Hypercallia fasciatipedella (Zeller).?Meyrick, 1922b: 163.? Gaede, 1939:259. Male genitalia slide BM 12510. Harpe almost as broad as long, divided; costa divided distally, the costal arm much broader than inner arm and clothed with setae; sacculus very broadly expanded distally, terminating in an inner hook and short outer digitate process. Gnathos bluntly pointed distally, with a patch of setae posteriorly. Uncus broad basally, pointed distally. Vinculum U-shaped. NUMBER 273 49 FIGURE 35.?Dita phococara, new species: a, venation of right wings; b, ventral view of male genitalia with left harpe and aedeagus removed; c, aedeagus. Tegumen twice as broad as long. Anellus a narrow, folded plate with a long, curved median process. Aedeagus slender, curved, distal third much at- tenuated; vesica unarmed. Female genitalia unknown. TYPE.?British Museum (Natural History). TYPE-LOCALITY.?Valparaiso. The moth clearly keys to Dita and the genitalia correspond with those of D. phococara, new species. The species does not belong in Cryptolechia, Ma- chimia or Hypercallia in all of which genera it has previously been placed. Utilia, new genus TYPE-SPECIES.?Utilia florinda, new species (by present destination). The gender of this generic name is feminine. Labial palpus slender, recurved, exceeding ver- tex; second segment slightly roughened anteriorly; third segment slightly shorter than second, acute, slightly roughened anteriorly. Maxillary palpus well developed, free. Head slightly roughened, side tufts spreading; ocellus absent. Antenna ciliated, finely and sparsely so in female; pecten present. Fore- wing smooth, costa arched, apex falcate, 12 veins; 50 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY FIGURE 36.?Dita fasciatipedella (Zeller): a, ventral view of male genitalia with left harpe and and aedeagus removed; b, aedeagus. lb furcate; lc well preserved; 2 remote from 3; 3, 4, and 5 well separated (or connate); 7 and 8 stalked, 7 to termen; 9 much nearer to stalk of 7 and 8 than to 10; 10 much nearer to 9 than to 11; 11 from well before middle of cell; upper internal vein present. Hind wing with 8 veins; 2 remote from 3; 3 and 4 connate; 5 nearer to 6 than to 4; 5, 6, and 7 equidistant, subparallel. Hind tibia slightly roughened with short, stiff scales. Ab- dominal terga setose, the setae fine and easily de- ciduous. Male genitalia with well-developed uncus and gnathos. Aedeagus coiled proximally; vesica armed. Female genitalia with signum. Utilia is similar to Alynda but is broader winged and has the upper internal vein of forewing well developed. I place the following four species here. Utilia luridella (Zeller), new combination FIGURE 37 Cryptoechia luridella Zeller, 1874:438, pi. 12: fig. 8?Butler, 1883:81. Cryptolechia (Machimia) luridella