SMITHSONIAN CONTRIBUTIONS TO BOTANY NUMBER 22 Vascular Flora of the Northern Marianas Islands F. Raymond Fosberg, M. V. C. Falanruw, and Marie- Helene Sachet SMITHSONIAN INSTITUTION PRESS City of Washington 1975 ABSTRACT Fosberg, F. Raymond, M. V. C. Falanruw, and Marie-Helene Sachet. Vascular Flora of the Northern Marianas Islands. Smithsonian Contributions to Botany, number 22, 45 pages, 2 figures, 1975.--1-\ systematic catalog of the vascular plants known from the northern 10 islands in the Marianas Archipelago in Micronesia, with a short geographical introduction and, for each species, ecological and geographic data and specimen citations, or, where these are unavailable, citations of literature records. OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is recorded in the Institution's annual report, Srnithsonian Year. SI PRESS Nmmm 5217. SWES COVER DESIGN: Leaf clearing from the katsura tree Cercidiphyllurn japonicum Siebold and Zuccarini. Library of Congress Cataloging in Publication Data Fosberg, Francis Raymond, 1908- Vascular flora of the Northern Marianas Islands. (Smithsonian contributions to botany, no. 22) Bibliography: p. Includes index. Supt. of Docs. no.: SI 1.29:22 1. Botany-Mariana Islands. I. Falanruw, Marjorie V. C., joint author. 11. Sachet, Marie- Hblhne, joint author. 111. Title. IV. Series: Smithsonian Institution. Smithsonian contribu- tions to botany, no. 22. QKlS2747 no. 22 [QK473.M34] 581'.08s [581.9'96'7] 74-20631 Con tents Page Introduction ........................................................ 1 Annotated List of the Vascular Flora .................................. 6 References .......................................................... 43 Index to Families ................................................... 45 iii Vascular Flora of the Northern Marianas Islands F. Raymond Fosberg, M. V. C. Falanruw, and Marie-Helene Sachet Introduction Micronesia (Figure 1) consists of a group of archipelagoes in the western tropical Pacific Ocean, all made up of very small islands (hence the name Micronesia). With a few exceptions, they are all in the northern hemisphere. One of the archipela- goes to the west is the long north-south chain of the Marianas Islands. To date there is no flora of the Marianas, let alone of the smaller northern islands of the group, but enough information has now accumulated to make such a flora meaningful, phytogeographically. Very considerable scientific interest has developed in recent years concerning these small islands. They, especially the unin- habited ones, form excellent, somewhat inacessi- ble, natural ecological and evolutionary labora- tories. Several expeditions have been organized by members of the Biology Department and Marine Laboratory of the University of Guam. One of the authors of this paper, Mrs. Falanruw, heads a project for interpretation of vegetation changes in these islands from satellite and aerial imagery. The U. S. Navy is about to file an environmental im- pact statement for a project to use Farallon de Medinilla as a target range. Hence, and as a part of the vegetation interpre- tation project mentioned above, we have brought F. Raymond Fosberg and Marie-Helenee Sachet, Department of Botany, National Museum of h?atural History, Smithsonian Institution, Washington, D.C. 20560. M. V. C. Falanrum, Yap Institute of Natural Science, Caroline Islands 96943. together the information available to us and put it into the form of a systematic census of the vascu- lar plants of the islands north of Saipan. The paper, therefore, serves as a record of ?ground truth? for the Trust Territory of the Pacific Islands EROS Project: Vegetation Changes in the Northern Mari- anas. Additional interpretation of the vegetation of the Northern Marianas is planned using aerial and orbital imagery. Because two of the authors of this flora are en- gaged in preparing a major flora of Micronesia, which will have full synonymies, keys, descriptions, and literature citations, these features are not in- cluded in the present work. Verified names, eco- logical and distributional information, and specimen citations are provided. We feel that the distribution by islands is reasonably complete, though careful exploration may fill this in and even reveal new species. One, Terminalia rostrata, was found on Asuncion Island in 1972. Medinilla, the smallest island, is also by far the least bo- tanically well known. Much information on many of these species may be found in Stone?s compre- hensive Flora of Guam (1971). Guam is the south- ernmost and largest of the Marianas. The Northern Marianas, the chain of islands extending about 480 km from Medinilla north to Uracas (Figure 2), are, with the exception of tiny Medinilla, active or dormant volcanoes, Anatahan may be extinct, but probably is only dormant. Medinilla is a flat-topped limestone remnant about 0.85 sq km in area and 30 m high. Geologically it might perhaps better be associated with the 1 130' 135' I400 145' 150' 30' g* b RYUKYU * 0 100 200 300 400 500 Statute Miles I I I I I I BONIN 1 s L A N DS kbk inaw a -jima ISLANDS Ha ha -jimd *'Da i tb-jima Ki to-io-jima. _L Iwo Jima, VOLCANO ISLANDS 2S0 -- -Minami-i6- jima / c \ / Marcus. . 200 .15' PHIL1 PPlNE ISLANDS c / / A- / f / . Parece Vela I I I I / I I I I I I 6Uracas +aug Asuncion * Agrigan MARIANAS rPaaan Alamagan: - ISLANDS Griguan Y Sa r i pa n Saipon Agiguan An at ahan' .Med i ni I la Tin i a n$ 'Rota #Guam Santa Rosa Reef I550 160' 165' 170' 175' FIGURE I.-Micronesia. \ ig Wake \ ZPokak I I I I Aur ISLANDS I I I I I ,.,Orolu k Pakin Namu ''. ..: Jabwot ':.UPONAPE Majuro Ant A i I i n g I a pa I a p:' q., .::.*Ar no Mokil :.pingelap Jaluit "' hJg a t i k Namor ik" ''~i I i E boi OKUSAIE P+ i... ,, ,,Little Makin Nukuoro % GILBERT BUtarltarl(M&in) 1 '! Ka pi ngamarangi 0, ranu ka I I .Nauru Banaba- Nonouti:,, Beru (Oc?an) Tabiteuea\.> ' onotoo Nikunau\ Arorai I -- -- -z - -- -- --- -/ -- SOLOMON E LLICE IS LANDS 165' 170' 1750% ~ I I I 1 1800 3 0' 250 2 00. 15" 100. 50- 0 O- 5O. 180? 4 SMITHSONIAN CONTRIBUTIONS TO BOTANY E 145' NORTHERN MARIANAS ... ,. .. .. .'. '.. '. ,:. . A I am aga n: ::Q< ; , .. I ........ ....... .. ... ..... Gugua,n@;:.::. . ,. . . ,, ........ .. Sa rig an ::p,': I :. .... .. , ...... . ',:,. .......... '.:'. . ::.a!Anata han ......... ........ ,, ,' FIGURE 2.-The Northern Marianas. (One degree of latitude = 60 nautical miles = 111 km.) NUMBER 22 5 Southern Marianas. The others are both larger and higher, and limestone is only known from a very small area on Pagan. Otherwise the substratum is entirely of volcanic origin. The limitation in area and in substratum types is, at least in part, responsible for a limited flora. Isolation, and the extreme geological youth of at least the volcanic islands, exert an influence in the same direction, toward a small flora. A detailed history of botanical exploration will be reserved for the Micronesian flora. Only those responsible for the records and specimens cited here will be mentioned. Doubtless many of the early voyagers sighted, perhaps even visited, some 01 the 10 islands treated here. La Perouse (1797), who visited the islands in 1786, reported stunted coconuts on Asuncion at that time. Captain Beechey (1831, II:438), in H. M. S. Blossom, visited Asuncion in 1827 and gives a very contrasting ac- count from that of La Perouse. Whether any plants were collected we have not found out. None are reported in the botany of the voyage. De la Corte (1926, 1927) wrote an account mentioning some plants by their Chamorro names. The first collec- tions of herbarium specimens were probably those made in 1887 by Marche (1891). Governor Fritz, administrator under the German regime, made a trip visiting all of the islands in 1901, making some botanical observations and planting a number of useful plants (Fritz, 1902). Few of these persisted, but a number of weedy species may have been carried along by accident and established them- selves. The first significant collections were made by the Japanese, especially Professors Ryozo Kanehira and Takahide Hosokawa, during the 1930s. They pub- lished informal accounts with some listing of col- lections (Kanehira, 1934; Hosokawa, 1934) and cited these plants in appropriate taxonomic treat- ments. These records from the literature, or the specimens if we have been able to see them, are listed under the species below. More recent collections were made by Donald Anderson (Pacific Vegetation Project) in 1949; F. R. Fosberg (Pacific Vegetation Project) in 1950; L. D. Bonham and G. W. Corwin (U. S. Geological Survey) in 1954; Michael Evans (Pacific Vegetation Project) in 1966; M. Villa-Gomez (University of Guam) in 1970; Philip Moore (Guam Public Schools) in 1971. M. V. C. Falanruw (University of Guam and Smithsonian Institution), in 1970 and 1972, made collections from the motor sailing ship Wanderer (Robert Hervin, Master). The best sets of these recent collections are or will be deposited at the Smithsonian Institution (U. S. National Herbarium). The Japanese collec- tions are in the Kyushu and Taipei University Herbaria, with some duplicates in various U. S. and European herbaria. We have been able to study part of them. Recently two other sources, not documented by specimens, have become available and are included here especially because they provide more informa- Lion than hitherto available on the plants of Medinilla. They are an excerpt from a manuscript draft environmental impact statement, by Swed- berg, on Medinilla, mentioning a few plants, and a booklet on the Marianas by Lehne and Gabler (1972), mentioning some plants by Chamorro or German names. They include five names for which we have been unable to find botanical equivalents: Admahaian, Dumpkin, Kurbisse, Nok, Paypayge- busch. The principal botanical papers on or including Northern Marianas plants are listed in the litera- ture cited at the end of this account. Special men- tion may be made of four of these papers, not cited otherwise, but dealing with aspects of North- ern Marianas botany: Corwin et al., 1957; Fosberg, 1958; Fosberg and Corwin, 1958; and Fosberg, 1960. The floristic information in the present account has been accumulated in our Micronesian flora card files by Fosberg and Sachet over a period of many years. It was compiled from this source and from specimen labels into the present form by our efficient botanical technician, Mr. Royce Oliver, to whom our appreciation is very sincere. The ecological information has been put together from field notebooks by Falanruw and Fosberg. Plant identifications are largely by Fosberg and Sachet. Much of the work on the manuscript, as well as on the original card files is by Sachet, who has worked in the Southern Marianas, though not yet in the northern islands, and who has handled and studied many of the collections and checked most of the published records. Parts of the work have been supported by Smith- sonian Research Awards Grants, Office of Naval Research contracts, a National Institutes of Health 6 SMITHSONIAN CONTRIBUTIONS TO BOTANY contract, and a Janss Foundation Grant; our time and space have been provided, over the years, by the National Museum of Natural History (formerly in the U. S. National Museum), the U. S. Geolog- ical Survey, the Pacific Science Board NAS/NRC, and the Yap Institute of Science. Transportation to the islands, on several occasions, was provided by the U. S. Navy. To all of these organizations our thanks are here expressed, as well as to the authorities of the many herbaria where we have worked or who have loaned specimens. Herbarium symbols used in the citations are according to the Index Herbariorum (Lanjouw) system except that the symbol ?Fo? indicates material still in the authors? possession, not yet deposited formally. An asterisk (*) before a name indicates that the plant concerned is believed to be a post-European intro- duction. For further information on the circumstances of work on the Micronesian flora readers are referred to the first installment of this flora (Fosberg and Sachet, 1975) and to other installments as they appear. Annotated List of the Vascular Flora PSILOTACEAE Psilotum complanatum Swartz Rare; usually epiphytic. Indo-Pacific. Alamagan: Hosokawa, 1935a, p. 442. Uncommon; terrestrial, on steep slope of bare Pantropical. Pagan: Road to Talague Beach, Moore 334 (US); Anatahan: Northwest corner of island, 300 ft Psiloturn nudum (L.) Beauvois clay soil on Anatahan. Mt. Pagan, 1870 ft [570 m], Moore 360 (US). [90 m], Falanruw 1622 (US). LYCOPODIACEAE Lycopodium cernuum L. places. Uncommon; on lava and ash in open or brushy Pantropical. Pagan: Mt. Pagan, 1870 ft [570 m], Falanruw Alamagan: Hosokawa, 1935a, p. 439. 1996. MARATTIACEAE Angiopteris evecta (Forster) Hoffmann Occasional small colonies in deep moist shaded ravines; fronds ascending, 4-5 m tall, from a mas- sive barrel-shaped fleshy rhizome or corm. Indo-Pacific. Agrigan: Canyon on east side, Fosberg 31623 (US, BISH, Fo). Alamagan: Hosokawa 7941 (BISH); Kanehira 2176 (US); south southwest coast, 650-1000 ft [200-300 m], Falanruw 1931 (US); Anderson 427 Sarigan: Hosokawa 7894; above village, 350-375 Anatahan: Northern coast, below 250 ft [75 m], (US). m, Evans 2370 (US, BISH, Fo, NY, L, P). Falanruw 1711 (US). GLEICHENIACEAE Gleichenia linearis (Burman f.) C.B. Clarke scars. Common locally on lava and colonizing erosion Old World tropics. Pagan: Mt. Charlen, spatter cone east of Fresh Alamagan: Hosokawa, 1935a, p. 435. Water Lake, Fosberg 31408 (US). POLYPODIACEAE Acrostichum aureum L. Occasional in moist ravines and on wet banks on Anatahan; forming continuous belt of marsh veg- etation around lake on Pagan in 1950; generally an inhabitant of swampy places, especially near mangrove swamps on other islands; a tall, stiff, coarse fern. Pantropical. Pagan: Crater Lake, Anderson 524 (US, BISH, Fo), 523 (US, BISH, Fo, NY, L); Fresh Water Lake, Fosberg 31379 (US). Anatahan: Northwest corner of island below 150 ft [45 m], Falanruw 1657 (US); northern coast, be- NUMBER 22 7 low 250 ft [75 m], Falanruw 1970 (US). Asplenium nidus L. Common, both epiphytic and terrestrial in shaded moist areas; in rock crevices, and on volcanic tuff cliffs, some specimens on Uracas with divided tips. Pan tropical. Uracas: South coast, below 400 ft [120 m], Fal- anruw 2201 (US). Maug: South central East Island, 400 ft [120 m], Falanruw 224 I (US, BISH). Asuncion: Southwest slope, 150 ft [45 m], Falan- ruw 2256 (US). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31611 (US, BISH). Pagan: 2 miles [3.2 km] south of pier, 10 m, Anderson 590 (US, BISH, Fo, MICH, IJ); road to Talague beach, Moore 333 (US). Alamagan: Around Partido village, Fosberg 31660 (US); south southwest coast, 450 ft [140 m], Falan- ruw 1895 (US); 1025 ft [310 m] Falanruw 1991? (US); Anderson 417 (US, Fo, MICH, IJ). Guguan: West coast, below 50 ft [15 m], Falan- ruw 1802 (US). Sarigan: Above village, 250-300 m, Evans 2385 (US, BISH, Fo, NY). Anatahan: Beach area, sea level, Falanruw 1666 (US). Asplenium unilaterale Lamarck Terrestrial on shaded banks and moist soil. Agrigan: Canyon on east side of island, Fosberg 31622 (US, BISH, Fo, MICH, IJ). Pagan: Hosokawa, 1936, p. 232. Alamagan: 100 m, Anderson 405 (US, BISH, Fo, MICH, IJ); south southwest coast, Falanruw 1905 (US). Sarigan: Hosokawa, 1936, p. 232. Anatahan: Behind village, southwest corner of island, Falanruw 1624 (US). Athyrium sorzogonense (Presl) Milde Not seen by us. Alamagan: Hosokawa, 1936, p. 229; Wagner and Grether, 1948, p. 63. Athyrium sp. Rare, on scoria cliff. Agrigan: Canyon on east side of island, Fosberg 31626 (US, Fo). Belvisia spicata (L. f.) Mirbel Epiphytic on coconut trunk in moist ravine. Indo-Pacific. Anatahan: South east of village, under 1000 ft [300 m], Falanruw 1972 (US). Bolbitis quoyana (Gaudichaud) Ching low to moderate elevations. Terrestrial, shaded banks and in deep ravines, Indo-Pacific. Agrigan: Trail around south side of island, 10- Alamagan: South southwest coast, 1025 ft [310 Sarigan: Hosokawa, 1936, p. 248. Anatahan: Western coast, below 200 m, Falanruw 1698 (US); northwest corner of island, 300 ft [90 m], Falanruw 1619 (US). Cheilanthes tenuifolia (Burman f.) Swartz widely distributed species in open habitats. 20 m, Fosberg 31624 (US). m], Falanruw 1924 (US). Terrestrial in savannas on volcanic ash soil. A Indo-Pacific. Pagan: Hosokawa 7963 (US); northwest, sea level, Falanruw 1853 (US); Mt. Pagan, Moore 392 (US), 388 (US); 900 ft [275 m], 372 (US); east of Fresh Water Lake, Fosberg 31399 (US); Mt. Charlen, spat- ter cone, east of Fresh Water Lake, Fosberg 31405 Ctenitis subglandulosa (Hance) Ching Rare on scoria cliffs and in shaded rich moist soil. Agrigan: East side of island, Fosberg 31625 (US); midwest coast, 150 ft [45 m], Falanruw 2337 (US, BISH). Alamagan: Anderson 428 (US); south southwest coast, 1025 ft [310 m], Falanruw 1992 (US). Anatahan: Northwest corner of island, 200-300 ft [60-90 m], Falanruw 1613 (US). Davallia solida (Forster f.) Swartz Ubiquitous, epiphytic on tree trunks and ter- restrial, on rocks, cliffs and even on young lava flows, rhizomes creeping extensively. In hot dry barren caldera of Mt. Pagan. Dominant plant on upper slopes of Asuncion above forest margin from 250 to 600 m or higher, forming a deep humus mat. Abundant also on Guguan. Possibly represents the original vegetation on these young volcanic slopes. Widespread in Pacific Islands in diverse habitats. Very variable in size and texture of fronds. Asuncion: Southwest slope, 1500 ft [460 m], (US). 8 SMITHSOKIAN CONTRIBUTIOKS TO BOTANY Falanruw 2291 (US); 250 ft [75 m], 2271 (US, BISH). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31610 (US, BISH). Pagan: Top of Mt. Pagan, Villagomez JV-X-12 (US), JV-7K (US); isthmus, Anderson 571 (US, BISH, Fo, NY, L); east of village, Moore 401 (US); Caldera Wall, Moore 325 (US); Mt. Charlen, east of Fresh Water Lake, Fosberg 31409 (US, BISH, Fo) . Alamagan: South southwest coast, 450 ft [I40 m], Falanrziw 1903 (US); around Partido village, Fos- berg 31676 (US, BISH, Fo, NY, L). Guguan: 200-300 ft [60-90 m], Falanruw 1842 Sarigan: Behind village, 50-100 m, Evans 2367 (US). (US, BISH, Fo, NY). Dryoptcris dissecta (Forster f.) 0. Kuntze Growing on cool shaded wet clay wall. Alamagan: South southwest coast, 500 ft [I50 m], Falanruw 1916 (US). Histiopteris incisa (Thunberg) J. Smith Alamagan: Hosokawa, 1936, p. 227. Humata repens (L. f.) Diels Epiphytic. Alamagan: Hosokawa, 1936, p. 121. Lindsaea ensifolia Swartz Characteristic of savannas and other open situa- tions, found in Miscanthus grassland on Sarigan, in lava crevasses and around steam vents on Pagan. Indo-Pacific. Pagan: Mt. Pagan, Moore 390 (US); Mt. Charlen, Sarigan: Above village, 375 m, Evans 2377 (US, east of Fresh Water Lake, Fosberg 31403 (US). BISH, Fo, NY). Lindraea obtusa J. Smith Not seen by us. Alamagan: Kramer, 1970, p. 171; Hosokawa, 1936, p. 117. -I9 -&A L- Lindsaea repens var. pectinata (Blume) Mettenius ex Kuhn Sarigan: Hosokawa, 1936, p. 116. Nephrolepis biserrata (Swartz) Schott Terrestrial in very shaded areas. Pagan: Mt. Pagan, 800 ft [240 m], Moore 376 (US). Alamagan: South southwest coast, 250 ft [75 m], Falanruw 1881 (US); 500 ft [150 m], Falanruw 1911 (US). hrephrolepis exaltata (L.) Schott Could this really be one or more of the other species misdetermined? We have not seen any of the cited material. Pagan: Hosokawa, 1936, p. 69. Alamagan: Hosokawa, 1936, p. 69. Nep hro lepis hirsu tula (Fors ter f.) Presl Terrestrial, common component of herbaceous vegetation in edges of forest on Asuncion. Asuncion: Southwestern slope, 750 ft [230 m], Falanruw 2288 (US). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31593 (US). Pagan: Mt. Pagan, 1870 ft [570 m], Moore 370 (US); Moore 387? (US), 389? (US); Caldera Wall, Moore 329 (US), 328 (US), near Crater Lake, An- derson 520 (US); east of Fresh Water Lake, 2-50 m, Fosberg 31406 (US), 31410 (US); south of Fresh Water Lake, 2-50 m, Fosberg 31391 (US); 150 ft [45 m], Bonham 7 (US). Alamagan: Around Partido village, Fosberg 31699 (US). Sarigan: Above village, 375 m, Evans 2374 (US, BISH, Fo, NY); 50-100 m, 2401 (US, BISH, Fo, NY). Anatahan: Northwest tip of island, 0-10 m, Evans 2455? (US, BISH, Fo, NY). Nep hro Eep is mu It iflora (Rox burgh) Jarret t ex Morton Terrestrial, in shaded areas. Pagan: East of Lake Laguna, Moore 352 (US), 350 (US); summit Mt. Pagan, Villagomez JV-X-16 Guguan: Southwest part of island, 175 ft [50 m], Anatahan: Northwest corner of island, 200-300 (US). Falanruw 1844 (US). ft [60-90 m], Falanruw 1618 (US). Nephrolepis sp. species, but too young to identify. ruw 1753 (US). [90 m], Falanruw 1626 (US). Undoubtedly belongs to one or more of other Sarigan: Northwest coast, 400 ft. [120 m], Falan- Anatahan: Northwest corner of island, 300 ft NUMBER 22 9 Polypodium nigrescens Blume Anatahan: Hosokawa, 1936, p, 246. Polypodium punctatum (L.) Swartz Sarigan: Above village, 250-300 m, Evans 2383 (US, BISH, Fo, NY); 50-100 m, Evans 2403 (US, BISH, Fo, NY). Polypodium scolopendria Burman f. Very common fern, on all islands south of Maug with the possible exception of Medinilla. It grows in a variety of habitats, varying from small pioneer plants on steep cinder slopes on Guguan to very large specimens growing in shaded ravines on Agrigan, fruiting under both harsh and favorable conditions. It may be either terrestrial or, more rarely, epiphytic. Asuncion: Southwest slope, 150 ft [45 m], Falan- ruw 2257 (US). Agrigan: 10-20 m, Fosberg 31592 (US, BISH); midwest coast, 150 ft [45 m], Falanruw 2345 (US). Pagan: 150 ft [45 m], Bonham 42 (US); Fresh Water Lake, Fosberg 31385 (US, BISH); Mt. Pagan 1700 ft [520 m], Moore 377 (US). Alamagan: Partido village, Fosberg 31659 (US); south southwest coast, 450 ft [I40 m], Falanruw 1893 (US); 250 ft [75 m], 1878 (US); Songsong [=Asongsong] village, 15 ft [5 m], Kondo 13 (BISH). Guguan: West coast, 200-225 ft [60-70 m], Falan- ruw 1824 (US); 200 ft [60 m], 1820a (US). Sarigan: Northwest coast, 250 ft [75 m], Falanruw 1727 (US), 1734 (US); near village, 10-100 m, Evans 2354 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, below 150 ft [45 m], Falanruw 1653 (US); western coast, be- low 200 m, Falanruw 1710 (US); northwest tip of island, 0-10 m, Evam 2456 (US, BISH). Pteris quadriaurita Retzius (Pteris pacifica Hieronymus) Very common, especially in semishade, on Anata- han and Sarigan. Small but fertile plants occur in full sun on bare black lava bluffs of Asuncion. Maug: Central East Island, 500 ft [I50 m], Falan- ruw 2236 (US), 2242 (US). Asuncion: Southwest slope, 250 ft [75 m], Falan- ruw 2276 (US, BISH), 2275 (US, BISH); 15 ft [4.5 m], 2283 (US). Agrigan: Midwest coast, 150 ft [45 m], Falanruw 2338 (US), 2339 (US); trail around south side of island, 10-20 m, Fosberg 31579 (US, BISH). Pagan: Fossil, Corwin PC-79-la to If; Caldera Wall, Moore 326 (US); Talague beach, Moore 339 (US); Fresh Water Lake, 2-50 m, Fosberg 31388 (US, BISH, Fo, MICH); 50 ft [15 m], Bonham 40 Alamagan: Around Partido village, Fosberg 31665 (US, BISH, Fo, MICH); 170 m, Anderson 406 (US); south southwest coast, 450 ft [140 m], Falanruw 1892 (US). Sarigan: Northwest coast, 250 ft [75 m], Falanruw 1742 (US); 200 ft [60 m], 1780 (US); near village, 10-100 m, Evans 2336 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, 200-300 ft [60-90 m], Falanruw 1617 (US); 150 ft [45 m], 1654 (US); west end of north coast, below 250 ft [75 m], Falanruw 1680 (US); northwest tip of island, 0-10 m, Evans 2444 (US); Kanehira 2145 (US). Pteris tripartita Swartz Four clusters of large specimens growing in semi- shaded crevasse on large exposure of older volcanic rock. A widespread species in the Pacific but curi- ously restricted in the Northern Marianas. Uracas: South coast, below 400 ft [I20 m], Falan- ruw 2202 (US). Pteris vittata L. Mt. Pagan, Moore 364 (US). Sphenomeris biflora (Kaulfuss) Tagawa Terrestrial in moist shaded areas and on bare pyroclastic material. Agrigan: In canyon on east side of island, Fos- berg 31630 (US, BISH). Pagan: Mt. Pagan, Moore 373 (US), 362 (US); near Salt Water Lake, 2-50 m, Fosberg 31344 (US); east of Fresh Water Lake, 2-50 m, Fosberg 31407 Alamagan: South southwest coast, 500 ft [150 m], Anatahan: West end of north coast, below 250 (US). Pagan: East of Lake Laguna, Moore 349 (US); (US). Falanruw 1909 (US). fL [75 m], Falanruw 1687 (US). Sphenonieris chinensis (L.) Maxon Pagan: Hosokawa, 1936, p. 115. Sarigan: Above village, 375 m, Evans 2376 (US, BISH, Fo, NY). Thelypteris opulenta (Kaulfuss) Fosberg & Sachet Terrestrial, occasional in shaded moist areas; bases of cliffs. A widespread Indo-Pacific species, 10 SMITHSONIAN CONTRIBUTIONS TO BOTANY reaching its northern limit here. Agrigan: East side of island, Fosberg 31633 (US, BISH, Fo, MICH); midwest coast, 150 ft [45 m], Falanruw 2336 (US, BISH, Fo). Alamagan: 60 m, Anderson 412 (US, BISH, Fo, MICH, IJ, NY); south southwest coast, 500 ft [I50 m], Falanruw 1913 (US). Thelypteris ornata (Wallich ex Beddome) Ching Alamagan: Wagner, 1948, p. 214; Wagner and Grether, 1948, pp. 58-59; Hosokawa, 1936, p. 233. Sarigan: Wagner, 1948, p. 214; Hosokawa, 1936, p. 233; Wagner and Grether, 1948, pp. 58-59. Anatahan: Wagner, 1948, p. 214; Hosokawa, 1936, p. 233; Wagner and Grether, 1948, pp. 58-59. Thelypteris torresiana (Gaudichaud) Alston Terrestrial, occasional in moist shaded or partly shaded areas. Pagan: Near Crater Lake, 25 m, Anderson 519 (US); pass between the two lakes, 2-50 m, Fosberg 31417 (US). Alamagan: South southwest coast, 500 ft [150 m], Falanruw 1910 (US), 1912 (US). Anatahan: Northwest corner of island, 300 ft [90 m], Falanruw 1627 (US). Thelypteris unita (L.) Morton Terrestrial, common locally in exposed places. Agrigan: Hosokawa, 1936, p. 76. Pagan: Summit Mt. Pagan, Villagomez JV-X- 11 (US): east of Lake Laguna, Moore 351 (US); 150 ft [45 m], Bonham 41 (US): pass between the two lakes, 2-50 m, Fosberg 31416 (US). Alamagan: Hosokawa, 1936, p. 76. Vittaria incurvata Cavanilles Epiphytic, rare. Sarigan: Above village, 250-300 m, Evans 2382 (US, BISH, Fo, NY). CYATHEACEAE Cyathea aramaganensis Kanehira slopes above forest limit. A common tree fern on high open volcanic Endemic. Alamagan: South southwest coast, 1025 ft [310 m], Falanruw 1927 (US, BISH); west slope of moun- tain, Anderson 424 (US). Sarigan: Hosokawa, 1936, p. 50; Wagner and Grether, 1948, p. 46. but not collected. Anatahan: Seen at higher elevations by Falanruw PANDAN ACEAE Freycinentia reineckei Warburg (F. mariannensis Merrill) Rooted in ground but climbing trees and earth cliffs in moist shaded areas, also in ravines, locally common. Endemic to Marianas. Agrigan: Midwest coast, 150 ft [45 m], Falanruw Pagan: Near Crater Lake, Anderson 522 (US). Alamagan: South southwest coast, 350 ft [I00 m], Falanruw 1882 (US); 60 m, Anderson 411 (US); vicinity of Asongsong village, Fosberg 31746 (US). Sarigan: Above village, 250-300 m, Evans 2384 (US, BISH, Fo, NY), Evans 2380 (US, BISH, Fo, NY). Anatahan: Northwest coast, below 200 ft [60 m], Falanruw 1611 (US). Pandanus tectorius Parkinson, sensu lato (incl. P. pseudo-menne Hosokawa) Probably present on all islands except Uracas, generally common, sea level to 200 m, occurring both in open and in sparse thickets, also on steep scoria canyon walls. In open the trees are rather low, with leaves touching ground: gregarious. Widespread variable species, which has been vari- ously split up. Maug: East Island, common on upper slopes, seen but not collected by Falanruw. Asuncion: Southwestern slope, 25 ft [7 m], Fa- lanruw 2279 (US). Agrigan: East side of island, Fosberg 31638 (US, BISH); midwest coast, 150 ft [45 m], Falanruw 2346 Pagan: 150-600 ft [45-180 m], Bonham 16 (US, BISH). Alamagan: Vicinity of Asongsong village, 3-1 5 m, Fosberg 31704 (US, BISH); south southwest coast, 0-200 ft [O-60 m], Falanruw 1955 (US). Guguan: West coast, below 50 ft [I5 m], Falan- ruw 1811a (US). Anatahan: Beach area, sea level, Falanruw 1671 2341' (US). (US). (US). NUMBER 22 11 GRAMINEAE WBambusa vulgaris Schrader ex Wendland forest at about 300 m, not seen elsewhere. [SO0 m], Falanruw 1975 (US). "Cenchrus brownii Roemer & Schultes Weed in disturbed places, coconut groves and along trails. Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31587 (US, BISH, Fo). Pagan: Marine camp, 4 m, Anderson 601 (US, BISH, Fo, NY). Alamagan: Vicinity of Asongsong village, Fos- berg 31715 (US, BISH, Fo.) "Cenchrus echinatus L. Weed of disturbed, usually open places, as around villages. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2158 (US). Pagan: Isthmus, 3 m, Anderson 557 (US, BISH, Fo, NY, L); village, Mome 396 (US). Alamagan: South southwest coast, 0-200 ft [O- 60 m], Falanruw 1954b (US); vicinity of Asongsong village, 3-15 m, Fosberg 31715 (US). Centorheca Zappacea (L.) Desvaux Common under forest canopy, seen only on Ana t a han. Anatahan: Hosokawa 7833 (Fo); northwest corner of island, 200-300 ft [60-90 m], Falanruw 1631 (US); southwest of village under 1000 ft [300 m], Falanruw 1968 (US). 'Chloris inflata Link Common in open areas, especially around villages and old human habitations, not seen on unin- habited islands. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2162 (US), 2192 (US). Pagan: Volcanic ash plain, 50 ft [15 m], Bonham 33 (US, Fo); village, Moore 415 (US). Sarigan: Northwest coast, 200 ft [60 m], Falan- ruw 1772 (US); village, 10-50 m, Evans 2437 (US, BISH, Fo, NY). Anatahan: Northwest corner, below 200 ft [60 m], Falanruw 1634 (US); northwest tip of island, 0-10 m, Evans 2449 (US, BISH, Fo, NY). "Chrysopogon aciculatus (Retzius) Trinius Two colonies, only seen near upper edge of Anatahan: Southwest of village, under 1000 ft Common pioneer species on volcanic ash and cinders from low elevations to at least 450 m (on Asuncion); in old village sites. Asuncion: Southwestern slope, 1500 ft [450 m], Falanruw 2296 (US). Pagan: Marine camp, 20 m, Anderson 516 (US, BISH, Fo, NY, L); volcanic ash plain, 50 ft [15 m], Bonham 30 (US, Fo); east of Fresh Water Lake, Fosberg 34412 (US); village, Moore 411 (US). Alamagan: Hosokawa, 1935b, p. 313. Guguan: West coast, 50 ft [15 m], Falanruw 1809 (US). "Cynodon dactylon (L.) Persoon Common in open areas around human habita- tions; sod-forming. Widespread. Agrigan: Southwest coast, below 300 ft 190 m], Falanruw 2163 (US), 2187 (US). Pagan: Marine camp, 20 m, Anderson 512 (US, BISH); isthmus, 10 m, Anderson 583 (US, BISH, Fo, NY, L, P). Alamagan: South southwest coast, 0-200 ft [O- 60 m], Falanruw 1961 (US). Sarigan: Village, 10-50 m, Evans 2438 (US, BJSH, Fo, NY). Medinilla: Swedberg ms. 1974. "Dactyloctenium aegypt ium (L.) Willdenow Occasional to common along paths and in open somewhat bare areas around human habitation. Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31583 (US). Pagan: Isthmus, Anderson 5.58 (US); volcanic ash soil, 50 ft [15 m], Bonham 28 (US, Fo); west side near Salt Lake, Fosberg 31351 (US); village, Moore 413 (US). Alamagan: Vicinity of Asongsong village, 3-15 m, Fosberg 31710 (US, BISH, Fo); south southwest coast, 0-200 ft [O-60 m], Falanruw 1953 (US). "Digitaria ciliaris (Retzius) Koeler Occasional in open places, on roads and paths, also on open lava and ash. Agrigan: Trail along south side of island, 10-20 m, Fosberg 31606 (US, BISH, Fo, NY, L). Pagan: Fresh Water Lake, 2-50 m, Fosberg 31401 (US, BISH, Fo); near Salt Water Lake, west side of island, Fosberg 31366 (US, BISH, Fo). Sarigan: Hosokawa, 1935b, p. 314. Anatahan: Hosokawa, 1935b, p. 315. *Digitaria ciliaris var. 12 SMITHSONIAN CONTRIBUTIONS TO BOTANY One plant seen on volcanic cinder sand at low Guguan: West coast, below 50 ft [15 m], Falan- elevation, among pioneer vegetation. ruw 1801 (US). Digitaria gaudickaudii (Kunth) Henrard pears very hardy. ruw 1812 (US); 200-300 ft [60-90 m], 1851 (US). Digitaria mariannensis Merrill Occasional pioneer on volcanic cinder sand; ap- Guguan: West coast, below 50 ft [I5 m], Falan- Occasional on hot dry banks. Maug: North end of East Island, below 400 ft [120 m], Falanruw 2212 (US). Digitaria mezii Kanehira Sarigan: Mez in Diels, 1924, p. 25. Digitaria radicosa (Presl) Miquel (Digitaria timorensis (Kunth) Balansa) Common in coconut groves and open areas from low elevations to 500 m (on Asuncion). Asuncion: Southwest slope, 1500 ft [450 m], Fa- lanruw 2311 (US); 250 ft [75 m], 2269 (US). Alamagan: Asongsong village, 3-1 5 m, Fosberg 31711 (US, BISH, Fo); Anderson 415 (US, BISH, Fo, NY). Sarigan: Northwest coast, 200 ft [60 m], Falanruw 1773 (US). Digitaria setigera Roth (Digitaria pruriens (Fisher ex Trinius) Biise) In open areas. Alamagan: Vicinity of Asongsong village, 3-15 m, Fosberg 31713 (US, BISH, Fo); Fosberg 31712 (US). Digitaria setigera var. (Digitaria pruriens var.) BISH, Fo, NY). Evans 2451 (US, BISH, Fo, NY). *Eleusine indica (L.) Gaertner Sarigan: Below village, 0-20 m, Evans 2431 (US, Anatahan: Northwest tip of island, 0-10 m Weed in villages and open areas. Cosmopolitan. Agrigan: Village, Fosberg 31431 (US); southwest coast, below 300 ft, Falanruw 2164 (US); below 200 ft [SO m], 1649 (US); midwest coast, below 200 ft [GO m], Falanruw 2328 (US, BISH). Pagan: Isthmus, 5 m, Anderson 560 (US, BISH, Fo, NY, L); near Salt Lake, west side of island, Fosberg 31353 (US). Alamagan: Around Partido village, Fosberg 31651 Sarigan: Near village, 10-100 m, Euans 2332 (US, BISH, Fo, NY). Anatahan: Northwest tip of island, Evans 2458 (US, BISH); northwest corner of island, below 75 m, Falanruw 1694 (US); 60 m, Falanruw 1649 (US). *Eragrostis tenella (L.) Beauvois (Eragrostis amabilis Wight & Arnott) Common on open bare ground, along paths, and in villages. Widespread. Agrigan: Trail around south side of island, Fos- berg 31572 (US); midwest coast, 200 ft [60 m], Fa- lanruw 2335 (US). Pagan: Isthmus, Anderson 556 (US, BISH, Fo, NY, L); Salt Lake, west side of island, 2-50 m, Fosberg 31367 (US, BISH). Alamagan: Around Partido village, Fosberg 31644 (US, BISH). Sarigan; Near anchorage, 150 ft [45 m], Falan- ruw 1723 (US). Anatahan: Village area northwest corner of is- land, below 200 ft [60 m], Falanruw 1645 (US); northwest tip of island, 0-10 m, Euans 2452 (US). Heteropogon contortus (L.) Beauvois Common on bare ash slopes, from low elevations to 500 m (on Asuncion). Asuncion: Southwest slope, 1500 ft [450 m], Fa- lanruw 2302 (US), 2308 (US). Agrigan: Hosokawa, 1935b, p. 313. Pagan: West side of island, near Salt Lake, Fosberg 31355 (US, BISH); Talague beach, Moore 335 (US); isthmus, 6 m, Anderson 561 (US, BISH, Fo, NY, L); west side of Lake Laguna, Moore 353 Alamagan: Near Asongsong village, Fosberg Guguan: West coast, below 50 ft [I5 m], Falan- Medinilla: Swedberg, MS 1974. Occasional on upper slopes of volcanoes. Asuncion: Upper southwestern slope, 500 m, Fa- Pagan: Hosokawa 7952 (US). Alamagan: Hosokawa, 1935b, p. 31 1. Anatahan: Hosokawa, 1935b, p. 311. (US) a (US). 31745 (US). ruw 1814 (US); 200-300 ft [90 m], 1850 (US). Ischaemum longisetum Merrill lanruw 2298 (US); 2310 (US). Ischaemum sp. NUMBER 22 13 Sarigan: Behind village, 50-100 m, Euans 2365 (US, BISH, Fo, NY) (sterile). Lepturus repens (Forster) R. Brown Small patch on top of bluff over ocean. Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31584 (US, BISH). Pagan: Villagomez JV-03 (US) (sterile, doubtful); Mt. Pagan, Moore 379 (US). Miscanthus floridulus (Labillardikre) Warburg ex Schumann & Lauterbach Common or abundant on all islands except Uracas, least so on Maug, Asuncion, and Guguan, which have been least subject to burning or graz- ing by goats. On the other islands it is the dominant vegetation on upper slopes, more locally so on the three mentioned. It is also occasionally found down to near sea level. On upper slopes it forms matted brakes to 3 m deep, with harsh cutting leaf mar- gins. It recovers readily from burning. Widespread in the high islands of the west and south Pacific. In Micronesia, found only in the Marianas chain and on Ponape, reaching its northern limit on Maug. Maug: Fritz, 1902, p. 104. Asuncion: Southwest slope, 1000 ft [300 m], Fa- lanruw 2303 (US). Agrigan: Marche 294 (P, Fo); east side of island, Fosberg 31628 (US); midwest coast, 200 ft [60 m], Falanruw 2334 (US). Pagan: Mt. Pagan, Villagomez JV-X-13 (US), JV-2K (US); Hosokawa 7950 (Fo); isthmus, 10 m, Anderson 517 (US, BISH, NY, L); 150 ft [45 m], Bonham 4 (Fo); Kanehira 2198 (KY); east end of air strip, Moore 301 (US). Alamagan: Vicinity of Asongsong village, 3-15 m, Fosberg 31744 (US). Sarigan: Northwest coast above anchorage, 0-75 ft [0-20 m], Falanl-uw 1788 (US). Anatahan: Hosokawa 7830 (US); northwestern corner of island, below 250 ft [75 m], Falanruw 1693 (US); sea level, 1663 (US). Oplismenus composittis (L.) Beauvois Common on ground in shade in semi-open for- ests. Not found on the most active volcanoes. A widespread IndePacific species. Asuncion: Southwestern slope, 400 ft [I20 m], Falanruw 2313 (US). Agrigan: Marche 299 (P, Fo); trail around south side of island, 10-20 m, Fosberg 31605 (US). Alamagan: South southwest coast, below 500 ft [I50 m], Falanruw 1936 (US) (sterile); vicinity of Asongsong village, 3-15 m, Fosberg 31735 (US). Sarigan: Above village, 150-250 m, Evans 2390 (US, BISH, Fo, NY). Anatahan: Northern coast, below 200 ft [60 m], Falanruw 1964 (US); northwest comer of island, 200-300 ft [60-90 m], Falanruw 1615 (US). Oplismenus undulatifolius var. imbecillis (Roemer & Schultes) Hackel Seen only growing abundantly in open area on slope above sea. Indo-Pacific. Sarigan: Northwest coast, 0-75 ft [0-20 m], Fa- lanruw 1789 (US). "Panicurn ambiguum Trinius (Brachiaria paspaloides (Presl) Hubbard) Rare in 1950 on roadsides near landing. Indo-Pacific. Pagan: Near landing, Fosberg 31639 (US, BISH, Fo) . "Panicurn distachyon L. Not seen by us. Indo-Pacific. Anatahan: Hosokawa, 1935b, p. 318. Panicum reptans L. Growing along open path in village area. Pantropical. Agrigan: Southwest coast village area, below 300 ft [90 m], Falanruw 2179 (US). "Paspalurn commersonii Lamarck (Paspalum scrobiculatum auct. non L.) Agrigan: Hosokawa, 1935b, p. 313. Pagan: Air strip, Moore 302 (US). Alamagan: Hosokawa, 193513, p. 313. "Paspalurn conjugatum Bergius Occasional to common along paths and in semi- open areas, abundant in moist area near Fresh Water Lake on Pagan, not known from the islands less disturbed by human activily and from the less- wooded islands. Pantropical, originally from tropical America. Agrigan: Trail around south side of island, 10- Pagan: Fresh Water Lake, 2-50 m, Fosberg 31394 Alamagan: South southwest coast, 750 ft [230 20 m, Fosberg 31568 (US). (US). 14 SMITHSONIAN CONTRIBUTIONS TO BOTANY m], Falanruw 1922 (US). Sarigan: Near anchorage, 100 ft [30 m], Falanruw 1719 (US); near village, 1-100 m, Evans 2335 (US, BISH, Fo, NY). Paspalum distichum L. (Paspalum uaginatum Swartz) Forming mats in saline areas near beaches and margins of Salt Water Lake. Pan tropical. Pagan: West side of Lake Laguna, Moore ?57 Anatahan: Beach area, sea level, Falanruw 1469 Anatahan: Falanruw 1987 (US). (US) * (US) * Paspalum orbiculare Forster A pioneer species on volcanic cinders and in soil pockets on lava, reaching 500 m elevation on Asun- cion, also around villages. A widely distributed weedy species, distinguished with difficulty from P. cartalagineum Presl. Asuncion: Southwest slope, 1500 ft [460 m], Fa- lanruw 2299 (US); 2307 (US). Pagan: Fosberg 31398 (US); isthmus, 5 m, Ander- son 559 (US, BISH, Fo, NY, L); village, Moore 412 Guguan: West coast, 200-225 ft [60-65 m], Falan- (US). ruw 1852 (US). *Pennisetum purpureum Schumacher near Salt Lake in 1950. Forming dense tangled brakes locally along road From Africa. Pagan: Near Salt Lake, west side of island, Fosberg 31350 (US). "Saccharum oficinarum L. Not collected in the area since 1900. Indo-Pacific. Anatahan: Fritz, 1902, p. 98; Lehne and Gabler, 1972, p. 22, as "Zuckerrohr." "Setaria lutescens (Weigel) Hubbard Not seen by us. Pan tropical. Agrigan: Hosokawa, 193513, p. 319. Alamagan: Hosokawa, 1935b, p. 319. Growing in open area near dwelling. Pantropical. Anatahan: Northwest corner of island, below 200 (W) "Sporobolus diander,Beauvois ft [60 m], Falanruw 1644 (US). "Sporobolus fertilis (Steudel) Clayton On west side of Salt Lake. Tropical Asia. Pagan: West side of Lake Laguna, Moore 354 (US), 355 (US). Sporobolus virginicus (L.) Kunth Very abundant along beach on Pagan and on upper slopes of exposure of older lava on Uracas. A pantropical strand grass, surprisingly localized in Northern Marianas. Uracas: West side of south coast, below 400 ft [120 m], Falanruw 220? (US, BISH, Fo). Pagan: Trail to Pansantati, below 100 ft [30 m], Falanruw 1842 (US); back of beach, 3 m, Anderson 611 (US). Stenotaphrum micranthum (Desvaux) Hubbard Rare in crevices in lava rock on hot dry lee slope. Widespread, but usually not abundant Indo- Maug: North end of East Island, below 400 ft Pacific species, known from Southern Marianas. [120 m], Falanruw 2218 (US). Thuarea involuta (Forster f.) R. Brown ex Roemer & Schultes Widespread Indo-Pacific strand species, surpris- ingly rare in the Northern Marianas, but elsewhere usually found on coral sand, which is not abundant here. Sarigan: Hosokawa, 193513, p. 320. *Zea mays L. Does not ordinarily persist long uncultivated. American. Pagan: Corte, 1927, p. 330. Zoysia matrella (L.) Merrill var. matrella This species forms a dense, very low turf or mat locally in openings and especially on cliff tops, seashores and other exposed places. The relation- ship between the two varieties merits further study. The difference is chiefly in leaf width and degree of involution of the blades. In cultivation at least certain clones seem to maintain their character- istics. How much simple fluctuation in response to environmental severity is involved and how much ecotypic variation merit much further study. To be conclusive this should involve careful field ob- servation on islands where both varieties occur, as well as cultivation experiments with living ma- NUMBER 22 15 terial brought from the same islands but different habitats. Eastern Asiatic and Micronesian. Uracas: Seen by Falanruw from a distance form- ing mats on exposures of older lava, but variety not determined. Maug: Northern end of East Island, below 400 ft [120 m], Falanruw 2215 (US). Asuncion: Southwestern slope, 50 ft [I5 m], Fa- lanruw 2285 (US). Agrigan: Southwest coast village area, below 300 ft [90 m], Falanruw 2155 (US). Alamagan: Hosokawa 7919 (US); south south- west coast, 0-200 ft [O-60 m], Falanruw 1954 (US). Sarigan: Northwest coast, 0-75 ft [O-20 m], Fa- Zanruw 1774 (US). Anatahan: Hosokawa, 1935b, p. 314. Zoysia matrella var. tenuifolia (Willdenow ex Thiele) Sasaki (Zoysia tenuifolia Willdenow ex Thiele) Generally from more severe habitats, exposed cliff-tops and other saline places. Asuncion: Southwest slope, 750 ft [230 m], Falan- ruw 2287 (US). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31581 (US, BISH, Fo). Pagan: Near Salt Lake, Fosberg 31345 (US, BISH, Fo); near landing, 3 m, Anderson 514 (US, BISH, Fo, NY, L). Alamagan: Asongsong village, 15 m, Fosberg 31709 (US, BISH, Fo). Sarigan: Seashore, 0-20 m, Evans 2430 (US, BlSH, Fo, NY, L). Anatahan: Hosokawa 7857 (Fo); north coast, below 200 ft [60 m], Falanruw 1963 (US). CY PERACEAE * Cyperus brevifolius (Rottboell) Hasskarl Found in disturbed, usually open places, un- doubtedly carried accidentally by man; present distribution pantropical, but origin now obscure. Strangely rare in Northern Marianas. Agrigan: Southwest coast village area, below 300 ft [90 m], Falanruw 2179 (US). *Cyperus compressus L. places. A weed of villages and other disturbed open Pantropical and warm-temperate. Pagan: 50 ft [15 m], Anderson 34 (US, Fo); isth- Alamagan: South southwest coast, village area, Anatahan: Northwest coast village area, below mus, Anderson 553 (US). 0-200 ft [O-60 m], Falanruw 1959 (US). 200 ft [60 m], Fulanruw 1636 (US). Cyperus cyperinus (Retzius) Suringar A weedy tufted sedge growing in semi-open and open areas from low elevations to 500 m (on Asun- cion), on cinder banks and slopes on Agrigan and Guguan. Widespread in warm-temperate and sub- tropical zones, origin obscure. Asuncion: Southwest slope, 1500 ft [460 m], Fa- lanruw 2300 (US). Agrigan: Midwest coast, 200 ft [60 m], Falanruw 2326 (US); trail around south side of island, 10-20 m, Fosberg 31571 (US). Pagan: Air strip, Moore 303 (US); isthmus, 6 m, Anderson 532 (US, BISH, Fo, NY, L, P), 554 (US); Bonham 6 (US, BISH). Alamagan: South southwest coast, 450 ft [I40 m], Falanruw 1900 (US). Guguan: 350 ft [lo0 m], Falanruw 1837 (US); 200-300 ft [60-90 m], 1830 (US). Sarigan: Northwest coast, 0-75 ft [O-20 m], Fa- lanruw 1796 (US); north of village 50-175 m, Evans 2420 (US, BISH, Fo, NY). Anatahan: Koyama, 1964, p. 97. Cyperus digitatus Roxburgh In tall grass and under semi-open forest canopy. Pan tropical. Anatahan: West end of north coast, below 250 ft [75 m], FaZanruw 1962 (US); northwest corner of island, below 200 ft [60 m], Falanruw 1600 (US). Cyperus javanicus Houttuyn A coarse tufted sedge, abundant and widespread in many habitats on all islands visited; from vol- canic cinder sands on Guguan to soil pockets on recent lava on Pagan and rich volcanic soil on Anatahan, one of the most abundant plants on Uracas, holding the soil and providing anchorage and materials for booby nests on the loose pyro- clastic slopes. A widespread and common Indo-Pacific species usually from saline habitats, generally rather close to the sea. Its behavior in the Northern Marianas is unusual, in that it seems not at all confined to saline places, and reaches high elevations. A pos- 16 SMITHSONIAN CONTRIBUTIONS TO BOTANY sible explanation is that the small flora in these islands provides only reduced competition, per- mitting the species to show its real tolerance for a much wider range of conditions than is possible in most of its geographic range. Uracas: South coast, below 400 ft [120 m], Falan- ruiu 2207 (US), 2206 (US, BISH, Fo). Maug: North end of East Island, below 400 ft [I20 m], Falanruw 2231 (US). Asuncion: Southwest slope, 150 ft [45 m], Falan- rztw 2262 (US). Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2169 (US); Marche 296 (P, Fo); Fosberg 31575 (US, BISH, Fo, NY). Pagan: Anderson 554 (US, BISH, Fo, NY); Cal- dera Wall, Moore 330 (US); Fresh Water Lake, Fosberg 31397 (US); Laguna village, Villagomez Alamagan: Near Partido village, Fosberg 31 641 (US, BISH, Fo, NY); south southwest coast, 450 ft [I40 m], Falanruw 1901 (US). Guguan: West coast, below 50 ft [I5 m], Falan- ruw 1816 (US), 1807 (US). Sarigan: Above anchorage northwest coast, 0-75 ft [O-20 m], Falanruw 1793 (US); near village, 10- 100 m, Evans 2357 (US). Anatahan: Northwest corner of island, below 150 ft [45 m], Falanruw 1656 (US); northwest tip of island, Evans 2457 (US, BISH). Medinilla: Swedberg, MS 1974, as ?Cyperus spp.? (probably this species). ?Cyperus kyllingia Endlicher Found in semishaded and moist open, usually disturbed areas, often about villages. A common pantropical species. Agrigan: Midwest coast, 30 ft [lo m] Falanruw Alamagan: South southwest coast, 450 ft [I40 Anatahan: Northwest corner of island, below 250 JV-X-8 (US). 2315 (US). m], Falanruw 1902 (US). ft [75 m], Fulanruw 1691 (US), 1994 (US). Cyperus odoratus L. in moist or wet places. A pantropical and warm-temperate species found Anatahan: Koyama, 1964, p. 100. Cyperus polystachyos Rottboell places. A pantropical weed, found in open usually moist Agrigan: Hosokawa 8004 (Fo). Pagan: Fresh Water Lake, Fosberg 31387 (US, BISH, Fo, NY); Caldera Wall, Moore 331 (US). Fimbristylis boninensis Hayata (Fimbristylis urakasiana Kukenthal) A pioneer species, scattered in harsh environ- ments, on crater rims in Pagan, barren cinder sands on Guguan, to 500 m on Asuncion; the only species on expanses of pyroclastic slopes on Uracas. Endemic to the Bonins and Northern Marianas. Uracas: West side of south coast, below 400 ft [I20 m], Falanruw 2208 (US, BISH, Fo), Falanruw 2204 (US), Falanruw 2205 (US). Maug: Northern end of East Island, below 400 ft [I20 m], Falanrziw 2214 (US, BISH), Falanruw 2213 (US, BISH), Falanruw 2216 (US). Asuncion: Southwest slope, 1500 ft [460 m], Fa- lunruw 2301 (US). Pagan: Trail to Pansantati, below 100 ft [30 m], Falanruw 1863 (US); Fosberg 31415 (US). Guguan: West coast, below 50 ft [I5 m], Falan- ruw 1804 (US). Anatahan: Koyama, 1964, p. 88. Fimbristylis cynzosa R. Brown A small densely tufted sclerophyllous sedge, growing in pioneer habitats often of extreme sever- ity, but also, where competition is not too severe, in more mesic places, where it reaches a much greater stature. Found on all islands visited except Uracas. Survives on cliffs and very steep slopes. Pantropical, extremely variable, no attempt made here to separate varieties. Maug: North end of East Island, below 400 ft [I20 m], Falanruw 2230 (US, BISH), Falanruw 2222 Asuncion: Southwest slope, 20 ft [6 m], Falan- Agrigan: Midwest coast, 20 ft [6 m], Falanruw Pagan: Mt. Pagan, Moore 363 (US); 900 ft [275 m], Moore 369 (US); Talague beach, Villagomez JV-10-11 (US); Talague beach, Moore 344 (US). Alamagan: South southwest coast, 0-200 ft [O- 60 m], Falanruw 1952 (US); vicinity of Asongsong, village, 3-15 m, Fosberg 31703 (US). Guguan: West coast, 200 ft [60 m], Falanrw 1817 (US); below 50 ft [I5 m], 1803 (US), Falanruw 1806 (US), Falanruw 1810 (US). Sarigan: Northwest coast, above anchorage, 0- (US). ruw 2278 (US, BISH, Fo). 2321 (US). NUMBER 22 17 75 ft [O-20 m], Falanruw 1792 (US); shore, 0-20 m, Evans 2404 (US, BISH). Anatahan: Kanehira 2150 (P, NY, FU); beach area, sea level, Falanruw 1661 (US). Fimbristylis dichotoma ssp. podocarpa (Nees & Meyen) T. Koyama Widespread weedy species, apparently uncom- mon in this area. Agrigan: Koyama, 1964, p. 87. Pagan: Anderson 564 (US, BISH, Fo). Alamagan: Koyama, 1964, p. 87. Sarigan: Kanehira 2166 (KY). Anatahan: Koyama, 1964, p. 87. Schoenus philippinensis (Palla) Kiikenthal ex Merrill Rare in our area. Alamagan: Koyama, 1964, p. 105; Hosokawa, 1935a, p. 263. Scleria lithosperma (L.) Swartz in semi-open places. Occasional large patches in forest and in thickets Pantropical. Asuncion: Southwestern slope, 150 ft [45 m], Agrigan: Koyama, 1964, p. 71. Pagan: Anderson 584 (US, BISH, Fo). Alamagan: Around Partido village, Fosberg 31670 Sarigan: Above village, 375 m, Evans 2368 (US, Fa 1 anru w 2261 (US). (US) * BISH, Fo, NY). PALMAE "Areca catechu L. Small palm growing in jungle and cultivated in semi-open area on Anatahan, where common enough that nuts are exported to Saipan. Culti- vated trees are usually robust and short, with inflorescences within easy reach. Indo-Pacific economic species. Agrigan: Seen but not collected by Falanruw. Alamagan: South southwest coast, 750 ft [230 m], Falanruw 1921 (US), 1921 b (US). Anatahan: Beach area, Falanruw 1989 (US); west coast, below 200 ft [60 m], Falanruw 1705 (US); northern coast, below 250 ft [75 m], Falanruw 1695 *Cocas nucifera L. (US). Generally extensively planted coconut groves are present on all inhabited islands, plus Maug, with copra exported at least from Anatahan. One stunted yellowing tree seen on Guguan. Coconuts were planted by Fritz in 1900 on Medinilla and Uracas but did not persist. Pantropical economic species. Uracas: Planted by Fritz in 1900 (Fritz, 1902, Maug: Fritz, 1902, p. 114. Asuncion: Seen by Falanruw, 1972, not collected. 100 stunted trees reported by La Pkrouse (1797, II:309) as growing on the lava in 1786. Agrigan: Corte, 1927, p. 345; Fritz, 1902, p. 109. Pagan: Bonham 25 (US, BISH); fossil, Corwin Alamagan: South southwest coast, sea level, Fa- lanruw 1993 (US). Guguan: 320 ft [lo0 m], Falanruw 1841 (US). Coconuts planted by Fritz in 1900 (Fritz, 1902, p. p. 116), not seen in 1972. P6-79-6 (USNM). 101). Sarigan: Corte, 1927, p. 328. Anatahan: Village area, below 250 ft [75 m], Fa- Medinilla: Fritz, 1902, p. 97. lanruw 1697 (US). ARACEAE "Alocasia macrorrhiza (L.) Schott Found in shaded moist areas, gorge bottom (Agrigan), and around lake (Pagan); undoubtedly spread by man. Fossil occurrence on Pagan (Fos- berg and Corwin, 1958, p. 10) regarded as evidence of presence of man in the Northern Marianas prior to the major eruption that formed the caldera on Pagan. Agrigan: Midwest coast, 150 ft [45 m], Falanruw 2331 (US); trail around south side of island, 10- 20 m, Fosberg 31618 (US). Pagan: Fossil, Corwin PC43-2A, PC433a, PC-43-5 (USNM); Fresh Water Lake, 2-50 m, Fos- berg 31375 (US). Alamagan: South southwest coast, 650-1000 ft [ZOO-300 m], Falanruw 1932 (US). Sarigan: North of village, 50-175 m, Evans 2424 (US, BISH, Fo, NY). "Colocasia esculenta (L.) Schott less brought by man. Planted and persisting in marshy places, doubt- Pantropic food plant. 18 SMITHSONIAN CONTRIBUTIONS TO BOTANY Pagan: Fresh Water Lake, 2-50 m, Fosberg 31376 Alamagan: Anderson 413 (US). Anatahan?: Lehne and Gabler, 1972, p. 22, (US). ?taro.? *Xanthosoma sagittifolium (L.) Schott Commonly planted in shaded and semishaded areas, especially under coconut trees. Food plant of American origin. Agrigan: Trail around south side of island, 10- Pagan: North of marine camp, Anderson 605 Alamagan: Around Partido village, Fosberg 31 694 Sarigan: Northwest coast, 250 ft [75 m], Falanruw Anatahan: West coast, below 200 ft [60 m], 20 m, Fosberg 31576 (US). (US). (US). 1738 (US). Falanruw 1699 (US). FLAGELLARIACEAE Flagellaria indica L. Extensive tough liana, climbing in trees by means of curling leaf-tips, occasional in thickets and woods in canyon bottoms. Indo-Pacific. Agrigan: East side of island, Fosberg 31635 (US); midwest coast, 200 ft [60 m], Falanruw 2318 (US). Sarigan: Above village, 150-250 m, Evans 2396 (US, BISH). BROMELIACEAE *Ananas comosus (L.) Merrill Common in cultivation and semiwild on Pagan. Food plant of American origin. Agrigan: Lehne and Gabler, 1972, p. 36. Pagan: 150-300 ft [45-90 m], Bonham 24 (US, BISH); north of marine camp, Anderson 604 (US, BISH); Fresh Water Lake, 2-50 m, Fosberg 31374 (US). Alamagan: Fritz, 1902, p. 104. COMMELINACEAE *Commelina benghalensis L. around villages. Common weed in open places and plantations Probably of Asiatic origin. Alamagan: Vicinity of Asongsong village, 3-15 Sarigan: Above village, 150-250 m, Evans 2397 ni, Fosberg 31723 (US). (US, BISH, Fo, NY). LILIACEAE, sensu latissimo (incl. ACAVACEAE and AMARYLLIDACEAE) *Agave americana L.? Persisting from planting. Ornamental of American origin. Pagan: Fosberg, 1958, p. 19. Persisting from planting. Fiber plant of American origin. Pagan: Villagomez JV-X-9 (US). ?Agave rigida var. sisalana (Perrine) Engelmann Pagan: Isthmus, Anderson 555 (US). Crinum asiaticum L.? Abundant, growing spontaneously on cliffs and lower seaward slopes; also planted around graves. Asiatic, widely planted as an ornamental, possi- bly native in Northern Marianas. Maug: Leeward slope of central East Island, 30 ft [lo m], Falanruw 2239 (US). Asuncion: Seen but not collected, Falanruw in 1972. Alamagan: Seen but not collected, Falanruw in ?Agave rigida Miller 1972. [75 m], Falanruw 1696 (US). Crinum macrantherum Engler Not seen by us. New Guinea to Micronesia. Anatahan: North northwest coast, below 250 ft Pagan: Hosokawa, 1935a, p. 266 (doubtful rec- ord). Curculigo orchioides Gaertner Usually found in grasslands, not seen by us. I ndo-Paci fic. Pagan: Kanehira, 1935a, p. 289. Dianella ensifolia L. Common in Southern Marianas in grassland on volcanic soil, not yet known living from Northern Marianas. Pagan: Fossil, Corwin PC-85-1, PC-85-2, Ter- mon dr Corwin ZD2-(F)-3, ZD-P(F)lf (USNM). NUMBER 22 19 (See Fosberg and Corwin, 1958, p. 10). ?Hymenocallis littoralis (Jacquin) Salisbury ing. Planted as an ornamental and sometimes persist- Tropical American strand species. Pagan: Isthmus, 3 m, Anderson 592 (US, BISH, Fo, NY, L); sea level, Bonham 22 (US, BISH); vil- lage, Moore 397 ? (US). Medinilla: Swedberg, MS 1974 as ?Pancratium 1 it t orale.? TACCACEAE * Tacca leontopetaloides (L.) 0. Kuntze Persisting from early introduction by Microne- sians. Growing in open rich volcanic soil. Indo-Pacific food plant. Pagan: 6 m, Anderson 529 (US, BISH). Sarigan: Vicinity of anchorage, 200 ft [60 m], Falanruw 1766 (US), 1765 (US). DIOSCOREACEAE ?Dioscorea alata L. Planted and persisting in moist wooded area. Pantropical food plant. Alamagan: South southwest coast, 500 ft [150 m], Falanruw 1906 ? (US). Sarigan: Above village, 150-250 m, Evans 2388 (US, BISH, Fo, NY). Dioscorea esculenta var. spinosa (Roxburgh) Prain & Burkill Probably spontaneous, not seen by us. Old World tropics. Agrigan: Corte, 1927, p. 344. Pagan: ?lehne and Gabler, 1972, p. 33, as ?Wilde yams.? #Dioscorea nummularia Lamarck Not seen by us. Pagan: Corte, 1927, p. 330. MUSACEAE ?Musa sapientum L. Wild, seedy, small-fruited bananas are common in upper ravines of Anatahan (1974), with many seedlings (1715, 1965). Cultivated plants at lower elevations (1714) had larger fruits with very few seeds. Asuncion: Seen but not collected by Falanruw. Agrigan: Fritz, 1902, p. 109. Pagan, Fritz, 1902, p. 104. Alamagan: Fritz, 1902, p. 102. Anatahan: East side of village, under 1000 ft [300 m], Falanruw 1974 (US); western coast, below 200 ft [60 m], Falanrziw 1714 (US): north of village, 400 ft [120 m], Falanruw 1965 (US) seedling; west- ern coast, below 200 ft [60 m], Falanruw 1715 (US) seedling. ?Mum textilis Nees Occasional groups of plants with small seedy fruits on ravine banks. Fiber is used to make lava- lavas by the inhabitants who are of Carolinian origin. This species may have been introduced by them for this purpose. Originally from Philippines. Agrigan: Midwest coast, Falanruw 2342 (US). ZINCIBERACEAE Curcuma australasica Hooker f.? Growing about graveyard and near house. Probably native to Micronesia or of very early introduction. Anatahan: Northwestern corner of island, below 200 ft [60 m], Falanruw 1713 (US); below 200 ft [60 m], 1642 (US) (both collections sterile). Zingiber zerumbet (L.) Roscoe areas. Occasional, growing along trails in moist forested Indo-Pacific. Anatahan: Northern coast, below 250 ft [75 m], Falanruw 1712 (US); south east of village, under 1000 ft [300 m], Falanruru 1973 (US). MARANTACEAE ?Maranta arundinacea L. In secondary forest in ravine. Alamagan: Anderson 429 (US). ORCHIDACEAE Calanthe furcata Bateman? slopes from 30 to 150 m. Terrestrial, frequent in sparse forest on lower Indo-Pacific. Asuncion: Southwest slope, 400 ft [ 120 m], Falan- ruw 2312 (US); 250 ft [75 m], 2277 (US). 20 SMITHSONIAN CONTRIBUTIONS TO BOTANY Eulophia macrostachya Lindley Not seen by us. Said to be usually terrestrial. Indo-Pacific. Alamagan: Hosokawa, 1942, p. 9. Sarigan: Hosokawa, 1942, p. 9. Terrestrial; in moist ravine bottoms and forest Indo-Pacific. Anatahan: Falanruw 1914, Falanruw 1966, Fa- Alamagan: Falanruw 1928. Spathoglottis sp. Rare, found living only once; reported fossil from Pagan as S. micronesiaca Schlechter in Fos- berg and Corwin (1958, p. 10) but not at all certain that it is that species. Neruilia aragoana Gaudichaud floors. lanruw 1967. Pagan: Fossil, Corwin lD2(F)7 (USNM). Alamagan: 200 m, Anderson 404 (US) (Sterile). Epiphytic. Endemic in Marianas. Alamagan: Above Partido Village, Fosberg 31 677 Sarigan: Falanruw 1756; above village, 150-250 Taeniophyllum mariannense Schlechter (US) * m, Evans 2400 (US). CASUARINACEAE Casuarina equisetifolia L. Becomes established on open mineral substrata; lava, scoria and ash, also in grassland, forms forests in a few years, quite susceptible to fire, rather uni- form stands of various ages on relatively recent volcanic areas, notably on Pagan. Large groves on east end of north coast of Anatahan and on east coast of Agrigan. In all likelihood native in the Northern Marianas, but this is by no means estab- lished. A more thorough search in consolidated volcanic ash deposits on all the islands might throw light on this question. No Casuarina was found in the small fossil plant collection made on Pagan (Fosberg and Corwin, 1958). Uracas: Planted by Fritz in 1900 (Fritz, 1902, p. 116). Maug: Small group seen on top of East Island by Falanruw in 1972. Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31578 (US, BISH, Fo, NY, L). Pagan: Hosokawa 7949 (US, Fo); Mt. Pagan, Moore 371 (US); Bonham 20 (US, Fo); Kondo in 1949 (BISH); isthmus near pier west side of island, Anderson 580 (US, BISH, Fo, NY, L); Mt. Charlen, east of Fresh Water Lake, Fosberg 31411 (US, BISH, Fo, NY, L); Laguna, Villagomez JV-02 (US). Alamagan: Around Partido village, Fosberg 31646 (US). Guguan: Planted by Fritz in 1900, Fritz, 1902, Anatahan: Beach area, Falanruw 1660 (US); northwest tip of island, 0-10 m, Euans 2447 (US, BISH, Fo, NY). p. 101. Medinilla: Fritz, 1902, p. 97. PIPERACEAE Peperomia mariannensis C. de Candolle f. mariannensis In semishaded rock crevices, on vertical scoria cliffs and wooded banks. Endemic to Marianas. Agrigan: Canyon east side of island, Fosberg Pagan: Hosokawa 7987 (A). Alamagan: Hosokawa 7908 (A); south southwest coast, 1025 ft [310 m], Falanruw 1923 (US); Partido village, Fosberg 31678 (US, BISH, Fo, DPU); An- derson 408 (US, BISH, NY, DPU, L); Fosberg 31674 (US, BISH, Fo, NY, DPU); Hosokawa 7908 (BISH). Sarigan: Near anchorage, 700 ft [215 m], Falan- ruw 1797 (US); 325 ft [I00 m], Falanruw 1746 (US); forest above village, 250-300 m, Euans 2381 (US). "Piper betle L. Climbing on trees in forest at higher elevations, probably of aboriginal introduction. Indo-Pacific, mostly associated with man, as the leaves are used in betel-chewing. Agrigan: Nosokawa 8009 (A); seen but not col- lected by Falanruw in 1972. Alamagan: South southwest coast, 500 ft [I50 m], Falanruw 1914 (US); 650-1000 ft [ZOO-300 m], 1929 (US); Anderson 418 (US); around Partido village, Fosberg 31667 (US). Sarigan: Above village, 350-375 m, Evans 2371 (US, BISH, Fo, NY). Piper guahamense C. de Candolle 31621 (US, BISH, Fo, NY, DPU). NUMBER 22 21 Found in gorge bottom. Endemic to Marianas. Agrigan: Hosokawa 8011 (BISH); canyon on east side of island, Fosberg 31620 (US, BISH, Fo, NY, DPU). ULMACEAE Trema orientalis var. argentea (Planchon) Lauterbach Generally distributed small to medium-sized tree in forests, thickets, semi-open ravines and even scattered in Miscanthus grassland on open slopes. India to Japan, Bonins and Marianas, in its extreme form in the Bonins and Marianas, inter- grading with other varieties in Asia. Maug: Seen but not collected by Falanruw in 1972. Asuncion: Southwest slope, 150 ft [45 m], Falan- ruw 2252 (US). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31602 (US). Pagan: Air strip, Moore 304 (US); Fresh Water Lake, Fosberg 31371 (US); Anderson 530 (US); 150 ft [45 m], Bonham 12 (US). Alamagan: South southwest corner of island, bebw 500 ft [150 m], Falanruw 1940 (US); around Partido village, Fosberg 31669 (US). Guguan: West coast, 200 ft [60 m], Falanruw I818 (US); 200-300 ft [60-90 m], 1843 (US). Sarigan: Near anchorage, 400 ft [120 m], Falan- ruw 1758 (US); above village, 250-300 m, Evans 2386 (US, BISH, Fo, NY, L, P); 10-100 m, Evans 2352 (US, BISH, Fo, NY, L, TI). Anatahan: Northwest corner of island, 350 ft [110 m], Falanruw 1625. Trema orientalis var. viridis Lauterbach Pagan: Caldera Wall, near airstrip, Moore 332 (US). MORACEAE ?Artocarpus altilis (Parkinson) Fosberg Found about villages and in forests in canyons. Putative hybrids with A. mariannensis seen by Falanruw on all inhabited islands from Agrigan southward. Artocarpus altilis, being sterile, must have been planted by people. The hybrid may have originated locally or may have been carried from island to island by fruit{ bats. Artocarpus altilis is pantropical in cultivation, possibly of New Guinea origin. Asuncion: Lehne and Gabler, 1972, p. 39, as ?Brotfruchtbaume,? Agrigan: Canyon on east side of island, Fosberg 31634 (US). Pagan: Isthmus, Anderson 579 (US, BISH, Fo, NY, L, Ca); 150 ft [45 m], Bonham 26 (US, Fo). Sarigan: Village, 10-50 m, Evans 2439 (US, BISH, Fo, NY). Alamagan: South southwest coast, 500 ft [I50 m], Falanruw 1935 (US). Artocarpus altilis X mariannensis ft [60-90 m], Falanruw 1630 (US). Artocarpus mariannensis Trecul In wooded canyon, probably found on most is- lands from Agrigan south, but no specimens avail- able. Found wild on Marianas and Palau, planted else- where. Alamagan: Vicinity of Asongsong village, 15 m, Fosberg 31736 (US). ?Ficus benghalensis L. Doubtless planted. South Asian. Pagan: Moore 347 (US). Small, stunted trees on volcanic cinder sand crest. Indo-Pacific. Guguan: West coast, 200-225 ft [60-70 m], Falan- ruw 1822 (US). Ficus prolixa Forster f. Both varieties of this are found together on some islands, possibly on all of the wooded islands, in woods and in open. Field studies are in order to ascertain whether these varieties are really distinct, or only phenotypes sorted out from a variable population. Anatahan: Northwest corner of island, 200-300 Ficus microcarpa var. Zatifolia (Miquel) Corner Pacific; varieties Micronesian. Pagan: Corner, 1965, p. 10. Asuncion: Southwest slope, 150 ft [45 m], Falan- Agrigan: Marche 291 (P, Fo). Pagan: 150 ft [45 m], Bonham I7 (Fo). Ficus prolixa var. carolinensis (Warburg) Fosberg TUW 2255 (US). 22 SMITHSONIAN CONTRIBUTIONS TO BOTANY Alamagan: Near Asongsong village, Fosberg 31738 (US, BISH, Fo, NY, CA), Fosberg 31737 (US, BISH, Fo, CA); near Partido village, Fosberg 31684 (US, BISH, CA). Guguan: 320 ft [lo0 m], Falanruw 1836 (US); 200-300 ft [60-90 m], Falanruw 1828 (US). Sarigan: Near anchorage, 250 ft [75 m], Falanruw 1729 (US); near village 10-100 m, Evans 2351 (US, BISH, Fo, NY). Medinilla: ?Lehne and Gabler, 1972, p. 20, as ?Nunok.? Ficus prolixa var. subcordata Corner Pagan: Anderson 508 (US, BISH, CA); near crater lake [=Freshwater Lake], 25 m, Anderson 521 (US, BISH, Fo, NY, CA). Alamagan: South southwest coast, 250 ft [75 m], Falanruw 1877 (US). Sarigan: Near anchorage, 425 ft [130 m], Falan- ruw 1755 (US); 250 ft [75 m], Falanruw 1734 (US). Ficus safordii Merrill Pagan: South of Fresh Water Lake, Fosberg 31392 (US, BISH, CA). Ficus tinctoria Forster f. Small tree in open and semi-open areas, climbing in forest and on rocks. Plants with acuminate leaves called var. neo-ebudarum. This distinction should be studied in the field to see if it holds up. Pacific. Maug: North end of East Island, below 400 ft [120 m], Falanruw 2211 (US). Agrigan: Midwest coast, 200 ft [60 m], Falanruw 2320 (US); trail around south side of island, 10-20 m, Fosberg 31612 (US). Pagan: Talague beacl., Moore 343 (US); Caldera Wall, Moore 316 (US). Alamagan: Vicinity of Asongsong village, 3-1 5 m, Fosberg 31733 (US). Sarigan: Near village, 10-100 m, Evans 2355 (US? BJSH, Fo, NY); Evans 2339 (US, BISH, Fo, NY). Anatahan: West coast, 200 m, Fulanruw 1701 (US); northwest corner of island, 200-300 ft [60- 90 m], Falanruw 1629 (US), 1603 (US). Ficus tinctoria var. neo-ebudarum (Summerhayes) Fosberg Agrigan: Hosokawa 7997 (A); trail around north side of island, 10-20 m, Fosberg 31612 (US, BISH, Fo, NY, L). Pagan: Hosokawa 7962 (Fo); Bonham 14 (US, BISH); Fresh Water Lake, Fosberg 31384 (US, BISH, Fo); isthmus, Anderson 537 (US, BISH, Fo, NY, L). Alamagan: Hosokawa 7920 (Fo); around Partido village, Fosberg 31656 (US, BISH); near Asongsong village, 3-5 m, Fosberg 3173? (US, BISH). URTICACEAE Boehmeria celebica Blume one of the 2 following species. Boehmeria densiflora Hooker & Arnott and stunted specimens in caldera of Pagan. Not seen by us. Possibly should be referred to Alamagan: Kanehira, 1935a, pp. 308-309. Occasional bushes in upper forest of Alamagan Indo-Pacific. Pagan: Inside Mt. Pagan, Villagomez in 1970 (US), JV4K (US); Mt. Pagan, 1870 ft [570 m], Moore 367 (US), Moore 382 (US), Moore 393 (US); summit of Mt. Pagan, Villagomez JV-X-14 (US). Alamagan: South southwest coast, 500 ft [150 m], Falanruw 1908 (US). Boehmeria virgula (Forster f.) Guillemin Frequent small tree in semishaded moist ravine. Indo-Pacific. Agrigan: East side of island, Fosberg 31619 (US). Alamagan: Anderson 425 (US). Anatahan: Northwest corner of island, below 200 ft [60 m], Falanruw 1605 (US). Pipturtis argenteus (Forster f.) Weddell Occasional in open and semi-open areas, com- mon, tending to be dominant understory shrub or small tree in forest and thickets. Indo-Pacific. Maug: North end of East Island, below 400 ft [120 m], Falanruw 2227 (US). Asuncion: Southwest slope, 150 ft [45 m], Falan- ruw 2253 (US). Agrigan: Trail along south side of island, 10- 20 m, Fosberg 31586 (US); midwest coast, 200 ft [60 m], Falanruw 2324 (US). Pagan: Isthmus, Anderson 534 (US, BISH, Fo, NY); on plain, 150 ft [45 m], Bonhum 31 (US, BISH); Mt. Pagan, 200 ft [60 m], Moore 386 (US). Alamagan: Around Partido village, Fosberg 31666 (US, BISH, Fo, NY); south southwest coast, 650-1000 ft [200-300 m], Falanruw 1930 (US). NUMBER 22 23 Guguan: West coast, 225 ft [70 m], Falanruw 1821 (US). Sarigan: Above village, 150-250 m, Evans 2393 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, below 150 ft [45 m], Falanruw 1655 (US); western coast, below 200 m, Falanruw 1700 (US), 1703 (US); south of northwest tip of island, 0-10 m, Evans 2448 (US, BISH, Fo, NY). BALANOPHORACEAE Balanophora fungosa var. mariannae (Hosokawa) Hosokawa A fleshy parasite on roots of other plants. Not seen by us but this may be because its flowering is seasonal and it is subterranean during the vegeta- tive stage. Species Indo-Pacific; variety endemic in Mari- anas. Alamagan?: Hosokawa, 1937, p. 202. Sarigan: Hosokawa, 1937, p. 202. POLYGONACEAE "Antigonon leptopus Wight & Arnott a climber. Planted and occasionally established in thickets; American. Agrigan: Southwest coast village area, below 300 ft [90 m], Falanruw 2154 (US). B ATIDACEAE "Batis spp." record!] Medinilla: Swedberg, MS 1974 [entirely unlikely AMARANTHACEAE 'Achyranthes aspera L. Established along trails and in disturbed places. Pantropical, probably of southeast Asiatic origin. Agrigan: Trail around south side of island, 10- Pagan: Talague beach, Moore 338 (US). Not seen by us. Southeast Asian. Sarigan: Kanehira 2169 (FU). 20 m, Fosberg 31596 (US). Achyrant hes b iden tutu Blume Achyranthes canescens R. Brown In forest and on dry open slopes. Australian-Pacific when taken in a broad sense. Maug: Northern end of East Island, below 40 Asuncion: Southwestern slope, 250 ft [75 m], Agrigan: Trail around south side of island, 10- ft [12 m], Falanruw 2210 (US). Falanruw 2272 (US). 20 m, Fosberg 31600 (US). "Amaranthus dubius Martius ex Thellung Weed around village. Pantropical. Alamagan: Around Partido village, Fosberg 31690 (US). *Amaranthus spinosus L. Weed in disturbed and open places. Pantropical. Agrigan: Village, Fosberg 31432 (US). Pagan: Near Salt Lake, west side of island, 2-50 Anatahan: Kanehira 2147 (K). Common weed in open and semi-open areas. Pantropical. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2160 (US). Alamagan: South southwest coast, 0-200 ft [O- 60 m], Falanruw 1947 (US); around Partido village, Fosberg 31653 (US). Sarigan: Near anchorage, 250 ft [75 m], Falanruw 1732 (US); below 25 ft [lo m], Falanruw 1786 (US); near village, 10-100 m, Evans 2358 (US, BISH, Fo, NY); near anchorage, 325 ft [lo0 m], Falanruw 1745 *Celosia argentea L. vation. m, Fosberg 31363 (US, BISH, Fo, NY). "Amaranthus viridis L. (US). Planted around dwellings. Widespread in culti- Agrigan: Around village, 10-20 m, Fosberg 31562 (US)* Deeringia amaranthoides (Lamarck) Merrill great lava boulders. Rare, climbing over other vegetation on bluff of Indo-Pacific. Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31598 (US, BISH, Fo, NY, L). "Gomphrena globosa L. Planted ornamental around dwellings. 24 SMITHSONIAN CONTRIBUTIOXS TO BOTANY Widely cultivated, probably of tropical Ameri- Agrigan: Around village, 10-20 m, Fosberg 31561 Sarigan: Village, 10-50 m, Evans 2436 (US, BISH, can origin. (US). Fo, NY). NYCTAGINACEAE ?Boerhavia diffusa L. Weed of disturbed open places. South Asian. Sarigan: North of village, 50-175 m, Evans 2421 (US, BISH, Fo, NY). Boerhavia repens L. Common on ledges and in rock crevices. Prob- ably also on other islands. Indo-Pacific and African. Maug: North end of East Island, below 400 ft [I20 m], Falanruw 2219 (US). Boerhavia sp. bluffs, closely related to last species. [120 m], Falanruw 2225 (US). lanruw 1777 (US). identical with Maug and Sarigan plants]. Pisonia grandis R. Brown very soft. Creeper, fairly frequent on coastal slopes and Maug: North end of East Island, below 400 ft Sarigan: Northwest coast, 0-75 ft [O-20 m], Fa- Medinilla: Swedberg MS 1974 [not necessarily Common small to medium trees in thickets, wood Indo-Pacific. Maug: South central East Island, 500 ft [150 m], Falanruw 2238 (US), Falanruw 2237 (US). Sarigan: Near anchorage, below 25 ft [lo m], Falanruw 1787 (US); 250 ft [75 m], Falanruw 1735 (US), Falanruw 1725 (US); south of anchorage on northwest coast, 200 ft [60 m], Falanruw 1782 (US); north of village, 175-250 m, Evans 2411 (US, BISH, Fo, NY). AIZOACEAE Sesuuium portulacastrum L. Prostrate very fleshy halophyte forming mats in strand locations. On Maug seen only around aban- doned Japanese installation. Pan tropical. Maug: North end of East Island, below 400 ft Pagan: East coast, south of Togari Peninsula, [120 m], Falanruw 2228 (US). Moore 419 (US); isthmus, Anderson 548 (US). PORTULACACEAE Portulaca australis Endlicher (?Portulaca samoensis von Poellnitz) Small fleshy plant with tuberous root, growing ii1 open coastal areas. Indo-Pacific. Maug: North end of East Island, below 400 ft [120 m], Falanruw 2223 (US), Falanruw 2229 (US). Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2167 (US); trail around south side of island, 10-20 m, Fosberg 31609 (US). Sarigan: Seashore below village, 0-20 m, Evans 2432 (US, BISH, Fo, NY). Anatahan: Beach area, sea level, Falanruw 1662 (US); northwest tip of island, 0-10 m, Euans 2463 (US, BISH, Fo, NY). ?Portulaca grandifiora Hooker Cultivated. A widespread ornamental species. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2150 (US). Portulaca lutea Solander ex Forster f. Very fleshy herb, found only on Uracas and Maug, on rocks, most luxuriant where shaded, smaller, less fleshy, purple-tinged where more ex- posed. The species is usually associated with sea- bird colonies and may be dispersed as seeds cling- ing to birds feet or feathers. Micronesian and Polynesian, with a variety in the Galapagos. Uracas: South coast, below 400 ft [120 m], Falan- ruw 2200 (US). Maug: North end of East Island, sea level, Falanruw 2232 (US). Portulaca oleracea L. Common, fleshy weed in gardens and disturbed places, and especially in open coastal situations, often with P. australis. Pantropical and warm-temperate in many va- rieties, difficult to tell if indigenous or introduced in a particular area. Maug: South central East Island, 15 ft [5 m], Falanruw 2244 (US). NUMBER 22 25 Agrigan: Southwest coast village area, below 300 fc [90 m], Falanruw 2166 (US); trail around south side of island, 10-20 m, Fosberg ?1582 (US). Pagan: East coast, south of Togari Peninsula, Moore 422? (US); isthmus, Anderson 570 (US); near Salt Lake, west side of island, Fosberg 31?65 (US). Alamagan: Around Partido village, Fosberg ?1642 Sarigan: Northwest coast, 0-75 ft [O-20 m], Fa- lanruw 1795 (US); near anchorage, sea level, Falan- ruw 1990 (US); near dock, 0-20 m, Evans 2405 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, 25 ft [lo m], Falanruw 1672 (US). "Portulaca suflrutescens Engelman Frequent in cultivated land but only in one place; obviously a recent introduction, but puzzling as it is not known elsewhere in the Pacific. Mexican and southwest U.S., from desert and semi-arid areas. Sarigan: In village, 10-50 m, Evans 2435 (US, UH, Fo, MO, K). "Portulaca spp." (US). Medinilla: Swedberg MS 1974. ANNONACEAE "Annonu muricata L. Semicultivated near village. Widespread tropical cultivated species. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2171 (US). "Annonu reticulata L. Rare, in coconut plantation near village. Pantropical cultivated species. Alamagan: Vicinity of Asongsong village, 3-1 5 m, Fosberg ?17?4 (US). "Annonu squamosa L. Several trees near village. Pantropical cultivated species. Alamagan: South southwestern coast, 250 ft [75 m], Falanruw 1873 (US). Guamia mariannae (Safford) Merrill Small tree in native forest on lava. Endemic to Marianas. Pagan: Kanehira 2199 (FU, US); East of village, Moore 410 (US); road to Inae Dikiki or Togari rock area, below 100 ft [30 m], Falanruw 1855 (US); Anderson 507 (US). Alamagan: Around Partido village, Fosberg ?1662 (US) * LAURACEAE Cassytha filiformis L. String-like parasitic vine climbing and forming yellowish tangles over other plants, on open cinders on Guguan, found as high as 350 m on Asuncion. Pantropical. Asuncion: Southwestern slope, 1100 ft [335 m], Pagan: East of village, Moore 410 (US). Alamagan: Around Partido village, Fosberg 31685 Guguan: West coast, below 50 ft [15 m], Falan- Falanruw 2295 (US). (US, BISH, Fo). ruw 1805 (US). HERNANDIACEAE Hernandia sonora L. Infrequent medium-sized tree in ravine. Pantropical. Maug: Fritz, 1902, p. 114. Agrigan: Midwest coast, 200 ft [60 m], Falanruw 2323 (US); canyon on east side of island, Fosberg 31637 (US). Anatahan: Northwest corner of island, 200-300 ft [60-90 m], Falanruw 1628 (US). CAPPARIDACEAE Capparis cordifolia Lamarck Abundant in patches in rocky open coastal areas. Maug: North end of East Island, below 400 ft Pagan: East side of isthmus, Anderson 547 (US, Sarigan: Near anchorage, 250 ft [75 m], Falan- Anatahan: Beach area, sea level, Falanruw 1667 [120 m], Falanruw 2226 (US). BISH, Fo); east coast, Moore 421 (US). ruw 17?1 (US). (US). "Cleome viscosa L. tropical weed. 60 m], Falanruw 1948 (US). Occasional herb on rocky beach. Widespread Alamagan: South southwest coast, 0-200 ft [O- 26 SMITHSONIAN CONTRIBUTIONS TO BOTANY LECUMINOSAE *Abrus precatorlus L. shrubs in forest and in open areas. Common vine, tangled in and over trees and Pantropical. Agrigan: Trail around south side of island, Fos- berg 31616 (US); southwest coast, below 300 ft [90 m], Falanruw 2168 (US); midwest coast, 150 ft [45 m], Falanruw 2340 (US). Pagan: Marche 152 (P); Road to Talague beach, Moore 336 (US); Fresh Water Lake, in forest, Fos- berg 31389 (US): 100 m east of Japanese airfield, center of isthmus, 18 m, Anderson 574 (US, BISH, Fo, NY). Alamagan: Around Partido village, Fosberg 31 695 (US); south southwest coast, 250 ft [75 m], Falanruw 1872 (US); Songsong [Asongsong village], 100 ft [30 m], Kondo in 1949 (BISH). Anatahan: Northwest corner of island, below 200 ft [60 m], Falanruw 1608 (US). *Acacia confusa Merrill In village area, planted by Japanese. Native of Taiwan. Sarigan: Village, 10-50 m, Evans 2443 (US, BISH, Fo, NY). "Acacia farnesiana (L.) Willdenow Planted in village. Of Mexican origin. Agrigan: Village, Fosberg 31427 (US). Occasional herbaceous vine growing in open Indo-Pacific. Pagan: Air strip, Moore 305 (US). Alamagan: Around Partido village, Fosberg 31 689 Guguan: 320 ft [I00 m], Falanruw 1839 (US). Occasional tree along road. Pagan: Village, Moore 407 (US); road to Inae Dikiki, 100-200 ft [30-60 m], Falanruw 1869 (Us). 'Calopogonium mucunoides Desvaux Alylosia scarabaeoides (L.) Bentham area. (US, BISH, Fo). 'Bauhinia monandra Kurz Tangled vine, forming large patches. Sarigan: Near anchorage, 300 ft [90 m], Falan- ruw 1760f (US); near village 10-100 m, Evans 2350 (US, BISH, Fo, NY). Canavalia megalantha Merrill var. Vine climbing in forest and creeping on vol- Species endemic to Marianas, the variety to Pagan: Plain of volcanic ash soil, Bonham 37 Alamagan: Around Partido village, Fosberg Guguan: 200-300 ft [60-90 m], Falanruw 1829 Sarigan: Track leading north from village, 175- 250 m, Evans 2416 (US, BISH, K, MO), Evans 2418 (US, BISH, K, NY). "Cassia data L. canic soil, up to 100 m. Northern Marianas. (US). 31655 (US, BISH, Fo, WIS, L). (US). Planted about dwellings. Widely planted in tropics, of American origin. Agrigan: Cultivated in village, 10-20 m, Fosberg 31563 (US); southwest coast village area, below 300 ft [90 m], Falanruw 2172 (US). *Cassia Zeschenaultiana de Candolle Weed in open places. Pagan: s.L, Anderson 602 (US). Weed in open places. Pagan: s.L, Kanehira 2202 (NY, FU). Weed in disturbed places, roadsides. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2152 (US). Pagan: Isthmus, Anderson 526 (US, BISH, Fo, NY, L); ash beds of isthmus, Fosberg 31352 (US); east of Lake Laguna, Moore 348 (USj. "Cassia mimosoides L. "Cassia occidentalis L. Alamagan: 170 m, Anderson 410 (US). Sarigan: Village, 10-50 m, Evans 2441 (US, BISH, Fo, NY). *Cassia sophera L. Weed about villages. Alamagan: South southwest coast, 650 ft [200 m], Falanruw 1933 (US): vicinity of Asongsong village, 3-15 m, Fosberg 31716 (US). Anatahan: Northwest corner of island, below 250 ft [75 m], Falanruw 1675 (US). "Clitoris ternatea L. Vine tangled in vegetation about villages and disturbed places. Pan tropical. Alamagan: Above Asongsong village, 3-1 5 m, NUMBER 22 27 Fosberg 31727 (US); south southwest coast, landing and village area, 0-200 ft [0-60 m], Falanruw 1951 "Crotalaria pallida Aiton (Crotalaria striata de Candolle, Crotalaria mucronata Desvaux) Generally common in disturbed places, around villages, and on open volcanic ash beds, up to 150 m. (US). Pantropical weed. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2151 (US, BISH); trail around south side of island, 10-20 m, Fosberg 31589 (US). Pagan: Near crater lake, 15 m, Anderson 518 (US, BISH, Fo, NY); near Salt Lake west side of island, Fosberg 31356 (US); village, Moore 406 Alamagan: South southwest coast, 450 ft [140 m], E'alanruw 1897 (US); vicinity of Asongsong village, 3-15 m, Fosberg 31730 (US). Sarigan: Near anchorage, 500 ft [90 m], Falan- ruw 1759 (US). "Crotalaria trifoliastrum Willdenow (US). Not seen by us. Pagan: Kanehira, 1935a, p. 330; Hosokawa, 1935a, p. 18. Cynometra ramiflora L. material, trunks to over a meter in diameter. Grove of large trees at top of ridge of volcanic Indo-Pacific. Sarigan: Near anchorage, 400 ft [120 m], Falan- ruw 1751 (US). Medinilla: Lehne and Gabler, 1972, p. 20, as "Gulus." Dalbergia candenatensis (Dennstedt) Prain Climbing in vegetation in moist forested ravine at about 75 m elevation, elsewhere in Western Micronesia found at back of mangrove swamps. Indo-Pacific. Anatahan: West end of north coast, below 250 ft [75 m], Falanruw 1682 (US). *Delonix regia (Bojer) Rafinesque Infrequent tree near village areas. Native of Madagascar, cultivated throughout Pagan: Plain of volcanic ash, 150 ft [45 m], Bon- Anatahan: Northwest corner of island, below tropics. ham 18 (US, Fo). 200 ft [60 m], Falanruw 1633 (US). *Desmanthus virgatus (L.) Willdenow Occasional weed in areas behind beach. Pantropical weed, of American origin. Agrigan: Southwest coast, village area, below 300 ft [90 m], Falanruw 2196 (US). 'Desmodium canum (Gmelin) Schinz & Thellung Weed in village. Pantropical weed. Anatahan: Village area, northwest corner of is- land, below 200 ft [60 m], Falanruw 1643 (US). 'Desmodium gangeticum (L.) de Candolle Weed in village and along trails. Old World tropics. Anatahan: Northwest corner of island, below 250 ft [75 m], Falanruw 1681 (US); northern coast, be- low 250 ft [75 m], Falanruw 1692 (US). 'Desmodium heterophyllum de Candolle Small herb of open places, easily mistaken for the next species. Widespread. Alamagan: Kanehira 2183 (NY). *Desmodium triflorum (L.) de Candolle Abundant small prostrate herb of open sunny habitats; on Guguan a pioneer with Cassytha, on bare volcanic cinder sand. Pantropical. Asuncion: Southwest slope, 1500 ft [460 m], Fa- lanruw 2304 (US). Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2189 (US); trail around south side of island, 10-20 m, Fosberg 31573 (US). Pagan: Marine camp, 20 m, Andenon 511 (US, BISH, Fo, NY, L); near Salt Lake, Fosberg 31351 (US); Mt. Pagan, Villagomez JV-5K (US); Mt. Pagan, Moore 381 (US), 378 (US). Guguan: West coast, below 50 ft [I5 m], Falnn- ruw 1808 (US). Alamagan: Vicinity of Asongsong village, 3-15 m, Fosberg 31748 (US); south southwest coast, 450 ft [140 m], Falanrziw 1901 (US). "Dolichos lablab L. m, Fosberg 31728 (US). 200 ft [60 m], Falanruw 1674 (US, BISH). Erythrina uariegata var. orientalis (L.) Merrill Alamagan: Vicinity of Asongsong village, 3-1 5 Anatahan: Northwest corner of island, below Prickly tree, leafless or almost so at flowering 28 SMITHSONIAN CONTRIBUTIONS TO BOTANY time; occasional scattered in open or semi-open areas and in woods from inland of coast to higher elevations, reaching nearly 500 m on Asuncion. Indo-Paci fic. Asuncion: Southwest slope, 750 ft [225 m], Falan- ruw 2286 (US). Agrigan: Midwest coast, 200 ft [60 m], Falanruw 2317 (US); trail around south side of island, 10- 20 m, Fosberg 31570 (US). Pagan: Caldera Wall, Moore 428 (US); isthmus, 10 m, Anderson 577 (US). Alamagan: South southwest coast, below 500 ft [150 m], Falanruw 1937 (US); around Partido village, Fosberg 31671 (US). Guguan: 320 ft [lo0 m], Falanruw 1834 (US). Sarigan: Near anchorage, 400 ft [120 m], Falan- ruw 1748 (US). "Glycine clandestina Wendland Not seen by us. Pagan: Hosokawa, 1935a, p. 18; Kanehira, 1935a, p. 334. "Zndigofera tinctoria L.? Scattered weedy shrubs along village road. Pantropical. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2181 (US). "Leucuenu latisiliqua (L.) Gillis & Stearn (L. glauca sensu auct. plur., non (L.) Bentham; L. leucocephala (Lamarck) de Wit) This small tree or shrub is a recent introduction, not recorded from Pagan in 1958 but common about village area by 1972. On Agrigan said not to have been on the island before World War I1 but now common in areas that were bulldozed. Of American origin but now pantropical. Agrigan: Southwest coast, below 300 ft [90 m], Pagan: Village, Moore 399 (US). Sarigan: Village and vicinity, 20-50 m, Euans Falanruw 2188 (US). 2407 (US, BISH, Fo, NY). Mucuna gigantea (Willdenow) de Candolle seashore, to 250 m. Vine climbing in thickets, occurring well above Pantropical, distributed by currents. Agrigan: Midwest coast, 200 ft [60 m], Falanruw Pagan: Kanehira 2207 (FU, NY). Sarigan: North of village, 175-250 m, Euans 2325 (US). 2413 ? (US, BISH, NY). Mucuna platyphylla Gray tions. Large vine growing over trees at higher eleva- Pacific. Sarigan: Near anchorage, 600 ft, Falanruw 1757 Anatahan: Seen, but not collected by Falanruw. Common around edges of village, climbing over American, widely introduced in Old World Alamagan: Asongsong village, 3-1 5 m, Fosberg (US). "Pachyrrhizus erosus (L.) Urban other vegetation. tropics. 31717 (US, BISH, Fo). 'Phaseolus vulgaris L,? pp. 20, 27 as "Bohnen," planted in 1901. "Pithecellobiurn duke (Roxburgh) Bentham vations. Guguan and Medinilla: Lehne and Gabler, 1972, Planted and naturalized at low to moderate ele- Pantropical but of American origin. Asuncion: South southwest coast, 0-200 ft [O- 60 m], Falanruw 2293 (US). Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2195 (US); around village, 10-20 m, Fos- berg 31564 (US). Pagan: Vicinity of landing pier, 3 m, Anderson 594 (US, BISH, Fo); Caldera Wall, east of airstrip, Moore 312 (US); 50 ft [I5 m], Bonham 23 (US, Fo). Alamagan: Vicinity of Asongsong village, 3-15 m, Fosberg 31747 (US, BISH, Fo); south southwest coast, 0-200 ft [O-60 m], Falanruw 1945 (US). Sarigan: Near village, 10-100 m, Evans 2338 (US, BISH, Fo, NY). Vigna marina (Burman) Merrill Common vine, normally of sea-beaches but in Northern Marianas common inland and to higher elevations, 500 m on Asuncion. Indo-Pacific, possibly also in Caribbean. Asuncion: Southwestern slope of volcano, 1500 ft [450 m], Falanruw 2309 (US); 125 ft [40 m], Falanruw 2259 (US). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31604 (US): Marche 295 (P, Fo). Pagan: Kanehira 2204 (NY); isthmus, 10 m, An- derson 568 (US, BISH, Fo, NY); west side of Lake NUMBER 22 29 Laguna, Moore 356 (US); Inae Dikiki (Togari Rock Bay), below 100 ft [SO m], Falanruw 1857 (US). Anatahan: West end of north coast, below 250 ft [75 m], Falanruw 1676 (US); northwest corner of island, below 200 ft [60 m], Falanruw 1648 (US), Falanruw 1640 (US); northwest tip of island, 0- 10 m, Evans 2453 (US). OXALIDACEAE 'Oxalis corniculata L. In disturbed areas. Pantropical, in many forms and varieties. Maug: Northern end of East Island, below 400 Sarigan: Above village, 250-300 ft [75-90 m], ft "20 m], Falanruw 2224 (US). Evans 2387 (US). RUTACEAE "Citrus aurantifolia (Christmann) Swingle plantations. Planted or persisting in and around villages or Pantropical in cultivation of Old World origin. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2183 (US). Alamagan: Southwest coast, 750 ft [230 m], Fa- lanruw 1918 (US); around Partido village, Fosberg 31692 (US). Sarigan: Near village, 10-100 m, Evans 2334 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, below 200 ft [60 m], Falanruw 1601 (US). "Citrus reticulata Blanco Planted. Pagan: Lehne and Gabler, 1972, p. 33, as "Man- darinen." "Citrus sinensis (L.) Osbeck Small tree near village. Pantropical in cultivation, originally probably Agrigan: Southwest coast, below 300 ft [go m], from subtropical Asia. Falanruw 2181 (US). * Triphasia trifolia (Burman f.) P. Wilson Naturalized, forming thorny underlayer in forest. Tropical Asiatic, widely naturalized elsewhere Pagan: East of village, Moore 402 (US). in tropics. MELIACEAE Aglaia mariannensis Merrill Common small tree in thickets and forests, Endemic to Marianas. Agrigan: South southwest coast, 200 ft [60 m], Falanruw 2332 (US, BISH); canyon on east side of island, Fosberg 31631 (US). Pagan: Caldera Wall, Moore 315 (US); Anderson 504 (US); south of Fresh Water Lake, 2-50 m, Fos- berg 31386 (US); Anderson 603 (US). Alamagan: South southwest coast, 250 ft [75 m], Falanruw 1874 (US); around Partido village, Fos- berg 31679 (US); 200 m, Anderson 401 (US). Guguan: Southwest part of island, 175 ft [55 m], Falanruw 1847 (US). Sarigan: Northwest coast, 250 ft [75 m], Falan- ruw 1741 (US), Falanruw 1733 (US); 400-450 ft [120-140 m], Falanruw 1754 (US); north of village 175-250 m, Evans 2410 (US, BISH, Fo, NY); above village, 150-250 m, Evans 2399 (US, BISH, Fo, NY). EUPHORBIACEAE Drypetes dolichocarpa Kanehira Endemic to Marianas. Anatahan: Kanehira 2141 (US). Common weed in disturbed places, villages, and Pantropical weed. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2175 (US). Pagan: Isthmus, Anderson 552 (US, BISH, Fo); west side of island, near Salt Lake, Fosberg 31348 (US, BISH); village, Moore 395 (US). Alamagan: Around Partido village, Fosberg 31648 (US). Sarigan: Northwest coast, 75 ft [25 m], Falanruw 1790 (US); near village 10-100 m, Evans 2360 (US, RISH, Fo, NY). Anatahan: Northwest corner of island, below 200 ft [60 m], Falanruw 1639 (US). "Euphorbia prostrata Aiton Weed in path in former village. Pantropical weed. Pagan: Fresh Water Lake, 2-50 m, Fosberg 31380 (US, BISH, Fo, NY). "Euphorbia thymifolia L. Euphorb ia h irta L. gardens on inhabited islands. 30 SMITHSONIAN CONTRIBUTIONS TO BOTANY Weed in paths in and around villages. Pantropical weed. Pagan: Marine camp, 20 m, Anderson 515 (US); near Salt Lake, 2-50 m, Fosberg 31364 (US, BISH). Alamagan: South southwest coast, 0-200 ft [O- 60 m], Falanruw 1950 (US); vicinity of Asongsong village, 3-15 m, Fosberg 31714 (US). ?Euph orb ia spp. ? Medinilla: Swedberg, MS 1974. ?Jatropha curcas L. Common small tree or shrub about villages, said to have been brought by Japanese. Pantropical, associated with man. Agrigan: Southwest coast, below 300 ft [90 m], Pagan: Isthmus, 6 m, Anderson 528 (US, BISH, Falanruw 2180 (US). Fo, NY, L); Caldera Wall, Moore 322 (US). +]atropha gossypifolia L. Fleshy-stemmed shrub, leaves purplish, common about villages, dominant in some areas. Said to have been introduced by Japanese in 1930s. Native of tropical America, widely naturalized in Old World. Pagan: Isthmus, near shore, Anderson 566 (US); volcanic ash soil, 150 ft [45 m], Bonham 5 (US, Fo); near Salt Lake, west side of island, 2-50 m, Fosberg 31360 (US); village, Moore 405 (US). Sarigan: Village, 20-50 m, Evans 2406 (US, BISH, Fo, NY). Alamagan: South southwest coast, 0-200 ft [O- 60 m], Falanruw 1943 (US). Macaranga thompsonii Merrill Medium sized tree in moist forested area. Endemic in Marianas. Anatahan: Under 1000 ft [300 m], Falanruw 1969 (US); northwest corner of island, below 200 ft [60 1x1, Falanruw 1607 (US). ?Manihot esculenta Crantz (Manihot utilissima Pohl) Planted about villages for its edible tubers. Prob- ably introduced early by Spanish or Germans. Native of tropical America, widely cultivated. Agrigan: Fritz, 1902, p. 109. Pagan: Isthmus, Anderson 582 (US, BISH, Fo, Alamagan: Seen but not collected by Falanruw. Sarigan: Near village, 10-100 rn, Evans 2333 NY, L). (US). Anatahan: West coast, below 200 m, Falanruw 1707 (US). Melanolepis multiglandulosa var. glabrata (Mueller-Argoviensis) Fosberg Common in forest and thickets away from coast. Indo-Pacific, indigenous in Marianas. Asuncion: Southwest slope, 50 ft [I5 m], Falan- ruw 2274 (US). Agrigan: Midwest coast, 200 ft [60 m], Falanruw 2330 (US); trail around south side of island, 10-20 m, Fosberg 31603 (US). Pagan: Caldera Wall, Moore 323 (US); road to Inae Dikiki, 100-200 ft [30-60 m], Falanruw 1867 (US); isthmus, 6 m, Anderson 540 (US); Bonham August-September 1954 (US); Fresh Water Lake, 2-50 m, Fosberg 31381 (US). Alamagan: South southwest coast, 500 ft [150 m], Falanruw 1915 (US); around Partido village, Fosberg 31663 (US). Sarigan: Near anchorage, 250 ft [75 m], Falanruw 1736 (US); near village, 10-100 m, Evans 2344 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, below 200 ft (60 m], Falanruw 1606 (US). ?Phyllanthus amarus Schumacher & Thonning disturbed places, mostly at low elevations. Common weed in villages, roadsides, and other Pantropical weed. Maug: South central East Island, 100 ft [30 m], Falanruw 2233t (US). Asuncion: Southwest slope, 75 ft [25 m], Falan- ruw 2282 (US). Agrigan: Village, Fosberg 31433 (US, BISH); southwest coast, below 300 ft [90 m], Falanruw 2174 Pagan: Near Salt Lake, west side of island, Fos- berg 31349 (US); isthmus, Anderson 562 (US, BISH). Alamagan: Around Partido village, Fosberg 31650 (US); south southwest coast, 0-200 ft [O-60 m], Falanruw 1944 (US). Sarigan: Near anchorage, 1000 ft [300 m], FaZan- ruw 1720 (US); near village, 10-100 m, Evans 2359 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, below 200 ft [60 m], Falanruw 1635 (US). Phyllanthus marianus Mueller-Argoviensis (US). Common on lava in open or scrubby areas and NUMBER 22 31 edges of thickets. Endemic to Marianas, but with close relatives in Western Carolines. Maug: Central East Island, 100 ft [30 m], Falan- ruw 2243 (US, BISH). Asuncion: Southwest coast, 25 ft [lo m], Falan- ruw 2294 (US, BISH). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31608 (US). Pagan: Summit of Mt. Pagan, Villagomez JV- X-15 (US); Caldera Wall, Moore 319 (US); isthmus, 6-8 m, Anderson 542 (US); Fresh Water Lake, 0- 50 m, Fosberg 31402 (US). Alamagan: Around Partido village, Fosberg 31687 (US). Guguan: 320 ft [I00 m], Falanruw 1832 (US). Sarigan: Near village, 10-100 m, Evans 2348 (US, BISH, Fo, NY); above village, 375 m, Evans 2375 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, below 200 ft [60 m], Falanruw 1610 (US); west end of north coast, below 250 ft [75 m], Falanruw 1690 Phyllanthus safordii Merrill Not seen by us. Possibly a misdetermination of extreme forms of P. marianus. Except for these records, endemic to savanna areas of southern Marianas. Pagan: Hosokawa, 1935a, p. 21. Alamagan: Hosokawa, 1935a, p. 21. Anatahan: Hosokawa, 1935a, p. 21. Occasional in open village areas. Pantropical. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2153 (US); trail around south side of island, 10-20 m, Fosberg 31585 (US). Alamagan: South southwest coast, below 200 ft [60 m], Falanruw 1934 (US); vicinity of Asongsong village, 3-15 m, Fosberg 31729 (US). Sarigan: Near village, 10-100 m, Evans 2341 (US, BISH). Anatahan: West end of north coast, below 250 ft [75 m], Falanruw 1886 (US). (US). *Ricinus communis L. ANACARDIACEAE *Mangifera indica L. Large trees planted or persisting in village areas. Pantropical cultivated tree probably originally from Borneo according to Kostermans (pers. comm.). Pagan: Volcanic ash plain, 50 ft [15 m], Bonham 27 (US, Fo). Alamagan: Near Asongsong village, 3-15 m, Fosberg 31732 (US). Anatahan: Seen but not collected by Falanruw. SAPINDACEAE Dodonaea viscosa L. Small bushes scattered on open slope of volcano between 250 and 500 m, not seen lower down or on other islands of Northern Marianas. Pan tropical. Asuncion: Southwest slope, 900 ft [275 m], Falan- ruw 2292 (US, BISH). RHAMNACEAE Colubrina asiatica (L.) Brongniart In forest and thickets, coastal to over 150 m, reaching unusually high elevations compared with its occurrence elsewhere. Pan tropical. Maug: South central East Island, 500 ft [150 m], Falanruw 2234 (US). Agrigan: Trail around south side of island, 12- 20 m, Fosberg 31574 (US). Pagan: Isthmus, 8 m, Anderson 536 (US, BISH, Fo, NY, L); road to Talague beach, Villagomez in 1970 (US); Caldera Wall, east of air strip, Moore 318 (US). Alamagan: South southwest coast, 350 ft [ 110 m], Falanruw 1884 (US); around Partido village, Fosberg 31675 (US). Guguan: West coast, 200 ft [60 m], Falanruw 1819 (US); 200-300 ft [60-90 m], Falanruw 1827 Sarigan: Vicinity of village, 10-100 m, Evans 2349 (US, BISH, Fo, NY); northwest coast, 200 ft [60 m], Falanruw 1764 (US). (US). TILIACEAE Elaeocarpus joga Merrill Medium sturdy tree, scattered in forest at higher elevations on rough lava. Endemic in the Marianas and Palau. This has recently (Stone, 1971:402) been reduced to E. 32 SMITHSONIAN CONTRIBUTIONS TO BOTANY sphaericus (Gaertner) K. Schumann, but we have not yet had time to verify this reduction. Pagan: Kanehira, 1935a, p. 363. Alamagan: Trail from south southwest coast, 750 ft [225 m], Falanruw 1919 (US); vicinity of Asongsong village, Fosberg 31740 (US); Anderson 423 (US). Grewia crenala (L. f.) Schinz & Guillaumin Small tree in second story of forests and groves. Indo-Pacific. Pagan: Marche 151 (P, Foj; east coast, Moore 418 (US); Caldera Wall, Moore 426 (US); northwest Pagan, sea level, Falanruw 1854 (US); Anderson 503 (US); Fresh Water Lake, 2-50 m, Fosberg 31373 (US). ?Muntingia calabura L. Sarigan: Kanehira 2155 (FU). Common about village area. Tropical American, widely naturalized in Old Anatahan: Northwest corner of island, 300 ft World tropics. [90 m], Falanruw 1620 (US). Triumfetta procumbens Forster f. Creeper on beach sands. Indo-Pacific strand plant. Pagan: Inae Dikiki or Togari Bay, below 100 ft [30 m], Falanruw 1861 (US). *Triumfetta semitriloba Jacquin Tall weed in coconut plantations and in villages. Pan tropical. Agrigan: Fosberg 31419 (US, BISH, Fo); south- west coast, below 300 ft [90 m], Falanruw 2190 (US); 250 ft [75 m], 1879 (US). Pagan: Anderson 609 (US, BISH, Fo). Alamagan: Asongsong village, Fosberg 31726 (US); south southwest coast, 450 ft [140 m], Falan- ruw 1894 (US); around Partido village, Fosberg 31700 (US). M ALVACEAE ?Abutilon indicum Sweet Near village. Indo-Pacific. Sarigan: North of village 50-175 m, Evans 2425 (US, BISH, Fo, NY). +Gossypium hirsutum L. Persisting from planting. Pantropical, of American origin. Maug: Lehne and Gabler, 1972, p. 42, as ?Blii- Pagan: Isthmus, Anderson 581 (US). Occasional in open weedy village area. Pantropical. Anatahan: Northwest corner of island, below hende Baumwolle.? ?Hibiscus abelmoschus L. 200 ft [60 m], Falanruw 1646 (US). Hibiscus tiliaceus L. In thickets and ragged forests especially near sea, and around saltwater lake on Pagan: forming belt on slopes of Sarigan, very rare on Guguan. Pantropical. Maug: South central East Island, 200 ft [60 m], Falanruw 2240 (US). Asuncion: Lehne and Gabler, 1972, p. 39, as ?Hibiskus.? Agrigan: Trail around south side of island, Fos- berg 31569 (US, BISH). Pagan: Bonham 10 (US, Fo); isthmus near pier, Anderson 599 (US, Fo, BISH, NY, L); near lagoon, Anderson 593 (US, Fo, BISH, NY, L); near Salt Lake, west side of island, Fosberg 31343 (US); vil- lage, Moore 398 (US). Alamagan: Southwest coast, 450 ft [140 m], Fa- lanruw I899 (US); Partido village, Fosberg 13658 (US, BISH, Fo). Sarigan: Near anchorage, Falanruw 1785 (US, BISH); 250 ft [75 m], 1726 (US); village and vi- cinity, 20-50 m, Evans 2408 (US, BISH, Fo, NY); behind village, 50-100 m, Evans 2364 (US, BISH, Fo, NY). Anatahan: Beach area, sea level, Falanruw 1670 (US); Northwest tip of island, 0-10 m, Evans 2459 (US, BISH, Fo, NY). *Hibiscus ornamental hybrid Medinilla: Swedberg, his 1974. Planted, apparently did not persist. Agrigan: Fritz, 1902, p. 109. Alamagan: Fritz, 1902, p. 109. ?Malvastrum coromandelianum (L.) Garcke Common weed in open areas in villages. Pantropical, of American origin. Agrigan: Village, Fosberg 31436 (US). Alamagan: South southwest coast, below 200 ft [60 m], Falanruw 1938 (US); around Partido vil- lage, Fosberg 31691 (US, BISH, Fo). NUMBER 22 33 *Sida acuta Burman f. other disturbed places. Weed in plantations and around villages and Pan tropical. Pagan: Village, Moore 408 (US); near Salt Lake, west side of island, Fosberg 31362 (US): isthmus, Anderson 563 (US). Alamagan: South southwest corner of island, be- low 500 ft [150 m], Falanruw 1941 (US); around Partido village, Fosberg 31702 (US); vicinity of Asongsong village, 3-15 m, Fosberg 31719 (US). Sarigan: Near anchorage, 150 ft [45 m], Falan- ruw 1724 (US); village and vicinity, 10-50 m, Evans 2433 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, below 200 ft [60 m], Falanruw 1647 (US). *Sida rhombifolia L. Common weed in thickets and plantations. Pan tropical. Agrigan: Village area, Fosberg 31421 (US). Pagan: Village, Moore 400 (US), 403 (US): Marche 150 (P, Fo): near Salt Lake, west side of island, Fosberg 31361 (US). Sarigan: Near anchorage, 200 ft [60 m], Falan- ruw 1781 (US); near village, 10-100 m, Evans 2346 *Sida rhombifolia X acuta? (US). Along path in village area. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2185 (US). 'Sida spinosa L. Weed in village. Pan tropical. Alamagan: Vicinity of Asongsong village, 3-15 m, Fosberg 31720 (US). Thespesia populnea (L.) Solander ex Correa Pagan: Isthmus, Anderson 573 (US, BISH, NY); Talague Beach, Moore 342 (US). Alamagan: Around Partido village, Fosberg 31647 (US); south southwest coast, 0-200 ft [0-60 m], Falanruw I492 (US). Sarigan: Kanehira 2157 (FU, NY). Urena lobata L. Occasional in open and semi-open areas. Pan tropical. Sarigan: Near anchorage, 200 ft [60 m], Falanruw Alamagan: South southwest coast, 450 ft [140 m], I767 (US). Falanruw 1898a (US). ft [60 m], Falanruw 1636 (US). Anatahan: Northwest comer of island, below 200 BOMBACACEAE "Ceiba pentandra (L.) Gaertner Japanese. Persisting around villages from planting by Pantropical. Agrigan: Southwest coast, below 300 ft [90 m], Pagan: Anderson 610 (US, BISH, Fo, NY, L). Falanruw 2159 (US). STERCULIACEAE Melochia compacta Hochreutiner areas. At very low elevations in thickets and open Marianas and Carolines. Agrigan: Trail around south side of island, 2 m, Fosberg 31567 (US, BISH). Pagan: 8-10 m, Anderson 501 (US). Melochia villosissima (Presl) Merrill Along trails at low elevations. Marianas and Carolines. Pagan: Caldera Wall, Moore 427 (US): trail to Pansantati, below 100 ft [30 m], Falanruw 1864 (US). THEACEAE Eurya japonica var. nitida (Korthals) Thistelton- Dyer Eastern Asia to Micronesia. Agrigan: Hosokawa, 1935a, p. 31. Pagan?: Hosokawa, 1935a, p. 31. Alamagan: Kanehira 2190 (K). GUTTIFERAE Calophyllum inophyllum L. Uncommon. Indo-Pacific. Pagan: Near center of island, Anderson 597 (US, Alamagan: Fritz, 1902, p. 102. Medinilla: Swedberg, MS 1974. BISH, Fo, NY, L). 34 SMITHSONIAN CONTRIBUTIONS TO BOTANY PASSIFLORACEAE +PassifEma foetida var. hispida (de Candolle) Killip Weed in plantations and thickets, especially around disturbed areas and villages. Pantropical, but of American origin. Agrigan: Village, Fosberg 31424 (US, BISH, Fo); midwest coast, 200 ft [60 m], Falanruw 2327 (US). Pagan: Marine camp, 20 m, Anderson 510 (US, BISH, Fo); isthmus, Anderson 527 (US, BISH, Fo, NY); Bonham 3 (US, Fo); Fresh Water Lake, 2-50 m, Fosberg 31372 (US); east of village, Moore 404 (US); edge of air strip, Moore 309 (US). Alamagan: South southwest coast, 350 ft. [110 m], Falanruw 1885 (US). Sarigan: Near anchorage, 300 ft [90 m], Falan- ruw 1762 (US); 100 ft, 1717 (US) (sterile); north of village, 5b-175 m, Evans 2427 (US, BISH, Fo, NY). LYTHRACEAE CARICACEAE LCarica papaya L. anas except Uracas. Sparingly naturalized throughout Northern Mari- Pantropical, of American origin. Maug: South central East Island, 200-300 ft [60- 90 m], Falanruw 2349 (US). Asuncion: Southwestern slope, 150 ft [45 m], Falanruw 2260 (US). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31590 (US); narrow ravine, midwest coast, 150 ft [45 m], Falanruw 2347. Pagan: North of marine camp, Anderson 607 (US, BISH, Fo); Fresh Water Lake, Fosberg 31390 Alamagan: Around Partido village, Fosberg Guguan: Seen, but not collected by Falanruw. Anatahan: Western coast, below 200 m, Falan- Medinilla: Swedberg, MS 1974. (US). 31698 (US, BISH). ruw I704 (US). CUCURBITACEAE ?Benincasa hispida (Thunberg) Cogniaux Vine climbing in woody vegetation. Alamagan: South southwest coast, 500 ft [150 m], Falanruw 1917 (US); vicinity of Asongsong village, 3-15 m, Fosberg 31725 (US). Pemphis acidula Forster f. Locally very abundant on limestone area, prob- ably the only occurrence in the Northern Marianas except possibly on Medinilla. Pagan: East side of isthmus, Anderson 545 (US, BISH, Fo, NY, L); east coast, Moore 425 (US); Inae Dikiki or Togari Rock Bay, 100 ft [30 m], Falan- ruw I859 (US). These are probably all from the same locality, a very localized limestone outcrop. COMBRETACEAE *Terminalia catappa L. Large trees throughout Northern Marianas ex- cept Asuncion, Uracas, and possibly Medinilla. Dwarfed somewhat on Guguan. Generally in open country or in wooded ravines: provide roosting and nesting sites for boobies on Guguan and Maug. Pantropical. Maug: South central East Island, Falanruw 2235 Asuncion: Lehne and Gabler, 1972, p. 39, as ?Talisay? (possibly T. rostrata). Agrigan: Village area, southwest coast, below 300 ft [90 m], Falanruw 2173 (US); trail around south side of island, 10-20 m, Fosberg 31607 (US, BISH). Pagan: Isthmus, Anderson 533 (US, BISH, Fo, NY, L); Caldera Wall, east of air strip, Moore 311 Alamagan: Above Asongsong village, Fosberg Guguan: 200-300 ft [60-90 m], Falanruw 1825 Sarigan: Near anchorage, 250 ft [75 m], Falan- ruw I739 (US, BISH), I740 (US); village and vi- cinity, 20-50 m, Evans 2409 (US, BISH, Fo, NY). Anatahan: Northwest coast, Falanruw 1709 (US). Medinilla: Lehne and Gabler, 1972, p. 20, as (US). (US). 31743 (US). (US) * ?Talisay-Mandelbaum. ? Terminalia rostrata Fosberg & Falanruw One large tree seen. Apparently endemic on Asuncion. Asuncion: Southwestern slope, 400 ft [120 m], Falanruw 2290 (US). Terminalia samoensis Rechinger NUMBER 22 35 Common in low forest on lower slopes. Pacific, widely distributed usually strand species. Asuncion: Southwestern slope, 100 ft [30 m], Falanruw 2281 (US). LECYTHIDACEAE Barringtonia asiatica (L.) Kurz occurring more inland about village areas. Occasional to common tree in coastal areas, some Indo-Pacific strand trees. Agrigan: East side of island, Fosberg 31636 (US); Pagan: South of pier on west coast, 20 m, And- Alamagan: Partido village, Fosberg 31697 (US). Sarigan: Near anchorage, sea level, Falanruw 1783 (US); north of village, 50-175 m, Evans 2426 (US, BISH, Fo, NY). Anatahan: Beach area, sea level, Falanruw 1664 (US); northwest tip of island, 0-10 m, Evans 2462 (US, BISH). midwest coast, 200 ft [60 m], Falanruw 2344 (US). erson 589 (US, BISH, Fo, NY, L). MYRTACEAE Eugenia palumbis Merrill Possibly too close to E. reinwardtiana. Pagan: Hosokawa, 1940, p. 542. Eugenia reinwardtiana (Blume) de Candolle bluffs. Infrequent in thickets, low forests, and on coastal Indo-Pacific. Agrigan: Trail around south side of island, 10- Pagan: Caldera Wall, Moore 314 (US); Anderson Guguan: Southwest part of island, 175 ft [55 m], 20 m, Fosberg 31601 (US). 578 (US). Falanruw 1846 (US). *Psidium guajava L. village areas. Occasional groups of small trees about dry open Pantropical, of American origin. Pagan: Anderson 509 (US, BISH, Po). Sarigan: Behind village, 50-100 m, Evans 2366 Anatahan: Northwest corner of island, below (US, BISH, Fo, NY). 200 ft [60 m], Falanruw 1673 (US). MELASTOM ATACEAE Melastoma malabathricum L. var. (Melastoma niarianum Naudin). Scattered shrubs in open grassy areas. Indo-Pacific, variety endemic in Marianas. Pagan: Mt. Pagan, 1870 ft [570 m], Moore 365 (US), Moore 385 (US); Mt. Charlen spatter cone, east of Fresh Water Lake, Fosberg 31414 (US). Alamagan: Kanehira, 1935a, p. 383, Kanehira, Sarigan: Above village, 375 m, Evans 2378 (US, 1938, pp. 240-241. BISH, Fo, NY). ARALIACEAE Polyscias grandifolia Volkens In undergrowth in ravines and thickets. Micronesian. Pagan: Isthmus, 6-8 m, Anderson 541 (US, BISH); Caldera Wall, Moore 313 (US). Alamagan: Vicinity of Asongsong village, Fos- berg 31739 (US, BISH, Fo); Anderson 421 (US, BISH, Fo, NY, L); south southwest coast, 1025 ft [310 m], Falanruw 1925 (US). Sarigan: Kanehira, 1935a, pp. 293-294. #PoZyscias scutellaria (Burman f.) Fosberg Persisting around old house site. Widespread cultivated ornamental. Pagan: Near Salt Lake, west side of island, 2- 50 m, Fosberg 31346 (US). UMBELLIFEREAE Centella asiatica (L.) Urban Prostrate herb on moist shaded forest floor. Indo-Pacific, a very closely related species, some- times regarded as the same, in eastern America. Alamagan: South southwest coast, 750 ft [230 m], Falanruw 1920 (US). Sarigan: Above village, 375 m, Evans 2379 (US, BISH, Fo, NY). Anatahan: West end of coast, 250 ft [75 m], Falanruw 1688 (US). SAPOTACEAE Pouteria obovata (R. Brown) Baehni (Sideroxylon glomeratum Volkens) Common tree in forests and wooded ravines. 36 SMITHSONIAN CONTRIBUTIONS TO BOTANY Indo-Paci fic. Agrigan: Hosokawa, 1937, p. 282. Pagan: Road to Inae Dikiki, 100-200 ft [30-60 m], Falanruw 1868 (US), 1866 (US); Caldera Wall, Moore 324 (US); trail to Talague Beach, below 150 ft [45 m], Falanruw 1871 (US); isthmus, 20 m, Anderson 534 (US). Alamagan: South southwest coast, 1025 ft [310 m], Falanruw 1926 (US); Kanehira 2174 (NY); vi- cinity of Asongsong village, Fosberg 31742 (US); 31705 (US); Anderson 430 (US); around Partido village, Fosberg 31681 (US). Guguan: 320 ft [lo0 m], Falanruw 1840 (US). PRIMULACEAE Lysimachia mauritiana Lamarck Not found by us. Indo-Pacific. Pagan: Hosokawa, 1935a, p. 34; Kanehira, 1935a, p. 387. LOGANIACEAE Geniostoma micranthum var. (Geniostoma hoeferi var. glabrum Gilg) Alamagan: Anderson 419 (US); Kanehira 2180 (NV Geniostoma micranthum de Candolle var. micranthum and semi-open areas on lava. Rare to common small tree or shrub in open Marianas endemic. Agrigan: East side of island, Fosberg 31629 (US, Pagan: Kanehira 2209 (NY). Alamagan: Around Partido village, Fosberg 31683 (US, BISH, Fo, NY); Hosokawa 7903 (A, Anatahan: Northwest corner of island, below 200 ft [60 m], Falanruw 1609 (US); western end of north coast, below 250 ft [75 m], Falanruw 1683 Sarigan: Above village, 375 m, Evans 2372 (US, BISH); Hosokawa 7994 (US). US). (US). BISH, Fo, NY). OLEACEAE Jasminum marianum de Candolle Uncommon subscandent shrub. Marianas endemic. Pagan: Mt. Pagan, Moore 359 (US); isthmus, Anderson 538 (US). Alamagan: South southwest coast, 250 ft [75 m], Falanruw 1876 (US); vicinity of Asongsong village, 3-15 m, Fosberg 31721 (US); 200 m, Anderson 420 (US). APOCYNACEAE *Catharanthus roseus (L.) G. Don camps. Cultivated and naturalized around villages and Pantropical, native of Madagascar. Agrigan: Southwest coast village area, below 300 ft [90 m], Falanruw 2177 (US); village, 10-20 m, Fosberg 31565 (US). Pagan: Village, Moore 416 (US); near Salt Lake, west side of island, Fosberg 31358 (US), Alamagan: South southwest coast, 350 ft [110 m], Falanruw 1889 (US), 1890 (US); around Partido village, Fosberg 31652 (US, BISH). Sarigan: Village and vicinity, 10-50 m, Evans 2442 (US, BISH). Anatahan: Village area, northwest corner of island, below 250 ft [75 m], Falanruw 1678 (US). Cerbera dilatata Markgraf Marianas endemic. Pagan: Hosokawa 7969 (A). Neiosperma oppositifolia (Lamarck) Fosberg & Sachet Scattered small trees in thickets and forests. Indo-Pacific. Maug: South central East Island, 400 ft [120 m], Falanruw 2245 (US). Asuncion: Southwest slope, 150 ft [45 m], Falan- ruw 2254 (US). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31577 (US, BISH, Fo.). Pagan: Cinder plain, 150 ft [45 m], Bonham 15 (US, Fo); isthmus, Anderson 576 (US, BISH, Fo); east of village, Moore 414 (US). Alamagan: South southwest coast, 350 ft [110 m], Falanruw 1883 (US); Anderson 435 (US), 409 (US, BISH, Fo, NY); around Partido village, Fosberg 31657 (US); around Asongsong village, Fosberg 31741 (US). NUMBER 22 37 Sarigan: Near anchorage, 200 ft [60 m], Falanruw 1778 (US); 100 ft [30 m], 1716 (US); north of vil- lage, 175-250 m, Evans 2412 (US, BISH, Fo, NY). Ochrosia mariannensis A. de Candolle Occasional in open areas, more common in semi- open and low woods. Marianas endemic. Pagan: Kanehira 2201 (NY, FU); isthmus, 16 m, Anderson 535 (US, BISH, Fo, NY, L); Caldera Wall, Moore 320 (US); trail to Talague beach, below 150 ft [45 m], Falanruw 1870 (US, BISH). Sarigan: Near anchorage, 200 ft [60 m], Falan- ruw 1770 (US) seedling; below 250 ft [75 m], 1728 (US, BISH); 200 ft [60 m], 1769 (US, BISH); north of village, 175-250 m, Evans 2417 (US, BISH, Fo, NY). 'Plumeria rubra L. Planted in graveyard. Pantropical ornamental, of American origin. Anatahan: Village area, northwest corner of is- land, below 250 ft [75 m], Falanruw 1679 (US). 'Thevetia peruviana (Persoon) K. Schumann Small tree occasionally planted about houses. Pantropical ornamental of American origin. Agrigan: Village, Fosberg 31425 (US); southwest coast, village area, below 300 ft [90 m], Falanruw 2156 (US). Pagan: Anderson 598 (US, BISH, Fo, NY, L); near Salt Lake, Fosberg 31347 (US): between village and Lake Laguna, Moore 358 (US). CONVOLVULACEAE +Zpomoea aquatica Forskal Growing in low marshy area at east end of lake. Pantropical potherb. Pagan: Fresh Water Lake, Fosberg 31393 (US, BISH, Fo). *Zpomoea batatas (L.) Lamarck Cultivated and persisting about villages and former habitations. Widespread food plant, probably of American origin. Agrigan: Fritz, 1902, p. 102; Gaudichaud, 1826, p. 76. Pagan: Near Salt Water Lake, west side of island, Fosberg 31357 (US); 30 m, Anderson 595 (US, BISH, Fo), Anderson 596 (US, BISH, Fo). Alamagan: Fritz, 1902, p. 102. Sarigan: North of village, 50-175 m, Evans 2422 (US, BISH, Fo, NY). Zpomoea indica (Burman f.) Merrill Vine climbing over vegetation. Indo-Pacific. Alamagan: Vicinity of Asongsong village, 3-1 5 Guguan: Southwest part of island, 175 ft [55 m], m, Fosberg 31731 (US). Falanruw 1848 (US). Ipomoea pes-caprae ssp. brasiliensis (L.) van Ooststroom Common coastal species on all islands, occasional inland in pioneer situations, as on volcanic sands and rocks of Guguan and Uracas. Pantropical. Uracas: West side of south coast, below 400 ft [I20 m], Falanruw 2209 (US). Asuncion: Southwest coast, 100 ft [30 m], Falan- ruw 2280 (US). Agrigan: Marche 298 (P, Fo); Village, Fosberg 31429 (US); southwest coast, below 300 ft [90 m], Falanruw 2184 (US, BISH); midwest coast, 15 ft [5 m], Falanruw 2348 (US). Pagan: Bonham 21 (US, Fo); isthmus, Anderson 565 (US, BISH, Fo, NY, L); Fresh Water Lake, Fosberg 31396 (US); Marche 149 (P, Fo); Mt. Pagan, Moore 368 (US); air strip, Moore 308 (US); La- guna, Villagomez JV-05 (US). Alamagan: Partido village, Fosberg 31645 (US, BISH); south southwest coast, 0-200 ft [O-60 m], Falanruw 1956 (US). Guguan: West coast, below 50 ft [15 m], Falan- ruw 1800 (US). Sarigan: Coastal bluffs above anchorage, 0-75 ft [O-20 m], Falanruw 1775 (US); below village, 0-20 m, Evans 2429 (US, BISH, Fo, NY). Anatahan: Beach area, sea level, Falanruw 1665 (US), northwest tip of island, 0-10 m, Evans 2460 (US, BISH, Fo, NY). Medinilla: Swedberg, MS 1974. Zpomoea sp. only, seen. Falanruw 1708 (US). *OpercuZina ventricosa (Bertero) Peter Vine growing along goat trail in ravine, seedling, Anatahan: Western coast, below 200 ft [60 m], 38 SMITHSONIAN COKTRIBUTIOSS TO BOTANY Locally abundant vine climbing over thickets and edges of forests. Pantropical. Asuncion: Southwestern slope, 150 ft [45 m], Falanruw 2251 (US, BISH). Agrigan: Trail around south side of island, Fos- berg 31617 (US); midwest coast, 200 ft [60 m], Fa- lanruw 2329 (US). Pagan: Bonham 2 (US, Fo); Fresh Water Lake, 2-50 m, Fosberg ?l383 (US, BISH); Marche 148 (PI Fo); edge of air strip, Moore 306 (US). Alamagan: Vicinity of Asongsong village, Fos- berg 31722 (US, BISH); south southwest coast, be- low 500 ft [150 m], Falanruw 19?9 ? (US). Sarigan: Near village, 10-100 m, Evans 2??7? (US, BISH, Fo, NY). Anatahan: Falanruw 1988 ? (US). Stictocardia campanulata (L.) Merrill Large vine, growing over shrubs and trees in ravines and coconut plantations. Indo-Pacific. Alamagan: Around Partido village, Fosberg 31696 Sarigan: Near anchorage, 300 ft [90 m], Falan- ruw 1761 (US); Kanehira 2170 (NY). Anatahan: Western coast, below 200 m, Falan- ruw 1706 (US). (US). BORAGINACEAE "Heliotropium indicum L. Weed in villages. Pantropical. Sarigan: Village and vicinity, 10-50 m, Evans 2440 (US, BISH, Fo, NY). Heliotropium oualifolium var. depressum (Chamisso) Merrill Weed in villages. Species Indo-Pacific; variety supposedly Marianas Pagan: Village, Moore 394 (US). endemic. VERBEN ACE AE Callicarpa candicans var. (Callicarpa paucinervia Merrill) Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31599 (US). Pagan: Kanehira 2203 (US); south of pier, An- derson 585 (US); isthmus, 2 m, Anderson 546 (US); east coast, Moore 424 (US); Talague beach, Moore ?40 (US). Alamagan: Vicinity of Asongsong village, 3-1 5 ni, Fosberg 31706 (US). Anatahan: Northwest corner of island, below 150 ft 145 m], Falanruw 1652 (US). Callicarpa lamii Hosokawa camp, edge of forest, Anderson 606 (US). Pagan: Marche 15? (P, Fo); north of marine Alamagan: Anderson 416 (US). Clerodendron inerme (L.) Gaertner shade. Common component of undergrowth in semi- Indo-Pacific. Pagan: Caldera Wall, Moore 317 (US). Sarigan: Near anchorage, 150 m, Falanruw I730 (US). Premna obtusifolia R. Brown areas. Small tree, occasional in semi-open and forested Indo-Pacific. Asuncion: Southwest slope, 250 ft [75 m], Falan- ruw 2273 (US); 400 ft [120 m], 2289 (US). Agrigan: Village, Fosberg 31426 (US). Pagan: Volcanic ash soil, 150 ft [45 m], Bonham 8 (US, Fo); Caldera il'all, Moore 321 (US); Fresh Water Lake, Fosberg 31?82 (US); isthmus, 6 m, Anderson 531 (US, BISH, Fo). Alamagan: Around Partido village, Fosberg 31680 (US, BISH, Fo, NY, L); above village, 150- 250 m, Evans 2395 (US, BISH, Fo, NY). Guguan: 200-300 ft [60-90 m], Falanruw 1826 (US); 320 ft [I00 m], 1835 (US). Sarigan: Above village, 350-375 m, Evans 2369 (US, BISH, Fo, NY). Anatahan: Northern coast, below 250 ft [75 m], Falanruw 1684 (US), I685 (US). Medinilla: Lehne and Gabler, 1972, p. 20, as "Ahgau." "Stachytarpheta jamaicensis (L.) Vahl In disturbed places around village and landing. Pantropical weed of American origin. Pagan: East of pier, Anderson 612 (US, BISH, Fo); flats near landing, Fosberg 31368 (US); vil- lage, Moore 409 (US). Vitex negundo L. NUMBER 22 39 Infrequent, in clumps near coast. Indo-Pacific. Maug: South central East Island, 15 ft [5 m], Falanruw 2246 (US, BISH). Pagan: Inae Dikiki, below 100 ft [30 m], Falan- ruw 1858 (US, BISH); east coast, Moore 423 (US). LABIATAE "Hyptis pectinata (L.) Poiteau trails. nesia. Falanruw 2248 (US, BISH). 20 m, Fosberg 31594 (US). Occasional to common in weedy places and along Weed of American origin, common in Micro- Maug: South central East Island, 50 ft [15 m], Agrigan: Trail around south side of island, 10- Pagan: Fresh Water Lake, Fosberg 31395 (US). Alamagan: South southwest coast, 500 ft [150 m], Falanruw 1907 (US); vicinity of Asongsong vil- lage, 3-15 m, Fosberg 31718 (US). Sarigan: Near village, 10-100 m, Evans 2345 (US, BISH, Fo, NY). "Ocimum basilicum L. Cultivated in gardens and dooryards. Widespread cultivated species. Agrigan: Village, 10-20 m, Fosberg 31558 (US, BISH). "Ocimum sanctum L. Commonly cultivated in villages. Indo-Pacific. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2176 (US), 2186 (US); Cultivated, Fos- berg 31422 (US, BISH, Fo). Pagan: Road to Talague beach, Moore 346 (US); roadside, Anderson 608 (US, BISH, Fo). SOLANACEAE *Capsicum frutescens L. Common shrub planted in villages, naturalized along trails and in undergrowth in thickets and woods. Tropical American, now pantropical. Asuncion: Southwest slope, 150 ft [45 m], Falan- ruw 2250 (US), Falanruw 2265 (US), Falanruw 2267 (US). Agrigan: Trail around south side of island, Fos- berg 31588 (US). Alamagan: Cultivated, Fosberg 31701 (US, BISH); around Partido village, Fosberg 31668 (US, BISH, Fo); south southwest coast, 350 ft [110 m], Falanruw 1886 (US). Sarigan: Northwest coast, 400 ft [120 m], Falan- ruw 1750' (US); near village, 10-100 m, Evans 2343 (US); above village, 150-250 m, Evans 2389 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, 150 ft [45 m], Falanruw 1651 (US). *Datura mete1 L. One plant seen in garden. Widespread cultivated species. Agrigan: Village, 10-20 m, Fosberg 31566 (US). Planted around villages. Widespread cultivated species of American origin. Agrigan: Fritz, 1902, p. 109. Alamagan: Around Partido village, Fosberg 31693 (US). Sarigan: Village and vicinity, 10-50 m, Evans 2434 (US). Anatahan: Seen but not collected by Falanruw. "Physalis angulata L. Infrequent in semi-open area. Pantropical weed. Anatahan: Northwest corner of island, below 150 *Nicotiana tabacum L. ft [45 m], Falanruw 1650 (US). Physalis minima L. infrequent inland. Scattered at low elevations in open rocky areas, Old World tropics. Uracas: Large precaldera rock on south coast, less than 400 ft [120 m], Falanruw 2199 (US). Sarigan: Near village, 10-100 m, Evans 2361 (US, BISH, Fo, NY); volcanic ridge near anchorage, 325 ft [lo0 m], Falanruw 1743 (US); 0-75 ft [O-20 m], Falanruw 1776 (US); northwest coast, Falanruw 1749. Anatahan: Trail inland from village, less than 1000 ft [300 m], Falanruw 1971 (US). "Solanum melongena L. Sparingly cultivated. Widespread cultivated garden vegetable. Alamagan: Around Partido village, Fosberg 31654 (US). 40 SMITHSONIAN CONTRIBUTIONS TO BOTANY Sarigan: North of village, 50-175 (US). m, Evans 2423 SCROPHULARIACEAE *Angelonia angustifolia Bentham Rare, in vicinity of houses. Cultivated ornamental of tropical American Anatahan: Northwest corner of island, below origin. 200 ft [60 m], Falanruw 1640 (US). "Lindernia crustacea (L.) F. von Mueller haps from a shaded spot. Rare, on crater floor, a very slender form, per- Widespread pantropical weed. Pagan: Mt. Pagan, 1870 ft [570 m], Moore 361 (US). MYOPORACEAE hlyoporum boninense Koidzumi Endemic to Bonin and Marianas islands. Pagan: On cliff above shore, 4 m, Anderson 600 (US). ACANTHACEAE Blechum brownei f. puberulum Leonard thickets. form. ruw 2268, 2258 (US). Abundant along trails and in plantations and Pantropical; f. puberulum is common Old World Asuncion: Southwest slope, 150 ft [45 m], Falan- Agrigan: Fosberg 31423 (US, BISH, Fo, NY). Alamagan: Partido village, Fosberg 31672 (US, BISH, Fo, NY, L); south southwest coast, 350 ft [I10 m], Falanruw 1887 (US). Guguan: Southwest part of island, 175 ft [55 m], Falanruw 1845 (US). Sarigan: Northwest coast above anchorage, 200 ft [60 m], Falanruw 1771 (US); near village, 10- 100 m, Evans 2342 (US, BISH, Fo, NY); Kanehira 2167 (FIJ). Anatahan: Northwest corner of island, 200-300 ft [60-90 m], Falanruw 1616 (US); western coast, below 200 m, Falanruw 1702 (US). Ruellia sp. Anatahan: West coast, Falanruw 1702 (US). RUBIACEAE Hedyotis albido-punctata (Merrill) Fosberg On rocks not far from sea. Western Pacific and eastern Asia strand species. Sarigan: Northwest coast, sea level to 75 ft [20 m], Falanruw 1779 (US), Falanruw 1794 (US); 300 ft [90 m], Falanruw 1763 (US). Anatahan: Northwest tip of island, 0-10 m, Evans 2450 (US, BISH, Fo, NY). Hedyotis biflora (L.) Lamarck Small herbs growing in moist shaded or semi- open areas and along trails. Agrigan: Midwest coast, 150 ft [45 m], Falanruw 2343 (US); trail around south side of island, 10- 20 m, Fosberg 31580 (US). Alamagan: South southwest coast, 350 ft [110 m], Falanruw 1888 (US). Anatahan: Hosokawa 7836 (BISH); west end of north coast, below 250 ft [75 m], Falanruw 1689 (US); northwest corner of island, 200-300 ft [60- 90 m], Falanruw 1612 (US); northwest tip of island, 0-10 m, Evans 2454 (US). Hedyotis foetida var. mariannensis (Merrill) Fosberg Foetid erect shrub or suffrutescent herb growing back of beaches, on ledges, in rocky areas, and in open on black cinder sand inland on Guguan. Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31597 (US), Pagan: Mt. Pagan, Moore 366 (US); 12 m, And- erson 502 (US); isthmus, 4 m, Anderson 550 (US). Anatahan: Beach area, sea level, Falanruw 1668 (US); northwest tip of island, 0-10 m, Evans 2445 (US, BISH, Fo, NY). Guguan: 320 ft [loo m], Falanruw 1833 (US). Alamagan: Hosokawa 7898 (BISH); south south- west coast, 0-200 ft [0-60 m], Falanruw 1960 (US); vicinity of Asongsong village, 3-15 m, Fosberg 31707 (US); around Partido village, Fosberg 31682 (US); Anderson 414 (US). Hedyotis kanehirae (Hatusima) Fosberg Not seen by us. Alamagan: Hatusima, 1936, p. 218. Alamagan: Kanehira 2181 (NY). Not seen by us. Hedyotis laciniata Kanehira Hedyotis sariganensis Hosokawa NUMBER 22 41 Sarigan: Hosokawa, 1935a, p. 36; Hatusima, 1936, p. 226; Kanehira, 1935a, p. 419. Hedyotis scabrifolia Kanehira var. Pagan: Mt. Pagan, Moore 383 (US). Morinda citrifolia L. Glossy-leafed shrub or small tree found on all islands except Uracas in semi-open and open areas, thickets, and coconut plantations, on various sub- strata, dominant "small shrub-like tree" on Medi- nilla. Pantropical. Maug: North end of island, below 400 ft [120 m], Falanruw 2217 (US). Asuncion: Lower southwest slope, 150 ft [45 m], Falanruw 2263 (US). Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31591 (US); southwest coast, below 300 ft [90 m], Falanruw 2194 (US). Pagan: Isthmus, Anderson 575 (US, BISH, Fo, NY, L); 150 ft [45 m], Bonham 13 (US, Fo); vil- lage, Moore 417 (US); east coast, Moore 420' (US). Alamagan: Around Partido village, Fosberg 31661 (US); south southwest coast, 250 ft [75 m], Falan- ruw 1880 (US). Guguan: West coast, below 50 ft [15 m], Falan- ruw 1815 (US). Sarigan: Above village, 150-250 m, Evans 2394 (US, BISH, Fo, NY); northwest coast, 325 ft [I00 m], Falanruw 1747 (US). Anatahan: Vicinity of beach, northwest tip of island, 0-10 m, Evans 2461 (US, BISH, Fo, NY); behind village, northwest corner of island, below 200 ft [60 m], Falanruw 1604 (US). Medinilla: Swedberg, MS 1974. Psychotria hombroniana (Baillon) Fosberg (Psychotria mariannensis Kanehira) Marianas endemic. Alamagan: Kanehira 2186 (P, NY). Shrubs or small tree, common in undergrowth Endemic to Marianas and Yap Island. Agrigan: Marche 292 (P); canyon on east side of island, Fosberg 31632 (US, BISH, Fo); trail around south side of island, 10-20 m, Fosberg 31615 (US, BISH, Fo, NY, L); midwest coast, 200 ft [60 m], Falanruw 2333 (US, BISH). Pagan: Kanehira 2205 (NY); Fresh Water Lake, Psychotria mariana Bartling in forests, thickets, and semi-open areas. 2-50 m, Fosberg 31370 (US, BISH, Fo), Fosberg 31377 (US); Anderson 505 (US, BISH, Fo, NY); 150 ft [45 m], Bonham 9 (US, BISH); Caldera Wall, Moore 327 (US); Talague beach, Moore 337 (US); road to Inae Dikiki, below 100 ft [30 m], Falanruw 1856 (US); trail to Talague beach, below 150 ft [45 m], Falanruw 1871a (US). Alamagan: Asongsong, 100 ft [30 m], Kondo llb (BISH); 200 m, Anderson 403 (US, BISH, Fo), An- derson 420 (US, BISH, Fo, NY); south southwest coast, 350 ft [I10 m], Falanruw 1891 (US); 250 ft [75 m], Falanruw 1875 (US); around Partido village, Fosberg 31664 (US, BISH, Fo, NY, L). Sarigan: Near anchorage, 325 ft [lo0 m], Falan- ruw 1744 (US, BISH); above village, 150-250 m, Evans 2391 (US, BISH, Fo, NY); near village, 10- 100 m, Evans 2340 (US, BISH, Fo, NY). Randia cochinchinensis (Loureiro) Merrill Shrub, common in forests. Indo-Pacific. Pagan: Anderson 506 (US); isthmus, Anderson Alamagan: Around Partido village, Fosberg 31 673 539 (US). (US); Anderson 422 (US). CAPRIFOLIACEAE "Sambucus mexicana Presl ex de Candolle Cultivated in village. American, thought to be medicinal. Agrigan: Village, Fosberg 31435 (US). GOODENIACEAE Scaeuola taccada (Roxburgh) Gaertner Shrub found in open areas at about 100 m or below, in other regions mostly found on beaches and sea-facing rocks, here mostly inland; a tall and a depressed form present. Asuncion: Southwestern slope, 75 ft [20 m], Fa- lanruw 2284 (US, BISH, Fo). Agrigan: Fosberg 31428 (US). Pagan: Anderson 587 (US, BISH, NY, L); Ta- lague beach, Moore 341 (US); Inae Dikiki, below 100 ft [30 m], Falanruw 1860 (US); trail to Pan- santati, below 100 ft [30 m], Falanruw 1865 (US, BISH). Alamagan: Kanehira 2195 (FU); Partido village, Fosberg 31686 (US, BISH). 42 SMITHSONIAN CONTRIBUTIONS TO BOTANY Guguan: 320 ft [lo0 m], Falanruw 1838 (US, Sarigan: Above village, 375 m, Evans 2373 (US, Anatahan: Hosokawa 7844 (A). BISH). BISH, Fo, NY, P). COMPOSITAE *Ageratum conyzoides L. Common in openings, forest edges and especially in disturbed areas. Agrigan: Midwest coast, 20 ft [6 m], Falanruw 2322 (US). Pagan: East of Fresh Water Lake, Fosberg 31413 (US, BISH). Alamagan: Around Partido village, Fosberg 31688 (US, BISH); south southwest coast, sea level to 200 ft [60 m], Falanruw 1946 (US). Guguan: 200-300 ft [60-90 m], Falanruw 1849 Sarigan: Near anchorage, sea level, Falanruw 1784 (US); 150 ft [45 m], 1722 (US); north of vil- lage, 175-250 m, Evans 2414 (US): near village, 10-100 m, Evans 2356 (US, BISH, Fo, NY). Anatahan: Northwest corner of island, below 200 ft [60 m], Falanruw 1638 (US). "Artemisia vulgaris L. widely and becoming established. (US). Medicinal herb, abundant in graveyards. Planted Eurasian origin. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2157 (US). Anatahan: Northwest coast, below 250 ft [75 m], Falanruw 1677 (US). "Bidens sulphurea (Cavanilles) Schultz-Bipontinus (Cosmos sulphureus Cavanilles) Planted ornamental of American origin. Agrigan: Fosberg 31559 (US). Found in bare or disturbed places. Cosmopolitan. Pagan: Mt. Pagan, Moore 375 (US); east of air *Conyza bonariensis (L.) Cronquist strip, Moore 310 (US). "Conyza canadensis (L.) Cronquist turbed places. Found near airstrip; generally in bare or dis- Cosmopolitan. Pagan: East of air strip, Moore 307 (US). "Elephantopus mollis Humboldt, Bonpland, 8c Kunth Common weed along trails and in village areas, also on new volcanic ash. Widespread weed of American origin, often con- fused with the Asiatic E. scaber. Asuncion: Lower southwest slope, 150 ft [45 m], Falanruw 2264 (US, BISH). Agrigan: Fosberg 31420 (US, BISH); southwest coast, below 300 ft [90 m], Falanruw 2193 (US, BISH). Pagan: Salt Lake, Fosberg 31359 (US, BISH); isthmus, Anderson 572 (US, BISH, NY, L); Mt. Pagan, Moore 384 (US). Alamagan: South southwest coast, 450 ft [140 m], Falanruw 1898 (US). Anatahan: Northwest corner of island, 200-300 ft [60-90 m], Falanruw 1632 (US). "Emilia sonchifolia (L.) de Candolle ex Wight as in disturbed areas and around villages. Scattered in open areas of volcanic ash, as well Pantropical, of south Asian origin. Agrigan: Midwest coast, 30 ft [lo m], Falanruw 2316 (US); trail around south side of island, 10-20 m, Fosberg 31614 (US, BISH, Fo). Pagan: Village, Moore 429 (US); near Salt Lake, west side of island, Fosberg 31369 (US); east of pier, Anderson 613 (US). Alamagan: South southwest coast, sea level to 200 ft [60 m], Falanruw 1958 (US). Guguan: West coast, below 50 ft [I5 m], Falan- ruw 1813 (US); 200 ft [60 m], Falanruw 1820b (US). "Eupatorium odoratum L. A few patches of low bushes inland of beach. Pantropical weed of American origin. Agrigan: Southwest coast, below 300 ft [90 m], Falanruw 2191 (US). Glossogyne tenuifolia (Lessing) Cassini Scattered on open lava slopes, and open areas of cinder sand, reaching 500 m elevation on Asuncion. A widely distributed Indo-Pacific savanna spe- cies. Asuncion: Southwestern slope, 1500 ft [460 m], Falanruw 2297 (US); 1000 ft [300 m], 2306 (US). Agrigan: Hosokawa 8006 (A). Pagan: Hosokawa 7956 (Fo); Kanehira 2212 (P, NY, FU); east of Fresh Water Lake, Fosberg 31404 NUMBER 22 (US, BISH, NY, L); Mt. Pagan, 1870 ft [570 m], Moore 374 (US). Guguan: 200-300 ft [60-90 m], Falanruw 1831 (US, BISH). *Pseudelephantopus spicatus (Aublet) Gleason Occasional in weedy places. A pantropical weed. Agrigan: Trail around south side of island, 10- 20 m, Fosberg 31595 (US); southwest coast, below 300 ft [90 m], Falanruw 2165 (US). 'Synedrella nodiflora (L.) Gaertner weedy. Around villages, along trails and on open slopes, Pantropical, of tropical American origin. Agrigan: Village, Fosberg 31434 (US). Pagan: Kanehira 2215 (NY, FU). Alamagan: Asongsong village, 3-1 5 m, Fosberg 31749 (US). Sarigan: Northwest coast, 150 ft [45 m], Falan- ruw 1721 (US); 200 ft [60 m], Falanruw 1768 (US); near village, 10-100 m, Evans 2363 (US). +Vernonia cinerea (L.) Lessing open and semishaded areas generally. Common small weed in villages, gardens, and Pantropical. Maug: South central East Island, 100 ft [30 m], Falanruw 2249 (US). Asuncion: Southwestern slope, 250 ft [75 m], Falanruw 2270 (US). Agrigan: Village, Fosberg 31430 (US, BISH); southwestern coast, below 300 ft [90 m], Falanruw 2161 (US). Pagan: Isthmus, Anderson 569 (US, BISH, NY, L); Mt. Pagan, Moore 380 (US); Fresh Water Lake, Fosberg 31400 (US). Alamagan: Partido village, Fosberg 31649 (US); south southwest coast, sea level to 200 ft [60 m], Falanruw 1949 (US); 450 ft [140 m], Falanruw 1896 (US). 43 Sarigan: Near anchorage, 100 ft [30 m], Falan- ruw 1718 (US); near village, 10-100 m, Evans 2362 (US, BISH, Fo, NY). Anatahan: Hosokawa 7856; northwest corner of island, 300 ft [90 m], Falanruw 1623 (US). Wedelia biflora var. canescens (Gaudichaud) Fosberg Common extensively semi-creeping shrub or coarse herb on rocks and volcanic material, mostly near seacoast. Indo-Pacific. Maug: South central East Island, 50 ft [I5 m], Falanruw 2247 (US). Agrigan: Fosberg 31613 (US, BISH, NY, L); southwest coast below 300 ft [90 m], Falanruw 2170 Pagan: Landing, Fosberg 31418 (US, BISH); isthmus, Anderson 551 (US, BISH, NY); south of pier, Anderson 588 (US, BISH, NY, L); Laguna, Villagomez JV-04 (US); Talague beach, Moore Alamagan: Asongsong village, 3-15 m, Fosberg Guguan: 200-225 ft [60-70 m], Falanruw 1823 Sarigan: Kanehira 2165 (FU); northwest coast, sea level to 75 ft [20 m], Falanrziw 1791 (US, BISH); above village 150-250 m, Evans 2395 (US, BISH, Fo, NY). Anatahan: Beach area, sea level, Falanruw 1659 (US); northwest tip of island, 0-10 m, Evans 2446 (US, BISH, Fo, NY). (US). 345 (US). 31708 (US, BISH, Fo). (US). Medinilla: Swedberg, MS 1974. 'Zinnia elegans Jacquin Planted ornamental in village. Cosmopolitan in cultivation, of tropical Ameri- Agrigan: Village, Fosberg 31560 (US). can origin. References Beechey, F. W. Corte, F. de la 1831. 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The Vegetation of Micronesia, 1: General Descrip- tions, the Vegetation of the Marianas Islands, and a Detailed Consideration of the Vegetation of Guam. Bulletin of the American hluseuni of Natural His- Diels, L. 1924. Fosberg, F. R. 1998. 1960. tory, 11931-75. Fosberg, F. R., and G. Corwin 1938. A Fossil Flora from Pagan, Mariana Islands. Pacific Science, 12: 3-16. Flora of Micronesia, 1 : Gymnospermae. Smithsonian Contributions to Botany, 20: 1-13. Reise nach den nordlichen Marianen. Mittkeilungen aus den deutscken Schutzgebieten der Sudsee, 15: Fosberg, F. R., and M.-H. Sachet 1975. Fritz, G. 1902. 96-1 18. Gaudichaud, C. 1826. Botanique. In M. Louis de Freycinet, Voyage autour du monde, entrepris par ordre du roi . . . execute sur les corvettes de S. M. 1'Uranie et la Physicienne pendant les annees 1817, 1818, 1819, 1820. 522 pages. Paris. Hatusinia, S. 1936. Materials for a Flora of Micronesia, Rubiaceae I: Hedyotideae. 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Fosberg, Department of Botany, h'ational Museum of Katural History, Smithsonian Institution, Tt'ashington, D.C. Tt'agner, Tt'. H., Jr. 1948. '1. Sew Fern from Rota, Marianas Islands, Pacific Science, 2 :2 14-21 5. W'agner, Tt'. H., Jr., and D. H. Grether 1948. Pteridophytes of Guam. Occasional Papers of the Bernice P. Bishop ,Museum, 19:25-99. Piperaceae of Micronesia. Occasional Papers of the Bernice P. Bishop Museum, 22:83-108. Yuncker, T. G. 1959. Index to Families Acanthaceae, 40 Agavaceae. See Liliaceae Aizoaceae, 24 Amaranthaceae, 23 Amaryllidaceae. See Liliaceae Anacardiaceae, 31 Annonaceae, 25 Apocynaceae, 36 Araceae, 17 Araliaceae, 35 Balanophoraceae, 23 Batidaceae, 23 Bombacaceae, 33 Boraginaceae, 38 Bromeliaceae, 18 Capparidaceae, 25 Caprifoliaceae, 41 Caricaceae, 34 Casuarinaceae, 20 Combretaceae, 34 Commelinaceae, 18 Compositae, 42 Convolvulaceae, 37 Cucurbitaceae, 34 Cyatheaceae, 10 Cyperaceae, 15 Dioscoreaceae, 19 Euphorhiaceae, 29 Flagellariaceae, 18 Gleicheniaceae, 6 Goodeniaceae, 41 Gramineae, 11 Guttiferae, 33 Hernandiaceae, 25 Labiatae, 39 Lauraceae, 25 Lecythidaceae, 35 Leguminosae, 26 Liliaceae, 18 Loganiaceae, 36 Lycopodiaceae, 6 Lythraceae, 34 Malvaceae, 32 Marantaceae, 19 Marattiaceae, 6 Melastomataceae, 35 Meliaceae, 29 Musaceae, 19 hlyoporaceae, 40 hlyrtaceae, 35 Nyctaginaceae, 24 Oleaceae, 36 Orchidaceae, 19 Oxalidaceae, 29 Palmae, 17 Pandanaceae, 10 Passifloraceae, 34 Piperaceae, 20 Polygonaceae, 23 Polypodiaceae, 6 Portulacaceae, 24 Primulaceae, 36 Psilotaceae, 6 Rhamnaceae, 31 Rubiaceae, 40 Rutaceae. 29 Sapindaceae, 31 Sapotaceae, 35 Scrophulariaceae, 40 Solanaceae, 39 Sterculiaceae, 33 Taccaceae, 19 Theaceae, 33 Tiliaceae, 31 Ulmaceae, 21 Umbelliferae, 39 Urticaceae, 22 T?erbenaceae, 38 Zingiberaceae, 19 45 U.S. GOVERNMENT PRlNTlNG OFFICE: 1975--58&977/11