Species code habit beta alpha Ag bg Am bm lambda Acalypha diversifolia Jacq. acaldi shrub 1.17 ± 0.32 19.69 ± 9.1 1.92 ± 0.7 -0.21 ± 0.36 0.12 ± 0.04 0.02 ± 0.17 -1.80 ± 7.63 Acalypha macrostachya Jacq. acalma underst. 1.41 ± 0.38 36.28 ± 9.16 3.55 ± 4.23 -0.13 ± 0.74 0.11 ± 0.07 0.20 ± 0.5 -1.81 ± 4.41 Adelia triloba (Müll. Arg.) Hemsl. ade1tr underst. 1.14 ± 0.18 47.5 ± 7.86 0.83 ± 0.53 0.28 ± 0.27 0.14 ± 0.05 -0.47 ± 0.15 -3.09 ± 2.15 Aegiphila panamensis Moldenke aegipa midst. 1.53 ± 0.25 83 ± 18.15 0.85 ± 0.45 0.09 ± 0.27 0.2 ± 0.13 -0.44 ± 0.28 -4.21 ± 1.92 Alchornea costaricensis Pax & K. Hoffm. in Engl. alchco canopy 0.84 ± 0.38 102.1 ± 55.9 1.88 ± 1.25 0.24 ± 0.21 0.25 ± 0.04 -0.64 ± 0.07 -0.98 ± 4.36 Alibertia edulis (Rich.) A. Rich. alibed underst. 0.92 ± 0.19 13.92 ± 5.93 0.46 ± 0.15 0.19 ± 0.42 0.03 ± 0.01 0.03 ± 0.2 1.24 ± 1.71 Allophylus psilospermus Radlk. allops underst. 0.64 ± 0.14 19.72 ± 11.64 0.95 ± 0.76 0.41 ± 0.33 0.08 ± 0.03 -0.09 ± 0.3 -1.57 ± 1.99 Alseis blackiana Hemsl. alsebl canopy 0.24 ± 0.08 1.07 ± 1.95 0.32 ± 0.12 0.66 ± 0.05 0.03 ± 0.01 -0.31 ± 0.23 0.23 ± 0.97 Andira inermis (W. Wright) DC. andiin canopy 1.10 ± 0.14 53.71 ± 5.25 0.4 ± 0.12 0.24 ± 0.11 0.03 ± 0.01 0.22 ± 0.18 -0.77 ± 0.55 Annona acuminata Saff. annoac shrub 1.05 ± 0.45 13.48 ± 9.14 0.48 ± 0.15 -0.11 ± 0.54 0.04 ± 0.01 -0.02 ± 0.26 0.0 ± 1.24 Annona spraguei Saff. annosp midst. 0.55 ± 0.20 24.59 ± 28.61 3.65 ± 1.13 -0.13 ± 0.12 0.20 ± 0.10 -0.19 ± 0.2 3.5 ± 6.52 Apeiba membranacea Spruce ex Benth. apeime canopy 1.08 ± 0.19 268.5 ± 40.71 1.96 ± 0.59 0.20 ± 0.09 0.12 ± 0.07 -0.40 ± 0.19 -1.02 ± 2.12 Apeiba tibourbou Aubl. apeiti midst. 1.37 ± 0.44 162.5 ± 43.86 7.56 ± 6.32 -0.49 ± 0.47 0.77 ± 1.24 -0.15 ± 0.8 0.10 ± 4.27 Aspidosperma spruceanum Benth. ex Müll. Arg. aspicr canopy 0.26 ± 0.05 1.10 ± 1.53 0.27 ± 0.05 0.80 ± 0.08 0.02 ± 0.01 0.25 ± 0.23 0.2 ± 0.62 Astronium graveolens Jacq. ast2gr canopy 0.70 ± 0.27 133.9 ± 80.38 0.8 ± 0.59 0.46 ± 0.17 0.09 ± 0.05 0.04 ± 0.34 1.55 ± 2.64 Beilschmiedia pendula (Sw.) Hemsl. beilpe canopy 0.29 ± 0.07 3.27 ± 3.42 0.39 ± 0.14 0.64 ± 0.05 0.03 ± 0.01 -0.06 ± 0.15 -0.37 ± 2.19 Brosimum alicastrum Sw. brosal canopy 0.54 ± 0.06 37.61 ± 9.15 0.40 ± 0.08 0.56 ± 0.13 0.01 ± 0.00 0.33 ± 0.10 0.09 ± 0.61 Calophyllum longifolium Willd. calolo canopy 0.28 ± 0.12 2.89 ± 4.90 0.52 ± 0.14 0.65 ± 0.06 0.02 ± 0.0 0.41 ± 0.09 3.61 ± 1.09 Capparis frondosa Jacq. cappfr shrub 1.80 ± 0.14 27.76 ± 1.38 0.36 ± 0.17 -0.28 ± 0.26 0.05 ± 0.02 -0.66 ± 0.4 -1.24 ± 1.33 Casearia aculeata Jacq. caseac underst. 0.93 ± 0.18 18.04 ± 7.08 0.46 ± 0.18 0.24 ± 0.13 0.05 ± 0.02 -0.36 ± 0.27 -0.12 ± 1.01 Casearia arborea (Rich.) Urb. casear canopy 1.03 ± 0.22 188.1 ± 54.84 1.26 ± 0.52 0.29 ± 0.15 0.11 ± 0.03 -0.20 ± 0.14 -2.77 ± 2.17 Casearia sylvestris Sw. casesy midst. 0.93 ± 0.14 70.24 ± 18.34 0.83 ± 0.34 0.22 ± 0.16 0.09 ± 0.02 -0.20 ± 0.16 -2.31 ± 1.1 Cassipourea elliptica (Sw.) Poir. cassel midst. 0.47 ± 0.05 7.13 ± 3.23 0.48 ± 0.17 0.62 ± 0.10 0.01 ± 0.0 0.40 ± 0.19 1.44 ± 1.11 Cecropia insignis Liebm. cecrin canopy 0.87 ± 0.30 125.4 ± 76.92 15.34 ± 5.4 -0.31 ± 0.18 0.58 ± 0.19 -0.79 ± 0.10 1.55 ± 7.61 Cecropia obtusifolia Bertol. cecrob midst. 1.24 ± 0.31 115.1 ± 36.15 24.2 ± 23.6 -0.96 ± 0.33 0.37 ± 0.26 -0.33 ± 0.30 3.9 ± 11.35 Ceiba pentandra (L.) Gaertn. ceibpe canopy 0.63 ± 0.18 537 ± 218.15 2.58 ± 1.68 0.19 ± 0.24 0.3 ± 0.12 -0.40 ± 0.32 -0.7 ± 3.0 Celtis schippii Standl. celtsc midst. 1.46 ± 0.16 92.38 ± 17.13 0.78 ± 0.22 0.37 ± 0.13 0.06 ± 0.05 0.01 ± 0.41 -1.89 ± 0.53 Chamguava schippii (Standl.) Landrum cha2sc underst. 0.95 ± 0.35 16.31 ± 8.95 0.52 ± 0.12 0.41 ± 0.16 0.01 ± 0.00 0.33 ± 0.41 3.91 ± 1.68 Chrysochlamys eclipes L.O. Williams chr1ec shrub 1.26 ± 0.33 24.62 ± 6.78 0.44 ± 0.21 0.06 ± 0.40 0.03 ± 0.01 -0.32 ± 0.37 -0.47 ± 0.73 Chrysophyllum argenteum Jacq. chr2ar canopy 0.40 ± 0.13 9.96 ± 13.91 0.58 ± 0.11 0.32 ± 0.26 0.02 ± 0.01 0.09 ± 0.14 1.84 ± 2.87 Chrysophyllum cainito L. chr2ca canopy 0.24 ± 0.13 11.82 ± 20.2 0.33 ± 0.14 0.80 ± 0.15 0.04 ± 0.01 -0.12 ± 0.68 3.28 ± 1.83 Coccoloba coronata Jacq. coccco midst. 0.87 ± 0.64 15.79 ± 16.21 0.23 ± 0.13 0.64 ± 0.37 0.09 ± 0.05 -0.14 ± 0.45 -0.84 ± 3.6 Coccoloba manzinellensis Beurl. coccma underst. 1.52 ± 0.21 25.44 ± 3.29 0.63 ± 0.48 -0.11 ± 0.28 0.05 ± 0.03 -0.62 ± 0.46 -0.8 ± 1.23 Cordia alliodora (Ruiz & Pav.) Oken cordal canopy 0.83 ± 0.3 127.2 ± 75.89 2.72 ± 2.05 0.23 ± 0.25 0.13 ± 0.05 -0.22 ± 0.19 1.69 ± 4.51 Cordia bicolor A. DC. cordbi midst. 0.76 ± 0.15 84.33 ± 35.79 2.01 ± 0.91 0.18 ± 0.18 0.16 ± 0.08 -0.65 ± 0.25 -0.09 ± 3.88 Cordia lasiocalyx Pittier cordla midst. 1.30 ± 0.20 69.84 ± 11.65 1.88 ± 0.43 -0.39 ± 0.16 0.04 ± 0.01 -0.05 ± 0.14 -1.38 ± 1.4 Coussarea curvigemmia Dwyer cou2cu underst. 0.98 ± 0.16 26.61 ± 6.38 0.69 ± 0.25 0.16 ± 0.11 0.02 ± 0.01 0.02 ± 0.29 1.15 ± 1.66 Croton billbergianus Müll. Arg. crotbi underst. 0.67 ± 0.2 21.9 ± 16.28 3.34 ± 1.07 0.13 ± 0.19 0.25 ± 0.11 -0.22 ± 0.21 -0.12 ± 8.77 Cupania latifolia Kunth cupala canopy 0.50 ± 0.15 24.82 ± 16.13 0.75 ± 0.17 0.60 ± 0.12 0.07 ± 0.03 0.25 ± 0.40 -0.04 ± 3.06 Cupania rufescens Triana & Planch. cuparu canopy 0.50 ± 0.26 7.67 ± 7.55 0.51 ± 0.17 0.75 ± 0.03 0.09 ± 0.04 -0.47 ± 0.50 2.48 ± 3.98 Cupania seemannii Triana & Planch. cupasy underst. 0.79 ± 0.17 9.55 ± 4.06 0.53 ± 0.14 0.41 ± 0.17 0.04 ± 0.02 -0.58 ± 0.49 1.07 ± 0.88 Dendropanax arboreus (L.) Dec. & Planch dendar canopy 1.40 ± 0.19 240.1 ± 23.2 0.62 ± 0.21 0.43 ± 0.08 0.13 ± 0.11 -0.10 ± 0.42 -2.86 ± 0.61 Desmopsis panamensis (B.L. Rob.) Saff. des2pa underst. 1.14 ± 0.23 19.58 ± 6.91 0.68 ± 0.31 -0.15 ± 0.31 0.02 ± 0.00 0.47 ± 0.18 -0.15 ± 0.68 Diospyros artanthifolia Mart. ex Miq. dio2ar midst. 0.74 ± 0.14 37.1 ± 14.29 0.97 ± 0.33 0.20 ± 0.18 0.04 ± 0.02 0.42 ± 0.38 2.86 ± 1.26 Dipteryx oleifera Benth. diptpa canopy 0.87 ± 0.19 496.7 ± 89.63 0.4 ± 0.11 0.74 ± 0.11 0.17 ± 0.12 -0.01 ± 0.64 -0.4 ± 1.82 Drypetes standleyi G.L. Webster drypst canopy 0.67 ± 0.05 25.74 ± 2.69 0.35 ± 0.15 0.66 ± 0.08 0.02 ± 0.01 -0.23 ± 0.09 -0.06 ± 0.61 Erythrina costaricensis Micheli ery1co underst. 2.23 ± 0.17 84.83 ± 5.86 0.71 ± 0.34 -0.10 ± 0.17 0.13 ± 0.08 -0.31 ± 0.27 -5.19 ± 0.77 Erythroxylum macrophyllum Cav. ery2ma underst. 0.49 ± 0.19 6.86 ± 7.27 0.34 ± 0.09 0.62 ± 0.08 0.05 ± 0.01 0.15 ± 0.12 -0.54 ± 2.06 Eugenia coloradoensis Standl. eugeco midst. 0.55 ± 0.05 15.47 ± 5.06 0.39 ± 0.15 0.64 ± 0.12 0.03 ± 0.01 0.09 ± 0.20 -0.88 ± 1.60 Eugenia galalonensis (Wright) Krug & Urb. eugega underst. 0.46 ± 0.11 3.07 ± 4.18 0.36 ± 0.16 0.57 ± 0.13 0.01 ± 0.00 0.67 ± 0.21 2.69 ± 1.62 Eugenia nesiotica Standl. eugene midst. 0.34 ± 0.08 2.85 ± 3.54 0.26 ± 0.12 0.60 ± 0.09 0.02 ± 0.00 0.26 ± 0.13 -0.08 ± 0.93 Eugenia oerstediana O. Berg. eugeoe midst. 0.56 ± 0.1 13.22 ± 7.08 0.48 ± 0.20 0.66 ± 0.11 0.03 ± 0.01 0.13 ± 0.13 -0.34 ± 1.68 Faramea occidentalis (L.) A. Rich. faraoc underst. 1.17 ± 0.07 41.47 ± 3.47 0.85 ± 0.31 0.13 ± 0.09 0.01 ± 0.00 0.32 ± 0.11 0.33 ± 1.15 Garcinia intermedia (Pittier) Hammel gar2in midst. 0.71 ± 0.06 11.77 ± 2.85 0.44 ± 0.14 0.47 ± 0.06 0.01 ± 0.00 0.35 ± 0.30 1.06 ± 0.93 Garcinia madruno (H. B. K.) Hammel gar2ma midst. 0.77 ± 0.15 16.26 ± 8.17 0.58 ± 0.17 0.30 ± 0.12 0.02 ± 0.03 0.64 ± 0.36 -1.55 ± 3.60 Genipa americana L. geniam midst. 0.71 ± 0.05 61.48 ± 13.59 0.17 ± 0.10 0.97 ± 0.38 0.06 ± 0.03 0.08 ± 0.21 -1.06 ± 0.98 Guapira standleyana Woodson guapst canopy 0.89 ± 0.12 181.2 ± 40.76 0.66 ± 0.48 0.54 ± 0.23 0.12 ± 0.04 -0.35 ± 0.26 -1.49 ± 1.08 Guarea grandifolia DC. guargr canopy 0.37 ± 0.05 10.21 ± 4.18 0.42 ± 0.13 0.66 ± 0.08 0.07 ± 0.04 0.23 ± 0.61 0.71 ± 1.52 Guarea guidonia (L.) Sleumer guargu midst. 0.51 ± 0.09 22.1 ± 12.45 0.60 ± 0.16 0.34 ± 0.07 0.03 ± 0.01 -0.16 ± 0.14 0.21 ± 1.03 Guarea bullata Radlk. guarsp midst. 0.65 ± 0.06 15.7 ± 3.86 0.38 ± 0.14 0.21 ± 0.11 0.05 ± 0.02 -0.07 ± 0.15 -2.62 ± 1.21 Guatteria dumetorum R.E. Fr. guatdu canopy 0.64 ± 0.10 57.26 ± 19.16 0.58 ± 0.18 0.58 ± 0.12 0.05 ± 0.02 -0.07 ± 0.17 -2.06 ± 1.46 Guazuma ulmifolia Lam. guazul canopy 0.78 ± 0.20 130.5 ± 54.28 2.29 ± 1.14 0.37 ± 0.26 0.12 ± 0.05 -0.10 ± 0.23 0.84 ± 2.61 Guettarda foliacea Standl. guetfo underst. 0.91 ± 0.30 43.06 ± 12.04 0.91 ± 0.55 0.002 ± 0.4 0.03 ± 0.02 -0.04 ± 0.19 -1.41 ± 0.96 Gustavia superba (Kunth) O. Berg gustsu midst. 2.44 ± 0.17 193.5 ± 10.65 1.76 ± 0.77 -0.16 ± 0.19 0.47 ± 0.22 -1.37 ± 0.30 -0.91 ± 0.73 Hampea appendiculata (Donn. Sm.) Standl. hampap midst. 0.78 ± 0.51 91.49 ± 85.28 5.27 ± 3.41 0.17 ± 0.22 0.19 ± 0.10 -0.12 ± 0.35 2.18 ± 17.1 Hasseltia floribunda Kunth hassfl midst. 1.11 ± 0.18 81.44 ± 18.33 0.65 ± 0.20 0.39 ± 0.08 0.10 ± 0.03 -0.42 ± 0.07 -3.56 ± 1.0 Heisteria acuminata (Humb. & Bonpl.) Engl. heisac underst. 1.53 ± 0.20 54.5 ± 5.74 0.74 ± 0.23 0.06 ± 0.14 0.05 ± 0.02 -0.11 ± 0.44 -0.49 ± 0.77 Heisteria concinna Standl. heisco midst. 0.91 ± 0.14 71.77 ± 14.47 1.02 ± 0.18 0.15 ± 0.10 0.01 ± 0.00 0.08 ± 0.19 0.16 ± 1.16 Herrania purpurea (Pittier) R.E. Schult. herrpu underst. 2.13 ± 0.15 31.05 ± 1.66 0.94 ± 0.36 -0.48 ± 0.47 0.04 ± 0.01 -0.46 ± 0.31 0.16 ± 0.66 Hirtella triandra Sw. hirttr midst. 0.62 ± 0.08 25.45 ± 9.66 0.85 ± 0.51 0.24 ± 0.15 0.02 ± 0.01 -0.05 ± 0.22 0.38 ± 1.77 Hura crepitans L. huracr canopy 1.16 ± 0.19 704.6 ± 88.05 0.71 ± 0.36 0.56 ± 0.16 0.21 ± 0.16 -0.32 ± 0.15 -1.05 ± 0.65 Hybanthus prunifolius (Humb. & Bonpl. ex Roem. & Schult.) Schulze-Menz hybapr shrub 1.59 ± 0.19 20.63 ± 2.89 0.49 ± 0.18 -0.20 ± 0.18 0.02 ± 0.00 0.68 ± 0.11 -1.16 ± 1.3 Hyeronima alchorneoides Allemão hyeral canopy 0.49 ± 0.12 112.3 ± 60.86 0.45 ± 0.20 0.64 ± 0.17 0.11 ± 0.04 -0.14 ± 0.22 0.49 ± 3.38 Inga cocleensis Pittier ingaco midst. 0.62 ± 0.27 36.31 ± 29.6 0.77 ± 0.62 0.65 ± 0.33 0.08 ± 0.04 0.23 ± 0.25 -3.34 ± 4.01 Inga laurina (Sw.) Willd. ingafa canopy 0.61 ± 0.17 20.86 ± 14.59 0.67 ± 0.35 0.44 ± 0.22 0.06 ± 0.04 0.41 ± 0.85 -0.68 ± 3.08 Inga goldmanii Pittier ingago canopy 0.79 ± 0.04 39.94 ± 3.81 0.67 ± 0.25 0.50 ± 0.15 0.03 ± 0.01 0.27 ± 0.19 -1.83 ± 1.08 Inga marginata Willd. ingama canopy 0.27 ± 0.09 2.70 ± 4.30 1.54 ± 0.46 0.58 ± 0.11 0.10 ± 0.03 -0.10 ± 0.15 -0.73 ± 7.2 Inga pezizifera Benth. ingape canopy 0.76 ± 0.15 35.04 ± 11.44 0.87 ± 0.24 0.54 ± 0.14 0.05 ± 0.02 0.25 ± 0.25 -1.16 ± 2.77 Inga nobilis Willd. ingaqu underst. 1.33 ± 0.10 50.99 ± 8.49 0.85 ± 0.58 0.26 ± 0.33 0.03 ± 0.01 -0.06 ± 0.25 -1.12 ± 0.89 Inga acuminata Benth. ingas1 underst. 0.57 ± 0.11 11.13 ± 5.4 1.44 ± 0.55 0.32 ± 0.22 0.01 ± 0.01 0.46 ± 0.28 3.79 ± 2.07 Inga sapindoides Willd. ingasa midst. 1.16 ± 0.07 64.58 ± 8.53 0.88 ± 0.22 0.39 ± 0.08 0.07 ± 0.02 -0.28 ± 0.17 -2.49 ± 1.14 Inga thibaudiana DC. ingath midst. 0.99 ± 0.30 39.95 ± 18.61 3.02 ± 1.60 0.19 ± 0.2 0.13 ± 0.04 -0.53 ± 0.36 5.18 ± 2.49 Inga umbellifera (Vahl) Steud. ingaum midst. 1.41 ± 0.09 36.55 ± 2.08 0.68 ± 0.32 0.20 ± 0.25 0.03 ± 0.02 0.29 ± 0.39 -0.65 ± 1.65 Jacaranda copaia (Aubl.) D. Don jac1co canopy 1.28 ± 0.30 344.8 ± 69.48 2.5 ± 0.39 0.30 ± 0.11 0.27 ± 0.15 -0.61 ± 0.22 -0.30 ± 2.03 Lacistema aggregatum (P.J. Bergius) Rusby laciag underst. 0.59 ± 0.09 6.91 ± 3.75 0.43 ± 0.17 0.38 ± 0.22 0.03 ± 0.01 0.15 ± 0.14 -0.58 ± 1.79 Lacmellea panamensis (Woodson) Markgr. lacmpa midst. 1.43 ± 0.10 129.2 ± 17.47 1.13 ± 0.47 0.36 ± 0.16 0.13 ± 0.09 -0.18 ± 0.89 0.34 ± 0.81 Laetia thamnia L. laetth underst. 0.97 ± 0.17 28.6 ± 5.86 0.66 ± 0.23 0.37 ± 0.07 0.04 ± 0.01 0.003 ± 0.1 -0.73 ± 1.51 Licania hypoleuca Benth. licahy canopy 0.46 ± 0.12 11.04 ± 9.82 0.39 ± 0.20 0.33 ± 0.28 0.03 ± 0.01 0.45 ± 0.22 0.54 ± 1.34 Licania platypus (Hemsl.) Fritsch licapl canopy 0.39 ± 0.10 2.54 ± 2.43 0.33 ± 0.21 0.56 ± 0.22 0.02 ± 0.01 0.35 ± 0.53 0.11 ± 2.15 Lonchocarpus heptaphyllus (Poir.) DC. loncla canopy 0.52 ± 0.06 23.18 ± 10.99 0.62 ± 0.15 0.36 ± 0.11 0.03 ± 0.01 0.01 ± 0.17 -0.8 ± 1.45 Luehea seemannii Triana & Planch. luehse canopy 0.50 ± 0.13 81.09 ± 61.07 1.00 ± 0.52 0.44 ± 0.20 0.19 ± 0.05 -0.47 ± 0.10 0.42 ± 2.24 Macrocnemum roseum (Ruiz & Pav.) Wedd. macrgl midst. 0.61 ± 0.16 37.53 ± 22.89 0.36 ± 0.18 0.58 ± 0.15 0.06 ± 0.04 0.16 ± 0.47 -0.47 ± 1.55 Mosannona garwoodii Chatrou & Welzenis malmsp midst. 0.94 ± 0.24 27.95 ± 12.7 0.71 ± 0.23 -0.08 ± 0.19 0.01 ± 0.01 0.28 ± 0.29 2.25 ± 1.23 Maquira guianensis Aubl. maquco midst. 1.14 ± 0.18 46.89 ± 6.24 1.07 ± 0.36 -0.08 ± 0.15 0.01 ± 0.00 0.48 ± 0.24 -0.23 ± 0.77 Maytenus schippii Lundell maytsc midst. 0.74 ± 0.12 55.48 ± 19.04 0.36 ± 0.26 0.66 ± 0.26 0.06 ± 0.04 0.30 ± 0.67 -0.46 ± 1.15 Miconia affinis DC. micoaf underst. 1.23 ± 0.29 27.63 ± 6.84 1.81 ± 0.59 0.01 ± 0.29 0.04 ± 0.02 0.48 ± 0.29 0.69 ± 1.85 Miconia argentea (Sw.) DC. micoar midst. 0.32 ± 0.11 2.48 ± 4.08 2.06 ± 0.53 0.27 ± 0.09 0.22 ± 0.10 -0.37 ± 0.17 1.21 ± 5.56 Miconia hondurensis Donn. Sm. micoho underst. 0.69 ± 0.14 29.65 ± 19.16 1.70 ± 0.80 0.09 ± 0.24 0.07 ± 0.07 0.74 ± 0.83 2.74 ± 2.46 Mouriri myrtilloides (Sw.) Poir. mourmy canopy 1.44 ± 0.35 14.53 ± 3.14 0.34 ± 0.08 0.27 ± 0.24 0.03 ± 0.01 0.15 ± 0.31 0.14 ± 1.81 Myrcia gatunensis Standl. myrcga underst. 0.44 ± 0.17 4.45 ± 4.99 0.76 ± 0.44 0.37 ± 0.24 -0.03 ± 0.25 -0.50 ± 1.31 1.37 ± 3.39 Nectandra cissiflora Nees nectci canopy 0.94 ± 0.42 17.79 ± 13.53 0.97 ± 0.29 0.53 ± 0.15 0.11 ± 0.04 -0.40 ± 0.28 -2.32 ± 2.05 Nectandra lineata (Kunth) Rohwer nectgl midst. 0.28 ± 0.17 3.79 ± 9.00 0.78 ± 0.20 0.61 ± 0.13 0.10 ± 0.03 0.10 ± 0.17 2.06 ± 5.44 Nectandra purpurea (Ruiz & Pav.) Mez nectpu midst. 1.24 ± 0.50 20.82 ± 11.68 1.13 ± 0.66 0.40 ± 0.30 0.05 ± 0.02 0.21 ± 0.32 -0.37 ± 1.98 Ocotea cernua (Nees) Mez ocotce midst. 0.60 ± 0.21 16.69 ± 12.09 0.67 ± 0.28 0.56 ± 0.17 0.09 ± 0.04 -0.27 ± 0.14 -0.73 ± 3.34 Ocotea oblonga (Meisn.) Mez ocotob canopy 0.33 ± 0.15 11.46 ± 17.59 1.8 ± 0.81 0.47 ± 0.21 0.14 ± 0.05 -0.19 ± 0.23 0.06 ± 4.50 Ocotea puberula (Rich.) Nees ocotpu canopy 0.49 ± 0.21 19.81 ± 15.75 0.83 ± 0.26 0.73 ± 0.15 0.11 ± 0.03 -0.15 ± 0.21 -1.94 ± 3.55 Ocotea whitei Woodson ocotwh canopy 0.57 ± 0.21 43.02 ± 40.55 0.88 ± 0.13 0.57 ± 0.10 0.09 ± 0.02 -0.28 ± 0.13 -3.73 ± 2.53 Trophis caucana (Pittier) C.C. Berg olmeas underst. 1.52 ± 0.51 52.2 ± 16.9 1.57 ± 0.75 -0.06 ± 0.20 0.13 ± 0.03 -0.58 ± 0.10 -4.1 ± 1.45 Ormosia coccinea (Aubl.) Jacks. ormocr canopy 0.36 ± 0.10 4.84 ± 6.20 0.52 ± 0.24 0.38 ± 0.19 0.04 ± 0.03 0.47 ± 0.42 2.44 ± 1.38 Ormosia macrocalyx Ducke ormoma canopy 0.20 ± 0.05 0.05 ± 0.10 0.43 ± 0.23 0.47 ± 0.12 0.05 ± 0.05 -0.27 ± 1.42 1.48 ± 2.60 Ouratea lucens (Kunth) Engl. ouralu shrub 1.18 ± 0.25 15.0 ± 4.76 0.26 ± 0.10 0.16 ± 0.38 0.02 ± 0.01 -0.05 ± 0.42 0.38 ± 1.36 Palicourea guianensis Aubl. paligu shrub 1.21 ± 0.42 15.41 ± 6.96 4.91 ± 2.31 -0.56 ± 0.35 0.15 ± 0.06 0.40 ± 0.21 4.66 ± 10.4 Pentagonia macrophylla Benth. pentma underst. 1.40 ± 0.14 25.78 ± 3.15 0.48 ± 0.26 -0.12 ± 0.53 0.07 ± 0.02 -0.36 ± 0.37 -2.23 ± 1.84 Perebea xanthochyma H. Karst. perexa midst. 1.00 ± 0.22 35.52 ± 13.52 0.75 ± 0.15 0.08 ± 0.13 0.04 ± 0.03 -0.07 ± 0.52 -0.57 ± 0.65 Cinnamomum triplinerve (Kunth) Koesterm. phoeci canopy 0.50 ± 0.17 9.92 ± 7.52 0.86 ± 0.15 0.62 ± 0.09 0.16 ± 0.06 -0.36 ± 0.34 -0.70 ± 3.20 Picramnia latifolia Tul. picrla underst. 0.69 ± 0.26 13.99 ± 11.02 0.40 ± 0.16 0.45 ± 0.16 0.03 ± 0.01 0.05 ± 0.18 -0.27 ± 0.73 Piper reticulatum L. pipere underst. 1.45 ± 0.46 34.6 ± 11.42 2.19 ± 1.36 -0.47 ± 0.43 0.14 ± 0.09 -0.61 ± 0.61 -1.06 ± 1.66 Platymiscium pinnatum (Jacq.) Dugand pla1pi canopy 0.37 ± 0.20 45.61 ± 54.66 0.34 ± 0.12 0.56 ± 0.12 0.1 ± 0.02 -0.06 ± 0.13 -0.55 ± 2.01 Platypodium elegans Vogel pla2el canopy 0.40 ± 0.07 45.56 ± 33.55 0.63 ± 0.30 0.45 ± 0.21 0.09 ± 0.03 -0.09 ± 0.07 -1.73 ± 1.47 Posoqueria latifolia (Rudg.) Roem. & Schult. posola shrub 0.59 ± 0.10 30.07 ± 13.59 0.35 ± 0.23 0.48 ± 0.22 0.05 ± 0.02 0.26 ± 0.74 -0.04 ± 1.18 Poulsenia armata (Miq.) Standl. poular canopy 1.28 ± 0.20 121.5 ± 29.64 1.10 ± 0.43 0.40 ± 0.09 0.07 ± 0.01 -0.09 ± 0.10 -4.47 ± 1.38 Pourouma bicolor Mart. pourbi canopy 0.47 ± 0.23 33.72 ± 27.29 1.33 ± 0.57 0.47 ± 0.22 0.06 ± 0.03 0.06 ± 0.65 4.12 ± 3.86 Pouteria reticulata (Engl.) Eyma poutre canopy 0.25 ± 0.05 0.81 ± 1.06 0.33 ± 0.06 0.81 ± 0.07 0.04 ± 0.02 -0.19 ± 0.27 -1.35 ± 1.79 Pouteria stipitata Cronquist poutst midst. 0.89 ± 0.23 107.6 ± 26.39 0.42 ± 0.24 0.54 ± 0.21 0.10 ± 0.07 -0.14 ± 0.16 -0.26 ± 1.33 Prioria copaifera Griseb. pri2co canopy 0.40 ± 0.02 21.85 ± 7.22 0.36 ± 0.16 0.77 ± 0.09 0.01 ± 0.01 0.04 ± 0.28 0.02 ± 0.71 Protium costaricense (Rose) Engl. protco midst. 0.93 ± 0.07 42.8 ± 4.64 0.93 ± 0.24 0.20 ± 0.07 0.04 ± 0.01 -0.04 ± 0.27 -1.03 ± 0.9 Protium panamense (Rose) I.M. Johnst. protpa midst. 1.01 ± 0.06 23.58 ± 1.61 0.49 ± 0.12 0.33 ± 0.23 0.02 ± 0.01 0.44 ± 0.23 0.41 ± 0.97 Protium tenuifolium Engl. protte midst. 0.44 ± 0.03 9.26 ± 3.36 0.70 ± 0.23 0.39 ± 0.09 0.02 ± 0.01 -0.06 ± 0.18 0.46 ± 1.17 Psidium friedrichsthalianum (O. Berg) Nied. psidfr underst. 0.29 ± 0.08 1.25 ± 2.18 0.32 ± 0.13 0.35 ± 0.15 0.05 ± 0.03 0.27 ± 0.53 1.33 ± 1.17 Psychotria grandis Sw. psycg3 underst. 0.76 ± 0.22 11.64 ± 6.45 0.99 ± 0.49 0.04 ± 0.44 0.12 ± 0.05 -0.14 ± 0.50 -2.08 ± 5.28 Pterocarpus rohrii Vahl pterro canopy 0.59 ± 0.18 11.38 ± 7.26 0.44 ± 0.08 0.45 ± 0.09 0.03 ± 0.01 0.20 ± 0.30 -0.8 ± 1.42 Quararibea asterolepis Pittier quaras canopy 0.52 ± 0.04 60.05 ± 7.38 0.46 ± 0.09 0.61 ± 0.13 0.02 ± 0.00 -0.10 ± 0.09 -0.34 ± 0.39 Quassia amara L. quasam underst. 1.66 ± 0.12 56.6 ± 2.10 0.7 ± 0.23 -0.32 ± 0.23 0.05 ± 0.03 -0.29 ± 0.28 -1.09 ± 0.65 Randia armata (Sw.) DC. randar underst. 1.00 ± 0.12 56.12 ± 8.57 0.67 ± 0.40 0.18 ± 0.18 0.02 ± 0.01 -0.08 ± 0.16 -0.58 ± 0.89 Rinorea sylvatica (Seem.) Kuntze rinosy shrub 1.42 ± 0.22 17.47 ± 2.41 0.33 ± 0.11 0.18 ± 0.23 0.02 ± 0.01 0.18 ± 0.33 -0.41 ± 0.69 Sapium glandulosum (L.) Morong sapiau canopy 0.57 ± 0.13 91.97 ± 55.01 1.25 ± 0.70 0.23 ± 0.22 0.18 ± 0.13 -0.11 ± 0.30 -0.85 ± 4.30 Senna dariensis (Britton & Rose) H. Irwin & Barneby sennda shrub 1.52 ± 0.39 25.52 ± 7.92 3.06 ± 2.16 -0.58 ± 0.7 0.23 ± 0.05 -0.48 ± 0.24 -1.47 ± 9.53 Simarouba amara Aubl. simaam canopy 0.45 ± 0.16 19.45 ± 18.4 1.11 ± 0.27 0.54 ± 0.08 0.08 ± 0.02 -0.19 ± 0.06 0.81 ± 1.56 Siparuna pauciflora (Beurl.) A. DC. sipapa underst. 0.72 ± 0.51 18.57 ± 19.31 1.32 ± 0.61 -0.28 ± 0.32 0.12 ± 0.04 -0.53 ± 0.22 -0.91 ± 3.91 Sloanea terniflora (Sessé & Moc. ex DC.) Standl. sloate canopy 0.5 ± 0.04 19.74 ± 4.97 0.20 ± 0.11 0.64 ± 0.13 0.02 ± 0.01 0.21 ± 0.22 -1.10 ± 0.55 Solanum hayesii Fernald solaha midst. 0.91 ± 0.31 48.67 ± 26.79 7.58 ± 10.3 -0.11 ± 0.37 0.23 ± 0.10 -0.09 ± 0.20 -2.83 ± 7.75 Sorocea affinis Hemsl. soroaf shrub 1.53 ± 0.12 35.57 ± 2.45 0.77 ± 0.30 -0.20 ± 0.15 0.02 ± 0.01 0.24 ± 0.22 -1.18 ± 1.10 Spondias mombin L. sponmo canopy 0.53 ± 0.11 75.15 ± 47.5 2.07 ± 1.13 0.36 ± 0.16 0.17 ± 0.06 -0.23 ± 0.23 2.71 ± 3.98 Spondias radlkoferi Donn. Sm. sponra canopy 0.42 ± 0.12 27.35 ± 25.83 0.83 ± 0.31 0.45 ± 0.20 0.13 ± 0.06 -0.25 ± 0.21 0.74 ± 2.95 Sterculia apetala (Jacq.) H. Karst. sterap canopy 0.65 ± 0.10 181.7 ± 57.4 0.63 ± 0.20 0.52 ± 0.09 0.15 ± 0.09 -0.12 ± 0.56 -1.24 ± 3.88 Stylogyne turbacensis (Kunth) Mez stylst shrub 0.99 ± 0.32 12.3 ± 8.06 0.40 ± 0.11 0.03 ± 0.33 0.05 ± 0.01 -0.33 ± 0.35 -0.01 ± 0.86 Swartzia simplex (Sw.) Spreng. swars1 underst. 0.73 ± 0.05 18.96 ± 3.96 0.29 ± 0.09 0.45 ± 0.03 0.01 ± 0.00 -0.12 ± 0.18 0.60 ± 0.56 Symphonia globulifera L. f. sympgl canopy 0.41 ± 0.11 15.99 ± 22.28 0.54 ± 0.16 0.55 ± 0.15 0.03 ± 0.01 0.42 ± 0.13 -0.74 ± 0.68 Tabebuia guayacan (Seem.) Hemsl. tab1gu canopy 0.45 ± 0.04 66.16 ± 19.64 0.42 ± 0.25 0.58 ± 0.06 0.08 ± 0.04 0.41 ± 1.10 -0.63 ± 1.20 Tabebuia rosea (Bertol.) A. DC. tab1ro canopy 0.47 ± 0.07 33.43 ± 19.12 0.41 ± 0.13 0.56 ± 0.12 0.05 ± 0.01 0.05 ± 0.17 -1.08 ± 1.60 Tabernaemontana arborea Rose in Don. Sm. tab2ar canopy 0.40 ± 0.03 14.89 ± 6.84 0.69 ± 0.16 0.59 ± 0.13 0.02 ± 0.01 -0.07 ± 0.16 1.03 ± 0.55 Tachigali versicolor Standl. & L.O. Williams tachve canopy 0.40 ± 0.05 1.94 ± 0.88 0.41 ± 0.13 0.67 ± 0.19 0.04 ± 0.02 0.13 ± 0.24 -1.10 ± 1.59 Talisia nervosa Radlk. taline underst. 2.67 ± 0.30 35.23 ± 2.55 0.64 ± 0.19 -0.56 ± 0.40 0.02 ± 0.01 -0.32 ± 0.57 -0.46 ± 0.34 Talisia croatii Acev.-Rodr. talipr underst. 0.90 ± 0.10 16.55 ± 5.40 0.42 ± 0.16 -0.26 ± 0.43 0.03 ± 0.01 -0.19 ± 0.19 0.50 ± 0.92 Terminalia amazonia (J.F. Gmel.) Exell termam canopy 0.97 ± 0.15 200.3 ± 36.08 0.66 ± 0.19 0.57 ± 0.13 0.59 ± 1.08 -0.27 ± 0.41 -0.95 ± 1.21 Terminalia oblonga (Ruiz & Pav.) Steud. termob canopy 0.69 ± 0.06 146.2 ± 20.52 0.70 ± 0.21 0.46 ± 0.10 0.09 ± 0.04 -0.20 ± 0.64 -0.16 ± 1.32 Tetragastris panamensis (Engl.) Kuntze tet2pa canopy 0.21 ± 0.07 0.45 ± 0.80 0.43 ± 0.16 0.58 ± 0.04 0.02 ± 0.01 -0.11 ± 0.16 1.34 ± 1.38 Thevetia ahouai (L.) A. DC. thevah shrub 1.40 ± 0.33 38.63 ± 11.3 1.06 ± 0.62 -0.06 ± 0.45 0.11 ± 0.04 -0.41 ± 0.31 -0.96 ± 2.77 Trattinnickia aspera (Standl.) Swart tratas canopy 1.00 ± 0.13 186.5 ± 30.14 3.0 ± 2.57 0.18 ± 0.23 0.08 ± 0.08 0.06 ± 0.20 -1.48 ± 3.14 Trichilia pallida Sw. tri2pa midst. 0.76 ± 0.09 31.86 ± 8.22 0.73 ± 0.15 0.34 ± 0.11 0.04 ± 0.01 -0.14 ± 0.24 -0.54 ± 1.03 Trichilia tuberculata (Triana & Planch.) C. DC. tri2tu canopy 0.41 ± 0.10 10.87 ± 13.22 0.32 ± 0.07 0.63 ± 0.05 0.02 ± 0.00 0.13 ± 0.11 -0.57 ± 0.75 Triplaris cumingiana Fisch. & C.A. Mey. ex C.A. Mey. tripcu midst. 1.37 ± 0.18 130.5 ± 20.85 1.10 ± 0.51 0.23 ± 0.13 0.14 ± 0.07 -0.63 ± 0.34 -1.75 ± 0.60 Trophis racemosa (L.) Urb. tropra midst. 0.60 ± 0.07 20.42 ± 10.82 0.41 ± 0.10 0.17 ± 0.06 0.03 ± 0.01 0.30 ± 0.20 -1.04 ± 1.32 Turpinia occidentalis (Sw.) G. Don turpoc canopy 1.30 ± 0.50 139.3 ± 51.3 2.31 ± 1.34 0.24 ± 0.14 0.08 ± 0.06 0.06 ± 0.31 -2.24 ± 4.85 Unonopsis pittieri Saff. unonpi midst. 0.90 ± 0.09 56.63 ± 10.08 1.42 ± 1.60 0.19 ± 0.43 0.05 ± 0.02 -0.31 ± 0.28 -0.60 ± 1.21 Virola sebifera Aubl. virose midst. 0.93 ± 0.18 82.46 ± 26.69 0.69 ± 0.23 0.43 ± 0.06 0.03 ± 0.01 -0.13 ± 0.17 -2.21 ± 0.58 Virola multiflora (Standl.) A.C. Sm. virosp canopy 0.92 ± 0.21 147.4 ± 49.12 0.88 ± 0.41 0.51 ± 0.17 0.09 ± 0.06 0.13 ± 0.39 -1.29 ± 0.81 Virola surinamensis (Rol. ex Rottb.) Warb. virosu canopy 1.19 ± 0.23 307.2 ± 59.16 0.65 ± 0.17 0.63 ± 0.06 0.08 ± 0.03 -0.23 ± 0.10 -2.05 ± 1.23 Vismia baccifera (L.) Tr. & Pl. vismba underst. 1.22 ± 0.19 33.98 ± 10.1 1.60 ± 1.09 -0.15 ± 0.2 0.09 ± 0.04 0.09 ± 0.23 -0.93 ± 4.51 Xylopia macrantha Triana & Planch. xyl1ma midst. 0.76 ± 0.09 31.26 ± 7.48 1.29 ± 0.44 0.13 ± 0.04 0.01 ± 0.00 0.10 ± 0.21 2.50 ± 0.76 Zanthoxylum ekmanii (Urb.) Alain zantbe canopy 0.90 ± 0.17 193.9 ± 50.21 12.6 ± 11.2 0.05 ± 0.32 0.34 ± 0.06 -0.55 ± 0.09 0.83 ± 3.98 Zanthoxylum panamense P. Wilson zantp1 canopy 0.65 ± 0.19 58.86 ± 39.1 1.14 ± 0.70 0.50 ± 0.18 0.10 ± 0.04 -0.17 ± 0.14 -1.15 ± 5.21 Zanthoxylum acuminatum (Sw.) Sw. zantpr canopy 0.87 ± 0.16 58.85 ± 20.48 1.20 ± 1.06 0.49 ± 0.30 0.06 ± 0.03 0.08 ± 0.20 -2.72 ± 2.40 Zuelania guidonia (Sw.) Britton & Mills. zuelgu midst. 0.50 ± 0.21 26.83 ± 30.39 0.41 ± 0.20 0.49 ± 0.12 0.09 ± 0.07 0.45 ± 1.00 -1.03 ± 2.04