NOTES ON THE RHINOTRAGINE BEETLES OF THEFAMILY CERAMBYCIDAE, WITH DESCRIPTIONS OFNEW SPECIES By W. S. FisherOf the Bureau of Entomology, United States Department of Agriculture
In rearranging the material of this tribe in the United StatesNational Museum collection, it is necessary to describe a number ofnew forms, and it seems also necessary to record some of the noteswhich have accumulated with the specimens in the collection.This tribe contains quite a number of species and genera which areconfined to tropical America. They are remarkable both for theirgenerally abbreviated elytra, although this is not an essential char-acter for distinguishing the tribe, and for the mimetic resemblancethat many of the species bear to Hymenoptera and other Coleoptera.Although the species are not great favorites with the systematist onaccount of the difficulty of separating the genera, they are full of in-terest to the general entomologist, because of their mimicry and themany fine illustrations they offer of the mode in which divergentmodifications occur in nature. For example, it is most interesting toobserve in forms so closely allied that while some species have rudi-mentary elytra and wasplike bodies or beelike posterior tibiae, suchas the species of Odontocera^ Tomopterus^ Epimelitta^ etc., othershave the elytra developed to the opposite extreme, and, aided by theirgeneral aspects and colors, become analogous to other Coleoptera,such as Oxylymma resembling the Galerucinae, the Erythroflatysthe Hispinae, and the Ornistomus various species of Lycidae.If we believe in the origin of species by natural variation and selec-tion, the species of Rhinotragini have varied in many directions, andsince a protective disguise of one kind or another was necessary tothe species, these variations have been gradually extended in manydirections to mimic some species which it was advantageous to re-semble. In some of the very closely allied species the most abruptchanges are seen, as for instance, between Odontocera fasciata Olivier(which resembles a wasp) and Odontocera covi'pressi'pes WhiteNo. 2842.?Proceedings U. S. National Museum, Vol. 77, Art. 192604?30 1 1
2 PROCEEDINGS OP THE 1^ \TIONAL MUSEUM vol. 77(which resembles a bee, with a pollen-collecting brush of hairs oneach posterior tibia). The abruptness with which important struc-tural parts change from species to species renders the definition ofgenera almost impossible in this tribe, as almost every species offerssufficient characters to render generic separation plausible for theextremist.An almost universal character of this tribe is the large eyes, espe-cially the lower lobes, which in the males are nearly contiguous infront, but even this character disappears in a few species. The headis generally elongate below the eyes, but this character exists in someof the other tribes of Cerambycidae. The pronotun is cylindrical orovate and unarmed at the sides. The antennae are almost alwaysmore or less serrate from the sixth joint, and the intermediate jointsare armed with setae on their inner margins. The prosternal processis alwaj^s present between the anterior coxae, although it is verynarrow in a few species, and the episterna of the metasternum arealv\^ays triangular and very broad in front.The adults, whose short elytra and strongly developed membranouswings permit them to fly swiftly, frequent flowers in company withthe Hymenoptera, which many of them resemble.ODONTOCERA FASCIATA (Olivier)Necydalis fasciata Oliviek, Entomol., vol. 4, gen. 74, 1795, no. 9, p. 10, pi. 1,fig. 9.Odontocera vitrea Sesville, Ann. Soc. Ent. France, vol. 2, 1833, pp. 547-548.Odontocera chrysosone White, Cat. Col. British Museum, pt. 8, 1855, p. 192,pi. 5, fig. 5.This species was originally described by Olivier (1795) fromSouth America. Serville (1833) redescribed the same species fromCayenne under the name of vitrea and placed it as the first species inhis new genus Odontocera^ and White (1855) redescribed it fromthe Amazon under the name of chrysozone.This species seems to have a rather wide distribution. Bates ^writes that it is generally distributed throughout the Amazons, andis not uncommon in the dry season at sweet-smelling flowers. Likeall the other species among the beautifully varied and interestinglittle creatures of this tribe, it flies nimbly from flower to flower,deceiving the eye of the beholder by its strong resemblance to awasp. A large series has been examined by the writer and scarcelyany variations were observed except in size. P. J. Darlington, jr.,reared examples of both sexes from the dead wood of Triplarisspecies at Rio Frio, near Santa Marta, Colombia, during M.a.j, 1928.James Zetek reared it in numbers from avocado branches collected atSummit, Canal Zone, during 1927, and Ferd. Nevermann collected
1 Trans. Ent. Soc. London, 1870, p. 324.
ART. 19 I^TEW BEETLES OP THE FAMILY CEEAMBYCIDAE PISHEE 6
adults at Hamburg farm, Costa Rica, August 15, 1923, on fallen" Quararibea turhinatayODONTOCERA TRIPLARIS, new speciesMale.?Form similar to fasciata Olivier. Head and pronotumyellow, the latter with the anterior margin and base in front ofscutellum narrowly black, and the disk ornamented with reddishbrown as follows : An oblique spot on each side at lateral margin, alongitudinal median vitta extending from anterior margin to base,and arcuately expanded at anterior margin and middle, and anarcuate vitta on each side of middle, more or less interrupted atmiddle, and not extending to anterior margin or base; antennaereddish brown; elytra pale yellow, vitreous, with the margins nar-rowly reddish brown, and a spot of the same color on each elytronnear base; beneath uniformly pale yellow.Head with the front short, flat between the eyes, and with anarrow longitudinal groove extending from vertex to middle offront; surface glabrous, coarsely, vaguely punctate, becoming finelygranulose on the occiput. Eyes separated from each other on thefront by about the width of the labrum. Antennae short, extendingslightly beyond base of elytra; joint 1 shining and sparsely punctate,the following joints subopaque and densely pubescent, and some ofthe joints armed with long erect hairs on inner margin; joint 1robust, arcuate, and rather strongly clavate; joints 2 to 4 subcylin-drical, the third distinctly longer than the fourth, which is slightlyenlarged toward apex; joints 5 to 10 serrate; joint 11 oblong, andacute at apex.Pronotum about as wide as long; sides arcuately rounded andstrongly constricted toward base ; anterior margin and base stronglyelevated; surface with the median dark area slightly elevated atmiddle, finely, confiuently punctate or granulose, and clothed towardthe sides with short, inconspicuous yellow pubescence. Scutellumtriangular, and clothed with a few inconspicuous hairs.Elytra extending to apex of third abdominal segment, with thetips obliquely truncate and the exterior angles acute, the suturalmargins strongly elevated, slightly narrowed, and moderatelydehiscent posteriorly; surface with the reddish brown areas clearlydefined, glabrous, and densely, coarsely punctate, the vitreous areasfinely, very sparsely punctate, and each puncture with a short, erectblack hair.Abdomen beneath coarsely, vaguely punctate, except the first seg-ment, and rather densely clothed with inconspicuous, recumbentyellow hairs. Sternum sparsely clothed with long, semierect yellowhairs. Anterior and middle legs short, and the femora strongly,abruptly clavate. Posterior legs long, the femora strongly, abruptlyclavate, and the tibiae straight and cylindrical.
4 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 77Female.?Differs from the male in having the eyes separated fromeach other on the front by about twice the width of the labrum,and the antennae slightly shorter.Length, 12-16 mm. ; width, 2.6-3.6 mm.Tyfe locality.?Rio Frio, near Santa Marta, Colombia.Type.^ allotype, and paratypes.?Cat. No. 42820, U. S. N. M.Paratypes.?Darlington collection.Described from 48 examples (one type), all of which were rearedduring 1928 by P. J. Darlington, jr., from the dead wood ofTriplaris species. Darlington was unable to get the specific nameof this tree, but he writes that it is the only narrow-leaved speciesof this genus found commonly along the banks of brooks in thefoothills. In a large series of specimens of this species examined,very little variation was observed except in size. There is a slightvariation in the shape of the pronotum; in some examples it isslightly wider than long, whereas in others it is slightly longer thanwide. The reddish brown markings on the pronotum are alsosomewhat variable; in some examples the arcuate vittae are dis-tinctly separated in the middle, and in others they are connectedanteriorly to the oblique lateral spot.This species is allied to tridentifera Gounelle described fromArgentine Republic, but differs from that species in having the headentirely yellow in front, the antennae uniformly reddish brown,pronotum ornamented with three longitudinal reddish brown vittae,which are more or less interrupted at the middle, but not connectedto each other at the base, scutellum yellow and nearly glabrous, andin having the tips of the elytra obliquely truncate, with the exteriorangles acute. ODONTOCERA BUSCKI, new speciesMale.?Form similar to fa^sciata Olivier. Head yellow, with ablack spot on occiput between upper lobes of eyes ; antennae reddishbrown, with the dorsal surface of first and second joints black;pronotum yellow, with the anterior margin and base broadly mar-gined with black, the black margin on base becoming narrower atthe sides, and the disk ornamented with black markings as follows
:
A transverse sinuate fascia at middle extending nearly to coxalcavities, a longitudinal median vitta extending from anterior marginto base, and wider behind the transverse fascia; scutellum black;elytra pale yellow, vitreous, with the lateral and sutural marginsnarrowly brownish red posteriorly, becoming blackish toward thebase, and with an elongate arcuate black fascia behind the base ofeach extending from the lateral margin at basal fourth to the suturalmargin a short distance behind the scutellum; body beneath yellow,with the apical segments of the abdomen dark brown, and orna-
AET. 19 NEW BEETLES OF THE FAMILY CERAMBYCIDAE FISHER 5
merited with black markings as follows: A longitudinal vitta oneach side of the head below the eyes, a transverse fascia on anteriorpart of prosternum, posterior margin of prosternum, a small roundspot on each side of prosternum near base, an elongate spot on an-terior coxae, anterior margin of episternum, an arcuate fascia oneach side of sternum, margin of posterior coxae, and the marginsof posterior intercoxal process.Head with the front short, transversely depressed behind the epis-toma, flat between the eyes, and with a narrow longitudinal grooveextending from vertex to middle of front; surface glabrous, nearlysmooth in front, but becoming densely ocellate-punctate on the occi-put. Eyes separated from each other on the front by about one-half the width of the labrum. Antennae short, extending to base offirst abdominal segment ; joints 1 and 2 shining, and rather densely,coarsely punctate, the following joints subopaque, densely pubescent,and some of the joints armed with long, erect hairs on inner margin;joint 1 robust, straight, and slightly clavate; joints 2 to 4 subcylin-drical, the third twice as long as the fourth, which is slightly morerobust and feebly enlarged toward apex; joints 5 to 10 serrate, andthe joints becoming gradually shorter toward the tip of the antennae
;
joint 11 oblong, acute at apex, and strongly emarginate.Pronotum about as wide as long, with the sides regularly arcuatelyrounded, and the anterior margin and base strongly elevated; diskstrongly convex, without tubercles or carinae; surface densely ocel-late-punctate, and sparsely clothed with a few inconspicuous hairs.Scutellum triangular, and densely clothed with yellowish pubescence.Elytra extending to base of fourth abdominal segment, with thetips transversely truncate and vaguely emarginate, the sutural mar-gins strongly elevated, slightly narrowed, and moderately dehiscentposteriorly ; surface with the opaque areas coarsely, confluently punc-tate, and clothed with a few erect hairs in the basal region, the vitre-ous areas glabrous, and sparsely, obsoletely punctate.Abdomen beneath sparsely, coarsely punctate, except the basal seg-ments, and sparsely clothed with long erect and recumbent hairs in-termixed. Sternum sparsely clothed with long, erect yellow hairs.Anterior and middle legs short, and the femora strongly, abruptlyclavate. Posterior legs long, and sparsely clothed with long, stiff,erect black and yellow hairs intermixed, the femora strongly,abruptly clavate, and the tibiae straight and cylindrical.Female.?Differs from the male in having the eyes separated fromeach other on the front by nearly twice the width of the labrum, andthe antennae slightly shorter.Length, 11.5 mm.; width, 2.4 mm.
6 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 77Type locality.?Trinidad River. Panama (type) ; Tabernilla,Canal Zone (allotype).Tyye and allotype.~C2ii. No. 42821, U. S. N. M.Described from two examples, the male (type) collected at Trini-dad River, Panama, May 7, 1911, by August Busck, and the female(allotype) collected at Tabernilla, Canal Zone, July 23, 1907, by thesame collector.This species is allied to tridentifera Gounelle, but is easily distin-guished from that species by having the three transverse black fas-ciae on the pronotum connected to one another by a median longi-tudinal vitta of the same color.ODONTOCERA ZETEKI, new speciesMale.?Form similar to fasciata Olivier. Head pale yellow, witha transverse black fascia on the occiput between the eyes; antennaeyellowish brown, v/ith the dorsal surface of the basal joint black;pronotum pale yellow, with a broad, transverse black fascia at middle,and along the base and anterior margin; scutellum and elytra paleyellow, the latter vitreous, with the lateral and sutural margins nar^rowly brownish black, and with a narrow oblique black fascia on eachelytron extending from the humerus obliquely backward to thesutural margin behind the scutellum; beneath pale yellow, the legsslightly more brownish, and the body ornamented with black mark-ings as follows: A narrow fascia along anterior margin of proster-num, a transverse fascia at middle and along base of prosternum, butbroadly interrupted at middle, base of each abdominal segment, andnumerous irregular spots on each side of the mesosternum andmetasternum.Head with the front short, and transversely depressed behind theepistoma ; surface nearly smooth in front, becoming coarsely, densely''punctate on the occiput, and rather densely clothed with short,recumbent yellow pubescence. Eyes nearly contiguous in front.Antennae short, extending to middle of first abdominal segment, andrather strongly enlarged toward the apex; joint 1 shining, andcoarsely, sparsely punctate, the following joints subopaque, ratherdensely pubescent, and some of the joints armed with long, erect hairson inner margin; joint 1 robust and slightly arcuate; joints 2 to 4subcylindrical, the third twice as long as the fourth, which issligthly more robust and feebly enlarged toward apex; fifth jointslightly longer and more strongly enlarged toward apex than thefourth ; joints 6 to 10 compact, slightly serrate, and becoming shorterand broader toward the tip of the antennae; joint 11 oblong, andacute at apex.
ART. 19 NEW BEETLES OF THE FAMILY CEKAMBYCIDAE FISHER 7Pronotum distinctly longer than wide, with the sides feebly,arcuately rounded, slightly more strongly narrowed posteriorly, andthe base and anterior margin slightly elevated; disk moderatelyconvex, without tubercles or carinae; surface coarsely, confluentlyocellate-punctate, the yellow fasciae densely clothed with long,recumbent yellow pubescence, and the black fasciae with a few long,erect, inconspicuous hairs. Scutellum densely clothed with recumbentyellow pubescence.Elytra extending to middle of second abdominal segment, withthe tips transversely truncate, and the sutural margins feebly ele-vated, slightly narrowed, and moderately dehiscent posteriorly;surface with the dark margins coarsely, confluently punctate, thevitreous areas sparsely, obsoletely punctate in basal region, and thesurface clothed with a few inconspicuous erect hairs.Abdomen beneath coarsely, vaguely punctate, except the basalsegments, and rather densely clothed toward the sides with recum-bent yellow pubescence. Sternum rather densely clothed with long,erect and recumbent yellow hairs intermixed. Legs sparsely clothedwith long, erect hairs ; anterior and middle legs rather short, and thefemora strongly, abruptly clavate; posterior legs long, the femoragradually clavate, and the tibiae straight and cylindrical.Female.?Differs from the male in having a longitudinal blackvitta and a narrow longitudinal groove on front of head between theeyes, which are separated from each other on the front by nearlytwice the width of the labrum.Length, 11-14.5 mm. ; width, 2-2.6 mm.Type locality.?Barro Colorado Island, Canal Zone.Type^ allotype., and paratypes.?Cat. No. 42822, U. S. N. M.Described from eleven examples, four males and seven females(one type), all of which were reared August 8, 1927, by JamesZetek, from a small Jbranch of an unknown tree, which had beengirdled by a cerambycid, and collected at the type locality. In theseries examined scarcely any variation was observed except in size.This species resembles fasciata Olivier very closely, but can beeasily distinguished from that species because it has three black andtwo pale j^ellow transverse fasciae on the pronotum, whereas infasciata there are four black and two pale yellow fasciae and amedian brownish yellow fascia on the pronotum. Odontocera fas-ciata is also more robust, and the elytra are longer, extending to thebase of the fourth abdominal segment, the tips being acute andstrongly emarginate.
8 PEOCEEDIlSrGS OF THE NATIONAL MUSEUM vol. 7TODONTOCERA NEVERMANNI, new speciesMale.?Form similar to sangwmolenta Bates. Head, antennae,pronotum, and scutellum black; elytra pale yellow, vitreous, withthe lateral and sutural margins and the base broadly black ; beneathblack, with the bases of the posterior tibiae and femora broadly paleyellow.Head with the front rather long, and flat between the eyes; sur-face glabrous, coarsely, irregularly punctate, and longitudinallyrugose on the front, becoming coarsely, confluently punctate on theocciput. Eyes separated from each other on the front by about thewidth of the labrum. Antennae short, robust, extending to base offirst abdominal segment, and with the intermediate joints widest;joints 1 to 3 shining, coarsely punctate, and the third densely pubes-cent, the following joints subopaque, densely pubescent, and someof the joints armed with long, erect hairs on the inner margin; joint1 robust, slightly arcuate, and rather strongly clavate; joints 2 to4 subcylindrical, the third twice as long as the fourth, which isslightly more robust; joints 5 to 10 strotigly serrate, the outeranterior angle of the joints acute, and the joints becoming narrowerand shorter toward tip of antennae; joint 11 oblong and acute atapex.Pronotum distinctly longer than wide, the sides feebly arcuatelyrounded in front, and broadly arcuately constricted behind the middle,the anterior margin strongly elevated ; disk strongly convex, withouttubercles or carinae; surface coarsely, transversely rugose, coarsely,irregularly punctate between the rugae, and clothed with a fewinconspicuous erect hairs. Scutellum densely clothed at apex withlong, recumbent cinereous pubescence, which conceals the surface.Elytra extending to apex of third abdominal segment, the tipsbroadly subtruncate or obtusely rounded, the sutural margins feeblyelevated, slightly narrowed, and slightly dehiscent posteriorly;surface glabrous, with the black areas coarsely, confluently punctate,becoming somewhat scabrous in basal region, and the vitreous areacoarsely, obsoletely punctate.Abdomen beneath shining, very sparsely, finely punctate, andclothed with a few inconspicuous erect hairs. Sternum sparselyclothed with long, erect hairs on posterior half, and the anterior halfdensely clothed with short, recumbent cinereous pubescence. An-terior and middle legs short, the femora strongly abruptly clavate.Posterior legs moderately long, clothed with long, erect black hairs,the femora strongly clavate, and the tibiae straight and cylindrical.Length, 15 mm. ; width, 3.2 mm.Type locality.?Hamburg farm, which is situated on the Kaventa-zon River about midway betweeii Siquires and the coast in CostaRica.
AET. 19 NEW BEETLES OF THE FAMILY CEEAMBYCIDAE FISHER 9Type.?C^t. No. 42824, U. S. N. M.Described from a single male collected flying around cut downtrees at the type locality, May 21, 1925, by Ferd. Nevermann.This species is allied to sanguinolenta Bates, but it differs fromthat species in being larger, the antennae robust but with the inter-mediate joints widest, and the pronotum and underside of the bodyuniformly black, except the base of the posterior tibiae and femora,which are pale yellow.ODONTOCERA DARLINGTONI, new speciesFemale.?Form similar to that of fasciata Olivier. Head andpronotum yellow, the latter ornamented with black markings asfollows: A large, transverse triangular spot, including the anteriormargin; a large, straight, transverse fascia in front of scutellum,but not extending to the lateral margins ; a transverse sinuate fasciaat middle extending nearly to the coxal cavities, which is more orless interrupted externally ; and an elongate spot at middle connectedposteriorly to the median fascia, but not extending to the transversespot in front of scutellum ; antennae reddish brown ; scutellum yellow,with the base blackish ; elytra pale yellow, vitreous, with the lateraland sutural margins narrowly I'eddish brown, and each with anarcuate blackish fascia extending ffom the humeral region back-ward to the sutural margin a short distance behind the scutellum;body beneath uniformly yellow (except margin of each posteriorcoxa which is black), and with a more or less distinct brownishtinge.Head with the front short, transversely depresssed behind theepistoma, flat between the eyes, and with a longitudinal grooveextending from vertex to frontal depression; surface glabrous,coarsely, irregularly and vaguely punctate in front, becoming finelypunctate or granulose on the occiput. Eyes separated from eachother on the front by about one and one-half times the width of thelabrum. Antennae short, extending to base of first abdominal seg-ment; joints 1 and 2 shining, and rather densely, coarsely punctate,the following joints subopaque, densely pubescent, and some of thejoints armed with long, erect hairs on the inner margin; joint 1robust, slightly arcuate, and gradually clavate; joints 2 to 4 sub-cylindrical, the third twice as long as the fourth, which is slightlymore robust and feebly enlarged toward apex; joints 5 to 10 serrateand about subequal in length; joint 11 oblong, acute at apex, andfeebly emarginate.Pronotum vaguely wider than long ; sides feebly arcuately rounded,and slightly constricted at base; anterior margin strongly elevated;disk strongly convex, slightly depressed along base, but without2604?30 2
10 PROCEEDINGS OF THE NATIONAL, MUSEUM vol. 7Ttubercles or carinae; surface densely ocellate-punctate and sparselyclothed with long, inconspicuous hairs toward the sides. Scutellumtriangular, and rather densely clothed with long, recumbent yellowpubescence.Elytra extending to middle of third abdominal segment, with thetips transversely truncate and vaguely emarginate, the sutural mar-gins slightly elevated, slightly narrowed, and moderately dehiscentposteriorly; surface with the opaque areas coarsely, confluentlypunctate, and clothed with a few scattered hairs in the basal regions,the vitreous areas glabrous, and sparselj^, obsoletely punctate.Abdomen beneath sparsely, coarsely punctate, except the basalsegments, and sparsely clothed with long, recumbent yellow pubes-cence. Sternum sparsely clothed with long, semierect yellow hairs.Anterior and middle legs short, and the femora strongly, abruptlyclavate. Posterior legs long, and sparsely clothed with long, stiff,erect black and yellow hairs intermixed, the femora strongly, ab-ruptly clavate, and the tibiae nearly straight and cylindrical.Length, 11-14 mm. ; width, 2.6-3.2 mm.Type locality.?Eio Frio, near Santa Marta, Colombia.Type and paratypes.?C?it. No. 42823, U. S. N. M.Paratypes.?Darlington collection.Described from five females (one type) , all of which were rearedduring May, 1928, from the dead wood of Triplaris species collectedat the type locality by P. J. Darlington, jr. Scarcely any variationwas observed in the specimens examined, except that in two of theseexamples there are two small black spots between the eyes on theocciput.This species is very closely allied to buscki Fisher, but it differsfrom that species in being more robust, with the eyes more narrowlyseparated from each other on front of head, antennae entirely red-dish brown, and the anterior and median transverse fasciae on thepronotum not connected to each other by a longitudinal median blackvitta. With a larger series for study, this form may prove to besimply a color variety of that species, but since this form is connectedup with its host plant, it seems advisable to separate it from thatspecies, at least for the present.ACYPHODERES CRIBRICOLLIS BatesAcyphoderes cnbricollis Bates, Trans. Bnt. Soc. London, 1892, p. 160, pi. 6,fig. 6.This species was described from three males from Ventanas inDurango, Mexico. A small series including both sexes were col-lected by Ferd. Nevermann at Hamburg farm, Costa Rica, August15, 1923, on fallen " QuaraHhea turhinataP This is a very elongate
AKT. 19 ISTEW BEETLES OF THE FAMILY OEEAMBYCIDAE FISHER 1
1
slender species, with the elytra long and only slightly dehiscent pos-teriorly, in which respect it resembles some of the species ofOdontooera, but on account of the longitudinal elevations on thepronotum it is placed in the genus Acyphoderes. In tlie seriesexamined scarcely any variation was observed in the color andmarkings, but the females differ from the males in having the eyesseparated from each other on the front of the head by about oneand one-half times the width of the labrum, and the antennae areslightly shorter. ACYPHODERES BAERI GounelleAcyphoderes baeri Gouneille, Bull. Mus. Nat. Hist. Paris, vol. 19, 1913,pp. 222-224, fig.This species seems to be rather common and was described from alarge series of specimens from Argentine Republic, Paraguay, andBrazil. Material has been examined by the writer from Bolivia(locality not recorded), and the following localities in ArgentineRepublic : Tucuman, collected during December, 1917, and January,1918, by E.W. Rust, and from El Quemado, Jujuy, between Marchand May, 1926, by G. L. Harrington.The species shows great variation in color and markings. Thetypical form is uniformly black above, except for the vitreous areason the elytra, and this color varies to specimens in which the elytraare reddish brown, and the pronotum black, with yellow vittae.Gounelle ^ has described two of these aberrations in which the elytraare reddish brown and -the pronotum ornamented with yellow vittaeunder the names of flavonotata and hiannulata. In the materialcollected by Harrington at El Quemado, the typical form, aberra-tion hiannulata Gounelle, and a form with the elytra reddish brownand the pronotum entirely black were found. E. W. Rust in aletter writes of this species as follows
:
There are also a fly and a moth which resemble these beetles so closely thatyou can scarcely tell the difference when collecting, and all resemble so closelythe wasp Megacantliopus ater Olivier that I never know which one is goingto sting me when I collect them. All four frequent the flowers of Baccharissp. on hot days, and the appearance, habits of flight, and the attitude of thewings in repose or while walking over the flowers is almost identical in allfour cases. The beetle with the wings outspread is the nearly natural positionit assumes when feeding among the flowers.This species is frequently erroneously identified as jSphecomorphamurina Klug, which it resembles very closely. It can, however, beseparated from that species in having the pronotum biimpressedand the elytra gradually narrowed to the apices, each of which termi-nates in an acute spine, whereas in murina the pronotum is strongly
2 Bull. Mus. Nat. Hist. Paris, vol. 19, 1913, pp. 223-224, figs. 1-4.
12 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 77
convex, and the elytra are abruptly dehiscent at the middle, withthe posterior half of each narrow, parallel, and the tip not terminat-ing in a spine. ACYPHODERES AURULENTA (Kirby)Necydalis auruienta Kieby, Trans. Linn. Soc. London, vol. 12, 1818, p. 443.Acyphoderes sericinus White, Cat. C@l. British Museum, pt. 8, 1855, p. 195.This species was described by Kirby (1818) from Brazil, andWhite (1855) redescribed the same species under the name of seri-cirms from the same country. In the United States National Museumare two examples of this species which were identified as abdominaZisOlivier. One of these examples is labeled Mayaguez, Porto Eico,and the other one Cayey, Porto Rico, May 23, 1923, collected byG. N. Wolcott. These two specimens show a slight variation fromthe typical specimens from Brazil by not having the femora andtibiae black in the middle, and by the elytra being without a furcateblack vitta on each side, but these specimens seem to be the extremelight form of this species, and not sufficiently distinct to warrant anew name. In a series examined from Brazil, all of the exampleshave the black bands on the legs, but in a few of these examples thefurcate black vittae on the elytra are absent.This species is closely allied to dbdoTninalis Olivier, but that speciesdiffers from auruienta in having the pronotum more sparsely pubes-cent, and the elytra shining black, each with a short humeral vitta,a small spot on each side of the scutellum, and the vitreous areaspale yellow. The specimens listed as ahdominalis from Porto Ricoby Gahan,^ and Leng and Mutchler * are ^voh^lj auruienta (Kirby).ACTFHODERES RUFOFEMORATA. new speciesFemale.?Form rather slender. Head, antennae, pronotum, andscutellum black ; elytra pale yellow, vitreous, with well defined blackmargins, which are considerably narrower along the sutural marginsexcept for a short distance behind the scutellum. Body beneathblack; anterior and middle legs black, except the femora which arepale yellow at the base ; posterior legs black, the femora pale yellowon basal half, with a black ring at beginning of club, which is brightred.Head with the front long, feebly concave between the eyes, andwith a vague, narrow, longitudinal carina extending from occiput tomiddle of front ; surface coarsely, irregularly punctate, and sparsely,irregularly clothed with rather long, recumbent yellow hairs. Eyesseparated from each other on the front by about one-half the width
?Trans. Ent. Soc. London, 1895, p. 111.* BuU. Amer. Mus. Nat. Hist., New York, vol. 37, 1917, p. 146.
AET. 19 NEW BEETLES OF THE FAMILY CERAMBYCIDAE FISHEE 13
of the labrum. Antennae extending to base of first abdominal seg-ment; joints 1 to 3 shining and sparsely punctate, the followingjoints opaque and densely pubescent; joint 1 slightly clavate; joints2 and 3 slender and cylindrical, the second small, and the third aboutas long as the fourth and fifth united; joint 4 feebly enlarged towardapex; joints 5 to 10 compact and slightly serrate; joint 11 oblong,and acute at apex.Pronotum slightly longer than wide ; sides arcuately rounded, andfeebly constricted at base; surface depressed, somewhat uneven, butwithout distinct longitudinal ridges, densely, coarsely, and conflu-ently punctate, and rather densely, irregularly clothed with moder-ately long, recumbent black and yellow pubescence, with a few erecthairs of the same color intermixed. Scutellum triangular, anddensely clothed with yellow pubescence.Elytra extending to middle of third abdominal segment, ratherstrongly subulate, strongly dehiscent posteriorly, with the tipsnarrowly rounded ; surface glabrous, with the black margins densely,coarsely punctate, becoming tuberculous at the base, and the vitreousareas sparsely, obsoletely punctate.Abdomen beneath shining, rather coarsely, irregularly punctate,and irregularly clothed with recumbent and erect hairs. Sternumrather densely, irregularly clothed with long, recumbent pale yellowpubescence. Anterior and middle legs short, femora stronglyclavate, and the anterior tibiae densely clothed with short brownishpubescence on inner sides. Posterior legs long, femora ratherstrongly clavate near apex, and the tibiae clothed with erect blackhairs, which are denser and longer on the apical half.Length, 17-18 mm.; width, 3.25-3.5 mm.Type locality.?Bahia, Brazil.Type and paratype.?Coi. No. 42825, U. S. N. M.Described from two females (one type) collected at the typelocality.This species is closel}^ allied to femorata Klug and carinicollisBates. It differs from the former in having the abdomen and tarsiblack, and the club of the posterior femora bright red. It resemblescarinicollis Bates very closely, but rufofemorata is larger and morerobust than that species, the anterior and middle legs are blackexcept at the base, and the pronotum is not distinctly trisulcate.BROMIADES BRACHYPTERA (Chevrolat)Odontocera brachyptera Cheveolat, Rev. Zool., 1838, p. 285.This species was described from Cuba by Chevrolat, who recordedit as common in all parts of the island during June on the flowersof Jucaro. He also recorded it as living in the wood of Java
14 PROCEEDINGS OP THE NATIONAL MUSEUM vol.77{^Andira inermis). In the United States National Museum collec-tion is a specimen from Habana, Cuba, collected by C. F. Baker,another example from Central Constancia, Cuba, collected by J. F.Merrill, and two examples collected at Cayuga, Guatemala, duringSeptember and October by William Schaus. Another example hasbeen examined from the Ferd. Nevermann collection which wascollected at San Jose, Costa Rica, June 20, 1924, by E. Gonzora,Bates ^ records it from Cuba and Santa Marta, New Granada, butstates that the specimen from the latter locality differs from theCuban form in having the hind legs wholly tawny red, with theexception of the two apical joints of the tarsi, which are black. Thisspecimen is without any doubt the same as my new species,Bromiades Tneridionalis from Panama, so hrachyptera should notbe listed in the catalogues from New Granada.BROMIADES MERIDIONALIS, new speciesFemale.?Form similar to hrachyptera Chevrolat. Head, anten-nae, pronotum, scutellum, and elytra black, the latter each with abroad, median longitudinal pale yellow vitreous area extending frombase to near apex. Body beneath black; anterior and middle legsblack, with a more or less reddish yellow spot near the base of thefemora; posterior legs yellow, except the femora, which have theswollen parts slightly more brownish and the tips narrowly mar-gined with black, the three apical joints of the tarsi and the tarsalclaws black.Head with the front moderately long, flat between the eyes, andwith a smooth longitudinal carina extending from occiput to middleof front, the carina broadly expanded on the occiput; surfacecoarsely, irregularly punctate, and irregularly clothed with short,recumbent golden yellow hairs. Eyes separated from each other onthe front by nearly the width of the labrum. Antennae extendingnearly to apex of elytra; joints 1 to 4 shining and sparsely punctate;joints 5 to 11 opaque and densely pubescent, and all of the jointsarmed with a few long hairs at apex on inner margin; joint 1strongly clavate; joints 2 to 4 subcjdindrical ; joint 5 robust andfeebly serrate; joints 6 to 10 broadly triangular; joint 11 oblong, andfeebly emarginate at apex.Pronotum about as wide as long; sides arcuately rounded, andstrongly constricted near base; surface somewhat depressed, withthree broad, smooth, longitudinal elevations, the median one straightand strongly interrupted near base, the lateral ones arcuate, notextending to base or apex, and the surface coarsely rugose poste-riorly, the depressions coarsely, irregularly punctate, and densely
?Ann. Mag. Nat. Hist. ser. 4, vol. 11. 1873, p. 120.
ART. 19 NEW BEETLES OF THE FAMILY CEEAMBYCIDAE EISHEE, 15
clothed with long, recumbent golden yellow pubescence. Scutellumtriangular, densely clothed with yellow pubescence at apex.Elytra extending to just beyond base of first abdominal segment,cuneiform, slightly arcuately dehiscent posteriorly, with the tipsrather broadly rounded; surface with the black margins clearly de-fined, glabrous, and coarsely, densely punctate except in the humeralregions, the vitreous areas coarsely, obsoletely punctate, and clothedwith a few erect, inconspicuous yellow hairs.Abdomen beneath slightly flattened, finely, very sparsely punctate,the apical segments densely clothed with short, recumbent cinereouspubescence. Sternum rather densely clothed with short, recumbentcinereous pubescence, with a few long, semierect golden yellow hairstoward the sides. Anterior and middle femora short and stronglyclavate ; posterior femora long and slightly clavate toward the apices.Anterior and middle tibiae short, densely clothed with golden yel-low pubescence on the inner sides; posterior tibiae long, slightlyarcuate, clothed Vv^ith long reddish yellow hairs, which are denserand longer on the apical halves. Tarsi clothed with black hairsexcept the two basal joints of the posterior legs, which are denselyclothed with long reddish yellow hairs, similar to those of the tibiae.Length, 19 mm. ; width, 4 mm.Ty'pe locality.?Cabima, Panama.Type.?QdX. No. 42826, U.S.N.M.Described from a single female collected at the type locality. May26, 1911, by August Busck.This species is closely allied to hrachyptera Chevrolat, but differsfrom that species in having the antennae entirely black, the elytramore arcuately dehiscent toward the tips, and the posterior legs yel-low, with the exception of the tips of the femora and the threeapical joints of the tarsi, which are black. It also resemblesAcyphoderes dbdo^ninalis Olivier very closely, but can be readilyseparated from that species by its short cuneiform elytra, whichbarely pass the base of the first abdominal segment.PHYGOPODA MANNI, new speciesMale.?Head, pronotum, and scutellum black; antennae reddishbrown, becoming slightly paler on the basal segments; elytra paleyellow, somewhat vitreous, with the lateral and sutural marginsbrownish black, the brownish-black areas very broad along the lat-eral margins, and with an oblique spot of the same color on eachelytron behind the scutellum; body beneath black except the lastabdominal segment, which is reddish; anterior and middle legs red-dish brown, the femora slightly darker; posterior legs black, withthe tarsi and bases of femora and tibiae pale yellow.
16 PROCEEDINGS OF THE NATIONAL, MUSEUM vol. 77Head with the front rather short, flat, and with a narrow longi-tudinal groove between the eyes ; surface rather coarsely, irregularlypunctate, and rather densely clothed with long, recumbent cinereouspubescence. Eyes narrowly separated on the front. Antennae ex-tending nearly to apex of elytra; joint 1 shining and sparselypunctate, the following joints opaque, densely pubescent, and witha few long, stiff hairs at apex on inner margin; joint 1 slightlyclavate ; joints 2 to 4 slender and cylindrical, the third only slightlylonger than fourth, which is subequal in length to the fifth; joints 5to 10 triangular, the joints becoming shorter and broader toward tipof antennae; joint 11 oblong, and feebly emarginate at apex.Pronotum about as wide as long; sides slightly arcuately rounded,and rather strongly constricted at base ; surface moderately convex,without tubercles or longitudinal ridges, rather coarsely reticulate-punctate, each puncture with a small round elevation in the middle,and the surface rather densely, irregularly clothed with long, recum-bent cinereous pubescence, which is denser along the anterior partof the lateral margins. Scutellum triangular, densely clothed withcinereous pubescence at apex.Elytra extending to just beyond base of first abdominal segment,abruptly subulate behind the middle, strongly dehiscent posteriorly,the tips narrowly rounded ; surface glabrous, with the brownish-blackareas coarsely, confluently punctate, and the vitreous areas with afew coarse, inconspicuous punctures.Abdomen beneath, except last segment, densely clothed with long,recumbent cinereous pubescence, which nearly conceals the surface.Sternum finely, densely punctate, and clothed with pubescence sim-ilar to that on the abdomen. Anterior and middle legs short, thefemora strongly clavate. Posterior legs very long, the femora feebly,gradually clavate, coarsely scabrous, and sparsely clothed with short,stiff, semierect white hairs; tibiae armed on apical halves with arather dense tuft of long, erect white and black hairs intermixed,the white hairs predominating apically.Female.?Differs from the male in having the eyes separated fromeach other on the front by about the width of the labrum, the an-tennae shorter, extending only to middle of elytra, and the posteriorfemora yellowish brow^n, becoming dark brown on the upper margina,nd at middle of outer surfaces.Length, 8.5-9 mm. ; width, 2.4 mm.Ty'pe locality.?Kurrenabaque and Ganamina, Bolivia.Ty^e and allotype.?C2Lt. No. 42827, U. S. N. M.Described from two examples, the type (male) collected at Kurre-nabaque (Beni River), Bolivia, December, 1921, and the femalecollected at Canamina, Bolivia, July, 1921. Both specimens were
ART. 19 NEW BEETLES OF THE FAMILY CEEAMBYCIDAE FISHER 17
collected by William M, Mann, with the Mulford Biological Explora-tion in Bolivia during 1921-1922.This species is allied to albitarsis Klug and fugws Thomson, butdiffers from albitarsis in not having the pronotum longitudinallycarinate, and from fugax by having the elytra abruptly subulate andstrongly dehiscent posteriorlj^TOMOPTERUS VESPOIDES WhiteToniopterus vespoides White, Cat. Col. British Museum, pt. 8, 1855, pp.176-177, pi. 5, fig. 8.This interesting little species was described by White from Gua-/ temala, and has been collected in numbers by Ferd. Nevermann atHamburg farm, Costa Rica, during August, 1925, flying like waspsaround freshly cut timber, apparently " Sapotaceae," and he hasobserved a female depositing eggs in a hole in the dry bark. Never-mann writes that the females seem to be very rare, as out of 35examples collected, only two were females.The sexes of this species are quite different ; the males have the eyesnearly contiguous in front, and are dull black above, with the an-terior and posterior margins of the pronotum, two spots on themetanotum, last dorsal segment of abdomen, and the posterior mar-gins of the ventral surface of the abdominal segments clothed withgolden yellow pubescence. The elytra have the anterior part of thelateral margins and a narrow, oblique vitta on each elytron reddishyellow, the vitta extending from the humerus to middle of elytron atapex. The females have the abdomen more robust, the eyes dis-tinctly separated from each other on the front, the surface sparselyclothed with yellowish pubescence, and in addition to the yellowpubescence mentioned on the males, have the median part of the pro-notum sparsely, and the scutellum densely clothed with golden yellowpubescence. The reddish yellow margins and vittae on the elytraare distinctly broader, and in some examples these unite and theentire lateral half of the elytron is reddish yellow, which is the formfigured by White. TOMOPTERUS VITTIPENNIS, new speciesMale.?Form similar to qioadratipennis Bates. Above uniformlyblack, each elytron with the lateral margin and a narrow obliquevitta brownish yellow, the color becoming paler posteriorly, and thevitta extending from the humeral region to apex at middle of ely-tron; beneath brownish black, with the base of posterior femoraand the inner surface of posterior femora and posterior tibiae paler.Head with the front moderately long, and with a narrow longi-tudinal carina on the occiput; surface finely, densely punctate, and
18 PROCEEDINGS OF THE NATIOISTAL MUSEUM vol.77densely clothed between and below the eyes with long, recumbentsilvery-white pubescence. Eyes nearly contiguous in front. An-tennae short, extending to base of pronotum, and strongly expandedtoward apex; joint 1 long and strongly clavate; joints 2 to 4 cylin-drical, and the third slightly longer than fourth; joint 5 subequalin length to the third, and strongly enlarged toward apex; joints6 to 10 compact, short, about subequal in length, and only feeblyserrate; joint 11 slightly oblong, emarginate, and acute at apex.Pronotum slightly wider than long, and distinctly narrower at basethan at apex; sides arcuately rounded in front of middle, and ob-liquely narrowed posteriorly to the base; disk strongly convex, witha narrow transverse depression along anterior margin and base ; sur-face coarsely, confluently ocellate-punctate, densely clothed with veryshort, inconspicuous silvery-white pubescence, which is visible incertain lights only, with a few erect black hairs intermixed, andornamented with a narrow fascia of long, recumbent yellow hairsalong the anterior margin. Scutelium very large, elongate, tri-angular, and the surface coarsely, confluently punctate.Elytra very short, not extending to base of abdomen, quadran-gular, not dehiscent posteriorly, the tips broadly rounded exteriorly,more or less obliquely toward the sutural margins; surface verycoarsely, confluently punctate, and sparsely clothed with fine, incon-spicuous, erect hairs in the basal region.Abdomen beneath rather densely, coarsely punctate, the posteriormargin of segments broadly clothed with long, recumbent yellowpubescence. Sternum coarsely, densely punctate, finely granulose,and rather densely clothed with moderately long, recumbent silvery-white pubescence, with a few semierect hairs of the same color inter-mixed. Prosternum clothed with a few very long, erect black hairs.Anterior legs short ; anterior femora robust, gradually clavate ; mid-dle femora feebly arcuate, and strongly abruptly clavate. Posteriorlegs long; femora strongly arcuate, strongly clavate on apical two-thirds; tibiae cylindrical, slightly sinuate.Length, 8.8 mm. ; width, 2.6 mm.Type locality.?Buenavista, Bolivia.Type.?Q^t. No. 42828, U. S. N. M.Described from a single male collected at Buenavista, Bolivia,between February and April, 1925, and labeled " Provincia d'Sara,1,700 ft."This species is allied to quadratipennis Bates, but quardratipennisdiffers from mttipennis in having the antennae reddish brown, thebase and anterior margin of pronotum narrowly margined withyellow pubescence, a narrow transverse pubescent fascia on each sideat lateral margin, the oblique reddish vitta on each elytron extend-
ART. 19 NEW BEETLES OP THE FAMILY CEEAMBYCIDAE FISHER 19ing from the humeral angle to the apex at sutural margin, the ab-dominal segments not distinctly margined with yellow pubescence,and the legs reddish brown.TOMOPTERUS SIMILIS. new speciesM(de.?Form similar to quadratipennis Bates. Above uniformlyblack, antennae brownish black, with the base of the first jointreddish, and each elytron with the lateral margin yqtj narrovviyreddish brown and a short, oblique, smooth ivory white vitta, whichdoes not extend to base or apex of elytron ; beneath brownish black,the legs reddish brown, with the base of the posterior femora ivorywhite.Head with the front moderately long, broadly, deeply concavebetween the antennae, and with a short, longitudinal carina on theocciput; surface jQnely, densely punctate, densely clothed below theeyes with long, recumbent silvery white joubescence, and with a rowof similar hairs between the eyes, which are nearly contiguous infront. Antennae short, extending to base of pronotum and stronglyexpanded toward apex; joint 1 long, arcuate, and strongly clavate;joints 2 to 4 cylindrical, the third distinctly longer than the fourth
;
joint 5 subequal in length to the third, and strongly enlargedtoward the apex; joints 6 to 10 compact, short, subequal in length,and strongly serrate; joint 11 slightly oblong, broadly truncate andemarginate at apex.Pronotum distinctly wider than long, nearly equal in width at baseand apex; sides broadly, arcuately rounded, more strongly towardthe base ; disk moderately convex, vaguely flattened on top, and witha deep narrow, transverse depression along anterior margin andbase; surface coarsely, confiuently ocellate-punctate, densely clothedwith very short, inconspicuous silvery white pubescence, which is onlyvisible in certain lights, with numerous long, erect black hairs inter-mixed, and ornamented in the basal depression with long, recum-bent golden yellow pubescence. Scutellum very large, elongate, tri-angular, the surface coarsely, densely, and irregularly punctate.Elytra very short, not extending to base of abdomen, quadrangular,not dehiscent posteriorly, the tips broadly rounded externally, moreor less obliquely toward the sutural margins; surface densely, con-fluently punctate, the punctures very coarse in basal region, but be-coming finer toward the apex, and sparsely clothed with fine, incon-spicuous, erect hairs in basal region.Abdomen beneath finely, densely punctate, the posterior marginof segments densely clothed with long, recumbent pubescence, whichis silvery v/hite on basal segments and golden yellow on apical seg-ments. Sternum closely clothed with long, erect and recumbent
20 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 77
silvery white pubescence intermixed. Prosternum sparsely clothedwith long, erect black hairs. Anterior leg's short; anterior femorarobust and gradually clavate. Middle femora strongly arcuate, andstrongly, abruptly clavate. Posterior legs long ; femora strongly ar-cuate, and strongly clavate on apical two-thirds; tibiae cylindricaland slightly sinuate.Length, 8.4 mm. ; width, 2.8 mm.Type locality.?Rio Negro, Bolivia.7V??e.?Cat.'jSTo. 42829, U.S.N.M.Described from a single male collected at the type locality duringJanuary, 1922, by William M. Mann with the Mulford BiologicalExploration in Bolivia during 1921-1922.This species is allied to quadratifennis Bates, but differs from thatspecie's in having the antennae brownish black, the transverse basaldepression on pronotum densely clothed with golden yellow pubes-cence, the oblique ivory white vitta on each elytron not extendingto the base or apex, and the underside of the abdominal segmentsbroadly clothed along the posterior margins with silvery white orgolden yellow pubescence. o