A XEAV CKAB FROM THE EOCENE OF FLORIDA
By ]\1ai{y fJ. IIatiibun
.l.s?^or?/?^? in Zoolof/if. Tutted Sfatcs \'<iil(>t\<iJ Mu^ictim
The material here described was collected in part by the UnitedStates Geological Survey many j^ears ago, and in greater numbers morerecently by the Florida State Geological Survey, which has presentedto the United States National Museum the specimen designated asholotype. Family XANTHIDAEOCALINA, new genusGenotype.?Ocalhia foi'idana^ new species.Carapace broadly suboval. margin lobulate, surface rough. Frontbroad, consisting of 4 lobes which are distinct from the tooth at theinner angle of the orbit. Orbit in an almost vertical plane, subcircu-lar, closed, the inner margin of the inner lower angle being closelyapplied to the lower side of the inner upper angle. Basal articleof antenna remote from orbit terminating not directly below theorbital tooth but below the sinus separating that tooth from the frontproper. Third and fourth segments of male abdomen fused. Chelaeresembling those of Cancel^; longitudinal rows of tubercles onpalm; fingers elongate.This genus is nearest to the Recent genus Carpi/iii.s Leach ' ofwhich one species, Carpilius coralliinis^^ a crab of large s'ze. is notuncommon in the West Indies and the Bahamas. It differs fromOcalhia in its narrower front without a median emargination, in theabsence of a distinct inner orbital tooth, in the inner oi'bital gap,which though narrow is always well defined and occui)ied by theantenna, in the smooth carapace and margins, and in the shorterchelae witli swollen palms and short, stout fingers. In Palaeocar-pU'ms A. Milne Edwards -' (Eocene) the basal article of the antennais long and reaches to the orbit, the carapace is smooth, nontuber-culate. except on the lateral margins, and the fingers are short.
1 In Desmarest. Diet. Sci. Xnt.. vol. 28. 1823, p. 228.
= Ilurbst, Natur. Krabben u. Krebsc, vol. 1, 1783, p. 133, pi. 5, fig. 40.3 Ann. Sci. Nat., Zool., ser. 4, vol. 18, 1862, p. 51.No. 2786?Proceedings U. S. National Museum, Vol. 75, Art. 15.33429?29 1
2 PEOCEEDIXGS OF THE XATIOXAL MUSEUM vol. 75Caloxanthus A. Milne Edwards,^ a Cretaceous genus in France,type C. formosus A. Milne Edwards,^ has a form and rough surfacesimilar to those of Ocalina, but the front is straight and undivided,the circular orbit has a small inner gap, the basal article of theantenna reaches tlie front, the chelipeds are as rough as the carapace.OCALINA FLORIDANA, new speciesPlates 1-3Description of Jiolofi/pe.?Carapace broadly suboval. very convex,more so longitudinally than transversely, antero-lateral margin alittle longer than postero-lateral margin. Front more than one-third width of carapace, orbit subcircular. Antero-lateral marginwith ten thick, similar, closely placed lobules, including that at outerend of orbit. Dorsal surface rough except in the middle or meso-gastric region with Ioav conical tubercles which are largest about theanterior and antero-lateral regions, smallest and more or less coales-cent across the posterior third of the carapace, highest and mostacute near the lateral angle (see also paratype c). Surface punc-tate, especially between tubercles, punctae visible to the naked eye.Orbit circular, closed, ornamented above with ten or eleven tuber-cles, all small, except those at inner and outer angles; below fourlow tubercles. Basal article of antenna obliquely-transverselyplaced, its extremity at a considerable distance from the orbit(paratype h). The broad front is most prominent near the middleand has a deep median V-shaped emargination; each half is sub-divided into two lobes; the inner lobe is triangular, its edge shallow-trilobulate or tuberculate; the outer lobe is transverse and bilobu-late; in the sinus between the lobes there is a lobule.Paratype a shows part of the ventral surface of the body andappendages of a male. The sternum, abdomen, and lower surfaceof carapace are smooth and punctate. The outer maxilliped resem-bles that of (JarpUhis. The left one is preserved but is detachedand turned at a right angle to its normal position; the merus isalso bent at an acute angle with the ischium. The ischium is longerthan broad, the merus is broader than long and widens inwardly,inner margin angled; exognath broad and long, reaching end ofmerus of endognath. Merus of ambulatory legs broad, compressed.The fourth, fifth, and sixth segments of the male abdomen are eachbroader than long; the fifth twice as broad as long, the sixth longer,the fourth still longer ; the cavity formei-ly occupied by the terminalsegment shows it to have been triangular and more than half aslong as broad.
*Ann. Sci. Nat., Zool., Ser. 4, vol. 20. 1863, p. 282; ser. 5, vol. 1, 1864, p. 43.
"5 Idem. vol. 20, 1863, p. .??26. pi. 0. figs. 1-ld : ser. 5. vol. 1, 1864, p. 44.
ART. 15 A NEW FOSSTT. CRAB FROM FLORIDA RATHBUN 3Pamtype d.?Chelipeds stout, siiuilai-. iincciiial. Mcnis nearly aswide as its greatest len?i-tli. smooth, lar?>Lly iiiolllcd willi short trans-verse color marks. Carpus covered with low squamose tubercles.Propodus increasing- in liciaht from proximal to distal end: lowermargin nearly straight, slightly sinuous; upijcr margin arcuate: (Uitersurface very convex transversely, inner surface less so. Outer sur-face, except in the lower part, rough with tubercles arranged largelyin longitudinal rows, and with their acute tii)s pointing distad. Ashort deep transverse furrow outside, ojjjiosite tlie articulation of thedactylus. Fingers stout, comi)resse(l. meeting when closed, tips acute
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l^rehensile edges subacute and furnished with a feAV low tubercles.Propodal finger bluntly carinate, with two longitudinal rows oftubercles and punctae outside; dactylus (paratype e) with four rowsof tubercles on the upper half, two close together above, one on innersurface, the other on outer surface.Measut'emeufs.?Holotype (sex unknown), total length of carapace74, width of same 108, fronto-orbital width 52, width of orbit 5.7,width between orbits 40.G, width of front exclusive of orbital tooth32.5 mm. Paratype c/, length of major propodus about 72, greatestheight of same 35, length of dactylus above 39.3, below 28.5 mm.Occurrence.?Florida ; Eocene series
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Clark, Alachua County; nummulitic limestone 50 feet below thesurface; " Vicksburgian "; 1896; L. ^L Everett, United States Geo-logical Survey; one carapace, dorsal view; paratype c\ Cat. Xo.137871, Ignited States National Museum.Hales Siding, 5 miles east of Xewberry, Alachua County ; in quarryof Ocala Road Material Corporation; depth, 35 feet from top ofOcala limestone: Thomas Sexton, collector; donated to Florida StateGeological Survey by A. T. Thomas; one pair of chelipeds;paratype d.About 2 miles north of Williston, Levy County; in quarry ofThompson "Williston Co.; 30 feet from surface; Ocala limestone;July 6, 1928; G. M. Ponton, Florida State Geological Survey; onespecimen very incomplete, lacking the dorsal surface of the carapace;paratype /.One mile south of Williston. Levy County; in quarry of Ocala-Tampa Lime Rock Co. ; Ocala limestone, 30 feet below surface ; Oc-tober 3, 1928; H. Gunter and G. M. Ponton. Florida State Geologi-cal Survey; parts of three specimens: paratypes /, f/, and h.Ocala, Marion County: 1891; J. Kort. United States GeologicalSurvey; one chela, incomplete, showing part of palm and dactylus;])aratype e; Cat. No. 370957. United States National Museum.Southwest City Limits, Ocala, Marion County; in ])it of FloridaLime Co.; procured from S. Phillips, Janiun-y. 1917: submitted byFlorida State Geoloaical Survev. thiouoh Dr. E. H. Sellards; one
4 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 75specimen is a ruck containing a carapace (holotype. Cat. Xo. 370956,United States National Museum) and another individual showingthe ventral surface of body and appendages (paratype a) ; a secondspecimen (paratype h) shows part of the ventral surface.EXPLANATION OF PLATESOcalina floridaiiaPlate 1Figure 1. Dorsal view of carapace, paratype c, X-%.2. Front view of carapace of holotype, X%.3. Dorsal view of carapace of holotype, X^-,.Plate 2Figure 1. Ventral view of paratype 6, X%. a. Basal article of antenna, o.Inner lower angle of orbit.2. Chelipeds of paratype d, showing outer view of chelae, X7J..Plate 3Figure 1. Ventral view of paratype a, and merus of a larger specimen. Xtu.2. Incomplete chela, paratype e, upper view, Xin.3. Same, outer Aiew, Xra. o
U. S. NATIONAL MUSEUM PROCEEDINGS, VOL. 75, ART. 15 PL. 1
OCALINA FLORIDANA, EOCENE CRABFOR EXPLANATION OF PLATE SEE PAGE 4
U. S. NATIONAL MUSEUM PROCEEDINGS, VOL. 75, ART. 15 PL. 2
OCALINA FLORIDANA, EOCENE CRABFor explanatjon of plate see page 4
U. S. NAT AL MUSEUM PROCEEDINGS, VOL. 75, ART. 15 PL. 3
OCALINA FLORIDANA, EOCENE CRABFor explanation of plate see page 4