DESCRIPTIONS OF SEVENTEEN NEW SPECIES OFRECENT CRINOIDS.
By Austin Hobart Clark,Collaborator, Division of Marine Innrtrhratcn, U. 8. National Museum.
The authorities of the Indian Museum, Calcutta, have recentlydone me the honor of intrusting to me for study the very importantcollections of recent crinoids brought together by the steamer Inves-tigator during the course of her work in the Indian Ocean. Manyof the new species are represented by a considerable number of speci-mens, and of these cotypes have been retained and deposited in theU. S. National Museum. The types themselves are in the IndianMuseum. The completed report on the collection will l)e publishedas one of the series of hivestigafor monographs.I wish here to record my appreciation of the kindness shown meby the authorities of the Indian Museum through the superintendent.Dr. N. Annandale, and by Dr. F. A. Bather, at whose suggestion thecollections were sent to me.Family ZYGOMETRTD.^.Genus EUDIOCRINUS P. H. Carpenter.EUDIOCRINUS ORNATUS, new species.Centro-dorsal a thin disk, the bare polar area flat, 2.5 mm, in diam-eter, the cirri arranged in a single marginal row.Cirri XVIII, 17-18, 10 mm. long; first joint twice as broad as long,second nearly or quite as long as broad, third to fifth twice as longas the proximal diameter, sixth slightly shorter, a more or less markedtransition joint; following joints gradually decreasing in length, theterminal joints being only slightly longer than broad; penultimatejoint about as long as broad. The third to the sixth joints are verystrongly " dice-box shaped," with the distal edge all around produced,except on the dorsal side; from the seventh onward both these fea-tures become less marked, and the cirrus becomes somewhat com-Proceedinqs U. S. National Museum, Vol. XXXVI?No. 1 691
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6.33
634 PROCEEDINGS OF THE NATIONAL MUSEUM. vol. xxxvi.pressed laterally. There are no dorsal spines; opposing spine sharp,prominent, arising from the entire dorsal surface of the penidtimatejoint, equal to about half the diameter of that joint in height; ter-minal claw equal in length to the penultimate joint, stout, andstrongly curved.Disk with a few rather large plates along the ambulacra, and wellplated in the anal area.Ends of the basal rays visible as small tubercles in the angles ofthe calyx; radials projecting slightly beyond the centro-dorsal, gentlyconcave distally; IBr^ and IBro united by syzygy, forming an ob-long syzygial pair from one-third to one-half again as broad as long,the lateral edges straight, barely in apposition basally, the ventro-lateral border slightly produced.Five arms, 85 mm. long; first brachial oblong, about three times asbroad as long; second slightly wedge-shaped, about the same size;third and fourth (syzygial pair) slightly longer on one side than onthe other, half again as broad as the median length; next threebrachials approximately oblong, twice and one-half as broad as long,the following becoming triangular, as broad as long, and after theproximal fourtli of the arm wedge-shaped, as long as l)road, and inthe terminal portion somewhat longer. The lower brachials haveon each side, as far as the lowest pinnule on that side, a slightlyproduced ventro-lateral edge, corresponding with that on the IBrseries; the brachials have a somewhat concave dorsal surface andvery prominent distal ends, everted on the proximal, strongly over-lapping on the distal, which gives the animal a curiously ornate ap-pearance. Syzygies occur between the third and fourth brachials,again between the eighth and ninth, and distally at intervals ofthree, more rarely four, oblique muscular articulations.Pc 5.5 mm. long, moderately stout basally, tapering evenly to thetip, rather strongly prismatic, with twelve joints, the first short, thesecond not quite so long as broad, the third and fourth squarish, thefollowing gradually increasing in length, being nearly or quite twiceas long as broad terminally; P^ similar to Pc, with the same number ofjoints, but somewhat stouter and not tapering so rapidly ; Pa 8.5 mm.long, much stouter than Pc, gradually tapering from the base to thetip with twelve or fifteen joints, the first three about as long asbroad, the following verj^ gradually becoming elongated and abouttwice as long as broad distally; the pinnule is rounded-prismatic;Po similar to Pq ; P?, G mm. long, slender, cylindrical, less stout basallythan Pc, gradually tapering and becoming very delicate in the termi-nal portion, with fifteen or sixteen joints, the first short, the secondand third about as long as broad, the following gradually increasingin length and becoming nearly or quite three times as long as broadin the terminal portion; P3 similar to P&; following pinnules similar.
NO. IGOl. S^EVFINTEEN NEW SFECFES OF CRINOWB?CLARK. 635
^Tadiially decreasing in length to 5 mm., then very slowly increasing,reaching a length of 10 mm. distally. The distal ends of the jointsof the lower pinnules are more or less produced and spinous.Type-,spen7>u'?.?Q.'Ai. No. TC, Indian Museum; lat. 14? 04' 30"N.;"long. J);i? 51' 00" E. ; 41 fathoms.Cotype.?Cut No. 25478, U.S.N.M., from the same locality.Family IITMEROMETRID^.Genus AMPHIMETRA A. H. Clark.AMPHIMETRA MORTENSENI, new species.Centro-dorsal thick-discoidal, the bare polar area flat, 4 mm. or5 nnn. in diameter; cirrus sockets arranged in two closely crowdedalternating marginal rows.'Cirri XVIII-XX, 30^2 (usually about 35), 25 mm. to 30 mm.long; first joint short, about three times as broad as long, second andthird about twice as broad as long, the following gradually increas-ing in length to the ninth or tenth, which is nearly, though neverquite, as long as broad ; next five to seven joints similar, the followinggradually decreasing in length, in almost the whole of the terminalhalf of the cirrus being about one-half again as broad as long; fromthe twelfth or fourteenth onward sharp median tubercles or smallspines are developed on the dorsal side of each joint, those on thelast few^ joints occupying a position slightly jDroximal to median;opposing spine much larger than the processes on the precedingjoints, triangular, the apex median, arising from very nearly thewhole of the dorsal surface of the penultimate joint, equal to abouthalf the diameter of that joint in height; terminal claw longer thanthe penultimate joint, moderately stout basally, but gradually be-coming slender distally, moderately curved.Radials concealed, or just visible beyond the centro-dorsal; IBr^oblong, very short, in close lateral apposition; IBr^ (axillary) verybroadly pentagonal, almost triangular, the lateral edges not quiteso long as those of the IBr^, about two and one-half times as broadas long; IIBr 4 (3-1-4); IIIBr 4 (3-|-4) ; division series and firsttW'O brachials in close lateral apposition and laterally flattened, thedorsal carination only of Pd being visible exteriorly; synarthrialtubercles usually prominent.Twenty to twenty-five arms 150 mm. long; first brachial slightlywedge-shaped, short, about three times as broad as its exterior length,almost entirely united interiorly; second about the same size, butmore pronouncedly wedge-shaped; third and fourth (syzygial pair)oblong, half again as broad as long; next five or six brachials oblong,nearly or quite four times as broad as long, then slowly becomingwedge-shaped and then almost triangular, four times as broad aslong, soon becoming wedge-shaped again, and in the outer half of
636 PROCEEDINGS OF THE NATIONAL MUSEUM. tol.xxxvi.the arm oblong and very short, though somewhat longer againterminally. The proximal discoicli\l brachials are somewhat swollen,and most of the brachials have slightly overlapping distal ends.Syzygies occur between the third and fourth brachials, again be-tween the thirteenth and fourteenth to thirty-first and thirty-second(usually somewhere between the sixteenth and twenty-fifth, withsometimes an extra one from two to four or five brachials beyondthe first), and distally at intervals of two to thirteen (usually eightto twelve) oblique muscular articulations.Pd T mm. long, moderately stout basally, but tapering rapidly andIjccoming slender in its distal half, with about twenty-five joints, atfirst three times as broad as long, becoming twice as broad as long atthe sixth, and squarish in the terminal portion; some of the lowerjoints are bluntly carinate; P^ 10 mm. long, with 30 joints, slightlyless stout basally than Pd and tapering somewhat less rapidly; jointsat first about twice as broad as long, becoming as long as broad at aboutthe eighth, and somewhat longer than broad terminally; P? 15 mm.long, stouter than P,, tapering evenly to a delicate tip, with 30joints, at first about half again as broad as long, becoming squarishat the eighth or ninth, and about twice as long as broad at the tip;P.; 22 mm. long, stouter than the preceding, with 30 joints, at firstbroader than long, becoming squarish about the tenth, and longerthan broad terminally; the pinnule is more or less carinate in itsproximal half and has a moderate supplementary ridge on the distalhalf of the outer side; P4 resembling P., but very slightly longer andslightly stouter and more carinate; Pr, like P;j; P^ 10 mm. long, re-sembling Pj, but somewhat more strongly carinate proximally; fol-lowing pinnules gradually decreasing to 7 mm. in length and losingthe basal carination, then increasing to 12 mm. distally. On somearms Pr, is small as described for P,.,, and again P^ may also be small,while occasionally P. and P^ are similar and P;, is greatly enlarged;sometimes PP., o, and ^ are as described for PP.,, 4, and 5. On oneor more of the inner arms of each ray P.j is often much larger thanon the outer, while the adjacent ])innules are reduced.Type-Specimen.?Cat. No. 42B., Indian Museum; Port Blair,Andaman Islands.rofype.?Q'Ai. No. 25470, TT.S.N.M.; from the same locality.T have dedicated this species to Dr. Th. Mortensen, of Copenhagen,in recognition of his valuable contributions to the knowledge of theEchinoderms.Genus HETEROMETRA A. H. Clark.HETEROMETRA COMPTA, new species.Centro-dorsal discoidal, the bare polar area flat, slightly convex orslightly concave, about 5 mm. in diameter;. cirrus sockets arrangedin a single more or less irregular marginal row.
NO. 1U91. .SEVENTEEN NEW SPECIES OF CRIN0ID8?CLARK. 637Cirri XVIII-XXIII, 31-35, 23 mm. to 25 mm. long; first jointvery sliort, the next three nearly two and one-half times as broad aslong, the following gradually increasing in length to the sixth orseventh, which is about as long as broad; next five to seven jointsusually slightly longer than broad (sometimes squarish), the follow-ing gradually decreasing in length, the terminal fifteen or ratheruiore being half again to twice as broad as long; at about the fifteenthjoint dorsal tubercles are devdoped, at first involving only the distall)ortion of the dorsal surface, later arising in a slightly convex linefrom near the proximal end, the apex being subterminal ; these tuber-cles are narrow, laterally occupying only a small portion of themedian part of each joint, and are slightly rounded dorsally; on thelast three joints the tubercles become somewhat sharper, more erect,and move to a median position; opposing spine small (though largerthan the tubercle on the preceding joint), blunt, arising from theentire dorsal surface of the joint, the apex median or submedian inposition, in height equal to about one-third the diameter of the penul-timate joint; terminal claw somewhat longer than the penultimatejoint, rather stout and strongly curved.Ends of the basal rays and radials concealed ; IBr^ very short andband-like; IBr^ (a:^illary) short, almost triangular, two and one-halftimes as broad as long; IIBr 4(3+4), in apposition laterally, thoughnot laterally flattened; IIBr^ entirely united interiorly; IIlBr 2,rarely 4(3+4) ; IVBr 2, but rarely present.Sixteen to twenty-five arms 110 mm. long; first two brachialswedge-shaj)ed, three times as broad as long exteriorly, the first in-teriorly united; following four or five brachials oblong, about fourtimes as broad as long, then gradually becoming wedge-shaped, almosttriangular, about three times as broad as long, and less oblique andsomewhat longer on the outer portion of the arms. The dorsal sur-face of the arms is perfectly smooth. Syzygies occur between thethird and fourth brachials, again between the thirteenth and four-teenth to twentieth and twenty-first (usually in the vicinity of thefifteenth) and distally at intervals of seven to eleven (most commonlyeight or nine) oblique muscular articulations.Pd ''^?5 nnn. long, moderately stout basally, but tapering ratherrapidly in the proximal half and slender distally, with twenty-fivejoints, at first twice as broad as long, becoming squarish after thetenth; the first four joints are strongly carinate, this carination de-creasing from this point onward and disapi)earing after the middleof the pinnule; P^ 13 mm. long, slightly stouter than Pd basally,tapering gradually, and becoming slender in its distal third, withtwenty-six joints, at first twice as broad as long, becoming squarishafter the ninth and somewhat longer than broad in the terminal por-tion ; the first seven or eight joints are rather strongly carinate and in
638 PROCEEDINGS OF THE NATIONAL MUSEUM. vol. xxsvt.addition have a low sharp ridge running along their exterior surfaceat the base of the carinate processes; Po similar to P^ and of the samelength, but the low ridge just described may be traced to about thetwelfth joint; Pg 9 mm. long with nineteen joints, similar to the twopreceding j^innules, but slightly less stout; P^ small, 6 mm. long,tapering rapidly in the proximal half and becoming very slenderdistally, with sixteen joints, at first twice as broad as long, becoming-squarish about the ninth, and longer than broad distally, the first sixjoints carinate ' like those of the preceding pinnules; Pg similar,5.5 mm. or 5 mm. long; P^ and the following pinnules G mm. longwith seventeen joints, at first twice as broad as long, becoming squar-ish about the eighth and twice as long as broad terminally; the pin-nules are about as stout basally as the two preceding, tapering rapidlyin the proximal half and becoming very slender distally; the carina-tion of the proximal joints is slightly marked on the first four; thiscarination later becomes restricted to the second and third joint only,and disappears entirely in the outer half of the arm.7'ype-specimen.?Cat. No. 4 F.=^ V * i Indian Museum ; Pedro Shoal,north of the Laccadive Islands.Cotype.?Cat. No. ^25480, U.S.N.M. ; from the same locality.HETEROMETRA SINGULARIS, new species.Centro-dorsal discoidal, the bare polar area flat, 1.5 mm. in diam-eter; cirrus sockets arranged in a single crowded, more or less irreg-lar marginal row.Cirri XVII, 21-25, 12 mm. long; first joint short, second abouttwice as broad as long, third somewhat longer, fourth about as longas broad, next two slightly longer than broad, the following grad-ually decreasing in length, the terminal fifteen being one-third orone-half again as broad as long; at the seventh subterminal dorsalspines begin to develop which soon become long and prominent;opi)osing spine large and long, much larger than the spines on thepreceding joints, triangular, the apex terminal, arising from thewhole surface of the penultimate joint and about equal to the diam-eter of that joint in length; terminal claw nearly twice as long asthe penultimate joint, slender, abruptly curved proximally, becomingnearly straight distally.Disk with a few calcareous granules in the anal area, especially onthe anal tube.Radials short, oblong, the dorsal surface with numerous prominentrounded tubercles; IBr^ short, oblong, slightly over four times asbroad as long, in close lateral apposition; IBr, (axillary) broadlypentagonal, almost triangular, twice as broad as long, the lateraledges shorter than those of the IBrj ; IIBr 4(3-|-4) ; joints up toand including the second brachial exteriorly and the fourth interiorly.
NO. 1G91. SEVENTEEN NEW SPECIES OP CRINOIDS?CLARK. 639as Avell as the first two joints of the first throe pinnules, in close appo-sition and sharply flattened, the lateral edi2:es somewhat j)roduced.Eleven arms (in the type), 40 nun. long; first two brachials sub-equal, wedge-shaped, about twice as broad as the exterior length, thefirst interiorly united; third and fourth (sy/ygial paii') slightlylonger interiorly than exteriorly, nearly three times as broad as theinterior length; next four brachials oblong, nearly four times asbroad as long, then becoming almost triangular, about three times asbroad as long, then gradually lengthening (though remaining almosttriangular) to about twice as broad as long, and at a point somewhatbeyond the end of the proximal third rather quickly becoming wedge-shaped, almost oblong, about two and one-half times as broad as long.From about the ninth onward the brachials have prominent distalends, though they do not overlap the bases of the succeeding joints.Syzygies occur betAveen the third and fourth, ninth and tenth, andfifteenth and sixteenth brachials (the second sometimes omitted), antldistally at intervals of seven to ten oblicpie muscular articulations,Pd 4.5 mm. long, moderately stout basally, but tapering rapidlyin the proximal half, and slender distally, with 20 joints, at first abouttwice as broad as long, becoming squarish after the eighth; the secondto the seventh joints are rather strongly carinate; P^ similar, veryslightly longer and stouter; Po G mm. long, considerably stouter andstiffer than the preceding, and rather more strongly carinate basally,with about 20 joints, the first 7 (except for the carinate processes)squarish, the remainder slightly longer than broad, becoming abouthalf again as long as broad distally; the ridge in the distal half ofthe outer side is but little marked ; Pg 3 mm. long, much smaller thanany of the preceding, with about 12 joints, at first broad, becomingsquarish about the fifth, and nearly twice as long as broad distally
;
the second-fifth joints are carinate; following pinnules similar andabout the same length, the joints becoming gradually longer andthe basal carination gradually less ; distal pinnules, 5 mm. long. Onthe arms arising from a IBr axillary, P^, Pj, and Pg are usually asdescribed for Pd, Pi, and Po, and P^ is much smaller, as describedfor Pg ; but occasionally Pj is enlarged and similar to Pg, as described,instead of being small like P^.Type-specimen.?Cat. No. 7A, Indian Museum; southern por-tion of Malacca Strait.Genus STEPHANOMETRA A. H. Clark.STEPHANOMETRA CORONATA, new species.This species is most closely related to S. tenuipinna.Cirri XXII-XXIII, 25-30, 22 mm. long, resembling those ofS. teimipimia,' the longest joints are about one-third longer thanbroad; the ninth, tenth, or eleventh is a well-marked transition joint.
640 PROCEEDINGS OF THE NATIONAL MUSEUM. vol. xxxvi.Radials projecting slightly beyond the edge of the centro-dorsal
;
IBi\ oblong, short, about three and one-half or four times as broad aslong, not in lateral apposition, with a rounded ventro-lateral processin the proximal half; IBr^ (axillary) broadly pentagonal, twice asbroad as long, the lateral edges about half as long as those of theIBr.,, produced into a rounded prominent ventro-lateral process;synarthrial tubercles rather prominent ; IIBr, IIIBr, and IVBr(when present) 2; elements of division series and first brachials withprominent rounded ventro-lateral processes.Thirty-three or thirty-four arms 120 nun. long, in general resem-bling those of S. teiiuipirnin.Pj 14 mm. long, stout, stiff, and spine-like, with fourteen joints, thefirst two somewhat broader than long, the third to the fifth squarish,the remainder becoming gradually elongated and twice as long asbroad distally ; Po and P3 exactly like P^ ; P4 10 mm. long with tenjoints, resembling the preceding; P^ 7 mm. long, spine-like as the pre-ceding, but somewhat more slender, with eight joints; following pin-nules decreasing gradually in length, P^ being 5 mm. long with eightjoints; subsequent pinnules remaining of similar length, but de-creasing in stiffness and increasing in the numl)er of joints, V^., being5 mm. long with twelve joints, the third squarish, the distal twice aslong as broad, only slightly stiffened proximally ; distal pinnulesslender, 9 nun. long.Type-specimen.?Cat. No. 18 H= ^^ijj^, Indian Museum; "India."Cotype.?Qvii. No. 25481, U.S.N.Iil. ; from 'r India."Family COLOBOMETEIDzE.Genus COLOBOMETRA A. H. Clark.COLOBOMETRA DISCOLOR, new species.Cirri XVIII-XXII, 20-40 (usually about 35), 25 mm. to 80 mm.long, slender, resembling those of C. i^ersp'moHa^ but with the distalends of the joints not so strongly spinous.liadials projecting slightly beyond the centro-dorsal; IBr^ oblong,slightly over twice as broad as long, the ventro-lateral borders slightl}'produced into a thin border, by which they are in ap]iosition; YWv.,(axillary) broadly pentagonal, twice as broad as long, the lateral edgessomewhat more than half the length of those of the IBr^, makingwith them a straight line, and with the same ventro-lateral projection;a slight constriction is usually present just below the lateral angles.Ten arms, 80 mm. long, rather slender, resembling in general thoseof C. suaois.Po absent; P^ 0.5 mm. long, small, tapering rapidly to a slenderand delicate tip, with 15 or 10 joints; first joint twice as broad as
NO. IGOl. SEVENTEEN NEW SPECIES OF CRIN0ID8?CLARK. Q/^llong, second somewhat longer, third about as long as broad, the fourthsimilar, Ihe following vory gradually increasing in lengtli to aboutiialf again as long as broad, and bcc;)ining scpiarish again in the(oruiinal -i or 5; 1\, 15 mm. long, moderately stout and very stiti' andspine-like, with about 20 joints, the first about twice as broad as long,the second slightly longer, the third nearly half again as long asbroad, the remainder about twice as Ion"' as broad: be<>:inninir on thesecond join.t there is a faintly indicated, broadlj'^ rounded keel runningalong the middle of the outer side, as on P, ; on the third and. follow-ing joints the distal dorsal edge projects in the line of this keel in anarrow fringe of spines, Avhich broadens on succeeding joints, thespines at the same time becoming longer, aiul is supplemented byadditional spines on the ventro-lateral angles of the joints; P.. similarto P.,, usually about 1 nnn. shorter; P^ 10 nnn. long, resembling P.and P.., though not quite so stiff, Avith 15 joints; P^ and followingpinnules very sloAvly decreasing in length and stiffness, at the sametime becoming more slender, with the spines on the distal ends of thejoints less and less pronounced; P,, is 8 nnn. long and Pj. is 7 nun.long, each with 15 joints; from this point the pinnules very graduallyincrease to 10 mm. in length distally, the distal pinnules being slender,comparatively little'stiffened, with 20 to 22 joints, which have moder-ately everted ends armed with fine spines; the distal pinnules aresomewhat compressed laterally.Type-specimen.?C'At No. 9C., Indian Museum; lat. 1-1? 04' 30"N.; long. 93? 51' 00" E. ; 11 fathoms.Cotijpe.?Cat No. 25-t82, U.S.N.INI. ; from the same locality.Genus CYLLOMETRA A. H. Clark.CYLLOMETRA TAPROBANES, new species.Centro-dorsal thin, discoidal, the bare polar area flat, 2 mm. to 3mm. in diameter; cirrus sockets arranged in a single, slightly irregu-lar, crowded marginal row.Cirri XX-XXI, 25-29, 12 mm. or 13 mm. long; flrst joint short,the next about two and one-half times as broad as long, the followingslowly increasing in length to the fifth or sixth, which is twice asbroad as long, and the tenth or twelfth, which is half again as broadas long, and still further increasing, so that the antepemdtimateand one or tAvo of the preceding joints are about as long as broad;fifth to seventh and succeeding joints with the distal dorsal edgeprominent, forming a low transverse ridge which sloAvly moves ante-riorly, attaining a median position on about the twelfth, and grad-ually narrows distally, becoming reduced to a small median tubercleon the last tAvelve; opposing spine prominent, rather slender, median,equal in height to about one-half the diameter of the penultimate
642 PROCEEniNG^ OF THE NATIONAL MUSEUM. vol. xxxvi.joint; terminal claw slightly longer than the penultimate joint, mod-erately slender, and moderately curved, rather more proximallythan distally.Radials projecting very slightly beyond the centro-dorsal, slightlyseparated distally ; IBr^ oblong or slightly trapezoidal, four times asbroad as long; IBr^ (axillary) broadly pentagonal, tAvice as broadas long; synarthrial tubercles moderately developed.Arms 10, about 80 mm. long, resembling those of C. studeri; dis-tal ends of the brachials very slightly, if at all, produced.Pa absent; P^ 4.5 mm. long, small and slender, Avith about 1-t joints,the first short, the second slightly longer, the third squarish, thosein the distal portion being half again as long as broad; P^ 8 mm.long, stouter and still'er than P^, though not especially enlarged, with15 to 17 joints, the first short, the second and third squarish, the re-mainder one-third to one-half again as long as l)road, becoming againsomewhat shorter at the extreme tip; the joints in the distal half haveslightly enlarged distal ends; P^ G nun. long, less stout than P., butsimilar to it, with 1-1 joints; P^ 5 nun. long, slightly less stout than P.5,but similar, with 12 joints; P^ and following pinnules 4 mm. long,about as stout as P^, but not stiffened, with 12 joints, the third squar-ish, the remainder longer than broad, becoming half again as long asbroad in the distal half; the distal ends of the component joints areslightly everted and spinous ; distal pinnules slender, 7 mm. long, thejoints smooth.Type-specimen.?Cat. No. ^\*^, Indian Museum; off" ColomboLight House, Ceylon; 2C| fathoms.Cotype.?Ciit. No. 25483, U.8.N.M. ; from the same locality.Family THALASSOMETRIDiE.Siibfainily TIIA.1L,ASSOlMEa^RIISr^S;.Genus CROTALOMETRA A. H. Clark.CROTALOMETRA ANNANDALEI, new species.Centro-dorsal columnar, the tip truncated conical as in Af^tero-metra., 5 mm. long by about 5 mm. broad at the base ; cirrus socketsarranged in ten columns of usually three each, the columns of adja-cent radial areas being closely crowded and more or less alternating,the two columns of each radial area being separated by a slightly con-cave median area of about half their width; polar area Avith fivemore or less marked interradial ridges which terminate in five smalltubercles about the apex.Cirri comparatively slender, XXX, 62-79, 65 mm. long; first threejoints approximately equal, short, about twice as broad as long, thefollowing gradually increasing in length, becoming squarish on thefifth or sixth and half again or nearly tAvice as broad as long on the
NO. K.Ol. SEVENTEEN NEW SPECIES OF CRIXOIDS?CLARK. ^43eighth or ninth; next three or four joints simihir, the length thenvery slowly decreasing, the joints in the middle of the cirrus beingsquarish and those in the distal part about twice as broad as long;eighth, ninth, or tenth a transition joint; shortly after the transitionjoint the median part of the distal dorsal edge l)egins to becomeprominent; this very slowly increases in height, arising from pro-gressively more and more of the dorsal surface of the joints, whi(;hbecome progressively more and more carinate, so that in the terminalforty-five or fifty the dorsal surface is produced into a sharp, thin keel,straight in front, convex posteriorly, the outer edge parallel w^ith themedian line of the cirrus, in height equal to about one-third the diam-eter of the joints which bear them; opposing spine small and blunt,arising from the entire surface of the penultimate joint, the apexsubterminal or central, in height equal to about one-third the diam-eter of the penultimate joint; terminal claw small, about equal inlength to the penultimate joint, stout, and moderately curved. Thecirri are rounded in the basal third, then becoming strongly com-pressed laterally and, when viewed from the side, somewhat broader.Ends of the basal rays visible as dorso-ventrally elongated tuber-cles in the angles of the calyx; a deep and narrow cleft betAveen theradials and the centro-dorsal ; radials very narrow, convex proxi-mally, concave distally, with a small, sharp tubercle in the medianpart of the proximal border; IBr^ about three times as broad as long,the proximal border convex, the distal concave, in close lateral appo-sition, and extending rather well up into the angles of the calyx;the lateral edges are more or less denticulate, and there is a low,though sharp, serrate median keel; IBr^ (axillary) slightly longerthan broad, shield-shaped, the posterior border produced into arounded projection incising the IBr,, the anterior edges concave, theanterior angle somewdiat produced, the lateral edges rather stronglydenticulate; it bears a sharp serrate median keel in the proximaltwo-thirds; IIBr 4 (S-j-l), strongly convex dorsally, in close appo-sition and sharply flattened like the IBr series, the lateral edgessomewhat produced and strongly denticulate; IIBr3+4 centrally con-stricted Avith the lateral angles produced as in the other species.Twenty arms, 115 mm. long; first braciiial short, slightly longerexteriorly than interiorly, interiorly united, somewhat incised by thesecond, which is nearly twice as large and has a rounded posteriorprojection; these two brachials, like the IBr^ and o, have a slightlymarked median carination ; third and fourth brachials (syzygial pair)not quite so long as broad, somewhat constricted centrally; next fiveor six brachials almost oblong, about twice as broad as long, thesurface rather strongly concave, then becoming wedge-shaped andsoon triangular, nearly as long as broad, and after the middle of thearm wedge-shaped again and about as long as broad. The arms are
644 PROCEEDINGS OF THE NATIONAL MUSEUM. vol. xxxvi.
at first evenly rounded dorsally, but after the basal third they gradu-ally become compressed and more sharply rounded dorsally, and inthe outer half very narrow and very sharply rounded dorsally, thoughnot really carinate; after the basal third of the arm the brachialsdevelop slightly proje(;ting and finely spinous distal edges. Thedorsal (but not the dorso-lateral) side of the fourth and followingbrachials is covered with fine short spines, which gradually becomecoarser after the proximal third of the arm and tend to arrangethemselves into longitudinal lines; joints of the division series andarm bases with strongly denticulate borders. Syzygies occur betweenthe third and fourth brachials, again between the twenty-fifth andtwenty-sixth to thirty-fifth and thirty-sixth (usually in the vicinityof the twenty-ninth) and distally at intervals of five to seventeen(usually seven to ten) oblicjue nniscular articulations.Pd 1'2 nun. long, moderately stout in the proximal half, but becom-ing slender distally, with about twenty joints, all of which are approx-imately as long as broad, and the basal two-thirds of which arestrongly carinate; P, 10 nun. long, similar to Pn, but less stoutbasally; ]*o C mm. long, nnich more slender than 1\, tapering evenlyfrom the base to the tip, with fifteen joints, the proximal four or fives(iuarish, then longer than broad, and about twice as long as broadterminally. P., similar, (5 nnn. long; P^ and following pinnules 5 mm.long with about thirteen joints, less slender distally than the preced-ing; the joints have slight overlapping spines developed on the distaledge along the dorsal crest ; distal pinnules 10 nnn. long, ratherslender, with about twenty joints, the first short and crescentic, thesecond trapezoidal, about as broad distally as its median length, thefollowing half again as long as broad, the terminal four or five dis-]n'oportionately small; the dorsal crest is sharp and somewhatspinous.TyjH'-spcc'tinen.?Cwi. No. 20A.= ^V'- , Indian Museum; MalayArchipelago; yo fathoms.Cotijpe.?Ko. 2r)4S4, U.S.N.M.; from the same locality.This species is named for Dr. N. Annandale, the superintendent ofthe Indian Museum, thi'ough wlioses courtesy the exceptionally inter-esting collections of that institution have been sent to me for study.Subfamily CHA.KI'rOlVIET'JRIJSrJE;.Genus CRINOMETRA A. H. Clark.CRINOMETRA PULCHRA, new species.Cirri XX-XXIV, 18-20, moderately slender, 30 nnn. to 40 mm.long.Ends of the basal rays visible as rather large tubercles in theangles of the calyx; radials concealed, or at most forming a/X-shaped ridge over the ends of the basal rays; IBr^ very nar-
NO. Id!)!. .SEVEXTEEN NEW SPECIES OF CRiyOIDS?CEARK. 645row, chevron-shaped or crescentic, or entirely concealed; IBr. (axil-lary) larc:e, rhombic, half again to twice as long as broad, the edgesall around smooth and ])rominent, with a moderate rounded mediancarination; IIBr 2, the first very short, the axillary rhombic, abouttwice as broad as long; IIIBr 2, similar to the IIBr, developedinteriorly in 1,2,2,1 order. The division series are perfectly smoothdorsally, in close lateral apposition and sharpl}^ flattened; the edgesof the component joints are slightly prominent, and the axillarieshave a slight broadly rounded median ridge, most pronounced onthe first. One specimen has one IIBr series, and one IIIBr series 4(34-4).Thirty arms, 150 mm. long, resembling, except in ornamentation,those of \he other species of the genus; after the third or fourthbrachial strongly overlapping distal ends are developed, the middleof which is swollen into a broad tubercle which may extend back-ward to the proximal end of the joint; after the thirtieth brachialthis gradually disappears.The pinnules are essentially as in the other species of the genus.Type-specimen.?Q?ii. No. 25473, U.S.N.M.; from Albatross Sta-tions Nos. 2319-2350, off Havana, Cuba; depth between 33 and 279fathoms. CRINOMETRA MARGARITACEA, new species.Cirri XX, 13-15, 20 mm. long.Ends of the basal rays visible in the angles of the calyx, bearing oneor more long tubercles; radials concealed; IBr^ very short, five or sixtimes as broad as long, the edges parallel and slightly curved; IBr?(axillary) rhombic, about two and one-half times as broad as long;IIBr 2; IIIBr 2, developed interiorly, but never present in the fullseries. The division series and first two brachials are slightly convexdorsally and are in close lateral apposition and sharply flattened later-ally ; the first eighteen or twenty brachials are also sharply flattenedlaterally. The axillaries and preceding joints are separated in theouter part of their contiguous surfaces, forming rhombic water pores
;
the first and second brachials are similarly separated interiorly.The ornamentation consists of moderately large blunt tubercles dis-tributed evenly over the surface of the division series, becominggradually less marked after the second brachial and disappearingaltogether at about the end of the proximal fourth of the arm. TheIIBr and IIIBr series and the first two brachials have a low butprominent rounded narrow median carination ; this is continued ontothe arm bases in the shape of prominent median tubercles on eachjoint which disappear at about the end of the proximal fourth ofthe arm.Twenty-one to twenty-nine arms, resembling in structure those ofother species of the genus.Proc.N.M.vol.xxxvi?09 45
546 PROCEEDINGS OF THE NATIONAL MUSEUM. vol. xxxvi.The pinnules are of the type common to most of the species of thegenus, but are somewhat more slender, the genital pinnules not beingso much expanded.Type-specimem.?Cat. No. 25-172, U.S.N.M. ; from Albatross Sta-tion No. 2154, off Havana, Cuba; 310 fathoms.CRINO^ETRA CONCINNA, new species.Cirri XX, 14-18 (usually 15 or 16) 25 mm. to 30 mm. long.Ends of the basal rays visible in the angles of the calyx, bearingone or more long tubercles; radials concealed; IBr^ very shortoften more or less concealed by the centro-dorsal, curved and band-like or narrowly crescentic; IBr. (axillary) rhombic tci approxi-mately triangular, two and one-half times as broad as long, thelateral edges as long as those of the IBr^, and often, like them, re-duced to a point ; IIBr 2 (once 4 (3 -)- 4) and twice 4 united in twosynarthrial pairs in eight sj^ecimens) ; IIBr 2, developed interiorly;edges of the joints to the third brachial everted and raised, usuallybroken up into high blunt tubercles which intermingle more or lesswith similar high blunt, more or less confluent tubercles on the dorsalsurface of the joints; division series (except the IBr) and first twobrachials usually with a high, rather narrow, median ridge, higherthan the tubercles on the dorsal surface of the joint ; this is sometimesi:)artially or entii^ely broken up into two or three dorso-ventrallyelongate tubercles, larger than any of the others on the joints. Theproximal edge of the axillaries and the inner proximal edge of thesecond brachial are curved upward, while the distal lateral anglesof the joints preceding the axillaries, and the inner distal angle ofthe first brachials, are cut away, leaving prominent openings, whichserve as water pores. The division series are only very slightlyconvex dorsally, and are in very close lateral apposition; the firstsixteen brachials are flattened laterally.Thirty arms 150 mm. long, resembling those of other species ofthe genus; the lower brachials to about the fifteenth have stronglyeverted distal ends, which are usually more or less crenulate, or maybe tubercular; there is usually a prominent central tubercle, dorso-ventrally elongate, and also some more or less obsolete tubercles onthe dorsal surface; from the fifteenth onward the brachials arealmost perfectly smooth dorsally.Type-speciinen.?Cat. No. 25476, U.S.N.M.; from Albatross stationNo. 2342, off Havana, Cuba ; 201 fathoms.CRINOMETRA INSCULPTA, new species.Cirri XX, 15-18, 25 mm. to 30 mm. long.Ends of the basal rays visible in the interradial angles as a clusterof high tubercles, with difficulty separable from the similarly modified
NO. 1691. SEVENTEEN NEW SPECIES OF CRINOIDS?CLARK. (547
siirfju'o of the surrounding skeletal elements; radials concealed; H'r^usually concealed excei^t in the angles of the calyx; very short; IBro(axillary) triangular, three oi' four times as broad as long; IIT5r
?4 (3-|-4) and 2, usually both in the same specimen, but the formeralways in the majority; IIIBr 2 (1 + 2), or 2 after a IIBr 2 series(rarely, when developed exteriorly, 4 (3 + 4) or 4 (1 + 2; 3 + 4) ),developed interiorly in 1, 2, 2, 1 order, but never present in the fullseries. The elements of the division series are in close apposition,no water-pores being present. The division series and lower brachialsare but slightly convex dorsally, and are in close lateral appositionand sharply flattened. The elements of the IBr series are thicklyand evenly covered with prominent tubercles resembling those on thedorsal pole of the centro-dorsal. These sometimes arrange themselvesin a more or less linear series in the median line, or there may be amore or less distinct median keel, which, however, is never very wellmarked. This evenly tubercular ornamentation may encroach some-what upon the lower elements of the IIBr series, and always extends aconsiderable distance up into the angles of the calyx and between theIIBr series, narrowing to a point anteriorly, as does the somewhatsimilar ornamentation in Marlanietra suhcarinata. The elements ofthe IIBr and IIIBr series and the lower brachials have more or less(usually strongly) crenulate or tubercular edges, and the dorsah sur-face usually bears a few small scattered tubercles; along the medianline they bear large and prominent, dorso-ventrally elongate, narrow^,dorsally rounded tubercles, which form a conspicuous narrow cari-nation. The lower brachials have very strongly tubercular or dentatedistal ends, in the center of Avhich is a single large tubercle, theselarge tubercles forming a median line of prominent tubercles, whichcontinues the carination of the division series out onto the arms,gradually dying away and disappearing at about the end of theproximal fourth. The prominent eversion of the distal edges of thebrachials becomes distally less and less strongly dentate, at the sametime becoming less and less erect, until at about the twentieth brachialit becomes merely a moderately marked, finely spinous overlap, andso continues to the ends of the arms. The brachials to about thetwentieth are sharply flattened laterally.The pinnules are as in other species of the genus.Type-specimen.?Cat. No. 25477, U.S.N.M. ; from Albatross sta-tion No. 2753, off the windward coast of St. Vincent; 281 fathoms.CRINOMETRA GEMMATA, new species.Cirri XX, 12-15, 20 mm. to 25 mm. long.Ends of the basal rays visible as elongate tubercles in the anglesof the calyx, usually covered with short, fine spines; radials con-cealed, or just visible over the ends of the basal rays; IBr^ very
648 PROCEEDINGS OF THE NATIONAL MUSEUM. vol. xxxvi.short and band-like, of uniform height, strongly curved, the proximaledge everted and dentate, and with a row of small pointed tubercles,sometimes more or less confluent, midway between the anterior andposterior borders; IBr^ (axillary) rhombic, twice and one-half asbroad as long, the anterior and posterior angles approximately equal,the lateral edges about equal to those of the IBr^; IIBr 4 (3+4) (inone specimen twice 2) ; IIIBr 2 (l-|-2), but only present in a singleinstance, developed interiorly. The division series are in closelateral apposition and are sharply flattened laterall}^; they arestrongly convex dorsally, so that the dorsal portion of Pd is exposed.The division series and arms to the fourteenth or eighteenth brachialare thickly covered with numerous uniform, small, sharp, conicaltubercles, which exhibit a tendency to arrange themselves in hori-zontal rows; these are more numerous and more slender along theedges of the division series. Seen without a glass, the proximal por-tion of the animal has the appearance of being finely and evenlygranulated.Nineteen to twenty-one arms, 100 mm. to 125 mm. long, resembling,except for the basal ornamentation as described, those of otherspecies of the genus.The pinnules are essentially as in the other species.Type-speeimen.?Cat. No. 25474, U.S.N.M. ; from Albatross stationNo. 2330; off Havana, Cuba; 121 fathoms.Family ANTEDONID.E.Genus PSATHYROMETRA A. H. Clark.PSATHYROMETRA MIRA, new species.Centro-dorsal conical, rounded at the apex, 4 mm. broad at thebase and 4 mm. high, divided into five radial areas by five shallowinterradial furrow^s, each equal in width to nearly or quite the diame-ter of the adjacent cirrus sockets; cirrus sockets closely crowded,regularly arranged in two converging columns in each radial area,with a single socket, the remnant of a third column, between thedistal ends of the first sockets of the outer columns, which come to-gether just beneath it.Cirri XL, lacking in both specimens.Ends of the basal rays visible as small tubercles in the angles ofthe calyx, but with difficulty separable from the general surface ofthe centro-dorsal and radials; radials even with the edge of the cen-tro-dorsal in the median line, but extending up in the angles of thecalyx and entirely separating the bases of the IBr^; IBr^ oblong,slightly over twice as broad as long, evenly rounded dorsally andlaterally; IBro (axillary) broadly pentagonal, about as long asbroad, the lateral edges not quite so long as those of IBr^, convex,the lateral angles somewhat produced outward.
NO. 1691. SEVENTEEN NEW SPECIES OF CRIN0ID8?CLARK. 649Ten arms, all broken off at the base in the two specimens at hand
;
first brachial slightly Avedge-shaped, about twice as broad as its ex-terior length, entirely free interiorly; second brachial considerablylarger, approximately oblong, not quite so long as broad; third andfourth brachials (syzygial pair) not quite so long as broad; the re-mainder of the arms and the pinnules, so far as can be judged fromthe fragments, are similar to those in other species of the genus. Thesynarthrial tubercles are very slightly marked.Type-specimen.?Cat. No. 9G=^V-^, Indian Museum; hit. 11?31' 40" K, long. 92? 4G' 40'' E.; 188-220 fathoms.Ootype.?Csit. No. 25485, U.S.N.M. ; from the same locality.Genus MASTIGOMETRA A. H. Clark.MASTIGOMETRA MICROPODA, new species.Centro-dorsal low hemispherical, 4 mm. in diameter at the base,the polar area slightlj^ convex or flattened ; cirrus sockets closelycrowded, very numerous, in four or five alternating rows.Cirri L-XC, 10, about 10 nnn.^long; first two joints short, ratherover twice as broad as long, third as long as broad to about one-thirdlonger than broad, fourth and fifth slightly longer; succeeding jointssubequal, about as long as broad; third to sixth joints slightly " dice-box shaped," the remainder with the ventral surface practicallystraight and the dorsal with a slight median concavity (in lateralview) ; no trace of dorsal spines or overlap; cirri becoming somewhatcompressed in the distal two-thirds, and therefore appearing veryslightly broader in lateral view ; opposing spine represented by aslight tubercle, terminally situated, which may be obsolete.Scattered calcareous granules are present along the disk ambulacra,and single interradial plates may be present between the IBr^.Radials even with the edge of the centro-dorsal; IBr^ very short,five or six times as broad as long, of uniform height, not quitein apposition basally, the lateral edges diverging distally; IBrj(axillary) triangular, about half again as broad as long, the anteriorangle somewhat produced, the proximal border as long as the proxi-mal edge of the IBrj.Ten arms, probably about 80 mm, long, their structure being thesame as in M. flageUlfera. The distal intersyzygial interval is threeoblique muscular articulations.Pj 15 mm. long, much stouter basally than the succeeding, thoughtapering to an exceedingly slender and delicate flagellate tip ; Po9 mm. long; following pinnules gradually decreasing in length. Thepinnules are of the same proportions and structure as are those ofM. ffagellifera.Type-specimen.?Cat. No. 14 H., Indian Museum; " ?India."Cotype.?Csit No. 25486, U.S.N.M.; " ?India."
650 PROCEEDINGS OF THE NATIONAL MUSEUM. vol. xxxvi.The only specimen in the collection with a definite locality is asmall and much broken one, which was dredged off Colombo Light,Ceylon, in 26| fathoms.Family PENTACRINITID^.Genus HYPALOCRINUS A. H. Clark.HYPALOCRINUS SPRINGERI, new species.Stem slender, 4 mm. in diameter, rounded-pentagonal in crosssection, the sides smooth, flat, or very slightly convex; interarticularpores extending to the eighth node; internodals, 10 (rarely 9 or 11)of equal size, each face slightly over twice as broad as high ; nodalsslightly longer than the internodals, the small transversely oval cirrussockets touching the distal (lower) border and extending upward tothe proximal fourth of the joint face; neither the supra- nor infra-noglals are modified in any 'way.Cirri slender and delicate, twelve times the diameter of the stem(48 mm.) in length, with 50 joints; first joint very short, the followinggradually increasing in length to the fourth, which is twice as broadas long, and further increasing to the sixth, which is about as longas broad; following joints slightly longer than broad, but in the ter-minal fourth becomiug again about as long as broad; from the twen-tieth or twenty-third joint onward small but prominent median dor-sal tubercles are developed; terminal claw small and blunt, conical,twice as long as broad at the base, slightly longer than the precedingjoint.Infrabasals present, resembling those of Isocrinvs decoriis; basalsprominent externally, rhombic in outline, just contiguous by theirlateral angles, strongly convex exteriorly, bearing from one to threeprominent tubercles; in dorsal view the basals form a figure similarto that made l)v the basals of Isoerinus decoru.^; radials large, stronglyconvex proximally, slightly concave distally, about half again asbroad as long, ornamented with a few coarse, high, tubercles, irregu-larly placed ; IBr, oblong, about twice as broad as long, without orna-mentation; the lateral edges are just in apjjosition, but are not flat-tened ; they are cut away somewhat anteriorly and posteriorly, form-ing small rhombic pores on the lines of articulation between the IBr^and the radials, and the IBr^ and ..\ IBr. (axillary) short and broad,triangular, twice and one-half as broad as long, the anterior edgeseverted and produced into a high scalloped ridge ; IBr^ and o unitedby syzygy; IIB 2, the distal edges of the joints standing out in highprominent scalloped ridges; IIIBr 4 (3+4), the distal edges of theIIIBr^, ., and 4 forming high scalloped vertical ridges.About twenty-five arms 140 mm. long, the terminal 80 mm. beingslender and with only very rudimentary pinnules, as in Metacrinus
NO. 1691. SEVENTEEX NEW SPECIES OF CRICOIDS?CLARK. 551and in 11. naresianiis; first brachial ver}^ obliquely wedge-shaped, thedistal edges forming a straight line with those of adjacent firstbrachials, and standing out in a high scalloped vertical ridge or bear-ing two or throe high tubercles, the intei-ior edges entirely united;
.second brachial smaller, wedge-shaped, about twice as long out-wardly as inwardly, the distal ed^e everted as in the preceding;following brachials obliquely wedge-shaped, about twice as broad aslong, after about the tw^elfth becoming oblong, at first half again asbroad as long, gradually increasing in length, after about the middleof the arm being about as long as broad, and in the terminal portionhalf again as long as broad ; the great eversion of the brachialsgradually dies away as the joints become oblong, giving place to aslight prominence of the distal edge of the brachials, which in theterminal portion of the arm becomes a rather strong overlap.Syzygies occur between the second and third or third and fourthbrachials (more rarely between the fourth and fifth), again betweenthe fifteenth and sixteenth to thirty-first and thirty-second (usuallyin the vicinity of the twentieth), and distally at intervals of fromfour to nineteen oblique muscular articulations, the interval beinglong in the proximal, short in the distal part of the arm.The pinnules are in general like those of H. 7iaresianus.Tyfe-specimen.?Cat. No. - -^- , Indian Museum; lat. 13? 47' 49"K, long. 73? 07' 00" E. ; 636 fathoms.Cotype.?Cat. No. 25487, U.S.N.M.; from the same locality.This species is dedicated to Mr. Frank Springer, the eminentauthority on the Crinoidea.HYPALOCRINUS ORNATUS, new species.In general like II. f^pringet'l^ but a smaller and more delicatespecies.Stem as in H. spritigeri, but only 3 mm. in diameter; cirri propor-tionately more slender, 30 mm. long (ten times the stem diameter)with 40 joints, the dorsal tubercles commencing at about the seven-teenth ; basals as in Isoerinus decorus, without ornamentation ; radialswithout dorsal ornamentation, but with the distal edges everted andproduced into a high, thin, scalloped overlapping ridge; IIBr 4(3+4).Eighteen to twenty arms, about 95 mm. long from the radials.Type-specimen.?Cat. No. ^V^, Indian Museum; Andaman Sea;200 fathoms.Cotype.?Cat. No. 25488, U.S.N.M. ; from the same locality.