The Type-Material of North American Clearwing Moths (Lepidoptera: Sesiidae) W. DONALD DUCKWORTH and THOMAS D. EICHLIN m wtu. SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY ? NUMBER 148 SERIAL P U B L I C A T I O N S OF T H E S M I T H S O N I A N INSTITUTION The emphasis upon publications as a means of diffusing knowledge was expressed by the first Secretary of the Smithsonian Institution. In his formal plan for the Insti- tution, Joseph Henry articulated a program that included the following statement: "It is proposed to publish a series of reports, giving an account of the new discoveries in science, and of the changes made from year to year in all branches of knowledge." This keynote of basic research has been adhered to over the years in the issuance of thousands of titles in serial publications under the Smithsonian imprint, com- mencing with Smithsonian Contributions to Knowledge in 1848 and continuing with the following active series: Smithsonian Annals of Flight Smithsonian Contributions to Anthropology Smithsonian Contributions to Astrophysics Smithsonian Contributions to Botany Smithsonian Contributions to the Earth Sciences Smithsonian Contributions to Paleobiology Smithsonian Contributions to Zoology Smithsonian Studies in History and Technology In these series, the Institution publishes original articles and monographs dealing with the research and collections of its several museums and offices and of professional colleagues at other institutions of learning. These papers report newly acquired facts, synoptic interpretations of data, or original theory in specialized fields. These pub- lications are distributed by mailing lists to libraries, laboratories, and other interested institutions and specialists throughout the world. Individual copies may be obtained from the Smithsonian Institution Press as long as stocks are available. S. DILLON RIPLEY Secretary Smithsonian Institution S M I T H S O N I A N C O N T R I B U T I O N S T O Z O O L O G Y ? N U M B E R 1 4 8 The Type-Material of North American Clearwing Moths (Lepidoptera: Sesiidae) W. Donald Duckworth and Thomas D. Eichlin SMITHSONIAN INSTITUTION PRESS City of Washington 1973 ABSTRACT Duckworth, W. Donald, and Eichlin, Thomas D. The Type-Material of North American Clearwing Moths (Lepidoptera: Sesiidae). Smithsonian Contributions to Zoology, number 148, 34 pages, 1973.?The type-material of North American clearwing moths of the family Sesiidae is reviewed by bringing together all the available data lor the 255 species group names applied to this fauna area. Of these 255 names, 72 required lectotype designations, 152 were described from single specimens, and the remaining 31 are lacking in sufficient data to determine the number of specimens included in the original type series. The names are arranged alphabetically by specific name preceded by the genus in which it was originally described. The following information sequence is presented for each name: original combination; reference to original description; pertinent com- ments from the original description concerning the number of specimens, sex, locality, collection, and source of the types; exact label data on types, their present location, and number of syntypes examined; and subsequent pertinent comments and actions. OFFICIAL PI'BLICATION DATE is handstamped in a limited number of initial copies and is re- corded in the Institution's annual report, Smithsonian Year. SI PRESS NUMBER 4778. SERIES COVER DESIGN: The coral Montaslrea cai Douglas Co., Kansas, 900 feet, Prof. Snow." HOLOTYPE: Unknown. 12. Cissuvora ampelopsis Engelhardt, 1946:134 "Type.?U.S.N.M., No. 56843. From Victoria, Tex." "Remarks.?Described from male type, female allotype, 3 male and 3 female paratypes from the type locality; 4 male and 10 female paratypes from San Antonio, Tex." SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY HOLOTYPE: Male, in the NMNH: "Paranthrene ampelopsis Engelhardt $ , Holotype"; "U.S.N.M. Type No. 24866,"; "from Cissus incisa"; "J. D. Mitchell, Collector"; "Victoria, Tex., 6-21-19"; "on Cissus incisa"; "Hunter, No. 3918." DISCUSSION: The catalog of USNM type numbers indicates that this species was entered twice. The erroneously recorded 24866 was canceled from the catalog in February, 1944. 13. Pyrrhotaenia animosa Hy. Edwards, 1883:156 "4 5 . 2 $ . Arizona. Coll. B. Neumoegen and G. H. French." SYNTYPES EXAMINED: 2 (both in NMNH): (1) "Pyrrohotaenia animosa Hy. Ed., $ , Type"; "Typi- cum specimen"; "TYPE"; "Arizona" (male). (2) "Pyrrhotaenia animosa Hy. Edw., 9, Type"; "Typicum specimen"; "TYPE"; "Arizona" (fe- male). DISCUSSION: The other four specimens supposed to be in the type series were not found. The male syntype in the NMNH is in good condition and has been selected, labeled, and designated as the lectotype. 14. Sesia anthracipennis Boisduval, 1874:392 "Elle se trouve en Georgia ou, selon Abbot, elle vir dans une espece de Salix. Nous l'avons recue de John Leconte." HOLOTYPE: Female, in the NMNH: "Type, an- thracipennis Bdv., a/c Hofer"; "Sesia anthracipen- nis Bdv., Amer. bor., Sp. G. Sphing. 392"; "Oberthur Collection"; "j'ai envoys en Jly 1882 la peinture a M. Edwards, Ch. Ob."; "Typicum Speci- men"; "Ex Musaeo, Dris Boisduval"; "anthracipen- nis, Am. B." 15. Carmenta apache Engelhardt, 1946:54 "Types?U.S.N.M. No. 56830, female." "Remarks.?Described from three specimens, a perfect female and two imperfect males, the latter not designated as paratypes, in all probability rep- resenting the same species, but without knowledge of the food plant and habits the evidence is not conclusive. The three specimens are from Prescott, Ariz., the males collected by H. Dyar, August 20, 1917, and the female from the Barnes collection, dated July 1-7." HOLOTYPE: Female, in NMNH: "Conopia apache Engel. 9 "; "Fig."; "Type No., U.S.N.M."; "Barnes Coll."; "Prescott, Ariz."; "July, 1-7." 16. Sphinx apiformis Clerck, 1759: pi. 9, fig. 2 TYPE: Lost (Naumann, 1971). DISCUSSION: The original description is based on a figure. Apparently the location of the specimen or series from which the drawing was made is un- known and may in fact be lost or destroyed. The species was redescribed by Linnaeus in 1761, and according to Naumann (1971), there is a syntype in the Linnean Society London labeled "apiformis" in Linne's own handwriting. 17. Sesia arctica Beutenmuller, 1900:208 "Habitat.?Kodiak, Alaska, July 20th, 1899." "Type.?One male, No. 5175, Coll. U. S. Nat. Mus.; collected by Mr. Trevor Kincaid. . . ." HOLOTYPE: Male, in the NMNH: "Sesia arctica Beut., Type, <} "; "U. S. N. M. Type No. 5175"; " s genitalia on slide, AB, Dec. 2, 1938." 18. Sesia arizonae Beutenmuller, 1898:240 "1 $ , Summit of Mt. Union, 9,000 feet, Arizona, July 3, 1887, flying about scrub oak (G. D. Hulst). Coll. Hy. Edwards." "1 9, Texas. Col. U. S. Nat. Mus." SYNTYPES EXAMINED: 2 (1 in AMNH, 1 in NMNH): (1) "Sesia arizonae Beut., $, Type"; "Ariz."; "Summit of Mt. Union, about scrub oak, July 3.87, alt. 9000 ft."; "No. 15871, Collection Hy. Edwards"; "Genitalia mounted on slide No. 0010" (female in AMNH). (2) "Sesia arizonae Beut., $ , Type"; "U.S.N.M. Type No. 4355"; "Tex."; "TDE Slide No. 76033" (female in NMNH). DISCUSSION: The female syntype in the NMNH is not conspecific with the female syntype in the AMNH, based particularly on the genitalia. The NMNH syntype is conspecific with refulgens Hy. Edwards. Therefore, the female syntype in the AMNH has been selected, labeled, and designated as the lectotype. 19. Gaea arizonesis Beutenmuller, 1916:372 "Habitat.?Pinal Mts., Arizona." "Described from a single female. Type: collec- tion Dr. William Barnes." HOLOTYPE: Female, in the NMNH: "Gaea ari- zonensis Beutm., Type, 9 "; "TYPE" "Pinal Mts., Ariz." 20. Albuna artemisiae Hy. Edwards, 1881:187 "1 s . Sier. Nev. Calif. (H. E.) on flowers of Artemisia vulgaris. L." NUMBER 148 HOLOTYPE: Male, in the AMNH: "Albuna arte- misiae Hy. Ed., Type"; "5041, Sierra Nev., Cali."; "No. 15875, Collection Hy. Edw."; "mellinipennis Bdv., det. G. P. E." 21. Sesia asilipennis Boisduval, 1829:496 "3a. Sa chenille, 3b. Sa crysalide.?Hab. l'Am?- rique septentrionale." HOLOTYPE: Male, in the NMNH: "Type, asili- pennis Bdv., a/c Hofer"; "Sesia asilipennis Bdv., Amer. bor., Sp. G. Sphing. p. 391"; "Sesia asilipen- nis Bdv., Sp. general, page 391"; "Oberthur Collec- tion"; "j'ai envoye en Jly 1882 la peinture a M. Edwards, Ch. Ob."; "Typicum Specimen"; "Ex. Musaeo, Dris Boisduval"; "Am. B." 22. Aegeria aureola Hy. Edwards, 1881:194 "1 $. Nevada. (Morrison.)" "Type. Coll. E. L. Graef." HOLOTYPE: Female, in the NMNH: "Aegeria aureola Hy. Ed., Type"; "TYPE"; "Col., E. L. Graef"; "Nev." (abdomen missing). 23. Aegeria aureopurpura Hy. Edwards, 1880:72 "Expanse, 0.60 inch. Texas, J. Boll." "Type, coll. Mus. Comp. Zool., Cambridge." HOLOTYPE: Female, in the MCZ: "Aegeria (?) aureopurpura Hy. Edw., (Type)"; "Dallas, Tex., Boll"; "Type 928." 24. Synanthedon auritincta Engelhardt, 1925c:216 "Habitat: Baboquivari Mts., Pima Co., Arizona, August 1-15, 1923 and 1924. O. C. Poling, collec- tor, Described from eleven specimens, two males and nine females, kindly submitted for determina- tion by Dr. Barnes and Mr. Benjamin." "Type, female; allotype, male, and six paratypes, females, William Barnes Collection; one paratype, male and two paratypes, females, Geo. P. Engel- hardt Collection at the Brooklyn Museum." HOLOTYPE: Female, in the NMNH: "Synanthe- don auritincta G.P.E. $, Type"; "Fig."; "14"; "Baboquivari Mts., Pima Co., Ariz., 1-15 Aug. 1923, O. C. Poling, Coll." 25. Carmenta austini Engelhardt, 1946:57 "Type.?U.S.N.M. No. 56832." "Remarks.?Represented only by the types; the male holotype captured inside a window, Biologi- cal Department, University of Texas, Austin, Tex., November 4, 1922 (H. B. Park), and the female allotype collected at Ephraim, Sanpete County, Utah, September 29, 1929 (H. B. Park)." HOLOTYPE: Male, in NMNH: "USNM Type No. 56832, austini Engelh."; "austini, $ genitalia on slide, AB, Jan. 22, 1940"; "Fig."; "Chamaesphecia austini Engelh. s "', "Chamaesphecia giliae austini Engelh. $ "; "GPEngelhardt Coll."; "H. B. Parks, Jr., Collector"; "Austin, Tex., 11-9-32." DISCUSSION: There is a discrepancy as to the date given in the publication and the date on the label. There is little doubt that the male in the NMNH is the actual type specimen, and an error in tran- scribing the date must have occurred at some point. 26. Alcathoe autumnalis Engelhardt, 1946:105 "Type.?U.S.N.M. No. 56840. From San An- tonio, Tex." "Remarks.?Described from the male type and 31 male and 27 female paratypes all from San An- tonio, Tex." HOLOTYPE: Male in NMNH; "Alcathoe au- tumnalis Engelh. s , Type"; "Coll. G. P. Engel- hardt"; "San Antonio, Tex., X.20.1930." 27. Melittia barnesi Dalla Torre, 1925:138, new name pro M. superba B. & L., 1922 HOLOTYPE: Ipso facto type of superba B. & L. 28. Aegeria bassiformis Walker, 1856:39 "a. United States. Presented by E. Doubleday, Esq." HOLOTYPE: Male, in the BMNH: "64, Aegeria bassiformis"; "46.110, U.S."; "Type." 29. Melittia beckeri Druce, 1892:276 "Expanse I1/2 inch." "Hab. Mexico, near Durango city (Becker)." HOLOTYPE: Male, in the BMNH: "B.C.A. Lep. Het. Beckeri Druce"; "Near Durango city, Mexico, Becker"; "Godman-Salvin Coll. 97.-52"; "Type, Sp. figured"; "Type." 30. Pyrrohotaenia behrensii Hy. Edwards, 1882c: 123 "2 i Soda Springs, Shasta Co., Calif. (J. Beh- rens)." SYNTYPES EXAMINED: 2 (both in AMNH): (1) "Pyrrhotaenia behrensii Hy. Edw. Type"; "Soda Spr., June 16"; "No. 15958, Collection Hy. Edw." (male). (2) "Pyrrhotaenia behrensii Hy. Edw."; "Soda Spr., June 16"; "No. 15960, Collection Hy. Edw." (male). DISCUSSION: The first male syntype in the AMNH listed above has been selected, labeled, and desig- nated as the lectotype. SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY 31. Albuna beutenmulleri Skinner, 1903:126 " 2 ? . . Described from one specimen taken at Stockton, Utah, May 24, 1902, by Mr. Thomas Spalding." HOLOTYPE: Female, in the Academy of Natural Sciences of Philadelphia; "A. beutenmulleri Skin- ner, TYPE"; "Stockton, Utah, V.24.02, T. Spald- ing"; "Type No. 7077, Albuna beutenmulleri Henry Skinner." 32. Sesia bibionipennis Boisduval, 1869:64 HOLOTYPE: Male, in the NMNH: "Type, bibio- nipennis Bdv., a/c Hofer."; "Sesia bibionipennis, Bdv. Sp.G. Sphing, California, p.421"; "bibioni- pennis, Calif."; "Oberthur Collection"; "Typicum specimen"; "Ex. Musaeo, Dris Boisduval"; "j'ai envoye en Jly 1882, La peinture a M. Edwards, Ch. Ober." DISCUSSION: Engelhardt (1946) states, "The spe- cific name bibionipennis Boisduval, 1869, has not been used in earlier check lists because of insuf- ficient description and the supposed loss of the type. This type, however, has been found in a part of the Oberthur Collection, acquired by Wil- liam Barnes, and is now at the United States Na- tional Museum." 33. Aegeria bolli Hy. Edwards, 1881:191 "1 $ . Texas. (J. Boll.)" "Type. Coll. B. Neumoegen." HOLOTYPE: Male, in the NMNH: "Aegeria bolli Hy. Edw., Type"; "TYPE"; "Typicum specimen"; "Tex."; "bolli, Type HE, $ genitalia on slide, AB, fan. 10, 1937." 34. Aegeria bolteri Hy. Edwards, 1883:155 "1. i N. Illinois. Collected by my friend, Mr. A. Bolter, to whom I dedicate the species. Type Coll. Hy. Edwards." HOLOTYPE: Female, in the AMNH: "Aegeria bolteri Hy. Edw., Type"; "N. 111."; "No. 15894, Collection, Hy. Edw." DISCUSSION: The type is a female and not a male as indicated by Edwards in the original descrip- tion, an error he has made more than once. 35. Tarsa bombyciformis Walker, 1856:61 "Male." "a. ?" HOLOTYPE: Male, in the BMNH: "Tarsa bomby- ciformis Wlk. Type, ? "; "1., Tarsa bombycifor- mis"; "Type"; "F3/13." 36. Penstemonia brevifolia Engelhardt, 1946:21 "Type.?U.S.N.M. No. 56825. Holotype male and allotype female; one male and one female para- types. Collected in the Green Horn Mountains, Calif." HOLOTYPE: Male, in the NMNH: "Penstemoni brevifloris Engelh., <$ "; "USNM Type No. 56825, brevifolia Engelh." "Out 8-28-39, $ , 1-Greenhorn Mts., 7-9-39, P. brevifl."; " $ genitalia on slide. May 10, 1940, J.F.G.C, 2963." 37. Aegeria brunneipennis Hy. Edwards, 1881:191 "1 $ . Georgia. (Morrison.)" "Type. Coll. F. Tepper." HOLOTYPE: Female, in MSU: "Aegeria brunnei- pennis Hy. Edw., Type"; "Ga."; "F. T." (abdomen missing). 38. Sesia brunneri Busck, 1914:143 "Foodplant: Pinus ponderosa." "Habitat: Camas, Montana, Josef Brunner, col- lector." "Type: Cat. No. 18238, U.S.N.M." SNYTYPES EXAMINED: 2 (both in NMNH): (1) "Sesia brunneri Busck, 2 , Type"; "U.S.N.M. Type No. 18238"; "Camas, Mont., P. ponderosa"; "on Pinus ponderosa, Camas, Mont., Josef Brunner"; "11530a" (male). (2) "Sesia brunneri Busck, Type" "Wissadula lozani"; "Brownsville, Tex."; "Diven, Coir."; "reared, May 7/19"; "Diven, 41, F.H.B., rearing." 254. Pyrrhotaenia wittfeldii Hy. Edwards, 1883:156 "2. i Indian River, Florida. Dr. Wittfield." "Type. Coll. Hy. Edwards." SYNTYPES EXAMINED: 2 (both in AMNH): (1) "Pyrrhotaenia wittfeldii Hy. Edw., Type"; "Indian River, Florida"; "No. 15985, Collection Hy. Edw." 32 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY (female). (2) "Pyrrhotaenia wittfeldii Hy. Edw."; "Indian River, Florida"; "No. 15986, Collection Hy. Edw." (female). DISCUSSION: It is assumed here that Edwards erred in the determination of the sex of the types. The first female syntype in the AMNH labeled type has been selected, labeled, and is presently designated as the lectotype. 255. Sesia xiphiaeformis Boisduval, 1874:409 "Nous avons recu cette remarquable Sesie de feu John Leconte, sans aucune indication sur la lo- calite qu'elle habite aux ?tats-Unis." HOLOTYPE: Female, in the NMNH: "Type xiphiaeformis Bdv., a/c Hofer"; "Sesia xiphiae- pennis Bdv., Amer. bor., Sp. G. Sphing. p. 409"; "Oberthur Collection"; "j'ai envoye" en Jly 1882 la peinture a M. Edwards, Ch. Ob."; "Typicum Speci- men"; "Ex Musaeo, Dris Boisduval"; "xiphiaefor- mis Bdv., Am. B."; "Barnes Collection"; "Synan- thedon exitiosa Say, $, det. G.P.E." Literature Cited Barnes, Wm., and F. H. Benjamin 1925. Change of a Preoccupied Name (Lepidoptera: Aegeriidae). Proceedings of the Entomological So- ciety of Washington, 27(1): 14. Barnes, Wm., and A. W. Lindsey 1922. Descriptions of Two New Species of Aegeriidae (Lep.). Bulletin of the Brooklyn Entomological Society, 18 (4): 122-123. Behrens, James 1889. In French, Some Texas, Arizona and California Moths. Canadian Entomologist, 21 (9): 161-163. Beutenmiiller, William 1893. Notes on Some North American Moths, with De- scriptions of New Species. Bulletin of the Ameri- can Museum of Natural History, 5:19-26. 1894. Studies of Some Species of North American Moths. Bulletin of the American Museum of Natural His- tory, 6:87-89. 1896. Critical Review of the Sesiidae Found in Amer- ca. North of Mexico. 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On the Insects, Colopterous, Hymenopterous and Dipterous, Inhabiting the Galls of Certain Species of Willows. Part 2nd and Last. Proceedings of the Entomological Society of Philadelphia, 6:223-288. Westwood, John O. 1848. The Cabinet of Oriental Entomology. Pages 1-88. London. 1854. The American Oak-gall Clear-Winged Sphinx. Gardener's Chronicle and Agricultural Gazette, 47: 757-758. ?Ct U.S. GOVERNMENT PRINTING OFFICE; 1973 818-820, II Publication in Smithsonian Contributions to Zoology Manuscripts for serial publications are accepted by the Smithsonian Institution Press, sub- ject to substantive review, only through departments of the various Smithsonian museums. Non- Smithsonian authors should address inquiries to the appropriate department. If submission is invited, the following format requirements of the Press will govern the preparation of copy. 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