Latin Status Current Mnemonic Family Authority Notes Abarema macradenia Accepted Abarema macradenia pit1ma Fabaceae-mimosoideae (Pittier) Barneby & J.W. Grimes Formerly Pithecellobium macradenium and appeared in 1996 paper under that old name. Pithecellobium was an enormously broad mix of of many taxa. Barnebey and Grimes (1996) did a detailed phylogeny of Ingeae, many of which (except Inga and Calliandra) were once Pithecellobium. The genus revisions do not alter species boundaries here. Acacia melanoceras Obsolete Vachellia melanoceras acacme Fabaceae-mimosoideae Beurl. See Acacia discussion with Vachellia melanoceras. Acalypha diversifolia Accepted Acalypha diversifolia acaldi Euphorbiaceae Jacq. Acalypha macrostachya Accepted Acalypha macrostachya acalma Euphorbiaceae Jacq. Adelia triloba Accepted Adelia triloba ade1tr Euphorbiaceae (Müll.Arg.) Hemsl. Aegiphila panamensis Accepted Aegiphila panamensis aegipa Lamiaceae Moldenke Alchornea costaricensis Accepted Alchornea costaricensis alchco Euphorbiaceae Pax & K.Hoffm. Alchornea latifolia Accepted Alchornea latifolia alchla Euphorbiaceae Sw. Alibertia dwyeri Accepted Alibertia dwyeri boropa Rubiaceae Delprete & C.H.Perss. Formerly B. panamensis. Entire genus of Borojoa, with 3 species, collapsed into Alibertia, without any changes in species boundaries. The name dwyeri was necessary because there is another A. panamensis. There was a single sapling in this taxon since 1982 (died in 2015), but the name is not in Croat nor the 1996 paper. It must have been identified later? Alibertia edulis Accepted Alibertia edulis alibed Rubiaceae (Rich.) A.Rich. ex DC. Allophylus psilospermus Accepted Allophylus psilospermus allops Sapindaceae Radlk. Alseis blackiana Accepted Alseis blackiana alsebl Rubiaceae Hemsl. Amaioua corymbosa Accepted Amaioua corymbosa amaico Rubiaceae Kunth Anacardium excelsum Accepted Anacardium excelsum anacex Anacardiaceae (Bertero & Balb. ex Kunth) Skeels Anaxagorea panamensis Accepted Anaxagorea panamensis anaxpa Annonaceae Standl. Andira inermis Accepted Andira inermis andiin Fabaceae-papilionoideae (W. Wright) Kunth ex DC. Annona acuminata Accepted Annona acuminata annoac Annonaceae Saff. Annona hayesii Accepted Annona hayesii annoha Annonaceae Saff. Annona spraguei Accepted Annona spraguei annosp Annonaceae Saff. Apeiba aspera Obsolete Apeiba membranacea apeime Malvaceae Aubl. Much confusion about identification of Apeiba glabra/membranacea/aspera. A. aspera is invalid. A. glabra is only Brazil and Guiana Shield. The BCI species is A. membrancea, but we mistakenly took this name from 2004 through 2015. Apeiba hybrid Accepted Apeiba hybrid apeihy Malvaceae Apeiba membranacea Accepted Apeiba membranacea apeime Malvaceae Spruce ex Benth. This is Croat's name and is the accepted name for a widespread Central and South American species. But A. membranacea has regularly been misidentified as A. aspera, leading us to use that incorrect name for a period. We used A. membranaceae correctly through the 1996 paper, but then the incorrect A. aspera in the 2004 paper. We have now restored the correct A. membranacea. Apeiba tibourbou Accepted Apeiba tibourbou apeiti Malvaceae Aubl. Aphelandra sinclairiana Accepted Aphelandra sinclairiana aphesi Acanthaceae Nees Appunia seibertii Accepted Appunia seibertii appuse Rubiaceae Standl. Not in Croat but identified in 1982 census, and we used in the 1996 paper; now we have reverted to it as of 2017. Steyermark (1972) and Taylor and Steyermark (Flora of NG Shield) moved Appunia within a broad Morinda, but others, however, maintained Appunia separate. Later DNA phylogeny (Razafimandimbison 2009, 2011) shows Morinda to be polyphyletic. They decided to restore Appunia, and Tropicos follows this latest. Ardisia bartlettii Accepted Ardisia bartlettii ardiba Myrsinaceae Lundell Ardisia fendleri Obsolete Ardisia standleyana ardife Myrsinaceae Lundell This taxon in Ardisia is in a state of confusion. Croat used this name, and we published it in both 1996 and 2004 papers. However, Pipoly reidentified one BCI specimen as A. standleyana in the 1980s, Fl. Nic. and Man CR use A. standleyana. However, Ricketson and Pipoly reverted most BCI specimens to A. fendleri since 2006. Ardisia guianensis Accepted Ardisia guianensis ardigu Myrsinaceae (Aubl.) Mez Ardisia standleyana Accepted Ardisia standleyana ardife Myrsinaceae P.H.Allen This was formely A. fendleri, and appeared as that old name in 1996 and 2004 papers. We switched to A. standleyana subsequently. There is confusion, though, as Ricketson and Pipoly reverted most BCI specimens to A. fendleri after 2006. Moreover, A. copeyana appears on one BCI specimen, though it is otherwise only at high elevation. Must check Man. CR and Fl. Nic. Aspidosperma cruenta Obsolete Aspidosperma desmanthum aspicr Apocynaceae This was Croat’s name. It is a misspelling of A. cruentum, and we repeated the error in 1996 paper but corrected in 2004 paper. Aspidosperma cruentum Obsolete Aspidosperma desmanthum aspicr Apocynaceae Woodson This is the correct spelling of Croat’s name (he wrote A. cruenta), and we had it correct in 2004 paper. This became A. spruceanum for a time, then A. desmanthum, but returns to A. cruentum as of 2018. Aspidosperma desmanthum Accepted Aspidosperma desmanthum aspicr Apocynaceae This genus is now in a state of confusion, largely because a PhD dissertation finished in 1999 (K. Potgieter), which makes many revisions, has not been published, so multiple names are still around. Potgieter advises that A. desmanthum should be used for the common BCI species. In the past, we used three other names: A. cruenta, A. cruentum, and A. spruceanum. The first is a misspelling, following Croat, and we published it in both the 1996 and 2004 papers. To put a cap on this, confusion goes full circle in 2018, with Morales restoring A. cruentum to Panama records, including BCI, contra Potgieter. Aspidosperma spruceanum Obsolete Aspidosperma desmanthum aspicr Apocynaceae Benth. ex Müll.Arg. From as early as 1989, this name replaced A. cruentum for BCI specimens, mostly by Morales (see also Fl. Nic.). We started using it after 2004 in our database, but both 1996 and 2004 papers used the old misspelling, A. cruenta. Astrocaryum standleyanum Accepted Astrocaryum standleyanum ast1st Arecaceae L.H.Bailey Astronium graveolens Accepted Astronium graveolens ast2gr Anacardiaceae Jacq. Attalea butyracea Obsolete Attalea rostrata sch1zo Arecaceae (Mutis ex L.f.) Wess.Boer We originally called this Scheelea zonensis, following Croat, and used that name in the 1996 paper. Henderson subsumed Scheelea under Attalea, and also merged the C. America species (A. rostrata) with the S. American A. butyracea; we used the latter in the 2004 paper. Foster (pers. comm.) believes they are distinct in habitat, but otherwise similar. Garwood (2009) treats the BCI species as A. rostrata, but Tropicos treats all as butyracea Attalea rostrata Accepted Attalea rostrata sch1zo Arecaceae Oerst. Formerly Scheelea zonensis, and we published under that name in 1996 paper, then A. butyracea, which we used in 2004 paper. We are following Garwood, who used A. rostrata, the Central American form, distinct from S. American A. butyracea. Note that Tropicos still uses A. butyracea. Bactris barronis Accepted Bactris barronis bactba Arecaceae L.H.Bailey Bactris coloniata Accepted Bactris coloniata bactc1 Arecaceae L.H.Bailey Bactris coloradonis Accepted Bactris coloradonis bactc2 Arecaceae L.H.Bailey Bactris major Accepted Bactris major bactma Arecaceae Jacq. Banara guianensis Accepted Banara guianensis banagu Salicaceae Aubl. Croat includes this taxon, but it did not appear in the 50-ha plot until the 1990 census, and it subsequently died out. Beilschmiedia pendula Obsolete Beilschmiedia tovarensis beilpe Lauraceae (Sw.) Hemsl. Croat's identification, and the name we used for 30 years, including 1996 and 2004 paper. Reidentified as B. tovarensis in Nishida (1999) but we didn’t discover the change until later. Beilschmiedia tovarensis Accepted Beilschmiedia tovarensis beilpe Lauraceae (Klotzsch & H.Karst. ex Meisn.) Sachiko Nishida We called this B. pendula for 30 years, and it appears under that name in 1996 and 2004 papers. But Nishida (1999) lists BCI specimens under B. tovarensis. He explains details of the differences at length, and they are subtle. Bertiera guianensis Accepted Bertiera guianensis bertgu Rubiaceae Aubl. Bombacopsis quinata Obsolete Pochota quinata pochqu Malvaceae This was Croat’s name and we used it at the outset. Bombacopis has been in use for 100 years, but it has been debated about whether it is a synonym of Pochota or later, Pachira. Now this is back to Pochota quinata. Bombacopsis sessilis Obsolete Pachira sessilis pochse Malvaceae This was Croat’s name and we used it in 1982. Bombacopsis has been in use for 100 years, but it has been debated about whether it is a synonym of Pochota or later, Pachira. Now this is Pachira sessilis. See notes with Pachira quinata. Borojoa panamensis Obsolete Alibertia dwyeri boropa Rubiaceae Dwyer Entire genus of Borojoa, with 3 species, collapsed into Alibertia, without any changes in species boundaries. This species has 1 tree in 50-ha plot, but it was not in 1996 paper. Croat does not include Borojoa. Brosimum alicastrum Accepted Brosimum alicastrum brosal Moraceae Sw. Brosimum guianense Accepted Brosimum guianense brosgu Moraceae (Aubl.) Huber Note misspelling in Appendix, but not main table, of 1996 paper. Calophyllum longifolium Accepted Calophyllum longifolium calolo Clusiaceae Willd. Capparidastrum frondosum Accepted Capparidastrum frondosum cappfr Capparaceae (Jacq.) Cornejo & Iltis Formerly Capparis frondosa. See notes under that name. Capparis frondosa Obsolete Capparidastrum frondosum cappfr Capparaceae Jacq. This was Croat's name, and we used in 1996 and 2004 papers. Currently Capparidastrum frondosum. Hall (2008) gives a thorough explanation of phylogeny in Capparis. The old-world species are a separate clade, not close to American. So the American Capparis have been divided into 4 genera. See also Cornejo and Iltis (2008). Carica cauliflora Obsolete Vasconcellea cauliflora carica Caricaceae Croat’s name for the wild species of papaya, but now moved to genus Vasconcellea (Badillo 2000 in Ernstia). It was not found in the 50-ha until 2010, but we identified in other plots in the 1990s and used this old name then. Casearia aculeata Accepted Casearia aculeata caseac Salicaceae Jacq. Casearia arborea Accepted Casearia arborea casear Salicaceae (Rich.) Urb. Casearia commersoniana Accepted Casearia commersoniana caseco Salicaceae Cambess. Casearia guianensis Accepted Casearia guianensis casegu Salicaceae (Aubl.) Urb. Casearia sylvestris Accepted Casearia sylvestris casesy Salicaceae Sw. Cassia fruticosa Obsolete Senna dariensis sennda Fabaceae-caesalpinioideae Croat's name, and we used it in 1982. Irwin and Barneby 1982 split Cassia into 3 genera, including Senna, and we had switched to S. dariensis by the late 1980s. Reviewed in Marazzi, with DNA phylogeny confirming monophyly of Senna. Croat indicates two forms on BCI but not distinct, one named C. f. var. gatunensis. Cassipourea elliptica Accepted Cassipourea elliptica cassel Rhizophoraceae (Sw.) Poir. Cavanillesia platanifolia Accepted Cavanillesia platanifolia cavapl Malvaceae (Bonpl.) Kunth Cecropia insignis Accepted Cecropia insignis cecrin Urticaceae Liebm. Cecropia longipes Accepted Cecropia longipes cecrlo Urticaceae Pittier Included in Croat but did not appear in the 50-ha plot until 1995 census, so does not appear in 1996 paper. Cecropia obtusifolia Accepted Cecropia obtusifolia cecrob Urticaceae Bertol. Cedrela odorata Accepted Cedrela odorata cedrod Meliaceae L. Ceiba pentandra Accepted Ceiba pentandra ceibpe Malvaceae (L.) Gaertn. Celtis schippii Accepted Celtis schippii celtsc Cannabaceae Standl. Cespedesia macrophylla Obsolete Cespedesia spathulata cespma Ochnaceae Croat's used this name, but he misspelled it Cespedezia macrophylla. Now lumped under C. spathulata. BCI specimens at Tropicos reidentified as C. spathulata in 1992 (Liesner). Cespedesia spathulata Accepted Cespedesia spathulata cespma Ochnaceae (Ruiz & Pav.) Planch. Formerly C. macrophylla. Cespedesia now considered monotypic, with nine names united under one widespread species. See Fl. Nic. We used the old species name, macrophylla, and Croat’s misspelling of the genus, so published this under Cespedezia macrophylla in both 1996 and 2004 papers. Cespedezia macrophylla Obsolete Cespedesia spathulata cespma Ochnaceae Croat used this incorrect spelling of Cespedesia macrophylla. We repeated the error in both 1996 and 2004 papers. On top of that, the species epithet was changed to C. spathulata (by 1992 at Tropicos, in 2001 Flora Nic), but we didn’t update our database until after 2004. Cestrum megalophyllum Obsolete Cestrum schlechtendahlii cestme Solanaceae Dunal Croat's name and we used in 1996 and 2004 papers. M. Nee (pers. comm.) and Hensold indicate this is a synonym of C. schlechtendahlii. Tropicos agrees. Must check published names (Knapp, Man CR??). Nee reidentified all Croat's BCI specimens as C. schlechtendahlii in 2013. Cestrum racemosum Accepted Cestrum racemosum cestra Solanaceae Ruiz & Pav. Croat included this taxon, but it did not appear in the 50-ha plot until the 2015 census. Cestrum schlechtendahlii Accepted Cestrum schlechtendahlii cestme Solanaceae G.Don We originally followed Croat and used C. megalophyllum in 1996 and 2004 papers. Nee reidentified all Croat's BCI specimens as C. schlechtendahlii in 2013. This is a question of name priority for same species circumpscription?? See Manual CR. Chamaedorea tepejilote Accepted Chamaedorea tepejilote cha1te Arecaceae Liebm. Formerly C. wendlandiana, but we switched by the 1996 paper. Lumped by Henderson (1997). Chamaedorea wendlandiana Obsolete Chamaedorea tepejilote cha1te Arecaceae Croat's original name, and we used it at the outset. By 1996 paper, we had changed to C. tepejilote. Henderson (1997) collapsed several tax into C. tepejilote. Chamguava schippii Accepted Chamguava schippii cha2sc Myrtaceae (Standl.) Landrum A small group of 3 rare C. American species, formerly Eugenia, moved to Psidium in 1963, and we initially used Croat’s name, Psidium anglohondurense. The new genus Chamguava was erected in 1991 by Landrum based on embryological details that separate it from Psidium; Chamguava is in a small group with Pimenta. The BCI population is the only one known s. of Guatemala, and one of 3 species only seen 3 times in the 1940s. No change in species boundaries. Huge DNA phylogeny (2015) does not include Chamguava, presumably due to rarity. Chimarrhis parviflora Accepted Chimarrhis parviflora chimpa Rubiaceae Standl. Chlorophora tinctoria Obsolete Maclura tinctoria maclti Moraceae Currently Maclura tinctoria, following Berg (2001). The two genus names, Chorophora and Maclura, were used in parallel during the 19th century for the species tinctoria. It was after 1962 that Chlorophora was finally and permanently moved under Maclura. Berg (2001) summarizes. Croat did not have this species under either name. D'Arcy used Chlorophora. Chrysochlamys eclipes Accepted Chrysochlamys eclipes chr1ec Clusiaceae L.O.Williams We originally followed Croat and used T. nicaraguensis, but now corrected to Chrysochlamys eclipses. We had switched to C. eclipes by 1996 paper, so no publication uses old name. See notes for T. nicaraguensis and Hammel (1999). Chrysophyllum argenteum Accepted Chrysophyllum argenteum chr2ar Sapotaceae Jacq. We used this name by 1996 and 2004 papers. In Croat, this was Cynodendron panamense, but Pennington (1990) lumped several species under C. argenteum as a subspecies. The name Chrysophyllum panamense has also been used, but never in our data. Chrysophyllum cainito Accepted Chrysophyllum cainito chr2ca Sapotaceae L. Pennington (1990) is certain this is not native to Panama. Chrysophyllum panamense Obsolete Chrysophyllum argenteum chr2ar Sapotaceae Synonym of Cynodendron panamense that we used briefly in 1980s. This became C. argenteum subsp. panamense in Pennington's monograph (1990), and we used Chr. argenteum in all publications. Cinnamomum triplinerve Accepted Cinnamomum triplinerve phoeci Lauraceae (Ruiz & Pav.) Kosterm. This was formerly Phoebe cinnamomifolia, and appeared as such in 1996 and 2004 papers. Croat called in P. mexicanum. See Fl. Nic. Clidemia dentata Accepted Clidemia dentata clidde Melastomataceae Pav. ex D. Don Clidemia is now moved within Miconia, but names are not yet transferred as of 2017. Clidemia octona Accepted Clidemia octona clidoc Melastomataceae (Bonpl.) L.O. Williams Clidemia is now moved within Miconia, but names are not yet transferred as of 2017. Clidemia septuplinervia Accepted Clidemia septuplinervia clidse Melastomataceae Cogn. Clidemia is now moved within Miconia, but names are not yet transferred as of 2017. Cnemidaria petiolata Obsolete Cyathea petiolata cnempe Cyatheaceae Genus Cnemidaria collapsed into much larger Cyathea. Coccoloba coronata Accepted Coccoloba coronata coccco Polygonaceae Jacq. Coccoloba manzanillensis Obsolete Coccoloba manzinellensis coccma Polygonaceae This is Croat's misspelling of C. manzinellensis. We propagated the misspelling in the 1996 paper, but corrected it by the 2004 paper. Coccoloba manzinellensis Accepted Coccoloba manzinellensis coccma Polygonaceae Beurl. We spelled this incorrectly in 1996, copying Croat's error C. manzanellensis. We propagated the misspelling in the 1996 paper, but corrected it by the 2004 paper. Cojoba rufescens Accepted Cojoba rufescens pit1ru Fabaceae-mimosoideae (Benth.) Britton & Rose Formerly Pithecellobium rufescens, and appears as such in 1996 paper. But Pithecellobium was an enormously broad mix of of many taxa. Barnebey and Grimes (1996) did a detailed phylogeny of Ingeae, many of which (except Inga and Calliandra) were once Pithecellobium. Cojoba is related to Inga and Calliandra according to Barneby and Grimes. The genus revisions do not alter species boundaries here. Colubrina glandulosa Accepted Colubrina glandulosa colugl Rhamnaceae Perkins Conostegia bracteata Accepted Conostegia bracteata conobr Melastomataceae Triana Conostegia cinnamomea Accepted Conostegia cinnamomea conoci Melastomataceae (Beurl.) Wurdack Cordia alliodora Accepted Cordia alliodora cordal Cordiaceae (Ruiz & Pav.) Oken Cordia bicolor Accepted Cordia bicolor cordbi Cordiaceae A.DC. Cordia lasiocalyx Accepted Cordia lasiocalyx cordla Cordiaceae Pittier Coussarea curvigemmia Accepted Coussarea curvigemmia cou2cu Rubiaceae Dwyer Coutarea hexandra Accepted Coutarea hexandra couthe Rubiaceae (Jacq.) K.Schum. Crematosperma garwoods Obsolete Mosannona garwoodii malmsp Annonaceae One name used in some early databases for the new Annonaceae, christened finally as a novel species, Mosannona garwoodii. Croat thought it was a novel Cremastosperma. Crematosperma sp. Obsolete Mosannona garwoodii malmsp Annonaceae Croat mentions this species as rare, citing a Garwood and Foster specimen, and likely new to science, but he misspelled the genus. It turned out to be very common in the 50-ha plot. We published as Malmea sp. nov. in 1996, then the final Mosannona garwoodii in 2004. Croton billbergianus Accepted Croton billbergianus crotbi Euphorbiaceae Müll.Arg. Cupania cinerea Accepted Cupania cinerea cupaci Sapindaceae Poepp. Cupania latifolia Accepted Cupania latifolia cupala Sapindaceae Kunth Cupania rufescens Accepted Cupania rufescens cuparu Sapindaceae Triana & Planch. Cupania seemannii Accepted Cupania seemannii cupasy Sapindaceae Triana & Planch. Seemann himself named it C. sylvatica in 1853 (Voyage of the Heald), however in 1862 Triana and Planch changed to C. seemanni saying C. sylvatica was already in use for a Brazilian specimen. But Croat used the old C. sylvatica, though he indicated C. seemannii as a synonym. Cupania sylvatica Obsolete Cupania seemannii cupasy Sapindaceae Croat used this name, and we used it in 1996 and 2004 papers. But this was wrong, and C. seemannii has been valid for 150 years. Cyathea petiolata Accepted Cyathea petiolata cnempe Cyatheaceae (Hook.) R.M. Tryon DNA phylogenies of tree ferns rearranged much traditional taxonomy. Cnemidaria is nested within Cyathea. No change species boundary. Cynodendron panamense Obsolete Chrysophyllum argenteum chr2ar Sapotaceae (Pittier) Aubrév. Croat's name, but we had switched to Chrysophyllum panamense by start of 50-ha plot. Yet another taxonomic revision followed, and this appears as Chrysophyllum argenteum in 1996 and 2004 papers. Pennington (1990) placed Cynodendron within Chrysophyllum. Older treatments of Sapotaceae have gone back and forth on this, indeed, Croat lists Chrysophyllum panamense as a synonym. Morever, Pennington moved C. panamense under the widely distributed Chrysophyllum argenteum. Cyphomandra hartwegii Obsolete Solanum circinatum cyphha Solanaceae Croat’s name, we used it through the 1996 paper. Now known as Solanum circinatum. Bohs (1995) collapsed entire genus Cyphomandra into Solanum with no changes in species boundaries. Damburneya umbrosa Accepted Damburneya umbrosa nectpu Lauraceae (Kunth) Trofimov One group of Nectandra now split into this genus, an old name that had been subsumed into Nectandra, now resurrected on molecular results. This was originally Nectandra purpurea, reidentified as N. umbrosa, but the latter never go into the database (for BCI specimens, but yes for one at Sherman) or any publication. Dendropanax arboreus Accepted Dendropanax arboreus dendar Araliaceae (L.) Decne. & Planch. What we currently call all Dendropanax in the 50-ha plot, but variation may require 4 species. Some D. arboreus at BCI now recognized as D. caucanus by Cannon, or D. concinnus by N. Hensold (the latter not valid according to Tropicos). See Cannon and Cannon (1989) about variability of widespread D. arboreus, and Hensold suggests hybridization or introgression might be blurring boundaries. The group requires revision, and individuals within the plot may need to be divided. Garwood supports a division of Croat's D. arboreus into D. caucanus and D. arboreus. See also notes with D. stenodontus about yet another possible taxon. Dendropanax stenodontus Obsolete Dendropanax arboreus dendst Araliaceae Croat's key indicates differences between this and D. arboreus (adult size, leaf dentition), and Foster originally separated many individuals under this name. In 1996 paper, both D. stenodontus and D. arboreus appear. After that, we lumped all under D. arboreus. Cannon and Cannon are inconclusive about whether there might be a small-stature species, but D. stenodontus would not be the correct name. Possibly D. maritimus. Desmopsis panamensis Accepted Desmopsis panamensis des2pa Annonaceae (B.L.Rob.) Saff. Didymopanax morototoni Obsolete Schefflera morototoni sch2mo Araliaceae The name in Croat, and we used this at the very beginning of the 50-ha censuses. Now Schefflera morotoni, and we had made the change by the late 1980s. The entire genus Didymopanax was collapsed into Schefflera (Frodin 1975), which is interestingly prior to the date of Croat's book. Diospyros artanthifolia Accepted Diospyros artanthifolia dio2ar Ebenaceae Mart. ex Miq. Dipteryx oleifera Accepted Dipteryx oleifera diptpa Fabaceae-papilionoideae Benth. Formerly D. panamensis. The two names appear to be used interchangeably for a species collected from Nicaragua through the Colombian choco, plus one in Ecuador. At Tropicos, de Lima named BCI specimens D. oleifera in 1986, but Klitgaard renamed D. panamensis in 2007. D. oleifera is the early name, but without specimens. The rules committee voted to accept it (Brummitt 2011). Dipteryx panamensis Obsolete Dipteryx oleifera diptpa Fabaceae-papilionoideae Croat's name, and we used it in 1996 paper. We have now switched to D. oleifera, though Fl Meso online is back to D. panamensis. Zamora (2010) uses D. panamensis but adds that D. oleifera might be correct. Drypetes standleyi Accepted Drypetes standleyi drypst Putranjivaceae G.L.Webster Elaeis oleifera Accepted Elaeis oleifera elaeol Arecaceae (Kunth) Cortés Enterolobium schomburgkii Accepted Enterolobium schomburgkii entesc Fabaceae-mimosoideae (Benth.) Benth. Erythrina costaricensis Accepted Erythrina costaricensis ery1co Fabaceae-papilionoideae Micheli Erythroxylum macrophyllum Accepted Erythroxylum macrophyllum ery2ma Erythroxylaceae Cav. Formerly E. multiflorum, a name from Canal Area in 1940. Plowman synonomized it and other local names under the widespread and variable E. macrophyllum. See also Hammel et al (2010) treating E. multiflorum under this. Appendix of 1996 paper failed to list E. multiflorum as Croat’s name for this taxa. Erythroxylum multiflorum Obsolete Erythroxylum macrophyllum ery2ma Erythroxylaceae Croat’s name, but we had switched to E. macrophyllum in 1996 and 2004 papers, following Plowman1991. But Plowman acknowledges that E. multiflorum is distinctive, close to E. macrophyllum skutchii in Costa Rica and two other large-leaved subspecies of E. macrophyllum in S. America. Erythroxylum panamense Accepted Erythroxylum panamense ery2pa Erythroxylaceae Turcz. Eugenia coloradensis Obsolete Eugenia coloradoensis eugeco Myrtaceae Croat's misspelling of E. coloradoensis. We repeated in 1996 paper but corrected in 2004. Eugenia coloradoensis Accepted Eugenia coloradoensis eugeco Myrtaceae Standl. Eugenia galalonensis Accepted Eugenia galalonensis eugega Myrtaceae (C.Wright ex Griseb.) Krug & Urb. Eugenia nesiotica Accepted Eugenia nesiotica eugene Myrtaceae Standl. Eugenia oerstedeana Obsolete Eugenia oerstediana eugeoe Myrtaceae Croat’s misspelling of E. oerstediana, which we repeated in the 1996 paper. Eugenia oerstediana Accepted Eugenia oerstediana eugeoe Myrtaceae O.Berg This was listed under Croat’s misspelling, E. oerstedeana, in the 1996 paper. Faramea occidentalis Accepted Faramea occidentalis faraoc Rubiaceae (L.) A.Rich. Ficus aurea Obsolete Ficus trigonata ficutr Moraceae Nutt. We used F. trigonata at least until 1996 paper, but F. aurea replaced it. This is a mistake; it's the Florida species and is not known on BCI (but is in western Panama). Ficus bullenei Obsolete Ficus matiziana ficubu Moraceae I.M.Johnst. Croat’s name, and we used it 1996 and 2004 papers. Berg (2009) synonomized with F. matiziana, a S. American species. Garwood (2009) continued with F. bullenei. Ficus citrifolia Accepted Ficus citrifolia ficuci Moraceae Mill. Ficus colubrinae Accepted Ficus colubrinae ficuc1 Moraceae Standl. Ficus costaricana Accepted Ficus costaricana ficuc2 Moraceae (Liebm.) Miq. Ficus insipida Accepted Ficus insipida ficuin Moraceae Willd. Ficus matiziana Accepted Ficus matiziana ficubu Moraceae Dugand Formerly F. bullenei, following Croat, and appearing under that name in 1996 and 2004 papers. Garwood (2009) retained F. bullenei. Ficus maxima Accepted Ficus maxima ficuma Moraceae Mill. Ficus obtusifolia Accepted Ficus obtusifolia ficuob Moraceae Kunth Ficus pertusa Accepted Ficus pertusa ficupe Moraceae L.f. Ficus popenoei Accepted Ficus popenoei ficupo Moraceae Standl. Ficus tonduzii Accepted Ficus tonduzii ficuto Moraceae Standl. Ficus trigonata Accepted Ficus trigonata ficutr Moraceae L. Croat’s name and ours originally for the 50-ha plot, appearing in 1996 paper (though mispelled as F. trigonta). We then mistakely listed as F. aurea for several years. Garwood uses trigonata, but indicates a complex of distinct types. However, now Berg says Panama specimens should be F. crocata and that F. trigonata is restricted to the Antilles; he describes variation in detail. Ficus trigonta Obsolete Ficus trigonata ficutr Moraceae A publishing typo for F. trigonata in our 1996 paper. Ficus yoponensis Accepted Ficus yoponensis ficuyo Moraceae Desv. Garcinia intermedia Obsolete Garcinia recondita gar2in Clusiaceae (Pittier) Hammel Croat used the old name Rheedia edulis. Became Garcinia intermedia after Hammel (1989), who summarized various studies showing that Rheedia cannot be separated from Garcinia. As of 2015, however, G. intermedia at BCI was moved to G. recondita (see notes there). Garcinia madruno Accepted Garcinia madruno gar2ma Clusiaceae (Kunth) Hammel Formerly R. acuminata. See notes for G. intermedia and Hammel (1989). We had switched before 1996, so this is in 1996 and 2004 papers. Garcinia recondita Accepted Garcinia recondita gar2in Clusiaceae Hammel Hammel described this species around 2015, though still not published (Hammel pers. comm.). All BCI specimens of R. edulis/G. intermedia now identified at Tropicos as G. recondita, but G. intermedia remains valid elsewhere in Panama. Genipa americana Accepted Genipa americana geniam Rubiaceae L. Geonoma interrupta Accepted Geonoma interrupta geonin Arecaceae (Ruiz & Pav.) Mart. Guapira standleyana Accepted Guapira standleyana guapst Nyctaginaceae Woodson We misspelled this G. standleyanum in the 1996 paper, copying the error from Croat. Guapira standleyanum Obsolete Guapira standleyana guapst Nyctaginaceae We used this misspelling from Croat for many years, and the name is spelled wrong in the 1996 paper. It should be G. standleyana. Guarea bullata Accepted Guarea bullata guarsp Meliaceae Radlk. We finally identified the fuzzy-leaved Guarea morphospecies as this after 2004. G. bullata has been known in other parts of Panama since at least the 1960s. Impressed venation is distinctive, otherwise very similar and closely related to G. glabra. Guarea fuzzy Obsolete Guarea bullata guarsp Meliaceae Our temporary name for a Guarea with pubescent leaves in the 50 ha plot, by far the most abundant of unnamed morphospecies in the initial census. It is not at all like the two other Guarea on BCI. Finally identified as G. bullata, Guarea grandifolia Accepted Guarea grandifolia guargr Meliaceae (L.) Sleumer BCI specimens of G. multiflora were renamed this before plot started (Pennington in 1979 on Mobot sheets, eg Croat 8443, 11300). Guarea guidonia Accepted Guarea guidonia guargu Meliaceae (L.) Sleumer Identified as G. glabra in Croat. Foster reidentified by 1981; G. glabra should have glabrous ovary and few lenticels, small flowers relative to G. guidonia. Guarea multiflora Obsolete Guarea grandifolia guargr Meliaceae Adr. Juss. Croat's name for the large-leaved Guarea we call G. grandifolia in the 50-ha plot. We never used G. multiflora because Pennington re-identified Croat's BCI specimens in 1979. G. multiflora was named in Juisseau (1830) without any specimen or location; one of two Mobot specimens is a liana!. It does not appear in Pennington (2013), even the appendix. Guarea sp. 1 Obsolete Guarea bullata guarsp Meliaceae Alternate name for G. fuzzy, later identified as G. bullata, used in Condit et al. (2004). Unfortunate, because Guarea sp.1 was applied to a species in plot 33 in 1996 and is still in use. Guarea sp. nov. Obsolete Guarea bullata guarsp Meliaceae Alternate name for G. fuzzy, later identified as G. bullata, used in Condit et al. (1996). This mistakenly suggests we thought it would be a newly named species, but it turned out not to be. Guatteria dumetorum Obsolete Guatteria lucens guatdu Annonaceae R.E.Fr. Croat's name, and we followed for 30 years, appearing in 1996 and 2004 papers.. Maas (2015) decided to lump with G. lucens. Guatteria lucens Accepted Guatteria lucens guatdu Annonaceae Standl. We knew this as G. dumetorum for 30 years, and see notes there. Maas (2015) provides details about slight leaf differences between the dumetorum type and lucens type, but indicates that in all other ways the two are indistringuishable. Both types also have the same range, in Costa Rica and Panama, so though the species circumscription was widened by lumping, the range was not. Guazuma ulmifolia Accepted Guazuma ulmifolia guazul Malvaceae Lam. Guettarda foliacea Accepted Guettarda foliacea guetfo Rubiaceae Standl. Gustavia superba Accepted Gustavia superba gustsu Lecythidaceae (Kunth) O.Berg Hamelia axillaris Accepted Hamelia axillaris hameax Rubiaceae Sw. Hamelia patens Accepted Hamelia patens hamepa Rubiaceae Jacq. Hampea appendiculata Accepted Hampea appendiculata hampap Malvaceae (Donn.Sm.) Standl. Handroanthus guayacan Accepted Handroanthus guayacan tab1gu Bignoniaceae (Seem.) S.O.Grose Formerly Tabebuia guayacan, and appeared as that name in 1996 and 2004 papers. Handroanthus is an old name, and morphological distinctions were known since the 19th century. Hasseltia floribunda Accepted Hasseltia floribunda hassfl Salicaceae Kunth Heisteria acuminata Accepted Heisteria acuminata heisac Erythropalaceae (Humb. & Bonpl.) Engl. Known as H. longipes in Croat, but we used this name from the very start, and it appears in both 1996 and 2004 paper use this. Sleumer explains that Heisteria are not identifiable based only on flowers or fruits and leaves are needed. He collapsed many local species under H. acuminata, and all Tropicos specimens formerly called H. longipes now identified as H. acuminata by S. Knapp, R. Liesner, H. Sleumer. Heisteria concinna Accepted Heisteria concinna heisco Erythropalaceae Standl. Heisteria longipes Obsolete Heisteria acuminata heisac Erythropalaceae Standl. Croat's name, following Standley for Panama specimens. He describes range as Colombia-Costa Rica. Sleumer monograph lumps under H. acuminata after viewing many BCI specimens. We never used this name. Herrania purpurea Accepted Herrania purpurea herrpu Malvaceae (Pittier) R.E. Schult. Hieronyma alchorneoides Accepted Hieronyma alchorneoides hyeral Euphorbiaceae Allemão Franco reidentified BCI specimens in 1989 and published a major revision, collapsing H. laxiflora and other widespread varieties. Major broadening of range, however many local names are still listed as valid at Tropicos. Franco recognizes two varieties of H. alchorneoides, both among Croat's BCI specimens, but this is stipule difference between adults and juveniles (Hensold). Hirtella americana Accepted Hirtella americana hirtam Chrysobalanaceae L. Hirtella triandra Accepted Hirtella triandra hirttr Chrysobalanaceae Sw. Hura crepitans Accepted Hura crepitans huracr Euphorbiaceae L. Hybanthus prunifolius Obsolete Pombalia prunifolia hybapr Violaceae (Humb. & Bonpl. ex Schult.) Schulze-Menz Croat's name, and we used for 35 years and published in 1996 and 2004. de Paula Souza and Ballard (2014) revised Hybanthus based on polyphyly shown by DNA phylogeny. Several new genera will be needed. Hyeronima alcheornoides Obsolete Hieronyma alchorneoides hyeral Euphorbiaceae This is an unfortunate typo in the 1996 paper. The species name should be alchorneoides. Separately, the genus should be Hieronyma, but this error follows Croat. See notes with Hyeronima laxiflora. Hyeronima laxiflora Obsolete Hieronyma alchorneoides hyeral Euphorbiaceae Croat's name, note misspelling of genus (an old problem due to two spellings in original description). We originally used this name in 50-ha plot. Franco1990 synonomized under H. alchorneoides. Inga acuminata Accepted Inga acuminata ingas1 Fabaceae-mimosoideae Benth. Originally censused as Inga skinny, an unidentified morphospecies. Croat did not include this species, and we finally got it identified in 1989 (Zamora). It was not (??) known in Panama until then. We had it as I. acuminata in the 1996 paper. Inga cocleensis Accepted Inga cocleensis ingaco Fabaceae-mimosoideae Pittier The identification of this species in the 50-ha plot has been reassessed. See detailed notes under I. thibaudiana and I. unidentified. Inga fagifolia Obsolete Inga laurina ingafa Fabaceae-mimosoideae We followed Croat and used this name at the outset, and it appears in the 1996 paper. Now synonomized with I. laurina, following Pennington's (1997) revision of the entire genus, and I. laurina appears in the 2004 paper. Tropicos specimens from BCI identified as I. laurina by Sousan in 1990 or Pennington in 1996. Inga goldmanii Accepted Inga goldmanii ingago Fabaceae-mimosoideae Pittier Inga laurina Accepted Inga laurina ingafa Fabaceae-mimosoideae (Sw.) Willd. Originally I. fagifolia, following Croat, but I. laurina was synonomized with I. fagifolia in Pennington's (1997) revision of the entire genus. We used I. fagifolia in the 1996 paper but I. laurina in the 2004 paper. Inga marginata Accepted Inga marginata ingama Fabaceae-mimosoideae Willd. Inga minutula Obsolete Inga oerstediana ingami Fabaceae-mimosoideae We followed Croat and used this name at the outset, and it appears in the 1996 paper. Now synonomized with I. oerstediana in Pennington's (1997). Tropicos specimens from BCI identified as I. oerstediana by Sousan in 1990. Inga mucuna Accepted Inga mucuna ingam1 Fabaceae-mimosoideae Walp. & Duchass. Inga nobilis Accepted Inga nobilis ingaqu Fabaceae-mimosoideae Willd. Originally I. quaternata, following Croat, but we now follow Pennington (1997), in which I. quaternata was synonomized with I. nobilis. In 2004 paper, we had changed to this. C. American specimens are subsp. quaternata, and S. American are subsp. nobilis, but Pennington explains that there are intermediates in between and sometime one plant mixes the two different flower forms. We used I. quaternata in the 1996 paper but I. nobilis in the 2004 paper. See notes on recent confusion under I. quaternata. Inga oerstediana Accepted Inga oerstediana ingami Fabaceae-mimosoideae Benth. Originally I. minutula, following Croat, and appeared as such in the 1996 paper. Pennington's (1997) synonomized I. minutula under I. oerstediana, and Tropicos BCI specimens were identified as I. oerstediana in 1990 (Sousa). I. oerstediana is closely related to and possibly conspecific with I. edulis. Inga pavoniana Obsolete Inga sapindoides ingasa Fabaceae-mimosoideae A name appearing at Tropicos now, along with I. marginata.. It is accepted by Sousa (1993), and C. Romero applies at Tropicos. But N. Hensold points out I. marginata has priority, and Pennington and Man CR both use the latter. Inga pezizifera Accepted Inga pezizifera ingape Fabaceae-mimosoideae Benth. Inga punctata Accepted Inga punctata ingapu Fabaceae-mimosoideae Willd. Inga quaternata Obsolete Inga nobilis ingaqu Fabaceae-mimosoideae We followed Croat and used this in 1996 paper. I. quaternata was synonomized with I. nobilis in Pennington's (1997) revision of the entire genus, and use I. nobilis in 2004 paper. Confusion remains, though, with the name I. quaternata still appearing: Tropicos specimens from BCI identified as I. quaternata by C. Romero in 2006. Note confusion also in Zamora (2010), mentioning both names I. nobilis subsp. quaternata and I. quaternata as if they are distinct taxa. Inga ruiziana Accepted Inga ruiziana ingaru Fabaceae-mimosoideae G.Don Inga sapindoides Accepted Inga sapindoides ingasa Fabaceae-mimosoideae Willd. Sousa (1993) argues that I. pavoniana should be used, because original specimen of I. sapindoides lacked flowers and is questionable. Tropicos specimens IDed by C. Romero in 2006 as I. pavoniana (also R. Gereult in 1994). Pennington synonomizes the latter under I. sapindoides and has no discussion of the problem. Inga semialata Obsolete Inga marginata ingama Fabaceae-mimosoideae A name in use at Tropicos because Sousa uses it, but Pennington and Man CR use I. marginata. N. Henshold says I. marginata has precedence. Inga skinny Obsolete Inga acuminata ingas1 Fabaceae-mimosoideae An Inga with narrow leaflets not identified in the first census. In 1989, identified as I. acuminata. Inga spectabilis Accepted Inga spectabilis ingasp Fabaceae-mimosoideae (Vahl) Willd. Inga thibaudiana Accepted Inga thibaudiana ingath Fabaceae-mimosoideae DC. Included in Croat, but not recognized in the 50-ha plot until 2005, when it was separated from I. coclensis for the first time. Many trees previously known as I. coclensis had been alive since 1982, so we know that I. thibaudiana has always been part of the 50-ha plot. Inga umbellifera Accepted Inga umbellifera ingaum Fabaceae-mimosoideae (Vahl) Steud. Inga unidentified Accepted Inga unidentified ingaun Fabaceae-mimosoideae The Inga species I. coclensis and I. thibaudiana were reassessed in 2005, and many individuals of I. coclensis were reidentified as I. thibaudiana. Any tree known as I. coclensis that had died by 2005 could not be revisited, and all those had to be switched to I. unidentified. Jacaranda copaia Accepted Jacaranda copaia jac1co Bignoniaceae (Aubl.) D.Don Jacaratia spinosa Accepted Jacaratia spinosa jac2sp Caricaceae (Aubl.) A.DC. Croat included this taxon, but it did not appear in the 50-ha plot until the 2015 census. Koanophyllon wetmorei Accepted Koanophyllon wetmorei koanwe Asteraceae (B.L.Rob.) R.M.King & H.Rob. Lacistema aggregatum Accepted Lacistema aggregatum laciag Lacistemataceae (P.J.Bergius) Rusby Lacmellea panamensis Accepted Lacmellea panamensis lacmpa Apocynaceae (Woodson) Markgr. Laetia procera Accepted Laetia procera laetpr Salicaceae (Poepp.) Eichler Laetia thamnia Accepted Laetia thamnia laetth Salicaceae L. Lafoensia punicifolia Accepted Lafoensia punicifolia lafopu Lythraceae DC. Leandra dichotoma Accepted Leandra dichotoma leandi Melastomataceae (Pav. ex D. Don) Cogn. Leandra is now moved within Miconia, but names are not yet transferred as of 2017. Licania hypoleuca Accepted Licania hypoleuca licahy Chrysobalanaceae Benth. Licania platypus Accepted Licania platypus licapl Chrysobalanaceae (Hemsl.) Fritsch Lindackeria laurina Accepted Lindackeria laurina lindla Achariaceae C. Presl Lonchocarpus heptaphyllus Accepted Lonchocarpus heptaphyllus loncla Fabaceae-papilionoideae (Poir.) DC. The species appears as L. latifolia in the 1996 paper then L. latifolius in the 2004 paper (the latter is correct). Croat knew this as L. pentaphyllus. Zamora2010a synonomized L. latifolius under this. Lonchocarpus latifolia Obsolete Lonchocarpus heptaphyllus loncla Fabaceae-papilionoideae An incorrect spelling for L. latifolius that we used in the 1996 paper. But that name is now obsolete, and this goes by L. heptaphyllus. Lonchocarpus latifolius Obsolete Lonchocarpus heptaphyllus loncla Fabaceae-papilionoideae This was our first name for the Lonchocarpus species in the 50-ha plot. Croat knew it as L. pentaphyllus. Zamora (2010) synonomized L. latifolius and L. pentaphyllus under L. heptaphyllus. Lonchocarpus pentaphyllus Obsolete Lonchocarpus heptaphyllus loncla Fabaceae-papilionoideae (Poir.) DC. Croat’s name, now considered a synonym of L. heptaphyllus (Zamora 2010). Lopimia dasypetala Obsolete Pavonia dasypetala lopida Malvaceae This was Croat's name, and we used in both 1996 and 2004 papers. Now considered Pavonia dasypetala. Lopimia was described in 1823, but frequently treated under Pavonia since the 19th century. Differs in mucilage on fruits. No Pavonia DNA phylogeny is available. Lozania pittieri Accepted Lozania pittieri lozapi Lacistemataceae (S.F.Blake) L.B.Sm. Luehea seemannii Accepted Luehea seemannii luehse Malvaceae Triana & Planch. Lycianthes maxonii Accepted Lycianthes maxonii lycima Solanaceae Standl. Maclura tinctoria Accepted Maclura tinctoria maclti Moraceae (L.) D.Don ex Steud. This was listed as Chlorophora tinctoria in the 2004 paper, but M. tinctoria in the 1996 paper. Neither name appears in Croat, and D'Arcy used Chlorophora. Our brief use of Chorophora was erroneous, since Berg's (2001) monograph clearly used Maclura. Macrocnemum glabrescens Obsolete Macrocnemum roseum macrgl Rubiaceae Croat's name and our original name, appearing in 1996 paper. Changed to M. roseum after 2000. Macrocnemum roseum Accepted Macrocnemum roseum macrgl Rubiaceae (Ruiz & Pav.) Wedd. Formerly M. glabrescens, but that C. American form was lumped under this widespread S. American species (Taylor pers. comm.). Appears as M. glabrescens in 1996 paper. [I need Lorence 1999 on C. American Rubiaceae.] Malmea garwood's Obsolete Mosannona garwoodii malmsp Annonaceae One name used in some early databases for the new Annonaceae, christened finally as a novel species, Mosannona garwoodii. Malmea Nancy?? Obsolete Mosannona garwoodii malmsp Annonaceae One name used in some early databases for the new Annonaceae, christened finally as a novel species, Mosannona garwoodii. For a period in the early censuses, we thought it might be a novel Malmea. Malmea sp. nov. Obsolete Mosannona garwoodii malmsp Annonaceae This was the name we used in the 1996 paper for the novel Annonaceae, later described as Mosannona garwoodii. Malpighia romeroana Accepted Malpighia romeroana malpro Malpighiaceae Cuatrec. Maquira costaricana Obsolete Maquira guianensis maquco Moraceae We took this name from Croat and used through the 1996 paper, but switched to M. guianensis for 2004 paper, following Berg (1972). With more specimens of M. guianensis from Ecuador, Berg could no longer separate M. costaricensus (Ecuador and north) and M. guianensis (across S. America). Now, Berg (2001) considers the former a subspecies of the latter. Maquira guianensis Accepted Maquira guianensis maquco Moraceae Aubl. Formerly M. costaricana. We switched after Berg (2001), and the 2004 paper used M. guianensis. See notes with M. costaricana. Margaritaria nobilis Accepted Margaritaria nobilis margno Phyllanthaceae L.f. Croat includes this taxon on BCI, but we did not identify in the 50-ha plot until 1990. A few individuals were in the census from the start, but remained unidentified until then. Marila laxiflora Accepted Marila laxiflora mar1la Clusiaceae Rusby Maytenus schippii Accepted Maytenus schippii maytsc Celastraceae Lundell Miconia affinis Accepted Miconia affinis micoaf Melastomataceae DC. Miconia argentea Accepted Miconia argentea micoar Melastomataceae (Sw.) DC. Miconia dorsiloba Accepted Miconia dorsiloba micodo Melastomataceae Gleason Originally called morphospecies Miconia sp.15 2000, identified as M. dorsiloba in 2005. Does not appear in Croat. Miconia elata Accepted Miconia elata micoel Melastomataceae (Sw.) DC. Miconia hondurensis Accepted Miconia hondurensis micoho Melastomataceae Donn. Sm. Miconia impetiolaris Accepted Miconia impetiolaris micoim Melastomataceae (Sw.) D. Don ex DC. Miconia nervosa Accepted Miconia nervosa micone Melastomataceae (Sm.) Triana Miconia prasina Accepted Miconia prasina micopr Melastomataceae (Sw.) DC. Miconia sp.15 Obsolete Miconia dorsiloba mico15 Melastomataceae A Miconia new to the 50-ha plot recruited in 2000. It remained until 2005, when another recruited. It was then identified as M. dorsiloba. Morinda seibertii Obsolete Appunia seibertii appuse Rubiaceae (Standl.) Steyerm. We originally used Appunia seibertii, but then followed Steyermark (1972) in moving Appunia within the broad Morinda. But recent DNA phylogeny reinstated Appunia. See more notes there. Mosannona garwoodii Accepted Mosannona garwoodii malmsp Annonaceae Chatrou & Welzenis The official species name, maintaining Nancy Garwood's, described by Chatrou (1998). This correct version appeared in our 2004 paper. The species was thus first described from the BCI 50-ha plot. It is very distinctive, and common in the plot. It had several morphospecies names before the final description, including the genera Malmea and Cremastosperma. Croat called in Crematosperma sp., misspelling the genus. Mosannona garwoods Obsolete Mosannona garwoodii malmsp Annonaceae One briefly used name for the new Annonaceae, after P. Maas saw it and placed it in a new Neotropical genus. Chatrou (1998) eventually christened it as a novel species, Mosannona garwoodii. Mouriri myrtilloides Accepted Mouriri myrtilloides mourmy Melastomataceae (Sw.) Poir. Myrcia gatunensis Obsolete Myrcia splendens tip. gatunensis myrcga Myrtaceae Standl. The name from Croat, and we used in 1996 paper and onward until 2017. Tropicos, citing Flora Mesoamerica and Flora Costa Rica, merges into M. splendens, a very widespread neotropical species. FMNH specimens also use M. splendens, det by Kawasaki 2005. But we have M. splendens separate (formerly myrc3), including many individuals of M. gatunensis and M. splendens together in plot 6. Kawasaki (pers. comm.) acknowledges that M. splendens is variable and needs revision. For now, we maintain M. gatunensis. Myrcia splendens tip. gatunensis Accepted Myrcia splendens tip. gatunensis myrcga Myrtaceae (Sw.) DC. See sp.3, the morphospecies eventually identified as this. See also M. gatunensis, whose BCI specimens are now identified as M. splendens. However, plot 6 has many individuals of both this species and M. gatunensis growing side by side. Myrospermum frutescens Accepted Myrospermum frutescens myrofr Fabaceae-papilionoideae Jacq. Not in Croat but we identified in 1982 census. Nectandra cissiflora Accepted Nectandra cissiflora nectci Lauraceae Nees Nectandra fuzzy Accepted Nectandra fuzzy nects1 Lauraceae An unidentified Lauraceae, tentatively placed in genus Nectandra. We have most consistently called it Nectandra fuzzy, but it was listed as Nectandra sp. nov. 1 in the 1996 paper, though we are not certain it's a novel species. The 50-ha plot has 11 individuals assigned the name. Nectandra globosa Obsolete Nectandra lineata nectgl Lauraceae This is Croat's name, and we originally used it. But Croat's BCI specimens are now identified as N. lineata (Rohwer 1993). N. globosa is valid but only occurs in Guyana. We used N. globosa in the 1996 paper, but the corrected N. lineata in the 2004 paper. Nectandra lineata Accepted Nectandra lineata nectgl Lauraceae (Kunth) Rohwer Our current identification for this Nectandra, following Rohwer (1993) who explains the source of Croat's mistaken identifcation as N. globosa. Nectandra purpurascens Obsolete Damburneya umbrosa nectpu Lauraceae (Ruiz & Pav.) Mez Croat used this name, but it is an error. Authority given by Croat matches that of N. purpurea, so Rowher assumed that's what Croat meant. Nectandra purpurea Obsolete Damburneya umbrosa nectpu Lauraceae (Ruiz & Pav.) Mez We used this in 1996 and 2004 papers, following but correcting Croat's name (he mispelled it as N. purpurascens). But Rohwer (1993) identified BCI specimens as N. umbrosa (N. purpurea is valid, and similar, but not on BCI). As of 2016, though, this section of Nectandra was moved into genus Damburneya, so now we call it D. umbrosa. Note, however, that van der Werff in 2016 IDed Croat 5053 from BCI as D. purpurea at same time as he named all other BCI specimens as D. umbrosa. Nectandra savannarum Obsolete Damburneya umbrosa nectpu Lauraceae (Standl. & Steyerm.) C.K. Allen We never used this in 50-ha plot data. Croat used it for a single specimen from BCI (Foster 960 at Tropicos), matching it to a species described in Guatemalan savanna (Allen 1945). That one specimen was later determined (on Tropicos specimen) to be the same as the other common Nectandra (now Damburneya umbrosa, what Croat called N. purpurascens). Nectandra sp._4 Obsolete Nectandra sp.4_(tiny_leaf) nects3 Lauraceae Nectandra sp.4_(tiny_leaf) Accepted Nectandra sp.4_(tiny_leaf) nects3 Lauraceae This extremely rare Lauraceae, with just one individual in the 1982 census of the 50-ha plot, was never identified, and we are not certain it is a Nectandra. The single individual died after 1985 and we have never seen it elsewhere, so it will probably never be identified. It had very small leaves, hence the morphospecies name, but it was listed as Nectandra sp. nov. 3 in the 1996 paper. The combination of sp. 4 and sp. 3 as parts of the name for the same morphospecies resulted from confusion caused by several other unidentified Nectandra in our Panama database. Nectandra sp.nov.1 Obsolete Nectandra fuzzy nects1 Lauraceae This is an alternative name for the unidentified morphospecies we call Nectandra fuzzy. It was coined for the 1996 paper to sound more professional than N. fuzzy. Nectandra sp.nov.3 Obsolete Nectandra sp.4_(tiny_leaf) nects3 Lauraceae This is an alternative name for the unidentified morphospecies we refer to as Nectandra sp.4_(tiny_leaf) in all databases. It was coined for the 1996 paper. The combination of sp. 4 and sp. 3 as parts of the name for the same morphospecies resulted from confusion caused by several other unidentified Nectandra in our Panama database. Nectandra umbrosa Obsolete Damburneya umbrosa nectum Lauraceae (Kunth) Mez Rolando used this name in Santa Rita transect, but he says it is same as BCI specimens with code NECTPU. So all taxa we once called N. purpurea are now Damburneya umbrosa, maintaining code NECTPU. At BCI we never used N. umbrosa, because by the time we discovered Rowher's reidentification of N. purpurea, it had been transferred to the genus Damburneya. Neea amplifolia Accepted Neea amplifolia neeaam Nyctaginaceae Donn.Sm. Ochroma pyramidale Accepted Ochroma pyramidale ochrpy Malvaceae (Cav. ex Lam.) Urb. Ocotea cernua Accepted Ocotea cernua ocotce Lauraceae (Nees) Mez Ocotea oblonga Accepted Ocotea oblonga ocotob Lauraceae (Meisn.) Mez Ocotea puberula Accepted Ocotea puberula ocotpu Lauraceae (Rich.) Nees Croat called this O. pyramidata, but we changed to O. puberula by the late 1980s. Van der Weff had already named BCI specimens O. puberula in 1983. See Van der Werff (2002). Ocotea pyrimidata Obsolete Ocotea puberula ocotpu Lauraceae Croat's named, and we used at the start of the 50-ha plot. We had switched to O. puberula before 1990, and both 1996 and 2004 papers list it as O. puberula. See notes with O. puberula. Ocotea skutchii Obsolete Ocotea whitei ocotwh Lauraceae Croat used this name, and we followed Croat at the very beginning. By the late 1980s, we switched to O. whitei, and both 1996 and 2004 paper used O. whitei. Van der Werff (2002) published the synonomy. Ocotea whitei Accepted Ocotea whitei ocotwh Lauraceae Woodson Croat used O. skutchii, but that name was synonomized soon after his book. See van der Werff (2002), who acknowledged variability of O. whitei. Oenocarpus mapora Accepted Oenocarpus mapora oenoma Arecaceae H.Karst. Formerly O. panamanus, in Croat and in the first census. Oenocarpus mapoura Obsolete Oenocarpus mapora oenoma Arecaceae This is a misspelling of O. mapora that appeared in our 1996 paper. We spelled it correctly in 2004 paper. Oenocarpus panamanus Obsolete Oenocarpus mapora oenoma Arecaceae Croat's name, a name used only in Costa Rica and Panama. We used it at the start of the 50-ha plot but had switched to O. mapora by the late 1980s. Henderson1997 officially lumped this with the widespread O. mapora, along with several other local names in the genus. Olmedia aspera Obsolete Trophis caucana olmeas Moraceae Our early name, from Croat, now Trophis caucana. Berg (1972) in early Moraceae monograph separated this as a monotypic genus from its earlier inclusion in Trophis. In Berg (1988, 2001), he reversed, saying that Trophis morphology had expanded with T. involucrata, and the wider variation accommodates Olmedia. Oreopanax capitatus Accepted Oreopanax capitatus oreoca Araliaceae (Jacq.) Decne. & Planch. Croat includes this, but it did not appear in the 50-ha plot until 2010. The sole individual then died by 2015. Ormosia amazonica Accepted Ormosia amazonica ormoam Fabaceae-papilionoideae Ducke Not in Croat but identified in 1982 census. Ormosia coccinea Accepted Ormosia coccinea ormocr Fabaceae-papilionoideae (Aubl.) Jacks. Current identification for a moderately common tree in the plot. Croat listed this form as O. croatii, an unofficial name, but it has been identified at Tropicos in 1978 (MO Dillon). N. Zamora confirmed in 2007 (pers. Comm. To N. Hensold). Ormosia croatii Obsolete Ormosia coccinea ormocr Fabaceae-papilionoideae Croat used this name, but it is not a published name. Later, MO Dillon and N. Zamora identified this as O. coccinea. Ormosia macrocalyx Accepted Ormosia macrocalyx ormoma Fabaceae-papilionoideae Ducke Ouratea lucens Accepted Ouratea lucens ouralu Ochnaceae (Kunth) Engl. Pachira quinata Obsolete Pochota quinata pochqu Malvaceae (Jacq.) W.S.Alverson Croat used Bombacopsis quinata, but by 1996 paper we had updated to Pochota quinata, then we changed to Pachira quinata by 2004 paper. But now we have to go back to Pochota quinata, following a recent DNA phylogeny. This makes the nomenclatural history of this taxon the most confusing of all in our database. The genera Bombacopsis, Pochota, and Pachira have been variously used due to taxonomic disagreement. They were coined in 1916, 1909, and 1775 respectively, and there has been a decades long argument about whether Bombacopsis and Pochota refer to the same plants and which takes precedence. At BCI, we originally followed Croat and used Bombacopsis, but Stevens (1987) argued in favor of Pochota as the earlier name. Subsequently, Alverson (1994) argued that they should all fall within Pachira, because none of the characters used to separate Pochota from Pachira were consistent; he acknowledged they may be separate but more evidence was needed. Sure enough, a DNA phylogeny from 2011 does not support Pachira, unless Eriotheca and Bombax are subsumed, and the genus Pochota was again resurrected for P. quinata (but not P. sessilis, which belongs in Pachira). Pachira sessilis Accepted Pachira sessilis pochse Malvaceae Benth. Bombacopsis sessilis according to Croat. In 1996 paper, it appeared as Pochota sessilis, but we updated to Pachira sessilis for 2004 paper. See genus discussion under P. quinata Palicourea acuminata Accepted Palicourea acuminata psycac Rubiaceae (Benth.) Borhidi Formerly Psychotria acumintata. Several psychotria moved to Palicourea due to improved character understanding (see Psych. acuminata). Palicourea brachiata Accepted Palicourea brachiata psycb1 Rubiaceae (Sw.) Borhidi Formerly Psychotria brachiata, and appearing under that old name in 1996 paper. Palicourea cuspidata Obsolete Palicourea acuminata psycac Rubiaceae (Benth.) Borhidi This was an error and is now corrected to P. acuminata. The mistake traces to misidentification of Psychotria acuminata as P. cuspidata in Panama (Taylor pers. comm.). When moving the name under Palicourea, we carried the incorrect new combination for a short time. Croat has the correct Psychotria acuminata and mentions the confusion with Psychotria cuspidata. Palicourea cyanococca Accepted Palicourea cyanococca psycpi Rubiaceae (Seem. ex Dombrain) Borhidi Palicourea deflexa Accepted Palicourea deflexa psycde Rubiaceae (DC.) Borhidi Formerly Psychotria deflexa, and appearing under that old name in 1996 and 2004 papers. Palicourea guianensis Accepted Palicourea guianensis paligu Rubiaceae Aubl. Palicourea hoffmannseggiana Accepted Palicourea hoffmannseggiana psycfu Rubiaceae (Willd. ex Roem. & Schult.) Borhidi Psychotria furcata in Croat and the 1996 paper. Later Psychotria hoffmannseggiana before it was moved under Palicourea with many other Psychotria. Palicourea racemosa Accepted Palicourea racemosa psycra Rubiaceae (Aubl.) Borhidi Formerly Psychotria racemosa, and appearing under that old name in 1996 and 2004 papers. Pavonia dasypetala Accepted Pavonia dasypetala lopida Malvaceae Turcz. Formerly Lopimia dasypetala, following Croat, but this is the widely accepted classification, placing Lopimia within Pavonia (Fryxell). Hammel et al VI (2007) uses P. dasypetala without even mentioning Lopimia as synonym. This appears under the old L. dasypetala in 1996 and 2004 papers. Pentagonia macrophylla Accepted Pentagonia macrophylla pentma Rubiaceae Benth. Perebea xanthochyma Accepted Perebea xanthochyma perexa Moraceae H.Karst. Phoebe cinnamomifolia Obsolete Cinnamomum triplinerve phoeci Lauraceae We used this in 1996 and 2004 papers for what Croat called P. mexicanum. We now consider Cinnamomum triplinerve the valid name. Phoebe cinnamomifolium Obsolete Cinnamomum triplinerve phoeci Lauraceae (Kunth) Nees This is a misspelling that showed up briefly in our database. We spelled it correctly, P. cinnamomifolia, in 1996 and 2004 papers. We now use Cinnamomum triplinerve for this taxon. Phoebe mexicana Obsolete Cinnamomum triplinerve phoeci Lauraceae Meisn. Croat used this name, but we changed to Phoebe cinnamomifolium for the 1996 and 2004 papers. That too is obsolete though, and we now use is Cinnamomum triplinerve. The combination C. cinnamomifolium has also been used, but never in our database. Picramnia latifolia Accepted Picramnia latifolia picrla Picramniaceae Tul. Piper aequale Accepted Piper aequale pipeae Piperaceae Vahl Piper arboreum Accepted Piper arboreum pipea1 Piperaceae Aubl. Piper carrilloanum Obsolete Piper schiedeanum pipeca Piperaceae This is Croat's name, but Tebbs synonomized with P. schiedeanum in 1990. Most of Croat's BCI specimens are named P. schiedeanum now. We used P. carrilloanum in the 1996 paper. According to Callejas, P. carrilloanum remains a valid species, and in Panama, but not on BCI. Piper colonense Accepted Piper colonense pipecu Piperaceae C.DC. Piper cordulatum Accepted Piper cordulatum pipeco Piperaceae C. DC. Piper culebranum Obsolete Piper colonense pipecu Piperaceae Croat's BCI specimens at Tropicos identified (R. Callejas) as P. culebranum in 2006. However, Tebbs identified them as P. colonense in 1990. Callejas (and Tropicos) consider P. colonense and P. culebranum both valid species, both in central Panama, but BCI specimens are P. culebranum. Our 2004 paper has both species!! Piper imperiale Accepted Piper imperiale pipeim Piperaceae (Miq.) C.DC. Croat had this correct, and our 1996 paper had it correct. At one point, the misspelling P. imperialis got in our database. Piper imperialis Obsolete Piper imperiale pipeim Piperaceae (Miq.) C. DC. Correctly spelled P. imperiale. This bad spelling got into our database at one point, but never in any publicatcion. Piper longispicum Accepted Piper longispicum pipelo Piperaceae C.DC. First identified in 50-ha plot in 2015. Not in Croat nor in 1996 or 2004 papers. Piper perlasense Accepted Piper perlasense pipepe Piperaceae Yunck. Piper reticulatum Accepted Piper reticulatum pipere Piperaceae L. Piper schiedeanum Accepted Piper schiedeanum pipeca Piperaceae Steud. Formerly P. carrilloanum in our data, and the 1996 paper, but we now follow Tebbs and use P. schiedeanum. However, a substantial confusion has arisen because Callejas identified many of Croat's BCI specimens as P. paulowniifolium. Flora Nicaragua by Callejas?? Pithecellobium macradenium Obsolete Abarema macradenia pit1ma Fabaceae-mimosoideae Currently Abarema macradenia. We used the old name in 1996 paper. Barnebey and Grimes (1996) detailed phylogeny of Ingeae reduced Pithecelllobium enormously, and this taxon was transferred to Abarema, as A. macradenia. Pithecellobium rufescens Obsolete Cojoba rufescens pit1ru Fabaceae-mimosoideae This is Croat’s name, and we used in name in 1996 paper but had updated to Cojoba rufescens in 2004 paper. Barnebey and Grimes (1996) detailed phylogeny of Ingeae reduced Pithecelllobium enormously. Platymiscium pinnatum Accepted Platymiscium pinnatum pla1pi Fabaceae-papilionoideae (Jacq.) Dugand Platypodium elegans Accepted Platypodium elegans pla2el Fabaceae-papilionoideae Vogel Pochota quinata Accepted Pochota quinata pochqu Bombacaceae This has been our most convoluted series of name changes. This was first Bombacopsis quinata, then Pochota quinata (which appears in 1996 paper), then Pachira quinata (in the 2004 paper), but now back to Pochota quinata. However, Pachira sessilis remains. See notes under Pachira quinata. Pochota sessilis Obsolete Pachira sessilis pochse Bombacaceae Current known as Pachira sessilis. See notes under Pachira quinata. Pombalia prunifolia Accepted Pombalia prunifolia hybapr Violaceae (Humb. & Bonpl. ex Willd.) Paula-Souza Originally Hybanthus prunifolius following Croat. Several new genera will be needed for Hybanthus (de Paula Souza and Balland 2014). See notes with H. prunifolius. Posoqueria latifolia Accepted Posoqueria latifolia posola Rubiaceae (Rudge) Schult. Poulsenia armata Accepted Poulsenia armata poular Moraceae (Miq.) Standl. Pourouma bicolor Accepted Pourouma bicolor pourbi Urticaceae Mart. This was P. guianensis in Croat, but we followed Berg and changed to this by the late 1980s. P. bicolor appears in 1996 and 2004 papers. A Tropicos specimen now carries the name P. chocoana, but Berg used P. bicolor subsp. chocoana and this is still accepted everywhere (see Flora Mes, CR). Pourouma guianensis Obsolete Pourouma bicolor pourbi Urticaceae Croat's name for the BCI Pourouma, and we used it at the outset of the 50-ha plot. Berg identified BCI specimens as P. bicolor subsp. chocoana. P. guianensis remains a valid, S. American taxa. P. guianensis never appeared in publications. Pouteria fossicola Accepted Pouteria fossicola poutfo Sapotaceae Cronquist Pouteria reticulata Accepted Pouteria reticulata poutre Sapotaceae (Engl.) Eyma We originally followed Croat and called this P. unilocularis, a C. American taxon. Pennington lumped it and 4 other species, and the combined group is widespread and variable. We switched before any publicacions, so this appears as P. reticulata in both 1996 and 2004 papers. Pouteria stipitata Accepted Pouteria stipitata poutst Sapotaceae Cronquist Pouteria unilocularis Obsolete Pouteria reticulata poutre Sapotaceae Croat used this C. American name, and we followed at the outset of the 50-ha plot. Pennington (1990) recognizes distinctions, but lumped 5 species, including this C. American form, under P. reticulata, and we made the switch by 1990. This appears as P. reticulata in both 1996 and 2004 papers. Prioria copaifera Accepted Prioria copaifera pri2co Fabaceae-caesalpinioideae Griseb. Protium confusum Accepted Protium confusum protsp Burseraceae (Rose) Pittier This Protium could not be identified from Croat and was censused as a morphospecies until 2000, called Protium skinny; in the 1996 paper it was called Protium sp. nov. It was finally matched to P. confusum, not a new species. Protium costaricense Accepted Protium costaricense protco Burseraceae (Rose) Engl. Protium panamense Accepted Protium panamense protpa Burseraceae (Rose) I.M.Johnst. Protium skinny Obsolete Protium confusum protsp Burseraceae A morphospecies name for a distinct but unidentified Protium with narrow leaflets. Finally identified as P. confusum. Protium sp. nov. Obsolete Protium confusum protsp Burseraceae The name appearing in our 1996 paper for the unidentified Protium, known in our early databases as Protium skinny, later identified as P. confusum. Protium tenuifolium Obsolete Protium tenuifolium subsp. sessiliflorum protte Burseraceae Engl. A subspecies distinct from the BCI species, known only at two distant plots. Protium tenuifolium subsp. sessiliflorum Accepted Protium tenuifolium subsp. sessiliflorum protte Burseraceae Engl. A subspecies distinct from the BCI species, known only at two distant plots. Pseudobombax septenatum Accepted Pseudobombax septenatum pse1se Malvaceae (Jacq.) Dugand Psidium anglohondurense Obsolete Chamguava schippii cha2sc Myrtaceae Croat used this name, and we used this during the 1980s. We switched to Chamguava schippii by the 1996 and 2004 papers, following Landrum’s (1991) revision. See notes with the latter. Psidium friedrichsthalianum Accepted Psidium friedrichsthalianum psidfr Myrtaceae (O.Berg) Nied. Psychotria acuminata Obsolete Palicourea acuminata psycac Rubiaceae Benth. This is name that Croat used. C.M. Taylor et al (2015) moved several Psychotria to Palicourea. Taylor (pers. comm.) indicates many past collectors did not correctly understand character differences between genera. Many more Psychotria will be moved. See also Borhidi (2011) about the transfer. Psychotria brachiata Obsolete Palicourea brachiata psycb1 Rubiaceae Sw. This is Croat’s name, and we used in 1996 paper. Moved to Psychotria following Borhidi2011. Psychotria chagrensis Accepted Psychotria chagrensis psycch Rubiaceae Standl. Psychotria cyanococca Obsolete Palicourea cyanococca psycpi Rubiaceae Seem. ex Dombrain Currently P. cyanococca, a prior name. No change in species circumscription (Taylor pers. comm.). Psychotria deflexa Obsolete Palicourea deflexa psycde Rubiaceae DC. This is Croat’s name, and we used in 1996 and 2004 papers. Moved to Psychotria following Borhidi2011. Psychotria furcata Obsolete Palicourea hoffmannseggiana psycfu Rubiaceae Croat's name and the name appearing in 1996 paper, but we switched to P. hoffmannseggiana, following Burger and Taylor (Flora Costaricensis). This lumps the C. American/Choco P. furcata into a widespread species. As of 2011, this and many other Psychotria were moved under Palicourea. Psychotria graciliflora Accepted Psychotria graciliflora psycg1 Rubiaceae Benth. Not in Croat but identified in 1982 census. Psychotria granadensis Obsolete Psychotria tenuifolia psycg2 Rubiaceae Croat's name and our original name, and we used in the 1996 paper. We later switched to P. tenuifolia following Hamilton (1989) Psychotria of C. America (Taylor pers. comm.). Psychotria grandis Accepted Psychotria grandis psycg3 Rubiaceae Sw. Psychotria hoffmannseggiana Obsolete Palicourea hoffmannseggiana psycfu Rubiaceae (Willd. ex Schult.) Müll.Arg. Formerly P. furcata, but Burger and Taylor lumped C. American group under this. To further confusion, this was switched under Palicourea by Taylor (pers. comm.) and Borhidi2011 in the big revision of New World Psychotria. Psychotria horizontalis Accepted Psychotria horizontalis psycho Rubiaceae Sw. Psychotria limonensis Accepted Psychotria limonensis psycli Rubiaceae K.Krause Psychotria marginata Accepted Psychotria marginata psycma Rubiaceae Sw. Psychotria pittieri Obsolete Palicourea cyanococca psycpi Rubiaceae Standl. Croat’s name, and we used in 1996 paper. Later switched to P. cyanococca, an earlier name for the same species found in Flora Nicaragua. Psychotria psychotriifolia Accepted Psychotria psychotriifolia psycps Rubiaceae (Seem.) Standl. Croat includes this, but it did not appear in the 50-ha plot until 1995 and is thus not in the 1996 paper. Psychotria racemosa Obsolete Palicourea racemosa psycra Rubiaceae Rich. This is Croat’s name, and we used in 1996 paper. Moved to Psychotria following Borhidi2011. Psychotria tenuifolia Accepted Psychotria tenuifolia psycg2 Rubiaceae Sw. Formerly P. granadensis and published under that name in 1996 paper. Hamilton (1989) synonomized under this widespread S. American form, and Taylor (pers. comm.) agrees. Pterocarpus belizense Obsolete Pterocarpus officinalis pterbe Fabaceae-papilionoideae An old misspelling of P. belizensis briefly in our database, never appearing in print. See notes with P. belizensis. Pterocarpus belizensis Obsolete Pterocarpus officinalis pterbe Fabaceae-papilionoideae Standl. The 1996 paper uses this name for the rare BCI species, ie P. officianalis. Tropicos shows this as synonym of P. officianalis. But Henshold lists it as synonym of P. rohrii=P. hayesii, though that appears to be a misunderstanding. Robin believes he found a single specimen of a distinct Pterocarpus on BCI, ie a third species, and tentatively considered it was P. belizensis. Pterocarpus hayesii Accepted Pterocarpus hayesii pterro Fabaceae-papilionoideae Hemsl. Formerly P. rohrii, and that old name was used in 1996 and 2004 papers. Pterocarpus officinalis Accepted Pterocarpus officinalis pterof Fabaceae-papilionoideae Jacq. A swamp species, rare on BCI, only 9 in plot and dying off. We listed it as P. belizensis in 1996 paper, but that appears to be a miscommunication. This has always been P. officianalis since Croat's identification. Pterocarpus rohrii Obsolete Pterocarpus hayesii pterro Fabaceae-papilionoideae Vahl Croat’s name, and we followed in 1996 and 2004 papers. Later, following N. Zamora, we consider this P. hayesii, contra Tropicos. According to Zamora, P. hayesii and P. rohrii are different species, both valid, but P. rohrii is only in South America. Quararibea asterolepis Accepted Quararibea asterolepis quaras Malvaceae Pittier Whether BCI specimens belong in Q. asterolepis subsp. stenophylla, Q. stenophylla, or Q. asterolepis, is unresolved at Tropicos. Quassia amara Accepted Quassia amara quasam Simaroubaceae L. Randia armata Accepted Randia armata randar Rubiaceae (Sw.) DC. Randia formosa Obsolete Rosenbergiodendron formosum randfo Rubiaceae Croat's and our original name, and we used it in 1996 paper. This, with a small set of Randia species, were split off into a new genus, and from 2004 paper this became Rosenbergiodendron formosum Rauvolfia littoralis Accepted Rauvolfia littoralis rauvli Apocynaceae Rusby Does not appear in Croat. Discovered on BCI in 50-ha plot in 1995 census. Rheedia acuminata Obsolete Garcinia madruno gar2ma Clusiaceae Croat’s name, now Garcinia madruno. See notes with R. edulis and Hammel (1989). We never published this name because we had changed before 1996. Rheedia edulis Obsolete Garcinia recondita gar2in Clusiaceae Croat’s name, but Garcinia intermedia in the 1996 paper, now Garcinia recondita. We never published Rheedia because we had changed before 1996. Need for moving Rheedia to Garcinia was noted as early as 1958 by Robson (on African flora). Rheedia had been New World, but traits separating it from Garcinia are inconsistent. Hammel (1989) assigned new combination G. intermedia, since G. edulis was not available. No change in species boundary. Hammel’s reidentification of BCI specimens as G. recondita still not published as of 2017 (Hammel pers. comm.) Rinorea sylvatica Accepted Rinorea sylvatica rinosy Violaceae (Seem.) Kuntze Rosenbergiodendron formosum Accepted Rosenbergiodendron formosum randfo Rubiaceae (Jacq.) Fagerl. Formerly Randia formosa and appearing as such in 1996 paper, changed to Rosenbergiodendron formosum by 2004 paper. This and 3 other species were shown to be outside Randia (DNA and morphology). The species circumscription was nearly unchanged, though a Guyana segregate was split, so the new R. formosum has a slightly smaller range (Taylor pers. comm.). Sapium aucuparium Obsolete Sapium glandulosum sapiau Euphorbiaceae We used this name in 1996 paper for what Croat called S. caudatum. Then Krujit (1996) moved S. aucuparium under S. glandulosum, which we have adopted. Specimens at Tropicos identified by R. Liesner as S. glandulosum. See notes under S. caudatum for further confusion about Croat’s names. Sapium broadleaf Accepted Sapium broadleaf sapisp Euphorbiaceae Sapium caudatum Obsolete Sapium glandulosum sapiau Euphorbiaceae Pitt. There has been much confusion about Sapium names. Croat used this name, but Webster and Huft (1988) moved S. caudatum under S. aucuparium and discussed problems. We followed Croat and used S. caudatum when the 50-ha censuses started, but switched to S. aucuparium by the late 1980s. Croat also used S. aucuparium for a different taxon, but his description matches S. laurifolium. The latter, however, has never otherwise observed or collected at BCI, and that specimen should be S. glandulosum. Sapium glandulosum Accepted Sapium glandulosum sapiau Euphorbiaceae (L.) Morong Krujit synonomized S. aucuparium with this. R. Liesner identifed BCI specimens as S. glandulosum. Sapium sp 1 Obsolete Sapium broadleaf sapisp Euphorbiaceae We used this name in the 2004 paper for the Sapium in the Sherman plot. They match Sapium broadleaf, the yet-to-be-described new species at BCI. Sapium sp. nov. Obsolete Sapium broadleaf sapisp Euphorbiaceae The name we used in 1996 paper for what we call Sapium broadleaf. We believe it is a new species, yet undescribed, and M. Huft (pers. comm.) confirms. The Foster BCI specimen from the 50-ha plot remains unnamed at Tropicos (Foster 2942). Scheelea zonensis Obsolete Attalea rostrata sch1zo Arecaceae Croat’s name, and this appears in 1996 paper. Henderson eliminated Scheelea and moved into Attalea. This became A. butyracea in the 2004 paper. Schefflera morototoni Accepted Schefflera morototoni sch2mo Araliaceae (Aubl.) Maguire, Steyerm. & Frodin We originally used Didymopanax morototoni for this, following Croat. But Frodin (1975) moved the entire genus Didymopanax into Schefflera, as new species added blurred distinctions. A DNA phylogeny confirmed this (Fiaschi & Plunkett 2011). We had the correct S. morototoni in the 1996 and 2004 papers. Schizolobium parahyba Accepted Schizolobium parahyba schipa Fabaceae-caesalpinioideae (Vell.) S.F. Blake This is the correct spelling of Croat's S. parahybum. We had the bad spelling in 1996 paper but corrected in database afterwards. Schizolobium parahybum Obsolete Schizolobium parahyba schipa Fabaceae-caesalpinioideae Croat used this misspelling of S. parahyba. We repeated the error in the 1996 paper. Senna dariensis Accepted Senna dariensis sennda Fabaceae-caesalpinioideae (Britton & Rose) H.S. Irwin & Barneby Formerly Cassia fruticosa, but we had corrected to Senna by the 1996 and 2004 papers. Irwin and Barneby 1982 split Cassia into 3 genera, including Senna. Reviewed in Marazzi, with DNA phylogeny confirming monophyly of Senna. Known as S. dariensis var. gatunensis in Tropicos. Simarouba amara Accepted Simarouba amara simaam Simaroubaceae Aubl. Siparuna cristata Accepted Siparuna cristata sipacr Siparunaceae (Poepp. & Endl.) A. DC. On BCI, this was formerly identified as S. guianensis, following Croat. Renner re-identified the Tropicos on BCI specimens as S. cristata in 1999. See notes with S. guianensis. Siparuna guianensis Obsolete Siparuna cristata sipagu Siparunaceae Aubl. Croat used S. guianensis for the less common BCI species of Siparuna, but those in the BCI plot are now reidentified S. cristata. We used the old name in early 50-ha plot censuses and it appears as S. guianensis in the 1996 paper. Though no longer known on BCI, S. guianensis remains a valid species in Panama, with many Canal Area specimens at Tropicos still called S. guianensis. We currently retain the name in 3 other plots, though those specimens may yet be re-identified. Siparuna pauciflora Accepted Siparuna pauciflora sipapa Siparunaceae (Beurl.) A. DC. Sloanea terniflora Accepted Sloanea terniflora sloate Elaeocarpaceae (Moc. & Sessé ex DC.) Standl. Socratea durissima Obsolete Socratea exorrhiza socrex Arecaceae Croat used this name, a Central American taxon, and we followed in the 1982 census. But Croat acknowledged the possibility it is not distinct from the South American, S. exorrhiza, and we had switched by the late 1980s and used the S. exorrhiza in 1996 and 2004 papers. Socratea exorrhiza Accepted Socratea exorrhiza socrex Arecaceae (Mart.) H.Wendl. We used this in 1996 and 2004 papers, changing from Croat, who called this S. durissima. See notes there. Henderson (1997) lumped S. durissima with the S. American S. exorrhiza. Solanum arboreum Accepted Solanum arboreum solaar Solanaceae Dunal Solanum argenteum Obsolete Solanum lepidotum solast Solanaceae Dunal This is Croat’s name, and we used it in 1982. It is, however, an Incorrect identification, now S. lepidotum. We mistakenly called it S. steyemarkii in 1996 paper, which is wrong taxon and misspelled (should be S. steyermarkii). Solanum asperum Accepted Solanum asperum solaas Solanaceae Rich. Solanum circinatum Accepted Solanum circinatum cyphha Solanaceae Bohs Bohs (1995) collapsed entire genus Cyphomandra into Solanum. DNA phylogeny shows both it and Lycopersicon nested within Solanum. No change in species circumscription. Solanum hayesii Accepted Solanum hayesii solaha Solanaceae Fernald Solanum lepidotum Accepted Solanum lepidotum solast Solanaceae Dunal Nee says Panama and Andes specimens should be this, while E. Brazil specimens are different. So Brazil specimens were first argenteum, changed to lepidotum by Soto, changed back to argenteum by Nee Solanum steyemarkii Obsolete Solanum lepidotum solast Solanaceae This is a misspelling of S. steyermarkii that appeared in our 1996 BCI paper. But the valid name is now S. lepidotum. Solanum steyermarkii Obsolete Solanum lepidotum solast Solanaceae Carvalho Carvalho's describes this species as new and attributes all BCI specimens (including Robin's) to it. But there is much doubt: elevation range is otherwise given as > 500 m, and height range to 35 m. Knapp et al (2005) and Tropicos now reject it. Nee gives this as alt name for FMNH specimens of lepidotum. Sorocea affinis Accepted Sorocea affinis soroaf Moraceae Hemsl. Spachea membranacea Accepted Spachea membranacea spacme Malpighiaceae Cuatrec. Spondias mombin Accepted Spondias mombin sponmo Anacardiaceae L. Spondias radlkoferi Accepted Spondias radlkoferi sponra Anacardiaceae Donn.Sm. Stemmadenia grandiflora Obsolete Tabernaemontana grandiflora stemgr Apocynaceae (Jacq.) Miers Croat's name, widely accepted across S. America, and we used it in 1996 and 2004 papers. Morales moved it, collapsing the entire genus under Tabernaemontana Sterculia apetala Accepted Sterculia apetala sterap Malvaceae (Jacq.) H.Karst. Stylogyne standleyi Obsolete Stylogyne turbacensis stylst Myrsinaceae Croat used this name, a taxon described from BCI. We followed Croat, and this name appears in 1996 and 2004 papers. Ricketson and Pipoly (1997) synonomized under the wide-ranging S. turbacensis, saying that across the range, characters of several local taxa overlap with S. turbacensis. Stylogyne turbacensis Accepted Stylogyne turbacensis stylst Myrsinaceae (Kunth) Mez Formerly S. standleyi, in both 1996 and 2004 paper. We since switched to S. turbacensis, following the monograph of Ricketson and Pipoly (1997). Swartzia simplex var. continentalis Accepted Swartzia simplex var. continentalis swars2 Fabaceae-papilionoideae (Sw.) Spreng. Formerly S. s. var. ochnaceae, and appearing as the latter in 1996 and 2004 papers. Swartzia simplex var. grandiflora Accepted Swartzia simplex var. grandiflora swars1 Fabaceae-papilionoideae (Sw.) Spreng. Formerly S. s. var. ochnaceae, and appearing as the latter in 1996 and 2004 papers. Swartzia simplex var. ochnacea Obsolete Swartzia simplex var. continentalis swars2 Fabaceae-papilionoideae (Sw.) Spreng. Formerly S. s. var. ochnaceae, and appearing as the latter in 1996 and 2004 papers. Symphonia globulifera Accepted Symphonia globulifera sympgl Clusiaceae L.f. Tabebuia guayacan Obsolete Handroanthus guayacan tab1gu Bignoniaceae (Seem.) Hemsl. DNA phylogeny shows the genus Tabebuia to be polyphyletic. and the yellow-fllowered taxa were moved into Handroanthus. Tabebuia rosea Accepted Tabebuia rosea tab1ro Bignoniaceae (Bertol.) DC. Tabernaemontana arborea Accepted Tabernaemontana arborea tab2ar Apocynaceae Rose ex J.D.Sm. Tabernaemontana grandiflora Accepted Tabernaemontana grandiflora stemgr Apocynaceae Jacq. Formerly Stemmadenia grandiflora, appearing as such in 1996 and 2004 papers. We are following Morales and placing it in Tabernaemontana. Tachigalia versicolor Obsolete Tachigali panamensis tachve Fabaceae-caesalpinioideae This is Croat’s misspelling of T. versicolor. We had it corrected by the 1996 paper, but used it in our own database for sometime before that. Tachigali panamensis Accepted Tachigali panamensis tachve Fabaceae-caesalpinioideae van der Werff & N.Zamora Formerly T. versicolor, following Croat, but van der Werff in 2010 erected this new name for Panama specimens. This appears as T. versicolor in 1996 and 2004 papers. Tachigali versicolor Obsolete Tachigali panamensis tachve Fabaceae-caesalpinioideae Standl. & L.O.Williams Croat’s name, and we used through 2015, including 1996 and 2004 papers. We now follow van der Werff et al. (2010) and use the new name T. panamensis. T. versicolor is valid but only in CR. It lacks pinnate stipule. Talisia croatii Accepted Talisia croatii talipr Sapindaceae Acev.-Rodr. Formerly T. princeps, but Acevedo-Rodriguez described in 2003, Fl. Neotrop. Talisia nervosa Accepted Talisia nervosa taline Sapindaceae Radlk. Talisia princeps Obsolete Talisia croatii talipr Sapindaceae Oliv. Croat's original name, and we used in 1996 paper. Currently T. croatii. Terminalia amazonia Accepted Terminalia amazonia termam Combretaceae (J.F.Gmel.) Exell We followed Croat’s error and spelled this incorrectly as T. amazonica through the 1996 paper. It was corrected in 2004 paper. Terminalia amazonica Obsolete Terminalia amazonia termam Combretaceae Croat’s incorrect spelling of T. amazonia; it appears in our 1996 paper but was corrected by the 2004 paper. Terminalia chiriquensis Obsolete Terminalia oblonga termob Combretaceae This was Croat's name, but it was obsolete before the plot started. By 1981, Robin's specimens were identified as T. oblonga by Stace. Terminalia oblonga Accepted Terminalia oblonga termob Combretaceae (Ruiz & Pav.) Steud. Our name since the plot began. Croat used T. chiriquensis, but that was synonomized before 1982. See Fl. Nic. Ternstroemia tepezapote Accepted Ternstroemia tepezapote ternte Pentaphylacaceae Schltdl. & Cham. Tetragastris panamensis Accepted Tetragastris panamensis tet2pa Burseraceae (Engl.) Kuntze Tetrathylacium johansenii Accepted Tetrathylacium johansenii tet4jo Salicaceae Standl. Theobroma cacao Accepted Theobroma cacao theoca Malvaceae L. Thevetia ahouai Accepted Thevetia ahouai thevah Apocynaceae (L.) A.DC. Tocoyena pittieri Accepted Tocoyena pittieri tocopi Rubiaceae (Standl.) Standl. Tovomitopsis nicaraguensis Obsolete Chrysochlamys eclipes chr1ec Clusiaceae Oerst., Planch. & Triana Croat used this name, but it is a double error. First, Tovomitopsis has been synonomized with Chrysoclamys more often than not for over 100 years; Hammel (1999) cleared this up in favor of Chrysoclamys. Second, C. nicaraguensis remains a valid species, but quite distinct, and not on BCI, where C. eclipes is correct. Hammel (1999) provides a clear explanation of all. Trattinnickia aspera Accepted Trattinnickia aspera tratas Burseraceae (Standl.) Swart Trema integerrima Accepted Trema integerrima tremin Cannabaceae (Beurl.) Standl. Croat mentioned this taxon, but did not consider it present on BCI. In 2010, we reassessed the genus and recognized several individuals in the 50-ha plot as belonging to T. integerrima, formerly identified as T. micrantha. See notes with Trema unidentified. Trema micrantha Accepted Trema micrantha tremmi Cannabaceae (L.) Blume All Trema in the 50-ha plot were identified as this species until 2010. But several individuals that had been alive prior to 2010 (one back to 1990) were reidentified, forcing us to reassess the genus. See notes with Trema unidentified. Trema unidentified Accepted Trema unidentified tremsp Cannabaceae Because we reassessed the genus in 2010, any individuals of Trema that had died earlier could not be reidentified, and all are now called Trema unidentified. Trichanthera gigantea Accepted Trichanthera gigantea tri1gi Acanthaceae (Bonpl.) Nees Trichilia cipo Obsolete Trichilia tuberculata tri2tu Meliaceae (A. Juss.) C. DC. Croat's misidentification of T. tuberculata, but Pennington corrected by 1981, also in Pennington (2016). T. cipo is valid but only in S. America. We had changed to T. tuberculata before the plot census began, following Pennington (1981). Tropicos still has Panama specimens, including Croat’s from BCI, under the erroneous T. cipo. Trichilia montana Obsolete Trichilia pallida tri2pa Meliaceae Croat's name, but we had changed to T. pallida before the plot census began, following Pennington in Fl. Neotrop. Trichilia pallida Accepted Trichilia pallida tri2pa Meliaceae Sw. Listed as T. montana in Croat, but we had changed to T. pallida before the plot began (Pennington 1981). Trichilia tuberculata Accepted Trichilia tuberculata tri2tu Meliaceae (Triana & Planch.) C. DC. Listed as T. cipo in Croat, but we had corrected to T. tuberculata before the plot began (Pennington 1981). Trichospermum galeottii Accepted Trichospermum galeottii tri4ga Malvaceae (Turcz.) Kosterm. Croat listed as T. mexicanum, but that name was obsolete by the early 1980s. BCI specimens renamed by Meijer in 1980 (see Fl. Nic.). Trichospermum mexicanum Obsolete Trichospermum galeottii tri4ga Malvaceae (DC.) Baill. Croat's name, but we had changed to T. galeottii by 1982, and both 1996 and 2004 papers list T. galeottii. Triplaris cumingiana Accepted Triplaris cumingiana tripcu Polygonaceae Fisch. & C.A.Mey. Trophis caucana Accepted Trophis caucana olmeas Moraceae (Pittier) C.C. Berg Originally Olmedia aspera, and that older name appeared in 1996 paper. Updated to Trophis caucana, following Berg (1988). See notes under Olmedia. Trophis racemosa Accepted Trophis racemosa tropra Moraceae (L.) Urb. Turpinia occidentalis Accepted Turpinia occidentalis turpoc Tapisciaceae (Sw.) G.Don Unidentified species Accepted Unidentified species uniden Unknown Unonopsis pittieri Accepted Unonopsis pittieri unonpi Annonaceae Saff. Urera baccifera Accepted Urera baccifera urerba Urticaceae (L.) Gaudich. ex Wedd. Not in Croat but identified in 1982 census. Vachellia melanoceras Accepted Vachellia melanoceras acacme Fabaceae (Beurl.) Seigler & Ebinger Formerly Acacia melanoceras through 2015, following Croat’s originally name. A long-running and heated debate finally ended around 2010 with the American Acacia settling in two new genera, Vachellia and Senegalia. Vasconcellea cauliflora Accepted Vasconcellea cauliflora carica Caricaceae (Jacq.) A.DC. This species appears in Croat as Carica cauliflora, but the species was not found in the 50-ha plot until 2010. We identified it in other plots in the 1990s, though, using the old name, but it is now updated to V. califlora in our database. (Badillo 2000 in Ernstia). Verbesina gigantea Accepted Verbesina gigantea verbgi Asteraceae Jacq. Croat included this taxon, but it was not identified in the 50-ha plot until the 2000 census and thus does not appear in 1996 paper. Virola bozo Obsolete Virola multiflora virosp Myristicaceae Our original name in 1982 for a Virola not matching any known BCI species. In the 1996 paper, it appears as Virola sp. nov., as if it was a novel species. But it was later identified as V. multiflora Virola multiflora Accepted Virola multiflora virosp Myristicaceae (Standl.) A.C.Sm. Labeled as unidentified morphospecies V. bozo in original BCI census, and listed as Virola sp. nov. in 1996 paper. By the 2004 paper, it was identified and appears as V. mutliflora there. Virola nobilis Accepted Virola nobilis virosu Myristicaceae A.C. Sm. Croat published V. surinamensis for the tallest BCI Virola, but lists V. nobilis as a synonym, and originally identified specimens as V. nobilis. Later, Gentry incorrectly reidentified as V. surinamensis. Rodriguez in 2007 identified BCI specimens as V. nobilis, but Tropicos has both names in Panama and Costa Rica. Virola sebifera Accepted Virola sebifera virose Myristicaceae Aubl. Virola sp. nov. Obsolete Virola multiflora virosp Myristicaceae This is the name used in the 1996 paper for the Virola morphospecies, given the less stodgy name of Virola bozo in our databases. Later identified as V. multiflora, not a new species at all. Virola surinamensis Obsolete Virola nobilis virosu Myristicaceae (Rol. ex Rottb.) Warb. This was Croat’s name, following an early misidentification of Gentry’s; we published it in the 1996 and 2004 papers (Croat listed V. nobilis as a synonym). As of 2007, however, Rodriguez advised that BCI specimens are V. nobilis. V. surinamensis remains valid in S. America. See more comments under V. nobilis. Vismia baccifera Accepted Vismia baccifera vismba Clusiaceae (L.) Triana & Planch. Vismia billbergiana Accepted Vismia billbergiana vismbi Clusiaceae Beurl. Vismia macrophylla Accepted Vismia macrophylla vismma Clusiaceae Kunth Vochysia ferruginea Accepted Vochysia ferruginea vochfe Vochysiaceae Mart. Xylopia macrantha Accepted Xylopia macrantha xyl1ma Annonaceae Triana & Planch. Xylosma chlorantha Accepted Xylosma chlorantha xyl2ch Salicaceae Donn.Sm. Croat misspelled this X. chloranthum and we repeated the error in the 1996 paper. Xylosma chloranthum Obsolete Xylosma chlorantha xyl2ch Salicaceae Croat's misspelling of X. chorantha. We used it in the 1997 paper. As for X. oligandra, silly Latin gender. Xylosma oligandra Accepted Xylosma oligandra xyl2ol Salicaceae Donn. Sm. Croat misspelled this X. oligandrum and we repeated the error in the 1996 paper. Xylosma oligandrum Obsolete Xylosma oligandra xyl2ol Salicaceae Croat's misspelling of X. oligandrum. We used it in the 1997 paper. As for X. chlorantha, silly Latin gender. Zanthoxylum acuminatum Accepted Zanthoxylum acuminatum zantpr Rutaceae (Sw.) Sw. Originally Z. procerum in Croat, and in our 1996 paper under that name. We then used Z. juniperinum for some time, and that's how it appeared in 2004 paper. Reynel identified all BCI specimens at Tropicos as this in 1993, with subsp. juniperinum. Zanthoxylum belizense Obsolete Zanthoxylum ekmanii zantbe Rutaceae Croat's name, and we used in 1996 paper. Reynel reidentified BCI specimens as Z. ekmanii in 1993. Reynel (1995) PhD dissertation not yet consulted. Also see Manual CR. Zanthoxylum ekmanii Accepted Zanthoxylum ekmanii zantbe Rutaceae (Urb.) Alain Formerly Z. belizense, reidentified by Reynel in 1993 at Tropicos. Zanthoxylum juniperinum Obsolete Zanthoxylum panamense zantpr Rutaceae Poepp. This name is used for Z. procerum/Z. acuminatum subsp. juniperinum by several treatments. See Manual CR. We used it in 2004 paper, but Z. procerum (following Croat) in the 1996 paper. Zanthoxylum panamense Accepted Zanthoxylum panamense zantp1 Rutaceae P.Wilson Zanthoxylum procerum Obsolete Zanthoxylum acuminatum zantpr Rutaceae Name used by Croat, and we used through 1996 paper. Reynel identified Croat's BCI specimen as Z. acuminatum subsp. juniperinum in 1995. Reynel (1995) PhD dissertation not seen yet. Zanthoxylum setulosum Accepted Zanthoxylum setulosum zantse Rutaceae P.Wilson Zuelania guidonia Accepted Zuelania guidonia zuelgu Salicaceae (Sw.) Britton & Millsp.