ZOOTAXA ISSN 1175-5326 (print edition) ISSN 1175-5334 (online edition)Copyright © 2017 Magnolia Press Zootaxa 4348 (1): 001–125 http://www.mapress.com/j/zt/ Monograph https://doi.org/10.11646/zootaxa.4348.1.1 http://zoobank.org/urn:lsid:zoobank.org:pub:63C5E0BC-6F55-440D-83EA-4BA9B9D31EAC ZOOTAXA Deep-water bony fishes collected by the B/O Miguel Oliver on the shelf edge of Pacific Central America: an annotated, illustrated and DNA-barcoded checklist D. ROSS ROBERTSON1,7, ARTURO ANGULO2,3,4, CAROLE C. BALDWIN5, DIANE PITASSY5, AMY DRISKELL5, LEE WEIGT5 & IGNACIO J.F. NAVARRO6 1Smithsonian Tropical Research Institute, Balboa, Republic of Panama 2 Museo de Zoología, Universidad de Costa Rica. 11501–2060, San Pedro de Montes de Oca, San José, Costa Rica. 3Centro de Investigación en Ciencias del Mar y Limnologia (CIMAR), Universidad de Costa Rica. 11501–2060, San Pedro de Montes de Oca, San José, Costa Rica. 4UNESP, Universidade Estadual Paulista “Júlio de Mesquita Filho”, Laboratório de Ictiologia, Departamento de Zoologia e Botâni- ca. Rua Cristóvão Colombo, 2265, CEP 15054–000, São José do Rio Preto, SP, Brazil. 5National Museum of Natural History, Smithsonian Institution, Washington, DC 20560. 6Instituto Español de Oceanografía, Centro Oceanográfico de Murcia, Varadero 1. 30740, San Pedro del Pinatar, Murcia, Spain. 7Corresponding author. E-mail: drr@stri.org Magnolia Press Auckland, New Zealand 4348Accepted by E. Hilton: 14 Aug. 2017; published: 15 Nov. 2017 Licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0 D. ROSS ROBERTSON, ARTURO ANGULO, CAROLE C. BALDWIN, DIANE PITASSY, AMY DRISKELL, LEE WEIGT & IGNACIO J.F. NAVARRO Deep-water bony fishes collected by the B/O Miguel Oliver on the shelf edge of Pacific Central America: an annotated, illustrated and DNA-barcoded checklist (Zootaxa 4348) 125 pp.; 30 cm. 15 Nov. 2017 ISBN 978-1-77670-262-6 (paperback) ISBN 978-1-77670-263-3 (Online edition) FIRST PUBLISHED IN 2017 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: magnolia@mapress.com http://www.mapress.com/j/zt © 2017 Magnolia Press ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition)ROBERTSON ET AL .2 · Zootaxa 4348 (1) © 2017 Magnolia Press Table of contents Abstract . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 Materials and methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 Results and discussion . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 Annotated and illustrated catalog . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 NOTACANTHIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 Halosauridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 Notacanthidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 29 ANGUILLIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 Muraenidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30 Congridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 Synaphobranchidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32 Ophichthidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33 Nemichthyidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35 Serrivomeridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 Nettastomatidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 OSMERIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 37 Argentinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 37 Bathylagidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 38 Alepocephalidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 38 Platytroctidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 41 STOMIIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 42 Sternoptychidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 42 Phosichthyidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 43 Stomiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 43 AULOPIFORMES. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 44 Chlorophthalmidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 44 Ipnopidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 45 Scopelarchidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 46 Synodontidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 47 Paralepididae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 48 MYCTOPHIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 49 Neoscopelidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 49 Myctophidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 49 GADIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 50 Bregmacerotidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 50 Macrouridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 51 Moridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 55 Merlucciidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 56 OPHIDIIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 56 Ophidiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 56 LOPHIIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 62 Lophiidae. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 62 Antennariidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 63 Ogcocephalidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 64 Caulophrynidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 68 Melanocetidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 69 Oneirodidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 69 STEPHANOBERYCIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 71 Melamphaidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 71 BERYCIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 72 Trachichthyidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 72 Anoplogastridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 72 SCORPAENIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 74 Sebastidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 74 Scorpaenidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 75 Triglidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 76 Peristediidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 78 Psychrolutidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79 Liparidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 80 PERCIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 81 Serranidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 81 Zootaxa 4348 (1) © 2017 Magnolia Press · 3TEP DEEP FISHES Malacanthidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84 Carangidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84 Haemulidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 86 Sciaenidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 87 Labridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88 Zoarcidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 89 Chiasmodontidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92 Uranoscopidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93 Callionymidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 94 Gobiidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 95 Trichiuridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 95 Nomeidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 96 Stromateidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 98 PLEURONECTIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 99 Paralichthyidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 99 Bothidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 101 Cynoglossidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 102 TETRAODONTIFORMES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 103 Balistidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 103 Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 104 References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 104 Abstract An annotated and photographically illustrated checklist with DNA barcodes of the species of bony fishes collected during a month-long research cruise of the Spanish Research vessel B/O Miguel Oliver is presented. The vessel made trawls on the continental shelf of the Pacific coast of Central America, in November-December 2010, at depths of 108–1625 m. This list, based on 707 specimens (of a total of 876 specimens collected during the whole expedition), includes 129 species belonging to 15 orders, 61 families, and 97 genera. New information is presented on the geographical distributions of more than a third of those species, with 29 species (22.4%) representing new records from Central American waters and 17 spe- cies (13.2%) having expanded latitudinal ranges. Data on capture depths demonstrate increased depth ranges due to new minimum and/or maximum known depths for 31 species, i.e. 24% of those captured. Tissue samples from frozen speci- mens were used to obtain DNA barcodes of 682 (96.5%) individuals belonging to 118 species (91.4% of those recorded here), which have been made publically available in Genbank. Those data include barcodes for 84 species (65.1% of the total collected, and 77.1% of those for which barcodes were obtained) and 30 genera (30.9% of those collected) for which no species barcodes have been previously published. Barcodes of 54 species represent the first genetic sequences of any type published for those species. The abundance of new data indicate that there is still much to learn about the composition and geographical and depth distributions of the fish fauna of the shelf edge and continental slope of this region. Key words: Deep sea, ichthyofauna, diversity, new records, tropical eastern Pacific Introduction Between 2007 and 2011 a series of research cruises were made by the Spanish research vessel B/O Miguel Oliver (Figure 1) along the continental shelves and slopes of the Pacific coast of Central and South America. Those cruises, aimed at assessing both the biodiversity of benthic organisms and fishery resources by deploying bottom trawls at depths down to ~1600 m, began with expeditions to Peru, Patagonia and the Pacific coast of Panama. There were further cruises to those three areas as well as to Ecuador in 2008 and 2009, and El Salvador in 2010. The last 2010 cruise sampled the outer edge of the continental shelf and the continental slope of Pacific Central America, between Panama and Guatemala, at depths of 108–1625 m. One of us (DRR) was invited to participate in this cruise and decided to acquire fish specimens and photographs thereof for scientific study. A companion cruise with similar objectives was made on the Caribbean shelf of Central America, between Panama and Belize in early 2011. ROBERTSON ET AL .4 · Zootaxa 4348 (1) © 2017 Magnolia Press The bony fishes collected during the 2010 expedition on the Pacific side of Central America were recently reviewed by us, providing the basis for the annotated and photographically illustrated checklist presented here. In addition to increasing knowledge of the deep-water fish fauna of the tropical eastern Pacific, we hope that this work will serve as an educational tool to deep-sea fish researchers and to the interested public-at-large, and will act as a guide to the pertinent (taxonomic and ecologic) literature and to the numerous name changes and additions of species since the most recent comprehensive works on the deep-water fish fauna of that region were published (see Fischer et al. 1995, De La Cruz-Agüero et al. 1997, Chirichigno & Velez 1998, Nakaya et al. 2009). Materials and methods Sampling effort and study site. Samples were collected on board of the Spanish research vessel, B/O Miguel Oliver (Fig 1) operated by the Spanish Ministerio de Agricultura y Pesca, Alimentación y Medio Ambiente (MAPAMA; http://www.mapama.gob.es). The B/O Miguel Oliver is a 70 m long, 2495 gross tons, stern trawler that began operation in 2007. It ships a crew of 45 personnel, and can accommodate up to 22 scientists. FIGURE 1. The Research Vessel B/O Miguel Oliver; photo MAPAMA. Sampling was done using a Lofoten bottom trawl equipped with a pair of 1,300 kg oval steel doors, each attached to the net by a 100 m long warp. The trawl itself was 32 m long, with a mouth opening 23 m wide and 3.5 m high when fully deployed on the bottom, and had a cod-end mesh of 35 mm. Rubber rollers along the bottom of the mouth of the net reduced its contact with the substratum. The ship’s side-scan sonar was used to locate a suitable area of relatively level, unobstructed bottom prior to each trawl. Bottom times for trawls were typically 30 mins, at speeds of ~3–3.5 knots. A total of 107 localities were sampled between southeastern Panama (7.4138º N, - 78.1242º W) and southwestern Guatemala (13.7593º N, -91.8780º W), between November 7 and December 10, 2010, at depths ranging from 108 to 1625 m (Fig. 2; Table 1). Zootaxa 4348 (1) © 2017 Magnolia Press · 5TEP DEEP FISHES FIGURE 2. Location of trawl stations made by the B/O Miguel Oliver during late 2010; black stars indicate trawls from which fish specimens discussed here were retained; white stars indicate trawls from which no specimens were kept. Base map: Google Earth. TABLE 1. Trawl stations (numbered from 1 to 107) of the research vessel B/O Miguel Oliver on the Pacific coast of Central America in November-December 2010. SN = Station number; DR = Depth range (m). Stations with data in bold yielded specimens discussed herein. SN Date Coordinates DR Initial Final 001 12-11-2010 7.4138º, -78.1242º 7.4322º, -78.1452º 165–183 002 12-11-2010 7.3435º, -78.1125º 7.3578º, -78.1360º 407–412 003 12-11-2010 7.6230º, -78.6715º 7.6232º, -78.6997º 116–118 004 12-11-2010 7.6268º, -78.6933º 7.6268º, -78.6653º 115–116 005 12-11-2010 7.5713º, -78.7945º 7.5742º, -78.8043º 430–500 006 13-11-2010 7.7080º, -78.9635º 7.6978º, -78.9373º 109 007 13-11-2010 7.6327º, -79.1225º 7.6337º, -79.1445º 470–478 008 13-11-2010 7.5478º, -79.2138º 7.5518º, -79.1867º 757–781 009 13-11-2010 7.5120º, -79.2107º 7.5118º, -79.2353º 987–1013 010 13-11-2010 7.5155º, -79.1968º 7.5152º, -79.1715º 1020–1024 011 14-11-2010 7.3935º, -79.4890º 7.3795º, -79.5103º 1292 012 14-11-2010 7.3950º, -79.5390º 7.3777º, -79.5590º 1021–1025 013 14-11-2010 7.4322º, -79.5612º 7.4448º, -79.5367º 711–733 014 14-11-2010 7.5432º, -79.4480º 7.5437º, -79.4202º 432–435 015 14-11-2010 7.8482º, -79.6457º 7.8697º, -79.6283º 113–114 016 15-11-2010 7.2443º, -79.6677º 7.2558º, -79.6460º 1446–1454 ......continued on the next pageROBERTSON ET AL .6 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 1. (Continued) SN Date Coordinates DR Initial Final 017 15-11-2010 7.2712º, -79.6645º 7.2602º, -79.6878º 1072–1141 018 15-11-2010 7.3492º, -79.6677º 7.3675º, -79.6487º 720–733 019 15-11-2010 7.3857º, -79.7310º 7.3728º, -79.7548º 480–485 020 15-11-2010 7.4480º, -79.8400º 7.4632º, -79.8172º 108–109 021 16-11-2010 7.1622º, -80.1342º 7.1695º, -80.1152º 764–987 022 16-11-2010 7.1893º, -80.1697º 7.1747º, -80.1922º 416–454 023 16-11-2010 7.2467º, -80.2307º 7.2492º, -80.2560º 103–108 024 17-11-2010 7.0298º, -80.9660º 7.0275º, -80.9407º 1364–1464 025 17-11-2010 7.0890º, -80.9212º 7.0912º, -80.9468º 1101–1113 026 17-11-2010 7.1473º, -81.1248º 7.1558º, -81.1498º 929–954 027 17-11-2010 7.2192º, -81.0955º 7.2248º, -81.1217º 134–138 028 18-11-2010 6.9033º, -81.3120º 6.9190º, -81.2940º 1467–1472 029 18-11-2010 7.0802º, -81.3682º 7.0820º, -81.3958º 1113–1126 030 18-11-2010 7.0945º, -81.4622º 7.0840º, -81.4372º 864-1026 031 18-11-2010 7.1637º, -81.4083º 7.1632º, -81.4353º 611-617 032 18-11-2010 7.2542º, -81.3957º 7.2542º, -81.4240º 108 033 19-11-2010 7.0105º, -81.7333º 7.9872º, -81.7472º 716-842 034 19-11-2010 6.9502º, -81.7455º 6.9332º, -81.7663º 1033-1214 035 19-11-2010 7.0707º, -81.7105º 7.0638º, -81.6843º 671-677 036 19-11-2010 7.1817º, -81.6668º 7.1832º, -81.7018º 138-143 037 20-11-2010 7.5852º, -82.0828º 7.6118º, -82.0848º 453-456 038 20-11-2010 7.7235º, -82.0570º 7.7143º, -82.0838º 144-154 039 20-11-2010 7.7427º, -82.4213º 7.7185º, -82.4075º 237-243 040 20-11-2010 7.6408º, -82.4512º 7.6680º, -82.4510º 977-1185 041 21-11-2010 8.0327º, -82.5642º 8.0042º, -82.5678º 115-152 042 21-11-2010 7.9820º, -82.7405º 7.9955º, -82.7632º 1115-1119 043 21-11-2010 7.9455º, -82.8083º 7.9693º, -82.8190º 860-951 044 21-11-2010 7.8910º, -82.8422º 7.8777º, -82.8652º 1093-1302 045 22-11-2010 8.2675º, -83.1892º 8.2662º, -83.1633º 1362-1376 046 22-11-2010 8.2318º, -83.1762º 8.2247º, -83.1500º 1157-1260 047 22-11-2010 8.3060º, -83.1608º 8.3053º, -83.1587º 416-421 048 22-11-2010 8.3107º, -83.1228º 8.3243º, -83.1483º 128-143 049 23-11-2010 8.4503º, -83.7362º 8.4640º, -83.7612º 128-136 050 23-11-2010 8.8020º, -84.0280º 8.8310º, -84.0325º 119-125 051 23-11-2010 8.7722º, -84.0987º 8.7843º, -84.0738º 359-477 052 23-11-2010 8.7675º, -84.2622º 8.7430º, -84.2745º 836-1030 053 24-11-2010 9.0260º, -84.5930º 9.0127º, -84.5703º 1359-1443 054 24-11-2010 9.0813º, -84.5750º 9.0958º, -84.5972º 1188-1257 055 24-11-2010 9.1472º, -84.5613º 9.1345º, -84.5382º 902-924 056 24-11-2010 9.2018º, -84.4948º 9.1920º, -84.5203º 656-668 057 24-11-2010 9.2403º, -84.5700º 9.2342º, -84.5975º 134-162 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 7TEP DEEP FISHES TABLE 1. (Continued) SN Date Coordinates DR Initial Final 058 25-11-2010 9.3918º, -85.1712º 9.3773º, -85.1935º 1181-1270 059 25-11-2010 9.4280º, -85.1613º 9.4258º, -85.1880º 841-920 060 25-11-2010 9.4500º, -85.1500º 9.4468º, -85.1792º 516-672 061 25-11-2010 9.5070º, -85.1503º 9.5128º, -85.1772º 137-140 062 25-11-2010 9.3670º, -85.4537º 9.3507º, -85.4355º 1414-1457 063 3-12-2010 9.7298º, -85.8315º 9.7427º, -85.8502º 1530-1625 064 3-12-2010 9.6805º, -85.7273º 9.6997º, -85.7215º 950-1105 065 3-12-2010 9.6650º, -85.7318º 9.6468º, -85.7453º 1178-1310 066 3-12-2010 9.7878º, -85.6773º 9.8125º, -85.6983º 127-128 067 4-12-2010 9.9925º, -85.9738º 9.9925º, -85.9738º 936 068 4-12-2010 10.1498º, -85.9432º 10.1498º, -85.9432º 145 069 4-12-2010 10.6927º, -86.5030º 10.6693º, -86.4860º 470-490 070 4-12-2010 10.6843º, -86.4147º 10.7045º, -86.4373º 237-243 071 5-12-2010 10.6875º, -86.5522º 10.7050º, -86.5743º 827-965 072 5-12-2010 10.4913º, -86.4583º 10.4702º, -86.4518º 1212-1330 073 5-12-2010 11.0842º, -86.7460º 11.0583º, -86.7375º 153-176 074 6-12-2010 11.0700º, -86.8317º 11.5375º, -86.8165º 417-422 075 6-12-2010 11.0173º, -86.8652º 10.9923º, -86.8507º 784-844 076 6-12-2010 10.8393º, -86.7483º 10.8477º, -86.7220º 926-1145 077 7-12-2010 11.4332º, -87.4430º 11.3997º, -87.4217º 1320-1370 078 7-12-2010 11.4257º, -87.4013º 11.4105º, -87.3782º 1076-1087 079 7-12-2010 11.4467º, -87.2937º 11.3948º, -87.2735º 825-842 080 7-12-2010 11.4812º, -87.2493º 11.4988º, -87.2703º 460-479 081 7-12-2010 11.7030º, -87.0635º 11.7212º, -87.0863º 122-124 082 8-12-2010 11.7275º, -87.6672º 11.7417º, -87.6890º 1471-1527 083 8-12-2010 11.8460º, -87.7090º 11.8715º, -87.7063º 965-1116 084 8-12-2010 11.9095º, -87.6427º 11.8912º, -87.6198º 404-510 085 8-12-2010 11.9318º, -87.5585º 11.9095º, -87.5410º 181-186 086 9-12-2010 12.1210º, -88.2688º 12.1453º, -88.2797º 1353-1389 087 9-12-2010 12.1712º, -88.2493º 12.1792º, -88.2745º 1100-1223 088 9-12-2010 12.2013º, -88.1990º 12.2273º, -88.1885º 570-767 089 9-12-2010 12.2278º, -88.1718º 12.2013º, -88.1620º 450-542 090 9-12-2010 12.2900º, -88.0747º 12.3158º, -88.0852º 129-131 091 10-12-2010 12.7440º, -88.9943º 12.7478º, -89.0233º 105-107 092 10-12-2010 13.0198º, -89.4582º 13.0027º, -89.4362º 103-104 093 10-12-2010 12.8307º, -89.5342º 12.8407º, -89.5588º 401-405 094 11-12-2010 12.8877º, -90.1585º 12.8897º, -90.1323º 1368-1406 095 11-12-2010 12.9578º, -90.2545º 12.9717º, -90.2792º 787-856 096 11-12-2010 12.9610º, -90.3173º 12.9808º, -90.3303º 1100-1129 097 11-12-2010 13.0417º, -90.2222º 13.0508º, -90.2375º 407-411 098 11-12-2010 13.3133º, -90.0032º 13.3273º, -90.0173º 111-114 ......continued on the next pageROBERTSON ET AL .8 · Zootaxa 4348 (1) © 2017 Magnolia Press Specimens were selected by DRR in an attempt to obtain relatively undamaged individuals representative of species caught during the cruise, rather than a representative sample of each trawl’s catch. Hence only a few specimens of common species were retained from a few trawls, even when they were present in many more trawls. No specimens were retained from 32 trawls. A pdf of a PowerPoint presentation made at the end of the cruise (by IJFN) that provides information about the ship and summarizes cruise activities and results is available at http:// hdl.handle.net/10508/1555. This pdf provides more information about catches, particularly of common species. The fishes retained as specimens by DRR were photographed by him and then frozen in the ship’s cold room. Small fishes were photographed in a 30 cm long, 25 cm high and 5 cm wide aquarium full of water that contained an angled glass plate to hold the fish suspended ~horizontally in midwater. Larger fishes were photographed in air laid out on a foam rubber (yoga-mat) background glued to a rigid plywood base. When needed, insect pins were used to hold fins erect against that background. Due to time limitations photo specimens were not measured, or individually tagged or kept separate from other specimens retained from the same trawl. At the end of the cruise the frozen specimens were stored in freezers at the Naos Laboratory of the Smithsonian Tropical Research Institute (STRI) in Panama. Subsequently a group of technicians and ichthyologists from the Smithsonian National Museum of Natural History in Washington, DC (formerly United States National Museum; USNM) went to Panama, defrosted, photographed, and tissue-sampled most of these specimens, which were then preserved in formalin. Several months later the formalized specimens were shipped to that museum and accessioned into its collection. Analysis and presentation of data. The checklist is arranged phylogenetically at the ordinal and familial levels following Eschmeyer & Fong (2017). Genera and species within families are arranged in alphabetical order. Number of families (F), genera (G) and species (S) are indicated in each more inclusive group (Order and Families), when appropriate. Popular, technical and/or vernacular names, in English (En) and Spanish (Sp), are included for each family following Robertson & Allen (2015) and Nelson et al. (2016). A considerable number of comprehensive taxonomic papers and guides were helpful in the identification of the specimens in the laboratory (e.g. Fitch & Lavenberg 1968, Eschmeyer et al. 1983, Cohen et al. 1990, Allen & Robertson 1994, Bussing & López 1994, Fisher et al. 1995, Nielsen et al. 1999, Nayaka et al. 2009, Robertson & Allen 2015). In addition to these general works, other more circumscribed taxonomic publications were employed. For each species listed below the following information is provided: valid scientific name with author(s) and year of description, following Eschmeyer et al. (2017), except when noted; popular, technical and/or vernacular names in English (En) and Spanish (Sp), when available, following Page et al. (2013), Robertson & Allen (2015) and Froese & Pauly (2016); collection data (localities—MOP stations—see Table 1; and depth range); voucher specimens (number of specimens examined, with their catalog numbers); current known distribution; literature used in its identification; and other pertinent information (e.g. substantial range extensions in the“Remarks” section for different species). When it was possible to identify the USNM catalog number of the photographed specimen of any species that number is given in the figure legend, together with the length of the specimen (determined from inspection of the thawed-fish photographs, each of which was taken with the specimen positioned above a 20-cm ruler). TABLE 1. (Continued) SN Date Coordinates DR Initial Final 099 12-12-2010 13.0830º, -90.6020º 13.1008º, -90.6215º 1054-1188 100 12-12-2010 13.1135º, -90.5943º 13.1270º, -90.5715º 697-798 101 12-12-2010 13.1793º, -90.5318º 13.2017º, -90.5205º 346-399 102 12-12-2010 13.2927º, -90.4045º 13.3213º, -90.4013º 149-165 103 13-12-2010 13.7140º, -91.8352º 13.7187º, -91.8647º 117-120 104 13-12-2010 13.5465º, -91.3480º 13.5313º, -91.3232º 114-116 105 13-12-2010 13.4770º, -91.3972º 13.4905º, -91.4220º 137 106 13-12-2010 13.6578º, -91.2283º 13.6667º, -91.2565º 57 107 14-12-2010 13.7593º, -91.8780º 13.7642º, -91.9058º 109-112 Zootaxa 4348 (1) © 2017 Magnolia Press · 9TEP DEEP FISHES The “Literature” section also includes other references arbitrarily considered by us to have taxonomic or distributional value and not mere usage of the specific names herein treated. In addition, for most species taken on the cruise, photographs of the fresh specimens as well as DNA barcodes (see below) are provided. DNA barcoding. DNA was extracted using an Autogen Prep 965 pheno-chloroform automated extractor. The target 652 base pair (bp) region of the 5' end of the cytochrome oxidase subunit 1 gene (COI) was amplified using primers FISHCO1LBC and FISHCO1HBC and PCR and sequencing conditions following Baldwin et al. (2011). Electropherograms were processed using Sequencher 5.0.1 (gene codes). Only trimmed fragments greater than 500 bp in length and with overall "confidence" above 90% (as calculated by Sequencher) were used to construct the final sequences. Passing sequences were visually examined for errors. Barcode sequences are deposited in Genbank with the accession numbers MF956433 - MF957114 (see Table 3). Detailed specimen information and associated photographs are available in the public project “Fish of the 2010 Miguel Oliver Pacific Cruise” in the Barcode of Life Database (BOLD; www.boldsystems.org). Results and discussion A total of 876 specimens of bony fishes collected on the 2010 Miguel Oliver cruise along the Pacific coast of Central America were accessioned into the USNM. That collection also included elasmobranchs, chimaeras and cyclostomes as well as the bony fishes treated here. Fifteen orders, 61 families, 97 genera and 129 species of bony fishes are listed herein (Table 2). Perciformes (33 species), Lophiiformes (13), Scorpaeniformes (12), Anguilliformes (11) and Gadiformes (11) were the most speciose orders, accounting for 62.0% of the total number of species recorded. Ophidiidae (9 species), Zoarcidae (9), Macrouridae (8), Alepocephalidae (6) and Ogcocephalidae (6) were the most speciose families, accounting for 29.5% of the total number of species recorded. Thirty-two families (52.4% of the total number of families recorded) were represented by a single species. Twenty- nine species (22.5%), in 16 families, are added to the Central American Pacific ichthyofauna. Seventeen species (13.3%), in 13 families (21.3%), showed expanded latitudinal (geographical) ranges, and 31 species (24 %), in 23 (37.7%) families, showed expanded depth ranges. The DNA barcodes that resulted from this collection (those of 682 individuals) include barcodes from 118 species belonging to 95 genera and 61 families of bony fishes (Table 3). These include members of some genera that could not be identified to species. Those DNA barcodes represent the first flagged barcodes in Genbank for 84 species and the first for 30 genera. In addition the barcodes of 54 of those 84 species are the first genetic sequences of any type published for those species in Genbank (Table 3). Our analysis of this collection of fishes clearly demonstrates how little is known about the species diversity, geographic distributions, and depth ranges of the deep-water fish fauna inhabitating the edge of the continental shelf and continental slope of the tropical eastern Pacific. In addition, the results of the DNA barcoding not only show how few species from this area have genetic information available, but also how few members of many genera have genetic samples available for public use. A similar level of new knowledge can be expected when the specimens of elasmobranchs, chimaeras and cyclostomes collected during the same cruise are fully analyzed. The description of two new species of elasmobranchs (see Vásquez et al. 2015 and Concha et al. 2016), based in whole or part on specimens collected during the present cruise, clearly indicates that this will be the case. To date only one new species of a genus not previously recorded in the eastern Pacific has been added to the regional bony-fish fauna (Angulo et al. 2016). However, the DNA barcode data also point to taxonomic issues that need resolution in a variety of genera, including Bathypterois, Bollmannia, Melamphaes, Notacanthus, Paraliparis, Synodus, and some Zoarcids (i.e. Lycodapus, Lycenchelys, Lycodes, Ophthalmolycus and Pachycara). Continued ocean exploration of the fish fauna of this area involving a broader range of collection methods is needed. Sampling with, for example, more maneuverable, smaller-mesh bottom trawls, mid-water trawls, and long- lines is likely to yield much new information on other members of the fish fauna of the tropical eastern Pacific that were not susceptible to capture by the large, coarse-mesh bottom trawl used by the B/O Miguel Oliver in 2010. A return of a ship like the B/O Miguel Oliver to the eastern Pacific would greatly benefit knowledge of the deepwater fish fauna of the tropical west coast of the Americas.ROBERTSON ET AL .10 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 2. Number of families (F), genera (G) and species (Sp), by order, of bony fishes collected by the research vessel B/O Miguel Oliver on the Pacific coast of Central America in November-December 2010. NR = number of species representing new records for the study area; RE = number of species with extended geographical ranges; NDR = number of species with extended depth ranges; No = total number of individuals examined for this study. TABLE 3. Specimens barcoded as part of this study. The status column indicates the significance of the new sequence (1=first barcode-flagged sequence for this species; 2=first barcode-flagged sequence and first public sequence for this species; 3= first barcode-flagged sequence for this genus; 4 = first barcode-flagged sequence and first public sequence of this genus). An asterisk in the Voucher Number column indicates that only a tissue sample of this specimen is cataloged at the USNM, not the physical specimen. Order F G Sp NR RE NDR No Notacanthiformes 2 2 4 4 2 3 22 Anguilliformes 7 9 11 1 1 4 62 Osmeriformes 4 10 10 7 2 1 63 Stomiiformes 3 3 3 0 0 0 23 Aulopiformes 5 5 7 2 0 1 36 Myctophiformes 2 3 3 0 0 1 17 Gadiformes 4 6 11 3 2 6 73 Ophidiiformes 1 8 9 4 4 5 67 Lophiiformes 6 7 13 4 3 5 66 Stephanoberyciformes 1 2 3 1 0 0 12 Beryciformes 2 2 2 0 0 0 10 Scorpaeniformes 6 7 12 2 2 2 57 Perciformes 14 27 33 1 1 2 163 Pleuronectiformes 3 5 7 0 0 1 35 Tetraodontiformes 1 1 1 0 0 0 1 Total 61 97 129 29 17 31 707 Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Alepocephalus tenebrosus MOP033-12 USNM 422642 MF956437 Alepocephalus tenebrosus MOP098-12 USNM 422463 MF956439 Alepocephalus tenebrosus MOP099-12 USNM 422398 MF956442 Alepocephalus tenebrosus MOP187-12 USNM 422396 MF956443 Alepocephalus tenebrosus MOP195-12 USNM 422612 MF956445 Alepocephalus tenebrosus MOP196-12 USNM 422457 MF956446 Alepocephalus tenebrosus MOP344-12 USNM 421560 MF956438 Alepocephalus tenebrosus MOP345-12 USNM 422529 MF956433 Alepocephalus tenebrosus MOP380-12 USNM 421555 MF956441 Alepocephalus tenebrosus MOP393-12 USNM 422556 MF956435 Alepocephalus tenebrosus MOP417-12 USNM 421518 MF956449 Alepocephalus tenebrosus MOP418-12 USNM 421451 MF956447 Alepocephalus tenebrosus MOP443-12 USNM 422619 MF956448 Alepocephalus tenebrosus MOP518-12 USNM 421282 MF956450 Alepocephalus tenebrosus MOP519-12 USNM 421280 MF956451 Alepocephalus tenebrosus MOP637-12 USNM 421573 MF956444 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 11TEP DEEP FISHES TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Alepocephalus tenebrosus MOP729-12 USNM 422616 MF956440 Alepocephalus tenebrosus MOP770-12 USNM 421471 MF956434 Alepocephalus tenebrosus MOP813-12 USNM 421242 MF956436 Anoplogaster cornuta MOP350-12 USNM 422533 MF956455 Anoplogaster cornuta MOP503-12 USNM 421537 MF956454 Anoplogaster cornuta MOP545-12 USNM 421295 MF956453 Anoplogaster cornuta MOP581-12 USNM 422626 MF956452 Argentina aliceae 4 MOP094-12 USNM 422405 MF956457 Argentina aliceae 4 MOP309-12 USNM 422575 MF956460 Argentina aliceae 4 MOP310-12 USNM 422548 MF956459 Argentina aliceae 4 MOP695-12 USNM 421411 MF956461 Argentina aliceae 4 MOP696-12 USNM 421424 MF956456 Argentina aliceae 4 MOP787-12 USNM 421222 MF956458 Ariosoma prorigerum 2 MOP263-12 USNM 422481 MF956468 Ariosoma prorigerum 2 MOP264-12 USNM 422480 MF956463 Ariosoma prorigerum 2 MOP342-12 USNM 423213 MF956462 Ariosoma prorigerum 2 MOP574-12 USNM 421209 MF956467 Ariosoma prorigerum 2 MOP575-12 USNM 421208 MF956465 Ariosoma prorigerum 2 MOP718-12 USNM 422482 MF956466 Ariosoma prorigerum 2 MOP806-12 USNM 421362 MF956469 Ariosoma prorigerum 2 MOP818-12 USNM 423212 MF956464 Avocettina bowersii 2 MOP476-12 USNM 421196 MF956472 Avocettina bowersii 2 MOP524-12 USNM 421290 MF956470 Avocettina bowersii 2 MOP645-12 USNM 421195 MF956471 Bajacalifornia burragei MOP034-12 USNM 422437 MF956473 Bajacalifornia burragei MOP766-12 USNM 421485 MF956474 Baldwinella eos 4 MOP013-12 USNM 422436 MF956475 Baldwinella eos 4 MOP014-12 USNM 422325 MF956477 Baldwinella eos 4 MOP015-12 USNM 422333 MF956476 Baldwinella eos 4 MOP284-12 USNM 421225 MF956478 Baldwinella eos 4 MOP325-12 USNM 422589 MF956480 Baldwinella eos 4 MOP326-12 USNM 422558 MF956479 Balistes polylepis MOP238-12 USNM 421241 MF956481 Barbantus curvifrons 3 MOP517-12 USNM 421294 MF956483 Barbantus curvifrons 3 MOP704-12 USNM 422531 MF956482 Bathypterois atricolor 3 MOP077-12 USNM 422418 MF956489 Bathypterois atricolor 3 MOP078-12 USNM 422395 MF956490 Bathypterois atricolor 3 MOP178-12 USNM 422427 MF956488 Bathypterois atricolor 3 MOP428-12 USNM 421538 MF956485 Bathypterois atricolor 3 MOP617-12 USNM 423598 MF956487 Bathypterois atricolor 3 MOP764-12 USNM 421469 MF956486 ......continued on the next pageROBERTSON ET AL .12 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Bathypterois sp. MOP101-12 USNM 422410 MF956484 Bathypterois ventralis 3 MOP398-12 USNM 422561 MF956496 Bathypterois ventralis 3 MOP399-12 USNM 422569 MF956499 Bathypterois ventralis 3 MOP429-12 USNM 421536 MF956493 Bathypterois ventralis 3 MOP466-12 USNM 421506 MF956501 Bathypterois ventralis 3 MOP467-12 USNM 421529 MF956497 Bathypterois ventralis 3 MOP468-12 USNM 421531 MF956495 Bathypterois ventralis 3 MOP521-12 USNM 421343 MF956500 Bathypterois ventralis 3 MOP522-12 USNM 421304 MF956494 Bathypterois ventralis 3 MOP546-12 USNM 421286 MF956491 Bathypterois ventralis 3 MOP742-12 USNM 422503 MF956498 Bathypterois ventralis 3 MOP765-12 USNM 421484 MF956492 Bathytroctes microlepis 3 MOP601-12 USNM 421563 MF956503 Bathytroctes microlepis 3 MOP778-12 USNM 421350 MF956502 Bellator loxias 3 MOP217-12 USNM 422360 MF956504 Bellator loxias 3 MOP218-12 USNM 421214 MF956506 Bellator loxias 3 MOP219-12 USNM 421274 MF956505 Bentartia pusillum MOP044-12 USNM 435808 MF956509 Bentartia pusillum MOP045-12 USNM 435800 MF956510 Bentartia pusillum MOP106-12 USNM 422425 MF956511 Bentartia pusillum MOP117-12 USNM 423184 MF956512 Bentartia pusillum MOP379-12 USNM 422560 MF956513 Bentartia pusillum MOP407-12 USNM 423199 MF956508 Bentartia pusillum MOP489-12 USNM 435805 MF956516 Bentartia pusillum MOP584-12 USNM 435791 MF956507 Bentartia pusillum MOP591-12 USNM 421466 MF956515 Bentartia pusillum MOP674-12 USNM 423210 MF956514 Benthosema panamense 1 MOP388-12 USNM 422523 MF956521 Benthosema panamense 1 MOP389-12 USNM 422541 MF956518 Benthosema panamense 1 MOP390-12 USNM 422484 MF956522 Benthosema panamense 1 MOP492-12 USNM 421449 MF956525 Benthosema panamense 1 MOP493-12 USNM 421453 MF956517 Benthosema panamense 1 MOP494-12 USNM 421450 MF956519 Benthosema panamense 1 MOP641-12 USNM 421419 MF956523 Benthosema panamense 1 MOP756-12 USNM 422486 MF956520 Benthosema panamense 1 MOP758-12 USNM 422536 MF956524 Bollmannia sp. 3 MOP084-12 USNM 422322 MF956536 Bollmannia sp. 3 MOP087-12 USNM 422381 MF956526 Bollmannia sp. 3 MOP278-12 USNM 422339 MF956529 Bollmannia sp. 3 MOP619-12 USNM 421381 MF956528 Bollmannia sp1 3 MOP085-12 USNM 422310 MF956535 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 13TEP DEEP FISHES TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Bollmannia sp1 3 MOP088-12 USNM 422401 MF956527 Bollmannia sp1 3 MOP691-12 USNM 421428 MF956537 Bollmannia sp1 3 MOP693-12 USNM 421397 MF956531 Bollmannia sp2 3 MOP086-12 USNM 422308 MF956534 Bollmannia sp2 3 MOP660-12 USNM 421387 MF956532 Bollmannia sp2 3 MOP661-12 USNM 421403 MF956533 Bollmannia sp2 3 MOP692-12 USNM 421380 MF956530 Bothrocara molle MOP063-12 USNM 422430 MF956540 Bothrocara molle MOP064-12 USNM 422331 MF956539 Bothrocara molle MOP107-12 USNM 422424 MF956538 Bothrocara molle MOP198-12 USNM 423188 MF956545 Bothrocara molle MOP346-12 USNM 422570 MF956542 Bothrocara molle MOP378-12 USNM 422584 MF956544 Bothrocara molle MOP408-12 USNM 423200 MF956541 Bothrocara molle MOP490-12 USNM 423202 MF956543 Bregmaceros bathymaster 2 MOP156-12 USNM 422379 MF956547 Bregmaceros bathymaster 2 MOP157-12 USNM 422387 MF956548 Bregmaceros bathymaster 2 MOP676-12 USNM 421278 MF956546 Bregmaceros bathymaster 2 MOP677-12 USNM 421377 MF956550 Bregmaceros bathymaster 2 MOP678-12 USNM 421416 MF956549 Caulolatilus affinis MOP710-12 USNM 422632 MF956552 Caulolatilus affinis MOP747-12 USNM 422646 MF956551 Caulophryne pelagica 3 MOP605-12 USNM 422598 MF956553 Cherublemma emmelas 4 MOP136-12 USNM 422407 MF956561 Cherublemma emmelas 4 MOP137-12 USNM 422374 MF956564 Cherublemma emmelas 4 MOP337-12 USNM 422581 MF956562 Cherublemma emmelas 4 MOP338-12 USNM 422549 MF956565 Cherublemma emmelas 4 MOP339-12 USNM 422557 MF956560 Cherublemma emmelas 4 MOP384-12 USNM 422567 MF956563 Cherublemma emmelas 4 MOP385-12 USNM 422544 MF956555 Cherublemma emmelas 4 MOP555-12 USNM 421337 MF956558 Cherublemma emmelas 4 MOP671-12 USNM 421565 MF956556 Cherublemma emmelas 4 MOP672-12 USNM 421429 MF956557 Cherublemma emmelas 4 MOP673-12 USNM 421492 MF956559 Cherublemma emmelas 4 MOP803-12 USNM 421218 MF956554 Chiasmodon subniger 4 MOP141-12 USNM 435803 MF956566 Chiasmodon subniger 4 MOP709-12 USNM 422491 MF956568 Chiasmodon subniger 4 MOP768-12 USNM 435801 MF956567 Chlorophthalmus mento MOP783-12 USNM 421332 MF956569 Citharichthys platophrys MOP299-12 USNM 422576 MF956572 Citharichthys platophrys MOP538-12 USNM 421298 MF956571 ......continued on the next pageROBERTSON ET AL .14 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Citharichthys platophrys MOP539-12 USNM 421283 MF956570 Coelorinchus canus 2 MOP553-12 USNM 421292 MF956576 Coelorinchus canus 2 MOP554-12 USNM 421291 MF956573 Coelorinchus canus 2 MOP751-12 USNM 422487 MF956574 Coelorinchus canus 2 MOP752-12 USNM 422495 MF956575 Coryphaenoides anguliceps 2 MOP039-12 USNM 422467 MF956581 Coryphaenoides anguliceps 2 MOP040-12 USNM 422447 MF956578 Coryphaenoides anguliceps 2 MOP171-12 USNM 422610 MF956579 Coryphaenoides anguliceps 2 MOP175-12 USNM 422456 MF956582 Coryphaenoides anguliceps 2 MOP396-12 USNM 422648 MF956584 Coryphaenoides anguliceps 2 MOP400-12 USNM 423198 MF956577 Coryphaenoides anguliceps 2 MOP430-12 USNM 422620 MF956580 Coryphaenoides anguliceps 2 MOP431-12 USNM 421564 MF956583 Coryphaenoides capito 2 MOP162-12 USNM 422370 MF956588 Coryphaenoides capito 2 MOP253-12 USNM 421235 MF956586 Coryphaenoides capito 2 MOP254-12 USNM 421273 MF956587 Coryphaenoides capito 2 MOP257-12 USNM 421312 MF956595 Coryphaenoides capito 2 MOP258-12 USNM 422355 MF956585 Coryphaenoides capito 2 MOP412-12 USNM 421461 MF956592 Coryphaenoides capito 2 MOP413-12 USNM 421456 MF956593 Coryphaenoides capito 2 MOP469-12 USNM 421480 MF956589 Coryphaenoides capito 2 MOP470-12 USNM 421444 MF956590 Coryphaenoides capito 2 MOP471-12 USNM 421460 MF956591 Coryphaenoides capito 2 MOP571-12 USNM 421341 MF956594 Coryphaenoides carminifer 2 MOP038-12 USNM 422441 MF956601 Coryphaenoides carminifer 2 MOP067-12 USNM 422446 MF956603 Coryphaenoides carminifer 2 MOP174-12 USNM 422433 MF956599 Coryphaenoides carminifer 2 MOP177-12 USNM 422419 MF956600 Coryphaenoides carminifer 2 MOP432-12 USNM 422649 MF956597 Coryphaenoides carminifer 2 MOP433-12 USNM 421475 MF956598 Coryphaenoides carminifer 2 MOP434-12 USNM 421557 MF956604 Coryphaenoides carminifer 2 MOP610-12 USNM 421432 MF956596 Coryphaenoides carminifer 2 MOP722-12 USNM 422634 MF956605 Coryphaenoides carminifer 2 MOP723-12 USNM 422555 MF956602 Cubiceps pauciradiatus 1 MOP243-12 USNM 422357 MF956608 Cubiceps pauciradiatus 1 MOP505-12 USNM 421513 MF956610 Cubiceps pauciradiatus 1 MOP633-12 USNM 421436 MF956609 Cubiceps pauciradiatus 1 MOP634-12 USNM 421415 MF956606 Cubiceps pauciradiatus 1 MOP738-12 USNM 422496 MF956607 Cynoscion nannus 2 MOP150-12 USNM 422364 MF956614 Cynoscion nannus 2 MOP151-12 USNM 422404 MF956613 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 15TEP DEEP FISHES TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Cynoscion nannus 2 MOP152-12 USNM 422403 MF956612 Cynoscion nannus 2 MOP295-12 USNM 421314 MF956620 Cynoscion nannus 2 MOP495-12 USNM 421517 MF956615 Cynoscion nannus 2 MOP496-12 USNM 421498 MF956616 Cynoscion nannus 2 MOP497-12 USNM 421499 MF956621 Cynoscion nannus 2 MOP498-12 USNM 421500 MF956627 Cynoscion nannus 2 MOP499-12 USNM 421446 MF956622 Cynoscion nannus 2 MOP527-12 USNM 421462 MF956631 Cynoscion nannus 2 MOP528-12 USNM 421483 MF956629 Cynoscion nannus 2 MOP529-12 USNM 421455 MF956634 Cynoscion nannus 2 MOP542-12 USNM 421285 MF956617 Cynoscion nannus 2 MOP543-12 USNM 421288 MF956611 Cynoscion nannus 2 MOP544-12 USNM 421303 MF956632 Cynoscion nannus 2 MOP664-12 USNM 421400 MF956623 Cynoscion nannus 2 MOP665-12 USNM 421409 MF956624 Cynoscion nannus 2 MOP666-12 USNM 421437 MF956626 Cynoscion nannus 2 MOP680-12 USNM 422586 MF956633 Cynoscion nannus 2 MOP681-12 USNM 421420 MF956635 Cynoscion nannus 2 MOP688-12 USNM 421568 MF956618 Cynoscion nannus 2 MOP689-12 USNM 421569 MF956619 Cynoscion nannus 2 MOP745-12 USNM 422513 MF956637 Cynoscion nannus 2 MOP746-12 USNM 422521 MF956636 Cynoscion nannus 2 MOP810-12 USNM 421255 MF956625 Cynoscion nannus 2 MOP811-12 USNM 421221 MF956628 Cynoscion nannus 2 MOP812-12 USNM 421251 MF956630 Decapterus macrosoma MOP304-12 USNM 421558 MF956638 Decapterus macrosoma MOP785-12 USNM 421355 MF956639 Decodon melasma 3 MOP305-12 USNM 422622 MF956642 Decodon melasma 3 MOP324-12 USNM 422591 MF956641 Decodon melasma 3 MOP687-12 USNM 421393 MF956643 Decodon melasma 3 MOP711-12 USNM 422522 MF956640 Dibranchus erinaceus 4 MOP334-12 USNM 422525 MF956648 Dibranchus erinaceus 4 MOP335-12 USNM 422554 MF956647 Dibranchus erinaceus 4 MOP336-12 USNM 422572 MF956646 Dibranchus erinaceus 4 MOP686-12 USNM 421396 MF956645 Dibranchus erinaceus 4 MOP801-12 USNM 421226 MF956644 Dibranchus erinaceus 4 MOP804-12 USNM 421253 MF956650 Dibranchus erinaceus 4 MOP805-12 USNM 421256 MF956649 Dibranchus hystrix 4 MOP186-12 USNM 422413 MF956652 Dibranchus hystrix 4 MOP561-12 USNM 422601 MF956654 Dibranchus hystrix 4 MOP586-12 USNM 421279 MF956651 ......continued on the next pageROBERTSON ET AL .16 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Dibranchus hystrix 4 MOP734-12 USNM 422606 MF956655 Dibranchus hystrix 4 MOP735-12 USNM 421467 MF956653 Dibranchus nudivomer 4 MOP363-12 USNM 422534 MF956658 Dibranchus nudivomer 4 MOP364-12 USNM 422578 MF956656 Dibranchus nudivomer 4 MOP733-12 USNM 422524 MF956657 Dibranchus spinosus 4 MOP104-12 USNM 422385 MF956661 Dibranchus spinosus 4 MOP506-12 USNM 421349 MF956665 Dibranchus spinosus 4 MOP507-12 USNM 421340 MF956666 Dibranchus spinosus 4 MOP585-12 USNM 422595 MF956659 Dibranchus spinosus 4 MOP602-12 USNM 422625 MF956660 Dibranchus spinosus 4 MOP684-12 USNM 421395 MF956663 Dibranchus spinosus 4 MOP685-12 USNM 421386 MF956664 Dibranchus spinosus 4 MOP720-12 USNM 422618 MF956662 Dibranchus spinosus 4 MOP732-12 USNM 421470 MF956667 Dibranchus velutinus 4 MOP125-12 USNM 422459 MF956671 Dibranchus velutinus 4 MOP126-12 USNM 422409 MF956669 Dibranchus velutinus 4 MOP127-12 USNM 422391 MF956673 Dibranchus velutinus 4 MOP159-12 USNM 422453 MF956672 Dibranchus velutinus 4 MOP160-12 USNM 422420 MF956668 Dibranchus velutinus 4 MOP161-12 USNM 422363 MF956674 Dibranchus velutinus 4 MOP453-12 USNM 421509 MF956670 Dicrolene filamentosa 4 MOP074-12 USNM 422442 MF956679 Dicrolene filamentosa 4 MOP111-12 USNM 422406 MF956680 Dicrolene filamentosa 4 MOP272-12 USNM 422643 MF956685 Dicrolene filamentosa 4 MOP409-12 USNM 421523 MF956689 Dicrolene filamentosa 4 MOP419-12 USNM 421497 MF956688 Dicrolene filamentosa 4 MOP439-12 USNM 421533 MF956691 Dicrolene filamentosa 4 MOP440-12 USNM 421526 MF956690 Dicrolene filamentosa 4 MOP472-12 USNM 421495 MF956675 Dicrolene filamentosa 4 MOP473-12 USNM 421520 MF956687 Dicrolene filamentosa 4 MOP474-12 USNM 421496 MF956686 Dicrolene filamentosa 4 MOP548-12 USNM 421482 MF956684 Dicrolene filamentosa 4 MOP582-12 USNM 421488 MF956683 Dicrolene filamentosa 4 MOP607-12 USNM 422588 MF956682 Dicrolene filamentosa 4 MOP611-12 USNM 421561 MF956681 Dicrolene filamentosa 4 MOP743-12 USNM 422494 MF956678 Dicrolene filamentosa 4 MOP763-12 USNM 422613 MF956677 Dicrolene filamentosa 4 MOP772-12 USNM 421477 MF956676 Diplectrum euryplectrum 2 MOP227-12 USNM 421315 MF956692 Diplectrum euryplectrum 2 MOP228-12 USNM 421266 MF956693 Engyophrys sanctilaurentii 4 MOP237-12 USNM 421316 MF956696 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 17TEP DEEP FISHES TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Engyophrys sanctilaurentii 4 MOP316-12 USNM 422537 MF956695 Engyophrys sanctilaurentii 4 MOP317-12 USNM 422515 MF956694 Fowlerichthys avalonis 3 MOP286-12 USNM 422343 MF956697 Gymnothorax equatorialis 2 MOP052-12 USNM 421373 MF956698 Gymnothorax phalarus 2 MOP698-12 USNM 421374 MF956699 Halosaurus attenuatus 4 MOP367-12 USNM 422539 MF956703 Halosaurus attenuatus 4 MOP368-12 USNM 421547 MF956701 Halosaurus attenuatus 4 MOP510-12 USNM 421271 MF956702 Halosaurus attenuatus 4 MOP511-12 USNM 421258 MF956700 Halosaurus radiatus 4 MOP259-12 USNM 421311 MF956706 Halosaurus radiatus 4 MOP260-12 USNM 421310 MF956709 Halosaurus radiatus 4 MOP261-12 USNM 421275 MF956708 Halosaurus radiatus 4 MOP483-12 USNM 421521 MF956707 Halosaurus radiatus 4 MOP484-12 USNM 421516 MF956710 Halosaurus radiatus 4 MOP485-12 USNM 421379 MF956705 Halosaurus radiatus 4 MOP775-12 USNM 421534 MF956704 Hemanthias signifer MOP012-12 USNM 422478 MF956714 Hemanthias signifer MOP223-12 USNM 422359 MF956717 Hemanthias signifer MOP224-12 USNM 421237 MF956711 Hemanthias signifer MOP225-12 USNM 422361 MF956719 Hemanthias signifer MOP226-12 USNM 421252 MF956718 Hemanthias signifer MOP306-12 USNM 422573 MF956713 Hemanthias signifer MOP307-12 USNM 422535 MF956712 Hemanthias signifer MOP327-12 USNM 422633 MF956715 Hemanthias signifer MOP328-12 USNM 421551 MF956716 Hippoglossina bollmani 2 MOP019-12 USNM 422468 MF956724 Hippoglossina bollmani 2 MOP234-12 USNM 421327 MF956722 Hippoglossina bollmani 2 MOP235-12 USNM 421239 MF956721 Hippoglossina bollmani 2 MOP236-12 USNM 421313 MF956720 Hippoglossina bollmani 2 MOP296-12 USNM 421351 MF956726 Hippoglossina bollmani 2 MOP536-12 USNM 421489 MF956723 Hippoglossina bollmani 2 MOP537-12 USNM 421339 MF956725 Hippoglossina tetrophthalma 2 MOP534-12 USNM 422637 MF956728 Hippoglossina tetrophthalma 2 MOP535-12 USNM 422597 MF956727 Hoplostethus mento 2 MOP239-12 USNM 421306 MF956730 Hoplostethus mento 2 MOP240-12 USNM 421317 MF956729 Hoplostethus mento 2 MOP241-12 USNM 421260 MF956733 Hoplostethus mento 2 MOP569-12 USNM 421268 MF956732 Hoplostethus mento 2 MOP570-12 USNM 421284 MF956731 Hyporthodus niphobles 1 MOP046-12 USNM 423181 MF956734 Hyporthodus niphobles 1 MOP048-12 USNM 422641 MF956735 ......continued on the next pageROBERTSON ET AL .18 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Ilyophis sp. MOP731-12 USNM 421372 MF956736 Kathetostoma averruncus MOP022-12 USNM 422305 MF956740 Kathetostoma averruncus MOP203-12 USNM 421261 MF956742 Kathetostoma averruncus MOP204-12 USNM 421220 MF956739 Kathetostoma averruncus MOP279-12 USNM 421231 MF956738 Kathetostoma averruncus MOP280-12 USNM 421232 MF956743 Kathetostoma averruncus MOP281-12 USNM 421228 MF956737 Kathetostoma averruncus MOP784-12 USNM 421335 MF956741 Lampanyctus parvicauda 1 MOP035-12 USNM 422304 MF956744 Lamprogrammus niger 3 MOP115-12 USNM 422477 MF956745 Lamprogrammus niger 3 MOP347-12 USNM 422593 MF956746 Lamprogrammus niger 3 MOP706-12 USNM 435786 MF956747 Lestidiops neles 2 MOP245-12 USNM 421259 MF956748 Leuroglossus urotranus 2 MOP249-12 USNM 421262 MF956749 Lophiodes caulinaris MOP090-12 USNM 422366 MF956754 Lophiodes caulinaris MOP091-12 USNM 422412 MF956752 Lophiodes caulinaris MOP092-12 USNM 422388 MF956753 Lophiodes caulinaris MOP205-12 USNM 421345 MF956755 Lophiodes caulinaris MOP206-12 USNM 421346 MF956750 Lophiodes caulinaris MOP786-12 USNM 421347 MF956751 Lophiodes spilurus MOP550-12 USNM 421230 MF956761 Lophiodes spilurus MOP551-12 USNM 421281 MF956760 Lophiodes spilurus MOP552-12 USNM 421229 MF956759 Lophiodes spilurus MOP632-12 USNM 421427 MF956757 Lophiodes spilurus MOP759-12 USNM 422527 MF956758 Lophiodes spilurus MOP802-12 USNM 421348 MF956756 Luciobrotula sp. 3 MOP383-12 USNM 422550 MF956762 Luciobrotula sp. 3 MOP488-12 USNM 421528 MF956765 Luciobrotula sp. 3 MOP705-12 USNM 421491 MF956763 Luciobrotula sp. 3 MOP782-12 USNM 421356 MF956764 Luciobrotula sp. 3 MOP814-12 USNM 421217 MF956766 Lycenchelys sp. MOP376-12 USNM 422609 MF956769 Lycenchelys sp. MOP441-12 USNM 421515 MF956767 Lycenchelys sp. MOP730-12 USNM 422617 MF956768 Lycodapus sp. MOP075-12 USNM 422332 MF956771 Lycodapus sp. MOP643-12 USNM 421188 MF956770 Lycodes sp. MOP381-12 USNM 421556 MF956773 Lycodes sp. MOP382-12 USNM 422538 MF956772 Macrouridae MOP721-12 USNM 440227* MF956774 Melamphaes sp. MOP037-12 USNM 422342 MF956777 Melamphaes sp. MOP167-12 USNM 422386 MF956780 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 19TEP DEEP FISHES TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Melamphaes sp. MOP168-12 USNM 422390 MF956782 Melamphaes sp. MOP250-12 USNM 422348 MF956778 Melamphaes sp. MOP251-12 USNM 421265 MF956776 Melamphaes sp. MOP454-12 USNM 421505 MF956785 Melamphaes sp. MOP455-12 USNM 421504 MF956775 Melamphaes sp. MOP456-12 USNM 421503 MF956784 Melamphaes sp. MOP523-12 USNM 421267 MF956783 Melamphaes sp. MOP635-12 USNM 421421 MF956781 Melamphaes sp. MOP776-12 USNM 422532 MF956779 Melanocetus johnsonii MOP030-12 USNM 422319 MF956787 Melanocetus johnsonii MOP105-12 USNM 422465 MF956786 Melanocetus johnsonii MOP403-12 USNM 423214 MF956788 Melanocetus johnsonii MOP404-12 USNM 423215 MF956789 Melanocetus johnsonii MOP405-12 USNM 423216 MF956790 Melanocetus johnsonii MOP423-12 USNM 421459 MF956791 Melanocetus johnsonii MOP424-12 USNM 421457 MF956792 Melanocetus johnsonii MOP598-12 USNM 422628 MF956793 Melanocetus johnsonii MOP618-12 USNM 421406 MF956794 Melanocetus johnsonii MOP739-12 USNM 440228* MF956795 Merluccius angustimanus 1 MOP027-12 USNM 422438 MF956798 Merluccius angustimanus 1 MOP028-12 USNM 422470 MF956797 Merluccius angustimanus 1 MOP029-12 USNM 422450 MF956802 Merluccius angustimanus 1 MOP486-12 USNM 421524 MF956799 Merluccius angustimanus 1 MOP640-12 USNM 422587 MF956800 Merluccius angustimanus 1 MOP748-12 USNM 422528 MF956796 Merluccius angustimanus 1 MOP749-12 USNM 421468 MF956804 Merluccius angustimanus 1 MOP750-12 USNM 422498 MF956803 Merluccius angustimanus 1 MOP800-12 USNM 421331 MF956801 Monolene asaedae 4 MOP020-12 USNM 422328 MF956805 Monolene asaedae 4 MOP021-12 USNM 422323 MF956812 Monolene asaedae 4 MOP753-12 USNM 422501 MF956811 Monolene asaedae 4 MOP754-12 USNM 421486 MF956808 Monolene asaedae 4 MOP755-12 USNM 422493 MF956807 Monolene asaedae 4 MOP760-12 USNM 422488 MF956810 Monolene asaedae 4 MOP761-12 USNM 422516 MF956809 Monolene asaedae 4 MOP762-12 USNM 422490 MF956806 Monolene maculipinna 4 MOP297-12 USNM 422356 MF956814 Monolene maculipinna 4 MOP298-12 USNM 422351 MF956813 Monolene maculipinna 4 MOP313-12 USNM 421550 MF956815 Monolene maculipinna 4 MOP314-12 USNM 422517 MF956816 Monolene maculipinna 4 MOP315-12 USNM 422565 MF956817 ......continued on the next pageROBERTSON ET AL .20 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Monomitopus malispinosus 4 MOP247-12 USNM 421240 MF956825 Monomitopus malispinosus 4 MOP248-12 USNM 421249 MF956826 Monomitopus malispinosus 4 MOP475-12 USNM 421441 MF956829 Monomitopus malispinosus 4 MOP487-12 USNM 421530 MF956823 Monomitopus malispinosus 4 MOP576-12 USNM 421464 MF956822 Monomitopus malispinosus 4 MOP577-12 USNM 422624 MF956828 Monomitopus malispinosus 4 MOP578-12 USNM 421300 MF956820 Monomitopus malispinosus 4 MOP609-12 USNM 435794 MF956824 Monomitopus malispinosus 4 MOP715-12 USNM 422552 MF956821 Monomitopus malispinosus 4 MOP716-12 USNM 422509 MF956819 Monomitopus malispinosus 4 MOP717-12 USNM 422492 MF956827 Monomitopus sp. 3 MOP773-12 USNM 422497 MF956818 Nemichthys scolopaceus MOP080-12 USNM 421357 MF956833 Nemichthys scolopaceus MOP108-12 USNM 421358 MF956831 Nemichthys scolopaceus MOP129-12 USNM 421185 MF956840 Nemichthys scolopaceus MOP130-12 USNM 421180 MF956841 Nemichthys scolopaceus MOP131-12 USNM 421184 MF956842 Nemichthys scolopaceus MOP170-12 USNM 421183 MF956843 Nemichthys scolopaceus MOP197-12 USNM 421191 MF956838 Nemichthys scolopaceus MOP255-12 USNM 421179 MF956830 Nemichthys scolopaceus MOP256-12 USNM 421182 MF956832 Nemichthys scolopaceus MOP374-12 USNM 421186 MF956836 Nemichthys scolopaceus MOP375-12 USNM 421189 MF956835 Nemichthys scolopaceus MOP479-12 USNM 421187 MF956839 Nemichthys scolopaceus MOP562-12 USNM 421181 MF956834 Nemichthys scolopaceus MOP644-12 USNM 421388 MF956837 Neobythites stelliferoides 2 MOP024-12 USNM 422341 MF956846 Neobythites stelliferoides 2 MOP025-12 USNM 422448 MF956847 Neobythites stelliferoides 2 MOP026-12 USNM 422312 MF956848 Neobythites stelliferoides 2 MOP629-12 USNM 421439 MF956849 Neobythites stelliferoides 2 MOP630-12 USNM 421392 MF956844 Neobythites stelliferoides 2 MOP631-12 USNM 421399 MF956845 Nezumia convergens 2 MOP068-12 USNM 422302 MF956856 Nezumia convergens 2 MOP176-12 USNM 422400 MF956850 Nezumia convergens 2 MOP391-12 USNM 421546 MF956854 Nezumia convergens 2 MOP392-12 USNM 422577 MF956855 Nezumia convergens 2 MOP557-12 USNM 421344 MF956851 Nezumia convergens 2 MOP657-12 USNM 421572 MF956852 Nezumia convergens 2 MOP659-12 USNM 421402 MF956853 Nezumia latirostrata 2 MOP573-12 USNM 421481 MF956858 Nezumia latirostrata 2 MOP708-12 USNM 422545 MF956859 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 21TEP DEEP FISHES TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Nezumia latirostrata 2 MOP781-12 USNM 421257 MF956857 Nezumia liolepis 1 MOP138-12 USNM 422421 MF956874 Nezumia liolepis 1 MOP139-12 USNM 422462 MF956871 Nezumia liolepis 1 MOP163-12 USNM 422375 MF956870 Nezumia liolepis 1 MOP164-12 USNM 422383 MF956869 Nezumia liolepis 1 MOP172-12 USNM 422460 MF956865 Nezumia liolepis 1 MOP173-12 USNM 422458 MF956864 Nezumia liolepis 1 MOP394-12 USNM 422590 MF956867 Nezumia liolepis 1 MOP401-12 USNM 421474 MF956872 Nezumia liolepis 1 MOP402-12 USNM 422568 MF956868 Nezumia liolepis 1 MOP411-12 USNM 421476 MF956863 Nezumia liolepis 1 MOP416-12 USNM 421544 MF956866 Nezumia liolepis 1 MOP457-12 USNM 421508 MF956873 Nezumia liolepis 1 MOP458-12 USNM 421440 MF956861 Nezumia liolepis 1 MOP459-12 USNM 422582 MF956862 Nezumia liolepis 1 MOP460-12 USNM 421458 MF956860 Nezumia stelgidolepis 1 MOP165-12 USNM 422368 MF956879 Nezumia stelgidolepis 1 MOP252-12 USNM 421238 MF956875 Nezumia stelgidolepis 1 MOP572-12 USNM 421297 MF956877 Nezumia stelgidolepis 1 MOP703-12 USNM 421490 MF956878 Nezumia stelgidolepis 1 MOP780-12 USNM 421330 MF956876 Notacanthus cf. chemnitzii MOP414-12 USNM 421522 MF956880 Notacanthus cf. chemnitzii MOP589-12 USNM 422600 MF956882 Notacanthus cf. chemnitzii MOP590-12 USNM 421354 MF956881 Notacanthus spinosus 2 MOP112-12 USNM 422417 MF956890 Notacanthus spinosus 2 MOP113-12 USNM 422423 MF956888 Notacanthus spinosus 2 MOP114-12 USNM 422461 MF956889 Notacanthus spinosus 2 MOP246-12 USNM 421321 MF956886 Notacanthus spinosus 2 MOP377-12 USNM 421559 MF956887 Notacanthus spinosus 2 MOP415-12 USNM 421532 MF956883 Notacanthus spinosus 2 MOP525-12 USNM 423204 MF956891 Notacanthus spinosus 2 MOP771-12 USNM 421487 MF956884 Notacanthus spinosus 2 MOP779-12 USNM 422583 MF956885 Oneirodes luetkeni 1 MOP072-12 USNM 422335 MF956896 Oneirodes luetkeni 1 MOP166-12 USNM 422393 MF956895 Oneirodes luetkeni 1 MOP269-12 USNM 421353 MF956897 Oneirodes luetkeni 1 MOP270-12 USNM 421223 MF956892 Oneirodes luetkeni 1 MOP351-12 USNM 422511 MF956894 Oneirodes luetkeni 1 MOP420-12 USNM 421514 MF956893 Oneirodes rosenblatti 2 MOP451-12 USNM 423219 MF956898 Ophichthus frontalis 2 MOP333-12 USNM 422483 MF956899 ......continued on the next pageROBERTSON ET AL .22 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Ophichthus remiger 2 MOP719-12 USNM 421369 MF956901 Ophichthus remiger 2 MOP797-12 USNM 421371 MF956902 Ophichthus remiger 2 MOP798-12 USNM 421370 MF956903 Ophichthus remiger 2 MOP799-12 USNM 421210 MF956900 Ophidiiformes MOP707-12 USNM 440226* MF956904 Ophthalmolycus sp. MOP109-12 USNM 422428 MF956911 Ophthalmolycus sp. MOP110-12 USNM 422452 MF956910 Ophthalmolycus sp. MOP116-12 USNM 422367 MF956905 Ophthalmolycus sp. MOP274-12 USNM 421318 MF956907 Ophthalmolycus sp. MOP275-12 USNM 421307 MF956908 Ophthalmolycus sp. MOP276-12 USNM 421309 MF956909 Ophthalmolycus sp. MOP446-12 USNM 421542 MF956906 Ophthalmolycus sp. MOP526-12 USNM 421473 MF956912 Otophidium indefatigabile 4 MOP491-12 USNM 421519 MF956913 Otophidium indefatigabile 4 MOP540-12 USNM 421302 MF956914 Otophidium indefatigabile 4 MOP541-12 USNM 421299 MF956915 Otophidium indefatigabile 4 MOP712-12 USNM 422512 MF956916 Pachycara sp. MOP194-12 USNM 422432 MF956919 Pachycara sp. MOP592-12 USNM 421417 MF956917 Pachycara sp. MOP593-12 USNM 421562 MF956920 Pachycara sp. MOP724-12 USNM 422599 MF956918 Paraliparis sp. MOP041-12 USNM 422607 MF956922 Paraliparis sp. MOP042-12 USNM 422316 MF956923 Paraliparis sp. MOP061-12 USNM 422466 MF956925 Paraliparis sp. MOP103-12 USNM 422426 MF956927 Paraliparis sp. MOP188-12 USNM 422454 MF956929 Paraliparis sp. MOP189-12 USNM 422455 MF956930 Paraliparis sp. MOP273-12 USNM 421269 MF956928 Paraliparis sp. MOP372-12 USNM 422585 MF956924 Paraliparis sp. MOP373-12 USNM 421548 MF956926 Paraliparis sp. MOP445-12 USNM 422592 MF956921 Peprilus medius 1 MOP220-12 USNM 421322 MF956934 Peprilus medius 1 MOP221-12 USNM 422337 MF956933 Peprilus medius 1 MOP222-12 USNM 422358 MF956932 Peprilus medius 1 MOP794-12 USNM 421325 MF956935 Peprilus medius 1 MOP796-12 USNM 421328 MF956931 Peprilus snyderi 2 MOP016-12 USNM 422315 MF956937 Peprilus snyderi 2 MOP017-12 USNM 422330 MF956938 Peprilus snyderi 2 MOP018-12 USNM 422303 MF956939 Peprilus snyderi 2 MOP795-12 USNM 421333 MF956936 Peristedion barbiger 2 MOP001-12 USNM 422320 MF956941 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 23TEP DEEP FISHES TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Peristedion barbiger 2 MOP002-12 USNM 422314 MF956943 Peristedion barbiger 2 MOP003-12 USNM 422329 MF956945 Peristedion barbiger 2 MOP214-12 USNM 421319 MF956940 Peristedion barbiger 2 MOP215-12 USNM 421211 MF956946 Peristedion barbiger 2 MOP792-12 USNM 421334 MF956944 Peristedion barbiger 2 MOP793-12 USNM 421212 MF956942 Physiculus nematopus 2 MOP330-12 USNM 422543 MF956951 Physiculus nematopus 2 MOP331-12 USNM 422571 MF956950 Physiculus nematopus 2 MOP332-12 USNM 422559 MF956948 Physiculus nematopus 2 MOP533-12 USNM 421305 MF956947 Physiculus nematopus 2 MOP697-12 USNM 421413 MF956949 Platytroctes apus 3 MOP062-12 USNM 422451 MF956953 Platytroctes apus 3 MOP066-12 USNM 422449 MF956955 Platytroctes apus 3 MOP370-12 USNM 422500 MF956954 Platytroctes apus 3 MOP558-12 USNM 421296 MF956952 Platytroctes apus 3 MOP559-12 USNM 422630 MF956956 Pomadasys branickii 1 MOP233-12 USNM 422650 MF956957 Pontinus furcirhinus 1 MOP321-12 USNM 422640 MF956973 Pontinus sierra 2 MOP004-12 USNM 422471 MF956975 Pontinus sierra 2 MOP005-12 USNM 422474 MF956976 Pontinus sierra 2 MOP006-12 USNM 422473 MF956974 Pontinus sierra 2 MOP008-12 USNM 422443 MF956982 Pontinus sierra 2 MOP290-12 USNM 422336 MF956978 Pontinus sierra 2 MOP291-12 USNM 422349 MF956981 Pontinus sierra 2 MOP292-12 USNM 422350 MF956979 Pontinus sierra 2 MOP318-12 USNM 422611 MF956980 Pontinus sierra 2 MOP319-12 USNM 422580 MF956984 Pontinus sierra 2 MOP320-12 USNM 421552 MF956977 Pontinus sierra 2 MOP622-12 USNM 421376 MF956983 Pontinus sp. MOP007-12 USNM 422440 MF956972 Pontinus sp. MOP009-12 USNM 422306 MF956971 Pontinus sp. MOP010-12 USNM 422327 MF956970 Pontinus sp. MOP011-12 USNM 422309 MF956969 Pontinus sp. MOP140-12 USNM 422382 MF956968 Pontinus sp. MOP147-12 USNM 422389 MF956967 Pontinus sp. MOP149-12 USNM 422369 MF956965 Pontinus sp. MOP229-12 USNM 421247 MF956961 Pontinus sp. MOP230-12 USNM 421323 MF956960 Pontinus sp. MOP231-12 USNM 422340 MF956958 Pontinus sp. MOP232-12 USNM 421270 MF956959 Pontinus sp. MOP667-12 USNM 421398 MF956962 ......continued on the next pageROBERTSON ET AL .24 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Pontinus sp. MOP668-12 USNM 421430 MF956963 Pontinus sp. MOP669-12 USNM 421384 MF956964 Pontinus sp. MOP148-12 USNM 422414 MF956966 Porogadus sp. 3 MOP438-12 USNM 421448 MF956987 Porogadus sp. 3 MOP512-12 USNM 435796 MF956986 Porogadus sp. 3 MOP599-12 USNM 421549 MF956985 Poromitra crassiceps MOP070-12 USNM 422334 MF956988 Poromitra crassiceps MOP071-12 USNM 422301 MF956989 Prionotus ruscarius 1 MOP216-12 USNM 422300 MF956993 Prionotus ruscarius 1 MOP282-12 USNM 421224 MF956991 Prionotus ruscarius 1 MOP283-12 USNM 421264 MF956990 Prionotus ruscarius 1 MOP791-12 USNM 421213 MF956992 Prionotus stephanophrys MOP211-12 USNM 421216 MF956994 Prionotus stephanophrys MOP212-12 USNM 422347 MF956997 Prionotus stephanophrys MOP213-12 USNM 421234 MF956996 Prionotus stephanophrys MOP807-12 USNM 421215 MF956995 Prionotus stephanophrys MOP809-12 USNM 421243 MF956998 Psenes cyanophrys MOP700-12 USNM 421567 MF956999 Psenes sio 2 MOP036-12 USNM 422472 MF957013 Psenes sio 2 MOP132-12 USNM 422378 MF957008 Psenes sio 2 MOP133-12 USNM 422411 MF957006 Psenes sio 2 MOP134-12 USNM 422377 MF957005 Psenes sio 2 MOP200-12 USNM 422373 MF957001 Psenes sio 2 MOP386-12 USNM 422526 MF957007 Psenes sio 2 MOP387-12 USNM 422547 MF957002 Psenes sio 2 MOP588-12 USNM 421342 MF957000 Psenes sio 2 MOP646-12 USNM 421375 MF957009 Psenes sio 2 MOP647-12 USNM 421382 MF957004 Psenes sio 2 MOP648-12 USNM 421410 MF957003 Psenes sio 2 MOP654-12 USNM 421570 MF957010 Psenes sio 2 MOP655-12 USNM 421434 MF957011 Psenes sio 2 MOP656-12 USNM 421405 MF957012 Pseudoscopelus lavenbergi 4 MOP369-12 USNM 422507 MF957014 Psychrolutes sio 2 MOP271-12 USNM 422353 MF957015 Psychrolutes sio 2 MOP371-12 USNM 422651 MF957016 Psychrolutes sio 2 MOP410-12 USNM 422603 MF957017 Psychrolutes sio 2 MOP504-12 USNM 421511 MF957018 Psychrolutes sio 2 MOP587-12 USNM 423597 MF957019 Rouleina attrita 3 MOP059-12 USNM 422434 MF957020 Scopelarchoides nicholsi 3 MOP365-12 USNM 422520 MF957021 Scopelarchoides nicholsi 3 MOP682-12 USNM 421401 MF957022 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 25TEP DEEP FISHES TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Scopelarchoides nicholsi 3 MOP725-12 USNM 422485 MF957023 Scopelengys tristis 3 MOP142-12 USNM 435799 MF957025 Scopelengys tristis 3 MOP169-12 USNM 435797 MF957024 Scopelengys tristis 3 MOP463-12 USNM 421443 MF957028 Scopelengys tristis 3 MOP464-12 USNM 421512 MF957029 Scopelengys tristis 3 MOP465-12 USNM 421447 MF957026 Scopelengys tristis 3 MOP642-12 USNM 421425 MF957027 Sebastolobus altivelis MOP452-12 USNM 423201 MF957030 Selene brevoortii MOP146-12 USNM 422365 MF957031 Selene brevoortii MOP663-12 USNM 421378 MF957032 Selene peruviana 1 MOP145-12 USNM 422399 MF957034 Selene peruviana 1 MOP679-12 USNM 421404 MF957033 Serranus aequidens 1 MOP287-12 USNM 421248 MF957038 Serranus aequidens 1 MOP288-12 USNM 421277 MF957035 Serranus aequidens 1 MOP289-12 USNM 422346 MF957037 Serranus aequidens 1 MOP308-12 USNM 422566 MF957036 Serrivomer sector MOP032-12 USNM 421364 MF957046 Serrivomer sector MOP190-12 USNM 421363 MF957041 Serrivomer sector MOP192-12 USNM 421198 MF957040 Serrivomer sector MOP360-12 USNM 421199 MF957039 Serrivomer sector MOP436-12 USNM 421197 MF957043 Serrivomer sector MOP480-12 USNM 421200 MF957045 Serrivomer sector MOP563-12 USNM 421287 MF957042 Serrivomer sector MOP564-12 USNM 421201 MF957044 Serrivomer sector MOP608-12 USNM 421336 MF957047 Sternoptyx diaphana MOP352-12 USNM 422506 MF957052 Sternoptyx diaphana MOP450-12 USNM 421510 MF957049 Sternoptyx diaphana MOP701-12 USNM 422505 MF957048 Sternoptyx diaphana MOP702-12 USNM 422542 MF957051 Sternoptyx diaphana MOP815-12 USNM 421254 MF957050 Stomias atriventer MOP073-12 USNM 435793 MF957059 Stomias atriventer MOP100-12 USNM 435787 MF957058 Stomias atriventer MOP158-12 USNM 435792 MF957054 Stomias atriventer MOP242-12 USNM 435789 MF957057 Stomias atriventer MOP366-12 USNM 421554 MF957063 Stomias atriventer MOP461-12 USNM 421535 MF957053 Stomias atriventer MOP462-12 USNM 421527 MF957064 Stomias atriventer MOP508-12 USNM 435790 MF957061 Stomias atriventer MOP509-12 USNM 435788 MF957060 Stomias atriventer MOP579-12 USNM 421465 MF957056 Stomias atriventer MOP580-12 USNM 422629 MF957055 ......continued on the next pageROBERTSON ET AL .26 · Zootaxa 4348 (1) © 2017 Magnolia Press TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Stomias atriventer MOP638-12 USNM 435798 MF957065 Stomias atriventer MOP769-12 USNM 422614 MF957062 Symphurus leei 2 MOP500-12 USNM 421454 MF957070 Symphurus leei 2 MOP501-12 USNM 421507 MF957069 Symphurus leei 2 MOP626-12 USNM 421245 MF957067 Symphurus leei 2 MOP627-12 USNM 421442 MF957066 Symphurus leei 2 MOP628-12 USNM 421418 MF957068 Synchiropus atrilabiatus 1 MOP311-12 USNM 422564 MF957072 Synchiropus atrilabiatus 1 MOP312-12 USNM 422551 MF957071 Synodus evermanni 2 MOP208-12 USNM 422338 MF957076 Synodus evermanni 2 MOP209-12 USNM 421246 MF957074 Synodus evermanni 2 MOP210-12 USNM 421320 MF957075 Synodus evermanni 2 MOP788-12 USNM 421250 MF957073 Synodus evermanni 2 MOP790-12 USNM 421329 MF957077 Talismania bifurcata MOP606-12 USNM 421553 MF957078 Trichiurus nitens 2 MOP049-12 USNM 422445 MF957079 Trichiurus nitens 2 MOP050-12 USNM 422317 MF957086 Trichiurus nitens 2 MOP051-12 USNM 422326 MF957087 Trichiurus nitens 2 MOP153-12 USNM 422372 MF957080 Trichiurus nitens 2 MOP154-12 USNM 422380 MF957081 Trichiurus nitens 2 MOP155-12 USNM 422376 MF957082 Trichiurus nitens 2 MOP623-12 USNM 421433 MF957083 Trichiurus nitens 2 MOP624-12 USNM 421407 MF957084 Trichiurus nitens 2 MOP625-12 USNM 421385 MF957085 Umbrina bussingi 1 MOP530-12 USNM 422631 MF957090 Umbrina bussingi 1 MOP531-12 USNM 421289 MF957088 Umbrina bussingi 1 MOP532-12 USNM 421301 MF957089 Venefica sp. 3 MOP435-12 USNM 421203 MF957091 Venefica tentaculata 4 MOP081-12 USNM 421365 MF957092 Venefica tentaculata 4 MOP082-12 USNM 421366 MF957098 Venefica tentaculata 4 MOP191-12 USNM 421367 MF957096 Venefica tentaculata 4 MOP353-12 USNM 423194 MF957099 Venefica tentaculata 4 MOP354-12 USNM 421202 MF957097 Venefica tentaculata 4 MOP594-12 USNM 422623 MF957093 Venefica tentaculata 4 MOP595-12 USNM 421204 MF957094 Venefica tentaculata 4 MOP675-12 USNM 421368 MF957095 Xenomystax atrarius 4 MOP244-12 USNM 421360 MF957105 Xenomystax atrarius 4 MOP262-12 USNM 421361 MF957100 Xenomystax atrarius 4 MOP340-12 USNM 421206 MF957102 Xenomystax atrarius 4 MOP341-12 USNM 421207 MF957101 Xenomystax atrarius 4 MOP567-12 USNM 421359 MF957103 ......continued on the next page Zootaxa 4348 (1) © 2017 Magnolia Press · 27TEP DEEP FISHES Annotated and illustrated catalog NOTACANTHIFORMES 2 F, 2 G, 4 S. Halosauridae Halosaurs (En); Halosauro (Sp). 1 G, 2 S. (1) Halosaurus attenuatus Garman, 1899. Attenuated halosaur (En); Halosauro atenuado (Sp) MOP stations. 2; 028 and 029; Depth range: 1113–1472 m; USNM voucher specimens: 4; 421258, 421271, 421547 and 422539. Distribution. Eastern Pacific (Gulf of California, Gulf of Panama and Galapagos Islands) and eastern Atlantic (Gulf of Guinea), occurring at depths between 1113 and 2500 m (see Remarks). Literature. Garman (1899: 296), McDowell (1973: 57), Sulak (1977a: 11; 1990: 130; 1995: 1174), Castro- Aguirre & Balart (1996: 69) and McCosker & Rosenblatt (2010: 188). Remarks. These specimens represent the first documented record of the species in lower Central American waters (Sulak 1977a, Bussing & López 1994, 2009, 2011). Previous records of this species in eastern Pacific waters were from the Gulf of California (Castro-Aguirre & Balart 1996) and the Galápagos Islands (Garman 1899, McCosker & Rosenblatt 2010). In addition, specimens collected at 1113 m represent a new upper limit to its known depth range (Sulak 1977a; Froese & Pauly 2016). FIGURE 3. Halosaurus attenuatus. Top: entire specimen; Bottom: detail of head; USNM 421271, 280 mm. TL. TABLE 3. (Continued) Status BOLD Process ID NMNH Voucher Number Genbank Accession No. Xenomystax atrarius 4 MOP568-12 USNM 421205 MF957104 Yarrella argenteola 4 MOP481-12 USNM 421493 MF957109 Yarrella argenteola 4 MOP482-12 USNM 421494 MF957108 Yarrella argenteola 4 MOP652-12 USNM 421422 MF957107 Yarrella argenteola 4 MOP653-12 USNM 421423 MF957106 Zalieutes elater 3 MOP201-12 USNM 422354 MF957111 Zalieutes elater 3 MOP202-12 USNM 421227 MF957112 Zalieutes elater 3 MOP294-12 USNM 422352 MF957113 Zalieutes elater 3 MOP322-12 USNM 422510 MF957114 Zalieutes elater 3 MOP323-12 USNM 422502 MF957110ROBERTSON ET AL .28 · Zootaxa 4348 (1) © 2017 Magnolia Press (2) Halosaurus radiatus Garman, 1899. Eastern Pacific halosaur (En); Halosauro del Pacífico oriental, Anguila radiante (Sp) MOP stations. 2; 004 and 075; Depth range: 115–844 m; USNM voucher specimens: 6; 421275, 421310, 421311, 421379, 421477 and 421516. FIGURE 4. Halosaurus radiatus. Top: Entire specimen; Bottom Detail of head; USNM 421379, 250 mm. TL. Distribution. Eastern Pacific (from Costa Rica to Chile), occurring at depths between 115 and 935 m (see Remarks). Literature. Garman (1899: 298), Bussing (1965: 212), McDowell (1973: 56), Pequeño & Revuelta (1980: 892), Pequeño (1989: 27), Chirichigno & Vélez (1998: 172), Pequeño (2000: 62), and Yabe (2009a: 74). Remarks. USNM 421275, 421310 and 421311 specimens represent the first documented record of the species in Costa Rican waters (Bussing & López 1994, 2009, 2011), and a northern extension of about 800 km in this species known distribution (Garman 1899, Pequeño 1989, Chirichigno & Vélez 1998, Yabe 2009a). The northernmost documented record for this species in eastern Pacific waters was previously in the Gulf of Panama (Garman 1899). In addition, specimens collected at 115 m represent a new maximum depth record for the species (Garman 1899, Bussing 1965, Yabe 2009a, Froese & Pauly 2016). Notacanthidae Spiny eels (En); Anguilas espinosas (Sp). 1 G, 2 S. (3) Notacanthus spinosus Garman, 1899. Eastern Pacific spiny eel (En); Anguila espinosa del Pacífico Oriental (Sp) MOP stations. 5; 003, 029, 078, 083, and 094; Depth range: 116–1527 m; USNM voucher specimens: 9; 421321, 421532, 421559, 422417, 422423, 422461, 422532, 423204 and 435801. Distribution. Eastern Pacific (from El Salvador to Peru), occurring at depths between 116 and 1527 m (see Remarks). Literature. Garman (1899: 301), McDowell (1973: 190), Chirichigno & Vélez (1998: 172), Yabe (2009b: 75), Mundy et al. (2011: 269) and Kanehira et al. (2012: 2). Remarks. USNM 421354, 421522, 421532, 422417, 422423, 422461, 422600 and 423204 specimens represent the first documented record of the species in upper Central American waters (Bussing & López 1994, 2009, 2011; Yabe 2009b), and a northern range extension of about 1400 km in the species’ known distribution (Garman 1899, Chirichigno & Vélez 1998, Yabe 2009b). Previously the northernmost documented record for this species in eastern Pacific waters was in the Gulf of Panama (Garman 1899). In addition, specimens collected at 1527 m represent a new minimum depth record for the species (Yabe 2009b, Froese & Pauly 2016). Zootaxa 4348 (1) © 2017 Magnolia Press · 29TEP DEEP FISHES FIGURE 5. Notacanthus spinosus. Top: entire specimen; Bottom: detail of head. (4) Notacanthus cf. chemnitzii Bloch 1788. Snubnosed spiny eel (En). MOP stations. 2; 082 and 087; Depth range: 1100–1527 m; USNM voucher specimens: 3; 421354, 421522 and 422600. Distribution. Circumglobal, occurring at depths between 125 and 3285 m. In the eastern Pacific this species has been previously recorded in Canada, the Gulf of California, Mexico, and Chile Literature. Bloch (1788: 278), McDowell (1973: 191), Nakamura et al. (1986: 62), Whitehead et al. (1986: 600), Lea & Rosenblatt (1987: 51), Matarese et al. (1989: 34), Paxton et al. (1989: 150), Pequeño (1989: 27), McAllister (1990: 45), McEachran & Fechhelm (1998: 217), Paepke (1999: 106), Trunov (1999: 461), Mecklenburg et al. (2002: 119), Hutchins (2001: 18), Moore et al. (2003: 172), Mundy et al. (2011: 269) and Parin et al. (2014: 46). Remarks. These specimens represent the first documented record of the species in Central American waters (Lea & Rosenblatt 1987, Pequeño 1989, Bussing & López 1994, 2009, 2011). However, Mundy et al. (2011) and David G. Smith (pers. comm.) have indicated that N. chemnitzii likely represents a species complex in need of further investigation. The type locality is Iceland. FIGURE 6. Notacanthus cf. chemnitzii. Top: entire specimen; Bottom: detail of head; USNM 421532, 255 mm. TL. ANGUILLIFORMES 7 F, 9 G, 11 S. Muraenidae Moray eels (En); Anguilas morena (Sp). 1 G, 2 S.ROBERTSON ET AL .30 · Zootaxa 4348 (1) © 2017 Magnolia Press (5) Gymnothorax equatorialis (Hildebrand, 1946). Equatorial moray, Spotted-tail moray, Spotted moray (En); Morena cola pintada, Morena del Ecuador (Sp) FIGURE 7. Gymnothorax equatorialis; USNM 421373, 435 mm. TL. MOP stations. 1; 036; Depth range: 138–143 m; USNM voucher specimens: 1; 421373. Distribution. Eastern Pacific (from the Gulf of California to Peru), occurring at depths between 5 and 143 m. Literature. Hildebrand (1946: 134), Allen & Robertson (1994: 46), Bussing & López (1994: 40; 2009: 443), Smith (1994: 23), McCosker & Rosenblatt (1995: 1310), Charter & Moser (1996: 88), De La Cruz Agüero et al. (1997: 226), Chirichigno & Vélez (1998: 130), Bearez (1996: 733), Bussing (1998: 444), Thomson et al. (2000: 44), Böhlke (2001: 3), Böhlke & Smith (2002: 108), Rodríguez-Romero et al. (2008: 1770), Smith (2012: 14), Del Moral-Flores et al. (2013: 190), Gómez et al. (2014: 392), Fuentes et al. (2015: 614) and Robertson & Allen (2015: 338). (6) Gymnothorax phalarus Bussing, 1998. Small-spotted moray (En); Morena cola pintada falsa (Sp) FIGURE 8. Gymnothorax phalarus; USNM 421374, 230 mm. TL. MOP stations. 1; 003; Depth range: 116–118 m; USNM voucher specimens: 1; 421374. Distribution. Eastern Pacific (from the mouth of the Gulf of California to Peru), occurring at depths between 23 and 339 m. Zootaxa 4348 (1) © 2017 Magnolia Press · 31TEP DEEP FISHES Literature. Bussing (1998: 439; 2009: 443), Böhlke (2001: 5), Böhlke & Smith (2002: 137), McCosker (2009a: 76), Smith (2012: 23), Gómez et al. (2014: 394), Fuentes et al. (2015: 611) and Robertson & Allen (2015: 2552). Congridae Conger eels, Garden eels (En); Congrios, Anguilas de jardín (Sp). 2 G, 2 S. (7) Japonoconger proriger (Gilbert, 1891). Slope conger (En); Congrio del talud (Sp) FIGURE 9. Japonoconger proriger. Top: Entire specimen; Bottom: Detail of head. MOP stations. 5; 003, 031, 059, 060 and 075; Depth range: 118–920 m; USNM voucher specimens: 8; 421208, 421209, 421362, 422480, 422481, 422482, 423212, and 423213. Distribution. Eastern Pacific (from the Gulf of Tehuantepec to Peru), occurring at depths between 55 and 920 m (see Remarks). Literature. Gilbert (1891a: 350), Peden (1972:1), Cervigón et al. (1980: 317), Raju (1985: 1), Bussing & López (1994: 38; 2009: 444), Smith (1994: 6, 1995: 1026), Grove & Lavenberg (1997: 180), Chirichigno & Vélez (1998: 131), McCosker (2009e: 84), McCosker & Rosenblatt (2010: 189) and Robertson & Allen (2015: 446). Remarks. Valid as Ariosoma prorigerum (Gilbert, 1891) in Eschmeyer et al. (2017), but provisionally placed in Japonoconger Asano, 1958, based on a personal communication of David G. Smith to DRR in 2016, until a detailed taxonomic study of the genus is carried out. Specimens collected at 920 m represent a new minimum depth record for the species (Robertson & Allen 2015, Froese & Pauly 2016). (8) Xenomystax atrarius Gilbert, 1891. Deepwater conger (En); Congrio de aguas profundas, Congrio de altura (Sp) FIGURE 10. Xenomystax atrarius. Top: Entire specimen; Bottom: Detail of head. MOP stations. 3; 003, 060 and 075; Depth range: 116–844 m; USNM voucher specimens: 6; 421205, 421206, 421207, 421359, 421360 and 421361.ROBERTSON ET AL .32 · Zootaxa 4348 (1) © 2017 Magnolia Press Distribution. Eastern Pacific (from British Columbia, Canada, to Chile), occurring at depths between 116 and 1050 m (see Remarks). Literature. Gilbert (1891a: 348), Bussing (1965: 211), Peden (1972: 1), De Lachica-Bonilla & Castro-Aguirre (1973: 1), Seigel (1987: 57), Pequeño (1989: 25), Smith (1989a: 559, 1994: 12, 1995: 1026), McAllister (1990: 49), Castro-Aguirre & Balart (1996: 69), Charter (1996a: 100), Sielfeld & Vargas (1996: 4), Bussing & López (2009: 444), McCosker (2009e: 86), Del Moral-Flores et al. (2013: 191), Hanke & Roias (2013: 210) and Hanke et al. (2014: 63). Remarks. Specimens collected at 116 m represent a new maximum depth record for the species (McCosker 2009, Froese & Pauly 2016). Synaphobranchidae Cutthroat eels, Deep sea eels (En); Anguilas garganta cortada, Anguilas branquias bajas, Anguilas de aguas profundas (Sp). 1 G, 1 S. (9) Ilyophis brunneus Gilbert, 1891. Muddy arrowtooth eel (En); Anguila fangosa dientes de flecha (Sp) MOP stations. 1; 025; Depth range: 1101–1113 m; USNM voucher specimens: 1; 421372. Distribution. Circumtropical, except the northeast Pacific, occurring at depths between 450 and 3120 m. In the eastern Pacific this species has been previously recorded only in the Galápagos Islands and Peru (see Remarks). FIGURE 11. Ilyophis brunneus. Top: entire specimen; Bottom: detail of head; USNM 421372, 420 mm. TL Literature. Gilbert (1891a: 352), Blache et al. (1973: 253), Masuda et al. (1984: 26), Okamura & Kitajima (1984: 83), Okamura & Machida (1987: 109), Scott & Scott (1988: 88), Robins & Robins (1989: 235), Paulin et al. Zootaxa 4348 (1) © 2017 Magnolia Press · 33TEP DEEP FISHES (1989: 71), McAllister (1990: 47), Smith & Castle (1990: 195), Smith (1994: 37, 1999a: 1661, 2003: 723), Shinohara et al. (1996: 157), Arruda (1997: 59), Sulak & Shcherbachev (1997: 1161), McEachran & Fechhelm (1998: 247), Chen & Mok (1999: 60), Karmovskaya & Parin (1999: 316), Nakabo (2000: 212; 2002: 212), Chen & Mok (2001: 79), Shinohara et al. (2001: 292), Moore et al. (2003: 173), Shinohara et al. (2005: 400), Melo (2007: 315), McCosker (2009b: 77) and Melo et al. (2009: 13). Remarks. This specimen represents the first documented record of the genus in lower Central American waters (Sulak & Shcherbachev 1997; Bussing & López 1994, 2009, 2011; McCosker 2009b) and a northeastern range extension of about 1400 km in the species’ known distribution (Gilbert 1891a, Sulak & Shcherbachev 1997, McCosker 2009b). The previous northernmost record for this genus in eastern Pacific waters, represented by the species I. brunneus and I. arx Robins, 1976, was in the Galápagos Islands (Gilbert 1891, Sulak & Shcherbachev 1997). Ophichthidae Snake eels, Finless eels, Ridgefin eels, Worm eels, Sailfin eels, Sand eels, Spoon-nose eels, Viper eels (En); Tiesos, Anguilas Culebra, Serpentones (Sp). 1 G, 2 S. (10) Ophichthus frontalis Garman, 1899. Deathbanded snake-eel (En); Tieso funebre, Tieso mortuorio (Sp) MOP stations. 1; 004; Depth range: 115–116 m; USNM voucher specimens: 1; 422483. Distribution. Eastern Pacific (two allopatric populations: (1) central Gulf of California and (2) from Costa Rica to Ecuador), occurring at depths between 35 and 760 m. Literature. Garman (1899: 309), McCosker (1977: 81), Smith (1994: 34), McCosker & Rosenblatt (1995: 1338), McCosker & Rosenblatt (1998: 404), Chirichigno & Vélez (1998: 124), Tang & Zhang (2002: 855), Bussing & López (2009: 444), Del Moral-Flores et al. (2013: 190), Gómez et al. (2014: 392), Murase et al. (2014: 1403) and Robertson & Allen (2015: 403). FIGURE 12. Ophichthys frontalis. Top: entire specimen; Bottom: detail of head; USNM 422483, 400 mm. TL.ROBERTSON ET AL .34 · Zootaxa 4348 (1) © 2017 Magnolia Press (11) Ophichthus remiger (Valenciennes, 1837). Punctuated snake-eel, Common snake eel (En); Tieso punteado, Tieso común, Anguila punto blanco (Sp) MOP stations. 3; 004, 007 and 081; Depth range: 115–478 m; USNM voucher specimens: 5; 421210, 421369, 421370, 421371 and 421571. FIGURE 13. Ophichthus remiger. Top: entire specimen; Bottom: detail of head. Distribution. Eastern Pacific (from Nicaragua to Chile), occurring at depths between 0 and 478 m (see Remarks). Literature. McCosker (1977: 81), Pequeño (1989: 25), Bauchot et al. (1993: 111), McCosker & Rosenblatt (1995: 1338), Bearez (1996: 733), Chirichigno & Vélez (1998: 120), Tang & Zhang (2002: 855), Bussing & López (2009: 445), McCosker (2009c: 78), Murase et al (2014: 1403), Fuentes et al. (2015: 614) and Robertson & Allen (2015: 405). Remarks. Specimens collected at 478 m represent a new minimum depth record for the species (Bussing & López 2009, McCosker 2009c, Froese & Pauly 2016). Nemichthyidae Snipe eels (En); Anguilas tijera (Sp). 2 G, 2 S. (12) Avocettina bowersii (Garman, 1899). Smalleye snipe eel (En); Anguila tijera de ojo pequeño (Sp) FIGURE 14. Avocettina bowersii.Top: entire specimen; Bottom: detail of head; USNM 421290, 420 mm. TL. Zootaxa 4348 (1) © 2017 Magnolia Press · 35TEP DEEP FISHES MOP stations. 4; 067, 076, 078 and 083; Depth range: 926–1145 m; USNM voucher specimens: 6; 421192, 421193, 421194, 421195, 421196 and 421290. Distribution. Eastern Pacific (from California, United States of America (USA), to Chile), occurring at depths between 0 and 4000 m. Literature. Garman (1899: 323), Robison (1972: 451), Pequeño (1989: 25), Smith & Nielsen (1989: 448), Castro-Aguirre & Balart (1996: 69), Charter (1996b: 122), De La Cruz Agüero et al. (1997: 239), Chirichigno & Vélez (1998: 126), Mecklenburg et al. (2002: 125), Rodríguez-Romero et al. (2008: 1770), McCosker (2009d: 80), Mundy (2005: 137), Reyes-Bonilla et al. (2011: 6), Hanke & Roias (2013: 210) and Cruz-Mena & Angulo (2016: 1). (13) Nemichthys scolopaceus Richardson, 1848. Slender snipe eel (En); Anguila tijera esbelta (Sp) MOP stations. 8; 001, 029, 065, 067, 071, 075, 078 and 088; Depth range: 165–1310 m; USNM voucher specimens: 15; 421179, 421180, 421181, 421182, 421183, 421184, 421185, 421186, 421187, 421189, 421190, 421191, 421357, 421358 and 421388. Distribution. Worldwide, occurring at depths between 0 and 2500 m. In the eastern Pacific this species has been recorded from Alaska, USA, to Chile. Literature. Richardson (1848: 25), Bussing (1965: 212), Fitch & Lavenberg (1968: 62), Robison (1972: 451), Larsen (1973: 231), Uyeno et al. (1983: 102), Masuda et al. (1984: 22), Nielsen (1986: 553), Pequeño (1989: 25), Smith & Nielsen (1989: 454), McAllister (1990: 47), Bossé (1991: 115), Bearez (1996: 733), Castro-Aguirre & Balart (1996: 69), Charter (1996b: 122), Shinohara et al. (1996: 158), De La Cruz Agüero et al. (1997: 239), Nakabo (2000: 237; 2002: 237), Hutchins (2001: 18), Mecklenburg et al. (2002: 126), Moore et al. (2003: 174), Smith (2003: 742), Mundy (2005: 138), Shinohara et al. (2005: 401), McCosker (2009d: 81), Reyes-Bonilla et al. (2011: 6), Hanke & Roias (2013: 210), Larson et al. (2013: 29), Hanke et al. (2014: 63), Parin et al. (2014: 49), Wirtz et al. (2014: 3) and Cruz-Mena & Angulo (2016: 1). FIGURE 15. Nemichthys scolopaceus. Top: entire specimen; Bottom: detail of head. Serrivomeridae Sawtooth eels (En); Anguilas dientes de sierra (Sp). 1 G, 1 S. (14) Serrivomer sector Garman, 1899. Sawtooth eel (En); Anguila dientes de sierra (Sp) MOP stations. 6; 030, 044, 067, 073, 077 and 086; Depth range: 153–1389 m; USNM voucher specimens: 9; 421197, 421198, 421199, 421200, 421201, 421287, 421336, 421363 and 421364. Distribution. Indo-west (Seychelles, Japan, and New Caledonia), central (Hawaiian Islands) and eastern Pacific (from San Francisco, California, USA, to Chile), occurring at depths between 0 and 3243 m. Literature. Garman (1899: 320), Bauchot (1959: 134), Bussing (1965: 212), Robison (1972: 448), Pequeño ROBERTSON ET AL .36 · Zootaxa 4348 (1) © 2017 Magnolia Press (1989: 25), Smith (1994: 37, 1999b: 1692), Castro-Aguirre & Balart (1996: 69), Charter (1996c: 131), Shinohara et al. (1996: 158), Nakabo (2000: 238; 2002: 235), Mundy (2005: 144), Shao et al. (2008: 239), McCosker (2009g: 88) Reyes-Bonilla et al. (2011: 6) and Fricke et al. (2011: 354). FIGURE 16. Serrivomer sector. Top: entire specimen; Bottom: detail of head; USNM 421199, 495 mm. TL. Nettastomatidae Duckbill eels (En); Anguilas pico de pato, Serpentinas (Sp). 1 G, 1 S. (15) Venefica tentaculata Garman, 1899. Duckbill eel (En); Anguila pico de pato (Sp) MOP stations. 5; 001, 002, 046, 082 and 086; Depth range: 183–1527 m; USNM voucher specimens: 9; 421202, 421203, 421204, 421365, 421366, 421367, 421368, 422623 and 423194. Distribution. Western (off Honshu and the sea of Okhotsk, Japan) and eastern (from British Columbia, Canada, to Peru) Pacific, occurring at depths between 183 and 1790 m (see Remarks). Literature. Garman (1899: 319), Amaoka et al. (1983: 67), Masuda et al. (1984), Smith (1989b: 610), Charter (1996d: 135), Shinohara et al. (1996: 158), Nakabo (2000: 239), Mundy (2005: 144), McCosker (2009f: 88), Hanke & Roias (2013: 210), Hanke et al. (2014: 63) and Parin et al. (2014: 50). Remarks. Specimens collected at 183 m represent a new maximum depth record for the species (Masuda et al. 1984, McCosker 2009f, and Froese & Pauly 2016). FIGURE 17. Venefica tentaculata. Top: entire specimen; Bottom: detail of head; USNM 421366, 630 mm. TL. OSMERIFORMES 4 F, 10 G, 10 S. Argentinidae Argentines, Herring smelts (En); Argentinas, Pejerreyes (Sp). 1 G, 1 S. Zootaxa 4348 (1) © 2017 Magnolia Press · 37TEP DEEP FISHES (16) Argentina aliceae Cohen & Atsaides, 1969. Alice's argentine (En); Argentina de Alicia (Sp) MOP stations. 4; 003, 005, 006 and 048; Depth range: 109–500 m; USNM voucher specimens: 9; 421222, 421411, 421424, 421438, 422384, 422392, 422405, 422548 and 422575. Distribution. Eastern Pacific (from Nicaragua to Peru), occurring at depths between 71 and 500 m (see Remarks). FIGURE 18. Argentina aliceae. Top: entire specimen; Bottom: detail of head. Literature. Cohen & Atsaides (1969: 19), Nielsen (1974: 12), Böhlke (1984: 22), Ibarra & Stewart (1987: 11), Bussing & López (1994: 54, 2009: 446), Schneider (1995: 858), Bearez (1996: 734), Castro-Aguirre & Balart (1996: 69), Chirichigno & Vélez (1998: 168), Kobyliansky (2004: 607), Bussing & López (2009: 446), Shinohara (2009: 94), McCosker & Rosenblatt (2010: 189) and Robertson & Allen (2015: 599). Remarks. Specimens collected at 500 m represent a new minimum depth record for the species (Cohen & Atsaides 1969, Bussing & López 2009, Froese & Pauly 2016, Robertson & Allen 2015). Bathylagidae Deep-sea smelts (En); Esperlanes, Capellanes (Sp). 1 G, 1 S. (17) Leuroglossus urotranus Bussing, 1965. Tropical eastern Pacific smoothtongue (En); Lengualisa del Pacífico tropical oriental (Sp) MOP stations. 1; 003; Depth range: 105–107 m; USNM voucher specimens: 1; 421262. Distribution. Eastern Pacific (from Panama to Chile), occurring at depths between 105 and 2830 m (see Remarks). Literature. Bussing (1965: 194) and Pequeño (1989: 29). Remarks. This specimen represents the first documented record of the species in lower Central American waters (Bussing 1965, Bussing & López 1994, 2009, 2011) and a northern range extension of about 1200 km in the species’ known distribution (Bussing 1965). The previous northernmost documented record for this species in eastern Pacific waters was in the Gulf of Guayaquil (Bussing 1965). In addition, this specimen represents a new upper depth record for the species (Bussing 1965). Alepocephalidae Slickheads (En); Cabezas lisas (Sp). 6 G, 6 S. (18) Alepocephalus tenebrosus Gilbert, 1892. California slickhead (En); Cabeza lisa californiana (Sp)ROBERTSON ET AL .38 · Zootaxa 4348 (1) © 2017 Magnolia Press MOP stations. 14; 004, 025, 026, 029, 034, 059, 064, 065, 073, 077, 078, 083, 086 and 087; Depth range: 115– 1389 m; USNM voucher specimens: 18; 421242, 421280, 421282, 421451, 421471, 421518, 421555, 421560, 421573, 422396, 422398, 422463, 422529, 422556, 422612, 422616, 422619 and 422642. Distribution. North and eastern Pacific (from the Bering Sea to Chile), occurring at depths between 46 and 5500 m. FIGURE 19. Alepocephalus tenebrosus. Top: entire specimen; Bottom: detail of head. Literature. Gilbert (1892: 545), Böhlke (1953: 13), Fitch & Lavenberg (1968: 17), Eschmeyer et al. (1983: 89), Matarese et al. (1989: 76), Sazonov et al. (1993: 43), Ambrose (1996a: 224), Castro-Aguirre & Balart (1996: 69), Mecklenburg et al. (2002: 167), Grinols, & Heyamoto (2011: 1151), Reyes-Bonilla et al. (2011: 6) and Maslenikov et al. (2013: 6). Remarks. These specimens represent the first documented record of the species in Central American waters (Eschmeyer et al. 1983; Bussing & López 1994, 2009, 2011; Ambrose 1996; Froese & Pauly 2016). (19) Bajacalifornia burragei Townsend & Nichols, 1925. Sharpchin slickhead (En); Cabeza lisa de barbilla afilada (Sp) MOP stations. 2; 077 and 094; Depth range: 1320–1406m; USNM voucher specimens: 2; 421485 and 422437. Distribution. Western Indian (Red Sea) and western (South China Sea) and eastern Pacific (from the Gulf of California to Chile), occurring at depths between 865 and 1406 m (see Remarks). Literature. Townsend & Nichols (1925: 8), Fowler (1956: 22), Robison (1972: 451), Markle & Krefft (1985: 348), Pequeño (1989: 33), Miya & Markle (1993: 744), Ambrose (1996a: 224), Castro-Aguirre & Balart (1996: 69), Chirichigno & Vélez (1998: 180), Yeh et al. (2006: 265), Sazonov & Williams (2001: S11) and Reyes-Bonilla et al. (2011: 6). Remarks. These specimens represent the first documented record of the species in Central American waters (Fowler 1956; Bussing & López 1994, 2009, 2011; Froese & Pauly 2016) and a new maximum depth record for the species (Townsend & Nichols 1925, Fowler 1956, Froese & Pauly 2016). Zootaxa 4348 (1) © 2017 Magnolia Press · 39TEP DEEP FISHES FIGURE 20. Bajacalifornia burragei. Top: entire specimen; Bottom: detail of head; USNM 422437, 170 mm.SL. (20) Bathytroctes microlepis Günther, 1878. Smallscale smooth-head (En); Cabeza lisa de escama pequeña (Sp) MOP stations. 1; 063; Depth range: 1530–1625 m; USNM voucher specimens: 2; 421350 and 421563. Distribution. Circumglobal, occurring at depths between 1097 and 4900 m. In the eastern Pacific this species has been previously recorded only in Peru and Chile (see Remarks). Literature. Günther (1878: 249), Markle & Quèro (1984: 228), Markle & Sazonov (1990: 246), Arruda (1997: 26), McEachran & Fechhelm (1998: 388), Porteiro et al. (1999: 27), Sazonov (1999: 737), Sazonov & Williams (2001: S32) and Moore et al. (2003: 179). Remarks. These specimens represent the first documented record of the species in Central American waters (Markle & Quèro 1984; Bussing & López 1994, 2009, 2011; Froese & Pauly 2016) and a northern range extension of about 1500 km in the species’ known distribution (Markle & Quèro 1984, Froese & Pauly 2016). The previous northernmost record for this species in eastern Pacific waters was off Peru (Markle & Quèro 1984, Froese & Pauly 2016). FIGURE 21. Bathytroctes microlepis. Top: entire specimen; Bottom: detail of head. (21) Leptoderma ospesca Angulo, Baldwin & Robertson, 2016. East Pacific smooth-head (En); Cabeza lisa del Pacífico Oriental (Sp)ROBERTSON ET AL .40 · Zootaxa 4348 (1) © 2017 Magnolia Press MOP stations. 1; 094; Depth range: 1368–1406 m; USNM voucher specimens: 1; 421478 (Holotype). Distribution. Eastern Pacific (known only from El Salvador), occurring at depths between 1368 and 1406 m. Literature. Angulo et al. (2016: 493). FIGURE 22. Leptoderma ospesca. Top: entire specimen; Bottom: detail of head; USNM 421478, 195 mm. SL. (22) Rouleina attrita (Vaillant, 1888). Softskin smooth-head (En); Cabeza lisa de piel suave (Sp) MOP stations. 1; 042; Depth range: 1115–1119 m USNM voucher specimens: 1; 422434. Distribution. Circumglobal, occurring at depths between 450 and 2300 m. In the eastern Pacific this species has been previously recorded off the states of Alaska, Washington and Oregon, USA, Peru and Chile. Literature. Vaillant (1888: 158), Paxton et al. (1989: 210), Markle & Sazonov (1990: 261), Arruda (1997: 28), Pequeño (1997: 80), McEachran & Fechhelm (1998: 398), Kenaley et al. (2006: 200), Zeballos (2009: 115), Møller et al. (2010:25) and Parin et al. (2014: 112). Remarks. This specimen represents the first documented record of the species in lower Central American waters (Markle & Sazonov 1990; Bussing & López 1994, 2011; Zeballos 2009; Froese & Pauly 2016). (23) Talismania bifurcata (Parr, 1951). Threadfin slickhead (En); Cabeza lisa de aletas filamentosas (Sp) MOP stations. 2; 030 and 087; Depth range: 864–1223 m; USNM voucher specimens: 2; 421479 and 421553. Distribution. Western (New South Wales, Australia) and eastern Pacific (from British Columbia, Canada, to Chile), occurring at depths between 300 and 2000 m. Literature. Parr (1951: 10), Bussing (1965: 192), Paxton et al. (1989: 211), McAllister (1990: 73), Ambrose (1996a: 224), Castro-Aguirre & Balart (1996: 69), Pequeño (1997: 80), Hutchins (2001: 19), Zeballos (2009: 117), Grinols & Heyamoto (2011: 1151) and Reyes-Bonilla et al. (2011: 6). Remarks. Specimen USNM 421479 represents the first documented record of the species in middle Central American waters (Parr 1951; Bussing & López 1994, 2009, 2011; Zeballos 2009; Froese & Pauly 2016). Zootaxa 4348 (1) © 2017 Magnolia Press · 41TEP DEEP FISHES FIGURE 23. Talismania bifurcata. Top: entire specimen; Bottom: detail of head; USNM 421479, 245 mm. SL. Platytroctidae Tubeshoulders, Searsids (En); Hombros luminosos (Sp). 2 G, 2 S. (24) Barbantus curvifrons (Roule & Angel, 1931). Palebelly searsid (En); Hombro luminoso de vientre claro (Sp) MOP stations. 2; 001 and 083; Depth range: 165–1116 m; USNM voucher specimens: 2; 421294 and 422531. Distribution. Circumtropical, occurring at depths between 0 and 4500 m. In the eastern Pacific this species has been recorded from the Gulf of California to Chile. Literature. Roule & Angel (1931: 6), Matsui & Rosenblatt (1987: 108), Pequeño (1989: 33), Castro-Aguirre & Balart (1996: 70), McEachran & Fechhelm (1998: 404), Sazonov & Merrett (2001: S38), Moore et al. (2003: 180) and Møller et al. (2010: 25). FIGURE 24. Barbantus curvifrons; USNM 421294, 104 mm. SL. Remarks. These specimens represent the first documented record of the species in Central American waters (Roule & Angel 1931; Bussing & López 1994, 2009, 2011; Froese & Pauly 2016). (25) Platytroctes apus Günther, 1878. Legless searsid (En); Hombro luminoso sin pies (Sp) MOP stations. 3; 001, 029 and 042; Depth range: 165–1126 m; USNM voucher specimens: 6; 421296, 421391, 422449, 422451, 422500 and 422630. Distribution. Circumglobal, occurring at depths between 385 and 5393 m. In the eastern Pacific this species has been recorded from Panama to Chile. Literature. Günther (1878: 249), Rohr (1968: 624), Paxton et al. (1989: 213), Pequeño (1989: 34), Matsui & ROBERTSON ET AL .42 · Zootaxa 4348 (1) © 2017 Magnolia Press Rosenblatt (1987: 104), Pakhorukov (1999: 656), Hutchins (2001: 19), Romero (2009a: 105), Møller et al. (2010: 26) and Fricke et al. (2011: 356). FIGURE 25. Platytroctes apus. Top: entire specimen; Bottom: detail of head; USNM 422500, 141 mm. SL. STOMIIFORMES 3 F, 3 G, 3 S. Sternoptychidae Marine hatchetfishes (En); Peces hacha marinos (Sp). 1 G, 1 S. 26) Sternoptyx diaphana Hermann, 1781. Diaphanous hatchet fish (En); Pez hacha transparente (Sp) MOP stations. 4; 001, 046, 058 and 059; Depth range: 183–1270 m; USNM voucher specimens: 5; 421254, 421510, 422505, 422506 and 422542. Distribution. Circumtropical, occurring at depths between 400 and 3676 m. In the eastern Pacific this species has been recorded from southern California, USA, to Chile. Literature. Hermann (1781: 33), Bussing (1965: 197), Baird (1971: 75), Matarese et al. (1989: 103), Paxton et al. (1989: 193), Pequeño (1989: 31), Scott & Scott (1988: 172), McAllister (1990: 90), Shinohara et al. (1996: 161), Arruda (1997: 34), De La Cruz Agüero et al. (1997: 291), Santos et al. (1997: 33), McEachran & Fechhelm (1998: 437), Porteiro et al. (1999: 29), Hutchins (2001: 19), Shinohara et al. (2001: 297), Nakabo (2002: 314), Moore et al. (2003: 184), Mundy (2005: 170), Castellanos-Galindo et al. (2006b: 251), Fricke et al. (2011: 357), Reyes-Bonilla et al. (2011: 6) and Parin et al. (2014: 146). Zootaxa 4348 (1) © 2017 Magnolia Press · 43TEP DEEP FISHES FIGURE 26. Sternoptyx diaphana; USNM 422505, 40 mm. SL. Phosichthyidae Lightfishes (En); Peces linterna, Linternas (Sp). 1 G, 1 S. (27) Yarrella argenteola (Garman, 1899). Panamanian lightfish (En); Linterna panameña (Sp) MOP stations. 2; 004 and 037; Depth range: 115–456 m; USNM voucher specimens: 4; 421422, 421423, 421493 and 421494. Distribution. Eastern Pacific (from the Gulf of Panama to northern Peru), occurring at depths between 384 and 1400 m. Literature. Garman (1899: 244), Bussing (1965: 197), Castro-Aguirre & Balart (1996: 70), Castellanos- Galindo et al. (2006b: 251) and Vélez (2009a: 124). FIGURE 27. Yarrella argenteola. Top: entire specimen; Bottom: detail of head. Stomiidae Barbeled dragonfishes (En); Peces demonio, Peces dragón con barbillas (Sp). 1 G, 1 S.ROBERTSON ET AL .44 · Zootaxa 4348 (1) © 2017 Magnolia Press (28) Stomias atriventer Garman, 1899. Black-belly dragonfish (En); Pez dragón de vientre oscuro (Sp) MOP stations. 10; 001, 003, 004, 028, 029, 034, 044, 067, 071 and 078; Depth range: 115–1472 m; USNM voucher specimens: 14; 421465, 421527, 421535, 421554, 422504, 422614, 422629, 435787, 435788, 435789, 435790, 435792, 435793 and 435798. Distribution. Eastern Pacific (from southern California to Chile, including the Galapagos Islands), occurring at depths between 100 and 1500 m. Literature. Garman (1899: 277), (Gibbs 1969: 3), Robison (1972: 451), Pequeño (1989: 31), Castro-Aguirre & Balart (1996: 70), Moser (1996a: 301), Rodríguez-Romero et al. (2008: 1771), Vélez (2009b: 130) and Reyes- Bonilla et al. (2011: 7). FIGURE 28. Stomias atriventer. Top: entire specimen; Bottom: detail of head; USNM 435798, 215 mm. SL. AULOPIFORMES 5 F, 5 G, 7 S. Chlorophthalmidae Greeneyes (En); Ojiverdes (Sp). 1 G, 1 S. (29) Chlorophthalmus mento Garman, 1899. Robust greeneye (En); Ojiverde robusto (Sp) FIGURE 29. Chlorophthalmus mento. Top: entire specimen; Bottom: detail of head; USNM 421332, 166 mm. SL. Zootaxa 4348 (1) © 2017 Magnolia Press · 45TEP DEEP FISHES MOP stations. 1; 033; Depth range: ok m; USNM voucher specimens: 1; 421332. Distribution. Eastern Pacific (from Mexico to Peru, including the Cocos and Galapagos Islands), occurring at depths between 17 and 1260 m. Literature. Garman (1899: 253), Bussing & López (1994: 54, 2005: 44; 2009: 447), Sulak (1995: 1005), Castellanos-Galindo et al. (2006b: 252), Sato (2009b: 135), McCosker & Rosenblatt (2010: 189) and Robertson & Allen (2015: 5404). Ipnopidae Deepsea tripod fishes, Spider fishes (En); Peces trípode, Peces araña (Sp). 1 G, 3 S. (30) Bathypterois atricolor Alcock, 1896. Attenuated spider fish (En); Pez araña atenuado (Sp) MOP stations. 5; 001, 053, 073, 086 and 094; Depth range: 153–1443 m; USNM voucher specimens: 7; 421469, 421538, 422395, 422418, 422427, 422431 and 423598. Distribution. Circumglobal, excluding the western Atlantic, occurring at depths between 250 and 5150 m. In the eastern Pacific this species has been recorded from the Gulf of California to central Chile. Literature. Alcock (1896: 306), Sulak (1977b: 87), Pequeño (1989: 34), Adam et al. (1998: 9), Nakabo (2000: 360), Shinohara et al. (2001: 300), Mundy (2005: 195), Castellanos-Galindo et al. (2006b: 252), Shao et al. (2008:241) and McCosker & Rosenblatt (2010: 189). Remarks. According to Sulak´s (1977b) key, B. atricolor and B. ventralis can be separated by the number of lower pectoral fin rays (9–11 vs. 12–14, respectively); however, other “diagnostic” characters listed by this author (i.e. numbers of lateral line scales and vertebrae, shape of subcaudal notch) overlap between the species, making their separation difficult. COI sequences for specimens identified as B. atricolor and B. ventralis based on number of lower pectoral-fin rays form a single genetic lineage (Figure 31). Further studies are necessary to resolve the taxonomy of these species. USNM 421469 and 421538 specimens represent the first documented record of B. atricolor in upper Central American waters (Sulak 1977; Bussing & López 1994, 2009, 2011; Froese & Pauly 2016). FIGURE 30. Bathypterois atricolor. Top: entire specimen; Bottom: detail of head; USNM 421538, 132 mm. SL.ROBERTSON ET AL .46 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 31. Neighbor-joining tree based on raw sequence divergence of Bathypterois specimens. Sequence divergence among specimens of B. ventralis and B. atricolor is less than 1.25%. Scale bar is equivalent to 1% raw sequence divergence. (31) Bathypterois sp. MOP stations. 1; 078; Depth range: 1320–1370 m; USNM voucher specimens: 1; 422410. Remarks. The COI sequence for a specimen identified only to the genus Bathypterois differs by about 4% from COI sequences of the B. atricolor/ventralis lineage (Figure 31). (32) Bathypterois ventralis Garman, 1899. Ventrad spiderfish (En); Pez araña (Sp) MOP stations. 9; 005, 055, 067, 078, 083, 086, 094, 096 and 099; Depth range: 165–1406 m; USNM voucher specimens: 16; 421272, 421276, 421286, 421304, 421343, 421484, 421506, 421529, 421531, 421536, 422345, 422402, 422415, 422503, 422561 and 422569. Distribution. Indo-Pacific, occurring at depths between 165 and 1735 m (see Remarks). In the eastern Pacific this species has been recorded from the Gulf of California to central Chile. Literature. Garman (1899: 256), Sato (2009c: 137), Sulak (1977b: 87), Pequeño (1997: 81), Hutchins (2001: 21), Castellanos-Galindo et al. (2006b: 252) and Angulo et al. (2015: 546). Remarks. USNM 421272, 421276, 421484, 421536, 422345, 422561 and 422569 represent the first documented record of the species in upper Central American waters (Sulak 1977; Bussing & López 1994, 2009, 2011; Angulo et al. 2015; Froese & Pauly 2016). In addition, specimens collected at 165 m represent a new minimum depth record for the species (Castellanos-Galindo et al. 2006b, Angulo et al. 2015, Froese & Pauly 2016). See also Figure 31 and remarks about B. atricola and Bathypterois sp. Zootaxa 4348 (1) © 2017 Magnolia Press · 47TEP DEEP FISHES FIGURE 32. Bathypterois ventralis. Top: entire specimen; Bottom: detail of head; USNM 422503, 125 mm. SL. Scopelarchidae Pearleyes (En); Ojos de perla (Sp). 1 G, 1 S. (33) Scopelarchoides nicholsi Parr, 1929. Nichols’s pearl eye (En); Ojo de perla de Nichols (Sp) MOP stations. 3; 029, 040 and 052; Depth range: 836–1185 m; USNM voucher specimens: 3; 421401, 422485 and 422520. Distribution. Eastern Pacific (from Baja California, Mexico, to Chile) and western central Atlantic (Cuba), occurring at depths between 94 and 3294 m. Literature. Parr (1929: 16), Bussing (1965: 211), Watson & Sandknop (1996a: 332), Johnson (1974: 136; 1982:165), Pequeño (1989: 34), Sato (2009d: 139) and Davis (2015: 67).ROBERTSON ET AL .48 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 33. Scopelarchoides nicholsi. Top: entire specimen; Bottom: detail of head; USNM 422485, 105 mm. SL. Synodontidae Lizardfishes (En); Garrobos (Sp). 1 G, 1 S. (34) Synodus evermanni Jordan & Bollman, 1890. Sharpnose lizardfish, Spotted lizardfish, Inotted lizardfish (En); Lagarto cadena, Garrobo, Lagarto camotillo (Sp) MOP stations. 2; 005 and 015; Depth range: 109–114 m; USNM voucher specimens: 7; 421246, 421250, 421308, 421320, 421324, 421329 and 422338. Distribution. Eastern Pacific (from central Baja California, México, to Peru), occurring at depths between 25 and 300 m. Literature. Allen & Robertson (1994: 72), Bussing & López (1994: 54; 2009: 446), Bussing & Lavenberg (1995a: 1626), Bearez (1996: 734), Stevens & Moser (1996a: 348), De La Cruz Agüero et al. (1997: 298), Chirichigno & Vélez (1998: 171), Castellanos-Galindo et al. (2006b: 25), Rodríguez-Romero et al. (2008: 1770), Sato (2009a: 134), Del Moral-Flores et al. (2013: 192), Fuentes et al. (2015: 615) and Robertson & Allen (2015: 621). Remarks. The seven specimens collected and sequenced form two genetic lineages based on their DNA barcodes, which differ by 10.0–10.2%. An individual of each lineage was present in Trawl 15, indicating they likely are not allopatric FIGURE 34. Synodus evermanni. Zootaxa 4348 (1) © 2017 Magnolia Press · 49TEP DEEP FISHES Paralepididae Barracudinas (En); Barracudinas (Sp). 1 G, 1 S. (35) Lestidiops neles (Harry, 1953). Tropical eastern Pacific barracudina (En); Barracudina del Pacífico Tropical Oriental (Sp) MOP stations. 1; 003; Depth range: 116–118 m; USNM voucher specimens: 1; 421259. Distribution. Eastern Pacific (from the vicinity of Cabo San Lucas, Baja California, México, to the Gulf of Panama), occurring at depths between 22 and 384 m. Literature. Harry (1953: 199), Ambrose (1996b: 352) and Castellanos-Galindo et al. (2006b: 254). FIGURE 35. Lestidiops neles. Top: entire specimen; Bottom: detail of head; USNM 421259, 157 mm. SL. MYCTOPHIFORMES 2 F, 3 G, 3 S. Neoscopelidae Blackchins, Neoscopelids (En); Linternillas, Barbillas oscuras (Sp). 1 G, 1 S. (36) Scopelengys tristis Alcock, 1890. Pacific blackchin (En); Barbilla negra del Pacífico (Sp) MOP stations. 4; 067, 071, 076 and 100; Depth range: 697–1145 m; USNM voucher specimens: 6; 421425, 421443, 421447, 421512, 435797 and 435799. Distribution. Cosmopolitan, occurring at depths between 400 and 1830 m. Literature. Alcock (1890: 303), Bussing (1965: 200), Fitch & Lavenberg (1968: 42), Robison (1972: 451), Nielsen (1974: 40), Menon & Rama-Rao (1975: 35), Butler & Ahlstrom (1976: 143), Pequeño (1989: 38), Becker & Shcherbachev (1990: 845), Bala11 & Fedorov (1996: 345), Castro-Aguirre & Balart (1996: 70), Moser (1996b: 383), Nakabo (2000: 377; 2002: 377), Mecklenburg et al. (2002: 243), Menezes et al. (2003: 52), Castellanos- Galindo et al. (2006b: 254), Romero (2009b: 143), Reyes-Bonilla et al. (2011: 7) and Parin et al. (2014: 154). ROBERTSON ET AL .50 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 36. Scopelengys tristis. Top: entire specimen; Bottom: detail of head; USNM 421425, 115 mm. SL. Myctophidae Lanternfishes, Myctophids (En); Peces linterna, Linternillas, Mictófidos (Sp). 2 G, 2 S. (37) Benthosema panamense (Tåning, 1932). Panama lanternfish (En); Pez linterna de Panamá (Sp) MOP stations. 4; 039, 047, 074 and 103; Depth range: 117–422 m; USNM voucher specimens: 10; 421419, 421449, 421450, 421453, 422484, 422486, 422499, 422523, 422536 and 422541. Distribution. Eastern Pacific (from the Gulf of California to northern Peru), occurring at depths between 13 and 600 m. Literature. Tåning (1932: 129), Robison (1972: 448), Nafpaktitis (1973: 10), Nielsen (1974: 40), Paxton (1979: 6), Paxton et al. (1995: 315), Castro-Aguirre & Balart (1996: 70), Moser & Ahlstrom (1996: 389), De La Cruz Agüero et al. (1997: 229), Chirichigno & Vélez (1998: 165), Castellanos-Galindo et al. (2006b: 254), Romero (2009c: 144), Del Moral-Flores et al. (2013: 192) and Santana-Iturríos et al. (2013: 387). (38) Lampanyctus parvicauda Parr, 1931. Slimtail lampfish (En); Pez linterna de cola delgada (Sp) MOP stations. 1; 077; Depth range: 1320–1370 m; USNM voucher specimens: 1; 422304. Distribution. Eastern Pacific (from the Gulf of California to Peru), occurring at depths between 9 and 1370 m (see Remarks). Literature. Parr (1931: 26), Bussing (1965: 207), Robison (1972: 451), Paxton (1979: 14), Pequeño (1989: 38), Paxton et al. (1995: 315), Castro-Aguirre & Balart (1996: 70), Moser & Ahlstrom (1996: 389), Mundy (2005: 225), Castellanos-Galindo et al. (2006b: 255) and Romero (2009c: 149). Remarks. This specimen represents a new maximum depth record for the species (Castellanos-Galindo et al. 2006b, Romero 2009c, Froese & Pauly 2016). Zootaxa 4348 (1) © 2017 Magnolia Press · 51TEP DEEP FISHES FIGURE 37. Benthosema panamense. Top & Bottom: two entire specimens. GADIFORMES 4 F, 6 G, 11 S. Bregmacerotidae Codlets, Codlings (En); Bacaletes (Sp). 1 G, 1 S. (39) Bregmaceros bathymaster Jordan & Bollman, 1890. East Pacific codlet (En); Bacalete del Pacífico oriental (Sp) FIGURE 38. Bregmaceros bathymaster. MOP stations. 2; 085 and 102; Depth range: okm; USNM voucher specimens: 5; 421278, 421377, 421416, 422379 and 422387. Distribution. Eastern Pacific (from the Gulf of California to Peru, including all oceanic islands), occurring at depths between 0 and 1246 m. Literature. Jordan & Bollman (1890: 173), Böhlke (1953: 56), D'Ancona & Cavinato (1965: 43), Robison (1972: 451), Masuda et al. (1986: 393), Bussing & López (1994: 72; 2009: 449), Bearez (1996: 734), Castro- Aguirre & Balart (1996: 70), Stevens & Moser (1996b: 477), De La Cruz Agüero et al. (1997: 75), Grove & Lavenberg (1997: 207), Chirichigno & Vélez (1998: 232), Castellanos-Galindo et al. (2006a: 195), Endo (2009a: 158), Del Moral-Flores et al. (2013: 192) and Robertson & Allen (2015: 1702).ROBERTSON ET AL .52 · Zootaxa 4348 (1) © 2017 Magnolia Press Macrouridae Grenadiers, Rattails (En); Granaderos, Colas de rata (Sp). 3 G, 8 S. (40) Caelorinchus canus (Garman, 1899). Clearsnout grenadier (En); Granadero cano, Granadero de hocico claro (Sp) MOP stations. 1; 057; Depth range: 134–162 m; USNM voucher specimens: 2; 421291 and 421292. Distribution. Eastern Pacific (from Costa Rica to northern Peru, including the Galápagos Islands), occurring at depths between 70 and 460 m. Literature. Garman (1899: 217), Cohen et al. (1990: 150), Iwamoto & Schneider (1995: 1246), Bearez (1996: 734), Chirichigno & Vélez (1998: 235), Iwamoto & McCosker (2001: 27), Castellanos-Galindo et al. (2006b: 195), Iwamoto (2009: 159) and McCosker & Rosenblatt (2010: 189). FIGURE 39. Caelorinchus canus. Top: entire specimen; Bottom: detail of head (41) Coryphaenoides anguliceps (Garman, 1899). Loosescale grenadier (En); Granadero anguloso (Sp) MOP stations. 3; 073, 086 and 099; Depth range: 153–1389 m; USNM voucher specimens: 5; 421564, 422456, 422610, 422620 and 423198. Distribution. Eastern Pacific (from the Gulf of California-northern Peru, including the Cocos and Galapagos Islands), occurring at depths between 700 and 2400 m. Literature. Garman (1899: 212), Iwamoto & Sazonov (1988: 45), Cohen et al. (1990: 203), Iwamoto & Schneider (1995: 1258), Ambrose (1996c: 483), Castro-Aguirre & Balart (1996: 70), McMillan (1999: 488), Iwamoto & McCosker (2001: 27), Iwamoto (2009: 161) and McCosker & Rosenblatt (2010: 189). Zootaxa 4348 (1) © 2017 Magnolia Press · 53TEP DEEP FISHES FIGURE 40. Coryphaenoides anguliceps. Top: entire specimen; Bottom: detail of head; USNM 421564, 63 mm. Head Length. (42) Coryphaenoides capito (Garman, 1899). Bighead grenadier (En); Granadero cabezón (Sp) MOP stations. 5; 003, 067, 071, 075 and 087; Depth range: 116–1223 m; USNM voucher specimens: 12; 421235, 421273, 421312, 421341, 421444, 421456, 421460, 421461, 421480, 422355 and 422370. Distribution. Eastern Pacific (from the Gulf of California to northern Peru), occurring at depths between 300 and 1223 m (see Remarks). Literature. Garman (1899: 200), Iwamoto & Sazonov (1988: 62), Iwamoto & Schneider (1995: 1262), Chirichigno & Vélez (1998: 236), Castellanos-Galindo et al. (2006a: 196) and Iwamoto (2009: 165). Remarks. Specimens collected at 1223 m represent a new maximum depth record for the species (Castellanos- Galindo et al. 2006a, Iwamoto 2009, Froese & Pauly 2016). FIGURE 41. Coryphaenoides capito. Top: entire specimen; Bottom: detail of head; USNM 421456, 46 mm. Head Length. (43) Coryphaenoides carminifer (Garman, 1899). Carmine grenadier (En); Granadero carminífero (Sp) MOP stations. 6; 001, 040, 043, 073, 077 and 086; Depth range: 183–1389 m; USNM voucher specimens: 10; 421432, 421475, 421557, 422419, 422433, 422441, 422446, 422555, 422634 and 422649. Distribution. Eastern Pacific (from Nicaragua to northern Peru, including the Cocos Island), occurring at depths between 183 and 1870 m (see Remarks). ROBERTSON ET AL .54 · Zootaxa 4348 (1) © 2017 Magnolia Press Literature. Garman (1899: 204), Iwamoto & Sazonov (1988: 42), Pequeño (1989: 45, 1997: 81), Cohen et al. (1990: 206) and Iwamoto & Schneider (1995: 1263), Castellanos-Galindo et al. (2006a: 196) and Iwamoto (2009: 166). Remarks. USNM 421475, 421557, 422419, 422433, 422441 and 422649 specimens represent the first documented record of the species in middle Central American waters (Garman 1899; Iwamoto & Sazonov 1988; Bussing & López 1994, 2009, 2011; Yabe 2009b), and a northwestern range extension of about 1000 km in the species’ known distribution (Garman 1899, Iwamoto & Sazonov 1988, Iwamoto 2009). The previous northernmost documented record for this species in eastern Pacific waters was in the Gulf of Panama (Garman 1899). In addition, specimens collected at 183 m represent a new minimum depth record for the species (Castellanos-Galindo et al. 2006a, Iwamoto 2009, Froese & Pauly 2016). FIGURE 42. Coryphaenoides carminifer. Top: entire specimen; Bottom: detail of head; USNM 422649, 70 mm. Head Length. (44) Nezumia convergens (Garman, 1899). Peruvian grenadier (En); Granadero peruano (Sp) FIGURE 43. Nezumia convergens. Top: entire specimen; Bottom: detail of head; USNM 422577, 36 mm. Head Length. Zootaxa 4348 (1) © 2017 Magnolia Press · 55TEP DEEP FISHES MOP stations. 4; 001, 064, 072 and 073; Depth range: 153–1330 m; USNM voucher specimens: 9; 421338, 421344, 421402, 421435, 421546, 421572, 422302, 422400 and 422577. Distribution. Eastern Pacific (from the Gulf of California to southern Chile, including the Cocos and Galápagos Islands), occurring at depths between 153 and 1870 m (see Remarks). Literature. Garman (1899: 210), Iwamoto (1979: 171, 2009: 168), Cohen et al. (1990: 270), Iwamoto & Schneider (1995: 1264), Ambrose (1996c: 483), Castro-Aguirre & Balart (1996: 70), Pequeño (1997: 82), Chirichigno & Vélez (1998: 237), Iwamoto & McCosker (2001: 27), Wilson (2001: 32), Castellanos-Galindo et al. (2006a: 196) and McCosker & Rosenblatt (2010:189). Remarks. Specimens collected at 153 m represent a new minimum depth record for the species (Castellanos- Galindo et al. 2006a, Iwamoto 2009, Froese & Pauly 2016). (45) Nezumia latirostrata (Garman, 1899). Broadsnout grenadier (En); Granadero hocicón (Sp) MOP stations. 3; 001, 003 and 033; Depth range: 116–842 m; USNM voucher specimens: 3; 421257, 421481 and 422545. Distribution. Eastern Pacific (from Panama to northern Peru), occurring at depths between 116 and 1865 m (see Remarks). Literature. Garman (1899: 211), Iwamoto (1979:168, 2009: 169), Cohen et al. (1990: 272), Iwamoto & Schneider (1995: 1259), Chirichigno & Vélez (1998: 238) and Castellanos-Galindo et al. (2006a: 196). Remarks. Specimens collected at 116 m represent a new minimum depth record for the species (Castellanos- Galindo et al. 2006a, Iwamoto 2009, Froese & Pauly 2016). (46) Nezumia liolepis (Gilbert, 1890). Smooth grenadier (En); Granadero liso (Sp) FIGURE 44. Nezumia liolepis. Top: entire specimen; Bottom: detail of head; USNM 422462, 65 mm. Head Length. MOP stations. 6; 064, 067, 071, 073, 099 and 100; Depth range: 153–1223 m; USNM voucher specimens: 13; 421440, 421458, 421474, 421476, 421544, 422375, 422421, 422458, 422460, 422462, 422568, 422582 and 422590. Distribution. Eastern Pacific (from north California, USA, to Costa Rica), occurring at depths between 153 and 1660 m (see Remarks). Literature. Gilbert (1890: 117), Iwamoto & Stein (1974: 46), Iwamoto (1979: 157), Cohen et al. (1990: 274), ROBERTSON ET AL .56 · Zootaxa 4348 (1) © 2017 Magnolia Press Iwamoto & Schneider (1995: 1265), Ambrose (1996c: 483), Castro-Aguirre & Balart (1996: 70), Hoff (1999: 115), Hoff et al. (2000: 662) and Rodríguez-Romero et al. (2008: 1771). Remarks. These specimens represent the first documented record of the species in Central American waters (Iwamoto 1979; Cohen et al. 1990; Bussing & López 1994, 2009, 2011) and a southeastern range extension of about 1600 km in the species’ known distribution (Iwamoto 1979, Cohen et al. 1990). The previous southernmost documented record for this species in eastern Pacific waters was off Guerrero, México (Cohen et al. 1990). In addition, specimens collected at 153 m represent a new minimum depth record for the species (Cohen et al. 1990, Froese & Pauly 2016). (47) Nezumia stelgidolepis (Gilbert, 1890). California grenadier (En); Granadero californiano (Sp) MOP stations. 4; 003, 033, 071 and 075; Depth range: 116–965 m; USNM voucher specimens: 5; 421238, 421297, 421330, 421490 and 422368. Distribution. Eastern Pacific (from British Columbia, Canada, to southern Peru, including the Galapagos Islands), occurring at depths between 116 and 965 m (see Remarks). Literature. Gilbert (1890: 116), Fitch & Lavenberg (1968: 73), Iwamoto & Stein (1974: 47), Iwamoto (1979: 160, 2009: 172), Pequeño (1989: 46), McAllister (1990: 123), Cohen et al. (1990: 286), Iwamoto & Schneider (1995: 1260), Ambrose (1996c: 483), Chirichigno & Vélez (1998: 237), Hoff et al. (2000: 662), Iwamoto & McCosker (2001: 27), Castellanos-Galindo et al. (2006a: 196), Rodríguez-Romero et al. (2008: 1771) and McCosker & Rosenblatt (2010: 189). Remarks. Specimens collected at 165 and 965 m represent new minimum and maximum depth records for the species (Cohen et al. 1990, Castellanos-Galindo et al. 2006a, Iwamoto 2009, Froese & Pauly 2016). Moridae Morid cods, Moras (En); Moras, Carboneros (Sp). 1 G, 1 S. (48) Physiculus nematopus Gilbert, 1890. Charcoal mora (En); Mora carbon, Mora de fango, Carbonero de fango (Sp) FIGURE 45. Physiculus nematopus. MOP stations. 3; 003, 004 and 040; Depth range: 115–1185 m; USNM voucher specimens: 5; 421305, 421413, 422543, 422559 and 422571. Distribution. Eastern Pacific (from Central Baja California, USA, to Peru, including the Galápagos Islands), occurring at depths between 0 and 1272 m. Literature. Gilbert (1890: 114), Böhlke (1953: 55), Matarese et al. (1989: 183), Paulin (1989: 119), Cohen et al. (1990: 370), Bussing & López (1994: 72), Paulin (1995: 1286), Bearez (1996: 734), Ambrose (1996d: 500), Castro-Aguirre & Balart (1996: 70), Grove & Lavenberg (1997: 204), McCosker et al. (1997: 25), Chirichigno & Vélez (1998: 239), Castellanos-Galindo et al. (2006a: 196), Rodríguez-Romero et al. (2008: 1771), Endo (2009b: 179), McCosker & Rosenblatt (2010: 189), López-Martínez et al. (2012: 351), Del Moral-Flores et al. (2013: 193) and Robertson & Allen (2015: 564). Zootaxa 4348 (1) © 2017 Magnolia Press · 57TEP DEEP FISHES Merlucciidae Merluccid hakes (En); Merluzas (Sp). 1 G, 1 S. (49) Merluccius angustimanus Garman, 1899. Panama hake (En); Merluza panameña (Sp) MOP stations. 5; 004, 007, 047, 061 and 068; Depth range: 115–478 m; USNM voucher specimens: 9; 421331, 421468, 421524, 422438, 422450, 422470, 422498, 422528 and 422587. Distribution. Eastern Pacific (from southern California, USA, to the Ensenada de Tumaco, Colombia), occurring at depths between 80 and 523 m. Literature. Garman (1899: 183), Robison (1972: 451), Cohen et al. (1990: 331), Bussing & López (1994: 72; 2009: 449), Balart et al. (1995: 82), Inada (1995: 1272), Ambrose (1996e: 508), Castro-Aguirre & Balart (1996: 70), Grant & Leslie (2001: 700), Lloris et al. (2005: 19), Castellanos-Galindo et al. (2006a: 196), Rodríguez- Romero et al. (2008: 1771), López-Martínez et al. (2012: 351), Del Moral-Flores et al. (2013: 193) and Robertson & Allen (2015: 2596). FIGURE 46. Merluccius angustimanus. Top: entire specimen; Bottom: detail of head; USNM 422587, 168 mm. SL. OPHIDIIFORMES 1 F, 7 G, 8 S. Ophidiidae Cusk-eels, Brotulas (En); Brotulas, Congriperlas (Sp). 7 G, 8 S. (50) Cherublemma emmelas (Gilbert 1890). Black brotula (En); Brotula negra (Sp) MOP stations. 4; 007, 060, 070 and 074; Depth range: 243–672 m; USNM voucher specimens: 12; 421218, 421337, 421429, 421492, 421565, 422374, 422407, 422544, 422549, 422557, 422567 and 422581. Distribution. Eastern Pacific (from Baja California, Mexico, including the Gulf of California, to northern Chile), occurring at depths between 102 and 1010 m. Literature. Gilbert (1890: 110), Böhlke (1953: 102), Pequeño (1989: 48), Castro-Aguirre et al. (1993: 80), Bussing & López (1994: 76; 2009: 449), Lea (1995: 1343), De La Cruz Agüero et al. (1997: 244), Ambrose (1996f: 513), Chirichigno & Vélez (1998: 224), Nielsen et al. (1999: 29), Castellanos-Galindo et al. (2006a: 197), Aguirre-Villaseñor & Castillo-Velázquez (2011: 713), López-Martínez et al. (2012: 351), and Robertson & Allen (2015: 5406).ROBERTSON ET AL .58 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 47. Cherublemma emmelas. Top: entire adult specimen; Center: detail of head of adult; Bottom: entire juvenile specimen. (51) Dicrolene filamentosa Garman, 1899. Deepwater thread brotula, Filamentous brotula (En); Brotula filamentosa (Sp) FIGURE 48. Dicrolene filamentosa. Top: entire specimen; Bottom: detail of head. MOP stations. 17; 001, 002, 004, 005, 030, 043, 053, 055, 063, 067, 077, 078, 086, 087, 094, 096 and 099; Depth range: 115–1625 m; USNM voucher specimens: 23; 421233, 421414, 421445, 421477, 421482, 421488, 421495, Zootaxa 4348 (1) © 2017 Magnolia Press · 59TEP DEEP FISHES 421496, 421497, 421520, 421523, 421526, 421533, 421561, 421566, 422406, 422442, 422494, 422588, 422613, 422643, 435796 and 435802. Distribution. Eastern Pacific (from southern Mexico to central Peru), occurring at depths between 780 and 1865 m. Literature. Garman (1899: 149), Bearez (1996: 734), Ambrose (1996f: 513), Castro-Aguirre & Balart (1996: 70), Chirichigno & Vélez (1998: 231), Nielsen et al. (1999: 62), Castellanos-Galindo et al. (2006a: 197) and Nielsen & Nishioka (2009: 187). (52) Lamprogrammus cf. niger Alcock, 1891. Paperbone brotula, Black cusk-eel (En); Brotula oscura, Brotula caverna (Sp) FIGURE 49. Lamprogrammus cf. niger. Top: entire specimen; Bottom: detail of head; USNM 435786, 270 mm. TL. MOP stations. 3; 001, 026 and 078; Depth range: 165–1087 m; USNM voucher specimens: 3; 422477, 422593 and 435786. Distribution. Circumtropical, sometimes found in subtropical areas, occurring at depths between 165 and 2615 m (see Remarks). In the eastern Pacific this species has been recorded previously only from Ecuador to northern Peru. Literature. Alcock (1891: 33), Menon and Rama-Rao (1970: 377), Cohen & Nielsen (1978: 34), Nielsen (1990: 564), Cohen et al. (1991), Cohen & Rohr (1993: 472), Ambrose (1996f: 513), Castro-Aguirre & Balart (1996: 70), McEachran & Fechhelm (1998: 715), Nielsen et al. (1999: 72), Hutchins (2001: 23), Moore et al. (2003: 207), Mundy (2005: 244), Castellanos-Galindo et al. (2006a: 198), Garrido-Linares & Acero (2006: 292), Nielsen & Nishioka (2009: 190) and Reyes-Bonilla et al. (2011: 7). Remarks. These specimens were tentatively identified by us as Lamprogrammus cf. niger as they share with this species the following combination of distinctive characters, sensu Cohen & Rohr (1993) and Nielsen et al. (1999): body depth 13.10–16.00% of SL; maxillary sheathed posterodorsally; opercular spine slender, flexible and sharp-pointed; and posterior margin of opercle smooth to weakly fimbriate. However, our specimens differ from L. niger, sensu stricto, by having fewer dorsal-fin (100–101, vs. 108–112) and anal-fin (79–81, vs. 85–90) elements, counts not reported until now in other Lamprogrammus species (Cohen et al. 1991, Cohen & Rohr 1993, Nielsen et al. 1999). Further, three specimens tentatively identified as Lamprogrammus cf. niger, deposited at the UCR (Velero 19073; two specimens) and FMNH (113549; one specimen), also have low dorsal- and anal-fin counts, and have a small median basibranchial tooth patch. This tooth patch is absent in L. niger according to Cohen & Rohr (1993), and was not observed in our specimens. COI sequences for our specimens and one specimen identified as L. niger from the Eastern North Pacific that is available in BOLD (UW150575) are less than 1% divergent and about 10% divergent to two other available species of the genus [i.e. L. brunswigi (Brauer 1906) and L. exutus Nybelin & Poll 1958]. Based on the above, and assuming intraspecific variation, our specimens represent the first documented record of the L. niger in Central American waters (Bussing & López 1994, 2009, 2011; Nielsen & Nishioka 2009) and a ROBERTSON ET AL .60 · Zootaxa 4348 (1) © 2017 Magnolia Press northwest range extension of about 1400 km in the species’ known distribution (Nielsen & Nishioka 2009). The previous northernmost record for L. niger in the eastern Pacific was off Ecuador (Nielsen & Nishioka 2009). In addition, specimens collected at 165 m will represent a new minimum depth record for the species (Nielsen 1990, Nielsen & Nishioka 2009, Froese & Pauly 2016). More detailed studies are suggested to validate the species assignation of these Eastern Pacific specimens. (53) Luciobrotula coheni Nielsen, 2009. Cohen’s brotula (En); Brotula de Cohen (Sp) MOP stations. 5; 001, 004, 033, 056 and 059; Depth range: 115–920 m; USNM voucher specimens: 5; 421217, 421356, 421491, 421528 and 422550. Distribution. Eastern Pacific (from Costa Rica to Peru), occurring at depths between 115 and 975 m (see Remarks). Literature. Nielsen (2009: 151). Remarks. USNM 421217 and 422550 specimens represent the first documented record of the species in Costa Rican waters (Bussing & López 1994, 2009, 2011; Nielsen 2009) and a northwestern range extension of about 600 km in the species’ known distribution (Nielsen (2009). The previous northernmost record for this species in the eastern Pacific was off Panama (see Nielsen 2009). In addition, specimens collected at 115 m represent a new minimum depth record for the species (Nielsen 2009, Froese & Pauly 2016). FIGURE 50. Luciobrotula coheni. Top: entire specimen; Bottom: detail of head; USNM 422550, 100 mm. SL. (54) Monomitopus malispinosus (Garman, 1899) FIGURE 51. Monomitopus malispinosus. Top: entire specimen; Bottom: detail of head. Zootaxa 4348 (1) © 2017 Magnolia Press · 61TEP DEEP FISHES MOP stations. 4; 003, 004, 043 and 067; Depth range: 116–951 m; USNM voucher specimens: 11; 421240, 421249, 421300, 421441, 421464, 421530, 422492, 422509, 422552, 422624 and 435794. Distribution. Eastern Pacific (from Costa Rica to northern Peru, including the Galápagos Islands), occurring at depths between 840 and 1865 m (see Remarks). Literature. Garman (1899: 158), Nielsen et al. (1999: 78), Castellanos-Galindo et al. (2006a: 198), Nielsen & Nishioka (2009: 195) and McCosker & Rosenblatt (2010: 190). Remarks. USNM 421441 specimen represent the first documented record of the species in Costa Rican waters (Bussing & López 1994, 2009, 2011; Nielsen & Nishioka 2009) and a northwestern range extension of about 600 km in the species’ known distribution (Garman 1899, Castellanos-Galindo et al. 2006, Nielsen & Nishioka 2009). The previous northernmost documented record for this species in the eastern Pacific was from the Gulf of Panama (Garman 1899). In addition, specimens collected at 116 m represent a new minimum depth record for the species (Nielsen et al. 1999, Nielsen & Nishioka 2009, Froese & Pauly 2016). (55) Monomitopus torvus Garman, 1899. Congrio cara con hueco (Sp) MOP stations. 1; 004; Depth range: 115–116 m; USNM voucher specimens: 1; 422497. Distribution. Eastern Pacific (from the Gulf of Panama to nothern Peru), occurring at depths between 115 and 1260 m (see Remarks). Literature. Garman (1899:157), Carter & Cohen (1985: 87), Pequeño (1989: 48), Nielsen et al. (1999: 78), Castellanos-Galindo et al. (2006a: 198) and Nielsen & Nishioka (2009: 196). Remarks. This specimen represents the first documented record of the species in Panamanian waters (Nielsen et al. 1999, Nielsen & Nishioka 2009) and a northern range extension of about 150 km in the species’ known distribution (Garman 1899, Castellanos-Galindo et al. 2006a, Nielsen & Nishioka 2009). The previous northernmost documented record for this species in the eastern Pacific was from north Colombia (Castellanos- Galindo et al. 2006a, Nielsen & Nishioka 2009). In addition, specimens collected at 115 m represents a new minimum depth record for the species (Nielsen et al. 1999, Nielsen & Nishioka 2009, Froese & Pauly 2016). (56) Neobythites stelliferoides Gilbert, 1890. Thread brotula (En); Brótula de hebra (Sp) FIGURE 52. Neobythites stelliferoides. MOP stations. 2; 038 and 068; Depth range: 144–154 m; USNM voucher specimens: 6; 421392, 421399, 421439, 422312, 422341 and 422448. Distribution. Eastern Pacific (from southern Baja California Sur, Mexico, including the Gulf of California to northern Peru), occurring at depths between 85 and 516 m. Literature. Gilbert (1890: 112), Bussing & López (1994: 76; 2009: 449), Balart et al. (1995: 82), Castro- Aguirre & Balart (1996: 70), Nielsen et al. (1999: 83), Nielsen (2002: 88), Castellanos-Galindo et al. (2006a: 198), Nielsen & Nishioka (2009: 197), Del Moral-Flores et al. (2013: 193) and Robertson & Allen (2015: 709).ROBERTSON ET AL .62 · Zootaxa 4348 (1) © 2017 Magnolia Press (57) Otophidium indefatigabile Jordan & Bollman 1890. Barred cusk-eel, Big-headed cusk-eel, Panamic cusk- eel (En); Congriperla cabezona, Congriperla listada, Congriperla mancha café (Sp) MOP stations. 3; 032, 074 and 098; Depth range: 108–422 m; USNM voucher specimens: 4; 421299, 421302, 422512 and 421519. Distribution. Eastern Pacific (from Baja California and the Gulf of California, Mexico, to Panama, including the Galápagos Islands), occurring at depths between 5 and 422 m (see Remarks). Literature. Jordan & Bollman (1890: 172), Grove & Lavenberg (1997: 213), Nielsen et al. (1999: 43), Castellanos-Galindo et al. (2006a: 198), McCosker & Rosenblatt (2010: 190) and Robertson & Allen (2015: 705). Remarks. Specimens collected at 422 m represent a new maximum depth record for the species (Nielsen et al. 1999, Castellanos-Galindo et al. 2006a, Robertson & Allen 2015, Froese & Pauly 2016). FIGURE 53. Otophidium indefatigabile. Top: entire specimen; Bottom: detail of head. (58) Porogadus sp. MOP stations. 3; 028, 063 and 086; Depth range: 1353–1625 m; USNM voucher specimens: 2; 421448, 421549 and 435796. Literature. Nielsen et al. (1999: 49). FIGURE 54. Porogadus sp. Top: entire specimen; Bottom: detail of head. Remarks. As noted by Nielsen et al. (1999), no formal review of Porogadus Goode & Bean 1885 has been published up to this point. Moreover, an identification key currently is available only for Western Atlantic species (Carter & Sulak 1984). The taxonomy of the genus is complicated and the validity of several forms is questionable given the absence of sound diagnostic characters. Accordingly, we were not able to make a specific identification Zootaxa 4348 (1) © 2017 Magnolia Press · 63TEP DEEP FISHES of these specimens. However, based on the fact that our specimens have very weak head spines (cf Nielsen et al. (1999)) and considering the collection sites and the absence of genetic structure (based on COI sequences), they possibly correspond to one of the following three species: Porogadus atripectus Garman 1899, P. catena (Goode & Bean 1885) or P. longiceps Garman 1899. LOPHIIFORMES 6 F, 7 G, 13 S. Lophiidae Goosefishes, Anglerfishes (En); Rapes, Peces pescadores (Sp). 1 G, 2 S. (59) Lophiodes caulinaris (Garman, 1899). Pacific anglerfish, spottedtail angler (En); Rape de rabo manchado (Sp) MOP stations. 3; 006, 015 and 048; Depth range: 109–143 m; USNM voucher specimens: 6; 421345, 421346, 421347, 422366, 422388 and 422412. Distribution. Eastern Pacific (from southern California, USA, to Peru, including the Cocos and Malpelo Islands), occurring at depths between 15 and 380 m. Literature. Garman (1899: 79), Caruso (1981: 535), Lea et al. (1984: 250), Allen & Robertson (1994: 81), Bussing & López (1994: 68, 2005: 44; 2009: 448), Balart et al. (1995: 82), Caruso (1995: 1229), Bearez (1996: 734), Castro-Aguirre & Balart (1996: 71), Watson (1996a: 553), De La Cruz Agüero et al. (1997: 210), Grove & Lavenberg (1997: 230), Chirichigno & Vélez (1998: 222), Castellanos-Galindo et al. (2006a: 199), Rodríguez- Romero et al. (2008: 1772), Hooker (2009a: 204), López-Martínez et al. (2012: 352), Del Moral-Flores et al. (2013: 193), Fuentes et al. (2015: 615) and Robertson & Allen (2015: 761). FIGURE 55. Lophiodes caulinaris. Dorsal view. (60) Lophiodes spilurus (Garman, 1899). Threadfin angler (En); Rape de rabo delgado (Sp) MOP stations. 4; 007, 038, 039 and 057; Depth range: 144–478 m; USNM voucher specimens: 6; 421229, 421230, 421281, 421348, 421427 and 422527. Distribution. Eastern Pacific (from southern California, USA, to Chile, including the Cocos and Galápagos Islands), occurring at depths between 80 and 850 m.ROBERTSON ET AL .64 · Zootaxa 4348 (1) © 2017 Magnolia Press Literature. Garman (1899: 77), Caruso (1981: 536), Lea et al. (1984: 250), Pequeño (1989: 43), van der Heiden & González (1990: 147), Bussing & López (1994: 68, 2005: 44; 2009: 448), Balart et al. (1995: 82), Caruso (1995: 1230), Bearez (1996: 734), Castro-Aguirre & Balart (1996: 71), Watson (1996a: 553), De La Cruz Agüero et al. (1997: 211), Grove & Lavenberg (1997: 231), Chirichigno & Vélez (1998: 222), Castellanos-Galindo et al. (2006a: 199), Cortés & Blum (2008: 194), Rodríguez-Romero et al. (2008: 1772), Hooker (2009a: 205), López-Martínez et al. (2012: 352), Del Moral-Flores et al. (2013: 193), Fuentes et al. (2015: 615), and Robertson & Allen (2015: 2598). FIGURE 56. Lophiodes spilurus. Top: dorsal view; Bottom: detail of head, lateral view. Antennariidae Frogfishes (En); Pejes sapo, Ranisapos, Zanahorias (Sp). 1 G, 1 S. (61) Fowlerichthys avalonis (Jordan & Starks, 1907). Roughbar frogfish (En); Ranisapo ocelado (Sp) MOP stations. 1; 049; Depth range: 128–136 m; USNM voucher specimens: 1; 422343. Distribution. Eastern Pacific (from southern California to Peru, including the Cocos and Galápagos Islands), occurring at depths between 0 and 300 m. Literature. Jordan & Starks (1907: 76), Böhlke (1953: 147), Eschmeyer et al. (1983: 112), Pietsch (1984: 36), Pietsch & Grobecker (1987: 125), Allen & Robertson (1994: 82), Bussing & López (1994: 68; 2009: 449), Schneider & Lavenberg (1995: 856), Bearez (1996: 734), Watson (1996: 359), De La Cruz Agüero et al. (1997: 52), Grove & Lavenberg (1997: 234), Watson (1998: 219), Chirichigno & Vélez (1998: 221), Thomson et al. (2000: 55), Castellanos-Galindo et al. (2006a: 199), Rodríguez-Romero et al. (2008: 1772), Hooker (2009b: 206), McCosker & Rosenblatt (2010: 190), López-Martínez et al. (2012: 352), Del Moral-Flores et al. (2013: 193), Murase et al. (2014: 1405) and Robertson & Allen (2015: 767). Zootaxa 4348 (1) © 2017 Magnolia Press · 65TEP DEEP FISHES FIGURE 57. Fowlerichthys avalonis; USNM 422343, 80 mm. SL. Ogcocephalidae Batfishes (En); Peces murcielago (Sp). 2 G, 6 S. (62) Dibranchus erinaceus (Garman, 1899). MOP stations. 4; 007, 031, 052 and 060; Depth range: 470–1030 m; USNM voucher specimens: 7; 421226, 421253, 421256, 421396, 422525, 422554 and 422572. Distribution. Eastern Pacific (from Costa Rica to Panama, and the Cocos and Galápagos Islands), occurring at depths between 470 and 1150 m (see Remarks). Literature. Garman (1899: 103), Bradbury (1962: 3; 1967: 414; 1999: 290; 2003: 3), Bradbury et al. (1999: 81), Castellanos-Galindo et al. (2006a: 200), Ho et al. (2009: 394) and McCosker & Rosenblatt (2010: 190). FIGURE 58. Dibranchus erinaceus. Dorsal view; USNM , 80 mm. SL. Remarks. USNM 421396, 422525, 422554 and 422572 specimens represent the first documented record of the species in Costa Rican waters (Bussing & López 1994, 2009, 2011; Bradbury 1999) and a northwestern range ROBERTSON ET AL .66 · Zootaxa 4348 (1) © 2017 Magnolia Press extension of about 520 km in the species’ known distribution (Garman 1899, Bradbury 1999). The previous northernmost documented record for this species in the eastern Pacific was from off the Peninsula de Azuero, Panama (Garman 1899, Bradbury 1999). In addition, specimens collected at 470 m represent a new minimum depth record for the species (Bradbury 1999, Froese & Pauly 2016). (63) Dibranchus hystrix Garman, 1899 MOP stations. 3; 062, 073 and 082; Depth range: 153–1527 m; USNM voucher specimens: 5; 421279, 421467, 422413, 422601 and 422606. Distribution. Eastern Pacific (from the mouth of Gulf of California, Mexico, to Peru, including the Cocos and Galápagos Islands) and South China Sea, occurring at depths between 153 and 2323 m (see Remarks). Literature. Garman (1899: 92), Bradbury (1962: 2; 1999: 280), Castro-Aguirre & Balart (1996: 71), Bradbury et al. (1999: 86), Bradbury (2003: 3), Castellanos-Galindo et al. (2006a: 200), Shimazaki (2009a: 208) and McCosker & Rosenblatt (2010: 190). Remarks. Specimens collected at 153 m represent a new minimum depth record for the species (Bradbury 1999, Shimazaki 2009a, Froese & Pauly 2016). FIGURE 59. Dibranchus hystrix. Top: dorsal view; Bottom: ventral view. Zootaxa 4348 (1) © 2017 Magnolia Press · 67TEP DEEP FISHES (64) Dibranchus nudivomer (Garman, 1899) MOP stations. 2; 025 and 029; Depth range: 1101–1314 m; USNM voucher specimens: 3; 422524, 422534 and 422578. Distribution. Eastern Pacific (from southern Baja California, Mexico to Peru), occurring at depths between 605 and 1457 m. Literature. Garman (1899: 99), Bradbury (1962: 2; 1967: 414; 1999: 292; 2003: 4), Castro-Aguirre & Moncayo-López (1976: 307), Balart et al. (1995: 82), Castro-Aguirre & Balart (1996: 71), Castellanos-Galindo et al. (2006a: 200) and Shimazaki (2009a: 209). FIGURE 60. Dibranchus nudivomer. Top: dorsal view; Bottom: ventral view; USNM 422534, 113 mm. SL. (65) Dibranchus spinosus (Garman, 1899) MOP stations. 7; 025, 028, 040, 052, 063, 078 and 082; Depth range: 836–1625 m; USNM voucher specimens: 9; 421340, 421349, 421386, 421395, 421470, 422385, 422595, 422618 and 422625. Distribution. Eastern Pacific (from the mouth of Gulf of California, Mexico, to off the southern tip of Peru) occurring at depths between 366 and 2324 m. Literature. Garman (1899: 104), Bradbury (1962: 4; 1967: 414; 1999: 289; 2003: 4), Bussing (1965: 222), Bearez (1996: 734), Pequeño (1997: 82), Shimazaki (2009a: 211), Castellanos-Galindo et al. (2006a: 200) and Ho et al. (2009: 394).ROBERTSON ET AL .68 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 61. Dibranchus spinosus. Top: dorsal view; Bottom: ventral view; USNM 422625, 140 mm. SL. (66) Dibranchus velutinus Bradbury, 1999 MOP stations. 3; 067, 071 and 088; Depth range: 570–965 m; USNM voucher specimens: 7; 421509, 422363, 422391, 422409, 422420, 422453 and 422459. Distribution. Eastern Pacific (from Nicaragua to Peru), occurring at depths between 450 and 965 m (see Remarks). Literature. Bradbury (1999: 285). Remarks. These specimens represent the first documented record of the species in Central American waters (Bussing & López 1994, 2009, 2011; Bradbury 1999) and a northern range extension of about 2000 km in the species’ known distribution (Bradbury 1999). The previous northernmost documented record for this species in the eastern Pacific was from northern Peru (Bradbury 1999). In addition, specimens collected at 965 m represent a new maximum depth record for the species (Bradbury 1999, Froese & Pauly 2016). Zootaxa 4348 (1) © 2017 Magnolia Press · 69TEP DEEP FISHES FIGURE 62. Dibranchus velutinus. Top: dorsal view; Bottom: ventral view; USNM 422363, 60 mm. SL. (67) Zalieutes elater (Jordan & Gilbert, 1882). Roundel batfish (En); Murcielago biocelado (Sp) MOP stations. 3; 004, 015 and 049; Depth range: 113–136 m; USNM voucher specimens: 6; 421219, 421227, 422352, 422354, 422502 and 422510. Distribution. Eastern Pacific (from California, USA, to Peru, Including the Cocos Island), occurring at depths between 18 and 160 m. Literature. Jordan & Gilbert (1882: 365), Bradbury (1967: 417; 2003: 11), Eschmeyer et al. (1983: 112), Allen & Robertson (1994: 84), Bussing & López (1994: 68), Bearez (1996: 734), Watson (1996c: 563), De La Cruz Agüero et al. (1997: 241), Chirichigno & Vélez (1998: 222), Bussing & López (2005: 48; 2009: 449), Castellanos- Galindo et al. (2006a: 200), Cortés & Blum (2008: 194), Rodríguez-Romero et al. (2008: 1772), Shimazaki (2009a: 212), Del Moral-Flores et al. (2013: 194) and Robertson & Allen (2015: 779).ROBERTSON ET AL .70 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 63. Zalieutes elater. Dorsal view. Caulophrynidae Fanfins, Anglerfishes (En); Peces pescadores (Sp). 1 G, 1 S. (68) Caulophryne pelagica (Brauer, 1902) MOP stations. 1; 030; Depth range: 864–1026 m; USNM voucher specimens: 1; 422598. Distribution. Circumglobal, known from widely scattered records across the Indo-Pacific, with a single record from the Atlantic (Cape Verde Islands), occurring at depths between 500 and 3500 m. In the eastern Pacific this species has been recorded in the Guadalupe Island, México, the Gulf of Panama and off Peru. Literature. Brauer (1902: 295), Pietsch (1979:14; 2009a: 213; 2009d: 445), Stewart & Pietsch (1998: 5), Nakabo (2000: 467; 2002: 467), Mundy (2005: 269), Castellanos-Galindo et al. (2006a: 200) and Reyes-Bonilla et al. (2011: 7). Melanocetidae Black seadevils (En); Peces pescadores, Demonios de mar (Sp). 1 G, 1 S. (69) Melanocetus johnsoni Günther, 1864. Humpback anglerfish, Blackdevil (En); Rape jorobado, Diablo negro (Sp) MOP stations. 6; 053, 077, 078, 082, 086 and 099; Depth range: 1054–1527 m; USNM voucher specimens: 8; 421406, 421457, 421459, 422319, 422465, 422628, 423214 and 423215. Distribution. Circumglobal in tropical through cold temperate seas, occurring at depths between 100 and 4500 m. In the eastern Pacific this species has been recorded from off Canada, USA, México, Panama, Colombia, Peru and Chile. Zootaxa 4348 (1) © 2017 Magnolia Press · 71TEP DEEP FISHES Literature. Günther (1864: 302), Fitch & Lavenberg (1968: 127), Bussing (1965: 222), Paxton et al. (1989: 287), Pequeño (1989: 43), Paulin et al. (1989: 140), McAllister (1990: 232), Castro-Aguirre & Balart (1996: 71), Watson (1996d: 571), Santos et al. (1997: 64), Arruda (1997: 126), Stewart & Pietsch (1998: 5), McEachran & Fechhelm (1998: 865), Fricke (1999: 110), Tru11 (1999: 464), Nakabo (2000: 474; 2002: 474), Anderson & Leslie (2001:4), Moore et al. (2003: 214), Menezes et al. (2003: 65), Mundy (2005: 269), Castellanos-Galindo et al. (2006a: 201), Pietsch (2009b: 214; 2009d: 361), Møller et al. (2010: 45) and Parin et al. (2014: 186). Remarks. These specimens represent the first documented record of the species in upper Central American waters (Bussing & López 1994, 2009, 2011; Pietsch 2009b, 2009d; Froese & Pauly 2016). FIGURE 64. Melanocetus johnsoni. Top: entire specimen; USNM 421406, 50 mm. SL; Bottom: detail of head; USNM 423216, 60 mm. SL. Oneirodidae Dreamers (En); Peces pescadores, Soñadores (Sp). 1 G, 2 S. (70) Oneirodes luetkeni (Regan, 1925). Lütken’s dreamer (En); Soñador de Lütken (Sp) MOP stations. 5; 001, 046, 071, 087 and 096; Depth range: 165–1157 m; USNM voucher specimens: 6; 421223, 421353, 421514, 422335, 422393 and 422511.ROBERTSON ET AL .72 · Zootaxa 4348 (1) © 2017 Magnolia Press Distribution. Eastern Pacific (from the Gulf of California, Mexico, to Peru), occurring at depths between 165 and 1750 m (see Remarks). FIGURE 65. Oneirodes luetkeni; USNM 421514, 50 mm. SL. Literature. Regan (1925: 562), Nielsen (1974: 94), Orr (1991: 1026), Castro-Aguirre & Balart (1996: 71), Castellanos-Galindo et al. (2006a: 202) and Pietsch (2009c: 224; 2009d: 397). Remarks. Specimens collected at 165 m represent a new minimum depth record for the species (Castellanos- Galindo et al. 2006a; Pietsch 2009c, 2009d; Froese & Pauly 2016). (71) Oneirodes rosenblatti Pietsch, 1974. Rosenblatt’s dreamer (En); Soñador de Rosenblatt (Sp) MOP stations. 1; 058; Depth range: 1181–1270 m; USNM voucher specimens: 1; 423219. Distribution. Eastern Pacific (from off Costa Rica to Chile), occurring at depths between 750 and 3000 m (see Remarks). Literature. Pietsch (1974: 41; 2009c: 226; 2009d: 399), Orr (1991: 1027) and Castellanos-Galindo et al. (2006a: 202). Remarks. This specimen represents the first documented record of the species in Costa Rican waters (Bussing & López 1994, 2009, 2011; Pietsch 2009d) and a northwestern range extension of about 700 km in the species’ known distribution (Pietsch 2009d). The previous northernmost documented record for this species in the eastern Pacific was from the Gulf of Panama (Castellanos-Galindo et al. 2006a, Pietsch 2009d). STEPHANOBERYCIFORMES 1 F, 2 G, 3 S. Melamphaidae Bigscale fishes (En); Peces de escama grande (Sp). 2 G, 3 S. (72) Melamphaes sp.1 MOP stations. 4; 067, 071, 075 and 083; Depth range: 784–1116 m; USNM voucher specimens: 8; 421265, 421267, 421503, 421504, 421505, 422348, 422386 and 422390. Literature. Kotlyar (1996: 272). Remarks. We were not able to make a specific identification of these specimens; however, despite the apparent absence of external diagnostical features between them, two different forms (Melamphaes sp.1 and Melamphaes sp.2) are herein listed based on molecular differences (2.67–2.99% divergence in COI sequences). (73) Melamphaes sp.2 MOP stations. 1; 077; Depth range: 1320–1370 m; USNM voucher specimens: 2; 422342 and 422532. Literature. Kotlyar (1996: 272). Remarks. see remarks about Melamphaes sp 1. Zootaxa 4348 (1) © 2017 Magnolia Press · 73TEP DEEP FISHES FIGURE 66. Melamphaes sp. 2. (74) Poromitra crassiceps (Günther, 1878). Crested bigscale (En); Escama-grande crestado (Sp) MOP stations. 1; 001; Depth range: 165–183 m; USNM voucher specimens: 2; 422301 and 422334. Distribution. Possibly circumglobal in tropical through cold temperate seas, occurring at depths between 164 and 2370 m. In the eastern Pacific this species has been recorded from off Canada, USA, including Alaska, México, including the Gulf of California, and Chile. Literature. Chapman (1939: 535), Bussing (1965: 213), Fitch & Lavenberg (1968: 93), Robison (1972: 451), Castro-Aguirre & Balart (1996: 71), Sandknop & Watson (1996: 692), Pequeño (2000: 69), Mecklenburg et al. (2002: 319) and Kotlyar (2008: 490). Remarks. Valid, in the eastern Pacific as Poromitra rugosa (Chapman, 1939) following Kotlyar (2008) and Eschmeyer et al. (2017), but listed here as P. crassiceps (Günther, 1878; restricted to the Atlantic Ocean by the latter authors), following Bussing (1965), Fitch & Lavenberg (1968), Robison (1972), Castro-Aguirre & Balart (1996), Sandknop & Watson (1996), Pequeño (2000) and Mecklenburg et al. (2002). This decision was made due to a lack of significant differences in morphometric and meristic characters between the present specimens and Atlantic specimens of P. crassiceps examined by the senior author. Additionally, COI sequences for both eastern Pacific (present study) and eastern Atlantic populations available in BOLD (BOLD process ID ME-7222 and ME- 7812) are not significantly different (typically less than 1.5% difference). This combination of morphological and genetic data suggests the existence of a single lineage comprising both “species.” The USNM 422301 and 422334 specimens represent the first documented record of P. crassiceps (or P. rugosa) in Central American waters (Bussing & López 1994, 2009, 2011; Kotlyar 2008; Froese & Pauly 2016). FIGURE 67. Poromitra crassiceps. BERYCIFORMES 2 F, 2 G, 2 S. Trachichthyidae Roughies, Slimy heads (En); Relojes (Sp). 1 G, 1 S.ROBERTSON ET AL .74 · Zootaxa 4348 (1) © 2017 Magnolia Press (75) Hoplostethus mento Garman, 1899. Slimy head (En); Cabeza viscosa (Sp) MOP stations. 1; 003; Depth range: 116–118 m; USNM voucher specimens: 5; 421260, 421268, 421284, 421306 and 421317. Distribution. Eastern Pacific (Costa Rica to Chile), occurring at depths between 300 and 1350 m. Literature. Garman (1899: 58), Bussing (1965: 214), Kotlyar (1986: 136; 1996:171), Pequeño (1989: 53), Iwasaki (2009b: 236), Angulo (2014: 1) and Robertson & Allen (2015: 5310). FIGURE 68. Hoplostethus mento. Anoplogastridae Fangtooths. (En); Colmilludos (Sp). 1 G, 1 S. (76) Anoplogaster cornuta (Valenciennes, 1833). Common fangtooth (En); Pez abisal con colmillos (Sp) FIGURE 69. Anoplogaster cornuta. Top: entire specimen; Bottom: detail of head; USNM 421537, 105 mm. SL. Zootaxa 4348 (1) © 2017 Magnolia Press · 75TEP DEEP FISHES MOP stations. 5; 002, 003, 005, 028 and 046; Depth range: 116–1472 m; USNM voucher specimens: 5; 421295, 421537, 422533, 422546 and 422626. Distribution. Circumglobal, occurring at depths between 75 and 5000 m. In the eastern Pacific this species has been previously recorded from off Alaska, USA, to Chile. Literature. Valenciennes (1833b: 470), Bussing (1965: 213), Fitch & Lavenberg (1968: 94), Bauchot (1970: 10), Uyeno et al. (1983:271), Masuda et al. (1984: 109), Nakamura et al. (1986: 166), Whitehead et al. (1986: 767), Scott & Scott (1988: 324), Matarese et al. (1989: 250), Paulin et al. (1989: 154), Pequeño (1989: 53), Castro- Aguirre & Balart (1996: 71), Kotlyar (1996: 238), Shinohara et al. (1996: 174), Watson (1996e: 683), Arruda (1997: 78), Santos et al. (1997: 71), McEachran & Fechhelm (1998: 998), Porteiro et al. (1999: 41), Konishi (1999: 33), Nakabo (2000: 504; 2002: 504), Mecklenburg et al. (2002: 32), Menezes et al. (2003: 69), Moore et al. (2003: 220), Mundy (2005: 303), Iwasaki (2009a: 235), Møller et al. (2010: 49), Fricke et al. (2011: 372), Reyes-Bonilla et al. (2011: 8), Maslenikov et al. (2013: 8) and Parin et al. (2014: 202). SCORPAENIFORMES 6 F, 7 G, 12 S. Sebastidae Scorpionfishes (En); Peces roca; Chancharros (Sp). 1 G, 1 S. (77) Sebastolobus alascanus Bean, 1890. Shortspine thornyhead (En); Chancharro alacrán (Sp) MOP stations. 1; 058; Depth range: 1181–1270 m; USNM voucher specimens: 1; 423201. Distribution. North Pacific (from the sea of Okhotsk north to the Navarin Canyon in the Bering Sea and from Stalemate Bank and Ulm Plateau in the Aleutian Islands southeast to Cedros Island, Baja California, Mexico; also reported from Japan); vagrants to Costa Rica; occurring at depths between 17 and 1600 m. FIGURE 70. Sebastolobus alascanus. Top: entire specimen; Bottom: detail of head; USNM 423201, 280 mm. SL. ROBERTSON ET AL .76 · Zootaxa 4348 (1) © 2017 Magnolia Press Literature. Bean (1890: 44), Eschmeyer et al. (1983: 152), Amaoka et al. (1983: 279), Masuda et al. (1984: 316), McAllister (1990: 184), Orr et al. (2000: 20), Nakabo (2000: 570, 2002: 570), Love et al. (2002: 115), Mecklenburg et al. (2002: 347), Fedorov et al. (2003: 60), Parin et al. (2014: 223) and Robertson & Allen (2015: 5437) Remarks. This specimen represents the first documented record of the genus and the species in Central American waters (Bussing & López 1994, 2009, 2011; Orr et al. 2000; Love et al. 2002; Froese & Pauly 2016) and a southeastern range extension of about 2900 km in the species’ known distribution (Orr et al. 2000, Love et al. 2002). The previous southernmost documented record for this genus and species in the eastern Pacific was from off Baja California, Mexico (Orr et al. 2000, Love et al. 2002). Scorpaenidae Scorpionfishes (En); Peces escorpion, Peces piedra (Sp). 1 G, 3 S. (78) Pontinus furcirhinus Garman, 1899. Red scorpionfish (En); Rascacio richichi, Lapón rojo, Pez escorpión rojo (Sp) MOP stations. 1; 004; Depth range: 115–116 m; USNM voucher specimens: 1; 422640. Distribution. Eastern Pacific (from southern Baja California, Mexico, to Peru, including the Cocos, Malpelo and Galápagos Islands) occurring at depths between 50 and 390 m. Literature. Garman (1899: 51), Poss (1995: 1547), Bussing & López (1994: 82; 2009: 452), Bearez (1996: 735), Moser (1996c: 738), Grove & Lavenberg (1997: 295), Chirichigno & Vélez (1998: 272), Rodríguez-Romero et al. (2008: 1773), Imamura (2009: 240), López-Martínez et al. (2012: 352), Del Moral-Flores et al. (2013: 195), Gómez et al. (2014: 394) and Robertson & Allen (2015: 1007). FIGURE 71. Pontinus furcirhinus; USNM 422640, 215 mm. SL. (79) Pontinus sierra (Gilbert, 1890). Speckled scorpionfish (En); Lapón manchado (Sp) MOP stations. 4; 004, 038, 049 and 068; Depth range: 115–154 m; USNM voucher specimens: 11; 421376, 421552, 422336, 422349, 422350, 422443, 422471, 422473, 422474, 422580 and 422611. Distribution. Eastern Pacific (from Baja California, Mexico, to Peru), occurring at depths between 48 and 351 m. Literature. Gilbert (1890: 82), Bussing & López (1994: 82; 2009: 452), Poss (1995: 1547), Bearez (1996: 735), Castro-Aguirre & Balart (1996: 71), Moser (1996c: 738), Grove & Lavenberg (1997: 295), Chirichigno & Vélez (1998: 271), Pedraza et al. (2002: 69), Rodríguez-Romero et al. (2008: 1773), Imamura (2009: 241), Zootaxa 4348 (1) © 2017 Magnolia Press · 77TEP DEEP FISHES Palacios-Salgado & Ramirez-Valdez (2011: 93), Del Moral-Flores et al. (2013: 195) and Robertson & Allen (2015: 4432). FIGURE 72. Pontinus sierra. Top: adult; Bottom: juvenile. (80) Pontinus sp. Rosy scorpionfish (En); Puñal rosado (Sp) MOP stations. 5; 015, 068, 081, 101 and 102; Depth range: 113–399 m; USNM voucher specimens: 15; 421247, 421270, 421323, 421384, 421398, 421430, 422306, 422309, 422327, 422340, 422369, 422382, 422389, 422414 and 422440. FIGURE 73. Pontinus sp. Distribution. Eastern Pacific (from southern Baja and the mouth of the Gulf of California to northern Peru), occurring at depths between 50 and 399 m (see Remarks).ROBERTSON ET AL .78 · Zootaxa 4348 (1) © 2017 Magnolia Press Literature. Robertson & Allen (2015: 1011, as Pontinus sp. A). Remarks. This appears to be an undescribed species currently under investigation by the authors. Specimens collected at 399 m represent a new maximum depth record for the species (Robertson & Allen 2015). Triglidae Gurnards, Searobins (En); Cabros, Malarmados, Rubios, Cabrillas (Sp). 2 G, 3 S. (81) Bellator loxias (Jordan, 1897). Barred searobin, Chevron searobin (En); Vaca angelita (Sp) MOP stations. 1; 015; Depth range: 113–114 m; USNM voucher specimens: 3; 421214, 421274 and 422360. Distribution. Eastern Pacific (southern Baja California, Mexico, including the Gulf of California and the Revillagigedos Islands to Isla San Lorenzo, Callao, Peru, including the Cocos and Galápagos Islands), occurring at depths between 20 and 191 m. Literature. Jordan (1897: 452), Garman (1899: 108), Gruchy (1970: 526), Chirichigno (1974: 207), Miller & Richards (1991a: 645, 1991b: 761), Bussing & López (1994: 86; 2009: 452), Castro-Aguirre et al. (1993:89), Bussing (1995: 1644), Richards (1996: 798), Bearez (1996: 735), Grove & Lavenberg (1997: 299), Chirichigno & Vélez (1998: 264), Rodríguez-Romero et al. (2008: 1773), Del Moral-Flores et al. (2013: 196) and Robertson & Allen (2015: 1050). FIGURE 74. Bellator loxias. (82) Prionotus ruscarius Gilbert & Starks, 1904. Common searobin, Rough searobin (En); Cabro áspero, Rubio gallineta, Vaca rasposa (Sp) MOP stations. 3; 006, 015 and 049; Depth range: 109–136 m; USNM voucher specimens: 4; 421213, 421224, 421264 and 422300. Distribution. Eastern Pacific (from southern Baja California, Mexico, including the Gulf of California to Chile), occurring at depths between 4 and 136 m (see Remarks). Literature. Gilbert & Starks (1904: 165), Böhlke (1953: 125), Pequeño (1989: 57), Miller & Richards (1991b: 760), Castro-Aguirre et al. (1993: 88), Bussing & López (1994: 86; 2009: 453), Balart et al. (1995: 82), Bussing (1995: 1645), Bearez (1996: 735), Richards (1996: 797), De La Cruz Agüero et al. (1997: 300), Chirichigno & Vélez (1998: 265), Castro-Aguirre et al. (1999: 246), Lucano-Ramírez et al. (2005: 1), Raymundo-Huizar et al. (2008: 7), Rodríguez-Romero et al. (2008: 1773), López-Martínez et al. (2012: 352), Del Moral-Flores et al. (2013: 196), Murase et al (2014: 1405), Fuentes et al. (2015: 615) and Robertson & Allen (2015: 1058). Remarks. Specimens collected at 136 m represent a new maximum depth record for the species (Bussing & López 1994, Robertson & Allen 2015). Zootaxa 4348 (1) © 2017 Magnolia Press · 79TEP DEEP FISHES FIGURE 75. Prionotus ruscarius. Top: lateral view; Bottom: dorsal view. (83) Prionotus stephanophrys Lockington, 1881. Lumptail searobin, Blackfin searobin (En); Vaca voladora (Sp) MOP stations. 2; 015 and 091; Depth range: 105–114 m; USNM voucher specimens: 6; 421215, 421216, 421234, 421243, 421244 and 422347. Distribution. Eastern Pacific (from off Washington, USA to Chile, but rare north of Baja California, Mexico, including the Malpelo and Galápagos Islands), occurring at depths between 2 and 255 m. Literature. Lockington (1881: 529), Eschmeyer et al. (1983: 154), Pequeño (1989: 57), Miller & Richards (1991: 760), Schmitter-Soto & Castro-Aguirre (1996: 23; 2015: 213), Castro-Aguirre et al. (1993: 89), Allen & Robertson (1994: 103), Bussing & López (1994: 84; 2009: 453), Bussing (1995: 1644), Bearez (1996: 735), Franke & Acero (1996: 766), Richards (1996: 797), De La Cruz Agüero et al. (1997: 300), De la Cruz-Agüero & Cota- Gómez (1998: 357), Richards & McCosker (1998: 940), Chirichigno & Vélez (1998: 264), Castro-Aguirre et al. (1999: 245), Lea & Rosenblatt (2000: 121), Rodríguez-Romero et al. (2008: 1773), Kawai (2009a: 247), McCosker & Rosenblatt (2010: 191), López-Martínez et al. (2012: 352), Del Moral-Flores et al. (2013: 196), Gómez et al. (2014: 394), Murase et al (2014: 1405), Fuentes et al. (2015: 615) and Robertson & Allen (2015: 1059).ROBERTSON ET AL .80 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 76. Prionotus stephanophrys. Peristediidae Armored Sea-Robins, Armored-Gurnards (En); Peces Cocodrilos, Malarmados, Cabros, Caballitas (Sp). 1 G, 1 S. (84) Peristedion barbiger Garman, 1899. Barbel’s crocodile, Bearded armored-searobin (En); Pez cocodrilo, caballito moro (Sp) MOP stations. 3; 006, 015 and 068; Depth range: 109–145 m; USNM voucher specimens: 7; 421211, 421212, 421319, 421334, 422314, 422320 and 422329. Distribution. Eastern Pacific (from Costa Rica to northern Peru), occurring at depths between 50 and 295 m. Literature. Garman (1899: 110), Bussing & López (1994: 70; 2009: 452), Richards (1996: 798), Richards & McCosker (1998: 941), Chirichigno & Vélez (1998: 263), Kawai (2008: 24; 2009b: 248), Bussing (2010: 1149), Tenda & Kawai (2012: 135) and Robertson & Allen (2015: 2635). FIGURE 77. Peristedion barbiger. Top: lateral view; Lower left: dorsal view of head; Lower right: ventral view of head. Psychrolutidae Fathead sculpins (En); Cabezas gordas (Sp). 1 G, 1 S. Zootaxa 4348 (1) © 2017 Magnolia Press · 81TEP DEEP FISHES (85) Psychrolutes cf. sio Nelson, 1980. South American sculpin (En); Pez chancho (Sp) MOP stations. 6; 028, 029, 030, 082, 087 and 096; Depth range: 864–1527 m; USNM voucher specimens: 6; 421511, 422353, 422602, 422603, 422651 and 423597. Distribution. Eastern Pacific (from off El Salvador, to Chile), occurring at depths between 768 and 1527 m (see Remarks). Literature. Nelson (1980: 444; 1995: 69), Pequeño (1989: 57), Fricke (1990: 407) and Yabe (2009c: 250). Remarks. These specimens were tentatively identified by us as Psychrolutes cf. sio because they shared with this species the following combination of distinctive characters, sensu Nelson 1980: jaws equal; cirri on head and scattered; lateral line pores in small tubes; length of two inner pelvic-fin rays about equal, outer only slightly shorter; and dorsal-fin elements VIII, 17–18. Our specimens, however, differ from P. sio, sensu stricto, by having a shallower body (20.29–24.41% of SL, vs. 28.2–33.2% of SL); a shorter eye diameter (5.17–6.68% of SL, vs. 7.90– 9.90% of SL); a shorter pectoral fin (22.17–25.76% of SL, vs. 28.20–28.30% of SL); a shorter pelvic fin (7.12– 8.53% of SL, vs. 14.10–14.40% of SL); a greater inter-pelvic distance (3.12–5.11% of SL, vs. 2.70–2.40% of SL); fewer anal-fin elements (12–13, vs. 14); and a different coloration pattern (greyish with small pale spots, vs. uniformly brown). COI sequences for our specimens of P. cf. sio and five specimens of P. macrocephalus (Gilchrist 1904) from South Africa and available in BOLD (ADC11_160.4 #1 to #5) are less than 2% divergent. Assuming these differences represent intraspecific variation in P. sio, our specimens represent the first documented record of the species in Central American waters (Bussing & López 1994, 2009, 2011; Nelson 1980,1995; Yabe 2009) and a northward range extension of about 2100 km in the species’ known distribution (Nelson 1980, Yabe 2009). The previous northernmost documented record for this species in the eastern Pacific was from northern Peru (Nelson 1980, Yabe 2009). In addition, specimens collected at 1527 m represent a new maximum depth record for the species (Nelson 1980, Froese & Pauly 2016). FIGURE 78. Psychrolutes cf. sio. Top: entire specimen; Bottom: detail of head; USNM 422603, 205 mm. SL. Liparidae Snailfishes (En); Peces babosos (Sp). 1 G, 3 S. (86) Paraliparis sp.1 MOP stations. 4; 029, 062, 073 and 077; Depth range: 153–1457 m; USNM voucher specimens: 5; 421548, 422455, 422585, 422607 and 422647. Literature. Stein (1978: 37), Andriashev (1986: 1) and Stein & Chernova (2002: 153). Remarks. We were not able to make a specific identification of these specimens; however, despite the ROBERTSON ET AL .82 · Zootaxa 4348 (1) © 2017 Magnolia Press apparent absence of external diagnostical features between them, three different forms (Paraliparis sp.1, Paraliparis sp.2 and Pachycara sp. 3) are herein listed based on molecular differences (2.52–3.66% divergence in COI sequences). FIGURE 79. Paraliparis sp.1. Top: entire specimen; Bottom: detail of head; USNM 422647, 305 mm. SL. (87) Paraliparis sp.2 MOP stations. 5; 042, 073, 078, 086 and 096; Depth range: 153–1353 m; USNM voucher specimens: 6; 421269, 422426, 422429, 422454, 422466 and 422592. Literature. Stein (1978: 37), Andriashev (1986: 1) and Stein & Chernova (2002: 153). Remarks. See Remarks about Paraliparis sp 1. FIGURE 80. Paraliparis sp.2. Top: entire specimen; Bottom: detail of head; USNM 422426, 190 mm. SL. Zootaxa 4348 (1) © 2017 Magnolia Press · 83TEP DEEP FISHES (88) Paraliparis sp.3 MOP stations. 1; 077; Depth range: 1320–1370 m; USNM voucher specimens: 1; 422316. Literature. Stein (1978: 37), Andriashev (1986: 1) and Stein & Chernova (2002: 153). Remarks. See remarks about Paraliparis sp 1. PERCIFORMES 14 F, 25 G, 32 S. Serranidae Sea basses (En); Meros, Cabrillas (Sp). 4 G, 4 S. (89) Baldwinella eos Gilbert, 1890. Bigeye bass (En); Serrano ojón (Sp) MOP stations. 3; 004, 049 and 068; Depth range: 115–145 m; USNM voucher specimens: 7; 421225, 422325, 422333, 422344, 422436, 422558 and 422589. Distribution. Eastern Pacific (from Baja California, Mexico, including the Gulf of California, to Peru, including the Cocos Island) and western Atlantic (from the Gulf of Mexico to Suriname), occurring at depths between 80 and 325 m. Literature. Gilbert (1890: 62), Uyeno et al. (1983: 313), Baldwin (1990: 92), Bussing & López (1994: 96; 2009: 454), Balart et al. (1995: 81), Heemstra (1995: 1572), Castro-Aguirre & Balart (1996: 71), Watson (1996f: 878), Grove & Lavenberg (1997: 338), Anderson & Baldwin (2000: 379), Rodríguez-Romero et al. (2008: 1774), Romero & Vélez (2009a: 261), Palacios-Salgado & Ramirez-Valdez (2011: 93), Anderson & Heemstra (2012: 112), Del Moral-Flores et al. (2013: 197) and Robertson & Allen (2015: 1107). FIGURE 81. Baldwinella eos. (90) Hemanthias signifer (Garman, 1899). Damsel bass (En); Cabrilla doncella (Sp) MOP stations. 4; 001, 004, 015 and 068; Depth range: 113–183 m; USNM voucher specimens: 6; 421237, 421551, 422478, 422535, 422573 and 422633. Distribution. Eastern Pacific (from southern California, USA, including the Gulf of California, to northern Peru, including the Malpelo Island), occurring at depths between 23 and 265 m. Literature. Garman (1899: 48), Eschmeyer et al. (1983: 199), Baldwin (1990: 936), Allen & Robertson (1994: 110), Bussing & López (1994: 96; 2009: 454), Heemstra (1995: 1572), Bearez (1996: 736), Castro-Aguirre & Balart (1996: 71), Watson (1996f: 878), De La Cruz Agüero et al. (1997: 287), Chirichigno & Vélez (1998: 353), Anderson & Baldwin (2000: 379), Thomson et al. (2000: 77), Romero & Vélez (2009a: 258), Anderson & Heemstra (2012: 56), López-Martínez et al. (2012: 352), Del Moral-Flores et al. (2013: 196) and Robertson & Allen (2015: 1105).ROBERTSON ET AL .84 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 82. Hemanthias signifer. (91) Hyporthodus niphobles (Gilbert & Starks, 1897). Star-studded grouper (En); Mero gris, Mero manchado (Sp) MOP stations. 2; 036 and 066; Depth range: 127–143 m; USNM voucher specimens: 3; 422476, 422641 and 423181. Distribution. Eastern Pacific (from southern California, USA, including the Gulf of California, to northern Peru), occurring at depths between 1 and 450 m. Literature. Gilbert & Starks (1897: 442), Heemstra & Randall (1993: 203), Allen & Robertson (1994: 109), Bussing & López (1994: 92; 2009: 454), Heemstra (1995: 1576), Bearez (1996: 736), Watson (1996f: 878), De La Cruz Agüero et al. (1997: 285), Grove & Lavenberg (1997: 323), Chirichigno & Vélez (1998: 354), Smith-Vaniz et al. (1999: 211), Lea & Rosenblatt (2000: 121), Thomson et al. (2000: 83), Craig & Hastings (2007: 17), Romero & Vélez (2009a: 257), McCosker & Rosenblatt (2010: 191), Craig et al. (2011: 278), Del Moral-Flores et al. (2013: 196), Murase et al. (2014: 1405) and Robertson & Allen (2015: 1127). FIGURE 83. Hyporthodus niphobles, USNM 423181, 265 mm. SL. (92) Serranus aequidens Gilbert, 1890. Deepwater serrano (En); Serrano de aguas profundas (Sp) MOP stations. 2; 001 and 049; Depth range: 128–183 m; USNM voucher specimens: 4; 421248, 421277, 422346 and 422566. Zootaxa 4348 (1) © 2017 Magnolia Press · 85TEP DEEP FISHES FIGURE 84. Serranus aequidens. Top: adult; Bottom: Juvenile. Distribution. Eastern Pacific (from southern California, USA, to the western Gulf of California and central Mexico to western Panama, including the Cocos and Galápagos Islands), occurring at depths between 75 and 265 m. Literature. Gilbert (1890: 61), Castro-Aguirre et al. (1993: 83), Bussing & López (1994: 94; 2009: 454), Balart et al. (1995: 82), Heemstra (1995: 1572), Castro-Aguirre & Balart (1996: 71), Pondella (1999: 130), Thomson et al. (2000:74), Rodríguez-Romero et al. (2008: 1774), McCosker & Rosenblatt (2010: 191), Del Moral- Flores et al. (2013: 197) and Robertson & Allen (2015: 2582). Malacanthidae Tilefishes (En); Matajuelos, Blanquillos (Sp). 1 G, 1 S. (93) Caulolatilus affinis Gill, 1865. Bighead tilefish (En); Blanquillo cabezón, Pierna (Sp) MOP stations. 2; 032 and 061; Depth range: 108–140 m; USNM voucher specimens: 2; 422632 and 422646. Distribution. Eastern Pacific (from California, USA, including the Gulf of California and the Revillagigedos Islands, to northern Peru including the Cocos, Malpelo and Galápagos Islands), occurring at depths between 30 and 200 m. Literature. Gill (1865: 68), Dooley (1978: 25), Allen & Robertson (1994: 124), Bussing & López (1994: 98; 2009: 455), Schneider & Krupp (1995: 1266), Bearez (1996: 736), Franke & Acero (1996: 766), Moser (1996d: 990), De La Cruz Agüero et al. (1997: 221), Grove & Lavenberg (1997: 350), Elorduy-Garay & Ruiz-Cordova (1998: 259), Chirichigno & Vélez (1998: 335), Thomson et al. (2000: 114), Walker et al. 2002:139), Rodríguez- Romero et al. (2008: 1774), Romero & Vélez (2009b: 264), McCosker & Rosenblatt (2010: 191), Reyes-Bonilla et al. (2011: 9), López-Martínez et al. (2012: 353), Del Moral-Flores et al. (2013: 198), Lea & Feeney (2013: 1), Gómez et al. (2014: 394) and Robertson & Allen (2015: 1218).ROBERTSON ET AL .86 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 85. Caulolatilus affinis; USNM 422632, 190 mm. SL. Carangidae Jacks, Pompanos (En); Jureles, Caballas, Palometas (Sp). 2 G, 3 S. (94) Decapterus macrosoma Bleeker, 1851. Shortfin scad (En); Macarela alicorta, Caballa fina (Sp) MOP stations. 2; 005 and 006; Depth range: 109–500 m; USNM voucher specimens: 2; 421355 and 421558. Distribution. Eastern (from southern Baja California, Mexico, including the Gulf of California to Chile, including the Galápagos Islands), Central (Hawai) and Indo-West Pacific (from East Africa, including the Red Sea, to Malaysia, Indonesia, the Arafura Sea, northern Line Islands (Kiribati) and Gambier Islands, north to southern Japan, south to Port Hedland (Western Australia), New South Wales (Australia), New Caledonia and Tonga), occurring at depths between 12 and 170 m. Literature. Bleeker (1851: 358), Allen & Swainston (1988: 72), Pequeño (1989: 60), Randall et al. (1990: 163), Kuiter (1993: 179), Allen & Robertson (1994: 126), Bussing & López (1994: 104), Goren & Dor (1994: 34), Suzuki & Hosokawa (1994: 4), Smith-Vaniz (1995: 940), Watson et al (1996: 917), Bearez (1996: 736), Allen (1997: 114), De La Cruz Agüero et al. (1997: 84), Grove & Lavenberg (1997: 376), Chirichigno & Vélez (1998: 274), Nakabo (2000: 796; 2002: 796), Mundy (2005: 368), Randall (2007: 233), McCosker & Rosenblatt (2010: 192), Psomadakis et al. (2015: 223) and Robertson & Allen (2015: 1257). Remarks. The specimens of this shallow water pelagic species (see Bussing & López (1994), Mundy (2005), Robertson & Allen (2015) and Froese & Pauly (2016), most likely were caught in shallow water during deployment or retrieval of the net. Hence we do not record an increase in maximum depth for this species. FIGURE 86. Decapterus macrosoma. Top: entire specimen; Bottom: detail of head. Zootaxa 4348 (1) © 2017 Magnolia Press · 87TEP DEEP FISHES (95) Selene brevoortii (Gill, 1863). Hairfin lookdown, Mexican lookdown, Pacific lookdown (En); Jorobado antena, Palometa jorobada, Jorobado mexicano (Sp) MOP stations. 2; 081, 102; Depth range: 124–165 m; USNM voucher specimens: 2; 421378 and 422365. FIGURE 87. Selene brevoortii; USNM 421378, 39 mm. SL. Distribution. Eastern Pacific: From southern California, USA, including the Gulf of California, to Chile), occurring at depths between 1 and 50 m. Literature. Gill (1863: 83), Pequeño (1989: 61), Allen & Robertson (1994: 128), Bussing & López (1994: 100; 2009: 456), Lea & Walker (1995: 90), Smith-Vaniz (1995: 940), Bearez (1996: 736), Watson et al (1996: 917), De La Cruz Agüero et al. (1997: 91), Chirichigno & Vélez (1998: 282), Castro-Aguirre et al. (1999: 285), Lea & Rosenblatt (2000: 122), Thomson et al. (2000: 289), Reed et al. (2001: 468), Del Moral-Flores et al. (2013: 198), Murase et al (2014: 1406), Fuentes et al. (2015: 616) and Robertson & Allen (2015: 1267). Remarks. The specimens of this shallow water pelagic species (see Bussing & López (1994), Mundy (2005), Robertson & Allen (2015) and Froese & Pauly (2016), most likely were caught in shallow water during deployment or retrieval of the net. Hence we do not record an increase in maximum depth for this species. (96) Selene peruviana (Guichenot, 1866). Peruvian moonfish, Pacific moonfish (En); Jorobado papelillo (Sp) MOP stations. 2; 085 and 102; Depth range: 149–186 m; USNM voucher specimens: 2; 421404 and 422399. Distribution. Eastern Pacific (from southern California, USA to Chile, including the Galápagos Islands), occurring at depths between 0 and 50 m. Literature. Guichenot (1866: 38), Eschmeyer et al. (1983: 209), Pequeño (1989: 61), Allen & Robertson (1994: 128), Bussing & López (1994: 100; 2009: 456), Smith-Vaniz (1995: 940), Bearez (1996: 736), Watson et al (1996: 917), De La Cruz Agüero et al. (1997: 91), Grove & Lavenberg (1997: 369), Chirichigno & Vélez (1998: 282), Castro-Aguirre et al. (1999: 284), Reed et al. (2001: 468), Rodríguez-Romero et al. (2008: 1775), McCosker & Rosenblatt (2010: 192), Del Moral-Flores et al. (2013: 198), Murase et al. (2014: 1406), Fuentes et al. (2015: 616) and Robertson & Allen (2015: 1269). Remarks. The specimens of this shallow water pelagic species (see Bussing & López (1994), Mundy (2005), Robertson & Allen (2015) and Froese & Pauly (2016), most likely were caught in shallow water during deployment or retrieval of the net. Hence we do not record an increase in maximum depth for this species.ROBERTSON ET AL .88 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 88. Selene peruviana; USNM 422399, 41 mm. SL. Haemulidae Grunts (En); Roncadores (Sp). 1 G, 1 S. (97) Pomadasys branickii (Steindachner, 1879). Sand grunt (En); Roncacho arenero (Sp) MOP stations. 1; 015; Depth range: 113–114 m; USNM voucher specimens: 1; 422650. Distribution. Eastern Pacific (from southern Baja California, Mexico, including the Gulf of California, to Peru), occurring at depths between 0 and 165 m. Literature. Steindachner (1879: 33), Bussing (1993: 245), Allen & Robertson (1994: 15), Bussing & López (1994: 107; 2009: 458), Bearez (1996: 737), McKay & Schneider (1995: 1140), Chirichigno & Vélez (1998: 376), Castro-Aguirre et al. (1999: 342), Rodríguez-Romero et al. (2008: 1775), Tavera et al. (2012: 6), Angulo et al. (2013: 1002), Del Moral-Flores et al. (2013: 200), Murase et al (2014: 1406), Fuentes et al. (2015: 616) and Robertson & Allen (2015: 1382). FIGURE 89. Pomadasys branickii; USNM 422650, 230 mm SL. Zootaxa 4348 (1) © 2017 Magnolia Press · 89TEP DEEP FISHES Sciaenidae Drums, Croakers (En); Corvinas (Sp). 2 G, 2 S. (98) Cynoscion nannus Castro-Aguirre & Arvizu-Martinez, 1976. Dwarf weakfish (En); Corvina de profundidad, Corvinata enana (Sp) MOP stations. 10; 003, 041, 049, 061, 074, 081, 085, 091, 098 and 102; Depth range: 105–1185 m; USNM voucher specimens: 26; 421251, 421255, 421285, 421288, 421303, 421314, 421400, 421409, 421420, 421437, 421446, 421455, 421462, 421483, 421498, 421499, 421500, 421517, 421568, 421569, 422364, 422403, 422513, 422521, 422586 and 422627. FIGURE 90. Cynoscion nannus. Top: entire specimen; Bottom: detail of head. Distribution. Eastern Pacific (from the Gulf of Mexico to Panama), occurring at depths between 100 and 815 m. Literature. Castro-Aguirre & Arvizu-Martínez (1976: 323), Castro-Aguirre (1991: 80), Bussing & López (1994: 128), Chao (1995: 1436), Castro-Aguirre & Balart (1996: 71), De La Cruz Agüero et al. (1997: 266), Béarez (2001: 61), Chao (2002: 11), Guzmán (2011: 173), Fuentes et al. (2015: 616) and Robertson & Allen (2015: 1521). (99) Umbrina bussingi López, 1980. Big eye drum, Big eye croaker, Bussing's drum (En); Polla ojona, Verrugato prieto, Corvina de Bussing (Sp) MOP stations. 1; 041; Depth range: 115–152 m; USNM voucher specimens: 3; 421289, 421301 and 422631. Distribution. Eastern Pacific (from southern Baja California, Mexico, to Colombia), occurring at depths between 30 and 185 m. Literature. López (1980: 203), Walker & Radford (1992: 575), Bussing & López (1994: 130; 2009: 458), Chao (1995: 1443), De La Cruz Agüero et al. (1997: 270), Béarez (2001: 61), Rodríguez-Romero et al. (2008: 1776) and Robertson & Allen (2015: 1487). ROBERTSON ET AL .90 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 91. Umbrina bussingi. Labridae Wrasses (En); Damiselas, Señoritas, Viejas (Sp). 1 G, 1 S. (100) Decodon melasma Gomon, 1974. Black-spot wrasse, Blotched hogfish (En); Viejita manchada (Sp) MOP stations. 4; 001, 003, 004, and 032; Depth range: 108–183 m; USNM voucher specimens: 4; 421393, 422522, 422591 and 422622. FIGURE 92. Decodon melasma. Distribution. Eastern Pacific (from southern California, USA, including the Gulf of California, to northern Peru, including the Cocos and Galápagos Islands), occurring at depths between 40 and 220m. Literature. Gomon (1974: 207), Bussing (1987: 95), Allen & Robertson (1994: 198), Bussing & López (1994: 116; 2009: 462), Bearez (1996: 738), Watson (1996g: 1090), Gomon (1995: 1203), Gomon (1997: 819), De La Cruz Agüero et al. (1997: 197), Chirichigno & Vélez (1998: 330), Parenti & Randall (2000: 17), Lea & Rosenblatt (2000: 125), Thomson et al. (2000: 172), Allen & Groce (2001:131), Baldwin & McCosker (2001: 91), Rodríguez- Romero et al. (2008: 1776), McCosker & Rosenblatt (2010: 193), Del Moral-Flores et al. (2013: 202) and Robertson & Allen (2015: 1646). Zoarcidae Eelpouts (En); Viruelas (Sp). 7 G, 9 S. (101) Bentartia pusillum (Bean 1890). Alaska eelpout (En); Viruela de Alaska (Sp) Zootaxa 4348 (1) © 2017 Magnolia Press · 91TEP DEEP FISHES MOP stations. 6; 002, 029, 077, 078, 082 and 099; Depth range: 407–1527 m; USNM voucher specimens: 10; 421466, 422425, 422560, 423184, 423199, 423210, 435791, 435800, 435805 and 435808. Distribution. North and eastern Pacific (from the eastern Bering Sea to off southern British Columbia and the Gulf of Panama), occurring at depths between 221 and 2189 m. Literature. Bean (1890: 39), McAllister (1990: 157), Anderson (1994: 116; 2006: 3), Springer & Anderson (1997: 21), Mecklenburg et al. (2002: 730), Anderson et al. (2009: 186) and (Balushkin et al. 2011: 1026). Remarks. These specimens represent the first documented record of the species in Central American waters (Bussing & López 1994, 2009, 2011; Anderson 1994; Anderson et al. 2009) and a southeast range extension of about 6800 km in the species’ known distribution (Anderson 1994, Anderson et al. 2009). The southernmost documented record for this species in the eastern Pacific prior to our collection was off British Columbia, Canada (Bean 1890, Anderson 1994, Anderson et al. 2009). FIGURE 93. Bentartia pusillum; USNM 423210, 160 mm. TL. (102) Bothrocara molle Bean 1890. Soft eelpout (En); Viruela suave (Sp) MOP stations. 6; 026, 029, 042, 065, 078 and 099; Depth range: 929–1310 m; USNM voucher specimens: 8; 422331, 422424, 422430, 422570, 422584, 423188, 423200 and 423202. Distribution. North, Eastern Pacific (from Okhotsk, Japan and the Bering Sea to Chile) and western South Atlantic (South Georgia, United Kingdom), occurring at depths between 60 and 2688 m. Literature. Bean (1890: 39), McAllister (1990: 156), Anderson (1994: 116; 2006: 2; 2009: 274), Nakabo (2000: 1041, 2002: 1041), Mecklenburg et al. (2002: 732), Anderson et al. (2009: 182), Balushkin et al. (2011: 973) and Parin et al. (2014: 385). FIGURE 94. Bothrocara molle. Top: entire specimen; Bottom: detail of head; USNM 422570, 178 mm. TL.ROBERTSON ET AL .92 · Zootaxa 4348 (1) © 2017 Magnolia Press (103) Lycenchelys sp.1 MOP stations. 2; 025 and 029; Depth range: 1101–1126 m; USNM voucher specimens: 3; 422609 and 422617. Literature. Anderson (1994: 1; 2009: 276). Remarks. Up to this point, no formal review of eastern Pacific Lycenchelys Gill, 1884 has been published. The taxonomy of the genus, as well as from other genera of the family (e.g. Lycodes, Ophthalmolycus and Pachycara, among others; see below) is complicated and the validity of several forms is questioned given the absence, in most cases, of reliable diagnostic characters (Anderson 1994). Accordingly, we were not able to make a specific identification of these specimens; however, despite the apparent absence of external diagnostical features between them, two different forms (Lycenchelys sp.1 and Lycenchelys sp.2) are herein listed based on molecular differences (6% divergence in COI sequences). FIGURE 95. Lycenchelys sp.1 (104) Lycenchelys sp.2 MOP stations. 1; 086; Depth range: 1353–1389 m; USNM voucher specimens: 1; 421515. Literature. Anderson (1994: 64; 2009: 276). Remarks. see remarks about Lycenchelys sp 1. (105) Lycodapus dermatinus Gilbert, 1896. Looseskin eelpout (En); Viruela pielfloja (Sp) MOP stations. 2; 001 and 076; Depth range: 165–1145 m; USNM voucher specimens: 2; 421188 and 422332. Distribution. Eastern Pacific (from the Welker Seamount off Alaska, USA, to Peru), occurring at depths between 165 and 1145 m. Literature. Gilbert (1896: 471), Springer & Anderson (1997: 17), Peden & Anderson (1978: 1948; 1989: 152; 1994:118), Mecklenburg et al. (2002: 740) and Balushkin et al. (2011: 985). Remarks. Specimens collected at 165 and 1145 m represent a new maximum depth record for the species (Anderson 1978, 1989, 1994; Froese & Pauly 2016). Zootaxa 4348 (1) © 2017 Magnolia Press · 93TEP DEEP FISHES FIGURE 96. Lycodapus dermatinus. Top: entire specimen; Bottom: detail of head; USNM 422332, 124 mm. SL. (106) Lycodes sp. MOP stations. 1; 056; Depth range: 656–668 m; USNM voucher specimens: 2; 421556 and 422538. Literature. Anderson (1994: 68). FIGURE 97. Lycodes sp. Top: entire specimen; Bottom: detail of head. (107) Ophthalmolycus sp. MOP stations. 4; 058, 078, 083 and 096; Depth range: 965–1270 m; USNM voucher specimens: 8; 421307, 421309, 421318, 421473, 421542, 422367, 422428 and 422452. Literature. Anderson (1992: 1; 1994: 80).ROBERTSON ET AL .94 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 98. Ophthalmolycus sp. Top: entire specimen; Bottom: detail of head. (108) Pachycara sp.1 MOP stations. 2; 040 and 073; Depth range: 153–1185 m; USNM voucher specimens: 2; 422432 and 422599. Literature. Anderson (1994: 82). Remarks. We were not able to make a specific identification of these specimens; however, despite the apparent absence of external diagnostical features between them, two different forms (Pachycara sp.1 and Pachycara sp.2) are herein listed based on molecular differences (3.81% divergence in COI sequences). (109) Pachycara sp.2 MOP stations. 1; 082; Depth range: 1471–1527 m; USNM voucher specimens: 2; 421417 and 421562. Literature. Anderson (1994: 82). Remarks. see remarks about Pachycara sp 1. FIGURE 99. Pachycara sp.2. Top: entire specimen; Bottom: detail of head. Chiasmodontidae Swallowers (En); Engullidores (Sp). 2 G, 2 S. (110) Chiasmodon subniger Garman, 1899. Eastern Pacific black swallower (En); Engullidor negro del Pacifico oriental (Sp) Zootaxa 4348 (1) © 2017 Magnolia Press · 95TEP DEEP FISHES MOP stations. 3; 001, 094 and 100; Depth range: 183–1406 m; USNM voucher specimens: 3; 422491, 435801 and 435803. Distribution. Eastern (from Canada to northern Chile) and Central Pacific (Hawai), occurring at depths between 183 and 4568 m. Literature. Garman (1899:73), Pequeño (1989: 64), Watson & Sandknop (1996b: 1131), McEachran & Fechhelm (2005: 554), Prokofiev (2008: 212; 2010: 507), Melo (2009: 589), Prokofiev & Kukuev (2009: 901) and Reyes-Bonilla et al. (2011: 10). Remarks. Prokofiev & Kukuev (2009) and Prokofiev (2010) suggested that C. subniger is a synonym of C. niger Johnson 1864 based on the absence of “significant distinctions between specimens from the equatorial and southeastern Pacific and from other regions of the World Ocean.” In contrast, Melo (2009) used several morphometric and meristic characters to clearly separate C. subniger (eastern Pacific), from C.niger (Atlantic Ocean). DNA barcodes for our specimens of C. subniger match those for C. subniger from the eastern Pacific in the BOLD database. DNA barcodes in BOLD for C.niger from the Atlantic are approximately 7.5% different, which provides empirical support for the validity of C. subniger as proposed by Melo (2009). Specimens collected at 183 m represent a new minimum depth record for the species (Melo 2009, Froese & Pauly 2016). FIGURE 100. Chiasmodon subniger; USNM 422491, 130 mm. SL. (111) Pseudoscopelus lavenbergi Melo, Walker & Klepadlo 2007. MOP stations. 1; 029; Depth range: 1113–1126 m; USNM voucher specimens: 1; 422507. Distribution. Eastern Pacific (from California, USA, to Chile, including the Hawaiian Islands), occurring at depths between 200 and 2300 m. Literature. Melo et al. (2007: 33), Melo et al. (2010: 65) and Prokofiev (2014: 647). FIGURE 101. Pseudoscopelus lavenbergi. Top: entire specimen; Bottom: detail of head; USNM 422507, 160 mm. SL. Uranoscopidae Stargazers (En); Miracielos (Sp). 1 G, 1 S.ROBERTSON ET AL .96 · Zootaxa 4348 (1) © 2017 Magnolia Press (112) Kathetostoma averruncus Jordan & Bollman, 1890. Smooth stargazer (En); Miracielo buldog (Sp) MOP stations. 4; 006, 015, 049 and 068; Depth range: 109–145 m; USNM voucher specimens: 8; 421220, 421228, 421231, 421232, 421261, 421335, 422305 and 422321. Distribution. Eastern Pacific (from California, USA, to Isla Lobos de Tierra, Peru, including the Galápagos Islands), occurring at depths between 13 and 600 m. Literature. Jordan & Bollman (1890: 163), Fierstine & Werner (1963: 50), Eschmeyer et al. (1983: 240), Kirsch (1989: 260), Pietsch (1989: 296), Bussing & López (1994: 140; 2009: 463), Bussing & Lavenberg (1995b: 1649), Bearez (1996: 738), Fitch & Lavenberg (1968: 102), Castro-Aguirre & Balart (1996: 71), Watson (1996h: 1144), De La Cruz Agüero et al. (1997: 306), Grove & Lavenberg (1997: 512), Chirichigno & Vélez (1998: 240), Rodríguez-Romero et al. (2008: 1776), Odani (2009: 291), McCosker & Rosenblatt (2010: 193), López-Martínez et al. (2012: 353), Vilasri (2013: 89), Fuentes et al. (2015: 616) and Robertson & Allen (2015: 1712). FIGURE 102. Kathetostoma averruncus. Callionymidae Dragonets (En); Dragoncillos (Sp). 1 G, 1 S. (113) Synchiropus atrilabiatus (Garman, 1899). Antler dragonet, Blacklip dragonet, Sleepy dragonet (En); Dragoncito de asta, Gobio-adorno vistoso (Sp) MOP stations. 1; 001; Depth range: 165–183 m; USNM voucher specimens: 1; 422551. Distribution. Eastern Pacific (from the mouth of the Gulf of California, to Paita, Peru, including the Cocos, Malpelo and Galapagos Islands), occurring at depths between 3 and 235 m. Literature. Garman (1899: 122), Fricke (1981: 33; 1982: 75; 1983: 567; 2002:52), Castro-Aguirre (1991: 82), Castro-Aguirre et al. (1993: 86), Bussing & López (1994: 142; 2009: 465), Watson (1996i: 1205), Grove & Lavenberg (1997: 545), Lea & Rosenblatt (2000: 126), Rodríguez-Romero et al. (2008: 1776), Hooker (2009c: 294), McCosker & Rosenblatt (2010: 194), López-Martínez et al. (2012: 353), Fuentes et al. (2015: 616) and Robertson & Allen (2015: 1965). FIGURE 103. Synchiropus atrilabiatus; USNM 422551, 120 mm. SL. Zootaxa 4348 (1) © 2017 Magnolia Press · 97TEP DEEP FISHES Gobiidae Gobies (En); Gobios, Chupapiedras, Guarasapas (Sp). 1 G, 2 S. (114) Bollmannia sp 1. MOP stations. 3; 003, 038 and 048; Depth range: 116–154 m; USNM voucher specimens: 6; 421390, 421397, 421408, 421428, 422310 and 422401. Literature. Hoese (1995: 1132) and Van Tassel et al. (2012: 61). Remarks. As noted by Van Tassel et al. (2012), no formal review of eastern Pacific Bollmannia Jordan, 1890 has been published up to this point. The taxonomy of the genus is complicated and the validity of several forms is questioned given the absence of real diagnostic characters. Additionally, several of the characters described as diagnostic for the most of the “valid” species are of little value as these characters are commonly destroyed when specimens are trawled or dredged from considerable depths, as in the present case. Accordingly, we were not able to make a specific identification of these specimens; however, despite the apparent absence of external diagnostical features between them, two different forms (Bollmannia sp.1 and Bollmannia sp.2) are herein listed based on molecular differences (6.1–6.8% divergence in COI sequences). (115) Bollmannia sp 2. MOP stations. 5; 003, 038, 048, 081 and 082; Depth range: 116–154 m; USNM voucher specimens: 6; 421380, 421381, 421387, 421403, 421412 and 422308. Literature. Hoese (1995: 1132) and Van Tassel et al. (2012: 61). Remarks. see remarks about Bollmannia sp 1. FIGURE 104. Bollmannia sp 2. Two different specimens to demonstrate color variation. Trichiuridae Cutlassfishes (En); Cintas, Fajas (Sp). 1 G, 1 S. (116) Trichiurus nitens Garman 1899. Largehead hairtail (En); Pez cinta, Pez sable (Sp) MOP stations. 4; 004, 036, 038 and 102; Depth range: 115–165 m; USNM voucher specimens: 10; 421385, 421407, 421433, 422317, 422326, 422372, 422376, 422380, 422445 and 422553. Distribution. Eastern Pacific (from California, USA, to Peru, including the Galápagos Islands), occurring at depths between 3 and 500 m. Literature. Garman (1899: 69), Eschmeyer & Herald (1983: 267), Nakamura & Parin (1993: 106), Allen & Robertson (1994: 275), Bussing & López (1994: 148; 2009: 468), Nakamura (1995: 1639), Bearez (1996: 739), De La Cruz Agüero et al. (1997: 299), Grove & Lavenberg (1997: 577), Chirichigno & Vélez (1998: 117), Castro-ROBERTSON ET AL .98 · Zootaxa 4348 (1) © 2017 Magnolia Press Aguirre et al. (1999: 461), Lea & Rosenblatt (2000: 126), Hutchins (2001: 45), Menezes et al. (2003: 102), Parin (2003: S27), McEachran & Fechhelm (2005: 770), Chakraborty et al. (2006: 93), Burhanuddin & Parin (2008: 825), McCosker & Rosenblatt (2010: 194), Psomadakis et al. (2015: 313) and Robertson & Allen (2015: 4507). FIGURE 105. Trichiurus nitens. Top: entire specimen; Bottom: detail of head. Nomeidae Driftfishes, Cigarfishes (En); Derivantes, Flotadores (Sp). 2 G, 3 S. (117) Cubiceps pauciradiatus Günther, 1872. Bigeye cigarfish (En); Derivante ojón (Sp) MOP stations. 4; 003, 028, 034 and 062; Depth range: 116–1472 m; USNM voucher specimens: 5; 421415, 421436, 421513, 422357 and 422496. FIGURE 106. Cubiceps pauciradiatus. Distribution. Circumglobal in tropical through warm temperate seas, occurring at depths between 58 and 1000 m. In the eastern Pacific this species has been previously recorded from Baja California, Mexico, to Chile. Literature. Günther (1872: 423), Haedrich (1967: 81; 1995: 1323), Ahlstrom et al. (1976: 285), Butler (1979: 236), Scott & Scott (1988: 473), Pequeño (1989: 68), McAllister (1990: 181), Gomon et al. (1994: 839), Bearez (1996: 739), Watson (1996j: 1301), Castro-Aguirre & Balart (1996: 71), Grove & Lavenberg (1997: 603), Chirichigno & Vélez (1998: 292), Nakabo (2000: 964; 2002:964), Hutchins (2001: 46), McEachran & Fechhelm (2005: 805), Mundy (2005: 512), Shimazaki (2009b: 302), Ho et al. (2010: 257), McCosker & Rosenblatt (2010: 195), Del Moral-Flores et al. (2013: 208) and Robertson & Allen (2015: 2266). Remarks. The specimens of this pelagic species (see Bussing & López 1994, Mundy 2005), Robertson & Zootaxa 4348 (1) © 2017 Magnolia Press · 99TEP DEEP FISHES Allen 2015, Froese & Pauly 2016), could have been caught in shallow water during deployment or retrieval of the net. Hence we do not record an increase in maximum depth for this species. (118) Psenes cyanophrys Valenciennes, 1833. Freckled driftfish, Striped driftfish (En); Flotador derivante, Derivante rayado (Sp) MOP stations. 1; 021; Depth range: 764–987 m; USNM voucher specimens: 1; 421567. Distribution. Circumglobal in tropical through warm temperate seas, occurring at depths between 0 and 550 m. In the eastern Pacific this species has been previously recorded from Costa Rica to Peru, including the Cocos, Malpelo and Galápagos Islands. Literature. Valenciennes (1833a: 260), Haedrich (1967: 84; 1995: 1323), Ahlstrom et al. (1976: 285), Uyeno et al. (1983: 423), Allen & Robertson (1994: 284), Grove & Lavenberg (1997: 605), Chirichigno & Vélez (1998: 291), Nakabo (2000: 964; 2002: 964), Hutchins (2001: 46), Menezes et al. (2003: 104), Parin & Piotrovsky (2004: S53), McEachran & Fechhelm (2005: 808), Mundy (2005: 512), Bussing & López (2009: 469), McCosker & Rosenblatt (2010: 195) and Robertson & Allen (2015: 2271). Remarks. The specimens of this pelagic species (see Bussing & López (1994), Mundy (2005), Robertson & Allen (2015) and Froese & Pauly (2016), could have been caught in shallow water during deployment or retrieval of the net. Hence we do not record an increase in maximum depth for this species. FIGURE 107. Psenes cyanophrys; USNM 421567, 90 mm. SL. (119) Psenes sio Haedrich, 1970. Two-spine driftfish (En); Flotador dos espinas (Sp) MOP stations. 8; 037, 065, 070, 074, 077, 080, 082 and 103; Depth range: 117–1527 m; USNM voucher specimens: 14; 421342, 421375, 421382, 421405, 421410, 421434, 421570, 422373, 422377, 422378, 422411, 422472, 422526 and 422547. Distribution. Eastern Pacific (from Baja California, Mexico, to Chile, including the Galápagos Islands), occurring at depths between 0 and 500m (see Remarks). Literature. Haedrich (1970: 2; 1995: 1323), Pequeño (1989: 68), Fujita (1991: 323), Ahlstrom et al. (1976: 285), Bussing & López (2009: 469), Castro-Aguirre et al. (1993: 85), Bearez (1996: 739), Watson (1996j: 1300), Castro-Aguirre & Balart (1996: 71), Grove & Lavenberg (1997: 606), Chirichigno & Vélez (1998: 291), Shimazaki (2009b: 303), McCosker & Rosenblatt (2010: 195), Del Moral-Flores et al. (2013: 208) and Robertson & Allen (2015: 2271). Remarks. The specimens of this pelagic species (see Bussing & López (1994), Mundy (2005), Robertson & Allen (2015) and Froese & Pauly (2016), could have been caught in shallow water during deployment or retrieval of the net. Hence we do not record an increase in maximum depth for this species.ROBERTSON ET AL .100 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 108. Psenes sio. Top: entire specimen; Bottom: detail of head. Stromateidae Butterfishes (En); Palometas (Sp). 1 G, 2 S. (120) Peprilus medius (Peters, 1869). Pacific harvestfish (En); Pampanito aleta larga, Palometa cometrapo, Palometa común (Sp) MOP stations. 2; 006 and 015; Depth range: 109–114 m; USNM voucher specimens: 5; 421322, 421325, 421328, 422337 and 422358. FIGURE 109. Peprilus medius. Distribution. Eastern Pacific (From Southern Baja California Sur, Mexico, including the Gulf of California, to northern Chile, including the Galápagos Islands), occurring at depths between 4 and 220 m. Literature. Peters (1869: 707), Horn (1970: 175), Allen & Robertson (1994: 285), Bussing & López (1994: 116; 2009: 469), Balart et al. (1995: 83), Haedrich & Schneider (1995: 1623), Sielfeld et al. (1995: 25), Bearez (1996: 739), Grove & Lavenberg (1997: 598), Chirichigno & Vélez (1998: 141), Rodríguez-Romero et al. (2008: Zootaxa 4348 (1) © 2017 Magnolia Press · 101TEP DEEP FISHES 1776), McCosker & Rosenblatt (2010: 195), Del Moral-Flores et al. (2013: 208), Murase et al (2014: 1408) and Robertson & Allen (2015: 2280). (121) Peprilus snyderi Gilbert & Starks, 1904. Salema butterfish (En); Palometa salema (Sp) MOP stations. 2; 006 and 068; Depth range: 109–145 m; USNM voucher specimens: 4; 421333, 422303, 422315 and 422330. Distribution. Eastern Pacific (from southern Baja California Sur, Mexico, including the Gulf of California, to northern Peru), occurring at depths between 0 and 235 m. Literature. Gilbert & Starks (1904: 87), Böhlke (1953: 105), Haedrich (1967: 107), Horn (1970: 189), Allen & Robertson (1994: 285), Bussing & López (1994: 116; 2009: 469), Balart et al. (1995: 83), Haedrich & Schneider (1995: 1623), Bearez (1996: 739), Chirichigno & Vélez (1998: 142), Rodríguez-Romero et al. (2008: 1776), López-Martínez et al. (2012: 353) and Robertson & Allen (2015: 2283). FIGURE 110. Peprilus snyderi. PLEURONECTIFORMES 3 F, 5 G, 7 S. Paralichthyidae Sand flounders (En); Lenguados, Peces Hoja (Sp). 2 G, 3 S. (122) Citharichthys platophrys Gilbert, 1891. Small sandab (En); Lenguado frentón (Sp) MOP stations. 2; 041 and 049; Depth range: 115–152 m; USNM voucher specimens: 4; 421283, 421298, 422562 and 422576. Distribution. Eastern Pacific (from southern Baja California, Mexico, including the Gulf of California, to northern Peru, including the Cocos Island), occurring at depths between 10 and 152 m (see Remarks). Literature. Gilbert (1891b: 454), Castro-Aguirre et al. (1992: 100), Bussing & López (1994: 150; 2009: 470), Hensley (1995b: 1349), Bearez (1996: 739), Moser & Sumida (1996: 1325), De La Cruz Agüero et al. (1997: 249), Grove & Lavenberg (1997: 613), Chirichigno & Vélez (1998: 210), Rodríguez-Romero et al. (2008: 1777), Amaoka (2009a: 310), van der Heiden et al. (2009: 814), Del Moral-Flores et al. (2013: 208), Murase et al (2014: 1408), Fuentes et al. (2015: 617) and Robertson & Allen (2015: 2312). Remarks. Specimens collected at 152 m represent a new maximum depth record for the species (Bussing & López 1994, 2009; Hensley 1995b; Robertson & Allen 2015; Froese & Pauly 2016).ROBERTSON ET AL .102 · Zootaxa 4348 (1) © 2017 Magnolia Press FIGURE 111. Citharichthys platophrys. (123) Hippoglossina bollmani Gilbert, 1890. Spotted flounder (En); Lenguado pintado (Sp) MOP stations. 4; 015, 041, 049 and 068; Depth range: 113–152 m; USNM voucher specimens: 7; 421239, 421313, 421327, 421339, 421351, 421489 and 422468. Distribution. Eastern Pacific (from southern Baja California, Mexico, including the Gulf of California, to Peru, including the Cocos and Galápagos Islands), occurring at depths between 18 and 191 m. Literature. Gilbert (1890: 122), Norman (1934: 68), Bussing & López (1994: 154; 2009: 470), Hensley (1995b: 1355), Bearez (1996: 739), Moser & Sumida (1996: 1325), De La Cruz Agüero et al. (1997: 252), Chirichigno & Vélez (1998: 200), Rodríguez-Romero et al. (2008: 1777), Amaoka (2009a: 312), McCosker & Rosenblatt (2010: 195), López-Martínez et al. (2012: 353), Del Moral-Flores et al. (2013: 208) and Robertson & Allen (2015: 2326). FIGURE 112. Hippoglossina bollmani. (124) Hippoglossina tetrophthalma (Gilbert, 1890). Fourspot flounder (En); Lenguado cuatro ojos (Sp) MOP stations. 1; 041; Depth range: 115–152 m; USNM voucher specimens: 2; 422597 and 422637. Distribution. Eastern Pacific (from southern Baja California, Mexico, including the Gulf of California, to Peru), occurring at depths between 23 and 233 m. Literature. Gilbert (1890: 122), Allen & Robertson (1994: 288), Bussing & López (1994: 152; 2009: 470), Hensley (1995b: 1355), Bearez (1996: 739), Moser & Sumida (1996: 1325), De La Cruz Agüero et al. (1997: 253), Chirichigno F. & Vélez D. (1998: 200), Rodríguez-Romero et al. (2008: 1777), López-Martínez et al. (2012: 354), Del Moral-Flores et al. (2013: 208) and Robertson & Allen (2015: 2331). Zootaxa 4348 (1) © 2017 Magnolia Press · 103TEP DEEP FISHES FIGURE 113. Hippoglossina tetrophthalma. Bothidae Lefteye flounders (En); Lenguados, Lenguados Izquierdos (Sp). 2 G, 3 S. (125) Engyophrys sanctilaurentii Jordan & Bollman, 1890. Speckled-tail flounder (En); Lenguado colimanchada (Sp) MOP stations. 2; 001and 015; Depth range: 113–183 m; USNM voucher specimens: 3; 421316, 422515 and 422537. Distribution. Eastern Pacific (from southern Baja California, Mexico, including the Gulf of California, to Peru, including the Malpelo Island), occurring at depths between 40 and 235 m. Literature. Jordan & Bollman (1890: 176), Böhlke (1953: 139), Castro-Aguirre et al. (1992: 102), Bussing & López (1994: 152; 2009: 469), Hensley (1995a: 931), Castro-Aguirre & Balart (1996: 71), Moser & Charter (1996: 1357), De La Cruz Agüero et al. (1997: 74), Chirichigno & Vélez (1998: 203), Lea & Rosenblatt (2000: 126), Allen & Groce (2001: 137), Rodríguez-Romero et al. (2008: 1777), Murase et al (2014: 1408), Fuentes et al. (2015: 617) and Robertson & Allen (2015: 2294). (126) Monolene dubiosa Garman, 1899. Acapulco flounder (En); Lenguado acapulqueño, Lenguado carbón (Sp) FIGURE 114. Monolene dubiosa.ROBERTSON ET AL .104 · Zootaxa 4348 (1) © 2017 Magnolia Press MOP stations. 3; 039, 061 and 068; Depth range: 137–237 m; USNM voucher specimens: 8; 421486, 422323, 422328, 422488, 422490, 422493, 422501 and 422516. Distribution. Eastern Pacific (from southern Baja California, Mexico, including the Gulf of California, to Peru), occurring at depths between 68 and 255 m. Literature. Garman (1899: 227), Bussing & López (1994: 152; 2009: 469), Hensley (1995a: 931), Castro- Aguirre & Balart (1996: 71), Moser & Charter (1996: 1357), De La Cruz Agüero et al. (1997: 74), Evseenko (1999: 625), Rodríguez-Romero et al. (2008: 1777), Amaoka (2009b: 315), Fuentes et al. (2015: 617) and Robertson & Allen (2015: 2296). Remarks. The USNM 422493 specimen differs from the others by having a relatively shorter eye (about 17% of head length, vs. about 21%), a feature that, according to some authors, e.g. Clark (1936) and Hensley (1995), distinguishes M. assaedae Clark, 1936 from M. dubiosa. However, other authors, e.g. Evseenko (1999) have questioned the validity of M. assaedae given the apparent absence of other more robust, distinctive characters, and have considered it to be a junior synonym of M. dubiosa. COI sequences for both the multiple M. dubiosa specimens and the single M. “assaedae” specimen in our collection are very similar (less than 1% sequence divergence), which supports Evseenko’s (1999) proposal. Hence all these specimens are listed here as M. dubiosa. (127) Monolene maculipinna Garman, 1899. Pacific deepwater flounder (En); Lenguado de profundidad (Sp) MOP stations. 2; 001 and 049; Depth range: 128–183 m; USNM voucher specimens: 5; 421550, 422351, 422356, 422517 and 422565. Distribution. Eastern Pacific (from Nicaragua to Peru, including the Galapagos Islands), occurring at depths between 65 and 385 m. Literature. Garman (1899: 226), Bussing & López (1994: 152; 2009: 470), Hensley (1995a: 931), Moser & Charter (1996: 1357), Sánchez (1997: 7), Chirichigno & Vélez (1998: 202), Hoshino et al. (2000: 293), Amaoka (2009b: 316), McCosker & Rosenblatt (2010: 195) and Robertson & Allen (2015: 2637). FIGURE 115. Monolene maculipinna. Cynoglossidae Tonguefishes (En); Lengüetas, Lenguas (Sp). 1 G, 1 S. (128) Symphurus leei Jordan & Bollman 1890. Black-tail tonguefish, Lee's tonguefish (En); Lengua colinegra, Lengua de Lee (sp) MOP stations. 2; 038 and 073; Depth range: 144–176 m; USNM voucher specimens: 6; 421245, 421418, 421442, 421452, 421454 and 421507. Distribution. Eastern Pacific (from the Gulf of California, Mexico, to Panama), occurring at depths between 10 and 176 m (see Remarks). Literature. Jordan & Bollman (1890: 178), Böhlke (1953: 143), Munroe & Nizinski (1990: 994), Allen & Robertson (1994: 291), Bussing & López (1994: 146), Munroe et al. (1995: 1051), De La Cruz Agüero et al. (1997: 116), Chirichigno & Vélez (1998: 215) and Robertson & Allen (2015: 2382). Zootaxa 4348 (1) © 2017 Magnolia Press · 105TEP DEEP FISHES Remarks. Specimens collected at 176 m represent a new maximum depth record for the species (Bussing & López 1994, 2009; Munroe et al. 1995; Robertson & Allen 2015; Froese & Pauly 2016). FIGURE 116. Symphurus leei. Top and Bottom: two specimens to illustrate variation. TETRAODONTIFORMES 1 F, 1 G, 1 S. Balistidae Triggerfishes (En); Peces Chanchos, Calafates (Sp). 1 G, 1 S. (129) Balistes polylepis Steindachner, 1876. Finescale triggerfish (En); Pejepuerco coche, Cochi (Sp) MOP stations. 1; 091; Depth range: 105–107 m; USNM voucher specimens: 1; 421241. FIGURE 117. Balistes polylepis; USNM 421241, 35 mm. SL.ROBERTSON ET AL .106 · Zootaxa 4348 (1) © 2017 Magnolia Press Distribution. Eastern (from northern California, USA, including the Gulf of California, to Chile, including all the offshore islands) and Central Pacific (Marquesas Islands and Hawaii), occurring at depths between 2 and 512 m. Literature. Steindachner (1876: 69), Berry & Baldwin (1966: 436), Eschmeyer et al. (1983: 295), Pequeño (1989: 77), Allen & Robertson (1994: 294), Bussing & López (1994: 158; 2009: 471), Bearez (1996: 739), Watson (1996k: 1417), De La Cruz Agüero et al. (1997: 64), Grove & Lavenberg (1997: 628), Randall & Mundy (1998: 324), Chirichigno & Vélez (1998: 137), Thomson et al. (2000: 268), Richards & Engle (2001: 18), Mecklenburg et al. (2002: 851), Mundy (2005: 523), Randall (2007: 470), Rodríguez-Romero et al. (2008: 1778), McCosker & Rosenblatt (2010: 195), Del Moral-Flores et al. (2013: 209), Murase et al (2014: 1408), Fuentes et al. (2015: 617), Delrieu-Trottin et al. (2015: 8) and Robertson & Allen (2015: 2396). Acknowledgements DRR participated in the 2010 survey by the B/O Miguel Oliver along the Pacific coast of Central America at the invitation of Mario Gonzalez Recinos of OSPESCA, which organization obtained the necessary government permits for the cruise activities. A trip to Panama by a team of investigators from the Smithsonian’s National Museum of Natural History (NMNH) in Washington, DC to process the frozen fish samples and subsequently barcode the tissue samples was made possible by a Smithsonian Institution Federal Barcode grant to LW, CB and AD in 2011. A trip to Panama by Cristina Castillo to prepare the shipping of specimens to NMNH was sponsored by the Herbert R and Evelyn Axelrod Endowment Fund for Systematic Ichthyology at the NMNH Division of Fishes. A trip to Washington, DC by AA to examine and determine the specific identification of the specimens was sponsored by the Leonard P. Schultz Endowment Fund for Systematic Ichthyology at the NMNH Division of Fishes; Lynne Parenti and Carol Youmans make this trip possible. The laboratory and computer work were conducted in and with the support of the Laboratories of Analytical Biology facilities of the NMNH or its partner labs. 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