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SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY • NUMBER 33

A Checklist of the American Bidessini (Coleoptera: Dytiscidae-Hydroporinae)

Frank N. Young

SMITHSONIAN INSTITUTION PRESS

CITY OF WASHINGTON

1969

A B S T R A C T

Young, Frank N. A Checklist of the American Bidessini (Coleoptera: Dytiscidae-Hydroporinae). Smithsonian Contributions to Zoology, 33:1–5. 1969.—A list of the American water beetles of tribe Bidessini (Dytiscidae-Hydroporinae). Type-species are listed for each genus with references to the original designations.

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Contribution No. 809 from the Zoological Laboratories of Indiana University aided by GB 2768 from the National Science Foundation

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Official publication date is handstamped in a limited number of initial copies and is recorded in the Institution's annual report, Smithsonian Year.

UNITED STATES GOVERNMENT PRINTING OFFICE
WASHINGTON : 1969

For sale by the Superintendent of Documents, U.S. Government Printing Office Washington, D.C. 20402 - Price 20 cents (paper cover)

Contents

A Checklist of the American Bidessini (Coleoptera: Dytiscidae-Hydroporinae)

Amarodytes Régimbart 00–524

Anodocheilus Babington 41–15

Bidessodes Régimbart 00–528

Bidessonotus Régimbart 95–331

Brachyvatus Zimmermann 19–134 (as subgenus of Bidessus)

Hemibidessus Zimmermann 21–196 (as subgenus of Bidessus)

Hydrodessus J. Balfour-Browne 53–55!

Brinkius Guignot 57–38!

Hypodessus Guignot 39–52, 54!

Brachybidessus Gschwendtner 54–111!

Liodessus Guignot 39–53! (as subgenus of Bidessonotus, sensu Guignot)

Microdessus Young 67–79!

Neobidessus Young 67–79!

Neoclypeodytes Young 67–77, 78

Trogloguignotus Sanfilippo 58–160!

Uvarus Guignot 39–53! (as subgenus of Amarodytes)

Literature Cited

Publication in Smithsonian Contributions to Zoology

A Checklist of the American Bidessini (Coleoptera: Dytiscidae-Hydroporinae)

Frank N. Young

____________

Frank N. Young, Department of Zoology, Indiana University, Bloomington, Indiana 47401.

This checklist attempts to bring together all of the names of water beetles of the family Dytiscidae, sub-family Hydroporinae, tribe Bidessini, which have been proposed for species from the Americas and to assign them as far as possible to currently accepted genera. In large part, reliance has been placed upon the original descriptions of the species, but wherever possible specimens have been examined. The latter include material in the British Museum (Natural History), United States National Museum, California Academy of Science, Museum of Comparative Zoology, University of Michigan Museum of Zoology, American Museum of Natural History, Zoologische Sammlung des Bayerischen Staates (Munich), and the collection of the writer.

An asterisk (*) preceding a species name indicates that I have personally examined the type or authentic cotype or paratype material and confirmed the generic assignment or synonymy. Those preceded by a question mark (?) are tentatively assigned pending further study. Those names considered to be synonyms, some of which probably represent geographical subspecies, are indicated by an “s-” preceding the name.

The arrangement adopted is that of the Leng Catalogue (Leng, 1920; Leng and Mutchler, 1927, 1933; Blackwelder, 1939) and Blackwelder's Checklist of Coleopterous Insects of Mexico, Central America, the West Indies, and South America (1944–1957). The citations to the literature are in the abbreviated form of these works and refer to the bibliographies in them with the exception of those followed by an exclamation mark (!), which are cited in full at the end of the present paper.

The references have been cross checked with the Coleopterorum Catalogus (Zimmermann, 1920), Zoological Record, and the original books or papers.

The shortcomings of such a list as this are evident, but some arrangement of the American species is necessary to provide a basis for further investigation. There are dozens, possibly hundreds of Bidessines awaiting study, particularly from Mexico and South America. This work has been seriously hampered by premature publication of incomplete descriptions. The male genitalia prove to be especially valuable in the diagnosis of species in many of the genera, and I urge that all future descriptions of new species be accompanied by figures of these structures.

Platydessus perforatus Guignot (55–4), described as a Bidessine, is not included because it was found to be a Macrovatellus (Vatellini) and was transferred to that genus by Spangler (66–57).

Amarodytes Régimbart 00–524

(Type-species Amarodytes percosioides Régimbart, 1900, subsequent designation of Guignot 39:53!)

boggianii Régimbart 00–526 (Paraguay)

duponti (Aubé) 38–568 (Brazil)

guidi Guignot 57–37! (Brazil)

oberthüri Régimbart 00–526 (Brazil)

percosioides Régimbart 00–525 (Brazil)

plaumanni Gschwendtner 35–152 (Brazil)

pulchellus Guignot 55–273! (Colombia)

segrix Guignot 50–7! (Brazil)

testaceo-pictus Régimbart 00–527 (Brazil)

undulatus Gschwendtner 54–111! (Peru)

Anodocheilus Babington 41–15

(Type-species Anodocheilus maculatus Babington, 1841, monobasic)

exiguus (Aubé) 38–490 (U.S.A.)

germanus (Sharp) 82–20 (Guatemala)

guatemalensis Zaitzev 10–223 (Guatemala)

*maculatus Babington 41–16 (Brazil)

silvestrii Régimbart 03–49 (Argentina)

Bidessodes Régimbart 00–528

(Type-species Bidessodes semistriatus Régimbart, 1900, present designation.)

*elongatus (Sharp) 82–25 (Guatemala)

fragilis Régimbart 00–530 (Paraguay)

knischi Zimmermann 21–198 (Brazil)

obscuripennis Zimmermann 21–198 (Brazil)

plicatus Zimmermann 21–198 (Brazil)

semistriatus Régimbart 00–529 (Brazil)

subsignatus Zimmermann 21–199 (Brazil)

Bidessonotus Régimbart 95–331

(Type-species Bidessonotus obtusatus Régimbart, 1895, subsequent designation of J. Balfour-Browne 47–427! to replace Hydroporus adumbratus Clark, 1862, species incognitus designated Guignot 46–114!)

bicolor Guignot 57–36! (Brazil)

*browneanus Balfour-Browne 47–441! (Jamaica)

*caraibus (Chevrolat) 63–199 (Cuba)

*championi Balfour-Browne 47–436! (Guatemala)

*fallax Balfour-Browne 47–429! (Cuba)

*inconspicuus (LeConte) 55–290! (Louisiana)

*longovalis (Blatchley) 19–310! (Florida)

melanocephalus Régimbart 95–332 (Brazil)

mexicanus Régimbart 95–333 (Mexico)

*mobilis Balfour-Browne 47–427! (Mexico)

*morosus Balfour-Browne 47–446! (Mexico)

*nepotinus Balfour-Browne 47–442! (Trinidad)

obtusatus Régimbart 95–336 (Brazil, Paraguay)

*peregrinus Balfour-Browne 47–444! (Panama)

pulicarius (Aubé) 38–494 (U.S.A.)

*régimbarti Balfour-Browne 47–430! (Mexico)

*sobrinus Balfour-Browne 47–445! (Panama)

tibialis Régimbart 95–337 (Brazil)

*truncatus Balfour-Browne 47–440! (Brazil)

*vicinus Balfour-Browne 47–428! (British Honduras)

Brachyvatus Zimmermann 19–134 (as subgenus of Bidessus)

(Type-species Hyphydrus acuminatus Steinheil, 1869, subsequent designation of Young 54–58!)

acuminatus (Steinheil) 69–249! (Argentina)

*apicatus (Clark) 62–184 (Mexico)

*hydrovatoides (Sharp) 82–20 (Guatemala)

*seminulum (LeConte) 78–377 (Florida)

*borrei (Sharp) 82–346 (Brazil)

Hemibidessus Zimmermann 21–196 (as subgenus of Bidessus)

(Type-species Bidessus [Hemibidessus] conicus Zimmermann, 1921, subsequent designation of Guignot 46–114!)

bifasciatus Zimmermann 21–197 (Brazil)

celinoides Zimmermann 21–197 (Brazil)

*conicus Zimmermann 21–196 (Brazil)

plaumanni Gschwendtner 35–374; 36–19 (Brazil)

Hydrodessus J. Balfour-Browne 53–55!

(Type-species Hydrodessus siolii Balfour-Browne, 1953, monobasic.)

Brinkius Guignot 57–38!

(Type-species Brinkius biguttatus Guignot, 1957, original designation.)

*amazonensis Spangler 66–380! (Peru)

biguttatus (Guignot) 57–39! (Brazil)

brasiliensis (Guignot) 57–40! (Brazil)

*nanayensis Spangler 66–382! (Peru)

octospilus (Guignot) 57–39! (Brazil)

pereirai (Guignot) 57–41! (Brazil)

*siolii Balfour-Browne 53–56! (Brazil)

Hypodessus Guignot 39–52, 54!

(Type-species Bidessus cruciatus Régimbart, 1903, original designation.)

Brachybidessus Gschwendtner 54–111!

(Type-species Amarodytes [Brachybidessus] titschacki Gschwendtner, 1954, monobasic)

cruciatus (Régimbart) 03–50 (Argentina)

crucifer Guignot 39–60! (Cayenne)

cruvilineatus (Zimmermann) 21–195 (Brazil)

dasythrix Guignot 54–40! (Cayenne)

*frustrator Spangler 66–378! (Peru)

titschacki (Gschwendtner) 54–112! (Peru)

Liodessus Guignot 39–53! (as subgenus of Bidessonotus, sensu Guignot)

(Type-species Hydroporus affinis Say, 1823, original designation.)

*abjectus (Sharp) 82–24 (Mexico)

acollensis Guignot 55–272! (Peru)

affinis (Say) 23–104 (North America)

*charlotti (Clark) 62–182 (Mexico)

*emilianus (Clark) 62–183 (Mexico)

s-erythrostomus (Mannerheim) 52–305 (Alaska)

*macularis (LeConte) 52–206! (Colorado)

nanus (Aubé) 38–496 (U.S.A.)

nigrinus (Casey) 84–80 (California)

*obscurellus (LeConte) 52–206! (California)

microreticulatus (Hatch) 28–219 (Washington)

andinus Guignot 58–3! (Peru)

atomus Guignot 57–36! (Brazil)

bogotensis Guignot 53–111! (Bolivia)

bonariensis (Steinheil) 69–249! (Argentina)

*cantralli (Young) 53–111! (Michigan)

chilensis (Solier) 49–189 (Chili)

*crassus (Sharp) 82–365 (Brazil)

dilatatus (Régimbart) 95–330 (Brazil)

delfini (Régimbart) 99–1 (Chili)

emaciatus Guignot 53–112! (Brazil)

flavofasciata (Steinheil) 69–249! (Argentina)

*flavicollis (LeConte) 55–291 (New York)

*fuscatus (Crotch) 73–391 (Lake Superior to Florida)

*hobbsi (Young) 50–4! (Florida)

involucer (Brinck) 48–4! (Tristan da Cunha)

microscopicus (Zimmermann) 21–93 (Brazil)

miersi (White) 47–33 (Brazil)

burrus (Guignot) 53–111 (Brazil)

*nitidus (Babington 41–14 (Brazil)

*obscurus (Babington) 41–14 (Brazil)

patagonicus (Zimmermann) 22–35 (Patagonia)

strobeli (Steinheil) 69–249! (Chili)

lusculentus Guignot 48–164! (Argentina)

*uruguensis (Sharp) 82–365 (Uruguay)

Microdessus Young 67–79!

(Type-species Bidessus atomarius Sharp, 1882, original designation)

*atomarius (Sharp) 82–366 (Brazil)

Neobidessus Young 67–79!

(Type-species Hydroporus pullus LeConte, 1855, original designation)

alternatus (Régimbart) 89–384 (Venezuela)

*pescheti (Zimmermann) 21–194 (Brazil)

brasiliensis (Régimbart) 95–238 (Brazil)

?cancellosus (Guignot) 57–34! (Peru)

*corumbensis (Zimmermann) 21–195 (Brazil)

curticornis (Régimbart) 03–54 (Argentina)

*discoidalis (Sharp) 82–369 (Brazil)

duplicatus (Guignot) 53–111! (Brazil)

liliputanus (Aubé) 38–591 (Brazil)

*obtusus (Sharp) 82–24 (Mexico)

persimilis (Régimbart) 95–329 (Mexico)

*pullus (LeConte) 55–290 (Louisiana)

s-*floridanus (Fall) 17–168 (Florida)

*shermani (Mutchler) 18–82 (Florida)

*subvittatus (Zimmermann) 21–193 (Brazil)

surinamensis (Régimbart) 89–390 (Surname)

trilineatus (Zimmermann) 25–255 (Brazil)

*vittatipennis (Zimmermann) 21–194 (Brazil)

*youngi (Leech) 48–392! (Lower California)

Neoclypeodytes Young 67–77, 78

(Type-species Hydroporus plicipennis Crotch, 1873, original designation.)

americanus (Guignot) 36–144 (Mexico)

*centralis (Sharp) 87–753 (Mexico)

*cinctellus (LeConte) 52–206 (Arizona)

*curtulus (Sharp) 87–753 (Mexico)

*decoratus (Fall) 17–167 (Arizona)

*discedens (Sharp) 82–19 (Mexico)

*discretus (Sharp) 82–350 (Texas)

*fryi (Clark) 62–421 (Mexico)

*latifrons (Sharp) 82–22 (Guatemala)

*leachi (Leech) 48–11! (California)

luctuosus (Guignot) 49–43! (Central America)

*lynceus (Sharp) 82–368 (Guatemala)

*obesus (Sharp) 82–349 (Texas)

*ornatellus (Fall) 17–167 (Oregon)

*pictodes (Sharp) 82–348 (North America)

*plicipennis (Crotch) 73–388 (California)

*quadrinotatus (Sharp) 82–21 (Mexico)

*quadrisignatus (Sharp) 82–21 (Mexico, Guatemala)

rugulosus (Guignot) 36–146 (Mexico)

*substriatus (Sharp) 82–23 (Mexico)

Trogloguignotus Sanfilippo 58–160!

(Type-species Trogloguignotus concii Sanfilippo, 1958, monobasic)

concii Sanfilippo 58–161! (Venezuela)

Uvarus Guignot 39–53! (as subgenus of Amarodytes)

(Type-species Hydroporus lacustris Say, 1823, original designation)

?adumbratus (Clark) 62–183 (Mexico)

*amandus (LeConte) 52–207! (Arizona)

costaricensis (Guignot) 39–59! (Costa Rica)

*falli (Young) 40–30! (Florida)

granarius (Aubé) 38–501 (U.S.A.)

*inflatus (Young) 50–1! (Florida)

lacustris (Say) 23–103 (U.S.A.)

*magensis (Clark) 62–182 (Mexico)

?omichlodes (Guignot) 57–37! (Brazil)

*rogersi (Young) 41–29! (Florida)

*spretus (Sharp) 82–24 (Mexico)

?subtilis (LeConte) 52–206 (California)

*subornatus (Sharp) 82–23 (Mexico)

?subplicatus (Hatch) 53–197! (Oregon)

*suburbanus (Fall) 17–169 (New York, Pennsylvania)

*texanus (Sharp) 82–366 (Texas)

Literature Cited

Balfour-Browne, J.

1947. A Revision of the Genus Bidessonotus Régimbart (Coleoptera: Dytiscidae). Transactions of the Royal Entomological Society of London, 98:425–448, 12 figures.

1953. Hydrodessus Gen. N. Hydroporinarum (Coleoptera: Dytiscidae). Proceedings of the Royal Entomological Society of London, (B) 22:55–56, 1 figure.

Blackwelder, R. E.

1939. Catalogue of the Coleoptera of America, North of Mexico. Fourth Supplement, 1933–1938:16–18.

1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. United States National Museum Bulletin, 185 (1):72–80.

1957. Ibid. (Bibliography), 185 (6):927–1492.

Blatchley, W. S.

1919. Insects of Florida. Va. Supplementary Notes on the Water Beetles. Bulletin of the American Museum of Natural History, 41:305–322.

Brinck, P.

1948. Coleoptera of Tristan da Cunha, Results of the Norwegian Scientific Expedition to Tristan da Cunha, 1937–1938, 17:1–122.

Gschwendtner, L.

1954. Dytiscidae (Col.). In Beitrage zur Fauna Peru: Nach der Ausbeute der Hamburger Südperu-Expedition 1936, anderer Sammlungen, wie auch auf Grund von Literaturangaben. 4:109–115, 24 figures. Jena: Fischer.

Guignot, F.

1939. Contribution à l'Étude des Bidessus. Bulletin Societé d'Étude Science Naturel de Vaucluse, 1939:51–61, 10 figures.

1946. Génotypes des Dytiscoidea et des Gyrinoidea. Revue Française d'Entomologie, 13:112–118.

1948. Vingt-septième Note sur les Hydrocanthans. Bulletin Mensuel de la Société Linneenne de Lyon, 17:163–171, 1 figure.

1949. Vingt-huitième Note sur les Hydrocanthares. Bulletin de la Société Entomologique de France, 54:42–47.

1950. Trente-troisième Note sur les Hydrocanthares. Bulletin Institute Science Naturel Belgique, Brussels, 26:1–10, 3 figures.

1953. Trente-neuvième Note sur les Hydrocanthares. Revue Française d'Entomologie, 20:109–117, 2 figures.

1954. Quarantième Note sur les Hydrocanthares. Bulletin et Annals Société Entomologique de Belgique, 90:40–45.

1955. Quarante-deuxième Note sur les Hydrocanthares. Revue Francaise d'Entomologie, 22:270–276, 1 figure.

1955. Description de Nouveaux Dytiscidae Principalement de l'Amérique du Sud. Bulletin Institute Royale Naturel Belgique (Brussels), 31:1–12.

1957. Contribution à la Connaissance des Dytiscides Sud-Américains (Coleoptera). Revue Française d'Entomologie, 24:33–45, 10 figures.

1957. Beiträge zur Kenntnis der Insektenfauna Boliviens, Teil II. Coleoptera I. Dytiscidae. (2. Contribution a l'ètude des Dytiscides Sud-Américains), Opuscula Zoologica, 6:1–10.

Leech, H. B.

1948. Contributions Toward a Knowledge of the Insect Fauna of Lower California, No. 4, Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Limnebiidae. Proceedings of the California Academy of Sciences, 24 (4) 375–483, 2 plates.

1948. Bidessus Leachi, a New Species of Dytiscid Water Beetle from Northern California. Proceedings of the Entomological Society of British Columbia, 44:11–12, 3 figures.

Leng, C. W.

1920. Catalogue of the Coleoptera of America, North of Mexico. Pages 75–86. New York: J. D. Sherman.

Leng, C. W. and A. J. Mutchler.

1927. Ibid. Supplement, 1919–1924:16–19.

1933. Ibid. Second and third supplements, 1925–1932:14–16 and 81.

Sanfilippo, N.

1958. Viaggio in Venezuela di Nino Sanfilippo, V: Descrizione di Trogloguignotus Concii N. Gen. N. Sp. di Dytiscidae Freatobio. Annali del Museo Civico di Storia Naturale di Genova, 70:159–164, 2 figures.

Spangler, P. J.

1966. Changes in Nomenclature and Reassignment of Platydessus Perforatus (Coleoptera: Dytiscidae) Coleopterists Bulletin, 20:57–58, 1 figure.

1966. The Catherwood Foundation Peruvian-Amazon Expedition XIII—Aquatic Coleoptera (Dytiscidae; Noteridae; Gyrinidae; Hydrophilidae; Dascillidae; Helodidae; Psephenidae; Elmidae). Monographs of the Academy of Natural Science, Philadelphia, 14:377–443, 8 plates.

Hatch, M. H.

1953. The Beetles of the Pacific Northwest, Part I: Introduction and Adephaga. University of Washington Publications in Biology, 16:1–340, 37 plates.

LeConte, J. L.

1852. Descriptions of New Species of Coleoptera from California. Annals of the Lyceum of Natural History of New York, 5:185–216.

Steinheil, E.

1869. Symbolae ad Historiam Coleopterorum Argentiniae Meridionalis. …Atti della Società Italiana di Scienze Naturali, 12:238–260 (Centuria I).

Young, F. N.

1940. Description of a New Dytiscid from Florida (Coleoptera: Dytiscidae). Florida Entomologist, 23:30.

1941. Bidessus Rogersi, New Water Beetle from Florida (Coleoptera: Dytiscidae). Florida Entomologist 24:29–31, 2 figures.

1950. Two New Species of Bidessus from the Apalachicola Flatwoods of Florida (Coleoptera: Dytiscidae). Occasional Papers of the Museum of Zoology, University of Michigan, 526:1–6, 2 figures.

1953. A New Species of Bidessus from Southern Michigan (Coleoptera: Dytiscidae). Bulletin of the Brooklyn Entomological Society, 48:111–112.

1954. The Water Beetles of Florida. University of Florida Biology Series, 5:1–238, illustrated.

1967. A Key to the Genera of American Bidessine Water Beetles, with Descriptions of Three New Genera (Coleoptera: Dytiscidae, Hydroporinae). Coleopterists Bulletin, 21:75–84.

Zimmermann, A.

1920. Coleopterorum Catalogus, Pars 71: Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae, pages 1–426.

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