PROCEEDINGS OF THE UNITED STATES NATIONAL MUSEUM
issued iMfii\A,^M iy the
SMITHSONIAN INSTITUTIONU. S. NATIONAL MUSEUMWashington: 1934
Vol. 83 No. 2973AMERICAN MUSCOID FLIES OF THE GENERA CERA-TOMYIELLA AND PARADIDYMA
By H. J. ReinhardTexas Agricultural Experiment Station, College Station, Tex.
This paper contains a discussion of the generic characters of thetachinid genera Geratomyiella and Paradidymal keys for separatingthe species in both sexes, and descriptions of 24 species, of which 15are new to science. The material used is preserved in the UnitedStates National Museum, the Kansas University Museum, and myown collection.I am under great obligations to the late Dr. J. M. Aldrich for theprivilege of studying the material in the National Museum collection,which he kindly assembled and forwarded to me, and for carryingon considerable correspondence, in which \qvj helpful notes on thegenotype of Paradidyma were supplied through the generous coop-eration of Miss Daphne Aubertin, of the British Museum. To Dr.R. H. Beamer I am indebted for permission to examine the typespecimens of Lachnomma magnicornis Townsend and Atrophopodahraueri Williston, in addition to other material, in the KansasUniversity Museum.The genera here under consideration may be readily recognizedby the row of bristles extending down the inner margin of the para-facial and the bare first vein of the wing. In the female the foreclaws and pulvilli are small or atrophied. There are a number ofgenera sharing this combination of characters except that the firstvein of the wing is beset with hairs. Among approximately 200specimens of Geratomyiella and Paradidyma examined in the pres-73007?34 1 a
10 PROCEEDINGS OF THE NATIONAL MUSEUM vol.83
ent study, I have seen but one specimen {P. singularis) in which thefirst vein is not entirely bare; this specimen has only one or twohairs present on the vein. When the persistently bare first vein isconsidered throughout the group, the character seems to be ofgeneric importance, and I have included in the present genera onlythose forms that agree in this respect.Genus CERATOMYIELLA TownsendCeratotnyiella Townsend, Ti*ans. Amer. Ent. Soc, vol. 18, p. 379, 1891. (Geno-type, C conlca, new species.)
?
Brauek and Bekgenstamm, Die Zweifliiglerdes Ivaiserliclien Museums zu Wien, no. 6, p. 189, 1S93.
?
Aldrich, Catalogueof Nortlj American Diptera, p. 427, 1905.AtropJiopalpus Townsend, Ent. News, vol. 3, p. 130, 1892. (Genotype, A.august icornis, new species.)
?
Coquillett, Revision of the Tachinidae ofAmerica, p. 12C, 1897.
?
Aldrich, Catalogue of North American Diptera,p. 475, 1905.Oedemapcza Townsend, Smithsonian Misc. Coll., vol. 51, p. 65, 1908. (Genotype.Atrophopoda townsc^idi Williston.)All of the type species concerned have been examined in arrangingthe above synonymy. The principal character, listed among othersin the description of Atrophopalpus^ is the reduced size of the palpi
;
although somewhat smaller than in the genotype of Geratomyiella^they are nevertheless distinctly developed, and the relative differencein size can hardly be considered of more than specific importance.Oedemapeza was established by the mere citation of a species as thetype. The genus Geratomyiella is closely related to Paradidymalfrom which it differs most obviously in having the eyes bare.Generic characters (from the type species).?Eyes bare. Frontnot prominent at antennae, where the length of head distinctlyexceeds the vibrissal axis. Antennae nearl}^ as long as face, insertedabove middle of eyes, basal segments short. Arista with short basalsegments. Face receding, the depression broad and deep. Facialridges practically bare, weakly divergent below. Parafacial with arow of bristles on the inner margin extending from the lowermostfrontals to level with lower edge of eye. Vibrissae situated on thefront edge of mouth. Proboscis short, labella fleshy. Palpi rathershort and slender. Cheek one-third to two-fifths the eye height.Male with one pair, female with two pairs, of proclinate orbitalbristles. Frontals extending below middle of second antennal seg-ment, uppermost larger, reclinate. Ocellars present, proclinate.Inner verticals developed, outer pair moderately developed in female,vestigial in male.Thoracic chaetotaxy: Acrostichal, 2, 1 (postsutural pair hairlike,situated in transverse line with hindmost dorsocentrals) ; humeral, 2;
CERATOMYIELLA AND PARADIDYMA EEINHARD Hposthumeral, 1; presutural, 1; dorsocentral, 2, 3; notopleural, 2;intraaiar, 2; supraaiar, 3; postalar, 2; hypopleural, 3 or 4; ptero-pleural, 1 (small); sternopleural, 1, 1; scutellum with two lateralsbesides one smaller decussate apical pair. Postscutellum normallydeveloped. Infrasquamal hairs absent.Abdomen rather narrow and slightly elongate, without discalbristles.Legs long and slender ; fore tarsi in female compressed and swollen,the basal segment nearly as long as tibia, claws and pulvilli minute
;
in male the fore tarsi normal with short but distinct claws andpulvilli.Wings normal in shape; veins bare except the third, which issetulose almost to small cross vein; last section of fifth vein short;apical cell closed with a short petiole reaching costa shortly beforewing tip ; costal spine developed.
KETT TO SPECIES OF CEBATOMYIEXLA
1. Apical cell closed and usually petiolate 3.Apical cell open 2.2. Male with orbitals ; epaulets black ; last three abdominal seg-ments largely gray pollinose, the narrow hind margins snb-shining, male only (New Jersey) (5) orbitalis, new species.Male without orbitals; epaulets red; last three abdominal seg-ments shining black on apical half; third antennal segmentin female very slender (Florida) (4) angusticomis (Townsend).3. Legs black 4.Femora reddish yellow (United States, widespread) (1) conica Towusend.4. Parafacial bristles reduced to small hairs in upper half ofrow; fourth abdominal segment polished black, usually with-out pollen ; petiole of apical cell shorter than small crossvein (Texas, New Mexico, Arizona) (3) bicincta, new species.Parafacial bristles not noticeably reduced in size above; fourthabdominal segment thinly pollinose on basal third; petiole ofapical cell about one-third the length of apical cross vein,male only (West Indies) (2) townsendi (Williston).(1) CERATOMYIELLA CONICA TownsendCeratomyiella conica Townsend, Trans. Amer. Ent. Soc, vol. 18, p. 380, 1891.Male.?Front at vertex 0.3 and 0.31 of the head width (two speci-mens), gradually widening to antennae; median stripe reddishbrown, hardly more than half the parafrontal width on entire length
;
parafrontals black and subshining, viewed from the side thinly polli-nose; face and parafrontals thinly gray pollinose; antennae reddishblack, third segment broader than parafacial, six or seven timeslonger than second; arista brown, thickened about to middle; palpi
12 PROCEEDINGS OF THE NATIONAL MUSEUM vol.83pale yellow, slender to tip ; back of head shining black above, thinlygray pollinose and sparsely pale haired downward.Thorax black, lightly dusted with gray pollen ; dorsal black stripespoorly defined behind suture; scutellum black, indistinctly polli-nose, without discals; infrasquamal hairs absent; calypters trans-parent, front lobes colorless, hind ones tawny.Abdomen shining black, last three segments pruinose on basalthird ; first and second segments bearing a pair of median marginalbristles ; third and fourth each with a marginal row of six or eight
;
no discals.Legs reddish yellow, tibiae more or less infuscated, tarsi black.Wings with a tawny tinge, paler on hind margins ; hind cross veinperpendicular to fourth, which it joins slightly nearer bend thansmall cross vein ; epaulets red.Female,?Front at vertex 0.27 and 0.25 of the head width in twospecimens; third antennal segment slender, about five times longerthan second ; arista thickened near the base, clothed with short hairscheek about one-third the eye height; front tarsi as noted undergeneric description.Length.?5 mm.Remarks.?Redescribed from 2 males and 2 females: 1, CollegeStation, Tex., December 4, 1932 (H. J. Reinhard) ; 1, A. and M.College, Miss. (F. M. Hull) ; 1, Opelousas, La., without collector'slabel; and the other, Dead Run, Fairfax County, Va. (R. C. Shan-non). The type locality is Carlinville, 111. The species is easilyrecognized by the red femora.
(2) CERATOMYIELLA TOWNSENDI (Williston)Atrophopoda tounsendi Williston, Trans. Ent. Soc. Loudon, 189G, p. 356, pi. 11,fig. 93.Paradidyma toumsendl Aldkich, Catalogue of North American Diptera, p. 474,1905.Oedemapeza townsendi Townsend, Smithsonian Misc. Coll., vol. 51, p. 65, 1908.Male.?Front at vertex 0.28 and 0.27 of the head width in twospecimens, widening uniformly downward; parafrontals blackish,thinly dusted with white pollen; median stripe reddish brown,slightly narrower than one parafrontal on entire length; frontalbristles about five in number, extending to level with apex of sec-ond antennal segment, uppermost pair of moderate length, reclinate,others directed inward; ocellars small but distinct, proclinate; oneproclinate orbital bristle situated at middle of front; verticals onepair (inner) developed; face moderately excavated, receding, graypollinose, its ridges bare; parafacials narrow, gray pollinose, bear-ing a row of moderate-sized bristles on inner margin, bare outside
CERATOMYIELLA AND PAEADIDYMA EEINHARD 13the main row ; vibrissae situated on oral margin ; antennae as long asface, basal segments yellow, third brownish, thickly covered withdense pale pubescence and about seven times longer than secondsegment; arista about as long as third antennal segment, thickenedon proximal third, reddish, basal segments darker, short but dis-tinct; eyes bare; cheek in profile about one-fifth the eye height;proboscis short ; palpi slender, pale yellow ; back of head subshiningabove, gray pollinose and pale haired below.Thorax and scutellum black, dusted with gray pollen ; mesonotummarked with two heavy black stripes, which extend from the anteriormargin to base of scutellum without interruption at suture ; chaeto-taxy as in C. conica; postscutellum thinly gray pollinose; no infra-squamal hairs ; calypters transparent, faintly tawny.Abdomen rather slender, shining black; intermediate segmentswith bluish-white pollen bands on basal fourth above, becomingwider on the sides and venter ; fourth segment pruinose on proximalthird; two basal segments each with a pair of median marginalbristles; third and fourth with a marginal row; no discals; genitalsegments black, small, and retracted.Legs rather long and slender, blackish; front claws and pulvillishort but distinct.Wings tinged with brown on anterior margin, grayish hyalinebehind ; veins bare except third, which is haired to small cross vein
;
hind cross vein perpendicular to fourth, joining it midway betweenbend and small cross vein ; apical cell closed, the petiole nearly one-third the length of the broadly concave apical cross vein, reachingcosta shortly before tip of wing; costal spine small.Length.?i mm.Reviarks.?Redescribed from two males in my collection from theWest Indies, donated by D. G. Hall ; labeled " Mustique Island,May."The species varies considerably in the degree of infuscation of thewings. Four specimens in the United States National Museum, ac-cording to Dr. J. M. Aldrich, all have the wings more distinctlyinfuscated than described above, agreeing better in this respect withWilliston's description. The species is closely related to conica^from which it is readily distinguished by the black legs, longerpetiole of apical cell, and other characters.I have not seen any specimens of the female. The type localityis St. Vincent, British West Indies.
(3) CERATOMYIELLA BICINCTA, new speciesMale.?Front at vertex 0.297 of the head width (one specimen),not prominent at antennae; parafrontals gray pollinose to vertex,
14 PROCEEDINGS OF THE NATIONAL MUSEUM vol.83bearing a few scattered inconspicuous hairs outside of frontal rows;median stripe short, brownish black, narrower than one parafrontalon entire length; verticals two pairs, outer ones about half as longas inner, curving outward and backward; ocellars present, procli-nate; frontals about five in the row, extending about to middle ofsecond antennal segment, uppermost pair larger and reclinate, thepair next in front of these erect, decussate at tip, others directedinward; one proclinate orbital bristle situated midway between theanterior ocellus and base of antennae; face rather long and deep,gray pollinose on reddish ground color, its ridges hardly divergentdownward, practically bare; vibrissae on the front edge of themouth ; parafacials narrow, gray pollinose, bearing a row of bristleswhich are reduced to small hairs on the upper half; antennae red-dish yellow, as long as the face, third segment darker, about seventimes longer than second, which bears one long and numerous shorterbristles on front side; arista about as long as third antennal seg-ment, finely pubescent, thickened and yellow on proximal two-fifths,black beyond, penultimate segment about twice as long as broad;proboscis short, labella fleshy; palpi slender to tip, pale yellow,bearing a few short black hairs near apex; cheek bare, gray polli-nose on red ground color, about one-fourth the eye height; pos-terior orbits broad to middle, thence narrowed upward, thickly cov-ered with gray pollen; back of head sparsely pale haired, gray pol-linose; eyes bare.Thorax black, gray pollinose; mesonotum marked with threebroad pale gray and two slightly narrower opaque black stripes,which extend from the anterior margin to base of scutellum; thelatter black, dusted with changeable gray pollen, bearing two lat-erals (with a large supernumerary bristle on one side), apical pairstrongly decussate; other details of chaetotaxy as in cornea; calyp-ters transparent, white; postscutellum normal; no infrasquamalhairs.Abdomen shining black; intermediate segments with silvery bandson basal fourth to third, extending on venter to median line; firstand second segments each with a pair of median marginal bristles;third with four marginals, none below the lateral pair; fourth witha complete marginal row of about 12; no discals; genitalia small,retracted; inner forceps blackish, short and united, moderately wideat base, which is haired behind, tapering sharply to middle, slenderand shining beyond, tip acute; outer forceps largely yellow, convexon outer side, tips acute and blackish, slightly longer than innerones; penis short, black, apex bordered with a narrow white mem-brane; fifth sternite narrowly and deeply incised, the lobes shiningblack, sparsely clothed behind with short, black hairs.
CERATOMYIELLA AND PARADIDYMA?REINHAED 15Legs largely black, trochanters yellow, coxae less distinctly so;fore tarsal segments normal, the claws and pulvilli short but dis-tinct; mid tibia with a whorl of three bristles near middle, the oneon outer front side stout; hind tibia with only three strong bristleson outer posterior edge.Wings with a brownish tinge on broad anterior margin, some-what paler behind; venation bare except third vein, which is setu-lose almost to small cross vein ; fourth vein with a broadly roundedstumpless bend, concave beyond; hind cross vein perpendicular tofourth, joining it midway between small cross vein and bend; api-cal cell closed, petiole short, reaching costa shortly before the wingtip; epaulets blackish; costal spine developed but not very strong.Female.?Front at vertex 0.296 of the head width (average of five
:
0.3; 0.29; 0.29; 0.29; 0.29), widening uniformly to antennae; theusual two proclinate orbitals present; verticals two pairs; antennaea little shorter than face, third segment narrow but wider than para-facial below, about five times longer than the second; arista thick-ened on proximal fourth, pubescent to tip; cheek about one-fourththe eye height; fore tarsi comjDressed, the segments slightly swollen,claws and pulvilli minute.Length.?Male, 6 mm ; female, 5.5 to 7 mm.Type.?MalQ, U.S.N.M. no. 44758, from College Station, Tex.Remarks.?Described from eight specimens. In my collection, 1male and 3 females, College Station, Tex., September 25 and October11, 1930, August 24, 1931, and October 19, 1933 (H. J. Reinhard).In the United States National Museum, 4 females as follows: 1,Brownsville, Tex., June (C. H. T. Townsend) ; 1, Yuma, Ariz., June26, 1917 (J. M. Aldrich) ; 2, Las Cruces, N.Mex., one labeled " Ckll.2293, Aug. 1894 ", the other without collector's label. The specimencollected by Cockerell also bears Coquillett's determination label,Paradldyma siiigularis Townsend.The species is strictl}^ congeneric with the type species, conica^from which it differs in having black legs and broad, well-definedthoracic stripes; in being more robust in build; and in othercharacters. (4) CERATOMYIELLA ANGUSTICORNIS (Townsend)AtropTiopalpus angusticornis Townsend, Eut. News, vol. 3, p. 130, 1892.Male.?Front at extreme vertex 0.271 of the head width (one speci-men), widening gradually downward to antennae; sides of front,face, and cheeks gray pollinose ; median stripe red, narrower than oneparafrontal; outer verticals and orbitals absent; ocellars proclinate;uppermost frontal reclinate, others directed inward, extendingbelow middle of second antennal segment; face rather deeply exca-vated, receding and concave above mouth in profile; facial ridges
16 PROCEEDINGS OF THE NATIONAL MUSEUM vol.83moderately divergent downward, practically bare; vibrissae on levelwith front edge of mouth; antennae as long as the face, basal seg-ments yellow, second distinctly longer than the first and about one-sixth the length of third, which is black except at extreme base;arista brown, thickened on basal fourth, slender beyond middle, sec-ond segment short; parafacial bearing a row of bristles, which be-come longer and stronger downward, a few hairs outside the largebristles on lower extremity; cheek nearly two-fifths the eye height;palpi but little longer than thickness of proboscis at point of attach-ment, pale yellow, bearing two black hairs near apex ; eyes practicallybare.Thorax and scutellum black, gray pollinose; four black stripeson mesonotum, outer ones broader, stopping shortly before base ofscutellum. Chaetotaxy as in conica; scutellum with two laterals(posterior pair large and divergent), one decussate apical and aweak discal pair ; postscutellum thinly pollinose ; infrasquamal hairsabsent; calypters tawny.Abdomen shining black on broad hind margins of last three seg-ments, basal segment without, second with one pair of large medianmarginal bristles; third and fourth each with a complete row ofabout 12; no discals; inner forceps united, slender on apical half,in profile slightly bowed forward at tip; penis simple, terminatingin a short pale membrane ; fifth sternite with a narrow deep incision,the lobes blackish.Legs reddish black; claws and pulvilli moderately elongated.Wings subhyaline ; venation normal ; third vein with hairs extend-ing almost to small cross vein ; apical cell open shortly before the wingtip; costal spine strong; epaulets red.Female.?Front at vertex 0.292 of the head width (one specimen)
;
third antennal segment very slender, four to five times longer thansecond ; outer verticals not developed ; two pairs of proclinate orbit-als; fore tarsi rather slender, compressed and tapering outward,claws and pulvilli minute.Length.?6.5 to 7.5 mm.Remarks.?Redescribed from one male and one female in theUnited States National Museum from Miami, Fla., October 8 and 15(C. H. T. Townsend).
(5) CERATOMYIELLA ORBITALIS, new gpedesMale.?Front at vertex 0.25 of the head width (one specimen),widening gradually downward to antennae ; parafrontals gray polli-nose to vertex, sparsely haired; median stripe reddish brown, aswide as one parafrontal except at antennae ; one pair of weak orbitalspresent; outer verticals not developed; ocellars proclinate; frontal
CEEATOMYIELLA AND PARADIDYMA REINHARD 17bristles about seven in number, descending below middle of secondantennal segment, uppermost pair reclinate, hardly larger than thenext preceding ones in the row; face moderately receding, ratherlong and deeply excavated, its ridges not strongly divergent below,bare except one or two bristles above vibrissae, which are situatedon the front edge of mouth ; antennae as long as face, basal segmentshort, second wholly yellow, about one-fifth the length of third,which is blackish beyond the narrow base; arista brown, thickenedon basal fourth, second segment short; parafacials gray pollinose,bearing a row of strong bristles along the inner margin, those inlower part of row larger than any of the frontals, a few hairs extend-ing outside the large bristles on the lower extremity; cheek abouttwo-fifths the eye height; palpi small, yellow, bearing two slenderblack hairs near tip; labella pale yellow, fleshy; eyes bare; back ofhead gray pollinose ; thinly clothed with pale hairs.Thorax and scutellum black, gray pollinose; mesonotum markedwith four narrow black stripes; chaetotaxy as in conica; postscutel-lum normally developed ; infrasquamal hairs absent ; calypters white.Abdomen black, subshining; the pollen gray, apparent on sidesof first segment, thicker on the basal margins of last three and ex-tending rather thinly past the middle on each; first segment witha weak pair of median marginals ; second with one pair (broken off,scars indicating strong bristles) ; third and fourth each with a com-plete marginal row ; no discals on anal segment.Legs black, basal segments reddish; claws and pulvilli elongate.Wings subhyaline; veins bare except third, which is setulose two-thirds of the distance to small cross vein; last section of fifth veinshort; fourth vein with a broadly rounded bend, beyond slightlyconcave to costa; apical cell open just before the extreme wing tip:epaulets black ; costal spine developed.Length.?8 mm.Female.?Unknown.r^/pe.?Male, U.S.N.M. no. 44759.Remarks.?Described from one male in the United States NationalMuseum from Hammonton, N.J., August 23, 1903; no collector'slabel. Genus PARADIDYMA Brauer and BergenstammParadidyma Bbaueii and Bergenstamm, Die Zweifliigler des kaiserlichenMuseums zu Wien, no. 5, p. 382, 1891; no. 6, p. 184, 1893. (Genotype,Dldyma validinervis "Van der Wulp.)
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Coquillett, Revision of theTachinidae of America, p. 126, 1897.
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Aldrich, Catalogue of North Amer-ican Diptera, p. 474, 1905.Atrophopoda Townsend, Trans. Amer. Ent. Soc, vol. 18, p. 373, 1891 [Geno-type, A. singularis, new species (female only)]; Smithsonian Misc. Coll.,vol. 51, p. 66, 1908.73007?34 2
18 PEOCEEDINGS OF THE NATIONAL MUSEUM vol.83Lachnomma Townsend, Trans. Amer. Ent. Soc, vol. 19, p. 103, 1892. [Geno-type, L. magnicornis, new species (male only) ^Atrophojjoda singularisTownsend. I have examined the type specimen.]Microchira Br.\ueb and Bekgenstamm, Die Zweifliigler des kaiserlichen Mus-eums zu Wien, no. 6, p. 188, 1893. [Genotype, M. mexicana, new species(male only) ?Paradidjjma apcrta Brauer and Bergenstaram, loc. cit., p,187.]Lachnommopsis Townsend, Proc. U.S. Nat. Mus., vol. 49, p. 421, 1915. [Geno-type, L. armata, new species (male only).]Phytoadmontia Townsend, Proc. U.S. Nat. Mus., vol. 49, p. 626. 1916. (Geno-type, Admontia sctigera Coquillett.)Ill arranging the foregoing synonymy, I have examined all thetype species involved. Atrophopoda hraiceri Williston, listed as asynonym of Paradidyma by Aldrich ^ and Coquillett,^ does not comewithin the limits of the genus as restricted herein. I have examineda male type specimen in the Kansas University Museum. It hasthe eyes practically bare, and the first vein of the wing is setuloseon almost the entire length. For this species Townsend ^ estab-lished Diaphoropeza^ which Coquillett ^ also placed in synonymywith Paradidyvia. Under the rules of the International Code thegenus is valid, although no description of the generic characters wasgiven.Generic characters (from the type species).?Eyes distinctly hairy.Front in male narrowed behind and rather prominent at base ofantennae. Face receding, moderately excavated, its ridges normallydivergent downward, in profile concave above mouth with the frontedge of latter slightly prominent between vibrissae. Antennae in-serted about on level with middle of eye, extending almost to oralmargin, basal segments subequal in length in male. Second seg-ment of arista short. Parafacial with a row of macrochaetae on theinner margin extending from lowermost frontal almost to loweredge of eye. Vibrissae situated on level with oral margin. Facialridges bearing a few bristles and hairs above the vibrissae. Pro-boscis shorter than height of head, distal segment moderately slender,labella fleshy. Palpi normal in size, slender, tips hardly thickened.Frontal rows moderately divergent beneath base of antennae, extend-ing to base of third segment. Ocellars present, proclinate. Orbitalsabsent in male. Cheek about one-half the eye height. Back ofhead densely pale haired, with a row of black hairs below the orbitalfringe.Thoracic chaetotaxy: Acrostichal, 2, 1 (postsutural pair well de-veloped, situated in transverse line with posterior dorsocentral pair) ;
* A catalogue of North American Diptera. Smithsonian Misc. Coll., vol. 46, p. 474,1905.2 The type-species of the North American genera of Diptera. Proc. U.S. Nat. Mus.,vol. 37, p. 532, 1910.8 Smithsonian Misc. Coll., vol. 51, no. 2, p. 64, 1908.
CEEATOMYIELLA AND PAEADIDYMA REINHAED 19dorsocentral, 3, 3; humeral, 2; posthnmeral, 1; presutiiral, 1; noto-pleural, 2 ; intraalar, 2 ; supraalar, 2 ; postalar, 2 ; hypopleural, 4 or5; pteropleiiral, 1 (^mall) ; sternopleural, 2, 1 (lower anterior onesmall). Scutellum with two lateral, one decussate apical and asmall discal pair. Postscutellum normally developed; infrasquamalhairs present.Abdomen without discal bristles on intermediate segments.Legs rather long and slender, claws and pulvilli elongate in male.Wing3 with first vein bare; third bristled nearly to small crossvein. Costal spine developed. Last section of fifth vein less thanhalf as long as the preceding one. Hind cross vein oblique to fourth,which it joins a little nearer bend than small cross vein. Apicalcell narrowly open, reaching costa well before extreme tip of wing.KEY TO SPECIES OF PABADIDYMAMALES1. Apical cell closed and petiolate 2.Apical cell oi>en 4.2. Mesonotum gray pollinose, the black stripes couspicuous andusually fused into a single liroad pair, wliich extends to baseof scutellum 3.Mesonotum subshining, at most lightly dusted with pollen, theblack stripes poorly defined or entirely obliterated behindsuture : third antennal segment ordinary ; arista thickenedon basal two-fifths (Indiana, Illinois, Maryland, Virginia).(23) petiolata, new species.3. Last three abdominal segments: with silvery bands on basalthird, the remainder of these segments including the firstshining black; third antennal segment entirely black(United States, widespread) (22) singularis (Townsend).Pollen on abdomen not in defined basal bands, first segmentconspicuously pollinose on the sides above, the second withpollen extending to hind margin ; base of third antennalsegment yellow to insertion of arista (Brazil).(21) brasiliana, new species.4. Last section of fifth vein one-half the length of preceding section 5.Last section of fifth vein distinctly less than one-half as long aspreceding section 8.5. Acrostichals only two pairs well developed before suture 6.Three pairs of strong presutural acrostichal bristles; palpibrownish ; abdomen almost wholly covei-ed with gray pollen
;
fourth segment bearing a row of strong discal bristles (NewMexico) (8) neomexicana, new species.6. Second antennal segment distinctly longer than first 7.Ba.s.il segments of antennae subequal in length; pollen onthorax and abdomen tinged with brown; postsutural acros-tichals one pair; calypters tawny (Mexico)^- (9) derelicta, new species.7. Last three abdominal segments with defined silvery bandson basal fourth, the remainder of these segments includingthe first shining black; mesonotum thinly pollinose, sub-shining; palpi pale yellow; two sternopleural bristles (Utah).(11) retracta, new species.
20 PKOCEEDINGS OF THE NATIONAL MUSEUM vol.83Pollen on last three abdominal segments not in defined crossbands, the first pollinose on the sides; mesonotum coveredwith dense cinereous pollen, the black vittae very distinctbehind suture; palpi brownish black; three sternopleurals(Utah) (10) cinerescens, new species.8. Front narrowed behind the middle; orbital bristles absent 9.Front almost uniform in width to vertex ; one pair of orbitalspresent; arista thickened on proximal tiiree-fourths ; palpiand second antennal segment yellow ; verticals two pairsdeveloped; fourth abdominal segment without discals (Peru).(24) armata (Townsend).9. Sides of face bare below level of arista outside main row ofbristles 11.Sides of face with hairs extending on lower half outside ofparafacial row 10.10. Bristles on lower half of face distinctly longer than thoseabove and approximating the frontals in size; arista bare,thickened on proximal two-thirds; face strongly receding:palpi dark brown; small cross vein infuscated; epauletsblack (Mexico, New Mexico) (12) crassiseta, new species.Parafacial bristles about equal in length throughout entirerow, and less than half the size of frontals ; arista pubes-cent, tliickene<l hardly to middle; palpi pale yellow; abdo-men wholly gray pollinose; wings hyaline; epaulets red(New Mexico) (13) aristalis, new species.11. Fourth abdominal segment with discal bristles above 12.Fourth abdominal segment without discals above ; first segmentbearing a pair of median marginals; front about one-half eyewidth; arista distinctly pubescent (Texas, North Carolina,Guatemala) (IS) apicalis, new species.12. First abdominal segment with a pair of median marginalbristles; arista pubescent; front narrow, 0.22 of head width;third antennal segment four times length of second(Mexico) (20) validinervis (Van der Wulp).First abdominal segment without median marginals ; aristabare; front rather wide, about 0.29 of head width; thirdantennal segment six to seven times longer than second(United States, widespread) (19) affinis, new speciea
1. Fore tarsal segments compressed and swollen, claws andpulvilli minute or atrophied 4.Fore tarsal segments ordinary, the claws and pulvilli welldeveloped 2.2. Last section of fifth vein one-half as long as preceding section 3.Last section of fifth vein about one-third length of precedingsection ; third antennal segment as wide as parafacialand three times length of second; abdomen thinly graypollinose, subshining; costal spine longer than small crossvein; wings hyaline (California, Arizona) (6) setigera (Coquillett).3. Abdomen shining black, last three segments with pollen indefined bands on basal fifth ; palpi pale yellow ; sterno-pleurals, 2; no infrasquamal hairs; small species, length4 mm (Idaho) (7) obliqua, new species.
OEEATOMYIELLA AND PARADIDYMA?EEINHAKD 21Abdomen wholly gray poUinose ; palpi dark brown ; sterno-pleurals, 3 ; infrasquamal hairs present ; larger species,length 8 mm (8) neomexicana, new species.4. Sides of face bare below level of arista outside main row ofbristles 5.Sides of face outside main row of bristles with coarse blackhairs extending almost to cheeks ; pollen on head golden
;
abdomen wholly pollinose (Mexico).(14) aperta Brauer and Bergenstamm.5. Fourth abdominal segment with a row of discals extendingacross the top 9.Fourth abdominal segment without median discal bristles above 6.6. Outer verticals not developed 7.Outer vertical bristles almost as large as inner ones ; abdomenwholly gray pollinose; arista thickened beyond middle(Peru) (24) armata (Townsend).7. Venter of abdomen clothed with only short black hairs 8.Venter bearing long pale or whitish hairs; palpi black;wings strongly infuscated ; antennae wholly black (Peru).(17) piliventris, new species.8. Arista thickened on proximal two-thirds, bare ; third antennalsegment yellow on basal half, five to six times longer thansecond ; parafacial at narrowest part but slightly widerthan third antennal segment ; pollen on intermediate abdomi-nal segments extending thinly to hind margin (Peru).(16) peruana (Townsend).Arista thickened at base, very slender on apical two-thirds,clothed with short hairs to tip; third antennal segmentblack, narrowed toward base, about two and one-half timeslength of second ; intei-mediate abdominal segments withdefined silvery bands on basal third (Central America,Vera Oruz) (15) aldrichi. new species.9. Apical cell of wing closed and petiolate; outer vertical bristlesabsent 11.Apical cell open ; outer verticals nearly half as long as innerpair 10.10. Epaulets reddish yellow; arista thickened on proximal fourth,distinctly short haired to tip ; mesonotum covered with thicklusterless yellowish gray pollen, the dark stripes very in-conspicuous and visible only in a flat rear view; abdomenlargely pollinose, third and fourth segments at most sub-shining on narrow hind margins (18) apicalis, new species.Epaulets black ; arista thickened almost to middle, pubescent
;
mesonotum gray pollinose, the black stripes distinct, not in-terrupted at suture; last three abdominal segments shiningblack on apical half or more (19) affinis, new species.11. Mesonotum subshining, lightly dusted with pollen, the darkstripes hardly apparent ; arista thickened on basal fifth tofourth ; abdomen black and shiny, basal fourth of segments2 and 3 thinly gray pollinose at most (23) petiolata, new species.Mesonotum densely gray pollinose and vittate, the black siripesusually fused into a single broad pair, which extends tobase of scutelluni ; arista tlilokened about to middle; lastthree abdominal segments conspicuously pollinose on basalthird to half (22) singularis (Townsend).
22 PROCEEDINGS OF THE NATIONAL MUSEUM vol.83(6) PARADIDYMA SETIGERA (CoquUlett)Admontia setiyera Coquiixett, Invertebrata Pacifica, vol. 1, p. 36, 1904.Phytoadmontia setigera Townsend, Proc. U. S, Nat. Mus., vol. 49, p. 626, 1916.Female.?Closely resembles P. obliqua, from which it differs inthe following characters : Front at extreme vertex 0.294 of the headwidth in one specimen; median stripe distinctly narrower than oneparafrontal on entire length; outer verticals weakly developed;third antennal segment three times longer than second; cheek one-half the eye height. Thorax and scutellum densely gray poUinose;mesonotum marked with four black stripes, which are distinct be-hind suture; three sternopleural bristles. Abdomen subshining, thepollen gray and without definite pattern, in certain lights extendingthinly to hind margin of last three segments, the first conspicuouslypollinose above. Fore claws and pulvilli almost normal in size.Hind cross vein of wing not unusually oblique ; last section of fifthvein one-third as long as preceding section; costal spine exceedingthe length of small cross vein.Length.?5.5 mm.Male.?Unknown.Type.?Female, in the United States National Museum, fromCalifornia.Remarks.?Redescribed from one female. East Verde River. Ariz.,4,500 feet; without collector's label.Besides the type and the present specimen, kindly loaned me forstudy by Dr. J. M. Aldrich, no additional specimens have apparentlybeen taken since the species was described 30 years ago.
(7) PARADIDYMA OBLIQUA, new speciesFemale.?Front at vertex 0.304 of the head width in one specimenmeasured; median stripe reddish brown, about equal the width ofone parafrontal;
?cellars small, proclinate; verticals broken off, butthe scars indicating a good-sized inner pair; orbital bristles two,proclinate ; frontals about five in number, extending about to middleof second antennal segment, uppermost bristle reclinate, rather short;parafrontals gray pollinose to vertex, almost devoid of hairs outsideof frontal rows ; face receding, moderately excavated, in profile con-cave above mouth, its ridges normally divergent, bearing only oneor two bristly hairs next to vibrissae; the latter situated on oralmargin but well above the lovrer edge of head; parafacials gray pol-linose, bare outside of the main row of bristles, which extend alonginner margin from lowest frontals to level with apex of third an-tennal segment; cheek sparsely haired on lower margin, gray pol-linose, about two-fifths the eye height; antennae a little shorter than
CERATOMYIELLA AND PARADIDYMA EEINHARD 23face, third segment black, about as wide as parafacial below andonly slightly more than twice the length of second which is yellow;arista blackish, thickened on about basal fourth, penultimate segmentas wide as long; palpi pale yellow, slender; proboscis short, labellafleshy ; eyes sparsely short haired ; back of head gray poUinose, mod-erately clothed with pale hairs.Thorax black, thinly gray pollinose; mesonotum with four blackstripes obliterated behind suture where the surface is subshining inmost views; scutellum black, almost shining but lightly dusted withuniform grayish pollen; chaetotaxy as in validine7'vis, but with onlytwo sternopleural bristles; postscutellum normal; no infrasquamalhairs; calypters white.Abdomen black, rather broad and flat above; last three segmentswith silvery bands on basal fifth, the remainder of these segmentsincluding the first polished or shining; first segment without medianmarginals; second with one pair, small; third also with a medianpair, a wide space intervening between these and the next ones sit-uated near the sides of the segment; fourth bearing a row of ratherstrong discals besides a row of somewhat weaker marginal bristles;hairs on intermediate segments depressed.Legs black, trochanters yellow, coxae less distinctly so ; front tarsalsegments not laterally compressed, the claws and pulvilli small butdistinct.Wings grayish hyaline; third vein bristly two-thirds the distanceto small cross vein; hind cross vein unusually oblique to fourth, join-ing it nearer bend than small cross vein ; apical section of fifth veinone-half the length of preceding section ; fourth vein with a roundedobtuse stumpless bend, curving outward shortly beyond, thencealmost straight to costa; apical cell very narrowly open shortlybefore extreme tip of wing; costal spine small.Length.?4 mm.Male.?Unknown.Type.?Female, U.S.N.M. no. 44760.Remarks.?Described from one female specimen in the UnitedStates National Museum, collected by W. Carter, labeled " S. pestifer,Salmon River Crossing, Idaho, August 31, 1927."This species, like P. setigera, has almost normally developed foretarsal claws and pulvilli, but differs in the more oblique hind crossvein and the extremely narrow pollen bands on last three abdominalsegments. There are other minor differences.
(8) PARADIDYMA NEOMEXICANA, new speciesMale.?Front before triangle 0.258 of the head width in the onespecimen ; parafrontals gray pollinose to vertex, clothed with moder-
24 PKOCEEDINGS OF THE NATIONAL. MUSEUM vol.83
ately long black hairs, -which extend downward below base ofantennae; frontals about six in number, descending almost to levelwith apex of second antennal segment, directed inward, the upper-most pair reclinate, not very long; ocellars strongly divergent, pro-clinate; orbitals absent; inner verticals strong, outer ones not devel-oped; face gray pollinose, its ridges normally divergent downward,bare except a few bristly hairs above vibrissae, which are on a levelwith the protruding front edge of mouth ; sides of face below baseof third antennal segment without any hairs between the row ofbristles and margin of eye; cheeks gray pollinose, sparsely hairedbelow, about two-fifths the eye height ; antennae black, third segmentreddish near base, about four times the length of second ; arista black,thickened to middle; palpi brown, bearing several long black hairsnear tip ; eyes distinctly hairy ; back of head clothed with pale hairs.Thorax and scutellum black, covered with dense gray pollen;mesonotum with four very distinct black stripes, outer pair inter-rupted at suture and stopping before base of scutellum; chaetotaxyas in Valldinervis ^ except that there are three pairs of well-developedacrostichal bristles before the suture; postscutellum normal, graypollinose; one hair present on each side of postnotum beneath thecalypters ; the latter semitransparent, white.Abdomen black, covered with gray changeable pollen, which inmost views extends to the hind margins of last three segments ; basalsegment without median marginals; second with one pair; thirdbearing a marginal row, the intermediate bristles poorly developed;fourth with a discal and a marginal row of rather stout bristles;genital segments blackish, subshining.Legs black ; claws and pulvilli elongated ; the latter dark or grayishin color.Wings subhyaline; hind cross vein sinuous, oblique to fourth,joining it nearer bend than small cross vein; apical section of fifthvein one-half the length of preceding section; apical cell narrowlyopen well before tip of wing; third vein setulose as usual; costalspine developed ; epaulets black.Female.?Front at extreme vertex 0.291 of the head width (onespecimen); two pairs of strong proclinate orbitals; outer verticalsabout half as large as inner pair; third antennal segment moderatelywide to tip, about three and one-half times as long as second; foretarsal segments normal, the claws and pulvilli distinctly developed.Length.?Male, 7.5 mm ; female, 8 mm.Type.?MoXe, U.S.N.M. no. 44761.ReTnarks.?Described from one male and one female. Las Vegas,N.Mex., July (Cockerell).
CEEATOMYIELLA AND PARADIDYMA REINHARD 25From the other forms having the hind cross vein retracted, thepresent species may be separated by the following characters : Malewith three pairs of well-developed presutural acrostichals; femalewith distinct fore tarsal claws and piilvilli; and the abdomen moreextensively pollinose in both sexes.
(9) PARADIDYMA DERELICTA, new speciesMale.?Front narrowed before vertex, at narrowest part 0.238 ofthe head width in one specimen; parafrontals densely gray pollinoseto vertex, sparsely haired outside of frontal rows; median stripereddish brov/n, at antennae about as wide as one parafrontal, verti-cals one pair (inner) developed; frentals ordinary in size, uppermostbristle reclinate and hardly stouter than the preceding one ; sides offace densely gray pollinose, bare outside the main row of bristles,which become longer and stouter downward; antennae black, thirdsegment five times the length of second; arista thickened hardly tomiddle; palpi dark brown; cheek nearly one-half the eye height^thickly clothed with hairs on lower margin.Thorax black, covered with gray pollen, which on the mesonotumhas a brownish sheen in certain lights; dorsal vittae four, outerpair interrupted at the suture; scutellum black, grajnsh pollinose;calypters tawny.Abdomen black ; pollen on intermediate segments with a brownishtinge apparent in most views; narrow hind margin of third andapical half of fourth segment subshining; first segment withoutmedian marginals; second with one pair; third bearing a marginalrow of about eight ; fourth with a row of discals, which are obviouslystouter than the marginals ; genital segments black.Legs black; pulvilli tawny.Wings grayish hyaline; hind cross vein noticeably retracted andsinuous; last section of fifth vein one-half as long as the precedingsection; third vein setulose almost to small cross vein; apical cellnarrowly open, reaching costa well before extreme wing tip; costalspine developed; epaulets black.Length.?7 mm.Female
.
?Unknowm
.
Ty^^.?Male, U.S.N.M. no. 44762.Remarks.?Described from one male in the United States NationalMuseum taken at Mound Valley, Chihuahua, Mexico, August 23,1909, by Dr. C. H. T. Townsend.Differs from P. validinervis, which it closely resembles, in the ab-sence of median marginals on first abdominal segment, and in havingthe hind cross vein of wing noticeably retracted. There are otherminor differences.73007?34 3
26 PROCEEDINGS OF THE NATIONAL MUSEUM vol.83(10) PARADIDYMA CINERESCENS, new speciesMale.?Very closely resembles P. derelicfa., from which it diflfersin the following characters: Front at narrowest part (before vertex)0.263 of the head width in one specimen measiired ; second antennalsegment distinctly longer than first and nearly one-fourth the lengthof third; cheeks clothed with sparse black hairs on lower half;palpi brownish black. Thorax and scutellum covered Avith dullcinereous pollen ; postscutellum membranous above ; calypters white.Abdomen subshining, lightly sprinkled with gray pollen, which inmost Adews extends to the hind margins of last three segments.Pulvilli grayish, about as long as last tarsal segment.Length.?6 mm.FeTnale.?Unknown
,
Type.?Mvile, U.S.N.M. no. 44763.Remarks.?One male. Promontory Point, Utah, August 5, 1929(G. F. I^owlton).The species has a general pale-gray appearance, in contrast withthe decidedly blacker aspect of P. derelicta., to which it is closelyallied. The slight structural differences are mentioned in the key.(11) PARADIDYMA RETRACTA, new speciesHind cross vein noticeably retracted; last section of fifth veinmore than half the length of preceding; apical cell open; sterno-pleurals two; last three abdominal segments shining black on apicalthree-fourths or more.Male.?Front at vertex 0.228 of the head width in one specimenmeasured, hardly widened to middle, and not very prominent atantennae ; cheeks, face, and sides of front gray pollinose ; medianstripe brownish, as wide as one parafrontal on entire length; ocellarsproclinate ; inner verticals developed ; frontals extending about toapex of second antennal segment, uppermost pair rather weak, recli-nate ; antennae nearly as long as face, black, second segment reddish,one-fourth the length of third; arista blackish, thickened on proxi-mal two-fifths; facial ridges strongly diverging downward, bareexcept a few hairs next to vibrissae, which are situated on oral mar-gin; parafacials bare outside of the main row of bristles; cheekssparsely black haired below, about two-fifths the eye height; palpipale yellow; eyes hairy; beard white.Thorax black, gray pollinose; mesonotum marked with four dis-tinct black vittae; scutellum black, subshining, lightly dusted withchangeable gray pollen; infrasquamal hairs absent; calypters semi-transparent, Avhit-e; postscutellum normally developed.Abdomen mostly shining black, with silvery basal bands on lastthree segments, which in a favorable angle extend at most over the
CEEATOMYIELLA AND PARADIDYMA REINHARD 27basal third of intermediate segments; first segment bearing a veryslender or hairlike pair of median marginals; second with a well-developed pair; third bearing a marginal row of about six with awide space between tlie median pair and the lateral one ; fourth witha row of strong discals besides the usual marginal row ; genital seg-ments shining blade, retracted ; fifth sternite black, the lobes promi-nent, narrowly and deeply incised.Legs slender, black; claws and pulvilli nearly as long as the apicaltarsal segment.Wings subhyaline; hind cross vein very oblique to fourth, whichit joins almost midway between bend and small cross vein; apical cellnarrowly open well before the tip of wing ; third vein setulose three-fourths the distance to small cross vein; costal spine small; epauletsblack.Length-?6 mm.Female.?(Unknown.Ty^^.?Male, U.S.N.M. no. 44764.Remarks.?Described from one male specimen taken at Smithfield,Utah, August 24, 1925, by G. F. Knowlton.The narrower front, retracted hind cross vein, two sternopleuralbristles, and the more defined and nan-ower pollen bands on lastthree abdominal segments distinguish the species from afflnis.,described herein. The two species agree in most other essentialcharacters. (12) PARADroYMA CRASSISETA, new speciesMale.?Front at vertex 0.23 and 0.21 of the head width in twospecimens, very prominent at antennae; face strongly receding, itsridges rather flattened below, bearing bristly hairs and one or twostrong macrochaetae next to vibrissae; parafrontals black, coveredwith reflecting grayish pollen, narrow on upper part behind middle
;
median stripe red, wider than one parafrontal except at antennae;frontals six to eight in number, directed inward, the uppermostnearly erect, not longer than preceding pair; orbitals absent; innerverticals strong, outer ones not developed; ?cellars ordinary in size,proclinate ; antennae black, about as long as face ; first segment ex-tending considerably above the front and as long as the second ; thirdsegment unusually broad to tip and about equal the width of para-facial at narrowest part, six to seven times as long as the secondsegment; arista black, thickened on proximal two-thirds, secondsegment short ; parafacials black, the pollen dark gray with a distinctluster, clothed with black hairs outside the row of bristles, whichbecome successively longer and stronger downward; palpi dark
-,,^_^ brown; cheek one-half the eye height; eyes distinctly hairy;beard white.
28 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 83Thorax and scutellum black, o-ray pollinose ; mesonotum markedwith four bUick stripes in front and five behind suture, outer pairstopping before base of scuteHum ; infrasquamal hairs present^calypters ^Yhite.Abdomen black, all segments with gray pollen, which in someviews extends thinly past the middle of the last three ; basal segmentseach with a pair of median marginals (smaller on the first) ; thirdwith a marginal row of 10 to 12 ; fourth bearing a complete row ofdiscals as large as those in the marginal roAV ; genital segments black,I'etracted.Legs black, reddish near base; claws and pulvilli longer thanapical tarsal segment; hind tibia with a row of uneven wide-spacedbristles on outer posterior edge.Wings with a brownish tinge on the anterior margin, small crossvein infuscated ; venation normal ; third vein setulose two-thirds thedistance to small cross vein ; apical cell open well before exact wingtip ; epaulets blackish ; costal spine ordinary in size.Length.?9 mm.Female.?Unknown.Type.?Male, U.S.N.M. no. 44765.Remarks.?Described from two male specimens in the UnitedStates National Museum: One (the type) labeled Sanchez, Chi-huahua, Mexico, September 2, 1909 (C. H. T. Townsend) ; the other,Las Vegas, New Mexico, August 19, 1901 (H. S. Barber).The prominent front, more strongly receding face, longer andbroader third antennal segment, and haired parafacials readily sep-arate the species from P. derelicta.^ to which it is closel}^ related.Another difference is the presence of median marginals on the firstabdominal segment.(13) PARADIDYMA ARISTALIS, new speciesArista densely pubescent; parafacial bristles uniform in lengthdownward, smaller than usual; sides of face cinereous, with blackhairs extending below middle outside of the main row of bristles;mesonotum with four black stripes in front and five behind suture.Male.?Front at vertex 0.26 of the head width (one specimen),rather prominent at antennae; parafrentals cinereous pollinose tovertex, rather sparsely clothed with short black hairs and consider-ably widened before middle; median stripe reddish brown, slightlynarrowed behind ; inner verticals large ; ocellars proclinate ; orbitalsabsent; frontals about six in the row, the lowermost nearly on levelwith apex of second antennal segment; face receding, moderatelydeep, and concave above the mouth; vibrissae large, on level withoral margin ; facial ridges divergent downward, bearing a few bristly
CEEATOMYIELLA AND PAEADIDYMA EEINHARD 29hairs on lower extremity; cheek bare above the lower margin, graypollinose, about two-fifths the eye height; antennae as long as face,first segment rather prominently elevated above the front, third seg-ment about five times longer than second; arista black, thickenednearly to middle, proximal segments short but distinct; palpi slen-der, pale yellow ; beard white ; eyes distinctly hairy.Thorax and scutellum black, densely gray pollinose; three pairsof acrostichal bristles behind the suture, the median ones very weakor hairlike; other details of chaetotaxy as in validinervis ; infra-squamal hairs present; postscutellum gray pollinose, membranousabove; calypters semitransparent, white.Abdomen black, wholly gray pollinose; first segment withoutmedian marginals; second with a stout closely spaced pair; thirdwith a marginal row of about 10; fourth bearing a complete rowof good-sized discals, besides a row of still larger marginals ; genitalsegments reddish black, lightly pollinose, fifth sternite deeplydivided, the lobes blackish, clothed with fine hairs.Legs mostly black, the basal segments and the hind tibiae obvi-ously reddish (front pair missing) ; claws and pulvilli elongate.Wings subhyaline; venation normal; apical cell open well beforetip of wing; costal spine large; epaulets red.Length.?8.5 mm.Female.?Unknown.Type.?^inlQ, U.S.N.M. no. 44766.Remarks.?Described from one male labeled "Animas Park, N.Mex.,6,500 feet (Townsend)."(14) PARADIDYMA APERTA Brauer and BergenstammParadidyma aperta Brauer and BnaiGENSTAMM, Die Zweifliigler des kaiser-lichen Museums zu Wien, no. 6, p. 187, 1893.Microchira mexicana Br.vx;er and Bergenstamm, Ibid., p. 188.Readily distinguished from all other known species by the goldenpollen on the head. I have not seen any male specimens.Female.?Front at extreme vertex 0.312 of the head width (onespecimen) ; pollen deep golden-yellow on parafrontals, parafacials,cheeks, and posterior orbits; face yellowish gray; outer verticalsabout half the size of inner ones; orbitals, two pairs; ocellars pro-clinate; frontals extending to apex of second antennal segment,directed inward, except the uppermost, which is reclinate ; parafacialon narrowest part about one-half the width of facial depression,bearing a row of rather weak bristles and with black hairs on entirelength outside the main row ; vibrissae situated on front edge ofmouth; facial ridges with one or two bristles and fine hairs nextto vibrissae; cheek fully one-half the eye height; palpi yellow;
30 PKOCEEDINGS OF THE NATIONAL MUSEUM vol.83antennae shorter than face, mostly black, third segment ratherslender and about two and one-half times the length of second;arista black, thickened on basal fourth, clothed with short hairs,penultimate segment short; eyes distinctly hairy; back of head graypollinose, moderately pale haired below.Thorax and scutellum black, gray pollinose ; mesonotum with fourdistinct black stripes before the suture and five behind, the outerpair not interrupted at middle and stopping well in front of scutel-lum; acrostichals, 3, 3 (first two behind suture small) ; postscutellumnormally developed, gray pollinose; infrasquamal hairs present;calypters white with yellow margins.Abdomen broadly ovate, black, wholly covered with thick yellow-ish-gray pollen; first segment without median marginals; secondwith a closely spaced pair; third bearing a marginal row of abouteight; fourth with a discal row well behind the middle besides amarginal row of weaker bristles.Legs black, trochanters red; front legs with the tarsal segmentslaterally compressed and swollen, claws and pulvilli minute; midtibia with one strong bristle on outer front side near middle; hindtibia with a scattering row of uneven bristles on outer posterioredge.Wings grayish hyaline ; venation normal ; third vein bristly two-thirds the distance to small cross vein ; apical cell open far before tipof wing; costal spine long; epaulets red.Length.?10 mm.Remarks.?Redescribed from one female specimen in the UnitedStates National Museum, Atzcopotzalco, D. F., Mexico, August 31,1922 (E. G. Smyth).(15) PARADmYMA ALDRICHI, new speciesFemale.?Front at extreme vertex 0.25 to 0.27 of the head widthin three specimens measured; parafrontals black, thinly gray pol-linose; median stripe reddish brown, about equal the width of oneparafrontal on entire length; the usual two proclinate orbitalspresent;
?cellars rather weak, proclinate; inner verticals large,outer ones not developed ; frontals about five in the row, descendingto middle of second antennal segment, uppermost one stouter andreclinate; antennae somewhat shorter than face, third segmentblack, rather narrow at base and about two and one-half timeslonger than the second, which is largely yellow; arista brown,thickened at base and very slender on apical two-thirds, clothed withshort hairs to tip, second segment short; face moderately excavated,hardly receding but concave above mouth in profile, its ridges bear-ing a few bristly hairs next to vibrissae, which are situated on oral
CEKATOMYIELLA AND PARADIDYMA REINHAED 31margin; parafacials blackish, covered with feebly shining graypollen, bare below arista, except a row of weak bristles along innermargin which are reduced in size to small hairs on the upper part;palpi yellow, slender to tip; cheek one-third to two-fifths the eyeheight; eyes sparsely short haired; back of head gray pollinose,moderately clothed with whitish hairs.Thorax black, gray pollinose; mesonotum with four black stripes,which are sometimes indistinctly separated behind the suture; scu-tellum black, lightly sprinkled with changeable gray pollen, post-scuteilum normal, thinly pollinose ; infrasquamai hairs absent in twospecimens and three hairs present in the other; calypters paleyellowish white.Abdomen shining black ; intermediate segments with silvery bandson basal third, the fourth thinly pollinose almost to apex; first seg-ment without median marginal bristles ; second with one pair ; thirdand fourth each bearing a marginal row ; fourth without discals, thebroad basal margin above destitute of hairs.Legs black ; front tarsi compressed and swollen, the claws and pul-villi very minute.Wings brown, paler on the posterior margin; apical cell openshortly before wing tip; venation normal; third vein setulose halfto three-fourths the distance to small cross vein ; costal spine longerthan small cross vein; epaulets black.Length.?7 mm.Male.?Unknown.Type.?Female, U.S.N.M. no. 44767.Remarks.?Described from three female specimens in the UnitedStates National Museum as follows: 1 (the type), taken at IngenioR.R. Station, Guatemala, April 28, 1926, by Dr. J. M. Aldrich, inwhose honor the species is named ; 1, labeled San Rafael, Vera Cruz(C. H. T. Townsend), and the other, Higuito, San Mateo, CostaRica (Pablo Schild).The species has the eyes less distinctly haired and smaller para-facial bristles than any other member of the genus. It is provision-ally included here. The accumulation of better preserved speci-mens, including the male sex, seems necessary to decide the questionof proper generic allocation. The relationship with Ceratomyiellaseems close, but the type species of that genus has the eyes entirelybare. (16) PARADroYMA PERUANA (Townsend)Diaphoropeza peruana Townsend, Proc. U. S. Nat. Mus., vol. 43, p. 308, 1912.Very similar to P. singularis, from which it differs in the followingcharacters
:
32 PROCEEDINGS OF THE NATIONAL MUSEUM vol.83Female.?Front rather narrow, at extreme vertex 0,26 of the headwidth (one specimen) ; parafacial at narrowest about as wide asthird antennal segment; cheek barely one-third the eye height.Infrasquamal hairs present; calypters with rims pale tawny. Abdo-men without defined silvery basal bands on last three segments ; firstthinly gray pollinose above and on sides ; intermediate segments withthe pollen extending thinly beyond the middle when viewed frombehind; anal segment almost entirely gray pollinose, without discalbristles; venter largely covered with gray pollen. Legs brownishblack. Wings with a distinctly yellow tinge on costal margin andalong the veins, hind margins grayish hyaline; apical cell closed atcosta, not petiolate; costal spine about as long as small cross vein;epaulets reddish black.Length.?7.5 mm.Male.?Unknown.ry;?e.?Female, U.S.N.M. no. 15147.Remarks.?Redescribed from one paratype female from Sullana,Peru, October 1, 1910, TD 3942 (C. H. T. Townsend) . In the UnitedStates National Museum there are three additional female typespecimens from the same locality and one female from Piura, Peru,all collected by Dr. C. H. T. Townsend.The narrower cheeks and parafacials and the absence of discalbristles on the fourth abdominal segment readily separate the speciesfrom P. singularis.(17) PARADIDYMA PILIVENTRIS, new speciesDistinguished from all others of this group by the presence of palehairs on venter of abdomen.Female.?Front at vertex 0.29 and 0.27 of the head width in thetwo specimens ; parafrontals black, thinly gray pollinose, with morenumerous black hairs on the lower part extending on the parafacialsabout to level with arista; outer verticals not developed; ocellarsproclinate ; orbitals two proclinate pairs ; frontals about six in num-ber, distinctly larger than parafacial bristles, uppermost pair ratherstout and reclinate, the others directed inward descending below mid-dle of second antennal segment ; face blackish, gray pollinose, mod-erately receding and concave above mouth in profile ; facial ridges notvery prominent, bearing bristly hairs on about the lowest fourth;parafacials largely black, covered with satiny gray pollen, a row ofbristles along the reddish inner margin, outside of these bare onlower half; antennae shorter than face, wholly black, third segmentnarrowed toward base and about two and one-half times the lengthof second ; arista black, thickened on proximal fourth, basal segmentsshort ; vibrissae near the front edge of the mouth ; cheeks two-fifthsthe eye height; eyes distinctly hairy; palpi black, bearing several
CERATOMYIELLA AND PAKADIDYMA EEINHARD 33long pale hairs on under side near the tip; back of head blackish,thinly gray pollinose, rather sparsely clothed with pale hairs.Thorax and scutelliim black, dusted with gray pollen; mesonotummarked with four black vittae, outer pair not interrupted at suture
;
chaetotaxy as in validinerms; infrasquamal hairs absent; postscu-tellum normal; calypters pale yellowish white.Abdomen black and shiny ; last three segments with dense grayish-white pollen bands, which are wider on the sides of the intermediatesegments and narrowed at the middle above, especially on the secondwhere the pollen is confined on the basal margin; first segmentwithout median marginals; second with one pair; third and fourtheach bearing a marginal row of about 10; fourth segment withoutdiscal bristles; venter pale haired.Legs black, basal segments reddish ; front tarsal segments swollen,the claws and pulvilli minute or atrophied; hind tibia with threelarge and several small bristles on outer posterior edge.Wings infuscated, a little paler along the hind border; venationnormal ; third vein with hairs extending almost to small cross veinapical cell open shortly before wing tip; costal spine strong;epaulets black.Length.?7 mm.Male.?Unknown
,
T'y;?^.?Female, U.S.N.M. no. 44768.Remarks.?Described from two females in the United StatesNational Museum, collected at Huariaca, Peru, December, 1921, byDr. C. H. T. Townsend.(18) PARADIDYMA APICALIS, new apeciesMale.?Front at vertex 0.258 of the head width (average of five:0.24; 0.26; 0.26; 0.27; 0.26), hardly wider to middle, thencediverging to antennae where it is rather prominent in profile;median stripe reddish brown, nearly as wide as one parafrontal onmost of its length; ocellars present; verticals one pair (inner)developed ; orbitals absent ; frontals five or six in number, the rowsmoderately divergent beneath antennae extending to apex of secondsegment, uppermost bristle stouter, reclinate; parafrontals coveredwith dull gray pollen to vertex, moderately clothed with rathercoarse black hairs; face receding and concave above the mouth inprofile, its ridges not very prominent, haired on lower fourth orless; parafacials densely gray pollinose, a row of bristles along theinner margin becoming longer and stronger downward, the largestapproximating the frontals in size, bare outside the main row ofbristles below level of arista; vibrissae situated on oral margin;antennae about as long as face, basal segments subequal in length.
34 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 83third black, five or six times longer than second which is largelyyellow ; arista blackish, finely pubescent, thickened on proximal two-fifths, basal segments short; palpi slender, pale yellow, bearing afew long black hairs beneath on apical half ; cheek bare above lowermargin, gray pollinose, clothed with pale hairs; eyes distinctlyhairy.Thorax and scutelliim black, covered with thick gray pollen;mesonotum with four narrow dark stripes, often poorly defined,interrupted at suture and stopping far before base of scutellum;two or three acrostichal bristles usually developed behind the su-ture, other details of chaetotaxy as in validwervis] postscutellumthinly pollinose and membranous above; infrasquamal hairs pres-ent; calypters semitransparent, white with a tawny tinge.Abdomen black, largely gray pollinose; hind margins of lastthree segments black and subshining when viewed from above, butpollinose or subpollinose in most other views, first segment withconspicuous pollen on the sides above ; one pair of median marginalbristles present on first and second segments, the third and fourtheach with a marginal row, no discals on dorsal surface of fourthsegment; genital segments small, black, retracted; inner forcepsunited, slender, clothed with brownish hairs behind near base,flattened and shining beyond middle, apex minutely notched ; outerforceps shorter than inner ones, tapering uniformly to a blunt tip,tinged with yellow^; penis simple, short, blackish, apex bearing ashort pale membrane.Legs black, basal segments and knees yellowish ; mid tibia witha whorl of three bristles near middle, the one on outer front sidestout ; hind tibia with a scattering row of irregular bristles on outerposterior edge ; claws and pulvilli elongate.Wings subhyaline; veins yellow, bare except third, which hashairs extending almost to small cross vein; fourth vein with arounded almost rectangular stumpless bend, beyond rather deeplyconcave to costa; last section of fifth vein about ono-fourth as longas preceding section; apical cell open well before the wing tip;costal spine strong; epaulets red.Female.?Front at vertex 0.282 of the head width (average offive: 0.28; 0.28; 0.28; O.ai; 0.2G) ; pollen on para frontals, meso-notum and abdomen with a distinct pale brassy tinge; outer verti-cals developed, orbitals two pairs; antennae shorter than face,third segment slender, j^ellow at base, about four times the lengthof second; arista short haired to the tip; first abdominaj segmentwithout median marginals, the fourth with discals above; foretarsal segments compressed and swollen, claws and pulvilli atrophied.Length.?G to 8 mm.
CERATOMYIELLA AND PARADIDYMA REINHARD 35Type.?Male, U.S.N.M. no. 44769, from College Station, Tex.Remarks.?Described from 14 males and 11 females in my collec-tion. All taken at College Station, Tex., September-November1920-1933 (H. J. Keinhard), and one male from Hidalgo County,Tex., May 18, 1932 (S. W. Clark). In the United States NationalMuseum, one female from La Providencia, Obispo, Guatemala (C.M. Kouillard). (19) PARADIDYMA AFFINIS, new speciesMale.?Front at vertex 0.288 of the head width (average of five:0.29; 0.28; 0.29; 0.29; 0.29), hardly widening to middle thence rap-idly so to antennae where it is moderately prominent; parafrentalsgray pollinose to vertex, moderately clothed with rather coarse blackhairs outside the frontal bristles ; median stripe red, occupying aboutone-third the frontal width; inner verticals large, outer pair notdeveloped; orbitals absent; frontal bristles extending below middleof second antennal segment, uppermost one reclinate ; ocellars pres-ent, proclinate ; antennae about as long as face, basal segments sub-equal in length, third segment black, six or seven times longer thansecond which is largely yellow; arista black, thickened about tomiddle, finely pubescent, second segment short; face gray pollinose,moderately excavated, receding with the lower border slightly prom-inent between vibrissae; facial ridges weakly divergent downward,bearing a few bristly hairs next to vibrissae, which are situated onthe oral margin ; palpi slender, pale yellow, with several long blackhairs on lower edge beyond the middle; parafacial gray pollinose,bare except a row of bristles along the inner margin, those in lowerpart of row approximating the frontals in size; cheek reddish inground color, thinly gray pollinose and bare above, about two-fifthsthe eye height; eyes distinctly haired; back of head densely graypollinose and thickly clothed with pale or whitish hairs.Thorax black, gray pollinose; mesonotum marked with four nar-row black stripes, which are not very conspicuous especiall}'^ behindthe suture ; scutellum black, covered with dull gray pollen, which isthinner on middle of disk; chaetotaxy as in validinervis but withthree postsutural acrostichals usually present ; postscutellum normal,thinly pollinose; sides of postnotum beneath calypters bearing atuft of small black hairs ; calypters semitransparent, white.Abdomen black, dusted with gray pollen, which in certain viewsextends thinly behind the middle on the intermediate segments;viewed from above the first segment pollinose on sides and the threefollowing ones shining black on posterior third to half; first seg-ment without median marginals ; second with one pair ; third bearinga marginal row of 10 to 12 ; fourth with a discal and a marginal
36 PROCEEDINGS OP THE NATIONAL MUSEUM vol.83row of stronger bristles ; genitalia as in P. singularis, but the outerforceps more broadly rounded at the apex.Legs rather slender, black ; claws and pulvilli elongated.Wings grayish hyaline; venation bare except third vein, which issetulose almost to small cross vein; fourth vein with a roundedstumpless bend and broadly concave beyond to costa; last sectionof fifth vein about one-fourth the length of preceding section ; apicalcell narrowly open far before wing tip ; costal spine strong ; epauletsblack.Female.?Front at vertex 0.288 of the head width (five measuredas follows: 0.3; 0.27; 0.29; 0.28; 0.3) ; outer verticals developed butsmaller than inner ones; orbitals two proclinate pairs; antennaeshorter than face, third segment slender, four to five times the lengthof second ; first abdominal segment usually without median marginalbristles (two specimens with a pair present) ; front tarsi laterallycompressed and swollen, the claws and pulvilli minute or atrophied.Length.?5.5 to 9.5 mm.Type.?Mfi\Q, U.S.N.M. no. 44770, from College Station, Tex.Remarks.?Described from 50 specimens. In the United StatesNational Museum : 2 males, Knoxville, Tenn., Ma}^ 25 (J. M. Aldrich) ;1 male, Birmingham, Ala., June 4, 1917 (J. M. Aldrich) ; 1 female,Lafayette, Ind., September 12, 1918 (J. M. Aldrich) ; 1 female,Riley County, Kans., May 29 (Popenoe) ; 1 female, Manhattan,Kans., May 30, 1928 (R. C. Smith) ; 1 female, labeled " Ga." with-out additional data; 1 male and 1 female, Clemson College, S.C.,October 10, 1908 (C. H. T. Townsend) ; 1 female. Holly Springs,Miss., September 7, 1890 (F. AV. Mally) ; 1 male, Miami, Fla.,September 6 (C. H. T. Townsend) ; 1 female, Washington, D.C.,September 18, 1921 (J. M. Aldrich) ; 1 female, Rock Creek, D.C.,flowers chrysanthemum, May 30, 1917 (C. H. T. Townsend); 1male. Eastern Branch near Bennings, D.C., August 29, 1915 (W. L.McAtee) ; 1 male, Washington, D.C., June 27 (Townsend) ; 1 fe-male, Washington, D.C., October 2, 1917 (W. L. McAtee) ; 1 female,Anacostia, D.C., September 24, 1914 (R. C. Shannon) ; 1 female, atlight, Plummers Island, Md., September 2, 1914 (R. C. Shannon)
;
1 female, Chesapeake Beach, Md., October 14, 1926 (J. M. Aldrich)1 female, Arlington, Va., October 6, 1913 (R. H. Hutchison) ; 1male, Lincoln, Nebr., July 7, 1922 (O. C. Bradbury) ; 1 female,Marfa, Tex., June 13, 1917 (J. M. Aldrich) ; 1 female, Raleigh,N.C. (C. S. Brimley). In the Kansas University Museum :1 male,Las Cruces, N.Mex., September 25, no collector's label. In my col-lection: 9 males and 16 females, College Station, Tex., April toOctober, 1917-1930 (H. J. Reinhard). In the collection of D. G.Hall, 1 female, Manhattan, Kans., May 10, 1929 (D. G. Hall).
CERATOMYIELLA AND PARADIDYMA REINHAED 37In the material examined I have noted but one specimen (female)having the first posterior cell closed and short petiolate. In singu-laris the first posterior cell is invariably closed and usually shortpetiolate. Other differences may be noted in the present species:Four narrow thoracic stripes; fourth abdominal segment withdiscals above in male; outer verticals present in female.(20) PARADroYMA VALIDINERVIS (Van der Wulp)Didyma validinervis Van der WrrLP, Biologia Centrali-Americaua, Diptera,vol. 2, p. 164, 1890.Paradidyma validinervis Braxjer and Beirgbnstamm, Die Zweifliigler deskaiserlichen Museums zu Wien, uo. 5, p. 404, 1891; no. 6, p. 127, 1893.Besides the characters mentioned in the generic description, thetype species has the following additional characters:Male.?Front narrowed before vertex, at narrowest part 0.22 ofthe head width in one specimen; parafrontals covered with densegray pollen to vertex, sparsely clothed with hairs outside of frontalrows ; median stripe reddish brown, at middle wider than one para-frontal; inner verticals developed; frontals ordinary in size, theuppermost bristle subreclinate and hardly stouter than the precedingone; parafacials densely gray pollinose, bare below level of aristaoutside of the main bristles, which increase in size downward in therow; antennae black, third segment about four times the length ofsecond; arista pubescent, thickened about on proximal half; palpidark brown ; cheek clothed with black hairs on lower margin, abouttwo-fifths the eye height; eyes hairy.Thorax black, gray pollinose ; mesonotum marked with four blackstripes, outer pair interrupted at suture; scutellum black, thinlygray pollinose; calypters whitish, the hind lobe translucent andwith a slight brownish tinge.Abdomen black; segments two to four shining beyond the basalsilvery band, only the second showing thin pollen more extensivelyin a flat rear view; basal segments each with a pair of median mar-ginals; third bearing a marginal row of six, large; fourth with arow of six discals, which are slightly stouter than the marginals;genital segments black.Legs black; claws and pulvilli elongate.Wings grayish hyaline; first vein bare, third setulose almost tosmall cross vein; apical cell narrowly open; last section of fifthvein less than half as long as preceding section ; costal spine devel-oped.Length.?7 mm.Reinarhs.?Redescribed from one cotype male specimen in theUnited States National Museum from Guerrero, Mexico. I havenot seen any specimens of the female.
38 PROCEEDINGS OF THE NATIONAL MUSEUM vol.83(21) PARADIDYMA BRASILIANA, new speciesMale.?Front moderately wide, at vertex 0.29 of the head widthin one specimen ; parafrentals covered with gray pollen tinged withyellow along the inner margins; parafacials, cheeks, and posteriororbits cinereous; third antennal segment largely black, the baseyellow to the insertion of the arista. Sides of postnotum beneathcalypters with a few small inconspicuous hairs. Abdomen slender,tapering to a rather narrow apex ; first segment conspicuously graypollinose on the sides above, second almost entirely covered withchangeable gray pollen, third and fourth shining black on apicalthird to half; genital segments black, small and retracted. Other-wise as in P. singularis.Length.?7 mm.Female.?Unknown.Type.?Mvile, U.S.N.M. no. 44771.Remarks.?Described from one male specimen collected at Itaqua-quecetuba, Sao Paulo. Brazil, September 26, by Dr. C. H. T.Townsend.I am unable to note any structural differences between the presentspecies and P. singularis; both have the third antennal segmentstrikingly elongated with the anterior margin concave below theinsertion of the arista. The rather slight characters separating thespecies are mentioned above and in the key. A second closely relatedspecies is P. peruana, known only in the female; it differs fromhrasiliana in having the cheeks distinctly narrower, the wings andcalypters obviously tinged with yellow, and apical cell barely closedat costa. (22) PARADmYMA SINGULARIS (Townsend)Atrophopoda singularis Townsend, Trans. Amer. Ent. Soc, vol. 18, p. 373, 1891.Lachnomma magnicarnis Townsend, Trans. Amer. Ent. Soc, vol. 19, p. 103, 1892.Paradidyma singularis Coquillett, Revision of the Tachinidae of America,p. 126, 3897.?.^XDl:ICH, Catalogue of North American Diptern, p. 174, 1905.Male.?Front at vertex 0.306 of the head width (five measured asfollows: 0.31; 0.31; 0.3; 0.3; 0.31), rather prominent at antennae;parafrontals, parafacials, and cheeks cinereous pollinose; face longand deeply excavated, its ridges weakly divergent downward, bear-ing a few bristles and hairs on lower extremity; antennae strikinglyelongate, first segment longer than second and extending consider-ably above level of the front, third segment black, about equal thelength of face with the front edge concave below insertion of arista;the latter thickened on proximal three-fourths; cheek two-fifths theeye height; parafacial bare outside the main row of bristles; palpiyellow; eyes distinctly hairy.
CEEATOMYIELLA AND PAEADIDYMA REINHARD 39Thorax black ; mesonotum densely gray pollinose, the black stripesusually fused into a single broad pair, which extends to base ofscutellum; chaetotaxy as in validinervis ; sides of postnotum beneathcalypters usually bare but sometimes with a few small hairs present;calypters white.Abdomen black; last three segments with silvery bands on basalthird, remainder of these segments including the first polished orshining; basal segment without median marginal bristles; secondwith one pair ; third and fourth each with a marginal row, the fourthoccasionally with one or two discals on the sides but none at middleabove; genital segments blackish, retracted; inner forceps black,with a slight median keel behind, moderately broad at base, theapical half narrowed terminating in an acutely tipped shining beak;outer forceps yellow, slightly shorter than inner ones, the sidesbulged and clothed with short brownish hairs, tips blunt; fifthsternite with a narrow deep incision, the lobes black.Legs black; claws and pulvilli elongate.Wings grayish hyaline; third vein with hairs extending almostto small cross vein ; last section of fifth vein one-third the length ofpreceding section; apical cell closed and usually short petiolate;costal spine well developed.Female.?Front at vertex 0.328 of the head width (average of five:0.35; 0.32; 0.35; 0.31; 0.31) ; the usual two pairs of proclinate orbit-als present; outer verticals not developed; antennae shorter thanface, third segment slender, about one-half as wide as parafacialand four or five times longer than second; cheek two-fifths the eyeheight; fourth abdominal segment with a row of discals behind themiddle above; fore tarsi compressed, the claws and pulvilliatrophied.Length.?5.5 to 8.5 mm.Remarks.?Redescribed from a long series of both sexes from allsections of the United States, including the type male of Lachnommamagnicornis Townsend, in the Kansas University Museum.A common North American species described from Carlinville,111. Readily distinguished from most other members of the genus bythe two broad black thoracic stripes, which in well-preserved speci-mens are sharply contrasted on entire length by a median and lateralpale gray pollen bands. CatemofliTys sequens Townsend presentsabout the same general appearance but can readily be separated byits bare eyes and parafacials.(23) PARADmYMA PETIOLATA, new speciesMesonotum subshining, the vittae poorly defined; arista thickenedon proximal half or less; apical cell closed, the petiole about twicethe length of small cross vein.
40 PROCEEDINGS OP THE NATIONAL MUSEUM vol.83Male.?Front 0.252 of the head width (average of five: 0.26; 0.25;0.25; 0.25; 0.25), hardly widening to middle, thence rapidly so down-ward ; parafrontals thinly gray pollinose often blackish before vertex,clothed with sparse short hairs; median stripe reddish brown, hardlynarrowed behind and extending on each side of triangle to vertex;inner verticals moderately strong, directed posteriorly, outer pairvestigial; ocellar bristles small, proclinate, divergent; orbitals absent;frontal bristles about five in each row, descending to level with apexof second antennal segment, uppermost pair reclinate, the othersdirected inward; face rather long and receding, deeply excavated,ground color black, gray pollinose, its ridges bare except a fewbristly hairs above vibrissae; parafacial covered with shining grayor almost silvery pollen, bearing a row of bristles along the innermargin which become longer and stouter downward; vibrissae sit-uated on level with front edge of mouth; antennae as long as face;third segment black, with the anterior edge practically straight,about seven times the length of second; basal segments tinged withred, the first longer than second and extending well above the levelof the front; arista slightly shorter than third antennal segment,black, thickened on proximal two-fifths, basal segments short butdistinct; cheek about two-fifths the eye height; proboscis short,labella fleshy ; palpi slender, hardly thickened apically, yellow ; eyesdistinctly hairy; beard white.Thorax black, thinly dusted with gray pollen ; mesonotum sub-shining, the vittae poorly defined; chaetotaxy as in validinervis^except that there are usually three pairs of acrostichals before thesuture; scutellum black, subshining; infrasquamal hairs present;postscutellum normally developed; calypters white.Abdomen shining black, with narrow silvery basal bands on inter-mediate segments, the fourth faintly pruinose at most; first segmentwithout median marginals; second with a single pair; third andfourth with marginal rows, the latter also with discals at the sidesbut none on the middle above; genital segments black; inner forcepsunited, flat behind, tapering from base to an acute ti]5 ; outer forcepsyellowish, shorter than inner pair, tapering outward, the tips rathernarrow and darker.Legs black; claws and pulvilli elongate; wings subhyaline; veinsbare except third, which is setulose almost to small cross vein; lastsection of fifth vein at most one-third the length of preceding sec-tion; apical cell closed, Avith petiole longer than small cross vein;costal spine small.Female.?Front at vertex 0.252 of the head width (average offive: 0.25; 0.25; 0.2C; 0.25; 0.25), widening gradually to base ofantennae; parafrontals blackish, subshining; orbitals two pairs, pro-
CEBATOMYIELLA AND PAEADIDYMA REINHAED 41
clinate; outer verticals not developed; third antennal segment slen-der, yellow at base, four to five times as long as second; abdomenmostly polished black, intermediate segments silvery on the narrowbasal margin, fourth with discal bristles above ; fore tarsal segmentscompressed and swollen, the claws and pulvilli minute.Length.?5 to 6.5 mm.ry;>e.?Male, U.S.N.M. no. 44772.Remarks.?Described from 6 males and 11 females. In the UnitedStates National Museum : 2 males and 3 females (including the type),Lafayette, Ind., August and September 1917-1921 (J. M. Aldrich") ;2 males and 4 females, Plummers Island, Md., September 24, 1902(Barber and Schwarz), August 1903 (A. Busck), August 3, 1912(J. K. Malloch), a females labeled "at light", September 7, 1912,without collector's label; 1 female, Chesapeake Beach, Md., Septem-ber 19, 1915 (W. L. McAtee) ; 1 male. Dead Run, Fairfax County,Va., September 30, 1915 (R. C. Shannon) ; 2 females. Difficult Run,Va., September 19, 1916, and October 28, 1917 (W. L. McAtee). Inthe Kansas University Museum : 1 pair labeled " Ills. Forbes."The species is closely related to P. singularis., from which it maybe readily separated by the shining black mesonotum ; longer petioleof the apical cell ; and narrower front in both sexes.(24) PARADmYMA ARMATA (Townsend)Lachnommopsis armata Townsend, Proc. U.S. Nat. Mus., vol. 49, p. 421, 1915.Front uncommonly broad to vertex and orbitals present in the malesex ; parafacial bristles about uniform in length throughout the row
;
abdomen densely gray pollinose.Male.?Front at extreme vertex 0.33 and 0.35 of the head widthin two specimens, not much wider at base of antennae ; parafrontalsgray pollinose to vertex, sparsely haired outside of frontal rows;median stripe red, narrower than one parafrontal on entire length;inner and outer verticals developed; orbitals one pair, proclinate;ocellars present; frontals extending to apex of second antennal seg-ment, the uppermost stronger and reclinate ; face rather strongly re-ceding and concave above the mouth in profile view, not very deep
;
facial ridges divergent below and haired on lower fourth or less;parafacials gray pollinose, a row of bristles along the inner marginthat are noticeably smaller than the lowermost frontals, bare outsidethe main rows below base of third antennal segment; vibrissae onlevel with front edge of mouth ; antennae slightly shorter than face,third segment black, about four times length of second which ismostly yellow and distinctly longer than the first; arista thickenedto apical fourth, basal segments short; cheek fully one-third the eyeheight ; palpi slender, pale yellow ; eyes distinctly hairy.
42 PKOCEEDINGS OF THE NATIONAL MUSEUM vol. 8SThorax and sciitellum black, gray pollinose; mesonotum markedwith four narrow bhick stripes, outer ones interrupted at suture andstopping before base of scutelhnn; chaetotaxy as in valklinervls;postscutelhim gra}- pollinose, pale membranous above; infrasquamalhairs present; calypters tawnj^, paler at middle.Abdomen black, wholly covered with gray pollen; first segmentwithout median marginals; second with one pair; third and fourtheach with a marginal row ; no discals on anal segment ; genital seg-ments reddish black retracted; fifth sternite prominent, narrowlyand deeply incised, the lobes pale yellow.Legs black, trochanters j'ellow, coxae less distinctly so; claws andpulvilli shorter than the apical tarsal segment.Wings subhyaline; venation normal; third vein haired about half-way to small cross vein; apical cell open a little before the exacttip of wing; costal spine developed; epaulets red.Female.?Front at vertex 0.349 of the head width (one specimen)
;
fore tarsal segments compressed, the claws and pulvilli minute oratrophied, otherwise very similar to male.Length.?6 mm.Type.?Male, U.S.N.M. no. 19442.Remarks.?Redescribed from three specimens in the United StatesNational Museum. Two paratypes (male and female), Chosica,Peru, May 25, 1913 (C. H. T. Townsend), and one male, Matucana,Peru, April 22, 1914 (C. H. T. Townsend).There appear to be no characters of generic importance, commonto both sexes, that distinguish the species from Paradidyma. Thesecondary sexual characters in the male, viz, the wide front andpresence of orbital bristles, at once separate it from all other knownforms. The female, however, agrees in the essential characters ofthe present genus. As usual the front tarsi are compressed andswollen, with the claws and pulvilli minute or atrophied.
CEEATOMYIELLA AND PARADIDYMA REINHARD 43UNRECOGNIZED SPECIESThe following species apparently belong to Paradidyma but havenot been identified in the material contained in the United StatesNational Museum. Both species were characterized in abbreviateddescriptive terms, which were kindly transcribed for me by the lateDr. J. M. Aldrich. The types are located in the Experiment StationCollection, Lima, Peru.Atrophopoda peruana TownsendAtrophopoda peruana Townsend, Eev. Chil. Hist. Nat., vol. 31, p. 159, 1927.Body length, 5 mm. ; wing length, 4iy4 mm. 1 male, Cacaturo, Piura Province,Peru, May 22 on herbage.Blackish ; head silvery white, facial plate and facial ridges gray ; parafrontalsblackish by direct view, thinly pollinose ; frontal stripe dark brown ; first an-tennal joint brown; second joint and palpi very pale fulvous; third jointblackish; pleura silvery, mesoscutellum and scutellum less thickly so; twoheavy wide black thoracic vittae unbroken and reaching scutellum ; abdomenshining; median vittae and narrow bases of intermediate segments thinlysilvery, fourth segment more widely on base. Legs black. Wings pale smokyyellowish on costa and veins. Squamae glassy-whitish.Apparently quite similar to Paradidyma {Diaphoropeza) peruana(Townsend), which was also described (female only) from Peru.The present form may be the male of the last mentioned species, butit seems impossible to decide without specimens available forcomparison. Paradidyma PEmxrviANA TownsendParadidyma peruviana Townsend, Rev. Chil. Hist. Nat., vol. 31, p. 159, 1927.Body length, 7 mm. ; wing length, 6 mm. One female, Chosica, Peru, 3,000 ft,Oct. 18, indoors.Differs from P. validincrvis by female vertical width well over one-thirdhead width ; frontals two below base of arista ; width of frontal stripe two-thirds of one parafrontal at middle ; ocellars of same strength as hind proclinatefronto-orbital ; facio-orbitals eight or nine in row along inner edge of para-facials ; cheek two-fifths the eye length ; third vein bristly halfway to anteriorcross vein ; apical cell closed considerably before wing tip ; hind cross veinmuch nearer bend of fourth and hardly its own length from same; palpi yel-lowish or fulvous ; four moderately wide, equal black thoracic vittae not veryheavy ; wing veins yellowish ; " nos " * infuscate.According to the description the species is distinct from all othermembers of the genus by the frontal bristles descending beneath thebase of the arista. This character, the wide front, and the strongocellars should make the species easily recognizable.
* Meaning unknown, probably a misprint.
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