THE CENTIPEDS OF CENTRAL AMERICA.
By Ralph V. Chamberlin,Of the Museum of Comparative Zoology, Cambridge, Massachusetts.
This paper is primarily a report upon the centipeds from theCentral American region contained in the collections of the UnitedStates National Museum. Surprisingly little has been done uponthe centiped and milliped fauna of this region; and that ourknowledge is still far from complete is amply attested by the per-centage of new forms appearing in each additional collection studied,this being especially noticeable in the case of the much more numer-ous millipeds with their more restricted distributions. The mil-lipeds will be treated in a separate report to follow. The centipedsnow known from the several countries of the region under consider-ation are listed separately below.PANAMA.Scolopocryptops miersii Newport.Rhysida nuda (Newport).Cupipes tmgulatus Meinert.Scolopendra subspinipes Leach.Scolopendra morsitans Linnaeus.PselHodes nigrovittattis (Meinert).Orphnaeus brevilabiatus (Newport),COSTA BICA.Cryptops bivittatus Pocock.Otocryptops melanostomus (Newport).Netcportia longitarsis (Newport).Newportia rogersi Pocock.Otostigmus denticulatus (Pocock).Otostigmvs scabHcaudus (Humbert and Saussure).Rhysida nuda (Newport).Rysida longipes (Newport).Scolopendra viridis Say.Scutigera linceci Wood.Scutigera nubila, new species.Labrobius costaricensis (Brolemann).Notiphilides maximiliani (Humbert and Saussure).No. 2402?Proceedings U. S. National Museum, Vol. 60, Art. 7.
PROCEEDIIsrGS OF THE NATIONAL MUSEUM. vor.. 60.NICAKAGUA.Otocnjptops melanostonius (Newport).Rhysida nuda (Newport).Rhysida celeris (Humbert and Saussure).Scolopendra viridis Say.Scutigera linceci Wood.Orphnaeus brevilabiatus (Newport).SAN SALVADOR.No centipeds recorded. HONDURAS.Cryptops pugtmns, new species.Otocryptops ferrugineus (Linnaeus).Otocryptops melanostonius (Newport).Neicportia stolli (Pocock).Newportia mimetica, new species.Neivportia sulana, new species.Scolopendra viridis Say.Scolopendra gigantea Linnaeus.Scolopendra polymorpha gaumeri Pocock.Ityphilns ceibanus, new species.Tanophilus hondurasanus, new species.Orphnaeus brevilabiatus (Newport).Suturodes tardus, new species.BEITISH HONDURAS.Rhysida nuda (Newport).Scolopendra sumichrasti Saussure.Scolopendra polymorpha gaumei-i Pocock.Scolopendra morsitans Linnaeus.GUATEMALA.Cryptops micrus, new species.Otocryptops ferrugineus (Linnaeus).Otocryptops melanostomus (Newport).Newportia stolli Pocock.Newportia divergens, new species.Otostigmus denticulatus (Pocock).Rhysida nuda (Newport).Cormocephalus aurantiipes (Newport).Scolopendra sumichrasti Saussure.Scolopendra viridis Say.Scutigera linceci Wood.Labrobius vulcafii (Pocock).Labrobius cobulcanus, new species.Soivubius stolli (Pococlv).Sotimpius decodontus (Pocock).Sogolabis scapheus Cliamberlln.Schcndylelius hoditcs Chamberlin.Sogodes difflcilis, new species.Orphnaeus brevilabiatus (Newport).Notiphilides maximiliani (Humbert and Saussure)Suturodes guatemalae, new species.Suturodes stolli (Pocock).
AET. 7. CENTIPEDS OF CENTRAL. AMERICA CHAMBERLIN. 3The United States National Museum material studied is chieflyfrom Honduras, collected by Dr. W. M. Mann, and from Guatemala,collected by Dr. O. F. Cook and G. P. Goll. Eleven of the 12 newforms described are from these two countries. The total number ofspecies listed is 43.Order SCOLOPENDROMORPHA.Family CRYPTOPIDAE.Genus CRYPTOPS Leach.
1. CRYPTOPS BIVITATTUS Pocock,Cryptops divitattus Pocock, Joiirn. Linn. Soc. London, 1894, voL 24, p. 462.
?
Kkaepelin, Revision der Scolopend., 1903, p. 50.Locality.?Costa Kica.
2. CRYPTOPS MICRUS, new species.Plate 1, fig. 2.In the type of this species the basal plate overlaps the cephalic.Cephalic plate with paired longitudinal sulci. Basal plate withsemicircular cervical groove and paired longitudinal sulci. Anteriormargin of prosternum gently doublj'^ bowed. Sulci of anterior platescomplete. Last ventral plate with sides straight and convergingcaudad. Coxopleurae not produced caudad; pores few, only aboutfour on each side in the type.Characterized particularly by features of the anal legs. In thesethe femur wholly lacks teeth and true spines and the tibia has onlya single tooth at distal end beneath. Both femur and tibia bearventrally numerous stout setae, with sparse fine hairs above. Meta-tarsus with four teeth below and the first tarsal joint with two.Tibia and first tarsal joint furrowed and notched at distal end above*Length, about 10 mm.Locality.?Guatemala : Trece Aguas, one specimen, June, 1907(O.F.Cook).Type.?C2it. No. 24118, U.S.N.M.
3. CRYPTOPS PUGNANS, new species.Among American forms similar to the Cu1>un C. cornifer Cham-berlin and the Argentinian G. galatheae Meinert in having pairedsulci extending over the full length of the cephalic plate. It agreeswith C. cornifer in having the first dorsal plate with a transversesulcus that is evenly curved, not at all angled at the middle, whichthe paired and parallel longitudinal sulci attain but do not cross.Prosternal margin doubly convex; bristles 7+7. Tarsi distinctlybiarticulate, excepting a few of the most anterior. Last ventral
4 PROCEEDINGS OF THE NATIONAL MUSEUM. vol.60.plate with sides converging, its caudal margin widely incurved.Coxopleurae not produced caudally; pores moderate in number.Femur of anal legs armed below and laterally with numerous spinesexcepting a median longitudinal naked area below, as in G. cornifer;sulcata above, at distal end, with a stout curved spine on mesal side.Tibia also with slender spines beneath ; at distal end above with twostout spines. Metatarsus similarly with two stout spines at distalend above and with seven teeth beneath. First tarsal joint withthree teeth beneath. Proximal joints of penult and preceding legsalso with numerous spines beneath. Tibiae and metatarsi of penultlegs subdensely setose beneath, the former also with some spines.Length, 15.5 mm.Locality.?Honduras: Progreso, one specimen (W. M. Mann).Type.?C^t. No. 24119, U.S.N.M.A smaller form than the Cuban C. cormifer Chamberlin, to whichit seems most closely related. It differs in its biarticulate tarsi,the presence of two stout spines at distal end of tibia of anal legsinstead of one, of seven teeth below on metatarsus instead of ten, etc.Genus OTOCRYPTOPS Haase.4. OTOCRYPTOPS FERRUGINEUS (Linnaens).Scolopendra ferruginca Linnaeus, Syst. Nat., ed. 12, 1766, p. 1063.Otocryptops ferrugineus Kraepelin, Revision der. Scolopend., 1903, p. 72.Localities.?Honduras: Choloma, one specimen (W. M. Mann);Guatemala. 5. OTOCRYPTOPS MELANOSTOMUS (Newport).Scolopocryptops melanostomus Newport, Trans. Linn. Soc. Loudon, 1844,vol. 19, p. 406.Otocryptops nielanostoma Pocock, Journ. Linn. Soc. London, 1894, vol. 24,p. 464.Localities.?Costa Eica: Rio Eeventazon (P. P. Calvert), LaPalma, and Surubres ; Nicaragua ; Honduras : San Juan Pueblo, threespecimens (W. M. Mann) ; Guatemala : near Guatemala city.Genus SCOLOPOCRYPTOPS Newport.6. SCOLOPOCRYPTOPS MIERSII Newport.Scolopocryptops miersii Newport, Trans. Linn. Soc. London, 1845, vol.19, p. 405.Locality.?Panama.Genus NEWPORTIA Gervais.7. NEWPORTIA STOLLI (Pocock).Scolopendrides stolli Pocock, Biol, centr. Amer., 1896, p. 31, pi. 3, flg. 4.Netvportia stolli Kraepelin, Revis. der Scolopend., 1903, p. 85.Localities.?Honduras : Progreso, one specimen (W. M. Mann)
;
Guatemala : Quetzaltenango.
ABT.7. CENTIPEDS OF CENTRAL AMERICA?CHAMBERLIN". 5The specimen from Honduras is referred with some doubt to thisspecies, as the anal legs are missing. It may possibly, instead, repre-sent the form N. sulana, new species, described below.
8. NEWPORTIA LONGITARSIS (Newport).Scolopocryptopa longitarsis Newport, Trans. Linn. Soc. London, 1845, vol.19, p. 407, pi. 40, fig. 10.Netcpoi'tia longitarsis Gervais, in Ins. Apt., 1847, vol. 4, p. 298.Locality.?Costa Eica: Eeventazon Valley, Juan Vinas (P. P.Calvert).
?. NEWPORTIA ROGERSI Pocock.Newportia rogersi Pocock, Biol. Centr. Amer., Chilopoda, 1896, p. 33, pi. 3,figs. QSd.Localities.?Costa Bica : Volcan de Irazu, Cocos, San Jose.
10. NEWPORTIA MIMETICA, new species.This species is close to the Guatemalan N. stolli (Pocock). As inthat species, the head has paired longitudinal sulci extending fromthe caudal margin forward to or a little beyond the middle; thesesulci not connected posteriorly by any transverse line. Cervicalsulcus evenly curved, in the types covered by the cephalic plate orfree only at the middle. Paired sulci of first plate parallel and un-branched, extending forward in front of transverse sulcus. Pairedsulci extending across anterior third of second tergite, but completeon the others to the twenty-second, inclusive. Last tergite withouta median sulcus. Prosternum with dental plates short and broad,convex. Last tergite without a median sulcus. Tarsi undividedexcepting in last two pairs of legs. Tibiae of anterior legs armedbeneath at distal end but not laterally. Last ventral plate with sidesconverging caudad, caudal margin mesally incurved, corners rounded.Coxopleural processes nearly glabrous, lacking the pilosity on exter-nal side present in N. stolli and N. sulana. Anal and penult legsalso with but very few hairs on mesal surface in contrast with thecondition in N. stolli.The general form of the anal legs as in N. stolli^ the tarsal divisionsimilarly proportioned and without claw. Femur of anal legs flat-tened within; lower edge typically with five distally curved teeth,of which the two anterior ones are smaller ; upper edge with a num-ber of points or spinules. and below them, on mesal surface, somestill more minute points or prickles. Differing from N. stolli inhaving the two ventral teeth of the tibia situated one well behindthe middle and one in front of it instead of both being in front of it.Length, 15-19 mm.
6 PROCEEDINGS OF THE NATIONAL. MUSEUM. vol.60.Locality.?Honduras: Lombardia, three specimens (W. M. Mann).TypG.?C2d,. No. 24120, U.S.N.M.In one of the paratypes the head plate is abnormally widely over-lapped by the first tergite.
11. NEWPORTIA DIVERGENS, new species.
? Plate 1, fig. 1.Head with paired sulci extending forward to middle, these not con-nected across base. The first dorsal plate differs from that of therelated N. stolli and N. sulana and N. mimetica, as well as the lessclosely related forms, in not having the cervical or transverse sulcuscross the middle; that is, the segment of the sulcus between thepaired sulci is missing, the paired sulci being thus, in effect, bifur-cate, the main branch continuing forward and the other curvingectocephalad.The characters of the posterior legs essentially as in zV. stoUi,the spining and hair being the same, or nearly so, though the minutespinules on the mesal face of the femur appear to be more numerousand more evenly distributed. The spinules or teeth on the tibiavary considerably. While typically two in number on the proximalhalf, a third may be present on the distal half and the spinuleslarger in size, and, on the other hand, in one specimen there is buta single spine on one of the legs.Length, about 27 mm.Locality.?Guatemala : Joyabaj, San Rafael, nine specimens (O. F.Cook, 1906 and 1914).Type.?Cat. No. 24121, U.S.N.M.
12. NEWPORTIA SULANA, new speciei.Plate 1, fig. 3.Agreeing in general with N. stolli, thus far with certainty knownfrom only a single specimen taken at Quetzaltenango, Guatemala,excepting that the tarsus of the anal legs ends in a distinct, thoughstraight, claw, as in N. amazonica Brolemann. The posterior pairsof legs are densely pilose over the mesal (caudal) surface and thecoxo-pleural processes are also pilose ectally and above.Length, 21.5 mm.Locality.?Honduras : San Pedro, Sula, one complete specimen andtwo specimens lacking anal legs (W. M. Mann).Type.?^o. 24122, U.S.N.M.Analogy with N. amazonica Brolemann and N. unguifer Cham-berlin, both known from numerous specimens, indicates that theclaw in the present form is likewise a constant character.
ABT. 7. CENTIPEDS OF CENTRAL AMERICA?CHAMBERLIN. 7Family OTOSTIGMIDAE.Genus OTOSTIGMUS Porat.X?. OTOSTIGMUS DENTICULATUS (Pocock).Parotostigmua denticulatus Pocock, Biol. Centr. Amer., Cliilopoda, 1896,p. 25.Otostigmus denticulatus Kbaepelin, Bevls. der Scolopend,, 1903, p. 124.Localities.?Costa Rica : San Jose ; Guatemala.14. OTOSTIGMUS SCABRICAUDUS (Humbert and Saussure).-Branchiostoma scahricauda Humbeet and Saussuee, Rev. et Mag. Zool.,1870, vol. 22, p. 203.Otostigmus scaltricaudus Kbaepelin, Revis. der Scolopend., 1903, p. 123,fig. 61.Localities.?Costa Rica: San Mateo, Cocos,Genus RHYSIDA Wood.15. RHYSIDA NUDA (Newport).Branehiostonm nudum NEwroET, Trau.s. IJnn. Soc. London, 1845, vol. 19,p. 412.Rhysida nuda Kbaepelin, Revis. der Scolopend., 1903, p. 144,Localities.?Panama: Volcan de Chiriqui, Oriental region; Nica-ragua : Greytown ; Costa Rica : San Jose : British Honduras : StannCreek, Belize (Robertson)
.
IG. RHYSIDA LONGIPES (Newp?rt).Branchiostoma, longipes Newpobt, Trans. Linn. Soc. London, 1845, vol. 19,p. 411.Rhysida longipes Pocock, Biol. Centr. Amer., Chilopoda, 1896, p. 27.Tvocality.?" Central America."17. RHYSIDA CELERIS (Hnmbert and Saussnre).Branchtostoma celer Humbeet and Saussuee, Rev. et Mag. Zool., ser. 2,vol. 22, p. 202.Rhysida celeris Kbaepelin, Revis. der Scolopend., 1903, p. 149.Locality.?Nicaragua (McNeil coll.).Family SCOLOPENDRIDAE.Genus CUPIPES Kohlrausch.18. CUPIPES UNGULATUS Meinert.Cupipes ungulatus Proc. Amer. Philos. Soc. 1886, vol. 23, p. 187.Locality.?Panama.Genus CORMOCEPHALUS Newport.19. CORMOCEPHALUS AURANTIIPES (Newport).{^colopendra aurantiipes Newpobt, Ann. and Mag. Nat. Hist., 1844, vol. 13,p. 99.Cormocephalus aurantiipes Newport, Trans. Linn. Soc. London, 1845, vol.19, p. 420.Locality.?Guatemala (F. M. Miiller).
8 PROCEEDINGS OF THE NATIONAL MUSEUM. vol.60.A single specimen in the collection of the Museum of Compara-tive Zoology labeled as coming from Guatemala is this form. Ithad probably escaped from some ship, as the species is charac-teristically Australian.Genus SCOLOPENDRA Linnaeus.20. SCOLOPENDRA GIGANTEA Linnaeus. 'Scolopendra gigantea Linnaeus, Syst. Nat, ed. 10, 1758, p. 638.Locality.?Honduras.21. SCOLOPENDRA SUMICHRASTI SansBure.Scolopendra suniichrasti Saussuke, M6m. Soc. Phys. Genfeve, 1860, vol. 15,p. 385, pi. 7, fig. 46.Localities.?British Honduras : Belize ; Guatemala : Tucuru, inVera Paz, Livingston.22. SCOLOPENDRA POLYMORPHA GAUMERI Pocock.Scolopendra polymorpha gaumeri Pocock, Biol. Centr. Amer., Chllopoda,1895, p. 20.Localities.?Honduras: Bonaca Island, in Bay of Honduras;British Honduras. 23. SCOLOPENDRA VIRIDIS Say.Scolopendra viridis Say, Proc. Acad. Nat. Sci. Philadelphia, 1821, p. 110.Scolopendra nicaraguensis Bollman, Bull. U. S. Nat. Mus., 1893, No. 46,p. 198.Localities.?Costa Rica: Juan Viiias (P. P. Calvert), Cache;Nicaragua: Ocotal (W. B. Richardson, Dr. Bransford) ; Honduras(C. H. Townsend) ; Guatemala: Quetzaltenango, Volcan de Pacaya,Guatemala city, Antigua, San Miguel Upsantan.24. SCOLOPENDRA MORSITANS Linnaeus.Scolopendra morsitans Linnaeus, Syst. Nat., ed. 10, 1758, p. 638.Localities.?Panama; British Honduras: Belize (W. A. Stanton),St. Johns College (Wm. Bennett).25. SCOLOPENDRA SUBSPINIPES L?ach.Scolopendra subspinipes Leach, Trans. Linn. Soc, 1814-15, vol. 11, p. 383.Locality.?Panama: Canal Zone, Tabernilla.Order SCUTIGEROMORPHA.Family SCUTIGERIDAE.Genus PSELLIODES Chamberlin.26. PSELLIODES NIGROVITTATA (Meinert).Scutigera nigroiHttata Meinebt, Proc. Amer. Philos. Soc, 1886, vol. 28,p. 173.Locality.?Panama. This is the type locality for the species.
AET.7, CENTIPEDS OF CENTRAL AMERICA CHAMBERLIN. 9Genus SCUTIGERA Lamarck.27. SCUTIGERA UNCECI Wood.Scutiffera linceci Wood, Proc. Acad. Sci. Philadelphia, 1867, p. 42.
28. SCUTIGERA NUBILA, new spedeg.Plate 1, tigs. 4 and .5.The dorsum is fuscous throughout, without any distinct medianpaler stripe, though the tergites appear a little darker on the sidesthan along the middle. The legs are blackish with an indistinct andincomplete paler annulus over middle of femur and tibia and witha vaguer annulus at base of latter on some legs. Articles of antennaevery short, much broader than long, first division of flagellum con-sisting of 79 articles. Head deeply depressed over posterior medianregion, the depression somewhat furcate between eyes. A shallowermedian longitudinal furrow on anterior part of head. The stoma-bearing tergites notched or incurved at middle behind, the stomaslightly projecting caudad. Stoma saddles moderately elevated, thestomata of moderate length, on caudal slope of saddles. Last ter-gite with caudal margin flattened at middle, but not at all excised.Dorsal surface bearing numerous short, slender spines or spiniformsetae, all unaccompanied by finer hairs. The slender spines numer-ous on margins, more sparse over the general surface.The body of the type is injured at the caudal end, so that the geni-tal forceps cannot be described.First division of tarsus I composed of 14 articles, the second of32. First tarsus of seventh legs with 10 articles, the second with 30.First tarsus unspined distally. Second tarsus of anterior legs withpegs beneath in typical manner. Spines present at distal end oftarsus I in median and posterior legs.Length, 17.5 mm.Locality.?Costa Eica: Turrialba, one specimen (C. Lankester).Type.?Cat. No. 24123, U.S.N.M.Order LITHOBIOMORPHA.Family GOSIBIIDAE.Genus LABROBIUS Chamberlin.29. LABROBIUS VULCANI (Pocock).LithoMus vulcani Pocock, Biol. Centr. Amer., Chilopoda, 1895, p. 8, pi. 1,figs. 8-8&.Lahrobius vulcani Chamberlin, Bull. Mus. Conip. Zool., 1915, vol. 59, p. 536.Locality.?Guatemala : Volcan de Agua.
10 PROCEEDINGS OF THE NATIONAL MUSEUM. tol. 60.
30. LABROBIUS COSTARICENSIS (Brolemann).Lithobius vvlcani costaricenais Beolemann, Ann. Soc. Ent. France, 1905,p. 339.Locality.?Costa Rica: El Reventado.81. LABROBIUS COBULCANUS. new species.Plate 1, figs. 6 and 7.This form is separated chiefly on the characters of the prosternum.This differs from L. vulcani (Pocock) in lacking the stout outerthird tooth on each side of the prosternum. The prosternal marginextends out horizontally considerably beyond the ectal tooth, muchas in L. minor Chamberlin ; the edge a little ectad of the outer toothis produced cephahid, the special ectal seta being borne upon theprominence, as shown in figure 6. Articles of antennae, 33 to 39.Ventral spines of penult legs, 0, 1, 3, 3, 1-0, 1, 3, 3, 2, with threeclaws. Claw of female genital forceps entire; basal spines 2-{-2,rather stout (fig. 7). Color, chestnut.Length, 16 mm.Locality.?Guatemala: Joyabaj, lower slope of Cobulco Mountain,two females, both of which have lost their anal legs (O. F. Cook).Type.?Cat. No. 24124, U.S.N.M.Genus SOWUBIUS Chamberlin.32. SOWUBIUS STOLU (Pocock).Lithobius stolU Pocock, Biol, Centr, Amer., Chilopoda, 1895, p, 9, pi. 1,figs, 10-lOc,Soiotibius stolli Chamberlin, Canad, Ent., 1912, p. 178.Locality.?Guatemala: Volcan de Agua,Genus SOTIMPIUS ChamberUn.33. SOTIMPIUS DECODONTUS (Pocock).Lithobius decodontus Pocock, Biol, Centr, Axuer., Chilopoda, 1895, p. 9,pi. 1, figs. 9-9&,Sotiinpius decodontus Chamberlin, Canad, Ent.. 1912, p, 177,Locality.?Guatemala : Volcan de Acatenango.Order GEOPHILOMORPHA.Family SCHENDYLIDAE.Genus SOGOLABIS Chamberlin.34. SOGOLABIS SCAPHEUS Chamberlin.Sogolabis scapheus Chamberlin, Psyche, 1920, vol. 27, p. 64.Locality.?Guatemala : Coban,The type of this species was taken actually at Washington. Dis-trict of Columbia, but from soil about roots of the pacaya or saladpalm (Chamaedorea) at quarantine, im.ported from Coban.
AST. 7. CENTIPEDS OF CENTBAL AMERICA?CHAMBERLIN. 11Genus SCHENDYLELLUS Chamberlin.35. SCHENDYLELLUS HODITES Chamberlin.Schemlylellus hodites Chambkrlin, Psyche, 1920, vol. 27, p. 65.Locality.?Guatemala : Coban.The type was also taken at quarantine in Washington, District ofof Columbia, from about roots of the Chamaedorea imported fromCoban. SOGODES, new genus.Labrum having the wide median arc armed with chitinous teeth.Claw of palpus of second maxillse smooth. No ventral pores.Coxal glands of last pediferous segment two on each side, thesesimple or homogeneous. Last ventral plate wide. Anal leg com-posed of a total of seven articles, the last ending in a well-developedclaw.Genotype.?Sogodes di^cUis, new species.In the presence of a claw on the anal legs resmbling Nyctunguvi,the species of which occur along the Californian coast, but differingin lacking ventral pores and in having the claw of the second maxillsesmooth. 36. SOGODES DIFFICILIS, new species.Plate 2, figs. 1 and 2.Color, pale fulvous. Head slightly wider than long; widest be-hind middle, from where narrowed to caudal end and also forward
;
anterior margin weakly angular; caudal margin wide, slightly in-curved (fig. 1). Antennae short, filiform; joints mostly short, espe-cially the more distal ones, excepting the ultimate, which equals thethree preceding ones in length. Prebasal plate exposed at middle.Basal plate four times as wide as long. Claws of prehensors, whenclosed, much short of attaining anterior margin of head; the pre-hensors being very short, the mesal margin of femuroid extendingbut little beyond margin of prosternum ; all joints unarmed. Pros-ternum unarmed; chitinous lines present, fine. First legs muchshorter than the second, the second but little shorter than the third.Dorsum bisulcate. Coxopleurae of last legs with two simple glandson each side, the anterior pore smaller than the posterior (fig. 2).Last ventral plate broad, narrowed caudad. Pairs of legs, 63.Leng-th, 13 mm.Locality.?Guatemala: San Rafael, one specimen, June 4, 1914(O.F.Cook).Type.?Cat No. 24125, U.S.N.M.3136?22?Proc.N.M.Vol.60 16
12 PROCEEDINGS OF THE NATIONAL, MUSEUM. vol.60.Family BALLOPHILIDAE.Genus ITYPHILUS Cook.37. ITYPHILUS CEIBANUS, new species.Plate 3, fig. 1.Distinguishable from /. guianensis Cliamberlin, of British Guiana,in the much larger number of pairs of legs, though the total num-ber of pairs of legs is not ascertainable because the caudal end ofthe type is missing. Sixty-nine pairs are now present in the incom-plete specimen, as against a total of from 49 to 55 in guianensis.From /. lilacinus Cook, of the Florida keys, the only other knownspecies, it differs in the form of the cephalic plate, this being longerthan wide instead of wider than long. The second tergite is broaderand rather shorter as compared with the first, these two beingslightly wider than the cephalic plate and obviously wider than theimmediately following tergites. The antennae more strongly clavateand flattened than in lilacinus, with the ultimate article less acumi-nate; strongly geniculate. Prosternum essentially as in lilacinus.Length of incomplete type (head+69 segments), 20 mm.Locality.?Honduras: La Ceiba (W. M. Mann).Type.?No. 24126, U.S.N.M.TANOPHILUS, new genus.Agrees with BallopMlus and differs from Ityphilus in having theventral pores in a definite, transversely elliptical area which is morestrongly chitinized and elevated above the general surface of thesternite. May be distinguished from BallopMlus and Prionothal-thyhius in having only a single large pit on each coxopleura of lastpediferous segment, several smaller pores opening into each pit.Genotype.?Tanophilus hondurasanus, new species.38. TANOPHILUS HONDURASANUS, new species.Plate 2, figs. 3 and 4.The general color in alcohol is ferruginous, but in life may havehad the dark-violet pigmentation typical for BaUophilus, Headwith no frontal suture, longer than wide, widest near middle, thesides and anterior margin convex, the caudal margin truncate. An-tennae strongly clavate, distally flattened, conspicuously geniculate(fig. 3). Basal plate narrowed caudad, two and a half times widerthan long. Prosternum with chitinous lines distinct and complete;anterior margin angularly excised. Prehensors small, not attainingthe front margin of the head. Dorsal plates strongly bisulcate,laterally somewhat roughened. Ventral pores beginning on firststernite. Last ventral plate strongly narrowed caudad. the sides
AUT.7. CENTIPEDS OF CENTRAL AMERICA CHAMBERLIN. 13
straight or nearly so, partly overlapping the large coxal pit on eachside. Anal legs strongly thickened proximally and conically nar-rowing distad, the last joint pointed and clawless (fig. 4). Pairs oflegs, 79.Length, 31 mm.Z(?caZ%.?Honduras : Cecilia, one specimen (W. M. Mann).Type.?Q^X. No. 24127, U.S.N.M.Family ORYIDAE.Genus ORPHNAEUS Meinert.89. ORPHNAEUS BREVILABIATUS (Newport).Geophilus brevilaUatus Newpobt, Trans. Linn. Soc. London, 1845, vol. 19p. 436.Orphnaeus hrasiliensis Meinert, Naturh Tidsskr., 1870-71, ser. 3, vol. 7.p. 20.Orphnaeus Uvidus Meinebt, Naturh. Tidsskr., 1870-71, ser. 3, vol. 7, p. 19.Orphaneus brevilabiatus Pocock, Biol. Centr. Amer., Chilopoda, 1896, p. 40.Localities.?Panama ; Nicaragua : Polvon, Occidental Department(McNeil) ; Honduras: La Ceiba, one specimen; Choloma, two speci-mens ; San Pedro, Sula, one specimen (W. M. Mann) ; Guatemala
:
Cacao, Trece Aguas (G. P. Goll), Patulul (W. M. Wheeler).Genus NOTIPHILIDES Latzel.
40. NOTIPHILIDES MAXIMILIANI (Humbert and Sanssure).Notiphilus maximiliani Humbert and Saussure, Rev. et Mag. Zool., 1870,ser. 2, vol. 22, p. 205.NotipMUdes maximiliani Latzel, Die Myriop. Osterr.?Ungar. Monarch.,1880, vol. 1, p. 20.Localities.?Costa Rica: San Mateo; Guatemala.Family CHILENOPHILIDAE.This group is used to include all genera formerly embraced in theGeophilidae which have a coxopleural suture present and conspicu-ously chitinized in the second maxillae. While in some genera ofthe group the coxae of the second maxillae are but weakly unitedwith each other, this character shows considerable variation anddoes not hold for most of the West Indian and Central Americanforms placed here by the author. Because of indications of transi-tion the group may ultimately prove untenable ; but pending furtherextension of our knowledge it is retained for its obvious convenience.SUTURODES, new genus.Head without frontal suture or this but vaguely indicated. An-tennae short, filiform. Basal plate wide, trapeziform, overlapped by
14 PROCEEDINGS OF THE NATIONAL MUSEUM. vol. 60.the cephalic plate. Labrum free, tripartite. Median piece large,protruding caudad, and armed with a series of long teeth. Lateralpieces fringed with pale spinescent processes. First maxillae withinner and outer branches distinctly set off from coxal plate. Outerbranch biarticulate, with two long membranous lappets. Secondmaxillae with coxae completel^y and broadly united at middle, as inNesidij)hilus^ though the median part is commonly more membra-nous. Coxopleural suture strongly chitinized, anteriorly curvingabout ectal edge of pore, where it may be only weakly indicated.Ventral sclerites strongly developed, extending far forward andunited at anterior end by a small median piece. Palpus triarticulate,terminating in a simple smooth claw; none of joints with a process.Prehensors large, exposed from above and extending beyond frontmargin of head. Claw and femuroid armed within and prosternumarmed anteriorl3\ Prosternum without chitinous lines. Ventralpores normally in four areas, one on each quarter of plate, as inNesidiphilus, etc. Last ventral plate narrow. Coxopleurae of lastlegs elongate, usually considerably exposed from above, and bearingvery numerous small pores. Anal legs with six joints beyond coxae,clawless.Genotype.?Suturodes tardus^ new species.This genus differs from the West Indian genus Nesidiphilus in theposition of the coxopleural suture, this in the latter genus alwaysending at the margin mesad of or caudomesad of the pore, as also itdoes in the closely related Telocriciis (pi. 4, fig. 5), whereas in thepresent group this suture curves about the ectal edge of the pore inthe more usual manner (pi. 3, fig. 4).
41. SUTURODES TARDUS, new species.Plate 2, fig. 5 ; plate 3, figs. 2, 3, and 4 ; plate 4, figs, 8 and 4.Flavous, the head and prehensors and a few anterior tergites typ-ically of a dilute chestnut tinge. Head about 1.68 times longer thanwide; widest at caudal end of frontal region, from where the sidesare parallel or converge but slightly caudad to the oblique caudalcorners and more strongly forward. iVnterior margin a little con-vex at middle. Caudal margin straight (pi. 4, fig. 3). Articles ofantennae moderately long, the ultimate clearly shorter than the twopreceding taken together. No distinct clypeal areas excepting thesmall, pale spots from which the principal, more median setae arise.Ectad of each of these setae of the median pair a series of three orfour setae and in front of them and between the antennae a secondpair. Caudad of the main transverse series of setae a second seriesof smaller setae, these 3+3 or 3-f 4. Caudad of this second seriestwo setae on the median line, one behind the other, with sometimes
-ART. 7. CENTIPEDS OF CENTBAL AMERICA?CHAMBERLIN. 15
a much reduced setae each side of the first of these. Median pieceof labrum i3rojecting well caudad of the lateral pieces ; bearing typ-ically six long teeth (pi. 4, fig. 4). Coxal plate of first maxillaewith a pair of long setae at anterior border, and three or four lessee-setae. Setae of coxae of second maxillae few, consisting typicallyof a submarginal series of four on each side and a group of four atmiddle. For other features see plate 3, figure 4. Exposed part ofbasal plate about four times as broad as long. (In the holotype thecephalic plate has been shifted abnormally forward.) Claws of pre-hensors when closed extending beyond distal end of first antennalarticle, but not attaining the distal end of the second. Tooth at baseof claw black, conical, that of the femuroid similar but larger. Teethof prosternum short, blunt. See further plate 3, figure 2. Dorsalplates distinctly bisulcate, the paired sulci also traversing the basalplate. ^Vnterior ventral plates w^ith a deep, median longitudinalsulcus. First spiracle large, vertically oval, the second of similarshape and but little smaller. The third more abruptly reduced, thesucceeding ones soon becoming circular and small. Last dorsal platenarrow and long, its sides nearly parallel (pi. 2, fig. 5). Last ven-tral plate very narrow, its sides a little converging caudad, towardcaudal end rounding in to caudal margin. Coxopleural pores smalland very numerous ventrally, laterally, and dorsally (pi. 3, fig. 3).Pairs of legs, 69 to 71.Length, 42 mm.Locality.?Honduras: San Juan Pueblo, two specimens (W. M.Mann).Type.?C^i. No. 24128, U.S.N.M.
42. SUTURODES GUATEMALAE. new species.Plate 3, fig. 5 ; plate 4, figs. 1 and 2.Resembling S. stolli (Pocock) in general. Frontal plate not dis-crete, but vaguely indicated in part only as a pale line. Headwidest near level of this line, the sides moderately converging caudadinstead of being parallel. The anterior corners are not widelyrounded, as represented in the figure of S. stolli, but are a littleobtusely angular, the anterior margin of the head being in the formof a very obtuse angle. The length of the exposed portion of thebasal plate is much less than half its anterior width, being over-lapped by head and by first tergite; it bears a single transverseseries of setae. It differs from stolli, according to notes on thelatter given by Attems,^ in having two small clypeal areas, eachbearing a seta instead of a single nonsetigerous area. In frontof the two setae of these areas in the present species is a second pair
I Schulze, Porschung.sr. in West, u. Zentr. SUdafrilta, vol. 2, Abt. 1, p. 35.
16 PROCEEDINGS OF THE NATIONAL, MUSEUM. vol.60.
of setae, and caudad and laterad of them are four or five setae oneach side in two series, the total number of setae being thus 12 or13; all are very short (pi. 4, fig. 1). Median piece of labrum large,bearing six teeth. Coxae of second maxillae broadly united, poresopening through mesal edge. The chitinous coxopleural suture isstrongly marked in its caudal half, becoming weaker anteriorlywhere it curves around the ectal side of the pore. The lappets ofthe first maxillae are long and conspicuous, but that of the secondjoint is smaller proportionately to that of the first than in tardus.Anterior margin of presternum with two reduced, almost obsoleteteeth. Femuroid of prehensors armed within at distal end with arounded tooth, claw at base with a dark, conical tooth. Otherjoints unarmed (pi, 4, fig, 2), First spiracle vertically elliptic,much larger than the second. The second and following spiraclescircular. Last ventral plate narroAv; sides converge caudad, morestrongly so toward caudal ends; caudal margin straight. Coxo-pleural pores numerous, but fewer above than in tardus. Last dorsalplate broader than in the latter species (pi. 3, fig. 5). Anal legsin male but little thickened; with numerous very short hairs onventral surface of proximal joints. Pairs of legs (male), 59,Length, about 21 mm.Locality?Guatemala: San Rafael, June 4, 1914. (O. F. Cook),Type?1^0. 24129, U,S,N.M.
43. SUTURODES STOLLI (Pocock).Geopliilvs stolli Pocock, Biol. Centr. Amer. Chilopoda, 1896, p. 38, pi. 3,figs. 9-9c.Pachymeriuni stolli Attems, Scluilze Forscliungsr. im West. u. Zentr.Siidafrika, 1909, vol. 2, Abt. 1, p. 35, fig, 5,This species is listed under Suturodes with but little doubt, al-though I have not seen specimens of the form. Attems places thespecies in Pachymeinum, of the Geophilidae proper; but his figureshows a coxopleural suture as developed posteriorly. This suggeststhe condition in the preceding species, in which the posterior partof the suture is strongly chitinous and conspicuous, wliile the anteriorpart is weaker and quite likely to be overlooked in the unmountedmaxillae. EXPLANATION 0? PLATES.The plates were drawn by the author.Plate 1.Neivportia divergcns.Fig. 1. Head and first segments, dorsal viewX 17.Cryptops micrus.2. Last leg, mesal viewX 65.
ABT.7. CENTIPEDS OF CENTRAL AMERICA CHAMBERLIN. 17Newportia sulana.3. Distal eud of tarsus of last legX 65.Scutigera nubila.4. Caudal portion of fifth tergiteX45.5. Last tergite and caudal border of preceding one, in outlineX45,Lahrohius cobulcanus.6. Anterior portion of prosternumX65.7. Basal joint and spines of left genital forceps of female, ventral viewX 65.Plate 2.Sogodes difflcilis.Fig. 1. Head, dorsal viewX 48.2. Glands of left coxopleura of anal legs, in outlineX 220.Tanophilus hondurasanus,3. Head and first tergite, dorsay viewX 54.4. Caudal end of body, ventral viewX 56.Suturodes tardus.5. Caudal end, dorsal viewX 38.Plate 3.Ityphilus ceihanus.Fig. 1. Head and first plateX 76.Suturodes tardus. ,2. Presternum and prehensorsX^S.3. Caudal end, ventral viewX 33.4. Maxillae, the right palpus of second pair omittedX 63.Suturodes guatemalae.5. Caudal end, dorsal view. PlATE 4.Suturodes guatemalae.FtG. 1. Anterior portion of head, with maxillae removed, ventral view X 49.2. Prosterniim and prehensorsX22.Suturodes tardus.3. Head and adjacent tergites, dorsal viewX15.4. LabrumX220. Telocricus marginalis.5. Coxosternum of second maxillae, showing coxopleural line, etc. X63.

U. S. NATIONAL MUSEUM PROCEEDINGS. VOL. 60, ART. 7 PL. I
Centipeds of Central America.For fxplanatjon of plate see pages 16 and 17.
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Centipeds of Central AmericaFor explanation of plate see page 17.
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Centipeds of Central AmericaFor explanation of plate see paoe 17.
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Centipeds OF Central America
-OR EXPLANATION OF PLATE SEE PAGE 17