NAMES APPLIED TO THE NORTH AMERICAN BEES OFTHE GENERA LITHURGUS, ANTHIDIUM, AND ALLIES.^By T. D. A. COCKERELL,Of the University of Colorado.The Anthidiine and Lithurgine bees, representing two groups ofMegachiiidae, are interesting on account of their habits, and fre-quently well-marked or even very peculiar structural characters.The Anthidiines, nearly always spotted or banded with yellow onthe abdomen, are very well represented in our fauna, and are evento be found in the IMiocene deposits of Colorado. Lithurgus, on theother hand, has only a few species in our fauna, although it is anold genus, found in the European Miocene, and widely spread overthe earth, even to Australia and the islands of the Pacific.LIST OF SPECIES.Genus ANTHIDIUM Fabricius,A genus of many species, found in most parts of the world, butabsent from Australia and New Zealand.The females use cottony fibers in making their nests, and are calledby Fabre "Cottoniers."For a discussion of the Palearctic genera and groups included byauthors in Anthidium, see Entomologist's Record, vol, 21, No. 12.Tables.(1) Cockerell, Bull. So. Cal. Acad. Sci., vol. 3, 1904, pp. 56-58. (Males.)(2) Cockerell, Proc. U. S. Nat. Mus., vol. 40, 1911, p. 249.(3) Friese, Daa Tierreich, Megachilinae, Uef. 28, 1911, pp. 379-381.(4) Friese, Das Tierreich, Megachilinse, Uef. 28, 1911, pp. 391-392.(5) Cockerell, University of Colorado Studies, vol. 4, 1907, pp. 249-250.(6) Swenk, University [of Nebraska] Studies, vol. 14, no. 1, 1913, pp. 9-11.americanum Friese, 1911. Tab. 4.New name for A. maculatum Smith, preoccupied.
1 The asterisk (*) indicates that the species is in the collection of the United States National
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Some species sent to the United States National Museum by Prof. T. D. A. Cockerell labeled "cotypes,"are herelisted as "paratypes," since Professor Cockerell ases the name "cotype" in the sense in whichthe museum uses "paratype." The museum has considerable material in the Anthidiine group andwhen this material is all determined the number of species in its collection will be considerably increased.Of the 107 names (fossil species omitted) in this list in the Anthidiine bees, 47 are represented by speci-mens in the collection. Of this 47, 20 are types or paratypes.?J. C. Crawford.Proceedings U. s. National Museum, Vol. 47?No. 2045. 87
PROCEEDINGS OF THE NATIOXAL MUSEUM.
*angelanim Trrus, 1906. Los Angeles County, California (Coquillett).Female 9| mm.; markings deep yellow, scopa very white, sixth abdominal seg-ment with large yellow spots.Type.?Cat. No. 9034, U.S.N.M.*astragali Swenk, 1913. Bad Lands at mouth of Monroe canyon, Sioux County,Nebraska. (M. Cary). Tab. 6.Paratype.?Cat. No. 15259, U.S.N.M.atrlfrons Cresson, 1868. New Mexico (S. Lewis).Same as emarginatum.atriventre Cresson, 1878. California (Hy. Edwards). Tab. 3.Female with ventral scopa and hair of face fuscous or black,atriventre Smith. 1S79. Orizaba, Mexico.Name preoccupied: ^=orizabx.aztecum Cresson, 1878. Mexico (Sumichrast). Tab. 4.Allied to cognatum.*banningense Cockerell. 1904. Banning, California (Davidson). Tab. 1.Male 14J mm.*bernardinum. Cockerell, 1904. Strawberry Valley, California (Davidson). Tab. 1.Male about 14 mm.; allied to A. pecosense.Paratype.?Cat. No. 13666, U.S.N.M.*bernar(iinum aridum Cockerell, 1904. Rock Creek, California (Davidson). Tab. 1.Scape of male yellow in front.*bernardinuin fragariellum Cockerell, 1904. Strawberry Valley, California (David-son). Tab. 1.Scape of male all black,bernardinum wilsoni Cockerell, 1904. Mount Wilson, California (Davidson). Tab. 1.Male about 11 mm.blanditum Cresson, 1879. Nevada (Morrison). Tab. 2, 3.Coxae and trochanters (female) all black.blanditum praedentatum Cockerell, 1907. Boulder, Colorado (G. Hite). Tab. 5.Known (female) from A. montivagum and A. porterx by the bright lemon-yellowsixth abdominal segment, its margin evidently notched in the middle, and theelongated marks over the eyes. Referred later to A. placitum as a variety.californicum Cresson, 1879. California (H. Edwards). Tab. 1, 3.Fowler describes the female from Berkeley and Redlands, California. Fox describesa variety from Lower California.clypeodentatum Swenk, 1913. Sioux county, Nebraska. Tab. 6.*cognatum Cresson, 1878. Georgia (Morrison). Tab. 3.Characters: Trans. Amer. Ent. Soc, vol. 29, p. 175. IlUnois (Robertson),compactum Provancher, 1896. Los Angeles, California (Coquillett).Name preoccupied: =collectuin.conspicuum Cresson, 1879. Nevada (Morrison). Tab. 3, 5.Male: Entom. News, 1909, p. 262.Also in Colorado.collectTim HuARD, 1896. Tab. 1, 3.Allied to A. emarginatum; see Bull. Southern California Acad. Sci., May 1904, p. 73.New name for A. compactum.coUectum ultrapictum Cockerell, 1904. Tehachapi, California (Davidson).Male a little larger; scape with a yellow stripe; abdominal bands very bright yellow,only that on first segment divided into four spots.crassipes Cresson, 1878. Florida (Tatnall). Tab. 3.Ventral scopa yellowish ; legs very robust.edwardsii Cresson, 1878. California (Hy. Edwards). Tab. 3.Dorsal hair of head and thorax dense and fulvous.
NO. 2045. NORTH AMERICAN BEES?COCKERELL. 89*einarginatum Say, 1824. Tab. 3, 5, 6.Face of female black.Cresson redescribes it from specimens collected in Kansas.emarginatum atripes Cresson, 1879. Nevada (Morrison).Male with legs entirely black,exhumatum Cockerell, 1906. Fossil in the Miocene shales of Florissant (Scudder).flavolineatum Smith, 1879. Oajaca, Mexico=Dianthidiumflavolineatum.Female 8 mm., a short narrow line behind the eyes yellow.*harbecki Crawford, 1910. Wenonah, New Jersey (H. S. Harbeck).Has a pulvillus, "Related according to the swollen legs to larrex and to crassipes.Outside of that one character it does not seem close to any others." (Crawford inletter.)Type.?G&t. No. 13454, U.S.N.M.This is to be referred to Heteranthidium on the authority of Mr. Crawford,hesperium Swenk, 1913. Palo Alto, California.hesperium dentipygum Swenk, 1913. Laramie, Wyoming.*illustre Cresson, 1879. Nevada (Morrison). Tab. 3.Fowler described the male from Redlands, California.Nesting habits; Entom. News, 1904, p. 284.This species is to be referred to Dianthidium.illustre consonum Cresson, 1879. Nevada (Morrison).impatiens Smith, 1879=Dianthidiu7n impatiens. Tab. 4.Male 8^ mm,incurvatum Swenk, 1913. Ute creek, Costilla County, Colorado (H. S. Smith).jocosum Cresson, 1878. Colorado (Ridings). Tab. 3."A pretty little species, with the bands on apex of abdomen broad and uninter-rupted." (Cresson.)*lupinellum Cockerell, 1904. Pecos, New Mexico (W. P. Cockerell). Tab. 1.Male about 11 mm. ; clypeus all yellow. Visits Lupinus.Also in Texas. ?maculatum Smith, 1854. Mexico.Notes on type: Trans. Amer. Ent. Soc, vol. 21, p. 336.Apex of male abdomen with long straight parallel blunt spines, the outer ones verymuch longer than the middle one; subapical lateral spines hooked.Renamed A. americanum Friese, 1911.maculifrons Smith, 1854. "United States." Tab. 3.*maculosuin Cresson, 1678. " Utah (Putnam); California (H. Edwards)." Tab.2,3.Male; Ann. Mag. Nat. Hist., May 1900, p. 412.montivagum Cresson, 1878. Colorado (Ridings). Tab. 2, 3.mormonum Cresson, 1878. Utah (Ulke). Tab. 3.nebrascense Swenk, 1913. Sowbelly canyon, Sioux County, Nebraska (R. W. Daw-son). Tab. 6. Also in Wyoming.orizabae Dalla Torre=Dianthidium orizabx. Tab. 4.Abdominal scopa black,palliventre Cresson, 1878. California (Hy. Edwards). Tab. 1, 3.Supposed male; Bull. Southern California Acad. Sci., April, 1904, p. 60.palmarum Cockerell, 1904. Palm Spring, California (Davidson). Tab. 1.Tegulre apricot color,paroselae Cockerell, 1898. Mesilla, Nex Mexico (Cockerell). Tab. 1.Male: Ann. Mag. Nat. Hist., May 1900, p. 412.Nesting habits; Minnie Newberry, Psyche, 1900, p. 94.*pecosense Cockerell, 1904. Pecos, New Mexico (Cockerell). Tab. 1, 2.Also at Flagstaff, Arizona (F. C. Pratt).
90 PROCEEDINGS OF THE NATIONAL MUSEUM. tol. 47.
*perplexum Smith, 1854. Georgia. Tab. 3.Legs and margin of thorax ferruginous.This is to be referred to Dianthidium subgenus Anthidiellum on the authority of Mr.Cra^vford.placitum Cressoon, 1879. Nevada (Morrison). Tab. 2, 3.Female clypeus yellow except a median stripe; a yellow band behind ocelli, in-terrupted in middle,placitum praedentatum Cockerell, 1907. Boulder, Colorado (G. Hite). Tab. 2, 5.*porter?e Cockerell, 1900. Las Vegas, New Mexico (W. Porter). Tab. 1, 5.Trior to 1900 confused with A. macuKfrons.Also in Texas, and north to Calgary in western Canada.Type.?Cat. No. 5812, U.S.N.M.*porterffi amabile Cockerell, 1904. Boulder, Colorado (Cockerell). Tab. 5.Ground color of abdomen red.Type.?Cat. No. 9655, U.S.N.M.*porter? personulatum Cockerell, 1907. Boulder, Colorado. (W. P. Cockerell).Tab. 5, 6.Female clypeus entirely black; male with apical lobes and spine of abdomen black.*poudreum Trrus, 1902. Fort Collins, Colorado (Titus). Tab. 2.'^pondreum,'^ in the original publication, is a misprint.Male (11 mm.) differs from atrifrons by the yellow markings, scape entirely black,and last abdominal segment deeply notched, with a blunt black tooth in middle.Variety (? hybrid with tenuifiorsc); Ent. News, 1909, p. 262.Also at Flagstaff, Arizona (Pratt).praedentatum trianguliferum Swenk, 1913. Fort Garland, Costilla County, Colorado(L. Bruner).*psoraleffi Robertson, 1902. Near Carlinville, Illinois (Robertson) Tab 6.Characters: Trans. Amer. Ent. Soc, vol. 29, p. 175.Face of female black.p*ridingsii Cresson, 1878. Georgia (J. Ridings). Tab. 3."A very distinct species, easily recognized by the abdominal fascise being verynarrow and regular." (Cresson.)This is to be referred to Heteranthidium on the authority of Crawford.rodriguezi Cockerell, 1912. Guatemala (Rodriguez).sagittipictum Swenk, 1913. Pullman, Washington (C. V. Piper),saxorum Cockerell, 1904. Rock Creek, California (Davidson). Tab. 1.Abdomen shining, with sparse punctures, the markings orange,scudderi Cockerell, 1906. Fossil in the Miocene shales of Florissant (Scudder).serranum Cockerell, 1904. Rock Creek, California (Davidson) Tab 6.Male 15J mm. Comparison with illustre and conspicuum; Ent. News, 1909, p. 262;also Bull. Southern California Acad. Sci., Feb. 1904, p. 23.*tenuiflor8e Cockerell, 1907. Boulder, Colorado (W. P. Cockerell). Tab. 5, 6.Allied to emarginahan; female ventral scopa sepia brown, pale at sides; male withdark tubercles and scutellum.Also in Montana and western Canada.titusi Cockerell, 1904. Fort Collins, Colorado (E. S. G. Titus). Tab. 1, 5.Allied to emargmatum.transversum Swenk, 1913. Ute creek, Costilla County, Colorado (H. S. Smith).*tricuspidum Provancher, 1896. Los Angeles, California (Coquillett). Tab. 1,3.Titus (1906) gives a new description.*utaliense Swenk, 1913. Logan, Utah.Paratype.?Cat. No. 15260, U.S.N.M.wallisi Cockerell, 1913. Peachland, BritLsh Columbia (J. B. Wallis).Described in Canadian Entomologist, 1913, p. 13.
NO, 2045. NORTH AMERICAN BEES?COCKERELL. 91Genus DIANTHIDIUM Cockerell.A very widespread genus, rather more so than Anthidium, since there is one speciesin Australia.The females use resin in making their nests, and are called by Fabre "Resiniers.
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Pulvilli are present on the feet of this genus, but absent in true Anthidium.Subgenera and Sections.Paranthidium T. and W. Cockerell, 1901.Type.?Perpictum Cockerell.Anthidiellum Cockerell, 1904.Type.?Strigatum Panzer (European).Tables.(1) Cockerell, Bull. Southern California Acad. Sci., vol. 3, 1904, pp. 3-4.(2) Friese, Das Tierreich, Lief. 28, Megacliilinae, 1911, pp. 379-381.(3) Friese, Das Tierreich, Lief. 28, Megachilinte, 1911, pp. 391-392.(4) Swenk, University [of Nebraska] Studies, vol. 14, No. 1, 1913, pt. 25-6.agnatum Cresson, 1878. Mexico (Sumichrast). Tab. 3.Wings fuliginous.apicale Cresson, 1878. Mexico (Sumichrast). Tab. 3.This, bivittatum and toltecum "are closely related, and have a short, broad, robustform somewhat like that of perplexum and notatum" (Cresson).balli Titus, 1902. Ridgeway, Colorado, July 31 (E. D. Ball).Length 14 mm., markings dark rich yellow; allied to venustum.*bivittatum Cresson, 1878. Mexico (Sumichrast). Tab. 3.Mesothorax with two yellow stripes.Guatemala (Barber and Schwarz). Female described; Ann. Mag. Nat. Hist., July,1913, p. 108.boreale Robertson, 1902. Near Carlinville, Illinois (Robertson).Allied to D. notatum; male 8 mm.; legs red, marked with yellow.Characters: Trans. Amer. Ent. Soc, vol. 29, p. 175.*concinnum Cresson, 1872. Texas (Belfrage). Tab. 2."Much like simile, differing principally by the color of the legs. " (Cresson.)The legs are fulvo-ferruginous.Cotype.?Cat. No. 1773, U.S.N.M.*conslin.ile Ashmead, 189G. Near Lcs Angeles, California (Davidson). Tab. 1.Female 7 mm.Parasite; Torymus anlMdii kshmend.The nest is figured and described in Ent. News, 1896, p. 23.Type.?Cat. No. 1GG98, U.S.N.M.cressonii Dalla Torre, 1896. Tab. 2.New name for venustum Cresson, preoccupied.Nest: Bull. Amer. Mus. Nat. Hist., vol. 12, (1906), p. 444, fig. 1.curvatum Smith, 1854. Georgia. Tab. 1.Tliis has been confused with sayi {interruptum Say).*davidsoni Cockerell, 1904. Bear Valley, California (Davidson). Tab. 1.Allied to parvum.Type.?Cat. No. 9653, U.S.N.M.*ehrhorni Cockerell, 1900. Mojave Desert, California (Ehrhorn). Tab. ].Belongs to subgenus Anthidiellum.Type.?Cat. No. 5811, U.S.N.M.*eiseni Cockerell, 1913. San Jose del Cabo, Lower California (Eisen). Belongs toAnthidiellum.Type.?C3it. No. 16225, U.S.N.M.
92 PROCEEDINGS OF THE NATIONAL MUSEUM. vol.47.flavolineatum Smith, 1879. Oajaca, Mexico. Tab. 3.formosum Cresson, 1878. Colorado (Ridings). Tab. 2.Possibly the male of cressonii.gabbii Cresson, 1878. Costa Rica (W. M. Gabb). Tab. 3.*gilense Cockerell, 1897. West Fork of Gila River, New Mexico (Townsend).Tab. 1, 2.Also in Texas.Male: Ann. Mag. Nat. Hist., May 1900, p. 413.Type?Cat. No. 5810, U.S.N.M.gualanense Cockerell, 1912. Gualan, Guatemala (W. P. Cockerell).illustre Cresson (see under Anthidium). Nevada (Monison). Tab. 2.Impatiens Smith, 1879. "South Mexico." Tab. 3.interruptum Say, 1824. "Missouri."Name preoccupied : = snyi.*jugatoriuni Say 1824. "Missouri." Tab. 4.Very close to perpictum, differing in color of tegulfe.*larreae Cockerell, 1897. Near Agricultural College, Mesilla Valley, New Mexico(Cockerell). Tab. 2.Legs red. Relationships; Bull. Southern California Acad. Sci., Feb., 1904, p. 23.Type.?C&t. No. 3507, U.S.N.M.lepidum Cresson, 1878. Georgia (Morrison). Tab. 2.''Resemhles jugatorium in ornamentation of abdomen" (Cresson).*inacruruin Cockerell, 1913. Federal District, Mexico (J. R. Inda).Type.?Cat. No. 16226, U.S.N.M.mexicanum Cresson, 1878. Mexico (Sumichrast). Tab. 3.\Vings fuscous.*iiotatuiaLATREiLLE, 1809. "Carolina." Tab. 2.Redescribed by Cresson, Proc. Ent. Soc. PMla., vol. 2, 1864, p. 376.orizabse Dalla Torre, 1896. Tab. 3.New name for A. atriventre Smith, preoccupied.*parvum Cresson, 1878. Colorado (Ridings). Tab. 1, 2."Marked very much like simile, but smaller" (Cresson).*perpictuni Cockerell, 1898. Ruidoso Creek, New Mexico (Wooton). Tab. 1.Type.?Cat. No. 5807, U.S.N.M.I have recently (Aug. 8, 1913) taken this at (lowers of Helianthus annuus, at Buulder,Colorado.perpictum coloradense Swenk, 1913. Colorado Springs, Colorado,perplexum Smith. See under Antliidium.*provancheri Titus, 1906. Los Angeles County, California (Coquillett)."Closely allied to ulkei, ehrhorni, and simile" (Titus).Type.?Cat. No. 9033, U.S.N.M.pudens Cresson, 1879. Nevada (Morrison). Tab. 2."This may prove to be the female of pudtcum." (Cresson).*pudicum Cresson, 1879. Nevada (Morrison). Tab. 2.Resembles D. parvum, but markings paler; perhaps not specifically distinct.Female; Entomologist, May, 1907, p. 99.*robertsoni Cockerell, 1904. Rock Creek, California (Davidson). Tab. 1.Belongs to subgenus Anthidiellum.Type.?Cat. No. 9654, U.S.N.M.*sayi Cockerell, 1907. Tab. 4.New name for interruptum Say, preoccupied.sayi xerophilum Cockerell, 1907. Mesilla, New Mexico (Cockerell).A very red form
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*simile Cresson, 1864. "Mass., Conn." Tab. 2.
NO. 2045. NORTH AMERICAN BEE8?C0CKERELL. 93Varietal fonn or race in Texas; Proc. Ent. Soc. Wasliington, IX, p. 72.singulare Cresson, 1879. Nevada (Morrison). Tab. 2."Easily recognized (female) by the dilated lateral margin of abdominal segments 2to 4" (Cresson). First two abdominal segments with the yellow bands dividedinto spots.singulare perluteumT. and W. Cockerell, 1904. Wilson's Peak, California (Davidson).First two abdominal segments with the yellow bands deeply notched, but notdivided.subparvum Swenk, 1913. Pullman, Washington (C. V. Piper),tertiarium Cockerell, 1906. Fossil in the Miocene ehalea of Florissant (Scudder.)*texanuni Cresson, 1878. Texas (Belrage). Tab. 2.Nest and female; Melander, Biol. Bull., 1902.Allied to concinnum.toltecum Cresson, 1878. Mexico (Sumichrast). Tab. 3.*ulkei Cresson, 1878. Utah (Hy. Ulke). Tab. 2,4.Comparison with parimm; Entom. News, 1909, p. 261.*ulkei perterritum Cockerell, 1913. Santa Fe, New i^Iexico (Cockerell).venustum Cresson, 1878. Colorado (Ridings, Morrison).Name preoccupied : = crcssonii.The following South American species are known to belong to Dianthidium:D. inerme Friese. D. gregarium Schrottky.D. hertonii Schrottky. D. gregarium continuifasciatum Strand.D. zehratwn Scln-ottky. D. tigrinum Schrottky.D. bicoloratum Smith. D. anisilsi Schrottky.D. bicoloratum tucumanum Vachal. D. paraguaycnse Schrottky.D. steloides Spinola. D. muUiplicatum Smith.D. confusuvi Smith. D. lunatum Smith.D. indescriptum Dalla Torre. D. joergensem Friese (bricchi Schrottky).D. nudum Schrottky. D. brcthesi Schrottky.D. vemonise Schrottky. D. catamarcense Schrottky.D. megachiloides Holmberg. D. autumnale Schrottky.D. heathi Cockerell. D. nectarinioides Sclu-ottky.D. iheringi Schrottky. D. multifasciatum Strand.Anihidium rubripes Friese, which Brethes (1909) referred to Dianthidium, is anAnlkidium. Genus HYPANTHIDIUM Cockerell.Second recurrent nervure passing well beyond second submarginal cell, as inDianthidium, but feet with no pulvilli.aureocinctum Cockerell, 1912. Mexico.*panamense Cockerell, 1913. Gatun, Canal Zone, Panama (A. II. Jennings.)Type.?C&t. No. 16227, U.S.N.M.The type of Eypanthidium is the South American E. fiavomarginatum Smith.Genus HETERANTHIDIUM Cockerell. iSecond recurrent nervure meeting second transverse cubital, as in Anthidium; feetwith pulvilli; end of male abdomen pointed. See also Ent. News, 1909, p. 261*cliippewaens8 Graenicher, 1910. Indian village at juncture of the Lower Tama-rack and St. Croix Rivers, Minnesota (Graenicher).Parafype.?Cat. No. 14915, U.S.N.M.cockerelli Titus, 1902. Rocky Ford (Gillette) and Virginia Dale (Titus), Colorado.Described under Protanthidium.dorsale Lepeletier, 1841. Georgia. This is the type of the genus.Legs ferruginous; hair of thorax and head above rufous.Entomological News, 1904, p. 292.
94 PROCEEDINGS OF THE NATIONAL MUSEUM. vol.47.harbecki Crawford. Sec under Anthidium.*occidentale Cresson, 1868. New Mexico (S. Lewis).Colorado (cf. Cresson).ridingsi Cresson. See under Anthidium..*zebratum Cresson, 1872. Texas (Heiligbrodt).Genus PROTANTHIDIUM T. and W. Cockerell.This is an Asiatic genus; the two American species (cockerelli and chippewacnse)whicii have been referred to it belong to Eeteranthidium.Genus TRACHUSA Jurine.A genus of Europe, represented in California by a single species, which departs fromtypical Trachitsa, and approaches Dianthidium, in the venation. In general, however,it is remarkably like the European T. serratulx.perdita Cockerell, 1904. Tehachapi, California (Davidson).Male about 12 mm.; abdomen black without light marldngs; clypeus and lateralface marks cream color.Genus LITHANTHIDIUM Cockerell.pertriste Cockerell^ 1911. Fossil at Florissant, Colorado.Genus LITHURGUS Berthold.Fox (1902) has proposed to separate the North American species as a genus Lithur-gopsis, but this can hardly be maintained. For an account of the characters of variousspecies of Lithurgits, see Trans. Amer. Ent. Soc, vol. 31 (1905), p. 333.Typical Lithurgus has no pulvillus in either sex ; the males of Lithurgopsis have adistinct pulvillus.*apicalis Cresson, 1875. "Colorado, New Mexico (H. C. Yarrow, C. E. Aiken)."Hair at end of abdomen fulvou.s.*apicalis opuntiae Cockerell, 1902. Mesilla Valley, New Mexico (Cockerell).Hair at end of abdomen dark. Formerly confused with, gibbosus.*compressus Smith, 1853. "United States" (Doubleday).Apparently the male of gibbosus.*echinocacti Cockerell, 1898. La Cueva, Organ Mountains, New Mexico (Townsend.)Visits Echinocactus wislizenii.*gibbosus Smith, 1853. "United States."Female with the facial prominence not at all bilobed.oblongus Fox= Megachilclongula.Lithurgus is known from the Eurppean Miocene (L. adamiticiis Ileer). It is repre-sented in South America by the following species:L. corumbse Cockerell. L. rvfiventris Frieso.L. dubius Sichel. L. pygmacus Friese.L. laticeps Friese. L. planifrons Friese.L. albiceps Friese. L. neoqueenensis Friese.L. huberi Ducke. L. osvwides Friese.L.friesci Ducke. L. macroglossa Friese.