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In these series, the Institution publishes small papers and full-scale monographs that report the research and collections of its various museums and bureaux or of professional colleagues in the world of science and scholarship. The publications are distributed by mailing lists to libraries, universities, and similar institutions throughout the world.

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Robert McC. Adams

Secretary

Smithsonian Institution

Corrections to Smithsonian Contributions to the Marine Sciences Number 27:

Page 69. Polysiphonia beaudettei. First entry should be spelled the same as the centered heading.

Page 142. Delete van den Hoek from the authorship of van den Hoek, Vannajan, and Trono, then transfer the two references to page 153, preceding Velasquez.

Page 157. Change explanation of Index to read: (Primary entries in roman, synonyms and names mentioned incidentally in italic.)

SMITHSONIAN CONTRIBUTIONS TO THE MARINE SCIENCES • NUMBER 27

Catalog of the Benthic Marine Algae
of the Philippines

Paul C. Silva,
Ernani G. Meñez,
and Richard L. Moe

SMITHSONIAN INSTITUTION PRESS
Washington, D.C.
1987

ABSTRACT

Silva, Paul C., Ernani G. Meñez, and Richard L. Moe. Catalog of the Benthic Marine Algae of the Philippines. Smithsonian Contributions to the Marine Sciences, number 27, 179 pages, 2 figures, 1987.—All published records of benthic marine algae from the Philippines are assembled in a catalog with the taxa arranged according to an assumed phylogenetic scheme to the rank of family. The taxonomic framework takes into consideration recently published opinions. Each taxonomic synonym is accompanied by a citation of the author who first proposed the synonymy. Additional taxonomic and nomenclatural notes are provided where deemed useful. Type localities are indicated for all accepted names and taxonomic synonyms.

Cyanophyceae (blue-green algae) comprise 19 genera with 61 species. They are arranged according to the Geitlerian system, accompanied by a reconciliation with the Drouetian system. Their nomenclature is based on a 1753 Linnaean starting point rather than the later starting points specified by the International Code of Botanical Nomenclature.

Rhodophyceae (red algae) comprise 130 genera with 506 specific or infraspecific taxa, of which 35 have Philippine type localities.

Phaeophyceae (brown algae) comprise 23 genera with 154 species or infraspecific taxa, of which 27 have Philippine type localities.

Chlorophyceae (green algae) comprise 37 genera with 251 species or infraspecific taxa, of which 20 have Philippine type localities.

Portieria Zanardini 1851 is adopted in place of Chondrococcus Kützing 1847, Eupogodon Kützing 1845 in place of Dasyopsis (Montagne) Montagne 1847, and Hincksia J.E. Gray 1864 in place of Giffordia Batters 1893. In addition, new binomials are proposed in Gelidium, Halymenia, Callophyllis, Sporolithon, Gracilaria, Ceramium, Polysiphonia, and Hormophysa.

The catalog is preceded by a brief history of Philippine phycology.

OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is recorded in the Institution’s annual report, Smithsonian Year. SERIES COVER DESIGN: Seascape along the Atlantic coast of eastern North America.

Library of Congress Cataloging in Publication Data

Silva, Paul C.

Catalog of the benthic marine algae of the Philippines.

(Smithsonian contributions to the marine sciences ; no. 27)

Bibliography: p.

Includes index.

Supt. of Docs, no.: SII.41:27

1. Marine algae—Philippines—Bibliography. 2. Benthos—Philippines—Bibliography. I. Meñez, Ernani G. II. Moe, Richard Lee. III. Title. IV. Series.   Z5356.A6S54   1987   016.589392′571   87-600124   [QK575.P6]

Contents

Introduction

Acknowledgments

History of Philippine Phycology

Scope of Catalog

Format of Catalog

Catalog

Cyanophyceae

Reconciliation with the Drouetian System

Rhodophyceae

Phaeophyceae

Chlorophyceae

Records Determined Only to Genus

Nomenclatural Notes (by Paul C. Silva)

Literature Cited

Index

We dedicate this work to Dr. Gregorio T. Velasquez, Professor Emeritus, University of the Philippines, devoted champion of phycology and phycologists in the Philippines.

Catalog of the Benthic Marine Algae
of the Philippines

Paul C. Silva, Ernani G. Meñez,
and Richard L. Moe

Paul C. Silva, Herbarium, Department of Botany, University of California, Berkeley, California 94720. Ernani G. Meñez, Smithsonian Oceanographic Sorting Center, Smithsonian Institution, Washington, D.C. 20560. Richard L. Moe, Herbarium, Department of Botany, University of California, Berkeley, California 94720.

Introduction

During May–June 1978 and April–May 1979, Ernani G. Meñez participated in the Smithsonian Institution Philippines Expedition (SIPHILEXP), which was officially based at the Silliman University Marine Laboratory in Dumaguete City, Negros Island. The primary objective was to collect marine organisms from the Central Visayas for studies on taxonomy and distribution. A large collection of benthic marine algae was obtained and subsequently studied.

In preparation for SIPHILEXP, Meñez initiated a compilation of the species of benthic marine algae reported from the Philippines and a list of references pertaining to them. Because of the existence of unique phycological resources in the Herbarium of the University of California at Berkeley, specifically the Index Nominum Algarum and the Bibliographia Phycologica Universalis (both in the form of card files compiled and maintained by Paul C. Silva), he took his preliminary draft there for checking. The utility of collaboration was perceived and an arrangement was made whereby Meñez would be responsible for the citations of species while Silva would be responsible for providing the taxonomic framework and correct nomenclature. In the five years that have passed since the cooperative effort was begun, the number of publications dealing with Philippine seaweeds has increased markedly, and both Meñez and Silva have been involved in keeping pace with the output. In the summer of 1984, Richard L. Moe was asked to join the project, in view of his competency in word-processing and broad phycological experience.

The purpose of this catalog is to integrate published information on the taxonomy and distribution of Philippine benthic marine algae and thus provide a foundation for future exploration and research. Rather than make an uncritical compilation of records (such as Velasquez, Trono, and Doty, 1975), we decided to incorporate as much information as was pertinent to understanding the taxonomic structure and nomenclature adopted in this catalog. The continued relevance of this information obviously depends upon the reliability of the determinations that underlie the published records. Considering that very little critical work has been done on Philippine marine algae, it is reasonable to assume that a certain portion of the incorporated information will eventually prove irrelevant. Nonetheless, we believe that this catalog will facilitate and perhaps stimulate future work by pointing out geographic and taxonomic areas most in need of attention. Unlike publications that are intended to endure like monoliths, this catalog is intended to become obsolete, with the rate of obsolescence an index of the progress in Philippine phycology. Because tens of thousands of data are incorporated in this catalog, the possibility of making errors is great, but we hope that we have initiated far fewer than we have encountered in the literature and corrected.

ACKNOWLEDGMENTS.—For assistance in obtaining copies of pertinent literature, we are especially grateful to Ms. Hilconida Calumpong, Silliman University Marine Laboratory, Dumaguete City, Dr. Paciente A. Cordero, Junior, Division of Botany, Philippine National Museum, Manila, Mrs. Edna T. Fortes-Ganzon, Marine Science Institute, University of the Philippines, Quezon City, Mr. Lawrence M. Liao, Department of Biology, University of San Carlos, Cebu City, and Dr. Milagrosa R. Martinez, University of the Philippines at Los Baños. We are indebted to Dr. Thomas O. Duncan, Director of the Herbarium of the University of California at Berkeley, for his efforts in obtaining a computer system for the Herbarium and for encouraging us to use it in preparing the electronic manuscript of this catalog.

History of Philippine Phycology

Detailed accounts of the history of Philippine phycology have been provided by Velasquez (1962b; 1985) and Cordero (1972; 1977a), but it seems useful to review its most salient features, especially with regard to marine algae. Unlike many areas that were explored biologically by colonial powers in the late eighteenth and early nineteenth centuries, the Philippines were not so favored by Spain, and the earliest information on its plants and their uses came from resident Augustinian monks (M. Blanco, 1837). Unfortunately, there are no extant specimens to document this work. In October 1836, Hugh Cuming, an English naturalist especially interested in conchology, arrived at Manila to begin three years of exploration that would take him to nearly every large island in the archipelago (Merrill, 1926). He made a small but important collection of marine algae, which was reported on by Montagne (1844a) and also served as the basis of Galaxaura fastigiata and Liagora caenomyce, both described by Decaisne (1842).

During the last half of the nineteenth century, four expeditions returned to Europe with varying amounts of Philippine algae. The Preussische Expedition nach Ost-Asien contributed collections made at Manila and Zamboanga in 1861 by its zoologist, Eduard von Martens; these collections were the subject of a publication by Eduard’s father, Georg von Martens (1868). In 1874–1875, H.N. Moseley, the naturalist of the Challenger Expedition, collected algae at the Gigantes Islands (Iloilo), Mactan I. (Cebu), and Zamboanga, which were recorded by Dickie (1874a, 1876a, 1876b, 1877). Cesare Marcacci, lieutenant on board the Vettor Pisani, collected algae at Ticao Island (Masbate) and Cavite during the circumnavigation of that ship in 1884. They were published on by Piccone (1886, 1889). The most important expedition, by far, was the Siboga Expedition to the Indonesian region, which made extensive collections in 1899 in the Sulu Archipelago. These collections contributed significantly to monographs of Halimeda (Barton, 1901), the Codiaceae (Gepp and Gepp, 1911), and the Corallinaceae (Foslie, 1904; Weber-van Bosse, 1904b). The entire phycological harvest of the expedition was treated by Weber-van Bosse (1913a, 1921, 1923, 1928).

In the Commonwealth period, Elmer Drew Merrill was impartial in his messianic pursuit of Philippine botany during 20 years of service to the Bureau of Agriculture, the Bureau of Forestry, the Bureau of Science, and the University of the Philippines (Quisumbing, 1957). Although primarily interested in vascular plants, he caused the Bureau of Science to assemble a large collection of algae. These were sent for study in 1911–1913 to M.A. Howe at the New York Botanical Garden and to W.A. Setchell at the University of California at Berkeley. No systematic study was made of these specimens at that time, but in 1951 the Chlorophyceae and Phaeophyceae at Berkeley were loaned to W.R. Taylor of the University of Michigan, who ultimately published on them (Taylor, 1966b). The all-important University of Michigan connection was established in 1935–1936, when H.H. Bartlett, Chairman of its Department of Botany, was an exchange professor at the University of the Philippines (Voss, 1961). A man of wide interests and great energy, he made massive collections of plants, including many algae, throughout the archipelago. These were sent to Ann Arbor, where the algae were studied by Taylor and his students. Bartlett returned to the Philippines under the auspices of the United States Department of Agriculture in 1940–1941, making further collections of algae in Mindanao. His collections have been cited in numerous papers, including Chou (1945, 1947) on Galaxaura, Gilbert (1942, 1943, 1947) on Chlorophyceae, Taylor (1964, 1966b) on Chlorophyceae and Phaeophyceae, and Tseng and Gilbert (1942) on Codium.

In the period of the Republic, Philippine phycology has come to share the dramatic surge of interest and activity in the science that initially was centered in the industrial nations. The leader and catalyst has been Gregorio T. Velasquez, Professor Emeritus of Botany, University of the Philippines, who received his doctorate at the University of Michigan in 1939, working on the viability of algae obtained from the digestive tract of a fresh-water fish. Although primarily interested in Cyanophyceae, Velasquez has had a profound influence on all aspects of Philippine phycology. The productivity of his academic offspring has created an urgent need for this catalog.

Awareness of benthic marine algae in the Philippines has been greatly heightened by the successful efforts to grow certain of them commercially (Eucheuma, Gracilaria, and Caulerpa). Not only has the welfare of many coastal communities been improved, but the nation as a whole has benefited from increased exports. Most phycological publications in the past two decades have been floristic, and critical taxonomic studies must now be given high priority. It is essential that the rapidly accumulating information on algal ecology, physiology, and biochemistry, much of which is of potential economic importance, be attached to correct names.

Scope of Catalog

To avoid the difficulty of having to assess each paper for its worthiness of inclusion, we have been comprehensive rather than selective, citing every reference to Philippine benthic marine algae that we could find, regardless of information content. Publications dealing with biochemical, physiological, ecological, and economic features supplement the basic core of taxonomic and morphological literature. Previous compilations were included. Twenty species that were erroneously cited by Velasquez, Trono, and Doty (1975) have been omitted from this catalog. Most of these erroneous records were taken from various reports of Siboga Expedition and based on collections from the Talud Islands and Kawio Islands, both groups belonging to Indonesia. Species were also erroneously included on the basis of collections from Borneo, Java, and Mangaia (Cook Islands).

FIGURE 1.—Map of the Philippines showing major islands.

FIGURE 2.—Map of the Philippines with the provinces designated by number. See Table 1 for a key to the numbers. (Redrawn from Vreeland et al., (second edition) 1976:25, fig. 4.)

The following is a list of species excluded from this catalog:

Cyanophyceae

Oscillatoria brevis Kützing

Rhodophyceae

Cheilosporum spectabile Harvey

Coriophyllum setchellii Weber-van Bosse

Cruoriella lemoineae Weber-van Bosse

Eucheuma dichotomum Weber-van Bosse

Galaxaura sibogae Weber-van Bosse

Gelidium anthoninii Lamouroux ex C. Agardh

Herposiphonia prorepens (Harvey) Schmitz

Laurencia pinnatifida (Hudson) Lamouroux

Liagora pulverulenta C. Agardh

Mastophora macrocarpa Montagne

Nitophyllum tongatense Grunow

Polycoelia vanhoevelii Weber-van Bosse

Polysiphonia pentamera Hollenberg

Phaeophyceae

Sphacelaria pulvinata J. Hooker and Harvey

Chlorophyceae

Boodlea vanbosseae Reinbold

Chaetomorpha clavata (C. Agardh) Kützing

Cladophora aegagropila [error in citation for Cladophora (Aegagropila)

sibogae Reinbold]

Microdictyon umbilicatum (Velley) Setchell

Microdictyon vanbosseae Setchell

Special problems relating to the compilation of records of Cyanophyceae are discussed in the introduction to that group. Brackish-water records from lagoons, canals, and swamps connecting with the sea or adjacent to it were included for all groups.

Format of Catalog

The sequence of taxa follows an assumed phylogenetic scheme to the rank of family. Genera within a family are listed alphabetically, as are species within a genus. Appended to the catalog is a list of citations of undescribed or undetermined species, assigned only to genus. An attempt was made to include in this list only those citations that are accompanied by at least some description or collection data.

In the process of constructing the taxonomic framework of this catalog, the need for various nomenclatural changes arose. Documentation for these changes is given in an appendix to avoid disruption of the catalog.

For each accepted name, synonym, and basionym at the rank of species or below, the author and the place of valid publication are given, followed by a chronological list of references to Philippine records under that name. The type locality of each basionym is indicated. When the location of the holotype or lectotype specimen is given (as in the reconciliation of the Geitlerian and Drouetian classifications of Cyanophyceae), the herbarium is indicated by the code prescribed by the Index Herbariorum (Holmgren et al., 1981). If the original publication of a name includes a Philippine record, an appropriate annotation is added unless the name has a Philippine type, in which case the name is preceded by an asterisk (*).

Nomenclatural synonyms are listed before taxonomic synonyms. The authority for taxonomic synonymy is given in an accompanying note. This innovative feature enables the user to consult and assess a taxonomic opinion as originally proposed.

Unusual nomenclatural situations are also explained in accompanying notes. A particularly troublesome and frequently encountered situation arises when the intended basionym of a binomial is shown to be a later homonym. Because, according to Article 45.3 of the International Code of Botanical Nomenclature (ICBN; Voss et al., 1983), only legitimate names are taken into consideration for purposes of priority, a later homonym (which is illegitimate according to Article 64.1 of the ICBN) cannot serve as a basionym and its author should not be placed in parentheses. The first transfer of the basionym to a different genus, however, results in a binomial that is treated as a nomen novum in accordance with Article 72, Note I. The nomen novum serves as the basionym (unless it too is a later homonym). Gracilaria lichenoides, for example, is usually attributed to (Turner) Greville, but Fucus lichenoides Turner, the intended basionym, is a later homonym, and hence not priorable. The first generic placement outside Fucus was to Gigartina by Lamouroux. Gigartina lichenoides Lamouroux thus is treated as a nomen novum and serves as the basionym of Sphaerococcus lichenoides and Gracilaria lichenoides.

TABLE 1.—List of Philippine provinces with numerical designation.

No.

Province

1

Batanes

2

Ilocos Norte

3

Cagayan

4

Abra

5

Kalinga-Apayao

6

Ilocos Sur

7

Mountain

8

Isabela

9

Ifugao

10

La Union

11

Benguet

12

Nueva Vizcaya

13

Quirino

14

Pangasinan

15

Nueva Ecija

16

Quezon

17

Zambales

18

Tarlac

19

Pampanga

20

Bulacan

21

Bataan

22

Rizal

23

Cavite

24

Laguna

25

Camarines Norte

26

Batangas

27

Camarines Sur

28

Catanduanes

29

Occidental Mindoro

30

Oriental Mindoro

31

Marinduque

32

Albay

33

Romblon

34

Sorsogon

35

Masbate

36

Northern Samar

37

Aklan

38

Samar

39

Eastern Samar

40

Capiz

41

Antique

42

Iloilo

43

Leyte

44

Cebu

45

Southern Leyte

46

Negros Occidental

47

Bohol

48

Surigao del Norte

49

Palawan

50

Negros Oriental

51

Siquijor

52

Agusan del Norte

53

Camiguin

54

Surigao del Sur

55

Misamis Oriental

56

Zamboanga del Norte

57

Misamis Occidental

58

Bukidnon

59

Agusan del Sur

60

Zamboanga del Sur

61

Lanao del Norte

62

Lanao del Sur

63

Davao

64

Davao Oriental

65

Maguindanao

66

North Cotabato

67

Basilan

68

Sultan Kudarat

69

Davao del Sur

70

South Cotabato

71

Sulu

72

Tawitawi

Geographic distribution is indicated primarily by island or island group (Figure 1), secondarily by province within the island or island group (Figure 2). The islands or island groups are listed in four successive northwest-southeast sweeps, thus: BATANES, LUZON, CATANDUANES; MINDORO, MARINDUQUE, ROMBLON, MASBATE, SAMAR, LEYTE; PANAY, GUIMARAS, NEGROS, CEBU, SIQUIJOR, BOHOL, CAMIGUIN, MINDANAO, BASILAN; PALAWAN, SULU. Within Luzon, the provinces are listed from north to south, first on the west coast (from Cagayan to Batangas), then on the east coast (from Isabela to Sorsogon). When appropriate, certain minor islands (arbitrarily limited to those labeled on United States Hydrographic Office Chart 14,200) are indicated parenthetically within provinces.

Cordero (1977a) has classified the Philippine Archipelago into four marine floristic regions, namely, the South China Sea region (Batanes, northwestern Luzon, and Mindoro), the Pacific Ocean region (northeastern Luzon, Marinduque, and Romblon), the Sulu Sea–Celebes Sea region (Palawan, Sulu Archipelago, Basilan, and the southern tip of Zamboanga del Sur), and the inland water region (all Visayan provinces and most of Mindanao). While we appreciate this attempt to bring phytogeographical order to the physiographic chaos presented by the 7,100 islands of the Philippine Archipelago, we believe that the purposes of this catalog are better served by an essentially linear sequence of localities. This sequence obviates the complication of listing Luzon in three regions and Mindanao in two regions. (In fact, the east coast of Mindanao, not specified by Cordero, should be included in the Pacific Ocean region, giving a total of three regions for that island.) It also avoids the arbitrary placement of Cagayan Province in the Pacific Ocean region. (The Babuyan Islands, which are politically part of Cagayan Province, are influenced equally by the South China Sea and the Pacific Ocean, as are the Batan Islands, which Cordero places in the South China Sea region.)

Catalog

Class CYANOPHYCEAE

Blue-green algae stand at a conspicuous distance from red, brown, and green seaweeds, both taxonomically and ecologically. Nonetheless, because they frequently are encountered by marine phycologists and preserved as herbarium specimens, they are included in this catalog. Most marine floristic accounts omit them, reflecting either insufficient interest on the part of the author or a desire to avoid numerous uncertainties peculiar to the group. When dealing with organisms that are tolerant over extremely broad physico-chemical ranges, a collector should document each collection with adequate data—including salinity or pH (whichever is appropriate) and temperature. Most herbarium specimens of blue-green algae are accompanied by meager ecological information, if any, and such indications as “along Narvacan Beach” are ambiguous when the user is faced with deciding whether the alga should be considered marine. It could have been growing in a freshwater pool next to the beach, on rocks far above high-tide mark and splashed by seawater with uncertain frequency, or on intertidal rocks. Alternatively, it could have lain free on the mud at low tide, having originated in a nearby freshwater stream or in offshore plankton. Even when critical information is given on the specimen, it is frequently omitted by the recording author.

Deciding which records of blue-green algae should be considered marine and benthic is only the first obstacle. This group is plagued with other uncertainties. Its taxonomy has been polarized between the Geitlerian and Drouetian schools. Geitler (1942), building on the monographs of Bornet and Flahault (1886–1888) and Gomont (1892–1893), recognized 4 orders, 22 families, 140 genera, and more than 1200 species, while Drouet (1981) recognized 2 orders, 6 families, 24 genera, and 61 species. Recently, microbiologists have proposed a system of classification based solely on characters revealed in pure cultures (Rippka et al., 1979).

Having delineated the taxonomic units, one is then faced with the problem of determining their correct names. This is a critical process that is complicated in the case of blue-green algae by the provision of the International Code of Botanical Nomenclature that nomenclature of the Nostocales begins with the monographs of Bornet and Flahault and of Gomont. For reasons discussed in the appended Nomenclatural Notes, later starting points are ignored in this catalog.

All names recorded for the Philippines except the few published by Dickie (1876a, 1877) essentially follow Bornet and Flahault for heterocystous Nostocales and Gomont for homocystous Nostocales. For this reason, the records have been arranged according to those monographs rather than being integrated into the Drouetian system. A few records appear under two names, one given by Velasquez or another Filipino worker, the other by Drouet. These instances of duplication are noted. The catalog of blue-green algal records is followed by a reconciliation of the two systems.

As though the foregoing ecological, taxonomic, and nomenclatural uncertainties were not enough, recently the inclusion of these organisms in the algae has been questioned. Placing greater importance on the lack of membrane-bounded organelles than on the presence of oxygen-evolving photosynthesis, certain workers (led by the late Roger Stanier) treat these organisms as bacteria (Cyanobacteria). Some of these workers appear to have little faith in their decision, however, as they publish articles on Cyanobacteria in phycological journals!

Order CHROOCOCCALES

Family CHROOCOCCACEAE

Anacystis Meneghini

Anacystis aeruginosa (Zanardini) Drouet and Daily

Palmogloea aeruginosa Zanardini, 1872:162, pl. IX.D [type locality: Cape Datu (Tandjoeng Datoek), boundary between Sarawak, Malaysia and West Kalimantan, Indonesia].

Anacystis aeruginosa (Zanardini) Drouet and Daily, 1948:77.—Velasquez, 1950:312.—Drouet and Daily, 1956:77.—Velasquez and Soriano, 1957:486.—Velasquez, 1962a:279, pl. 1: fig. 7.—Velasquez, Trono, and Doty, 1975:127.—Cordero, 1976c:10, 12.—Reyes, 1978:139, pl. 1: figs. 1, 2.—Martinez, 1984:24.

PHILIPPINE DISTRIBUTION.—BATANES. MINDORO: Oriental Mindoro. SIQUIJOR. PALAWAN.

Family ENTOPHYSALIDACEAE

Entophysalis Kützing

Entophysalis conferta (Kützing) Drouet and Daily

Palmella conferta Kützing, 1845:149 [type locality: Cuxhaven, West Germany].

Entophysalis conferta (Kützing) Drouet and Daily, 1948:79; 1956:118.—Velasquez, 1962a:281, pl. 1: fig. 9.—Velasquez, Trono, and Doty, 1975:143.—Martinez, 1984:36.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal, Manila.

Order NOSTOCALES

Family NOSTOCACEAE

Anabaina Bory de Saint-Vincent

This generic name is usually spelled Anabaena, an orthographic variant used by Bornet and Flahault (1888 [1886–1888]:224).

Anabaina pseudoscillatoria Bory de Saint-Vincent

Anabaina pseudoscillatoria Bory de Saint-Vincent, 1822:308 [“pseudo-oscillatoria”] [type locality: not specified; Everquem, near Gent, Belgium fide Drouet, 1978:157, 186].

Anabaina oscillarioides Bory de Saint-Vincent, 1831:29 [explanation of pl. LII: figs. 7a–c].—Drouet, 1978:195.—Cordero, 1981d:62, fig. 1.—Martinez, 1984:22.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. PANAY: Aklan. PALAWAN (incl. Cuyo I.).

NOTE.—Bory de Saint-Vincent changed the epithet of this specific name without explanation. He has been followed in this regard by all subsequent authors. The Oriental Mindoro record (Velasquez 1045) was previously published under Hormothamnium enteromorphoides (Velasquez, 1962a:351), while the Palawan records (Velasquez 3019 and Merrill 9158) were previously published under H. solutum (Velasquez, 1955:175 and 1962a:351, respectively).

Hormothamnium Grunow

The variant spelling Hormothamnion was initiated by Bornet and Flahault (1888 [1886–1888]:259) and has been used by all subsequent authors.

Hormothamnium enteromorphoides Grunow

Hormothamnium enteromorphoides Grunow, 1867:31, pl. I: fig. 25 [type locality: Guadeloupe].—Velasquez, 1950:323; 1962a:350, pl. 10: fig. 120.—Reyes, 1978:141, pl. 2: figs. 5, 6.—Martinez, 1984:39.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. SIQUIJOR.

NOTE.—Drouet (1978:165, 195), after examining the holotype of this species (in W), assigned it to Anabaina oscillarioides Bory de Saint-Vincent (= A. pseudoscillatoria Bory de Saint-Vincent).

Hormothamnium solutum Bornet and Grunow

Hormothamnium solutum Bornet and Grunow in Bornet and Flahault, 1888 [1886–1888]:259 [lectotype locality: Nouméa, New Caledonia fide Drouet, 1978:166, 198].—Velasquez, 1955:175, pl. [I] fig. 45; 1962a:351, pl. 10: figs. 121, 121a.—Martinez, 1984:39.

PHILIPPINE DISTRIBUTION.—PALAWAN (incl. Cuyo I.).

NOTE.—Drouet, after choosing and examining the lectotype of this species (in PC), assigned it to Anabaina oscillarioides Bory de Saint-Vincent (= A. pseudoscillatoria Bory de Saint-Vincent).

Family OSCILLATORIACEAE

Hydrocoleum Kützing

Hydrocoleum cantharidosmum (Montagne) Gomont

Lyngbya? cantharidosma Montagne, 1841 [1839–1842]:188 [type locality: Islas Canarias].

Hydrocoleum cantharidosmum (Montagne) Gomont, 1890:353.—Velasquez, 1950:319; 1962a:335, pl. 7: fig. 93.—Martinez, 1984:40.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:266, 297), after examining the holotype of this species (in PC), assigned it to Microcoleus lyngbyaceus (Kutzing) P. Crouan and H. Crouan.

Hydrocoleum comoides (Harvey) Gomont

Calothrix comoides Harvey, 1863:lxii [lectotype locality: Port Phillip Heads, Victoria, Australia fide Drouet, 1968:276, 308; 1973:207].

Hydrocoleum comoides (Harvey) Gomont, 1892[1892–1893]:335.—Velasquez, 1962a:335.—Martinez, 1984:40.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—Although Harvey gave Cape Riche [Western Australia] as the only locality for Calothrix comoides, he cited two of his collections, numbers 597 and 598. The latter is from Port Phillip Heads and that specimen (in BM) was chosen as lectotype by Drouet, who assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Hydrocoleum glutinosum (C. Agardh) Gomont

Lyngbya glutinosa C. Agardh, 1824:73 [type locality: “In sinu Codano” (Kattegat, between Denmark and Sweden)].

Hydrocoleum glutinosum (C. Agardh) Gomont, 1890:353.—Velasquez, 1950:319; 1962a:334, pl. 7: fig. 92.—Martinez, 1984:40.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:263), incorrectly treating Lyngbya glutinosa as a nomenclatural synonym of Conferva majuscula Dillwyn (1809 [1802–1809]:40, suppl. pl. A), referred this species to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan on the basis of the Dillwyn type. In fact, C. Agardh (1824:73, 74) cited “Osc. majuscula, Lyngb. & fl. D. t. 1549. f. 1” under Lyngbya glutinosa, intending this citation to refer to a circumscription (Lyngbye, 1819:91) that excluded the type of Conferva majuscula, which he cited in the synonymy of Lyngbya crispa (C. Agardh) C. Agardh, as “Oscill. majuscula, Dillw. t. A.”

Hydrocoleum lyngbyaceum Kützing

Hydrocoleum lyngbyaceum Kützing, 1849:259 [lectotype locality: Cherbourg, France fide Drouet, 1968:272, 296].—Velasquez, 1950:318.—Agor, 1962:34.—Velasquez, 1962a:336, pl. 7: fig. 94.—Vannajan and Trono, 1977:36, fig. 4.—Fortes, 1981b:396.—Martinez, 1984:40.

Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan, 1867:114.—Drouet, 1968:306–307.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), Ilocos Sur, Pangasinan, Manila, MINDORO: Oriental Mindoro. CEBU. SULU: Tawitawi.

NOTE.—Drouet (1968:224–225) placed Hydrocoleum and Lyngbya, among other genera, in the synonymy of Microcoleus. The records for Ilocos Sur, Cebu, and Tawitawi, which were added by Drouet, are based on collections previously reported under Phormidium ambiguum (Velasquez, 1962a:306), Lyngbya confervoides (Velasquez, 1940:271; 1962a:321), and L. majuscula (Velasquez, 1955:167), respectively.

Lyngbya C. Agardh

Lyngbya aestuarii (Mertens) Liebman

Conferva aestuarii Mertens in Jürgens, 1816: decade 2, no. 8 [type locality: “Dynastia Jeverana” (Oldenburg, West Germany)].

Lyngbya aestuarü (Mertens) Liebman, 1839:492.—Velasquez, 1941:193; 1955:166, pl. II: fig. 25; 1962a:317, pl. 3: fig. 65.—Cordero, 1976c:14; 1979b:278; 1981d:62, fig. 2.—Martinez, 1984:41.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Pangasinan, Rizal. PANAY: Aklan. PALAWAN (Cuyo I.).

NOTE.—Drouet (1968:263, 292) chose a specimen in L as the lectotype of this species and assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Lyngbya confervoides C. Agardh

Lyngbya confervoides C. Agardh, 1824:73 [lectotype locality: Spain (probably Cádiz) fide Drouet, 1968:264].—Velasquez, 1940:271; 1950:318.—Agor, 1962:33; Velasquez, 1962a:321, pl. 3: fig 67.—Reyes, 1978:139, pl. 1: figs. 9, 10.—Saraya and Trono, 1980:6.—Velasquez, 1980:127, table 1.—Fortes, 1981b:396.—Martinez, 1984:41.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro. CEBU. SIQUIJOR.

NOTE.—Drouet (1968:264, 296) chose a specimen in LD as the lectotype of this species and assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Lyngbya ferruginea C. Agardh

Lyngbya ferruginea C. Agardh, 1824:73 [type locality: “In sinu ad Hofmansgave Fioniae” (Fyn, Denmark)].—Dickie, 1877:489.—Velasquez, Trono, and Doty, 1975:152.—Martinez, 1984:42.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

NOTE.—Drouet (1968:264, 292), after examining the holotype of this species (in LD) assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Lyngbya infixa Frémy

Lyngbya infixa Frémy, 1932:1414 [type locality: Banyuls-sur-Mer, Pyrénées-Orientales, France].—Cordero, 1976c:14.

PHILIPPINE DISTRIBUTION.—BATANES.

NOTE.—Drouet (1968:52, 73) chose a specimen in the personal herbarium of J. Feldmann as the lectotype of this species and assigned it to Schizothrix calcicola (C. Agardh) Gomont.

Lyngbya lutea (C. Agardh) J.E. Areschoug

Oscillatoria lutea C. Agardh, 1824:68 [type locality: Hälsingborg, Sweden].

Lyngbya lutea (C. Agardh) J.E. Areschoug, 1850:443.—Fortes and Trono, 1980:53.—Martinez, 1984:42.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:185, 188), after examining the holotype of this species (in LD), retained its original disposition, as Oscillatoria lutea C. Agardh.

Lyngbya majuscula (Dillwyn) Harvey

Conferva majuscula Dillwyn, 1809 [1802–1809]:40, suppl. pl. A [lectotype locality: England fide Drouet, 1968:263].

Lyngbya majuscula (Dillwyn) Harvey, 1833:370.—Dickie, 1876a:245.—Velasquez, 1940:271; 1950:317; 1955:167, pl. II: fig. 26.—Velasquez and Soriano, 1957:486.—Agor, 1962:33.—Velasquez, 1962a:319, pl. 4: fig. 68; 1971:428, fig. 3—Cornejo and Velasquez, 1972:171, pl. 1: fig. 1; pl. 4: fig. 28.—Reyes, 1972:135.—Velasquez, Trono, and Doty, 1975:152.—Cordero, 1976c:10, 14, fig. A.—Vannajan and Trono, 1977:37.—Reyes, 1978:139, pl. 1: figs. 7, 8.—Trono, 1978:2.—Saraya and Trono, 1980:6.—Liao and Sotto, 1980:94.—Chan, 1981:387, 389.—Fortes, 1981b:396.—Trono and Ang, 1982:2.—Martinez, 1984:42.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Pangasinan, Bataan, Cavite, Batangas. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. NEGROS: Negros Oriental, CEBU. SIQUIJOR. MINDANAO: Zamboanga. PALAWAN (incl. Bugsuk I., Cuyo I.). SULU: Sulu (Tapul Group), Tawitawi.

NOTE.—Drouet (1968:263, 295) chose a specimen in LD as the lectotype of this species and assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Lyngbya martensiana Meneghini

Lyngbya martensiana Meneghini, 1837:330 [type locality: Euganean thermal springs, Veneto, Italy].—Agor, 1962:33.—Velasquez, 1980:127, table 1.—Martinez, 1984:43.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:266, 295), after examining the holotype of this species (in FI), assigned it to Microcoleus lyngbyaceus (Kutzing) P. Crouan and H. Crouan.

Lyngbya mesotricha Skuja

Lyngbya mesotricha Skuja, 1949:54 [type locality: Burma (Rangoon fide Drouet, 1968:54, 87)]—Fortes and Trono, 1980:53.—Martinez, 1984:43.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:54, 87) chose a specimen in PH as the lectotype of this species and assigned it to Schizothrix calcicola (C. Agardh) Gomont.

Lyngbya rosea W.R. Taylor

Lyngbya rosea W.R. Taylor, 1928:45, pl. 1: fig. 22 [type locality: Dry Tortugas, Florida, USA].—Vannajan and Trono, 1977:37.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

NOTE.—Drouet (1968:91, 95), after examining the holotype of this species (in the personal herbarium of W.R. Taylor), assigned it to Schizothrix mexicana Gomont.

Lyngbya semiplena (C. Agardh) J. Agardh

Calothrix semiplena C. Agardh, 1827:634 [type locality: Trieste, Italy].

Lyngbya semiplena (C. Agardh) J. Agardh, 1842:11.—Agor, 1962:33.—Velasquez, 1962a:319, pl. 3: fig. 66.—Reyes, 1978:140, pl. 1: figs. 11, 12.—Martinez, 1984:44.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro. SIQUIJOR.

NOTE.—Drouet (1968:264–265, 295), after examining the holotype of this species (in LD), assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Lyngbya sordida Gomont

Lyngbya sordida Gomont, 1893[1892–1893]:126, pl. 2: fig. 21 [lectotype locality: Dalmatian coast of Yugoslavia; see Note].—Velasquez, 1940:270; 1962a:316, pl. 3: fig. 64.—Cordero, 1976c:14, fig. B.—Martinez, 1984:44.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), MINDORO: Oriental Mindoro. CEBU.

NOTE.—Drouet (1968:88) treated Lyngbya sordida Gomont as a nomenclatural synonym of Leibleinia capillacea Kützing (1843b:221), which he referred to Schizothrix mexicana Gomont. When establishing Lyngbya sordida, however, Gomont listed Leibleinia capillacea as a species inquirenda, so that the two names do not share the same type. For the distribution of Lyngbya sordida, Gomont gave several localities in the North Atlantic Ocean and Mediterranean Sea as well as the Antilles and Tonga. When later starting points in Cyanophyceae are abandoned, as in the present catalog, L. sordida is seen to be a superfluous name, since Gomont included in its synonymy several legitimate names, the earliest being Leibleinia polychroa and L. violacea, both published by Meneghini (1844:304) and both transferred to Lyngbya by Rabenhorst (1847:83 and 1865:144, respectively). The type specimens of the Meneghini species should be in his herbarium in FI, but Drouet chose specimens distributed by Meneghini and housed in L. We herewith choose the type of Leibleinia polychroa as the lectotype of Lyngbya sordida. Gomont did not purposely by-pass Meneghini’s names; rather, he thought that Calothrix sordida Zanardini (1843:63) was an earlier available name. That name was published without a description, however, and is thus invalid. Moreover, Lyngbya sordida Gomont is a later homonym of L. sordida P. Crouan and H. Crouan (in Mazé and Schramm, 1878:21), based on Leibleinia sordida Kűtzing (1845:179), an illegitimate substitute name for Calothrix luteofusca C. Agardh (1827:635).

Microcoleus Desmazières

Microcoleus acutissimus Gardner

Microcoleus acutissimus Gardner, 1927a:55, pl. II: fig. 2 [type locality: Sabana Grande, Puerto Rico].—Velasquez, 1962a:332, pl. 6: figs. 89, 89a.—Fortes and Trono, 1980:54.—Fortes, 198lb:396.—Martinez, 1984:47.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:135, 139), after examining the holotype of this species (in NY), assigned it to Schizothrix tenerrima (Gomont) Drouet.

Microcoleus tenerrimus Gomont

Microcoleus tenerrimus Gomont, 1892[1892–1893]:355, pl. 14: figs. 9–11[lectotype locality: Guadeloupe fide Drouet, 1968:135, 139].

Schizothrix tenerrima (Gomont) Drouet, 1968:135 [including Philippine record on page 140].

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—This record (Velasquez 723) had previously been published under Microcoleus acutissimus (Velasquez, 1962a:332).

Oscillatoria Vaucher

Oscillatoria bonnemaisonii (P. Crouan and H. Crouan)
P. Crouan and H. Crouan

Oscillaria bonnemaisonii P. Crouan and H. Crouan in Desmazieres, 1858: no. 537 [type locality: passage de Plougastel, Finistère, France].

Oscillatoria bonnemaisonii; (P. Crouan and H. Crouan) P. Crouan and H. Crouan, 1860:371.—Velasquez, 1950:315; 1962a:288, pl. 1: fig. 16.—Cordero, 1976c:10, 15, figs, C, D.—Velasquez, 1980:127, table 1.—Martinez, 1984:57.

PHILIPPINE DISTRIBUTION.—BATANES. MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:274, 296) chose a specimen in PC as the lectotype of this species and assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Oscillatoria corallinae (Kützing) Gomont

Leibleinia corallinae Kützing, 1849:276 [type locality: “ad oras gallicas boreales” (Arromanches-les-Bains, Calvados, France fide Drouet, 1968:272, 296)].

Oscillatoria corallinae (Kützing) Gomont, 1893[1892–1893]:218.—Velasquez, 1962a:288, pl. 1: fig. 17.—Fortes, 198lb:396.—Martinez, 1984:59.

PHILIPPINE DISTRIBUTION.—LUZON: Bataan. MINDANAO: Zamboanga.

NOTE.—Drouet (1968:272, 296), after examining the holotype of this species (in L), assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Oscillatoria curviceps C. Agardh

Oscillatoria curviceps C. Agardh, 1824:68 [type locality: “In stagnis Scaniae” (near Lund, Sweden fide Drouet, 1968:264, 291)].—Fortes and Trono, 1980:52.—Fortes, 1981b:396.—Martinez, 1984:59.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:264, 291), after examining the holotype of this species (in LD), assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Oscillatoria margaritifera (Kützing) Gomont

Oscillaria margaritifera Kützing, 1847b:31, pl. 43: fig. X [type locality:Calvados, France].

Oscillatoria margaritifera (Kützing) Gomont, 1893[1892–1893]:216.—Velasquez,1962a:293, pl. 2: fig. 23.—Vannajan and Trono, 1977:36, fig.1b.—Saraya and Trono, 1980:6.—Chan, 1981:387, 389.—Martinez, 1984:62.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite. MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:270, 296), after examining the holotype of this species (in L), assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Oscillatoria nigroviridis Thwaites

Oscillatoria nigroviridis Thwaites in Harvey, 1849 [1847–1851]: pl. CCLI:A [“nigro-viridis”] [type locality: Shirehampton near Bristol, England].—Velasquez, 1962a:293, pl. 2: fig. 24.—Fortes and Trono, 1980:52.—Fortes, 1981b:396.—Martinez, 1984:63.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal. MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:145, 154) chose a specimen in K (now in BM) as the lectotype of this species and assigned it to Porphyrosiphon notarisii (Meneghini) Kützing.

Oscillatoria sancta (Kützing) Gomont

Oscillaria sancta Kützing, 1847b:30, pl. 42: fig. VII [type locality: “In der ‘Aequa santa’” (near Roma, Italy fide Drouet, 1968:270, 294)].

Oscillatoria sancta (Kützing) Gomont, 1893[1892–1893]:209.—Fortes and Trono, 1980:53, fig. 1.—Fortes, 1981b:396.—Martinez, 1984:65.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:270, 294), after examining the holotype of this species (in L), assigned it to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Phormidium Kützing

Phormidium ambiguum Gomont

Amphithrix amoena Kützing, 1843b:220 [type locality: Nordhausen, East Germany].

Phormidium ambiguum Gomont, 1893[1892–1893]:178, pl. 5: fig. 10.—Velasquez, 1962a:306, pl. 3: fig. 50.—Martinez, 1984:68.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Sur.

NOTE.—Phormidium ambiguum was proposed as a substitute name for Amphithrix amoena Kützing in consideration of the prior existence of P. amoenum Kützing (1843b:192). Drouet (1968:232, 246), after examining the holotype of A. amoena (in L), assigned it to Microcoleus vaginatus (Vaucher) Gomont. The Ilocos Sur record, however, was referred by Drouet (1968:307) to Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan.

Phormidium crosbyanum Tilden

Phormidium crosbyanum Tilden, 1909: no. 645 [type locality: Waianae, Oahu, Hawaiian Is.].—Velasquez, 1940:271; 1950:316; 1955:165, pl. 1: fig. 20; 1962a:304, pl. 2: fig. 46; 1971:427, fig. 2.—Reyes, 1972:135.—Velasquez, Trono, and Doty, 1975:159.—Cordero, 1976c:10, 15; 1981d:63, fig. 3.—Martinez, 1984:68.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, MINDORO: Oriental Mindoro. PANAY: Aklan. NEGROS: Negros Oriental, SULU: Sulu (Siasi I.), Tawitawi.

NOTE.—Drouet (1968:48, 83), after examining the holotype of this species (in MIN), assigned it to Schizothrix calcicola (C. Agardh) Gomont.

Phormidium laysanense Lemmermann

Phormidium laysanense Lemmermann, 1905:619, pl. VII: figs. 4, 5 [type locality: Laysan Islet, Hawaiian Is.].—Agor, 1962:33.—Martinez, 1984:69.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—Drouet (1968:213), on the basis of the original description, assigned this species to Oscillatoria erythraea (Ehrenberg) Drouet. Although the latter species (better known as Trichodesmium erythraeum Ehrenberg) is generally planktonic, Agor (l.c.) reported his material as being epiphytic on brown algae.

Phormidium penicillatum Gomont

Phormidium penicillatum Gomont, 1893:LXXXVIII, pl. IV: figs. 5–7; in Jadin, 1893:CLIX [“penicellatum”] [type locality: Saint-Gilles, Réunion].—Velasquez, 1962a:306.—Martinez, 1984:70.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:204, 212), after examining the holotype of this species (in PC), assigned it to Oscillatoria submembranacea (Ardissone and Strafforello) Drouet.

Phormidium persicinum (Reinke) Gomont

Lyngbya persicina Reinke, 1889:91 [type locality: Kieler Bucht, West Germany].

Phormidium persicinum (Reinke) Gomont, 1893[1892–1893]:164.—Velasquez, 1962a:301, pl. 2: fig. 41—Reyes, 1978:139, pl. 1: figs. 5, 6.—Martinez, 1984:70.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. SIQUIJOR.

NOTE.—Drouet (1968:43, 68) chose a specimen in PC as the lectotype of this species and assigned it to Schizothrix calcicola (C. Agardh) Gomont.

Phormidium submembranaceum

(Ardissone and Strafforello) Gomont

Oscillaria submembranacea Ardissone and Strafforello, 1877:65 [type locality: Porto Maurizio, Liguria, Italy].

Phormidium submembranaceum (Ardissone and Strafforello) Gomont, 1893[1892–1893]:180.

Oscillatoria submembranacea (Ardissone and Strafforello) Drouet, 1968:203 [including Philippine records on page 212].

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), MINDORO: Oriental Mindoro.

NOTE.—These records were previously published as Phormidium penicillatum Gomont by Velasquez (1962a:306).

Phormidium tinctorium Kützing

Phormidium tinctorium Kützing, 1847b:35, pl. 49: fig. Ill[type locality: Falaise, Calvados, France].—Cornejo and Velasquez, 1972:171, pl. 1: fig. 2.—Velasquez, Trono, and Doty, 1975:159—Martinez, 1984:71.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

NOTE.—Drouet (1968:100, 105), after examining the holotype of this species (in L), assigned it to Schizothrix friesii (C. Agardh) Gomont.

Phormidium valderiae (Delponte) Schmidle

Leptothrix valderiae Delponte in Garelli. 1857:35 [“valderia”] [type locality: Valdieri, Piemonte, Italy].

Phormidium valderiae (Delponte) Schmidle, 1901:100, footnote.

Phormidium valdenanum Gomont, 1893[1892–1893]:167—Velasquez, 1955:164, pl. I: fig. 19; 1962a:305, pl. 3: fig. 48.—Martinez, 1984:72.

PHILIPPINE DISTRIBUTION.—PALAWAN.

NOTE.—Phormidium valderianum was proposed as a substitute name for Leptothrix valderiae. When later starting points are abandoned, the correct name for this species becomes P. valderiae. Drouet (1968:38, 71) chose a specimen in PC as the lectotype of this species and assigned it to Schizothrix calcicola (C. Agardh) Gomont.

Schizothrix Kützing

Schizothrix mexicana Gomont

Schizothrix mexicana Gomont, 1892[1892–1893]:304 [type locality: Rio Guatulco, Oaxaca, Mexico).—Drouet, 1968:98.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. CEBU.

NOTE.—The Mindoro records (Velasquez 871, 988) had previously been published under Lyngbya majuscula (Velasquez, 1962a:320) and L. sordida (Velasquez, 1962a:317), respectively.

Sirocoleum Kützing

Sirocoleum kurzii (Zeller) Gomont

Chthonoblastus kurzii Zeller, 1873:178 [type locality: Elephant Point, Pegu, Burma].

Sirocoleum kurzii (Zeller) Gomont, 1892[1892–1893]:349.

Porphyrosiphon kurzii (Zeller) Drouet, 1968:162 [including Philippine record on page 164].

PHILIPPINE DISTRIBUTION.—LUZON: Rizal.

Spirulina Turpin

Spirulina major Kützing

Spirulina major Kützing, 1843b:183 [lectotype locality: Weissenfels, East Germany fide Drouet, 1968:17, 20].—Velasquez, 1980:127, table 1.—Martinez, 1984:82.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1968:17, 20) chose a specimen in L as the lectotype of this species and assigned it to Spirulina subsalsa Oersted.

Symploca Kützing

Symploca howei Gardner

Symploca howei Gardner, 1932:283, fig. 1, pl. 2: fig. 9 [type locality: Cayo Don Luis near Point Montalva, Puerto Rico].—Velasquez and Soriano, 1957:486.—Reyes, 1972:135.—Velasquez, Trono, and Doty, 1975:163.—Reyes, 1978:140, pl. 2: fig. 1, 2.—Martinez, 1984:85.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental. SIQUIJOR.

NOTE.—Drouet (1968:92, 95), after examining the holotype of this species (in NY), assigned it to Schizothrix mexicana Gomont.

Symploca hydnoides (Harvey) Kützing

Calothrix hydnoides Harvey, 1833:368 [type locality: Appin, Argyll, Scotland].

Symploca hydnoides (Harvey) Kützing, 1849:272.—Velasquez, 1962a:312, pl. 3: fig. 59.—Reyes, 1972:136.—Velasquez, Trono, and Doty, 1975:164.—Reyes, 1978:140, pl. I: figs. 13, 14.—Saraya and Trono, 1980:7.—Liao and Sotto, 1980:94.—Martinez, 1984:85.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), Pangasinan. MINDORO: Oriental Mindoro. NEGROS: Negros Oriental, CEBU. SIQUIJOR.

NOTE.—Drouet (1968:310; 1973:177, 185) chose a specimen in LD as the lectotype of this species and assigned it to Calothrix crustacea Thuret.

Symploca laeteviridis Gomont

Symploca laeteviridis Gomont, 1893[1892–1893]:109, pl. 2: figs. 6–8 [type locality: Key West, Florida, USA].—Velasquez, 1955:168, pl. II: fig. 28; 1962a:311, pl. 3: fig. 58.—Martinez, 1984:85.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. SULU: Tawitawi.

NOTE.—Drouet (1968:111, 118), after examining the holotype of this species (in PC), assigned it to Schizothrix arenaria (Berkeley) Gomont.

Family RIVULARIACEAE

Calothrix C. Agardh

Calothrix aeruginea (Kützing) Thuret

Leibleinia aeruginea Kützing, 1843b:221 [type locality: Trieste, Italy].

Calothrix aeruginea (Kützing) Thuret, 1875:381.—Fortes and Trono, 1980:54, fig. 2.—Fortes, 1981b:396.—Martinez, 1984:28.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1973:168, 184), after examining the holotype of this species (in L), assigned it to Calothrix crustacea Thuret.

Calothrix confervicola (Dillwyn) C. Agardh

Conferva confervicola Dillwyn, 1802 [1802–1809]: pl. 8 [lectotype locality: Aberystwyth, Wales fide Drouet, 1973:164].

Calothrix confervicola (Dillwyn) C. Agardh, 1824:70.—Velasquez, 1962a:353, pl. 10: fig. 125.—Martinez, 1984:28.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.).

NOTE.—Drouet (1973:163–164) chose a specimen in OXF as the lectotype of this species and assigned it to Calothrix crustacea Thuret.

Calothrix contarenii (Zanardini) Bornet and Flahault

Rivularia contarenii Zanardini, 1840a:134 [lectotype locality: Venezia, Italy fide Drouet, 1973:167, 184].

Calothrix contarenii (Zanardini) Bornet and Flahault, 1886a [1886–1888]:355.—Saraya and Trono, 1980:7.—Martinez, 1984:29.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—Drouet (1973:166–167, 184) chose a specimen in PC as the lectotype of this species and assigned it to Calothrix crustacea Thuret.

Calothrix crustacea Thuret

Calothrix crustacea Thuret in Bornet and Thuret, 1876:13, pl. IV: figs. 1–6 [lectotype locality: Tangiers, Morocco fide Drouet, 1973:175, 186].—Fan, 1956:176.—Drouet, 1973:194.—Martinez, 1984:29.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), Ilocos Sur. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. PALAWAN, SULU: Tawitawi.

NOTE.—These records have also been reported as follows: Cagayan under Calothrix confervicola (Velasquez, 1962a:353), Ilocos Sur under C. parietina (Fan, 1956:168), Oriental Mindoro (Velasquez 1076) under Brachytrichia maculans (Velasquez, 1962a:356), Palawan under Rivularia bullata (Velasquez, 1962a:355), and Tawitawi under Calothrix pilosa (Velasquez, 1955:181).

Calothrix epiphytica W. West and G.S. West

Calothrix epiphytica W. West and G.S. West, 1897:240 [type locality: Mossâmedes, Angola].—Cornejo and Velasquez, 1972:172, pl. 1: fig. 6.—Velasquez, Trono, and Doty, 1975:130.—Martinez, 1984:29.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

NOTE.—Drouet (1973:121,150) chose a specimen in UC as the lectotype of this species and assigned it to Calothrix parietina (Nägeli ex Kützing) Thuret.

Calothrix parietina (Nägeli ex Kützing) Thuret

Schizosiphon parietinus Nägeli ex Kützing, 1849:327 [type locality: Zürich, Switzerland].

Calothrix parietina (Nägeli ex Kützing) Thuret, 1875:381.—Fan, 1956:168.—Martinez, 1984:29.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Sur.

NOTE.—This record has also been reported under Phormidium ambiguum (Velasquez, 1962a:306) and Calothrix crustacea (Drouet, 1973:194).

Calothrix pilosa Harvey

Calothrix pilosa Harvey, 1858:106, pl. XLVIII.C [type locality: Key West, Florida, USA].—Velasquez, 1940:269; 1950:323; 1955:181, pl. V: figs. 57a, 57b.—Fan, 1956:171–172.—Velasquez, 1962a:353, pl. 10: figs. 123, 123a.—Martinez, 1984:29.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon, MINDORO: Oriental Mindoro. LEYTE (Biliran I.), SULU: Tawitawi.

NOTE.—Drouet (1973:39, 77) chose a specimen in K (in BM) as the lectotype of this species and assigned it to Scytonema hofmannii C. Agardh [= S. hofman-bangii C. Agardh].

Calothrix robusta Setchell and Gardner

Calothrix robusta Setchell and Gardner in Gardner, 1918:473, pl. 40: fig. 22 [type locality: Cypress Point, Monterey County, California, USA].—Velasquez, 1950:324.—Martinez, 1984:30.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1973:177, 190), after examining the holotype of this species (in UC), assigned it to Calothrix crustacea Thuret.

Calothrix scopulorum (Weber and Mohr) C. Agardh

Conferva scopulorum Weber and Mohr, 1804:195, pl. 3: fig. 3 [type locality: Varberg, Sweden].

Calothrix scopulorum (Weber and Mohr) C. Agardh, 1824:70.—Fortes and Trono, 1980:54.—Fortes, 1981b:396.—Martinez, 1984:30.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1973:164, 183) chose a specimen in K (in BM) as the lectotype of this species and assigned it to Calothrix crustacea Thuret.

Calothrix viguieri Frémy

Calothrix viguieri Frémy, 1930:252, fig. 226 [type locality: Gabon].—Cornejo and Velasquez, 1972:172, pl. 1: fig. 5.—Velasquez, Trono, and Doty, 1975:1 30.—Martinez, 1984:30.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

NOTE.—Drouet (1973:131), on the basis of the original description, assigned this species to Calothrix parietina (Nägeli ex Kützing) Thuret.

Dichothrix Zanardini

Dichothrix gypsophila (Kützing) Bornet and Flahault

Schizosiphon gypsophilus Kützing, 1843b:234, pl. 6: fig. II [type locality: “An Gypswänden des südlichen Harzes” (Sachswerfen, near Nordhausen, East Germany fide Drouet, 1973:104)].

Dichothrix gypsophila (Kützing) Bornet and Flahault, 1886a [1886–1888]:377.—Cornejo and Velasquez, 1972:172, pl. 1: fig. 4.—Velasquez, Trono, and Doty, 1975:140.—Martinez, 1984:35.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

NOTE.—Drouet (1973:103–104, 145), after examining the holotype of this species (in L), assigned it to Calothrix parietina (Nägeli ex Kützing) Thuret.

Gardnerula De Toni f.

Gardnerula corymbosa (Harvey) De Toni f.

Microcoleus corymbosus Harvey, 1858:109, pl. XLVIII:B [type locality: Key West, Florida, USA].

Gardnerula corymbosa (Harvey) De Toni f., 1936:[5].—Velasquez, 1940:270; 1950:327; 1962a:355, pl. 10: figs. 128, 128a, 128b.—Reyes, 1978:142, pl. 2: figs. 14–17.—Martinez, 1984:37.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon, MINDORO: Oriental Mindoro. SIQUIJOR.

NOTE.—Drouet (1973:173, 189), after examining the holotype of this species (in TCD), assigned it to Calothrix crustacea Thuret.

Rivularia C. Agardh

Rivularia bullata (Poiret) Berkeley

Ulva bullata Poiret, 1808:175 [type locality: Siriac, Bretagne, France].

Rivularia bullata (Poiret) Berkeley, 1833:8.—Velasquez, 1962a:355, pl. 10: fig. 127.—Reyes, 1978:141, pl. 2: figs. 12, 13.—Martinez, 1984:75.

PHILIPPINE DISTRIBUTION.—SIQUIJOR. PALAWAN.

NOTE.—Drouet (1973:165, 185), after examining the holotype of this species (in PC), assigned it to Calothrix crustacea Thuret.

Rivularia mesenterica (Kützing) Thuret

Heteractis mesenterica Kützing, 1843b:236 [type locality: Pula, Yugoslavia].

Rivularia mesenterica (Kützing) Thuret, 1875:382.—Reyes, 1978:141, pl. 2: figs. 7–9.—Martinez, 1984:76.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

NOTE.—Drouet (1973:168–169, 184), after examining the holotype of this species (in L), assigned it to Calothrix crustacea Thuret.

Rivularia nitida C. Agardh

Rivularia nitida C. Agardh, 1812 [1810–1812]:44 [type locality: Stockholm, Sweden].—Fortes and Trono, 1980:55, fig. 3.—Fortes, 1981b:396.—Martinez, 1984:76.

PHILIPPINE DISTRIBUTION—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1973:165, 183), being unable to find original material of this species, chose a specimen in LD (from Båstad, Sweden) as a neotype and assigned it to Calothrix crustacea Thuret.

Rivularia polyotis (J. Agardh) Hauck

Diplotrichia polyotis J. Agardh, 1842:10 [type locality: Pag I., Yugoslavia].

Rivularia polyotis (J. Agardh) Hauck, 1884 [1883–1885]:495.—Reyes, 1978:141, pl. 2: figs. 10, 11.—Martinez, 1984:76.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

NOTE.—Drouet (1973:167–168, 184), after examining the holotype of this species (in LD), assigned it to Calothrix crustacea Thuret.

Family SCYTONEMATACEAE

Scytonema C. Agardh

Scytonema hofman-bangii C. Agardh

Scytonema hofman-bangii C. Agardh, 1812 [1810–1812]:39 [“hoffman-bangii”] [type locality: Jäder, Västmanland, Sweden].

Scytonema hofmanii C. Agardh, 1817:117 [“hofmannii”].—Velasquez, 1950:325; 1962a:363, pl. 12: fig. 138—Drouet, 1973:91–92—Martinez, 1984:79.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon, MINDORO: Oriental Mindoro.

NOTE.—Scytonema hofmannii was proposed as a substitute name for S. hoffman-bangii. Both epithets commemorate the Danish botanist, Niels Hofman Bang (for whom the red alga Bangia is also named), so that neither is correctly spelled. When later starting points are abandoned, the correct name for this species becomes Scytonema hofman-bangii. The Quezon record was previously published under Calothrix pilosa (Velasquez, 1940:270).

Order STIGONEMATALES

Family MASTIGOCLADACEAE

Brachytrichia Zanardini

Brachytrichia maculans Gomont

Brachytrichia maculans Gomont, 1901:210, pl. 5: figs. 5–7 [type locality: Lem Dan, Ko Chang Archipelago, Thailand].—Velasquez, 1962a:356.—Martinez, 1984:28.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Drouet (1973:177, 194), after examining the holotype of this species (in PC), assigned it to Calothrix crustacea Thuret.

Brachytrichia quoyi (C. Agardh) Bornet and Flahault

Nostoc quoyi C. Agardh, 1824:22 [“quoji”] [type locality: Mariana Is.]

Brachytrichia quoyi (C. Agardh) Bornet and Flahault, 1886b [1886–1888]:373.—Velasquez, 1940:270; 1950:325; 1955:180, pl. V: fig 55.—Velasquez and Soriano, 1957:486.—Velasquez, 1962a:356, pl 11: fig. 129.—Villones and Magdamo, 1968:11, fig. 2.—Velasquez 1971:428, fig. 4.—Reyes, 1972:135.—Velasquez, Trono, and Doty, 1975:130.—Drouet, 1981:107.—Fortes, 1981b:396.—Cordero 1984a:69.—Marcos-Agngarayngay, 1983:65, fig. 3.—Cordero 1984b:57.—Martinez, 1984:28.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Ilocos Sur, Batangas, Quezon, Albay. MINDORO: Oriental Mindoro. NEGROS: Negros Oriental. PALAWAN (incl. Cuyo I.).

RECONCILIATION WITH THE DROUETIAN SYSTEM

Philippine Cyanophyceae arranged according to the Drouetian classification (synonyms indented beneath accepted names):

Order Chroococcales

Family Chamaesiphonaceae

Entophysalis conferta (Kützing) Drouet and Daily

Family Chroococcaceae

Anacystis aeruginosa (Zanardini) Drouet and Daily

Order Hormogonales

Family Nostocaceae

Anabaina oscillarioides Bory de Saint-Vincent

Anabaina pseudoscillatoria Bory de Saint-Vincent

Hormothamnium enteromorphoides Grunow

H. solutum Bornet and Grunow

Calothrix crustacea Thuret

Brachytrichia maculans Gomont

Calothrix aeruginea (Kützing) Thuret

C. confervicola (Dillwyn) C. Agardh

C. contarenii (Zanardini) Bornet and Flahault

C. robusta Setchell and Gardner

C. scopulorum (Weber and Mohr) C. Agardh

Gardnerula corymbosa (Harvey) De Toni f.

Rivularia bullata (Poiret) Berkeley

R. mesenterica (Kützing) Thuret

R. nitida C. Agardh

R. polyotis (J. Agardh) Hauck

Symploca hydnoides (Harvey) Kützing

Calothrix parietina (Nägeli ex Kützing) Thuret

Calothrix epiphytica W. West and G.S. West

C. viguieri Frémy

Dichothrix gypsophila (Kützing) Bornet and Flahault

Scytonema hofmannii C. Agardh [= S. hofman-bangii C. Agardh]

Calothrix pilosa Harvey

Family Oscillatoriaceae

Microcoleus lyngbyaceus (Kützing) P. Crouan and H. Crouan

Hydrocoleum lyngbyaceum Kützing

H. cantharidosmum (Montagne) Gomont

H. comoides (Harvey) Gomont

H. glutinosum (C. Agardh) Gomont

Lyngbya aestuarii (Mertens) Liebman

L. confervoides C. Agardh

L. ferruginea C. Agardh

L. majuscula (Dillwyn) Harvey

L. martensiana Meneghini

L. semiplena (C. Agardh) J. Agardh

Oscillatoria bonnemaisonii (P. Crouan and H. Crouan)

P. Crouan and H. Crouan

O. corallinae (Kützing) Gomont

O. curviceps C. Agardh

O. margaritifera (Kützing) Gomont

O. sancta (Kützing) Gomont

M. vaginatus (Vaucher) Gomont

Phormidium ambiguum Gomont

Oscillatoria erythrea (Ehrenberg) Drouet

Phormidium laysanense Lemmermann

O. lutea C. Agardh

Lyngbya lutea (C. Agardh) J.E. Areschoug

O. submembranacea (Ardissone and Strafforello) Drouet

Phormidium submembranaceum (Ardissone and Strafforello) Gomont

P. penicillatum Gomont

Porphyrosiphon kurzii (Zeller) Drouet

Sirocoleum kurzii (Zeller) Gomont

P. notarisii (Meneghini ex Kützing) Kützing

Oscillatoria nigroviridis Thwaites

Schizothrix arenaria (Berkeley) Gomont

Symploca laeteviridis Gomont

S. calcicola (C. Agardh) Gomont

Lyngbya infixa Frémy

L. mesotricha Skuja

Phormidium crosbyanum Tilden

P. persicinum (Reinke) Gomont

P. valderiae (Delponte) Schmidle

S. friesii (C. Agardh) Gomont

Phormidium tinctorium Kützing

S. mexicana Gomont

Lyngbya rosea W.R. Taylor

L. sordida Gomont [uncertain; see entry in Catalog]

Symploca howei Gardner

S. tenerrima (Gomont) Drouet

Microcoleus tenerrimus Gomont

M. acutissimus Gardner

Spirulina subsalsa Oersted

Spirulina major Kützing

Family Stigonemataceae

Brachytrichia quoyi (C. Agardh) Bornet and Flahault

Class RHODOPHYCEAE

Subclass BANGIOPHYCIDAE

Order GONIOTRICHALES

Family GONIOTRICHACEAE

Chroodactylon Hansgirg

Chroodactylon ornatum (C. Agardh) Basson

Conferva ornata C. Agardh, 1824:104 [type locality: Lake Mälaren, Stockholm, Sweden].

Asterocytis ornata (C. Agardh) Hamel, 1924:451.—Meñez and Calumpong, 1981:380 [“Asterocystis”].—Saraya and Trono, 1982:25.

Chroodactylon ornatum (C. Agardh) Basson, 1979:67.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CENTRAL VISAYAS.

Goniotrichum Kützing

Goniotrichum alsidii (Zanardini) Howe

Bangia alsidii Zanardini, 1840a:136 [type locality: Trieste, Italy].

Goniotrichum alsidii (Zanardini) Howe, 1914:75.—Meñez and Calumpong, 1981:380.

Bangia elegans Chauvin, 1842:33 [type locality: Arromanches-les-Bains, Calvados, France].

Goniotrichum elegans (Chauvin) Zanardini, 1847:249.—Weber-van Bosse, 1921:187.—Velasquez, Trono, and Doty, 1975:146.—Chan, 1981:387.—Saraya and Trono, 1982:25, pl. I: fig. 1.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CENTRAL VISAYAS. SULU: Tawitawi (Sangasiapu I.)

NOTE.—The conspecificity of Bangia alsidii and Goniotrichum elegans was proposed by Zanardini (1873:457), who, however, considered B. alsidii a variety of G. elegans. The varietal distinction is not usually recognized, but if it is, G. elegans must be subordinated to G. alsidii.

Order BANGIALES

Family ERYTHROPELTIDACEAE

Erythrocladia Rosenvinge

Erythrocladia irregularis Rosenvinge

Erythrocladia irregularis Rosenvinge, 1909:72, figs. 11, 12 [type locality: Møllegrund off Hirshals, Denmark].

Erythrocladia subintegra Rosenvinge, 1909:73, figs. 13, 14 [syntype localities: two, off Hirshals, Denmark].—Vannajan and Trono, 1978:15, fig. 14.—Saraya and Trono, 1982:26.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Manila.

NOTE.—The conspecificity of Erythrocladia irregularis and E. subintegra was proposed by Heerebout (1968:141). The latter species was treated as a forma of E. irregularis by Garbary, Hansen, and Scagel (1981:154).

Erythrocladia pinnata W.R. Taylor

Erythrocladia pinnata W.R. Taylor, 1942:75, pl. 2: figs. 1, 2 [type locality: Tobago I., West Indies].—Meñez and Calumpong, 1981:380.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Erythrotrichia J.E. Areschoug

Erythrotrichia bangioides Levring

Erythrotrichia bangioides Levring, 1955:410, fig. 1 [syntype localities: various, all in New Zealand].—Saraya and Trono, 1982:26.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Erythrotrichia biseriata Tanaka

Erythrotrichia biseriata Tanaka, 1944:86, fig. 8 [syntype localities: Kasho-to, Taiwan; Hachijo-jima, Japan].—Cordero, 1977a:33, figs. 1, 2.—Saraya and Trono, 1982:26.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas.

Erythrotrichia parietalis Tanaka

Erythrotrichia parietalis Tanaka, 1952:18, fig. 10 [type locality: Takamatsu, Japan].—Cordero, 1977a:34, fig. 3.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:380.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite (Corregidor I.), CENTRAL VISAYAS.

Family BANGIACEAE

Bangia Lyngbye

Bangia atropurpurea (Roth) C. Agardh

Conferva atropurpurea Roth, 1806:208, pl. VI [type locality: Bremen, West Germany].

Bangia atropurpurea (Roth) C. Agardh, 1824:76.

Conferva fuscopurpurea Dillwyn, 1807 [1802–1809]: pl. 92 [“fusco-purpurea”] [type locality: Dunraven Castle, Glamorganshire, Wales].

Bangia fuscopurpurea (Dillwyn) Lyngbye, 1819:83, pl. 24:C.—Meñez and Calumpong, 1981:380.—Calumpong, 1982:145.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental (Apo I.).

NOTE.—The conspecificity of Bangia fuscopurpurea (marine) and B. atropurpurea (freshwater) was proposed by den Hartog (1972) and Geesink (1973).

Bangia yamadae Tanaka

Bangia yamadae Tanaka, 1944:84, figs. 6, 7 [“yamadai”] [type locality: Hoko-to (P’eng-hu), Taiwan].—Cordero, 1977a:34, figs. 4, 5.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite (Corregidor I.).

NOTE.—In forming an epithet intended to commemorate a personal name ending in the letter a, the letter e is added regardless of gender. Thus Yamada yields yamadae and Okamura yields okamurae (used in several species in this catalog).

Porphyra C. Agardh

Porphyra atropurpurea (Olivi) De Toni

Ulva atropurpurea Olivi, 1794:153, pls. I—III [“atro-purpurea”] [type locality: Venezia, Italy].

Porphyra atropurpurea (Olivi) De Toni, 1897:17.—Zaneveld, 1956:24; 1959:105.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos.

Porphyra crispata Kjellman

Porphyra crispata Kjellman, 1897a:15, pl. 1: figs. 4, 5; pl. 3: figs. 5–7; pl. 5: fig. 15 [type locality: Goto, Japan].—Domantay, 1962:293.—Galutira and Velasquez, 1964:501, pl. 3: figs. 8a-c; pl. 7: figs. 27a,b.—Velasquez, 1968a:121, fig. 6.—De Leon and Domantay, 1971:5, 8.—Velasquez, 1971:446, fig. 25; 1972:63.—Cordero, 1974c:138, fig. 2.—Velasquez, Trono, and Doty, 1975:161—Cordero, 1977a:36, figs. 6–8; 1979a:21, fig. 1.—Velasquez, 1979b:230.—Cordero, 1980b:38, fig. 4.—Trono and Fortes, 1980:67.—Trono, Velasquez, and Guevarra, 1980:78.—Ganzon-Fortes, 1981:22—Trono and Fortes, 1982:149.—Cordero, 1984a:86.—Marcos-Agngarayngay, 1984a:17; 1984b:124.—Tungpalan, 1984:140.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Pangasinan, Batangas.

Porphyra denticulata Levring

Porphyra denticulata Levring, 1953:467, figs. 5, 6A-G [syntype localities: various, all in Queensland, Australia].—Quisumbing, 1951:1011.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Norte.

*Porphyra marcosii Cordero

Porphyra marcosii Cordero, 1976b:15, figs, A-H, pl. 1 [type locality: Dirique Bay, Burgos, Ilocos Norte Prov., Luzon].—Cordero, 1974c:139, fig. 4 [Porphyra sp.]; 1977a:36, figs. 9, 9a, pl. I:A; 1979a:22, fig. 2; 1980b:38, fig. 5, pl. 25; 1984a:87.—Marcos-Agngarayngay, 1984a:19; 1984b:124.—Tungpalan, 1984:140.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Porphyra suborbiculata Kjellman

Porphyra suborbiculata Kjellman, 1897a:10, pl. 1: figs. 1–3; pl. 2: figs. 5–9; pl. 5: figs. 4–7 [type locality: Goto, Japan].—Cordero, 1974c:138, fig. 3; 1977a:37, figs. 10, 11; 1979a:23, fig. 3; 1980b:39, fig. 6, pls. 26, [55].—Trono and Fortes, 1980:67.—Ganzon-Fortes, 1981:22.—Trono and Fortes, 1982:149.—Cordero, 1984a:86.—Marcos-Agngarayngay, 1984a:19, fig. 14; 1984b:124.—Tungpalan, 1984:140.

PHILIPPINE DISTRIBUTION. Norte, Cavite (Corregidor I.).

Porphyra variegata (Kjellman) Kjellman

Diploderma variegatum Kjellman, 1889:33, pl. II: figs. 1–4 [type locality: Bering I., USSR].

Porphyra variegata (Kjellman) Kjellman in Hus, 1900:69.—Velasquez et al., 1973:25, pl. 9: fig. 43.

PHILIPPINE DISTRIBUTION.—LUZON: Bataan.

Subclass FLORIDEOPHYCIDAE

Order ACROCHAETIALES

Family ACROCHAETIACEAE

Deliberations leading to the choice of the classification of acrochaetioid algae employed here are given in the appended Nomenclatural Notes.

Acrochaetium Nägeli

Acrochaetium gracile Børgesen

Acrochaetium gracile Børgesen, 1915:26, figs. 19, 20 [type locality: St. Thomas, Virgin Is.].—Abbott, 1962:115, fig. 17.—Velasquez, Trono, and Doty, 1975:126.—Cordero, 1977a:38, fig. 12.—Vannajan and Trono, 1978:15.—Chan, 1981:387. —Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite. PALAWAN (Balabac I.).

Acrochaetium hallandicum (Kylin) Hamel

Chantransia hallandica Kylin, 1906:123, fig. 8 [type locality: Halland coast of Sweden].

Acrochaetium hallandicum (Kylin) Hamel, 1927:20, 82.

Acrochaetium sargassi Børgesen, 1915:17, figs. 7–10 [type locality: St. Thomas, Virgin Is.].—Meñez and Calumpong, 1981:380.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

NOTE.—The conspecificity of Chantransia hallandica and Acrochaetium sargassi was proposed by Woelkerling (1973:84).

Acrochaetium hancockii (Dawson) Papenfuss

Rhodochorton hancockii Dawson, 1944:255, pl. 41: figs. 4–6 [type locality: Puerto Refugio, Isla Angel de la Guarda, Baja California Norte, Mexico].

Acrochaetium hancockii (Dawson) Papenfuss, 1945:306.—Cornejo and Velasquez, 1972:178, pl. 2: fig. 16; pl. 4: fig. 33.—Velasquez, Trono, and Doty, 1975:126.—Saraya and Trono, 1982:27, pl. I: fig. 2.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas.

Acrochaetium liagorae Børgesen

Acrochaetium liagorae Børgesen, 1915:57, figs. 60–62 [type locality: St. Croix, Virgin Is.].—Abbott, 1962:100, figs. 8a,d,j, 9.—Velasquez, Trono, and Doty, 1975:126.

PHILIPPINE DISTRIBUTION.—PALAWAN (Balabac I.), SULU: Sulu (Cagayan Sulu Is.).

*Acrochaetium nitidulum Abbott

Acrochaetium nitidulum Abbott, 1962:95, fig. 6 [type locality: Gnat Reef, Balabac I., Palawan Prov.].—Velasquez, Trono, and Doty, 1975:126.

PHILIPPINE DISTRIBUTION.—As above.

*Acrochaetium papenfussii Abbott

Acrochaetium papenfussii Abbott, 1962:97, fig. 7 [type locality: Cagayan Sulu I., Palawan Prov.].—Velasquez, Trono, and Doty, 1975:126.

PHILIPPINE DISTRIBUTION.—As above.

Acrochaetium robustum Børgesen

Acrochaetium robustum Børgesen, 1915:40, figs. 38–40 [type locality: St. Thomas, Virgin Is.].—Cordero, 1977a:38, figs. 13, 14.—Liao and Sotto, 1980:98.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. CEBU (Mactan I.).

Acrochaetium seriatum Børgesen

Acrochaetium seriatum Børgesen, 1915:32, figs. 25–28 [syntype localities: various, all in Virgin Is.].—Abbott, 1962:113, fig. 15.—Velasquez, Trono, and Doty, 1975:126.

PHILIPPINE DISTRIBUTION.—PALAWAN (Balabac I.).

Acrochaetium sinicola (Dawson) Papenfuss

Rhodochorton sinicola Dawson, 1944:256, pl. 41: figs. 1, 2 [type locality: Isla Turner, near Isla Tiburón, Sonora, Mexico].—Cornejo and Velasquez, 1972:177, pl. 2: fig. 15.—Velasquez, Trono, and Doty, 1975:161.

Acrochaetium sinicola (Dawson) Papenfuss, 1945:317 [“sinicolum”].

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

Acrochaetium trichogloeae Børgesen

Acrochaetium trichogloeae Børgesen, 1952:13, figs. 6, 7 [type locality: Barkly I., Mauritius].—Abbott, 1962:93, figs. 5b-i.—Velasquez, Trono, and Doty, 1975:126.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Cagayan Sulu Is.).

Acrochaetium tuticorinense Børgesen

Acrochaetium tuticorinense Børgesen, 1937:30, figs. 15, 16 [type locality: Tuticorin, India].—Abbott, 1962:114, fig. 16.—Velasquez, Trono, and Doty, 1975:126.

PHILIPPINE DISTRIBUTION.—PALAWAN (Balabac I.).

Order PALMARIALES

Family PALMARIACEAE

Palmaria Stackhouse

Palmaria palmata (Linnaeus) Kuntze

Fucus palmatus Linnaeus, 1753:1162 [type locality: “in Oceano”].

Palmaria palmata (Linnaeus) Kuntze, 1891:909.—Cordero, 1977a:173; 1978a:43.

PHILIPPINE DISTRIBUTION.—BATANES.

Order NEMALIALES

Family NEMALIACEAE

Trichogloea Kützing

Trichogloea requienii (Montagne) Kützing

Batrachospermum requienii Montagne, 1843b:355 [type locality: Red Sea].

Trichogloea requienii (Montagne) Kützing, 1847a:54.—Cordero, 1977a:48, fig. 26, pl. I:C; 1984a:87.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Quezon.

Family DERMONEMATACEAE

Dermonema Harvey ex Heydrich

Dermonema frappieri (Montagne and Millardet) Børgesen

Cladosiphon frappieri Montagne and Millardet, 1862:20, pl. XXVI: fig. 1 [type locality: Réunion].

Dermonema frappieri (Montagne and Millardet) Børgesen, 1942:42.—Cordero, 1977a:40.

PHILIPPINE DISTRIBUTION.—BATANES.

Liagoropsis Yamada

Liagoropsis schrammii (Børgesen) Doty and Abbott

Nemalion schrammii Børgesen, 1909:4, fig. 3, pl. I [“schrammi”] [type locality: St. Croix, Virgin Is.] [Helminthocladia schrammi Crouan freres in Mazé and Schramm, 1878:177, nomen subnudum].

Liagoropsis schrammii (Børgesen) Doty and Abbott, 1964:443 [including Philippine record, pages 445–448].—Trono, 1973a:129, fig. 14.—Velasquez, Trono, and Doty, 1975:152.

PHILIPPINE DISTRIBUTION.—LUZON: Albay, Sorsogon.

Yamadaella Abbott

*Yamadaella caenomyce (Decaisne) Abbott

Liagora caenomyce Decaisne, 1842:119 [type locality: Manila, Luzon].—Montagne, 1844a:659.—Martens, 1868:88–89.—J. Agardh, 1876:518.—Villones and Magdamo, 1968:29, fig. 32—Cornejo and Velasquez, 1972:178.—Reyes, 1972:151.—Velasquez et al., 1973:25, pl. 9: fig. 44.—Velasquez, Trono, and Doty, 1975:151.—Reyes, 1980:128, pl. 5: fig. 1.—Marcos-Agngarayngay, 1984b:125.

Yamadaella caenomyce (Decaisne) Abbott, 1970:117 [“cenomyce”] [including Philippine records].—Cordero, 1977a:48, fig. 16; 1979b:287; 1980b:43, fig. 7.—Meñez and Calumpong, 1981:380.—Hurtado-Ponce, 1983:127.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Batangas, Quezon, Albay. CATANDUANES. MINDORO: Oriental Mindoro. PANAY: Aklan. NEGROS: Negros Oriental. SIQUIJOR. MINDANAO (Agonoy I.).

Family HELMINTHOCLADIACEAE

Helminthocladia J. Agardh

Helminthocladia australis Harvey

Helminthocladia australis Harvey, 1863: pl. CCLXXII [type locality: Fremantle, Western Australia, Australia].—Cordero, 1977a:40, fig. 15.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan.

Liagora Lamouroux

Liagora boergesenii Yamada

Liagora boergesenii Yamada, 1938a:11, figs. 5, 6, pl. 2 [syntype localities: Yonakuni-jima, Ryukyu-retto, Japan; Kasho-to, Taiwan].—Reyes, 1972:151.—Velasquez, Trono, and Doty, 1975:151.—Cordero, 1977a:43, pl. II:A; 1980b:41.—Reyes, 1980:128, pl. 5: fig. 4.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Panagasinan, Quezon, CATANDUANES. NEGROS: Negros Oriental, SIQUIJOR. PALAWAN. SULU.

Liagora canariensis Børgesen

Liagora canariensis Børgesen, 1927:48, figs. 25–29 [syntype localities: Orotava and Santa Cruz, Isla Tenerife, Islas Canarias].—Cordero, 1977a:43, fig. 18, pl. II:B.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Liagora ceranoides Lamouroux

Liagora ceranoides Lamouroux, 1816:239 [type locality: St. Thomas, Virgin Is.].—Reyes, 1972:151.—Trono, 1973d:14, pl. 7: fig. 27.—Velasquez et al., 1973:25, pl. 9: fig. 46.—Velasquez, Trono, and Doty, 1975:151.—Cordero, 1976c:6; 1977a:44, figs. 17, 20, 21, pl. I:B; 1978a:13; 1979b:287.—Puig and Cordero, 1979:34 [“charoides”].—Cordero, 1980b:41, fig. 8.—Liao and Sotto, 1980:98.—Reyes, 1980:128, pl. 5: fig. 3.—Meñez and Calumpong, 1981:380.—Trono and De Lara, 1981:11.—Saraya and Trono, 1982:28.—Hurtado-Ponce, 1983:127.—Cordero, 1984a:87.—Hurtado-Ponce, 1984:180.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Batangas, Quezon, MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SAMAR: Eastern Samar. PANAY: Aklan. LEYTE (Biliran I.). NEGROS: Negros Oriental, CEBU (Mactan I.), SIQUIJOR.

Liagora ceranoides Lamouroux f. leprosa (J. Agardh) Yamada

Liagora leprosa J. Agardh, 1847:8 [type locality: Veracruz, Mexico].

Liagora ceranoides Lamouroux f. leprosa (J. Agardh) Yamada, 1938a:21.—Domantay, 1962:273.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Liagora divaricata Tseng

Liagora divaricata Tseng, 1941:268, figs. 2–4 [type locality: Tsinglan-Kang, Wenchang, Hainan, China].—Domantay, 1962:292.—Velasquez, Trono, and Doty, 1975:152.—Cordero, 1977a:45, fig. 19, pl.III:B; 1980b:42, fig. 9.—Saraya and Trono, 1982:29, pl. I: fig. 3.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas. MINDORO: Oriental Mindoro. SIQUIJOR.

Liagora farinosa Lamouroux

Liagora farinosa Lamouroux, 1816:240 [type locality: Suez, Egypt].—Reyes, 1972:151.—Cordero, 1973b:27.—Velasquez et al., 1973:25, pl. 9: fig. 45.—Westernhagen, 1973a:65.; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:152.—Cordero, 1976c:8; 1977a:45, figs. 23–25, pl. III:A; 1978a:14.—Trono and Tuason, 1978:13.—Cordero, 1979b:287; 1980b:43, fig. 10, pl. 27.—Reyes, 1980:128, pl. 5: fig. 2.—Trono and Fortes, 1980:69.—Ganzon-Fortes, 1981:22.—Trono and De Lara, 1981:11, pl. VII: fig. 4.—Trono and Fortes, 1982:149.—Saraya and Trono, 1982:27, pl. II: fig. 4.—Hurtado-Ponce, 1983:126.—Marcos-Agngarayngay, 1984a:19, fig. 15; 1984b:125.

Ganonema farinosum (Lamouroux) Fan and Wang, 1974:492 [“farinose”].—Meñez and Calumpong, 1981:380 [“farinose”].

Liagora cheyneana Harvey, 1855:552 [“cheyniana”] [type locality: Cape Riche, Western Australia, Australia].—Seale, 1911:309.—Wester, 1916:159; 1921:224; 1924:21.—G. Blanco, 1938:513.—Quisumbing, 1951:1011.—Montilla and Blanco, 1953:166.

Liagora farinosa Lamouroux var. cheyneana (Harvey) Zaneveld, 1956:45 [including Philippine records].—Zaneveld, 1959:105.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Pangasinan, La Union, Bataan, Batangas, Quezon, CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. LEYTE (incl. Biliran I.), PANAY: Aklan. NEGROS: Negros Oriental. CEBU (Mactan I.), SIQUIJOR. PALAWAN.

NOTE.—Liagora farinosa was placed in its own genus, Ganonema, by Fan and Wang (1974) on the basis of the carpogonial branch in that species, which was said to be borne on a special short filament rather than directly on an ordinary vegetative filament. Abbott (1984), pointing out the inconstancy of this character, returned the species to Liagora. Liagora cheyneana is included as a synonym on the authority of Howe (1920:554).

Liagora hawaiiana Butters

Liagora hawaiiana Butters, 1911:169, pl. XXIV: figs. 8, 9 [type locality: Laie Point, Koolauloa, Oahu, Hawaiian Is.].—Meñez, 1961:69, pl. 7: figs. 81, 82.—Velasquez, Trono, and Doty, 1975:152.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Liagora japonica Yamada

Liagora japonica Yamada, 1938a:16, figs. 9, 10, pl. 4: fig. 1 [syntype localities: various, all in Japan].—Domantay, 1962:292.—Velasquez, Trono, and Doty, 1975:152.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Liagora orientalis J. Agardh

Liagora orientalis J. Agardh, 1896:99 [type locality: Sri Lanka].—Cordero, 1977a:47.—Meñez and Calumpong, 1981:380.—Saraya and Trono, 1982:27.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CATANDUANES. CENTRAL VISAYAS.

Liagora robusta Yamada

Liagora robusta Yamada, 1938a:8, figs. 3, 4, pl. 12: fig. 1 [syntype localities: Chichi-jima and Haha-jima, Ogasawara-gunto (Bonin Is.), Japan].—Cordero, 1977a:47, fig. 20.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. LEYTE.

Liagora segawae Yamada

Liagora segawae Yamada, 1938a:18, figs. 11, 12, pl. 5, [“segawai”] [syntype localities: Chichi-jima, Ogasawara-gunto (Bonin Is.), and Okinawa-jima, Ryukyu-retto, Japan].—Cordero, 1973b:27.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. LEYTE (Biliran I.).

Liagora setchellii Yamada

Liagora setchellii Yamada, 1938a:13, figs. 7, 8, pl. 3: fig. 2 [syntype localities: Ogasawara-gunto (Bonin Is.) and Ryukyu-retto, Japan; Kasho-to, Taiwan].—Saraya and Trono, 1982:28, pl. 1: fig. 4.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Liagora tenuis J. Agardh

Liagora tenuis J. Agardh, 1896:101 [syntype localities: West Indies; Florida, USA].—Meñez and Calumpong, 1981:380.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Liagora valida Harvey

Liagora valida Harvey, 1853:138, pl. XXXI A [type locality: Sand Key, Florida, USA].—Velasquez et al., 1973:26, pl. 9: fig. 47.—Meñez and Calumpong, 1981:380.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. CENTRAL VISAYAS.

Order BONNEMAISONIALES

Family BONNEMAISONIACEAE

Asparagopsis Montagne

Asparagopsis taxiformis (Delile) Trevisan

Fucus taxiformis Delile, 1813:151, 295, pl. 57: fig. 2 [type locality: Alexandria, Egypt].

Asparagopsis taxiformis (Delile) Trevisan, 1845:45.—Velasquez, 1971:446, fig. 27.—Trono and Biña, 1973:2.—Cordero, 1976c:8; 1977a:69, pl. XI:B; 1978a:20; 1979b:276; 1980b:44.—Trono and Fortes, 1980:69.—Ganzon-Fortes, 1981:22.—Trono and De Lara, 1981:14, pl. IX: fig.2.—Calumpong, 1982:145.—Saraya and Trono, 1982:32, pl. III: fig. 3.—Trono and Fortes, 1982:149.—Cordero, 1984a:92.

Dasya delilei Montagne, 1841 [1839–1842]:166, pl. 8: fig. 6.

Asparagopsis delilei (Montagne) Montagne, 1841 [1839–1842]:xIv.—Montagne, 1844a:662.—Kützing, 1849:802.—J. Agardh, 1852 [1851–1863]:776.—Montagne, 1856:428.—Martens, 1868:98–99.—Dickie, 1874a:192.—Velasquez, Trono, and Doty, 1975:128.

Asparagopsis sanfordiana Harvey, 1855:544 [syntype localities: Garden I. and Rottnest I., Western Australia, Australia].—Quisumbing, 1951:1007.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan nan, Quezon, MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. PANAY: Aklan, Antique, NEGROS: Negros Oriental (incl. Apo I.), CEBU (Sumilon I.). PALAWAN.

NOTE.—The conspecificity of Asparagopsis sanfordiana and A. taxiformis was proposed by J. Feldmann and G. Feldmann (1943:81). Dasya delilei Montagne is a superfluous and hence illegitimate substitute name (or Fucus taxiformis Delile.

Family GALAXAURACEAE

An explanation of the replacement of Chaetangiaceae by Galaxauraceae is given in the appended Nomenclatural Notes.

Actinotrichia Decaisne

Actinotrichia fragilis (Forsskål) Børgesen

Fucus fragilis Forsskål, 1775:190 [type locality: Mokha, Yemen].

Actinotrichia fragilis (Forsskål) Børgesen, 1932:6.—Villones and Magdamo, 1968:28, fig. 24.—Reyes, 1972:152.—Trono and Biña, 1973:2.—Cordero, 1973b:28.—Westernhagen, 1973a:64.—Ortega, Alcala, and Reyes, 1974:186–188 (tables 2–4) [“fragilissima”].—Trono, 1974b:84.—Westernhagen, 1974:112 (table I).—Cordero, 1975:272, figs. 2, 3.—Velasquez, Trono, and Doty, 1975:126.—Cordero, 1977a:50, figs. 28, 202, pl. IV:A.—Trono and Young, 1977:59.—Trono, 1978:15.—Trono and Tuason, 1978:14.—Cordero, 1979b:287.—Puig and Cordero, 1979:35.—Liao and Sotto, 1980:98.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:380.—Trono and De Lara, 1981:13.—Saraya and Trono, 1982:31, pl. III: fig. 2.—Hurtado-Ponce, 1983:127.—Cordero, 1984a:88; 1984b:63; 1984c:54.—Marcos-Agngarayngay, 1984a:22, fig. 17.

Galaxaura rigida Lamouroux, 1816:265, pl. VIII: fig. 4 [type locality: “la mer des Indes”].

Actinotrichia rigida (Lamouroux) Decaisne, 1842:118.—Montagne, 1844a:659.—Dickie, 1874a:196.—Dickie, 1876a:244.—Velasquez, Trono, and Doty, 1975:126.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Norte, Pangasinan, Bataan, Cavite (Corregidor I.), Batangas, Quezon, Manila Bay. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. MASBATE. SAMAR: Western Samar, Eastern Samar. LEYTE (incl. Biliran I.), PANAY: Aklan. NEGROS: Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. S: Zamboanga, Davao. PALAWAN, SULU: Sulu (Siasi I.).

NOTE.—The synonymy was proposed by Børgesen (1932:6).

Galaxaura Lamouroux

Kjellman (1900), using anatomy as a primary criterion, recognized 62 species of Galaxaura, 47 of which were new. Certain sections and subsections contained only tetrasporophytes or only sexual plants. Efforts to match tetrasporophytes with sexual plants and otherwise to reduce the number of recognized species have been made by subsequent workers, notably Howe, Børgesen, Chou, and Papenfuss. The greatest reduction of species is to be found in the monograph by Papenfuss, Mshigeni, and Chiang (1982). Many taxonomic decisions incorporated in that paper do not agree with those of previous workers. In none of these investigations has static comparative morphology been supported by culture studies. In a recent paper, Magruder (1984) determined by culture studies that G. oblongata had a life cycle involving a small filamentous tetrasporophyte. Magruder’s discovery, while not affecting this catalog directly, emphasizes the need to broaden the scope of taxonomic study in Galaxaura. Meanwhile, the taxonomic scheme of Papenfuss, Mshigeni, and Chiang provides a useful framework.

Galaxaura apiculata Kjellman

Galaxaura apiculata Kjellman, 1900:74, pl. 12: figs. 13–26; pl. 20: fig. 36 [type locality: Ski, Japan].—Chou, 1945:51, pl. V: figs. 13–19; pl. IX: fig. 1.—Meñez, 1961:71, pl. 7: figs. 69–72.—Trono and Santiago, 1970:73. pl. III: figs. 3–7.—Cornejo and Velasquez, 1972:178.—Trono, 1974b:85.—Velasquez, Trono, and Doty, 1975:144.—Saraya and Trono, 1982:29, pl. II: fig. 2.

Galaxaura acuminata Kjellman ex Butters, 1911:180, pl. XXIV: figs. 17–19 [type locality: Waianae, Oahu, Hawaiian Is.].—Cordero, 1977a:52, fig. 31, pl. IX:A.—Puig and Cordero, 1979:36.—Cordero, 1984a:88.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Batangas. MINDORO: Oriental Mindoro. SAMAR: Eastern Samar. LEYTE (incl. Biliran I.), SULU: Sulu (Siasi I.).

NOTE.—The synonymy was proposed by Chou (1945:51).

Galaxaura arborea Kjellman

Galaxaura arborea Kjellman, 1900:72, pl. II: figs. 1–11; pl. 20: fig. 39 [type locality: Australia].—Chou, 1945:50, pl. V: figs. 1–5; pl. X: figs. 1, 2.—Meñez, 1961:70, pl. 7: figs. 76, 78.—Velasquez, Trono, and Doty, 1975:144.—Cordero, 1976c:8; 1977a:54, fig. 29, pl. X:A; 1978a:15.—Meñez and Calumpong, 1981:380.—Hurtado-Ponce, 1983:128.—Cordero, 1984a:88.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Dalupiri I.), Ilocos Norte, Pangasinan, MINDORO: Occidental Mindoro. CENTRAL VISAYAS.

Galaxaura contigua Kjellman

Galaxaura contigua Kjellman, 1900:78, pl. 17: figs. 1–14; pl. 20: fig. 23 [type locality: Hawaiian Is.].—Cordero, 1977a:54, fig. 36, pl. VIII:C; 1984a:89.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, SAMAR: Eastern Samar.

Galaxaura falcata Kjellman

Galaxaura falcata Kjellman, 1900:73, pl. II: figs. 12–21; pl. 12: figs. 1–4; pl. 20: fig. 33 [type locality: Enoshima, Kanagawa Prefecture, Japan].—Westernhagen, 1973a:65; 1974:112 (table I).—Cordero, 1977a:57, fig. 32, pl. X:B.—Hurtado-Ponce, 1983:128.—Cordero, 1984a:89.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, MINDORO: Oriental Mindoro. CEBU (Mactan I.).

Galaxaura fasciculata Kjellman

Galaxaura fasciculata Kjellman, 1900:53, pl. 5: figs. 1–9. pl. 20: fig. 14 [type locality: Celebes, Indonesia].—Chou, 1945:44, pl. II: fig. 2; pl. VIII: fig. 1.—Trono and Santiago, 1970:73, pl. III: fig. 2; pl. IV: fig. 1.—Reyes, 1972:1 52.—Ortega, Alcala, and Reyes, 1974:186, 188.—Velasquez, Trono, and Doty, 1975:144.—Cordero, 1976c:10; 1977a:57, fig. 33, pl. VI:B.—Cordero, 1978a:16.—Trono, 1978:15.—Trono and Tuason, 1978:14—Cordero, 1979b:288.—Liao and Sotto, 1980:98.—Meñez and Calumpong, 1981:380.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Manila, Quezon, CATANDUANES. MINDORO: Oriental Mindoro. MARINDUQUE. LEYTE. PANAY: Aklan. NEGROS: Negros Oriental, CEBU (incl. Mactan I.). MINDANAO: Davao. PALAWAN (incl. Culion I.).

Galaxaura filamentosa Chou

Galaxaura filamentosa Chou in W.R. Taylor, 1945:139 [type locality: Isla Clarión, Islas Revillagigedo, Mexico].—Cordero, 1976c:10; 1977a:58, fig. 35, pl. IV:B; 1978a:17; 1984a:90.

Galaxaura rudis [misapplied name].—Meñez and Calumpong, 1981:380.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Quezon, SAMAR: Eastern Samar. CENTRAL VISAYAS. SIQUIJOR.

NOTE.—Chou (1945:39) noted that certain records of Galaxaura rudis from China and Japan were representative of her new species, G. filamentosa. Cordero (1977a:58; 1978a:17), apparently misinterpreting Chou, listed G. rudis Kjellman (1900:43, pl. 2: figs. 1–9; pl. 20: fig. 11) as a taxonomic synonym of G. filamentosa. According to Papenfuss, Mshigeni, and Chiang (1982:407), G. rudis is a taxonomic synonym of G. lapidescens (Ellis and Solander) Lamouroux.

*Galaxaura kjellmanii Weber-van Bosse

Galaxaura kjellmanii Weber-van Bosse, 1921:217, fig. 66 [type locality: North Ubian I., Sulu Archipelago].—Cordero, 1973b:29.—Velasquez, Trono, and Doty, 1975:145.—Cordero, 1977a:60, fig. 38, pl. IX:C.—Puig and Cordero, 1979:36.—Cordero, 1984a:90.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Norte, LEYTE (Biliran I.), SULU: Sulu (North Ubian I.).

NOTE.—According to Puig and Cordero (1979:36), part or all of the treatment of Galaxaura elongata by Cordero (1977a) applies to G. kjellmanii.

Galaxaura marginata (Ellis and Solander) Lamouroux

Corallina marginata Ellis and Solander, 1786:115, pl. 22: fig. 6 [type locality: Bahama Is.].

Galaxaura marginata (Ellis and Solander) Lamouroux, 1816:264.—Papenfuss, Mshigeni, and Chiang, 1982:411, 415.

Galaxaura clavigera Kjellman, 1900:76, pl. 13: figs. 1–13; pl. 20: fig. 25 [type locality: Lasgori (Las Khoreh), Somalia].—Cordero, 1976c:9; 1978a:15.

Galaxaura tenera Kjellman, 1900:77, pl. 14: figs. 10–19; pl. 20: fig. 32 [type locality: Mombasa, Kenya].—Cordero, 1977a:67, fig. 45, pl. IX:B; 1984a:91.

Galaxaura veprecula Kjellman, 1900:80, pl. 16: figs. 17–33; pl. 20: fig. 32 [type locality: Malagasy Republic].—Chou, 1947:16, pl. VI: figs. 1–8; pl. XII: fig. 1.—Velasquez, Trono, and Doty, 1975:145.—Cordero, 1976c:8, 9; 1978a:19.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Dalupiri I.). MINDORO: Oriental Mindoro. MINDANAO: Surigao.

NOTE.—This synonymy was proposed by Papenfuss, Mshigeni, and Chiang (1982:41).

Galaxaura oblongata (Ellis and Solander) Lamouroux

Corallina oblongata Ellis and Solander, 1786:114, pl. 22: fig. 1 [type locality: West Indies].

Galaxaura oblongata (Ellis and Solander) Lamouroux, 1816:262.—Chou, 1947:7, pls. II, III, IX.—Villones and Magdamo, 1968:28, fig. 26.—Trono and Santiago, 1970:76, pl. I: figs. 1–12; pl. IV: figs. 4–6.—Velasquez, 1971:446, fig. 26.—Reyes, 1972:152.—Velasquez et al., 1973:26, pl. 10: fig. 48.—Westernhagen, 1973a:65.—Ortega, Alcala, and Reyes, 1974:187, 188.—Trono, 1974b:85.—Westernhagen, 1974:112 (table I).—Velasquez, Trono, and Doty, 1975:145.—Cordero, 1976c:9, 10; 1977a:60, fig. 34, pls. V:B, VII:B.—Trono and Young, 1977:59.—Trono, 1978:15.—Trono and Tuason, 1978:14.—Cordero, 1979b:276, 288.—Puig and Cordero, 1979:36.—Liao and Sotto, 1980:98.—Reyes, 1980:128, pl. 5: fig. 5.—Trono and Ganzon-Fortes, 1980:61, fig. [s.n.].—Meñez and Calumpong, 1981:380.—Trono and De Lara, 1981:12, pl. VIII: figs. 1, 3.—Montaño, Laserna, and Cajipe, 1982:39.—Papenfuss, Mshigeni, and Chiang, 1982:415, 418.—Saraya and Trono, 1982:29, pl. II: fig. 3.—Hurtado-Ponce, 1983:129.—Cordero, 1984a:90; 1984b:64.—Marcos-Agngarayngay, 1984a:24, fig. 18.

Corallina cylindrica Ellis and Solander, 1786:114, pl. 22: fig. 4 [type locality: West Indies].

Galaxaura cylindrica (Ellis and Solander) Lamouroux, 1821:22.—Chou, 1947:5, pl. 1; pl. VIII: fig. 1.—Velasquez et al., 1973:26, pl. 10: fig. 50.—Velasquez, Trono, and Doty, 1975:144.—Cordero, 1979b:276.

*Galaxaura fastigiata Decaisne, 1842:116 [lectotype locality: Manila, Luzon fide Svedelius, 1945:32.].—Montagne, 1844a:659.—Martens, 1868:86–87.—Dickie, 1874a:195.—Grunow, 1874:37.—Kjellman, 1900:64, pl. 9: figs. 1–3.—De Toni, 1924:126.—Howe, 1932:169.—Domantay, 1962:289.—De Leon and Domantay, 1971:5, 8.—Velasquez et al., 1973:26, pl. 10: fig. 49.—Cordero, 1973b:28.—Velasquez, Trono, and Doty, 1975:145.—Cordero, 1976c:8; 1978a:16.—Reyes, 1980:129, pl. 5: fig. 6.—Cordero, 1984a:89.

Galaxaura constipata Kjellman, 1900:63, pl. 8: figs. 29–33; pl. 20: fig. 5 [type locality: Veracruz, Mexico].—Meñez, 1961:70.—Velasquez, Trono, and Doty, 1975:144.

Galaxaura dimorpha Kjellman, 1900:63, pl. 8: figs. 23–28; pl. 20: fig. 3 [type locality: Timor, Indonesia].—Meñez, 1961:70, pl. 7: figs. 65–68, 73–75.—Velasquez, Trono, and Doty, 1975:144.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Ilocos Sur, Pangasinan, Zambales, Bataan, Batangas, Quezon, Albay, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. SAMAR: Northern Samar, Western Samar, Eastern Samar. LEYTE (incl. Biliran I.). PANAY: Aklan. NEGROS: Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Surigao, Misamis Oriental. PALAWAN, SULU: Sulu (Siasi I.).

NOTE.—The synonymy was proposed by Papenfuss, Mshigeni, and Chiang (1982:415). Svedelius (1945:32) selected Cuming 2241 as the lectotype collection of Galaxaura fastigiata. Kjellman (1900:65) stated that this specimen was from the Moluccas, an error that originated with Decaisne (1842:116), who wrote “Hab. in Moluccis (Manilla)”.

Galaxaura obtusata (Ellis and Solander) Lamouroux

Corallina obtusata Ellis and Solander, 1786:113, pl. 22: fig. 2 [type locality: Bahama Is.].

Galaxaura obtusata (Ellis and Solander) Lamouroux, 1816:262.—Trono and Santiago, 1970:74, pl. II; pl. IV: fig. 2.—Cordero, 1977a:62, fig. 37, pl. VIII:A,B; 1978a:17.—Vannajan and Trono, 1978:15, fig. 15.—Cordero, 1984a:90.

Tubularia umbellata Esper, [1805–1812]:125, pl. Tubularia XVII [type locality: probably West Indies; see Note].

Galaxaura umbellata (Esper) Lamouroux, 1816:262.—Chou, 1947:14, pl. V; pl. XI: fig. 1.—Trono and Santiago, 1970:75, pl. I: figs. 13–20.—Velasquez, Trono, and Doty, 1975:145.

Galaxaura robusta Kjellman, 1900:85, pl. 18: figs. 19–32; pl. 20: fig. 42 [type locality: Nosy-Bé, Malagasy Republic].—Cordero, 1978a:19.—Liao and Sotto, 1980:98.—Meñez and Calumpong, 1981:380.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Manila, Cavite, Batangas. MINDORO: Oriental Mindoro. CEBU (Mactan I.).

NOTE.—The synonymy was proposed by Papenfuss, Mshigeni, and Chiang (1982:418). For the provenance of Tubularia umbellata, Esper wrote: “gleichfalls in den ostindischen Gewässern.” Because “gleichfalls” (i.e., “likewise”) refers to T. obtusata, which is from the West Indies, it seems likely that “ostindischen” was an error for “westindischen.”

Galaxaura rugosa (Ellis and Solander) Lamouroux

Corallina rugosa Ellis and Solander, 1786:115, pl. 22: fig. 3 [type locality: Jamaica].

Galaxaura rugosa (Ellis and Solander) Lamouroux, 1816:263.—Chou, 1947:13, pl. IV: figs. 12, 13; pl. X: fig. 2.—Cordero, 1977a:63, fig. 39, pl. VI:A.

Galaxaura elongate J. Agardh, 1876:529 [syntype localities: Tonga; northeastern Australia].—Cordero, 1973b:28; 1976c:9, 10.; 1977a:54, figs. 30, 37, pl. V:A, pl. VI:C; 1978a:15.—Liao and Sotto, 1980:98.—Hurtado-Ponce, 1983:128.—Cordero, 1984a:89; 1984b:64.—Marcos-Agngarayngay, 1984a:24, fig. 19.

Galaxaura squalida Kjellman, 1900:55, pl. 6: figs. 1–12; pl. 20: fig. 9 [type locality: St. Croix, Virgin Is.].—Chou, 1947:9, pl. IV: figs. 1–11; pl. VIII: fig. 2.—Domantay, 1962:290.—Velasquez, Trono, and Doty, 1975:145.

Galaxaura glabriuscula Kjellman, 1900:56, pl. 7: figs. 1, 2; pl. 20: fig. 26 [type locality: Tahiti).—Puig and Cordero, 1979:36.

Galaxaura cuculligera Kjellman, 1900:58, pl. 6: figs. 22–30; pl. 20: fig. 30 [type locality: Goto, Japan].—Cordero, 1976c:9.

Galaxaura pacifica Tanaka, 1935:55, figs. 5, 6, pl. 17: fig. 2 [syntype localities: Haha-jima, Ogasawara-gunto [Bonin Is.], Japan; Garan-bi, Taiwan].—Cordero, 1976c:8, 9; 1977a:63; 1978a:18.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Pangasinan. MINDORO: Occidental Mindoro. SAMAR: Eastern Samar. LEYTE (Biliran I.). CEBU (incl. Mactan I.). PALAWAN.

NOTE.—The synonymy was proposed by Papenfuss, Mshigeni, and Chiang (1982:421–422). According to Puig and Cordero (1979:36), part or all of the treatment of Galaxaura elongata by Cordero (1977a) applies to G. kjellmanii

Galaxaura striata Kjellman

Galaxaura striata Kjellman, 1900:66, pl. 9: figs. 18–38; pl. 20: fig. 7 [type locality: Marquesas Is.].—Cordero, 1977a:64, figs. 40–42, pl. IV:C.

PHILIPPINE DISTRIBUTION.—BATANES.

Galaxaura subfruticulosa Chou

Galaxaura subfruticulosa Chou in W.R. Taylor, 1945:140 [type locality: Isla Clarión, Islas Revillagigedo, Mexico].—Domantay, 1962:290.—Cordero, 1973b:29; 1976c:9; 1977a:65, fig. 44, pl. V:C; 1978a:18; 1979b:289.—Puig and Cordero, 1979:36.—Meñez and Calumpong, 1981:380.—Saraya and Trono, 1982:30, pl. III: fig. 1.—Marcos-Agngarayngay, 1984a:25, fig. 20.

Galaxaura fruticulosa Kjellman, 1900:51, pl. 4: figs. 4–16; pl. 20: fig. 19 [type locality: Nomo-zaki, Nagasaki Prefecture, Japan] [intended nomenclatural synonym].—Velasquez, Trono, and Doty, 1975:145.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Quezon, SAMAR: Western Samar, Eastern Samar. LEYTE (Biliran I.), PANAY: Aklan.

NOTE.—Chou intended to propose a new name for Galaxaura fruticulosa Kjellman, a later homonym of G. fruticulosa (Ellis and Solander) Lamouroux (1816:264), but designated a new type and thus effectively published a new species. Whether G. fruticulosa and G. subfruticulosa are conspecific remains to be determined.

Galaxaura subverticillata Kjellman

Galaxaura subverticillata Kjellman, 1900:48, pl. 3: figs. 12–14; pl. 20: fig. 17 [type locality: St. Croix, Virgin Is.].—Chou, 1945:45, pl. II: fig. 1; pl. VIII: fig. 2—Trono and Santiago, 1970:74, pl. III: fig. 1; pl. IV: fig. 3.—Velasquez, Trono, and Doty, 1975:145.—Cordero, 1977a:65, fig. 47, pl. VII:A.—Puig and Cordero, 1979:37.—Trono and De Lara, 1981:13, pl. VIII: fig. 2.—Saraya and Trono, 1982:30, pl. II: fig. 1.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SAMAR: Eastern Samar. LEYTE (Biliran I.).

Scinaia Bivona-Bernardi

Scinaia hormoides Setchell

Scinaia hormoides Setchell, 1914:106, 125, pl. 12: figs. 33–35; pl. 13: figs. 36, 37 [including Philippine record] [type locality: Haleiwa, Oahu, Hawaiian Is.].—De Toni, 1924:101.—Velasquez, Trono, and Doty, 1975:163.—Trono and Fortes, 1980:69.—Saraya and Trono, 1982:31.

PHILIPPINE DISTRIBUTION.—LUZON: La Union, Pangasinan.

Scinaia latifrons Howe

Scinaia latifrons Howe, 1911:500, fig. 1, pl. 28 [type locality: La Paz, Baja California Sur, Mexico].—Vannajan and Trono, 1978:16.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

Scinaia moniliformis J. Agardh

Scinaia moniliformis J. Agardh, 1885:72 [type locality: Port Phillip, Victoria, Australia].—Cordero, 1977a:68, fig. 46, pl. XI:A; 1980b:44, pl. 28; 1982a:61, fig. 9; 1984a:91.—Marcos-Agngarayngay, 1984a:21, fig. 16.—Tungpalan, 1984:140.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Cagayan, Quezon.

Order GELIDIALES

Family GELIDIACEAE

Beckerella Kylin

*Beckerella scalaramosa Kraft

Beckerella scalaramosa Kraft, 1976:85, figs. 1–15 [type locality: Bulusan, Sorsogon Prov., Luzon].

PHILIPPINE DISTRIBUTION.—As above.

NOTE.—The type collection of Beckerella scalaramosa had previously been recorded as Beckerella [“Becherella”] sp. by Trono (1973a:130, figs. 18, 19).

Gelidiella Feldmann and Hamel

Gelidiella acerosa (Forsskål) Feldmann and Hamel

Fucus acerosus Forsskål, 1775:190 [type locality: Mokha, Yemen].

Gelidiella acerosa (Forsskål) Feldmann and Hamel, 1934:533.—Domantay, 1962:290.—Galutira and Velasquez, 1964:502, pl. 3: figs. 9a,b; pl. 7: fig. 28.—Velasquez, 1971:446, fig. 28.—Cornejo and Velasquez, 1972:179.—Reyes, 1972:152.—Trono, 1972a:103.—Velasquez, 1972:63.—Cordero, 1973b:29.—Trono, 1973d:15, pl. 7: fig. 26.—Trono and Biña, 1973:3.—Velasquez et al., 1973:27.—Westernhagen, 1973a:65.—De Leon, 1974:31, 33, photo [s.n.], fig. [s.n.].—Ortega, Alcala, and Reyes, 1974:187, 188.—Westernhagen, 1974:112 (table I).—Velasquez, Trono, and Doty, 1975:145.—Cordero, 1976c:9, 10; 1977a:70; 1978a:20.—Tahil, 1978:52.—Cordero, 1979b:276.—Garcia, 1979:45 (table 1).—Puig and Cordero, 1979:37.—Velasquez, 1979b:230.—Cordero, 1980b:45, pl. 29.—Fortes and Trono. 1980:59.—Liao and Sotto, 1980:98.—Reyes, 1980:129, pl. 6: fig. 1.—Trono and Fortes, 1980:70.—Trono and Ganzon-Fortes, 1980:63, fig. [s.n.].—Velasquez, 1980:127.—Chan, 1981:387.—Ganzon-Fortes, 1981:22.—Guzman, 1981:42, 45, 50.—Laserna et al., 1981:443.—Meñez and Calumpong, 1981:380.—Trono and De Lara, 1981:14, pl. IX: fig. 4.—Calumpong, 1982:145.—Cordero, 1982a:60, 61, fig. 12.—Saraya and Trono, 1982:32.—Trono and Fortes, 1982:150.—Hurtado-Ponce, 1983:129.—Cordero, 1984a:92; 1984c:54.—Hurtado-Ponce, 1984:180.—Marcos-Agngarayngay, 1984a:27, fig. 21 (captions for figs. 21 and 22 interchanged); 1984b:125.—Tungpalan, 1984:140.—Trono and Ganzon-Fortes, 1985:67.

Fucus rigidus Vahl, 1802:46 [type locality: St. Croix, Virgin Is.].

Sphaerococcus rigidus C. Agardh, 1822a:285.

Gelidium rigidum (C. Agardh) Greville, 1830:lvii.—Martens, 1868:92–93.—Dickie, 1876a:243, 244.—Howe, 1932:169.—Zaneveld, 1956:34.—Velasquez, Trono, and Doty, 1975:145.

Gelidiopsis rigida (C. Agardh) Weber-van Bosse, 1904a:104 [“rigidum”].—Zaneveld, 1959:116.—Cordero, 1976c:9, 10; 1977a:122; 1978a:22.

Fucus spiniformis Lamouroux, 1805:77, pl. XXXVI: figs. 3, 4 [“spinaeformis”] [syntype localities: Malagasy Republic; Mauritius].

Gelidium spiniforme (Lamouroux) Lamouroux, 1813:129.—Montagne, 1844a:662.—Velasquez, Trono, and Doty, 1975:145.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Bataan, Batangas, Quezon, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MASBATE. SAMAR: Western Samar, Eastern Samar. LEYTE (Biliran I.), PANAY: Aklan, Capiz. GUIMARAS. NEGROS: Negros Occidental (incl. Ilacaon I., Suyac I.), Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Surigao del Sur, Zamboanga. PALAWAN, SULU: Tawitawi.

NOTE.—Fucus rigidus Vahl, the intended basionym of Gelidium rigidum (and other combinations), is a later homonym of F. rigidus Turra (1780:68) and hence not priorable. Sphaerococcus rigidus C. Agardh is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN. Fucus spiniformis is included as a synonym on the authority of C. Agardh (1822a:285), Sphaerococcus rigidus on the authority of Børgesen (1932:5).

Gelidiella adnata Dawson

Gelidiella adnata Dawson, 1954b:4 [type locality: Nha Trang, Vietnam].—Fortes and Trono, 1980:58.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Gelidiella taylorii Joly

Gelidiella taylorii Joly, 1957:102, fig. 1, pl. IX: figs. 5, 5a; pl. XII: fig. 3 [“taylori”] [type locality: São Vicinte, São Paulo, Brazil].—Westernhagen, 1973a:65.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Gelidium Lamouroux

Gelidium amansii (Lamouroux) Lamouroux

Fucus amansii Lamouroux, 1805:48, pl. XXVI: figs. 2–5 [syntype localities: Malagasy Republic; Mauritius].

Gelidium amansii (Lamouroux) Lamouroux, 1813:129.—Martens, 1868:94–95.—Zaneveld, 1956:34; 1959:115.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Gelidium capense (S.G. Gmelin) P.C. Silva, new combination

Fucus capensis S.G. Gmelin, 1768:157, pl. XVII: fig. 1 [type locality: Cape of Good Hope, South Africa].

Gelidium cartilagineum [misapplied name fide Dixon, 1967].—J. Agardh, 1852 [1851–1863]:473.—De Toni, 1897:152

PHILIPPINE DISTRIBUTION.—Locality not specified.

NOTE.—This species, which is common at wave-swept sites on cold-water shores of South Africa, probably does not occur in the Philippines. Dixon (1967) showed that Fucus cartilagineus Linnaeus (1753:1161), a name traditionally applied to this species, should be typified with a specimen referable to Plocamium. Fucus capensis and F. versicolor S.G. Gmelin (1768:158, pl. XVII: fig. 2) have long been considered conspecific with the alga traditionally called Gelidium cartilagineum (Linnaeus) Gaillon (see Turner, 1809:138 and J. Agardh 1852 [1851–1863]:473). With the removal of Fucus cartilagineus from consideration, Dixon (1967:58) adopted Gelidium versicolor (S.G. Gmelin) Lamouroux (1813:129) as the correct name for the South African alga. Fucus versicolor, however, is a superfluous name for F. abrotanifolius Linnaeus (1753:1161), which Gmelin cited as a synomym, and hence is illegitimate. (Fucus abrotanifolius is referable to Cystoseira.) The earliest available basionym for the South African alga thus appears to be Fucus capensis S.G. Gmelin.

Gelidium coulteri Harvey

Gelidium coulteri Harvey, 1853:117 [type locality: Monterey Peninsula, California, USA].—Olea and De Leon, 1956:104.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Gelidium crinale (Turner) Gaillon

Fucus crinalis Turner, 1815 [1815–1819]:4, pl. 198: figs, a-c, e-g [syntype localities: England; Northern Ireland].

Gelidium crinale (Turner) Gaillon, 1828:362.—Trono and Fortes, 1980:70.—Fortes, 1981b:396.—Ganzon-Fortes, 1981:22.—Calumpong, 1982:145.—Trono and Fortes, 1982:150.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Occidental (Ilacaon I.), Negros Oriental (incl. Apo I.), CEBU (incl. Pescador I.).

Gelidium crinale (Turner) Gaillon var. perpusillum Piccone et Grunow

Gelidium crinale (Turner) Gaillon var. perpusillum Piccone et Grunow in Piccone, 1884a:317 [“perpusilla”] [type locality: Massawa, Ethiopia].—Trono, 1973a:131.—Cordero, 1977a:72, fig. 49, pl. XXV:C—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. CENTRAL VISAYAS.

Gelidium divaricatum Martens

Gelidium divaricatum Martens, 1868:30, pl. VIII: fig. 4 [type locality: Hong Kong].—Cordero, 1977a:73; 1978a:23.

PHILIPPINE DISTRIBUTION.—BATANES.

Gelidium isabelae W.R. Taylor

Gelidium isabelae W.R. Taylor, 1945:154, pl. 5: figs. 8–12 [type locality: Isla Isabela, Galápagos].—Cordero, 1977a:73, fig. 50; 1978a:23.

PHILIPPINE DISTRIBUTION.—BATANES

Gelidium kintaroi Yamada

Gelidium clavatum Okamura, 1934:61, pl. 28; pl. 32: figs. 4–6 [syntype localities: Hoko-to (P’eng-hu), Taiwan; Amoy, China] [replaced name].—Cordero, 1977a:72, fig. 48.

Gelidium kintaroi Yamada, 1941a:201.

PHILIPPINE DISTRIBUTION.—CATANDUANES. MINDORO: Oriental Mindoro.

NOTE.—Gelidium kintaroi is a substitute name for G. clavatum Okamura, a later homonym of G. clavatum (Lamouroux) Lamouroux (1813:129).

Gelidium pulchellum (Turner) Kützing

Fucus corneus Hudson var. pulchellus Turner, 1819 [1815–1819]:146, pl. 257: fig. p [syntype localities: Bantry Bay, Eire; Sidmouth and Isle of Portland, England].

Gelidium pulchellum (Turner) Kützing, 1868:18.—Domantay, 1962:290.—De Leon and Domantay, 1971:5, 10.—Velasquez, Trono, and Doty, 1975:145.—Trono and Fortes, 1980:70.—Ganzon-Fortes, 1981:22.—Trono and Fortes, 1982:150.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas.

Gelidium pusillum (Stackhouse) Le Jolis

Fucus pusillus Stackhouse, 1795 [1795–1801]:16, pl. vi [type locality: Sidmouth, Devonshire, England].

Acrocarpus pusillus (Stackhouse) Kützing, 1849:762.—Martens, 1868:30, 92–93.—Velasquez, Trono, and Doty, 1975:126.

Gelidium pusillum (Stackhouse) Le Jolis, 1863:139.—Trono, 1973d:15.—Cordero, 1977a:74, fig. 51.—Vannajan and Trono, 1978:16.—Reyes, 1980:130, pl. 6: fig. 5.—Trono and Fortes, 1980:70.—Fortes, 1981b:396.—Ganzon-Fortes, 1981:22.—Meñez and Calumpong, 1981:381.—Calumpong, 1982:145.—Saraya and Trono, 1982:32.—Trono and Fortes, 1982:150.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite. MINDORO: Oriental Mindoro. NEGROS: Negros Occidental, Negros Oriental (incl. Apo I.), SIQUIJOR. MINDANAO: Zamboanga. PALAWAN.

Gelidium pusillum (Stackhouse) Le Jolis var. pacificum W.R. Taylor

Gelidium pusillum (Stackhouse) Le Jolis var. pacificum W.R. Taylor, 1945:153, pl. 5: fig. 7; pl. 33: fig. 1 [type locality: Isla Santa Maria, Galápagos].—Cordero, 1977a:74.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Gelidium rigens (C. Agardh) Greville ex Kützing

Sphaerococcus rigens C. Agardh, 1822a:332 [type locality: Sea of Japan].

Gelidium rigens (C. Agardh) Greville ex Kützing, 1849:767.—Martens, 1868:94–95.—Velasquez, Trono, and Doty, 1975:145.—Trono and Fortes, 1980:70.—Ganzon-Fortes, 1981:22.—Trono and Fortes, 1982:150.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Pterocladia J. Agardh

Pterocladia caloglossoides (Howe) Dawson

Gelidium caloglossoides Howe, 1914:96, pl. 34: fig. 7; pl. 35 [type locality: Isla San Lorenzo, Peru].

Pterocladia caloglossoides (Howe) Dawson, 1953:76, pl. 6: fig. 1.

Pterocladia parva Dawson, 1953:77, pl. 6: fig. 2 [type locality: San Felipe, Baja California Norte, Mexico).—Meñez and Calumpong, 1981:381.—Calumpong, 1982:147.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental (incl. Apo I.), CEBU (Sumilon I.).

NOTE.—The conspecificity of Pterocladia parva and P. caloglossoides was proposed by Stewart and Norris (1981:281). The generic position of P. caloglossoides, however, is uncertain. Cystocarpic plants have not been reported from the type locality (Peru). On the basis of cystocarpic material from Mexico, Dawson transferred the species to Pterocladia, but abandoned this stance when treating the seaweeds of Peru (Dawson, Acleto, and Foldvik, 1964:37).

Pterocladia capillacea (S.G. Gmelin) Bornet

Fucus capillaceus S.G. Gmelin, 1768:146, pl. XV: fig. 1 [type locality: Mediterranean Sea].

Pterocladia capillacea (S.G. Gmelin) Bornet in Bornet and Thuret, 1876:57, pl. XX: figs. 1–7.—Cordero, 1977a:75.—Hurtado-Ponce, 1983:130.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Pterocladia densa Okamura

Pterocladia densa Okamura, 1934:63, pl. 30: figs. 1, 2; pl. 33: figs. 4–8 [syntype localities: various, all in Japan].—Hurtado-Ponce, 1983:130.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Pterocladia nana Okamura

Pterocladia nana Okamura, 1931 [1929–1932]:53, pl. CCLXXVIII: figs. 1–14 [syntype localities: various, all in Japan].—Cordero, 1977a:75, fig. 52.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan.

Order CRYPTONEMIALES

Family ENDOCLADIACEAE

Gloiopeltis J. Agardh

Gloiopeltis complanata (Harvey) Yamada

Endocladia complanata Harvey, 1860a:333 [type locality: Shimoda, Shizuoka Prefecture, Japan.

Goiopeltis complanata (Harvey) Yamada, 1932a:117.—Cordero, 1977a:114.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Gloiopeltis tenax (Turner) Decaisne

Fucus tenax Turner, 1806:376, pl. 13 [type locality: China Sea].

Gloiopeltis tenax (Turner) Decaisne, 1842:360.—De Leon, 1974:31, 32, photo [s.n.].—Cordero, 1977a:114, fig. 90.

PHILIPPINE DISTRIBUTION.—PALAWAN.

NOTE.—This species is traditionally accredited to (Turner) J. Agardh, the combination supposedly having been made at the time the generic name was proposed (J. Agardh, 1842:68, footnote). Agardh, however, merely referred to “Typo Fuco tenaci Turn. tab. 125,” and the combination was not made until a few months later, by Decaisne.

Family PEYSSONNELIACEAE

Peyssonnelia Decaisne

Peyssonnelia calcea Heydrich

Peyssonnelia calcea Heydrich, 1897a:10 [type locality: Tami I., Papua New Guinea].—Domantay, 1962:293.—Velasquez, Trono, and Doty, 1975:159.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Peyssonnelia conchicola Piccone and Grunow

Peyssonnelia conchicola Piccone and Grunow in Piccone, 1884a:317, pl. VII: figs. 5–8 [type locality: Massawa, Ethiopia].—Weber-van Bosse, 1921:272.—Velasquez, Trono, and Doty, 1975:159.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I.).

Peyssonnelia distenta (Harvey) Yamada

Galaxaura distenta Harvey, 1860a:331 [type locality: O-shima, Amami-gunto, Japan].

Peyssonnelia distenta (Harvey) Yamada, 1930b:30, pl. VI: fig. 7.—Cordero, 1976c:9; 1977a:78, figs. 57, 58, pl. XIII:A; 1978a:24; 1984a:93.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte.

*Peyssonnelia evae Weber-van Bosse

Peyssonnelia evae Weber-van Bosse, 1921:279, fig. 95 [type locality: Pearl Bank, Tawitawi Prov., Sulu Archipelago].—De Toni, 1924:593.—Velasquez, Trono, and Doty, 1975:159.

PHILIPPINE DISTRIBUTION.—As above.

*Peyssonnelia foveolata (Weber-van Bosse) Denizot

Cruoriella foveolata Weber-van Bosse, 1921:294, figs. 105, 106 [syntype localities: Muaras Reef, East Kalimantan, Indonesia; North Ubian I. and Pearl Bank, Sulu Archipelago].—De Toni, 1924:588.—Velasquez, Trono, and Doty, 1975:138.

Peyssonnelia foveolata (Weber-van Bosse) Denizot, 1968:113, 310.

PHILIPPINE DISTRIBUTION.—As above.

*Peyssonnelia indica (Weber-van Bosse) Denizot

Cruoriella indica Weber-van Bosse, 1921:283, figs. 97 [syntype localities: various in Indonesia; Tongquil I. and Capual I., Sulu Prov., Sulu Archipelago].—Velasquez, Trono, and Doty, 1975:138.

Peyssonnelia indica (Weber-van Bosse) Denizot, 1968:123, 310.

PHILIPPINE DISTRIBUTION.—As above.

*Peyssonnelia luzonensis Cordero

Peyssonnelia luzonensis Cordero, 1977a:80, fig. 61, pl. XII:B [type locality:Quezon I., Hundred Is., Pangasinan Prov., Luzon].

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan.

*Peyssonnelia mariti (Weber-van Bosse) Denizot

Cruoriella mariti Weber-van Bosse, 1921:288, fig. 101 [syntype localities:Sebangkaitan I., Little Paternoster Is. and Flores, Indonesia; Tongquil I., Sulu Prov., Sulu Archipelago].—De Toni, 1924:587.—Velasquez, Trono, and Doty, 1975:138.

Peyssonnelia mariti (Weber-van Bosse) Denizot, 1968:122, 310.

PHILIPPINE DISTRIBUTION.—As above.

*Peyssonnelia obscura Weber-van Bosse

Peyssonnelia obscura Weber-van Bosse, 1921:274, fig. 92 [syntype localities: various in Indonesia; Capual I. and North Ubian I., Sulu Prov., Sulu Archipelago.—De Toni, 1924:592.—Velasquez, Trono, and Doty, 1975:159.

PHILIPPINE DISTRIBUTION.—As above.

Peyssonnelia rubra (Greville) J. Agardh

Zonaria rubra Greville, 1827:340, pl. 3: fig. 3 [type locality: Ionian Islands, Greece].

Peyssonnelia rubra (Greville) J. Agardh, 1851 [1851–1863]:502.—Dickie, 1876a:245.—Velasquez, Trono, and Doty, 1975:159.—Liao and Sotto, 1980:98.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.). MINDANAO: Zamboanga.

*Peyssonnelia rubra (Greville) J. Agardh f. orientalis Weber-van Bosse

Peyssonnelia rubra (Greville) J. Agardh f. orientalis Weber-van Bosse, 1921:270, 272, fig. 89 [syntype localities: various in Indonesia; North Ubian I., Sulu Prov., Sulu Archipelago].—Cordero, 1976c:10 [var. orientalis]; 1977a:78, fig. 56, pl. XIII:C [var. orientalis].—Saraya and Trono, 1982:34, pl. IV: fig. 2 [var. orientalis].

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. PALAWAN, SULU: Sulu (North Ubian I.).

Peyssonnelia squamaria (S.G. Gmelin) Decaisne

Fucus squamarius S.G. Gmelin, 1768:171, pl. XX: fig. 1 [type locality: Mediterranean Sea].

Peyssonnelia squamaria (S.G. Gmelin) Decaisne, 1842:360.—Cordero, 1977a:80; 1978a:25.—Marcos-Agngarayngay, 1984a:30, fig. 24.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte.

Polystrata Heydrich

Polystrata dura Heydrich

Polystrata dura Heydrich, 1905:35, pl. I [type locality: Tami I., Papua New Guinea].

Cruoriella dura (Heydrich) Weber-van Bosse, 1921:293, pl. VIII: fig. 3 [including Philippine record].—De Toni, 1924:585.—Velasquez, Trono, and Doty, 1975:138.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Tongquil I.).

Family CRYPTONEMIACEAE

Carpopeltis Schmitz

Carpopeltis affinis (Harvey) Okamura

Gigartina affinis Harvey, 1860a:332 [type locality: Hakodate, Hokkaido, Japan].

Carpopeltis affinis (Harvey) Okamura, 1934 [1933–1942]:30, pl. 316: figs. 4–11.—Cordero, 1976c:8; 1977a:100, fig. 79; 1978a:31.—Hurtado-Ponce, 1983:135; 1984:180.—Marcos-Agngarayngay, 1984a:37, fig. 30

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, SIQUIJOR. PALAWAN.

Carpopeltis angusta (Harvey) Okamura

Gymnogongrus ligulatus Harvey ex Kützing var. angustus Harvey, 1860a:332 [type locality: Shimoda, Shizuoka Prefecture, Japan].

Carpopeltis angusta (Harvey) Okamura, 1909 [1909–1912]:66, pl. LXVII—Cordero, 1976c:8, 10; 1977a:100, fig. 78, pl. XVI:A; 1978a:32.—Hurtado-Ponce, 1983:135.—Marcos-Agngarayngay, 1984a:39, fig. 31.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte.

Carpopeltis articulata (Okamura) Okamura

Prionitis articulata Okamura, 1899a:5, pl. 1: figs. 3, 4 [syntype localities: various, all in Japan].

Carpopeltis articulata (Okamura) Okamura, 1909 [1909–1912]:70, pl. LXVIII—Cordero, 1977a:101, pl. XV:B.—Hurtado-Ponce, 1983:135.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, MINDORO: Oriental Mindoro. PALAWAN.

Carpopeltis capitellata (Sonder) Schmitz

Cryptonemia capitellata Sonder, 1871:62 [type locality: Cape York, Queensland, Australia].

Carpopeltis capitellata (Sonder) Schmitz in Schmitz and Hauptfleisch, 1897 [1896–1897]:514.—Weber-van Bosse, 1921:246.—Velasquez, Trono, and Doty, 1975:130.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

Carpopeltis crispata Okamura

Carpopeltis crispata Okamura, 1934 [1933–1942]:32, pl. 317: figs. 6–11 [syntype localities: various, all in Japan].—Cordero, 1976c:8, 10; 1977a:101; 1978a:32 [C. crispata prox.]; 1984b:65.—Tungpalan, 1984:140.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte. PALAWAN.

Carpopeltis divaricata Okamura

Carpopeltis divaricata Okamura, 1934 [1933–1942]:31, pl. 317: figs. 1–5 [syntype localities: various, all in Japan].—Cordero, 1977a:103, fig. 77, pl. XV:C; 1978a:32.—Hurtado-Ponce, 1983:137.—Cordero, 1984a:96.—Tungpalan, 1984:140.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, MINDORO: Oriental Mindoro. SAMAR: Eastern Samar.

Carpopeltis formosana Okamura

Carpopeltis formosana Okamura, 1931:110, pl. 12 [type locality: Kotosho (Hung-t’ou), Taiwan].—Cordero, 1976c:6, 8, 9, 10; 1977a:104, fig. 80, pl. XV:A; 1978a:33.

PHILIPPINE DISTRIBUTION.—BATANES.

Carpopeltis maillardii (Montagne and Millardet) Chiang

Phyllophora maillardii Montagne and Millardet, 1862:8, pl. XXIV [type locality: Réunion].

Carpopeltis maillardii (Montagne and Millardet) Chiang, 1970:68, pl. 5c—Reyes, 1980:132, pl. 8: fig. 2.

Cryptonemia rigida Harvey ex J. Agardh, 1876:163 [type locality: Sri Lanka].

Carpopeltis rigida (Harvey ex J. Agardh) Schmitz, 1895:168.—Cordero, 1976c:9; 1978a:33.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—BATANES. SIQUIJOR.

NOTE.—The conspecificity of Phyllophora maillardii and Cryptonemia rigida was proposed by Schmitz (1895:167), who, however, incorrectly adopted the later of the two names.

Carpopeltis prolifera (Hariot) Kawaguchi and Masuda

Gigartina prolifera Hariot, 1891:220 [type locality: Yokosuka, Kanagawa Prefecture, Japan]

Carpopeltis prolifera (Hariot) Kawaguchi and Masuda, 1984:232.

Grateloupia fiabellata Holmes, 1896:254, pl. 9: figs. 3a-c [type locality: Enoshima, Kanagawa Prefecture, Japan].

Carpopeltis fiabellata (Holmes) Okamura, 1934 [1933–1942]:39, pl. 321: figs. 1–6.—Cordero, 1976c:10, 1977a:103, pl. XVI:B; 1978a:33

PHILIPPINE DISTRIBUTION.—BATANES.

NOTE.—The conspecificity of Gigartina prolifera and Grateloupia fiabellata was proposed by Kawaguchi and Masuda (1984.)

Cryptonemia J. Agardh

Cryptonemia crenulata (J. Agardh) J. Agardh

Phyllophora crenulata J. Agardh, 1841:18 [type locality: Bahia, Brazil].

Cryptonemia crenulata (J. Agardh) J. Agardh, 1851 [1851–1863]:225.—Cordero, 1977a:104, fig. 81, pl. XVI:C.—Meñez and Calumpong, 1981:381.—Hurtado-Ponce, 1983:136.—Cordero, 1984a:97.—Tungpalan, 1984:140.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Batangas CENTRAL VISAYAS

Cryptonemia luxurians (C. Agardh) J. Agardh

Sphaerococcus lactuca C. Agardh var. luxurians C. Agardh, 1822a:232 [type locality: Brazil].

Cryptonemia luxurians (C. Agardh) J. Agardh, 1851 [1851–1863]:228.—Cordero, 1976c:8, 9, 10; 1977a:105; 1978a:34.

PHILIPPINE DISTRIBUTION.—BATANES.

Cryptonemia schmitziana (Okamura) Okamura

Prionitis schmitziana Okamura, 1899a:6 [syntype localities: various, all in Japan].

Cryptonemia schmitziana (Okamura) Okamura, 1910 [1909–1912]:77.—Cordero, 1978a:34.

PHILIPPINE DISTRIBUTION.—BATANES.

Grateloupia C. Agardh

Grateloupia conferta (Kützing) Kützing

Grateloupia filicina (Lamouroux) C. Agardh var. conferta Kützing, 1849:730 [type locality: Java, Indonesia].—Martens, 1868:30, 90–91.

Grateloupia conferta (Kützing) Kützing, 1867:7, pl. 23: figs, a, b.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Grateloupia dichotoma J. Agardh

Grateloupia dichotoma J. Agardh, 1842:103 [syntype localities: various, all in Mediterranean France].—Velasquez et al., 1973:29.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

Grateloupia divaricata Okamura

Grateloupia divaricata Okamura, 1895:482, pl. IX: figs. 1, 2 [syntype localities: various, all in Japan].—Cordero, 1977a:106; 1982a:61.—Hurtado-Ponce, 1983:137; 1984:180.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Norte.

Grateloupia doryphora (Montagne) Howe

Halymenia (?) doryphora Montagne, 1839:21 [type locality: Callao, Peru].

Grateloupia doryphora (Montagne) Howe, 1914:169 [“doryophora”].

Grateloupia californica Kylin, 1941:9, fig. 2B, pl. 1: figs. 1, 2 [type locality: La Jolla, California, USA].—Cordero, 1977a:106, pl. XVII:A; 1980b:46.—Hurtado-Ponce, 1983:136; 1984:180.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

NOTE.—The synonymy was proposed by Hollenberg and Abbott (1966:67).

Grateloupia filicina (Lamouroux) C. Agardh

Fucus filicinus Wulfen in Jacquin, 1789:157, pl. 15: fig. 2 [type locality: Trieste, Italy].

Delesseria filicina Lamouroux, 1813:125.

Grateloupia filicina (Lamouroux) C. Agardh, 1822a:223.—Martens, 1868:30, 46, 88–89.—Zaneveld, 1956:40; 1959:120.—Reyes, 1972:154.—Trono, 1973a:131, figs. 10–12.—Velasquez, Trono, and Doty, 1975:147.—Cordero, 1977a:107.—Vannajan and Trono, 1978:17, fig. 23.—Cordero, 1980b:47, pl. [47].—Trono and Fortes, 1980:75.—Ganzon-Fortes, 1981:23.—Cordero, 1982a:61.—Trono and Fortes, 1982:154.—Hurtado-Ponce, 1983:137; 1984:180.—Marcos-Agngarayngay, 1984b:125.—Tungpalan, 1984:140.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Rizal, Sorsogon. NEGROS: Negros Oriental. MINDANAO: Zamboanga.

NOTE.—Fucus filicinus Wulfen, the intended basionym of Grateloupia filicina, is a later homonym of F. filicinus Hudson (1762:473) and hence not priorable. Delesseria filicina Lamouroux is treated as a nomen novum in accordance with Article 72, Note I, of the ICBN.

Grateloupia ramosissima Okamura

Grateloupia ramosissima Okamura, 1913 [1913–1915]:60, pl. CXVII: figs. 1–11 [syntype localities: various, all in Japan].—Cordero, 1977a:107; 1980b:47; 1982a:61.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan. SIQUIJOR.

Halymenia C. Agardh

Halymenia acuminata (Holmes) Okamura

Grateloupia acuminata Holmes, 1896:254, pl. 10: figs. 2a-c [type locality: Enoshima, Kanagawa Prefecture, Japan].

Halymenia acuminata (Holmes) Okamura, 1908 [1907–1909]:174, pl. XXXV: figs. 6–12.—Cordero, 1977a:108, fig. 83.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. PANAY: Iloilo. PALAWAN.

Halymenia dilatata Zanardini

Halymenia dilatata Zanardini, 1851:35 [type locality: Red Sea].—Domantay, 1962:291.—Velasquez, Trono, and Doty, 1975:149.—Cordero, 1977a:109, figs. 84, 85, pl. XVIII:B.—Vannajan and Trono, 1978:18.—Puig and Cordero, 1979:39.—Meñez and Calumpong, 1981:381.—Calumpong, 1982:145.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Bataan, Cavite, Batangas. MINDORO: Oriental Mindoro. LEYTE (Biliran I.). PANAY: Capiz. NEGROS: Negros Oriental (Apo I.). CEBU (Sumilon I.).

Halymenia durvillaei Bory de Saint-Vincent

Halymenia durvillaei Bory de Saint-Vincent, 1828 [1827–1829]:180, pl. 15 [type locality: New Ireland, Papua New Guinea].—Martens, 1868:29, 88–89.—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Galutira and Velasquez, 1964:504, pl. 3: fig. 10; pl. 8: fig. 29.—Domantay, 1968:26.—De Leon and Domantay, 1971:5, 10.—Velasquez, 1971:448, fig. 30.—Reyes, 1972:154.—Velasquez, 1972:63.—Velasquez et al., 1973:28, pl. 12: fig. 56.—Ortega, Alcala, and Reyes, 1974:188.—Velasquez, Trono, and Doty, 1975:149.—Cordero, 1977a:III, fig. 86, pl. XVII:B. —Trono and Young, 1977:59.—Cordero, 1979b:276, 290, fig. 1.—Velasquez, 1979b:231.—Cordero, 1980b:48, fig. 11A, pl. 30.—Guzman, 1981:43, 51.—Liao and Sotto, 1980:99.—Reyes, 1980:133, pl. 8: fig. 5.—Trono and Fortes, 1980:75.—Ganzon-Fortes, 1981:23.—Meñez and Calumpong, 1981:381.—Calumpong, 1982:145.—Cordero, 1982a:60, fig. 13.—Trono and Fortes, 1982:154.—Hurtado-Ponce, 1983:137.—Cordero, 1984a:96; 1984b:65.—Hurtado-Ponce, 1984:180.—Marcos-Agngarayngay, 1984a:36, fig. 29; 1984b:126.—Tungpalan, 1984:141.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Bataan, Cavite (Corregidor I.), Batangas, Quezon, Sorsogon. CATANDUANES. MINDORO: Oriental Mindoro. SAMAR: Western Samar, Eastern Samar. PANAY: Aklan. GUIMARAS. NEGROS: Negros Occidental, Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Zamboanga. PALAWAN, SULU.

Halymenia floresia (Clemente y Rubio) C. Agardh

Fucus floresius Clemente y Rubio, 1807:312 [type locality: Sanlúcar de Barrameda, Spain].

Halymenia floresia (Clemente y Rubio) C. Agardh, 1817:XIX.—Tungpalan, 1984:141.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Halymenia formosa Harvey ex Kützing

Halymenia formosa Harvey ex Kützing, 1866:33, pl. 91: figs. g,h [syntype localities: Tonga; Australia].—Seale, 1911:309.—Wester, 1916:159; 1921:224; 1924:21.—G. Blanco, 1938:513.—Quisumbing, 1951:1010 [“formosana”].—Montilla and Blanco, 1953:fig. 5:2.—Domantay, 1962:291.—Velasquez, Trono, and Doty, 1975:149.—Cordero, 1982a:60.

Halymenia durvillaei Bory de Saint-Vincent var. formosa (Harvey ex Kützing) Weber-van Bosse, 1921:235.—Zaneveld, 1956:41; 1959:120.—Cordero, 1976c:8; 1978a:31.—Cordero, 1984a:96 [“var. formosana Yamada”].

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, La Union, Pangasinan, Manila Bay.

Halymenia harveyana J. Agardh

Halymenia harveyana J. Agardh, 1892:55 [type locality: Port Phillip Heads, Victoria, Australia].—Meñez, 1961:72.—Velasquez, Trono, and Doty, 1975:149.—Cordero, 1977a:112, pl. XVIII:C.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan.

Halymenia japonica

Halymenia japonica.—Domantay, 1968:26.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—Domantay did not give an author for this name and we have been unable to find its place of publication.

Halymenia maculata J. Agardh

Halymenia maculate J. Agardh, 1885:12 [type locality: Mauritius].—Domantay, 1962:291.—Velasquez, Trono, and Doty, 1975:149.—Trono and Ang, 1982:16.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. PALAWAN (Bugsuk I.).

*Halymenia microcarpa (Montagne) P.C. Silva, new combination

Mesogloia microcarpa Montagne, 1844a:660 [“Mesogloea”] [type locality: “in insulis Philippinensibus”].—J. Agardh, 1848:59.—Kützing, 1849:546.—Montagne, 1856:400.—Martens, 1868:68–69.—Velasquez, Trono, and Doty, 1975:153.

Halymenia ceylanica Harvey ex Kützing, 1866:33, pl. 93: figs, a, b [type locality: Sri Lanka].—Grunow, 1874:32.

Halymenia durvillaei Bory de Saint-Vincent var. ceylanica (Harvey ex Kützing) Weber-van Bosse, 1921:235.—Trono, 1973d:17.—Trono and Bina, 1973:6.—Trono, 1974b:87.—Vannajan and Trono, 1978:17, fig. 22.—Trono and Ganzon-Fortes, 1980:71, fig. [s.n.].—Trono and De Lara, 1981:17, pl. XI: fig. 2.—Saraya and Trono, 1982:36.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite, Batangas, Quezon, MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MINDANAO: Zamboanga. SULU: Sulu (Siasi I.).

NOTE.—The conspecificity of Mesogloia microcarpa and Halymenia ceylanica was proposed by Grunow (1874:32).

Prionitis J. Agardh

Prionitis cornea (Okamura) Dawson

Grateloupia cornea Okamura, 1913 [1913–1915]:63, pl. CXVIII [syntype localities: various, all in Japan].

Carpopeltis cornea (Okamura) Okamura, 1942 [1933–1942]:100.—Cordero, 1977a:101.

Prionitis cornea (Okamura) Dawson, 1958:71.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Thamnoclonium Kützing

*Thamnoclonium procumbens Weber-van Bosse

Thamnoclonium procumbens Weber-van Bosse, 1921:251, figs. 78, 79 [syntype localities: various in Indonesia; Capual I., Sulu Prov., Sulu Archipelago].—Velasquez, Trono, and Doty, 1975:165.

PHILIPPINE DISTRIBUTION.—As above.

*Thamnoclonium treubii Weber-van Bosse

Thamnoclonium treubii Weber-van Bosse, 1910:587, pl. XVI: fig. 1 [type locality: North Ubian I., Sulu Prov., Sulu Archipelago].—Weber-van Bosse, 1921:250.—Velasquez, Trono, and Doty, 1975:165.

PHILIPPINE DISTRIBUTION.—As above.

Family KALLYMENIACEAE

Callophyllis Kützing

Callophyllis adhaerens Yamada

Callophyllis adhaerens Yamada, 1932b:273, pl. VI:a [syntype localities: various, all in Japan].—Velasquez et al., 1973:29, pl. 12: fig. 57.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

Callophyllis adnata Okamura

Callophyllis adnata Okamura, 1932 [1929–1932]:79, pl. CCLXXXIX [type locality: Enoshima, Kanagawa Prefecture, Japan].—Velasquez et al., 1973:29, pl. 12: fig. 58.—Cordero, 1980c:71, fig. 3.—Hurtado-Ponce, 1983:138; 1984:180.—Marcos-Agngarayngay, 1984b:126.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Bataan.

Callophyllis okamurae P.C. Silva, new name

Pugetia japonica Kylin, 1941:16 [type locality: Chiba Prefecture, Japan].

Callophyllis chilensis [misapplied name fide Kylin, 1941:15–16].—Cordero, 1977a:115.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

NOTE.—Norris (1957:298) suggested that Pugetia japonica was referable to Callophyllis. Because of the existence of C. japonica Okamura (in De Toni and Okamura, 1895:77, pl. 16: figs. 13–17), a new epithet must be chosen. Pugetia japonica is validated by Kylin’s reference to Howe (1914:117), who distinguished the Japanese plant from Callophyllis chilensis without giving it a name.

Kallymenia J. Agardh

Kallymenia callophylloides Okamura and Segawa

Kallymenia callophylloides Okamura and Segawa in Segawa, 1935:78, fig. 1., pl. 19: fig. 1 [type locality: Susaki, Shizuoka Prefecture, Japan].—Cordero, 1977a:116, fig. 94, pl. XXV:A.—Marcos-Agngarayngay, 1984a:39, fig. 33.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Kallymenia pacifica Kylin

Meredithia californica ]. Agardh, 1899:49 [type locality: San Diego, California, USA].

Kallymenia pacifica Kylin, 1956:233 [“Callymenia”].—Cordero, 1981b:24, fig. 1.

PHILIPPINE DISTRIBUTION.—PANAY: Iloilo.

NOTE.—When transferring Meredithia californica to Kallymenia, Kylin was obliged to change the epithet by the prior existence of K. californica Farlow (1877:241).

Kallymenia sessilis Okamura

Kallymenia sessilis Okamura, 1934 [1933–1942]:20, pl. 312 [type locality: Tateyama, Japan].—Velasquez et al., 1973:29, pl. 12: fig. 59.—Cordero, 1981b:26, fig. 2.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas, Bataan. PANAY: Iloilo.

Order CORALLINALES

Family CORALLINACEAE

Amphiroa Lamouroux

Amphiroa anastomosans Weber-van Bosse

Amphiroa anastomosans Weber-van Bosse, 1904b:91, pl. XVI: figs. 3, 4 [syntype localities: Borneo Bank and Flores, Indonesia].—Cordero, 1977a:82.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Sorsogon. MINDORO: Oriental Mindoro. SAMAR: Western Samar. CENTRAL VISAYAS.

Amphiroa anceps (Lamarck) Decaisne

Corallina anceps Lamarck, 1815:238 [type locality: “les mers Australes ou de la Nouv. Holl.”].

Amphiroa anceps (Lamarck) Decaisne, 1842:125.—Cordero, 1977a:81, pl. XIV:A.—Hurtado-Ponce, 1983:131.—Cordero, 1984a:94.

Amphiroa dilatata Lamouroux, 1816:299 [type locality: “Australasie”].—Garcia, 1979:44 (table 1).—Cordero, 1984a:94.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, siquijor. MINDANAO.

NOTE.—The synonymy was proposed by Weber-van Bosse (1904b:93).

Amphiroa annulata Lemoine

Amphiroa annulata Lemoine, 1929:78, fig. 34, pl. IV: fig. 1 [type locality: James I. [Isla San Salvador or Santiago], Galápagos].—Domantay, 1962:288—Velasquez, Trono, and Doty, 1975:126.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Amphiroa beauvoisii Lamouroux

Amphiroa beauvoisii Lamouroux, 1816:299 [type locality: Portugal].—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

*Amphiroa cumingii Montagne

Amphiroa cumingii Montagne, 1844a:662 [type locality: “in insulis Philippinensibus”].—Kützing, 1849:702.—Areschoug, 1852:541.—Montagne, 1856:430.—Martens, 1868:86–87.—Dickie, 1876a:244.—Velasquez, Trono, and Doty, 1975:126.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Amphiroa ephedraea (Lamarck) Decaisne

Corallina ephedraea Lamarck, 1815:238 [type locality: “les mers Australes ou de la Nouv. Holl.”].

Amphiroa ephedraea (Lamarck) Decaisne, 1842:124.—Cordero, 1973b:30 [A. ephedraea prox.]; 1976c:8; 1977a:82; 1978a:27.—Puig and Cordero, 1979:38.—Cordero, 1984c:54.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. MASBATE. LEYTE (Biliran I.), MINDORO: Oriental Mindoro. PALAWAN.

Amphiroa foliacea Lamouroux

Amphiroa foliacea Lamouroux in Quoy and Gaimard, 1824:628, pl. 93: figs. 2, 3 [type locality: Mariana Is.].—Cornejo and Velasquez, 1972:181.—Trono and Biña, 1973:4.—Velasquez et al., 1973:27, pl. II: fig. 52.—Cordero, 1977a:84.—Trono and Young, 1977:59.—Puig and Cordero, 1979:38.—Liao and Sotto, 1980:98.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.—Trono and De Lara, 1981:15, pl. X: fig. 4.—Saraya and Trono, 1982:34a, pl. IV: fig. 1.—Hurtado-Ponce, 1983:131.—Cordero, 1984a:94; 1984b:64; 1984c:54.—Marcos-Agngarayngay. 1984a:32, fig. 26.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Bataan, Batangas, Quezon, Albay. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MASBATE. SAMAR: Western Samar, Eastern Samar. LEYTE (incl. Biliran I.), CEBU (Mactan I.). PALAWAN.

*Amphiroa foliacea Lamouroux f. erecta Weber-van Bosse

Amphiroa foliacea Lamouroux f. erecta Weber-van Bosse, 1904b:93, pl. XIV: figs. 2–8 [syntype localities: various in Indonesia; North Ubian I., Sulu Prov., and Sangasiapu I., Tawitawi Prov., Sulu Archipelago].—Velasquez, Trono, and Doty, 1975:127 [without designation of forma].—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—As above.

Amphiroa fragilissima (Linnaeus) Lamouroux

Corallina fragilissima Linnaeus, 1758:806 [type locality: Jamaica].

Amphiroa fragilissima (Linnaeus) Lamouroux, 1816:298.—Dickie, 1876a:243.—Weber-van Bosse, 1904b:90 [f. fragilissima].—Howe, 1932:170.—Domantay, 1962:288.—Cornejo and Velasquez, 1972:182.—Reyes, 1972:153.—Trono, 1972a:103; 1973d:16.—Trono and Biña, 1973:4 [f. fragilissima].—Velasquez et al., 1973:28, pl. II: fig. 53.—Ortega, Alcala, and Reyes, 1974:185–188 (tables 1–4, 7).—Trono, 1974b:86 [f. fragilissima].—Velasquez, Trono, and Doty, 1975:127.—Cordero, 1976c:8; 1977a:85, figs. 62, 63; 1977a:85, pl. XIV:B [var. fragilissima]; 1978a:28.—Trono, 1978:16.—Vannajan and Trono, 1978:16, fig. 16.—Cordero, 1979b:289.—Puig and Cordero, 1979:38.—Liao and Sotto, 1980:98.—Reyes, 1980:130, pl. 6: fig. 6.—Trono and Ganzon-Fortes, 1980:67, fig. [s.n.].—Chan, 1981:387.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.—Trono and De Lara, 1981:16, pl. X: fig. 3.—Saraya and Trono, 1982:34a, pl. V: fig. 2.—Hurtado-Ponce, 1983:132.—Cordero, 1984a:95; 1984c:54.—Marcos-Agngarayngay, 1984a:34, fig. 27.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Bataan, Cavite, Batangas, MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. MASBATE. LEYTE (Biliran I.), PANAY: Aklan. NEGROS: Negros Occidental, Negros Oriental, CEBU (Mactan I.). SIQUIJOR. PALAWAN. SULU: Sulu (Siasi I.), Tawitawi (incl. Sangasiapu I.)

Amphiroa fragilissima (Linnaeus) Lamouroux f. cyathifera (Lamouroux) Weber-van Bosse

Amphiroa cyathifera Lamouroux, 1824:627 [type locality: Moluccas, Indonesia].

Amphiroa fragilissima (Linnaeus) Lamouroux f. cyathifera (Lamouroux) Weber-van Bosse, 1904b:90 [including Philippine record].—Trono and Biña, 1973:4.—Trono, 1974b:86.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. SULU: Sulu (Siasi I.), Tawitawi (Sangasiapu I.).

Amphiroa hancockii W.R. Taylor

Amphiroa hancockii W.R. Taylor, 1942:95, pl. 13 [type locality: Caledonia, Atlantic Panama].—Velasquez et al., 1973:28, pl. II: fig. 54.

PHILIPPINE DISTRIBUTION.—LUZON: Bataan.

Amphiroa pacifica Kützing

Amphiroa pacifica Kützing, 1858:20. pl. 43: fig. 1 [type locality: Peru].—Martens, 1868:28, 84–85.—Velasquez, Trono, and Doty, 1975:127.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Amphiroa rigida Lamouroux

Amphiroa rigida Lamouroux, 1816:297, pl. XI: fig. 1 [type locality: Mediterranean Sea].—Cordero, 1977a:86.—Meñez and Calumpong, 1981:381.—Meñez, Phillips, and Calumpong, 1983:23.—Meñez and Calumpong, 1984:105.

PHILIPPINE DISTRIBUTION.—SAMAR: Western Samar. CENTRAL VISAYAS. PALAWAN.

Amphiroa subcylindrica Dawson

Amphiroa subcylindrica Dawson, 1953:139, pl. 29: fig. 1 [type locality: Guaymas, Sonora, Mexico].—Domantay, 1962:288.—Velasquez, Trono, and Doty, 1975:127.—Saraya and Trono, 1982:34b, pl. V: fig. 1.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Amphiroa tribulus (Ellis and Solander) Lamouroux

Corallina tribulus Ellis and Solander, 1786:124, pl. 21: fig. e [type locality: West Indies].

Amphiroa tribulus (Ellis and Solander) Lamouroux, 1816:302.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Amphiroa valonioides Yendo

Amphiroa valonioides Yendo, 1902:5, pl. I: figs. 1–3; pl. IV: fig. 1 [syntype localities: various, all in Japan].—Cordero, 1977a:86, fig. 60, pl. XIII: B.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon, SAMAR: Eastern Samar. LEYTE.

Amphiroa zonata Yendo

Amphiroa zonata Yendo, 1902:10, pl. 1: figs. 11–14; pl. IV: fig. 9 [syntype localities: various, all in Japan].—Cordero, 1977a:86.—Hurtado-Ponce, 1983:132.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, MINDORO: Oriental Mindoro. SIQUIJOR.

Cheilosporum (Decaisne) Zanardini

Cheilosporum cultratum (Harvey) J.E. Areschoug

Amphiroa cultrata Harvey, 1849 [1847–1849]:102, pl. XXXIX [in part] [lectotype locality: Port Natal (Durban), South Africa fide Johansen, 1977:175].

Cheilosporum cultratum (Harvey) J.E. Areschoug, 1852:545.—Domantay, 1962:288.—Meñez, 1961:71.—Trono, 1973a:133, fig. 9.—Velasquez, Trono, and Doty, 1975:134.—Cordero, 1977a:88, figs. 64, 65.—Saraya and Trono, 1982:36, pl. V: fig. 4.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Sorsogon. SAMAR: Western Samar, Eastern Samar.

Cheilosporum jungermannioides Ruprecht ex J.E. Areschoug

Cheilosporum jungermannioides Ruprecht ex J.E. Areschoug, 1852:546 [type locality: Tahiti]; 1976c:9; 1977a:88, figs. 66, 68; 1978a:28 [C. jungermannioides prox.].—Hurtado-Ponce, 1983:132.—Cordero, 1984a:95; 1984b:64.—Marcos-Agngarayngay, 1984a:34, fig. 28.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan. MINDORO: Oriental Mindoro. LEYTE. PALAWAN.

Cheilosporum sagittatum (Lamouroux) J.E. Areschoug

Corallina sagittata Lamouroux in Quoy and Gaimard, 1824:625, pl. 95: figs. 11, 12 [type locality: Mauritius].

Cheilosporum sagittatum (Lamouroux) J.E. Areschoug, 1852:545.—Trono and De Lara, 1981:16, pl. X: fig. 2.

PHILIPPINE DISTRIBUTION.—MINDORO: Occidental Mindoro (Lubang Is.).

Corallina Linnaeus

Corallina frondescens Postels and Ruprecht

Corallina frondescens Postels and Ruprecht, 1840:20, pl. XL: fig. 103 [type locality: Unalaska I., Aleutian Is.].

Corallina pinnatifolia (Manza) Dawson var. digitata Dawson, 1953:125, pl. 9: figs. 14–20; pl. 30: fig. 1 [type locality: Guaymas, Sonora, Mexico].—Cordero, 1976c:8, 9, 10; 1977a:90; 1978a:28.

PHILIPPINE DISTRIBUTION.—BATANES.

NOTE.—The synonymy was proposed by Johansen (1976:403).

Fosliella Howe

Fosliella farinosa (Lamouroux) Howe

Melobesia farinosa Lamouroux, 1816:315, pl. XII: fig. 3 [type locality: not specified (Mediterranean Sea fide Chamberlain, 1983:343)].—Piccone, 1889:51, 59.—Foslie, 1904:55.—Domantay, 1962:293.—Velasquez, Trono, and Doty, 1975:153.

Fosliella farinosa (Lamouroux) Howe, 1920:587.—Cordero, 1977a:90.—Trono, 1978:17.—Reyes, 1980:132.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MARINDUQUE. SAMAR: Eastern Samar. SIQUIJOR. SULU: Sulu (North Ubian I.).

Haliptilon (Decaisne) Lindley

Haliptilon cubense (Montagne ex Kützing) Garbary and Johansen

Jania cubensis Montagne ex Kützing, 1849:709 [type locality: Cuba].

Corallina cubensis (Montagne ex Kützing) Kützing, 1858:37, pl. 77: fig. II.—Meñez and Calumpong, 1981:381.

Haliptilon cubense (Montagne ex Kützing) Garbary and Johansen, 1982:218.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Hydrolithon (Foslie) Foslie

Hydrolithon reinboldii (Weber-van Bosse and Foslie) Weber-van Bosse

Lithophyllum reinboldii Weber-van Bosse and Foslie in Foslie, 1901c:5 [lectotype locality: Muaras Reef, East Kalimantan, Indonesia fide Adey and Lebednik, 1967:32].

Goniolithon reinboldii (Weber-van Bosse and Foslie) Weber-van Bosse and Foslie in Foslie, 1904:49 [including Philippine records].—Velasquez, Trono, and Doty, 1975:146.

Hydrolithon reinboldii (Weber-van Bosse and Foslie) Weber-van Bosse and Foslie in Foslie, 1909:55.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.), Tawitawi (Sangasiapu I.).

Jania Lamouroux

Jania adhaerens Lamouroux

Jania adhaerens Lamouroux, 1816:270 [type locality: “Méditerranée?”].—Cordero, 1976c:9, 10; 1977a:91, fig. 73; 1978a:29.—Chan, 1981:387, 389.—Meñez and Calumpong, 1981:381.—Hurtado-Ponce, 1983: 133.—Meñez, Phillips, and Calumpong, 1983:23.—Meñez and Calumpong, 1984:105.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, SAMAR: Eastern Samar. NEGROS: Negros Oriental. SIQUIJOR. PALAWAN.

Jania capillacea Harvey

Jania capillacea Harvey, 1853:84 [type locality: Bahia Honda, Florida, USA].—Trono, 1972a:103; 1973d:16.—Trono and Biña, 1973:5.—Westernhagen, 1973a:65.—Trono, 1974b:85.—Westernhagen, 1974:112 (table 1).—Cordero, 1976c:9; 1977a:91, figs. 67, 69; 1978a:29.—Trono, 1978:17.—Fortes and Trono, 1980:60, 66.—Liao and Sotto, 1980:99.—Trono and Ganzon-Fortes, 1980:69, fig. [s.n.]—Chan, 1981:387, 389.—Fortes, 198lb:396.—Trono and De Lara, 1981:16, pl. X: fig. 1.—Saraya and Trono, 1982:35.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Batangas. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. LEYTE. PANAY: Iloilo. NEGROS: Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. SULU: Sulu (Siasi I.), Tawitawi.

Jania decussato-dichotoma (Yendo) Yendo

Corallina decussato-dichotoma Yendo, 1902:25, pl. III: figs. 1–3: pl. VII: figs. 3, 4 [syntype localities: various, all in Japan].

Jania decussato-dichotoma (Yendo) Yendo, 1905b:37.—Cordero, 1973b:31 [J. decussato-dichotoma prox.]; 1976c:8; 1978a:29.—Fortes, 1981b:396.—Saraya and Trono, 1982:35.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. LEYTE (Biliran I.).

NOTE.—Cordero (1976c) recognized both J. adhaerens and J. decussato-dichotoma from the Batan Islands, but later (1977a) he merged the two species. While all Philippine reports under the two names may be referable to a single species, it seems likely that this species will prove to be distinct from J. adhaerens, in which case it would be called J. decussato-dichotoma.

Jania longiarthra Dawson

Jania longiarthra Dawson, 1953:119, pl. 9: fig. 4; pl. 27: fig. 4 [type locality: Isla Espiritu Santo, Baja California Sur, Mexico].—Trono, 1978:17.

PHILIPPINE DISTRIBUTION.—MARINDUQUE.

Jania micrarthrodia Lamouroux

Jania micrarthrodia Lamouroux, 1816:271, pl. 9: fig. 5 [type locality: “Australasie”].

Jania tenuissima Sonder, 1848 [1846–1848]:186 [type locality: Western Australia, Australia (mouth of Swan River fide Harvey, 1849 [1847–1849]:106)].—Piccone, 1886:67, 90.—Velasquez, Trono, and Doty, 1975:151.

PHILIPPINE DISTRIBUTION.—MASBATE (Ticao I.).

NOTE.—The synonymy was proposed by Areschoug (1852:555).

Jania pacifica J.E. Areschoug

Jania pacifica J.E. Areschoug, 1852:556 [type locality: Guatulco, Oaxaca, Mexico].

Jania mexicana W.R. Taylor, 1945:197, pl. 60 [type locality: Bahía Petatlán, Guerrero, Mexico].—Cordero, 1977a:92, fig. 74.—Chan, 1981:387.—Saraya and Trono, 1982:36, pl. V: fig. 3.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. PALAWAN.

NOTE.—The conspecificity of Jania mexicana and J. pacifica became obvious to one of us (PCS) after collecting in Pacific tropical Mexico, the type locality for both species.

Jania pumila Lamouroux

Jania pumila Lamouroux, 1816:269, pl. IX: fig. 2 [syntype localities: Red Sea; “Indes Orientales”].—Velasquez et al., 1973:28, pl. II: fig. 55.—Cordero, 1977a:92, figs. 70–72.—Chan, 1981:387, 389.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Bataan, Batangas.

Jania rubens (Linnaeus) Lamouroux

Corallina rubens Linnaeus, 1758:806 [type locality: Europe].

Jania rubens (Linnaeus) Lamouroux, 1816:272, pl. IX: fig. 6, 7.—Meñez, 1961:71, pl. 8: figs. 89, 90.—Reyes, 1972:153.—Velasquez, Trono, and Doty, 1975:150.—Reyes, 1980:131, pl. 7: figs. 4a,b.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. NEGROS: Negros Oriental, SIQUIJOR.

Jania tenella (Kützing) Grunow

Corallina tenella Kützing, 1858:41, pl. 85: fig. II [syntype localities: Napoli, Italy; Mexico].

Jania tenella (Kützing) Grunow, 1874:42.—Cornejo and Velasquez, 1972:183, pl. 3: fig. 27 [var. tenella].—Velasquez, Trono, and Doty, 1975:150.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

Jania tenella (Kützing) Grunow var. zacae Dawson

Jania tenella (Kützing) Grunow var. zacae Dawson, 1953:121, pl. 8: fig. 3; pl. 31: fig. 1 [type locality: Bahía Piedra Blanca, Pacific Costa Rica].—Domantay, 1962:292.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Jania ungulata (Yendo) Yendo

Corallina ungulata Yendo, 1902:26, pl. III: figs. 7, 8; pl. VII: fig. 8 [syntype localities: various, all in Japan].

Jania ungulata (Yendo) Yendo, 1905b:38.—Vannajan and Trono, 1978:17, fig. 17.—Chan, 1981:387, 389.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite.

Jania ungulata (Yendo) Yendo f. brevior (Yendo) Yendo

Corallina ungulata Yendo f. brevior Yendo, 1902:27, pl. III: fig. 9; pl. VII: fig. 9 [type locality: Chiba Prefecture, Japan].

Jania ungulata (Yendo) Yendo f. brevior (Yendo) Yendo, 1905b:38.—Trono, 1973d:16 [var. brevior].—Cordero, 1977a:94, figs. 75, 76, pl. XIV:c [var. brevior]; 1978a:30 [var. brevior].—Liao and Sotto, 1980:99 [var. brevior].—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381 [var. brevior].—Saraya and Trono, 1982:35 [var. brevior].

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. SIQUIJOR. CEBU (Mactan I.).

Lithophyllum Philippi

Lithophyllum byssoides (Lamarck) Foslie

Nullipora byssoides Lamarck, 1801:374 [type locality: Mediterranean Sea].

Lithothamnion byssoides (Lamarck) Philippi, 1837:388.—Dickie, 1877:489.—Velasquez, Trono, and Doty, 1975:152.

Lithophyllum byssoides (Lamarck) Foslie, 1900d:20.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Lithophyllum moluccense (Foslie) Foslie

Lithothamnion moluccense Foslie, 1897:12 [type locality: Moluccas, Indonesia].

Lithophyllum moluccense (Foslie) Foslie, 1901a:12 [including Philippine record].—Foslie, 1904:67, pl. XII: fig. 8 [f. typica].—Velasquez, Trono, and Doty, 1975:152.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I.).

Lithophyllum pallescens (Foslie) Foslie

Lithothamnion pallescens Foslie, 1895:4, pl. 1: figs. 11–13 [type locality:Isla Espíritu Santo, Baja California Sur, Mexico].

Lithophyllum pallescens (Foslie) Foslie, 1900d:20.

Lithophyllum okamurae Foslie, 1900c:4 [“okamurai”] [type locality: Kanagawa Prefecture, Japan].—Foslie, 1904:59, pl. XI: fig. 17.—Domantay, 1962:292.—Velasquez, Trono, and Doty, 1975:152 [“okamurai”] and [“okamura”]

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. SULU: Sulu (North Ubian I., Tongquil I.), Tawitawi (Sangasiapu I.).

NOTE.—The synonymy was proposed by Adey, Townsend, and Boykins (1982:40).

Lithoporella (Foslie) Foslie

Lithoporella indica (Foslie) Adey

Litholepis indica Foslie, 1907:21 [type locality: Réunion].

Lithoporella indica (Foslie) Adey, 1970:15.

PHILIPPINE DISTRIBUTION.—See “Litholepsis indica Foslie f. philippinensis Foslie.”

TAXON OF UNCERTAIN VALUE

Philippine records for Lithoporella indica are under the following infraspecific taxon, which has not been formally transferred to it.

*Litholepsis indica Foslie f. philippinensis Foslie

Litholepsis indica Foslie f. philippinensis Foslie, 1908:9 [type locality: Adiagñao, Camarines Sur Prov., Luzon].—De Toni, 1924:655.

PHILIPPINE DISTRIBUTION.—As above.

Lithoporella melobesioides (Foslie) Foslie

Mastophora melobesioides Foslie, 1903:24 [type locality: Maldives].—Foslie, 1904:73.—Velasquez, Trono, and Doty, 1975:153.

Lithoporella melobesioides (Foslie) Foslie, 1909:59.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Capual I., Tongquil I.), Tawitawi (Sangasiapu I.).

Lithothamnion Heydrich

Woelkerling (1983b:193), after discovering that none of the five original species of Lithothamnium Philippi (1837:387) conforms to any modern concept of the genus, proposed the conservation of that generic name as treated by Heydrich (1897c:412), lectotypified by L. muelleri Lenormand ex Rosanoff. Heydrich spelled the name Lithothamnion, the spelling that has been proposed for conservation.

*Lithothamnion australe Foslie f. brachiatum Foslie

Lithothamnion australe Foslie f. brachiatum Foslie, 1904:24, pl. II: figs. 25–38 [“brachiata”] [syntype localities: Semau I. and Timor, Indonesia; North Ubian I. and Tongquil I., Sulu Prov., Sulu Archipelago].—Foslie, 1929: pl. XVII: figs. 51–54.

PHILIPPINE DISTRIBUTION.—As above.

*Lithothamnion australe Foslie f. minutulum Foslie

Lithothamnion australe Foslie f. minutulum Foslie, 1904:24, pl. II: figs. 39–62 [“minutula”] [syntype localities: Tual, Kai Is., Moluccas, Indonesia; Tongquil I., Sulu Prov., Sulu Archipelago].—Foslie, 1929: pl. XVII: figs. 59–70.

PHILIPPINE DISTRIBUTION.—As above.

*Lithothamnion australe Foslie f. ubianum Foslie

Lithothamnion australe Foslie f. ubianum Foslie, 1904:24, fig. 12, pl. II: figs. 18–24 [“ubiana”] [syntype localities: Celebes, Indonesia; Capual I., North Ubian I., and Tongquil I., Sulu Prov., Sulu Archipelago].—Foslie, 1929: pl. XVII: figs. 40–44.

PHILIPPINE DISTRIBUTION.—As above.

NOTE.—The typification of Lithothamnion australe is discussed by Adey, Townsend, and Boykins (1982:57).

Lithothamnion indicum Foslie f. subtile Foslie

Lithothamnion indicum Foslie f. subtile Foslie, 1907:7 [“subtilis”] [syntype localities: Indian and Pacific oceans].—Foslie, 1929: pl. XIII: fig. 26.

Lithothamnion fruticulosum (Kützing) Foslie f. clavulatum [misapplied name fide Foslie, 1907:7].—Foslie, 1904:20, pl. II: figs. 6, 7, 9 [“clavulata”].—Velasquez, Trono, and Doty, 1975:152 [without designation of forma].

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Capual I., Tongquil I.), Tawitawi (Sangasiapu I.).

Mastophora Decaisne

Mastophora rosea (C. Agardh) Setchell

Zonaria rosea C. Agardh, 1824:264 [type locality: Guam, Mariana Is.].

Mastophora rosea (C. Agardh) Setchell, 1943:129.—Domantay, 1962:292.—Velasquez, 1971:453, fig. 39.—Reyes, 1972:153.—Trono, 1972a:103.—Cordero, 1973b:31.—Trono, 1973d:16, pl. 9: fig. 2.—Trono and Biña, 1973:5.—Westernhagen, 1973a:65.—Ortega, Alcala, and Reyes, 1974:187, 188.—Trono, 1974b:87.—Westernhagen, 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:153.—Cordero, 1976c:8, 9, 10; 1977a:94, pl. XIV:D; 1978a:30.—Trono, 1978:18.—Cordero, 1979b:276, 289.—Garcia, 1979:45 (table 1).—Puig and Cordero, 1979:38.—Liao and Sotto, 1980:99.—Reyes, 1980:131, pl. 7: fig. 5.—Meñez and Calumpong, 1981:381.—Trono and De Lara, 1981:15, pl. IX: fig. 1.—Saraya and Trono, 1982:34, pl. IV: fig. 4.—Hurtado–Ponce, 1983:133.—Cordero, 1984a:93; 1984b:65.—Marcos–Agngarayngay, 1984a:32, fig. 25.

*Mastophora licheniformis Decaisne, 1842:359, pl. 17: fig. 11 [type locality: Manila, Luzon].—Montagne, 1844a:662.—Velasquez, Trono, and Doty, 1975:153.

Mastophora decaisnei Kützing, 1849:697.—Martens, 1868:84–85.—Velasquez, Trono, and Doty, 1975:152.

Melobesia foliacea Kützing, 1843b:385 [type locality: Mariana Is.].—Dickie, 1876a:244 [with query].—Velasquez, Trono, and Doty, 1975:153.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Pangasinan, Manila, Batangas, Quezon, CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. SAMAR: Western Samar. LEYTE (Biliran I.), PANAY: Aklan, Capiz, Iloilo. GUIMARAS. NEGROS: Negros Oriental, CEBU (Mactan 1.). SIQUIJOR. MINDANAO: Zamboanga. PALAWAN, SULU: Sulu (Siasi I.), Tawitawi

NOTE.—The synonymy was proposed by Setchell (1943:129). Mastophora decaisnei Kützing is a superfluous and hence illegitimate substitute name for Mastophora licheniformis Decaisne.

Mesophyllum Lemoine

Mesophyllum erubescens (Foslie) Lemoine

Lithothamnion erubescens Foslie, 1900a:9 [type locality: Ilha Fernando de Noronha, Brazil].—Foslie, 1904:31.—Velasquez, 1971:448, fig. 29.—Velasquez et al., 1973:27.

Mesophyllum erubescens (Foslie) Lemoine, 1928:252.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. MINDORO: Oriental Mindoro. MINDANAO: Davao. SULU: Tawitawi (Sangasiapu I.).

Mesophyllum imbricatum (Dickie) Adey

Lithothamnion imbricatum Dickie, 1877:486 [type locality: Tahiti].

Mesophyllum imbricatum (Dickie) Adey, 1970:24.

Lithothamnion dickiei Foslie, 1900a:7 [including Philippine record].—Foslie, 1904:30.

Lithothamnion polymorphum [misapplied name fide Foslie, 1900a:7].—Dickie, 1876b:455.—Piccone, 1889:45, 53.—Velasquez, Trono, and Doty, 1975:152.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

NOTE.—Lithothamnion dickiei Foslie is a superfluous and hence illegitimate substitute name for L. imbricatum Dickie.

Mesophyllum pulchrum (Weber-van Bosse and Foslie) Lemoine

Lithothamnion pulchrum Weber-van Bosse and Foslie in Foslie, 1901c:3 [type locality: Sailus Besar, Great Paternoster Is., Indonesia].—Foslie, 1904:36, pl. IV: fig. 11.—Velasquez, Trono, and Doty, 1975:152.

Mesophyllum pulchrum (Weber-van Bosse and Foslie) Lemoine, 1928:252.

PHILIPPINE DISTRIBUTION.—SULU.

Mesophyllum siamense (Foslie) Adey

Lithothamnion siamense Foslie, 1901b:19 [type locality: various, all in Ko Chang Archipelago, Thailand].—Foslie, 1904:10, pl. I: figs. 1, 2.—De Toni, 1924:611.—Velasquez, Trono, and Doty, 1975:152.

Mesophyllum siamense (Foslie) Adey, 1970:26.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Tongquil I.).

NOTE.—Foslie (1901b:19) originally described Lithothamnion siamense as comprising two formae, f. minutum and f. simulans. Upon elevating the latter to specific rank, Foslie (1904:16) lectotypified L. siamense with f. minutum, which he then called f. typicum. In the same work, Foslie (1904:10) described L. siamense f. pseudoramosum. Tongquil I. is in the list of localities for the species, but to which forma it pertains is not indicated.

Mesophyllum simulans (Foslie) Lemoine

Lithothamnion siamense Foslie f. simulans Foslie, 1901b:19 [type locality: Ko Sarlak, Ko Chang Archipelago, Thailand].

Lithothamnion simulans (Foslie) Foslie, 1904:16, pl. I: figs. 21–25 [including Philippine record].—Velasquez, Trono, and Doty, 1975:152.

Mesophyllum simulans (Foslie) Lemoine, 1928:252.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I.).

Neogoniolithon Setchell and Mason

Neogoniolithon frutescens (Foslie) Setchell and Mason

Goniolithon frutescens Foslie, 1900b:9 [type locality: Funafuti, Ellice Is. (Tuvalu)].—Foslie, 1904:53.

Neogoniolithon frutescens (Foslie) Setchell and Mason, 1943:91.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I.).

Neogoniolithon megalocystum (Foslie) Setchell and Mason

Goniolithon megalocystum Foslie, 1904:48, fig. 20, pl. IX: figs. 8, 9 [including Philippine record] [type locality: Karkaralong Is., Indonesia].—De Toni, 1924:663.

Neogoniolithon megalocystum (Foslie) Setchell and Mason, 1943:90.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I.).

Phymatolithon Foslie

Phymatolithon has earlier taxonomic synomyms, but has been proposed for conservation by Woelkerling and Irvine (1986).

Phymatolithon calcareum (Pallas) Adey and McKibbin

Millepora calcarea Pallas, 1766:265 [syntype localities: North Atlantic Ocean and Mediterranean Sea],

Lithothamnion calcareum (Pallas) J.E. Areschoug, 1852:523.—Velasquez, 1953b:206.—Velasquez, Trono, and Doty, 1975:152.

Phymatolithon calcareum (Pallas) Adey and McKibbin, 1970:100.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Phymatolithon purpureum (P. Crouan and H. Crouan) Woelkerling and L. Irvine

Lithothamnion purpureum P. Crouan and H. Crouan, 1867:150, pl. 20: fig. 133 bis:1–5 [lectotype locality: Mingant, near Brest, Finistère, France fide Woelkerling and Irvine, 1986:69, 71].

Phymatolithon purpureum (P. Crouan and H. Crouan) Woelkerling and L. Irvine, 1986:71.

Lithothamnion polymorphum [misapplied name; see Note].—Piccone, 1889:51, 59.

PHILIPPINE DISTRIBUTION.—Locality not specified.

NOTE.—Millepora polymorpha Linnaeus (1767:1285) is a superfluous and hence illegitimate substitute name for M.calcarea Pallas (1766:265). Nonetheless, the two names have traditionally been applied to separate species. According to Woelkerling and Irvine (1986), the species traditionally but incorrectly known as Lithothamnion polymorphum (Linnaeus) J.E. Areschoug (1852:524) or Phymatolithon polymorphum (Linnaeus) Foslie (1898:4) is conspecific with L. purpureum P. Crouan and H. Crouan.

Porolithon (Foslie) Foslie

Porolithon onkodes (Heydrich) Foslie

Lithothamnion onkodes Heydrich, 1897a:6, pl. I: fig. 11 [type locality: Tami I., Papua New Guinea].

Lithophyllum onkodes (Heydrich) Heydrich, 1901:533.—Foslie, 1904:57.

Porolithon onkodes (Heydrich) Foslie, 1909:57 [“oncodes”].

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I.).

Sporolithon Heydrich

Sporolithon erythraeum (Rothpletz) Kylin

Lithothamnion erythraeum Rothpletz, 1893:5 [type locality: Red Sea].

Sporolithon erythraeum (Rothpletz) Kylin, 1956:205.

Sporolithon ptychoides Heydrich f. durum Heydrich, 1897b:67, pl. III figs. 20–23 [“dura”] [type locality: Tor, Sinai Peninsula, Egypt].

Archaeolithothamnion erythraeum (Rothpletz) Foslie f. durum (Heydrich) Foslie, 1900d:8 [“dura”].—Foslie, 1904:38, pl. V: figs. 8, 9.—Velasquez, Trono, and Doty, 1975:128 [without designation of forma].

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

TAXON OF UNCERTAIN VALUE

The following infraspecific taxon is related to Sporolithon erythraeum but has not been formally transferred to it.

Archaeolithothamnion erythraeum (Rothpletz) Foslie f. molle (Heydrich) Foslie

Sporolithon ptychoides Heydrich f. molle Heydrich, 1897b:67, pl. III: figs. 15–19 [“mollis”] [type locality: Tor, Sinai Peninsula, Egypt].

Archaeolithothamnion erythraeum (Rothpletz) Foslie f. molle (Heydrich) Foslie, 1900d:8 [“mollis”].—Foslie, 1904:38, pl. VI: figs. 2–4, 7, 9.—Velasquez, Trono, and Doty, 1975:128 [without designation of forma].

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I., Tongquil I.), Tawitawi (Pearl Bank, Sangasiapu I.).

NOTE.—When establishing Sporolithon ptychoides, Heydrich divided the species into two formae, durum and molle. Later, Heydrich (1897c:415–416) elevated f. molle to specific rank and thus lectotypified S. ptychoides with f. durum. Foslie (1900d:8) proposed the conspecificity of Lithothamnion erythraeum and both forms of Sporolithon ptychoides. We are not convinced of the usefulness of giving taxonomic recognition to forma molle.

Sporolithon schmidtii (Foslie) Gordon, Masaki, and Akioka

Archaeolithothamnion schmidtii Foslie, 1901b:16 [type locality: Ko Kahdat, Ko Chang Archipelago, Thailand].—Foslie, 1904:43, pl. VIII: fig. 16.—De Toni, 1924:603.—Foslie, 1929:pl. XLIV: fig. 16.—Velasquez, Trono, and Doty, 1975:128.

Sporolithon schmidtii (Foslie) Gordon, Masaki, and Akioka, 1976:250.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Pearl Bank).

*Sporolithon sibogae (Weber-van Bosse and Foslie) P.C. Silva, new combination

Archaeolithothamnion sibogae Weber-van Bosse and Foslie in Foslie, 1901c:3 [type locality: Pearl Bank, Tawitawi Prov., Sulu Archipelago].—Foslie, 1904:41, pl. VII.—De Toni, 1924:604.—Foslie, 1929:pl. XLIII: figs. 5–15.—Velasquez, Trono, and Doty, 1975:128.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.) Tawitawi (Pearl Bank).

Sporolithon timorense (Foslie) P.C. Silva, new combination

Archaeolithothamnion timorense Foslie, 1904:42, pl. VIII: figs. 1–14 [lectotype locality: Sailus Besar, Great Paternoster Is., Indonesia fide Adey and Lebednik, 1967:85] [including Philippine record].—Velasquez, Trono, and Doty, 1975:128.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

Order GIGARTINALES

Family RHIZOPHYLLIDACEAE

Portieria Zanardini

The complicated nomenclature of this genus is discussed in the appended Nomenclatural Notes.

Portieria hornemannii (Lyngbye) P.C. Silva, new combination

Desmia hornemannii Lyngbye, 1819:35, pl. 7:C [“hornemanni”] [type locality: probably Red Sea].—Reyes, 1972:154.—Trono, 1973d:15, pl. 10: fig. 6.—Trono and Biña, 1973:5.—Trono, 1974b:88.—Velasquez, Trono, and Doty, 1975:140.—Cordero, 1976c:8.—Trono and Tuason, 1978:15.—Cordero, 1979b:276.—Reyes, 1980:132, pl. 8: fig. 1.—Trono and Ganzon-Fortes, 1980:65, fig. [s.n.].—Trono and De Lara, 1981:15, pl. IX: fig. 3.—Saraya and Trono, 1982:33, pl. III: fig. 4.

Chondrococcus hornemannii (Lyngbye) Schmitz, 1895:170 [“hornemanni”].—Meñez, 1961:73.—Villones and Magdamo, 1968:28, fig. 25.—Cornejo and Velasquez, 1972:182.—Westernhagen, 1973a:64.—Ortega, Alcala, and Reyes, 1974:188.—Westernhagen, 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:135.—Cordero, 1976c:8, 10, 11; 1977a:96, fig. 53; 1978a:26; 1979b:289.—Garcia, 1979:44 (table 1).—Puig and Cordero, 1979:39.—Liao and Sotto, 1980:98.—Meñez and Calumpong, 1981:381.—Hurtado-Ponce, 1983:133.—Cordero, 1984a:92; 1984b:66.—Marcos-Agngarayngay, 1984a:28, fig. 22 [captions for figs. 21 and 22 interchanged].

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Cavite, Batangas, Quezon, CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SAMAR: Eastern Samar. LEYTE (Biliran I.), PANAY: Aklan, Iloilo. NEGROS: Negros Oriental. CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO. PALAWAN, SULU: Sulu (Siasi I.).

Portieria japonica (Harvey) P.C. Silva, new combination

Desmia japonica Harvey, 1860a:331 [type locality: Shimoda, Shizuoka Prefecture, Japan].

Chondrococcus japonicus (Harvey) De Toni, 1895a:39.—Cordero, I977a:97; 1984a:93.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Bataan, Cavite, Quezon, MINDORO: Oriental Mindoro. NEGROS: Negros Oriental.

Family POLYIDACEAE

Rhodopeltis Harvey

Rhodopeltis borealis Yamada

Rhodopeltis borealis Yamada, 1931a:75, pl. XIX: fig. 1 [syntype localities: Ryukyu-retto, Japan; Kotosho (Hung-t’ou), Taiwan].—Meñez, 1961:72.—Domantay, 1962:293.—Velasquez, Trono, and Doty, 1975:161.—Cordero, 1976c:8; 1977a:98, fig. 59, pl. XII: c; 1978a:25.—Saraya and Trono, 1982:33, pl. IV: fig. 3.—Hurtado-Ponce, 1983:134.—Marcos-Agngarayngay, 1984a:28, fig. 23.

PHILIPPINE DISTRIBUTION—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan.

Rhodopeltis gracilis Yamada and Tanaka

Rhodopeltis gracilis Yamada and Tanaka in Yamada, 1935:30, fig. 1, pl. 15: fig. 2 [syntype localities: Chichi-jima, Ogasawara-gunto (Bonin Is.), Japan; Kotosho (Hung-t’ou), Taiwan].—Meñez, 1961:72, pl. 7: figs. 78–80.—Velasquez, Trono, and Doty, 1975:161.—Cordero, 1976c:9, 10; 1977a:98, figs. 54, 55; 1978a:26.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan LEYTE.

Family NEMASTOMATACEAE

Titanophora (J. Agardh) J. Feldmann

Titanophora incrustans (J. Agardh) Børgesen

Halymenia incrustans J. Agardh, 1885:15 [type locality: Florida, USA].

Titanophora incrustans (J. Agardh) Børgesen, 1949:4.—Velasquez et al., 1973:30, pl. 13: fig. 60.

PHILIPPINE DISTRIBUTION.—LUZON: Bataan, Batangas.

Titanophora weberae Børgesen

Titanophora weberae Børgesen, 1943:39, fig. 13 [type locality: Salee Strait, Irian Barat, Indonesia].—Domantay, 1962:293.—Velasquez, Trono, and Doty, 1975:165.—Cordero, 1977a:118, figs. 91–93, 95, pl. XIX:A.—Meñez and Calumpong, 1981:381.—Hurtado-Ponce, 1983:139.—Cordero, 1984b:65.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas, Quezon, MINDORO: Oriental Mindoro. SAMAR: Eastern Samar. CENTRAL VISAYAS. PALAWAN.

Family SEBDENIACEAE

Sebdenia (J. Agardh) Berthold

Sebdenia limensis (Kützing) Howe

Euhymenia limensis Kützing, 1849:743 [type locality: Callao, Peru].—Martens, 1868:90–91 [with query].—Velasquez, Trono, and Doty, 1975:143.

Sebdenia limensis (Kützing) Howe, 1914:160.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Sebdenia yamadae Okamura and Segawa

Sebdenia yamadae Okamura and Segawa in Segawa, 1938:144, fig. 6, pl. 34: fig. 1 [“yamadai”] [syntype localities: various, all in Japan].—Cordero, 1977a:150, fig. 143, pl. XXI:C.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Family GRACILARIACEAE

Ceratodictyon Zanardini

Ceratodictyon spongiosum Zanardini

Ceratodictyon spongiosum Zanardini, 1878:37 [type locality: Aru Is., Indonesia].—Trono, 1972a:104; 1973d:19—Trono and Biña, 1973:9.—Cordero, 1979b:291.—Puig and Cordero, 1979:41.—Liao and Sotto, 1980:99.—Trono and Ganzon-Fortes, 1980:85, fig. [s.n.].—Meñez and Calumpong, 1981:381.—Saraya and Trono, 1982:42.—Trono and Ang, 1982:20.—Cordero, 1984a:100.—Marcos-Agngarayngay, 1984a:51, fig. 42.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Batangas, Quezon, MINDORO: Oriental Mindoro. PANAY: Aklan. LEYTE (Biliran I.), CEBU (Mactan I.). PALAWAN (Bugsuk I.), SULU: Tawitawi.

Gelidiopsis Schmitz

Gelidiopsis intricata (C. Agardh) Vickers

Sphaerococcus intricatus C. Agardh, 1822a:333 [syntype localities: Mauritius; Hawaiian Is.; “Ravak” (Lawak), Waigeo I., Moluccas, Indonesia].

Gelidiopsis intricata (C. Agardh) Vickers, 1905:61.—Reyes, 1972:152.—Trono, 1972a:104; 1973d:19.—Trono and Biña, 1973:3.—Velasquez, Trono, and Doty, 1975:145.—Cordero, 1976c:10; 1977a:l19, figs. 99, 100; 1978a:21.—Trono and Ganzon-Fortes, 1980:87, fig. [s.n.].—Meñez and Calumpong, 1981:381.—Trono and De Lara, 1981:18, pl. XI: fig. 1—Calumpong, 1982:145.—Saraya and Trono, 1982:30.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Batangas, Quezon, MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. NEGROS: Negros Oriental, SIQUIJOR. SULU: Tawitawi.

Gelidiopsis repens (Kützing) Weber-van Bosse

Gelidium repens Kützing, 1868:21, pl. 60: figs, a, b [type locality: New Caledonia].

Gelidiopsis repens (Kützing) Weber-van Bosse, 1928:425.—Cordero, 1976c:8, 9, 10; 1977a:121, figs. 98, 267; 1978a:22, 53; 1980b:54, fig. 19B, pl. 36; 1984C:55.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Zambales. MINDORO: Oriental Mindoro. MASBATE. SAMAR: Western Samar, Eastern Samar. PANAY: Aklan. NEGROS: Negros Oriental, SIQUIJOR. PALAWAN.

Gelidiopsis variabilis (J. Agardh) Schmitz

Gelidium variabile J. Agardh, 1851 [1851–1863]:468 [type locality: India].

Gelidiopsis variabilis (J. Agardh) Schmitz, 1895:148.—Cordero, 1977a:122 [“Gelidiopsis repens” in error].

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Gracilaria Greville

Gracilaria arcuata Zanardini

Gracilaria arcuata Zanardini, 1858:265, pl. V: fig. 2 [type locality: Aqaba, Jordan].—Weber-van Bosse, 1928:429, fig. 173.—Velasquez, Trono, and Doty, 1975:146.—Cordero, 1977a:124, fig. 101, pl. XX:B; 1980b:55, fig. 18, pl. [49].—Trono and Fortes, 1980:73.—Trono and Ganzon-Fortes, 1980:95, fig. [s.n.].—Ganzon-Fortes, 1981:22.—Meñez and Calumpong, 1981:381.—Calumpong, 1982:145.—Trono and Fortes, 1982:152.—Hurtado-Ponce, 1983:140.—Cordero, 1984a:100; 1984c:55.—Hurtado-Ponce, 1984:180.—Marcos-Agngarayngay, 1984b:126.—Tungpalan, 1984:138.—Abbott, 1985:89, fig. 17.—Trono and Ganzon-Fortes, 1985:66.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Norte, Batangas. MINDORO: Oriental Mindoro. MASBATE. NEGROS: Negros Occidental (incl. Ilacaon I., Suyac I.), Negros Oriental (incl. Apo I.), SIQUIJOR. MINDANAO: Davao. PALAWAN, SULU: Sulu (Tongquil I.).

Gracilaria arcuata Zanardini var. snackeyi Weber-van Bosse

Gracilaria arcuata Zanardini var. snackeyi Weber-van Bosse, 1928:430, fig. 173 [type locality: Macassar, Celebes, Indonesia].—Trono, Azanza-Corrales, and Manuel, 1983:32, fig. 7.—Abbott, 1985:89.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas.

Gracilaria blodgettii Harvey

Gracilaria blodgettii Harvey, 1853:111 [type locality: Key West, Florida, USA].—Cordero, 1977a:124, figs. 102, 103, 105, 112, pl. XIX:B; 1979b:292; 1980b:56, fig. 19A.—Meñez and Calumpong, 1981:381.—Calumpong, 1982:145.—Meñez, Phillips, and Calumpong, 1983:18.—Cordero, 1984c:55.—Marcos-Agngarayngay, 1984a:46, fig. 38.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, MASBATE. SAMAR: Western Samar. PANAY: Aklan. NEGROS: Negros Oriental (Apo I.), CEBU. SIQUIJOR. PALAWAN.

Gracilaria bursa-pastoris (S.G. Gmelin) P.C. Silva

Fucus bursa-pastoris S.G. Gmelin, 1768:121, pl. VIII: fig. 3 [type locality: Mediterranean Sea].

Gracilaria bursa-pastoris (S.G. Gmelin) P.C. Silva, 1952a:265.—Cordero, 1977a:125, fig. 106.

Sphaerococcus compressus C. Agardh, 1822a:308 [syntype localities: Cádiz, Spain; Oneglia, Liguria, Italy].

Gracilaria compressa (C. Agardh) Greville, 1830:liv.—Howe, 1932:169.—Meñez, 1961:74.—Velasquez, Trono, and Doty, 1975:146.—Trono and Fortes, 1980:73.—Ganzon-Fortes, 1981:22.—Trono and Fortes, 1982:152.—Abbott, 1985:89.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas, Quezon, Albay. MINDORO: Oriental Mindoro. SAMAR: Eastern Samar. PANAY. NEGROS: Negros Oriental. PALAWAN.

NOTE.—The synonymy was proposed by J. Agardh (1852 [1851–1863]:593).

Gracilaria canaliculata Sonder

Sphaerococcus canaliculars Kützing, 1868:29, pl. 82: figs, d, e [type locality: Wagap, New Caledonia].

Gracilaria canaliculata Sonder, 1871:56.—Cantoria, Valenzuela, and Velasquez, 1951:187, fig. 2.—Cantoria, Velasquez, and Valenzuela, 1951:297, 298.—Velasquez, Trono, and Doty, 1975:146.—Abbott, 1985:89, fig. 43.

Corallopsis opuntia]. Agardh, 1872:40 [type locality: Sri Lanka].—Westernhagen, 1973a:65; 1974:112 (table 1).

Gracilaria crassa Harvey ex J. Agardh, 1876:417 [type locality: Sri Lanka].—Seale, 1911:309.—Wester, 1916:159; 1921:224; 1924:21.—G. Blanco, 1938:513.—Quisumbing, 1951:1010.—Montilla and Blanco, 1953:166.—Zaneveld, 1956:38; 1959:118.—Domantay, 1962:291.—Velasquez, Trono, and Doty, 1975:146.—Cordero, 1977a:127, figs. 108–110; 1978a:36.—Trono and Azanza–Corrales, 1979:26.—Cordero, 1980b:56, fig. 20, pl. [48].—Trono and Fortes, 1980:73.—Ganzon-Fortes, 1981:22.—Meñez and Calumpong, 1981:381.—Calumpong, 1982:145.—Cordero, 1982a:60, 61.—Saraya and Trono, 1982:41.—Trono and Fortes, 1982:152.—Marcos-Agngarayngay, 1984b:127.—Abbott, 1985:89.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, La Union, Pangasinan, Manila Bay. MINDORO: Oriental Mindoro. SAMAR: Western Samar. NEGROS: Negros Oriental (incl. Apo I.), CEBU (Mactan I.). SIQUIJOR. MINDANAO: Davao.

NOTE.—The synonymy was proposed by Newton (1953:412). Sphaerococcus canaliculatus Kützing, the intended basionym of Gracilaria canaliculata, is a later homonym of S. canaliculatus C. Agardh (1822a:260) and hence not priorable. Gracilaria canaliculata Sonder is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Gracilaria coronopifolia J. Agardh

Gracilaria coronopifolia J. Agardh, 1852 [1851–1863]:592 [type locality: Oahu, Hawaiian Is.].—Galutira and Velasquez, 1964:508, pl. 5: fig. 14; pl. 9: fig. 34.—Velasquez, 1972:63.—Cordero, 1977a:127, figs. 104, 104a, pl. XX:A; 1978a:35.—Trono and Azanza-Corrales, 1979:26.—Velasquez, 1979b:231.—Liao and Sotto, 1980:99.—Trono and Fortes, 1980:73.—Trono and Ganzon-Fortes, 1980:89, fig. [s.n.].—Ganzon- Fortes, 1981:22.—Guzman, 1981:42, 44 [“coronipifolia”].—Laserna et al., 1981:443.—Trono, 1981b:55 (table 1).—Trono and Azanza-Corrales, 1981:743.—Trono and De Lara, 1981:18.—Cordero, 1982a:60, 61.—Luistro, Cajipe, and Laserna, 1982:46.—Trono and Ang, 1982:19.—Trono and Fortes, 1982:152.—Hurtado-Ponce, 1983:140.—Trono, Azanza-Corrales, and Manuel, 1983:20, figs. 2, 9b.—Cordero, 1984a:101; 1984b:66; 1984c:55.—Hurtado-Ponce, 1984:180.—Marcos-Agngarayngay, 1984b:127.—Tungpalan, 1984:138.—Abbott, 1985:89, fig. 46.

Gracilaria lichenoides (Lamouroux) Greville f. coronopifolia (J. Agardh) May, 1948:37.—Zaneveld, 1956:39; 1959:119.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Ilocos Sur, Pangasinan, Cavite, Batangas, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. MASBATE. SAMAR: Eastern Samar. NEGROS: Negros Oriental, CEBU. SIQUIJOR. MINDANAO: Zamboanga. PALAWAN (incl. Bugsuk I.), SULU: Sulu (Lapac I., Siasi I.).

Gracilaria cylindrica Børgesen

Gracilaria cylindrica Børgesen, 1920:375, figs. 364, 365 [type locality: St. Jan (St. John), Virgin Is.].—Westernhagen, 1973a:65; 1974:112 (table 1).—Reyes, 1980:135, pl. 10, fig. 1.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.), SIQUIJOR.

Gracilaria damaecornis J. Agardh

Gracilaria damaecornis J. Agardh, 1852 [1851–1863]:597 [type locality: Atlantic North America].—Westernhagen, 1973a:65; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Gracilaria disticha (J. Agardh) J. Agardh

Sphaerococcus distichus J. Agardh, 1837:172 [type locality: Ethiopia].

Gracilaria disticha (J. Agardh) J. Agardh, 1852 [1851–1863]:594.—Abbott, 1985:89, fig. 56.

PHILIPPINE DISTRIBUTION.—ROMBLON (Sibuyan I.).

Gracilaria edulis (S.G. Gmelin) P.C. Silva

Fucus edulis S.G. Gmelin, 1768:113 [syntype localities: various, all in “India Orientalis” (Indonesia)].

Gracilaria edulis (S.G. Gmelin) P.C. Silva, 1952a:293.—Cordero, 1977a:128, fig. 111.—Hurtado-Ponce, 1983:140.—Cordero, 1984a:101.—Hurtado-Ponce, 1984:180.—Abbott, 1985:89, figs. 55, 58.

Fucus lichenoides Turner, 1809:124, pl. II8: figs, a–c, e–i [superfluous name].

Gigartina lichenoides Lamouroux, 1813:137.

Sphaerococcus lichenoides (Lamouroux) C. Agardh, 1817:XVII.—Martens, 1868:31, 94–95.—Velasquez, Trono, and Doty, 1975:163.

Gracilaria lichenoides (Lamouroux) Greville, 1830:liv.—Seale, 1911:309.—Wester, 1916:159; 1921:224; 1924:21.—Collado, 1926:129 [with query].—Howe, 1932:169.—G. Blanco, 1938:513.—Cantoria, Valenzuela, and Velasquez, 1951:187, fig. 3.—Cantoria, Velasquez, and Valenzuela, 1951:297, 298 [f. lichenoides].—Quisumbing, 1951:50.—Montilla and Blanco, 1953:166.—Zaneveld, 1956:38; 1959:119.—Domantay, 1962:290.—De Leon and Domantay, 1971:5.—Velasquez, Trono, and Doty, 1975:146.—Trono and Fortes, 1980:73.—Ganzon-Fortes, 1981:22.—Cordero, 1982a:60, 61.—Trono and Fortes, 1982:153.—Abbott, 1985:89.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (incl. Babuyan Is.), Ilocos, La Union, Pangasinan, Manila Bay, Sorsogon. MINDORO: Oriental Mindoro. PANAY. MINDANAO: Zamboanga.

NOTE.—Fucus lichenoides Turner, the intended basionym of Gracilaria lichenoides (and other combinations), is a later homonym of F. lichenoides S.G. Gmelin (1768:120, pl. VIII: figs. 1, 2) and hence not priorable. Gigartina lichenoides Lamouroux is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN. Fucus lichenoides Turner is not only a later homonym, but also superfluous, since its protologue included the citation of F. edulis S.G. Gmelin.

Gracilaria eucheumoides Harvey

Gracilaria eucheumoides Harvey, 1860a:331 [“eucheumioides”] [syntype localities: various, all in Ryukyu-retto, Japan].—Dickie, 1876a:243.—Seale, 1911:309.—Wester, 1916:159; 1921:224; 1924:21.—G. Blanco, 1938:513.—Quisumbing, 1951:1010.—Montilla and Blanco, 1953 166.—Zaneveld, 1956:38; 1959:118.—Domantay, 1962:290.—De Leon and Domantay, 1971:5, 9.—Cornejo and Velasquez, 1972 182, 183.—Reyes, 1972:155.—Trono, 1972a:104.—Bersamin et al. 1973:187.—Trono, 1973d:18, pl. 9: fig. 1.—Trono and Biña 1973:9.—Velasquez et al., 1973:31, pl. 13: fig. 63.—Westernhagen 1973a:65—Ortega, Alcala, and Reyes, 1974:186, 187, 188.—Trono 1974b:90; 1974e:18.—Westernhagen, 1974:112 (table 1).—Velasquez Trono, and Doty, 1975:146.—Cordero, 1976c:10; 1977a:129, figs 113, 113a, 114, pls. X, XX; 1978a:37; 1979b:292, fig. 4.—Puig and Cordero, 1979:40.—Cordero, 1980b:57, fig. 21, pl. 37.—Liào and Sotto, 1980:99.—Reyes, 1980:135, pl. 9: fig. 5.—Trono and Fortes, 1980:73.—Trono and Ganzon-Fortes, 1980:91, fig. [s.n.].—Chan, 1981:387.—Ganzon-Fortes, 1981:22.—Guzman, 1981:43, 51.—Meñez and Calumpong, 1981:381.—Trono and De Lara, 1981:18.—Calumpong, 1982:145.—Cordero, 1982a:60, 61.—Saraya and Trono, 1982:40, pl. VI: fig. 4.—Trono and Ang, 1982:19.—Trono and Fortes, 1982:153.—Hurtado-Ponce, 1983:141.—Trono, Azanza-Corrales, and Manuel, 1983:27, fig. 5.—Cordero, 1984a:101.—Hurtado-Ponce, 1984:180.—Marcos-Agngarayngay, 1984a:48, fig. 39; 1984b:127.—Tungpalan, 1984:138.—Abbott, 1985:89.—Trono and Ganzon-Fortes, 1985:66.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Ilocos Sur, La Union, Pangasinan, Bataan, Batangas, Quezon, Sorsogon. MARINDUQUE. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SAMAR: Northern Samar, Western Samar, Eastern Samar. LEYTE (incl. Biliran I.), PANAY: Aklan. NEGROS: Negros Occidental (incl. Ilacaon I.), Negros Oriental. CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.), Misamis Occidental. PALAWAN (incl. Bugsuk I.), SULU: Sulu (Siasi I.), Tawitawi.

Gracilaria foliifera (Forsskål) Børgesen

Fucus foliifer Forsskål, 1775:191 [type locality: Mokha, Yemen].

Gracilaria foliifera (Forsskål) Børgesen, 1932:7.

Fucus lacinulatus Vahl, 1802:39 [type locality: St. Croix, Virgin Is.].

Gracilaria lacinulata (Vahl) Howe, 1920:562.—Weber-van Bosse, 1928:434.—Velasquez, Trono, and Doty, 1975:146.—Trono and Fortes, 1980:73.—Ganzon-Fortes, 1981:22.—Trono and Fortes, 1982:153.—Abbott, 1985:89.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

NOTE.—The synonymy was proposed by Børgesen (1932:7).

Gracilaria foliifera (Forsskål) Børgesen f. aeruginosa (Turner) Børgesen

Fucus aeruginosus Turner, 1811:29, pl. 147 [syntype localities: Portugal; Red Sea].

Gracilaria foliifera (Forsskål) Børgesen f. aeruginosa (Turner) Børgesen, 1938:226.

Gracilaria multipartita (Clemente y Rubio) Harvey var. aeruginosa (Turner) J. Agardh, 1852 [1851–1863]:601.—Grunow, 1867:83.

PHILIPPINE DISTRIBUTION.—LUZON.

Gracilaria gigas Harvey

Gracilaria gigas Harvey, 1860a:331 [type locality: Shimoda, Shizuoka Prefecture, Japan].—Kraft, 1972:332.—Trono, Azanza-Corrales, and Manuel, 1983:31, figs. 6, 11a.—Abbott, 1985:89, fig. 39.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Sorsogon.

Gracilaria incurvata Okamura

Gracilaria incurvata Okamura, 1931 [1929–1932]:41, pl. CCLXXIII: figs. 1–6 [syntype localities: various, all in Japan].—Cordero, 1984a:101.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Gracilaria minor (Sonder) Durairatnam

Corallopsis salicornia (C. Agardh) Greville var. minor Sonder, 1871:56, pl. III: figs. 6–11 [type locality: Cape York, Queensland, Australia].

Corallopsis minor (Sonder) J. Agardh, 1876:409.—Piccone, 1886:68, 90.—Velasquez, Trono, and Doty, 1975:138.

Gracilaria minor (Sonder) Durairatnam, 1961:64.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

Gracilaria papenfussii Abbott

Gracilaria papenfussii Abbott, 1983:562, figs. 1–3 [type locality: La Jolla, California, USA].

Gracilaria andersonii [misapplied name fide Abbott, 1983:562].—Cordero, 1977a:123, figs. 96, 97.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

*Gracilaria salicornia (C. Agardh) Dawson

Sphaerococcus salicornia C. Agardh, 1820a: pl. VIII [type locality: said to be Unalaska, Aleutian Is., but probably Manila, Luzon].

Corallopsis salicornia (C. Agardh) Greville, 1830:liii.—Cantoria, Velasquez, and Valenzuela, 1951:296, footnote.

Gracilaria salicornia (C. Agardh) Dawson, 1954a:4, fig. 3 [including Philippine record].—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Galutira and Velasquez, 1964:506, pl. 5: figs. 15a, 15b; pl. 8: figs. 32a, 32b.—Velasquez, 1968a:121, fig. 7.—Villones and Magdamo, 1968:28, fig. 28.—Velasquez, 1971:448, fig. 31.—Reyes, 1972:155.—Trono, 1972a:104.—Velasquez, 1972:63.—Trono, 1973d:18, pl. 9: fig. 3.—Trono and Biña, 1973:9.—Velasquez et al., 1973:30, pl. 13: fig. 62.—De Leon, 1974:31, 33, photo [s.n.], fig. [s.n.].—Ortega, Alcala, and Reyes, 1974:185.—Trono, 1974b:90; 1974e:18.—Velasquez, Trono, and Doty, 1975:146.—Cordero, 1977a:132, figs. 116–118, pl. XIX:C.—Tahil, 1978:52.—Vannajan and Trono, 1978:22, fig. 24.—Cordero, 1979b:276.—Puig and Cordero, 1979:41.—Trono and Azanza-Corrales, 1979:26.—Velasquez, 1979b:231.—Cordero, 1980b:57, fig. 22, pl. 38.—Fortes and Trono, 1980:60.—Liao and Sotto, 1980:99.—Reyes, 1980:135, pl. 9: fig. 4.—Trono and Fortes, 1980:73.—Trono and Ganzon-Fortes, 1980:93, fig. [s.n.].—Trono, Velasquez, and Guevarra, 1980:78.—Ganzon-Fortes, 1981:22.—Guzman, 1981:43.—Meñez and Calumpong, 1981:381.—Trono, 1981b:55 (table 1).—Trono and Azanza-Corrales, 1981:743.—Trono and De Lara, 1981:18.—Calumpong, 1982:145.—Cordero, 1982a:60, 61.—Trono and Fortes, 1982:153.—Hurtado-Ponce, 1983:141.—Trono, Azanza-Corrales, and Manuel, 1983:23, figs. 3, 4b.—Cordero, 1984a:102; 1984c:55.—Hurtado-Ponce, 1984:180.—Marcos-Agngarayngay, 1984a:48, fig. 40; 1984b:127.—Abbott, 1985:89, fig. 42.—Trono and Ganzon-Fortes, 1985:66.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Norte, Pangasinan, Zambales, Bataan, Rizal, Cavite, Quezon, Batangas, Sorsogon. MINDORO: Occidental Mindoro (incl. Lubang Is.), Oriental Mindoro. MASBATE. SAMAR: Western Samar, Eastern Samar. LEYTE (incl. Biliran I.). PANAY: Aklan, Iloilo. NEGROS: Negros Occidental (incl. Ilacaon I., Suyac I.), Negros Oriental (incl. Apo I.), CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.), Misamis Occidental, Surigao del Sur, Davao (incl. Samal I.). PALAWAN (incl. Cuyo I.), SULU: Sulu (Siasi I.).

NOTE.—The type specimen of Sphaerococcus salicornia was said by Agardh to have been collected by Chamisso at Unalaska (during the voyage of the Rurik), but Dawson (1954a:5) showed that the provenance probably was Manila. The Rurik remained at Manila from 17 December 1817 to 29 January 1818, following its voyage from Unalaska by way of Hawaii and Guam.

Gracilaria spinigera Dawson

Gracilaria spinigera Dawson, 1949a:24, pl. 8: figs. 1–3, 5; pl. 9: figs. 1–3 [type locality: Guaymas, Sonora, Mexico].—Cordero, 1977a:134, fig. 119, pl. XXI:A; 1980b:58, fig. 23, pl. [52]; 1984a:102.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Gracilaria spinulosa (Okamura) Chang and Xia

Rhodymenia spinulosa Okamura, 1934 [1933–1942]:33, pl. 318: figs. 1–6 [type locality: Tainan, Taiwan].—Meñez, 1961:78.—Velasquez, Trono, and Doty, 1975:162—Meñez and Calumpong, 1981:381.

Gracilaria spinulosa (Okamura) Chang and Xia, 1976:148.

Gracilaria purpurascens J. Agardh, 1885:63 [type locality: Sri Lanka].—Cordero, 1977a:130, fig. 115.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. NEGROS: Negros Oriental, CENTRAL VISAYAS.

NOTE.—Rhodymenia spinulosa was referred to Gracilaria by Yamada (1941a:204) as a form of G. purpurascens J. Agardh. The latter name, however, is a later homonym of G. purpurascens Greville (1830:liv) and must be replaced.

Gracilaria textorii (Suringar) De Toni

Sphaerococcus textorii Suringar, 1867:259 [type locality: Japan].

Gracilaria textorii (Suringar) De Toni, 1895a:27.—Cordero, 1977a:135; 1980b:59, pl. [58] [G. textorii prox.]; 1984a:102.—Marcos-Agngarayngay, 1984b:128 [G. textorii prox.].—Tungpalan, 1984:138.—Abbott, 1985:90, fig. 26.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Gracilaria turgida Dawson

Gracilaria turgida Dawson, 1949a:14, pl. 21; pl. 24: fig. 1 [type locality: Newport-Balboa, Orange County, California, USA].—Saraya and Trono, 1982:40, pl. VI: fig. 3.—Abbott, 1985:90.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Gracilaria venezuelensis W.R. Taylor

Gracilaria venezuelensis W.R. Taylor, 1942:110, pl. 3: fig. 10[type locality: Isla Cubagua, Venezuela].—Westernhagen, 1973a:65; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Gracilaria verrucosa (Hudson) Papenfuss

Fucus verrucosus Hudson, 1762:470 [type locality: England].

Gracilaria verrucosa (Hudson) Papenfuss, 1950:195.—Galutira and Velasquez, 1964:507, pl. 4: fig. 13; pl. 8: fig. 33.—Velasquez, 1968a:121, fig. 8.—Villones and Magdamo, 1968:29, fig. 29.—De Leon and Domantay, 1971:5, 9.—Velasquez, 1971:448, fig. 32.—Cornejo and Velasquez, 1972:180.—Reyes, 1972:155.—Velasquez, 1972:63.—Cordero, 1973b:33.—Velasquez et al., 1973:30, pl. 13: fig. 61.—Westernhagen, 1973a:65.—Trono, 1974e:18.—Westernhagen, 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:146.—Cordero, 1976c:8, 10; 1977a:135; 1978a:37.—Vannajan and Trono, 1978:22, fig. 18.—Laserna et al., 1978:111.—Cordero, 1979b:276, 292.—Garcia, 1979:45 (table 1).—Puig and Cordero, 1979:41.—Trono and Azanza-Corrales, 1979:26.—Velasquez, 1979b:231.—Cordero, 1980b:59, pl. 39.—Liao and Sotto, 1980:99.—Reyes, 1980:135, pl. 9: fig. 6.—Trono and Fortes, 1980:74.—Trono, Velasquez, and Guevarra, 1980:78.—Ganzon-Fortes, 1981:22.—Guzman, 1981:42, 46, 47.—Laserna et al., 1981:443.—Meñez and Calumpong, 1981:381.—Trono, 1981b:55 (table 1).—Trono and Azanza-Corrales, 1981:743.—Calumpong, 1982:145.—Cordero, 1982a:60, 61.—Luistro, Cajipe, and Laserna, 1982:46.—Trono and Fortes, 1982:153.—Hurtado-Ponce, 1983:142.—Trono, Azanza-Corrales, and Manuel, 1983:17, figs. 1, 4a.—Cordero, 1984a:102.—Hurtado-Ponce, 1984:180.—Marcos- Agngarayngay, 1984a:50, fig. 41; 1984b:128.—Tungpalan, 1984:138.—Abbott, 1985:90, fig. 9.—Trono and Ganzon-Fortes, 1985:64, 66.

Fucus confervoides Linnaeus, 1763:1629 [type locality: England].

Ceramium confervoides Wiggers, 1780:91.

Sphaerococcus confervoides (Wiggers) Stackhouse, 1797 [1795–1801: xxiv.—Martens, 1868:94–95.—Velasquez, Trono, and Doty, 1975:163.

Gracilaria confervoides (Wiggers) Greville, 1830:liv.—Montagne, 1844a:662.—Seale, 1911:309.—Wester, 1916:158; 1921:224; 1924:21.—G. Blanco, 1938:512.—De Leon, Leyva, and Martinez-Pesigan, 1947:381.—Abagon, Buñag, and De Vera, 1951:41.—Quisumbing, 1951:1010.—Montilla and Blanco, 1953:166.—Velasquez, 1953a:100; 1953b:206.—Villaluz, 1953:186, 187.—Sulit, Salcedo, and Panganiban, 1956:280.—Zaneveld, 1956:36; 1959:117.—Domantay, 1968:25.—De Leon and Domantay, 1971:5, 9.—Bersamin et al., 1973:186.—Sulit, Salcedo, and Panganiban, 1973:177–182.—Westernhagen, 1973a:65; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:146.—Abbott, 1985:89.

*Fucus gulaman M. Blanco, 1837:839 [syntype localities: Tambobo (“Tambobon”) and Parañaque, Rizal Prov., Luzon] [probable taxonomic synonym].—Montilla and Blanco, 1953:166.—Velasquez, Trono, and Doty, 1975:144.

Fucus edulis [probable misapplied name fide Merrill, 1918:41].—M. Blanco, 1845:580; 1879:260.—Velasquez, Trono, and Doty, 1975:144.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Pangasinan, Bataan, Pampanga, Bulacan, Rizal, Manila, Cavite, Batangas, Quezon, Albay (Batan I.), MINDORO: Occidental Mindoro, Oriental Mindoro. SAMAR: Western Samar, Eastern Samar. LEYTE (Biliran I.), PANAY: Aklan. NEGROS: Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Zamboanga. PALAWAN, SULU.

NOTE.—The synonymy was proposed by Papenfuss (1950:195). Fucus confervoides Linnaeus, the intended basionym of Gracilaria confervoides (and other combinations), is a later homonym of F. confervoides Hudson (1762:474) and hence not priorable. Ceramium confervoides Wiggers is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN. Although the Tagalog word “gulaman” is applied to various red algae that are rich in hydrocolloids, Blanco’s description of F. gulaman appears to apply, at least in part, to the stringy Gracilaria currently called G. confervoides or G. verrucosa. Martens (1868:45–46) thought that it applied to two species, Sphaerococcus gelatinus (=Eucheuma gelatinum) and S. lichenoides (=Gracilaria edulis), while Seale (1911:309) and Quisumbing (1951:1007) thought that it applied to a species of Agardhiella. In the absence of authentic specimens, a definitive placement is impossible.

Gracilaria vieillardii P.C. Silva, new name

Sphaerococcus denticulatus Kützing, 1869:19, pl. 51: figs, e–g [type locality: New Caledonia].

Gracilaria denticulata (Kützing) Weber–van Bosse, 1928:432 [replaced name].—Cordero, 1976c:10; 1977a:128, fig. 109; 1978a:36.

PHILIPPINE DISTRIBUTION.—BATANES.

NOTE.—Gracilaria vieillardii is proposed as a substitute name for G. denticulata (Kützing) Weber-van Bosse, a later homonym of G. denticulata Schmitz ex Mazza (1907:138).

Polycavernosa Chang and Xia

Polycavernosa debilis (Forsskål) Fredericq and J. Norris

Fucus debilis Forsskål, 1775:191 [type locality: Mokha, Yemen].

Gracilaria debilis (Forsskål) Børgesen, 1932:7.—Taylor, 1977b:13.

Polycavernosa debilis (Forsskål) Fredericq and J. Norris, 1985:152.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

NOTE.—Xia and Abbott (1985) support the segregation of Polycavernosa from Gracilaria on the basis of the origin of the compound spermatangial conceptacles from inner or medullary cells and the indirect origin of the gonimoblast from the fusion cell.

Family PLOCAMIACEAE

Plocamium Lamouroux

Plocamium costatum (C. Agardh) J. Hooker and Harvey

Delesseria plocamium (S.G. Gmelin) C. Agardh var. costata C. Agardh, 1822a:181 [type locality: Western Australia, Australia].

Plocamium costatum (C. Agardh) J. Hooker and Harvey, 1847:404.—Cordero, 1977a:140, fig. 127, pl. XXI: B.

PHILIPPINE DISTRIBUTION.—BATANES.

*Plocamium patens Martens

Plocamium patens Martens, 1868:32 [type locality: Zamboanga, Mindanao]—Velasquez, Trono, and Doty, 1975:160.

PHILIPPINE DISTRIBUTION.—As above.

*Plocamium serrulatum Okamura var. pectinatum Cordero

Plocamium serrulatum Okamura var. pectinatum Cordero, 1977a:140, figs. 128, 129 [type locality: Batan I., Batanes Prov., Luzon].—Cordero, 1978a:38 (as unnamed variety).

PHILIPPINE DISTRIBUTION.—As above.

NOTE.—This name was previously published without a description by Cordero (1976c:8, 9, 11 [“pectinata”])

Plocamium telfairiae (W. Hooker and Harvey) Harvey ex Kützing

Thamnophora telfairiae W. Hooker and Harvey in Harvey, 1834a:149, pl. CXXV [type locality: Cap Malheureux, Mauritius].

Plocamium telfairiae (W. Hooker and Harvey) Harvey ex Kützing, 1849:885.—Cordero, 1977a:141; 1978a:38.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Bataan, Batangas. MINDORO: Oriental Mindoro. PALAWAN.

Family SARCODIACEAE

Sarcodia J. Agardh

Sarcodia montagneana (J. Hooker and Harvey) J. Agardh

Rhodymenia montagneana J. Hooker and Harvey, 1845b:544 [“Rhodomenia”] [type locality: Bay of Islands, New Zealand].

Sarcodia montagneana (J. Hooker and Harvey) J. Agardh, 1852 [1851–1863]:623.

Sarcodia ceylanica Harvey ex Kützing, 1869:12, pl. 33: figs, a, b [type locality: Sri Lanka].—Cordero, 1977a:116.

PHILIPPINE DISTRIBUTION.—BATANES.

NOTE.—The synonymy was proposed by Yendo (1917:82) and maintained by Børgesen (1954:28).

Family SOLIERIACEAE

Callophycus Trevisan

Callophycus serratus (Harvey ex Kützing) P.C. Silva

Thysanocladia serrata Harvey ex Kützing, 1869:10, pl. 29: figs, a, b [type locality: Tonga].

Callophycus serratus (Harvey ex Kützing) P.C. Silva, 1957:143.—Kraft, 1984:54, 56, figs. 3–5, 7, 10–16.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon.

Eucheuma J. Agardh

Eucheuma alvarezii Doty

Eucheuma alvarezii Doty, 1985:37, figs. 1–6 [including Philippine record] [type locality: Creagh Reef, Sabah, Malaysia].

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi.

*Eucheuma alvarezii Doty var. ajakii-assii Doty

Eucheuma alvarezii Doty var. ajakii-assii Doty, 1985:42, fig. 8 [“ajak-assi”] [type locality: Tumindao I., Tawitawi Prov.].

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi.

*Eucheuma alvarezii Doty var. tambalangii Doty

Eucheuma alvarezii Doty var. tambalangii Doty, 1985:41, fig. 7 [“tambalang”] [type locality: Calatagan, Batangas Prov.].

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. SULU: Tawitawi.

NOTE.—The varietal epithets of the two foregoing taxa commemorate three persons named Ajak, Assi, and Tambalang and therefore are to be written ajakii-assii and tambalangii in accordance with Article 73.10 of the ICBN. We doubt the wisdom or usefulness of giving formal taxonomic recognition to horticultural strains, which these varieties seem to be.

Eucheuma arnoldii Weber-van Bosse

Eucheuma arnoldii Weber-van Bosse, 1928:421, pl. XIII: fig. 1 [type locality: Gisser (Gesser) I., near Ceram, Indonesia].—Kraft, 1972:318–334, figs. 1–18.—Trono, 1973d:19; 1974c:328, 333; 1974d:4; 1974e:18.—Cordero, 1977a:151, fig. 144, pl. XXIV:D.—Smith and Pestaño-Smith, 1980:7.—Trono and Fortes, 1980:72.—Ganzon-Fortes, 1981:22.—Meñez and Calumpong, 1981:381.—Calumpong, 1982:145.—Saraya and Trono, 1982:38.—Trono and Ang, 1982:17.—Trono and Fortes, 1982:151.

Eucheuma cupressoideum Weber-van Bosse, 1928:421, pl. XIV: fig. 3 [type locality: Torres Strait].—Kraft, 1972:321, 323.—Cordero, 1977a:153, figs. 147, 150; 1984a:98.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan. MINDORO: Oriental Mindoro. NEGROS: Negros Occidental (Ilacaon I.), Oriental Mindoro. CEBU. BOHOL. PALAWAN (Bugsuk I.), SULU: Sulu (Siasi I.), Tawitawi.

NOTE.—The synonymy was proposed by Kraft (1972).

*Eucheuma arnoldii Weber-van Bosse var. alcyonida Kraft

Eucheuma arnoldii Weber-van Bosse var. alcyonida Kraft, 1972:323, figs. 5, 11 [type locality: Bulusan, Sorsogon Prov., Luzon].

PHILIPPINE DISTRIBUTION.—As above.

Eucheuma cottonii Weber-van Bosse

Eucheuma cottonii Weber-van Bosse, 1913b:115, pl. 12: fig. 2 [syntype localities: Saya de Malha and Cargados Carajos, Indian Ocean].—Doty, 1969:4, figs, A, D.—De Leon and Domantay, 1971:5, 11.—Velasquez, 1971:449.—Kraft, 1972:326, 327, 329, 332.—Cordero, 1973b:32.—Trono, 1973d:19, pl. 12: figs. 13, 15.—Trono and Biña, 1973:7.—Westernhagen, 1973a:65.—Trono, 1974b:92; 1974c:333; 1974d:4; 1974e:18, 19.—Westernhagen, 1974:112 (table 1).—Cordero, 1979b:276.—Ricohermoso and Deveau, 1979:525.—Smith and Pestaño-Smith, 1980:7.—Trono and Fortes, 1980:72.—Donaire, 1981:166.—Ganzon-Fortes, 1981:22.—Meñez and Calumpong, 1981:381.—Laite and Ricohermoso, 1981:595.—Calumpong, 1982:145.—Saraya and Trono, 1982:38.—Trono and Ang, 1982:17.—Trono and Fortes, 1982:151.—Doty and Norris, 1985:figs. 5,6.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Sorsogon. MINDORO: Oriental Mindoro. LEYTE (Biliran I.), PANAY: Aklan. NEGROS: Negros Occidental (Ilacaon I.), CEBU (Mactan I.), BOHOL. MINDANAO: Zamboanga. PALAWAN (Bugsuk I., Cabulauan I.), SULU: Sulu (Siasi I.), Tawitawi.

Eucheuma cottonii var. erecta

Eucheuma cottonii var. erecta—De Leon, 1974:30, 32, photo [s.n.].

PHILIPPINE DISTRIBUTION.—PALAWAN.

NOTE.—De Leon did not give an author for this name and we have been unable to find its place of publication.

Eucheuma crassum Zanardini

Eucheuma crassum Zanardini, 1878:36 [type locality: Aru Is., Indonesia].—Westernhagen, I973a:65.—Cordero, 1977a:151, fig. 145, pl. XXIII:A.—Meñez and Calumpong, 1981:381.—Calumpong, 1982:145.—Cordero, 1984a:97 [with query].

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, NEGROS: Negros Oriental, CEBU (Mactan I.).

Eucheuma crustiforme Weber-van Bosse

Eucheuma crustiforme Weber-van Bosse, 1928:415, fig. 165 [“crustaeforme”] [type locality: Great Sangir (Sangihe) I., Indonesia].—Westernhagen, 1973a:65; 1974:112 (table 1).—Cordero, 1977a:153, fig. 146.—Puig and Cordero, 1979:39.—Cordero, 1984a:98 [“(?) cristaeforme”].

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Batangas. LEYTE (incl. Biliran I.), CEBU (Mactan I.).

Eucheuma denticulatum (N.L. Burman) Collins and Hervey

Fucus denticulatus N.L. Burman, 1768:28 [bis] [type locality: said to be Cape of Good Hope, South Africa, but probably in error].—Velasquez, Trono, and Doty, 1975:144.

Eucheuma denticulatum (N.L. Burman) Collins and Hervey, 1917:106.—Pelayo and Lantican, 1981:3.

Fucus spinosus Linnaeus, 1771:313 [superfluous name].

Eucheuma spinosum J. Agardh, 1847:16 [“spinosa”].—Heydrich, 1894:293.—Seale, 1911:309.—Wester, 1916:159; 1921:224; 1924:21.—G. Blanco, 1938:512.—Quisumbing, 1951:1010.—Montilla and Blanco, 1953:166.—Domantay, 1962:289.—Doty, 1969:6, figs. B, C.—DeLeon and Domantay, 1971:5, 12.—Velasquez, 1971:449.—Trono, 1972a:105.—Westernhagen, 1973a:65.—Parker, 1974:434 (table 1).—Trono, 1974b:92.; 1974c:328, 333; 1974d:4; 1974e:18.—Westernhagen, 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:143.—Laserna et al., 1978:111.—Ricohermoso and Deveau, 1979:525.—Liao and Sotto, 1980:99.—Smith and Pestaño-Smith, 1980:7.—Trono and Fortes, 1980:72.—Veroy et al., 1980:59.—Donaire, 1981:166.—Ganzon-Fortes, 1981:22.—Guzman, 1981:43.—Laite and Ricohermoso, 1981:595.—Laserna et al., 1981:445.—Lim and Porse, 1981:603.—Trono, 1981b:53.—Uyenco, Saniel, and Jacinto, 1981a:69.—Uyenco [Uyengco], Saniel, and Jacinto, 1981b:627.—Saraya and Trono, 1982:37.—Trono and Fortes, 1982:152.—Trono and Ganzon-Fortes, 1985:62, 64, 65, fig. [s.n.].

Fucus muricatus S.G. Gmelin, 1768:111, pl. VI: fig. 4 [type locality: “Oceanus Indicus”].

Eucheuma muricatum (S.G. Gmelin) Weber-van Bosse, 1928:413 [including Philippine record].—Howe, 1932:169.—Zaneveld, 1956:31; 1959:111.—Domantay, 1962:289.—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Galutira and Velasquez, 1964:505, pl. 4: fig. 11; pl. 8: fig. 30.—Velasquez, 1968a:122, fig. 10; 1971:448, fig. 33; 1972:63.—Velasquez et al., 1973:31, pl. 13: fig. 64.—De Leon, 1974:30, 33, photo [s.n.], 2 figs. [s.n.].—Velasquez, Trono, and Doty, 1975:143.—Cordero, 1977a:156, fig. 154.—Velasquez, 1979b:231.—Cordero, 1980b:50, fig. 14, pls. 32, [54].—Trono, Velasquez, and Guevarra, 1980:79.—Cordero, 1982a:60, 61, fig. 11.—Trono and Ang, 1982:17.—Hurtado-Ponce, 1983:143.—Cordero, 1984a:98.—Hurtado-Ponce, 1984:180.—Marcos-Agngarayngay, 1984a:43, fig. 35; 1984b:129.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, La Union, Pangasinan, Bataan. MINDORO: Occidental Mindoro. PANAY: Aklan. CEBU (Mactan I.), SIQUIJOR. BOHOL. MINDANAO: Zamboanga, Davao. PALAWAN (Bugsuk I.). SULU: Sulu (Siasi I.), Tawitawi.

NOTE.—Fucus spinosus Linnaeus, the intended basionym of Eucheuma spinosum, is a later homonym of F. spinosus S.G. Gmelin (1768:161, pl. XVIII: fig. 3) and hence not priorable. Eucheuma spinosum J. Agardh is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN. Fucus spinosus Linnaeus is not only a later homonym, but also an illegitimate substitute for F. denticulatus N.L. Burman. The conspecificity of F. denticulatus and E. muricatum was first proposed by C. Agardh (1822a:271). Both names were published in 1768, with the Burman publication having priority (1 March–6 April vs. after 1 May). The specimen figured by Cordero (1980b: pl. 32) with the caption “E. muricatum var.” is listed under E. gelatinae in the text.

TAXA OF UNCERTAIN VALUE

The following two infraspecific taxa are probably related to Eucheuma denticulatum but have not been formally transferred to it.

Eucheuma muricatum (S.G. Gmelin) Weber-van Bosse f. depauperatum Weber-van Bosse

Eucheuma muricatum (S.G. Gmelin) Weber-van Bosse f. depauperatum Weber-van Bosse, 1928:415, pl. XII: fig. 5 [“depauperata”] [type locality: Ryukyu-retto, Japan].—Domantay, 1962:289.—Cordero, 1977a:156, fig. 155.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan. SIQUIJOR.

Eucheuma muricatum (S.G. Gmelin) Weber-van Bosse f. incrassatum Yamada

Eucheuma muricatum (S.G. Gmelin) Weber-van Bosse f. incrassatum Yamada, 1936a:124, figs. 4, 5, pl. 23: fig. 1 [“incrassata”] [syntype localities: Miyako-jima and Okinawa-jima, Ryukyu-retto, Japan].—Cordero, 1977a:158, figs. 156, 157, pl. XXIII:B; 1980b:50, fig. 15, pl. 33.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Cagayan.

Eucheuma edule (Kützing) Weber-van Bosse

Chondrus edulis Kützing, 1867:19, pl. 63: figs, c–e [type locality: New Caledonia].

Eucheuma edule (Kützing) Weber-van Bosse, 1926:136.—Meñez, 1961:73.—Westernhagen, 1973a:65; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:143.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CEBU (Mactan I.).

Eucheuma gelatinum (Esper) J. Agardh

Fucus gelatinus Esper, 1800:188, pl. Cl [type locality: not specified].

Sphaerococcus gelatinus (Esper) C. Agardh, 1822a:270.—Martens, 1868:46, 94–95.—Velasquez, Trono, and Doty, 1975:163.

Eucheuma gelatinum (Esper) J. Agardh, 1847:16 [“gelatinae”].—Zaneveld, 1956:31; 1959:111.—Domantay, 1962:289.—De Leon and Domantay, 1971:5, 11.—Westernhagen, 1973a:65 [with query]; 1974:112 (table 1) [with query].—Velasquez, Trono, and Doty, 1975:143.—Cordero, 1976c:10; 1977a:153, figs. 151, 152, pl. XXIV:C; 1978a:39; 1980b:49, fig. 13, pl. [53].—Guzman, 1981:44 [“galatinae”].—Cordero, 1984a:98.—Marcos-Agngarayngay, 1984a:41, fig. 34; 1984b:128.—Tungpalan, 1984:145.

Gigartina gelatinosa Endlicher, 1843:42 [illegitimate nomenclatural synonym].—Montagne, 1844a:662.—Velasquez, Trono, and Doty, 1975:145.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Rizal. CEBU (Mactan I.). MINDANAO: Zamboanga.

NOTE.—When establishing the genus Eucheuma, J Agardh (1847:16) treated the name as feminine. Later (J Agardh, 1852 [1851–1863]:624), he treated it as neuter the gender most often assigned to it by subsequent authors. The neuter ending of the adjective gelatinus is gelatinum. J Agardh’s incorrect spelling gelatinae has persisted to the present.

Eucheuma horridum J. Agardh

Eucheuma horridum J. Agardh, 1852 [1851–1863]:625 [type locality: Mauritius].—Cordero, 1977a:155, figs. 148, 149.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Eucheuma isiforme (C. Agardh) J. Agardh

Sphaerococcus isiformis C. Agardh, 1822a:271 [type locality: West Indies].

Eucheuma isiforme (C. Agardh) J. Agardh, 1847:16 [“isiformis”].—Reyes, 1972:157.—Velasquez, Trono, and Doty, 1975:143.—Cordero, 1977a:155, fig. 153, pl. XXIV:B; 1982a:60, 61.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental. SULU.

Eucheuma leeuwenii Weber-van Bosse

Eucheuma leeuwenii Weber-van Bosse, 1928:410, figs. 161, 162 [type locality: Nusa Kembangan, Java, Indonesia].—Westernhagen, 1973a:65; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Eucheuma okamurae Yamada

Eucheuma okamurae Yamada, 1936a:125, figs. 8, 9, pls. 26, 27 [“okamurai”] [type locality: Miyako-jima, Ryukyu-retto, Japan].—Domantay, 1962:289.—De Leon and Domantay, 1971:5, 10.—Velasquez, Trono, and Doty, 1975:143. —Cordero, 1977a:159, fig. 158, pl. XXII:B.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

*Eucheuma procrusteanum Kraft

Eucheuma procrusteanum Kraft, 1970:215, figs. 1–7 [type locality: Semirara Is., Antique Prov., Panay].—Trono, 1974c:328, 333; 1974d:4; 1974e:17, 18.—Velasquez, Trono, and Doty, 1975:143.—Trono and Fortes, 1980:72.—Ganzon-Fortes, 1981:22.—Trono and Fortes, 1982:151.

PHILIPPINE DISTRIBUTION.—PANAY: Antique (Semirara Is.). SULU: Sulu (Siasi I.).

Eucheuma serra (J. Agardh) J. Agardh

Sphaerococcus serra J. Agardh, 1841:17 [type locality: “in mari Indiae orientalis”].

Eucheuma serra (J. Agardh) J. Agardh, 1847:16.—Cordero, 1973b:32.—Trono, 1973d:19, pl. 12: fig. 14.—De Leon, 1974:30, 31, photo [s.n.], fig. [s.n.].—Trono, 1974c:333; 1974d:4; 1974e:17, 18.—Cordero, 1976c:10; 1977a:159, fig. 165; 1978a:39.—Puig and Cordero, 1979:40.—Trono and Fortes, 1980:72.—Ganzon-Fortes, 1981:22.—Trono and Ang, 1982:16.—Trono and Fortes, 1982:151.—Cordero, 1984c:55.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Sorsogon. MASBATE. LEYTE (Biliran I.). PALAWAN (Bugsuk I.).

Eucheuma striatum Schmitz

Eucheuma striatum Schmitz, 1895:151 [“striata”] [type locality: Zanzibar, Tanzania].—Domantay, 1962:289.—Trono, 1972a:105.—Doty and Alvarez, 1973: fig. 2.—Westernhagen, 1973a:65 [with query].—Colina, 1974:109.—De Leon, 1974:30, 31, photo [s.n.].—Parker, 1974:434 (table 1).—Trono, 1974b:91; 1974c:328, 333; 1974d:4; 1974e:18, 19.—Westernhagen, 1974:112 (table 1) [with query].—Colina, 1975:125.—Velasquez, Trono, and Doty, 1975:143.—Colina, 1976:50.—Cordero, 1976c:10; 1977a:160, fig. 159; 1978a:39.—Laserna et al., 1978:111.—Garcia, 1979:45(table 1).—Cordero, 1980b:51, fig. 16, pl. [59].—Liao and Sotto, 1980:99.—Trono and Fortes, 1980:72.—Trono and Ganzon-Fortes, 1980:75, fig. [s.n.].—Veroy et al., 1980:59.—Donaire, 1981:166.—Doty and Alvarez, 1981:688.—Ganzon-Fortes, 1981:22.—Guzman, 1981:43.—Laserna et al., 1981:445—Lim and Porse, 1981:603.—Pelayo and Lantican, 1981:3—Trono, 1981b:53.—Uyenco, Saniel, and Jacinto, 1981a:69.—Uyenco [Uyengco], Saniel, and Jacinto, 1981b:627.—Luistro, Cajipe, and Laserna, 1982:46.—Cordero, 1982a:60, 61.—Trono and Ang, 1982:17.—Trono and Fortes, 1982:152.—Cordero, 1984a:99; 1984c:55.—Marcos-Agngarayngay, 1984a:43, fig. 36; 1984b:129.—Tungpalan, 1984:145.—Trono and Ganzon-Fortes, 1985:62, 64, 65, fig. [s.n.].

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Batangas. MASBATE. CEBU (incl. Mactan I.), BOHOL. MINDANAO: Zamboanga. PALAWAN (Bugsuk I.). SULU: Sulu (Siasi I.), Tawitawi.

Meristotheca J. Agardh

Meristotheca papulosa (Montagne) J. Agardh

Kallymenia papulosa Montagne, 1850:246 [“Callymenia”] [type locality: Hodeida, Yemen].

Meristotheca papulosa (Montagne) J. Agardh, 1872:37.—Velasquez et al., 1973:31, pl. 14: fig. 56.

PHILIPPINE DISTRIBUTION.—LUZON: Bataan.

Solieria J. Agardh

Solieria dura (Zanardini) Schmitz

Rhabdonia dura Zanardini, 1858:278, pl. XI: fig. 1 [syntype localities: Hodeida and Mokha, Yemen].—Piccone, 1886:78, 90.—Velasquez, Trono, and Doty, 1975:161.

Solieria dura (Zanardini) Schmitz, 1895:139.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

Family CAULACANTHACEAE

Catenella Greville

Catenella caespitosa (Withering) L. Irvine

Ulva caespitosa Withering, 1776:735 [type locality: Anglesey, Wales].

Catenella caespitosa (Withering) L. Irvine, 1976:590.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.

Fucus opuntia Goodenough and Woodward, 1797:107, 219 [type locality: Tenby, Wales].

Catenella opuntia (Goodenough and Woodward) Greville, 1830:lxiii.—Post, 1936:73, 76, 78; 1939:34.—Cordero, 1977a:163, figs. 168, 169—Fortes and Trono, 1980:59.—Reyes, 1980:134, pl. 9: fig. 3.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon, MINDORO: Oriental Mindoro. SIQUIJOR. PALAWAN.

NOTE.—The synonymy was proposed by Irvine (1976).

Catenella impudica (Montagne) J. Agardh

Lomentaria impudica Montagne, 1840:197 [type locality: Cayenne, French Guiana].

Catenella impudica (Montagne) J. Agardh, 1852 [1851–1863]:701.—Post, 1936:66, 68; 1938:213.—Cordero, 1977a:163, fig. 161.—Vannajan and Trono, 1978:18, fig. 19.—Trono and Fortes, 1980:71.—Ganzon- Fortes, 1981:22.—Trono and Fortes, 1982:151.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Rizal, Manila. SULU: Sulu.

Catenella nipae Zanardini

Catenella nipae Zanardini, 1872:143, pl. VI:A [type locality: Sarawak, Malaysia].—Post, 1938:214.—Quisumbing, 1951:1007.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal. PANAY: Iloilo.

Family CYSTOCLONIACEAE

Rhodophyllis Kützing

Rhodophyllis peltata Grunow var. lacunosa Grunow

Rhodophyllis peltata Grunow var. lacunosa Grunow, 1874:34 [type locality: Tongatapu, Tonga].—Weber-van Bosse, 1928:403.—Velasquez, Trono, and Doty, 1975:161 [without designation of variety].

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Capual I.).

Family HYPNEACEAE

Hypnea Lamouroux

Hypnea boergesenii Tanaka

Hypnea boergesenii Tanaka, 1941:233, figs. 6–8, pl. 53: fig. 1 [syntype localities: Keelung (Chi-lung) and Tairi, Taiwan].—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Hypnea cenomyce J. Agardh

Hypnea cenomyce J. Agardh, 1851 [1851–1863]:452 [type locality: “ad oras Novae Hollandiae” (Australia)]. —Zaneveld, 1956:41; 1959:121.—Cordero, 1977a:143, fig. 130; 1979b:291; 1980b:52.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro. PANAY: Aklan. SIQUIJOR.

*Hypnea cenomyce J. Agardh var. tenuis Weber-van Bosse

Hypnea cenomyce J. Agardh var. tenuis Weber-van Bosse, 1928:456, fig. 194 [syntype localities: Muaras Reef, East Kalimantan, Indonesia; Sangasiapu I., Tawitawi Prov., Sulu Archipelago].

PHILIPPINE DISTRIBUTION.—As above.

Hypnea cervicornis J. Agardh

Hypnea cervicornis J. Agardh, 1851 [1851—1863]:451 [syntype localities: Brazil; West Indies; Mexico; Mauritius].—Weber-van Bosse, 1928:454.—Zaneveld, 1956:42; 1959:121.—Velasquez et al., 1973:31, pl. 14: fig. 66.—Westernhagen, 1973a:65; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:150.—Cordero, 1977a:143, figs. 131, 132; 1978a:40.—Trono, 1978:18.—Vannajan and Trono, 1978:21, fig. 21 [miscited as 20].—Cordero, 1979b:276, 290; 1980b:52,pl. 34.—Reyes, 1980:136, pl. 10: figs. 2a,b.—Trono and Fortes, 1980:70.—Trono and Ganzon-Fortes, 1980:77, fig. [s.n.].—Chan, 1981:387.—Cordero, 1981d:65, fig. 6.—Ganzon-Fortes, 1981:22.—Laserna et al., 1981:445.—Meñez and Calumpong, 1981:381.—Trono and De Lara, 1981:17.—Cordero, 1982a: fig. 10.—Saraya and Trono, 1982:39.—Trono and Ang, 1982:18.—Trono and Fortes, 1982:150.—Hurtado-Ponce, 1983:142; 1984:180.—Tungpalan, 1984:141.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Bataan, Cavite, Batangas, Quezon, MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. SAMAR: Eastern Samar. PANAY: Aklan. NEGROS: Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Surigao del Sur, Davao. PALAWAN (incl. Bugsuk I.), SULU: Sulu (Siasi I.), Tawitawi (Sangasiapu I.).

Hypnea charoides Sonder

Hypnea charoides Sonder, 1848 [1846–1848]:189 [type locality: Western Australia, Australia].—Meñez, 1961:74, pl. 8: fig. 91.—Galutira and Velasquez, 1964:505, pl. 4: fig. 12; pl. 8: fig. 31.—Villones and Magdamo, 1968:29. fig. 30.—Velasquez, 1971:449, fig. 33.—Reyes, 1972:156.—Velasquez, 1972:63.—Westernhagen, 1973a:65; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:150.—Cordero, 1976c:10; 1977a:144, fig. 133; 1978a:58.—Puig and Cordero, 1979:40.—Velasquez, 1979b:231.—Cordero, 1980b:53, fig. 12, pl. 35.—Liao and Sotto, 1980:99.—Guzman, 1981:42, 45, 49.—Cordero, 1982a:60.—Hurtado-Ponce, 1983:142.—Cordero, 1984a:99; 1984b:66.—Hurtado-Ponce, 1984:180.—Marcos-Agngarayngay, 1984a:45, fig. 37; 1984b:128.—Tungpalan, 1984:141.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Cavite, Batangas, Quezon, CATANDUANES. MINDORO: Oriental Mindoro. SAMAR: Eastern Samar. LEYTE (Biliran I.), NEGROS: Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. PALAWAN.

NOTE.—Hypnea charoides is traditionally accredited to Lamouroux (1813:132, pl. 10: figs. 1–3), but that author did not provide a validating description. C. Agardh (1828:141), after having examined a specimen sent to him by Lamouroux, assigned the name to Ceramium filamentosum (= Spyridia filamentosa). Nonetheless, Sonder (1848 [1846–1848]) apparently believed that Lamouroux’s illustration was representative of another alga since he applied the name to a plant collected by Preiss in Western Australia. Sonder provided a diagnosis so that the name must be ascribed to him and dated from 1848.

Hypnea cornuta (Kützing) J. Agardh

Chondroclonium cornutum Kützing, 1849:741 [type locality: “Locus natalis ignotus” (“ad oras Guineae” fide J. Agardh, 1852 [1851–1863]:449)].—Martens, 1868:90–91.—Velasquez, Trono, and Doty, 1975:135.

Hypnea cornuta (Kützing) J. Agardh, 1851 [1851–1863]:449.—Reyes, 1972:156.—Trono, 1973d:17; 1974e:18.—Velasquez, Trono, and Doty, 1975:150.—Meñez and Calumpong, 1981:381.—Trono and Ang, 1982:18.

PHILIPPINE DISTRIBUTION.—LUZON: Manila, NEGROS: Negros Oriental, SIQUIJOR. PALAWAN (Bugsuk I.).

*Hypnea cornuta (Kützing) J. Agardh var. stellulifera J. Agardh

Hypnea cornuta (Kützing) J. Agardh var. stellulifera J. Agardh, 1852 [1851—1863]:449 [syntype localities: “ad oras Cochinchinae” (Vietnam); Manila].

PHILIPPINE DISTRIBUTION.—LUZON: Manila.

Hypnea divaricata (C. Agardh) Greville

Fucus divaricatus Brown ex Turner, 1811:110, pl. 181 [type locality: “coast of New Holland” (Australia)].

Sphaerococcus divaricatus C. Agardh, 1817:XVII.

Hypnea divaricata (C. Agardh) Greville, 1830:lix.—Martens, 1868:30, 92–93.—Dickie, 1874a:194—Piccone, 1886:77, 90.—Zaneveld, 1956:42; 1959:121.—Velasquez, Trono, and Doty, 1975:150.—Cordero, 1977a:146, fig. 134.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite, Quezon. MINDANAO: Zamboanga. SULU.

NOTE.—Fucus divaricatus Brown ex Turner, the intended basionym of Hypnea divaricata, is a later homonym of F. divaricatus Linnaeus (1753:1159) and hence not priorable. Sphaerococcus divaricatus C. Agardh is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Hypnea musciformis (Wulfen) Lamouroux

Fucus musciformis Wulfen in Jacquin, 1789:154, pl. 14: fig. 3 [type locality: Trieste, Italy].

Hypnea musciformis (Wulfen) Lamouroux, 1813:131.—Howe, 1932:170.—Velasquez, 1953a:100; 1953b:206.—Zaneveld, 1956:42; 1959:121.—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Velasquez, 1971:449.—Westernhagen, 1973a:65; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:150.—Cordero, 1977a:148, fig. 136, pl. XXII:A.—Trono and Fortes, 1980:70.—Ganzon-Fortes, 1981:22.—Trono and Fortes, 1982:150.—Cordero, 1984a:100.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Rizal, Manila, NEGROS: Negros Occidental, CEBU (Mactan I.).

Hypnea musciformis (Wulfen) Lamouroux var. esperi J. Agardh

Hypnea musciformis (Wulfen) Lamouroux var. esperi J. Agardh, 1851 [1851–1863]:442 [syntype localities: Brazil; Chile; Australia; Mauritius].—Reyes, 1972:156.—Trono, 1972a:105.—Cordero, 1977a:146, fig. 135.—Meñez and Calumpong, 1981:381.—Trono and Ang, 1982:19.

Hypnea esperi [misapplied name; see Note].—Reyes, 1972:156.—Trono, 1973d:17.—Trono and Biña, 1973:8.—Velasquez, Trono, and Doty, 1975:150.—Cordero, 1980b:53.—Reyes, 1980:136, pl. 10: figs. 3a,b.—Trono and Fortes, 1980:70.—Trono and Ganzon-Fortes, 1980:79, fig. [s.n.].—Chan, 1981:387.—Saraya and Trono, 1982:38.—Trono and Fortes, 1982:150.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas, Quezon, MINDORO: Oriental Mindoro. NEGROS: Negros Oriental. SIQUIJOR. PALAWAN (Bugsuk I.). SULU: Tawitawi.

NOTE.—Hypnea esperi Bory de Saint-Vincent (1828 [1826–1829]:157) was superfluous when published, being a substitute name for Fucus nootkanus Esper (1802:30, pl. CXXV). If the entity currently passing under this name is recognized at the level of species, it must receive a different name. It was first given a valid name, at varietal level, by J. Agardh, who made it clear that he was describing the material in Bory de Saint-Vincent’s hands and excluding Fucus nootkanus.

Hypnea musciformis (Wulfen) Lamouroux var. hippuroides (Kützing) Weber-van Bosse

Hypnea hippuroides Kützing, 1868:7, pl. 21: figs, a, b [type locality: Celebes, Indonesia].

Hypnea musciformis (Wulfen) Lamouroux var. hippuroides (Kützing) Weber- van Bosse, 1928:445.—Cantoria, Valenzuela, and Velasquez, 1951:187, fig. 1.—Cantoria, Velasquez, and Valenzuela, 1951:296, 298.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal.

Hypnea nidulans Setchell

Hypnea nidulans Setchell, 1924:161, fig. 30 [type locality:Tutuila I., American Samoa].—Weber-van Bosse, 1928:454.—Meñez, 1961:74, pl. 7:figs. 85, 86.—Domantay, 1962:291.—Westernhagen, 1973a:65; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:150.—Cordero, 1977a:148, figs. 138, 139.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas. MINDORO: Oriental Mindoro. CENTRAL VISAYAS. CEBU (Mactan I.), SULU:Sulu (Jolo I.).

Hypnea pannosa J. Agardh

Hypnea pannosa J. Agardh, 1847:14 [type locality: “St. Augustin” (Oaxaca, Mexico)].—Trono, 1972a:105; 1973d:17, pl. 7: fig. 25; 1974e:18.—Trono and Biña, 1973:8.—Trono, 1978:19.—Liao and Sotto, 1980:99.—Trono and Fortes, 1980:71.—Trono and Ganzon-Fortes, 1980:81, fig. [s.n.].—Ganzon-Fortes, 1981:22.—Meñez and Calumpong, 1981:381.—Saraya and Trono, 1982:39.—Trono and Fortes, 1982:151.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas. MINDORO: Oriental Mindoro. MARINDUQUE. CEBU (Mactan I.), SULU: Tawitawi.

Hypnea saidana Holmes

Hypnea saidana Holmes, 1896:256, pl. II: figs. 3a,b [type locality: Enoshima, Kanagawa Prefecture, Japan].—Cordero, 1977a:149, fig. 140; 1979b:291; 1984c:55.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite (Corregidor I.), MINDORO: Oriental Mindoro. MASBATE. PANAY: Aklan.

Hypnea spinella (C. Agardh) Kützing

Sphaerococcus spinellus C. Agardh, 1822a:323 [type locality: West Indies].

Hypnea spinella (C. Agardh) Kützing, 1847a:23.—Dickie, 1876a:244.—Westernhagen, 1973a:65; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:150.—Trono and Fortes, 1980:70.—Trono and Fortes, 1982:150.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.). MINDANAO: Zamboanga.

Hypnea valentiae (Turner) Montagne

Fucus valentiae Turner, 1809:17, pl. 78 [type locality: Red Sea].

Hypnea valentiae (Turner) Montagne, 1841 [1839–1842]:161; 1844a:662.—Martens, 1868:92–93.—Dickie, 1874a:194.—Domantay, 1962:291.—Trono, 1972a:105; 1974e:18.—Velasquez, Trono, and Doty, 1975:150.—Vannajan and Trono, 1978:21, fig. 20 [miscited 21].—Trono and Fortes, 1980:71.—Trono and Ganzon-Fortes, 1980:83, fig. [s.n.].—Chan, 1981:387, 389.—Ganzon-Fortes, 1981:22.—Meñez and Calumpong, 1981:381.—Trono and De Lara, 1981:17, pl. XI: fig. 4—Saraya and Trono, 1982:39.—Trono and Ang, 1982:18.—Trono and Fortes, 1982:151.—Meñez, Phillips, and Calumpong, 1983:18, 23.—Meñez and Calumpong, 1984:105.

Hypnea hamulosa [misapplied name fide Papenfuss, 1958:105–106].—Cordero, 1976c:9, 10; 1977a:146, fig. 137; 1978a:40.—Meñez and Calumpong, 1981:381.—Trono and Ang, 1982:19.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Batangas, Cavite. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SIQUIJOR. MINDANAO: Zamboanga. PALAWAN (incl. Bugsuk I.), SULU: Tawitawi.

NOTE.—The name Hypnea hamulosa is traditionally attributed to (Turner) Montagne and applied in accordance with a concept derived from Turner’s treatment of Fucus hamulosus (1809: pl. 79). As pointed out by Papenfuss (I.c.), however, Turner did not describe his plant as new, but identified it with Fucus hamulosus Esper (1800:169 [129 misprint], pl. LXXXIX), based on a collection from the Malabar coast of India. Esper’s plant was referred by J. Agardh (1852 [1851–1863]:444), with a query, to Hypnea nigrescens Greville ex J. Agardh (1852 [1851–1863]:443) while Turner’s plant is generally considered to be referable to H. valentiae (see Kützing, 1849:758; Hauck, 1887 [1886–1887]:20; Børgesen, 1943:59).

Family PHYLLOPHORACEAE

Ahnfeltia E.M. Fries

Ahnfeltia concinna J. Agardh

Ahnfeltia concinna J. Agardh, 1851 [1851—1863]:312 [syntype localities: Hawaiian Is.; Chincha Is., Peru].—Cordero, 1977a:164.

PHILIPPINE DISTRIBUTION.—BATANES. SIQUIJOR. PALAWAN.

Ahnfeltia furcellata Okamura

Ahnfeltia furcellata Okamura, 1933 [1933–1942]:16, pl. 310: figs, 6–10 [syntype localities: various, all in Japan].—Cordero, 1977a:165; 1978a:42.

PHILIPPINE DISTRIBUTION.—BATANES.

Gymnogongrus Martius

Gymnogongrus dilatatus (Turner) J. Agardh

Fucus dilatatus Turner, 1819 [1815–1819]:57, pl. 219 [type locality: Cape of Good Hope, South Africa].

Gymnogongrus dilatatus (Turner) J. Agardh, 1851 [1851–1863]:326.—Dickie, 1876a:246.—Velasquez, Trono, and Doty, 1975:147.

PHILIPPINE DISTRIBUTION.—PANAY: Iloilo (Gigantes Is.).

Gymnogongrus divaricatus Holmes

Gymnogongrus divaricatus Holmes, 1896:255, pl. 8: figs. 3a,b [type locality: Shimoda, Shizuoka Prefecture, Japan].—Cordero, 1977a:165; 1978a:41.

PHILIPPINE DISTRIBUTION.—BATANES.

Gymnogongrus flabelliformis Harvey

Gymnogongrus flabelliformis Harvey, 1857:332 [type locality: Shimoda, Shizuoka Prefecture, Japan].—Cordero, 1977a:166, fig. 170.—Hurtado–Ponce, 1983:143; 1984:180.—Marcos-Agngarayngay, 1984a:51, fig. 43.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Norte.

Gymnogongrus pygmaeus J. Agardh

Gymnogongrus pygmaeus J. Agardh, 1851 [1851—1863]:317 [type locality: “ad oras Hindostaniae” (India)].—Martens, 1868:31, 96–97.—Velasquez, Trono, and Doty, 1975:147.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Phyllophora Greville

Phyllophora submaritima Dawson

Phyllophora submaritima Dawson, 1949b:6, figs. 17, 18 [“submaritimus”] [type locality: Cortes Bank, southern California, USA].—Cordero, 1977a:168, fig. 173.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental.

Family GIGARTINACEAE

Gigartina tenella Harvey

Gigartina tenella Harvey, 1860a:332 [type locality: Kikai-jima, Amami-gunto, Ryukyu-retto, Japan].—Cordero, 1977a:169.—Tahil, 1978:52.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. CEBU (Mactan I.).

Order RHODYMENIALES

Family RHODYMENIACEAE

Botryocladia (J. Agardh) Kylin

Botryocladia botryoides (Wulfen) J. Feldmann

Fucus botryoides Wulfen in Jacquin, 1789:146, pl. 13: fig. 1 [type locality: Adriatic Sea].

Botryocladia botryoides (Wulfen) J. Feldmann, 1941:90.

Chrysymenia uvaria J. Agardh, 1842:106.—Dickie, 1876a:245.—Velasquez, Trono, and Doty, 1975:135.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

NOTE.—The nomenclature of this species is discussed by Silva (1980:124). Chondria uvaria C. Agardh (1822a:347), the intended basionym of Chrysymenia uvaria, is a superfluous name for Fucus botryoides and hence not priorable. Chrysymenia uvaria J. Agardh, while treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN, is equally superfluous.

Botryocladia pyriformis (Børgesen) Kylin

Chrysymenia pyriformis Børgesen, 1910:187, figs. 8, 9 [type locality: St. Jan (St. John), Virgin Is.].

Botryocladia pyriformis (Børgesen) Kylin, 1931:18.—Reyes, 1980:137, pl. II: fig. 2.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Botryocladia skottsbergii (Børgesen) Levring

Chrysymenia skottsbergii Børgesen, 1924:307, figs. 49, 50 [type locality: Easter I.].

Botryocladia skottsbergii (Børgesen) Levring, 1941:645, footnote.

Chrysymenia kuckuckii Weber-van Bosse, 1928:466, fig. 199 [syntype localities: various, all in Indonesia].

Botryocladia kuckuckii (Weber-van Bosse) Yamada and Tanaka, 1938:77.—Meñez, 1961:78.—Velasquez, Trono, and Doty, 1975:130.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—The conspecificity of Chrysymenia kuckuckii and C. skottsbergii was proposed by G. Feldmann (1945:55–56) and Børgesen (1950:42–44).

Botryocladia uvarioides Dawson

Botryocladia uvarioides Dawson, 1944:306, pl. 45: figs. 8–10; pl. 75: fig. 1 [type locality: San Jose del Cabo, Baja California Sur, Mexico].—Trono, 1973a:134, figs. 6, 8; 1974b:88.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon. SULU: Sulu (Siasi I.).

Coelothrix Børgesen

Coelothrix irregularis (Harvey) Børgesen

Cordylecladia ? irregularis Harvey, 1853:156 [type locality: Key West, Florida, USA].

Coelothrix irregularis (Harvey) Børgesen, 1920:389.—Trono, 1972a:103; 1974b:89.—Saraya and Trono, 1982:43, pl. VII: figs. 1, 4.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. SULU: Sulu (Siasi I.), Tawitawi.

Erythrocolon J. Agardh

Erythrocolon podagricum J. Agardh

Erythrocolon podagricum J. Agardh in Grunow, 1874:33 [syntype localities: Tongatapu and Haafeva I., Tonga; Ovalau I., Fiji].—Cordero, 1977a:169, fig. 174, pl. XXIV:A.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas, Quezon.

Fauchea Montagne and Bory

Fauchea leptophylla Segawa

Fauchea leptophylla Segawa, 1941:264, fig. 10, pl. 58: fig. 1 [type locality: Kozu-shima, Izu-shoto, Japan].—Cordero, 1977a:170, fig. 171.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Gloiocladia J. Agardh

Gloiocladia ramellifera Hauck

Gloiocladia ramellifera Hauck, 1886 [1886–1887]:219 [type locality: Meith [Maidh], Somalia].—Weber-van Bosse, 1928:457, pl. XI: fig. 1.—Velasquez, Trono, and Doty, 1975:145.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Pearl Bank).

Hymenocladia J. Agardh

Hymenocladia dactyloides (Sonder) J. Agardh

Gracilaria dactyloides Sonder, 1845:55 [type locality: Western Australia, Australia].—Dickie, 1876a:245.—Velasquez, Trono, and Doty, 1975:146.—Abbott, 1985:89.

Hymenocladia dactyloides (Sonder) J. Agardh, 1870:454.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Rhodymenia Greville

Rhodymenia californica Kylin

Rhodymenia californica Kylin, 1931:21, pl. 9: fig. 22 [type locality: Pacific Grove, California, USA].—Cordero, 1977a:170, fig. 175

PHILIPPINE DISTRIBUTION.—PALAWAN.

Rhodymenia coacta Okamura and Segawa

Rhodymenia coacta Okamura and Segawa in Segawa, 1935:84, pl. 20: fig. 1 [type locality: Shikine-jima, Izu-shoto, Japan].—Cordero, 1977a:172, fig. 176.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. CENTRAL VISAYAS.

Rhodymenia decumbens W.R. Taylor

Rhodymenia decumbens W.R. Taylor, 1945:251, pl. 84: fig. 1 [type locality: Isla Baltra, Galápagos].—Cordero, 1977a:172 [R. procumbens, in error].

PHILIPPINE DISTRIBUTION.—BATANES.

Rhodymenia intricata (Okamura) Okamura

Phyllophora intricata Okamura, 1921:129, pl. CLXXXII: figs. 1–8 [syntype localities: various, all in Japan].—Cordero, 1977a:168, fig. 172, pl. XXV:B.

Rhodymenia intricata (Okamura) Okamura, 1930 [1929–1932]:31 [“23”], pl. CCLXVII—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Family LOMENTARIACEAE

Lomentaria Lyngbye

Lomentaria articulata (Hudson) Lyngbye

Ulva articulata Hudson, 1762:476 [type locality: Cornwall, England].

Lomentaria articulata (Hudson) Lyngbye, 1819:101.—Cordero, 1976c:10; 1977a:177; 1978a:44.

PHILIPPINE DISTRIBUTION.—BATANES.

Lomentaria baileyana (Harvey) Farlow

Chylocladia baileyana Harvey, 1853:185, pl. XX:C: fig. 1 [syntype localities: various, in Massachusetts, Rhode Island, and New York, USA].

Lomentaria baileyana (Harvey) Farlow, 1876:698.—Cordero, 1976c:6.

PHILIPPINE DISTRIBUTION.—BATANES.

Lomentaria hakodatensis Yendo

Lomentaria hakodatensis Yendo, 1920:6 [syntype localities: various, all in Japan].—Cordero, 1976c:9; 1977a:177; 1978a:44.

PHILIPPINE DISTRIBUTION.—BATANES.

Lomentaria pinnata Segawa

Lomentaria pinnata Segawa, 1938:148, fig. 7 [type locality: Miyake-jima, Izu-shoto, Japan].—Cordero, 1977a:177; 1984a:103.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Family CHAMPIACEAE

Champia Desvaux

Champia bifida Okamura

Champia bifida Okamura, 1901 [1900–1902]:67, pl. XXIV [syntype localities: Enoshima and Misaki, Kanagawa Prefecture, Japan].—Cordero, 1977a:173, fig. 178.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Champia caespitosa Dawson

Champia caespitosa Dawson, 1944:311, pl. 46: figs. 3, 4 [type locality: Isla Pond (Estanque), Baja California Norte, Mexico].—Cornejo and Velasquez, 1972:178, pl. 4: figs. 34, 34a.—Velasquez, Trono, and Doty, 1975:134.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

Champia compressa Harvey

Champia compressa Harvey, 1838:402 [type locality: Muizenberg, Cape Province, South Africa].—Dickie, 1876a:244.—Velasquez, Trono, and Doty, 1975:134.

Champia vieillardii Kützing, 1866:14, pl. 37: figs, e, f [type locality: not specified (New Caledonia fide Grunow, 1874:34)].—Saraya and Trono, 1982:42.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDANAO: Zamboanga.

NOTE.—The synonymy was proposed by Grunow (1874:34).

Champia disticha Dawson

Champia disticha Dawson, 1944:310, pl. 46: fig. 5 [type locality: Isla San Esteban, Sonora, Mexico].—Cordero, 1977a:174.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Champia japonica Okamura

Champia japonica Okamura, 1931 [1929–1932]:49, pl. CCLXXVI: figs. 1–6 [syntype localities: various, all in Japan].—Cordero, 1977a:174; 1978a:45.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Batangas.

Champia parvula (C. Agardh) Harvey

Chondria parvula C. Agardh, 1824:207 [type locality: Cádiz, Spain].

Champia parvula (C. Agardh) Harvey, 1853:76.—Reyes, 1972:157.—Trono and Biña, 1973:10.—Ortega, Alcala, and Reyes, 1974:187, 188.—Trono, 1974b:89.—Velasquez, Trono, and Doty, 1975:134.—Cordero, 1976c:10; 1977a:175, fig. 179.—Taylor, 1977b:13.—Cordero, 1978a:45.—Liao and Sotto, 1980:99.—Chan, 1981:387.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.—Saraya and Trono, 1982:42, pl. VI: figs. 1, 2.—Meñez, Phillips, and Calumpong, 1983:23.—Meñez and Calumpong, 1984:105.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Quezon, MINDORO: Oriental Mindoro. NEGROS: Negros Oriental, CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga. SULU: Sulu (Siasi I.). PALAWAN.

Champia salicornoides Harvey

Champia salicomoides Harvey, 1853:76, pl. XIX:B [type locality: Key West, Florida, USA].—Weber-van Bosse, 1928:477—Velasquez, Trono, and Doty, 1975:130.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS. SULU: Sulu (North Ubian I.), Tawitawi (Pearl Bank).

*Champia spathulata Weber-van Bosse

Champia spathulata Weber-van Bosse, 1928:477, pl. XVI: fig. 6 [type locality: North Ubian I., Sulu Prov., Sulu Archipelago].—Velasquez, Trono, and Doty, 1975:134.

PHILIPPINE DISTRIBUTION.—As above.

Order CERAMIALES

Family CERAMIACEAE

Anotrichium Nägeli

Anotrichium tenue (C. Agardh) Nägeli

Griffithsia tenuis C. Agardh, 1828:131 [“Griffitsia”] [type locality: Venezia, Italy].—Fortes and Trono, 1980:60.—Chan, 1981:387, 389.—Fortes, 198lb:396.—Saraya and Trono, 1982:47.

Anotrichium tenue (C. Agardh) Nägeli, 1862:399.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas.

Antithamnion Nägeli

Antithamnion antillanum Børgesen

Antithamnion antillanum Børgesen, 1917:226, figs. 213–216 [type locality: St. Thomas, Virgin Is.].—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Antithamnion lherminieri Nasr

Antithamnion lherminieri Nasr, 1941:66, figs. 9, 10 [type locality: Abu Sadaf Reef, Red Sea, Egypt].—Cordero, 1977a:178, figs. 181, 182.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS. PALAWAN.

NOTE.—Nasr appears to be the first author to have supplied a validating description for this species, which traditionally is accredited to (P. Crouan and H. Crouan) Bornet. Its morphology, taxonomy, and nomenclature are treated by Abbott (1979:215).

Balliella Itono and Tanaka

Balliella subcorticata (Itono) Itono and Tanaka

Antithamnion subcorticatum Itono, 1969:40, fig. 7 [type locality: Yoronjima, Okinawa-gunto, Ruykyu-retto, Japan].—Cordero, 1983:189, fig. 1.

Balliella subcorticata (Itono) Itono and Tanaka, 1973:250.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Centroceras Kützing

Centroceras apiculatum Yamada

Centroceras apiculatum Yamada, 1944:42 [type locality: Ant Atoll, near Ponape, Caroline Is.].—Saraya and Trono, 1982:44.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Centroceras clavulatum (C. Agardh) Montagne

Ceramium clavulatum C. Agardh, 1822b:2 [type locality: Callao, Peru].

Centroceras clavulatum (C. Agardh) Montagne, 1846b:140.—Reyes, 1972:158.—Ortega, Alcala, and Reyes, 1974:186, 187.—Velasquez, Trono, and Doty, 1975:133.—Cordero, 1976c:9; 1977a:179, figs. 183, 184; 1978a:47.—Vannajan and Trono, 1978:23, fig. 29.—Reyes, 1980:138, pl. II: figs. 5a,b.—Chan, 1981:387.—Meñez and Calumpong, 1981:381.—Trono and De Lara, 1981:19, pl. XII: fig. 4.—Cordero, 1984a:104.

Centroceras cryptacanthum Kützing, 1841:741 [type locality: Antilles].

Centroceras clavulatum (C. Agardh) Montagne var. cryptacanthum (Kützing) Grunow, 1867:65.—Piccone, 1886:54, 90.

Centroceras hyalacanthum Kützing, 1841:742 [type locality: “Wahrscheinlich aus Westindien”].—Martens, 1868:28, 84–85.—Velasquez, Trono, and Doty, 1975:133.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Manila, Cavite. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. NEGROS: Negros Oriental, SIQUIJOR. MINDANAO: Zamboanga. PALAWAN.

NOTE.—The synonymy was proposed by J. Agardh (1851 [1851–1863]:148–149).

Centroceras minutum Yamada

Centroceras minutum Yamada, 1944:42 [type locality: Ant Atoll, near Ponape, Caroline Is.].—Trono, 1973d:20.—Cordero, 1977a:180, fig. 185.—Fortes, 1981b:396.—Saraya and Trono, 1982:43, pl. VII: fig. 3.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. SIQUIJOR. MINDANAO: Zamboanga.

Ceramiella Børgesen

Hommersand (1963:238) retained Ceramiella within the circumscription of Ceramium, but Díaz-Piferrer (1969a:202) argued in support of its distinctness.

Ceramiella procumbens (Setchell and Gardner) Díaz-Piferrer

Ceramium procumbens Setchell and Gardner, 1924a:772, pl. 27: figs. 51–54 [type locality: Isla Partida, Baja California Norte, Mexico].—Fortes, 1981b:396.

Ceramiella procumbens (Setchell and Gardner) Díaz-Piferrer, 1969c:203.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Ceramium Roth

Ceramium affine Setchell and Gardner

Ceramium affine Setchell and Gardner, 1930:172 [type locality: Isla Guadalupe, Pacific Mexico].—Fortes and Trono, 1980:59 [var. affine].—Fortes, 1981b:396.—Saraya and Trono, 1982:44, pl. VIII: fig. 2 [var. affine].

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas.

Ceramium cruciatum Collins and Hervey

Ceramium cruciatum Collins and Hervey, 1917:144, pl. IV: figs. 27, 28 [type locality: Bermuda].—Westernhagen, 1973a:64; 1974:112 (table 1)

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Ceramium equisetoides Dawson

Ceramium equisetoides Dawson, 1944:320, pl. 51: fig. 1 [type locality: Puerto San Carlos, near Guaymas, Sonora, Mexico].—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Ceramium fastigiatum (Wulfen ex Roth) Harvey

Conferva fastigiata Wulfen ex Roth, 1800b:224 [type locality: Adriatic Sea].

Ceramium fastigiatum (Wulfen ex Roth) Harvey, 1834b:303.—Vannajan and Trono, 1978:24, fig. 31.—Chan, 1981:387.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Manila, Cavite.

NOTE.—Ceramium fastigiatum (Roth) Harvey is a later homonym of C. fastigiatum (Linnaeus) Wiggers (1780:91), C. fastigiatum Roth (1800a:463), and C. fastigiatum Roussel (1806:87). Its correct name probably will be found among taxonomic synonyms.

Ceramium flaccidum (Harvey ex Kützing) Ardissone

Hormoceras flaccidum Harvey ex Kützing, 1862:21, pl. 69: figs. a–d [type locality: Kilkee, County Clare, Eire].

Ceramium flaccidum (Harvey ex Kützing) Ardissone, 1871:40.

Ceramium byssoideum Harvey, 1853:218 [type locality: Key West, Florida, USA].

Ceramium gracillimum (Kützing) Zanardini var. byssoideum (Harvey) Mazoyer, 1938:323.—Cordero, 1976c:9; 1977a:180, fig. 186; 1978a:47.—

Vannajan and Trono, 1978:24.—Fortes and Trono, 1980:61.—Meñez and Calumpong, 1981:381.—Saraya and Trono, 1982:45, pl. VIII: fig. 1 [“byssoideure” p. 45].

Ceramium fimbriatum Setchell and Gardner, 1924a:777, pl. 26: figs. 43, 44 [type locality: near La Paz, Baja California Sur, Mexico].—Meñez and Calumpong, 1981:381.—Saraya and Trono, 1982:44.

Ceramium masonii Dawson, 1950c:126, figs. 11, 12 [type locality: Cabeza Ballena, Baja California Sur, Mexico].—Liao and Sotto, 1980:99.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Cavite. CEBU (Mactan I.).

NOTE.—The synonymy was proposed by Womersley (1978:234). Ceramium byssoideum Harvey is treated as a homonym of C. byssoides Ducluzeau (1805:66) in accordance with Article 64.2, Ex. 8, of the ICBN.

Ceramium gracillimum (Kützing) Zanardini

Hormoceras gracillimum Kützing, 1841:733 [type locality: Trieste, Italy].

Ceramium gracillimum (Kützing) Zanardini, 1847:223.—Chan, 1981:387, 389.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—Fortes cited the binomial without an author, and it is uncertain whether it should be referred to Hormoceras gracillimum. The combination is traditionally (but incorrectly) accredited to (Kützing) Griffiths and Harvey in Harvey (1848 [1847–1851]: pl. CCVI). According to Womersley (1978:234), Harvey had Ceramium flaccidum in hand. A further consideration is that Ceramium gracillimum (Kützing) Zanardini is a later homonym of C. gracillimum C. Agardh (1824:140), a name that has disappeared from the literature.

Ceramium loureiri C. Agardh

Ceramium loureiri C. Agardh, 1824:137 [type locality: “Ad rupes marinas Cochinchinae” (Vietnam)].—Martens, 1868:82–83.—Velasquez, Trono, and Doty, 1975:133.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Ceramium luetzelburgii O.C. Schmidt

Ceramium luetzelburgii O.C. Schmidt, 1924:98, fig. 6 [type locality: Cabo Branco, Paraiba, Brazil].—Fortes and Trono, 1980:59.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Ceramium marshallense Dawson

Ceramium marshallense Dawson, 1957:120, figs. 27a,b [type locality: Rigili I., Eniwetok Atoll, Marshall Is.].—Fortes and Trono, 1980:60.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Ceramium maryae Weber-van Bosse

Ceramium maryae Weber-van Bosse, 1923:324, figs. 117, 118 [type locality: Kawasa I., Paternoster Is. (Kepulauan Tengah), Indonesia].—Domantay, 1962:288.—Velasquez, Trono, and Doty, 1975:133.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Ceramium mazatlanense Dawson

Ceramium mazatlanense Dawson, 1950c:130, pl. 2: figs. 14, 15 [type locality: Mazatlan, Sinaloa, Mexico].—Cornejo and Velasquez, 1972:182, pl. 5: fig. 40.—Velasquez, Trono, and Doty, 1975:133.—Chan, 1981:387, 389.—Fortes, 198lb:396.—Meñez and Calumpong, 1981:381.—Saraya and Trono, 1982:45.—Meñez, Phillips, and Calumpong, 1983:23.—Meñez and Calumpong, 1984:105.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas. CENTRAL VISAYAS. PALAWAN.

Ceramium multijugum Jaasund

Ceramium multijugum Jaasund, 1970:68, figs. 1C–E,H, 2 [intended type locality: Tanzania].—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

NOTE.—Ceramium multijugum is an invalid name because Jaasund failed to indicate a type, as required by Article 37 of the ICBN.

Ceramium nitens (C. Agardh) J. Agardh

Ceramium rubrum (Hudson) C. Agardh var. nitens C. Agardh, 1824:136 [type locality: Antilles].

Ceramium nitens (C. Agardh) J. Agardh, 1851 [1851–1863]:130.—Westernhagen, 1973a:64; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Ceramium personatum Setchell and Gardner

Ceramium personatum Setchell and Gardner, 1930:171, pl. 6: figs. 21, 22 [type locality: Isla Guadalupe, Pacific Mexico].—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Ceramium serpens Setchell and Gardner

Ceramium serpens Setchell and Gardner, 1924a:775, pl. 27: fig. 58 [type locality: La Paz, Baja California Sur, Mexico].—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Ceramium sinicola Setchell and Gardner

Ceramium sinicola Setchell and Gardner, 1924a:773, pl. 25: figs. 40, 41; pl. 75 [type locality: Ensenada, Baja California Norte, Mexico].—Fortes and Trono, 1980:60 [var. sinicola].—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—PANAY: Iloilo.

Ceramium taylorii Dawson

Ceramium taylorii Dawson, 1950c:127, figs. 13, 31–33 [type locality: Cabeza Ballena, Baja California Sur, Mexico].—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Ceramium tenerrimum (Martens) Okamura

Hormoceras tenerrimum Martens, 1868:146, pl. VIII: fig. 2 [type locality: Nagasaki, Japan].

Ceramium tenerrimum (Martens) Okamura, 1921:112.—Cordero, 1977a:181, fig. 187.

PHILIPPINE DISTRIBUTION.—LUZON: Manila Bay.

Ceramium tenuissimum (Roth) J.E. Areschoug

Ceramium diaphanum (Lightfoot) Roth var. tenuissimum Roth, 1806:156 [type locality: not specified].

Ceramium tenuissimum (Roth) J.E. Areschoug, 1847:322.—Cornejo and Velasquez, 1972:182, pl. 3: fig. 25.—Velasquez, Trono, and Doty, 1975:133.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

NOTE.—This species is usually attributed either to (Lyngbye) J. Agardh (1851 [1851–1863]:120) or to (Roth) J. Agardh (1851 [1851–1863]:120). In the first instance, the basionym is cited as Ceramium diaphanum var. tenuissimum Lyngbye (1819:120, pl. 37:B.4), but Lyngbye accredited the variety to Roth. Areschoug was the first to elevate it to specific rank. Regardless, the name is untenable because it is a later homonym of C. tenuissimum Bonnemaison (1828:132) from Atlantic France, which is referable to Aglaothamnion. The correct name probably will be found among taxonomic synonyms.

Ceramium vagans P.C. Silva, new name

Ceramium vagabundum Dawson, 1957:121, fig. 27 e [“vagabunde”] [type locality: Parry I., Eniwetok Atoll, Marshall Is.] [replaced name].—Vannajan and Trono, 1978:24, fig. 32.—Chan, 1981:387.—Saraya and Trono, 1982:45, pl. VII: fig. 2 [“vugabunde” p. 45].

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite.

NOTE.—Ceramium vagans is proposed as a substitute name for C. vagabundum Dawson, a later homonym of C. vagabundum (Linnaeus) Roth (1800a:465).

Ceramium zacae Setchell and Gardner

Ceramium zacae Setchell and Gardner, 1937:89, pl. 8: figs. 22a–22c [type locality: Bahía San Bartolomé (Bahía Tortugas), Baja California Sur, Mexico].—Fortes and Trono, 1980:60, fig. 11.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Crouania J. Agardh

Crouania attenuata (C. Agardh) J. Agardh

Mesogloia attenuata C. Agardh, 1824:51 [type locality: “In mari Atlantico”].

Crouania attenuata (C. Agardh) J. Agardh, 1842:83.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Crouania minutissima Yamada

Crouania minutissima Yamada, 1944:40 [type locality: Ant Atoll, near Ponape, Caroline Is.].—Vannajan and Trono, 1978:25, fig. 33.—Fortes and Trono, 1980:65.—Fortes, 198lb:396.—Saraya and Trono, 1982:46.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite. PALAWAN.

Dasyphila Sonder

Dasyphila plumarioides Yendo

Dasyphila plumarioides Yendo, 1920:7 [type locality: Kotosho (Hung–t’ou), Taiwan].—Berdach, 1980:101, fig. 2.

PHILIPPINE DISTRIBUTION.—PANAY: Aklan.

Gordoniella Itono

Gordoniella yonakuniensis (Yamada and Tanaka) Itono

Spermothamnion yonakuniense Yamada and Tanaka, 1938:79, figs. 12, 13 [“yonakuniensis”] [type locality: Yonakuni–jima, Sakishima-gunto, Ryukyu-retto, Japan].—Cordero, 1976c:9; 1977a:183, figs. 189, 190; 1978a:46.

Gordoniella yonakuniensis (Yamada and Tanaka) Itono, 1977:54.

PHILIPPINE DISTRIBUTION.—BATANES.

Griffithsia C. Agardh

Griffithsia ovalis Harvey

Griffithsia ovalis Harvey, 1855:559 [type locality: King George Sound, Western Australia, Australia].—Cornejo and Velasquez, 1972:179, pl. 2: figs. 18, 18a.—Velasquez, Trono, and Doty, 1975:147.—Chan, 1981:387.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas. CENTRAL VISAYAS.

Griffithsia rhizophora Grunow ex Weber-van Bosse

Griffithsia rhizophora Grunow ex Weber-van Bosse, 1923:313 [syntype localities: various, in Indonesia and Sri Lanka].—Saraya and Trono, 1982:46, pl. VIII: fig. 4.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Gymnothamnion J. Agardh

Gymnothamnion elegans (Schousboe ex C. Agardh) J. Agardh

Callithamnion elegans Schousboe ex C. Agardh, 1828:162 [type locality: Tangier, Morocco].

Gymnothamnion elegans (Schousboe ex C. Agardh) J. Agardh, 1892:178 [“rlegans”].—Saraya and Trono, 1982:48.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Haloplegma Montagne

Haloplegma duperreyi Montagne

Haloplegma duperreyi Montagne, 1842d:258, pl. 7: fig. 1 [type locality: Martinique].—Berdach, 1980:101, fig. 1 [“Haploplegma”].—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—PANAY: Aklan.

Microcladia Greville

Microcladia elegans Okamura

Microcladia elegans Okamura, 1907 [1907–1909]:1, pl. 1: figs. 1–10 [syntype localities: various, all in Japan].—Cordero, 1976c:11; 1977a:181, pl. XII:A.; 1978a:48; 1984a:104.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan. CATANDUANES. MINDANAO: Davao.

Microcladia glandulosa (Solander ex Turner) Greville

Fucus glandulosus Solander ex Turner, 1808:81, pl. 38 [syntype localities: various, in England and Atlantic Spain].

Microcladia glandulosa (Solander ex Turner) Greville, 1830:1 [“el”].—Domantay, 1962:293.—Velasquez, Trono, and Doty, 1975:153.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Pleonosporium (Nägeli) Nägeli ex Hauck

Pleonosporium globuliferum Levring

Pleonosporium globuliferum Levring, 1941:647, fig. 19 [type locality: Más Afuera, Islas Juan Fernández, Chile].—Saraya and Trono, 1982:46.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Ptilothamnion Thuret

Ptilothamnion cladophorae (Yamada and Tanaka) Feldmann-Mazoyer

Spermothamnion cladophorae Yamada and Tanaka, 1934:342, figs. 1, 2 [syntype localities: Garan-bi (O-luan) and Kasho-to, Taiwan].

Ptilothamnion cladophorae (Yamada and Tanaka) Feldmann-Mazoyer, 1941:375, footnote.—Cordero. 1976c:9; 1977a:183, fig. 188; 1978a:46.

PHILIPPINE DISTRIBUTION.—BATANES

Spyridia Harvey

Spyridia filamentosa (Wulfen) Harvey

Fucus filamentosus Wulfen, 1803:64 [type locality: Adriatic Sea].

Spyridia filamentosa (Wulfen) Harvey, 1833:337.—Weber-van Bosse 1923:320.—Howe, 1932:170.—Reyes, 1972:158.—Trono, 1972a 106; 1973a:140; 1973d:20, pl. II: fig. 12.—Trono and Biña, 1973 11.—Trono, 1974b:95.—Velasquez, Trono, and Doty, 1975:164.—Cordero, 1977a:184, figs. 191, 192.—Reyes, 1980:138, pl. II:figs. 4a,b. —Chan, 1981:387.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.—Saraya and Trono, 1982:47.—Trono and Ang, 1982:20.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan. MINDORO: Oriental Mindoro. NEGROS: Negros Occidental, Negros Oriental, SIQUIJOR. MINDANAO: Zamboanga. PALAWAN (Bugsuk I.), SULU: Sulu (Capual I., Siasi I.), Tawitawi.

Spyridia velasquezii Trono

Spyridia velasquezii Trono, 1972b:53, pl. 9: figs. 1, 2; pl. 10: fig. 3 [type locality: Koror I., Palau Group (Belau), Caroline Is.].—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Wrangelia C. Agardh

Wrangelia argus (Montagne) Montagne

Griffithsia argus Montagne, 1841 [1839–1842]:176, pl. 8: fig. 4 [type locality: Roque del Gando, Islas Canarias].

Wrangelia argus (Montagne) Montagne, 1856:444.—Cordero, 1977a:186, figs. 195–198.—Saraya and Trono, 1982:47, pl. IX: fig. 3.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan. Pangasinan.

Wrangelia bicuspidata Børgesen

Wrangelia bicuspidata Børgesen, 1916:118, figs. 127–130 [type locality: between St. Jan (St. John) and St. Thomas, Virgin Is.].—Fortes, 1981b:396.—Trono and Ang, 1982:21.

PHILIPPINE DISTRIBUTION.—PALAWAN (Bugsuk I.).

Wrangelia penicillata (C. Agardh) C. Agardh

Griffithsia penicillata C. Agardh, 1824:143 [“Griffitsia”] [type locality: “Ad oras Italiae”].

Wrangelia penicillata (C. Agardh) C. Agardh, 1828:138.—Fortes and Trono, 1980:61, fig. 8.—Saraya and Trono, 1982:48, pl. VIII: fig. 3.—Trono and Ang, 1982:21.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CEBU (Mactan I.). PALAWAN (Bugsuk I.).

Family DELESSERIACEAE

Botryocarpa Greville

Botryocarpa prolifera Greville

Fucus botryocarpus Turner, 1819 [1815–1819]:115, pl. 246 [type locality: Cape of Good Hope, South Africa].

Botryocarpa prolifera Greville, 1830:xlix.—Dickie, 1876a:245.—Velasquez, Trono, and Doty, 1975:130.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

NOTE.—Greville, when establishing the genus Botryocarpa on the basis of Fucus botryocarpus Turner, was obliged to change the epithet in order to avoid the creation of a tautonym, which is proscribed by Article 23.4 of the ICBN.

Caloglossa (Harvey) J. Agardh

Caloglossa adnata (Zanardini) De Toni

Delesseria adnata Zanardini, 1872:141, pl. V:B [type locality: Sarawak, Malaysia].

Caloglossa adnata (Zanardini) De Toni, 1900:730.—Post, 1938:211.—Fortes and Trono, 1980:61.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal. MINDORO: Oriental Mindoro.

Caloglossa leprieurii (Montagne) J. Agardh

Delesseria leprieurii Montagne, 1840:196, pl. 5: fig. 1 [type locality: near Cayenne, French Guiana].

Caloglossa leprieurii (Montagne) J. Agardh, 1876:499.—Fortes and Trono, 1980:61, fig. 7.—Fortes, 198lb:396.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. CENTRAL VISAYAS.

Caloglossa leprieurii (Montagne) J. Agardh var. hookeri Post

Caloglossa leprieurii (Montagne) J. Agardh var. hookeri Post, 1936:53 [type locality: Bay of Islands, New Zealand].—Post 1938:211, 212.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal.

Caloglossa ogasawaraensis Okamura

Caloglossa ogasawaraensis Okamura, 1897:12, figs, A–D [type locality: Ogasawara-gunto (Bonin Is.), Japan].—Fortes and Trono, 1980:62.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Claudea Lamouroux

*Claudea batanensis Tanaka

Claudea batanensis Tanaka, 1967:18, figs. 6–8, pl. II:B [type locality: Batan I., Batanes Prov., Luzon].—Velasquez, Trono, and Doty, 1975:136.—Cordero, 1977a:198; 1978a:48.

PHILIPPINE DISTRIBUTION.—BATANES.

Claudea multifida Harvey

Claudea multifida Harvey, 1854:145, pl. VI [type locality: Belligam (Weligama), Sri Lanka].—Cordero, 1977a:199, fig. 215, pl. XXVI.B-Liao and Sotto, 1980:99.—Meñez and Calumpong, 1981:381.—Cordero, 1984a:106.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Batangas. MINDORO: Oriental Mindoro. PANAY: Aklan, Iloilo. GUIMARAS. CEBU (Mactan I.), SIQUIJOR.

Cottoniella Børgesen

Cottoniella filamentosa (Howe) Børgesen

Sarcomenia filamentosa Howe, 1905:571, pl. 27; pl. 29: figs. 1–11 [type locality: Biscayne Key, Florida, USA].

Cottoniella filamentosa (Howe) Børgesen, 1920:478.—Westernhagen, 1973a:65; 1974:112 (table I).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Hypoglossum Kützing

Hypoglossum attenuatum Gardner

Hypoglossum attenuatum Gardner, 1927b:104, pl. 20: fig. 3; pls. 35, 36 [type locality: Puerto Libertad, Sonora, Mexico].—Domantay, 1962:291.—Trono, 1973a:141, fig. 22.—Velasquez, Trono, and Doty, 1975:150.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Sorsogon.

Hypoglossum serrulatum J. Agardh

Hypoglossum serrulatum J. Agardh, 1898:186 [type locality: Port Jackson, New South Wales, Australia].—Weber-van Bosse, 1923:389.—Velasquez, Trono, and Doty, 1975:150.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

Hypoglossum spathulatum (Sonder) Kützing

Delesseria spathulata Sonder, 1845:57 [type locality: Western Australia, Australia].

Hypoglossum spathulatum (Sonder) Kützing, 1849:877.—Weber-van Bosse, 1923:389.—Velasquez, Trono, and Doty, 1975:150.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

Martensia Hering

Martensia flabelliformis Harvey ex J. Agardh

Martensia flabelliformis Harvey ex J. Agardh, 1863 [1851–1863]:826 [type locality: Tonga].—Westernhagen, 1973a:65; 1974:112 (table 1).—Cordero, 1977a:222.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.), SIQUIJOR.

Martensia speciosa Zanardini

Martensia speciosa Zanardini, 1874:488 [type locality: Lord Howe I.].—Weber-van Bosse, 1923:385.—Velasquez, Trono, and Doty, 1975:152.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Pearl Bank).

Opephyllum Schmitz

*Opephyllum martensii Schmitz

Opephyllum martensii Schmitz in Schmitz and Hauptfleisch, 1897 [1896–1897]:410 [type locality, “an der Küste der Philippinen” (Zamboanga)].—De Toni, 1900:620; 1924:321.—Kylin, 1956:449.—Velasquez, Trono, and Doty, 1975:158.

Pollexfenia pedicellata [misapplied name fide Schmitz, l.c.].—Martens, 1868:32, 100–101.—Velasquez, Trono, and Doty, 1975:160.

PHILIPPINE DISTRIBUTION.—As above.

Taenioma J. Agardh

Taenioma perpusillum (J. Agardh) J. Agardh

Polysiphonia perpusilla J. Agardh, 1847:16 [type locality: “St. Augustin” (Oaxaca, Mexico)].

Taenioma perpusillum (J. Agardh) J. Agardh, 1863 [1851–1863]:1257.—Fortes and Trono, 1980:65.—Fortes, 1981b:396.—Saraya and Trono, 1982:48, pl. IX: figs. 1, 4.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. PALAWAN (Bugsuk I.).

Vanvoorstia Harvey

Vanvoorstia spectabilis Harvey

Vanvoorstia spectabilis Harvey, 1854:144, pl. V [type locality: Belligam (Weligama), Sri Lanka].—Weber-van Bosse, 1923:390.—Trono, 1973d:20, pl. 10: fig. 5.—Ortega, Alcala, and Reyes, 1974:186.—Trono, 1974b:94.—Velasquez, Trono, and Doty, 1975:169.—Cordero, 1977a:230, pl. XXVIII:B.—Liao and Sotto, 1980:99.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon, NEGROS: Negros Oriental, SIQUIJOR. CEBU (Mactan I.), SULU: Sulu (North Ubian I., Siasi I.), Tawitawi.

Zellera Martens

Zellera tawallina Martens

Zellera tawallina Martens, 1868:33, pl. VIII: fig. 3 [type locality: Little Tawali 1., Moluccas, Indonesia].—Reyes, 1980:139, pl. 12: fig. 2.—Saraya and Trono, 1982:48.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. SIQUIJOR.

Family DASYACEAE

Dasya C. Agardh

Dasya adhaerens Yamada

Dasya adhaerens Yamada, 1944:43, pl. 7: fig. 1 [type locality: Ant Atoll, near Ponape, Caroline Is.].—Cordero, 1977a:231, fig. 261; 1978a:49.

PHILIPPINE DISTRIBUTION.—BATANES.

Dasya baillouviana (S.G. Gmelin) Montagne

Fucus baillouviana S.G. Gmelin, 1768:165 [type locality: Mediterranean Sea].

Dasya baillouviana (S.G. Gmelin) Montagne, 1841 [1839–1842]:165.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Dasya mollis Harvey

Dasya mollis Harvey, 1853:62 [type locality: Key West, Florida, USA].—Vannajan and Trono, 1978:25, fig. 37.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

Dasya ocellata (Grateloup) Harvey

Ceramium ocellatum Grateloup, 1806: pl. [1]: fig. II [type locality: Sète, Hérault, France].

Dasya ocellata (Grateloup) Harvey, 1833:335.—Cordero, 1977a:231, fig. 262; 1978a:49.

PHILIPPINE DISTRIBUTION.—BATANES.

Dasya punicea (Zanardini) Meneghini

Baillouviana punicea Zanardini, 1840b:204 [type locality: Chioggia, Italy]

Dasya punicea (Zanardini) Meneghini in Zanardini, 1841:168.—Cordero, 1977a:232, fig. 263; 1978a:49.

PHILIPPINE DISTRIBUTION.—BATANES.

Dasya sessilis Yamada

Dasya sessilis Yamada, 1928:524, fig. 19 [syntype localities: various, all in Japan].—Cordero, 1977a:234, figs. 264, 265; 1984a:105.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan.

Eupogodon Kützing

The reason for adopting Eupogodon for the genus currently known as Dasyopsis is given in the appended Nomenclatural Notes.

Eupogodon antillarum (Howe) P.C. Silva, new combination

Dasyopsis antillarum Howe, 1920:577 [type locality: Fort George Cay, Caicos Is., Bahama Is.].—Trono and Ang, 1982:22.

PHILIPPINE DISTRIBUTION.—PALAWAN (Bugsuk I.).

Eupogodon pilosus (Weber-van Bosse) P.C. Silva, new combination

Dasyopsis pilosa Weber-van Bosse, 1923:377, fig. 137 [syntype localities: Indonesia: Waroe, Ceram I., Moluccas; Atja Tuning (Attiatti Onin), Irian Barat].—Cordero, 1977a:234, fig. 266.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental.

Heterosiphonia Montagne

Heterosiphonia crispella (C. Agardh) Wynne

Callithamnion crispellum C. Agardh, 1828:183 [lectotype locality: near Cádiz, Spain fide Wynne, 1985b:86].

Heterosiphonia crispella (C. Agardh) Wynne, 1985b:87.

Dasya wurdemannii Bailey ex Harvey, 1853:64, pl. XV:C [“wurdemanni”] [type locality: Key West, Florida, USA].

Heterosiphonia wurdemannii (Bailey ex Harvey) Falkenberg in Schmitz and Hauptfleisch, 1897 [1896–1897]:473 [“wurdemanni”].—Vannajan and Trono, 1978:25, fig. 38.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

NOTE.—The conspecificity of Callithamnion crispellum and Dasya wurdemannii was proposed by Wynne (1985b).

Heterosiphonia crispella (C. Agardh) Wynne var. laxa (Børgesen) Wynne

Heterosiphonia wurdemannii (Bailey ex Harvey) Falkenberg var. laxa Børgesen, 1919:327, figs. 327, 328 [type locality: St. Croix, Virgin Is.].—Fortes, 1981b:396.

Heterosiphonia crispella (C. Agardh) Wynne var. laxa (Børgesen) Wynne, 1985b:87.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Heterosiphonia muelleri (Sonder) De Toni

Dasya muelleri Sonder, 1855:525 [type locality: Port Phillip, Victoria, Australia].

Heterosiphonia muelleri (Sonder) De Toni, 1903:1237.—Weber-van Bosse, 1923:380.—Velasquez, Trono, and Doty, 1975:150.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

NOTE.—Parsons (1975:627) expressed doubt that tropical records of Heterosiphonia muelleri (including Weber-van Bosse’s record from Sulu) are referable to this species.

Tapeinodasya Weber-van Bosse

*Tapeinodasya bornetii Weber-van Bosse

Tapeinodasya bornetii Weber-van Bosse, 1904a:96, figs. 1, 2 [“borneti”] [syntype localities: Saleyer (Selajar) I., Celebes, Indonesia; Sulu Archipelago].—Weber-van Bosse, 1923:381, figs. 138, 139, pl. X: figs. 1–4.—Velasquez, Trono, and Doty, 1975:165.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Pearl Bank, Sangasiapu I.).

Family RHODOMELACEAE

Acanthophora Lamouroux

Acanthophora aokii Okamura

Acanthophora aokii Okamura, 1934 [1933–1942]:35, pl. 318: figs. 15–17 [“aoki”] [syntype localities: Tainan and Kotosho (Hung-t’ou), Taiwan].—Cordero, 1977a:187, figs. 199, 200, 207; 1978a:48, 56; 1980b:60.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, MINDORO: Oriental Mindoro. SAMAR: Western Samar, Eastern Samar. PANAY: Aklan.

Acanthophora muscoides (Linnaeus) Bory de Saint-Vincent

Fucus muscoides Linnaeus, 1753:1161 [type locality: Ascension I.].

Acanthophora muscoides (Linnaeus) Bory de Saint-Vincent, 1828 [1826–1829]:156.—Westernhagen, 1973a:64; 1974:112 (table 1).—Cordero, 1977a:189, fig. 204; 1980b:61.—Trono and Ganzon-Fortes, 1980:97, fig. [s.n.].—Ganzon-Fortes, 1981:22.—Laserna et al., 1981:445.—Meñez and Calumpong, 1981:381.—Calumpong, 1982:145.—Cordero, 1984a:105.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Batangas, Quezon, NEGROS: Negros Oriental (Apo I.), CEBU (Mactan I.). SIQUIJOR.

Acanthophora spicifera (Vahl) Børgesen

Fucus spiciferus Vahl, 1802:44 [type locality: St. Croix, Virgin Is.].

Acanthophora spicifera (Vahl) Børgesen, 1910:201.—Weber-van Bosse, 1923:347.—Cantoria, Velasquez, and Valenzuela, 1951:296, footnote.—Domantay, 1962:287.—Galutira and Velasquez, 1964:510, pl. 5: fig. 18; pl. 9: figs. 37 a,b.—Velasquez, 1968a:122, fig. 9.—Villones and Magdamo, 1968:27, fig. 23.—Velasquez, 1971:449, fig. 36.—Reyes, 1972:158.—Trono, 1972a:106.—Velasquez, 1972:63.—Trono, 1973d:20, pl. II: fig. 10.—Trono and Biña, 1973:11.—Velasquez et al., 1973:32, pl. 14: fig. 67.—Westernhagen, 1973a:64.—Ortega, Alcala, and Reyes, 1974:185, 186, 187.—Trono, 1974b:96.—Westernhagen, 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:125.—Cordero, 1977a:189, figs. 201, 203, pl. XXVI:A; 1978a:56.—Vannajan and Trono, 1978:25, fig. 36.—Trono, 1978:19.—Cordero, 1979b:295.—Puig and Cordero, 1979:42.—Velasquez, 1979b:230.—Cajipe et al., 1980:69.—Cordero, 1980b:61, pl. 40.—Liao and Sotto, 1980:99.—Moreland, 1980:43, 46.—Reyes, 1980:139, pl. 12: fig. 3.—Trono and Fortes, 1980:74.—Trono and Ganzon-Fortes, 1980:99, fig. [s.n.].—Trono, Velasquez, and Guevarra, 1980:79.—Ganzon-Fortes, 1981:22.—Guzman, 1981:42.—Laserna et al., 1981:445.—Meñez and Calumpong, 1981:381.—Trono and Azanza-Corrales, 1981:744.—Trono and De Lara, 1981:19.—Calumpong, 1982:145.—Cordero, 1982a:60, 61.—Saraya and Trono, 1982:49.—Trono and Fortes, 1982:153.—Buchan-Antalan and Trono, 1983:19, fig. 2.—Hurtado-Ponce, 1983:144.—Cordero, 1984a:105; 1984b:66; 1984c:55.—Hurtado-Ponce, 1984:180.—Marcos-Agngarayngay, 1984a:58, fig. 48; 1984b:129.—Tungpalan, 1984:141.—Trono and Ganzon-Fortes, 1985:63.

Fucus acanthophorus Lamouroux, 1805:61, pl. XXX; pl. XXXI: fig. 1 [type locality: “Amérique septentrionale”].

Acanthophora thierryi Lamouroux, 1813:132 [“thierii”]—Montagne, 1844a:662.—Martens, 1868:100–101.—Velasquez, Trono, and Doty, 1975:125.

*Acanthophora orientalis J. Agardh, 1863 [1851–1863]:820 [syntype localities: Marianas; “ad Manillam,” Luzon].—Piccone, 1886:81, 90.—Heydrich, 1894:296.—Mazza, 1909:16—De Toni, 1924:380.—Collado, 1926:129.—Howe, 1932:169, 170.—G. Blanco, 1938:512.—Montilla and Blanco, 1953:166.—Domantay, 1962:288.—Westernhagen, 1973a:64; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:125.

Acanthophora spicifera (Vahl) Børgesen f. orientalis (J. Agardh) Weber-van Bosse, 1923:348.—Zaneveld, 1956:25 [var. orientalis]; 1959:106 [var. orientalis].—Bersamin et al., 1973:184 [var. orientalis].

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Ilocos Sur, Pangasinan, Rizal, Manila, Cavite, Batangas, Quezon, Albay. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. MASBATE. SAMAR: Western Samar, Eastern Samar. LEYTE (Biliran I.). PANAY: Aklan. GUIMARAS. NEGROS: Negros Occidental (incl. Ilacaon I.), Negros Oriental (incl. Apo I.). CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Zamboanga, Surigao del Sur, Davao. PALAWAN. SULU: Sulu (incl. Siasi I.) Tawitawi.

NOTE.—The conspecificity of Fucus spiciferus and Acanthophora thierryi was proposed by Børgesen (1910:201). Acanthophora orientalis was added as a synonym by Weber-van Bosse (1923:348). Lamouroux, when establishing the genus Acanthophora on the basis of Fucus acanthophorus Lamouroux, was obliged to change the epithet in order to avoid the creation of a tautonym, which is proscribed by Article 23.4 of the ICBN. The new epithet was chosen to commemorate “M. Thierry fils, professeur de physique et de chimie à…Caen”. Lamouroux’s incorrect original spelling is automatically correctable in accordance with Article 73.1 of the ICBN.

Acrocystis Zanardini

Acrocystis nana Zanardini

Acrocystis nana Zanardini, 1872:145, pl. VIII:A [type locality: “Tangion Datu” (Cape Datu), boundary between Sarawak, Malaysia and West Kalimantan, Indonesia].—Cordero, 1981a:173, fig. 2.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. CENTRAL VISAYAS.

Alsidium C. Agardh

Alsidium pusillum Dawson

Alsidium pusillum Dawson, 1963:20, pl. 3: figs. 1–4 [type locality: Academy Bay, Isla Santa Cruz, Galápagos].—Vannajan and Trono, 1978:28, fig. 34 [A. pusillum prox.].

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

Amansia Lamouroux

Amansia glomerata C. Agardh

Amansia glomerata C. Agardh, 1822a:194 [syntype localities: Hawaiian Is.; “Ravak” (Lawak), Waigeo I., Moluccas, Indonesia].—J. Agardh, 1863 [1851–1863]:1111.—Domantay, 1962:288.—Reyes, 1972:159.—Trono, 1972a:108; 1973a:139, fig. 21.—Westernhagen, 1973a:64; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:126.—Cordero, 1976c:8; 1977a:192, fig. 205, 268. 1978a:54.—Trono, 1978:19.—Trono and Tuason, 1978:15.—Liao and Sotto, 1980:99.—Reyes, 1980:140, pl. 12: fig. 5.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.—Saraya and Trono, 1982.49, pl. IX: fig. 2.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Quezon, Sorsogon. CATANDUANES. MARINDUQUE. NEGROS: Negros Oriental, CEBU (incl. Mactan I.). SIQUIJOR. BASILAN. PALAWAN, SULU: Tawitawi.

Bostrychia Montagne

Bostrychia binderi Harvey

Bostrychia binderi Harvey, 1849 [1847–1849]:68, pl. XXVIII [in part] [type locality: Port Natal (Durban), South Africa].—Post, 1936:28, 30, 33; 1939:23.—Villones and Magdamo, 1968:28, fig. 27.—Trono, 1973a:138.—Cordero, 1977a:193, figs. 206, 208, 209.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:381.—Saraya and Trono, 1982:49, pl. X: figs. 1–3.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Pangasinan, Batangas, Quezon. MINDORO: Oriental Mindoro. SIQUIJOR. PALAWAN.

Bostrychia calliptera (Montagne) Montagne

Rhodomela calliptera Montagne, 1840:197, pl. 5: fig. 2 [type locality: Cayenne, French Guiana].

Bostrychia calliptera (Montagne) Montagne, 1842b:661.—Post, 1938:206.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon.

Bostrychia intricata (Bory de Saint-Vincent) Montagne

Scytonema intricatum Bory de Saint-Vincent, 1829 [1826–1829]:225 [“intricata”] [lectotype locality: Falkland Is. fide Post, 1936:36, 40].

Bostrychia intricata (Bory de Saint-Vincent) Montagne, 1852:317.

Bostrychia mixta J. Hooker and Harvey, 1845a:270 [type locality: Bay of Islands, New Zealand].

NOTE.—The conspecificity of Bostrychia intricata and B. mixta was proposed by Post (1936:10, 36, 40), who adopted the later of the two names without comment. As pointed out by Papenfuss (1964:59), Scytonema intricatum is invalid because the generic name Scytonema is invalid (Articles 13.2 and 43 of the ICBN), being originally applied to a group of Cyanophyceae (Nostocaceae heterocysteae) with an 1886 starting point (Article 13.1(e)). With the abandonment of later starting points in the Cyanophyceae, however, Scytonema intricatum becomes available.

TAXON OF UNCERTAIN VALUE

Philippine records for Bostrychia intricata are under the following infraspecific taxon, which has not been formally transferred to it.

Bostrychia mixta J. Hooker and Harvey f. inermis Post

Bostrychia mixta J. Hooker and Harvey f. inermis Post, 1936:7, 42 [syntype localities: Indonesia; Queensland, Australia].—Post, 1938:211.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal.

Bostrychia kelanensis Grunow

Bostrychia kelanensis Grunow in Post, 1936:20 [type locality: Kelana, Papua New Guinea].—Post, 1936:20, 21, 22; 1938:205.—Tanaka, 1967:20, figs. 9, 10, pl. II:A.—Velasquez, Trono, and Doty, 1975:129.—Cordero, 1977a:193.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Rizal. SULU: Sulu.

Bostrychia moritziana (Sonder ex Kützing) J. Agardh

Polysiphonia (?) moritziana Sonder ex Kützing, 1849:838 [syntype localities: Antilles; Cayenne, French Guiana].

Bostrychia moritziana (Sonder ex Kützing) J. Agardh, 1863 [1851–1863]:862.—Post, 1938:204; 1939:15.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal, Quezon.

Bostrychia radicans (Montagne) Montagne

Rhodomela radicans Montagne, 1840:198, pl. 5: fig. 3 [type locality: Cayenne, French Guiana].

Bostrychia radicans (Montagne) Montagne, 1842b:661.—Post, 1938:205; 1939:15, 16.—Hamoy and Garciano, 1975:71.—Fortes and Trono, 1980:65, fig. 9.—Trono and Fortes, 1980:74.—Fortes, 1981b:396.—Ganzon-Fortes, 1981:22.—Trono and Fortes, 1982:153.

Bostrychia rivularis Harvey, 1853:57, pl. XIV:C [syntype localities: various, all in Atlantic USA].—Hamoy and Garciano, 1975:71.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal, Quezon. MINDORO: Oriental Mindoro. CEBU

NOTE.—The synonymy was proposed by Post (1936:13–19).

Bostrychia simpliciuscula Harvey ex J. Agardh

Bostrychia simpliciuscula Harvey ex J. Agardh, 1863 [1851–1863]:854 [type locality: Tonga].

Bostrychia tenuis Post f. simpliciuscula (Harvey ex J. Agardh) Post, 1936:23.—Post, 1938:205; 1939:16.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal, Quezon.

NOTE.—Post (1936:22) erred in establishing a new species, Bostrychia tenuis, to which she assigned two existing species, B. simpliciuscula and B. andoi Okamura 1907.

Bostrychia tenella (Lamouroux) J. Agardh

Fucus tenellus Vahl, 1802:45 [type locality: St. Croix, Virgin Is.].

Plocamium tenellum Lamouroux, 1813:138.

Bostrychia tenella (Lamouroux) J. Agardh, 1863 [1851–1863]:869.—Post, 1936:25, 26; 1938:206.—Trono, 1973a:138.—Cordero, 1976c:10; 1977a:195, fig. 210; 1978a:52; 1979b:295.—Reyes, 1980:140, pl. 12, fig. 6.—Meñez and Calumpong, 1981:381.—Cordero, 1984b:67.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Quezon. MINDORO: Oriental Mindoro. PANAY: Aklan. SIQUIJOR. PALAWAN.

NOTE.—Fucus tenellus Vahl, the intended basionym of Bostrychia tenella, is a later homonym of F. tenellus Esper (1800:197, pl. CIX) and hence not priorable. Plocamium tenellum Lamouroux is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Chondria C. Agardh

Chondria armata (Kützing) Okamura

Lophura armata Kützing, 1866:2, pl. 3: figs. a, b [type locality: New Caledonia].

Chondria armata (Kützing) Okamura, 1907 [1907–1909]:69.—Quisumbing, 1951:1008.—Cordero, 1977a:195, fig. 211, pl. XXVI:C; 1980b:62.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Quezon, Sorsogon. CENTRAL VISAYAS.

Chondria crassicaulis Harvey

Chondria crassicaulis Harvey, 1860a:330 [syntype localities: Shimoda and Hakodate, Japan].—Cordero, 1977a:196, fig. 212.

PHILIPPINE DISTRIBUTION.—PALAWAN.

Chondria curvilineata Collins and Hervey

Chondria curvilineata Collins and Hervey, 1917:120, pl. II: figs. 10, 11 [type locality: Heron Bay, Bermuda].—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Chondria dasyphylla (Woodward) C. Agardh

Fucus dasyphyllus Woodward, 1794:239, pl. 23: figs. 1–3 [syntype localities: Cromer and Yarmouth, Norfolk, England].

Chondria dasyphylla (Woodward) C. Agardh, 1817:xviii.—Weber-van Bosse, 1923:352.—Velasquez, Trono, and Doty, 1975:135.—Cordero, 1977a:196, fig. 213; 1978a:55.—Chan, 1981:387.—Cordero, 1984a:106.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan. LEYTE. SULU: Sulu (North Ubian I.).

Chondria polyrhiza Collins and Hervey

Chondria polyrhiza Collins and Hervey, 1917:121, pl. II: fig. 12 [type locality: Shelly Bay, Bermuda].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Chondria repens Børgesen

Chondria repens Børgesen, 1924:299, figs. 40, 41 [type locality: Isla de Pascua (Easter Island)].—Cordero, 1977a:198, fig. 214; 1980b:63—Saraya and Trono, 1982:50.—Cordero, 1984a:106.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Cavite (Corregidor I.), SIQUIJOR.

Chondria sedifolia Harvey

Chondria sedifolia Harvey, 1853:19, pl. XVIII:G [type locality: Key West, Florida, USA].—Reyes, 1980:141, pl. 13, fig. 5.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Chondria seticulosa (Forsskål) C. Agardh

Conferva seticulosa Forsskål, 1775:188 [type locality: Mokha, Yemen].

Chondria seticulosa (Forsskål) C. Agardh, 1822a:345.

Laurencia seticulosa (Forsskål) Greville, 1830:lii.—Quisumbing, 1951:1010.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Norte.

*Chondria sibogae Weber-van Bosse

Chondria sibogae Weber-van Bosse, 1923:350, fig. 134. [syntype localities: Lombok I. and Borneo Bank, Indonesia; North Ubian I., Sulu Prov., Sulu Archipelago].—Velasquez, Trono, and Doty, 1975:135.

PHILIPPINE DISTRIBUTION.—As above.

Digenea C. Agardh

Digenea simplex (Wulfen) C. Agardh

Conferva simplex Wulfen, 1803:17 [type locality: Trieste, Italy].

Digenea simplex (Wulfen) C. Agardh, 1822a:389.—G. Blanco, 1938 512.—Quisumbing, 1951:49.—Sulit, Navarro, and San Juan, 1952:167.—Montilla and Blanco, 1953:166, fig. 5:3.—Antonio, 1962:13.—Domantay, 1968:26.—Velasquez, 1971:449, fig. 35.—Trono, 1973a:140, fig. 16.—Cordero, 1976c:6, 8, 10; 1977a:200, figs. 216, 217; 1978a:52; 1980b:63, fig. 24, pl. 41.—Moreland, 1980:51.—Trono and Fortes, 1980:74.—Ganzon-Fortes, 1981:22.—Trono and Fortes, 1982:153.—Tungpalan, 1984:138.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Bataan, Manila, Batangas, Quezon, Sorsogon. PALAWAN.

Enantiocladia Falkenberg

Enantiocladia okamurae Yamada

Enantiocladia okamurae Yamada, 1930b:27 [“okamurai”] [syntype localities: various, all in Japan.]—Cordero, 1977a:201, fig. 220.—Marcos-Agngarayngay, 1984a:60, fig. 50.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Endosiphonia Zanardini

Endosiphonia spinuligera Zanardini

Endosiphonia spinuligera Zanardini, 1878:35 [type locality: Wokam I., Aru Is., Indonesia].—Weber-van Bosse, 1923:354.—Velasquez, Trono, and Doty, 1975:142.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Pearl Bank, Sangasiapu I.).

Exophyllum Weber-van Bosse

*Exophyllum wentii Weber-van Bosse

Exophyllum wentii Weber-van Bosse, 1911:29 [syntype localities: Savu I. and Borneo Bank, Indonesia; North Ubian I., Sulu Prov., Sulu Archipelago].—Weber-van Bosse, 1928:478.—Velasquez, Trono, and Doty, 1975:144

PHILIPPINE DISTRIBUTION.—As above.

NOTE.—This alga, long considered of uncertain taxonomic position, was assigned to the Rhodomelaceae by Hollenberg (1968d:81) on the basis of newly discovered spermatangial plants.

Herposiphonia Nägeli

Herposiphonia crassa Hollenberg

Herposiphonia crassa Hollenberg, 1968c:539, figs. 12, 13 [type locality: Kahakuloa Bay, Maui, Hawaiian Is.].—Saraya and Trono, 1982:51.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Herposiphonia delicatula Hollenberg

Herposiphonia delicatula Hollenberg, 1968c:540, figs. 1A,B, 2H, 3 [including Philippine record] [type locality: Falas I., Truk Is., Caroline Is.].—Velasquez, Trono, and Doty, 1975:149.

PHILIPPINE DISTRIBUTION.—MINDANAO: Davao.

Herposiphonia dendroidea Hollenberg

Herposiphonia dendroidea Hollenberg, 1968c:543, figs. 1C–E, 9 [type locality: Raroia Atoll, Tuamotu Archipelago].—Fortes and Trono, 1980:65, fig. 12.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LEYTE (Biliran I.).

Herposiphonia dendroidea var. minor Hollenberg

Herposiphonia dendroidea var. minor Hollenberg, 1968c:543, figs. 7, 24 [type locality: Raroia Atoll, Tuamotu Archipelago].—Fortes and Trono, 1980:66, fig. 14.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—SAMAR: Eastern Samar (Manicani I.), CEBU.

Herposiphonia nuda Hollenberg

Herposiphonia nuda Hollenberg, 1968c:548, figs. 2E,F, 8 [type locality: Ilio Point, Molokai, Hawaiian Is.].—Chan, 1981:387.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Herposiphonia obscura Hollenberg

Herposiphonia obscura Hollenberg, 1968c:549, fig. 25 [type locality: Ewa Beach, Oahu, Hawaiian Is.].—Fortes and Trono, 1980:66, fig. 13.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—CEBU.

Herposiphonia pacifica Hollenberg

Herposiphonia pacifica Hollenberg, 1968c:549, figs. 2A,B, 4, 19 [including Philippine record] [type locality: Maalea, Maui, Hawaiian Is.].—Velasquez, Trono, and Doty, 1975:149.

PHILIPPINE DISTRIBUTION.—MINDANAO: Surigao.

Herposiphonia parca Setchell

Herposiphonia parca Setchell, 1926:103, pl. 20: fig. 2 [type locality: Arue Reef, Tahiti].—Hollenberg, 1968c:552.—Velasquez, Trono, and Doty, 1975:149.—Fortes and Trono, 1980:66.—Chan, 1981:387.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. LEYTE. MINDANAO: Davao. PALAWAN (Balabac I.), SULU: Sulu (Cagayan Sulu I., Siasi I.).

Herposiphonia plumula (J. Agardh) Falkenberg

Polysiphonia plumula J. Agardh, 1885:99 [type locality: Santa Barbara, California, USA].

Herposiphonia plumula (J. Agardh) Falkenberg, 1901:728.—Marcos-Agngarayngay, 1984a:60, fig. 51 [miscited fig. 53].

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Herposiphonia secunda (C. Agardh) Ambronn

Hutchinsia secunda C. Agardh, 1824:149 [type locality: Mediterranean Sea].

Herposiphonia secunda (C. Agardh) Ambronn, 1880:197.

Herposiphonia tenella (C. Agardh) Ambronn f. secunda (C. Agardh) Hollenberg, 1968c:556.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS. MINDANAO: Surigao.

Herposiphonia secunda (C. Agardh) Ambronn f. tenella (C. Agardh) Wynne

Hutchinsia tenella C. Agardh, 1828:105 [type locality: Sicily].

Herposiphonia tenella (C. Agardh) Ambronn, 1880:197.—Hollenberg, 1968c:555.—Cornejo and Velasquez, 1972:181, pl. 3: fig. 22.—Velasquez, Trono, and Doty, 1975:149.—Vannajan and Trono, 1978:26.—Chan, 1981:387, 389.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:382.—Saraya and Trono, 1982:50, pl. X: fig. 4.

Herposiphonia secunda (C. Agardh) Ambronn f. tenella (C. Agardh) Wynne, 1985a:173.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite, Batangas. CENTRAL VISAYAS. PALAWAN (Balabac I.), SULU: Sulu (Siasi I.).

NOTE.—Hollenberg (1968c:556) reduced Herposiphonia secunda to the rank of a form of Herposiphonia tenella. Because Hutchinsia secunda C. Agardh (1824:149), the basionym of Herposiphonia secunda, is older than Hutchinsia tenella C. Agardh (1828:105), the basionym of Herposiphonia tenella, the relationship of the two taxa must be reversed.

Herposiphonia subdisticha Okamura

Herposiphonia subdisticha Okamura, 1899a:37, pl. 1: figs. 12–14 [syntype localities: Enoshima, Kanagawa Prefecture and “Boshu” (Chiba Prefecture), Japan].—Cordero, 1976c:8, 10; 1977a:202, figs. 218, 219; 1978a:53; 1984a:106.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte.

Herposiphonia trichia Hollenberg

Herposiphonia trichia Hollenberg, 1968c:557, fig. 1G [type locality: Yap I. Caroline Is.].—Fortes and Trono, 1980:66.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

Laurencia Lamouroux

Laurencia brongniartii J. Agardh

Laurencia brongniartii J. Agardh, 1841:20 [type locality: Martinique].—Cordero, 1977a:204, pl. XXVII:A.—Hurtado-Ponce, 1983:145; 1984:180.—Marcos-Agngarayngay, 1984a:55, fig. 45; 1984b:129.

Laurencia concinna Montagne, 1842a:6 [type locality: Toud I. [Warrior Islet], Torres Strait, Australia].—Dickie, 1876a:245.—Weber-van Bosse, 1923:346.—Velasquez, Trono, and Doty, 1975:151.

Laurencia grevilleana Harvey, 1855:545 [type locality: Rottnest I., Western Australia, Australia].—Tahil, 1978:52.—Guzman, 1981:44.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, CEBU (Mactan I.). MINDANAO: Zamboanga. SULU: Sulu (North Ubian I., Tongquil I.).

NOTE.—The synonymy was proposed by Saito and Womersley 1974:839).

Laurencia capituliformis Yamada

Laurencia capituliformis Yamada, 1931b:217, pl. 14 [type locality: O-shima, Aomori Prefecture, Japan].—Cordero, 1977a:205.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Laurencia caraibica P.C. Silva

Laurencia nana Howe, 1920:566 [type locality: Mariguana (Mayaguana) I., Bahama Is.] [replaced name].—Meñez and Calumpong, 1981:382.

Laurencia caraibica P.C. Silva, 1972:205.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

NOTE.—Laurencia caraibica is a substitute name for L. nana Howe, a later homonym of L. nana (C. Agardh) Greville (1830:lii).

Laurencia carolinensis Saito

Laurencia carolinensis Saito, 1969:154, figs. 6, 7 [type locality: Helen Reef, Caroline Is.].—Cordero, 1977a:205.

PHILIPPINE DISTRIBUTION.—PALAWAN.

Laurencia cartilaginea Yamada

Laurencia cartilaginea Yamada, 1931b:230, fig. O, pl. 19: fig. a [syntype localities: Chikuzen and Iyo provinces (Fukuoka and Ehime prefectures), Japan].—Meñez, 1961:76.—Saito, 1968:89; 1969:154.—Velasquez, 1971:453, fig. 38.—Cornejo and Velasquez, 1972:181, 182, 183.—Kraft, 1972:332.—Reyes, 1972:159.—Trono, 1972a:88; 1973d:22, pl. II: fig. 11.—Trono and Biña, 1973:11.—Velasquez et al., 1973:32, pl. 14: fig. 68.—Ortega, Alcala, and Reyes, 1974:187.—Trono, 1974b:97; 1974e:18.—Velasquez, Trono, and Doty, 1975:151.—Cordero, 1977a:206, fig. 221; 1978a:57.—Puig and Cordero, 1979:42.—Liao and Sotto, 1980:99.—Trono and Fortes, 1980:75.—Trono and Ganzon-Fortes, 1980:101, fig. [s.n].—Ganzon-Fortes, 1981:22.—Trono and De Lara, 1981:21, pl. XII: fig. 1.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Bataan, Batangas, Quezon, MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SIQUIJOR. SAMAR: Western Samar. LEYTE (incl. Biliran I.), PANAY: Iloilo. NEGROS: Negros Oriental. CEBU (Mactan I.). MINDANAO: Zamboanga (Sacol I.), Surigao. PALAWAN (incl. Cuyo I.), SULU: Sulu (Siasi I.), Tawitawi.

Laurencia ceylanica J. Agardh

Laurencia ceylanica J. Agardh, 1876:660 [type locality: Sri Lanka].—Meñez, 1961:76.—Velasquez, Trono, and Doty, 1975:151.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Laurencia chondrioides Børgesen

Laurencia chondrioides Børgesen, 1918:252, figs. 243–246 [type locality: St. Jan (St. John), Virgin Is.].—Westernhagen, 1973a:65; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Laurencia clavata Sonder

Laurencia clavata Sonder, 1853:694 [type locality: Lefevre Peninsula, South Australia, Australia].—Weber-van Bosse, 1923:346.—Velasquez, Trono, and Doty, 1975:151.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Tongquil I.).

Laurencia columellaris Børgesen

Laurencia columellaris Børgesen, 1945:53, figs. 28–30 [type locality: Réunion].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Laurencia composita Yamada

Laurencia composita Yamada, 1931b:236, figs. R, S, pl. 23 [syntype localities: Enoshima (Kanagawa Prefecture) and Mera (Chiba Prefecture), Japan].—Cordero, 1977a:206, pl. XXVIII:A.—Marcos-Agngarayngay, 1984a:55, fig. 46; 1984b:130.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte. PALAWAN.

Laurencia corallopsis (Montagne) Howe

Sphaerococcus corallopsis Montagne, 1842c:49, pl. III: fig. 1 [type locality: La Habana, Cuba].—Martens, 1868:31, 96–97.—Velasquez, Trono, and Doty, 1975:151.

Laurencia corallopsis (Montagne) Howe, 1918:519.—Cordero, 1977a:208, fig. 222; 1978a:57.

PHILIPPINE DISTRIBUTION.—BATANES. MINDANAO: Zamboanga.

Laurencia decumbens Kützing

Laurencia decumbens Kützing, 1863:18 [type locality: New Caledonia].—Cordero, 1977a:208.

PHILIPPINE DISTRIBUTION.—SAMAR: Western Samar.

Laurencia distichophylla J. Agardh

Laurencia distichophylla J. Agardh, 1852 [1851–1863]:762 [type locality: “in oceano australi?”].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Laurencia forsteri (Mertens ex Turner) Greville

Fucus forsteri Mertens ex Turner, 1809:15, pl. 77 [lectotype locality: Australia (probably King George Sound, Western Australia) fide Saito and Womersley, 1974:824].

Laurencia forsteri (Mertens ex Turner) Greville, 1830:lii.—Cordero, 1977a:208.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Laurencia glandulifera (Kützing) Kützing

Chondria glandulifera Kützing, 1845:329 [type locality: Trieste, Italy].

Laurencia glandulifera (Kützing) Kützing, 1849:855.—Westernhagen, 1973a:65; 1974:112 (table 1).—Cordero, 1977a:209, fig. 223, pl. XXVII:D; 1984a:107.—Marcos-Agngarayngay, 1984b:130.—Tungpalan, 1984:141.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, CEBU (Mactan I.), SULU.

Laurencia implicata J. Agardh

Laurencia implicata J. Agardh, 1852 [1852–1863]:745 [type locality: St. Croix, Virgin Is.].

Chondria intricata Montagne, 1842c:41 [type locality: Cuba].

Laurencia intricata (Montagne) Lamouroux ex J. Agardh, 1852 [1852–1863]:750 [footnote].—Cordero, 1976c:8; 1977a:210, figs. 225, 226; 1978a:58.—Trono and Ang, 1982:25.

PHILIPPINE DISTRIBUTION.—BATANES. PALAWAN (Bugsuk I.).

NOTE.—The name Laurencia intricata Lamouroux has been applied to this species, but it is not correct. Lamouroux (1813:131, pl. 9: figs. 8, 9) failed to provide a description, as did C. Agardh (1817:XVIII) when transferring the species to Chondria. The first description was provided by Montagne under the name Chondria intricata. The species was returned to Laurencia by J. Agardh, but the resulting binomial is a later homonym of L. intricata Suhr (1840:265). The conspecificity of L. implicate J. Agardh and L. intricata (Montagne) Lamouroux ex J. Agardh was proposed by Howe (1918:518).

Laurencia intermedia Yamada

Laurencia intermedia Yamada, 1931b:191, pl. 1: fig. c; pl. 2 [type locality: Enoshima, Kanagawa Prefecture, Japan].—Westernhagen, 1973a:65; 1974:112 (table 1).—Cordero, 1977a:209, fig. 224; 1978a:57.—Guzman, 1981:42, 46, 47, 48.—Hurtado-Ponce, 1983:145.—Cordero, 1984a:107.—Hurtado-Ponce, 1984:180.—Tungpalan, 1984:141.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, CEBU (Mactan I.).

Laurencia japonica Yamada

Laurencia japonica Yamada, 1931b:211, fig. L, pl. II [syntype localities: Amatsu and Emi, Chiba Prefecture, Japan].—Meñez, 1961:77.—Saito, 1968:86; 1969:152.—Velasquez, Trono, and Doty, 1975:151.—Trono and Ang, 1982:25.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Quezon. PALAWAN (incl. Bugsuk I.).

Laurencia majuscula (Harvey) Lucas

Laurencia obtusa (Hudson) Lamouroux var. majuscula Harvey, 1863: xxvi [syntype localities: Rottnest I., Western Australia and Cape Shank, Bass Strait, Australia].

Laurencia majuscula (Harvey) Lucas, 1935:223.—Saito, 1968:84; 1969:149.—Velasquez, Trono, and Doty, 1975:151.—Cordero, 1977a:210, fig. 227.—Puig and Cordero, 1979:42.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Albay. LEYTE (incl. Biliran I.), SIQUIJOR. PALAWAN.

Laurencia mariannensis Yamada

Laurencia mariannensis Yamada, 1931b:200, figs. F, G, pl. 5: fig. b [type locality: Saipan, Mariana Is.].—Saito, 1969:151.—Trono and Biña, 1973:11.—Velasquez, Trono, and Doty, 1975:151.—Saraya and Trono, 1982:52.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro.

Laurencia nidifica J. Agardh

Laurencia nidifica J. Agardh, 1852[1852–1863]:749 [type locality: Hawaiian Is.].—Cordero, 1977a:211, fig. 228.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Bataan.

Laurencia obtusa (Hudson) Lamouroux

Fucus obtusus Hudson, 1778:586 [syntype localities: Hastings, Sussex and Devonshire, England].

Laurencia obtusa (Hudson) Lamouroux, 1813:130.—Montagne, 1844a:662.—Martens, 1868:100–101.—Dickie, 1874a:192.—Cantoria, Velasquez, and Valenzuela, 1951:296, footnote.—Meñez, 1961:76.—Saito, 1968:84 [var. obtusa]; 1969:150 [var. obtusa].—Westernhagen, 1973a:65; 974:112 (table 1).—Velasquez, Trono, and Doty, 1975:151.—Cordero, 1977a:211, fig. 229; 1977a:213 [var. obtusa]; 1978a:58; 1979b:294; 1980b:64, pl. 42 [var. obtusa].—Trono and Fortes, 1980:75.—Fortes, 198lb:396.—Ganzon-Fortes, 1981:22.—Meñez and Calumpong, 1981:382.—Cordero, 1982a: fig. 14.—Saraya and Trono, 1982:53.—Trono and Ang, 1982:25.—Trono and Fortes, 1982:154.—Cordero, 1984a:107 [var. obtusa]; 1984c:55.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Albay. MASBATE. PANAY: Aklan. CEBU (Mactan I.). PALAWAN (Bugsuk I.).

Laurencia obtusa (Hudson) Lamouroux var. dendroidea (J. Agardh) Yamada

Laurencia dendroidea J. Agardh 1852 [1852–1863]:753 [type locality: Brazil].—Weber-van Bosse, 1923:343.—Velasquez, Trono, and Doty, 1975:151.

Laurencia obtusa (Hudson) Lamouroux var. dendroidea (J. Agardh) Yamada, 1931b:224.—Saito, 1968:84; 1969:150.

PHILIPPINE DISTRIBUTION.—MINDANAO, SULU: Sulu (North Ubian I.).

Laurencia obtusa (Hudson) Lamouroux var. densa Yamada

Laurencia obtusa (Hudson) Lamouroux var. densa Yamada, 1931b:226 [type locality: Daibanratsu, Taiwan].—Cordero, 1977a:213.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Laurencia obtusa (Hudson) Lamouroux var. snackeyi (Weber-van Bosse) Yamada

Laurencia paniculata (C. Agardh) J. Agardh var. snackeyi Weber-van Bosse, 1923:342 [type locality: Semau I., Indonesia].

Laurencia obtusa (Hudson) Lamouroux var. snackeyi (Weber-van Bosse) Yamada, 193lb:225.—Saito, 1968:85; 1969:151.

PHILIPPINE DISTRIBUTION.—MINDANAO.

Laurencia okamurae Yamada

Laurencia okamurae Yamada, 1931b:206, figs. J, K, pl. 7 [“okamurai”] [syntype localities: various, in Japan and China].—Galutira and Velasquez, 1964:509, pl. 5: fig. 17; pl. 9: fig. 36.—Velasquez, 1972:63.—Cordero, 1977a:213, fig. 231; 1979b:294.—Velasquez, 1979b:230.—Trono and Fortes, 1980:75.—Ganzon-Fortes, 1981:23.—Guzman, 1981:43, 49.—Cordero, 1982a:60.—Trono and Fortes, 1982:154.—Marcos-Agngarayngay, 1984b:130.—Tungpalan, 1984:141.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Quezon, PANAY: Aklan.

Laurencia palisada Yamada

Laurencia palisada Yamada, 193lb:196, figs. C, D, pl. 4: fig. a [syntype localities: Kotosho (Hung-t’ou) and Takao, Taiwan].—Cordero, 1977a:214, fig. 232, pl. XXVII: C, E; 1978a:59.

PHILIPPINE DISTRIBUTION.—BATANES. SIQUIJOR.

Laurencia papillosa (C. Agardh) Greville

Fucus papillosus Forsskål, 1775:190 [type locality: Mokha, Yemen].

Chondria papillosa C. Agardh, 1822a:344.

Laurencia papulosa (C. Agardh) Greville, 1830:lii.—Montagne, 1844a:662.—Bailey and Harvey, 1862:713.—Martens, 1868:100–101.—Dickie, 1876a:244.—Quisumbing, 1951:1011.—Zaneveld, 1956:44; 1959:123.—Domantay, 1962:292.—Galutira and Velasquez, 1964:509, pl. 5: fig. 16; pl. 9: fig. 35a,b.—Saito, 1968:92.—Villones and Magdamo, 1968:29, fig. 31.—Saito, 1969:158.—Velasquez, 1971:449, fig. 37.—Cornejo and Velasquez, 1972:172.—Kraft, 1972:328.—Reyes, 1972:159.—Trono, 1972a:107.—Velasquez, 1972:63.—Cordero, 1973b:33.—Bersamin et al., 1973:187.—Trono, 1973d:21, pl. II: fig. 9—Trono and Biña, 1973:12.—Velasquez et al., 1973:32, pl. 14: fig. 69.—Westernhagen, 1973a:65.—De Leon, 1974:31, 32 photo [s.n.], fig. [s.n.].—Ortega, Alcala, and Reyes, 1974:187.—Trono, 1974b:96; 1974e:18.—Westernhagen, 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:151.—Cordero, 1976c:10; 1977a:215, figs. 233–235; 1978a:58; 1979b:276.—Velasquez, 1979b:230.—Cordero, 1980b:64, fig. 25, pl. 43.—Liao and Sotto, 1980:100.—Reyes, 1980:139, pl. 12: fig. 4a,b.—Trono and Fortes, 1980:74.—Trono and Ganzon-Fortes, 1980:103, fig. [s.n.].—Ganzon-Fortes, 1981:23.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:21, pl. XII: fig. 2.—Calumpong, 1982:145.—Cordero, 1982a:60.—Saraya and Trono, 1982:51.—Trono and Fortes, 1982:154—Hurtado-Ponce, 1983:145.—Cordero, 1984a:107; 1984b:67; 1984c:56.—Hurtado-Ponce, 1984:180—Marcos-Agngarayngay, 1984a:57, fig. 47; 1984b:130.—Tungpalan, 1984:141.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Batangas, Quezon, Albay. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MASBATE. SAMAR: Western Samar, Eastern Samar. LEYTE (incl. Biliran I.), PANAY: Aklan, Antique (incl. Panagatan Cays), NEGROS: Negros Occidental (incl. Ilacaon I.), Negros Oriental (incl. Apo I.), CEBU (incl. Mactan I.). SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.), Surigao del Sur. PALAWAN (incl. Cuyo I.), SULU: Sulu (Siasi I.), Tawitawi.

NOTE.—Fucus papillosus Forsskål, the intended basionym of Laurencia papillosa, is a later homonym of F. papillosus S.G. Gmelin (1768:188) and hence not priorable. Chondria papillosa C. Agardh is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Laurencia parvipapillata Tseng

Laurencia parvipapillata Tseng, 1943:204, pl. IV [type locality: Cape d’Aguilar, Hong Kong].—Saito, 1968:93; 1969:159.—Velasquez, Trono, and Doty, 1975:151.—Meñez and Calumpong, 1981:382.—Saraya and Trono, 1982:52, pl. XI: fig. 2.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CENTRAL VISAYAS. PALAWAN.

Laurencia patentiramea (Montagne) Kützing

Chondria obtusa (Hudson) C. Agardh var. patentiramea Montagne, 1836:322, pl. 18: fig. 3 [type locality: Mèze, Hérault, France].

Laurencia patentiramea (Montagne) Kützing, 1849:854.

Chondria obtusa (Hudson) C. Agardh var. paniculata C. Agardh, 1822a:343 [type locality: Adriatic Sea].

Laurencia paniculata (C. Agardh) J. Agardh, 1852 [1852–1863]:755.—Cordero, 1977a:214, fig. 230; 1979b:294.—Meñez and Calumpong, 1981:382.—Saraya and Trono, 1982:52.—Trono and Ang, 1982:24.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CENTRAL VISAYAS. LEYTE (Biliran I.), PANAY: Aklan. PALAWAN (incl. Bugsuk I.), SULU: Sulu (Siasi I.)

NOTE.—The conspecificity of Laurencia patentiramea and L. paniculata was proposed by Ardissone (1883:329). As a species, L. patentiramea dates from 1843, as Chondria patentiramea (Montagne) Kützing (1843b:437), while L. paniculata dates from 1852. Moreover, L. paniculata (C. Agardh) J. Agardh is a later homonym of L. paniculata Kützing (1849:855).

Laurencia pinnata Yamada

Laurencia pinnata Yamada, 1931b:242, pl. 28 [type locality: Enoshima (Kanagawa Prefecture), Japan].—Ortega, Alcala, and Reyes, 1974:186, 187.—Velasquez, Trono, and Doty, 1975:151.—Cordero, 1976c:10; 1977a:217, fig. 236, pl. XXVII:B.—Cordero, 1978a:59 [L. pinnata prox.].—Hurtado-Ponce, 1983:145.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Batangas. NEGROS: Negros Oriental.

Laurencia poiteaui (Lamouroux) Howe

Fucus poiteaui Lamouroux, 1805:63, pl. XXX1: figs. 2, 3 [“poitei” ][type locality: Santo Domingo, Dominican Republic].

Laurencia poiteaui (Lamouroux) Howe, 1918:518 [“poitei”].—Westernhagen, 1973a:65; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

NOTE.—Lamouroux, in naming this species after its collector, Pierre Antoine Poiteau, incorrectly spelled the epithet poitei. This spelling is automatically correctable in accordance with Article 73.1 of the ICBN.

Laurencia subsimplex Tseng

Laurencia subsimplex Tseng, 1943:202, pl. III: figs. 4–6 [type locality: Big Wave Bay, Hong Kong].—Saito, 1968:87; 1969:153.—Velasquez, Trono, and Doty, 1975:151.

PHILIPPINE DISTRIBUTION.—CATANDUANES.

Laurencia surculigera Tseng

Laurencia surculigera Tseng, 1943:192, pl. I: figs. 4, 5 [type locality: Stanley Bay, Hong Kong].—Fortes and Trono, 1980:67.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

*Laurencia tronoi Ganzon-Fortes

Laurencia tronoi Ganzon-Fortes, 1983:404, figs. 1, 2 [type locality: Batangas Prov., Luzon].—Trono and Ganzon-Fortes, 1980:105, fig. [s.n.] [Laurencia sp.].—Ganzon-Fortes and Trono, 1982:27, figs. 1, 2 [Laurencia sp.].

PHILIPPINE DISTRIBUTION.—As above.

Laurencia tropica Yamada

Laurencia tropica Yamada, 1931b:233, figs. B, Q, pl. 20 [type locality: Saipan, Mariana Is.].—Cordero, 1977a:217, fig. 237.—Marcos-Agngarayngay, 1984b:130.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, SIQUIJOR.

Laurencia undulata Yamada

Laurencia undulata Yamada, 1931b:243, fig. T, pl. 29: fig. a [type locality: Enoshima, Kanagawa Prefecture, Japan].—Westernhagen, 1973a:65; 1974:112 (table 1).—Cordero, 1977a:219, figs. 238, 239; 1978a:59 [L. undulata prox.].—Tahil, 1978:52.—Puig and Cordero, 1979:42.—Cordero, 1980b:65, pl. 44; 1984a:108; 1984c:56.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Quezon. MASBATE. LEYTE (Biliran I.), CEBU (Mactan I.). SIQUIJOR.

Laurencia venusta Yamada

Laurencia venusta Yamada, 1931b:203, fig. H, pl. 6: fig. a [syntype localities: Koshiki-jima (Kagoshima Prefecture) and Goto (Nagasaki Prefecture), Japan].—Saraya and Trono, 1982:53.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Laurencia yamadana Howe

Laurencia yamadana Howe. 1934:37, fig. 4 [type locality: Kaneohe Bay, Oahu, Hawaiian Is.].—Trono, 1973d:22.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Leveillea Decaisne

Leveillea jungermannioides (Hering and Martens) Harvey

Amansia jungermannioides Hering and Martens in Martens and Hering, 1836:485, figs. 1–4 [type locality: Tor, Sinai Peninsula, Egypt].

Polyzonia jungermannioides (Hering and Martens) J. Agardh, 1841:25.—Piccone, 1886:84, 90.—Velasquez, Trono, and Doty, 1975:161.

Leveillea jungermannioides (Hering and Martens) Harvey, 1855:539 [“Leveillia”].—Meñez, 1961:75.—Reyes, 1972:159.—Cornejo and Velasquez, 1972:181, pl. 5: fig. 38.—Trono and Biña, 1973:12.—Trono, 1973d:21.—Velasquez, Trono, and Doty, 1975:151.—Cordero, 1976c:9; 1977a:221, fig. 244; 1978a:53.—Trono, 1978:20.—Trono and Tuason, 1978:16.—Vannajan and Trono, 1978:28, fig. 40.—Fortes and Trono, 1980:65.—Liao and Sotto, 1980:100.—Reyes, 1980:140, pl. 13: fig. 1.—Chan, 1981:387, 389.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:382.—Saraya and Trono, 1982:53, pl. XI: fig. 1.—Trono and Ang, 1982:22.—Marcos-Agngarayngay, 1984a:58, fig. 49.

Leveillea gracilis Decaisne, 1839:376 [type locality: Sri Lanka].—Martens, 1868:102–103.—Velasquez, Trono, and Doty, 1975:151.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Manila, Cavite, Batangas. CATANDUANES. MINDORO: Oriental Mindoro. MARINDUQUE. MASBATE (Ticao 1.), SAMAR: Western Samar. NEGROS: Negros Oriental, CEBU (Mactan I.), SIQUIJOR. MINDANAO: Zamboanga. PALAWAN (incl. Bugsuk I.).

NOTE.—The synonymy was proposed by Falkenberg (1901:392).

Lophocladia (J. Agardh) Schmitz

Lophocladia lallemandii (Montagne) Schmitz

Dasya lallemandii Montagne, 1849:289 (“lallemandi”] [type locality: Red Sea].

Lophocladia lallemandii (Montagne) Schmitz, 1893:223 [“lallemandi”].—Weber-van Bosse, 1923:362.—Velasquez, Trono, and Doty, 1975:152.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Pearl Bank).

Lophosiphonia Falkenberg

Lophosiphonia cristata Falkenberg

Lophosiphonia cristata Falkenberg, 1901:499, pl. 9: figs. 7–10 [type locality: Napoli, Italy].—Hollenberg, 1968d:80.—Velasquez, Trono, and Doty, 1975:152.

PHILIPPINE DISTRIBUTION.—CATANDUANES.

Murrayella Schmitz

Murrayella periclados (C. Agardh) Schmitz

Hutchinsia periclados C. Agardh, 1828:101 [type locality: St. Croix, Virgin Is.].

Murrayella periclados (C. Agardh) Schmitz, 1893:227.—Fortes and Trono, 1980:67, fig. 15.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Neurymenia J. Agardh

Neurymenia fraxinifolia (Mertens ex Turner) J. Agardh

Fucus fraxinifolius Mertens ex Turner, 1811:140, pl. 193 [type locality: “East Indies”].

Neurymenia fraxinifolia (Mertens ex Turner) J. Agardh, 1863 [1852–1863]:1135.—Dickie, 1876a:245.—Trono, 1973a:139, fig. 20.—Velasquez, Trono, and Doty, 1975:156.—Cordero, 1976c:10; 1977a:222, figs. 242, 245; 1978a:55.—Saraya and Trono, 1982:54.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Sorsogon. SAMAR: Eastern Samar. MINDANAO: Zamboanga.

Polysiphonia Greville

Polysiphonia apiculata Hollenberg

Polysiphonia apiculata Hollenberg, 1968a:61, figs. 1D, 8, 9 [including Philippine record] [type locality: Pokai Bay, Oahu, Hawaiian Is.].—Velasquez, Trono, and Doty, 1975:160.

PHILIPPINE DISTRIBUTION.—PALAWAN (Balabac I.).

Polysiphonia beaudettei Hollenberg

Polysiphonia beaudetti Hollenberg in Hollenberg and Dawson, 1961:348, pl. 1: fig. 4 [type locality: Isla Grande, Guerrero, Mexico].—Hollenberg, 1968a:62.—Velasquez, Trono, and Doty, 1975:160.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Siasi I.), Tawitawi (Turtle Is.).

Polysiphonia ferulacea Suhr ex J. Agardh

Polysiphonia ferulacea Suhr ex J. Agardh, 1863 [1852–1863]:980 [syntype localities: Atlantic Mexico; Guadeloupe; Australia; Marquesas Is.; Hawaiian Is.].—Weber-van Bosse, 1923:357.—Velasquez, Trono, and Doty, 1975:160.—Trono and Ang, 1982:23.

PHILIPPINE DISTRIBUTION.—PALAWAN (Bugsuk I.), SULU: Sulu (Jolo I.).

Polysiphonia flabellulata Harvey

Polysiphonia flabellulata Harvey, 1860a:330 [type locality: Tanega-shima, Osumi-gunto, Japan].—Vannajan and Trono, 1978:29.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal.

Polysiphonia forfex Harvey

Polysiphonia forcipata Harvey, 1855:541 [lectotype locality: Rottnest I., Western Australia, Australia fide Womersley, 1979:495] [replaced name].—Cordero, 1977a:223.

Polysiphonia forfex Harvey, 1859: pl. XCVI.

PHILIPPINE DISTRIBUTION.—BATANES.

NOTE.—Polysiphonia forfex is a substitute name for P. forcipata Harvey, a later homonym of P. forcipata J. Agardh (1842:127).

Polysiphonia fragilis Suringar

Polysiphonia fragilis Suringar, 1867:259 [type locality: “in mari Japonico”].—Cordero, 1978a:50.

PHILIPPINE DISTRIBUTION.—BATANES.

Polysiphonia gorgoniae Harvey

Polysiphonia gorgoniae Harvey, 1853:39 [type locality: Key West, Florida, USA].—Cornejo and Velasquez, 1972:173, 180, pl. 5: figs. 37, 37a, 37b.—Velasquez, Trono, and Doty, 1975:161.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas.

Polysiphonia hawaiiensis Hollenberg

Polysiphonia hawaiiensis Hollenberg, 1968a:66, figs. 2F, 16, 41 [including Philippine record] [type locality: Waikiki, Oahu, Hawaiian Is.].—Velasquez, Trono, and Doty, 1975:161.

PHILIPPINE DISTRIBUTION.—PALAWAN (Balabac I.), SULU: Tawitawi.

Polysiphonia howei Hollenberg

Polysiphonia howei Hollenberg in W.R. Taylor, 1945:302, fig. 3 [type locality: Whale Cay, Berry Is., Bahama Is.].—Hollenberg, 1968b:203.—Velasquez, Trono, and Doty, 1975:161.—Cordero, 1976c:10; 1977a:224, fig. 243; 1978a:50.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. MINDANAO: Davao.

Polysiphonia mollis J. Hooker and Harvey

Polysiphonia mollis J. Hooker and Harvey in Harvey, 1847 [1847–1849]:43 [type locality: Tasmania, Australia].—Weber-van Bosse, 1923:356 [with query].—Hollenberg, 1968a:69.—Velasquez, Trono, and Doty, 1975:161.—Cordero, 1977a:226, figs. 246–249; 1978a:51.—Meñez, Phillips, and Calumpong, 1983:23.—Meñez and Calumpong, 1984:105.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte. PALAWAN, SULU: Sulu (North Ubian I.).

Polysiphonia mollis J. Hooker and Harvey var. tongatensis (Harvey ex Kützing) Hollenberg

Polysiphonia tongatensis Harvey ex Kützing, 1864:14, pl. 41: figs, a–d [type locality: Tonga].

Polysiphonia mollis J. Hooker and Harvey var. tongatensis (Harvey ex Kützing) Hollenberg, 1968a:69 [including Philippine record].

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Siasi I.).

Polysiphonia pacifica Hollenberg

Polysiphonia pacifica Hollenberg, 1942:777, figs. 2, 3, 12, 13 [type locality: Santa Cruz, California, USA].—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Polysiphonia pacifica Hollenberg var. delicatula Hollenberg

Polysiphonia pacifica Hollenberg var. delicatula Hollenberg, 1942:778 [type locality: Monterey, California, USA].—Fortes and Trono, 1980:67.

PHILIPPINE DISTRIBUTION.—SAMAR: Eastern Samar (Manicani I.).

Polysiphonia poko Hollenberg

Polysiphonia poko Hollenberg, 1968a:70, fig. 3A [type locality: Johnson Atoll].—Chan, 1981:387.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Polysiphonia savatieri Hariot

Polysiphonia savatieri Hariot, 1891:226 [type locality: Yokosuka, Kanagawa Prefecture, Japan].—Hollenberg, 1968a:77.—Velasquez, Trono, and Doty, 1975:161.—Chan, 1981:387, 389.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDANAO: Surigao. PALAWAN (Balabac I.), SULU: Sulu (Cagayan Sulu I., Siasi I.).

Polysiphonia scopulorum Harvey

Polysiphonia scopulorum Harvey, 1855:540 [type locality: Rottnest I., Western Australia, Australia].—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Polysiphonia scopulorum Harvey var. villum (J. Agardh) Hollenberg

Polysiphonia villum J. Agardh, 1863 [1852–1863]:941 [type locality: “ad littus Americae tropicae” (Mexico)].

Polysiphonia scopulorum Harvey var. villum (J. Agardh) Hollenberg, 1968a:81 [including Philippine record].—Velasquez, Trono, and Doty, 1975:161 [without designation of variety].—Fortes and Trono, 1980:68.

PHILIPPINE DISTRIBUTION.—SAMAR: Eastern Samar (Manicani I.). MINDANAO: Davao.

Polysiphonia setacea Hollenberg

Polysiphonia setacea Hollenberg, 1968a:85, figs. 5A–C [including Philippine record] [type locality: Koko Head, Oahu, Hawaiian Is.].—Velasquez, Trono, and Doty, 1975:161.—Cordero, 1977a:226, fig. 253.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDANAO: Surigao, Davao.

Polysiphonia sparsa (Setchell) Hollenberg

Lophosiphonia sparsa Setchell, 1926:103, pl. 21: figs. 3, 4 [type locality: Arue Reef, Tahiti].

Polysiphonia sparsa (Setchell) Hollenberg, 1968a:87 [including Philippine record].—Velasquez, Trono, and Doty, 1975:161.

PHILIPPINE DISTRIBUTION.—PALAWAN (Balabac I.), SULU: Tawitawi.

Polysiphonia sphaerocarpa Børgesen

Polysiphonia sphaerocarpa Børgesen, 1918:271, figs. 267–271 [type locality: St. Thomas, Virgin Is.].—Cornejo and Velasquez, 1972:180, pl. 5: figs. 36, 36a,b.—Velasquez, Trono, and Doty, 1975:161.—Chan, 1981:387.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas.

Polysiphonia subtilissima Montagne

Polysiphonia subtilissima Montagne, 1840:199 [type locality: Cayenne, French Guiana].—Cordero, 1977a:226, figs. 250, 251.—Vannajan and Trono, 1978:28, fig. 35.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal, Manila, Cavite, Batangas.

Polysiphonia tepida Hollenberg

Polysiphonia tepida Hollenberg, 1958:65, fig. 1 [type locality: Beaufort, North Carolina, USA].—Cordero, 1977a:227, figs. 252, 254; 1978a:51.

PHILIPPINE DISTRIBUTION.—BATANES.

Polysiphonia triton P.C. Silva, new name

Polysiphonia tenuis Hollenberg, 1968a:92, figs. 6 B,C [type locality: Johnson Atoll] [replaced name],—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

NOTE.—Polysiphonia triton is proposed as a substitute name for P. tenuis Hollenberg, a later homonym of P. tenuis (C. Agardh) E.M. Fries (1836[1835–1837]:314).

Polysiphonia tsudana Hollenberg

Polysiphonia tsudana Hollenberg, 1968b:205, figs. 1F,G, 2C [type locality: Laysan Islet, Leeward Is., Hawaiian Is.].—Fortes and Trono, 1980:68, fig. 10.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—PANAY: Iloilo.

Polysiphonia upolensis Grunow

Polysiphonia upolensis Grunow, 1874:49 [type locality: Upolu I., Western Samoa].—Hollenberg, 1968a:94.—Velasquez, Trono, and Doty, 1975:161.—Cordero, 1977a:227, figs. 255–257; 1978a:51.—Chan, 1981:387.—Meñez and Calumpong, 1981:382.—Saraya and Trono, 1982:51, pl. XI: fig. 3.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Pangasinan. CENTRAL VISAYAS. MINDANAO: Davao. PALAWAN (Balabac I.). SULU: Tawitawi (Turtle Is.).

NOTE.—Grunow published this species as Polysiphonia (tongatensis var.?) upolensis, which was his way of indicating some doubt that the new entity should be recognized at specific rank. Nonetheless, the binomial should be attributed directly to Grunow rather than to (Grunow) Hollenberg (1968a:94).

Tolypiocladia Schmitz

Tolypiocladia calodictyon (Harvey ex Kützing) P.C. Silva

Polysiphonia calodictyon Harvey ex Kützing, 1864:16, pl. 46: figs, a–c [type locality: Tonga].

Tolypiocladia calodictyon (Harvey ex Kützing) P.C. Silva, 1952a:308.—Trono, 1972a:106; 1973a:137, fig. 17.—Trono and Ganzon-Fortes, 1980:107, fig. [s.n.].—Fortes, 1981b:396.—Saraya and Trono, 1982:54, pl. XI: fig. 4.—Trono and Ang, 1982:23.

Roschera africana Sonder, 1879:81, pl. I: figs. 5–11 [including Philippine record] [type locality: Zanzibar, Tanzania].

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas, Sorsogon. PALAWAN (Bugsuk I.), SULU: Tawitawi.

NOTE.—Roschera africana is included as a synonym on the authority of Weber-van Bosse (1923:359).

*Tolypiocladia condensata (Weber-van Bosse) P.C. Silva

Roschera condensata Weber-van Bosse, 1913a: pl. V: fig. 3; 1923:359 [syntype localities: various in Indonesia; Sangasiapu I., Tawitawi Prov., Sulu Archipelago].—Velasquez, Trono, and Doty, 1975:162.

Tolypiocladia condensata (Weber-van Bosse) P.C. Silva, 1952a:308.—Domantay, 1962:294.—Velasquez, Trono, and Doty, 1975:165.—Trono and Ang, 1982:24.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. PALAWAN (Bugsuk I.), SULU: Tawitawi (Sangasiapu I.).

Tolypiocladia glomerulata (C. Agardh) Schmitz

Hutchinsia glomerulata C. Agardh, 1824:158 [type locality: Shark Bay, Western Australia, Australia].

Tolypiocladia glomerulata (C. Agardh) Schmitz in Schmitz and Hauptfleisch, 1897 [1896–1897]:442.—De Toni, 1903:964.—Cornejo and Velasquez, 1972:182, pl. 3: fig. 24.—Trono, 1973a:137; 1973d:21; 1974b:95.—Velasquez, Trono, and Doty, 1975:165.—Cordero, 1977a:229.—Vannajan and Trono, 1978:26, fig. 39.—Liao and Sotto, 1980:100—Reyes, 1980:140, pl. 13, fig. 2a–c.—Chan, 1981:387.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:20, pl. XII: fig. 3.—Saraya and Trono, 1982:54.—Trono and Ang, 1982:23.—Meñez, Phillips, and Calumpong, 1983:18.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Cavite, Batangas. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SAMAR: Eastern Samar. NEGROS: Negros Oriental, CEBU (Mactan I.), SIQUIJOR. MINDANAO: Zamboanga, Davao. PALAWAN (Bugsuk I.), SULU: Sulu (Siasi I.).

Vidalia Lamouroux ex J. Agardh

Vidalia obtusiloba (C. Agardh) J. Agardh

Rytiphlaea obtusiloba C. Agardh, 1824:161 [type locality: Brazil].

Vidalia obtusiloba (C. Agardh) J. Agardh, 1863 [1852–1863]:1123.—Cordero, 1976c:8, 10; 1977a:230, fig. 260; 1978a:54.

PHILIPPINE DISTRIBUTION.—BATANES.

FAMILIA INCERTAE SEDIS

Family WURDEMANNIACEAE

Wurdemannia Harvey

Wurdemannia miniata (Sprengel) Feldmann and Hamel

Fucus miniatus Draparnaud ex De Candolle, 1815:6 [type locality: Montpellier, Hérault, France].

Sphaerococcus miniatus Sprengel, 1827:340.

Wurdemannia miniata (Sprengel) Feldmann and Hamel, 1934:544.—Domantay, 1962:294.—Cordero, 1976c:8; 1977a:77; 1978a:24.—Meñez and Calumpong, 1981:381.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. CENTRAL VISAYAS.

NOTE.—Fucus miniatus Draparnaud ex De Candolle, the intended basionym of Wurdemannia miniata, is a later homonym of F. miniatus O.F. Müller (1778:7, pl. 769) and hence not priorable. Sphaerococcus miniatus Sprengel is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Class PHAEOPHYCEAE

Order ECTOCARPALES

Family ECTOCARPACEAE

Feldmannia Hamel

Feldmannia columellaris (Børgesen) Islam

Ectocarpus columellaris Børgesen, 1936:71, fig. 4 [type locality: Galle, Sri Lanka].—Saraya and Trono, 1980:26.

Feldmannia columellaris (Børgesen) Islam, 1976:32.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Feldmannia formosana (Yamada) Itono

Ectocarpus formosanus Yamada, 1950:182, fig. 3 [type locality: Ryukyusho (Lambai I.), Taiwan].

Feldmannia formosana (Yamada) Itono, 1973:162.—Cordero, 1981d:64, fig. 5 [“formosanus”].

PHILIPPINE DISTRIBUTION.—PANAY: Aklan.

Feldmannia indica (Sonder) Womersley and Bailey

Ectocarpus indicus Sonder in Zollinger, 1854:2, 3, footnote [type locality: Bima Bay, Sumbawa I., Indonesia].—Agor, 1962:34.—Cornejo and Velasquez, 1972:175, pl. I: figs. 11, 11a.—Trono, 1972a:99.—Trono and Santos-Maranan, 1974a:2, fig. 1.—Velasquez, Trono, and Doty, 1975:141.—Vannajan and Trono, 1978:8, fig. 1.—Liao and Sotto, 1980:97.—Reyes, 1980:119, pl. 1 figs. 1a,b.—Chan, 1981:387,389.—Fortes, 1981b:396.

Feldmannia indica (Sonder) Womersley and Bailey, 1970:288.

Ectocarpus duchassaingianus Grunow, 1867:45, footnote, pl. IV: fig. 1 [type locality: Guadeloupe].

Giffordia duchassaingiana (Grunow) W.R. Taylor, 1960:207, pl. 29: fig. 10 [“duchassaigniana”].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite, Batangas. MINDORO: Oriental Mindoro. CEBU (Mactan I.). SIQUIJOR. SULU: Tawitawi.

NOTE.—The synonymy was proposed by Womersley and Bailey (1970:288).

Feldmannia irregularis (Kützing) Hamel

Ectocarpus irregularis Kützing, 1845:234 [type locality: Adriatic Sea].—Cornejo and Velasquez, 1972:175, pl. 1: fig. 10.—Trono and Santos-Maranan, 1974a:2, fig. 2.—Velasquez, Trono, and Doty, 1975:141.

Feldmannia irregularis (Kützing) Hamel, 1939:XVII.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. MINDORO: Oriental Mindoro.

Hincksia J.E. Gray

The reason for adopting Hincksia for the genus currently known as Giffordia is given in the appended Nomenclatural Notes.

Hincksia breviarticulata (J. Agardh) P.C. Silva, new combination

Ectocarpus breviarticulatus J. Agardh, 1847:7 [type locality: “St. Augustin” (Oaxaca, Mexico)].—Reyes, 1980:119, pl. 1: figs. 2a,b.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

NOTE.—This species was assigned to Feldmannia by Phm-Hoàng H (1969:298). As noted by Womersley and Bailey (1970:288), however, it does not have the long unbranched filaments with basal meristems that are characteristic of that genus. It seems best placed in Hincksia.

Hincksia mitchelliae (Harvey) P.C. Silva, new combination

Ectocarpus mitchelliae Harvey, 1852:142, pl. XII. [“mitchellae”] [type locality: Nantucket, Massachusetts, USA].—Trono and Santos-Maranan, 1974a:3, fig. 3.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Hincksia rallsiae (Vickers) P.C. Silva, new combination

Ectocarpus rallsiae Vickers, 1905:59 [type locality: Barbados].

Giffordia rallsiae (Vickers) W.R. Taylor, 1960:208.—Meñez and Calumpong, 1981:382.—Meñez, Phillips, and Calumpong, 1983:23.—Meñez and Calumpong, 1984:105.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS. PALAWAN.

Family RALFSIACEAE

Ralfsia Berkeley

Ralfsia fungiformis (Gunnerus) Setchell and Gardner

Fucus fungiformis Gunnerus, 1772:107 [type locality: Iceland].

Ralfsia fungiformis (Gunnerus) Setchell and Gardner, 1924b:11.—Agor, 1962:34.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Order CUTLERIALES

Family CUTLERIACEAE

Cutleria Greville

Cutleria cylindrica Okamura

Cutleria cylindrica Okamura, 1902 [1900-1902]:85, pl. XXVIII [type locality: various, all in Japan].—Agor, 1962:34.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Order SPHACELARIALES

Family SPHACELARIACEAE

Sphacelaria Lyngbye

Sphacelaria mucifera

Sphacelaria mucifera.—Velasquez, 1980:127.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

NOTE.—Velasquez did not give an author for this name and we have been unable to find its place of publication.

Sphacelaria novae-hollandiae Sonder

Sphacelaria novae-hollandiae Sonder, 1845:50 [type locality: Western Australia, Australia].—Liao and Sotto, 1980:97.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Sphacelaria rigidula Kützing

Sphacelaria rigidula Kützing, 1843b:292 [type locality: Red Sea].—Martens, 1868:23, 68-69.—Velasquez, Trono, and Doty, 1975:163.

Sphacelaria furcigera Kützing, 1855:27, pl. 90: fig. II [type locality: Karak (Khark) I., Iran].—Meñez, 1961:59, pl. 9: figs. 103, 104; pl. 10: figs. 105–108; pl. 12: figs. 126, 127.—Cornejo and Velasquez, 1972:175, 178, pl. 1: fig. 9; pl. 4: fig. 31.—Velasquez, Trono, and Doty, 1975:163.—Trono, 1976:214.—Liao and Sotto, 1980:97.—Chan, 1981:387.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:382.—Meñez, Phillips, and Calumpong, 1983:23.—Meñez and Calumpong, 1984:105.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas, Sorsogon. MINDORO. CEBU (Mactan I.). PALAWAN.

NOTE.—The synonymy was proposed by Prud’homme van Reine (1982:203).

Sphacelaria tribuloides Meneghini

Sphacelaria tribuloides Meneghini, 1840:[2] [type locality: Golfo di Spezia, Italy].—Meñez, 1961:59, pl. II: figs. 122–125.—Velasquez, Trono, and Doty, 1975:163.—Cordero, 1976c:9, 15, figs, E-I.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:26.—Chan, 1981:387.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:382.

Sphacelaria rigida Hering in Krauss, 1846:213 [type locality: “Natalbai” (Durban), South Africa].—Trono, 1978:9.—Liao and Sotto, 1980:97.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. MARINDUQUE. CEBU (Mactan I.).

NOTE.—The synonymy was proposed by Prud’homme van Reine (1982:179).

Order DICTYOTALES

Family DICTYOTACEAE

Dictyopteris Lamouroux

*Dictyopteris camiguinensis Tanaka

Dictyopteris camiguinensis Tanaka, 1967:16, fig. 5, pl. 1:A [type locality: San Pio Quinto, Camiguin I., Babuyan Is., Cagayan Prov., Luzon].—Velasquez, Trono, and Doty, 1975:140.

PHILIPPINE DISTRIBUTION.—As above.

Dictyopteris delicatula Lamouroux

Dictyopteris delicatula Lamouroux, 1809b:332, pl. 6: fig. 2B [type locality: Antilles].—Weber-van Bosse, 1913a:181.—Velasquez, Trono, and Doty, 1975:140.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS. SULU: Tawitawi (Pearl Bank).

Dictyopteris divaricata (Okamura) Okamura

Haliseris divaricata Okamura, 1907 [1907–1909]:57, pl. XIII; pl. XIV: fig. 5 [syntype localities: Cape Iwai, Miyage Prefecture and Hakodate, Hokkaido, Japan].

Dictyopteris divaricata (Okamura) Okamura, 1932:75.—Cordero, 1980c:71, fig. 2.—Fortes, 1981b:396.—Cordero, 1984a:83.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Dictyopteris jamaicensis W.R. Taylor

Dictyopteris jamaicensis W.R. Taylor, 1960:631, pl. 32: fig. 2 [type locality: St. Catharine Parish, Jamaica].—Vannajan and Trono, 1978:9, fig. 3.—Saraya and Trono, 1980:27, pl. VII: figs. 1, 3.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite. CENTRAL VISAYAS.

Dictyopteris polypodioides (De Candolle) Lamouroux

Fucus membranaceus Stackhouse, 1795 [1795–1801]:13, pl. 6 [type locality: Sidmouth, Devonshire, England].

Polypodoidea membranacea Stackhouse, 1809:97.

Dictyopteris membranacea (Stackhouse) Batters, 1902:54.—Cordero, 1984b:60.

Fucus polypodioides Desfontaines, 1799 [1798–1799]:421 [type locality: “mare Numidico” (Algeria)].

Ulva polypodioides De Candolle in Lamarck and De Candolle, 1805:15.

Dictyopteris polypodioides (De Candolle) Lamouroux, 1809b:332.

PHILIPPINE DISTRIBUTION.—PALAWAN.

NOTE.—The conspecificity of Fucus membranaceus Stackhouse and Fucus polypodioides Desfontaines was proposed by Turner (1809:41). Both names are later homonyms, however, of F. membranaceus N.L. Burman (1768:32 [28 in error]) and F. polypodioides S.G. Gmelin (1768:186), respectively. A legitimate basionym was created for F. membranaceus by Stackhouse (1809) in transferring it into a new genus, Polypodoidea, but in the meantime a legitimate basionym had been created for F. polypodioides by De Candolle (in Lamarck and De Candolle, 1805), who transferred it into Ulva. Both Ulva polypodioides and Polypodoidea membranacea are treated as legitimate new names in accordance with Article 72, Example 2, of the ICBN.

Dictyopteris repens (Okamura) Børgesen

Haliseris repens Okamura, 1916:8, fig. 3, pl. I: figs. 7–18 [type locality: Truk Is., Caroline Is.].

Dictyopteris repens (Okamura) Børgesen, 1924:265.—Trono, 1973a:129, fig. 3; 1974a:146.—Cordero, 1976c:9, 10, 16, figs. L,M.—Trono, 1978:9—Saraya and Trono, 1980:26, pl. VII: fig. 4.—Marcos-Agngarayngay, 1984a:5, fig.4.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Sorsogon. MARINDUQUE. SULU: Sulu (Siasi I.).

Dictyopteris undulata Holmes

Dictyopteris undulata Holmes, 1896:251, pl. VIII: fig. 1 [type locality: Misaki, Kanagawa Prefecture, Japan].—Flores-Sian, 1959:97.—Domantay, 1962:282.—Manapat, 1969:37.—Velasquez, Trono, and Doty, 1975:140.—Cordero, 1976c:18.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan.

Dictyota Lamouroux

Dictyota bartayresiana Lamouroux

Dictyota bartayresiana Lamouroux, 1809a:43 [type locality: Antilles].

Dictyota bartayresii Lamouroux, 1809b:331.—Taylor, 1966b:354.—Westernhagen, 1973a:65.—Trono, 1974a:143.—Westernhagen, 1974:113 (table 1).—Trono, 1976:216.—Garcia, 1979:44 (table 1).—Liao and Sotto, 1980:97.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon. MINDORO: Oriental Mindoro. CEBU (Mactan I.). MINDANAO. PALAWAN (Balabac I.). SULU: Sulu (Siasi I.), Tawitawi (Turtle Is.).

NOTE.—Lamouroux’s change of epithet from bartayresiana to bartayresii was unwarranted. Collins and Hervey (1917:90, footnote) erred in believing that Lamouroux’s paper in Desvaux’s Journal de Botanique appeared after his paper in the Nouveau Bulletin des Sciences. The respective dates of publication were April 1809 (see Stafleu, 1967:105) and May 1809, respectively.

Dictyota bartayresiana Lamouroux var. denticulata Kützing

Dictyota bartayresiana Lamouroux var. denticulata Kützing, 1859:8, pl. 16: fig. II [type locality: Ilhas do Cabo Verde].—Weber-van Bosse, 1913a:183, pl. III: fig. 4.—Velasquez, Trono, and Doty, 1975:141 [without designation of variety].

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Jolo I.).

NOTE.—Dictyota bartayresiana var. denticulata, which Kützing illustrated using material from the Cape Verde Is. but did not describe, was validated by the citation of D. crenulata J. Agardh (1847:7, type locality: “St. Augustin” [Oaxaca, Mexico]) as a synonym. J. Agardh (1882:99), however, doubted that Kützing’s plant was conspecific with D. crenulata. Moreover, D. crenulata is recognized as a species of Dilophus by Nizamuddin and Gerloff (1980:867), so that the identity of Weber-van Bosse’s material is very much in doubt.

*Dictyota bidentata Harvey and Bailey

Dictyota bidentata Harvey and Bailey, 1851:373 [type locality: Mindanao].—Bailey and Harvey, 1862:173, pl. VIII: figs. 6, 7.—Velasquez, Trono, and Doty, 1975:141.

PHILIPPINE DISTRIBUTION.—As above.

Dictyota cervicornis Kützing

Dictyota cervicornis Kützing, 1859:11, pl. 24: fig. 11 [type locality: Key West, Florida, USA].—Meñez, 1961:63, pl. 12: fig. 133.—Taylor, 1966b:354.—Manapat, 1969:38.—Reyes, 1972:145.—Cordero, 1973b:23.—Velasquez et al., 1973:19, pl. 6: fig. 30.—Ortega, Alcala, and Reyes, 1974:187, 188.—Velasquez, Trono, and Doty, 1975:141.—Puig and Cordero, 1979:30.—Reyes, 1980:119, pl. 1: fig. 4.—Saraya and Trono, 1980:27, pl. VIII: fig. 2.—Chan, 1981:387.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Bataan, Batangas, Quezon, LEYTE (Biliran I.), NEGROS: Negros Oriental, SIQUIJOR. PALAWAN (Balabac I.).

Dictyota ceylanica Kützing

Dictyota ceylanica Kützing, 1859:11, pl. 25: fig. 1 [type locality: Sri Lanka].—Martens, 1868:26, 80–81.—Velasquez, Trono, and Doty, 1975:141.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Dictyota ciliolata Kützing

Dictyota ciliolata Kützing, 1859:12, pl. 27: fig. I [type locality: La Guaira, Venezuela].—Garcia, 1979:44 (table 1).—Saraya and Trono, 1980:29, pl. VIII: fig. 3

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDANAO.

Dictyota dichotoma (Hudson) Lamouroux

Ulva dichotoma Hudson, 1762:476 [type locality: Walney I., Lancashire, England].

Dictyota dichotoma (Hudson) Lamouroux, 1809a:42.—Dickie, 1876a:243.—De Toni, 1895b:263.—Howe, 1932:170.—Meñez, 1961:62, pl. 12: fig. 136.—Agor, 1962:34.—Domantay, 1962:283.—Taylor, 1966b:355.—Manapat, 1969:37, 38.—Velasquez, 1971:441, fig. 19. —Reyes, 1972:145.—Cordero, 1973b:22.—Trono, 1973d:11, pl. 8: fig. 30.—Velasquez et al., 1973:18, pl. 6: fig. 29—Ortega, Alcala, and Reyes, 1974:186, 187.—Trono, 1974a:142.—Velasquez, Trono, and Doty, 1975:141.—Trono and Young, 1977:58.—Trono and Tuason, 1978:11.—Vannajan and Trono, 1978:9, fig. 4.—Cordero, 1979b:276, 284.—Puig and Cordero, 1979:30.—Liao and Sotto, 1980:97.—Reyes, 1980:119, pl. 1: fig. 3.—Saraya and Trono, 1980:28, pl. VIII: fig. 1.—Meñez and Calumpong, 1981:382.—Hurtado-Ponce, 1983:120.—Cordero, 1984a:82; 1984b:61; 1984c:52.—Marcos-Agngarayngay, 1984a:3, fig. 2.

Dictyota volubilis Kützing, 1849:554 [type locality: Arromanches-les-Bains, Calvados, France].—Westernhagen, 1973a:66; 1974:113 (table I).

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Bataan, Cavite, Batangas, Quezon, Albay. CATANDUANES. MINDORO: Oriental Mindoro. MASBATE. LEYTE (Biliran I.), PANAY: Aklan. NEGROS: Negros Occidental, Negros Oriental, CEBU (Mactan I.), SIQUIJOR. MINDANAO: Zamboanga. PALAWAN (incl. Balabac I.), SULU: Sulu (Siasi I.), Tawitawi.

NOTE.—The synonymy was proposed by J. Agardh (1882:92).

Dictyota divaricata Lamouroux

Dictyota divaricata Lamouroux, 1809a:43 [type locality: Mediterranean France].—Domantay, 1962:283.—Manapat, 1969:37, 38—Trono, 1972a:100.—Velasquez et al., 1973:19, pl. 7: fig. 31.—Westernhagen, 1973a:66.—Trono, 1974a:142.—Westernhagen, 1974:113 (table I).—Velasquez, Trono, and Doty, 1975:141.—Trono, 1978:10.—Cordero, 1979b:276.—Fortes and Trono, 1980:66.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:28, pl. VII: fig. 2.—Fortes, 1981b:396.—Trono and De Lara, 1981:8: pl. V: figs. 1, 2.—Trono and Ang, 1982:13.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas, Quezon, MINDORO: Occidental Mindoro (Lubang Is.). MARINDUQUE. PANAY: Aklan. CEBU (Mactan I.). PALAWAN (Bugsuk I.). SULU: Sulu (Siasi I.), Tawitawi.

Dictyota friabilis Setchell

Dictyota friabilis Setchell, 1926:91, pl. 13: figs. 4–7 [type locality: Tafaa Point, Tahiti]; pl. 20: fig. 1.—Trono, 1974a:144; 1976:216.—Trono and Young, 1977:58.—Trono and Tuason, 1978:12.—Saraya and Trono, 1980:28.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Sorsogon. CATANDUANES. SULU: Sulu (Siasi I.).

Dictyota indica Sonder ex Kützing

Dictyota indica Sonder ex Kützing, 1859:8, pl. 17: fig. 1 [type locality: La Habana, Cuba].—Meñez, 1961:62.—Velasquez, Trono, and Doty, 1975:141.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Dictyota lata Lamouroux

Dictyota lata Lamouroux, 1809a:43 [type locality: “Ind. orient.” (Malagasy Republic)].—Martens, 1868:80–81.—Dickie, 1876a:244.—De Toni, 1895b:281—Velasquez, Trono, and Doty, 1975:141.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Dictyota linearis (C. Agardh) Greville

Zonaria linearis C. Agardh, 1820b:134 [syntype localities: Cádiz, Spain; “ad oras Americae”].

Dictyota linearis (C. Agardh) Greville, 1830:xliii.—Montagne, 1844a 660.—Martens, 1868:80–81.—Domantay, 1962:283.—Taylor, 1966b:355.—Cordero, 1973b:23.—Velasquez, Trono, and Doty, 1975 141._Cordero, 1976c:10, 16, figs. J,K; 1979b:276, 285.—Puig and Cordero, 1979:30.—Saraya and Trono, 1980:29, pl. VIII: fig. 4.—Meñez and Calumpong, 1981:382.—Cordero, 1984a:83; 1984b:61.—Marcos-Agngarayngay, 1984a:5, fig. 3.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan. LEYTE (Biliran I.), PANAY: Aklan. CENTRAL VISAYAS. PALAWAN, SULU: Tawitawi (Turtle Is.).

Dictyota major W.R. Taylor

Dictyota major W.R. Taylor, 1945:88, pls. 8, 9 [type locality: Isla Santa María, Galápagos].—Saraya and Trono, 1980:29, pl. VIII: fig. 4.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Dictyota mertensii (Martius) Kützing

Ulva mertensii Martius, 1828:5, pl. 1 [type locality: Salvador, Brazil].

Dictyota mertensii (Martius) Kützing, 1859:15.

Dictyota dentata [misapplied name].—Vannajan and Trono, 1978:9, fig. 5.—Saraya and Trono, 1980:27.—Trono and Ganzon-Fortes, 1980:39, fig. [s.n.].

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Cavite, Batangas.

NOTE.—Dictyota dentata Lamouroux (1809a:42), the name usually applied to this species, is illegitimate because Lamouroux cited Fucus atomarius S.G. Gmelin (1768:125, pl. X: fig. 1) as a synonym and therefore should have retained the epithet atomaria. This epithet cannot be reinstated now, however, because two other species have subsequently been given the name Dictyota atomaria. Moreover, Lamouroux erred in his taxonomic opinion, since Fucus atomarius is representative of the red algal genus Odonthalia (see Papenfuss, 1950:189). The conspecificity of D. dentate sensu J. Agardh and D. mertensii was proposed by Hauck (1888:466).

Dictyota patens J. Agardh

Dictyota patens J. Agardh, 1882:93 [syntype localities: St. Thomas, Virgin Is.; Tonga].—Trono, 1974a:143.—Puig and Cordero, 1979:31.

PHILIPPINE DISTRIBUTION.—LEYTE (Biliran I.), SULU: Sulu (Siasi 1.).

Dilophus J. Agardh

Dilophus okamurae Dawson

Dictyota marginata Okamura, 1913 [1913–1915]:33, pl. CVIII: fig. 9; pl. Cl [syntype localities: various, all in Japan].—Cordero, 1973b:24 [D. marginata prox.].—Puig and Cordero, 1979:31.

Dilophus marginatus (Okamura) Okamura, 1915 [1913–1915]:154 [replaced name].

Dilophus okamurae Dawson, 1950a:86 [“okamurai”].

PHILIPPINE DISTRIBUTION.—LEYTE (Biliran I.).

NOTE.—Dilophus okamurae is a substitute name for D. marginatus (Okamura) Okamura, a later homonym of D. marginatus J. Agardh (1894:91).

Lobophora J. Agardh

Lobophora variegata (Lamouroux) Womersley

Dictyota variegata Lamouroux, 1809a:40 [type locality: Antilles].

Zonaria variegata (Lamouroux) C. Agardh, 1817:xx.—Dickie, 1876a:245.—Weber-van Bosse, 1913a:175.—Meñez, 1961:61, pl. 12: fig. 135.—Velasquez, Trono, and Doty, 1975:169.—Liao and Sotto, 1980:97.

Gymnosorus variegatus (Lamouroux) J. Agardh, 1894:11.—De Toni, 1895b:227.

Pocockiella variegata (Lamouroux) Papenfuss, 1943b:467.—Domantay, 1962:284.—Taylor, 1966b:357.—Trono, 1972a:100; 1973d:11.—Velasquez et al., 1973:19, pl. 7: fig. 32.—Trono and Santos-Maranan, 1974b:3, figs. 2, 3.—Velasquez, Trono, and Doty, 1975:160.—Reyes, 1980:120, pl. 2: fig. 3.—Cordero, 1984b:62.

Lobophora variegata (Lamouroux) Womersley, 1967:221.—Trono, 1978:10.—Vannajan and Trono, 1978:13, figs. 10, 13.—Fortes and Trono, 1980:60, 66.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:31, pl. IX: fig. 1.—Chan, 1981:387.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Bataan, Cavite, Batangas. MINDORO: Oriental Mindoro. MARINDUQUE. PANAY: Iloilo (Gigantes Is.), CEBU (Mactan I.), SIQUIJOR. PALAWAN (Balabac I.), SULU: Sulu (North Ubian I.), Tawitawi.

NOTE.—Papenfuss (1980) has proposed the generic name Pocockiella Papenfuss (1943b:467) for conservation over the earlier taxonomic synonym Lobophora J. Agardh (1894:21). The latter name has come into wide use since its identity was clarified by Womersley (1967:221), however, so there seems to be little or no merit in conserving Pocockiella.

Padina Adanson

Padina arborescens Holmes

Padina arborescens Holmes, 1896:251, pl. 12: fig. 1 [type locality: Enoshima, Kanagawa Prefecture, Japan].—Agor, 1962:34.—Manapat, 1969:38.—Garcia, 1979:44 (table 1).

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDANAO. PALAWAN.

Padina australis Hauck

Padina australis Hauck, 1887 [1886–1887]:44 [type locality: Cape York, Queensland, Australia].—Merrill, 1918:40.—Howe, 1932:170.—Zaneveld, 1952:131.—Abelardo and De Leon, 1956:102.—Olea and De Leon, 1956:104.—Zaneveld, 1956:17; 1959:100.—Domantay, 1962:283.—Taylor, 1966b:355.—Manapat, 1969:37, 38.—Reyes, 1972:145.—Trono, 1973d:11, pl. 8: fig. 29.—Ortega, Alcala, and Reyes, 1974:185, 186, 187, 188.—Velasquez, Trono, and Doty, 1975:158.—Trono, 1976:215; 1978:10.—Reyes, 1980:120, pl. 1: fig. 5.—Saraya and Trono, 1980:29, pl. IX: fig. 3.—Trono and Fortes, 1980:66.—Trono and Ganzon-Fortes, 1980:41, fig. [s.n.].—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:8, pl. V: fig. 4.—Laserna et al., 1982:52.—Trono and Ang, 1982:13.—Trono and Fortes, 1982:146.

Ulva umbilicalis [misapplied name fide Merrill, 1918:40].—M. Blanco, 1837:842; 1845:581; 1879:261.—Velasquez, Trono, and Doty, 1975:168.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, La Union, Pangasinan, Bataan, Batangas, Quezon, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. SAMAR: Eastern Samar. LEYTE. PANAY. NEGROS: Negros Occidental, Negros Oriental. SIQUIJOR. MINDANAO: Misamis Occidental. PALAWAN (incl. Balabac I., Bugsuk I.).

NOTE.—Martens (1868:47) identified Blanco’s plant as Zonaria gymnospora (= Padina gymnospora). In the absence of authentic specimens, no definitive placement can be made.

Padina boryana Thivy

Padina boryana Thivy in Taylor, 1966b:355, fig. 2 [including Philippine records] [type locality: Tonga].—Kraft, 1972:332.—Velasquez, Trono, and Doty, 1975:158.

Padina commersonii [misapplied name].—Domantay, 1962:284.—Velasquez, Trono, and Doty, 1975:158.—Cordero, 1976c:6, 19.—Puig and Cordero, 1979:32.—Trono and Fortes, 1980:66.—Ganzon-Fortes, 1981:21 —Trono and Fortes, 1982:146.

Padina tenuis [misapplied name].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Pangasinan, Quezon (incl. Polillo Is.), CATANDUANES. LEYTE (Biliran I), PANAY: Antique, CENTRAL VISAYAS. PALAWAN (Balabac I.), SULU: Sulu.

NOTE.—Padina commersonii Bory de Saint-Vincent (1828 [1826–1829]:144), a name that traditionally has been applied to the present species, is an illegitimate substitute for P. tenuis (C. Agardh) Bory de Saint-Vincent (1827:590), whose basionym, Zonaria pavonia var. tenuis C. Agardh (1824:264), has been shown by Papenfuss (1977:277) to be referable to Lobophora variegata. The material that Bory de Saint-Vincent had in hand was incorporated in a new species, P. boryana, by Thivy.

Padina crassa Yamada

Padina crassa Yamada, 1931a:67, pl. XVII: fig. 2 [syntype localities: various, all in Japan].—Manapat, 1969:37, 38.—Velasquez, 1971:441, fig. 20.—Cordero, 1973b:24.—Velasquez et al., 1973:19, pl. 7: fig. 33.—Cordero, 1979b:276, 284.—Puig and Cordero, 1979:31.—Hurtado-Ponce, 1983:120.—Cordero, 1984a:83; 1984b:61; 1984c:52.—Marcos-Agngarayngay, 1984a:3, fig. 1.—Tungpalan, 1984:142.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Bataan, Batangas, Quezon, MASBATE. LEYTE (Biliran I.). PANAY: Aklan. CEBU. PALAWAN.

Padina distromatica Hauck

Padina distromatica Hauck, 1887 [1886–1887]:43 [type locality: Meith (Maidh), Somalia].—Howe, 1932:169.—Taylor, 1966b:356.—Velasquez, Trono, and Doty, 1975:158.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro. PANAY. BOHOL.

Padina fraseri (Greville) Greville

Zonaria fraseri Greville, 1829:423, pl. XXVI: fig. 2 [type locality: “ad Novam Hollandiam” (probably near Fremantle, Western Australia, Australia fide Womersley, 1967:222)].

Padina fraseri (Greville) Greville, 1830:xliv.—Taylor, 1966b:356.—Velasquez, Trono, and Doty, 1975:159.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Sur, La Union.

Padina gymnospora (Kützing) Sonder

Zonaria gymnospora Kützing, 1859:29, pl. 71: fig. 11 [type locality: St. Thomas. Virgin Is.].—Martens, 1868:26, 47, 82–83.

Padina gymnospora (Kützing) Sonder, 1871:47.—Taylor, 1966b:356.—Velasquez, Trono, and Doty, 1975:159.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon, CENTRAL VISAYAS. MINDANAO: Zamboanga.

Padina japonica Yamada

Padina japonica Yamada, 1931a:69, pl. XIX: fig. 2 [syntype localities: various, all in Japan].—Domantay, 1962:284.—Taylor, 1966b:356.—Manapat, 1969:37.—Cornejo and Velasquez, 1972:174, 179, 180, 181.—Kraft, 1972:332.—Trono, 1972a:100.—Velasquez et al., 1973:20, pl. 7: fig. 34.—Velasquez, Trono, and Doty, 1975:159.—Trono, 1976:215.—Trono and Young, 1977:58.—Trono, 1978:11.—Cordero, 1979b:276.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:30, pl. IX: fig. 2.—Trono and Fortes, 1980:66; 1982:146.—Hurtado-Ponce, 1983:121.—Cordero, 1984c:52.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Bataan, Batangas, Quezon, Camarines Sur, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), MARINDUQUE. MASBATE. PANAY: Aklan. CEBU (Mactan I.), SULU: Tawitawi.

Padina minor Yamada

Padina minor Yamada, 1925b:251, fig. V [type locality: Garan-Bi (O-luan), Taiwan].—Manapat, 1969:37, 38.—Cornejo and Velasquez, 1972:175, 176, 182.—Trono, 1973d:12, pl. 6: fig. 21.—Velasquez et al., 1973:20, pl. 7: fig. 35.—Trono, 1974a:141; 1976:214; 1978:11.—Trono and Tuason, 1978:12.—Puig and Cordero, 1979:31.—Liao and Sotto, 1980:97.—Reyes, 1980:120, pl. 2: fig. 1.—Saraya and Trono, 1980:30.—Trono and Fortes, 1980:66.—Trono and Ganzon-Fortes, 1980:43, fig. [s.n.].—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:8, pl. V: fig. 3.—Trono and Fortes, 1982:146.—Hurtado-Ponce, 1983:121.—Cordero, 1984c:52.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Pangasinan, Bataan, Batangas, Quezon, Sorsogon. CATANDUANES. MARINDUQUE. MASBATE. LEYTE (Biliran I.), CEBU (Mactan I.), SIQUIJOR. SULU: Sulu (Siasi I.).

Padina pavonica (Linnaeus) Thivy

Fucus pavonicus Linnaeus, 1753:1162 [type locality: “In Mari Europae australis”].

Fucus pavonius Linnaeus, 1759:1345.

Padina pavonia (Linnaeus) Lamouroux, 1816:304.—Bailey and Harvey, 1862:173.—Dickie, 1876a:243.—Meñez, 1961:61, pl. 10: figs. 113–117; pl. II: figs. 118–120.—Velasquez, Trono, and Doty, 1975:159.

Padina pavonica (Linnaeus) Thivy in Taylor, 1960:234.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CEBU (Mactan I.). MINDANAO: Zamboanga.

NOTE.—We consider Fucus pavonius Linnaeus, the basionym of Padina pavonia, an illegitimate substitute for F. pavonicus. Some authors, however, consider the two epithets orthographic variants and hence accredit the binomial P. pavonica to (Linnaeus) Lamouroux (see Price, Tittley, and Richardson, 1979:3, footnote).

Padina sanctae-crucis Børgesen

Padina sanctae-crucis Børgesen, 1914b:45, figs. 27, 28 [type locality: St. Croix, Virgin Is.].—Meñez, 1961:62, pl. 10: figs. 109–112.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—According to Taylor (1960:237), Dictyerpa jamaicensis Collins (1901:251; type locality: Manchioneal, Jamaica) “is a growth stage of a Padina, perhaps of various species, but certainly of this one” (Padina sanctae-crucis). On the basis of this statement, Papenfuss (1977:272) made the combination P. jamaicensis (Collins) Papenfuss. We prefer to retain Børgesen’s name, however, because of the uncertainty as to which species is represented by the type material of D. jamaicensis.

Padina tetrastromatica Hauck

Padina tetrastromatica Hauck, 1887 [1886–1887]:43 [type locality: Meith (Maidh), Somalia].—Taylor, 1966b:356.—Velasquez, Trono, and Doty, 1975:159.—Vannajan and Trono, 1978:10, figs. 6–8.—Trono and Fortes, 1980:66.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:146.

PHILIPPINE DISTRIBUTION.—LUZON: Bataan, Rizal, Manila, Cavite.

Spatoglossum Kützing

Spatoglossum asperum J. Agardh

Spatoglossum asperum J. Agardh, 1894:36 [type locality: Sri Lanka].—Trono, 1974a:144.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Siasi I.).

Spatoglossum variabile Figari and De Notaris

Spatoglossum variabile Figari and De Notaris, 1853:153, fig. IV [syntype localities: Suez, Egypt and Aqaba, Jordan, Red Sea].—Weber-van Bosse, 1913a:181.—Velasquez, Trono, and Doty, 1975:163.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

Stypopodium (Kützing) J. Agardh

*Stypopodium flabelliforme Weber-van Bosse

Stypopodium flabelliforme Weber-van Bosse, 1913a:176, pl. II: fig. 10; pl. III: fig. 2 [syntype localities: Rotti I., Indonesia; Pearl Bank, Tawitawi Prov., Sulu Archipelago].—Domantay, 1962:287.—Velasquez, Trono, and Doty, 1975:164.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. SULU: Tawitawi (Pearl Bank).

Zonaria C. Agardh

Zonaria diesingiana J. Agardh

Zonaria diesingiana J. Agardh, 1841:443 [type locality: “Nov. Holl.” (probably near Sydney, Australia fide Allender and Kraft, 1983:81)].—Cordero, 1976c:6, 10, 18.

PHILIPPINE DISTRIBUTION.—BATANES.

NOTE.—Womersley (1967:228) rejected this name as a nomen dubium on the grounds that the type specimen (from Australia) represents the basal portion of an undeterminable Zonaria.

Zonaria flabellata (Okamura) Papenfuss

Homoeostrichus flabellatus Okamura, 1931 [1929–1932]:57, pl. CCLXXXIX: figs. 11–13 [syntype localities: Ryukyu-retto and Seto, Wakayama Prefecture, Japan].—Flores-Sian, 1959:97.

Zonaria fiabellata (Okamura) Papenfuss, 1944:341.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Order CHORDARIALES

Family CORYNOPHLAEACEAE

Leathesia S.F. Gray

Leathesia difformis (Linnaeus) J.E. Areschoug

Tremella difformis Linnaeus, 1755:429 [type locality: Sweden].

Leathesia difformis (Linnaeus) J.E. Areschoug, 1847:376.—Domantay, 1962:283.—Cordero, 1973b:24.—Velasquez, Trono, and Doty, 1975:151.—Marcos-Agngarayngay, 1984a:7, fig. 5.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan. LEYTE (Biliran I.).

Order DICTYOSIPHONALES

Family CHNOOSPORACEAE

Chnoospora J. Agardh

Chnoospora implexa J. Agardh

Chnoospora implexa J. Agardh, 1848:172 [type locality: Tor, Sinai Peninsula, Egypt].—Meñez, 1961:60, pl. 12: figs. 129–131.—Reyes, 1972:148.—Velasquez et al., 1973:21, pl. 8: fig. 38.—Trono and Santos-Maranan, 1974b:4, fig. 4.—Velasquez, Trono, and Doty, 1975:135.—Trono, 1976:217.—Liao and Sotto, 1980:98.—Reyes, 1980:122, pl. 3: fig. 4.—Meñez and Calumpong, 1981:382.—Cordero, 1982b:32.—Hurtado-Ponce, 1983:122.—Hurtado-Ponce, 1984:179.—Tungpalan, 1984:139.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Bataan, Sorsogon. MINDORO: Oriental Mindoro. SAMAR: Eastern Samar. NEGROS: Negros Oriental, CEBU (Mactan I.), SIQUIJOR.

Chnoospora minima (Hering) Papenfuss

Fucus minimus Hering, 1841:92 [type locality: “Port Natal” (Durban), South Africa].

Chnoospora minima (Hering) Papenfuss, 1956:69.—Taylor, 1966b:357. Cordero, 1982b:32, fig. 1.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. SAMAR: Eastern Samar.

Chnoospora pannosa J. Agardh

Chnoospora pannosa.J. Agardh, 1848:172 [type locality: Oahu, Hawaiian Is.].—Martens, 1868:26, 70–71.—Velasquez, Trono, and Doty, 1975:135.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Order SCYTOSIPHONALES

Family SCYTOSIPHONACEAE

Colpomenia (Endlicher) Derbès and Solier

Colpomenia sinuosa (Mertens ex Roth) Derbès and Solier

Ulva sinuosa Mertens ex Roth, 1806:327, pl. 12 [type locality: Cádiz, Spain].

Colpomenia sinuosa (Mertens ex Roth) Derbès and Solier in Castagne, 1851:95.—Velasquez, 1949:155 [“sinuate”].—Meñez, 1961:60, pl. 12: fig. 134.—Agor, 1962:34.—Domantay, 1962:282.—Galutira and Velasquez, 1964:500.—Velasquez, 1971:441, fig. 22.—Reyes, 1972:148.—Trono, 1973d:12.—Velasquez et al., 1973:20, pl. 7: fig. 36.—Westernhagen, 1973a:65.—Ortega, Alcala, and Reyes, 1974:186, 187.—Trono and Santos-Maranan, 1974b:4, figs. 5, 6.—Westernhagen, 1974:113 (table 1).—Velasquez, Trono, and Doty, 1975:138 [“sinuosa” and “sinuate”].—Cordero, 1979b:285.—Puig and Cordero, 1979:34.—Cordero, 1980b:34, pl. [50].—Reyes, 1980:121, pl. 3: fig. 1.—Saraya and Trono, 1980:31, pl. IX: fig. 4.—Chan, 1981:387.—Meñez and Calumpong, 1981:382.—Hurtado-Ponce, 1983:122.—Cordero, 1984a:84.—Hurtado-Ponce, 1984:179.—Marcos-Agngarayngay, 1984a:9, fig. 7.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Bataan, Batangas. MINDORO: Oriental Mindoro. LEYTE (Biliran I.), PANAY: Aklan. NEGROS: Negros Oriental. CEBU (Mactan I.), SIQUIJOR.

Hydroclathrus Bory

Hydroclathrus clathratus (C. Agardh) Howe

Encoelium clathratum C. Agardh, 1822a:412 [type locality: uncertain].—Martens, 1868:47, 70–71. —Velasquez, Trono, and Doty, 1975:141.

Hydroclathrus clathratus (C. Agardh) Howe, 1920:590.—Zaneveld, 1952:130; 1956:15; 1959:99.—Agor, 1962:34.—Domantay, 1962:283.—Galutira and Velasquez, 1964:500, pl. 2: fig. 7; pl. 7: fig. 26.—Taylor, 1966b:357.—Velasquez, 1968a:120, fig. 5.—Villones and Magdamo, 1968:26, fig. 18 [figure cited but not published].—Manapat, 1969:37, 38, 39.—Velasquez, 1971:441, fig. 21.—Reyes, 1972:148.—Velasquez, 1972:63.—Bersamin et al., 1973:187.—Trono, 1973d:12.—Velasquez et al., 1973:21, pl. 8: fig. 37.—Westernhagen, 1973a:66.—Trono, 1974a:147.—Trono and Santos-Maranan, 1974b:2, fig. 1.—Westernhagen, 1974:113 (table 1).—Velasquez, Trono, and Doty, 1975:150.—Trono, 1976:216.—Cordero, 1979b:276, 285.—Puig and Cordero, 1979:34.—Velasquez, 1979b:230.—Cordero, 1980b:33, pl. 21.—Liao and Sotto, 1980:97.—Moreland, 1980:43, 46, 47.—Reyes, 1980:121, pl. 3: figs. 2a,b.—Saraya and Trono, 1980:32.—Trono and Fortes, 1980:66.—Trono and Ganzon-Fortes, 1980:45, fig. [s.n.].—Trono, Velasquez, and Guevarra, 1980:77.—Ganzon-Fortes, 1981:21 .—Guzman, 1981:43, 49.—Meñez and Calumpong, 1981:382.—Laserna et al., 1981:447.—Trono and De Lara, 1981:9, pl. VI: fig. 4.—Calumpong, 1982:145.—Cordero, 1982a:60.—Trono and Ang, 1982:14.—Trono and Fortes, 1982:147.—Cordero, 1984a:85.—Hurtado-Ponce, 1983:122.—Cordero, 1984b:62; 1984c:53.—Hurtado-Ponce, 1984:179.—Marcos-Agngarayngay, 1984a:7, fig. 6; 1984b:122.—Tungpalan, 1984:139.—Ang, 1985b:298.

Hydroclathrus cancellatus Bory de Saint-Vincent, 1825:419.—Weber-van Bosse, 1913a:136.—Merrill, 1918:40.—Collado, 1926:129.—G. Blanco, 1938:513.—Velasquez, 1949:155.—Quisumbing, 1951:1010.—Velasquez, 1953b:206.—Montilla and Blanco, 1953:166, fig. 5:4.—Meñez, 1961:60.—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Cordero, 1973b:25.—Westernhagen, 1973a:66; 1974:113 (table 1).—Velasquez, Trono, and Doty, 1975:150.

Ulva reticulata [misapplied name fide Martens, 1868:47 and Collins in Merrill, 1918:40].—M. Blanco, 1837:842; 1845:582; 1879:262.—Martens, 1868:60-61 [Blanco record only].—Velasquez, Trono, and Doty, 1975:167 [Blanco and Martens records only].

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Ilocos Sur, La Union, Pangasinan, Bataan, Batangas, Quezon, Albay, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MASBATE. SAMAR: Eastern Samar. LEYTE (Biliran I.), PANAY: Aklan. NEGROS: Negros Oriental (incl. Apo I.), CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Zamboanga, Davao. PALAWAN (incl. Balabac I., Bugsuk I.), SULU: Sulu (Siasi I.), Tawitawi.

NOTE.—Encoelium clathratum C. Agardh (based on Fucus clathratus Bory de Saint-Vincent mscr.) and Hydroclathrus cancellatus Bory de Saint-Vincent share the same type collection. The provenance was stated to be Belle-Ile (Atlantic France), but Hamel (1937:202) explained how this was shown to be in error. The true provenance is unknown.

Rosenvingea Børgesen

Rosenvingea intricata (J. Agardh) Børgesen

Asperococcus intricatus J. Agardh, 1847:7 [type locality: Veracruz, Mexico].

Rosenvingea intricata (J. Agardh) Børgesen, 1914b:26.—Reyes, 1972:148.—Velasquez, Trono, and Doty, 1975:162.—Reyes, 1980:121, pl. 3: fig. 3.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental. SIQUIJOR.

*Rosenvingea orientalis (J. Agardh) Børgesen

Asperococcus orientalis J. Agardh, 1848:78 [type locality: Manila, Luzon].—De Toni, 1895b:495.

Encoelium orientate (J. Agardh) Kützing, 1849:551.—Martens, 1868:68–69.—Velasquez, Trono, and Doty, 1975:142.

Hydroclathrus orientalis (J. Agardh) Heydrich, 1894:286 [including Philippine record].—Velasquez, Trono, and Doty, 1975:150.

Rosenvingea orientalis (J. Agardh) Børgesen, 1914b:26.—Saraya and Trono, 1980:32, pl. X: fig. 2.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Manila.

Order FUCALES

Family CYSTOSEIRACEAE

Cystoseira C. Agardh

Cystoseira hakodatensis (Yendo) Fensholt

Cystophyllum hakodatense Yendo, 1907:32, pl. II: figs. 13–16 [lectotype locality: Shoya, Japan fide Yoshida, 1980:100].—Cordero, 1982b:33, fig. 2 [“Cestophyllum”].

Cystoseira hakodatensis (Yendo) Fensholt, 1952:563 [“hakotadense”].

PHILIPPINE DISTRIBUTION.—LEYTE (Leyte del Sur).

NOTE.—The name Cystophyllum J. Agardh (1848:228) was superfluous when published inasmuch as the circumscription of the genus included both of the original species of Myagropsis Kützing (1843a:57), the single original species of Sirophysalis Kützing (1843b:368), as well as several species previously assigned to Sargassum C. Agardh (1820b:1) and Cystoseira C. Agardh (1820b:50). According to Article 7.11 of the ICBN, the type of Cystophyllum is the type of Myagropsis, the earlier of the two Kützing generic names and therefore the one that J. Agardh should have adopted. Fucus myagroides Mertens ex Turner (1809:28, pl. 83) was designated lectotype of Myagropsis by Silva (1952a:279) and is thus the type of Cystophyllum. The lectotypification by De Toni (1891:175), who chose C. trinode (Forsskål) J. Agardh (1848:230), is irrelevant.

J. Agardh separated Cystophyllum from Cystoseira on the basis of vesicle position. In Cystophyllum the vesicles were said to be restricted to ultimate ramuli, while in Cystoseira they are produced within the axes. The diagnostic value of this character at the generic level was doubted by Gardner (1917:390), who found both conditions in the same species. Fensholt (1952) abandoned this distinction and treated as members of Cystoseira most species that had been placed in Cystophyllum by J. Agardh and subsequent authors. If Cystoseira hakodatensis and other species currently placed in Cystophyllum are segregated from Cystoseira, the segregate genus cannot bear the name Cystophyllum, which, as shown above, is a synonym of Myagropsis.

Hormophysa Kützing

Hormophysa cuneiformis (J.F. Gmelin) P.C. Silva, new combination

Fucus triqueter Linnaeus, 1771:312 [type locality: “in Mari Capensi”].

Cystoseira triquetra C. Agardh, 1820b:61.

Hormophysa triquetra (C. Agardh) Kützing, 1843b:359.—Taylor, 1966b:357.—Papenfuss, 1968:45.—Cornejo and Velasquez, 1972:179, 182.—Trono, 1972a:102.—Cordero, 1973b:25.—Velasquez et al., 1973:21, pl. 8: fig. 39.—Ortega, Alcala, and Reyes, 1974:187.—Trono, 1974a:147.—Velasquez, Trono, and Doty, 1975:150.—Trono, 1976:220.—Cordero, 1976c:20.—Trono, 1978:11.—Cordero, 1979b:286.—Puig and Cordero, 1979:32.—Liao and Sotto, 1980:98.—Reyes, 1980:123, pl. 3: fig. 5.—Saraya and Trono, 1980:33, pl. X: fig. 3.—Trono and Ganzon-Fortes, 1980:47, fig. [s.n.].—Guzman, 1981:43.—Laserna et al., 1981:447.—Meñez and Calumpong, 1981:382.—Laserna et al., 1982:52.—Trono and Ang, 1982:14.—Hurtado-Ponce, 1983:123.—Marcos-Agngarayngay, 1984a:11, fig. 8.—Marcos-Agngarayngay, 1984b:123.—Tungpalan, 1984:143.

Fucus articulatus Forsskål, 1775:191 [type locality: Suez, Egypt].

Cystoseira articulate J. Agardh, 1848:216.—Meñez, 1961:68.—Velasquez, Trono, and Doty, 1975:140.

Fucus cuneiformis J.F. Gmelin, 1792:1389.

Cystoseira prolifera J. Agardh, 1848:215 [type locality: Western Australia, Australia].—Westernhagen, 1973a:65; 1974:113 (table 1).

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, La Union, Bataan, Batangas, Sorsogon. MARINDUQUE. LEYTE (Biliran I.), PANAY: Aklan. NEGROS: Negros Oriental, CEBU (Mactan I.), SIQUIJOR. PALAWAN (Bugsuk I.), SULU: Sulu (Siasi I.), Tawitawi.

NOTE.—Cystoseira articulata is included as a synonym on the authority of Setchell (1935:264), C. prolifera on the authority of Womersley (1964:64). Various taxonomic synonyms are discussed by Papenfuss (1968), while the tortuous nomenclature is brought up to date in the appended Nomenclatural Notes.

Myagropsis Kützing

Myagropsis myagroides (Mertens ex Turner) Fensholt

Fucus myagroides Mertens ex Turner, 1809:28, pl. 83 [type locality: Nagasaki, Japan].

Myagropsis myagroides (Mertens ex Turner) Fensholt, 1955:319.

Fucus sisymbrioides Turner, 1809:150, pl. 129 [type locality: Korea Strait].

Cystophyllum sisymbrioides (Turner) J. Agardh, 1848:234.—Cordero, 1982b:35, fig. 3 [“Cestophyllum”].

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

NOTE.—While returning most members of Cystophyllum to Cystoseira, Fensholt (1952) retained the name Cystophyllum for a new generic concept distinguished from Cystoseira by a suite of developmental and cytological characters. Only two species were known by Fensholt to share these characters, namely, C. sisymbrioides (Turner) J. Agardh (1848:234) and C. turneri Yendo (1907:40, pl. III: figs. 7–11). Upon realizing that C. sisymbrioides included both of the original species of Myagropsis, Fensholt (1955) adopted that generic name for what she had previously designated Cystophyllum J. Agardh emend. Fensholt. Papenfuss and Jensen (1967) reviewed Fensholt’s work and concluded that the segregation of Myagropsis from Cystoseira was advantageous.

The conspecificity of Fucus myagroides and F. sisymbrioides was proposed by J. Agardh (1848:234). Both species were originally described in the second volume of Turner’s “Fuci.” If they had been published simultaneously, J. Agardh would have been free to adopt either epithet in preference to the other. Moreover, his choice (sisymbrioides) would have been imposed on subsequent workers (Article 57.2). In fact, however, the two species were not published simultaneously. Turner’s “Fuci” was issued serially, with 12 parts constituting a volume (see Price, 1984). Although exact dates have not been ascertained, it is certain that pl. 83 (F. myagroides) appeared prior to pl. 129 (F. sisymbrioides). Myagropsis myagroides is thus the correct name for this species.

Family SARGASSACEAE

Sargassum C. Agardh

Sargassum aemulum Sonder

Sargassum aemulum Sonder, 1853:672 [type locality: Holdfast Bay (Gulf of St. Vincent), South Australia, Australia].

Sargassum cristatum J. Agardh, 1889:84, pl. XXV: figs. 18–20 [type locality: South Australia, Australia].—Taylor, 1966b:357.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—The synonymy was proposed by Grunow (1915 [1915–1916]:376).

Sargassum agardhianum Farlow ex J. Agardh

Sargassum agardhianum Farlow ex J. Agardh, 1889:93 [type locality: San Diego, California, USA].—Cordero, 1976c:22.

PHILIPPINE DISTRIBUTION.—BATANES.

Sargassum baccularia (Mertens) C. Agardh

Fucus baccularia Mertens, 1819:177 [type locality: not specified (“Ad Novam Hollandiam” fide C. Agardh, 1824:304)].

Sargassum baccularia (Mertens) C. Agardh, 1824:304.—Ang, 1985b:295.

PHILIPPINE DISTRIBUTION.—Batangas.

*Sargassum belangeri Bory de Saint-Vincent

Sargassum belangeri Bory de Saint-Vincent, 1834:162 [syntype localities: Java and Sunda Strait, Indonesia; Philippines].—J. Agardh, 1848:345.—Martens, 1868:72-73.—De Toni, 1895b:113.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Sargassum berberifolium J. Agardh

Sargassum berberifolium J. Agardh, 1848:337 [syntype localities: Western Australia, Australia; Admiralty Is.].—Grunow, 1915 [1915–1916]:411.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Sargassum biforme Sonder

Sargassum biforme Sonder, 1845:51 [type locality: Western Australia, Australia].—Dickie, 1876a:245.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—PANAY: Iloilo (Gigantes Is.).

Sargassum cervicome Greville

Sargassum cervicome Greville, 1849:217, pl. IX: figs. 1–6 [type locality: “in mare Peninsulae Indiae Orientalis”].

*Sargassum binderi Sonder var. angustifolium Sonder, 1871:44 [syntype localities: various, Indian and Pacific oceans, including San Bernardino Strait, between Sorsogon and Northern Samar provinces, Philippines].

PHILIPPINE DISTRIBUTION.—San Bernardino Strait.

NOTE.—The synonymy was proposed by Grunow (1915 [1915–1916]:384).

*Sargassum cinctum J. Agardh var. gracilentum Grunow

Sargassum cinctum J. Agardh var. gracilentum Grunow, 1915 [1915–1916]:419 [“gracilenta”] [syntype localities: Australia: Goodie I., Ballina, and Rockingham; near Manila, Luzon].

PHILIPPINE DISTRIBUTION.—LUZON: Manila.

*Sargassum cinctum J. Agardh var.? mixtum Grunow

Sargassum cinctum J. Agardh var.? mixtum Grunow, 1915 [1915–1916]:419 [“mixta”] [type locality: near Manila, Luzon].

Sargassum siliquosum [misapplied name fide Grunow, l.c.].—Martens, 1868:27, 74–75.—Velasquez, Trono, and Doty, 1975:163 [Martens records only].

PHILIPPINE DISTRIBUTION.—LUZON: La Union, Manila. MINDANAO: Zamboanga.

NOTE.—In view of the fact that Grunow (1916 [1915–1916]:173) did not list the Philippines under Sargassum siliquosum J. Agardh, it appears that he intended to include in S. cinctum var. mixtum the Martens record of S. siliquosum from Zamboanga as well as the one from Manila, even though he cited only the latter locality.

Sargassum confusum C. Agardh

Sargassum confusum C. Agardh, 1824:301 [type locality: “In mari Japonico”].—Flores-Sian, 1959:97.—Agor, 1962:34.—Domantay, 1962:284.—Velasquez, Trono, and Doty, 1975:162.—Cordero, 1976c:8.—Garcia, 1979:44 (table 1).—Puig and Cordero, 1979:33.—Cordero, 1981c:408; 1982a:fig. 6; 1984c:53.—Marcos-Agngarayngay, 1984a:12, fig. 9; 1984b:123.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan. MASBATE. SAMAR: Western Samar, Eastern Samar. LEYTE (Biliran I.), PANAY: Antique, Iloilo. MINDANAO.

Sargassum crispifolium Yamada

Sargassum crispifolium Yamada, 1931a:72, pl. XX [syntype localities: various, all in Japan].—Velasquez, 1971:443, fig. 23.—Cordero, 1979b:276; 198lc:408.—Guzman, 1981:43, 50.—Cordero, 1982a:fig. 8; 1984b:63; 1984c:54.

PHILIPPINE DISTRIBUTION.—MASBATE. SAMAR: Western Samar, Eastern Samar. PANAY: Aklan, Antique, CEBU (Mactan I.). PALAWAN.

Sargassum cristaefolium C. Agardh

Sargassum cristaefolium C. Agardh, 1820b:13 [type locality: “Locus natalis ignotus”].—Montagne, 1844a:660; 1846a:42.—Martens, 1868:26,72–73.—De Toni, 1895b:51.—Velasquez, Trono, and Doty, 1975:162.—Trono, 1978:12.—Saraya and Trono, 1980:32.—Trono and Fortes, 1980:68.—Chan, 1981:385.—Ganzon-Fortes, 1981:21—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:10, pl. VI: figs. 1, 2.—Trono and Ang, 1982:14.—Trono and Fortes, 1982:147.

Sargassum ilicifolium (Turner) C. Agardh var. duplicatum J. Agardh, 1848:318 [type locality: Moluccas, Indonesia].

Sargassum duplicatum (J. Agardh) J. Agardh, 1889:90.—Sulit and San Juan, 1955:48 (table 1).—Meñez, 1961:65.—Domantay, 1962:286.—Taylor, 1966b:357.—Manapat, 1969:37, 38, 39.—Trono, 1974a:147.—Velasquez, Trono, and Doty, 1975:162.—Trono, 1976:219.—Cordero, 1976c:22; 1979b:276, 286, fig. 3.—Puig and Cordero, 1979:33.—Cordero, 1980a:29, 36, fig. 4; 1980b:35, pl. 22.—Laserna et al., 1981:447.—Cordero, 1984a:85.—Marcos-Agngarayngay, 1984a:12, fig. 10.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Manila, Batangas, Quezon, Sorsogon. MARINDUQUE. MINDORO: Occidental Mindoro (incl. Lubang Is.), SAMAR: Eastern Samar. LEYTE (Biliran I.), PANAY: Aklan. SIQUIJOR. MINDANAO: Zamboanga del Sur, Surigao del Sur. SULU: Sulu (Siasi I.). PALAWAN (incl. Bugsuk I.).

NOTE.—J. Agardh (1848:318) was uncertain whether his plant from the Moluccas was conspecific with Sargassum duplicatum Bory de Saint-Vincent (1828 [1826–1829]:127; type locality: Pacific Ocean between Tahiti and New Zealand) and later (1889:90, 123) recognized his S. duplicatum as a species distinct from that of Bory de Saint-Vincent. Womersley and Bailey (1970:296) treated the two species as one entity, which they merged into S. cristaefolium.

TAXON OF UNCERTAIN VALUE

The following infraspecific taxon is probably related to Sargassum cristaefolium but has not formally been transferred to it.

*Sargassum duplicatum (J. Agardh) J. Agardh var. rotundatum Grunow

Sargassum duplicatum (J. Agardh) J. Agardh var. rotundatum Grunow, 1915 [1915–1916]:392 [“rotundata”] [syntype localities: Philippines; Java, Indonesia; Seychelles].

PHILIPPINE DISTRIBUTION.—Locality not specified.

Sargassum cystocarpum C. Agard

Sargassum cystocarpum C. Agardh, 1820b:33 [type locality: “E mari Indico”].—Martens, 1868:26, 72–73.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

*Sargassum cystophyllum Montagne

Sargassum cystophyllum Montagne, 1842d:249 [type locality: “Manilla,” Luzon].—Montagne, 1846a:46.—J. Agardh, 1848:345.—Montagne, 1856:388.—De Toni, 1895b:113.—Grunow, 1915 [1915–1916]:414.

Carpacanthus cystophyllus (Montagne) Kützing, 1849:623.—Martens, 1868:76-77.—Velasquez, Trono, and Doty, 1975:130.

PHILIPPINE DISTRIBUTION.—As above.

Sargassum droserifolium Bory de Saint-Vincent

Sargassum droserifolium Bory de Saint-Vincent, 1828 [1826–1829]:129 [syntype localities: Port Praslin (Kambotorosh Harbor), New Ireland; New Zealand; Tahiti; New Guinea].—Grunow, 1915 [1915–1916]:410.

PHILIPPINE DISTRIBUTION.—Locality not specified.

*Sargassum droserifolium Bory de Saint Vincent var.? spathulatum Grunow

Sargassum droserifolium Bory de Saint-Vincent var.? spathulatum Grunow, 1915 [1915–1916]:411 [“spathulata”] [type locality: “in freto Bernardino” (San Bernardino Strait between Sorsogon Prov., Luzon, and Northern Samar Prov.)].

PHILIPPINE DISTRIBUTION.—As above.

Sargassum elongatum

Sargassum elongatum.—Manapat, 1969:38.

PHILIPPINE DISTRIBUTION.—PALAWAN.

NOTE.—Manapat did not give an author for this name and we have been unable to find its place of publication.

Sargassum esperi C. Agardh

Sargassum esperi C. Agardh, 1820b:9 [syntype localities: Brazil; Bengal, India].—Montagne, 1844a:660.—Martens, 1868:74-75.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Sargassum filicinum Harvey

Sargassum filicinum Harvey, 1860a:327 [type locality: “East coast of Japan”].—Garcia, 1979:44 (table 1).

PHILIPPINE DISTRIBUTION.—MINDANAO.

Sargassum filifolium C. Agardh

Sargassum filifolium C. Agardh, 1824:305 [type locality: Western Australia, Australia].—Grunow, 1916 [1915–1916]:2.

PHILIPPINE DISTRIBUTION.—“In freto Bernardino” [San Bernardino Strait between Sorsogon Prov., Luzon, and Northern Samar Prov.].

Sargassum filifolium C. Agardh var. aciculare (Grunow) Grunow

Sargassum aciculare Grunow, 1874:26 [type locality: Bribie I., Queensland, Australia].

Sargassum filifolium C. Agardh var. aciculare (Grunow) Grunow, 1916 [1915–1916]:3 [“acicularis”].

PHILIPPINE DISTRIBUTION.—“In freto Bernardino” [San Bernardino Strait between Sorsogon Prov., Luzon, and Northern Samar Prov.].

*Sargassum filiforme Montagne

Sargassum filiforme Montagne, 1844a:660 [type locality: “in insulis Philippinensibus”].—J. Agardh, 1848:346.—Kützing, 1849:614.—Montagne, 1856:383.—Martens, 1868:27, 74-75.—De Toni, 1895b:113.—Grunow, 1916 [1915–1916]:14.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

Sargassum fluitans (Børgesen) Børgesen

Sargassum hystrix J. Agardh var. fluitans Børgesen, 1914a:11, fig. 8 [type locality: Sargasso Sea].

Sargassum fluitans (Børgesen) Børgesen, 1914b:66, footnote.—Westernhagen, 1973a:66; 1974:113 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Sargassum fulvellum (Turner) C. Agardh

Fucus fulvellus Turner, 1808:147, pl. 66 [type locality: Korea Strait].

Sargassum fulvellum (Turner) C. Agardh, 1820b:34.—Domantay, 1962:287.—Cordero, 1973b:25 [S. fulvellum prox.].—Velasquez, Trono, and Doty, 1975:162.—Cordero, 1976c:23; 1979b:276; 1981c:407; 1982a:fig. 7; 1984c:53.

Sargassum enerve C. Agardh, 1820b:17 [type locality: “In Oceano” (“E mari Coreano” fide Yoshida, 1983:148, fig. 26c)].—Sulit and San Juan, 1955:48 (table 1) pl. 2: fig. 1.—Domantay, 1962:274.—Manapat, 1969:38.—Westernhagen, 1973a:66; 1974:113 (table 1).—Trono and Fortes, 1980:68—Ganzon-Fortes, 1981:21—Trono and Fortes, 1982:147.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Batangas. MASBATE. SAMAR: Western Samar. LEYTE (Biliran I.), PANAY: Aklan, Iloilo. CEBU (Mactan I.).

NOTE.—Sargassum enerve is included as a synonym on the authority of Yoshida (1983:148, 151).

Sargassum furcatum Kützing

Sargassum furcatum Kützing, 1843b:362 [type locality: St. Thomas, Virgin Is.].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

NOTE.—Grunow (1916 [1915–1916]:43), in agreement with J. Agardh (1889:108), considered this species a variety of S. vulgare, but Taylor (1960:277) recognized both species

*Sargassum gaudichaudii Montagne

Sargassum gaudichaudii Montagne, 1842d:249 [syntype localities: Mauritius; “Manilla,” Luzon].—Montagne, 1846a:48, pl. 141; 1856:384.—Grunow, 1915 [1915–1916]:446.

Carpacanthus gaudichaudii (Montagne) Kützing, 1849:623.—Martens, 1868:76–77.—Velasquez, Trono, and Doty, 1975:130.

PHILIPPINE DISTRIBUTION.—LUZON: Manila.

Sargassum giganteifolium Yamada

Sargassum giganteifolium Yamada in Okamura, 1925:105, pl. CCXXX [syntype localities: Enoshima, Kanagawa Prefecture and Futomi, Chiba Prefecture, Japan].—Sulit and San Juan, 1955:48 (table 1).—Domantay, 1962:284.—Manapat, 1969:37, 38.—Cordero, 1973b:26 [with query].—Velasquez, Trono, and Doty, 1975:162.—Cordero, 1981c:408; 1984b:63; 1984c:53.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.) Pangasinan, Batangas, Albay (Batan I.), MASBATE. SAMAR: Western Samar. LEYTE (Biliran I.), PANAY: Iloilo. CEBU (Mactan I.). PALAWAN.

NOTE.—Yoshida (1983:195) cites the holotype of this species as coming from Shichirigahama (Kanagawa Prefecture), Japan, but this locality is not mentioned in the protologue.

Sargassum gracile J. Agardh

Sargassum gracile J. Agardh, 1848:310 [syntype localities: “in Oceano Indico ad Javam” and “inter Borneo et Sumatram,” Indonesia].—Dickie, 1876a:243.—Grunow, 1916 [1915–1916]:5.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.). Basilan Strait (between Mindanao and Basilan I.).

Sargassum granuliferum C. Agardh

Sargassum granuliferum C. Agardh, 1820b:31 [type locality: “In mari Indico?”].—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Trono and Fortes, 1980:68.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:147.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Sargassum hemiphyllum (Turner) C. Agardh

Fucus hemiphyllus Turner, 1811:85, pl. 169 [type locality: Nagasaki, Japan].

Sargassum hemiphyllum (Turner) C. Agardh, 1820b:39.—Montagne, 1844a:661 [with query].—Sulit and San Juan, 1955:48 (table 1), pl. 2: fig. 2.—Domantay, 1962:285.—Manapat, 1969:37.—Velasquez, Trono, and Doty, 1975:162.—Cordero, 1976c:23.—Trono and Fortes, 1980:68.—Cordero, 1981c:406.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:147.—Cordero, 1984c:53.

Spongocarpus hemiphyllus (Turner) Kützing, 1849:632.—Martens, 1868:80–81.—Velasquez, Trono, and Doty, 1975:163.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. MASBATE. SAMAR: Western Samar, Eastern Samar. PANAY: Iloilo. CEBU (Mactan I.).

Sargassum heterocystum Montagne

Sargassum heterocystum Montagne, 1842d:250 [type locality: “In mari chinensi Cochinchinam alluente” (Vietnam)].—Grunow, 1915 [1915–1916]:409.

PHILIPPINE DISTRIBUTION.—Locality not specified.

*Sargassum hombronianum Montagne var.? manilense (Grunow) Grunow

Sargassum polycystum C. Agardh var. manilense Grunow in Piccone, 1886:44, 90 [“manilensis”] [type locality: Cavite Prov., Luzon].—De Toni, 1895b:104.

Sargassum hombronianum Montagne var.? manilense (Grunow) Grunow, 1916 [1915–1916]:8 [“manilensis”].

PHILIPPINE DISTRIBUTION.—As above.

Sargassum ilicifolium (Turner) C. Agardh

Fucus ilicifolius Turner, 1808:113, pl. 51 [type locality: Sunda Strait, Indonesia].

Sargassum ilicifolium (Turner) C. Agardh, 1820b:11.—Dickie, 1874a:190; 1876a:243.—Piccone, 1886:45, 90.—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Velasquez, Trono, and Doty, 1975:162.—Trono and Fortes, 1980:68.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:148.

Carpacanthus ilicifolius (Turner) Kützing, 1849:625.—Martens, 1868:27, 78–79.—Velasquez, Trono, and Doty, 1975:130.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Manila, Cavite. MINDORO. MINDANAO: Zamboanga.

*Sargassum ilicifolium (Turner) C. Agardh f. benkulense Grunow

Sargassum ilicifolium (Turner) C. Agardh f. benkulense Grunow, 1915 [1915–19I6]:407 [“benkulensis”] [syntype localities: Pulau Tikus near Bengkulu, Sumatra; Manila, Luzon].

PHILIPPINE DISTRIBUTION.—As above.

Sargassum ilicifolium (Turner) C. Agardh var. compactum (Bory de Saint-Vincent) Grunow

Sargassum compactum Bory de Saint-Vincent, 1828 [1826–1829]:126 [type locality: Concepción, Chile].

Sargassum ilicifolium (Turner) C. Agardh var. compactum (Bory de Saint-Vincent) Grunow, 1915 [1915–1916]:405.

Carpacanthus spinulosus [misapplied name fide Grunow, l.c.].—Martens, 1868:27, 78–79 [in part].—Velasquez, Trono, and Doty, 1975:130 [in part].

PHILIPPINE DISTRIBUTION.—LUZON: Manila

Sargassum ilicifolium (Turner) C. Agardh var.? pseudospinulosum Grunow

Sargassum ilicifolium (Turner) C. Agardh var.? pseudospinulosum Grunow, 1915 [1915–1916]:403 [“pseudospinulosa”] [type locality: near Atapupu Timor, Indonesia].

Carpacanthus spinulosus [misapplied name fide Grunow, l.c.].—Martens, 1868:27, 78–79 [in part].—Velasquez, Trono, and Doty, 1975:130 [in part].

PHILIPPINE DISTRIBUTION.—LUZON: Manila.

NOTE.—In view of the fact that Grunow (1915 [1915–1916]:399) attributed Carpacanthus spinulosus Kützing (as Sargassum hystrix var. spinulosus) solely to the West Indies, it appears that he intended to include in S. ilicifolium var.? pseudospinulosum the Martens record of C. spinulosus from Manila as well as the one from Timor, even though he cited only the latter locality.

Sargassum latifolium (Turner) C. Agardh

Fucus latifolius Turner, 1809:66, pl. 94 [type locality: Red Sea].

Sargassum latifolium (Turner) C. Agardh, 1820b:13.—Dickie, 1876a:243.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

*Sargassum microcystum J. Agardh

Sargassum microcystum J. Agardh, 1848:323 [syntype localities: Singapore; Moluccas; Manila, Luzon].

Carpacanthus microcystis (J. Agardh) Martens, 1868:28, 78–79 [including Philippine record].—Velasquez, Trono, and Doty, 1975:130.

PHILIPPINE DISTRIBUTION.—LUZON: Manila. MINDANAO: Zamboanga.

*Sargassum microcystum J. Agardh f. luzonense Grunow

Sargassum microcystum J. Agardh f. luzonense Grunow, 1915 [1915–1916]:413 [“luzonensis”] [type locality: near Manila, Luzon].

Sargassum carpophyllum [misapplied name fide Grunow, 1915 [1915–1916]:413].—Grunow, 1867:56.

PHILIPPINE DISTRIBUTION.—As above.

*Sargassum microcystum J. Agardh var. microtis Grunow

Sargassum microcystum J. Agardh var. microtis Grunow, 1915 [1915–1916]:414 [syntype localities: Java; Basilan Strait (between Mindanao and Basilan I.)].

PHILIPPINE DISTRIBUTION.—Basilan Strait.

Sargassum microphyllum C. Agardh

Sargassum microphyllum C. Agardh, 1820b:33 [type locality: “In India oriental”].—Grunow, 1915 [1915–1916]:446.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Sargassum miyabei Yendo

Sargassum miyabei Yendo, 1907:112, pl. XIV: figs. 13, 14 [lectotype locality: Cape Soya, Hokkaido, Japan fide Yoshida, 1978:122].

Sargassum kjellmanianum Yendo, 1907:102, pl. XV: figs. 1–4 [lectotype locality: Takashima, Hokkaido, Japan fide Yoshida, 1978:122].—Sulit and San Juan, 1955:48 (table 1).—Domantay, 1962:285.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. PALAWAN.

NOTE.—The synonymy was proposed by Yoshida (1978).

Sargassum myriocystum J. Agardh

Sargassum myriocystum J. Agardh, 1848:314 [syntype localities: Jakarta, Java, Indonesia; China; India].—Grunow, 1915 [1915–1916]:440.

PHILIPPINE DISTRIBUTION.—Locality not specified.

*Sargassum myriocystum J. Agardh var. euryphyllum (Grunow) Grunow

Sargassum polycystum J. Agardh var. euryphyllum Grunow in Piccone, 1886:44, 90 [“euryphylla”] [type locality: Ticao I., Masbate Prov.].—De Toni, 1895b:104.

Sargassum myriocystum]. Agardh var. euryphyllum (Grunow) Grunow, 1915 [1915–1916]:440 [“euryphylla”].

PHILIPPINE DISTRIBUTION.—As above.

Sargassum natans (Linnaeus) Gaillon

Fucus natans Linnaeus, 1753:1160 [type locality: “in Pelago libere natans”].—M. Blanco, 1845:579; 1879:259.—Velasquez, Trono, and Doty, 1975:144.

Sargassum natans (Linnaeus) Gaillon, 1828:355.

Fucus bacciferus Turner, 1802:xxxii, 55 [superfluous name].

Sargassum bacciferum C. Agardh, 1820b:6.—Montagne, 1844a:660.—Velasquez, Trono, and Doty, 1975:162.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos.

NOTE.—Fucus bacciferus Turner, the intended basionym of Sargassum bacciferum, is a superfluous and hence illegitimate name since F. natans was cited as a synonym in the protologue. In the first edition of Flora de Filipinas, M. Blanco (1837:839) described this entity as a new species, Fucus denticulatus (a name used previously for four different species!). Martens (1868:45) pointed out that Blanco’s description fits various species of Sargassum. In the absence of authentic specimens, no definitive placement can be made. Montagne’s record, on the other hand, is documented by extant collections (Cuming 2251, presumably at PC and possibly other herbaria) and could be re-evaluated.

Sargassum nigrifolium Yendo

Sargassum nigrifolium Yendo, 1907:127, pl. XVI: figs. 1–3 [lectotype locality: Misaki, Kanagawa Prefecture, Japan fide Yoshida, 1980:104].—Sulit and San Juan, 1955:48 (table 1), pl. 3.—Domantay, 1962:286.—Manapat, 1969:38.—Velasquez, Trono, and Doty, 1975:162.—Cordero, 1976c:24 [S. nigrifolium prox.]; 1981c:406.—Guzman, 1981:43, 51.—Cordero, 1984c:53.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. MASBATE. SAMAR: Western Samar. PANAY: Iloilo. PALAWAN.

Sargassum oligocystum Montagne

Sargassum oligocystum Montagne, 1845:67–69 [type locality: Lampung Bay, Sumatra, Indonesia].

Sargassum binderi Sonder in J. Agardh, 1848:328 [syntype localities: Sunda Strait and Java, Indonesia; China Sea].—Piccone, 1886:47, 90.—De Toni, 1895b:47.—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Velasquez, Trono, and Doty, 1975:162.—Cordero, 1976c:22.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Cavite.

NOTE.—The synonymy was proposed by Womersley and Bailey (1970:299).

*Sargassum oligocystum Montagne var.? bernardinum Grunow

Sargassum oligocystum Montagne var.? bernardinum Grunow, 1915 [1915–1916]:386 [“bernardina”] [type locality: “in freto Bernardino” (San Bernardino Strait between Sorsogon Prov., Luzon, and Northern Samar Prov.)].

PHILIPPINE DISTRIBUTION.—As above.

*Sargassum oligocystum Montagne var.? subflexuosum Grunow

Sargassum oligocystum Montagne var.? subflexuosum Grunow, 1915:385 [“subflexuosa”] [syntype localities: “Fretum Bernardinum, Macassar”].

PHILIPPINE DISTRIBUTION.—San Bernardino Strait between Sorsogon Prov., Luzon, and Northern Samar Prov.

TAXON OF UNCERTAIN VALUE

The following infraspecific taxon is probably related to Sargassum oligocystum, but has not been formally transferred to it.

Sargassum binderi Sonder var. vitiense (Grunow) Grunow

Sargassum echinocarpum J. Agardh var. vitiense Grunow, 1874:28 [type locality: Ovalau I., Fiji].

Sargassum binderi Sonder var. vitiense (Grunow) Grunow in Askenasy, 1888:28.—Piccone, 1889:31, 59.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Sargassum oocyste J. Agardh

Sargassum oocyste J. Agardh, 1848:97 [syntype localities: Chile; Moluccas, Indonesia].—Grunow, 1915 [1915–19I6]:437.

PHILIPPINE DISTRIBUTION.—LUZON.

*Sargassum oocyste J. Agardh var. bernardinum Grunow

Sargassum oocyste J. Agardh var. bernardinum Grunow, 1915 [1915–1916]:437 [“bernardina”] [type locality: “in freto Bernardino” (San Bernardino Strait between Sorsogon Prov., Luzon, and Northern Samar Prov.)].

PHILIPPINE DISTRIBUTION.—As above.

Sargassum paniculatum J. Agardh

Sargassum paniculatum J. Agardh, 1848:315 [type locality: “in mari Indico”].—Cordero, 1976c:24, fig. N.—Ang, 1984:548; 1985a:231; 1985b:294; 1985c:52.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Batangas.

Sargassum parvifolium (Turner) C. Agardh

Fucus parvifolius Turner, 1819 [1815–1819]:33, pl. 221 [type locality: not specified].

Sargassum parvifolium (Turner) C. Agardh, 1820b:30.—Dickie, 1876a:243.—Grunow, 1916 [1915–1916]:13.—Velasquez, Trono, and Doty, 1975:163.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Sargassum patens C. Agardh var. schizophyllum (Kützing) Yendo

Halochloa schizophylla Kützing, 1843a:56 [type locality: Japan].

Sargassum patens C. Agardh var. schizophyllum (Kützing) Yendo, 1905a:155 [“schizophylla”].—Sulit and San Juan, 1955: table 1.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

*Sargassum philippinense Grunow

Sargassum philippinense Grunow, 1916 [1915-1916]:166 [type locality: “in freto Bernardino” (San Bernardino Strait between Sorsogon Prov., Luzon, and Northern Samar Prov.)].

PHILIPPINE DISTRIBUTION.—As above.

Sargassum piluliferum (Turner) C. Agardh

Fucus pilulifer Turner, 1808:145, pl. 65 [type locality: Nagasaki, Japan].

Sargassum piluliferum (Turner) C. Agardh, 1820b:27.—Manapat, 1969:38.—Cordero, 1981c:407; 1984c:54.

PHILIPPINE DISTRIBUTION.—MASBATE. SAMAR: Western Samar. PANAY: Iloilo. CEBU (Mactan I.). PALAWAN.

Sargassum polyceratium Montagne

Sargassum polyceratium Montagne, 1837:356 [type locality: La Habana, Cuba].—Velasquez et al., 1973:22, pl. 8: fig. 41.—Hamoy and Garciano, 1975:71.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Bataan. CENTRAL VISAYAS. CEBU.

Sargassum polycystum C. Agardh

Sargassum polycystum C. Agardh, 1824:304 [type locality: Sunda Strait, Indonesia].—Montagne, 1844a:661.—Martens, 1868:72-73.—Dickie, 1874a:190.—De Toni, 1895b:103.—Howe, 1932:170.—Zaneveld, 1952:132; 1956:19; 1959:102.—Velasquez, Trono, and Doty, 1975:163.—Trono, 1976:219; 1978:12.—Trono and Tuason, 1978:13.—Fortes and Trono, 1980:61.—Trono and Fortes, 1980:68.—Trono and Ganzon-Fortes, 1980:49, fig. [s.n.].—Chan, 1981:385.—Ganzon-Fortes, 1981:21.—Laserna et al., 1981:447.—Trono and De Lara, 1981:9, pl. VI: fig. 3.—Laserna et al., 1982:52.—Trono and Ang, 1982:15.—Trono and Fortes, 1982:148.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), MARINDUQUE. NEGROS: Negros Occidental. PALAWAN (Bugsuk I.).

Sargassum pteropleuron Grunow

Sargassum pteropleuron Grunow, 1867:55, pl. V: fig. 1 [type locality: Nassau, New Providence I., Bahama Is.].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Sargassum sagamianum Yendo

Sargassum sagamianum Yendo, 1907:151, pl. XVII: figs. 6-10 [lectotype locality: Oosashi, Mie Prefecture, Japan fide Yoshida, 1980:105, pl. VII: fig. 13].—Domantay, 1962:274.—Manapat, 1969:37.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Sargassum sandei Reinbold

Sargassum sandei Reinbold in Weber-van Bosse, 1913a:158, fig. 47, pl. IV: figs. 1, 2 [type locality: Flores, Indonesia].—Taylor, 1966b:357.—Velasquez, Trono, and Doty, 1975:163.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Sargassum serratifolium (C. Agardh) C. Agardh

Fucus serratifolius C. Agardh, 1815:41 [type locality: “prope Satsuma,” Kagoshima Prefecture, Japan].

Sargassum serratifolium (C. Agardh) C. Agardh, 1820b:16.—Sulit and San Juan, 1955:48 (table 1), pl. 1: fig. 2.—Domantay, 1962:285.—Manapat, 1969:37, 38.—Velasquez, Trono, and Doty, 1975:163.—Trono and Fortes, 1980:68.—Cordero, 1981c:407.—Trono and Fortes, 1982:148.—Cordero, 1984c:53.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Sur, Pangasinan. MASBATE. SAMAR: Western Samar, Eastern Samar. PANAY: Iloilo. CEBU (Mactan I.). PALAWAN.

Sargassum siliquosum J. Agardh

Sargassum siliquosum J. Agardh, 1848:316 [syntype localities: Singapore; Jakarta, Java, Indonesia].—Piccone, 1886:45, 90.—De Toni, 1895b:107.—Seale, 1911:309.—Wester, 1916:159; 1921:224; 1924: 21.—Howe, 1932:170.—G. Blanco, 1938:513.—Quisumbing, 1951:1012.—Zaneveld, 1952:132.—Montilla and Blanco, 1953:166, fig. 5:6 [“siliguisum”].—Sulit and San Juan, 1955:48 (table 1), pl. 1: fig. 1.—Zaneveld, 1956:20; 1959:102.—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Manapat, 1969:37.—Bersamin et al., 1973:188.—Velasquez, Trono, and Doty, 1975:163.—Cordero, 1976c:24.—Trono and Fortes, 1980:68.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:148.—Ang, 1984:548; 1985a:231; 1985b:294; 1985c:52.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Ilocos Sur, La Union, Pangasinan, Cavite, Batangas. MASBATE (Ticao I.), NEGROS: Negros Occidental. MINDANAO: Zamboanga.

*Sargassum siliquosum J. Agardh var.? basilanicum Grunow

Sargassum siliquosum J. Agardh var.? basilanicum Grunow, 1916 [1915–1916]:175 [“basilanica” ] [type locality: Basilan Strait (between Mindanao and Basilan I.)].

PHILIPPINE DISTRIBUTION.—As above.

*Sargassum siliquosum J. Agardh var. bicornutum Grunow

Sargassum siliquosum J. Agardh var. bicornutum Grunow, 1916 [1915–1916]:175 [“bicornuta”] [syntype localities: China Sea; Ticao I., Masbate Prov.].

PHILIPPINE DISTRIBUTION.—MASBATE (Ticao I.).

*Sargassum siliquosum J. Agardh var. manipaense Grunow

Sargassum siliquosum J. Agardh var. manipaense Grunow, 1916 [1915–1916]:175 [“manipaensis” ] [syntype localities: “ad insulas Philippinas, Manipa”].

PHILIPPINE DISTRIBUTION.—As above.

Sargassum spinifex C. Agardh

Sargassum spinifex C. Agardh, 1820b:29 [type locality: “In mari Chinensi”].—Bailey and Harvey, 1862:173.—Quisumbing, 1951:1012.—Velasquez, Trono, and Doty, 1975:163.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.). MINDANAO.

Sargassum subspathulatum (Grunow) Grunow

Sargassum pulchellum Grunow var. subspathulatum Grunow in Askenasy, 1888:25, pl. VI: figs. 5, 6 [including Philippine record] [type locality: MacCluer Gulf (Berau Bay), Irian Barat, Indonesia].

Sargassum subspathulatum (Grunow) Grunow, 1916 [1915–1916]:10.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Sargassum tenerrimum J. Agardh

Sargassum tenerrimum J. Agardh, 1848:305 [type locality: Bombay, India].—Cordero, 1976c:25 [S. tenerrimum prox.].

PHILIPPINE DISTRIBUTION.—BATANES.

Sargassum vulgare C. Agardh

Fucus salicifolius S.G. Gmelin, 1768:98 [syntype localities: “Mare americanum?”; Sea of Marmara, Turkey].

Sargassum vulgare C. Agardh, 1820b:3 [superfluous name].—Montagne, 1846a:40.—Martens, 1868:74–75.—Westernhagen, 1973a:66.—Rivera, 1974:108.—Westernhagen, 1974:113 (table 1).—Velasquez, Trono, and Doty, 1975:163.

PHILIPPINE DISTRIBUTION.—LUZON: Manila, CEBU (incl. Mactan I.).

NOTE.—The name Sargassum vulgare is traditionally given to a common species of the tropical Atlantic Ocean (e.g., Taylor, 1960:272). At the time of its publication, however, this name was superfluous and hence illegitimate, since the species to which it was applied included Fucus salicifolius S.G. Gmelin 1768, which may or may not prove to be conspecific with S. vulgare as traditionally interpreted. Future research will undoubtedly turn up a taxonomic synonym that can serve as the correct name.

Sargassum vulgare C. Agardh var. indicum C. Agardh

Sargassum vulgare C. Agardh var. indicum C. Agardh, 1820b:4 [type locality: “E mari Indico” ].—Montagne, 1846a:40.

PHILIPPINE DISTRIBUTION.—LUZON: Manila.

Sargassum yendoi Okamura and Yamada

Sargassum yendoi Okamura and Yamada in Yamada, 1938b:121, figs. 1, 2, pls. 21, 22 [syntype localities: various, all in Japan].—Domantay, 1962:287.—Velasquez, Trono, and Doty, 1975:163.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Turbinaria Lamouroux

Turbinaria condensata Sonder

Turbinaria condensata Sonder in Kützing, 1860:25, pl. 69: fig. II [type locality: “In mari chinensi”].—Taylor, 1964:482, pl. 2: figs. 18–28.—Velasquez, Trono, and Doty, 1975:165.—Trono and Fortes, 1980:67; 1982:148.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte. MINDANAO: Surigao. BASILAN.

Turbinaria conoides (J. Agardh) Kützing

Turbinaria vulgaris J. Agardh var. conoides J. Agardh, 1848:267 [syntype localities: India; China; Sri Lanka; Western Australia, Australia].

Turbinaria conoides (J. Agardh) Kützing, 1860:24, pl. 66: fig. II.—Howe, 1932:169.—Zaneveld, 1952:133; 1956:22; 1959:104.—Domantay 1962:287.—Taylor, 1964:480, pl. 2: figs. 1–8 [var. conoides]; 1966b:358.—Trono, 1972a:100; 1973d:12.—Velasquez, Trono, and Doty, 1975:165.—Trono, 1976:218.—Liao and Sotto, 1980:98.—Saraya and Trono, 1980:33, pl. X: fig. 4 [var. conoides].—Trono and Fortes, 1980:67.—Trono and Ganzon-Fortes, 1980:55, fig. [s.n.].—Ganzon-Fortes, 1981:21.—Laserna et al., 1981:447.—Meñez and Calumpong, 1981:382.—Laserna et al., 1982:52.—Luistro, Cajipe, and Laserna, 1982:46.—Trono and Fortes, 1982:148.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Sur, Pangasinan, Batangas, Quezon (incl. Polillo Is.), Sorsogon. MINDORO: Oriental Mindoro. PANAY. CEBU (incl. Mactan I.). SIQUIJOR. MINDANAO: Zamboanga. BASILAN. PALAWAN (incl. Balabac I.), SULU: Tawitawi.

*Turbinaria conoides (J. Agardh) Kützing f. laticuspidata W.R. Taylor

Turbinaria conoides (J. Agardh) Kützing f. laticuspidata W.R. Taylor, 1964:481, pl. 2: figs. 9–12 [type locality: Cebu I.].—Trono, 1972a:100.

PHILIPPINE DISTRIBUTION.—CEBU. MINDANAO: Zamboanga, Surigao. PALAWAN, SULU: Tawitawi.

*Turbinaria conoides (J. Agardh) Kützing f. retroflexa W.R. Taylor

Turbinaria conoides (J. Agardh) Kützing f. retroflexa W.R. Taylor, 1964:481 [intended type locality: Tandayong I., Pangasinan Prov., Luzon].

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. NEGROS: Negros Occidental

NOTE.—The name of this form is invalid because the protologue lacks an illustration in contravention of Article 39 of the ICBN.

Turbinaria decurrens Bory de Saint-Vincent

Turbinaria decurrens Bory de Saint-Vincent, 1828 [1826-1829]:119 [type locality: floating between the Society Is. and New Guinea].—Taylor, 1964:477, pl. 1: figs. 22-28; 1966a:92; 1966b:358.—Kraft, 1972:329.—Velasquez, Trono, and Doty, 1975:166.—Trono, 1976:218.—Trono and Young, 1977:58.—Saraya and Trono, 1980:33, pl. X:fig. L—Trono and Fortes, 1980:67; 1982:148.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Sur, Pangasinan, Albay, Sorsogon. CATANDUANES. CEBU. MINDANAO: Misamis Oriental, Surigao del Sur, Davao. BASILAN. SULU: Tawitawi (Turtle Is.).

Turbinaria denudata Bory de Saint-Vincent

Turbinaria denudata Bory de Saint-Vincent, 1828 [1826-1829]:117 [syntype localities: various, in the Pacific and Indian oceans and the Caribbean Sea].—Montagne, 1844a:661.—Bailey and Harvey, 1862:173.—Velasquez, Trono, and Doty, 1975:166.

PHILIPPINE DISTRIBUTION.—MINDANAO.

NOTE.—According to J. Agardh (1848:266, 267) and Barton (1891:217, 219), Turbinata denudata as originally proposed by Bory de Saint-Vincent included specimens referable to T. ornata (Turner) J. Agardh and T. conoides (J. Agardh) Kützing. Bory de Saint-Vincent also included material from the West Indies that probably is referable to T. turbinata (Linnaeus) Kuntze, the commonest Turbinaria in that region. Turbinaria denudata should be lectotypified on the basis of authentic specimens at PC. Depending on the specimen chosen, T. denudata would become a synonym of T. turbinata or displace either T. ornata or T. conoides. Taylor (1964; 1966a) did not mention T. denudata.

Turbinaria filamentosa Yamada

Turbinaria filamentosa Yamada, 1925b:243, fig. I [type locality: Garan-bi (O-luan), Taiwan].—Cordero, 1976c:25, fig. O.

PHILIPPINE DISTRIBUTION.—BATANES.

*Turbinaria luzonensis W.R. Taylor

Turbinaria luzonensis W.R. Taylor, 1964:482, pl. 2: figs. 13-17. [type locality: Mulanay, Quezon Prov., Luzon].—Velasquez, Trono, and Doty. 1975:166.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon.

Turbinaria murrayana Barton

Turbinaria murrayana Barton, 1891:218, pl. LIV (54): fig. 2 [syntype localities: New Guinea; Macassar, Celebes, Indonesia].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Turbinaria ornata (Turner) J. Agardh

Fucus turbinatus Linnaeus var. ornatus Turner, 1808:50, pl. 24: figs. c,d [type locality: not specified].

Turbinaria ornata (Turner) J. Agardh, 1848:266.—Martens, 1868:27,47, 76-77.—Dickie, 1874a:190.—Zaneveld, 1952:132; 1956:21; 1959:103.—Agor, 1962:34.—Domantay, 1962:287.—Taylor, 1964:483, pl. 3: figs. 1–6 [var. ornata]; 1966a:98; 1966b:358.—Villones and Magdamo, 1968:27, fig. 22.—Manapat, 1969:37, 38.—Velasquez, 1971:443, fig. 24—Cornejo and Velasquez, 1972:171, 175, 183.—Kraft, 1972:328, 329, 332.—Reyes, 1972:147.—Trono, 1972a:100.—Cordero, 1973b:26.—Velasquez et al., 1973:22, pl. 8: fig. 40.—Ortega, Alcala, and Reyes, 1974:185, 186, 188.—Hamoy and Garciano, 1975:71—Velasquez, Trono, and Doty, 1975:166.—Cordero, 1976c:9, 26.—Trono, 1976:217; 1978:14.—Trono and Tuason, 1978:13.—Vannajan and Trono, 1978:14, fig. 11.—Cordero, 1979b:276.—Garcia, 1979:44 (table 1).—Puig and Cordero, 1979:33.—Liao and Sotto, 1980:98.—Reyes, 1980:122, pl. 4: fig. 1.—Saraya and Trono, 1980:33.—Trono and Fortes, 1980:67.—Trono and Ganzon-Fortes, 1980:57, fig. [s.n.].—Ganzon-Fortes, 1981:21.—Laserna et al., 1981:447.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:10, pl. VII: fig. 3.—Laserna et al., 1982:52.— Trono and Fortes, 1982:148.—Hurtado-Ponce, 1983:123.—Cordero, 1984b:62; 1984c:53.—Marcos-Agngarayngay, 1984a:14, fig. 12.—Tungpalan, 1984:143.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Ilocos Sur, Pangasinan, Bataan, Cavite, Batangas, Quezon, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. MASBATE. LEYTE (Biliran I.). PANAY: Aklan, Antique (Panagatan Cays). NEGROS: Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Zamboanga, Davao. BASILAN. PALAWAN (incl. Culion I., Balabac I.), SULU: Sulu, Tawitawi (Cagayan Sulu I.).

Turbinaria ornata (Turner) J. Agardh f. evesiculosa (Barton) W.R. Taylor

Turbinaria conoides (J. Agardh) Kützing var. evesiculosa Barton, 1891:217 [type locality: Edam (Damar Besar) I./Enkhuizen shoal, near Jakarta, Java, Indonesia].

Turbinaria ornata (Turner) J. Agardh f. evesiculosa (Barton) W.R. Taylor, 1964:485.—Trono, 1973d:12, pl. 7: fig. 28.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CEBU.

Turbinaria trialata (J. Agardh) Kützing

Turbinaria vulgaris]. Agardh var. trialata]. Agardh, 1848:268 [syntype localities: Brazil; West Indies].

Turbinaria trialata (J. Agardh) Kützing, 1860:24, pl. 67.—Domantay, 1962:287.—De Leon, Eufemio, and Pineda, 1963:82 (table 1).—Velasquez, Trono, and Doty, 1975:166.—Cordero, 1979b:286.—Trono and Fortes, 1980:67; 1982:148.—Cordero, 1984b:62; 1984c:53.—Marcos-Agngarayngay, 1984a:16, fig. 13.—Tungpalan, 1984:143.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan. MASBATE. PANAY: Aklan. MINDANAO: Zamboanga. PALAWAN.

Turbinaria trialata (J. Agardh) Kützing var. capensis Kützing

Turbinaria trialata (J. Agardh) Kützing var. capensis Kützing, 1860:24, pl. 67: fig. II [type locality: “Caput bonae spei” (Cape of Good Hope, South Africa)].—Meñez, 1961:68.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Turbinaria turbinata (Linnaeus) Kuntze

Fucus turbinatus Linnaeus, 1753:1160 [type locality: “ in Americae rupibus marinis” (Jamaica fide Howe, 1920:591)].

Turbinaria turbinata (Linnaeus) Kuntze, 1898:434.—Meñez, 1961:68.—Westernhagen, 1973a:66; 1974:113 (table 1).—Velasquez, Trono, and Doty, 1975:166.—Trono and Fortes, 1980:67; 1982:148.—Cordero, 1984b:62.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CEBU (Mactan I.). PALAWAN.

NOTE.—According to Taylor (1964:476), T. turbinata is an Atlantic-Caribbean species.

Turbinaria vulgaris J. Agardh

Turbinaria vulgaris J. Agardh, 1848:267 [type locality: floating between the Society Is. and New Guinea].—Dickie, 1876a:243, 244.—Velasquez, Trono, and Doty, 1975:166.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.). MINDANAO: Zamboanga.

NOTE.—Turbinaria vulgaris is a name used by J. Agardh to encompass four varieties, namely, α conoides, β decurrens, γ triquetra, and δ trialata. All but var. decurrens were newly described and were subsequently elevated to specific rank. Var. decurrens, however, was based on T. decurrens Bory de Saint-Vincent, so that according to the ICBN, T. vulgaris is a superfluous name (Article 63.1) and is automatically typified by the type of T. decurrens (Article 7.11). To which of the four component species the Dickie records should be referred can be determined only from an examination of the specimens.

Class CHLOROPHYCEAE

Order CTENOCLADALES

Family ULVELLACEAE

Phaeophila Hauck

Phaeophila dendroides (P. Crouan and H. Crouan) Batters

Ochlochaete dendroides P. Crouan and H. Crouan, 1852:no. 346 [type locality: Brest, Finistère, France].

Phaeophila dendroides (P. Crouan and H. Crouan) Batters, 1902:13.—Tanaka, 1967:13, fig. 1.—Velasquez, Trono, and Doty, 1975:159.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.).

Order ULVALES

Family MONOSTROMATACEAE

Monostroma Thuret

Monostroma latissimum Wittrock

Monostroma latissimum Wittrock, 1866:33, pl. 1: fig. 4 [syntype localities: various, including Europe, North Africa, and New Zealand].—Taylor, 1966b:343.—Velasquez, Trono, and Doty, 1975:154.

PHILIPPINE DISTRIBUTION.—PALAWAN.

Monostroma nitidum Wittrock

Monostroma nitidum Wittrock, 1866:41, pl. II: fig. 7 [syntype localities: Tonga; China].—Gilbert, 1961:415.—Trono, 1973c:5.—Velasquez, Trono, and Doty, 1975:154.—Cordero, 1980b:19, pl. 4.—Trono and Fortes, 1980:66.—Ganzon-Fortes, 1981:21.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. SIQUIJOR. PALAWAN.

Family ULVACEAE

Enteromorpha Link

The taxonomic arrangement of the species of this genus follows that proposed by Bliding (1963).

Enteromorpha aragoensis Bliding

Enteromorpha aragoensis Bliding, 1960:174, fig. 2 [syntype localities: various, in Atlantic France and Mediterranean Sea],—Gilbert and Doty, 1969:121.—Velasquez, Trono, and Doty, 1975:142.

PHILIPPINE DISTRIBUTION.—LUZON: Quezon.

Enteromorpha chaetomorphoides Børgesen

Enteromorpha chaetomorphoides Børgesen, 1911:149, fig. 12 [type locality: St. Thomas, Virgin Is.].—Fortes and Trono, 1980:55.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Enteromorpha clathrata (Roth) Greville

Conferva clathrata Roth, 1806:175 [type locality: Baltic Sea, Germany].

Enteromorpha clathrata (Roth) Greville, 1830:lxvi, 181.—Trono, 1973b:213.—Velasquez et al., 1973:9, pl. 1: fig. 3.—Trono, 1975:25.—Cordero, 1976c:10; 1977b:19, pl. 1: figs. 1, 2.—Reyes, 1978:146, pl. 3: figs. 2–5.—Calumpong, 1980:143 (table 3).—Puig and Cordero, 1979:18.—Cordero, 1980b:16, fig. 1A.—Liao and Sotto, 1980:96.—Trono and De Lara, 1981:2, pl. I: fig. 2.—Calumpong, 1982:145.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Batangas, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), LEYTE (Biliran I.), NEGROS: Negros Oriental (incl. Apo I.), CEBU (Mactan I.), SIQUIJOR. BOHOL. SULU: Sulu (Siasi I.).

NOTE.—J. Agardh (1883:153) is traditionally accredited with the combination Enteromorpha clathrata, but in fact it was first made by Greville.

Enteromorpha compressa (Linnaeus) Nees

Ulva compressa Linnaeus, 1753:1163 [type locality: Europe].

Enteromorpha compressa (Linnaeus) Nees, 1820: Index [2].—Galutira and Velasquez, 1964:494, pl. 1: fig. 2; pl. 6: fig. 20.—Cornejo and Velasquez, 1972:173, pl. 1: fig. 7.—Velasquez, 1972:63.—Westernhagen, 1973a:64; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:142.—Cordero, 1976c:9; 1977b:20.—Vannajan and Trono, 1977:39.—Puig and Cordero, 1979:18.—Velasquez, 1979b:230.—Cordero, 1980b:17, pl. 1.—Saraya and Trono, 1980:8.—Trono and Fortes, 1980:65.—Chan, 1981:387.—Fortes, 1981b:396.—Ganzon- Fortes, 1981:21.—Meñez and Calumpong, 1981:382.—Calumpong, 1982:145.—Cordero, 1982a:60, fig. 1.—Trono and Fortes, 1982:145.—Hurtado-Ponce, 1983:104.—Hurtado-Ponce and Modelo, 1983:146.—Marcos-Agngarayngay, 1983:66, fig. 4.—Cordero, 1984a:70.—Hurtado-Ponce, 1984:179.—Marcos-Agngarayngay, 1984b:l19.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Manila, Batangas. SAMAR: Eastern Samar. LEYTE (Biliran I.), PANAY: Aklan. NEGROS: Negros Occidental (incl. Ilacaon I.), Negros Oriental, CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Surigao del Sur.

NOTE.—Either Link (1820:5) or Greville (1830:lxvi, 180) is traditionally accredited with having first made the combination Enteromorpha compressa, but in fact it was Nees who did so in the index that he prepared for the collective work in which Link’s article was published.

Enteromorpha flexuosa (Wulfen) J. Agardh

Conferva flexuosa Roth, 1800b:188 [type locality: Trieste, Italy].

Ulva flexuosa Wulfen, 1803:1.

Enteromorpha flexuosa (Wulfen) J. Agardh, 1883:126 [including subsp. flexuosa].—Piccone, 1886:25, 89.—Gilbert, 1946:77; 1961:414.—Taylor, 1966b:343.—Gilbert and Doty, 1969:121, fig. 1.—Cornejo and Velasquez, 1972:173, pl. 4: fig. 30.—Trono, 1973b:213; 1975:25.—Velasquez, Trono, and Doty, 1975:142.—Vannajan and Trono, 1977:39, fig. 15.—Carumbana and Luchavez, 1980:189.

Enteromorpha intestinalis (Linnaeus) Link var. tubulosa Kützing, 1845:247 [type locality: fresh water, Germany].

Enteromorpha tubulosa (Kützing) Kützing, 1856:11, pl. 32: fig. II.—Sulit, Navarro, and San Juan, 1952:170 (table 5).—Rabanal and Montalban, 1953:143.—Villadolid and Villaluz, 1953:5–10.—Villaluz, 1953:103.—Gilbert, 1961:415.—Rabanal and Montalban, 1961:4.—Velasquez, Trono, and Doty, 1975:142.—Vannajan and Trono, 1977:39.—Trono and Ang, 1982:3.

Enteromorpha intestinalis (Linnaeus) Link f. tubulosa (Kützing) V.J. Chapman, 1937:229.—Reyes, 1972:138.

Enteromorpha intermedia Bliding, 1955:262, figs. 1–5 [syntype localities: various in northern Europe; USA].—Gilbert and Doty, 1969:121, fig. 2.—Velasquez, Trono, and Doty, 1975:142.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Sur, Rizal, Manila, Batangas, Quezon, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MASBATE (Ticao I.), NEGROS: Negros Oriental. MINDANAO: Zamboanga. BASILAN. PALAWAN (incl. Balabac I., Bugsuk I., Cuyo I.). SULU: Sulu (Siasi I.).

NOTE.—Conferva flexuosa Roth, the intended basionym of Enteromorpha flexuosa, is a later homonym of C. flexuosa O.F. Müller (1782:5, pl. 882) and hence not priorable. Ulva flexuosa Wulfen is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Enteromorpha flexuosa (Wulfen) J. Agardh subsp. paradoxa (C. Agardh) Bliding

Conferva paradoxa Dillwyn, 1809 [1802–1809]:70, suppl. pl. F [syntype localities: Bangor, Wales; Brighton, England].

Ulva paradoxa C. Agardh, 1817:XXII.

Enteromorpha flexuosa (Wulfen) J. Agardh subsp. paradoxa (C. Agardh) Bliding, 1963:79, figs. 42–45.

Scytosiphon erectus Lyngbye, 1819:65, pl. 15:C [type locality: Norway].

Enteromorpha erecta (Lyngbye) Carmichael in W. Hooker, 1833:314.—Gilbert, 1961:414.—Velasquez, Trono, and Doty, 1975:142.

Enteromorpha plumosa Kützing, 1843b:300, pl. 20:1 [type locality: Trieste, Italy],—Gilbert, 1961:415.—Galutira and Velasquez, 1964:494, pl. 1: figs. 1a–c;pl. 6: fig. 19.—Velasquez, 1972:63.—Velasquez, Trono, and Doty, 1975:142.—Velasquez, 1979b:230.—Trono and Fortes, 1980:65.—Ganzon-Fortes, 1981:21.—Cordero, 1982a:60.—Trono and Fortes, 1982:146.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Bulacan, Rizal.

NOTE.—Conferva paradoxa Dillwyn, the intended basionym of Enteromorpha flexuosa subsp. paradoxa, is a later homonym of C. paradoxa Roth (1806:172) and hence not priorable. Ulva paradoxa C. Agardh is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Enteromorpha flexuosa (Wulfen) J. Agardh f. submarina Collins and Hervey

Enteromorpha flexuosa (Wulfen) J. Agardh f. submarina Collins and Hervey, 1917:34 [type locality: Bermuda].—Taylor, 1966b:343.

PHILIPPINE DISTRIBUTION.—LUZON: Rizal.

Enteromorpha intestinalis (Linnaeus) Nees

Ulva intestinalis Linnaeus, 1753:1163 [type locality: “in Mari omni”].—M. Blanco, 1837:842; 1845:582; 1879:262.—Velasquez, Trono, and Doty, 1975:167.

Enteromorpha intestinalis (Linnaeus) Nees, 1820, Index [2].—Martens, 1868:47, 60–61.—Seale, 1911:309.—Wester, 1916:158.—Merrill, 1918:39.—Wester, 1921:224; 1924:21.—G. Blanco, 1938:512.—Gilbert, 1946:77.—Zaneveld, 1950:113.—Quisumbing, 1951:1009.—Montilla and Blanco, 1953:166.—Rabanal and Montalban, 1953:143.—Rabanal et al., 1953:156.—Velasquez, 1953a:100.—Zaneveld, 1956:11; 1959:97.—Gilbert, 1961:415.—Rabanal and Montalban, 1961:4.—Villones and Magdamo, 1968:25, fig. 12.—Velasquez, 1971:431, fig. 5.—Reyes, 1972:137.—Trono, 1973C:3, fig. 17.—Velasquez et al., 1973:9, pl. 1: fig. 2.—Westernhagen, 1973a:64; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:142.—Vannajan and Trono, 1977:38, fig. 16.—Reyes, 1978:146, 162, pl. 3: fig. 1.—Puig and Cordero, 1979:18.—Carumbana and Luchavez, 1980:189.—Cordero, 1980b:18, pl. 2.—Liao and Sotto, 1980:96.—Trono and Fortes, 1980:65.—Trono and Ganzon-Fortes, 1980:5, fig. [s.n].—Fortes, 1981b:396.—Ganzon-Fortes, 1981:21.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:1, pl. I: fig. 1.—Calumpong, 1982:145.—Cordero, 1982a:60.—Trono and Fortes, 1982:146.—Hurtado-Ponce, 1983:105.—Hurtado-Ponce and Modelo, 1983:146.—Marcos-Agngarayngay, 1983:69, fig. 5.—Cordero, 1984a:70.—Marcos-Agngarayngay, 1984b:120.—Tungpalan, 1984:144.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Pangasinan, Bataan, Rizal, Manila, Cavite, Batangas, Quezon, MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. LEYTE (Biliran I.). NEGROS: Negros Oriental (incl. Apo I.), CEBU (Mactan I.). SIQUIJOR.

NOTE.—Either Link (1820:5) or Greville (1830:lxvi, 179) is traditionally accredited with having first made the combination Enteromorpha intestinalis, but in fact it was Nees who did so in the index that he prepared for the collective work in which Link’s article was published.

Enteromorpha kylinii Bliding

Enteromorpha kylinii Bliding, 1948:199, figs. 1–3 [“kylini”] [type locality: Kristineberg, Sweden].—Trono, 1973b:213; 1973c:4.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. SULU: Sulu (Siasi I.).

Enteromorpha lingulata J. Agardh

Enteromorpha lingulata J. Agardh, 1883:143 [syntype localities: North Atlantic; Gulf of Mexico; New Zealand; Tasmania].—Howe, 1932:169.—Gilbert, 1946:77; 1961:415.—Taylor, 1966b:343.—Velasquez, Trono, and Doty, 1975:142.—Cordero, 1976c:10; 1977b:20.—Vannajan and Trono, 1977:38, fig. 14.—Cordero, 1980b:18, pl. 3.—Trono and Fortes, 1980:65.—Chan, 1981:387, 389.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:146.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Ilocos Sur, Pangasinan, Rizal, Manila, Quezon. PANAY. BASILAN.

Enteromorpha prolifera (O.F. Müller) J. Agardh

Ulva prolifera O.F. Müller, 1778:7, pl. 763(1) [type locality: Denmark].

Enteromorpha prolifera (O.F. Müller) J. Agardh, 1883:129.—Merrill, 1918:39.—Gilbert, 1946:77; 1961:415.—Cordero, 1973b:16.—Velasquez, Trono, and Doty, 1975:142.—Trono and Fortes, 1980:65.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:146.—Cordero, 1984a:70.

Enteromorpha salina Kützing, 1845:247 [type locality: near Hildesheim, West Germany].—Chan, 1981:387, 389.

Ulva compressa [misapplied name fide Collins in Merrill, 1918:39].—M. Blanco, 1837:842; 1845:581; 1879:261.—Velasquez, Trono, and Doty, 1975:167.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Ilocos Sur, Pangasinan, Manila Bay, Batangas. LEYTE (Biliran I.). MINDANAO: Zamboanga.

NOTE.—Martens (1868:47, 62–63) identified Blanco’s plant as Enteromorpha complanata Kützing var. crinita (Nees) Kützing, which Bliding (1963:119) referred to E. ramulosa. In the absence of authentic specimens, no definitive placement can be made.

Enteromorpha ramulosa (J.E. Smith) Carmichael

Ulva ramulosa J.E. Smith, 1810 [1809–1810]: pl. 2137 [type locality: Bantry Bay, Eire].

Enteromorpha ramulosa (J.E. Smith) Carmichael in W. Hooker, 1833:315.—Taylor, 1966b:343 [E. ramulosa prox.].—Gilbert and Doty, 1969:122, fig. 3.—Velasquez, Trono, and Doty, 1975:142.—Fortes, 1981b:396.

Conferva crinita Roth, 1797:162, pl. 1: fig. 3 [type locality: near Eckwarden, Oldenburg, West Germany].

Enteromorpha crinita Nees, 1820: Index [2].—Martens, 1868:62–63.—Weber-van Bosse, 1913a:55.—Gilbert, 1946:77; 1961:414.—Velasquez, Trono, and Doty, 1975:142.—Saraya and Trono, 1980:8.

Enteromorpha clathrata (Roth) Greville var. crinita (Nees) Hauck, 1885 [1883–1885]:429.—Meñez and Calumpong, 1981:382.

Enteromorpha ramulosa (J.E. Smith) Carmichael var. spinosa Kützing, 1845:247 [syntype localities: North Sea; Adriatic Sea].

Enteromorpha spinescens Kützing, 1856:12, pl 33: fig. III.—Taylor, 1966b:343.—Velasquez, Trono, and Doty, 1975:142.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Ilocos Sur, Pangasinan, Rizal, Batangas. MINDORO: Oriental Mindoro. CENTRAL VISAYAS. SULU: Sulu (Jolo I.).

NOTE.—Conferva crinita Roth, the intended basionym of Enteromorpha crinita, is a later homonym of C. crinita Withering (1776:751) and hence not priorable. Enteromorpha crinita Nees is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN. In elevating E. ramulosa var. spinosa to the status of species, Kützing altered the spelling of the epithet to spinescens.

Ulva Linnaeus

Ulva fasciata Delile

Ulva fasciata Delile, 1813:297, pl. 58: fig. 5 [type locality: Alexandria, Egypt].—Gilbert, 1961:415.—Reyes, 1972:138.—Rivera, 1974:108.—Velasquez, Trono, and Doty, 1975:167.—Trono and Fortes, 1980:65.—Ganzon-Fortes, 1981:21.—Cordero, 1982a:60, 61.—Trono and Fortes, 1982:145.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental. CEBU. SULU: Sulu (Jolo I.).

Ulva lactuca Linnaeus

Ulva lactuca Linnaeus, 1753:1163 [type locality: “in Oceano”].—Gilbert, 1946:77.—Velasquez, 1949:155.—Zaneveld, 1950:113.—Montilla and Blanco, 1953:166.—Zaneveld, 1956:12; 1959:98.—Gilbert, 1961:416.—Galutira and Velasquez, 1964:495, pl. 1, fig. 3; pl. 6: fig. 21.—Taylor, 1966b:344.—Velasquez, 1968a:120, fig. 3.—Villones and Magdamo, 1968:25, fig. 16.—Velasquez, 1971:431, fig. 6.—Reyes, 1972:138.—Trono, 1972a:87, 89.—Velasquez, 1972:63.—Trono, 1973a:127; 1973b:213; 1973c:5.—Velasquez et al., 1973:9, pl 1: fig. 1.—Westernhagen, 1974:112 (table 1).—Hamoy and Garciano, 1975:71.—Trono, 1975:26.—Velasquez, Trono, and Doty, 1975:167.—Vannajan and Trono, 1977:40, fig. 12, 13.—Reyes, 1978:146, 162, pl. 3: fig. 7.—Velasquez, 1979b:230.—Cordero, 1980b:19, pls. 5, [56].—Liao and Sotto, 1980:96.—Trono and Fortes, 1980:65.—Trono, Velasquez, and Guevarra, 1980:77.—Ganzon-Fortes, 1981:21.—Guzman, 1981:43, 50, 52.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:2, pl. 1: fig. 4.—Calumpong, 1982:145.—Cordero, 1982a:60, 61.—Luistro, Cajipe, and Laserna, 1982:46.—Trono and Fortes, 1982:145.—Hurtado-Ponce, 1983:105.—Hurtado-Ponce and Modelo, 1983:146.—Marcos-Agngarayngay, 1983:70, fig. 6.—Cordero, 1984a:71.—Hurtado-Ponce, 1984:179.—Marcos-Agngarayngay, 1984b:120.—Tungpalan, 1984:139.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Manila Bay, Rizal, Cavite, Batangas, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. NEGROS: Negros Occidental (Ilacaon I.), Negros Oriental (incl. Apo I.), CEBU (incl. Mactan I.), SIQUIJOR. MINDANAO: Zamboanga. BASILAN. PALAWAN (incl. Cuyo I.). SULU: Sulu (Siasi I.).

Ulva latissima auctorum

Ulva latissima [misapplied name].

Ulva lactuca Linnaeus var. latissima [misapplied name].—Weber-van Bosse, 1913a:50.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Pearl Bank).

NOTE.—The name Ulva latissima Linnaeus has traditionally been misapplied to representatives of the genus with very broad sheet-like fronds, whereas the type is referable to Laminaria saccharina Linnaeus) Lamouroux (see Silva, 1952a:295 and Papenfuss, 1960:303). While European records apparently are referable to Ulva gigantea (Kützing) Bliding (1969:558), Weber-van Bosse’s material probably belongs elsewhere.

Ulva pertusa Kjellman

Ulva pertusa Kjellman, 1897b:4, pl 1: figs. 1–5; pl. 3: figs. 1–8 [syntype localities: various, all in Japan].—Domantay, 1962:281.—Reyes, 1972:138.—Trono, 1975:26.—Velasquez, Trono, and Doty, 1975:167.—Cordero, 1976c:9, 10; 1977b:20; 1980b:20, pl. 6.—Trono and Fortes, 1980:65.—Ganzon-Fortes, 1981:21.—Cordero, 1982a: fig. 2.—Trono and Fortes, 1982:145.—Cordero, 1984a:71; 1984c:50.—Tungpalan, 1984:139.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Sorsogon. MASBATE. NEGROS: Negros Oriental. SIQUIJOR.

Ulva reticulata Forsskål

Ulva reticulata Forsskål, 1775:187 [syntype localities: “Gomfodae” (Al-Qunfidha), Saudi Arabia; Mokha, Yemen].—Montagne, 1844a:659.—Dickie, 1876a:243, 244.—piccone, 1886:22, 89.—Weber-van Bosse, 1913a:52.—Gilbert, 1946:77; 1961:416.—Meñez, 1961:47, pl. 1: figs. 14–16.—Domantay, 1962:281.—Taylor, 1966b:344.—Velasquez, 1971:431, fig. 7.—Reyes, 1972:138.—Trono, 1972a:94.—Cordero, 1973b:16.—Trono, 1973b:212; 1973d:4,pl. 1: fig. 3.—Westernhagen, 1973b:369 (table 3).—Ortega, Alcala, and Reyes, 1974:187.—Rivera, 1974:108.—Westernhagen, 1974:112 (table 1).—Trono, 1975:27.—Velasquez, Trono, and Doty, 1975:167 [excluding Blanco and Martens records].—Taylor, 1977b:8.—Reyes, 1978:147, pl. 3: figs. 8, 9.—Cordero, 1979b:279.—Puig and Cordero, 1979:19.—Calumpong, 1980:143 (table 3).—Cordero, 1980b:20, pl. [45].—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:7.—Cordero, 1981d:63, fig. 4.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:2, pl. 1: fig. 3.—Hurtado-Ponce, 1983:106.—Hurtado-Ponce and Modelo, 1983:146.—Marcos-Agngarayngay, 1983:70, fig. 7.—Cordero, 1984a:71; 1984b:57; 1984c:50.—Hurtado-Ponce, 1984:179.—Marcos-Agngarayngay, 1984b:120.—Tungpalan, 1984:139.

Phycoseris reticulata (Forsskål) Kützing, 1849:478.—Martens, 1868:23, 60–61 [excl. Blanco record].—Velasquez, Trono, and Doty, 1975:159.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos Norte, La Union, Pangasinan, Cavite, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.). MASBATE (incl. Ticao I.). SAMAR: Eastern Samar. LEYTE (Biliran I.). PANAY: Aklan. NEGROS: Negros Oriental, CEBU (Mactan I.). SIQUIJOR. BOHOL. MINDANAO: Zamboanga. BASILAN. PALAWAN. SULU: Sulu (Jolo I., Siasi I.), Tawitawi.

Ulva rigida C. Agardh

Ulva rigida C. Agardh, 1822a:410 [lectotype locality: Cádiz, Spain fide Papenfuss, 1960:305].

Ulva lactuca Linnaeus var. rigida (C. Agardh) Le Jolis, 1863:38.—Weber-van Bosse, 1913a:50.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (Jolo I.).

Order CLADOPHORALES

Family ANADYOMENACEAE

Anadyomene Lamouroux

Anadyomene brownii (J.E. Gray) J. Agardh

Calomena brownii J.E. Gray, 1866:46, pl. XLIV: fig. 3 [type locality: Australia].

Anadyomene brownii (J.E. Gray) J. Agardh, 1887:127.—Taylor, 1966b:349.—Velasquez, Trono, and Doty, 1975:128.

PHILIPPINE DISTRIBUTION.—PALAWAN (Balabac I.). SULU: Tawitawi (Turtle Is.).

*Anadyomene eseptata Gilbert

Anadyomene eseptata Gilbert, 1961:425, pl. I [type locality: Dalupiri I., Babuyan Is., Cagayan Prov., Luzon].—Velasquez, Trono, and Doty, 1975:128.—Cordero, 1977b:30, pl. III: fig. 12; pl. IV: fig. 15.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), CENTRAL VISAYAS.

*Anadyomene leclancheri Decaisne

Anadyomene leclancheri Decaisne, 1844:236 [type locality: “Sooloo”].—Gilbert, 1946:77; 1961:427.—Velasquez, Trono, and Doty, 1975:128.

Cystodictyon leclancheri (Decaisne) J.E. Gray, 1866:72, pl. XLIV: fig. 6 [“leclancherii”].—De Toni, 1889:373.

PHILIPPINE DISTRIBUTION.—Sulu Archipelago.

NOTE.—This species was made the type of a new genus, Cystodictyon, by J.E. Gray (1866:72). It was returned to Anadyomene by Heydrich (1894:283).

Anadyomene plicata C. Agardh

Anadyomene plicata C. Agardh, 1822a:400 [type locality: “Ravak” (Lawak), Waigeo I., Moluccas, Indonesia].—Gilbert, 1961:427.—Taylor, 1966b:349.—Trono, 1972a:94; 1973a:127, fig. 1; 1973c:9; 1975:29.—Velasquez, Trono, and Doty, 1975:128.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:10.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Zambales, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. CEBU (Mactan I.). PALAWAN. SULU: Sulu (Siasi I.), Tawitawi.

Anadyomene stellata (Wulfen) C. Agardh

Ulva stellata Wulfen in Jacquin, 1786:351 [type locality: Adriatic Sea].

Anadyomene stellata (Wulfen) C. Agardh, 1822a:400.—Gilbert, 1946:77; 1961:427.—Galutira and Velasquez, 1964:499.—Villones and Magdamo, 1968:12, fig. 4.—Velasquez, 1971:432, fig. 12.—Reyes, 1972:140.—Velasquez et al., 1973:13, pl. 3: fig. 15.—Ortega, Alcala, and Reyes, 1974:187.—Velasquez, Trono, and Doty, 1975:128.—Reyes. 1978:149, pl. 5: fig. 6.—Hurtado-Ponce, 1983:108.—Hurtado-Ponce and Modelo, 1983:147.—Marcos-Agngarayngay, 1983:75, fig. 12.

Anadyomene flabellata Lamouroux, 1816:366, pl. XIV: fig. 3 [type locality: “Dans la Mousse de Corse des pharmaciens” (Alsidium helminthochortos)].—Dickie, 1876a:245.—Velasquez, Trono, and Doty, 1975:128.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Batangas, Albay. MINDORO: Oriental Mindoro. NEGROS: Negros Oriental, SIQUIJOR. MINDANAO: Zamboanga. PALAWAN.

NOTE.—The synonymy was proposed by C. Agardh (1822a:400).

Anadyomene wrightii Harvey ex J.E. Gray

Anadyomene wrightii Harvey ex J.E. Gray, 1866:48, pl. XLIV: fig. 5 [type locality: Ryukyu-retto, Japan].—Gilbert, 1961:428.—Meñez, 1961:49.—Velasquez, Trono, and Doty, 1975:128.—Puig and Cordero, 1979:19.—Meñez and Calumpong, 1981:382.—Cordero, 1984a:74; 1984c:51.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan. MASBATE. LEYTE (Biliran I.). CENTRAL VISAYAS.

Microdictyon Decaisne

Microdictyon agardhianum Decaisne

Microdictyon agardhianum Decaisne, 1841:115 [type locality: Red Sea near Jidda, Saudi Arabia].—Reyes, 1972:140.—Velasquez et al., 1973:12, pl. 3: fig. 13.—Velasquez, Trono, and Doty, 1975:153.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. NEGROS: Negros Oriental.

Microdictyon boergesenii Setchell

Microdictyon boergesenii Setchell, 1925:106 [type locality: St. Jan (St. John), Virgin Is.].—Reyes, 1978:149, pl. 5: fig. 7.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

*Microdictyon clathratum Martens

Microdictyon clathratum Martens, 1868:25, 66–67, pl. IV: fig. 1. [syntype localities: Ceram and Flores, Indonesia; Zamboanga, Mindanao].—De Toni, 1889:363.—Velasquez, Trono, and Doty, 1975:153.

PHILIPPINE DISTRIBUTION.—As above.

Microdictyon curtissiae W.R. Taylor

Microdictyon curtissiae W.R. Taylor, 1955:69, figs. 1–8, pl. I: fig. 2; pl. III [type locality: southeast coast of Florida, USA].—Westernhagen, 1973a:64; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Microdictyon japonicum Setchell

Microdictyon japonicum Setchell, 1925:107 [type locality: Tateyama, Chiba Prefecture, Japan].—Cordero, 1977b:29, pl. III: fig. 14.—Hurtado-Ponce, 1983:109.—Hurtado-Ponce and Modelo, 1983:147.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte.

Microdictyon okamurae Setchell

Microdictyon okamurae Setchell, 1925:107 [“Okamurai”] [type locality: Ryukyu-retto, Japan].—Cordero, 1977b:29, pl. III: fig. 13.

PHILIPPINE DISTRIBUTION.—BATANES.

Family CLADOPHORACEAE

Chaetomorpha Kützing

Chaetomorpha aerea (Dillwyn) Kützing

Conferva aerea Dillwyn, 1806 [1802–1809]: pl. 80 [syntype localities: various, in England and Wales].

Chaetomorpha aerea (Dillwyn) Kützing, 1849:379.—Velasquez, 1953a:100.—Reyes, 1972:140.—Velasquez, Trono, and Doty, 1975:133.—Trono and Fortes, 1980:63.—Fortes, 1981b:396.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:144.

PHILIPPINE DISTRIBUTION.—LUZON (southern), NEGROS: Negros Oriental.

NOTE.—Arguments for and against the conspecificity of Chaetomorpha aerea and C. linum have been presented from time to time. We follow Kornmann (1972) in recognizing separate species.

Chaetomorpha antennina (Bory de Saint-Vincent) Kützing

Conferva antennina Bory de Saint-Vincent, 1804a:381; 1804b:161, footnote [type locality: Réunion].

Chaetomorpha antennina (Bory de Saint-Vincent) Kützing, 1847a:166.—Gilbert, 1961:430.—Meñez, 1961:47.—Velasquez, Trono, and Doty, 1975:133.

Conferva media C. Agardh, 1824:100 [type locality: West Indies].

Chaetomorpha media (C. Agardh) Kützing, 1849:380.—Marcos-Agngarayngay, 1983:74, fig. 9.—Cordero, 1984a:75.—Marcos-Agngarayngay, 1984b:121.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Quezon.

NOTE.—The conspecificity of Chaetomorpha antennina and C. media is discussed by Børgesen (1940:37) and Gilbert (1961:430).

Chaetomorpha brachygona Harvey

Chaetomorpha brachygona Harvey, 1858:87, pl. XLVI A [syntype localities: Key West, Florida, USA; mouth of Rio Bravo (Rio Grande), border of Mexico and Texas, USA].—Gilbert, 1961:430.—Meñez, 1961:47.—Taylor, 1966b:344.—Velasquez, Trono, and Doty, 1975:133.—Cordero, 1976c:10; 1977b:21.—Reyes, 1978:149, pl. 5: fig. 3.—Chan, 1981:387.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Rizal, Manila, SIQUIJOR.

Chaetomorpha crassa (C. Agardh) Kützing

Conferva crassa C. Agardh, 1824:99 [syntype localities: Trieste and Venezia, Italy; England].

Chaetomorpha crassa (C. Agardh) Kützing, 1845:204.—Seale, 1911:309.—Wester, 1916:158; 1921:224; 1924:21.—G. Blanco, 1938:512.—Zaneveld, 1950:111.—Quisumbing, 1951:1008.—Montilla and Blanco, 1953:166.—Zaneveld, 1956:8; 1959:94.—Gilbert, 1961:431.—Domantay, 1962:279.—Reyes, 1972:140.—Trono, 1972a:94.—Velasquez et al., 1973:10, pl. 1: fig. 4.—Westernhagen, 1973a:64; 1973b:369 (table 3); 1974:112 (table 1).—Trono, 1975:28.—Velasquez, Trono, and Doty, 1975:133.—Cordero, 1976c:9, 10; 1977b:21.—Taylor, 1977b:8.—Reyes, 1978:148, 162, pl. 5: figs. 1, 2.—Tahil, 1978:52.—Cordero, 1979b:279.—Puig and Cordero, 1979:19.—Cordero, 1980b:21, pl. [60].—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:8.—Trono and Fortes, 1980:63.—Trono and Ganzon-Fortes, 1980:7, fig. [s.n].—Chan, 1981:387, 389.—Cordero, 1981d:63.—Ganzon-Fortes, 1981:21.—Guzman, 1981:42, 48.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:3, pl. II: fig. 3.—Calumpong, 1982:145.—Cordero, 1982a:60.—Trono and Fortes, 1982:144.—Hurtado-Ponce, 1983:106.—Hurtado-Ponce and Modelo, 1983:146.—Marcos-Agngarayngay, 1983:73, fig. 8.—Meñez, Phillips, and Calumpong, 1983:18.—Cordero, 1984a:75.—Marcos-Agngarayngay, 1984b:121.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, La Union, Pangasinan, Bataan, Manila, Batangas, Albay, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SAMAR: Eastern Samar. LEYTE (Biliran I.). PANAY: Aklan. NEGROS: Negros Oriental (incl. Apo I.).(Mactan I.). SIQUIJOR. MINDANAO: Zamboanga. BASILAN. PALAWAN. SULU: Tawitawi.

Chaetomorpha gracilis Kützing

Conferva gracilis Kützing, 1843b:259 [type locality: Trieste, Italy].

Chaetomorpha gracilis Kützing, 1845:203.—Gilbert, 1961:432.—Velasquez et al., 1973:10, pl. 2: fig. 5.—Velasquez, Trono, and Doty, 1975:134.

PHILIPPINE DISTRIBUTION.—LUZON: Manila, Batangas. MINDORO: Occidental Mindoro (Lubang Is.).

NOTE.—Conferva gracilis Kützing, the intended basionym of Chaetomorpha gracilis, is a later homonym of Conferva gracilis Wulfen (1803:21), C. gracilis (Vaucher) De Candolle (in Lamarck and De Candolle, 1805:55), and C. gracilis Griffiths (in Harvey, 1834b:304) and hence is not priorable. Chaetomorpha gracilis Kützing is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN. This species was placed in the genus Lola by Chapman (1961:74).

Chaetomorpha inflata Kützing

Chaetomorpha inflata Kützing, 1849:378 [type locality: Java, Indonesia].—Martens, 1868:22, 58–59.—Gilbert, 1946:77; 1961:432.—Velasquez, Trono, and Doty, 1975:134.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga.

*Chaetomorpha kellersii Howe

Chaetomorpha kellersii Howe, 1932:169, fig. 1 [type locality: “Panay Island”].—Gilbert, 1946:77; 1961:432.—Velasquez, Trono, and Doty, 1975:134.

PHILIPPINE DISTRIBUTION.—As above.

Chaetomorpha ligustica (Kützing) Kützing

Conferva ligustica Kützing, 1845:259 [type locality: Golfo di Genova, Italy].

Chaetomorpha ligustica (Kützing) Kützing, 1849:376.

Chaetomorpha tortuosa [misapplied name].—Taylor, 1966b:344.—Velasquez, Trono, and Doty, 1975:134.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), Manila.

NOTE.—The name Chaetomorpha tortuosa (Dillwyn) Kützing, although often cited, is non-existent. In fact, Kützing (1845:205) transferred Conferva tortuosa Dillwyn (1805 [1802–1809]: pl. 46) to Rhizoclonium and established a new species, R. capillare Kützing (1847a:166), to accommodate Mediterranean populations that other workers had associated erroneously (in his opinion) with Dillwyn’s plant. Still later, Kützing (1849:376) transferred R. capillare to Chaetomorpha, unfortunately changing the epithet to tortuosa and thus setting the stage for serious confusion. Two taxonomic synonyms listed by Ardissone (1887 [1886–1887]:209) have equal priority, Conferva ligustica Kützing (1843:259) and Spongopsis mediterranea Kützing (1843b:261), of which the former is herewith chosen to serve as the basionym of the correct name for this species.

Chaetomorpha linum (O.F. Müller) Kützing

Conferva linum O.F. Müller, 1778:7, pl. 771:2 [syntype localities: Nakskov and Rødby, Denmark].

Chaetomorpha linum (O.F. Müller) Kützing, 1845:204.—Sulit, Navarro, and San Juan, 1952:170 (table 5).—Rabanal and Montalban, 1953:142.—Villadolid and Villaluz, 1953:5–10.—Villaluz, 1953:103.—Gilbert, 1961:432.—Rabanal and Montalban, 1961:3.—Taylor, 1966b:344.—Reyes, 1972:140.—Westernhagen, 1973a:64.—Ortega, Alcala, and Reyes, 1974:187.—Westernhagen, 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:134.—Meñez and Calumpong, 1981:382.—Calumpong, 1982:145.—Hurtado-Ponce, 1983:107.—Hurtado-Ponce and Modelo, 1983:146.—Hurtado-Ponce, 1984:179.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Ilocos Sur, Rizal, Manila Bay, Albay. NEGROS: Negros Oriental. CEBU (Mactan I.).

Chaetomorpha spiralis Okamura

Chaetomorpha spiralis Okamura, 1903:131 [type locality: Japan (Nemoto, Boshu Prov. [Chiba Prefecture] fide isotype in UC)].—Meñez and Calumpong, 1981:382.—Calumpong, 1982:145.

Chaetomorpha clavata Kützing var. torta Farlow ex Collins, 1909:323 [type locality: San Diego, California, USA].

Chaetomorpha torta (Farlow ex Collins) Yendo, 1914:264.—Gilbert, 1961:433.—Domantay, 1962:279.—Velasquez, Trono, and Doty, 1975:134.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. NEGROS: Negros Occidental, Negros Oriental. MINDANAO: Zamboanga. BASILAN.

NOTE.—The synonymy was proposed by Abbott (1972:259).

Cladophora Kützing

Cladophora albida (Nees) Kützing

Conferva albida Hudson, 1778:595 [type locality: England].

Annulina albida Nees, 1820: Index [1].

Cladophora albida (Nees) Kützing, 1843b:267.—Domantay, 1962:279.—Velasquez, Trono, and Doty, 1975:135.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—Conferva albida Hudson, the intended basionym of Cladophora albida, is a later homonym of Conferva albida Forsskål (1775:xii) and hence not priorable. Annulina albida Nees is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Cladophora aokii Yamada

Cladophora aokii Yamada, 1925a:85, fig. III [type locality: Byobi-to, Taiwan].—Gilbert, 1961:429.—Taylor, 1966b:345.—Velasquez, Trono, and Doty, 1975:135.—Saraya and Trono, 1980:9, pl. 1: fig. 4.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Bataan.

Cladophora catenata (Linnaeus) Kützing

Conferva catenata Linnaeus, 1753:1166 [syntype localities: “in mari Europam australem et Americam meridionalem”].

Cladophora catenata (Linnaeus) Kützing, 1843b:271.

Cladophora fuliginosa Kützing, 1849:415 [type locality: La Habana, Cuba].—Gilbert, 1961:429.—Meñez, 1961:46, pl. 2: figs. 17–19.—Westernhagen, 1973a:64; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:135.—Cordero, 1977b:22, pl. I: figs. 3, 4.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Pangasinan. CEBU (Mactan I.).

NOTE.—The synonymy was proposed by van den Hoek (1963:123, footnote; 1982:59). Philippine records of this species were cited by van den Hoek (1982:61), but with an expression of uncertainty.

Cladophora conferta P. Crouan and H. Crouan

Cladophora conferta P. Crouan and H. Crouan in Schramm and Mazé, 1865:37 [type locality: Guadeloupe].

Cladophora uncinata Børgesen, 1913:20, figs. 9, 10 [type locality: St. Croix, Virgin Is.].—Chan, 1981:387.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

NOTE.—The synonymy was proposed by van den Hoek (1982:174).

Cladophora crispula Vickers

Cladophora crispula Vickers, 1905:56 [type locality: Barbados].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Cladophora crucigera Grunow

Cladophora crucigera Grunow, 1867:38, footnote [type locality: Guadeloupe].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

NOTE.—Van den Hoek (1982:39) did not see the type of this species, but thinks that the description suggests a robust plant of C. vagabunda whose ultimate branch systems have disappeared by sporulation.

Cladophora crystallina (Roth) Kützing

Conferva crystallina Roth, 1797:196 [type locality: near Wismar, East Germany].

Cladophora crystallina (Roth) Kützing, 1843b:267.—Saraya and Trono, 1980:9.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CENTRAL VISAYAS.

Cladophora cymopoliae Børgesen

Cladophora cymopoliae Børgesen, 1925:69, figs. 27–29 [type locality: Las Palmas, Gran Canaria, Islas Canarias].—Vannajan and Trono, 1977:43.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

Cladophora dalmatica Kützing

Cladophora dalmatica Kützing, 1843b:268 [type locality: Split, Yugoslavia].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Cladophora filipendula

Cladophora filipendula.—Westernhagen, 1973a:64; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

NOTE.—Westernhagen did not give an author for this name and we have been unable to find its place of publication.

Cladophora inserta Dickie

Cladophora inserta Dickie, 1876b:454 [type locality: Honolulu, Hawaiian Is.].—Chan, 1981:387, 389.—Trono and Ang, 1982:3.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. PALAWAN (Bugsuk I.).

Cladophora japonica Yamada

Cladophora japonica Yamada, 1931a:65, pl. XVI [syntype localities: Sagami and Kazusa provinces (Kanagawa and Chiba prefectures), Japan].—Cordero, 1977b:22, pl. 1: fig. 5.

PHILIPPINE DISTRIBUTION.—BATANES.

Cladophora laetevirens (Dillwyn) Kützing

Conferva laetevirens Dillwyn, 1805 [1802–1809]: pl. 48 [“laete virens”] [type locality: Swansea, Wales].

Cladophora laetevirens (Dillwyn) Kützing, 1843b:267.

Conferva heteronema C. Agardh, 1824:114 [type locality: Venezia, Italy].

Cladophora heteronema (C. Agardh) Kützing 1843b:265.—Westernhagen, 1973a:64; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

NOTE.—The synonymy was proposed by van den Hoek (1963:129).

Cladophora liebetruthii Grunow

Cladophora liebetruthii Grunow in piccone, 1884b:53 [type locality: Gran Canaria, Islas Canarias].—Fortes and Trono, 1980:55, fig. 4.—Chan, 1981:387.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro.

Cladophora pellucida (Hudson) Kützing

Conferva pellucida Hudson, 1762:483 [type locality: Walney I., Lancashire, England].—Montagne, 1844a:658 [with query].—Velasquez, Trono, and Doty, 1975:138.

Cladophora pellucida (Hudson) Kützing, 1843b:271.—Martens, 1868:58–59 [with query].—Gilbert, 1946:77; 1961:429.—Domantay, 1962:279.—Velasquez, Trono, and Doty, 1975:135.

Conferva trichotoma C. Agardh, 1824:121 [type locality: Atlantic France].

Cladophora trichotoma (C. Agardh) Kützing, 1843b:271.—Cornejo and Velasquez, 1972:174, pl. 4: fig. 29.—Westernhagen, 1973a:64; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:136.—Trono and Young, 1977:54.—Vannajan and Trono, 1977:40.—Chan, 1981:387.—Fortes, 1981b:396.—Hurtado-Ponce, 1983:108.—Hurtado-Ponce and Modelo, 1983:146.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Rizal, Cavite, Batangas. CATANDUANES. CEBU (Mactan I.).

NOTE.—The synonymy was proposed by van den Hoek (1963:215).

Cladophora prolifera (Roth) Kützing

Conferva prolifera Roth, 1797:182, pl. III: fig. 2 [type locality: “in mare Corsicam”].

Cladophora prolifera (Roth) Kützing, 1843b:271.—Cordero, 1977b:22.—Meñez and Calumpong, 1981:382.

Cladophora rugulosa Martens, 1868:112, pl. II: fig. 3 [lectotype locality: Port Natal (Durban), South Africa fide Papenfuss, 1943a:80].—Marcos-Agngarayngay, 1983:75, fig. 10.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte. CENTRAL VISAYAS.

NOTE.—The synonymy was proposed by van den Hoek (1982:169).

*Cladophora quisumbingii Manza

Cladophora quisumbingii Manza, 1939:109 [type locality: Batan I., Batanes Prov., Luzon].—Taylor, 1966b:345, fig. 1.—Velasquez, Trono, and Doty, 1975:135.—Saraya and Trono, 1980:9, pl. I: fig. 1.

*Cladophora philippinensis Manza, 1939:109 [type locality: Bawa, Cagayan Prov., Luzon].

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Pangasinan.

NOTE.—Manza (1939:109) described Cladophora philippinensis and C. quisumbingii on the basis of specimens in the Bureau of Science, Manila. His publication was overlooked by Taylor, who studied Philippine algae including duplicates of the Bureau of Science collections in the Herbarium of the University of California, Berkeley. Seeing an apparently undescribed Cladophora labeled C. quisumbingii (without authorship) and believing that this name had not been published, Taylor (1966b:345) described it as a new species under that name. Because the holotype of C. quisumbingii Manza was destroyed during World War II, the isotype at Berkeley (UC 1402218) becomes the lectotype. The isotype of C. philippinensis Manza (UC 1402215), which is labeled with that name but without its author, similarly becomes the lectotype of its species. This specimen was included by Taylor in his circumscription of C. quisumbingii.

Cladophora rupestris (Linnaeus) Kützing

Conferva rupestris Linnaeus, 1753:1167 [type locality: “in Europae marinis rupibus copiosissima”].

Cladophora rupestris (Linnaeus) Kützing, 1843b:270.—Cordero, 1976c:8; 1977b:23.—Marcos-Agngarayngay, 1983:75, fig. 11.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte.

Cladophora sakaii Abbott

Cladophora densa Harvey, 1860a:334 [type locality: “Hakodadi Bay” (Hakodate), Japan] [replaced name].—Cordero, 1984a:75.

Cladophora sakaii Abbott, 1972:259.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

NOTE.—Cladophora sakaii is a substitute name for C. densa Harvey, a later homonym of C. densa (Roth) Kützing (1845:209).

Cladophora sericea (Hudson) Kützing

Conferva sericea Hudson, 1762:485 [type locality: Isle of Sheppey, Kent, England].

Cladophora sericea (Hudson) Kützing, 1843b:264.

Conferva rudolphiana C. Agardh, 1827:636 [type locality: Trieste, Italy].

Cladophora rudolphiana (C. Agardh) Kützing, 1843b:268.—Hurtado-Ponce, 1983:107.—Hurtado-Ponce and Modelo, 1983:146.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

NOTE.—The synonymy was proposed by van den Hoek (1963:77).

Cladophora sibogae Reinbold

Cladophora sibogae Reinbold, 1905.146 [syntype localities: Banda Is., Moluccas and Dongola (Donggala), Celebes, Indonesia].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Cladophora vagabunda (Linnaeus) van den Hoek

Conferva vagabunda Linnaeus, 1753:1167 [type locality: “in Mari Europaeo”].

Cladophora vagabunda (Linnaeus) van den Hoek, 1963:144.

Conferva fascicularis Mertens ex C. Agardh, 1824:114 [type locality: West Indies].

Cladophora fascicularis (Mertens ex C. Agardh) Kützing, 1843b:268.—Gilbert, 1961:429.—Reyes, 1972:141.—Ortega, Alcala, and Reyes, 1974:187.—Velasquez, Trono, and Doty, 1975:135.—Reyes, 1978:147, pl. 4: figs. 1, 2.

Cladophora mauritiana Kützing, 1849:399 [type locality: Mauritius].—Dickie, 1876a:244.—Gilbert, 1946:77; 1961:429 [with query].—Velasquez, Trono, and Doty, 1975:135.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Manila, NEGROS: Negros Oriental, SIQUIJOR. MINDANAO: Zamboanga. BASILAN.

NOTE.—The synonymy was proposed by van den Hoek (1982:137).

Cladophora wrightiana Harvey

Cladophora wrightiana Harvey, 1860a:333 [type locality: Shimoda, Japan].—Hurtado-Ponce, 1983:108.—Hurtado-Ponce and Modelo, 1983:146.—Cordero, 1984a:75.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Rhizoclonium Kützing

Rhizoclonium africanum Kützing

Rhizoclonium africanum Kützing, 1853:21, pl. 67: fig. II [type locality: “Senegambien” (Senegal or Gambia)].

Rhizoclonium hookeri [misapplied name fide Womersley and Bailey, 1970:265].—Taylor, 1966b:334.—Westernhagen, 1973a:64; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:161.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Bataan. CENTRAL VISAYAS. CEBU (Mactan I.).

Rhizoclonium crassipellitum W. West and G.S. West var. robustum G.S. West

Rhizoclonium crassipellitum W. West and G.S. West var. robustum G.S. West, 1904:283 [type locality: Barbados].—Taylor, 1966b:344.—Velasquez, Trono, and Doty, 1975:161 [without designation of variety].

PHILIPPINE DISTRIBUTION.—LUZON: Rizal.

Rhizoclonium grande Børgesen

Rhizoclonium grande Børgesen, 1935:14, figs. 5, 6 [type locality: Bombay, India].–Fortes and Trono, 1980:56.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Rhizoclonium kerneri Stockmayer

Rhizoclonium kerneri Stockmayer, 1890:582 [syntype localities: France; Scandinavia; North America].—Gilbert, 1961:430.—Westernhagen, 1973a:64; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:161.—Reyes, 1978:149, pl. 5: figs. 4, 5.—Carumbana and Luchavez, 1980:189.

PHILIPPINE DISTRIBUTION.—LUZON: Laguna. NEGROS: Negros Oriental, CEBU (Mactan I.). SIQUIJOR.

Rhizoclonium kochianum Kützing

Rhizoclonium kochianum Kützing, 1845:206 [syntype localities: North Sea, West Germany; Dubrovnik, Yugoslavia].—Westernhagen, 1973a:64; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Rhizoclonium riparium (Roth) Harvey

Conferva riparia Roth, 1806:216 [type locality: Norderney I., West Germany].

Rhizoclonium riparium (Roth) Harvey, 1849 [1847–1851]: pl. CCXXXVIII.—Westernhagen, 1973a:64; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Rhizoclonium setaceum Kützing

Rhizoclonium setaceum Kützing, 1847a:166 [type locality: Goes, Netherlands].—Gilbert, 1961:430.—Velasquez, Trono, and Doty, 1975:161.

PHILIPPINE DISTRIBUTION.—LUZON: Bataan.

Order SIPHONOCLADALES

Family SIPHONOCLADACEAE

Boergesenia J. Feldmann

Boergesenia forbesii (Harvey) J. Feldmann

Valonia forbesii Harvey, 1860a:333 [syntype localities: Ryukyu-retto, Japan; Sri Lanka].

Boergesenia forbesii (Harvey) J. Feldmann, 1938:1503.—Gilbert, 1961:420.—Meñez, 1961:49, pl. 1: fig. 13.—Taylor, 1966b:347.—Villones and Magdamo, 1968:12, fig. 5.—Reyes, 1972:139.—Trono 1972a:95; 1973b:216; 1973c:7, fig. 14.—Velasquez et al., 1973:13, pl. 4: fig. 17.—Trono, 1975:31.—Velasquez, Trono, and Doty 1975:129.—Cordero, 1976c:8, 9, 10; 1977b:26.—Reyes, 1978:152, pl. 6: fig. 14.—Trono, 1978:3.—Trono and Tuason, 1978:3.—Cordero 1979b:276.—Calumpong, 1980:143 (table 3).—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:12.—Trono and Ganzon-Fortes, 1980:9, fig. [s.n].—Chan, 1981:387.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:3, pl. II: fig. 1.—Hurtado-Ponce, 1983:111.—Hurtado-Ponce and Modelo, 1983:147.—Marcos-Agngarayngay, 1983:76, fig. 13.—Cordero, 1984a:76.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Ilocos Norte, Pangasinan, Bataan, Batangas, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. PANAY: Aklan. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.). BASILAN. PALAWAN. SULU: Sulu (Siasi I.), Tawitawi.

Boodlea Murray and De Toni

Boodlea coacta (Dickie) Murray and De Toni

Cladophora coacta Dickie, 1876b:451 [type locality: “Osima Harbour” (O-shima, Wakayama Prefecture), Japan].

Boodlea coacta (Dickie) Murray and De Toni in Murray, 1889:245.—Cordero, 1984a:73.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Boodlea composita (Harvey) Brand

Conferva composita Harvey, 1834a:157 [type locality: Mauritius].

Boodlea composita (Harvey) Brand, 1904:187.—Gilbert, 1961:421.—Meñez, 1961:48, pl. 3: figs. 40–42.—Reyes, 1972:139.—Trono, 1972a:95.—Cordero, 1973b:17.—Trono, 1973c:6, fig. 18.—Ortega, Alcala, and Reyes, 1974:186, 187.—Trono, 1975:30.—Velasquez, Trono, and Doty, 1975:129.—Cordero, 1976c:10; 1977b:28, pl. II: fig. 9.—Reyes, 1978:153, pl. 7: fig. 1.—Trono, 1978:4.—Trono and Tuason, 1978:2.—Puig and Cordero, 1979:20.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:13.—Trono and Ganzon-Fortes, 1980:15, fig. [s.n].—Chan, 1981:387.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:4.—Trono and Ang, 1982:4.—Marcos-Agngarayngay, 1983:80, fig. 15.—Cordero, 1984a:73.

Boodlea siamensis Reinbold, 1901:191 [type locality: Ko Kahdat, Ko Chang Archipelago, Thailand].—Saraya and Trono, 1980:13.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Ilocos Sur, Pangasinan, Batangas. Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. LEYTE (Biliran I.). NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga. BASILAN. PALAWAN (incl. Bugsuk I., Culion I.). SULU: Sulu (between Lapac I. and Siasi I.), Tawitawi.

NOTE.—Boodlea siamensis is included as a synonym on the authority of Børgesen (1946:16).

Boodlea montagnei (Harvey ex J.E. Gray) Egerod

Microdictyon montagnei Harvey ex J.E. Gray, 1866:69 [type locality: Tonga].—Gilbert, 1961:422, fig. 1.—Taylor, 1966b:349.—Velasquez, Trono, and Doty, 1975:153.

Boodlea montagnei (Harvey ex J.E. Gray) Egerod, 1952:332, footnote.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga. BASILAN. SULU: Sulu (Siasi I.), Tawitawi.

Boodlea struveoides Howe

Boodlea struveoides Howe, 1918:496 [type locality: Harrington Sound, Bermuda].—Saraya and Trono, 1980:12.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CENTRAL VISAYAS.

Chamaedoris Montagne

Chamaedoris orientalis Okamura and Higashi

Chamaedoris orientalis Okamura and Higashi in Okamura, 1931:98, pl. 10 [type locality: Kotosho (Hung-t’ou), Taiwan].—Gilbert and Doty, 1969:125, fig. 15.—Velasquez, Trono, and Doty, 1975:134.—Cordero, 1976c:8; 1977b:27, pl. II: figs. 10, 11.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Quezon.

Cladophoropsis Børgesen

Cladophoropsis dichotoma (Zanardini) Papenfuss

Spongodendron dichotomum Zanardini, 1878:38 [type locality: Wokam I., Aru Is., Indonesia].

Spongocladia dichotoma (Zanardini) Murray and Boodle, 1888:175.—Taylor, 1966b:348.—Velasquez, Trono, and Doty, 1975:163.—Meñez and Calumpong, 1984:105.

Cladophoropsis dichotoma (Zanardini) Papenfuss, 1958:104.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. PALAWAN.

Cladophoropsis fasciculata (Kjellman) Wille

Siphonocladus fasciculatus Kjellman, 1897b:36, pl. 7: figs. 10–17 [type locality: Yokohama, Japan].

Cladophoropsis fasciculata (Kjellman) Wille, 1910 [1909–1910]:116.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Cladophoropsis gracillima Dawson

Cladophoropsis gracillima Dawson, 1950b:149, figs. 12, 13 [type locality: Punta Palmilla, Baja California Sur, Mexico].—Trono, 1973c:6.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. CENTRAL VISAYAS.

Cladophoropsis membranacea (C. Agardh) Børgesen

Conferva membranacea C. Agardh, 1824:120 [type locality: “Ad insulam S. Crucis” (St. Croix, Virgin Is.)].

Cladophoropsis membranacea (C. Agardh) Børgesen, 1905:285, figs. 8–13.—Cordero, 1976c:10; 1977b:28.—Vannajan and Trono, 1977:43.—Reyes, 1978:151, pl. 6: figs. 5, 6.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Manila, SIQUIJOR.

Cladophoropsis neocaledonica

(Grunow ex Murray and Boodle) Papenfuss

Spongocladia neocaledonica Grunow ex Murray and Boodle, 1888:175 [type locality: Poro, New Caledonia].—Trono, 1973d:5, pl. 1: fig. 1.—Saraya and Trono, 1980:13, pl. II: fig. 1.

Cladophoropsis neocaledonica (Grunow ex Murray and Boodle) Papenfuss, 1958:104.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

*Cladophoropsis philippinensis W.R. Taylor

Cladophoropsis philippinensis W.R. Taylor, 1961:58, figs. 1–6 [type locality: Little Santa Cruz I., near Zamboanga (city), Mindanao].—Taylor, 1966b:348.—Velasquez, Trono, and Doty, 1975:136.—Taylor, 1977b:9.

PHILIPPINE DISTRIBUTION.—MINDANAO: Zamboanga del Sur. BASILAN.

Cladophoropsis sundanensis Reinbold

Cladophoropsis sundanensis Reinbold, 1905:147 [syntype localities: “Timor, Laut, etc.”, Indonesia].—Weber-van Bosse, 1913a:77.—Gilbert, 1946:77; 1961:424.—Velasquez, Trono, and Doty, 1975:136.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I.).

Cladophoropsis vaucheriaeformis (J.E. Areschoug) Papenfuss

Spongocladia vaucheriaeformis J.E. Areschoug, 1854:202, pl. II [type locality: Mauritius].—Gilbert, 1961:424.—Taylor, 1966b:348.—Velasquez, Trono, and Doty, 1975:163.—Saraya and Trono, 1980:14.

Cladophoropsis vaucheriaeformis (J.E. Areschoug) Papenfuss, 1958:104.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro. SULU: Tawitawi.

Cladophoropsis zollingeri (Kützing) Reinbold

Cladophora zollingeri Kützing, 1849:415 [type locality: Java, Indonesia].

Cladophoropsis zollingeri (Kützing) Reinbold, 1905:147.—Hurtado-Ponce, 1983:110.—Hurtado-Ponce and Modelo, 1983:147.—Cordero, 1984a:73.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Struvea Sonder

Struvea anastomosans (Harvey) Piccone and Grunow ex Piccone

Cladophora? anastomosans Harvey, 1859:pl. CI [type locality: Fremantle, Western Australia, Australia].

Struvea anastomosans (Harvey) Piccone and Grunow ex Piccone, 1884b:20.—Cordero, 1977b:30.—Reyes, 1978:153, pl. 7: fig. 2.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:13, pl. 1: fig. 3. Chan, 1981:387, 389.

Struvea delicatula Kützing, 1866:1, pl. 2: figs, e–g [type locality: New Caledonia].—Gilbert, 1961:421.—Velasquez, Trono, and Doty, 1975:164.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. CEBU (Mactan I.), SIQUIJOR. MINDANAO: Zamboanga. BASILAN.

NOTE.—The conspecificity of Struvea delicatula and S. anastomosans was proposed by Murray and Boodle (1888:281), who, however, erroneously adopted the later name.

Struvea ramosa Dickie

Struvea ramosa Dickie, 1874b:316 [type locality: Bermuda].—Westernhagen, 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Family VALONIACEAE

Dictyosphaeria Decaisne ex Endlicher

Dictyosphaeria cavernosa (Forsskål) Børgesen

Ulva cavernosa Forsskål, 1775:187 [syntype localities: “Gomfodae” (Al-Qunfidha), Saudi Arabia; Mokha, Yemen].

Dictyosphaeria cavernosa (Forsskål) Børgesen, 1932:2, pl. I: fig. 1.—Gilbert, 1946:77; 1961:417.—Meñez, 1961:48.—Domantay, 1962:280.—Taylor, 1966b:348.—Velasquez, 1971:432, fig. 11.—Reyes, 1972:139.—Trono, 1972a:95.—Cordero, 1973b:17.—Trono, 1973b:216; 1973c:7, fig. 13; 1973d:5, pl. 5: fig. 18.—Velasquez et al., 1973:12, pl. 3: fig. 11.—Ortega, Alcala, and Reyes, 1974:185, 186, 187, 188.—Trono, 1975:31.—Velasquez, Trono, and Doty, 1975:140.—Cordero, 1976c:8, 10, 11; 1977b:27.—Trono and Young, 1977:54.—Reyes, 1978:151, pl. 6: fig. 9.—Trono, 1978:2.—Trono and Tuason, 1978:2.—Cordero, 1979b:276, 279.—Puig and Cordero, 1979:21.—Cordero, 1980b:22, pl. 7.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:10.—Chan, 1981:387.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:3, pl. II: fig. 2.—Cordero, 1982a:61.—Trono and Ang, 1982:3.—Hurtado-Ponce, 1983:110.—Hurtado-Ponce and Modelo, 1983:147.—Marcos-Agngarayngay, 1983:79, fig. 14.—Cordero, 1984a:72; 1984b:57.—Marcos-Agngarayngay, 1984b:121.

Valonia favulosa C. Agardh, 1822a:432 [type locality: “Ravak” (Lawak), Waigeo I., Moluccas, Indonesia].

Dictyosphaeria favulosa (C. Agardh) Decaisne ex Endlicher, 1843:18.—Dickie, 1876a:244.—Velasquez, Trono, and Doty, 1975:141.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Pangasinan, Bataan, Batangas, Quezon, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. SAMAR: Eastern Samar. LEYTE (Biliran I.). PANAY: Aklan. GUIMARAS. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.), Surigao del Sur, Davao. BASILAN. PALAWAN (incl. Balabac I., Bugsuk I.). SULU: Sulu (Siasi I.), Tawitawi (incl. Sibutu I., Turtle Is.).

NOTE.—The synonymy was proposed by Børgesen (1932:2).

Dictyosphaeria cavernosa (Forsskål) Børgesen var. bullata Børgesen

Dictyosphaeria cavernosa (Forsskål) Børgesen var. bullata Børgesen, 1952:5, figs. 1, 2 [type locality: Riambel, Mauritius].—Cordero, 1977b:28.

PHILIPPINE DISTRIBUTION.—BATANES.

Dictyosphaeria intermedia Weber-van Bosse

Dictyosphaeria intermedia Weber-van Bosse, 1905:143 [type locality: Jangkar, Java, Indonesia].—Meñez, 1961:49.—Taylor, 1966b:348.—Velasquez, Trono, and Doty, 1975:141.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Pangasinan.

Dictyosphaeria ocellata (Howe) Olsen-Stojkovich

Valonia ocellata Howe, 1920:603 [type locality: Watling’s I., Bahama Is.].—Cordero, 1977b:25.

Dictyosphaeria ocellata (Howe) Olsen-Stojkovich, 1985b:62.

PHILIPPINE DISTRIBUTION.—BATANES.

NOTE.—Taylor (1960:111) considered the structure of this species to be intermediate between Valonia and Dictyosphaeria, “with more resemblance to solid forms of the latter genus.” Olsen-Stojkovich assigned it to Dictyosphaeria on the basis of immunological distance data.

Dictyosphaeria versluysii Weber-van Bosse

Dictyosphaeria versluysii Weber-van Bosse, 1905:144 [“versluysi”] [syntype localities: various, all in Indonesia].—Domantay, 1962:280.—Trono, 1973b:216; 1973c:8.—Velasquez, Trono, and Doty, 1975:141.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:11, pl. 1: fig. 2.

Dictyosphaeria vanbosseae Børgesen, 1912:256, figs. 7–9 [“van Bosseae”] [type locality: Cane Bay, St. Croix, Virgin Is.].—Velasquez et al., 1973:12, pl. 3: fig. 12.

Dictyosphaeria setchellii Børgesen, 1940:12, figs. 1, 2a, 3a [type locality: Mauritius].—Gilbert, 1961:418.—Velasquez, Trono, and Doty, 1975:141.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Bataan, Batangas. MINDORO: Oriental Mindoro. CEBU (Mactan I.). BASILAN. SULU: Sulu (Siasi I.).

NOTE.—The synonymy was proposed by Valet (1966).

Ernodesmis Børgesen

Ernodesmis verticillata (Kützing) Børgesen

Valonia verticillata Kützing, 1847a:165 [type locality: St. Croix, Virgin Is.].

Ernodesmis verticillata (Kützing) Børgesen, 1912:259, figs. 10–12.—Reyes, 1978:151, pl. 6: figs. 7, 8.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Valonia C. Agardh

Valonia aegagropila C. Agardh

Valonia aegagropila C. Agardh, 1822a:429 [lectotype locality: Venezia, Italy fide Egerod, 1952:348].—Gilbert, 1961:418.—Taylor, 1966b:347.—Reyes, 1972:139.—Trono, 1972a:95; 1973b:215; 1973c:8, fig. 16; 1973d:4, pl. 1: fig. 2.—Velasquez et al., 1973:11, pl. 3: fig. 10.—Ortega, Alcala, and Reyes, 1974:186, 188.—Trono, 1975:30.—Velasquez, Trono, and Doty, 1975:168.—Taylor, 1977b:8.—Reyes, 1978:152, pl. 6: fig. 13.—Trono, 1978:3.—Trono and Tuason, 1978:3.—Cordero, 1979b:276.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:11.—Trono and Ganzon-Fortes, 1980:11, fig. [s.n].—Chan, 1981:387.—Guzman, 1981:42, 45.—Meñez and Calumpong, 1981:382.—Trono and De Lara, 1981:4, pl. II: fig. 4.—Hurtado-Ponce, 1983:111.—Hurtado-Ponce and Modelo, 1983:147.—Cordero, 1984a:72.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Bataan, Batangas, Sorsogon. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. PANAY: Aklan. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.). BASILAN. PALAWAN, SULU: Sulu (Siasi I.), Tawitawi.

Valonia confervoides Harvey ex J. Agardh

Valonia confervoides Harvey ex J. Agardh, 1887:100 [syntype localities: Sri Lanka; Tonga].—Domantay, 1962:281.—Velasquez, Trono, and Doty, 1975:168.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Valonia fastigiata Harvey ex J. Agardh

Valonia fastigiata Harvey ex J. Agardh, 1887:101, pl. I: fig. 5 [syntype localities: Sri Lanka; Tonga].—Dickie, 1876a:245.—Gilbert, 1946:77; 1961:419.—Taylor, 1966b:347.—Velasquez, Trono, and Doty, 1975:168.—Saraya and Trono, 1980:11.—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro. CENTRAL VISAYAS. MINDANAO: Zamboanga. BASILAN. SULU: Sulu (Siasi I.), Tawitawi (Sibutu I.).

Valonia macrophysa Kützing

Valonia macrophysa Kützing, 1843b:307 [type locality: Hvar I., Yugoslavia].—Martens, 1868:25, 66–67.—Gilbert, 1946:77; 1961:420.—Velasquez, Trono, and Doty, 1975:168.—Meñez and Calumpong, 1981:382.—Cordero, 1984a:72.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, CENTRAL VISAYAS. MINDANAO: Zamboanga.

Valonia utricularis (Roth) C. Agardh

Conferva utricularis Roth, 1797:160, pl. I: fig. 1 [type locality: Mediterranean Sea].

Valonia utricularis (Roth) C. Agardh, 1822a:431.—Martens, 1868:25, 66–67.—Dickie, 1876a:244.—Weber-van Bosse, 1913a:60.—Gilbert, 1947:77; 1961:420.—Domantay, 1962:282.—Velasquez, Trono, and Doty, 1975:168.—Cordero, 1976c:8, 9, 10; 1977b:25.—Saraya and Trono, 1980:12.—Meñez and Calumpong, 1981:382.—Cordero, 1984a:72.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan. CENTRAL VISAYAS. MINDANAO: Zamboanga.

Valonia ventricosa J. Agardh

Valonia ventricosa J. Agardh, 1887:96 [syntype localities: St. Croix, Virgin Is.; Guadeloupe].—Meñez, 1961:48.—Domantay, 1962:282.—Taylor, 1966b:347.—Reyes, 1972:139.—Trono, 1972a:95.—Cordero, 1973b:17.—Trono, 1973c:8.—Ortega, Alcala, and Reyes, 1974:185, 186, 187, 188.—Trono, 1975:30.—Velasquez, Trono, and Doty, 1975:168.—Reyes, 1978:152, pl. 6: fig. 10.—Trono, 1978:3.—Trono and Tuason, 1978:3.—Puig and Cordero, 1979:20.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:11.—Trono and Ganzon-Fortes, 1980:13, fig. [s.n].—Meñez and Calumpong, 1981:382.—Hurtado-Ponce, 1983:111.—Hurtado-Ponce and Modelo, 1983:147.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Batangas, Sorsogon. MINDORO: Oriental Mindoro. MARINDUQUE. LEYTE (Biliran I.). NEGROS: Negros Oriental. CEBU (incl. Mactan I.). SIQUIJOR. SULU: Tawitawi (incl. Sibutu I.).

Valoniopsis Børgesen

Valoniopsis pachynema (Martens) Børgesen

Bryopsis pachynema Martens, 1868:24, pl. IV: fig. 2 [syntype localities: Bengkulu and Pulau Tikus, Sumatra, Indonesia].

Valoniopsis pachynema (Martens) Børgesen, 1934:10, figs. 1, 2.—Gilbert, 1961:428.—Taylor, 1961:58; 1966b:347.—Westernhagen, 1973a:64; 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:168.—Cordero, 1977b:26.—Meñez and Calumpong, 1981:382.—Cordero, 1984a:74.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, La Union. MINDORO: Oriental Mindoro. CENTRAL VISAYAS. CEBU (Mactan I.). MINDANAO: Zamboanga. BASILAN.

Order BRYOPSIDALES

Family BRYOPSIDACEAE

Bryopsis Lamouroux

Bryopsis corticulans Setchell

Bryopsis corticulans Setchell in Collins, Holden, and Setchell, 1899: no. 626 [syntype localities: Carmel Bay and Pacific Grove, California, USA].—Westernhagen, 1973a:64; 1974:112 (table 1).

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Bryopsis indica A. Gepp and E.S. Gepp

Bryopsis indica A. Gepp and E.S. Gepp, 1908:169, pl. 22: figs. 10, 11 [syntype localities: various, in Indian Ocean].—Gilbert, 1961:434.—Velasquez, Trono, and Doty, 1975:130.—Cordero, 1977b:31.—Vannajan and Trono, 1977:45, fig. 7.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cavite. MINDORO: Oriental Mindoro. CEBU (Mactan I.).

Bryopsis pennata Lamouroux

Bryopsis pennata Lamouroux, 1809b:333 [type locality: Antilles].—Trono, 1975:32.—Reyes, 1978:153, pl. 7: fig. 3.—Liao and Sotto, 1980:96.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon. CEBU (Mactan I.). SIQUIJOR.

Bryopsis pennata Lamouroux var. secunda (Harvey) Collins and Hervey

Bryopsis plumosa (Hudson) C. Agardh var. secunda Harvey, 1858:31, pl. XLV:A: figs. 1–3 [syntype localities: Key West and Sand Key, Florida, USA].

Bryopsis pennata Lamouroux var. secunda (Harvey) Collins et Hervey, 1917:62.—Gilbert, 1961:434.—Velasquez, Trono, and Doty, 1975:130 [without designation of variety].

PHILIPPINE DISTRIBUTION.—BASILAN.

Bryopsis plumosa (Hudson) C. Agardh

Ulva plumosa Hudson, 1778:571 [type locality: Exmouth, Devonshire, England].

Bryopsis plumosa (Hudson) C. Agardh, 1822a:448.—Velasquez et al., 1973:13, pl. 4: fig. 16.—Vannajan and Trono, 1977:44, figs. 5, 6.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:14, pl. II: fig. 4.—Meñez and Calumpong, 1981:383.—Cordero, 1984a:76.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Rizal, Manila, Cavite, Batangas. CEBU (Mactan I.).

Derbesia Solier

Derbesia attenuata Funk

Derbesia attenuata Funk, 1955:23 [type locality: Napoli, Italy].—Chan, 1981:387, 389.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Derbesia marina (Lyngbye) Solier

Vaucheria marina Lyngbye, 1819:79, pl. 22:A [sporophyte] [type locality: Kvivig, Strømø, Faeroes].

Derbesia marina (Lyngbye) Solier, 1846:453.

Gastridium ovale Lyngbye, 1819:72, pl. 18:B [gametophyte] [syntype localities: various, all in Faeroes].

Halicystis ovalis (Lyngbye) J.E. Areschoug, 1850:447.—Domantay, 1962:281.—Velasquez, Trono, and Doty, 1975:147.—Reyes, 1978:152.—Saraya and Trono, 1980:14, pl. II: fig. 2.

PHILIPPINE DISTRIBUTION (of gametophyte).—LUZON: Pangasinan. SIQUIJOR.

NOTE.—Kornmann (1938) was the first to show that Halicystis ovalis and Derbesia marina are alternating stages. The sporophyte has not yet been reported from the Philippines.

Derbesia tenuissima (Moris and De Notaris) P. Crouan and H. Crouan

Bryopsis tenuissima Moris and De Notaris, 1839:259, pl. VI: fig. III [type locality: Cabrera, Islas Baleares, Spain].

Derbesia tenuissima (Moris and De Notaris) P. Crouan and H. Crouan, 1867:133.—Cordero, 1976c:9; 1977b:31.

PHILIPPINE DISTRIBUTION.—BATANES.

NOTE.—J. Feldmann (1950) showed that Derbesia tenuissima alternates with Halicystis parvula Schmitz ex G. Murray (1893:50, pl. XIII: fig. 5). The gametophyte has not yet been reported from the Philippines.

Pedobesia MacRaild and Womersley

Pedobesia ryukyuensis (Yamada and Tanaka) Kobara and Chihara

Derbesia ryukyuensis Yamada and Tanaka, 1938:64, fig. 5 [type locality: Pinai, Yonakuni-shima, Sakishima-gunto, Ryukyu-retto, Japan].—Cordero, 1976c:9; 1977b:31.

Pedobesia ryukyuensis (Yamada and Tanaka) Kobara and Chihara, 1984:156.

PHILIPPINE DISTRIBUTION.—BATANES.

Family CAULERPACEAE

Caulerpa Lamouroux

Caulerpa ambigua Okamura

Caulerpa ambigua Okamura, 1897:4, pl. I: figs. 3–12 [type locality: Ogasawara-gunto (Bonin Is.) Japan].—Saraya and Trono, 1980:15.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Caulerpa arenicola W.R. Taylor

Caulerpa arenicola W.R. Taylor, 1950:55, pl. 28: fig. 2 [type locality: Rongelap Atoll, Marshall Is.].—Meñez and Calumpong, 1981:383; 1982:5, pl. 1:H.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental. SIQUIJOR.

Caulerpa brachypus Harvey

Caulerpa brachypus Harvey, 1860a:333 [type locality: Tanega-shima, Osumi-gunto, Japan].—Gilbert, 1961:435.—Domantay, 1962:279.—Taylor, 1966b:350.—Kraft, 1972:328.—Trono, 1972a:95; 1973b:217; 1973d:6, pl. 3: fig. 9.—Cordero, 1974b:6, fig. 1.—Velasquez, Trono, and Doty, 1975:130.—Cordero, 1977c:25, 27, fig. 2.—Taylor, 1977a:4.—Reyes, 1978:153, pl. 7: fig. 4.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:17.—Meñez and Calumpong, 1981:383; 1982:5, pl. 1:I.—Trono and Ang, 1982:4.—Cordero, 1984c:51.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MASBATE. SAMAR: Eastern Samar. PANAY: Antique (Panagatan Cays), NEGROS: Negros Occidental, Negros Oriental, CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga, Surigao del Norte, BASILAN. PALAWAN Bugsuk I., Cuyo I.). SULU: Sulu (incl. Siasi I.), Tawitawi (incl. Sibutu I., Tumindao I.).

Caulerpa brachypus Harvey f. parvifolia (Harvey) Cribb

Caulerpa parvifolia Harvey, 1860b: pl. CLXXII [type locality: Kiama, New South Wales, Australia].—Gilbert, 1942:11; 1961:437.—Velasquez, Trono, and Doty, 1975:131.

Caulerpa brachypus Harvey f. parvifolia (Harvey) Cribb, 1958:209.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.).

Caulerpa cupressoides (Vahl) C. Agardh

Fucus cupressoides Vahl, 1802:38 [type locality: St. Croix, Virgin Is.].

Caulerpa cupressoides (Vahl) C. Agardh, 1817:XXIII.—Domantay, 1962:278.—de los Reyes, 1967:231.—Villones and Magdamo, 1968:24, fig. 8.—Reyes, 1972:142.—Trono, 1972a:96.—Cordero, 1973b:19.—Trono, 1973c:9, fig. 22; 1973d:5, pl. 3: fig. 12; 1975:32.—Velasquez, Trono, and Doty, 1975:130.—Taylor, 1977a:4.—Reyes, 1978:155, pl. 8, fig. 7.—Trono, 1978:4.—Trono and Tuason, 1978:4.—Puig and Cordero, 1979:23.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:17.—Meñez and Calumpong, 1982:6, pl. 1:B,C.—Trono and Ang, 1982:4.—Marcos-Agngarayngay, 1983:83, fig. 16.—Cordero, 1984a:76.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Quezon, Sorsogon. CATANDUANES. MINDORO: Oriental Mindoro. MARINDUQUE. SAMAR: Eastern Samar. LEYTE (Biliran I.). NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga. PALAWAN (Bugsuk I.). SULU: Sulu (incl. Siasi I.), Tawitawi.

Caulerpa cupressoides (Vahl) C. Agardh var. ericifolia (Turner) Weber-van Bosse

Fucus ericifolius Turner, 1808:124, pl. 56 [type locality: Bermuda].

Caulerpa cupressoides (Vahl) C. Agardh var. ericifolia (Turner) Weber-van Bosse, 1898:335.—Domantay, 1962:278.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Caulerpa cupressoides (Vahl) C. Agardh var. lycopodium Weber-van Bosse

Caulerpa lycopodium J. Agardh, 1847:6 [syntype localities: Brazil; West Indies].

Caulerpa cupressoides (Vahl) C. Agardh var. lycopodium Weber-van Bosse, 1898:335.—Hurtado-Ponce, 1983:115.—Hurtado-Ponce and Modelo, 1983:148.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

NOTE.—Caulerpa lycopodium J. Agardh, the intended basionym of C. cupressoides var. lycopodium, is a later homonym of C. lycopodium C. Agardh (1817:XXIII) and hence not priorable. The Weber-van Bosse trinomial is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Caulerpa cupressoides (Vahl) C. Agardh var. lycopodium Weber-van Bosse f. amicorum (Harvey) Weber-van Bosse

Caulerpa amicorum Harvey, 1860a:333 [type locality: O-shima, Amami-gunto, Ryukyu-retto, Japan].

Caulerpa cupressoides (Vahl) C. Agardh var. lycopodium Weber-van Bosse f. amicorum (Harvey) Weber-van Bosse, 1898:337.—Gilbert, 1942:17.—Trono, 1973c:9.—Meñez and Calumpong, 1981:383.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. CENTRAL VISAYAS.

Caulerpa cupressoides (Vahl) C. Agardh var. lycopodium Weber-van Bosse f. disticha Weber-van Bosse

Caulerpa cupressoides (Vahl) C. Agardh var. lycopodium Weber-van Bosse f. disticha Weber-van Bosse, 1898:338, pl. XXVII: fig. 14 [syntype localities: Guadeloupe; Florida, USA].

Caulerpa cupressoides (Vahl) C. Agardh var. disticha Weber-van Bosse, 1898:327, 328.—Gilbert, 1961:436.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. MINDANAO: Zamboanga. BASILAN.

NOTE.—Caulerpa cupressoides var. lycopodium f. disticha and C. cupressoides var. disticha are alternative names for the same taxon, both being used by Weber-van Bosse in the original publication.

Caulerpa cupressoides (Vahl) C. Agardh var. lycopodium Weber-van Bosse f. elegans (P. Crouan and H. Crouan) Weber-van Bosse

Caulerpa plumaris (Forsskål) C. Agardh var. elegans P. Crouan and H. Crouan in Schramm and Mazé, 1865:39 [type locality: Guadeloupe].

Caulerpa cupressoides (Vahl) C. Agardh var. lycopodium Weber-van Bosse f. elegans (P. Crouan and H. Crouan) Weber-van Bosse, 1898:336.—Cordero, 1977b:32.

PHILIPPINE DISTRIBUTION.—BATANES.

Caulerpa elongata Weber-van Bosse

Caulerpa elongata Weber-van Bosse, 1898:271, pl. XXI: figs. 5, 6 [syntype localities: Tongatapu, Tonga; Macassar, Celebes, Indonesia].—Taylor, 1977a:5.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon.

Caulerpa fastigiata Montagne

Caulerpa fastigiata Montagne, 1837:353 [type locality: Cuba].—Gilbert, 1942:9; 1961:436.—Velasquez, Trono, and Doty, 1975:131.—Taylor, 1977a:5.—Reyes, 1978:154, pl. 8: figs. 1, 2.—Meñez and Calumpong, 1981:383; 1982:6, pl. 1:A.

PHILIPPINE DISTRIBUTION.—MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SAMAR: Eastern Samar. NEGROS: Negros Occidental (incl. Suyac I.), Negros Oriental (incl. Apo I.), SIQUIJOR. MINDANAO: Zamboanga. BASILAN.

Caulerpa fergusonii Murray

Caulerpa fergusonii Murray, 1891:212, pl. 53: figs. 1, 2 [type locality: Sri Lanka].—Taylor, 1977a:5.—Cordero, 1980b:23, pls. 8, [46].—Hurtado-Ponce, 1983:115.—Hurtado-Ponce and Modelo, 1983:148.—Hurtado-Ponce, 1984:179.—Tungpalan, 1984:139.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Sorsogon. SIQUIJOR.

Caulerpa filicoides Yamada

Caulerpa filicoides Yamada, 1936b:135, pl. 30: fig. 2 [syntype localities: Mikako and Naha, Ryukyu-retto, Japan].—Saraya and Trono, 1980:15, pl. III: figs. 1,4.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Caulerpa lentillifera J. Agardh

Caulerpa lentillifera J. Agardh, 1837:173 [type locality: Ethiopia].—Gilbert, 1942:23; 1961:436.—Domantay, 1962:277.—Taylor, 1966b:350 [var. lentillifera].—Villones and Magdamo, 1968:24, fig. 7.—Trono 1972a:96; 1973b:217; 1973c:10, fig. 1; 1973d:5, pl. 5: fig. 20; 1975:33.—Velasquez, Trono, and Doty, 1975:131.—Taylor, 1977a:5.—Reyes, 1978:155, 162, pl. 8: fig. 6.—Trono, 1978:4.—Trono and Tuason, 1978:3.—Cordero, 1979b:276.—Fortes, 1979:112, figs. 1, 2.—Liao and Sotto, 1980:96.—Moreland, 1980:43.—Saraya and Trono, 1980:16.—Trono and Fortes, 1980:63.—Trono and Ganzon-Fortes, 1980:17, fig. [s.n].—Ganzon-Fortes, 1981:21.—Meñez and Calumpong, 1981:383.—Trono, 1981a:49.—Calumpong, 1982:145.—Luistro, Cajipe, and Laserna, 1982:46.—Meñez and Calumpong, 1982:7, pl. 1:F,G.—Trono and Fortes, 1982:144.—Trono and Ganzon-Fortes, 1985:64, 66.

Caulerpa lentillifera J. Agardh var. longistipitata Weber-van Bosse in Reinbold, 1901:189 [syntype localities: Ko Chick, Ko Chang Archipelago, Thailand; various in Indonesia].—Taylor, 1966b:350.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Batangas, Camarines Norte, Sorsogon. CATANDUANES. MINDORO: Oriental Mindoro. MARINDUQUE. SAMAR: Eastern Samar. PANAY: Aklan. NEGROS: Negros Oriental. CEBU (incl. Mactan I.). SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.), Misamis Occidental, Davao. BASILAN. PALAWAN. SULU: Sulu (incl. Siasi I.), Tawitawi (incl. Sibutu I., Turtle Is.).

NOTE.—The synonymy was proposed by Nizamuddin (1967:158).

*Caulerpa lentillifera J. Agardh var. compacta Trono and Ang

Caulerpa lentillifera J. Agardh var. compacta Trono and Ang, 1982:7, figs. 1–4 [type locality: Bugsuk I., Palawan Prov.].

PHILIPPINE DISTRIBUTION.—As above.

Caulerpa lessonii Bory de Saint-Vincent

Caulerpa lessonii Bory de Saint-Vincent, 1828 [1826–1829]:193, pl. 22: fig. 3 [type locality: Oualan (Kosrae) I., Caroline Is.].—Taylor, 1977a:6.

PHILIPPINE DISTRIBUTION.—LUZON: La Union, Pangasinan.

Caulerpa mexicana Sonder ex Kützing

Caulerpa mexicana Sonder ex Kützing, 1849:496 [type locality: Mexico].—Taylor, 1966b:350.—Velasquez, Trono, and Doty, 1975:131.—Taylor, 1977a:7.—Trono and Ang, 1982:5.

Caulerpa taxifolia (Vahl) C. Agardh var. crassifolia C. Agardh, 1822a:436 [syntype localities: Indian Ocean; Red Sea; West Indies].

Caulerpa crassifolia (C. Agardh) J. Agardh, 1873:13.—Weber-van Bosse, 1913a:99 [f. typica].—Gilbert, 1942:12; 1946:78.—Gilbert, 1961:436.—Meñez, 1961:53.—Velasquez, Trono, and Doty, 1975:130.—Meñez and Calumpong, 1981:383; 1982:6, pl 1:D,E.

PHILIPPINE DISTRIBUTION.—LUZON: La Union, Pangasinan,Sorsogon. NEGROS: Negros Oriental. PALAWAN (incl. Balabac I., Bugsuk I.). SULU: Tawitawi (Sangasiapu I.).

NOTE.—The synonymy was proposed by Papenfuss (1956:65). Caulerpa crassifolia has priority only from 1873, the basionym being at varietal rank.

Caulerpa mexicana Sonder ex Kützing var. pluriseriata W.R. Taylor

Caulerpa mexicana Sonder ex Kützing var. pluriseriata W.R. Taylor, 1975:77, fig. 1 [including Philippine records] [type locality: Tatagan I., Malaysia].

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. SIQUIJOR.

Caulerpa microphysa (Weber-van Bosse) J. Feldmann

Caulerpa racemosa (Forsskål) J. Agardh var. clavifera (Turner) Weber-van Bosse f. microphysa Weber-van Bosse, 1898:361, pl XXXIII: fig. 5 [type locality: Macassar, Celebes, Indonesia].—Cordero, 1984c:52.

Caulerpa microphysa (Weber-van Bosse) J. Feldmann, 1955:430.—Taylor, 1966b:350.—Velasquez, Trono, and Doty, 1975:131.—Meñez and Calumpong, 1981:383.—Calumpong, 1982:145.—Meñez and Calumpong, 1982:7, pl 2:L.

PHILIPPINE DISTRIBUTION.—MASBATE. NEGROS: Negros Oriental (incl. Apo I.). CEBU (Pescador I., Sumilon I.). SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.). BASILAN.

Caulerpa prolifera (Forsskål) Lamouroux

Fucus prolifer Forsskål, 1775:193 [type locality: Alexandria, Egypt].

Caulerpa prolifera (Forsskål) Lamouroux, 1809b:332.—Ortega, Alcala, and Reyes, 1974:185.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental.

Caulerpa racemosa (Forsskål) J. Agardh

Fucus racemosus Forsskål, 1775:191 [type locality: Suez, Egypt].

Caulerpa racemosa (Forsskål) J. Agardh, 1873:35.—Gilbert, 1946:78.—Zaneveld, 1950:110.—Velasquez, 1953a:100.—Zaneveld, 1956:6; 1959:92.—Meñez, 1961:51.—Taylor, 1966b:350.—de los Reyes, 1967:231.—Velasquez, 1968a:119, fig. 1.—Villones and Magdamo, 1968:12, fig. 6.—Velasquez, 1971:435, fig. 13.—Kraft, 1972:328, 329.—Reyes, 1972:142.—Trono, 1972a:96.—Velasquez, 1972:63.—Bersamin et al., 1973:185.—Trono, 1973b:218.—Trono, 1973d:6, pl. 2: figs. 7, 8; pl. 4: figs. 13–16.—Velasquez et al., 1973:14, pl. 4: fig. 20.—Westernhagen, 1973a:64.—Ortega, Alcala, and Reyes, 1974:178.—Westernhagen, 1974:112 (table 1).—Trono, 1975:33.—Velasquez, Trono, and Doty, 1975:131.—Cordero, 1976c:6 [var. racemosa]; 1977b:34 [var. racemosa].—Taylor, 1977a:8.—Reyes, 1978:156, 162, pl. 8: fig. 8; pl. 9: figs. 1–4.—Sotto, 1978:109.—Trono, 1978:5.—Trono and Tuason, 1978:3.—Cordero, 1979b:276, 280.—Fortes, 1979:112, fig. 3.—Puig and Cordero, 1979:23.—Velasquez, 1979b:230.—Cordero, 1980b:24, pl. 10; 26, pls. 13, [57] [var. racemosa].—Liao and Sotto, 1980:96.—Moreland, 1980:43, 46.—Saraya and Trono, 1980:16.—Trono and Fortes, 1980:63.—Trono and Ganzon-Fortes, 1980:19, fig. [s.n].—Trono, Velasquez, and Guevarra, 1980:76.—Ganzon-Fortes, 1981:21.—Guzman, 1981:43.—Trono and De Lara, 1981:4.—Calumpong, 1982:145.—Meñez and Calumpong, 1982:7.—Trono and Ang, 1982:6.—Trono and Fortes, 1982:144.—Marcos-Agngarayngay, 1983:84, fig. 17.—Cordero, 1984a:77; 1984b:58; 1984c:51.—Marcos-Agngarayngay, 1984b:121.—Tungpalan, 1984:139.—Trono and Ganzon-Fortes, 1985:64, 66.

Fucus clavifer Turner, 1808:126, pl. 57 [type locality: Red Sea].

Caulerpa clavifera (Turner) C. Agardh, 1817:XXIII.—Dickie, 1874a:197; 1876a:244.—Howe, 1932:169.—Velasquez, Trono, and Doty, 1975:130.

Chauvinia clavifera (Turner) Bory de Saint-Vincent, 1829 [1826–1829]:207.—Martens, 1868:24, 64–65.—Velasquez, Trono, and Doty, 1975:134.

Caulerpa racemosa (Forsskål) J. Agardh var. clavifera (Turner) Weber-van Bosse, 1898:361, pl. XXXIII: figs. 1–3.—Gilbert, 1942:18; 1946:78.—Quisumbing, 1951:1007.—Gilbert 1961:437.—Galutira and Velasquez, 1964:496.—Taylor, 1966b:351.—Aguilar-Santos and Doty, 1968:174.—Taylor, 1977a:10.—Meñez and Calumpong,—1981:383.—Cordero, 1982a:60, fig. 5.—Meñez and Calumpong, 1982:7, pl. 2:A.—Hurtado-Ponce, 1983:115.—Hurtado-Ponce and Modelo, 1983:148.—Cordero, 1984c:52.—Hurtado-Ponce, 1984:179.

Fucus uvifer Turner, 1816 [1815–1819]:81, pl. 230 [type locality: Red Sea].

Caulerpa uvifera C. Agardh, 1817:XXIII.

Caulerpa racemosa (Forsskål) J. Agardh var. uvifera (C. Agardh) J. Agardh, 1873:35.—Collado, 1926:129.—G. Blanco, 1938:512.—Gilbert, 1942:20.—Zaneveld, 1950:110.—Quisumbing, 1951:1008.—Montilla and Blanco, 1953:166.—Zaneveld, 1956:7; 1959:93.—Gilbert, 1961:440.—Bersamin et al., 1973:185.—Cordero, 1980b:27, pl. 15.—Meñez and Calumpong, 1981:383; 1982:9, pl. 2:D.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Ilocos Sur, La Union, Pangasinan, Bataan, Batangas, Quezon, Albay, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. MASBATE. SAMAR: Western Samar, Eastern Samar. LEYTE (Biliran I.). PANAY: Aklan, Antique (incl. Panagatan Cays). NEGROS: Negros Occidental (Ilacaon I., Suyac I.), Negros Oriental (incl. Apo I.). CEBU (incl. Mactan I., Pescador I., Sumilon I.). SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.), Misamis Occidental, Surigao del Norte, Davao. BASILAN. PALAWAN (incl. Balabac I., Bugsuk I., Cuyo I.). SULU: Sulu (incl. Cagayan Sulu I., Siasi I.), Tawitawi (incl. Sibutu I, Tumindao I., Turtle Is.).

NOTE.—The synonymy was proposed by Papenfuss and Egerod (1957:88). Fucus uvifer Turner, the intended basionym of Caulerpa uvifera (and other combinations), is a later homonym of F. uvifer Forsskål (1775:192) and hence not priorable. Caulerpa uvifera C. Agardh is treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

Caulerpa racemosa (Forsskål) J. Agardh var. corynephora (Montagne) Weber-van Bosse

Caulerpa corynephora Montagne, 1842a:14 [type locality: Toud I. (Warrior Islet), Torres Strait, Australia].

Caulerpa racemosa (Forsskål) J. Agardh var. corynephora (Montagne) Weber-van Bosse, 1898:364, pl. XXXIII: figs. 10–13.—Gilbert, 1942:20; 1961:438.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Caulerpa racemosa (Forsskål) J. Agardh var. gracilis (Zanardini) Weber-van Bosse

Caulerpa clavifera (Turner) C. Agardh var. gracilis Zanardini, 1851:37 [type locality: Tor, Sinai Peninsula, Egypt].

Caulerpa racemosa (Forsskål) J. Agardh var. gracilis (Zanardini) Weber-van Bosse, 1898:370.—Gilbert, 1942:21; 1961:438.

PHILIPPINE DISTRIBUTION.—Locality not specified.

Caulerpa racemosa (Forsskål) J. Agardh var. laetevirens (Montagne) Weber-van Bosse

Caulerpa laetevirens Montagne, 1842a:13 [type locality: Toud I. (Warrior Islet), Torres Strait, Australia].—Piccone, 1886:29, 89.—Velasquez, Trono, and Doty, 1975:131.

Caulerpa racemosa (Forsskål) J. Agardh var. laetevirens (Montagne) Weber-van Bosse, 1898:366, pl. XXXII: figs. 16–22.—Gilbert, 1942:21; 1946:78.—Zaneveld, 1950:110; 1956:7; 1959:93.—Gilbert, 1961:438.—Taylor, 1966b:351.—Kraft, 1972:328.—Cordero, 1976c:6, 8; 1977b:33.—Taylor, 1977a:10.—Cordero, 1980b:25, pl. II; 1984c:52.

PHILIPPINE DISTRIBUTION.—BATANES. MASBATE (incl. Ticao I.), PANAY: Antique (Panagatan Cays). CEBU (incl. Olango I.). SIQUIJOR. MINDANAO: Zamboanga. BASILAN. SULU: Tawitawi (Bilitan I., Sibutu I.).

Caulerpa racemosa (Forsskål) J. Agardh var. lamourouxii (Turner) Weber-van Bosse

Fucus lamourouxii Turner, 1816 [1815–1819]:79, pl. 229 [type locality: Red Sea].

Caulerpa racemosa (Forsskål) J. Agardh var. lamourouxii (Turner) Weber-van Bosse, 1898:368, pl. XXXII: figs. 1–7; pl. XXXIII: fig. 15.—Gilbert, 1961:438.—Taylor, 1977a:10.—Cordero, 1978b:47, fig. 3; 1980b:26, pl. 12.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.). MINDANAO: Zamboanga, Surigao del Sur.

Caulerpa racemosa (Forsskål) J. Agardh var. macro Weber-van Bosse

Caulerpa racemosa (Forsskål) J. Agardh var. macro Weber-van Bosse, 1913a:106, fig. 26 [syntype localities: Moluccas, Indonesia: Fau Islet, Gebe I.; Majalibit Bay, Waigeo I.].—Gilbert, 1942:20; 1961:439.

PHILIPPINE DISTRIBUTION.—PALAWAN (Culion I.).

Caulerpa racemosa (Forsskål) J. Agardh var. macrophysa (Sonder ex Kützing) Taylor

Chauvinia macrophysa Sonder ex Kützing, 1857:6, pl. 15: fig. II [type locality: Central America].

Caulerpa racemosa (Forsskål) J. Agardh var. macrophysa (Sonder ex Kützing) Taylor, 1928:101, pl. 12: fig. 3; pl. 13: fig. 9.—Gilbert, 1942:19; 1961:439.—Domantay, 1962:277.—Taylor, 1966b:351.—Trono, 1973c:11, fig. 3.—Meñez and Calumpong, 1981:383; 1982:8, pl. 2:C.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDORO: Oriental Mindoro. NEGROS: Negros Oriental. CEBU. SIQUIJOR. MINDANAO: Zamboanga. BASILAN. SULU: Sulu (North Ubian I.).

Caulerpa racemosa (Forsskål) J. Agardh var. occidentalis (J. Agardh) Børgesen

Caulerpa chemnitzia (Esper) Lamouroux var. occidentalis J. Agardh, 1873:37 [type locality: upper Gulf of Mexico to Recife, Brazil].

Caulerpa racemosa (Forsskål) J. Agardh var. occidentalis (J. Agardh) Børgesen, 1907:379, figs. 28, 29.—Domantay, 1962:278.—Taylor, 1966b:351.—Taylor, 1977a:10.—Meñez and Calumpong, 1981:383; 1982:8, pl. 2:B.—Cordero, 1984c:51.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Sur, Pangasinan, Rizal, Sorsogon. CATANDUANES. MINDORO: Oriental Mindoro. MARINDUQUE. MASBATE. NEGROS: Negros Oriental, SIQUIJOR. MINDANAO: Zamboanga. BASILAN. PALAWAN, SULU: Sulu (Tapul Group).

Caulerpa racemosa (Forsskål) J. Agardh var. peltata (Lamouroux) Eubank

Caulerpa peltata Lamouroux, 1809b:332 [type locality: Antilles].—Dickie, 1876a:244.—Collado, 1926:129.—Gilbert, 1942:22 [var. typica]; 1946:78.—Taylor, 1966b:350 [var. peltata].—de los Reyes, 1967:231.—Reyes, 1972:142.—Velasquez et al., 1973:14, pl. 4: fig. 18.—Ortega, Alcala, and Reyes, 1974:186, 187.—Trono, 1975:33.—Velasquez, Trono, and Doty, 1975:131.—Cordero, 1976a:84; 1976c:8.—Taylor, 1977a:8.—Cordero, 1978b:45, fig. 2 [var. typica].—Reyes, 1978:156, pl. 9: fig. 5.—Cordero, 1979b:280, fig. 2 [var. typica]; 1980b:23, pl. 9 [var. typica].—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:15, pl. II: fig. 3.—Trono and Fortes, 1980:63.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:144.—Cordero, 1984a:77; 1984c:51.—Marcos-Agngarayngay, 1984b:121.

Caulerpa racemosa (Forsskål) J. Agardh var. peltata (Lamouroux) Eubank, 1946:421, fig. 2r,s.—Gilbert, 1961:439.—Cordero, 1977b:33.—Meñez and Calumpong, 1981:383; 1982:8, pl. 2:K.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Ilocos Sur, Pangasinan, Batangas, Albay, Sorsogon. CATANDUANES. MINDORO: Oriental Mindoro. MASBATE. SAMAR: Eastern Samar. LEYTE (Biliran I.), PANAY: Aklan. NEGROS: Negros Oriental (incl. Apo I.). CEBU (incl. Mactan I., Pescador I., Sumilon I.). SIQUIJOR. MINDANAO: Zamboanga, Surigao del Norte. BASILAN. PALAWAN (incl. Cuyo I.). SULU: Sulu (incl. North Ubian I., Siasi I.), Tawitawi (incl. Tumindao I.).

NOTE.—Shameel (1978) argues for retention of C. peltata as a distinct species.

TAXA OF UNCERTAIN VALUE

The following two infraspecific taxa are probably related to Caulerpa racemosa but have not been formally transferred to it.

Caulerpa peltata Lamouroux var. macrodisca (Decaisne) Weber-van Bosse

Caulerpa macrodisca Decaisne, 1842a:336 [type locality: Anambas Is., Indonesia].—Howe, 1932:169.—Velasquez, Trono, and Doty, 1975:131.—Taylor, 1977a:7.

Caulerpa peltata Lamouroux var. macrodisca (Decaisne) Weber-van Bosse, 1898:376.—Gilbert, 1942:23.—Zaneveld, 1950:110.—Montilla and Blanco, 1953:166.—Zaneveld, 1956:6; 1959:92.

PHILIPPINE DISTRIBUTION.—PANAY. CEBU.

Caulerpa peltata Lamouroux var. nummularia (Harvey ex J. Agardh) Weber-van Bosse

Caulerpa nummularia Harvey ex J. Agardh, 1873:38 [syntype localities: Tonga; Nukahiva, Marquesas Is.].

Caulerpa peltata Lamouroux var. nummularia (Harvey ex J. Agardh) Weber-van Bosse, 1898:376, pl. XXXII: fig. 9.—Cordero, 1976a:84, fig. 1; 1976c:6; 1977b:32.

PHILIPPINE DISTRIBUTION.—BATANES.

Caulerpa racemosa (Forsskål) J. Agardh var. turbinata (J. Agardh) Eubank

Caulerpa clavifera (Turner) C. Agardh var. turbinata J. Agardh, 1837:173 [type locality: near Tor, Sinai Peninsula, Egypt].

Caulerpa racemosa (Forsskål) J. Agardh var. turbinata (J. Agardh) Eubank, 1946:420, figs. 2o–q.—Taylor, 1977a:10.—Cordero, 1980b:27, pl. 14.

Fucus chemnitzia Esper, 1800:167 [given as “127”], pl. LXXXVIII [type locality: Malabar Coast, India].

Caulerpa racemosa (Forsskål) J. Agardh var. chemnitzia (Esper) Weber-van Bosse, 1898:370, pl. XXXI: figs. 5–8.—Gilbert, 1961:437.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon. SAMAR: Eastern Samar. BASILAN. SULU: Sulu.

NOTE.—The synonymy was proposed by Weber-van Bosse (1898:370)

*Caulerpa reyesii Meñez and Calumpong

Caulerpa reyesii Meñez and Calumpong, 1982:10, fig. 2, pl. 3:G–I [type locality: Solong-on, Siquijor I.].—Reyes, 1978:157, pl. 9: figs. 6, 7 [Caulerpa sp.].

PHILIPPINE DISTRIBUTION.—As above.

Caulerpa selago (Turner) C. Agardh

Fucus selago Turner, 1808:122, pl. 55 [type locality: Red Sea].

Caulerpa selago (Turner) C. Agardh, 1817:XXIII.—Weber-van Bosse, 1913a:101.—Gilbert, 1946:78; 1961:440.—Velasquez, Trono, and Doty, 1975:132.—Taylor, 1977a:10.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I., Tumindao I.).

Caulerpa serrulata (Forsskål) J. Agardh

Fucus serrulatus Forsskål, 1775:189 [type locality: Mokha, Yemen].

Caulerpa serrulata (Forsskål) J. Agardh, 1837:174.—Gilbert, 1942:14 [incl. var. typica f. serrulata]; 1946:78.—Zaneveld, 1950:111.—Montilla and Blanco, 1953:166.—Zaneveld, 1956:7; 1959:94.—Gilbert, 1961:440.—Meñez, 1961:53.—Domantay, 1962:278.—Taylor, 1966b:351.—Aguilar-Santos and Doty, 1968:174.—Velasquez, 1971:435, fig. 15.—Reyes, 1972:142.—Trono, 1972a:96.—Cordero, 1973b:19.—Bersamin et al., 1973:185.—Trono, 1973b:218; 1973c:11, figs. 2, 23; 1973d:7, pl. 5: fig. 19.—Velasquez et al., 1973:15, pl. 5: fig. 21.—Ortega, Alcala, and Reyes, 1974:186.—Trono, 1975:33.—Velasquez, Trono, and Doty, 1975:132.—Taylor, 1977a:10 [var. serrulata].—Reyes, 1978:155, pl. 8: fig. 4.—Trono, 1978:5.—Trono and Tuason, 1978:3.—Cordero, 1979b:276.—Fortes, 1979:112.—Garcia, 1979:44 (table 1).—Puig and Cordero, 1979:24.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:18.—Trono and Ganzon-Fortes, 1980:21, fig. [s.n].—Meñez and Calumpong, 1981:383.—Trono and De Lara, 1981:4, pl. III: fig. 1.—Cordero, 1982a:60.—Meñez and Calumpong, 1982:9, pl. 2:E.—Hurtado-Ponce, 1983:116.—Hurtado-Ponce and Modelo, 1983:148 [var. serrulata].—Marcos-Agngarayngay, 1983:84, fig. 18.—Cordero, 1984a:77; 1984b:58; 1984c:51.—Hurtado-Ponce, 1984:179.—Marcos-Agngarayngay, 1984b:122.

Caulerpa freycinetii C. Agardh, 1822a:446 [type locality: Mariana Is.].—Collado, 1926:129.—G. Blanco, 1938:512.—Quisumbing, 1951:1007 [var. typica].—Montilla and Blanco, 1953:166.—de los Reyes, 1967:231.—Cordero, 1982a:60.

Caulerpa hummii Díaz-Piferrer, 1969b:13, fig. 1 [type locality: Orquilla I. (cited Orchilla I. in error), Archiplélago Los Hermanos, Venezuela].—Trono, 1973c:9, fig. 19.—Taylor, 1977a:11–12.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Ilocos Sur, Pangasinan, Bataan, Batangas, Quezon, Albay, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. MASBATE. SAMAR: Eastern Samar. LEYTE (Biliran I.). PANAY: Aklan. NEGROS: Negros Oriental (incl. Apo I.). CEBU (incl. Mactan I., Sumilon I.). SIQUIJOR. MINDANAO: Zamboanga, Davao. BASILAN. PALAWAN (incl. Balabac I., Culion I.). SULU: Sulu (incl. Siasi I.), Tawitawi (incl. Tumindao I.).

NOTE.—The conspecificity of Caulerpa freycinetii and C. serrulata was proposed by Børgesen (1932:5).Caulerpa hummii is included as a synonym on the authority of Taylor (1977a:11–12).

Caulerpa serrulata (Forsskål) J. Agardh var. boryana (J. Agardh) Gilbert

Caulerpa boryana J. Agardh, 1873:20 [type locality: “ad insulas Oceani pacifici calidioris”].

Caulerpa serrulata (Forsskål) J. Agardh var. boryana (J. Agardh) Gilbert, 1942:15.—Taylor, 1977a:11.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MINDANAO: Davao.

*Caulerpa serrulata (Forsskål) J. Agardh var. boryana (J. Agardh) Gilbert f. longifolia Gilbert

Caulerpa serrulata (Forsskål) J. Agardh var. boryana (J. Agardh) Gilbert f. longifolia Gilbert, 1942:16, figs. 4, 5 [type locality: Siasi I., Sulu Prov., Sulu Archipelago].—Gilbert, 1961:441.—Hurtado-Ponce and Modelo, 1983:149.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, SULU: Sulu (Siasi I.).

Caulerpa serrulata (Forsskål) J. Agardh var. boryana (J. Agardh) Gilbert f. occidentalis (Weber-van Bosse) Yamada and Tanaka

Caulerpa freycinetii C. Agardh var. boryana f. occidentalis Weber-van Bosse, 1898:315, pl. XXV: fig. 11 [type locality: Guadeloupe].—Gilbert, 1942:15; 1961:440.

Caulerpa serrulata (Forsskål) J. Agardh var. boryana (J. Agardh) Gilbert f. occidentalis (Weber-van Bosse) Yamada and Tanaka, 1938:62.—Gilbert, 1942:15; 1961:441.—Meñez and Calumpong, 1981:383; 1982:9, pl. 2:F.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental. PALAWAN (Culion I.). SULU: Sulu (Siasi I.).

Caulerpa serrulata (Forsskål) J. Agardh f. lata (Weber-van Bosse) Tseng

Caulerpa freycinetii C. Agardh var. typica f. lata Weber-van Bosse, 1898:313, pl. XXV: fig. 5 [syntype localities: Red Sea; Mariana Is.; Tonga; Sumbawa I., Indonesia; Guadeloupe].—Gilbert, 1942:15; 1961:440.

Caulerpa serrulata (Forsskål) J. Agardh f. lata (Weber-van Bosse) Tseng, 1936:178.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Batangas. MINDORO: Oriental Mindoro. LEYTE (Biliran I.). CEBU (Mactan I.). MINDANAO: Zamboanga. BASILAN. PALAWAN (incl. Culion I.).

Caulerpa serrulata (Forsskål) J. Agardh var. pectinata (Weber-van Bosse) W.R. Taylor

Caulerpa freycinetii C. Agardh var. pectinata Weber-van Bosse, 1898:316 [type locality: Guadeloupe].—Weber-van Bosse, 1913a:102.—Velasquez, Trono, and Doty, 1975:131 [without designation of variety].

Caulerpa serrulata (Forsskål) J. Agardh var. pectinata (Weber-van Bosse) W.R. Taylor, 1960:146.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I.).

Caulerpa serrulata (Forsskål) J. Agardh f. spiralis (Weber-van Bosse) Gilbert

Caulerpa freycinetii C. Agardh var. typica f. spiralis Weber-van Bosse, 1898:314 [type locality: New Caledonia].

Caulerpa serrulata (Forsskål) J. Agardh f. spiralis (Weber-van Bosse) Gilbert, 1942:15 [var. typica f. spiralis].—Trono, 1973c:11, fig. 2.—Taylor, 1977a:11 [var. spiralis].

PHILIPPINE DISTRIBUTION.—LUZON: Batangas, Sorsogon. CATANDUANES. MINDORO: Oriental Mindoro. SULU: Sulu.

Caulerpa sertularioides (S.G. Gmelin) Howe

Fucus sertularioides S.G. Gmelin, 1768:151, pl. XV: fig. 4 [type locality: “in coralliis americanis”].

Caulerpa sertularioides (S.G. Gmelin) Howe, 1905:576.—Weber-van Bosse, 1913a:100 [forma typica].—Collado, 1926:129.—G. Blanco, 1938:512.—Gilbert, 1946:78.—Zaneveld, 1950:111.—Quisumbing, 1951:1008.—Montilla and Blanco, 1953:166.—Zaneveld, 1956:8; 1959:94.—Gilbert, 1961:441.—Meñez, 1961:52.—Domantay 1962:278.—Galutira and Velasquez, 1964:496.—Taylor, 1966b:351.—Aguilar-Santos and Doty, 1968:174.—Velasquez, 1968a:119, fig. 2.—Villones and Magdamo, 1968:24, fig. 9 [figure cited but not published].—Velasquez, 1971:435, fig. 14.—Reyes, 1972 143.—Trono, 1972a:96.—Bersamin et al., 1973:186.—Trono 1973b:219; 1973c:12, fig. 4; 1973d:7, pl. 3: fig. 11.—Velasquez et al., 1973:14, pl. 4: fig. 19.—Westernhagen, 1973a:64.—Ortega, Alcala and Reyes, 1974:187.—Westernhagen, 1974:112 (table 1).—Trono 1975:34.—Velasquez, Trono, and Doty, 1975:132.—Taylor 1977a:12.—Vannajan and Trono, 1977:44, fig. 8.—Reyes, 1978:154, 162, pl. 7: fig. 6.—Trono, 1978:5.—Trono and Tuason, 1978:5.—Cordero, 1979b:276, 280.—Fortes, 1979:112.—Garcia, 1979:44 (table 1).—Puig and Cordero, 1979:24.—Cordero, 1980b:28.—Saraya and Trono, 1980:17.—Trono and Fortes, 1980:64.—Trono and Ganzon-Fortes, 1980:23, fig. [s.n].—Trono, Velasquez, and Guevarra, 1980:77.—Ganzon-Fortes, 1981:21.—Meñez and Calumpong, 1981:383.—Trono and De Lara, 1981:5.—Calumpong, 1982:145.—Meñez and Calumpong, 1982:9, pl. 3:F.—Trono and Fortes, 1982:144.—Trono and Ang, 1982:5.—Hurtado-Ponce, 1983:116.—Hurtado-Ponce and Modelo, 1983:149.—Marcos-Agngarayngay, 1983:85, fig. 19.—Cordero, 1984a:78; 1984b:59; 1984c:51.—Hurtado-Ponce, 1984:179.—Marcos-Agngarayngay, 1984b:122.—Ang, 1985b:298.

Fucus plumaris Forsskål, 1775:190 [type locality: Mokha, Yemen].

Caulerpa plumaris (Forsskål) C. Agardh, 1822a:436.—Montagne, 1844a:659.—Martens, 1868:62–63.—Dickie, 1876a:245.—Velasquez, Trono, and Doty, 1975:131.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, Pangasinan, Bataan, Manila, Batangas, Quezon, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. MASBATE. LEYTE (Biliran I.). PANAY: Aklan. NEGROS: Negros Occidental, Negros Oriental. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga. PALAWAN (incl. Bugsuk I., Cuyo I.). SULU: Sulu (North Ubian I., Siasi I.), Tawitawi.

NOTE.—The conspecificity of Fucus sertularioides and F. plumaris was proposed by C. Agardh (1822a:436), who erroneously adopted the later of the two names. The situation was corrected by Howe (1905:576).

Caulerpa sertularioides (S.G. Gmelin) Howe f. brevipes (J. Agardh) Svedelius

Caulerpa plumaris (Forsskål) C. Agardh var. brevipes J. Agardh, 1873:15 [type locality: uncertain; see Note].

Caulerpa sertularioides (S.G. Gmelin) Howe f. brevipes (J. Agardh) Svedelius, 1906:114.—Gilbert, 1942:13; 1961:441.—Taylor, 1966b:351 [var. sertularioides f. brevipes].—Cordero, 1976c:8 [var. brevipes]; 1977b:34.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Rizal, Manila Bay. MINDORO: Oriental Mindoro. PALAWAN (Cuyo I.). SULU: Tawitawi (Pearl Bank, Turtle Is.).

NOTE.—In the protologue of C. plumaris var. brevipes, J. Agardh cited as a synonym Fucus sertularioides S.G. Gmelin, a species with the West Indies as its type locality, but indicated the provenance of the variety as “in pacifico calidiore.” An examination of pertinent specimens in the Agardh Herbarium (in LD) would probably resolve the conflict.

Caulerpa sertularioides (S.G. Gmelin) Howe f. farlowii (Weber-van Bosse) Børgesen

Caulerpa plumaris (Forsskål) C. Agardh f. farlowii Weber-van Bosse, 1898:295, pl. XXIV: figs. 5, 6 [“farlowi”] [syntype localities: Ilhas do Cabo Verde; Florida, USA; Flores, Indonesia].

Caulerpa sertularioides (S.G. Gmelin) Howe f. farlowii (Weber-van Bosse) Børgesen, 1907:365.—Trono and Ang, 1982:6.

PHILIPPINE DISTRIBUTION.—PALAWAN (Bugsuk I.).

Caulerpa sertularioides (S.G. Gmelin) Howe f. flagellata (Weber-van Bosse) Weber-van Bosse

Caulerpa plumaris (Forsskål) C. Agardh f. flagellata Weber-van Bosse, 1898:295, pl. XXIV: fig. 10 [type locality: Tonga].

Caulerpa sertularioides (S.G. Gmelin) Howe f. flagellata (Weber-van Bosse) Weber-van Bosse, 1913a:101 [including Philippine record].

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Sangasiapu I.).

Caulerpa sertularioides (S.G. Gmelin) Howe f. longipes (J. Agardh) Collins

Caulerpa plumaris (Forsskål) C. Agardh var. longipes J. Agardh, 1873:15 [type locality: West Indies].

Caulerpa sertularioides (S.G. Gmelin) Howe f. longipes (J. Agardh) Collins, 1909:415.—Weber-van Bosse, 1913a:101.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

Caulerpa sertularioides (S.G. Gmelin) Howe f. longiseta (Bory de Saint-Vincent) Svedelius

Caulerpa plumaris (Forsskål) C. Agardh var. longiseta Bory de Saint-Vincent, 1828 [1826–1829]:194, pl. 22: fig. 4 [type locality: not specified].

Caulerpa sertularioides (S.G. Gmelin) Howe f. longiseta (Bory de Saint-Vincent) Svedelius, 1906:114, fig. 10.—Gilbert, 1942:13; 1961:441.—Domantay, 1962:278.—Taylor, 1966b:351.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Ilocos Sur, Pangasinan, Bataan, Rizal, Manila Bay. MINDORO: Oriental Mindoro. MINDANAO: Zamboanga. BASILAN. PALAWAN.

Caulerpa subserrata Okamura

Caulerpa subserrata Okamura, 1897:3, pl. I: figs. 1,2 [type locality: Ogasawara-gunto (Bonin Is.), Japan].—Cordero, 1980c:70, fig. 1.—Hurtado-Ponce, 1983:117.—Hurtado-Ponce and Modelo, 1983:149.—Marcos-Agngarayngay, 1983:86, fig. 20.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Caulerpa taxifolia (Vahl) C. Agardh

Fucus taxifolius Vahl, 1802:36 [type locality: St. Croix, Virgin Is.].

Caulerpa taxifolia (Vahl) C. Agardh, 1817:xxII.—Montagne, 1844a:659.—Martens, 1868:62–63.—Gilbert, 1942:12; 1946:78; 1961:441.—Meñez, 1961:53.—de los Reyes, 1967:231.—Trono, 1975:34.—Velasquez, Trono, and Doty, 1975:133.—Cordero, 1976c:10; 1977b:34.—Taylor, 1977a:13.—Reyes, 1978:155, pl. 8: fig. 5.—Liao and Sotto, 1980:96.—Saraya and Trono, 1980:16.—Trono and Fortes, 1980:64.—Ganzon-Fortes, 1981:21.—Meñez and Calumpong, 1981:383.—Cordero, 1984a:78; 1984b:59; 1984c:51.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Sorsogon. MASBATE. LEYTE (Biliran I.). CEBU (Mactan I.). SIQUIJOR. BASILAN. PALAWAN. SULU: Sulu (North Ubian I.).

Caulerpa urvilliana Montagne

Caulerpa urvilliana Montagne, 1845:21 [type locality: Toud I. (Warrior Islet), Torres Strait, Australia].—Taylor, 1966b:352.—Trono, 1972a:97; 1973b:219; 1973d:7, pl. 2: fig. 5; 1975:35.—Velasquez, Trono, and Doty, 1975:133.—Taylor, 1977a:14 [var. urvilliana].—Reyes, 1978:154, pl. 7: fig. 5.—Meñez and Calumpong, 1981:383; 1982:10, pl. 3:D,E.—Trono and Ang, 1982:6.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon. NEGROS: Negros Occidental, Negros Oriental, SIQUIJOR. PALAWAN (incl. Bugsuk I., Cuyo I.). SULU: Sulu (incl. Siasi I.), Tawitawi (incl. Pearl Bank, Sibutu I.).

Caulerpa urvilliana Montagne var. vitiensis Weber-van Bosse

Caulerpa urvilliana Montagne var. vitiensis Weber-van Bosse, 1898:319, pl. XXVI: fig. 12 [syntype localities: Fiji; New Hebrides].—Taylor, 1977a:14.

PHILIPPINE DISTRIBUTION.—SULU: Sulu.

Caulerpa verticillata J. Agardh

Caulerpa verticillata J. Agardh, 1847:6 [type locality: not specified].—Gilbert, 1961:442.—Trono, 1973b:219; 1973d:8, pl. 3: fig. 10.—Velasquez, Trono, and Doty, 1975:133.—Taylor, 1977a:14.—Reyes, 1978:154, pl. 8: fig. 3 [forma typica].—Saraya and Trono, 1980:18.—Meñez and Calumpong, 1981:383; 1982:10, pl. 3:A–C.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. NEGROS: Negros Oriental (Apo I.). CEBU (Mactan I.). SIQUIJOR. BASILAN. PALAWAN. SULU: Sulu (Siasi I.).

Caulerpa vesiculifera (Harvey) Harvey

Caulerpa simpliciuscula (Turner) C. Agardh var. vesiculifera Harvey, 1859: pl. LXV: figs. 3, 4 [lectotype locality: Western Port, Victoria, Australia fide Womersley, 1984:272].

Caulerpa vesiculifera (Harvey) Harvey, 1863:lvi.—Taylor, 1966b:352.—Velasquez, Trono, and Doty, 1975:133.

PHILIPPINE DISTRIBUTION.—PALAWAN.

Caulerpa webbiana Montagne

Caulerpa webbiana Montagne, 1837:354 [type locality: Arrecife, Isla Lanzarote, Islas Canarias].—Meñez and Calumpong, 1981:383; 1982:10, pl. 2:G–J.—Cordero, 1984a:78.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte. NEGROS: Negros Oriental (Apo I.).

Caulerpa webbiana Montagne var. pickeringii (Harvey and Bailey) Eubank

Caulerpa pickeringii Harvey and Bailey, 1851:373 [type locality: Wilson’s I. (Manihi I.), Tuamotu Archipelago].—Trono, 1973a:128.

Caulerpa webbiana Montagne var. pickeringii (Harvey and Bailey) Eubank, 1946:416.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon.

Family CODIACEAE

Codium Stackhouse

A preliminary study (unpublished) of Philippine Codium was made by P.C. Silva in 1950–1951 as part of a worldwide survey of the genus, but with special reference to Hawaii (Silva, 1952b). It was concluded that the representation in the Indonesian-Philippine region posed many problems that could not be solved until the genus as a whole was better understood. Subsequently, the Codium flora has been worked out for five temperate regions—southern Australia (Silva and Womersley, 1956), South Africa (Silva, 1959), New Zealand, Japan, and Pacific Mexico (the last three unpublished)—each showing a high degree of endemism. In addition, the Codium flora of the troplcal Atlantic has been worked out (Silva, 1960), but Indonesian-Philippine collections that have become available since 1951 have been examined only superficially. At present, it is easier to say which species do not occur in the Philippines than it is to say which ones are there.

Codium arabicum Kützing

Codium arabicum Kützing, 1856:35, pl. 100: fig. II [type locality: Tor, Sinai Peninsula, Egypt].—Silva, 1952b:382, figs. 11–13, pl. 34b.—Meñez, 1961:54, pl. 6: figs. 58, 59.—Reyes, 1972:143.—Trono 1973b:221; 1973c:13, fig. 5.—Ortega, Alcala, and Reyes, 1974:185, 186.—Velasquez, Trono, and Doty, 1975:137.—Reyes, 1978:157, pl. 9: fig. 8.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:20, pl. IV: fig. 2.—Meñez and Calumpong, 1981:383.—Trono and De Lara 1981:5, pl. III: fig. 2.—Hurtado-Ponce and Modelo, 1983:149.—Meñez, Phillips, and Calumpong, 1983:23.—Meñez and Calumpong 1984:105.—Ang, 1985b:298.

Codium coronatum Setchell, 1926:82, pl. 10: figs. 2–5; pl. II: figs. 2, 3; pl.12: figs. 1, 5 [type locality: Arue Reef, Tahiti].—Gilbert, 1947:123; 1961:442.—Velasquez, Trono, and Doty, 1975:137.

Codium adhaerens [misapplied name].—Dickie, 1876a:243.—Gilbert 1946:78; 1947:123; 1961:442.—Villones and Magdamo, 1968:25, fig 11.—Velasquez, Trono, and Doty, 1975:136.—Cordero, 1979b:276, 281.—Puig and Cordero, 1979:26.—Cordero, 1984b:59; 1984c:52.

Codium setchellii [misapplied name].—Cordero, 1973b:20.—Westernhagen, 1973a:64; 1974:112 (table 1).—Cordero, 1976c:8; 1977b:42, pl. VII: figs. 39–42; 1980b:30, fig. 3A.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Batangas, Quezon, MASBATE. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. LEYTE (Biliran I.). PANAY: Aklan. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga. BASILAN. PALAWAN. SULU: Sulu (Siasi I.).

NOTE.—The synonymy was proposed by Silva (1952b:382).Codium adhaerens C. Agardh (1822a:457) is restricted to the Atlantic coast of southern Europe and adjacent North Africa, while C. setchellii Gardner (1919:489, pl. 42: figs. 10, 11) is restricted to the cool-temperate portion of the Pacific coast of North America.

*Codium bartlettii Tseng and Gilbert

Codium bartlettii Tseng and Gilbert, 1942:291, figs. 1, 2a [type locality: Puerto Galera, Oriental Mindoro Prov., Mindoro].—Gilbert, 1947:124; 1961:442.—Trono, 1973c:13, fig. 8.—Velasquez, Trono, and Doty, 1975:137.—Meñez and Calumpong, 1981:383.—Calumpong, 1982:145.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro. NEGROS: Negros Oriental.

Codium contractum Kjellman

Codium contractum Kjellman, 1897b:35, pl. 2: fig. 12; pl. 7: figs. 1–3 [type locality: Amakusa Is., Kumamoto Prefecture, Japan].—Gilbert, 1947:124; 1961:442.—Velasquez, Trono, and Doty, 1975:137.

PHILIPPINE DISTRIBUTION.—LUZON: Cavite.

Codium cylindricum Holmes

Codium cylindricum Holmes, 1896:250, pl. 7: figs. 1a,b [type locality: Misaki, Kanagawa Prefecture, Japan].—Cordero, 1980b:28; 1984a:79.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte. SAMAR: Eastern Samar.

Codium difforme auctorum

Codium difforme [misapplied name].—Gepp and Gepp, 1911:134.—Weber-van Bosse, 1913a:118.—Gilbert, 1946:78; 1961:443.—Velasquez, Trono, and Doty, 1975:137.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I.).

NOTE.—Codium difforme Kützing (1843b:309) is a taxonomic synonym of C. effusum (Rafinesque) Delle Chiaje (1829:14, pl. 37), a name that applies to a species restricted to the Mediterranean Sea and adjacent Atlantic Ocean shores. The Siboga record (Gepp and Gepp, 1911:134) probably is referable to C. spongiosum Harvey (1855:565), a widespread Indo-Pacific species. The specimen, which would permit confirmation, is not among other Siboga collections at Leiden (L).

Codium edule P.C. Silva

Codium edule P.C. Silva, 1952b:392, fig. 18, pl. 35b [type locality: Waikiki, Oahu, Hawaiian Is.].—Trono and Fortes, 1980:64.—Ganzon-Fortes, 1981:21.—Trono and Fortes, 1982:145.—Hurtado-Ponce, 1983:119.—Hurtado-Ponce and Modelo, 1983:149.—Marcos-Agngarayngay, 1983:94, fig. 27.—Hurtado-Ponce, 1984:179.—Marcos-Agngarayngay, 1984b:122.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte.

Codium elongatum auctorum

Codium elongatum [misapplied name].—Gepp and Gepp, 1911:136.—Weber-van Bosse, 1913a:120.—Gilbert, 1946:78; 1961:443.—Velasquez, Trono, and Doty, 1975:137.

PHILIPPINE DISTRIBUTION.—SULU: Tawitawi (Pearl Bank).

NOTE.—Codium elongatum (Turner) C. Agardh (1822a:454) is a nomenclatural synonym of C. decorticatum (Woodward) Howe (1911:494), a name that applies to a species restricted to warm waters of the Mediterranean Sea and Atlantic Ocean (cf. Silva, 1960:516–521). The Siboga record (Gepp and Gepp, 1911:136) may be referable to C. papillatum, but the voucher specimen is not among other Siboga collections at Leiden (L).

Codium fragile (Suringar) Hariot

Acanthocodium fragile Suringar, 1867:258 [type locality: Japan].

Codium fragile (Suringar) Hariot, 1889:32.—Westernhagen, 1973a:64.—Cordero, 1980b:29, pl. 17.—Tungpalan, 1984:139.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte. PANAY: Aklan. CEBU (Mactan I.). SIQUIJOR.

Codium geppii O.C. Schmidt

Codium divaricatum A. Gepp and E.S. Gepp, 1911:136, 145, pl. 22: figs. 195–199 [syntype localities: Great Kai I., Moluccas and Kwandang Bay, Celebes, Indonesia] [replaced name].

Codium geppii O.C. Schmidt, 1923:50, fig. 33.—Gilbert, 1947:123.—Quisumbing, 1951:1009.—Gilbert, 1961:443.—Domantay, 1962:280.—Velasquez, Trono, and Doty, 1975:137.—Meñez and Calumpong, 1981:383.—Calumpong, 1982:145.

Codium repens [misapplied name].—Cordero, 1977b:42, pl. VII: figs. 43–45; 1979b:276; 1980b:30, fig. 3B, pl. 18.—Marcos-Agngarayngay, 1983:97, fig. 28; 1984b:122.—Tungpalan, 1984:139.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, La Union, Pangasinan, Albay. CATANDUANES. MINDORO: Oriental Mindoro. PANAY: Aklan. NEGROS: Negros Oriental.

NOTE.—Codium repens P. Crouan and H. Crouan (in Vickers, 1905:56) in the Atlantic is the ecological and taxonomic counterpart of C. geppii in the Indo-Pacific. Each species encompasses a wide range of morphological variation.Codium geppii is a substitute name for C. divaricatum A. Gepp and E.S. Gepp, a later homonym of C. divaricatum (C. Agardh) Biasoletto (1841:237).

Codium intricatum Okamura

Codium intricatum Okamura, 1913:74, pl. CXX: figs. 9–13 [syntype localities: various, all in Japan].—Gilbert, 1947:123; 1961:443.—Meñez, 1961:54, pl. 6: figs. 63, 64.—Galutira and Velasquez, 1964:499, pl. 2: fig. 5; pl. 7: fig. 25.—Velasquez, 1968a:120, fig. 4; 1971:437, fig. 18; 1972:63.—Velasquez, Trono, and Doty, 1975:137.—Reyes, 1978:157, 162, pl. 9: figs. 9, 10.—Puig and Cordero, 1979:26.—Velasquez, 1979b:230.—Cordero, 1980b:29, pl. 16.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:20.—Trono and Fortes, 1980:64.—Trono, Velasquez, and Guevarra, 1980:77.—Ganzon-Fortes, 1981:21.—Guzman, 1981:43.—Cordero, 1982a:60.—Trono and Fortes, 1982:145.—Cordero, 1984a:79.—Tungpalan, 1984:139.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Quezon, LEYTE (Biliran I.). CEBU (Mactan I.). SIQUIJOR. BASILAN.

Codium muelleri auctorum

Codium muelleri [misapplied name].—Zaneveld, 1950:112; 1956:9; 1959:95.—Bersamin et al., 1973:186.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan, Ilocos, La Union, Pangasinan. LEYTE. CEBU. BOHOL.

NOTE.—Codium muelleri Kützing (1856:34, pl. 95: fig. II) is a distinctive species apparently restricted to southwestern and southern Australia. It seems likely that these records of C. muelleri and most (if not all) Philippine records of C. tenue apply to the same undescribed species.

Codium ovale Zanardini

Codium ovale Zanardini, 1878:37 [type locality: Sorong, Irian Barat, Indonesia].—Gepp and Gepp, 1911:134.—Weber-van Bosse, 1913a:118.—Gilbert, 1946:78; 1961:443.—Trono, 1973b:221.—Velasquez, Trono, and Doty, 1975:137.

PHILIPPINE DISTRIBUTION.—SULU: Sulu (North Ubian I., Siasi I.), Tawitawi (Pearl Bank, Sangasiapu I.).

*Codium papillatum Tseng and Gilbert

Codium papillatum Tseng and Gilbert, 1942:293, figs. 2b–d, 3 [type locality: Puerto Galera, Oriental Mindoro Prov.].—Gilbert, 1947:124; 1961:443.—Galutira and Velasquez, 1964:497, pl. 2: fig. 4; pl. 6: figs. 23a,b.—Velasquez, 1972:63.—Velasquez, Trono, and Doty, 1975:137.—Velasquez, 1979b:230.—Trono and Fortes, 1980:64.—Ganzon-Fortes, 1981:21.—Cordero, 1982a:60, fig. 4.—Trono and Fortes, 1982:145.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Bataan, Cavite. MINDORO: Oriental Mindoro.

Codium platyclados Jones and Kraft

Codium platyclados Jones and Kraft, 1984:266, figs. 8–13 [including Philippine record, page 273, figs. 10B, 12C] [type locality: Lord Howe I.].

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon (San Bernardino I.).

Codium pugniforme Okamura

Codium pugniforme Okamura, 1915:147, pl. CXXXV: figs. 6–9 [“pugniformis”] [syntype localities: various, all in Japan].—Cordero, 1973b:20 [C. pugniforme prox.].—Puig and Cordero, 1979:26.

PHILIPPINE DISTRIBUTION.—LEYTE (Biliran I.).

Codium tenue auctorum

Codium tenue [misapplied name].—Gepp and Gepp, 1911:136.—Seale, 1911:309.—Wester, 1916:158.—Weber-van Bosse, 1913a:120.—Wester, 1921:224; 1924:21.—G. Blanco, 1938:512.—Gilbert, 1946:78.—Zaneveld, 1950:112.—Quisumbing, 1951:1009.—Montilla and Blanco, 1953:166, fig. 5:5.—Zaneveld, 1956:10; 1959:96.—Gilbert, 1961:443.—Domantay, 1962:279.—Cordero, 1973b:20.—Velasquez et al., 1973:17, pl. 6: fig. 28.—Velasquez, Trono, and Doty, 1975:137.—Puig and Cordero, 1979:27.—Trono and Fortes, 1980:64.—Ganzon-Fortes, 1981:21.—Guzman, 1981:43.—Cordero, 1982a:60, fig. 3.—Trono and Fortes, 1982:145.—Cordero, 1984a:79.—Tungpalan, 1984:139.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, La Union, Pangasinan, Bataan. LEYTE (Biliran I.). SULU: Sulu (North Ubian I.).

NOTE.—Codium tenue (Kützing) Kützing (1856:33, pl. 95: fig. 1) appears to be restricted to estuaries of South Africa (cf. Silva, 1959:140–143). Placement of Philippine records awaits completion of a monographic study.

Codium tomentosum auctorum

Codium tomentosum [misapplied name].—Montagne, 1844a:659.—Martens, 1868:47, 64–65.—Dickie, 1876a:243.—Piccone, 1886:29, 89.—Schmidt, 1923:39.—Zaneveld, 1950:112; 1956:10; 1959:96.—Meñez, 1961:54, pl. 6: figs. 60–62.—Reyes, 1972:143.—Cordero, 1973b:20.—Velasquez, Trono, and Doty, 1975:137.—Reyes, 1978:157, 162, pl. 9: figs. 11, 12.—Liao and Sotto, 1980:97.—Trono and Fortes, 1980:64.

Codium dichotomum [misapplied name].—Gilbert, 1946:78; 1961:442.—Velasquez, Trono, and Doty, 1975:137.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. MASBATE (Ticao I.). LEYTE (Biliran I.). NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR.

NOTE.—Codium dichotomum S.F. Gray (1821:293) is a nomenclatural synonym of Codium tomentosum Stackhouse (1797 [1795–1801]:xxiv), a name that applies to a species restricted to the eastern North Atlantic (cf. Silva, 1955:569–571). Placement of the Philippine records awaits completion of a monographic study.

Family HALIMEDACEAE

Halimeda Lamouroux

Halimeda bikinensis W.R. Taylor

Halimeda bikinensis W.R. Taylor, 1950:87, 207, pl. 48: fig. 1 [type locality: Bikini Atoll, Marshall Is.].—Cordero and Tanaka, 1972:83, figs. 1A, 2E,F.—Saraya and Trono, 1980:22, pl. V: fig. 2.—Marcos-Agngarayngay, 1983:89, fig. 22.—Cordero, 1984a:80.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Pangasinan.

Halimeda copiosa Goreau and Graham

Halimeda copiosa Goreau and Graham, 1967:433, figs. 1–10 [type locality: Jamaica].

Halimeda opuntia (Linnaeus) Lamouroux f. hederacea Barton, 1901:21, pl. III: fig. 23 [lectotype locality: Tanah-Djampeah, Flores Sea, Indonesia fide Colinvaux, 1968:32].—Gilbert, 1947:130; 1961:444.

Halimeda opuntia (Linnaeus) Lamouroux var. hederacea (Barton) Hillis, 1959:360, pl. 5: fig. 4 [including Philippine records].—Cordero, 1977b:40.

Halimeda hederacea (Barton) Colinvaux, 1968:30, figs. 1, 2(1,4,5,6,8) [including Philippine record].

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Quezon. MINDANAO: Zamboanga. BASILAN. SULU: Sulu (near Lapac I.).

NOTE.—The synonymy was proposed by Hillis-Colinvaux (1980:118).

Halimeda cuneata Hering

Halimeda cuneata Hering in Krauss, 1846:214 [type locality: “Natalbai” (Durban), South Africa].—Martens, 1868:25, 66–67.—Meñez, 1961:58, pl. 4: figs. 43–46; pl. 5: figs. 54, 55.—Cordero, 1973b:21.—Velasquez, Trono, and Doty, 1975:147.—Cordero, 1977b:39, pl. VI: fig. 30; 1979b:281.—Puig and Cordero, 1979:27.—Cordero, 1980b:31, fig. 1B, pl. 19.—Marcos-Agngarayngay, 1983:90, fig. 23.—Cordero, 1984a:80 [with query].—Tungpalan, 1984:138.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan. SAMAR: Western Samar, Eastern Samar. LEYTE (Biliran I.). PANAY: Aklan. SIQUIJOR. MINDANAO: Zamboanga, Surigao del Sur.

Halimeda cylindracea Decaisne

Halimeda cylindracea Decaisne, 1842b:103 [type locality: Nosy-bé, Malagasy Republic].—Hillis, 1959:373, pl. 4: fig. 3; pl. 5: figs. 22, 23; pl. 6: fig. 19; pl. 7: fig. 13.—Taylor, 1966b:353 [“cylindrica”].—Trono, 1972a:97.—Cordero, 1973a:167, figs. 1–4.—Trono, 1973b:225; 1975:36.—Velasquez, Trono, and Doty, 1975:147 [“cylindracea” and “cylindrica”].—Trono and Ganzon-Fortes, 1980:25, fig. [s.n].—Trono and Ang, 1982:11.

Halimeda monile [misapplied name fide Hillis, 1959:374].—Gilbert, 1947:130; 1961:443.—Velasquez, Trono, and Doty, 1975:148.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas, Quezon, Sorsogon. SAMAR: Eastern Samar. PALAWAN (Bugsuk I., Culion I.). SULU: Sulu (Siasi I.), Tawitawi.

Halimeda discoidea Decaisne

Halimeda discoidea Decaisne, 1842b:102 [type locality: said to be “Kamtschatka” but true provenance unknown].—Martens, 1868:25, 66–67.—Gilbert, 1946:78; 1947:125 [f. typica].—Hillis, 1959:352, pl. 2: fig. 5; pl. 5: fig. 11; pl. 6: fig. 11; pl. 7: figs. 9, 10; pl. 8: figs. 5–8.—Gilbert, 1961:443 [incl. f. discoidea].—Meñez, 1961:57, pl. 4: figs. 47–50.—Taylor, 1966b:353 [var. discoidea].—Reyes, 1972:143.—Trono, 1972a:97.—Cordero, 1973b:21.—Trono, 1973b:225; 1973c:15, figs. 7, 11; 1973d:8, pl. 6: figs. 23, 24.—Ortega, Alcala, and Reyes, 1974:186, 187, 188.—Velasquez, Trono, and Doty, 1975:147.—Trono and Young, 1977:55.—Reyes, 1978:159, pl. II: figs. 5, 6.—Trono, 1978:6.—Trono and Tuason, 1978:4.—Garcia, 1979:44 (table 1).—Puig and Cordero, 1979:27.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:22, pl. IV: figs. 3, 4.—Hurtado-Ponce, 1983:117.—Hurtado-Ponce and Modelo, 1983:149.—Cordero, 1984c:52.—Tungpalan, 1984:138.

*Halimeda discoidea Decaisne f. intermedia Gilbert, 1947:126 [type locality: Dalupiri I., Babuyan Is., Cagayan Prov., Luzon].—Gilbert, 1961:443.

*Halimeda discoidea Decaisne f. subdigitata Gilbert, 1947:125 [type locality: Siasi Island, Sulu Prov., Sulu Archipelago].—Gilbert, 1961:443.—Taylor, 1966b:353 [var. discoidea f. subdigitata].

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Pangasinan. CATANDUANES. MINDORO: Oriental Mindoro. MARINDUQUE. MASBATE. SAMAR: Eastern Samar. LEYTE (Biliran I.). NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga. BASILAN. PALAWAN (incl. Balabac I.). SULU: Sulu (incl. Cagayan Sulu I., Siasi I.), Tawitawi (incl. Pearl Bank, Sibutu I.).

NOTE.—Hillis (1959:354) does not give taxonomic recognition to the formae of this species.

Halimeda fragilis W.R. Taylor

Halimeda fragilis W.R. Taylor, 1950:88, 207, pl. 48: fig. 2 [type locality: Eniwetok Atoll, Marshall Is.].—Cordero, 1976c:6; 1977b:39, pl. VI: fig. 34.—Trono and Young, 1977:56.—Trono and Tuason, 1978:5.—Saraya and Trono, 1980:23.—Meñez and Calumpong, 1981:383.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. CATANDUANES. CENTRAL VISAYAS.

Halimeda gigas W.R. Taylor

Halimeda gigas W.R. Taylor, 1950:84, 206, pl. 44: fig. 2 [type locality: Eniwetok Atoll, Marshall Is.].—Domantay, 1962:280.—Velasquez, Trono, and Doty, 1975:147.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Halimeda gracilis Harvey ex J. Agardh

Halimeda gracilis Harvey ex J. Agardh, 1887:82 [type locality: Sri Lanka].—Gilbert, 1946:78; 1961:443.—Velasquez, Trono, and Doty, 1975:147.—Hurtado-Ponce, 1983:117.—Hurtado-Ponce and Modelo, 1983:149.—Cordero, 1984a:80; 1984b:60.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte. PALAWAN.

Halimeda incrassata (Ellis) Lamouroux

Corallina incrassata Ellis, 1768:408, pl. XVII: figs. 20–27 [type locality: West Indies].

Halimeda incrassata (Ellis) Lamouroux, 1816:307.—Hillis, 1959:365, pl. 4: figs. 1, 2; pl. 5: fig. 21; pl. 6: figs. 21–24.—Gilbert, 1961:444.—Velasquez, 1971:437, fig. 17.—Trono, 1972a:98.—Velasquez, Trono, and Doty, 1975:148.—Saraya and Trono, 1980:23, pl. V: fig. 3.

Corallina tridens Ellis and Solander, 1786:109, pl. 20: fig.a [type locality: “coast of the new ceded Islands” (probably Dominica)].

Halimeda tridens (Ellis and Solander) Lamouroux, 1816:308.—Gilbert, 1947:130 [forma typica].—Velasquez, Trono, and Doty, 1975:148.

Halimeda incrassata (Ellis and Solander) Lamouroux f. lamourouxii J. Agardh, 1887:86 [syntype localities: West Indies; Australia].—Gilbert, 1961:444.

Halimeda tridens (Ellis and Solander) Lamouroux f. lamourouxii (J. Agardh) Gilbert, 1947:131.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Quezon. MINDANAO: Zamboanga. BASILAN. PALAWAN (incl. Culion I., Cuyo I.). SULU: Sulu (North Ubian I., Siasi I.), Tawitawi.

NOTE.—The synonymy was proposed by Hillis (1959:365).

Halimeda lacunalis W.R. Taylor

Halimeda lacunalis W.R. Taylor, 1950:91, 208, pl. 51 [type locality: Eniwetok Atoll, Marshall Is.].—Meñez and Calumpong, 1981:382.

PHILIPPINE DISTRIBUTION.—CENTRAL VISAYAS.

Halimeda macroloba Decaisne

Halimeda macroloba Decaisne, 1841:118 [type locality: Red Sea].—Montagne, 1844a:659.—Dickie, 1876a:243.—Barton, 1901:24, pl. III: figs. 33–38.—Weber-van Bosse, 1913a:122.—Gilbert, 1946:78; 1947:126.—Hillis, 1959:375, pl. 3: fig. 3; pl. 5: figs. 19, 20; pl. 6: fig. 17.—Gilbert, 1961:443.—Domantay, 1962:280.—Moul, 1964:6.—Taylor, 1966b:353.—Cordero and Tanaka, 1972:85, figs. 1B, 2C,D.—Cornejo and Velasquez, 1972:172, 179, 183.—Reyes, 1972:144.—Trono, 1972a:98; 1973b:226; 1973c:15, figs. 10, 12; 1973d:9.—Velasquez et al., 1973:17, pl. 6: fig. 27.—Ortega, Alcala, and Reyes, 1974:186.—Trono, 1975:37.—Velasquez, Trono, and Doty. 1975:148.—Trono and Young, 1977:56.—Reyes, 1978:159, pl. II: figs. 1, 2.—Trono, 1978:7.—Trono and Tuason, 1978:5.—Cordero, 1979b:281.—Puig and Cordero, 1979:28.—Cordero, 1980b:32, fig. 2B, pl. 20.—Fortes and Trono, 1980:68.—Saraya and Trono, 1980:20, pl. V: fig. 1.—Trono and Ganzon-Fortes, 1980:27, fig. [s.n].—Meñez and Calumpong, 1981:383.—Trono and De Lara, 1981:6, pl. IV: fig. 1.—Trono and Ang, 1982:11.—Hurtado-Ponce, 1983:118.—Hurtado-Ponce and Modelo, 1983:149.—Marcos-Agngarayngay, 1983:90, fig. 24.—Cordero, 1984a:81.—Cordero, 1984b:60.—Cordero, 1984c:52.—Tungpalan, 1984:138.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (incl. Babuyan Is.), Ilocos Norte, La Union, Pangasinan, Bataan, Batangas, Quezon, Camarines Norte, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. MASBATE. SAMAR: Eastern Samar. LEYTE (Biliran I.). PANAY: Aklan. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. BOHOL. MINDANAO: Zamboanga, Surigao del Sur. BASILAN. PALAWAN (incl. Balabac I., Bugsuk I., Culion I.). SULU: Sulu (Siasi I.), Tawitawi (incl. Pearl Bank, Sibutu I., Turtle Is.).

Halimeda macrophysa Askenasy

Halimeda macrophysa Askenasy, 1888:14, pl. IV: figs. 1–4 [type locality: Makutu I., Fiji].—Taylor, 1966b:353.—Trono, 1975:37.—Velasquez, Trono, and Doty, 1975:148.—Trono and Young, 1977:56.—Trono and Tuason, 1978:6.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon. CATANDUANES. SULU: Tawitawi (Turtle Is.).

Halimeda micronesica Yamada

Halimeda micronesica Yamada, 1941b:121, fig. 15 [type locality: Ant Atoll, near Ponape, Caroline Is.].—Hillis, 1959:364, pl. 3: fig. 1; pl. 5: fig. 13, 14; pl. 6: fig. 2.—Gilbert, 1961:444.—Taylor, 1966b:353.—Velasquez, Trono, and Doty, 1975:148.

*Halimeda orientalis Gilbert, 1947:126, fig. 1 [type locality: Tumakid, Basilan I.].

PHILIPPINE DISTRIBUTION.—BASILAN. SULU: Tawitawi.

NOTE.—The synonymy was proposed by Hillis (1959:364).

Halimeda opuntia (Linnaeus) Lamouroux

Corallina opuntia Linnaeus, 1758:805 [lectotype locality: Jamaica fide Hillis, 1959:361].

Halimeda opuntia (Linnaeus) Lamouroux, 1816:308.—Martens, 1868:25, 64–65.—Dickie, 1876a:243, 245 [f. typica].—Barton, 1901:18, pl. II: figs. 19–21.—Weber-van Bosse, 1913a:121 [f. typica].—Merrill, 1918:39.—Gilbert, 1946:78.—Gilbert, 1947:129 [f. typica].—Hillis, 1959:359, pl. 2: figs. 7, 8; pl. 5: fig. 3; pl. 6: fig. 6; pl. 7: fig. 3 [var. opuntia].—Gilbert, 1961:443 [incl. f. opuntia].—Meñez, 1961:59, pl. 5: figs. 51–53, 56, 57.—Domantay, 1962:280.—Moul, 1964:6 [var. opuntia].—Taylor, 1966b:353.—Villones and Magdamo, 1968:25, fig. 13.—Velasquez, 1971:437, fig. 16.—Reyes, 1972:144.—Trono, 1972a:98.—Cordero, 1973b:21.—Trono, 1973b:226; 1973c:16, fig. 9; 1973d:9.—Velasquez et al., 1973:16, pl. 5: fig. 24.—Westernhagen, 1973a:65.—Ortega, Alcala, and Reyes, 1974:185, 186, 187.—Trono, 1975:37.—Velasquez, Trono, and Doty, 1975:148.—Cordero, 1976c:8, 9, 10; 1977b:40, pl. VI: figs. 31–33 [var. opuntia].—Taylor, 1977b:10.—Trono and Young, 1977:55.—Reyes, 1978:158, pl. 10: figs. 5, 6.—Trono, 1978:7.—Trono and Tuason, 1978:6.—Puig and Cordero, 1979:28.—Calumpong, 1980:143 (table 3).—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:21, pl. VI: fig. 1.—Trono and Ganzon-Fortes, 1980:29, fig. [s.n].—Chan, 1981:387.—Meñez and Calumpong, 1981:383.—Trono and De Lara, 1981:6, pl. III: fig. 3. Trono and Ang, 1982:10.—Hurtado-Ponce, 1983:118.—Hurtado-Ponce and Modelo, 1983:149.—Cordero, 1984a:81; 1984b:60.

*Fucus prolifer M. Blanco, 1837:838 [type locality: “Punta Santiago” (Batangas Prov. ?)].—M. Blanco, 1845:579; 1879:259.—Velasquez, Trono, and Doty, 1975:144.

Halimeda triloba Decaisne, 1842:102 [lectotype locality: China Sea fide Barton, 1901 explanation of pl. II: fig. 20].—Bailey and Harvey, 1862:173.—Martens, 1868:64–65.—Velasquez, Trono, and Doty, 1975:149.

Halimeda opuntia (Linnaeus) Lamouroux f. triloba (Decaisne) J. Agardh, 1887:84.—Gilbert, 1947:129.

Halimeda cordata J Agardh, 1887:83 [type locality: Red Sea].

Halimeda opuntia (Linnaeus) Lamouroux f. cordata (J. Agardh) Barton, 1901:20, pl. II: fig. 21.—Gilbert, 1947:129; 1961:444.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, La Union, Pangasinan, Batangas, Quezon, Camarines Norte, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SAMAR: Eastern Samar. LEYTE (Biliran I.). GUIMARAS. NEGROS: Negros Occidental, Negros Oriental. CEBU (incl. Mactan I.). SIQUIJOR. BOHOL. MINDANAO: Zamboanga (incl. Sacol I.), Misamis Occidental, Davao. BASILAN. PALAWAN (incl. Balabac I., Bugsuk I. Culion I., Cuyo I.). SULU: Sulu (North Ubian I., Siasi I.), Tawitawi.

NOTE.—Hillis-Colinvaux (1980:110) does not give taxonomic recognition to formae cordata and triloba. According to Merrill (1918:39), Blanco’s description of Fucus prolifer is referable to Halimeda opuntia, but Martens (1868:45) thought that Blanco could have had other species in hand. In the absence of authentic specimens, a definitive placement is impossible. In any event, Fucus prolifer M. Blanco is a later homonym of F. prolifer Forsskål (1775:193) and F. prolifer Lightfoot (1777:949, pl. XXX).

Halimeda renschii Hauck

Halimeda renschii Hauck, 1886 [1886–1887]:167 [type locality: Johanna (Anjouan) I., Comoro Is.].—Hillis-Colinvaux, 1975:93.

Halimeda opuntia (Linnaeus) Lamouroux f. renschii (Hauck) Barton, 1901:21, pl. II: figs. 22, 22a.—Gilbert, 1947:130; 1961:444.

*Halimeda batanensis W.R. Taylor, 1973:34, figs. 1, 2 [type locality: Batan I., Batanes Prov., Luzon].

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.).

NOTE.—The synonymy was proposed by Hillis-Colinvaux (1975:93).

Halimeda simulans Howe

Halimeda simulans Howe, 1907:503, pl. 29 [type locality: Culebra I., Puerto Rico].—Trono, 1972a:94; 1975:38.—Trono and Ganzon-Fortes, 1980:31, fig. [s.n].—Trono and Ang, 1982:10.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas, Sorsogon. PALAWAN (Bugsuk I.). SULU: Tawitawi.

Halimeda stuposa W.R. Taylor

Halimeda stuposa W.R. Taylor, 1950:90, 207, pl. 43: fig. 1; pl. 49; pl. 50: fig. 2 [type locality: Rongelap Atoll, Marshall Is.].—Reyes, 1978:159, pl. II: figs. 3, 4.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Halimeda taenicola W.R. Taylor

Halimeda taenicola W.R. Taylor, 1950:86, 207, pl. 46: fig. 1 [type locality: Rongerik Atoll, Marshall Is.].—Trono and Young, 1977:56.—Trono and Tuason, 1978:7.

PHILIPPINE DISTRIBUTION.—CATANDUANES.

Halimeda tuna (Ellis and Solander) Lamouroux

Corallina tuna Ellis and Solander, 1786:111, pl. 20: fig.e [type locality Mediterranean Sea].

Halimeda tuna (Ellis and Solander) Lamouroux, 1816:309, pl. XI: fig 8.—Dickie, 1876a:244.—Gilbert, 1946:78; 1947:124 [f. typica].—Hillis, 1959:342, pl. 1: figs. 4, 5; pl. 5: fig. 9; pl. 6: fig. 7.—Gilbert 1961:444.—Meñez, 1961:57.—Domantay, 1962:280 [f. typica].—Taylor, 1966b:354 [var. tuna].—Trono, 1972a:98; 1973b:226; 1973c:17 fig. 21.—Velasquez et al., 1973:16, pl. 6: fig. 26.—Westernhagen 1973a:65.—Hamoy and Garciano, 1975:71.—Trono, 1975:38.—Velasquez, Trono, and Doty, 1975:149.—Cordero, 1976c:6; 1977b:41 pl. VI: fig. 37.—Trono and Young, 1977:55.—Reyes, 1978:160, pl 11: fig. 7.—Trono, 1978:7.—Trono and Tuason, 1978:7.—Cordero 1979b:276; 1980b:33, fig. 2B.—Saraya and Trono, 1980:21, pl. V: fig 4.—Trono and Fortes, 1980:64.—Trono and Ganzon-Fortes, 1980:33, fig. [s.n].—Chan, 1981:387.—Ganzon-Fortes, 1981:21.—Meñez and Calumpong, 1981:383.—Trono and Fortes, 1982:145.—Cordero, 1984a:81.—Tungpalan, 1984:138.

Halimeda platydisca Decaisne, 1842:102 [type locality: Islas Canarias].

Halimeda tuna (Ellis and Solander) Lamouroux f. platydisca (Decaisne) Barton, 1901:14, pl. I: fig. 2.—Gilbert, 1947:124; 1961:444.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Bataan, Batangas, Sorsogon. CATANDUANES. MINDORO: Oriental Mindoro. MARINDUQUE. PANAY: Aklan. CEBU (incl. Mactan I.). SIQUIJOR. MINDANAO: Zamboanga, Surigao del Sur. BASILAN. PALAWAN (Culion I.). SULU: Sulu (Siasi I.), Tawitawi.

NOTE.—Hillis-Colinvaux (1980:122) does not give taxonomic recognition to f. platydisca.

*Halimeda velasquezii W.R. Taylor

Halimeda velasquezii W.R. Taylor, 1962:177, figs. 8–14 [type locality: Santa Ana, Cagayan Prov., Luzon].—Taylor, 1966b:354.—Velasquez et al., 1973:16, pl. 5: fig. 25.—Trono, 1975:39.—Velasquez, Trono, and Doty, 1975:149.—Cordero, 1976c:6, 8; 1977b:41, pl. VI: figs. 35, 36; pl. VII: fig. 38.—Hillis-Colinvaux, 1980:117, fig. 32.

Halimeda opuntia (Linnaeus) Lamouroux f. intermedia Yamada, 1934:81 [syntype localities: Naha and Itoman, Ryukyu-retto, Japan].—Gilbert, 1947:130; 1961:444.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan, Bataan, Batangas, Sorsogon.

NOTE.—The synonymy was proposed by Hillis-Colinvaux (1980:117).

*Halimeda velasquezii W.R. Taylor var. robusta Cordero

*Halimeda velasquezii W.R. Taylor var. robusta Cordero, 1974a:115, fig. 1 [type locality: Camiguin I., Babuyan Is., Cagayan Prov., Luzon].—Cordero and Tanaka, 1972:86, figs, 1C, 2A,B [H. velasquezii].

PHILIPPINE DISTRIBUTION.—As above.

Family UDOTEACEAE

Avrainvillea Decaisne

*Avrainvillea erecta (Berkeley) A. Gepp and E.S. Gepp

Dichonema erectum Berkeley, 1842:157, pl. VII: fig. 11 [type locality: “Philippine Islands”].

Avrainvillea erecta (Berkeley) A. Gepp and E.S. Gepp, 1911:29, pl. IX: figs. 84, 85; pl. X: figs. 86–89 [including Philippine records].—Gilbert, 1946:78; 1947:122; 1961:442.—Meñez, 1961:56, pl. 3: figs. 26–30.—Domantay, 1962:276.—Taylor, 1966b:352.—Gilbert and Doty, 1969:123, figs. 7, 8.—Reyes, 1972:143.—Trono, 1973b:220; 1973c:18.—Velasquez et al., 1973:15, pl. 5: fig. 22.—Velasquez, Trono, and Doty, 1975:128.—Reyes, 1978:160, pl. II: fig. 8.—Puig and Cordero, 1979:25.—Saraya and Trono, 1980:19, pl. III: fig. 3.—Meñez and Calumpong, 1981:383.—Marcos-Agngarayngay, 1983:94, fig. 26.—Olsen-Stojkovich, 1985a:22.

*Udotea sordida Montagne, 1844a:659 [type locality: “Philippines”].—Kützing, 1849:503.—Montagne, 1856:451.—Martens, 1868:64–65.—Dickie, 1874a:198.—Velasquez, Trono, and Doty, 1975:167.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Bataan. CATANDUANES. MINDORO: Oriental Mindoro. SAMAR: Eastern Samar. LEYTE (Biliran I.). NEGROS: Negros Oriental. SIQUIJOR. BASILAN. PALAWAN (incl. Balabac I., Cuyo I.). SULU: Sulu (Siasi I.).

NOTE.—The synonymy was proposed by Gepp and Gepp (1911:29).

Avrainvillea lacerata Harvey ex J. Agardh

Avrainvillea lacerata Harvey ex J. Agardh, 1887:54 [type locality: Tonga].—Taylor, 1966b:352.—Trono, 1972a:97; 1973b:220; 1973d:10, pl. 5: fig. 17.—Velasquez, Trono, and Doty, 1975:129.—Reyes, 1978:160, pl. II: fig. 9.—Trono, 1978:6.—Trono and Tuason, 1978:4.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:18, pl. III: fig. 2.—Trono and Ang, 1982:9.—Olsen-Stojkovich, 1985a:33.

Avrainvillea lacerata Harvey ex J. Agardh var. robustior A. Gepp and E.S. Gepp, 1911:139, pl. XIII: figs. 108, 109 [type locality: Singapore].—Domantay, 1962:276 [f. robustior].—Meñez and Calumpong, 1981:383.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CATANDUANES. MARINDUQUE. SAMAR: Eastern Samar. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga (incl. Sacol I.). PALAWAN (incl. Bugsuk I.). SULU: Sulu (Siasi I.), Tawitawi.

NOTE.—Olsen-Stojkovich (1985a) does not give taxonomic recognition to var. robustior.

Avrainvillea longicaulis (Kützing) Murray and Boodle

Rhipilia longicaulis Kützing, 1858:13, pl. 28: fig. II [type locality: Antilles].

Avrainvillea longicaulis (Kützing) Murray and Boodle, 1889:70.—Olsen-Stojkovich, 1985a:29.

Avrainvillea levis Howe, 1905:565, pl. 23: fig. 1; pl. 26: figs. 8–10 [type locality: Cave Cays, Exuma Chain, Bahama Is.].

Avrainvillea sordida [misapplied name; see Note].—Meñez, 1961:56, pl. 3: figs. 35–39.—Gilbert and Doty, 1969:124, figs. 13, 14.—Trono, 1973b:221.—Velasquez, Trono, and Doty, 1975:129.—Cordero, 1976c:9; 1977b:36, pl. V: fig. 26.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan. CATANDUANES. SULU: Sulu (Siasi I.).

NOTE.—The taxonomic and nomenclatural complexities of this species are discussed by Olsen-Stojkovich (1985a:29). The conspecificity of Rhipilia longicaulis and Avrainvillea levis was suspected by Gepp and Gepp (1911:40), who, however, were reluctant to use the binomial A. longicaulis because Murray and Boodle, when making the combination, misapplied it to representatives of A. nigricans. Retention of A. longicaulis is required by Article 55.2 of the ICBN.

Avrainvillea nigricans Decaisne

Avrainvillea nigricans Decaisne, 1842:108 [type locality: West Indies near Guadeloupe].—Liao and Sotto, 1980:97.—Olsen-Stojkovich, 1985a:38.

PHILIPPINE DISTRIBUTION.—CEBU (Mactan I.).

Avrainvillea obscura (C. Agardh) J. Agardh

Anadyomene ? obscura C. Agardh, 1822a:401 [type locality: Guam, Mariana Is.].

Avrainvillea obscura (C. Agardh) J. Agardh, 1887:53.—Meñez, 1961:55, pl. 3: figs. 30–34.—Domantay, 1962:272, 273.—Gilbert and Doty, 1969:124.—Trono, 1972a:97.—Velasquez, Trono, and Doty, 1975:129.—Cordero, 1977b:36, pl. IV: figs. 21, 22.—Olsen-Stojkovich, 1985a:19.

*Avrainvillea capituliformis Tanaka, 1967:14, figs. 2, 3, pl. I:B [type locality: San Pio Quinto, Camiguin I., Babuyan Is., Cagayan Prov., Luzon].—Trono, 1973c:18; 1975:35.—Velasquez, Trono, and Doty, 1975:128.—Puig and Cordero, 1979:25.—Saraya and Trono, 1980:19.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Pangasinan, Sorsogon. CATANDUANES. MINDORO: Oriental Mindoro. LEYTE (Biliran I.). SULU: Tawitawi.

NOTE.—The synonymy was proposed by Olsen-Stojkovich (1985a).

Boodleopsis A. Gepp and E.S. Gepp

Boodleopsis pusilla (Collins) W.R. Taylor, Joly, and Bernatowicz

Dichotomosiphon pusillus Collins, 1909:431 [type locality: West Indies].

Boodleopsis pusilla (Collins) W.R. Taylor, Joly, and Bernatowicz, 1953:105.—Fortes and Trono, 1980:56, fig. 5.—Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Boodleopsis verticillata Dawson

Boodleopsis verticillata Dawson, 1960:32, figs. 1A–D [type locality: Isla del Rey, Islas Perlas, Panama].—Fortes and Trono, 1980:58, fig. 6. Fortes, 1981b:396.

PHILIPPINE DISTRIBUTION.—MINDORO: Oriental Mindoro.

Chlorodesmis Harvey and Bailey

Chlorodesmis caespitosa J. Agardh

Chlorodesmis caespitosa J. Agardh, 1887:49 [type locality: Sri Lanka].—Ducker, 1967:157, pls. 27, 36–38; pl. 43: fig. 2.

Chlorodesmis formosana Yamada, 1925a:92, fig. V [syntype localities: various, all in Taiwan].—Gilbert, 1947:122; 1961:422.—Velasquez, Trono, and Doty, 1975:135.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.).

NOTE.—The synonymy was proposed by Ducker (1967:157).

Chlorodesmis fastigiata (C. Agardh) Ducker

Vaucheria fastigiata C. Agardh, 1824:176 [type locality: Mariana Is.].

Chlorodesmis fastigiata (C. Agardh) Ducker, 1969:17.

Chlorodesmis comosa Harvey and Bailey, 1851:373 [type locality: Fiji].—Gilbert, 1947:122; 1961:442.—Meñez, 1961:55, pl. 2: figs. 20, 21.—Domantay, 1962:279.—Taylor, 1966b:352.—Ducker, 1967:160, pl. 28, 29, 39.—Gilbert and Doty, 1969:123.—Velasquez et al., 1973:13, pl. 3: fig. 14.—Trono, 1975:36.—Velasquez, Trono, and Doty, 1975:134.—Cordero, 1976c:6, 8; 1977b:35, pl. IV: figs. 18, 19.—Trono, 1978:6.—Trono and Tuason, 1978:4.—Cordero, 1979b:276.—Puig and Cordero, 1979:25.—Saraya and Trono, 1980:19, pl. IV: fig. 1.—Chan, 1981:387.—Trono and Ang, 1982:9.—Hurtado-Ponce, 1983:107.—Hurtado-Ponce and Modelo, 1983:146.—Marcos-Agngarayngay, 1983:86, fig. 21.—Cordero, 1984a:79; 1984b:59.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Pangasinan, Bataan, Quezon, Sorsogon. CATANDUANES. MARINDUQUE. LEYTE (Biliran I.). PANAY: Aklan. MINDANAO: Zamboanga, Davao. BASILAN. PALAWAN (incl. Bugsuk I., Cuyo I.)

NOTE.—The conspecificity of Vaucheria fastigiata and Chlorodesmis comosa was suggested by Gepp and Gepp (1911:15) and confirmed by Ducker (1969:17).

Chlorodesmis hildebrandtii A. Gepp and E.S. Gepp

Chlorodesmis hildebrandtii A. Gepp and E.S. Gepp, 1911:16, 137, pl. 8: figs. 74, 75 [lectotype locality: Johanna (Anjouan) I., Comoro Is. fide Ducker, 1967:165].—Gilbert, 1947:122; 1961:442.—Taylor, 1966b:352.—Tanaka, 1967:16, fig. 4, pl. II:C—Trono, 1973c:18.—Velasquez, Trono, and Doty, 1975:135.—Cordero, 1977b:35, pl. V: figs. 23–25.—Reyes, 1978:158, pl. 10: fig. 3.—Meñez and Calumpong, 1981:383.—Trono and De Lara, 1981:5. pl. III: fig. 4.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.). MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. SIQUIJOR. SULU: Tawitawi (Sibutu I.).

Chlorodesmis major Zanardini

Chlorodesmis major Zanardini, 1874:504 [type locality: Lord Howe I.].

Chlorodesmis torresiensis W.R. Taylor, 1945:66 [type locality: Murray Is., Torres Strait, Australia].—Taylor, 1966b:352.—Velasquez, Trono, and Doty, 1975:135.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan.

NOTE.—The synonymy was proposed by Ducker (1967:167).

Rhipiliopsis A. Gepp and E.S. Gepp

*Rhipiliopsis carolyniae Kraft

Rhipiliopsis carolyniae Kraft, 1986:51, figs. 8–11 [type locality: Bulusan, Sorsogon Prov., Luzon].

Rhipiliopsis peltata [misapplied name.—Gilbert and Doty, 1969:122, figs. 9–12.—Velasquez, Trono, and Doty, 1975:161.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon. MINDANAO: Surigao.

Tydemania Weber-van Bosse

Tydemania expeditionis Weber-van Bosse

Tydemania expeditionis Weber-van Bosse, 1901:139 [syntype localities: Macassar, Celebes and Sumbawa I., Indonesia].—Gilbert, 1947:122; 1961:444.—Taylor, 1966b:352.—Velasquez, 1971:432, fig. 10.—Trono, 1973c:17, fig. 15.—Trono, 1973d:10, pl. 6: fig. 22; 1975:40.—Velasquez, Trono, and Doty, 1975:166.—Trono and Young, 1977:55.—Reyes, 1978:158, pl. 10: fig. 4.—Trono and Tuason, 1978:8.—Cordero, 1979c:53, 57, 61, figs. 2, 3.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:24, pl. VI: fig. 4.—Meñez and Calumpong, 1981:383.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Sorsogon. CATANDUANES. MINDORO: Oriental Mindoro. SAMAR: Eastern Samar. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. MINDANAO: Zamboanga. PALAWAN (Balabac I.). SULU: Sulu (Siasi I.), Tawitawi.

Udotea Lamouroux

Farghaly (1980:245–262), in his treatment of Udotea, resurrected the genus Rhipidosiphon Montagne (1842a:14) and recognized three new segregate genera: Decaisnella, Geppina, and Howella. The new generic names are invalid because they lack Latin diagnoses. The species of Udotea in this catalog are distributed by Farghaly as follows:Udotea (U. argentea, U. flabellum, U. occidentalis); Decaisnella (U. indica); and Rhipidosiphon (U. glaucescens, U. javensis, U. orientalis). The generic placement of U. geppii was not indicated.

Udotea argentea Zanardini

Udotea argentea Zanardini, 1858:290, pl. XII: fig. 1 [type locality: Suez, Egypt].—Trono, 1972a:98.—Cordero, 1976c:8 [f. typica]; 1977b:37 [f. typica]; 1979b:278.—Trono and Ang, 1982:12.

PHILIPPINE DISTRIBUTION.—BATANES. PANAY: Aklan. PALAWAN (Bugsuk I.). SULU: Tawitawi.

*Udotea argentea Zanardini var. spumosa A. Gepp and E.S. Gepp

Udotea argentea Zanardini var. spumosa A. Gepp and E.S. Gepp, 1911:126, 144 [syntype localities: Tanah Djampeah and Saleyer (Selajar) I., Indonesia; Pearl Bank, Tawitawi Prov., Sulu Archipelago].—Weber-van Bosse, 1913a:117.—Gilbert, 1946:78; 1947:123; 1961:444.—Taylor, 1966b:352.—Gilbert and Doty, 1969:124, fig. 6.—Velasquez, Trono, and Doty, 1975:166 [without designation of forma].

PHILIPPINE DISTRIBUTION.—PALAWAN (incl. Balabac I., Culion I., Cuyo I.). SULU: Tawitawi (Pearl Bank, Turtle Is.).

Udotea flabellum (Ellis and Solander) Howe

Corallina flabellum Ellis and Solander, 1786:124, pl. 24 [type locality: West Indies].

Udotea flabellum (Ellis and Solander) Howe, 1904:94.—Gepp and Gepp, 1911:131, pl. III: figs. 26–28.—Weber-van Bosse, 1913a:118.—Gilbert, 1946:78; 1947:123; 1961:444.—Reyes, 1972:144.—Velasquez, Trono, and Doty, 1975:166.

PHILIPPINE DISTRIBUTION.—NEGROS: Negros Oriental. MINDANAO: Zamboanga. SULU: Sulu (North Ubian I.).

Udotea geppii Yamada

Udotea geppii Yamada, 1930a:141, figs. 1–3 [syntype localities: Palau (Belau), Caroline Is.; Tonga].—Reyes, 1978:161, pl. II: fig. 10.—Fortes and Trono, 1980:66.—Trono and Ang, 1982:12.

PHILIPPINE DISTRIBUTION.—SAMAR. SIQUIJOR. PALAWAN (Bugsuk I.).

Udotea glaucescens Harvey ex J. Agardh

Udotea glaucescens Harvey ex J. Agardh, 1887:70 [type locality: Tonga].—Gilbert and Doty, 1969:124.—Velasquez, Trono, and Doty, 1975:166.

PHILIPPINE DISTRIBUTION.—CATANDUANES.

Udotea indica A. Gepp and E.S. Gepp

Udotea indica A. Gepp and E.S. Gepp, 1911:121, pl. II: figs. 13, 14; pl. VI: figs. 52, 53 [type locality: Karachi, Pakistan].—Cordero, 1973b:22; 1976c:6, 11; 1977b:37, pl. IV: figs. 16, 17.—Puig and Cordero, 1979:29.—Cordero, 1984a:82.

Decaisnella indica (A. Gepp and E.S. Gepp) Farghaly, 1980:257 [including Philippine record].

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, LEYTE (Biliran I.).

NOTE.—The generic name Decaisnella was not validly published so that the binomial D. indica is also invalid.

Udotea javensis (Montagne) A. Gepp and E.S. Gepp

Rhipidosiphon javensis Montagne, 1842a:15 [type locality: Java, Indonesia].

Udotea javensis (Montagne) A. Gepp and E.S. Gepp, 1904:364.—Gilbert, 1947:122; 1961:445.—Villones and Magdamo, 1968:26, fig. 15 [figure cited but not published].—Gilbert and Doty, 1969:125.—Trono, 1973c:19.—Velasquez, Trono, and Doty, 1975:166.—Cordero, 1979c:55, 56, 61, fig. 1.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:23, pl. VI: fig. 3.—Meñez and Calumpong, 1981:383.—Cordero, 1984c:52.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Batangas. MINDORO: Oriental Mindoro. MASBATE. SAMAR: Western Samar, Eastern Samar. PANAY: Aklan. CEBU (Mactan I.). BASILAN. PALAWAN (incl. Cuyo I.).

Udotea occidentalis A. Gepp and E.S. Gepp

Udotea occidentalis A. Gepp and E.S. Gepp, 1911:127, 144, pl. II: figs. 18, 22a, b; pl. VII: figs. 63–65 [syntype localities: St. Thomas and St. Jan (St. John), Virgin Is.].—Trono, 1975:36.

PHILIPPINE DISTRIBUTION.—LUZON: Sorsogon.

*Udotea orientalis A. Gepp and E.S. Gepp

Udotea orientalis A. Gepp and E.S. Gepp, 1911:119, 142, pl. I: figs. 1, 4; pl. VI: figs. 47, 48 [syntype localities: various in Indian and Pacific oceans and Indonesia; North Ubian I., Sulu Prov., Sulu Archipelago].—Weber-van Bosse, 1913a:117.—Gilbert, 1946:78; 1947:122; 1961:445.—Meñez, 1961:56, pl. 2: figs. 22–25.—Taylor, 1966b:353.—Gilbert and Doty, 1969:125, fig. 4.—Reyes, 1972:144.—Cordero, 1973b:22.—Trono, 1973c:19, fig. 24.—Velasquez et al., 1973:15, pl. 5: fig. 23.—Velasquez, Trono, and Doty, 1975:167.—Cordero, 1976c:8; 1977b:38, pl. V: figs. 27–29.—Trono and Young, 1977:55.—Trono, 1978:8.—Trono and Tuason, 1978:8.—Cordero, 1979b:276.—Puig and Cordero, 1979:29.—Saraya and Trono, 1980:24, pl. VI: fig. 2.—Meñez and Calumpong, 1981:383.—Trono and De Lara, 1981:6, pl. IV: fig. 4.—Trono and Ang, 1982:12.—Hurtado-Ponce, 1983:119.—Hurtado-Ponce and Modelo, 1983:149.—Marcos-Agngarayngay, 1983:93, fig. 25.—Cordero, 1984a:82.

Rhipidosiphon orientalis (A. Gepp and E.S. Gepp) Farghaly, 1980:254 [including Philippine record].

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Pangasinan, Zambales, Bataan, Quezon, Albay, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. LEYTE (Biliran I.). PANAY: Aklan. NEGROS: Negros Oriental. MINDANAO: Surigao. PALAWAN (incl. Balabac I., Bugsuk I.). SULU: Sulu (North Ubian I.), Tawitawi (Turtle Is.).

Order DASYCLADALES

Family DASYCLADACEAE

Bornetella Munier-Chalmas

Bornetella nitida Munier-Chalmas ex Sonder

Bornetella nitida Munier-Chalmas ex Sonder, 1880:39 [type locality: Tonga].—Gilbert, 1943:25, fig. 1e,f; 1961:443.—Meñez, 1961:50, pl. 1: figs. 6–9.—Taylor, 1966b:347.—Valet, 1969:588.—Reyes, 1972:141.—Trono, 1972a:99.—Cordero, 1973b:18.—Trono, 1973a:128, fig. 2.—Ortega, Alcala, and Reyes, 1974:186, 188.—Trono, 1975:40.—Velasquez, Trono, and Doty, 1975:129.—Cordero, 1976c:8; 1977b:24, pl. II: figs. 7, 8.—Reyes, 1978:150, pl. 5: fig. 10.—Trono and Tuason, 1978:10.—Cordero, 1979b:276, 282.—Puig and Cordero, 1979:22.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:25.—Hurtado-Ponce, 1983:113.—Hurtado-Ponce and Modelo, 1983:148.—Marcos-Agngarayngay, 1983:100, fig. 31.—Cordero, 1984b:58; 1984c:51.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Sorsogon, CATANDUANES. MINDORO: Oriental Mindoro. MASBATE. LEYTE (Biliran I.). PANAY: Aklan. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. BOHOL. PALAWAN. SULU: Tawitawi.

NOTE.—This species was distributed by Harvey in his Friendly Islands exsiccatae (1857) under the name Neomeris nitida, without a description. Munier-Chalmas (1877:816) established the genus Bornetella on the basis of Neomeris nitida, but without publishing the binomial B. nitida. Sonder appears to be the first author to use that binomial, although he accredited it to Munier-Chalmas. Because Neomeris nitida remained a nomen nudum, Harvey’s name does not appear in the authorship.

Bornetella oligospora Solms-Laubach

Bornetella oligospora Solms-Laubach, 1892:87, pl. IX: figs. 1–4, 6, 7 [syntype localities: Macassar, Celebes and Bari, Flores, Indonesia].—Weber-van Bosse, 1913a:89.—Gilbert, 1943:26, fig. 1g–i; 1946:78; 1961:433 [“sphaerica” in error].—Domantay, 1962:277.—Valet, 1969:589.—Trono, 1973c:21.—Velasquez, Trono, and Doty, 1975:129.—Trono and Young, 1977:57.—Trono and Tuason, 1978:10.—Meñez and Calumpong, 1981:383.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CATANDUANES. MINDORO: Oriental Mindoro. CENTRAL VISAYAS. PALAWAN, SULU: Tawitawi (Sangasiapu I.).

Bornetella ovalis Yamada

Bornetella ovalis Yamada, 1933:277 [type locality: Ryukyu-retto, Japan].—Domantay, 1962:277.—Velasquez, Trono, and Doty, 1975:129.—Cordero, 1984c:51.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

MASBATE.

Bornetella sphaerica (Zanardini) Solms-Laubach

Neomeris ? sphaerica Zanardini, 1878:38 [type locality: Sorong, Irian Barat, Indonesia].

Bornetella sphaerica (Zanardini) Solms-Laubach, 1892:80, pl. IX: fig. 8.—Weber-van Bosse, 1913a:90.—Gilbert, 1943:22, fig. 1a,b; 1946:78; 1961:433.—Domantay, 1962:277.—Taylor, 1966b:347.—Reyes, 1972:141.—Trono, 1972a:99; 1973c:21; 1973d:10; 1975:41.—Velasquez, Trono, and Doty, 1975:129.—Trono and Young, 1977:57.—Reyes, 1978:150, pl. 6: fig. 1.—Trono, 1978:8.—Trono and Tuason, 1978:10.—Cordero, 1979b:276.—Liao and Sotto, 1980:97.—Meñez and Calumpong, 1981:383.—Trono and De Lara, 1981:7, pl. IV: fig. 2.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. SAMAR: Eastern Samar. PANAY: Aklan. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. BOHOL. BASILAN. SULU: Tawitawi (incl. Sangasiapu I.).

Chlorocladus Sonder

Chlorocladus australasicus Sonder

Chlorocladus australasicus Sonder, 1871:67, pl. V: figs. 1–6 [type locality: Cape York, Queensland, Australia].

Dasycladus australasicus (Sonder) De Toni, 1889:411.—Taylor, 1966b:345.—Velasquez, Trono, and Doty, 1975:140 [“australicus”].

PHILIPPINE DISTRIBUTION.—CATANDUANES.

NOTE.—A specimen in UC, collected in Aklan Province, Panay, and published as Dasycladus sp. by Cordero (1979b:282), was determined as Chlorocladus australasicus by I.R. Price, 27 January 1982.

*Chlorocladus philippinensis Gilbert

Chlorocladus philippinensis Gilbert, 1978:305, figs. 1–8 [type locality: Solong-on, Siquijor].—Meñez and Calumpong, 1981:383.

PHILIPPINE DISTRIBUTION.—CATANDUANES. SIQUIJOR.

NOTE.—Gilbert was unable to confirm Taylor’s record of Chlorocladus australasicus because he could not locate the cited specimen. Probably only one species of Chlorocladus is represented in the Philippines, but whether it is distinct from Sonder s species remains to be investigated (see Price, 1981:175).

Cymopolia Lamouroux

Cymopolia vanbosseae Solms-Laubach

Cymopolia vanbosseae Solms-Laubach, 1892:78, pl. VIII: figs. 9, 10, 14–16 [“van Bossei”] [type locality: Maumere, Flores, Indonesia].—Gilbert, 1943:20; 1961:433.—Gilbert and Doty, 1969:125, fig. 16.—Velasquez, Trono, and Doty, 1975:140.—Cordero, 1979b:276.—Meñez and Calumpong, 1981:383.—Hurtado-Ponce, 1983:113.—Hurtado-Ponce and Modelo, 1983:148.—Marcos-Agngarayngay, 1983:100, fig. 32.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Zambales. CATANDUANES. PANAY: Aklan. CEBU. MINDANAO: Surigao.

Dasycladus C. Agardh

Dasycladus vermicularis (Scopoli) Krasser

Spongia vermicularis Scopoli, 1771:412, pl. 64: no. 1454 [type locality: Adriatic Sea].

Dasycladus vermicularis (Scopoli) Krasser in Beck and Zahlbruckner, 1898:459.—Reyes, 1978:150, pl. 5: fig. 8.

PHILIPPINE DISTRIBUTION.—SIQUIJOR.

Neomeris Lamouroux

Neomeris annulata Dickie

Neomeris annulata Dickie, 1874a:198 [type locality: Mauritius].—Gilbert, 1943:19; 1961:434.—Meñez, 1961:50, pl. 1: figs. 1–3.—Cordero, 1973b:18.—Velasquez et al., 1973:10, pl. 2: fig. 6.—Velasquez, Trono, and Doty, 1975:156.—Cordero, 1976c:8; 1977b:24.—Vannajan and Trono, 1977:45, fig. 9.—Cordero, 1979b:276.—Puig and Cordero, 1979:21.—Liao and Sotto, 1980:97.—Meñez and Calumpong, 1981:383.—Hurtado-Ponce, 1983:113 [with query].—Hurtado-Ponce and Modelo, 1983:148 [with query].—Marcos-Agngarayngay, 1983:98, fig. 30.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Ilocos Norte, Pangasinan, Bataan, Cavite, Batangas. MINDORO: Oriental Mindoro. LEYTE (Biliran I.). PANAY: Aklan. CEBU (Mactan I.).

Neomeris vanbosseae Howe

Neomeris vanbosseae Howe, 1909:80, pl. 1: figs. 4, 7; pl. 5: figs. 17–19 [“van Bosseae”] [type locality: Sikka, Flores, Indonesia].—Gilbert, 1943:17; 1961:434.—Meñez, 1961:50, pl. 1: figs. 4, 5.—Domantay, 1962:281.—Taylor, 1966b:347.—Valet, 1969:596.—Reyes, 1972:141.—Trono, 1972a:99; 1973c:20, fig. 20.—Ortega, Alcala, and Reyes, 1974:187, 188.—Westernhagen, 1974:112 (table 1).—Velasquez, Trono, and Doty, 1975:156.—Cordero, 1976c:8; 1977b:24, pl. II: fig. 6.—Reyes, 1978:150, pl. 5: fig. 9.—Trono and Tuason, 1978:11.—Liao and Sotto, 1980:97.—Saraya and Trono, 1980:25.—Trono and Ganzon-Fortes, 1980:35, fig. [s.n].—Meñez and Calumpong, 1981:383.

PHILIPPINE DISTRIBUTION.—BATANES. LUZON: Pangasinan, Bataan, Batangas, Quezon, CATANDUANES. MINDORO: Oriental Mindoro. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. PALAWAN (Balabac I.). SULU: Tawitawi.

Family POLYPHYSACEAE

The families Dasycladaceae and Polyphysaceae were described simultaneously by Kützing (1843). The latter, which included Polyphysa and Acetabularia, was merged into the former by subsequent workers, but was resurrected by Hauck (1884 [1883–1885]:421) under the name Acetabulariaceae.

Acetabularia Lamouroux

Acetabularia calyculus Lamouroux

Acetabularia calyculus Lamouroux in Quoy and Gaimard, 1824:621, pl. 90: figs. 6, 7 [“caliculus”] [type locality: Shark Bay, Western Australia, Australia].—Dickie, 1877:489.—Gilbert, 1943:33; 1946:78; 1961:433.—Villones and Magdamo, 1968:12, fig. 3.—Velasquez et al., 1973:11, pl. 2: fig. 8.—Velasquez, Trono, and Doty, 1975:125.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. CEBU.

Acetabularia clavata Yamada

Acetabularia clavata Yamada, 1934:57, figs. 24, 25 [type locality: Tomari, Okinawa-gunto, Ryukyu-retto, Japan].—Trono and Santiago, 1972:43.—Trono, Santiago, and Ganzon-Fortes, 1978:87, fig. 5d.

PHILIPPINE DISTRIBUTION.—LUZON: Batangas. CATANDUANES.

NOTE.—Polyphysa Lamarck (1816:151), characterized by the lack of an inferior corona, is usually treated as a section of Acetabularia, following the monograph of Solms-Laubach (1895:21). Recently, however, it has been resurrected at the generic level (Bailey, Rezak, and Cox, 1976) and Acetabularia clavata transferred to it (Schnetter and Meyer, 1982:41).

Acetabularia crenulata Lamouroux

Acetabularia crenulata Lamouroux, 1816:249, pl. VIII:fig. 1 [type locality: Caribbean Sea].—Velasquez, 1971:432, fig. 8.—Vannajan and Trono, 1977:45, figs. 10, 11.—Reyes, 1978:151, pl. 6: figs. 3, 4.

PHILIPPINE DISTRIBUTION.—LUZON: Zambales, Cavite, Albay. PANAY: Iloilo. SIQUIJOR. PALAWAN.

Acetabularia dentata Solms-Laubach

Acetabularia dentata Solms-Laubach, 1895:23, pl. 1: fig. 11 [syntype localities: various, all in Indonesia].—Gilbert, 1943:30, fig.2d-f; 1961:433.—Gilbert and Doty, 1969:126, figs. 18, 19.—Reyes, 1972:141.—Trono and Santiago, 1972:44.—Trono, 1973c:21.—Velasquez, Trono, and Doty, 1975:125.—Trono, Santiago, and Ganzon-Fortes, 1978:84, fig. 3.—Liao and Sotto, 1980:97.—Fortes, 1981b:396.—Meñez and Calumpong, 1981:383.—Hurtado-Ponce, 1983:112.—Hurtado-Ponce and Modelo, 1983:148.

PHILIPPINE DISTRIBUTION.—LUZON: Cagayan (Babuyan Is.), Ilocos Norte, Batangas. CATANDUANES. MINDORO: Oriental Mindoro. NEGROS: Negros Oriental, CEBU (Mactan I.). MINDANAO: Surigao.

Acetabularia exigua Solms-Laubach

Acetabularia exigua Solms-Laubach, 1895:28, pl. 2: figs. 1, 4 [syntype localities: Macassar, Celebes and Sikka, Flores, Indonesia].—Trono and Santiago, 1972:42.—Trono, Santiago, and Ganzon-Fortes, 1978:84, 87, fig. 4b–f.—Saraya and Trono, 1980:25.—Meñez and Calumpong, 1981:383.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Batangas. CENTRAL VISAYAS.

Acetabularia major Martens

Acetabularia major Martens, 1868:25, pl. IV: fig. 3 [type locality: “Simaharadscha” (Ban Si Racha?), Thailand].—Howe, 1932:169.—Gilbert, 1943:29, fig. 2a–c; 1946:78.—Zaneveld, 1950:109.—Montilla and Blanco, 1953:166.—Zaneveld, 1956:5; 1959:92.—Gilbert, 1961:433.—Meñez, 1961:51, pl. 1: figs. 10–12.—Domantay, 1962:276, 277.—Gilbert and Doty, 1969:126, fig. 17.—Trono and Santiago, 1972:44.—Trono, 1973c:21.—Velasquez et al., 1973:11, pl. 2: fig. 9.—Velasquez, Trono, and Doty, 1975:125.—Trono and Young, 1977:57.—Trono, Santiago, and Ganzon-Fortes, 1978:79, fig. 1.—Trono and Tuason, 1978:9.—Cordero, 1979b:276.—Puig and Cordero, 1979:22.—Meñez and Calumpong, 1981:383.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan, Zambales, Batangas. CATANDUANES. MINDORO: Oriental Mindoro. LEYTE (Biliran I.). PANAY (incl. Aklan).

Acetabularia minutissima Okamura

Acetabularia minutissima Okamura, 1912 [1909–1912]:184, pl. c: figs. 7–11 [syntype localities: Futae, Amakusa Is. and Hyuga Prov. (Miyazaki Prefecture), Japan].—Meñez, 1961:51.—Velasquez, Trono, and Doty, 1975:125.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan.

Acetabularia parvula Solms-Laubach

Acetabularia parvula Solms-Laubach, 1895:29, pl. 2: figs. 3, 5 [syntype localities: “Tropical India”; Macassar, Celebes, Indonesia].

Acetabularia moebii Solms-Laubach, 1895:30, pl. 4: fig. 1 [type locality: Mauritius].—Trono and Santiago, 1972:43.—Cordero, 1973b:18.—Trono, Santiago, and Ganzon-Fortes, 1978:87, fig. 4a.—Puig and Cordero, 1979:22.—Saraya and Trono, 1980:24.—Meñez and Calumpong, 1981:383.—Hurtado-Ponce, 1983:112.—Hurtado-Ponce and Modelo, 1983:148.—Marcos-Agngarayngay, 1983:98, fig. 29.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan. CATANDUANES. LEYTE (Biliran I.). CENTRAL VISAYAS.

NOTE.—The synonymy was proposed by Valet (1969:621). This species was transferred to Polyphysa by Schnetter and Meyer (1982:42).

*Acetabularia roxasii Trono

Acetabularia roxasii Trono, Santiago, and Ganzon-Fortes, 1978:80, fig. 2. [type locality: Calatagan, Batangas Prov., Luzon].

PHILIPPINE DISTRIBUTION.—As above.

Acetabularia ryukyuensis Okamura and Yamada

Acetabularia ryukyuensis Okamura and Yamada in Okamura, 1932a [1929–1932]:71, pl. CCLXXXV: figs. 5–12 [type locality: Ryukyu-retto, Japan].—Cordero, 1979b;282.—Puig and Cordero, 1979:22.

PHILIPPINE DISTRIBUTION.—LEYTE (Biliran I.), PANAY: Aklan.

*Acetabularia ryukyuensis Okamura and Yamada var. philippinensis (Gilbert) Valet

Acetabularia philippinensis Gilbert, 1943:31, fig. 3 [type locality: Taytay, Palawan].—Gilbert, 1961:433.—Taylor, 1966b:347.—Velasquez, Trono, and Doty, 1975:125.—Trono and Tuason, 1978:9.

Acetabularia ryukyuensis Okamura and Yamada var. philippinensis (Gilbert) Valet, 1969:613, pl. 154 (43): fig. 12.

PHILIPPINE DISTRIBUTION.—LUZON: Pangasinan. CATANDUANES. PALAWAN.

*Acetabularia velasquezii Trono, Santiago, and Ganzon-Fortes

Acetabularia velasquezii Trono, Santiago, and Ganzon-Fortes, 1978:87, fig. 5a–c [type locality: Minabalay I., Barrio Danao, Baras, Catanduanes].

PHILIPPINE DISTRIBUTION.—As above.

Halicoryne Harvey

Halicoryne wrightii Harvey

Halicoryne wrightii Harvey, 1860a:333 [type locality: Ryukyu-retto, Japan].—Solms-Laubach, 1895:31.—Gilbert, 1943:27; 1946:78; 1961:433.—Meñez, 1961:51.—Domantay, 1962:281.—Taylor, 1966b:346.—Valet, 1969:603.—Velasquez, 1971:432, fig. 9.—Reyes, 1972:141.—Cordero, 1973b:19.—Velasquez et al., 1973:11, pl. 2: fig. 7.—Westernhagen, 1973a:64.—Ortega, Alcala, and Reyes, 1974:188.—Westernhagen, 1974:112 (table 1).—Trono, 1975:40.—Velasquez, Trono, and Doty, 1975:147, 149 [“Halycoryne wrigtii”].—Trono and Young, 1977:57.—Reyes, 1978:150, pl. 6: fig. 2.—Trono, 1978:8.—Trono and Tuason, 1978:10.—Cordero, 1980a:29, 34, fig. 3.—Liao and Sotto, 1980:97.—Meñez and Calumpong, 1981:383.—Trono and De Lara, 1981:7, pl. IV: fig. 3.—Cordero, 1984a:76.

Polyphysa spicata [misapplied name fide Solms-Laubach, 1895:31].—Dickie, 1876a:243.—Gilbert, 1961:434.—Velasquez, Trono, and Doty, 1975:160.

PHILIPPINE DISTRIBUTION.—LUZON: Ilocos Norte, Pangasinan, Zambales, Bataan, Albay, Sorsogon. CATANDUANES. MINDORO: Occidental Mindoro (Lubang Is.), Oriental Mindoro. MARINDUQUE. LEYTE (Biliran I.). PANAY. NEGROS: Negros Oriental. CEBU (Mactan I.). SIQUIJOR. BOHOL.

Records Determined Only To Genus

(Genera marked with an asterisk (*) have not been otherwise reported from the Philippines, or (in the case of Cyanophyceae) at least not from marine habitats.)

CYANOPHYCEAE

Anabaina

Reyes, 1972:136 [“Anabaena”]. NEGROS: Negros Oriental.

*Aphanocapsa

Agor, 1962:33. LUZON: Pangasinan.

Calothrix

Cornejo and Velasquez, 1972:172, pl. 1: fig. 3. LUZON: Batangas.

Vannajan and Trono, 1977:37–38. LUZON: Cavite.

*Coccochloris

Reyes, 1972:135. NEGROS: Negros Oriental.

Entophysalis

Reyes, 1978:139, pl. 1: figs. 3, 4. SIQUIJOR.

*Gloeotrichia

Velasquez, 1980:127 (table 1) [no description or collecting data]. MINDORO: Oriental Mindoro.

Hormothamnium

Villones and Magdamo, 1968:11. LUZON: Batangas.

Reyes, 1972:136. NEGROS: Negros Oriental.

Lyngbya

Vannajan and Trono, 1977:37. LUZON: Cavite.

Oscillatoria

Vannajan and Trono, 1977:36. LUZON: Cavite.

Rivularia

Domantay, 1962:276. LUZON: Pangasinan.

Schizothrix

Reyes, 1978:140, pl. 2: figs. 3, 4. SIQUIJOR.

Symploca

Domantay, 1962:276. LUZON: Pangasinan.

RHODOPHYCEAE

Acrochaetium

Cornejo and Velasquez, 1972:178, pl. 2: fig. 17. LUZON: Batangas.

*Agardhiella

Seale, 1911:309 [no description; see Gracilaria verrucosa in catalog].

Quisumbing, 1951:1007. LUZON: Manila Bay.

Zaneveld, 1956:26. LUZON: Manila Bay.

Zaneveld, 1959:107. LUZON: Manila Bay.

Ahnfeltia

Cordero, 1978a:42. BATANES.

Amansia

Meñez, 1961:75, pl. 8: figs. 92–94. LUZON: Pangasinan.

Amphiroa

Villones and Magdamo, 1968:30, fig. 33. LUZON: Batangas.

Reyes, 1972:153. NEGROS: Negros Oriental.

Trono and Biña, 1973:5. MINDORO: Oriental Mindoro.

Trono, 1974b:86. SULU: Sulu (Siasi I.).

Reyes, 1980:131, pl. 7: figs. 1, 2. SIQUIJOR.

Cordero, 1984a:95. LUZON: Ilocos Norte.

Antithamnion

Vannajan and Trono, 1978:23, figs. 26, 27. LUZON: Cavite.

Asterocytis

Vannajan and Trono, 1978:14. LUZON: Cavite.

*Calliarthron

Cordero, 1979b:276. PANAY: Aklan.

*Callithamnion

Vannajan and Trono, 1978:23, fig. 28. LUZON: Cavite.

Callophyllis

Cordero, 1979b:290. PANAY: Aklan.

Reyes, 1980:134, pl. 9: figs. 1, 2. SIQUIJOR.

Cordero, 1984a:97. LUZON: Ilocos Norte.

Carpopeltis

Meñez, 1961:73. LUZON: Pangasinan.

Reyes, 1972:154. NEGROS: Negros Oriental.

Cordero, 1973b:31. LEYTE (Biliran I.).

Trono and Biña, 1973:3. MINDORO: Oriental Mindoro.

Centroceras

Domantay, 1962:288. LUZON: Pangasinan.

Ceramium

Cornejo and Velasquez, 1972:183, pl. 3: fig. 26; pl. 5: fig. 39. LUZON: Batangas.

Trono, 1972:106. SULU: Tawitawi.

Trono, 1973d:20. MINDANAO: Zamboanga.

Reyes, 1980:138, pl. 12, figs. 1a,b. SIQUIJOR.

Trono and Ang, 1982:22. PALAWAN (Bugsuk I.).

Cordero, 1984a:104. LUZON: Ilocos Norte.

Champia

Meñez, 1961:78. LUZON: Pangasinan.

Cornejo and Velasquez, 1972:179, pl. 4: fig. 35. LUZON: Batangas.

Chondria

Cornejo and Velasquez, 1972:179–180, pl. 2: figs. 19, 20; pl. 3: fig. 21. LUZON: Batangas.

*Chondrus

Cordero, 1984a:103 [with query]. LUZON: Ilocos Norte.

*Chrysymenia

Reyes, 1972:157. NEGROS: Negros Oriental.

Trono, 1973a:135, fig. 7. LUZON: Sorsogon.

Corallina

Reyes, 1972:153. NEGROS: Negros Oriental.

Reyes, 1980:131, pl. 7: fig. 3. SIQUIJOR.

Cryptonemia

Reyes, 1980:133, pl. 8: fig. 4. SIQUIJOR.

Marcos-Agngarayngay, 1983b:39, fig. 32. LUZON: Ilocos Norte.

*Dicranema

Trono, 1974b:93. SULU: Sulu (Siasi I.).

Eupogodon

Reyes, 1972:158 [as Dasyopsis]. NEGROS: Negros Oriental.

Erythrotrichia

Cornejo and Velasquez, 1972:177, pl. 2: fig. 14; pl. 4: fig. 32. LUZON: Batangas.

Vannajan and Trono, 1978:14, fig. 12. LUZON: Cavite.

Eucheuma

Cordero, 1973b:31. LEYTE (Biliran I.).

Reyes, 1980:136, pl. 10: fig. 4. SIQUIJOR.

Cordero, 1984a:99. LUZON: Ilocos Norte.

Fauchea

Reyes, 1972:158. NEGROS: Negros Oriental.

Galaxaura

Trono and De Lara, 1981:13, pl. VIII: fig. 4. MINDORO: Occidental Mindoro (Lubang Is.).

Cordero, 1984a:91. LUZON: Ilocos Norte.

Gelidiella

Trono and Biña, 1973:3. MINDORO: Oriental Mindoro.

Gelidiopsis

Cordero, 1973b:30. LEYTE (Biliran I.).

Reyes, 1980:129–130, pl. 6: figs. 3, 4. SIQUIJOR.

Cordero, 1984a:100. LUZON: Ilocos Norte.

Gelidium

Cordero, 1973b:30. LEYTE (Biliran I.).

Cordero, 1978a:23. BATANES.

Gracilaria

Collado, 1926:129. LUZON: Cagayan, Ilocos.

Reyes, 1972:155–156. NEGROS: Negros Oriental.

Cordero, 1973b:33. LEYTE (Biliran I.).

Trono, 1973d:18, pl. 9: fig. 4. LUZON: Pangasinan. SAMAR: Eastern Samar. NEGROS: Negros Oriental. MINDANAO: Zamboanga.

Vannajan and Trono, 1978:22, fig. 25. LUZON: Rizal.

Saraya and Trono, 1982:41. LUZON: Pangasinan.

Trono and Ang, 1982:20. PALAWAN (Bugsuk I.).

Trono, Azanza-Corrales, and Manuel, 1983:34–41, figs. 8, 9a, 10, 11b.

LUZON: Rizal, Cavite. SULU: Sulu (Siasi I.).

Cordero, 1984a:103. LUZON: Ilocos Norte.

Grateloupia

Trono, 1973d:16. LUZON: Pangasinan.

Reyes, 1980:133, pl. 8: fig. 3. SIQUIJOR.

Marcos-Agngarayngay, 1983c:126. LUZON: Ilocos Norte.

Cordero, 1984a:97. LUZON: Ilocos Norte.

Griffithsia

Reyes, 1972:158. NEGROS: Negros Oriental.

Reyes, 1980:138, pl. II: figs. 6a,b. SIQUIJOR.

Gymnogongrus

Reyes, 1980:137, pl. II: fig. 1. SIQUIJOR.

Halymenia

Vannajan and Trono, 1978:18. LUZON: Cavite.

Cordero, 1980:49, fig. 11B, pl. 31. SAMAR: West Samar.

Hurtado-Ponce, 1983a:138. LUZON: Ilocos Norte.

Herposiphonia

Cornejo and Velasquez, 1972:181, pl. 3: figs. 23, 23a. LUZON: Batangas.

Trono, 1973d:21, pl. 10: fig. 8. MINDANAO: Zamboanga.

Trono and De Lara, 1981:20. MINDORO: Occidental Mindoro (Lubang Is.).

*Heteroderma

Saraya and Trono, 1982:35. LUZON: Pangasinan.

Hypnea

Collado, 1926:129. LUZON: Cagayan, Ilocos.

G. Blanco, 1938:513. LUZON: Cagayan (incl. Babuyan Is.).

Reyes, 1972:156. NEGROS: Negros Oriental.

Trono, 1972:105. SULU: Tawitawi.

Trono, 1973d:17. CEBU. MINDANAO: Zamboanga (incl. Sacol I.).

Cordero, 1978a:41. BATANES.

Jania

Cordero, 1984a:95. LUZON: Ilocos Norte.

Cordero, 1984c:54. MASBATE.

Kallymenia

Marcos-Agngarayngay, 1983c:126. LUZON: Ilocos Norte.

Laurencia

Meñez, 1961:77. LUZON: Pangasinan.

Reyes, 1972:159. NEGROS: Negros Oriental.

Trono, 1972:108. SULU: Tawitawi.

Trono, 1973d:22. LUZON: Pangasinan, Quezon. MINDANAO: Zamboanga.

Trono and Ganzon-Fortes, 1980:105, fig. [s.n.]. LUZON: Batangas.

Trono and Ang, 1982:26. PALAWAN (Bugsuk I.).

Cordero, 1984a:108. LUZON: Ilocos Norte.

*Leptofauchea

Reyes, 1972:157. NEGROS: Negros Oriental.

Ortega, Alcala, and Reyes, 1974:186, 187, 188 [no description or collecting data], NEGROS: Negros Oriental.

Liagora

Domantay, 1962:292. LUZON: Pangasinan.

Trono and De Lara, 1981:11–12, pl. VII: fig. 2. MINDORO: Occidental Mindoro (Lubang Is.).

Cordero, 1984a:87. LUZON: Ilocos Norte.

Lithophyllum

Howe, 1932:169. PANAY.

Reyes, 1980:132, pl. 7: figs. 6, 7. SIQUIJOR.

Lomentaria

Vannajan and Trono, 1978:23, fig. 30. LUZON: Manila, Cavite.

Lophosiphonia

Reyes, 1980:141, pl. 13: figs. 4a–c. SIQUIJOR.

Martensia

Trono, 1974b:94. SULU: Sulu (Siasi I.).

*Melobesia

Reyes, 1972:153. NEGROS: Negros Oriental.

Microcladia

Tahil, 1978:52 [no description or collecting data], CEBU (Mactan I.).

Murrayella

Trono, 1973a:136. MINDORO: Oriental Mindoro.

Peyssonnelia

Reyes, 1972:154. NEGROS: Negros Oriental.

Cordero, 1984a:93. LUZON: Ilocos Norte.

Polysiphonia

Domantay, 1962:293. LUZON: Pangasinan.

Reyes, 1972:158. NEGROS: Negros Oriental.

Trono, 1972:107. SULU: Tawitawi.

Trono, 1973d:21, pl. 10: fig. 7. MINDANAO: Zamboanga.

Reyes, 1980:141, pl. 13: figs. 3a,b. SIQUIJOR.

Porphyra

Bersamin et al., 1973:188. LUZON: Cagayan, Ilocos.

Pterocladia

Reyes, 1980:129, pl. 6: fig. 2. SIQUIJOR.

Rhodopeltis

Trono, 1973a:133, fig. 4. LUZON: Sorsogon.

Rhodymenia

Trono, 1973a:136, fig. 15. LUZON: Sorsogon.

Trono and Biña, 1973:10. MINDORO: Oriental Mindoro.

Cordero, 1978a:43. BATANES.

Cordero, 1980c:72, fig. 4. LUZON: Ilocos Norte.

Reyes, 1980:137, pl. II: fig. 3. SIQUIJOR.

Trono and De Lara, 1981:19, pl. XI: fig. 3. MINDORO: Occidental Mindoro (Lubang Is.).

Marcos-Agngarayngay, 1983b:53, fig. 44. LUZON: Ilocos Norte.

Cordero, 1984a:103. LUZON: Ilocos Norte.

Sarcodia

Trono, 1973d:18 [with query], NEGROS: Negros Oriental.

Cordero, 1978a:35. BATANES.

Scinaia

Trono, 1973a:130, fig. 5. LUZON: Sorsogon.

*Spermothamnion

Westernhagen, 1973b:368 [no description or collecting data], CEBU (Mactan I.).

Titanophora

Trono, 1973a:134, fig. 13. LUZON: Sorsogon.

Trono and Bina, 1973:7. MINDORO: Oriental Mindoro.

Trono, 1978:18. MARINDUQUE.

Trono and Ganzon-Fortes, 1980:73, fig. [s.n.]. LUZON: Batangas.

Saraya and Trono, 1982:37. LUZON: Pangasinan.

Tolypiocladia

Trono, 1972:107. SULU: Tawitawi.

Trono and De Lara, 1981:20. MINDORO: Occidental Mindoro (Lubang Is.).

Trichogloea

Cordero, 1978a:13. BATANES.

PHAEOPHYCEAE

Colpomenia

Cordero, 1984a:85. LUZON: Ilocos Norte.

Cutleria

Cordero, 1979b:284. PANAY: Aklan.

Cystoseira

Reyes, 1980:123, pl. 4: fig. 3. SIQUIJOR.

Dictyopteris

Meñez, 1961:64. LUZON: Pangasinan.

Dictyota

Meñez, 1961:63, pl. II: fig. 121; pl. 12: fig. 132. LUZON: Pangasinan.

Villones and Magdamo, 1968:26, fig. 17. LUZON: Batangas.

Trono, 1974a:144. SULU: Sulu (Siasi I.).

Cordero, 1984a:83. LUZON: Ilocos Norte.

Cordero, 1984b:61. MASBATE.

*Ectocarpus

Cornejo and Velasquez, 1972:176, pl. 2: figs. 12, 13. LUZON: Batangas.

Feldmannia

Cordero, 1979b:284. PANAY: Aklan.

Hydroclathrus

Villones and Magdamo, 1968:26, fig. 19. LUZON: Batangas.

Hormophysa

Reyes, 1972:147. NEGROS: Negros Oriental.

Padina

Meñez, 1961:62. LUZON: Pangasinan.

Villones and Magdamo, 1968:27, fig. 20. LUZON: Batangas.

Reyes, 1972:146. NEGROS: Negros Oriental.

Reyes, 1980:120, pl. 2: fig. 2. SIQUIJOR.

Cordero, 1984a:84. LUZON: Ilocos Norte.

*Punctaria

Cordero, 1984a:84. LUZON: Ilocos Norte.

Sargassum

Meñez, 1961:64–67, pl. 9: figs 96–102. LUZON: Pangasinan.

de los Reyes, 1967:231, 233, fig. 6. LEYTE (Biliran I.).

Villones and Magdamo, 1968:27, fig. 21. LUZON: Batangas.

Reyes, 1972:146–147. NEGROS: Negros Oriental.

Cordero, 1973b:26. LEYTE (Biliran I.).

Trono, 1973d:13, pl. 8: fig. 31, 32.

Velasquez et al., 1973:22. LUZON: Bataan.

Trono, 1974a:148. SULU: Sulu (Siasi I.).

Trono and Young, 1977:58. CATANDUANES.

Trono, 1978:12–14. MARINDUQUE.

Trono and Tuason, 1978:12. CATANDUANES.

Vannajan and Trono, 1978:13, fig. 9. LUZON: Cavite.

Cordero, 1980b:36, fig. 3c, pls. 23, 24, [51]. LUZON: Cagayan, Ilocos Norte, SAMAR: Western Samar, Eastern Samar. PANAY: Aklan. SIQUIJOR. MINDANAO: Surigao del Sur.

Reyes, 1980:123, pl. 4: fig. 4. SIQUIJOR.

Trono and Ganzon-Fortes, 1980:51–53, [2] figs. LUZON: Batangas.

Guzman, 1981:43, 52.

Trono and De Lara, 1981:10, pl. VII: fig. 1. MINDORO: Occidental Mindoro (Lubang Is.).

Trono and Ang, 1982:15–16. PALAWAN (Bugsuk I.).

Hurtado-Ponce, 1983a:125–126. LUZON: Ilocos Norte.

Marcos-Agngarayngay, 1983b:14, fig. 11. LUZON: Ilocos Norte.

Cordero, 1984a:86. LUZON: Ilocos Norte.

Cordero, 1984b:63. PALAWAN.

Cordero, 1984c:54. MASBATE.

Spatoglossum

Trono, 1974a:145–146. SULU: Sulu (Siasi I.).

Reyes, 1980:121, pl. 2: fig. 4. SIQUIJOR.

Sphacelaria

Trono, 1973d:11. LUZON: Pangasinan.

Vannajan and Trono, 1978:8, fig. 2. LUZON: Cavite.

Turbinaria

Meñez, 1961:68. LUZON: Pangasinan.

Reyes, 1972:147. NEGROS: Negros Oriental.

Trono, 1972:101. SULU: Tawitawi.

Trono, 1973d:12. SAMAR: Eastern Samar.

CHLOROPHYCEAE

Acetabularia

Domantay, 1962:277. LUZON: Pangasinan.

Anadyomene

Cordero, 1984a:74. LUZON: Ilocos Norte.

Boergesenia

Trono, 1973b:217. SULU: Sulu (Siasi I.).

Boodlea

Villones and Magdamo, 1968:26, fig. 14. LUZON: Batangas.

Trono, 1973c:6. MINDORO: Oriental Mindoro.

Trono, 1973d:4. SULU: Tawitawi.

Bryopsis

Reyes, 1972:142. NEGROS: Negros Oriental.

Caulerpa

Trono, 1973a:128. LUZON: Sorsogon.

Trono, 1973b:220. SULU: Sulu (Siasi I.).

Trono, 1973d:8, pl. 2: fig. 6. SIQUIJOR.

Cordero, 1976a:86, fig. 2. CEBU (Mactan I.).

Cordero, 1977b:26, 28, fig. 3 [Caulerpa brachypus? “false-type”], SIQUIJOR.

Cordero, 1984a:78. LUZON: Ilocos Norte.

Chaetomorpha

G. Blanco, 1938:512. LUZON: Cagayan (incl. Babuyan Is.).

Villones and Magdamo, 1968:24, fig. 10 [cited but not published].

LUZON: Batangas.

Reyes, 1972:140. NEGROS: Negros Oriental.

Trono, 1973b:214. SULU: Sulu (Siasi I.).

Reyes, 1978:149. SIQUIJOR.

*Cladocephalus

Taylor, 1966b:352. PALAWAN (Balabac I.).

Cladophora

Cornejo and Velasquez, 1972:174, pl. 1: fig. 8. LUZON: Batangas.

Trono, 1972:94. SULU: Tawitawi.

Trono, 1973b:214. SULU: Sulu (Siasi I.).

Trono, 1973d:4. SIQUIJOR.

Trono, 1975:29. LUZON: Sorsogon.

Cordero, 1977b:22. BATANES.

Vannajan and Trono, 1977:43, fig. 17. LUZON: Rizal, Manila.

Reyes, 1978:147–148, pl. 4: figs. 3–7. SIQUIJOR.

Cladophoropsis

Trono, 1973d:4, pl. 1: fig. 4. MINDANAO: Zamboanga.

Cordero, 1984a:73 [as Spongocladia]. LUZON: Ilocos Norte.

Codium

Galutira and Velasquez, 1964:498, pl. 2: fig. 6; pl. 7: fig. 24. LUZON: Ilocos Norte,

de los Reyes, 1967:231, 233, fig. 1. LEYTE (Biliran I.).

Trono, 1973b:223–225. SULU: Sulu (Siasi I.).

Trono, 1973c:14, fig. 6.

Trono, 1973d:9. MINDANAO: Zamboanga.

Reyes, 1978:158, pl. 10: figs. 1, 2. SIQUIJOR.

Cordero, 1979b:282. PANAY: Aklan.

Cordero, 1984a:79. LUZON: Ilocos Norte.

Dasycladus

Cordero, 1979b:282. PANAY: Aklan. [Specimen in UC determined as Chlorocladus australasicus by I.R. Price, 27.i. 1982.]

Dictyosphaeria

Reyes, 1978:152, pl. 6: figs. 11, 12. SIQUIJOR.

Enteromorpha

Trono, 1972:94. SULU: Tawitawi.

Trono, 1973c:4. MINDORO: Oriental Mindoro.

Trono, 1975:25. LUZON: Sorsogon.

Reyes, 1978:146. SIQUIJOR.

Saraya and Trono, 1980:8. LUZON: Pangasinan.

Guzman, 1981:42, 47, 51.

Cordero, 1984a:70. LUZON: Ilocos Norte.

Halimeda

Trono, 1973b:227–228. SULU: Sulu (Siasi I.).

Trono, 1973d:9. MINDANAO: Zamboanga.

Trono, 1975:39–40. LUZON: Sorsogon.

Trono and Young, 1977:57. CATANDUANES.

Cordero, 1984a:81. LUZON: Ilocos Norte.

Microdictyon

Trono, 1973b:215. SULU: Sulu (Siasi I.).

Cordero, 1984a:74. LUZON: Ilocos Norte.

Monostroma

Reyes, 1972:138. NEGROS: Negros Oriental.

Reyes, 1978:146, pl. 3: fig. 6. SIQUIJOR.

*Spongomorpha

Liao and Sotto, 1980:96 [no description or collecting data], CEBU (Mactan I.).

*Trichosolen

Gilbert and Doty, 1969:122, fig. 5. LUZON: Rizal. CATANDUANES. CEBU.

Ulva

Trono, 1975:28. LUZON: Sorsogon.

Reyes, 1978:147, pl. 3: fig. 10. SIQUIJOR.

Cordero, 1984a:71. LUZON: Ilocos Norte.

Nomenclatural Notes

by Paul C. Silva

Later Starting Points in CYANOPHYCEAE

The International Code of Botanical Nomenclature (ICBN; Voss et al., 1983) states that nomenclature for the Nostocales begins with the monographs of Bornet and Flahault (1886–1888) and Gomont (1892–1893). (The ICBN assigns an arbitrary date of 1886 and 1892, respectively, to these monographs.) An enormous body of taxonomic information about blue-green algae had been published before those distinguished French phycologists wrote their monographs. According to the ICBN, pre-starting point taxonomy may be retained, but pre-starting point nomenclature does not exist. This legalistic nonsense, rather than simplify the nomenclature of blue-green algae, has introduced hopeless confusion. Even those (like Drouet) who profess to follow the ICBN with regard to later starting points are inconsistent. Names treated as synonyms in the starting point monographs are invalid according to the ICBN, but Drouet (1968:14) pointed out that all such names presumably have been validated subsequently “by reference or by a new description.” Lacking the time to “search through thousands of local floras, check-lists of species, and morphological, ecological, and physiological papers”, Drouet treated those older names “as having been validly and autonomously published, not as synonyms, but by reference to their original descriptions” in the starting point monograph and subsequent publications.

In view of the facts that Drouet has located the types of the majority of blue-green algal species and that blue-green algal taxonomy is in an extraordinary state of flux, it seems pointless to attempt to accommodate their nomenclature to later starting points. Oscillatoria submembranacea is an example of the needless complications caused by the use of later starting points. This species was first described by Ardissone and Strafforello (1877) as Oscillaria submembranacea. In Gomont (1892), the starting point monograph for Oscillatoria, it was treated as Phormidium submembranaceum. In accordance with normal nomenclature, this name would be written Phormidium submembranaceum (Ardissone and Strafforello) Gomont, but Article 13.1 of the ICBN renders pre-starting point nomenclature invalid, so that the name must be ascribed solely to Gomont and dated from 1892, not 1877. Drouet (1968:203) transferred the species to Oscillatoria, but incorrectly attributed it directly to Ardissone and Strafforello (considering Oscillaria and Oscillatoria as orthographic variants, an equivocal interpretation). In accordance with later starting points, it should be written Oscillatoria submembranacea (Gomont) Drouet!

In the long course of compiling the Index Nominum Algarum (since 1948), I repeatedly struggled with the complications of later starting points and eventually concluded that it was impractical to continue to do so. The present catalog is my first opportunity to make the iconoclastic move of basing the nomenclature of blue-green algae uniformly on the Linnaean starting date, 1753.

The Taxonomy of Acrochaetioid Algae

The confused taxonomy of acrochaetioid algae was thoroughly reviewed by Woelkerling (1983a), who expressed personal preference for either of two classifications: a monogeneric scheme using the oldest available name, Audouinella Bory de Saint-Vincent (1823:340, “Auduinella”); or a polygeneric scheme based solely on the type of life history without regard for vegetative features. Woelkerling concluded: “In the short term, it seems to me that it is more important to accumulate detailed, trustworthy data on numerous species than to engage in endless manipulation of generic schemes. Use of either of the above two schemes would provide a workable taxonomic framework while new studies are being carried out.”

The decision as to which classification would best serve the purposes of this catalog was based on the following considerations. Papenfuss (1945), in trying to delineate the South African acrochaetioid algae, concluded that it was possible to distribute them among four genera on the basis of the number and form of the cloroplasts in each vegetative cell, as follows:

Rhodochorton, few to many small, discoid chloroplasts;

Acrochaetium, one parietal or laminate chloroplast;

Audouinella, one or more spiral chloroplasts; and

Chromastrum, one or more stellate chloroplasts.

It soon became apparent that the distinction drawn by Papenfuss between Acrochaetium and Chromastrum was difficult to maintain, partly because he (and many subsequent workers) confused form with position. A chloroplast with a stellate form may be axile or parietal in position. There are intermediate situations with regard to both form and position. Feldmann (1962) proposed that Acrochaetium include plants whose cells contain a single chloroplast of either form or position. Woelkerling (1971), believing that vegetative characters are of diagnostic value only at the species level, placed all acrochaetioid algae with sexual stages in one genus, Audouinella. He referred those without sexual stages to Colaconema Batters (1896:8), which he treated as a form genus. Chromastrum was re-established by Stegenga and Mulder (1979) to include species with one axile, stellate, pyrenoid-bearing chloroplast in each cell and a life history involving an alternation of heteromorphic phases—the gametophyte with a unicellular base and the tetrasporophyte with a multicellular base that develops from a septately germinating carpospore. Acrochaetium was circumscribed to include species with one to several parietal pyrenoid-bearing chloroplasts in each cell and a life history involving an alternation of isomorphic phases. Audouinella was retained for species with one to several parietal chloroplasts of spiral or irregular shape, without pyrenoids.

The nomenclature of acrochaetioid algae was muddled long ago by the selection of Acrochaetium daviesii (Dillwyn) Nägeli (1862:405, 412), based on Conferva daviesii Dillwyn (1809 [1802–1809]:73, pl. F), as the lectotype of its genus by Drew (1928:147), who erroneously believed (along with all other authors) that the genus was first described by Nägeli in 1862. In fact, it was described in 1858, and A. daviesii was not among the original species. Accordingly, Woelkerling (1983a:65) set aside the lectotype that, although untenable, had been accepted for more than a half-century, replacing it with A. secundatum (Lyngbye) Nageli (1858:532, footnote), based on Callithamnion daviesii (Dillwyn) Lyngbye var. secundatum Lyngbye (1819:129, pl. 41B: figs. 4–6). Although in both species each cell contains a single chloroplast with a pyrenoid, in A. daviesii the chloroplast is laminate and parietal, while in A. secundatum it is stellate and axile. Those schemes that are based on vegetative features, either wholly or partly, are thus severely affected. Chromastrum falls into synonymy under Acrochaetium, while some name other than Acrochaetium must be applied to those species considered by many workers through many decades to be congeneric with A. daviesii. Stegenga (1985) applied Colaconema to the latter circumscription. The coining of numerous binomials as a consequence of changing the lectotype of Acrochaetium could be precluded by the conservation of that generic name using the 1862 publication as protologue and A. daviesii as type. Any proposal for conservation, however, should await further clarification of the taxonomy of acrochaetioid algae.

Because any classification of acrochaetioid algae used at present either would be admittedly artificial (based on a single character) or, if allegedly natural (based on morphological, ontogenetic, and life history characters), would be criticized as being premature, I have employed a scheme that at least has the merit of not requiring new binomials, namely that of Feldmann (1962). According to this classification, all records of acrochaetioid algae for the Philippines are encompassed by Acrochaetium.

Galaxauraceae as a Replacement for Chaetangiaceae

When Chaetangium was established, Kützing (1843b:392) misidentified his material with Fucus ornatus Linnaeus, which has turned out to be representative of Suhria in the Gelidiaceae (see Papenfuss, 1952:173). For many decades the name Chaetangium has been applied on the basis of Kützing’s material, but a decision made at the International Botanical Congress at Sydney in 1981 makes it clear that the name must remain with Suhria unless conservation is invoked.

Parkinson (1983), after reviewing the nomenclatural history of Chaetangium, decided against conservation of that generic name, choosing instead to propose Suhria for conservation against Chaetangium and to resurrect Nothogenia Montagne (1843a:302) as a replacement for Chaetangium. [Nothogenia variolosa (Montagne) Montagne, the type of its genus, was placed in the synonymy of Chaetangium fastigiatum (Bory de Saint-Vincent) J. Agardh by Kylin (in Kylin and Skottsberg, 1919:6)]. Because the name Chaetangiaceae must be considered a synonym of Gelidiaceae, Parkinson simultaneously established the family Galaxauraceae.

Portieria as a Replacement for Chondrococcus

The genus of Rhizophyllidaceae currently called Desmia Lyngbye (1819) emend. J. Agardh (1852 [1851–1863]) or Chondrococcus Kützing (1847) has a troubled nomenclatural history. I have discussed it in detail previously (Silva, 1952a:304–306), but it is necessary to return to it in light of changes in the, ICBN made at Sydney pertaining to the typification of generic names.

Desmia was established by Lyngbye (1819:XXIX, 33) to receive three species with narrow, compressed branches. One species is the type of Herbacea Stackhouse (1809) and another is the type of Hippurina Stackhouse (1809). Both of these genera are referable to Desmarestia Lamouroux (1813), but the name Desmarestia has been conserved against the Stackhouse names. Desmia is thus a superfluous name for Desmarestia and illegitimate in accordance with Article 63.1 of the ICBN. Although Lyngbye did not give a reason for abandoning the name Desmarestia, which he cited in synonymy, he may have considered Desmia a better name because of its brevity or because it was descriptive (derived from the Greek word desmos, “band”) rather than commemorative (Desmarestia being named for Anselm Gaetan Desmarest, a French zoologist). The sharing of the first four letters by the two names perhaps is not purely coincidental.

The third species assigned by Lyngbye to Desmia was D. hornemannii (“hornemanni”), based on a specimen sent to him by Mertens under the name Fucus hornemanni. Mertens found this specimen among Forsskål’s collections and indicated that it came from Helsingør on the Øresund (strait between Denmark and Sweden), but the provenance probably was the Red Sea (see Rosenvinge, 1931:617). Lyngbye cited Fucus hornemanni as being published by Mertens in the Göttingische gelehrte Anzeigen for 1815, but at that place (page 633) it was a nomen nudum (see Papenfuss, 1940:216). Valid nomenclature for this species thus begins with Desmia hornemannii as described and illustrated by Lyngbye (1819:35, pl. 7:c). Subsequent to 1819 and prior to 1852, the name Desmia was applied either to a genus closely related to Desmarestia or to a section or subgenus of that genus.

J. Agardh (1852 [1851–1863]:639) emended the genus Desmia, retaining only D. hornemannii of Lyngbye’s original species and adding two other species, D. ambigua. J. Agardh from India and D. tripinnata (Hering) J. Agardh (Rhodymenia tripinnata Hering) from South Africa. He accredited the emended genus to himself and placed it (with a query) in the tribus Sphaerococcoideae of the ordo Sphaerococcoideae (a broadly defined group not equivalent taxonomically to the modern Sphaerococcaceae).

Meanwhile, Kützing (1847a:23) had established the genus Chondrococcus on the basis of Fucus abscissus Turner (1819 [1815–1819]:65, pl. 223) from New Zealand and F. lambertii Turner (1819 [1815–1810]:96, pl. 237) from Australia. The first species is currently placed in Melanthalia Montagne (1843) in the Gracilariaceae, while the second species, as defined by the type collection, is referable to Callophyllis Kützing (1843) in the Kallymeniaceae. Kützing applied the name Chondrococcus lambertii to a mixture of species, citing both Australian and South African specimens, the former being representative of Callophyllis, the latter of Chondrococcus as currently defined. At the time that I wrote my account of the nomenclatural history of Chondrococcus, typification of generic names was not legislated unequivocally by the ICBN, and the contemporary consensus was that in the event an author based a generic description on one entity but erroneously assigned it to a different entity, the generic name should apply to the material at hand. In the case of Chondrococcus, Kützing based the generic description on South African material, so that in 1952 I concluded that the correct name for the genus was Chondrococcus. At Sydney in 1981, however, the ICBN was changed to specify that the type of a generic name must be a cited species. Thus, the type of Chondrococcus is either Fucus abscissus or F. lambertii, making the generic name a later taxonomic synonym of Melanthalia Montagne (1843) or Callophyllis Kützing (1843).

J. Agardh (1852 [1851–1863]:639) cited Chondrococcus as a synonym of his emended Desmia, and during the period 1852–1895 only Kützing used that name. Schmitz (1895:168–172) reviewed the genus and adopted the name Chondrococcus in preference to Desmia. Most subsequent authors have followed Schmitz in this regard.

Portieria Zanardini (1851:33) appears to be the only available synonym. This genus was established to receive a single species, P. coccinea, based on material obtained in the Red Sea by Portier, a poorly known French collector (see Zanardini, 1858:210). This species was placed in the synonymy of Chondrococcus hornemannii by Schmitz (1895:170). The status of Portieria as the first generic name unequivocally applied to the rhizophyllidaceous genus under consideration supports the adoption of this name in preference to manipulating the conservation of either Desmia or Chondrococcus.

The genus Portieria includes the following species.

Portieria? dichotoma (Hauck) P.C. Silva, new combination (Desmia dichotoma Hauck, 1886 [1886–1887]:218. fig. [s.n.]).

P. harveyi (J. Agardh) P.C. Silva, new combination (Desmia harveyi J. Agardh, 1876:356).

P. hornemannii (Lyngbye) P.C. Silva, new combination (Desmia hornemannii Lyngbye, 1819:35, pl. 7:c).

P. japonica (Harvey) P.C. Silva, new combination (Desmia japonica Harvey, 1860a:331).

P. kilneri (J. Agardh) P.C. Silva, new combination (Desmia kilneri J. Agardh, 1876:355).

P. spinulosa (Kützing) P.C. Silva, new combination (Chondrococcus spinulosus Kützing, 1868:11, pl. 32: figs. a–c).

It may be noted that Papenfuss (1940:218) recognized Desmia tripinnata as a separate species, but D. Reid Wiseman, in a recent study of material in the Herbarium of the University of California at Berkeley, retained it within the circumscription of Chondrococcus hornemannii.

Eupogodon as a Replacement for Dasyopsis

The genus Dasyopsis is always attributed to Zanardini (1843:52). In the place cited, however, there is no description. Zanardini merely stated that he was persuaded to segregate Dasya plana C. Agardh and D. spinella C. Agardh into their own genus by the position, form, and disposition of the stichidia, without giving the stichidial characters.

Montagne (1844b:611) prepared a special article on Dasyopsis for d’Orbigny’s Dictionnaire Universelle d’Histoire Naturelle, but without giving a description. In his Flore d’Algérie, Montagne (1846b:85) reduced Dasyopsis to a subgenus of Dasya and validated the name at subgeneric rank by providing a diagnosis. The following year, Montagne (1847:54) recognized Dasyopsis as a genus.

Meanwhile, Kützing (1845:312), arriving independently at the taxonomic conclusion that Dasya plana should be segregated from Dasya, established the genus Eupogodon. He later (Kützing, 1849:801) added Dasya spinella and D. cervicornis J. Agardh to the genus. Kützing provided a diagnosis in his original publication of Eupogodon, and thus this name takes priority over Dasyopsis (Montagne) Montagne. Kützing originally treated Eupogodon (“true beard-tooth”) as a neuter name, but in fact it is masculine, as he himself recognized in subsequent publications.

The genus Eupogodon includes the following species.

Eupogodon anastomosans (Weber-van Bosse) P.C. Silva, new combination (Dasyopsis anastomosans Weber-van Bosse, 1921:309, pl. VII: fig. X).

E. antillarum (Howe) P.C. Silva, new combination (Dasyopsis antillarum Howe, 1920:577).

E. apertus (Weber-van Bosse) P.C. Silva, new combination (Dasyopsis aperta Weber-van Bosse, 1913b: 128, pl. 13: fig. 17; pl. 14: fig. 32).

E. apiculatus (C. Agardh) P.C. Silva, new combination (Bonnemaisonia apiculata C. Agardh, 1835: no. XXXIX, pl. 39).

E. cervicornis (J. Agardh) Kützing, 1849:802.

E. geppii (Weber-van Bosse) P.C. Silva, new combination (Dasyopsis geppii Weber-van Bosse, 1913b:130, pl. 13: figs. 18–20; pl. 14: fig. 33).

E. palmatifidus (Weber-van Bosse) P.C. Silva, new combination (Dasyopsis palmatifida Weber-van Bosse, 1913b: 130, pl. 13: fig. 21).

E. penicillatus (Zanardini) P.C. Silva, new combination (Dasya penicillata Zanardini, 1865:379, pl. XIV [XLI]).

E. pilosus (Weber-van Bosse) P.C. Silva, new combination (Dasyopsis pilosa Weber-van Bosse, 1923:377, fig. 137).

E. pinnatifolius (Suhr) P.C. Silva, new combination (Ptilota ? pinnatifolia Suhr, 1834:732, pl. II: figs. 18 and t).

E. planus (C. Agardh) Kützing, 1845:312.

E. pulchellus (Weber-van Bosse) P.C. Silva, new combination (Dasyopsis pulchella Weber-van Bosse, 1921:309, pl. VII: fig. VII).

E. spinellus (C. Agardh) Kützing, 1849:801.

E. stanleyi (Weber-van Bosse) P.C. Silva, new combination (Dasyopsis stanleyi Weber-van Bosse, 1913b: 128, pl. 13: fig. 16).

E. tenellus (Weber-van Bosse) P.C. Silva, new combination (Dasyopsis tenella Weber-van Bosse, 1921:309, pl. VII: fig. VI).

Hincksia as a Replacement for Giffordia

While John Edward Gray was Keeper of the Zoology Department of the British Museum, he prepared a catalog “merely to enable Mrs. Gray to arrange her collection of British Algae according to the most recent methods.” In this scholarly but generally overlooked work, Gray (1864) defined some common filamentous genera more narrowly than was customary at that time, creating new genera among blue-green, red, brown, and green algae. One of the genera segregated from Ectocarpus was Hincksia, to which Gray assigned a single species, H. ramulosa, based on Ectocarpus hincksiae Harvey (1841:40). This species was transferred to Giffordia Batters (1893:86) by Hamel (1939:XV). Gray (1864:12) characterized Hincksia as follows: “Frond secundly branched; fruit conical, sessile, produced along the inner face of the branches and ramuli, one arising from almost every joint, giving the branch a serrated appearance.” Although this diagnosis lacks two essential generic characters (discoid plastids and lack or irregular occurrence of meristematic regions), it is clear that Gray had a species of Giffordia in hand and that Hincksia should supersede Giffordia. Both generic names commemorate female British phycologists, Hannah Hincks (1798–1871) of Belfast and Isabella Gifford (1823?–1891) of Minehead, Somerset, respectively.

The genus Hincksia includes the following species.

Hincksia andamanensis (Krishnamurthy and Baluswami) P.C. Silva, new combination (Giffordia andamanensis Krishnamurthy and Baluswami, 1982:104, figs. 1–11; 1983:47).

H. bhimlipatnamensis (Krishnamurthy and Baluswami) P.C. Silva, new combination (Giffordia bhimlipatnamensis Krishnamurthy and Baluswami, 1982:105, figs. 12–15; 1983:47).

H. clavata (Krishnamurthy and Baluswami) P.C. Silva, new combination (Giffordia clavata Krishnamurthy and Baluswami, 1982:106, figs. 23–28; 1983:48).

H. fenestrata (Berkeley ex Harvey) P.C. Silva, new combination (Ectocarpus fenestratus Berkeley ex Harvey, 1849 [1847–1851]: pl. CCLVII; 1849:58).

H. fuscata (Zanardini) P.C. Silva, new combination (Ectocarpus fuscatus Zanardini in Meneghini, 1846:381).

H. ghardaqaensis (Nasr) P.C. Silva, new combination (Ectocarpus ghardaqaensis Nasr, 1939:59, figs. 8, 9).

H. granulosa (J.E. Smith) P.C. Silva, new combination (Conferva granulose J.E. Smith, 1811 [1811–1812]: pl. 2351).

H. hincksiae (Harvey) P.C. Silva, new combination (Ectocarpus hincksiae Harvey, 1841:40).

H. hincksiae var. californica (Hollenberg and Abbott) P.C. Silva, new combination (Giffordia hincksiae var. californica Hollenberg and Abbott, 1968:1238, fig. 3).

H. intermedia (Rosenvinge) P.C. Silva, new combination (Ectocarpus ovatus var. intermedius Rosenvinge in Rosenvinge and S. Lund, 1941:49, figs. 21, 22).

Hi mitchelliae (Harvey) P.C. Silva, new combination (Ectocarpus mitchelliae Harvey, 1852:142, pl. XII:G, “mitchellae”).

H. mitchelliae var. neilii (Krishnamurthy and Baluswami) P.C. Silva, new combination (Giffordia mitchelliae var. neilii Krishnamurthy and Baluswami, 1982:106, figs. 16–22; 1983:47).

H. onslowensis (Amsler and Kapraun) P.C. Silva, new combination (Giffordia onslowensis Amsler and Kapraun, 1985:94, figs. 1–3).

H. ovata (Kjellman) P.C. Silva, new combination (Ectocarpus ovatus Kjellman, 1877:35).

H. prolifera (Krishnamurthy and Baluswami) P.C. Silva, new combination (Giffordia prolifera Krishnamurthy and Baluswami, 1982:107, figs. 29–41; 1983:48).

H. rallsiae (Vickers) P.C. Silva, new combination (Ectocarpus rallsiae Vickers, 1905:59).

H. sandriana (Zanardini) P.C. Silva, new combination (Ectocarpus sandrianus Zanardini, 1843:41).

H. saundersii (Setchell and Gardner) P.C. Silva, new combination (Ectocarpus saundersii Setchell and Gardner, 1922:411).

H. secunda (Kützing) P.C. Silva, new combination (Ectocarpus secundus Kützing, 1847a:54).

H. sordida (Harvey) P.C. Silva, new combination (Ectocarpus sordidus Harvey, 1860c:294).

H. terminalis (Krishnamurthy and Baluswami) P.C. Silva, new combination (Giffordia terminalis Krishnamurthy and Baluswami, 1982:108, figs. 46–57; 1983:48).

H. thyrsoidea (Børgesen) P.C. Silva, new combination (Ectocarpus thyrsoideus Børgesen, 1937:12, figs. 6, 7).

The Correct Name for Hormophysa triquetra

The nomenclature of this species was investigated by Papenfuss (1968), who, after considering several synonyms, concluded that the correct name was Hormophysa triquetra (C. Agardh) Kützing. As will be shown, this name must be superseded.

When uncritically compiling the hitherto described species of Fucus, J.F. Gmelin (1792) confronted two pairs of homonyms, both now believed to involve Hormophysa triquetra. One pair was Fucus triqueter S.G. Gmelin (1768:122, pl. VIII: fig. 4) and F. triqueter Linnaeus (1771:312). Swartz (1788:148) had previously attempted to rectify the homonymy, but erred in proposing a substitute name, Fucus trifarius, for the senior homonym rather than the junior homonym. The first species is now known as Bryothamnion triquetrum (S.G. Gmelin) Howe (1915:222), a rhodomelaceous red alga from the Caribbean. Fucus triqueter Linnaeus was the first description of a species of Hormophysa. J.F. Gmelin (1792:1382, 1383) erred in the same manner as Swartz, retaining the junior homonym and substituting the name Fucus triangularis for the Caribbean alga (which was called Bryothamnion triangulare until Howe pointed out the illegitimacy of that name). On transferring F. triqueter Linnaeus to Cystoseira, C. Agardh (1820b:61) provided a binomial (C. triquetra) that may be treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN.

The second pair of homonyms was Fucus articulatus S.G. Gmelin (1768:77) and F. articulatus Forsskål (1775:191). The first species was described, but not illustrated (the reference to pl. II: fig. 1, apparently being erroneous, judging from the legend and a comparison of the cited figure with the description). The provenance was said to be “Oceanus Indicus.” In the absence of a type specimen, a definitive identification is not possible. The Forsskål species was united with F. triqueter Linnaeus in the genus Cystoseira by C. Agardh (1820b: 61), and subsequent authors have considered the two species as congeneric, but not always conspecific. J. Agardh (1848:215–216) recognized separate species, and on transferring F. articulatus Forsskål to Cystoseira, provided a binomial (C. articulata) that may be treated as a nomen novum in accordance with Article 72, Note 1, of the ICBN. On the basis of priority or perhaps purely by chance, J.F. Gmelin (1792:1381, 1389) retained the senior homonym and substituted the name Fucus cuneiformis for F. articulatus Forsskål. By an examination of Forsskål’s collection at Copenhagen (C), Børgesen (1932:11) confirmed that Fucus articulatus Forsskål is referable to Hormophysa. Hence, Fucus cuneiformis is the earliest legitimate name for this species and its correct name is Hormophysa cuneiformis (J.F. Gmelin) P.C. Silva, new combination.

Validation of the Green Algal Order Ctendocladales

The order Ctenocladales was proposed by Silva (1982:147) to accommodate those green algae previously placed in the Chaetophoraceae with ulvophycean rather than chlorophycean ultrastructure. The nature of the publication (an encyclopedia) precluded the provision of a Latin diagnosis essential to the validity of the name. Validation is effected herewith: Chlorophytae (algae virides) filamenta uniseriata ramosa aut sarcinas cellularum constantes; thallus erectus aut prostratus aut heterotrichus, filamentis prostratis libris aut cohaerentibus; cellulae omnes nucleum unum et chloroplastum singulum parietalem laminatem plerumque pyrenoidibus thylakoidiperductis (uno aut aliquot) continentes; cellulae motiles squamatae aut nudae cum uno aut multis flagellis; flagella apicaliter affixa, symmetrica radialiter, cum systemate radicum microtubulare cruciato, radicibus rotationem contra horologii habentibus; cytokinesis per sulco cum phycoplasto nihil consociato effectus.

Familia typificata: Ctenocladaceae (Borzi) Borzi (1892:48, “Ctenocladiaceae”).

Literature Cited

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1904. Über die Anheftung der Cladophoraceen und über verschiedene polynesische Formen dieser Familie. Beihefte zum Botanischen Centralblatt, 18(Abteilung I):165–193, plates V, VI.

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*1983. The Morphology, Growth and Seasonality in the Reproductive States of Acanthophora spicifera (Vahl) Boergesen in Bacoor Bay. Natural and Applied Science Bulletin [University of the Philippines], 35:17–27, 5 figures.

Burgan, B.G., K.A. Zseleczky, and F.J. Vande Vusse

*1980. An Illustrated Key to Philippine Rabbitfishes, Siganus Species, with Descriptive and Ecological Notes. Sillíman Journal, 26:99–114, 14 figures.

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1768. Flora indica: cui accedit series zoophytorum indicorum, nee non prodromus florae capensis. Pages [i–iv] + 1-241 + [1–4 + 1–11] + 1–28, 25[bis] –28[bis], plates 1–21, 21*, 22–67. Lugdunum Batavorum [Leiden] and Amstelodamum [Amsterdam].

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*1981. A Seaweed Primer. The Filipinos Journal of Science and Culture, 1:54–59.

Cajipe, G.J.B., E.C. Laserna, R.L. Veroy, and A.H. Luistro

*1980. On the Infrared Spectrum of a Polysaccharide Obtained by Alkaline Extraction of the Red Alga Acanthophora spicifera (Vahl) Boergesen. Botanica Marina, 23:69–70, 2 figures.

Calumpong, H.P.

*1980. Some Aspects of the Ecology of the Sea Hare, Dolabella auricularia (Lightfoot), in Central Visayas, Philippines. Silliman Journal, 26:134–146, 3 figures, 3 tables.

*1982. Economically Important Species of Benthic Marine Algae in the Central Visayas, Philippines. Silliman Journal, 28:143–148.

Cantoria, M.

*1948. Red Algae as Sources of Agar. Journal of the Philippine Pharmaceutical Association, 35:26–27. [Abstract.]

Cantoria, M., and P. Valenzuela

*1950. Agar from Philippine Hypnea and Gracilaria. Journal of the Philippine Pharmaceutical Association, 37:271.

Cantoria, M., P. Valenzuela, and G.T. Velasquez

*1951. Pharmacopeial Properties of Agar from Three Philippine Seaweeds. Journal of the Philippine Pharmaceutical Association, 38:187–190, 3 figures.

Cantoria, M., G.T. Velasquez, and P. Valenzuela

*1951. A Botanical Study of Three Philippine Agar-yielding Seaweeds. Journal of the Philippine Pharmaceutical Association, 38:295–300, 3 figures.

Carumbana, E.E., and J. A. Luchavez

*1980. A Comparative Study of the Growth Rates of Siganus canaliculatus, S. spinus, and S. guttatus Reared under Laboratory and Seminatural Conditions in Southern Negros Oriental, Philippines. Silliman Journal, 26:187–209, 10 figures.

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1851. Supplement au catalogue des plantes qui croissent naturellement aux environs de Marseille. 125 pages, plates 8–11. Aix.

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*1979. Taxonomic Study on the Marine Benthic Algae of Borongan, and Vicinities with Notes on Their Economic Potential. Journal of Graduate Research, Graduate School of the University of Santo Tomas, 9:91–92. [Abstract of master s thesis.]

Chamberlain, Y.M.

1983. Studies in the Corallinaceae with Special Reference to Fosliella and Pneophyllum in the British Isles. Bulletin of the British Museum (Natural History), Botany, 11:291–463, 89 figures.

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*1981. Species Diversity and Zonation of Epiphytes on Two Species of Sargassum. In Proceedings of the Fourth International Coral Reef Symposium [Manila, 1981], volume 2, pages 385–392, 9 figures.

Chang, C.F., and B.M. Xia

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1937. A Revision of the Marine Algae of Norfolk. Journal of the Linnean Society of London, Botany, 51:205–263, 37 figures, plate 22.

1961. The Marine Algae of Jamaica, Part 1: Myxophyceae and Chlorophyceae. Bulletin of the Institute of Jamaica, Science Series, 12(1): 159 pages, 178 figures.

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1842. Recherches sur l’organisation, la fructification et la classification de plusieurs genres d’algues. 132 pages. Caen.

Chiang, Y.M.

1970. Morphological Studies of Red Algae of the Family Cryptonemiaceae. University of California Publications in Botany, 58: vi + 83 pages, 34 figures, 10 plates.

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*1945. Pacific Species of Galaxaura, I: Asexual Types. Papers of the Michigan Academy of Science, Arts and Letters, 30:35–56, 2 figures, XI plates.

*1947. Pacific Species of Galaxaura, II: Sexual Types. Papers of the Michigan Academy of Science, Arts and Letters, 31:3–24, 3 figures, XIII plates.

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1807. Ensayo sobre las variedades de la vid comun que vegetan en Andalucía. xviii + 324 pages. Madrid.

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*1974. A Study of the Culture of Eucheuma striatum in the Visayan Area. Philippine Scientist [University of San Carlos], 11:109. [Abstract of thesis.]

*1975. Investigations on the Culture of Eucheuma striatum in the Philippines. Philippine Scientist [University of San Carlos], 12:125. [Abstract of thesis.]

*1976. Studies on the Culture of Eucheuma striatum. Philippine Scientist [University of San Carlos], 13:48–61, 4 figures.

Colinvaux, L.H.

1968. A New Species of Halimeda: A Taxonomic Reappraisal. Journal of Phycology, 4:30–35.

Collado, E.G.

*1926. Studies on the Nutritive Properties of Seaweeds. Philippine Agriculturist, 15:129–148, 3 figures.

Collins, F.S.

1901. The Algae of Jamaica. Proceedings of the American Academy of Arts and Sciences, 37:229–270, VI tables.

1909. The Green Algae of North America. Tufts College Studies (Science), 2:79–480, XVIII plates.

Collins, F.S., and A.B. Hervey

1917. The Algae of Bermuda. Proceedings of the American Academy of Arts and Sciences, 53:1–195, including VI plates.

Collins, F.S., I. Holden, and W.A. Setchell

1899. Phycotheca Boreali-Americana. Fascicle XIII, numbers 601–650. Maiden, Massachusetts. [Exsiccatae.]

Cordero, P. A., Jr.

*1972. Philippine Algology: Its Beginnings and Development. Leyte-Samar Studies, 6:16–47.

*1973a. A Noteworthy Halimeda Species from Eastern Samar. Kalikasan, Philippine Journal of Biology, 1:167–170, 4 figures.

*1973b. On the Marine Algae of Biliran Island (Leyte) and Vicinities, Central Philippines. Leyte-Samar Studies, 7:15–37.

*1974a. A New Variety of Halimeda velasquezii Taylor from Camiguin Island. Kalikasan, Philippine Journal of Biology, 2:114–117, 1 figure.

*1974b. [Distribution of Caulerpa brachypus Harvey (Chlorophyta) in Japan and Adjacent Regions.] Nankiseibutu [Journal of the Nanki Biological Society], 16:6–9, 2 figures. [In Japanese.]

*1974c. Phycological Observations, I: Genus Porphyra of the Philippines, Its Species and Their Occurrences. Bulletin of the Japanese Society of Phycology, 22:134–142, 4 figures.

*1975. Phycological Observations, III: On the Occurrence of Genus Actinotrichia in the Philippines. Publications of the Seto Marine Biological Laboratory, 22:267–276, 3 figures.

*1976a. Phycological Observations, IV: Two Noteworthy Species of Caulerpa (Chlorophyta) from the Philippines. Publications of the Seto Marine Biological Laboratory, 23:83–87, 2 figures.

*1976b. Phycological Observations, II: Porphyra marcosii, a New Species from the Philippines. Acta Manilana [University of Santo Tomas Research Center], series A, 15(24): 14–24, figures A–H, including I plate.

*1976c. The Marine Algae of Batan Island, Northern Philippines, I: Cyanophyceae and Phaeophyceae. Fisheries Research Journal of the Philippines, 1(2): 3–29, figures A–O.

*1977a. Studies on Philippine Marine Red Algae. Special Publications from the Seto Marine Biological Laboratory, 4: 258 pages, 268 figures, XXVIII plates.

*1977b. The Marine Algae of Batan Island, Northern Philippines, II: Chlorophyceae. Fisheries Research Journal of the Philippines, 2(1):19–55, including VII plates [= 45 figures].

*1977c. Phycological Observations, V: Gross Morphological Polymorphisms in Caulerpa brachypus (Caulerpales, Chlorophyta) from the Philippines, with Notes on Their Taxonomy. Bulletin of the Japanese Society of Phycology, 25(supplement):25–30, 3 figures.

*1978a. The Marine Algae of Batan Island, Northern Philippines, III: Rhodophyceae. Fisheries Research Journal of the Philippines, 3(1): 13–64.

*1978b. Phycological Observations, VIII: Additional Notes on Genus Caulerpa from the Philippines. Fisheries Research Journal of the Philippines, 3(2):44–51, 3 figures.

*1979a. Phycological Observations, IX: Additional Notes on the Occurrence and Seasonality of the Genus Porphyra in the Philippines. Acta Manilana [University of Santo Tomas Research Center], series A, 18:20–35, 4 figures.

*1979b. Phycological Observations, VI: Mangrove-associated Algae from Aklan, Philippines. Kalikasan, Philippine Journal of Biology, 7:275–296, 5 figures.

*1979c. Phycological Observations, X: On the Phyco-geographical Distribution of Two Codiaceous Marine Algae in the Philippines and Adjacent Regions. Fisheries Research Journal of the Philippines, 4(2):52–63, 3 figures.

1980a. Phycological Observations, VII: On the Occurrence and Phyco-geographical Distribution of Sargassum duplicatum and Halicoryne wrightii in the Philippines. Publications of the Seto Marine Biological Laboratory, 25:27-38, 4 figures.

*1980b. Taxonomy and Distribution of Philippine Useful Seaweeds. National Research Council of the Philippines, Bulletin, 81: [xiv +] 78 pages, figures 1–16, 18–25; 44 + [16] plates.

*1980c. Species of Marine Macro-algae New to the Philippines. Fisheries Research Journal of the Philippines, 5(2):69–72, 4 figures.

*1981a. Phycological Observations, XIV: Acrocystis nana Zanardini, a New Algal Record for the Philippines. Publications of the Seto Marine Biological Laboratory, 26:171–175, 2 figures.

*198lb. Phycological Observations, XIII: Two Unreported Marine Benthic Rhodophytans from Iloilo, Philippines. Acta Manilana [University of Santo Tomas Research Center], series A, 20:22–28, 2 figures.

*1981c. Eco-morphological Observation of the Genus Sargassum in Central Philippines, Including Notes on Their Biomass and Bed Determination. In Proceedings of the Fourth International Coral Reef Symposium [Manila, 1981], volume 2, pages 399–409, 1 figure.

*1981d. Some Epiphytic Marine Algae of Aklan, Philippines. Fisheries Research Journal of the Philippines, 6(2):59–67, 6 figures.

*1982a. A Preliminary Study on the “Ethnobotany of Philippine Edible Seaweeds, Especially from Ilocos Norte and Cagayan Provinces.” Acta Manilana [University of Santo Tomas Research Center], series A, 21:54–79, 14 figures.

*1982b. Phycological Observations, XI: Some Noteworthy Marine Benthic Phaeophyceans of the Philippines. Fisheries Research Journal of the Philippines, 7(2):31–37, 3 figures. [Printed X, changed by author to XI on reprint.]

*1983. Phycological Observations, XV: Occurrence of the Red Alga Balliella subcorticata in the Philippines. Kalikasan, Philippine Journal of Biology, 12:189–190, 1 figure.

*1984a. Phycological Observations, XX: Marine Algae in the Vicinity of the Bureau of Fisheries and Aquatic Resources Marine Station, Bobon Bay, Ilocos Norte. Ilocos Fisheries Journal, 1(2):67–117.

*1984b. Phycological Observations, XVII: Records of Western Palawan Algae Particularly from Lipuun Point, Quezon and Nearby Islets. Pulong [Divine Word University, Tacloban], 7:55–69.

*1984c. Phycological Observations, XVIII: Some Marine Macro-algae from Southern Masbate. Pulong [Divine Word University, Tacloban], 8:49–58. [Printed XXII, changed by author to XVIII on reprint.]

Cordero, P.A., Jr., and T. Tanaka

*1972. Genus Halimeda from Camiguin Island, Northern Philippines. Bulletin of the Japanese Society of Phycology, 20:83–89, 2 figures.

Cornejo, D.F., and G.T. Velasquez

*1972. Study on the Algal Epiphytes of Exposed and Protected Marine Waters of Batangas Province. Philippine Journal of Science, 99:165–190, 1 figure, 5 plates.

Cribb, A.B.

1958. Records of Marine Algae from South-eastern Queensland, IV: Caulerpa. University of Queensland, Department of Botany, Papers, 3:209–220, including 5 plates.

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1944. The Marine Algae of the Gulf of California. Allan Hancock Pacific Expeditions, 3:189–453, including plates 31–77.

1949a. Studies of Northeast Pacific Gracilariaceae. Allan Hancock Foundation Publications, Occasional Papers, 7: 105 pages, including 25 plates.

1949b. Contributions toward a Marine Flora of the Southern California Channel Islands, I–III. Allan Hancock Foundation Publications, Occasional Papers, 8: 57 pages, including 15 plates.

1950a. Notes on Some Pacific Mexican Dictyotaceae. Bulletin of the Torrey Botanical Club, 77:83–93, 3 figures.

1950b. Notes on Pacific Coast Marine Algae, IV. American Journal of Botany, 37:149–158, 29 figures.

1950c. A Review of Ceramium along the Pacific Coast of North America with Special Reference to Its Mexican Representatives. Farlowia, 4:113–138, including 4 plates.

1953. Marine Red Algae of Pacific Mexico, Part I: Bangiales to Corallinaceae subf. Corallinoideae. Allan Hancock Pacific Expeditions, 17:1–239, including plates 1–33.

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1954b. The Marine Flora of Isla San Benedicto following the Volcanic Eruption of 1952–1953. Allan Hancock Foundation Publications, Occasional Papers, 16: 25 pages, including 5 plates.

1957. An Annotated List of Marine Algae from Eniwetok Atoll, Marshall Islands. Pacific Science, 11:92–132, 31 figures.

1958. Notes on Pacific Coast Marine Algae, VII. Bulletin of the Southern California Academy of Sciences, 57:65–80, 12 figures.

1960. New Records of Marine Algae from Pacific Mexico and Central America. Pacific Naturalist, 1(19/20):31–52, 7 figures.

1963. New Records of Marine Algae from the Galapagos Islands. Pacific Naturalist, 4:3–23, including 5 plates.

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1964. The Seaweeds of Peru. Beihefte zur Nova Hedwigia, 13: 111 pages, 81 plates.

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1839. Note sur le genre Amansia. Annales des Sciences Naturelles, Botanique, series 2, 11:373–376.

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*1974. Studies on the Carrageenan Content of Various Species of Eucheuma and Other Allied Genera. Acta Manilana [University of Santo Tomas Research Center], series A, 12:30–47, [18] figures.

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*1971. Studies on Some Philippine Rhodophytes and Their Colloidal Contents. Acta Manilana [University of Santo Tomas Research Center], series A, 8:3–38, including 9 plates.

de Leon, W., J. Leyva, and L. Martinez-Pesigan

*1947. The Possibility of Agar-agar Industry in the Philippines. Journal of the Philippine Pharmaceutical Association, 34:381–384.

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1813. Flore d’Égypte, Explication des planches. In France, Commission d’Égypte, Description de l’Égypte: Histoire naturelle, volume 2, pages 145–320. [Atlas, 62 plates, 1826.] Paris.

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*1967. Observations on Some Economically Important Algae of Biliran Island. Leyte-Samar Studies, 1:228–235, 2 figures, [1] plate.

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1968. Les Algues Floridées encroutantes (à l’exclusion des Corallinacées). 310 pages, 227 figures. Paris: privately published.

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1798–1799. Flora atlantica. Volume 2, 458 pages, plates 121–261. Parisii [Paris]. [1798 = pages 1–160, plates 121–180; 1799 = pages 161–458, plates 181–261.]

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*1889. Chlorophyceae. In Sylloge algarum, volume I, cxxxix + 1315 pages. Patavium [Padua].

1891. Systematische Uebersicht der bisher bekannten Gattungen der echten Fucoideen. Flora, 74:171–182.

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1895. Neue Meeresalgen aus Japan. Berichte der Deutschen Botanischen Gesellschaft, 12:(72)–(78), plate XVI.

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1936. Noterelle di nomenclatura algologica, VIII: Terzo elenco di Missoficee omonime. [6] pages. Brescia: privately published.

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1969a. Ceramiella jolyi, a New Species of Rhodophyta from Puerto Rico. Caribbean Journal of Science, 8:199–205, including II plates.

1969b. Caulerpa hummii, a New Species of Marine Algae (Chlorophyta, Caulerpales) from Venezuela. Phycologia, 7:12–17, 1 figure.

1969c. The Genus Ceramiella in the Caribbean and Its Biological Significance. Caribbean Journal of Science, 9:53–58.

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*1874a. On the Algae of Mauritius. Journal of the Linnean Society of London, Botany, 14:190–202.

1874b. [Enumeration of the Algae Collected at St. Thomas and Bermuda by H.N. Moseley, M.A., Naturalist to H.M.S. “Challenger.”] Journal of the Linnean Society of London, Botany, 14:312–316.

*1876a. Contributions to the Botany of the Expedition of H.M.S. “Challenger:”—Algae, Chiefly Polynesian. Journal of the Linnean Society of London, Botany, 15:235–246.

*1876b. Notes on Algae Collected by H.N. Moseley, M.A., of H.M.S. “Challenger,” Chiefly Obtained in Torres Straits, Coasts of Japan, and Juan Fernandez. Journal of the Linnean Society of London, Botany, 15:446–455.

*1877. Supplemental Notes on Algae Collected by H.N. Moseley, M.A., of H.M.S. “Challenger,” from Various Localities. Journal of the Linnean Society of London, Botany, 15:486–489.

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*1981. The Culture of Seaweeds in the Philippines with Emphasis on Eucheuma Culture in Bohol Province. In G.C. Trono, Jr. and E.T. Ganzon-Fortes, editors, Report on the Training Course on Gracilaria Algae, pages 165–170. Manila. [United Nations, Food and Agriculture Organization, South China Sea Fisheries Development and Coordinating Programme, Work Plan Implementation (General), SCS/GEN/81/29.]

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*1973. Seaweed Farms: A New Approach for U.S. Industry. In Proceedings of the 9th Annual Conference of the Marine Technology Society [Washington, D.C., 1973], pages 701–708, 3 figures.

*1981. Eucheuma Farm Productivity. In Proceedings of the Eighth International Seaweed Symposium [Bangor, North Wales, 1974], pages 688–691.

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*1985. Eucheuma Species (Solieriaceae, Rhodophyta) That Are Major Sources of Carrageenan. In I.A. Abbott and J.N. Norris, editors, Taxonomy of Economic Seaweeds with Reference to Some Pacific and Caribbean Species, pages 47–67, 6 figures. [California Sea Grant College Program Report, T–CSGCP–011.]

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1928. A Revision of the Genera Chantransia, Rhodochorton, and Acrochaetium, with Descriptions of the Marine Species of Rhodochorton (Naeg.) gen. emend, on the Pacific Coast of North America. University of California Publications in Botany, 14:139–224, plates 37–48.

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*1968. Revision of the Classification of the Oscillatoriaceae. Academy of Natural Sciences of Philadelphia, Monographs, 15: [vi] + 370 pages, 131 figures.

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1901. Die Rhodomelaceen des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. Fauna und Flora des Golfes von Neapel, Monographie, 26: xvi + 754 pages, 10 figures, 24 plates. Berlin.

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*1956. Revision of Calothrix Ag. Revue Algologique, series 2, 2:154–178, 6 figures

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1876. List of the Marine Algae of the United States. Report of the United States Fish Commissioner, 1873/1875:691–718.

1877. On Some Algae New to the United States. Proceedings of the American Academy of Arts and Sciences, 12:235–245.

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1945. Révision du genre Botryocladia Kylin (Rhodophycées-Rhodyméniacées).Bulletin de la Societé d’Histoire Naturelle de l’Afrique du Nord, 35:49–61, 5 figures.

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1938. Sur un nouveau genre de Siphonocladacées. Comptes Rendus Hebdomadaires des Séances de l’Academie des Sciences [Paris], 206:1503–1504.

1941. Les algues marines de la Côte des Albères, IV: Rhodophycées (suite). Revue Algologique, 12:77–100, figures 26–34.

1950. Sur l’existence d’une alternance de générations entre l’Halicystis parvula Schmitz et le Derbesia tenuissima (De Not.) Crn. Comptes Rendus Hebdomadaires des Séances de l’Academie des Sciences [Paris], 230:322–323.

1955. Les plastes des Caulerpa et leur valeur systématique. Revue Generale de Botanique, 62:422–431, 2 figures.

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1943. Recherches sur les Bonnemaisoniacées et leur alternance de générations. Annales des Sciences Naturelles, Botanique, series 11, 3:75–175, 26 figures, 2 plates.

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1934. Observations sur quelques Gélidiacées. Revue Générale de Botanique, 46:528–549, 11 figures.

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1941. Recherches sur les Céramiacées de la Mediterranée occidentale. 510 pages, 191 figures, IV plates. [Printing finished 6 June 1940, but not used as thesis until 17 May 1941.]

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1952. A Revision of the Genus Cystophyllum. Summaries of Doctoral Dissertations… Submitted to the Graduate School of Northwestern University, 19:560–563.

1955. An Emendation of the Genus Cystophyllum (Fucales). American Journal of Botany, 42:305–322, 51 figures.

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1853. Nuovi materiali per l’algologia del mar Rosso. Memorie della Reale Accademia delle Scienze di Torino, series 2, 13:133–169, 1 plate.

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1775. Flora aegyptiaco-arabica: Post mortem auctoris edidit Carsten Niebuhr. 32 + cxxvi + 219 pages. Haunia [Copenhagen].

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*1980. Marine Algal Microphytes New to the Philippines. Kalikasan, Philippine Journal of Biology, 8:51–68, 15 figures.

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1895. New or Critical Lithothamnia. Kongelige Norske Videnskabers Selskabs Skrifter, 1895:1–10, 1 plate.

1897. On Some Lithothamnia. Kongelige Norske Videnskabers Selskabs Skrifter, 1897(1): 20 pages.

1898. Systematical Survey of the Lithothamnia. Kongelige Norske Videnskabers Selskabs Skrifter, 1898(2): 7 pages.

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1900b. Calcareous Algae from Funafuti. Kongelige Norske Videnskabers Selskabs Skrifter, 1900(1): 12 pages.

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*1929. Contributions to a Monograph of the Lithothamnia: After the Author’s Death Collected and Edited by Henrik Printz. 60 pages, LXXV plates. Trondhjem: Kongelige Norske Videnskabers Selskab Museet.

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1930. Les Myxophycées de l’Afrique equatoriale francaise. Archives de Botanique, Mémoires, 3(2): 508 pages, 362 figures.

1932. Cyanophycees vivant dans le thalle des Codium. Comptes Rendus Hebdomadaires des Séances de l’Académie des Sciences [Paris], 195:1413–1414.

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1828. Thalassiophytes. Dictionnaire des Sciences Naturelles [Levrault], 53:350–406.

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*1981. Introduction to the Seaweeds: Their Characteristics and Economic Importance. In G.C. Trono, Jr., and E.T. Ganzon-Fortes, editors, Report on the Training Course on Gracilaria Algae, pages 17–25. Manila. [United Nations, Food and Agriculture Organization, South China Sea Fisheries Development and Coordinating Programme, Work Plan Implementation (General), SCS/ GEN/81/29.]

*1983. Laurencia tronoi (Rhodophyta, Ceramiales), a New Species from Calatagan, Batangas, Philippines. Kalikasan, Philippine Journal of Biology, 11:404–409, 2 figures.

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*1982. Reproductive Morphology and Periodicity of Laurencia sp. at Calatagan, Batangas, Philippines. Kalikasan, Philippine Journal of Biology, 11:27–38, 4 figures.

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1981. The Marine Algae of British Columbia and Northern Washington: Division Rhodophyta (Red Algae), Class Bangiophyceae. Syesis, 13:137–195, 15 figures.

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1982. Scanning Electron Microscopy of Corallina and Haliptilon (Corallinaceae, Rhodophyta): Surface Features and Their Taxonomic Implications. Journal of Phycology, 18:211–219, 13 figures.

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*1979. A Preliminary Report on the Screening of Marine Algae for Antibacterial Properties. Mindanao Journal, Mindanao State University, 5(3):41–45.

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1917. New Pacific Coast Marine Algae, I. University of California Publications in Botany, 6:377–416, plates 31–35.

1918. New Pacific Coast Marine Algae, III. University of California Publications in Botany, 6:455–486, plates 38–41.

1919. New Pacific Coast Marine Algae, IV. University of California Publications in Botany, 6:487–496, plate 42.

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1927b. New Rhodophyceae from the Pacific Coast of North America, VI. University of California Publications in Botany, 14:99–138, plates 20–36.

1932. The Myxophyceae of Porto Rico and the Virgin Islands. In New York Academy of Sciences, Scientific Survey of Porto Rico and the Virgin Islands, volume 8 (part 2), pages 249–311, plates 1, 2.

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1973. Experimental Investigations on Marine and Freshwater Bangia (Rhodophyta) from the Netherlands. Journal of Experimental Marine Biology and Ecology, 11:239–247, 6 Figures.

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1904. Rhipidosiphon and Callipsygma. Journal of Botany, 42:363–366, plate 467.

1908. Marine Algae (Chlorophyceae and Phaeophyceae) and Marine Phanerogams of the “Sealark” Expedition, Collected by J. Stanley Gardiner, M.A., F.R.S., F.L.S. Transactions of the Linnean Society of London, Botany, 7:163–188, plates 22–24.

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*1942. Notes on Caulerpa from Java and the Philippines. Papers of the Michigan Academy of Science, Arts and Letters, 27:7–26, 5 figures.

*1943. Studies on Philippine Chlorophyceae, I: The Dasycladaceae. Papers of the Michigan Academy of Science, Arts and Letters, 28:15–35, 3 figures.

*1946. Studies on Philippine Chlorophyceae, II: Survey of Literature and List of Recorded Species Prior to 1940. Bulletin of the Torrey Botanical Club, 73:73–79.

*1947. Studies on Philippine Chlorophyceae, III: The Codiaceae. Bulletin of the Torrey Botanical Club, 74:121–132, 1 figure.

*1961. An Annotated Checklist of Philippine Marine Chlorophyta. Philippine Journal of Science, 88:413–451, 1 figure, plate 1.

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*1969. Some Additional Records of Philippine Marine Chlorophyta. Micronesica, 5:121–130, 19 figures.

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1864. Handbook of British Water-weeds or Algae: The Diatomaceae by W. Carruthers. iv + 123 pages. London.

1866. On Anadyomene and Microdictyon, with the Description of Three New Allied Genera, Discovered by Menzies in the Gulf of Mexico. Journal of Botany, 4:41–51, 65–72, plate XLIV.

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*1867. Algae. In E. Fenzl, editor, Reise der Österreichischen Fregatte Novara µm die Erde in den Jahren 1857, 1858, 1859: Botanischer Theil, Erster Band: Sporenpflanzen, pages 1–104, plates I, Ia, II–XI.

1874. Algen der Fidschi-, Tonga- und Samoa-Inseln, gesammelt von Dr. E. Graeffe. Journal des Museums Godeffroy [Hamburg], 3:23–50.

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1924. Floridées de France, II. Revue Algologique, 1:427–457, figures III–VII.

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1897c. Melobesiae. Berichte der Deutschen Botanischen Gesellschaft, 15:403–420, plate XVIII.

1901. Die Lithothamnien des Museum d’histoire naturelle in Paris. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie, 28:529–545, plate XI.

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*1980. Ecology and Taxonomy of Halimeda: Primary Producer of Coral Reefs. Advances in Marine Biology, 17:1–327, 104 figures.

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*1982. A Taxonomic Revision of the American Species of Cladophora (Chlorophyceae) in the North Atlantic Ocean and Their Geographic Distribution. Verhandelingen der Koninklijke Nederlandse Akademie van Wetenschappen, Afdeeling Natuurkunde, Tweede Reeks, 78: 236 pages, including 39 plates, 4 figures.

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*1978. The Marine Benthic Algae of Manila Bay, II: Phaeophyta and Rhodophyta: Kaliksan, Philippine Journal of Biology, 7:7–30, 40 figures.

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1958. Phycological Notes, II. Bulletin of the Torrey Botanical Club, 85:63–69, 2 figures.

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*1968b. An Account of the Species of the Red Alga Polysiphonia of the Central and Western Tropical Pacific Ocean, II. Polysiphonia. Pacific Science, 22:198–207, 5 figures.

*1968c. An Account of the Species of the Red Alga Herposiphonia Occurring in the Central and Western Tropical Pacific Ocean. Pacific Science, 22:536–559, 25 figures.

*1968d. Phycological Notes, III: New Records of Marine Algae from the Central Tropical Pacific Ocean. Brittonia, 20:74–82, 7 figures.

Hollenberg, G.J., and I.A. Abbott

1966. Supplement to Smith’s Marine Algae of the Monterey Peninsula, xii + 130 pages, 53 figures. Stanford: Stanford University Press.

1968. New Species of Marine Algae from California. Canadian Journal of Botany, 46:1235–1251, 14 figures.

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Index

(Valid taxa (centered heads in the text) appear in roman; invalid taxa in italic.)

abrotanifolius, Fucus, 26

abscissus, Fucus, 129

Acanthocodium fragile, 112

Acanthophora, 60

aokii, 60

muscoides, 60

orientalis, 61

spicifera, 60

spicifera f. orientalis, 61

thierryi, 61

acanthophorus, Fucus, 61

acerosa, Gelidiella, 25

acerosus, Fucus, 25

Acetabularia, 121

calyculus, 121

clavata, 121

crenulata, 121

dentata, 121

exigua, 121

major, 121

minutissima, 122

moebii, 122

parvula, 122

philippinensis, 122

roxasii, 122

ryukyuensis, 122

ryukyuensis var. philippinensis, 122

velasquezii, 122

aciculare, Sargassum, 84

aciculare, Sargassum filifolium var., 84

Acrocarpus pusillus, 25

Acrochaetium, 8, 123, 127

daviesii, 128

gracile, 18

hallandicum, 19

hancockii, 19

liagorae, 19

nitidulum, 19

papenfussii, 19

robustum, 19

sargassi, 19

secundatum, 128

seriatum, 19

sinicola, 19

trichogloeae, 19

tuticorinense, 19

Acrocystis, 61

nana, 61

Actinotrichia, 22

fragilis, 22

rigida, 22

acuminata, Galaxaura, 22

acuminata, Grateloupia, 30

acuminata, Halymenia, 30

acutissimus, Microcoleus, 10, 12

adhaerens, Callophyllis, 31

adhaerens, Codium, 111, 112

adhaerens, Dasya, 59

adhaerens, Jania, 34

adnata, Callophyllis, 31

adnata, Caloglossa, 58

adnata, Delesseria, 58

adnata, Gelidiella, 25

aegagropila, Cladophora, 5

aegagropila, Valonia, 102

aemulum, Sargassum, 82

aerea, Chaetomorpha, 95

aerea, Conferva, 95

aeruginea, Calothrix, 13

aeruginea, Leibleinia, 13

aeruginosa, Anacystis, 7

aeruginosa, Gracilaria foliifera f., 42

aeruginosa, Gracilaria multipartita var., 42

aeruginosa, Palmogloea, 7

aeruginosus, Fucus, 42

aestuarii, Conferva, 9

aestuarii, Lyngbya, 9

affine, Ceramium, 54

affinis, Carpopeltis, 28

affinis, Gigartina, 28

africana, Roschera, 71

africanum, Rhizoclonium, 99

agardhianum, Microdictyon, 95

agardhianum, Sargassum, 82

Agardhiella, 44, 123

Aglaothamnion, 56

Ahnfeltia, 50, 123

concinna, 50

furcellata, 51

ajakii-assii, Eucheuma alvarezii var., 45

albida, Annulina, 97

albida, Cladophora, 97

albida, Conferva, 97

alcyonida, Eucheuma arnoldii var., 45

alsidii, Bangia, 17

alsidii, Goniotrichum, 17

Alsidium, 61

helminthochortos, 95

pusillum, 61

alvarezii, Eucheuma, 45

alvarezii var. ajakii-assii, Eucheuma, 45

alvarezii var. tambalangii, Eucheuma, 45

Amansia, 61, 123

glomerata, 61

jungermannioides, 68

amansii, Fucus, 25

amansii, Gelidium, 25

ambigua, Caulerpa, 104

ambigua, Desmia, 129

ambiguum, Phormidium, 9, 11, 14

amicorum, Caulerpa, 105

amicorum, Caulerpa cupressoides var. lycopodium f., 105

amoena, Amphithrix, 11

amoenum, Phormidium, 11

Amphiroa, 32, 123

anastomosans, 32

anceps, 32

annulata, 32

beauvoisii, 32

cultrata, 34

cumingii, 32

cyathifera, 33

dilatata, 32

ephedraea, 32

foliacea, 33

foliacea f. erecta, 33

fragilissima, 33

fragilissima f. cyathifera, 33

hancockii, 33

pacifica, 33

rigida, 33

subcylindrica, 33

tribulus, 33

valonioides, 34

zonata, 34

Amphithrix amoena, 11

Anabaena, 123

Anabaina, 8, 123

oscillarioides, 8

pseudoscillatoria, 8

Anacystis, 7

aeruginosa, 7

Anadyomene, 94, 125

brownii, 94

eseptata, 94

flabellata, 95

leclancheri, 94

obscura, 117

plicata, 94

stellata, 94

wrightii, 95

anastomosans, Amphiroa, 32

anastomosans, Cladophora, 101

anastomosans, Dasyopsis, 129

anastomosans, Eupogodon, 129

anastomosans, Struvea, 101

anceps, Amphiroa, 32

anceps, Corallina, 32

andamanensis, Giffordia, 130

andamanensis, Hincksia, 130

andersonii, Gracilaria, 43

andoi, Bostrychia, 62

angusta, Carpopeltis, 28

angustifolium, Sargassum binderi var., 82

angustus, Gymnogongrus ligulatus var., 28

annulata, Amphiroa, 32

annulata, Neomeris, 121

Annulina albida, 97

Anotrichium, 53

tenue, 53

antennina, Chaetomorpha, 95

antennina, Conferva, 95

anthoninii, Gelidium, 5

antillanum, Antithamnion, 53

antillarum, Dasyopsis, 60, 129

antillarum, Eupogodon, 60, 129

Antithamnion, 53, 123

antillanum, 53

Iherminieri, 53

subcorticatum, 54

aokii, Acanthophora, 60

aokii, Cladophora, 97

aperta, Dasyopsis, 129

apertus, Eupogodon, 129

Aphanocapsa, 123

apiculata, Bonnemaisonia, 129

apiculata, Galaxaura, 22

apiculata, Polysiphonia, 69

apiculatum, Centroceras, 54

apiculatus, Eupogodon, 129

arabicum, Codium, 111

aragoensis, Enteromorpha, 91

arborea, Galaxaura, 22

arborescens, Padina, 77

Archaeolithothamnion

erythraeum f. durum, 38

erythraeum f. molle, 38

schmidtii, 39

sibogae, 39

timorense, 39

arcuata, Gracilaria, 40

arcuata var. snackeyi, Gracilaria, 41

arenaria, Schizothrix, 13

arenicola, Caulerpa, 104

argentea, Udotea, 118

argentea var. spumosa, Udotea, 119

argus, Griffithsia, 57

argus, Wrangelia, 57

armata, Chondria, 62

armata, Lophura, 62

arnoldii, Eucheuma, 45

arnoldii var. alcyonida, Eucheuma, 45

articulata, Carpopeltis, 29

articulata, Cystoseira, 81, 131

articulata, Lomentaria, 52

articulata, Prionitis, 29

articulata, Ulva, 52

articulatus, Fucus, 81, 131

Asparagopsis, 21

delilei, 21

sanfordiana, 22

taxiformis, 21

Asperococcus

intricatus, 80

orientalis, 80

asperum, Spatoglossum, 79

Asterocytis, 123

ornata, 17

atomaria, Dictyota, 76

atomarius, Fucus, 76

atropurpurea, Bangia, 17

atropurpurea, Conferva, 17

atropurpurea, Porphyra, 18

atropurpurea, Ulva, 18

attenuata, Crouania, 56

attenuata, Derbesia, 104

attenuata, Mesogloia, 56

attenuatum, Hypoglossum, 58

australasicus, Chlorocladus, 120, 126

australasicus, Dasycladus, 120

australe f. brachiatum, Lithothamnion, 36

australe f. minutulum, Lithothamnion, 37

australe f. ubianum, Lithothamnion, 37

australis, Helminthocladia, 20

australis, Padina, 77

Audouinella, 127, 128

Avrainvillea, 117

capituliformis, 117

erecta, 117

lacerata, 117

lacerata var. robustior, 117

levis, 117

longicaulis, 117

nigricans, 117

obscura, 117

sordida, 117

bacciferum, Sargassum, 86

bacciferus, Fucus, 86

baccularia, Fucus, 82

baccularia, Sargassum, 82

baileyana, Chylocladia, 52

baileyana, Lomentaria, 52

baillouviana, Dasya, 59

baillouviana, Fucus, 59

Baillouviana punicea, 59

Balliella, 54

subcorticata, 54

Bangia, 15, 17

alsidii, 17

atropurpurea, 17

elegans, 17

fuscopurpurea, 18

yamadae, 18

bangioides, Erythrotrichia, 17

bartayresiana, Dictyota, 75

bartayresiana var. denticulata, Dictyota, 75

bartayresii, Dictyota, 75

bartlettii, Codium, 112

basilanicum, Sargassum siliquosum var., 88

batanensis, Claudea, 58

batanensis, Halimeda, 116

Batrachospermum requienii, 19

beaudettei, Polysiphonia, 69

beauvoisii, Amphiroa, 32

Beckerella, 25

scalaramosa, 25

belangeri, Sargassum, 82

benkulense, Sargassum ilicifolium f., 85

berberifolium, Sargassum, 82

bernardinum, Sargassum oligocystum var., 86

bernardinum, Sargassum oocyste var., 87

bhimlipatnamensis, Giffordia, 130

bhimlipatnamensis, Hincksia, 130

bicornutum, Sargassum siliquosum var., 88

bicuspidata, Wrangelia, 57

bidentata, Dictyota, 75

bifida, Champia, 53

biforme, Sargassum, 82

bikinensis, Halimeda, 114

binderi, Bostrychia, 61

binderi, Sargassum, 86

binderi var. angustifolium, Sargassum, 82

binderi var. vitiense, Sargassum, 86

biseriata, Erythrotrichia, 17

blodgettii, Gracilaria, 41

Boergesenia, 100, 125

forbesii, 100

boergesenii, Hypnea, 48

boergesenii, Liagora, 20

boergesenii, Microdictyon, 95

Bonnemaisonia apiculata, 129

bonnemaisonii, Oscillaria, 11

bonnemaisonii, Oscillatoria, 11

Boodlea, 100, 125

coacta, 100

composita, 100

montagnei, 100

siamensis, 100

struveoides, 100

vanbosseae, 5

Boodleopsis, 117

pusilla, 117

verticillata, 117

borealis, Rhodopeltis, 39

Bornetella, 119

nitida, 119

oligospora, 120

ovalis, 120

sphaerica, 120

bornetii, Tapeinodasya, 60

boryana, Caulerpa, 109

boryana, Caulerpa serrulata var., 109

boryana f. longifolia, Caulerpa serrulata var., 109

boryana f. occidentalis, Caulerpa freycinetii var., 109

boryana f. occidentalis, Caulerpa serrulata var., 109

boryana, Padina, 77

Bostrychia, 61

andoi, 62

binderi, 61

calliptera, 61

intricata, 61

kelanensis, 62

mixta, 62

mixta f. inermis, 62

moritziana, 62

radicans, 62

rivularis, 62

simpliciuscula, 62

tenella, 62

tenuis, 62

tenuis f. simpliciuscula, 62

Botryocarpa, 58

prolifera, 58

botryocarpus, Fucus, 58

Botryocladia, 51

botryoides, 51

kuckuckii, 51

pyriformis, 51

skottsbergii, 51

uvarioides, 51

botryoides, Botryocladia, 51

botryoides, Fucus, 51

brachiatum, Lithothamnion australe f., 36

brachygona, Chaetomorpha, 95

brachypus, Caulerpa, 104, 125

brachypus f. parvifolia, Caulerpa, 104

Brachytrichia, 15

maculans, 15

quoyi, 15

breviarticulata, Hincksia, 73

breviarticulatus, Ectocarpus, 73

brevior, Corallina ungulata f., 36

brevior, Jania ungulata f., 36

brevipes, Caulerpa plumaris var., 110

brevipes, Caulerpa sertularioides f., 110

brevis, Oscillatoria, 5

brongniartii, Laurencia, 64

brownii, Anadyomene, 94

brownii, Calomena, 94

Bryopsis, 103, 125

corticulans, 103

indica, 103

pachynema, 103

pennata, 103

pennata var. secunda, 103

plumosa, 103

plumosa var. secunda, 103

tenuissima, 104

Bryothamnion

triangulare, 131

triquetrum, 130

bullata, Dictyosphaeria cavernosa var., 102

bullata, Rivularia, 13, 14

bullata, Ulva, 14

bursa-pastoris, Fucus, 41

bursa-pastoris, Gracilaria, 41

byssoides, Lithophyllum, 36

byssoides, Lithothamnion, 36

byssoides, Nullipara, 36

byssoideum, Ceramium, 54, 55

byssoideum, Ceramium gracillimum var., 54

caenomyce, Liagora, 20

caenomyce, Yamadaella, 20

caespitosa, Catenella, 48

caespitosa, Champia, 53

caespitosa, Chlorodesmis, 118

caespitosa, Ulva, 48

calcarea, Millepora, 38

calcareum, Lithothamnion, 38

calcareum, Phymatolithon, 38

calcea, Peyssonnelia, 27

calcicola, Schizothrix, 9, 10, 11, 12

californica, Giffordia hincksiae var., 130

californica, Grateloupia, 30

californica, Hincksia hincksiae var., 130

californica, Kallymenia, 32

californica, Meredithia, 32

californica, Rhodymenia, 52

Calliarthron, 123

calliptera, Bostrychia, 61

calliptera, Rhodomela, 61

Callithamnion, 123

daviesii var. secundatum, 128

elegans, 57

Callophycus, 45

serratus, 45

Callophyllis, 31, 123, 129

adhaerens, 31

adnata, 31

chilensis, 32

japonica, 32

okamurae, 32

callophylloides, Kallymenia, 32

calodictyon, Polysiphonia, 71

calodictyon, Tolypiocladia, 71

Caloglossa, 58

adnata, 58

leprieurii, 58

leprieurii var. hookeri, 58

ogasawaraensis, 58

caloglossoides, Gelidium, 27

caloglossoides, Pterocladia, 27

Calomena brownii, 94

Calothrix, 13, 123

aeruginea, 13

comoides, 8

confervicola, 13

contarenii, 13

crustacea, 13, 14, 15

epiphytica, 13

hydnoides, 13

luteofusca, 10

parietina, 13, 14

pilosa, 14, 13, 15

robusta, 14

scopulorum, 14

semiplena, 10

sordida, 10

viguieri, 14

calyculus, Acetabularia, 121

camiguinensis, Dictyopteris, 74

canaliculata, Gracilaria, 41

canaliculatus, Sphaerococcus, 41

canariensis, Liagora, 20

cancellatus, Hydroclathrus, 80

cantharidosma, Lyngbya, 8

cantharidosmum, Hydrocoleum, 8

capense, Gelidium, 26

capensis, Fucus, 26

capensis, Turbinaria trialata var., 90

capillacea, Jania, 35

capillacea, Leibleinia, 10

capillacea, Pterocladia, 27

capillaceus, Fucus, 27

capillare, Rhizoclonium, 96

capitellata, Carpopeltis, 29

capitellata, Cryptonemia, 29

capituliformis, Avrainvillea, 117

capituliformis, Laurencia, 65

caraibica, Laurencia, 65

carolinensis, Laurencia, 65

carolyniae, Rhipiliopsis, 118

Carpacanthus

cystophyllus, 83

gaudichaudii, 84

ilicifolius, 85

microcystis, 85

spinulosus, 85

Carpopeltis, 28, 123

affinis, 28

angusta, 28

articulata, 29

capitellata, 29

cornea, 31

crispata, 29

divaricata, 29

flabellata, 29

formosana, 29

maillardii, 29

prolifera, 29

rigida, 29

carpophyllum, Sargassum, 85

cartilaginea, Laurencia, 65

cartilagineum, Gelidium, 26

cartilagineus, Fucus, 26

catenata, Cladophora, 97

catenata, Conferva, 97

Catenella, 48

caespitosa, 48

impudica, 48

nipae, 48

opuntia, 48

Caulerpa, 104, 125

ambigua, 104

amicorum, 105

arenicola, 104

boryana, 109

brachypus, 104, 125

brachypus f. parvifolia, 104

chemnitzia var. occidentalis, 108

clavifera, 106

clavifera var. gracilis, 107

clavifera var. turbinata, 108

corynephora, 107

crassifolia, 106

cupressoides, 104

cupressoides var. disticha, 105

cupressoides var. ericifolia, 105

cupressoides var. lycopodium, 105

cupressoides var. lycopodium f. amicorum, 105

cupressoides var. lycopodium f. disticha, 105

cupressoides var. lycopodium f. elegans, 105

elongata, 105

fastigiata, 105

fergusonii, 105

filicoides, 105

freycinetii, 109

freycinetii var. boryana f. occidentalis, 109

freycinetii var. pectinata, 109

freycinetii var. typica f. lata, 109

freycinetii var. typica f. spiralis, 109

hummii, 109

laetevirens, 107

lentillifera, 105

lentillifera var. compacta, 106

lentillifera var. longistipitata, 106

lessonii, 106

lycopodium, 105

macrodisca, 108

mexicana, 106

mexicana var. pluriseriata, 106

microphysa, 106

nummularia, 108

parvifolia, 104

peltata, 108

peltata var. macrodisca, 108

peltata var. nummularia, 108

pickeringii, 111

plumaris, 110

plumaris f. farlowii, 110

plumaris f. flagellata, 110

plumaris var. brevipes, 110

plumaris var. elegans, 105

plumaris var. longipes, 110

plumaris var. longiseta, 110

prolifera, 106

racemosa, 106

racemosa var. chemnitzia, 108

racemosa var. clavifera, 107

racemosa var. clavifera f. microphysa, 106

racemosa var. corynephora, 107

racemosa var. gracilis, 107

racemosa var. laetevirens, 107

racemosa var. lamourouxii, 107

racemosa var. macra, 107

racemosa var. macrophysa, 107

racemosa var. occidentalis, 108

racemosa var. peltata, 108

racemosa var. turbinata, 108

racemosa var. uvifera, 107

reyesii, 108

selago, 108

serrulata, 108

serrulata f. lata, 109

serrulata f. spiralis, 109

serrulata var. boryana, 109

serrulata var. boryana f. longifolia, 109

serrulata var. boryana f. occidentalis, 109

serrulata var. pectinata, 109

sertularioides, 109

sertularioides f. brevipes, 110

sertularioides f. farlowii, 110

sertularioides f. flagellata, 110

sertularioides f. longipes, 110

sertularioides f. longiseta, 110

simpliciuscula var. vesiculifera, 111

subserrata, 110

taxifolia, 111

taxifolia var. crassifolia, 106

urvilliana, 111

urvilliana var. vitiensis, 111

uvifera, 107

verticillata, 111

vesiculifera, 111

webbiana, 111

webbiana var. pickeringii, 111

cavernosa, Dictyosphaeria, 101

cavernosa, Ulva, 101

cavernosa var. bullata, Dictyosphaeria, 102

cenomyce, Hypnea, 48

cenomyce var. tenuis, Hypnea, 48

Centroceras, 54, 123

apiculatum, 54

clavulatum, 54

cryptacanthum, 54

hyalacanthum, 54

minutum, 54

Ceramiella, 54

procumbens, 54

Ceramium, 54, 123

affine, 54

byssoideum, 54, 55

clavulatum, 54

confervoides, 44

cruciatum, 54

diaphanum, 56

diaphanum var. tenuissimum, 56

equisetoides, 54

fastigiatum, 54

filamentosum, 49

fimbriatum, 55

flaccidum, 54

gracillimum, 55

gracillimum var. byssoideum, 54

loureiri, 55

luetzelburgii, 55

marshallense, 55

maryae, 55

masonii, 55

mazatlanense, 55

multijugum, 55

nitens, 55

ocellatum, 59

personatum, 55

procumbens, 54

rubrum var. nitens, 55

serpens, 55

sinicola, 55

taylorii, 56

tenerrimum, 56

tenuissimum, 56

vagabundum, 56

vagans, 56

zacae, 56

ceranoides f. leprosa, Liagora, 20

ceranoides, Liagora, 20

Ceratodictyon, 40

spongiosum, 40

cervicome, Sargassum, 82

cervicornis, Desmia, 129

cervicornis, Dictyota, 75

cervicornis, Eupogodon, 130

cervicornis, Hypnea, 49

ceylanica, Dictyota, 75

ceylanica, Halymenia, 31

ceylanica, Halymenia durvillaei var., 31

ceylanica, Laurencia, 65

ceylanica, Sarcodia, 45

Chaetangium, 128

fastigiatum, 128

Chaetomorpha, 95, 125

aerea, 95

antennina, 95

brachygona, 95

clavata, 5

clavata var. torta, 97

crassa, 96

gracilis, 96

inflata, 96

kellersii, 96

ligustica, 96

linum, 96

media, 95

spiralis, 97

torta, 97

tortuosa, 96

chaetomorphoides, Enteromorpha, 91

Chamaedoris, 100

orientalis, 100

Champia, 53, 123

bifida, 53

caespitosa, 53

compressa, 53

disticha, 53

japonica, 53

parvula, 53

salicornoides, 53

spathulata, 53

vieillardii, 53

Chantransia hallandica, 19

charoides, Hypnea, 49

Chauvinia

clavifera, 107

macrophysa, 107

Cheilosporum, 34

cultratum, 34

jungermannioides, 34

sagittatum, 34

spectabile, 5

chemnitzia, Caulerpa racemosa var., 108

chemnitzia, Fucus, 108

chemnitzia var. occidentalis, Caulerpa, 108

cheyneana, Liagora, 21

cheyneana, Liagora farinosa var., 21

chilensis, Callophyllis, 32

Chlorocladus

australasicus, 120, 126

philippinensis, 120

Chlorodesmis, 118

caespitosa, 118

comosa, 118

fastigiata, 118

formosana, 118

hildebrandtii, 118

major, 118

torresiensis, 118

Chnoospora, 79

implexa, 79

minima, 79

pannosa, 80

Chondria, 62, 123

armata, 62

crassicaulis, 62

curvilineata, 62

dasyphylla, 63

glandulifera, 66

intricata, 66

obtusa var. paniculata, 67

obtusa var. patentiramea, 67

papillosa, 67

parvula, 53

patentiramea, 68

polyrhiza, 63

repens, 63

sedifolia, 63

seticulosa, 63

sibogae, 63

uvaria, 51

chondrioides, Laurencia, 65

Chondroclonium cornutum, 49

Chondrococcus, 129

hornemannii, 39, 129

japonicus, 39

lambertii, 129

spinulosus, 129

Chondrus, 123

edulis, 47

Chromastrum, 127, 128

Chroodactylon, 17

ornatum, 17

Chrysymenia, 123

kuckuckii, 51

pyriformis, 51

skottsbergii, 51

uvaria, 51

Chthonoblastus kurzii, 12

Chylocladia baileyana, 52

ciliolata, Dictyota, 75

cinctum var. gracilentum, Sargassum, 82

cinctum var. mixtum, Sargassum, 82

Cladocephalus, 125

Cladophora, 97, 125

aegagropila, 5

albida, 97

anastomosans, 101

aokii, 97

catenata, 97

coacta, 100

conferta, 97

crispula, 97

crucigera, 97

crystallina, 97

cymopoliae, 97

dalmatica, 97

densa, 98

fascicularis, 99

filipendula, 97

fuliginosa, 97

heteronema, 98

inserta, 98

japonica, 98

laetevirens, 98

liebetruthii, 98

mauritiana, 99

pellucida, 98

philippinensis, 98

prolifera, 98

quisumbingii, 98

rudolphiana, 99

rugulosa, 98

rupestris, 98

sakaii, 98

sericea, 99

sibogae, 99

trichotoma, 98

uncinata, 97

vagabunda, 99

wrightiana, 99

zollingeri, 101

cladophorae, Ptilothamnion, 57

cladophorae, Spermothamnion, 57

Cladophoropsis, 100, 125

dichotoma, 100

fasciculata, 100

gracillima, 101

membranacea, 101

neocaledonica, 101

philippinensis, 101

sundanensis, 101

vaucheriaeformis, 101

zollingeri, 101

Cladosiphon frappieri, 20

clathrata, Conferva, 91

clathrata, Enteromorpha, 91

clathrata var. crinita, Enteromorpha, 93

clathratum, Encoelium, 80

clathratum, Microdictyon, 95

clathratus, Fucus, 80

clathratus, Hydroclathrus, 80

Claudea, 58

batanensis, 58

multifida, 58

clavata, Acetabularia, 121

clavata, Chaetomorpha, 5

clavata, Giffordia, 130

clavata, Hincksia, 130

clavata, Laurencia, 65

clavata var. torta, Chaetomorpha, 97

clavatum, Gelidium, 25

clavifer, Fucus, 106

clavifera, Caulerpa, 106

clavifera, Caulerpa racemosa var., 107

clavifera, Chauvinia, 107

clavifera f. microphysa, Caulerpa racemosa var., 106

clavifera var. gracilis, Caulerpa, 107

clavifera var. turbinata, Caulerpa, 108

clavigera, Galaxaura, 23

clavulatum, Centroceras, 54

clavulatum, Ceramium, 54

coacta, Boodlea, 100

coacta, Cladophora, 100

coacta, Rhodymenia, 52

coccinea, Portieria, 129

Coccochloris, 123

Codium, 111, 125

adhaerens, 111, 112

arabicum, 111

bartlettii, 112

contractum, 112

coronatum, 111

cylindricum, 112

decorticatum, 112

dichotomum, 113

difforme, 112

divaricatum, 112

edule, 112

effusum, 112

elongatum, 112

fragile, 112

geppii, 112, 113

intricatum, 113

muelleri, 113

ovale, 113

papillatum, 113

platyclados, 113

pugniforme, 113

repens, 112

setchellii, 112

tenue, 113

tomentosum, 113

Coelothrix, 52

irregularis, 52

Colaconema, 127, 128

Colpomenia, 80, 125

sinuosa, 80

columellaris, Ectocarpus, 73

columellaris, Feldmannia, 73

columellaris, Laurencia, 65

commersonii, Padina, 77

comoides, Calothrix, 8

comoides, Hydrocoleum, 8

comosa, Chlorodesmis, 118

compacta, Caulerpa lentillifera var., 106

compactum, Sargassum, 85

compactum, Sargassum ilicifolium var., 85

complanata, Endocladia, 27

complanata, Gloiopeltis, 27

composita, Boodlea, 100

composita, Conferva, 100

composita, Laurencia, 65

compressa, Champia, 53

compressa, Enteromorpha, 91

compressa, Gracilaria, 41

compressa, Ulva, 91, 93

compressus, Sphaerococcus, 41

conchicola, Peyssonnelia, 27

concinna, Ahnfeltia, 50

concinna, Laurencia, 64

condensata, Roschera, 71

condensata, Tolypiocladia, 71

condensata, Turbinaria, 88

conferta, Cladophora, 97

conferta, Entophysalis, 7

conferta, Grateloupia, 30

conferta, Grateloupia filicina var., 30

conferta, Palmella, 7

Conferva

aerea, 95

aestuarii, 9

albida, 97

antennina, 95

atropurpurea, 17

catenata, 97

clathrata, 91

composita, 100

confervicola, 13

crassa, 96

crinita, 93

crystallina, 97

daviesii, 128

fascicularis, 99

fastigiata, 54

flexuosa, 92

fuscopurpurea, 17

gracilis, 96

granulosa, 130

heteronema, 98

laetevirens, 98

ligustica, 96

linum, 96

majuscula, 8, 9

media, 95

membranacea, 101

ornata, 17

paradoxa, 92

pellucida, 98

prolifera, 98

riparia, 99

rudolphiana, 99

rupestris, 98

scopulorum, 14

sericea, 99

seticulosa, 63

simplex, 63

tortuosa, 96

trichotoma, 98

utricularis, 103

vagabunda, 99

confervicola, Calothrix, 13

confervicola, Conferva, 13

confervoides, Ceramium, 44

confervoides, Fucus, 44

confervoides, Gracilaria, 44

confervoides, Lyngbya, 9

confervoides, Sphaerococcus, 44

confervoides, Valonia, 102

confusum, Sargassum, 82

conoides, Turbinaria, 88

conoides, Turbinaria vulgaris var., 88

conoides f. laticuspidata, Turbinaria, 89

conoides f. retroflexa, Turbinaria, 89

conoides var. evesiculosa, Turbinaria, 90

constipata, Galaxaura, 23

contarenii, Calothrix, 13

contarenii, Rivularia, 13

contigua, Galaxaura, 22

contractum, Codium, 112

copiosa, Halimeda, 114

Corallina, 34, 123

anceps, 32

cubensis, 34

cylindrica, 23

decussato-dichotoma, 35

ephedraea, 32

flabellum, 119

fragilissima, 33

frondescens, 34

incrassata, 115

marginata, 23

oblongata, 23

obtusata, 24

opuntia, 115

pinnatifolia var. digitata, 34

rubens, 35

rugosa, 24

sagittata, 34

tenella, 35

tribulus, 33

tridens, 115

tuna, 116

ungulata, 35

ungulata f. brevior, 36

corallinae, Leibleinia, 11

corallinae, Oscillatoria, 11

corallopsis, Laurencia, 65

Corallopsis

minor, 43

opuntia, 41

salicornia, 43

salicornia var. minor, 43

corallopsis, Sphaerococcus, 65

cordata, Halimeda, 116

cordata, Halimeda opuntia f., 116

Cordylecladia irregularis, 52

Coriophyllum setchellii, 5

cornea, Carpopeltis, 31

cornea, Grateloupia, 31

cornea, Prionitis, 31

corneus var. pulchellus, Fucus, 25

cornuta, Hypnea, 49

cornuta var. stellulifera, Hypnea, 49

cornutum, Chondroclonium, 49

coronatum, Codium, 111

coronopifolia, Gracilaria, 41

coronopifolia, Gracilaria lichenoides f., 41

corticulans, Bryopsis, 103

corymbosa, Gardnerula, 14

corymbosus, Microcoleus, 14

corynephora, Caulerpa, 107

corynephora, Caulerpa racemosa var., 107

costata, Delesseria plocamium var., 44

costatum, Plocamium, 44

Cottoniella, 58

filamentosa, 58

cottonii, Eucheuma, 46

cottonii var. erecta, Eucheuma, 46

coulteri, Gelidium, 25

crassa, Chaetomorpha, 96

crassa, Conferva, 96

crassa, Gracilaria, 41

crassa, Herposiphonia, 63

crassa, Padina, 78

crassicaulis, Chondria, 62

crassifolia, Caulerpa, 106

crassifolia, Caulerpa taxifolia var., 106

crassipellitum var. robustum, Rhizoclonium, 99

crassum, Eucheuma, 46

crenulata, Acetabularia, 121

crenulata, Cryptonemia, 29

crenulata, Dictyota, 75

crenulata, Phyllophora, 29

crinale, Gelidium, 25

crinale var. perpusillum, Gelidium, 25

crinalis, Fucus, 25

crinita, Conferva, 93

crinita, Enteromorpha, 93

crinita, Enteromorpha clathrata var., 93

crispa, Lyngbya, 9

crispata, Carpopeltis, 29

crispata, Porphyra, 18

crispifolium, Sargassum, 82

crispella, Heterosiphonia, 60

crispella var. laxa, Heterosiphonia, 60

crispula, Cladophora, 97

cristaefolium, Sargassum, 83

cristata, Lophosiphonia, 69

cristatum, Sargassum, 82

crosbyanum, Phormidium, 11

Crouania, 56

attenuata, 56

minutissima, 56

cruciatum, Ceramium, 54

crucigera, Cladophora, 97

Cruoriella

dura, 28

foveolata, 28

indica, 28

lemoinei, 5

mariti, 28

crustacea, Calothrix, 13, 14, 15

crustiforme, Eucheuma, 46

cryptacanthum, Centroceras, 54

Cryptonemia, 29, 123

capitellata, 29

crenulata, 29

luxurians, 29

rigida, 29

schmitziana, 29

crystallina, Cladophora, 97

crystallina, Conferva, 97

cubense, Haliptilon, 34

cubensis, Corallina, 34

cubensis, Jania, 34

cuculligera, Galaxaura, 24

cultrata, Amphiroa, 34

cultratum, Cheilosporum, 34

cumingii, Amphiroa, 32

cuneata, Halimeda, 114

cuneiformis, Fucus, 81, 131

cuneiformis, Hormophysa, 81, 131

cupressoides, Caulerpa, 104

cupressoides, Fucus, 104

cupressoides var. disticha, Caulerpa, 105

cupressoides var. ericifolia, Caulerpa, 105

cupressoides var. lycopodium, Caulerpa, 105

cupressoides var. lycopodium f. amicorum, Caulerpa, 105

cupressoides var. lycopodium f. disticha, Caulerpa, 105

cupressoides var. lycopodium f. elegans, Caulerpa, 105

cupressoideum, Eucheuma, 45

curtissiae, Microdictyon, 95

curviceps, Oscillatoria, 11

curvilineata, Chondria, 62

Cutleria, 74, 125

cylindrica, 74

cyathifera, Amphiroa, 33

cyathifera, Amphiroa fragilissima f., 33

cylindracea, Halimeda, 114

cylindrica, Corallina, 23

cylindrica, Cutleria, 74

cylindrica, Galaxaura, 23

cylindrica, Gracilaria, 41

cylindricum, Codium, 112

Cymopolia, 120

vanbosseae, 120

cymopoliae, Cladophora, 97

cystocarpum, Sargassum, 83

Cystodictyon, 94

leclancheri, 94

Cystophyllum, 81

hakodatense, 81

sisymbrioides, 81

trinode, 81

turneri, 81

cystophyllum, Sargassum, 83

cystophyllus, Carpacanthus, 83

Cystoseira, 81, 125, 131

articulata, 81, 131

hakodatensis, 81

prolifera, 81

triquetra, 81, 131

dactyloides, Gracilaria, 52

dactyloides, Hymenocladia, 52

dalmatica, Cladophora, 97

damaecornis, Gracilaria, 41

Dasya, 59

adhaerens, 59

baillouviana, 59

delilei, 21, 22

lallemandii, 69

mollis, 59

muelleri, 60

ocellata, 59

penicillata, 130

plana, 129

punicea, 59

sessilis, 59

spinella, 129

wurdemannii, 60

Dasycladus, 120, 126

australasicus, 120

vermicularis, 120

Dasyopsis, 60, 123, 129

anastomosans, 129

antillarum, 60, 129

aperta, 129

geppii, 130

palmatifida, 130

pilosa, 60, 130

pulchella, 130

stanleyi, 130

tenella, 130

Dasyphila, 56

plumarioides, 56

dasyphylla, Chondria, 63

dasyphyllus, Fucus, 63

daviesii, Acrochaetium, 128

daviesii var. secundatum, Callithamnion, 128

debilis, Fucus, 44

debilis, Gracilaria, 44

debilis, Polycavernosa, 44

decaisnei, Mastophora, 37

Decaisnella, 118

indica, 119

decorticatum, Codium, 112

decumbens, Laurencia, 65

decumbens, Rhodymenia, 52

decurrens, Turbinaria, 89

decussato-dichotoma, Corallina, 35

decussato-dichotoma, Jania, 35

Delesseria

adnata, 58

filicina, 30

leprieurii, 58

plocamium var. costata, 44

spathulata, 58

delicatula, Dictyopteris, 74

delicatula, Herposiphonia, 63

delicatula, Polysiphonia pacifica var., 70

delicatula, Struvea, 101

delilei, Asparagopsis, 21

delilei, Dasya, 21, 22

dendroidea, Herposiphonia, 64

dendroidea, Laurencia, 67

dendroidea, Laurencia obtusa var., 67

dendroidea var. minor, Herposiphonia, 64

dendroides, Ochlochaete, 91

dendroides, Phaeophila, 91

densa, Cladophora, 98

densa, Laurencia obtusa var., 67

densa, Pterocladia, 27

dentata, Acetabularia, 121

denticulata, Dictyota bartayresiana var., 75

denticulata, Gracilaria, 44

denticulata, Porphyra, 18

denticulatum, Eucheuma, 46

denticulatus, Fucus, 46, 86

denticulatus, Sphaerococcus, 44

denudata, Turbinaria, 89

depauperatum, Eucheuma muricatum f., 46

Derbesia, 104

attenuata, 104

marina, 104

ryukyuensis, 104

tenuissima, 104

Dermonema, 20

frappieri, 20

Desmarestia, 128, 129

Desmia, 128, 129

ambigua, 129

cervicornis, 129

dichotoma, 129

harveyi, 129

hornemannii, 39, 128, 129

japonica, 39, 129

kilneri, 129

tripinnata, 129

diaphanum, Ceramium, 56

diaphanum var. tenuissimum, Ceramium, 56

Dichonema erectum, 117

Dichothrix, 14

gypsophila, 14

dichotoma, Cladophoropsis, 100

dichotoma, Desmia, 129

dichotoma, Dictyota, 76

dichotoma, Grateloupia, 30

dichotoma, Portieria, 129

dichotoma, Spongocladia, 100

dichotoma, Ulva, 76

Dichotomasiphon pusillus, 117

dichotomum, Codium, 113

dichotomum, Eucheuma, 5

dichotomum, Spongodendron, 100

dickiei, Lithothamnion, 37

Dicranema, 123

Dictyerpa jamaicensis, 78

Dictyopteris, 74, 125

camiguinensis, 74

delicatula, 74

divaricata, 74

jamaicensis, 74

membranacea, 74

polypodioides, 74

repens, 75

undulata, 75

Dictyosphaeria, 101, 102, 126

cavernosa, 101

cavernosa var. bullata, 102

favulosa, 102

intermedia, 102

ocellata, 102

setchellii, 102

vanbosseae, 102

versluysii, 102

Dictyota, 75, 125

atomaria, 76

bartayresiana, 75

bartayresiana var. denticulata, 75

bartayresii, 75

bidentata, 75

cervicornis, 75

ceylanica, 75

ciliolata, 75

crenulata, 75

dichotoma, 76

divaricata, 76

friabilis, 76

indica, 76

lata, 76

linearis, 76

major, 76

marginata, 77

mertensii, 76

patens, 77

variegata, 77

volubilis, 76

diesingiana, Zonaria, 79

difforme, Codium, 112

difformis, Leathesia, 79

difformis, Tremella, 79

Digenea, 63

simplex, 63

digitata, Corallina pinnatifolia var., 34

dilatata, Amphiroa, 32

dilatata, Halymenia, 30

dilatatus, Fucus, 51

dilatatus, Gymnogongrus, 51

Dilophus, 77

marginatus, 77

okamurae, 77

dimorpha, Galaxaura, 23

Diploderma variegatum, 18

Diplotrichia polyotis, 15

discoidea f. intermedia, Halimeda, 114

discoidea f. subdigitata, Halimeda, 114

discoidea, Halimeda, 114

distenta, Galaxaura, 28

distenta, Peyssonnelia, 28

disticha, Caulerpa cupressoides var., 105

disticha, Caulerpa cupressoides var. lycopodium f., 105

disticha, Champia, 53

disticha, Gracilaria, 42

distichophylla, Laurencia, 66

distichus, Sphaerococcus, 42

distromatica, Padina, 78

divaricata, Carpopeltis, 29

divaricata, Dictyopteris, 74

divaricata, Dictyota, 76

divaricata, Grateloupia, 30

divaricata, Haliseris, 74

divaricata, Hypnea, 49

divaricata, Liagora, 20

divaricatum, Codium, 112

divaricatum, Gelidium, 25

divaricatus, Fucus, 49

divaricatus, Gymnogongrus, 51

divaricatus, Sphaerococcus, 49

doryphora, Grateloupia, 30

doryphora, Halymenia, 30

droserifolium, Sargassum, 83

droserifolium var. spathulatum, Sargassum, 83

duchassaingiana, Giffordia, 73

duchassaingiana, Hincksia, 73

duchassaingianus, Ectocarpus, 73

duperreyi, Haloplegma, 57

duplicatum, Sargassum, 83

duplicatum, Sargassum ilicifolium var., 83

duplicatum var. rotundatum, Sargassum, 83

dura, Cruoriella, 28

dura, Polystrata, 28

dura, Rhabdonia, 48

dura, Solieria, 48

durum, Archaeolithothamnion erythraeum f., 38

durum, Sporolithon ptychoides f., 38

durvillaei, Halymenia, 30

durvillaei var. ceylanica, Halymenia, 31

durvillaei var. formosa, Halymenia, 31

echinocarpum var. vitiense, Sargassum, 86

Ectocarpus, 125, 130

breviarticulatus, 73

columellaris, 73

duchassaingianus, 73

fenestratus, 130

formosanus, 73

fuscatus, 130

ghardaqaensis, 130

hincksiae, 130

indicus, 73

irregularis, 73

mitchelliae, 73, 130

ovatus, 130

ovatus var. intermedius, 130

rallsiae, 73, 130

sandrianus, 130

saundersii, 130

secundus, 130

sordidus, 130

thyrsoideus, 130

edule, Codium, 112

edule, Eucheuma, 47

edulis, Chondrus, 47

edulis, Fucus, 42, 44

edulis, Gracilaria, 42

effusum, Codium, 112

elegans, Bangia, 17

elegans, Callithamnion, 57

elegans, Caulerpa cupressoides var. lycopodium f., 105

elegans, Caulerpa plumaris var., 105

elegans, Goniotrichum, 17

elegans, Gymnothamnion, 57

elegans, Microcladia, 57

elongata, Caulerpa, 105

elongata, Galaxaura, 23, 24

elongatum, Codium, 112

elongatum, Sargassum, 83

Enantiocladia, 63

okamurae, 63

Encoelium

clathratum, 80

orientale, 80

Endocladia complanata, 27

Endosiphonia, 63

spinuligera, 63

enerve, Sargassum, 84

Enteromorpha, 91, 126

aragoensis, 91

chaetomorphoides, 91

clathrata, 91

clathrata var. crinita, 93

compressa, 91

crinita, 93

erecta, 92

flexuosa, 92

flexuosa f. submarina, 92

flexuosa subsp. paradoxa, 92

intermedia, 92

intestinalis, 92

intestinalis f. tubulosa, 92

intestinalis var. tubulosa, 92

kylinii, 92

lingulata, 93

plumosa, 92

prolifera, 93

ramulosa, 93

ramulosa var. spinosa, 93

salina, 93

spinescens, 93

tubulosa, 92

enteromorphoides, Hormothamnium, 8

Entophysalis, 7, 123

conferta, 7

ephedraea, Amphiroa, 32

ephedraea, Corallina, 32

epiphytica, Calothrix, 13

equisetoides, Ceramium, 54

erecta, Amphiroa foliacea f., 33

erecta, Avrainvillea, 117

erecta, Enteromorpha, 92

erecta, Eucheuma cottonii var., 46

erectum, Dichonema, 117

erectus, Scytosiphon, 92

ericifolia, Caulerpa cupressoides var., 105

ericifolius, Fucus, 105

Ernodesmis, 102

verticillata, 102

erubescens, Lithothamnion, 37

erubescens, Mesophyllum, 37

erythraea, Oscillatoria, 12

erythraeum, Lithothamnion, 38

erythraeum, Sporolithon, 38

erythraeum, Trichodesmium, 12

erythraeum f. durum, Archaeolithothamnion, 38

erythraeum f. molle, Archaeolithothamnion, 38

Erythrocladia, 17

irregularis, 17

pinnata, 17

subintegra, 17

Erythrocolon, 52

podagricum, 52

Erythrotrichia, 17, 123

bangioides, 17

biseriata, 17

parietalis, 17

eseptata, Anadyomene, 94

esperi, Hypnea, 49

esperi, Hypnea musciformis var., 49

esperi, Sargassum, 83

Eucheuma, 45, 123

alvarezii, 45

alvarezii var. ajakii-assii, 45

alvarezii var. tambalangii, 45

arnoldii, 45

arnoldii var. alcyonida, 45

cottonii, 46

cottonii var. erecta, 46

crassum, 46

crustiforme, 46

cupressoideum, 45

denticulatum, 46

dichotomum, 5

edule, 47

gelatinum, 47

horridum, 47

isiforme, 47

leeuwenii, 47

muricatum, 46

muricatum f. depauperatum, 46

muricatum f. incrassatum, 47

okamurae, 47

procrusteanum, 47

serra, 47

spinosum, 46

striatum, 47

eucheumoides, Gracilaria, 42

Euhymenia limensis, 40

Eupogodon, 60, 123, 129

anastomosans, 129

antillarum, 60, 129

apertus, 129

apiculatus, 129

cervicornis, 130

geppii, 130

palmatifidus, 130

penicillatus, 130

pilosus, 60, 130

pinnatifolius, 130

planus, 130

pulchellus, 130

spinellus, 130

stanleyi, 130

tenellus, 130

euryphyllum, Sargassum polycystum var., 86

euryphyllum, Sargassum myriocystum var., 86

evae, Peyssonnelia, 28

evesiculosa, Turbinaria conoides var., 90

evesiculosa, Turbinaria ornata f., 90

exigua, Acetabularia, 121

Exophyllum, 63

wentii, 63

expeditionis, Tydemania, 118

falcata, Galaxaura, 22

farinosa, Fosliella, 34

farinosa, Liagora, 20

farinosa, Melobesia, 34

farinosa var. cheyneana, Liagora, 21

farinosum, Ganonema, 21

farlowii, Caulerpa sertularioides f., 110

farlowii, Caulerpa plumaris f., 110

fasciata, Ulva, 93

fascicularis, Cladophora, 99

fascicularis, Conferva, 99

fasciculata, Cladophoropsis, 100

fasciculata, Galaxaura, 23

fasciculatus, Siphonocladus, 100

fastigiata, Caulerpa, 105

fastigiata, Chlorodesmis, 118

fastigiata, Conferva, 54

fastigiata, Valonia, 102

fastigiata, Vaucheria, 118

fastigiatum, Ceramium, 54

Fauchea, 52, 123

leptophylla, 52

favulosa, Dictyosphaeria, 102

favulosa, Valonia, 102

Feldmannia, 73, 125

columellaris, 73

formosana, 73

indica, 73

irregularis, 73

fenestrata, Hincksia, 130

fenestratus, Ectocarpus, 130

fergusonii, Caulerpa, 105

ferruginea, Lyngbya, 9

ferulacea, Polysiphonia, 69

filamentosa, Cottoniella, 58

filamentosa, Galaxaura, 23

filamentosa, Sarcomenia, 58

filamentosa, Spyridia, 49, 57

filamentosa, Turbinaria, 89

filamentosum, Ceramium, 49

filamentosus, Fucus, 57

filicina, Delesseria, 30

filicina, Grateloupia, 30

filicina var. conferta, Grateloupia, 30

filicinum, Sargassum, 83

filicinus, Fucus, 30

filicoides, Caulerpa, 105

filifolium, Sargassum, 83

filifolium var. aciculare, Sargassum, 84

filiforme, Sargassum, 84

filipendula, Cladophora, 97

fimbriatum, Ceramium, 55

flabellata, Anadyomene, 95

flabellata, Carpopeltis, 29

flabellata, Grateloupia, 29

flabellata, Zonaria, 79

flabellatus, Homoeostrichus, 79

flabelliforme, Stypopodium, 79

flabelliformis, Gymnogongrus, 51

flabelliformis, Martensia, 59

flabellulata, Polysiphonia, 69

flabellum, Corallina, 119

flabellum, Udotea, 119

flaccidum, Ceramium, 54, 55

flaccidum, Hormoceras, 54

flagellata, Caulerpa plumaris f., 110

flagellata, Caulerpa sertularioides f., 110

flexuosa, Conferva, 92

flexuosa, Enteromorpha, 92

flexuosa, Ulva, 92

flexuosa f. submarina, Enteromorpha, 92

flexuosa subsp. paradoxa, Enteromorpha, 92

floresia, Halymenia, 31

floresius, Fucus, 31

fluitans, Sargassum, 84

fluitans, Sargassum hystrix var., 84

foliacea, Amphiroa, 33

foliacea, Melobesia, 37

foliacea f. erecta, Amphiroa, 33

foliifer, Fucus, 42

foliifera, Gracilaria, 42

foliifera f. aeruginosa, Gracilaria, 42

forbesii, Boergesenia, 100

forbesii, Valonia, 100

forcipata, Polysiphonia, 69

forfex, Polysiphonia, 69

formosa, Halymenia, 31

formosa, Halymenia durvillaei var., 31

formosana, Carpopeltis, 29

formosana, Chlorodesmis, 118

formosana, Feldmannia, 73

formosanus, Ectocarpus, 73

forsteri, Fucus, 66

forsteri, Laurencia, 66

Fosliella, 34

farinosa, 34

foveolata, Cruoriella, 28

foveolata, Peyssonnelia, 28

fragile, Acanthocodium, 112

fragile, Codium, 112

fragilis, Actinotrichia, 22

fragilis, Fucus, 22

fragilis, Halimeda, 114

fragilis, Polysiphonia, 69

fragilissima, Amphiroa, 33

fragilissima, Corallina, 33

fragilissima f. cyathifera, Amphiroa, 33

frappieri, Cladosiphon, 20

frappieri, Dermonema, 20

fraseri, Padina, 78

fraseri, Zonaria, 78

fraxinifolia, Neurymenia, 69

fraxinifolius, Fucus, 69

freycinetii, Caulerpa, 109

freycinetii var. boryana f. occidentalis, Caulerpa, 109

freycinetii var. pectinata, Caulerpa, 109

freycinetii var. typica f. lata, Caulerpa, 109

freycinetii var. typica f. spiralis, Caulerpa, 109

friabilis, Dictyota, 76

friesii, Schizothrix, 12

frondescens, Corallina, 34

frutescens, Goniolithon, 38

frutescens, Neogoniolithon, 38

fruticulosa, Galaxaura, 24

fruticulosum, Lithothamnion, 37

Fucus

abrotanifolius, 26

abscissus, 129

acanthophorus, 61

acerosus, 25

aeruginosus, 42

amansii, 25

articulatus, 81, 131

atomarius, 76

bacciferus, 86

baccularia, 82

baillouviana, 59

botryocarpus, 58

botryoides, 51

bursa-pastoris, 41

capensis, 26

capillaceus, 27

cartilagineus, 26

chemnitzia, 108

clathratus, 80

clavifer, 106

confervoides, 44

corneus var. pulchellus, 25

crinalis, 25

cuneiformis, 81, 131

cupressoides, 104

dasyphyllus, 63

debilis, 44

denticulatus, 46, 86

dilatatus, 51

divaricatus, 49

edulis, 42, 44

ericifolius, 105

filamentosus, 57

filicinus, 30

floresius, 31

foliifer, 42

forsteri, 66

fragilis, 22

fraxinifolius, 69

fulvellus, 84

fungiformis, 73

gelatinus, 47

glandulosus, 57

gulaman, 44

hamulosus, 50

hemiphyllus, 84

hornemanni, 128

ilicifolius, 85

lacinulatus, 42

lambertii, 129

lamourouxii, 107

latifolius, 85

lichenoides, 42

membranaceus, 74

miniatus, 72

minimus, 79

muricatus, 46

musciformis, 49

muscoides, 60

myagroides, 81

natans, 86

nootkanus, 50

obtusus, 66

opuntia, 48

ornatus, 128

palmatus, 19

papillosus, 67

parvifolius, 87

pavonicus, 78

pavonius, 78

pilulifer, 87

plumaris, 110

poitei, 68

polypodioides, 74, 75

prolifer, 106, 115, 116

pusillus, 25

racemosus, 106

rigidus, 25

salicifolius, 88

selago, 108

serratifolius, 87

serrulatus, 108

sertularioides, 109, 110

sisymbrioides, 81

spiciferus, 60

spiniformis, 25

spinosus, 46

squamarius, 28

taxifolius, 111

taxiformis, 21, 22

tenax, 27

tenellus, 62

triangularis, 131

trifarius, 130

triqueter, 81, 131

turbinatus, 90

turbinatus var. ornatus, 89

uvifer, 107

valentiae, 50

verrucosus, 44

versicolor, 26

fuliginosa, Cladophora, 97

fulvellum, Sargassum, 84

fulvellus, Fucus, 84

fungiformis, Fucus, 73

fungiformis, Ralfsia, 73

furcatum, Sargassum, 84

furcellata, Ahnfeltia, 51

furcigera, Sphacelaria, 74

fuscata, Hincksia, 130

fuscatus, Ectocarpus, 130

fuscopurpurea, Bangia, 18

fuscopurpurea, Conferva, 17

Galaxaura, 22, 123

acuminata, 22

apiculata, 22

arborea, 22

clavigera, 23

constipata, 23

contigua, 22

cuculligera, 24

cylindrica, 23

dimorpha, 23

distenta, 28

elongata, 23, 24

falcata, 22

fasciculata, 23

filamentosa, 23

fruticulosa, 24

glabriuscula, 24

kjellmanii, 23

lapidescens, 23

marginata, 23

oblongata, 23

obtusata, 24

pacifica, 24

rigida, 22

robusta, 24

rudis, 23

rugosa, 24

sibogae, 5

squalida, 24

striata, 24

subfruticulosa, 24

subverticillata, 24

tenera, 23

umbellata, 24

veprecula, 23

Ganonema, 21

farinosum, 21

Gardnerula, 14

corymbosa, 14

Gastridium ovale, 104

gaudichaudii, Carpacanthus, 84

gaudichaudii, Sargassum, 84

gelatinae, Eucheuma, 46

gelatinosa, Gigartina, 47

gelatinum, Eucheuma, 47

gelatinus, Fucus, 47

gelatinus, Sphaerococcus, 44, 47

Gelidiella, 25, 124

acerosa, 25

adnata, 25

taylorii, 25

Gelidiopsis, 40, 124

intricata, 40

repens, 40

rigida, 25

variabilis, 40

Gelidium, 25, 124

amansii, 25

anthoninii, 5

caloglossoides, 27

capense, 26

cartilagineum, 26

clavatum, 25

coulteri, 25

crinale, 25

crinale var. perpusillum, 25

divaricatum, 25

isabelae, 25

kintaroi, 25

pulchellum, 25

pusillum, 25

pusillum var. pacificum, 27

repens, 40

rigens, 27

rigidum, 25

spiniforme, 25

variabile, 40

geppii, Codium, 112, 113

geppii, Dasyopsis, 130

geppii, Eupogodon, 130

geppii, Udotea, 119

Geppina, 118

ghardaqaensis, Ectocarpus, 130

ghardaqaensis, Hincksia, 130

Giffordia, 73

andamanensis, 130

bhimlipatnamensis, 130

duchassaingiana, 73

clavata, 130

hincksiae var. californica, 130

mitchelliae var. neilii, 130

onslowensis, 130

prolifera, 130

rallsiae, 73

terminalis, 130

gigantea, Ulva, 94

giganteifolium, Sargassum, 84

Gigartina, 51

affinis, 28

gelatinosa, 47

lichenoides, 42

prolifera, 29

tenella, 51

gigas, Gracilaria, 42

gigas, Halimeda, 114

glabriuscula, Galaxaura, 24

glandulifera, Chondria, 66

glandulifera, Laurencia, 66

glandulosa, Microcladia, 57

glandulosus, Fucus, 57

glaucescens, Udotea, 119

globuliferum, Pleonosporium, 57

Gloeotrichia, 123

Gloiocladia, 52

ramellifera, 52

Gloiopeltis, 27

complanata, 27

tenax, 27

glomerata, Amansia, 61

glomerulata, Hutchinsia, 71

glomerulata, Tolypiocladia, 71

glutinosa, Lyngbya, 8

glutinosum, Hydrocoleum, 8

Goniolithon

frutescens, 38

megalocystum, 38

reinboldii, 34

Goniotrichum, 17

alsidii, 17

elegans, 17

Gordoniella, 56

yonakuniensis, 56

gorgoniae, Polysiphonia, 69

Gracilaria, 40, 124

andersonii, 43

arcuata, 40

arcuata var. snackeyi, 41

blodgettii, 41

bursa-pastoris, 41

canaliculata, 41

compressa, 41

confervoides, 44

coronopifolia, 41

crassa, 41

cylindrica, 41

dactyloides, 52

damaecornis, 41

debilis, 44

denticulata, 44

disticha, 42

edulis, 42

eucheumoides, 42

foliifera, 42

foliifera f. aeruginosa, 42

gigas, 42

incurvata, 43

lacinulata, 42

lichenoides, 42

lichenoides f. coronopifolia, 41

minor, 43

multipartita var. aeruginosa, 42

papenfussii, 43

purpurascens, 43

salicornia, 43

spinigera, 43

spinulosa, 43

textorii, 43

turgida, 43

venezuelensis, 44

verrucosa, 44, 123

vieillardii, 44

gracile, Acrochaetium, 18

gracile, Sargassum, 84

gracilentum, Sargassum cinctum var., 82

gracilis, Caulerpa clavifera var., 107

gracilis, Caulerpa racemosa var., 107

gracilis, Chaetomorpha, 96

gracilis, Conferva, 96

gracilis, Halimeda, 114

gracilis, Leveillea, 68

gracilis, Rhodopeltis, 39

gracillima, Cladophoropsis, 101

gracillimum, Ceramium, 55

gracillimum, Hormoceras, 55

gracillimum var. byssoideum, Ceramium, 54

grande, Rhizoclonium, 99

granuliferum, Sargassum, 84

granulosa, Conferva, 130

granulosa, Hincksia, 130

Grateloupia, 30, 124

acuminata, 30

californica, 30

conferta, 30

cornea, 31

dichotoma, 30

divaricata, 30

doryphora, 30

filicina, 30

filicina var. conferta, 30

fiabellata, 29

ramosissima, 30

grevilleana, Laurencia, 65

Griffithsia, 56, 124

argus, 57

ovalis, 56

penicillata, 57

rhizophora, 56

tenuis, 53

gulaman, Fucus, 44

Gymnogongrus, 51, 124

dilatatus, 51

divaricatus, 51

flabelliformis, 51

ligulatus var. angustus, 28

pygmaeus, 51

Gymnosorus variegatus, 77

gymnospora, Padina, 77, 78

gymnospora, Zonaria, 77, 78

Gymnothamnion, 57

elegans, 57

gypsophila, Dichothrix, 14

gypsophilus, Schizosiphon, 14

hakodatense, Cystophyllum, 81

hakodatensis, Cystoseira, 81

hakodatensis, Lomentaria, 52

Halicoryne, 122

wrightii, 122

Halicystis

ovalis, 104

parvula, 104

Halimeda, 114, 126

batanensis, 116

bikinensis, 114

copiosa, 114

cordata, 116

cuneata, 114

cylindracea, 114

discoidea, 114

discoidea f. intermedia, 114

discoidea f. subdigitata, 114

fragilis, 114

gigas, 114

gracilis, 114

hederacea, 114

incrassata, 115

incrassata f. lamourouxii, 115

lacunalis, 115

macroloba, 115

macrophysa, 115

micronesica, 115

monile, 114

opuntia, 115

opuntia f. cordata, 116

opuntia f. hederacea, 114

opuntia f. intermedia, 116

opuntia f. renschii, 116

opuntia f. triloba, 116

opuntia var. hederacea, 114

orientalis, 115

platydisca, 116

renschii, 116

simulans, 116

stuposa, 116

taenicola, 116

tridens, 115

tridens f. lamourouxii, 115

triloba, 115

tuna, 116

tuna f. platydisca, 116

velasquezii, 116

velasquezii var. robusta, 116

Haliptilon, 34

cubense, 34

Haliseris

divaricata, 74

repens, 75

hallandica, Chantransia, 19

hallandicum, Acrochaetium, 19

Halochloa schizophylla, 87

Haloplegma, 57

duperreyi, 57

Halymenia, 30, 124

acuminata, 30

ceylanica, 31

dilatata, 30

doryphora, 30

durvillaei, 30

durvillaei var. ceylanica, 31

durvillaei var. formosa, 31

floresia, 31

formosa, 31

harveyana, 31

incrustans, 40

japonica, 31

maculata, 31

microcarpa, 31

hamulosa, Hypnea, 50

hamulosus, Fucus, 50

hancockii, Acrochaetium, 19

hancockii, Amphiroa, 33

hancockii, Rhodochorton, 19

harveyana, Halymenia, 31

harveyi, Desmia, 129

harveyi, Portieria, 129

hawaiiana, Liagora, 21

hawaiiensis, Polysiphonia, 70

hederacea, Halimeda, 114

hederacea, Halimeda opuntia f., 114

hederacea, Halimeda opuntia var., 114

helminthochortos, Alsidium, 95

Helminthocladia, 20

australis, 20

schrammi, 20

hemiphyllum, Sargassum, 84

hemiphyllus, Fucus, 84

hemiphyllus, Spongocarpus, 85

Herbacea, 128

Herposiphonia, 63, 124

crassa, 63

delicatula, 63

dendroidea, 64

dendroidea var. minor, 64

nuda, 64

obscura, 64

pacifica, 64

parca, 64

plumula, 64

prorepens, 5

secunda, 64

secunda f. tenella, 64

subdisticha, 64

tenella, 64

tenella f. secunda, 64

trichia, 64

Heteractis mesenterica, 14

heterocystum, Sargassum, 85

Heteroderma, 124

heteronema, Cladophora, 98

heteronema, Conferva, 98

Heterosiphonia, 60

crispella, 60

crispella var. laxa, 60

muelleri, 60

wurdemannii, 60

wurdemannii var. laxa, 60

hildebrandtii, Chlorodesmis, 118

Hincksia, 73

andamanensis, 130

bhimlipatnamensis, 130

breviarticulata, 73

clavata, 130

fenestrata, 130

fuscata, 130

ghardaqaensis, 130

granulosa, 130

hincksiae, 130

hincksiae var. californica, 130

intermedia, 130

mitchelliae, 73, 130

mitchelliae var. neilii, 130

onslowensis, 130

ovata, 130

prolifera, 130

rallsiae, 73, 130

ramulosa, 130

sandriana, 130

saundersii, 130

secunda, 130

sordida, 130

terminalis, 130

thyrsoidea, 130

hincksiae, Ectocarpus, 130

hincksiae, Hincksia, 130

hincksiae var. californica, Giffordia, 130

hincksiae var. californica, Hincksia, 130

Hippurina, 128

hippuroides, Hypnea, 50

hippuroides, Hypnea musciformis var., 50

hoffman-bangii, Scytonema, 15

hofman-bangii, Scytonema, 15

hofmanii, Scytonema, 15

hofmannii, Scytonema, 14

hombronianum var. manilense, Sargassum, 85

Homoeostrichus flabellatus, 79

hookeri, Caloglossa leprieurii var., 58

hookeri, Rhizoclonium, 99

Hormoceras

flaccidum, 54

gracillimum, 55

tenerrimum, 56

hormoides, Scinaia, 25

Hormophysa, 81, 125

cuneiformis, 81, 131

triquetra, 81, 130

Hormothamnium, 8, 123

enteromorphoides, 8

solutum, 8

hornemannii, Chondrococcus, 39, 129

hornemannii, Desmia, 39, 128, 129

hornemannii, Fucus, 128

hornemannii, Portieria, 39, 129

horridum, Eucheuma, 47

howei, Polysiphonia, 70

howei, Symploca, 13

Howella, 118

hummii, Caulerpa, 109

Hutchinsia

glomerulata, 71

periclados, 69

secunda, 64

tenella, 64

hyalacanthum, Centroceras, 54

hydnoides, Calothrix, 13

hydnoides, Symploca, 13

Hydroclathrus, 80, 125

cancellatus, 80

clathratus, 80

orientalis, 80

Hydrocoleum, 8

cantharidosmum, 8

comoides, 8

glutinosum, 8

lyngbyaceum, 9

Hydrolithon, 34

reinboldii, 34

Hymenocladia, 52

dactyloides, 52

Hypnea, 48, 124

boergesenii, 48

cenomyce, 48

cenomyce var. tenuis, 48

cervicornis, 49

charoides, 49

cornuta, 49

cornuta var. stellulifera, 49

divaricata, 49

esperi, 49

hamulosa, 50

hippuroides, 50

musciformis, 49

musciformis var. esperi, 49

musciformis var. hippuroides, 50

nidulans, 50

nigrescens, 50

pannosa, 50

saidana, 50

spinella, 50

valentiae, 50

Hypoglossum, 58

attenuatum, 58

serrulatum, 58

spathulatum, 58

hystrix, Sargassum, 85

hystrix var. fluitans, Sargassum, 84

hystrix var. spinulosus, Sargassum, 85

ilicifolium, Sargassum, 85

ilicifolium f. benkulense, Sargassum, 85

ilicifolium var. compactum, Sargassum, 85

ilicifolium var. duplicatum, Sargassum, 83

ilicifolium var. pseudospinulosum, Sargassum, 85

ilicifolius, Carpacanthus, 85

ilicifolius, Fucus, 85

imbricatum, Lithothamnion, 37

imbricatum, Mesophyllum, 37

implexa, Chnoospora, 79

implicata, Laurencia, 66

impudica, Catenella, 48

impudica, Lomentaria, 48

incrassata, Corallina, 115

incrassata, Halimeda, 115

incrassata f. lamourouxii, Halimeda, 115

incrassatum, Eucheuma muricatum f., 47

incrustans, Halymenia, 40

incrustans, Titanophora, 40

incurvata, Gracilaria, 43

indica, Bryopsis, 103

indica, Cruoriella, 28

indica, Decaisnella, 119

indica, Dictyota, 76

indica, Feldmannia, 73

indica, Litholepis, 36

indica, Lithoporella, 36

indica, Peyssonnelia, 28

indica, Udotea, 119

indica f. philippinensis, Litholepis, 36

indicum, Sargassum vulgare var., 88

indicum f. subtile, Lithothamnion, 37

indicus, Ectocarpus, 73

inermis, Bostrychia mixta f., 62

infixa, Lyngbya, 9

inflata, Chaetomorpha, 96

inserta, Cladophora, 98

intermedia, Dictyosphaeria, 102

intermedia, Enteromorpha, 92

intermedia, Halimeda discoidea f., 114

intermedia, Halimeda opuntia f., 116

intermedia, Hincksia, 130

intermedia, Laurencia, 66

intermedius, Ectocarpus ovatus var., 130

intestinalis, Enteromorpha, 92

intestinalis, Ulva, 92

intestinalis f. tubulosa, Enteromorpha, 92

intestinalis var. tubulosa, Enteromorpha, 92

intricata, Bostrychia, 61

intricata,. Chondria, 66

intricata, Gelidiopsis, 40

intricata, Laurencia, 66

intricata, Phyllophora, 52

intricata, Rhodymenia, 52

intricata, Rosenvingea, 80

intricatum, Codium, 113

intricatum, Scytonema, 61, 62

intricatus, Asperococcus, 80

intricatus, Sphaerococcus, 40

irregularis, Coelothrix, 52

irregularis, Cordylecladia, 52

irregularis, Ectocarpus, 73

irregularis, Erythrocladia, 17

irregularis, Feldmannia, 73

isabelae, Gelidium, 25

isiforme, Eucheuma, 47

isiformis, Sphaerococcus, 47

jamaicensis, Dictyerpa, 78

jamaicensis, Dictyopteris, 74

jamaicensis, Padina, 78

Jania, 34, 124

adhaerens, 34

capillacea, 35

cubensis, 34

decussato-dichotoma, 35

longiarthra, 35

mexicana, 35

micrarthodia, 35

pacifica, 35

pumila, 35

rubens, 35

tenella, 35

tenella var. zacae, 35

tenuissima, 35

ungulata, 35

ungulata f. brevior, 36

japonica, Callophyllis, 32

japonica, Champia, 53

japonica, Cladophora, 98

japonica, Desmia, 39, 129

japonica, Halymenia, 31

japonica, Laurencia, 66

japonica, Liagora, 21

japonica, Padina, 78

japonica, Portieria, 39, 129

japonica, Pugetia, 32

japonicum, Microdictyon, 95

japonicus, Chondrococcus, 39

javensis, Rhipidosiphon, 119

javensis, Udotea, 119

jungermannioides, Amansia, 68

jungermannioides, Cheilosporum, 34

jungermannioides, Leveillea, 68

jungermannioides, Polyzonia, 68

Kallymenia, 32, 124

californica, 32

callophylloides, 32

pacifica, 32

papulosa, 48

sessilis, 32

kelanensis, Bostrychia, 62

kellersii, Chaetomorpha, 96

kerneri, Rhizoclonium, 99

kilneri, Desmia, 129

kilneri, Portieria, 129

kintaroi, Gelidium, 25

kjellmanianum, Sargassum, 86

kjellmanii, Galaxaura, 23

kochianum, Rhizoclonium, 99

kuckuckii, Botryocladia, 51

kuckuckii, Chrysymenia, 51

kurzii, Chthonoblastus, 12

kurzii, Porphyrosiphon, 12

kurzii, Sirocoleum, 12

Kylinia rosulata, 19

kylinii, Enteromorpha, 92

lacerata, Avrainvillea, 117

lacerata var. robustior, Avrainvillea, 117

lacinulata, Gracilaria, 42

lacinulatus, Fucus, 42

lactuca, Ulva, 93

lactuca var. latissima, Ulva, 93

lactuca var. luxurians, Sphaerococcus, 29

lactuca var. rigida, Ulva, 94

lacunalis, Halimeda, 115

lacunosa, Rhodophyllis peltata var., 48

laetevirens, Caulerpa, 107

laetevirens, Caulerpa racemosa var., 107

laetevirens, Cladophora, 98

laetevirens, Conferva, 98

laeteviridis, Symploca, 13

lallemandii, Dasya, 69

lallemandii, Lophocladia, 69

lambertii, Chondrococcus, 129

lambertii, Fucus, 129

Laminaria saccharina, 94

lamourouxii, Caulerpa racemosa var., 107

lamourouxii, Fucus, 107

lamourouxii, Halimeda incrassata f., 115

lamourouxii, Halimeda tridens f., 115

lapidescens, Galaxaura, 23

lata, Caulerpa freycinetii var. typica f., 109

lata, Caulerpa serrulata f., 109

lata, Dictyota, 76

laticuspidata, Turbinaria conoides f., 89

latifolium, Sargassum, 85

latifolius, Fucus, 85

latifrons, Scinaia, 25

latissima, Ulva, 93

latissima, Ulva lactuca var., 93

latissimum, Monostroma, 91

Laurencia, 64, 124

brongniartii, 64

capituliformis, 65

caraibica, 65

carolinensis, 65

cartilaginea, 65

ceylanica, 65

chondrioides, 65

clavata, 65

columellaris, 65

composita, 65

concinna, 64

corallopsis, 65

decumbens, 65

dendroidea, 67

distichophylla, 66

forsteri, 66

glandulifera, 66

grevilleana, 65

implicata, 66

intermedia, 66

intricata, 66

japonica, 66

majuscula, 66

mariannensis, 66

nana, 65

nidifica, 66

obtusa, 66

obtusa var. dendroidea, 67

obtusa var. densa, 67

obtusa var. majuscula, 66

obtusa var. snackeyi, 67

okamurae, 67

palisada, 67

paniculata, 67

paniculata var. snackeyi, 67

papillosa, 67

parvipapillata, 67

patentiramea, 67

pinnata, 68

pinnatifida, 5

poiteaui, 68

seticulosa, 63

subsimplex, 68

surculigera, 68

tronoi, 68

tropica, 68

undulata, 68

venusta, 68

yamadana, 68

laxa, Heterosiphonia crispella var., 60

laxa, Heterosiphonia wurdemannii var., 60

laysanense, Phormidium, 12

Leathesia, 79

difformis, 79

leclancheri, Anadyomene, 94

leclancheri, Cystodictyon, 94

leeuwenii, Eucheuma, 47

Leibleinia

aeruginea, 13

capillacea, 10

corallinae, 11

polychroa, 10

sordida, 10

violacea, 10

lemoinei, Cruoriella, 5

lentillifera, Caulerpa, 105

lentillifera var. compacta, Caulerpa, 106

lentillifera var. longistipitata, Caulerpa, 106

leprieurii, Caloglossa, 58

leprieurii, Delesseria, 58

leprieurii var. hookeri, Caloglossa, 58

leprosa, Liagora, 20

leprosa, Liagora ceranoides f., 20

Leptofauchea, 124

leptophylla, Fauchea, 52

Leptothrix valderiae, 12

lessonii, Caulerpa, 106

Leveillea, 68

gracilis, 68

jungermannioides, 68

levis, Avrainvillea, 117

lherminieri, Antithamnion, 53

Liagora, 20, 124

boergesenii, 20

caenomyce, 20

canariensis, 20

ceranoides, 20

ceranoides f. leprosa, 20

cheyneana, 21

divaricata, 20

farinosa, 20

farinosa var. cheyneana, 21

hawaiiana, 21

japonica, 21

leprosa, 20

orientalis, 21

pulverentula, 5

robusta, 21

segawae, 21

setchellii, 21

tenuis, 21

valida, 21

liagorae, Acrochaetium, 19

Liagoropsis, 20

schrammii, 20

licheniformis, Mastophora, 37

lichenoides, Fucus, 42

lichenoides, Gigartina, 42

lichenoides, Gracilaria, 42

lichenoides, Sphaerococcus, 42, 44

lichenoides f. coronopifolia, Gracilaria, 41

liebetruthii, Cladophora, 98

ligulatus var. angustus, Gymnogongrus, 28

ligustica, Chaetomorpha, 96

ligustica, Conferva, 96

limensis, Euhymenia, 40

limensis, Sebdenia, 40

linearis, Dictyota, 76

linearis, Zonaria, 76

lingulata, Enteromorpha, 93

linum, Chaetomorpha, 96

linum, Conferva, 96

Litholepis

indica, 36

indica f. philippinensis, 36

Lithophyllum, 36, 124

byssoides, 36

moluccense, 36

okamurae, 36

onkodes, 38

pallescens, 36

reinboldii, 34

Lithoporella, 36

indica, 36

melobesioides, 36

Lithothamnion, 36

australe f. brachiatum, 36

australe f. minutulum, 37

australe f. ubianum, 37

byssoides, 36

calcareum, 38

dickiei, 37

erubescens, 37

erythraeum, 38

fruticulosum, 37

imbricatum, 37

indicum f. subtile, 37

moluccense, 36

muelleri, 36

onkodes, 38

pallescens, 36

polymorphum, 37, 38

pulchrum, 37

purpureum, 38

siamense, 37

siamense f. simulans, 38

simulans, 38

Lithothamnium, 36

Lobophora, 77

variegata, 77

Lola, 96

Lomentaria, 124, 52

articulata, 52

baileyana, 52

hakodatensis, 52

impudica, 48

pinnata, 52

longiarthra, Jania, 35

longicaulis, Avrainvillea, 117

longicaulis, Rhipilia, 117

longifolia, Caulerpa serrulata var. boryana f., 109

longipes, Caulerpa plumaris var., 110

longipes, Caulerpa sertularioides f., 110

longiseta, Caulerpa plumaris var., 110

longiseta, Caulerpa sertularioides f., 110

longistipitata, Caulerpa lentillifera var., 106

Lophocladia, 69

lallemandii, 69

Lophosiphonia, 69, 124

cristata, 69

sparsa, 70

Lophura armata, 62

loureiri, Ceramium, 55

luetzelburgii, Ceramium, 55

lutea, Lyngbya, 9

lutea, Oscillatoria, 9

luteofusca, Calothrix, 10

luxurians, Cryptonemia, 29

luxurians, Sphaerococcus lactuca var., 29

luzonense, Sargassum microcystum f., 85

luzonensis, Peyssonnelia, 28

luzonensis, Turbinaria, 89

lycopodium, Caulerpa, 105

lycopodium, Caulerpa cupressoides var., 105

lycopodium f. amicorum, Caulerpa cupressoides var., 105

lycopodium f. disticha, Caulerpa cupressoides var., 105

lycopodium f. elegans, Caulerpa cupressoides var., 105

Lyngbya, 9, 123

aestuarii, 9

cantharidosma, 8

confervoides, 9

crispa, 9

ferruginea, 9

glutinosa, 8

infixa, 9

lutea, 9

majuscula, 9, 12

martensiana, 10

mesotricha, 10

persicina, 12

rosea, 10

semiplena, 10

sordida, 10, 12

lyngbyaceum, Hydrocoleum, 9

lyngbyaceus, Microcoleus, 8, 9, 10, 11

macra, Caulerpa racemosa var., 107

macrocarpa, Mastophora, 5

macrodisca, Caulerpa, 108

macrodisca, Caulerpa peltata var., 108

macroloba, Halimeda, 115

macrophysa, Caulerpa racemosa var., 107

macrophysa, Chauvinia, 107

macrophysa, Halimeda, 115

macrophysa, Valonia, 103

maculans, Brachytrichia, 15

maculata, Halymenia, 31

maillardii, Carpopeltis, 29

maillardii, Phyllophora, 29

major, Acetabularia, 121

major, Chlorodesmis, 118

major, Dictyota, 76

major, Spirulina, 12

majuscula, Conferva, 8, 9

majuscula, Laurencia, 66

majuscula, Laurencia obtusa var., 66

majuscula, Lyngbya, 9, 12

manilense, Sargassum hombronianum var., 85

manilense, Sargassum polycystum var., 85

manipaense, Sargassum siliquosum var., 88

marcosii, Porphyra, 18

margaritifera, Oscillaria, 11

margaritifera, Oscillatoria, 11

marginata, Corallina, 23

marginata, Dictyota, 77

marginata, Galaxaura, 23

marginatus, Dilophus, 77

mariannensis, Laurencia, 66

marina, Derbesia, 104

marina, Vaucheria, 104

mariti, Cruoriella, 28

mariti, Peyssonnelia, 28

marshallense, Ceramium, 55

Martensia, 59, 124

flabelliformis, 59

speciosa, 59

martensiana, Lyngbya, 10

martensii, Opephyllum, 59

maryae, Ceramium, 55

masonii, Ceramium, 55

Mastophora, 37

decaisnei, 37

licheniformis, 37

macrocarpa, 5

melobesioides, 36

rosea, 37

mauritiana, Cladophora, 99

mazatlanense, Ceramium, 55

media, Chaetomorpha, 95

media, Conferva, 95

mediterranea, Spongopsis, 96

megalocystum, Goniolithon, 38

megalocystum, Neogoniolithon, 38

Melanthalia, 129

Melobesia, 124

farinosa, 34

foliacea, 37

melobesioides, Lithoporella, 36

melobesioides, Mastophora, 36

membranacea, Cladophoropsis, 101

membranacea, Conferva, 101

membranacea, Dictyopteris, 74

membranacea, Polypodoidea, 74, 75

membranaceus, Fucus, 74

Meredithia californica, 32

Meristotheca, 48

papulosa, 48

mertensii, Dictyota, 76

mertensii, Ulva, 76

mesenterica, Heteractis, 14

mesenterica, Rivularia, 14

Mesogloia

attenuata, 56

microcarpa, 31

Mesophyllum, 37

erubescens, 37

imbricatum, 37

pulchrum, 37

siamense, 37

simulans, 38

mesotricha, Lyngbya, 10

mexicana, Caulerpa, 106

mexicana, Jania, 35

mexicana, Schizothrix, 10, 12, 13

mexicana var. pluriseriata, Caulerpa, 106

micrarthodia, Jania, 35

microcarpa, Halymenia, 31

microcarpa, Mesogloia, 31

Microcladia, 57, 124

elegans, 57

glandulosa, 57

Microcoleus, 10

acutissimus, 10, 12

corymbosus, 14

lyngbyaceus, 8, 9, 10, 11,

tenerrimus, 10

vaginatus, 11

microcystis, Carpacanthus, 85

microcystum, Sargassum, 85

microcystum f. luzonense, Sargassum, 85

microcystum var. microtis, Sargassum, 85

Microdictyon, 95, 126

agardhianum, 95

boergesenii, 95

clathratum, 95

curtissiae, 95

japonicum, 95

montagnei, 100

okamurae, 95

umbilicatum, 5

vanbosseae, 5

micronesica, Halimeda, 115

microphyllum, Sargassum, 85

microphysa, Caulerpa, 106

microphysa, Caulerpa racemosa var. clavifera f., 106

microtis, Sargassum microcystum var., 85

Millepora

calcarea, 38

polymorpha, 38

miniata, Wurdemannia, 72

miniatus, Fucus, 72

miniatus, Sphaerococcus, 72

minima, Chnoospora, 79

minimus, Fucus, 79

minor, Corallopsis, 43

minor, Corallopsis salicornia var., 43

minor, Gracilaria, 43

minor, Herposiphonia dendroidea var., 64

minor, Padina, 78

minutissima, Acetabularia, 122

minutissima, Crouania, 56

minutulum, Lithothamnion australe f., 37

minutum, Centroceras, 54

mitchelliae, Ectocarpus, 73, 130

mitchelliae, Hincksia, 73, 130

mitchelliae var. neilii, Giffordia, 130

mitchelliae var. neilii, Hincksia, 130

mixta, Bostrychia, 62

mixta f. inermis, Bostrychia, 62

mixtum, Sargassum cinctum var., 82

miyabei, Sargassum, 86

moebii, Acetabularia, 122

molle, Archaeolithothamnion erythraeum f., 38

molle, Sporolithon ptychoides f., 38

mollis, Dasya, 59

mollis, Polysiphonia, 70

mollis var. tongatensis, Polysiphonia, 70

moluccense, Lithophyllum, 36

moluccense, Lithothamnion, 36

monile, Halimeda, 114

moniliformis, Scinaia, 25

Monostroma, 91, 126

latissimum, 91

nitidum, 91

montagneana, Rhodymenia, 45

montagneana, Sarcodia, 45

montagnei, Boodlea, 100

montagnei, Microdictyon, 100

moritziana, Bostrychia, 62

moritziana, Polysiphonia, 62

mucifera, Sphacelaria, 74

muelleri, Codium, 113

muelleri, Dasya, 60

muelleri, Heterosiphonia, 60

muelleri, Lithothamnion, 36

multifida, Claudea, 58

multijugum, Ceramium, 55

multipartita var. aeruginosa, Gracilaria, 42

muricatum, Eucheuma, 46

muricatum f. depauperatum, Eucheuma, 46

muricatum f. incrassatum, Eucheuma, 47

muricatus, Fucus, 46

murrayana, Turbinaria, 89

Murrayella, 69, 124

periclados, 69

musciformis, Fucus, 49

musciformis, Hypnea, 49

musciformis var. esperi, Hypnea, 49

musciformis var. hippuroides, Hypnea, 50

muscoides, Acanthophora, 60

muscoides, Fucus, 60

myagroides, Fucus, 81

myagroides, Myagropsis, 81

Myagropsis, 81

myagroides, 81

myriocystum, Sargassum, 86

myriocystum var. euryphyllum, Sargassum, 86

nana, Acrocystis, 61

nana, Laurencia, 65

nana, Pterocladia, 27

natans, Fucus, 86

natans, Sargassum, 86

neilii, Giffordia mitchelliae var., 130

neilii, Hincksia mitchelliae var., 130

Nemalion schrammii, 20

neocaledonica, Cladophoropsis, 101

neocaledonica, Spongocladia, 101

Neogoniolithon, 38

frutescens, 38

megalocystum, 38

Neomeris, 121

annulata, 121

nitida, 120

sphaerica, 120

vanbosseae, 121

Neurymenia, 69

fraxinifolia, 69

nidifica, Laurencia, 66

nidulans, Hypnea, 50

nigrescens, Hypnea, 50

nigricans, Avrainvillea, 117

nigrifolium, Sargassum, 86

nigroviridis, Oscillatoria, 11

nipae, Catenella, 48

nitens, Ceramium, 55

nitens, Ceramium rubrum var., 55

nitida, Bometella, 119

nitida, Neomeris, 120

nitida, Rivularia, 14

nitidulum, Acrochaetium, 19

nitidum, Monostroma, 91

Nitophyllum tongatense, 5

nootkanus, Fucus, 50

Nostoc quoyi, 15

Nothogenia, 128

variolosa, 128

notarisii, Porphyrosiphon, 11

novae-hollandae, Sphacelaria, 74

nuda, Herposiphonia, 64

Nullipora byssoides, 36

nummularia, Caulerpa, 108

nummularia, Caulerpa peltata var., 108

oblongata, Corallina, 23

oblongata, Galaxaura, 23

obscura, Anadyomene, 117

obscura, Avrainvillea, 117

obscura, Herposiphonia, 64

obscura, Peyssonnelia, 28

obtusa, Laurencia, 66

obtusa var. dendroidea, Laurencia, 67

obtusa var. densa, Laurencia, 67

obtusa var. majuscula, Laurencia, 66

obtusa var. paniculata, Chondria, 67

obtusa var. patentiramea, Chondria, 67

obtusa var. snackeyi, Laurencia, 67

obtusata, Corallina, 24

obtusata, Galaxaura, 24

obtusata, Tubularia, 24

obtusiloba, Rytiphlaea, 71

obtusiloba, Vidalia, 71

obtusus, Fucus, 66

occidentalis, Caulerpa chemnitzia var., 108

occidentalis, Caulerpa freycinetii var. boryana f., 109

occidentalis, Caulerpa racemosa var., 108

occidentalis, Caulerpa serrulata var. boryana f., 109

occidentalis, Udotea, 119

ocellata, Dasya, 59

ocellata, Dictyosphaeria, 102

ocellata, Valonia, 102

ocellatum, Ceramium, 59

Ochlochaete dendroides, 91

ogasawaraensis, Caloglossa, 58

okamurae, Callophyllis, 32

okamurae, Dilophus, 77

okamurae, Enantiocladia, 63

okamurae, Eucheuma, 47

okamurae, Laurencia, 67

okamurae, Lithophyllum, 36

okamurae, Microdictyon, 95

oligocystum, Sargassum, 86

oligocystum var. subflexuosum, Sargassum, 86

oligocystum var. bemardinum, Sargassum, 86

oligospora, Bornetella, 120

onkodes, Lithophyllum, 38

onkodes, Lithothamnion, 38

onkodes, Porolithon, 38

onslowensis, Giffordia, 130

onslowensis, Hincksia, 130

oocyste, Sargassum, 86

oocyste var. bemardinum, Sargassum, 87

Opephyllum, 59

martensii, 59

opuntia, Catenella, 48

opuntia, Corallina, 115

opuntia, Corallopsis, 41

opuntia, Fucus, 48

opuntia, Halimeda, 115

opuntia f. cordata, Halimeda, 116

opuntia f. hederacea, Halimeda, 114

opuntia f. intermedia, Halimeda, 116

opuntia f. renschii, Halimeda, 116

opuntia f. triloba, Halimeda, 116

opuntia var. hederacea, Halimeda, 114

orientate, Encoelium, 80

orientalis, Acanthophora, 61

orientalis, Acanthophora spicifera f., 61

orientalis, Asperococcus, 80

orientalis, Chamaedoris, 100

orientalis, Halimeda, 115

orientalis, Hydroclathrus, 80

orientalis, Liagora, 21

orientalis, Peyssonnelia rubra f., 28

orientalis, Rhipidosiphon, 119

orientalis, Rosenvingea, 80

orientalis, Udotea, 119

ornata, Asterocytis, 17

ornata, Conferva, 17

ornata, Turbinaria, 89

ornata f. evesiculosa, Turbinaria, 90

ornatum, Chroodactylon, 17

ornatus, Fucus, 128

ornatus, Fucus turbinatus var., 89

Oscillaria, 127

bonnemaisonii, 11

margaritifera, 11

sancta, 11

submembranacea, 12, 127

oscillarioides, Anabaina, 8

Oscillatoria, 11, 123, 127

bonnemaisonii, 11

brevis, 5

corallinae, 11

curviceps, 11

erythraea, 12

lutea, 9

margaritifera, 11

nigroviridis, 11

sancta, 11

submembranacea, 12, 127

ovale, Codium, 113

ovale, Gastridium, 104

ovalis, Bometella, 120

ovalis, Griffithsia, 56

ovalis, Halicystis, 104

ovata, Hincksia, 130

ovatus, Ectocarpus, 130

ovatus var. intermedius, Ectocarpus, 130

pachynema, Bryopsis, 103

pachynema, Valoniopsis, 103

pacifica, Amphiroa, 33

pacifica, Galaxaura, 24

pacifica, Herposiphonia, 64

pacifica, Jania, 35

pacifica, Kallymenia, 32

pacifica, Polysiphonia, 70

pacifica var. delicatula, Polysiphonia, 70

pacificum, Gelidium pusillum var., 27

Padina, 77, 125

arborescens, 77

australis, 77

boryana, 77

commersonii, 77

crassa, 78

distromatica, 78

fraseri, 78

gymnospora, 77, 78

jamaicensis, 78

japonica, 78

minor, 78

pavonia, 78

pavonica, 78

sanctae-crucis, 78

tenuis, 77

tetrastromatica, 79

palisada, Laurencia, 67

pallescens, Lithophyllum, 36

pallescens, Lithothamnion, 36

Palmaria, 19

palmata, 19

palmata, Palmaria, 19

palmatifida, Dasyopsis, 130

palmatifidus, Eupogodon, 130

palmatus, Fucus, 19

Palmella conferta, 7

Palmogloea aeruginosa, 7

paniculata, Chondria obtusa var., 67

paniculata, Laurencia, 67

paniculata var. snackeyi, Laurencia, 67

paniculatum, Sargassum, 87

pannosa, Chnoospora, 80

pannosa, Hypnea, 50

papenfussii, Acrochaetium, 19

papenfussii, Gracilaria, 43

papillatum, Codium, 113

papulosa, Chondria, 67

papillosa, Laurencia, 67

papillosus, Fucus, 67

papulosa, Kallymenia, 48

papulosa, Meristotheca, 48

paradoxa, Conferva, 92

paradoxa, Enteromorpha flexuosa subsp., 92

paradoxa, Ulva, 92

parca, Herposiphonia, 64

parietalis, Erythrotrichia, 17

parietina, Calothrix, 13, 14

parietinus, Schizosiphon, 13

parva, Pterocladia, 27

parvifolia, Caulerpa, 104

parvifolia, Caulerpa brachypus f., 104

parvifolium, Sargassum, 87

parvifolius, Fucus, 87

parvipapillata, Laurencia, 67

parvula, Acetabularia, 122

parvula, Champia, 53

parvula, Chondria, 53

parvula, Halicystis, 104

patens, Dictyota, 77

patens, Plocamium, 45

patens var. schizophyllum, Sargassum, 87

patentiramea, Chondria, 68

patentiramea, Chondria obtusa var., 67

patentiramea, Laurencia, 67

pavonia, Padina, 78

pavonia var. tenuis, Zonaria, 78

pavonica, Padina, 78

pavonicus, Fucus, 78

pavonius, Fucus, 78

pectinata, Caulerpa freycinetii var., 109

pectinata, Caulerpa serrulata var., 109

pectinatum, Plocamium serrulatum var., 45

Pedobesia ryukyuensis, 104

pedicellata, Pollexfenia, 59

pellucida, Cladophora, 98

pellucida, Conferva, 98

peltata, Caulerpa, 108

peltata, Caulerpa racemosa var., 108

peltata, Rhipiliopsis, 118

peltata, Udotea, 118

peltata var. lacunosa, Rhodophyllis, 48

peltata var. macrodisca, Caulerpa, 108

peltata var. nummularia, Caulerpa, 108

penicillata, Dasya, 130

penicillata, Griffithsia, 57

penicillata, Wrangelia, 57

penicillatum, Phormidium, 12

penicillatus, Eupogodon, 130

pennata, Bryopsis, 103

pennata var. secunda, Bryopsis, 103

pentamera, Polysiphonia, 5

periclados, Hutchinsia, 69

periclados, Murrayella, 69

perpusilla, Polysiphonia, 59

perpusillum, Gelidium crinale var., 25

perpusillum, Taenioma, 59

persicina, Lyngbya, 12

persicinum, Phormidium, 12

personatum, Ceramium, 55

pertusa, Ulva, 94

Peyssonnelia, 27, 124

calcea, 27

conchicola, 27

distenta, 28

evae, 28

foveolata, 28

indica, 28

luzonensis, 28

mariti, 28

obscura, 28

rubra, 28

rubra f. orientalis, 28

squamaria, 28

Phaeophila, 91

dendroides, 91

philippinense, Sargassum, 87

philippinensis, Acetabularia, 122

philippinensis, Acetabularia ryukyuensis var., 122

philippinensis, Chlorocladus, 120

philippinensis, Cladophora, 98

philippinensis, Cladophoropsis, 101

philippinensis, Litholepis indica f., 36

Phormidium, 11

ambiguum, 11, 14

amoenum, 11

crosbyanum, 11

laysanense, 12

penicillatum, 12

persicinum, 12

submembranaceum, 12, 127

tinctorium, 12

valderiae, 12

valderianum, 12

Phycoseris reticulata, 94

Phyllophora, 51

crenulata, 29

intricata, 52

maillardii, 29

submaritima, 51

Phymatolithon, 38

calcareum, 38

polymorphum, 38

purpureum, 38

pickeringii, Caulerpa, 111

pickeringii, Caulerpa webbiana var., 111

pilosa, Calothrix, 13, 14, 15

pilosa, Dasyopsis, 60, 130

pilosus, Eupogodon, 60, 130

pilulifer, Fucus, 87

piluliferum, Sargassum, 87

pinnata, Erythrocladia, 17

pinnata, Laurencia, 68

pinnata, Lomentaria, 52

pinnatifida, Laurencia, 5

pinnatifolia, Ptilota, 130

pinnatifolia var. digitata, Corallina, 34

pinnatifolius, Eupogodon, 130

plana, Dasya, 129

planus, Eupogodon, 130

platyclados, Codium, 113

platydisca, Halimeda, 116

platydisca, Halimeda tuna f., 116

Pleonosporium, 57

globuliferum, 57

plicata, Anadyomene, 94

Plocamium, 44

costatum, 44

patens, 45

serrulatum var. pectinatum, 45

telfairiae, 45

tenellum, 62

plocamium var. costata, Delesseria, 44

plumarioides, Dasyphila, 56

plumaris, Caulerpa, 110

plumaris, Fucus, 110

plumaris f. farlowii, Caulerpa, 110

plumaris f. flagellata, Caulerpa, 110

plumaris var. brevipes, Caulerpa, 110

plumaris var. elegans, Caulerpa, 105

plumaris var. longipes, Caulerpa, 110

plumaris var. longiseta, Caulerpa, 110

plumosa, Bryopsis, 103

plumosa, Enteromorpha, 92

plumosa, Ulva, 103

plumosa var. secunda, Bryopsis, 103

plumula, Herposiphonia, 64

plumula, Polysiphonia, 64

pluriseriata, Caulerpa mexicana var., 106

Pocockiella variegata, 77

podagricum, Erythrocolon, 52

poiteaui, Laurencia, 68

poitei, Fucus, 68

poko, Polysiphonia, 70

Pollexfenia pedicellata, 59

Polycavernosa, 44

debilis, 44

polyceratium, Sargassum, 87

polychroa, Leibleinia, 10

Polycoelia vanhoevelii, 5

polycystum, Sargassum, 87

polycystum var. euryphyllum, Sargassum, 86

polycystum var. manilense, Sargassum, 85

polymorpha, Millepora, 38

polymorphum, Lithothamnion, 37, 38

polymorphum, Phymatolithon, 38

polyotis, Diplotrichia, 15

polyotis, Rivularia, 15

Polyphysa, 121

spicata, 122

polypodioides, Dictyopteris, 74

polypodioides, Fucus, 74, 75

polypodioides, Ulva, 75

Polypodoidea, 75

membranacea, 74, 75

polyrhiza, Chondria, 63

Polysiphonia, 69, 124

apiculata, 69

beaudettei, 69

calodictyon, 71

ferulacea, 69

flabellulata, 69

forcipata, 69

forfex, 69

fragilis, 69

gorgoniae, 69

hawaiiensis, 70

howei, 70

mollis, 70

mollis var. tongatensis, 70

moritziana, 62

pacifica, 70

pacifica var. delicatula, 70

pentamera, 5

perpusilla, 59

plumula, 64

poko, 70

savatieri, 70

scopulorum, 70

scopulorum var. villum, 70

setacea, 70

sparsa, 70

sphaerocarpa, 70

subtilissima, 71

tenuis, 71

tepida, 71

tongatensis, 70

tongatensis var. upolensis, 71

triton, 71

tsudana, 71

upolensis, 71

villum, 70

Polystrata, 28

dura, 28

Polyzonia jungermannioides, 68

Porolithon, 38

onkodes, 38

Porphyra, 18, 124

atropurpurea, 18

crispata, 18

denticulata, 18

marcosii, 18

suborbiculata, 18

variegata, 18

Porphyrosiphon

kurzii, 12

notarisii, 11

Portieria, 39, 128

coccinea, 129

dichotoma, 129

harveyi, 129

hornemannii, 39, 129

japonica, 39, 129

kilneri, 129

spinulosa, 129

Prionitis, 31

articulata, 29

cornea, 31

schmitziana, 29

procrusteanum, Eucheuma, 47

procumbens, Ceramiella, 54

procumbens, Ceramium, 54

procumbens, Thamnoclonium, 31

prolifer, Fucus, 106, 115, 116

prolifera, Botryocarpa, 58

prolifera, Carpopeltis, 29

prolifera, Caulerpa, 106

prolifera, Cladophora, 98

prolifera, Conferva, 98

prolifera, Cystoseira, 81

prolifera, Enteromorpha, 93

prolifera, Giffordia, 130

prolifera, Gigartina, 29

prolifera, Hincksia, 130

prolifera, Ulva, 93

prorepens, Herposiphonia, 5

pseudoscillatoria, Anabaina, 8

pseudospinulosum, Sargassum ilicifolium var., 85

Pterocladia, 27, 124

caloglossoides, 27

capillacea, 27

densa, 27

nana, 27

parva, 27

pteropleuron, Sargassum, 87

Ptilota pinnatifolia, 130

Ptilothamnion, 57

cladophorae, 57

ptychoides f. durum, Sporolithon, 38

ptychoides f. molle, Sporolithon, 38

Pugetia japonica, 32

pugniforme, Codium, 113

pulchella, Dasyopsis, 130

pulchellum, Gelidium, 25

pulchellum var. subspathulatum, Sargassum, 88

pulchellus, Eupogodon, 130

pulchellus, Fucus corneus var., 25

pulchrum, Lithothamnion, 37

pulchrum, Mesophyllum, 37

pulverentula, Liagora, 5

pulvinata, Sphacelaria, 5

pumila, Jania, 35

Punctaria, 125

punicea, Baillouviana, 59

punicea, Dasya, 59

purpurascens, Gracilaria, 43

purpureum, Lithothamnion, 38

purpureum, Phymatolithon, 38

pusilla, Boodleopsis, 117

pusillum, Alsidium, 61

pusillum, Gelidium, 25

pusillum var. pacificum, Gelidium, 27

pusillus, Acrocarpus, 25

pusillus, Dichotomosiphon, 117

pusillus, Fucus, 25

pygmaeus, Gymnogongrus, 51

pyriformis, Botryocladia, 51

pyriformis, Chrysymenia, 51

quisumbingii, Cladophora, 98

quoyi, Brachytrichia, 15

quoyi, Nostoc, 15

racemosa, Caulerpa, 106

racemosa var. chemnitzia, Caulerpa, 108

racemosa var. clavifera, Caulerpa, 107

racemosa var. clavifera f. microphysa, Caulerpa, 106

racemosa var. corynephora, Caulerpa, 107

racemosa var. gracilis, Caulerpa, 107

racemosa var. laetevirens, Caulerpa, 107

racemosa var. lamourouxii, Caulerpa, 107

racemosa var. macra, Caulerpa, 107

racemosa var. macrophysa, Caulerpa, 107

racemosa var. occidentalis, Caulerpa, 108

racemosa var. peltata, Caulerpa, 108

racemosa var. turbinata, Caulerpa, 108

racemosa var. uvifera, Caulerpa, 107

racemosus, Fucus, 106

radicans, Bostrychia, 62

radicans, Rhodomela, 62

Ralfsia, 73

fungiformis, 73

rallsiae, Ectocarpus, 73, 130

rallsiae, Giffordia, 73, 130

rallsiae, Hincksia, 73, 130

ramellifera, Gloiocladia, 52

ramosa, Struvea, 101

ramosissima, Grateloupia, 30

ramulosa, Enteromorpha, 93

ramulosa, Ulva, 93

ramulosa var. spinosa, Enteromorpha, 93

reinboldii, Goniolithon, 34

reinboldii, Hydrolithon, 34

reinboldii, Lithophyllum, 34

renschii, Halimeda, 116

renschii, Halimeda opuntia f., 116

repens, Chondria, 63

repens, Codium, 112

repens, Dictyopteris, 75

repens, Gelidiopsis, 40

repens, Gelidium, 40

repens, Haliseris, 75

requienii, Batrachospermum, 19

requienii, Trichogloea, 19

reticulata, Phycoseris, 94

reticulata, Ulva, 84, 94

retroflexa, Turbinaria conoides f., 89

reyesii, Caulerpa, 108

Rhabdonia dura, 48

Rhipidosiphon

javensis, 119

orientalis, 119

Rhipilia longicaulis, 117

Rhipiliopsis, 118

carolyniae, 118

peltata, 118

Rhizoclonium, 99

africanum, 99

capillare, 96

crassipellitum var. robustum, 99

grande, 99

hookeri, 99

kerneri, 99

kochianum, 99

riparium, 99

setaceum, 99

rhizophora, Griffithsia, 56

Rhodochorton, 127

hancockii, 19

sinicola, 19

Rhodomela

calliptera, 61

radicans, 62

Rhodopeltis, 39, 124

borealis, 39

gracilis, 39

Rhodophyllis, 48

peltata var. lacunosa, 48

Rhodymenia, 52, 124

californica, 52

coacta, 52

decumbens, 52

intricata, 52

montagneana, 45

spinulosa, 43

tripinnata, 129

rigens, Gelidium, 27

rigens, Sphaerococcus, 27

rigida, Actinotrichia, 22

rigida, Amphiroa, 33

rigida, Carpopeltis, 29

rigida, Cryptonemia, 29

rigida, Galaxaura, 22

rigida, Gelidiopsis, 25

rigida, Sphacelaria, 74

rigida, Ulva, 94

rigida, Ulva lactuca var., 94

rigidula, Sphacelaria, 74

rigidum, Gelidium, 25

rigidus, Fucus, 25

rigidus, Sphaerococcus, 25

riparia, Conferva, 99

riparium, Rhizoclonium, 99

Rivularia, 14, 123

bullata, 13, 14

contarenii, 13

mesenterica, 14

nitida, 14

polyotis, 15

rivularis, Bostrychia, 62

robusta, Calothrix, 14

robusta, Galaxaura, 24

robusta, Halimeda velasquezii var., 116

robusta, Liagora, 21

robustior, Avrainvillea lacerata var., 117

robustum, Acrochaetium, 19

robustum, Rhizoclonium crassipellitum var., 99

Roschera

africana, 71

condensata, 71

rosea, Lyngbya, 10

rosea, Mastophora, 37

rosea, Zonaria, 37

Rosenvingea, 80

intricata, 80

orientalis, 80

rosulata, Kylinia, 19

rotundatum, Sargassum duplicatum var., 83

roxasii, Acetabularia, 122

rubens, Corallina, 35

rubens, Jania, 35

rubra, Peyssonnelia, 28

rubra, Zonaria, 28

rubra f. orientalis, Peyssonnelia, 28

rubrum var. nitens, Ceramium, 55

rudis, Galaxaura, 23

rudolphiana, Cladophora, 99

rudolphiana, Conferva, 99

rugosa, Corallina, 24

rugosa, Galaxaura, 24

rugulosa, Cladophora, 98

rupestris, Cladophora, 98

rupestris, Conferva, 98

Rytiphlaea obtusiloba, 71

ryukyuensis, Acetabularia, 122

ryukyuensis, Derbesia, 104

ryukyuensis, Pedobesia, 104

ryukyuensis var. philippinensis, Acetabularia, 122

saccharina, Laminaria, 94

sagamianum, Sargassum, 87

sagittata, Corallina, 34

sagittatum, Cheilosporum, 34

saidana, Hypnea, 50

sakaii, Cladophora, 98

salicifolius, Fucus, 88

salicornia, Corallopsis, 43

salicornia, Gracilaria, 43

salicornia, Sphaerococcus, 43

salicornia var. minor, Corallopsis, 43

salicornoides, Champia, 53

salina, Enteromorpha, 93

sancta, Oscillaria, 11

sancta, Oscillatoria, 11

sanctae-crucis, Padina, 78

sandei, Sargassum, 87

sandriana, Hincksia, 130

sandrianus, Ectocarpus, 130

sanfordiana, Asparagopsis, 22

Sarcodia, 45, 124

ceylanica, 45

montagneana, 45

filamentosa, 58

sargassi, Acrochaetium, 19

Sargassum, 82, 125

aciculare, 84

aemulum, 82

agardhianum, 82

bacciferum, 86

baccularia, 82

belangeri, 82

berberifolium, 82

biforme, 82

binderi, 86

binderi var. angustifolium, 82

binderi var. vitiense, 86

carpophyllum, 85

cervicome, 82

cinctum var. mixtum, 82

cinctum var. gracilentum, 82

compactum, 85

confusum, 82

crispifolium, 82

cristaefolium, 83

cristatum, 82

cystocarpum, 83

cystophyllum, 83

droserifolium, 83

droserifolium var. spathulatum, 83

duplicatum, 83

duplicatum var. rotundatum, 83

echinocarpum var. vitiense, 86

elongatum, 83

enerve, 84

esperi, 83

filicinum, 83

filifolium, 83

filifolium var. aciculare, 84

filiforme, 84

fluitans, 84

fulvellum, 84

furcatum, 84

gaudichaudii, 84

giganteifolium, 84

gracile, 84

granuliferum, 84

hemiphyllum, 84

heterocystum, 85

hombronianum var. manilense, 85

hystrix, 85

hystrix var. fluitans, 84

hystrix var. spinulosus, 85

ilicifolium, 85

ilicifolium f. benkulense, 85

ilicifolium var. compactum, 85

ilicifolium var. duplicatum, 83

ilicifolium var. pseudospinulosum, 85

kjellmanianum, 86

latifolium, 85

microcystum, 85

microcystum f. luzonense, 85

microcystum var. microtis, 85

microphyllum, 85

miyabei, 86

myriocystum, 86

myriocystum var. euryphyllum, 86

natans, 86

nigrifolium, 86

oligocystum, 86

oligocystum var. bernardinum, 86

oligocystum var. subflexuosum, 86

oocyste, 86

oocyste var. bemardinum, 87

paniculatum, 87

parvifolium, 87

patens var. schizophyllum, 87

philippinense, 87

piluliferum, 87

polyceratium, 87

polycystum, 87

polycystum var. euryphyllum, 86

polycystum var. manilense, 85

pteropleuron, 87

pulchellum var. subspathulatum, 88

sagamianum, 87

sandei, 87

serratifolium, 87

siliquosum, 82, 87

siliquosum var. basilanicum, 88

siliquosum var. bicornutum, 88

siliquosum var. manipaense, 88

spinifex, 88

subspathulatum, 88

tenerrimum, 88

vulgare, 84, 88

vulgare var. indicum, 88

yendoi, 88

saundersii, Ectocarpus, 130

saundersii, Hincksia, 130

savatieri, Polysiphonia, 70

scalaramosa, Beckerella, 25

schizophylla, Halockloa, 87

schizophyllum, Sargassum patens var., 87

Schizosiphon

gypsophilus, 14

parietinus, 13

Schizothrix, 12, 123

arenaria, 13

calcicola, 9, 10, 11, 12

friesii, 12

mexicana, 10, 12, 13

tenerrima, 10, 12

schmidtii, Archaeolithothamnion, 39

schmidtii, Sporolithon, 39

schmitziana, Cryptonemia, 29

schmitziana, Prionitis, 29

schrammi, Helminthocladia, 20

schrammii, Liagoropsis, 20

schrammii, Nemalion, 20

Scinaia, 25, 124

hormoides, 25

latifrons, 25

moniliformis, 25

scopulorum, Calothrix, 14

scopulorum, Conferva, 14

scopulorum, Polysiphonia, 70

scopulorum var. villum, Polysiphonia, 70

Scytonema, 15

hoffman-bangii, 15

hofman-bangii, 15

hofmanii, 15

hofmannii, 14

intricatum, 61, 62

Scytosiphon erectus, 92

Sebdenia, 40

limensis, 40

yamadae, 40

secunda, Bryopsis pennata var., 103

secunda, Bryopsis plumosa var., 103

secunda, Herposiphonia, 64

secunda, Herposiphonia tenella f., 64

secunda, Hincksia, 130

secunda, Hutchinsia, 64

secunda f. tenella, Herposiphonia, 64

secundatum, Acrochaetium, 128

secundatum, Callithamnion daviesii var., 128

secundus, Ectocarpus, 130

sedifolia, Chondria, 63

segawae, Liagora, 21

selago, Caulerpa, 108

selago, Fucus, 108

semiplena, Calothrix, 10

semiplena, Lyngbya, 10

seriatum, Acrochaetium, 19

sericea, Cladophora, 99

sericea, Conferva, 99

serpens, Ceramium, 55

serra, Eucheuma, 47

serra, Sphaerococcus. 47

serrata, Thysanocladia, 45

serratifolium, Sargassum, 87

serratifolius, Fucus, 87

serratus, Callophycus, 45

serrulata, Caulerpa, 108

serrulata f. lata, Caulerpa, 109

serrulata f. spiralis, Caulerpa, 109

serrulata var. boryana, Caulerpa, 109

serrulata var. boryana f. longifolia, Caulerpa, 109

serrulata var. boryana f. occidentalis, Caulerpa, 109

serrulata var. pectinata, Caulerpa, 109

serrulatum, Hypoglossum, 58

serrulatum var. pectinatum, Plocamium, 45

serrulatus, Fucus, 108

sertularioides, Caulerpa, 109

sertularioides, Fucus, 109, 110

sertularioides f. brevipes, Caulerpa, 110

sertularioides f. farlowii, Caulerpa, 110

sertularioides f. flagellata, Caulerpa, 110

sertularioides f. longipes, Caulerpa, 110

sertularioides f. longiseta, Caulerpa, 110

sessilis, Dasya, 59

sessilis, Kallymenia, 32

setacea, Polysiphonia, 70

setaceum, Rhizoclonium, 99

setchellii, Codium, 112

setchellii, Coriophyllum, 5

setchellii, Dictyosphaeria, 102

setchellii, Liagora, 21

seticulosa, Chondria, 63

seticulosa, Conferva, 63

seticulosa, Laurencia, 63

siamense, Lithothamnion, 37

siamense, Mesophyllum, 37

siamense f. simulans, Lithothamnion, 38

siamensis, Boodlea, 100

sibogae, Archaeolithothamnion, 39

sibogae, Chondria, 63

sibogae, Cladophora, 99

sibogae, Galaxaura, 5

sibogae, Sporolithon, 39

siliquosum, Sargassum, 82, 87

siliquosum var. basilanicum, Sargassum, 88

siliquosum var. bicornutum, Sargassum, 88

siliquosum var. manipaense, Sargassum, 88

simplex, Conferva, 63

simplex, Digenea, 63

simpliciuscula, Bostrychia, 62

simpliciuscula, Bostrychia tenuis f., 62

simpliciuscula var. vesiculifera, Caulerpa, 111

simulans, Halimeda, 116

simulans, Lithothamnion, 38

simulans, Lithothamnion siamense f., 38

simulans, Mesophyllum, 38

sinicola, Acrochaetium, 19

sinicola, Ceramium, 55

sinicola, Rhodochorton, 19

sinuosa, Colpomenia, 80

sinuosa, Ulva, 80

Siphonocladus fasciculatus, 100

Sirocoleum, 12

kurzii, 12

Sirophysalis, 81

sisymbrioides, Cystophyllum, 81

sisymbrioides, Fucus, 81

skottsbergii, Botryocladia, 51

skottsbergii, Chrysymenia, 51

snackeyi, Gracilaria arcuata var., 41

snackeyi, Laurencia obtusa var., 67

snackeyi, Laurencia paniculata var., 67

Solieria, 48

dura, 48

solutum, Hormothamnium, 8

sordida, Avrainvillea, 117

sordida, Calothrix, 10

sordida, Hincksia, 130

sordida, Leibleinia, 10

sordida, Lyngbya, 10, 12

sordida, Udotea, 117

sordidus, Ectocarpus, 130

sparsa, Lophosiphonia, 70

sparsa, Polysiphonia, 70

spathulata, Champia, 53

spathulata, Delesseria, 58

spathulatum, Hypoglossum, 58

spathulatum, Sargassum droserifolium var., 83

Spatoglossum, 79, 125

asperum, 79

variabile, 79

speciosa, Martensia, 59

spectabile, Cheilosporum, 5

spectabilis, Vanvoorstia, 59

Spermothamnion, 124

cladophorae, 57

yonakuniense, 56

Sphacelaria, 74, 125

furcigera, 74

mucifera, 74

novae-hollandae, 74

pulvinata, 5

rigida, 74

rigidula, 74

tribuloides, 74

sphaerica, Bornetella, 120

sphaerica, Neomeris, 120

sphaerocarpa, Polysiphonia, 70

Sphaerococcus

canaliculars, 41

compressus, 41

confervoides, 44

corallopsis, 65

denticulatus, 44

distichus, 42

divaricatus, 49

gelatinus, 44, 47

intricatus, 40

isiformis, 47

lactuca var. luxurians, 29

lichenoides, 42, 44

miniatus, 72

rigens, 27

rigidus, 25

salicornia, 43

serra, 47

spinellus, 50

textorii, 43

spicata, Polyphysa, 122

spicifera, Acanthophora, 60

spicifera f. orientalis, Acanthophora, 61

spiciferus, Fucus, 60

spinella, Dasya, 129

spinella, Hypnea, 50

spinellus, Eupogodon, 130

spinellus, Sphaerococcus, 50

spinescens, Enteromorpha, 93

spinifex, Sargassum, 88

spiniforme, Gelidium, 25

spiniformis, Fucus, 25

spinigera, Gracilaria, 43

spinosa, Enteromorpha ramulosa var., 93

spinosum, Eucheuma, 46

spinosus, Fucus, 46

spinuligera, Endosiphonia, 63

spinulosa, Gracilaria, 43

spinulosa, Portieria, 129

spinulosa, Rhodymenia, 43

spinulosus, Carpacanthus, 85

spinulosus, Chondrococcus, 129

spinulosus, Sargassum hystrix var., 85

spiralis, Caulerpa freycinetii var. typica f., 109

spiralis, Caulerpa serrulata f., 109

spiralis, Chaetomorpha, 97

Spirulina, 12

major, 12

subsalsa, 12

Spongia vermicularis, 120

spongiosum, Ceratodictyon, 40

Spongocarpus hemiphyllus, 85

Spongocladia, 125

dichotoma, 100

neocaledonica, 101

vaucheriaeformis, 101

Spongodendron dichotomum, 100

Spongomorpha, 126

Spongopsis mediterranea, 96

Sporolithon, 38

erythraeum, 38

ptychoides f. durum, 38

ptychoides f. molle, 38

schmidtii, 39

sibogae, 39

timorense, 39

spumosa, Udotea argentea var., 119

Spyridia, 57

filamentosa, 49, 57,

velasquezii, 57

squalida, Galaxaura, 24

squamaria, Peyssonnelia, 28

squamarius, Fucus, 28

stanleyi, Dasyopsis, 130

stanleyi, Eupogodon, 130

stellata, Anadyomene, 94

stellata, Ulva, 94

stellulifera, Hypnea cornuta var., 49

striata, Galaxaura, 24

striatum, Eucheuma, 47

Struvea, 101

anastomosans, 101

delicatula, 101

ramosa, 101

struveoides, Boodlea, 100

stuposa, Halimeda, 116

Stypopodium, 79

flabelliforme, 79

subcorticata, Balliella, 54

subcorticatum, Antithamnion, 54

subcylindrica, Amphiroa, 33

subdigitata, Halimeda discoidea f., 114

subdisticha, Herposiphonia, 64

subflexuosum, Sargassum oligocystum var., 86

subfruticulosa, Galaxaura, 24

subintegra, Erythrocladia, 17

submarina, Enteromorpha flexuosa f., 92

submaritima, Phyllophora, 51

submembranacea, Oscillaria, 12

submembranacea, Oscillatoria, 12

submembranaceum, Phormidium, 12, 127

suborbiculata, Porphyra, 18

subsalsa, Spirulina, 12

subserrata, Caulerpa, 110

subsimplex, Laurencia, 68

subspathulatum, Sargassum, 88

subspathulatum, Sargassum pulchellum var., 88

subtile, Lithothamnion indicum f., 37

subtilissima, Polysiphonia, 71

subverticillata, Galaxaura, 24

Suhria, 128

sundanensis, Cladophoropsis, 101

surculigera, Laurencia, 68

Symploca, 13, 123

howei, 13

hydnoides, 13

laeteviridis, 13

taenicola, Halimeda, 116

Taenioma, 59

perpusillum, 59

tambalangii, Eucheuma alvarezii var., 45

Tapeinodasya, 60

bornetii, 60

tawallina, Zellera, 59

taxifolia, Caulerpa, 111

taxifolia var. crassifolia, Caulerpa, 106

taxifolius, Fucus, 111

taxiformis, Asparagopsis, 21

taxiformis, Fucus, 21, 22

taylorii, Ceramium, 56

taylorii, Gelidiella, 25

telfairiae, Plocamium, 45

telfairiae, Thamnophora, 45

tenax, Fucus, 27

tenax, Gloiopeltis, 27

tenella, Bostrychia, 62

tenella, Corallina, 35

tenella, Dasyopsis, 130

tenella, Gigartina, 51

tenella, Herposiphonia, 64

tenella, Herposiphonia secunda f., 64

tenella, Hutchinsia, 64

tenella, Jania, 35

tenella f. secunda, Herposiphonia, 64

tenella var. zacae, Jania, 35

tenellum, Plocamium, 62

tenellus, Eupogodon, 130

tenellus, Fucus, 62

tenera, Galaxaura, 23

tenerrima, Schizothrix, 10, 12

tenerrimum, Ceramium, 56

tenerrimum, Hormoceras, 56

tenerrimum, Sargassum, 88

tenerrimus, Microcoleus, 10

tenue, Anotrichium, 53

tenue, Codium, 113

tenuis, Bostrychia, 62

tenuis, Griffithsia, 53

tenuis, Hypnea cenomyce var., 48

tenuis, Liagora, 21

tenuis, Padina, 77

tenuis, Polysiphonia, 71

tenuis, Zonaria pavonia var., 78

tenuis f. simpliciuscula, Bostrychia, 62

tenuissima, Bryopsis, 104

tenuissima, Derbesia, 104

tenuissima, Jania, 35

tenuissimum, Ceramium, 56

tenuissimum, Ceramium diaphanum var., 56

tepida, Polysiphonia, 71

terminalis, Giffordia, 130

terminalis, Hincksia, 130

tetrastromatica, Padina, 79

textorii, Gracilaria, 43

textorii, Sphaerococcus, 43

Thamnoclonium, 31

procumbens, 31

treubii, 31

Thamnophora telfairiae, 45

thierryi, Acanthophora, 61

thyrsoidea, Hincksia, 130

thyrsoideus, Ectocarpus, 130

Thysanocladia serrata, 45

timorense, Archaeolithothamnion, 39

timorense, Sporolithon, 39

tinctorium, Phormidium, 12

Titanophora, 40, 124

incrustans, 40

weberae, 40

Tolypiocladia, 71, 124

calodictyon, 71

condensata, 71

glomerulata, 71

tomentosum, Codium, 113

tongatense, Nitophyllum, 5

tongatensis, Polysiphonia, 70

tongatensis, Polysiphonia mollis var., 70

tongatensis var. upolensis, Polysiphonia, 71

torresiensis, Chlorodesmis, 118

torta, Chaetomorpha, 97

torta, Chaetomorpha clavata var., 97

tortuosa, Chaetomorpha, 96

tortuosa, Conferva, 96

Tremella difformis, 79

treubii, Thamnoclonium, 31

trialata, Turbinaria, 90

trialata, Turbinaria vulgaris var., 90

trialata var. capensis, Turbinaria, 90

triangularis, Fucus, 131

triangulare, Bryothamnion, 131

tribuloides, Sphacelaria, 74

tribulus, Amphiroa, 33

tribulus, Corallina, 33

trichia, Herposiphonia, 64

Trichodesmium erythraeum, 12

Trichogloea, 19, 125

requienii, 19

trichogloeae, Acrochaetium, 19

Trichosolen, 126

trichotoma, Cladophora, 98

trichotoma, Conferva, 98

tridens, Corallina, 115

tridens, Halimeda, 115

tridens f. lamourouxii, Halimeda, 115

trifarius, Fucus, 130

triloba, Halimeda, 115

triloba, Halimeda opuntia f., 116

trinode, Cystophyllum, 81

triqueter, Fucus, 81, 131

triquetra, Cystoseira, 81, 131

triquetra, Hormophysa, 81, 130

triquetrum, Bryothamnion, 130

tripinnata, Desmia, 129

tripinnata, Portieria, 129

triton, Polysiphonia, 71

tronoi, Laurencia, 68

tropica, Laurencia, 68

tsudana, Polysiphonia, 71

Tubularia

obtusata, 24

umbellata, 24

tubulosa, Enteromorpha, 92

tubulosa, Enteromorpha intestinalis f., 92

tubulosa, Enteromorpha intestinalis var., 92

tuna, Corallina, 116

tuna, Halimeda, 116

tuna f. platydisca, Halimeda, 116

Turbinaria, 88, 89, 125

condensata, 88

conoides, 88

conoides f. laticuspidata, 89

conoides f. retroflexa, 89

conoides var. evesiculosa, 90

decurrens, 89

denudata, 89

filamentosa, 89

luzonensis, 89

murrayana, 89

ornata, 89

ornata f. evesiculosa, 90

trialata, 90

trialata var. capensis, 90

turbinata, 90

vulgaris, 90

vulgaris var. conoides, 88

vulgaris var. trialata, 90

turbinata, Caulerpa clavifera var., 108

turbinata, Caulerpa racemosa var., 108

turbinata, Turbinaria, 90

turbinatus, Fucus, 90

turbinatus var. ornatus, Fucus, 89

turgida, Gracilaria, 43

turneri, Cystophyllum, 81

tuticorinense, Acrochaetium, 19

Tydemania, 118

expeditionis, 118

typica f. lata, Caulerpa freycinetii var., 109

typica f. spiralis, Caulerpa freycinetii var., 109

ubianum, Lithothamnion australe f., 37

Udotea, 118

argentea, 118

argentea var. spumosa, 119

flabellum, 119

geppii, 119

glaucescens, 119

indica, 119

javensis, 119

occidentalis, 119

orientalis, 119

peltata, 118

sordida, 117

Ulva, 93, 126

articulata, 52

atropurpurea, 18

bullata, 14

caespitosa, 48

cavernosa, 101

compressa, 91, 93

dichotoma, 76

fasciata, 93

flexuosa, 92

gigantea, 94

intestinalis, 92

lactuca, 93

lactuca var. latissima, 93

lactuca var. rigida, 94

latissima, 93

mertensii, 76

paradoxa, 92

pertusa, 94

plumosa, 103

polypodioides, 75

prolifera, 93

ramulosa, 93

reticulata, 94, 80

rigida, 94

sinuosa, 80

stellata, 94

umbilicalis, 77

umbellata, Galaxaura, 24

umbellata, Tubularia, 24

umbilicalis, Ulva, 77

umbilicatum, Microdictyon, 5

uncinata, Cladophora, 97

undulata, Dictyopteris, 75

undulata, Laurencia, 68

ungulata, Corallina, 35

ungulata, Jania, 35

ungulata f. brevior, Corallina, 36

ungulata f. brevior, Jania, 36

upolensis, Polysiphonia, 71

urvilliana, Caulerpa, 111

urvilliana var. vitiensis, Caulerpa, 111

utricularis, Conferva, 103

utricularis, Valonia, 103

uvaria, Chondria, 51

uvaria, Chrysymenia, 51

uvarioides, Botryocladia, 51

uvifer, Fucus, 107

uvifera, Caulerpa, 107

uvifera, Caulerpa racemosa var., 107

vagabunda, Cladophora, 99

vagabunda, Conferva, 99

vagabundum, Ceramium, 56

vagans, Ceramium, 56

vaginatus, Microcoleus, 11

valderiae, Leptothrix, 12

valderiae, Phormidium, 12

valderianum, Phormidium, 12

valentiae, Fucus, 50

valentiae, Hypnea, 50

valida, Liagora, 21

Valonia, 102

aegagropila, 102

confervoides, 102

fastigiata, 102

favulosa, 102

forbesii, 100

macrophysa, 103

ocellata, 102

utricularis, 103

ventricosa, 103

verticillata, 102

valonioides, Amphiroa, 34

Valoniopsis, 103

pachynema, 103

vanbosseae, Boodlea, 5

vanbosseae, Cymopolia, 120

vanbosseae, Dictyosphaeria, 102

vanbosseae, Microdictyon, 5

vanbosseae, Neomeris, 121

vanhoevelii, Polycoelia, 5

Vanvoorstia, 59

spectabilis, 59

variabile, Gelidium, 40

variabile, Spatoglossum, 79

variabilis, Gelidiopsis, 40

variegata, Dictyota, 77

variegata, Lobophora, 77

variegata, Pocockiella, 77

variegata, Porphyra, 18

variegata, Zonaria, 77

variegatum, Diploderma, 18

variegatus, Gymnosorus, 77

variolosa, Nothogenia, 128

Vaucheria

fastigiata, 118

marina, 104

vaucheriaeformis, Cladophoropsis, 101

vaucheriaeformis, Spongocladia, 101

velasquezii, Acetabularia, 122

velasquezii, Halimeda, 116

velasquezii, Spyridia, 57

velasquezii var. robusta, Halimeda, 116

venezuelensis, Gracilaria, 44

ventricosa, Valonia, 103

venusta, Laurencia, 68

veprecula, Galaxaura, 23

vermicularis, Dasycladus, 120

vermicularis, Spongia, 120

verrucosa, Gracilaria, 44, 123

verrucosus, Fucus, 44

versicolor, Fucus, 26

versicolor, Gelidium, 26

versluysii, Dictyosphaeria, 102

verticillata, Boodleopsis, 117

verticillata, Caulerpa, 111

verticillata, Ernodesmis, 102

verticillata, Valonia, 102

vesiculifera, Caulerpa, 111

vesiculifera, Caulerpa simpliciuscula var., 111

Vidalia, 71

obtusiloba, 71

vieillardii, Champia, 53

vieillardii, Gracilaria, 44

viguieri, Calothrix, 14

villum, Polysiphonia, 70

villum, Polysiphonia scopulorum var., 70

violacea, Leibleinia, 10

vitiense, Sargassum binderi var., 86

vitiense, Sargassum echinocarpum var., 86

vitiensis, Caulerpa urvilliana var., 111

volubilis, Dictyota, 76

vulgare, Sargassum, 84, 88

vulgare var. indicum, Sargassum, 88

vulgaris, Turbinaria, 90

vulgaris var. conoides, Turbinaria, 88

vulgaris var. trialata, Turbinaria, 90

webbiana, Caulerpa, 111

webbiana var. pickeringii, Caulerpa, 111

weberae, Titanophora, 40

wentii, Exophyllum, 63

Wrangelia, 57

argus, 57

bicuspidata, 57

penicillata, 57

wrightiana, Cladophora, 99

wrightii, Anadyomene, 95

wrightii, Halicoryne, 122

Wurdemannia, 72

miniata, 72

wurdemannii, Heterosiphonia, 60

wurdemannii, Dasya, 60

wurdemannii var. laxa, Heterosiphonia, 60

yamadae, Bangia, 18

yamadae, Sebdenia, 40

Yamadaella, 20

caenomyce, 20

yamadana, Laurencia, 68

yendoi, Sargassum, 88

yonakuniense, Spermothamnion, 56

yonakuniensis, Gordoniella, 56

zacae, Ceramium, 56

zacae, Jania tenella var., 35

Zellera, 59

tawallina, 59

zollingeri, Cladophora, 101

zollingeri, Cladophoropsis, 101

Zonaria, 79

diesingiana, 79

flabellata, 79

fraseri, 78

gymnospora, 77, 78

linearis, 76

pavonia var. tenuis, 78

rosea, 37

rubra, 28

variegata, 77

zonata, Amphiroa, 34

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Taxonomic keys in natural history papers should use the aligned-couplet form for zoology and may use the multi-level indent form for botany. If cross referencing is required between key and text, do not include page references within the key, but number the keyed-out taxa, using the same numbers with their corresponding heads in the text.

Synonymy in zoology must use the short form (taxon, author, yeanpage), with full reference at the end of the paper under “Literature Cited.” For botany, the long form (taxon, author, abbreviated journal or book title, volume, page, year, with no reference in “Literature Cited”) is optional.

Text-reference system (author, year:page used within the text, with full citation in “Literature Cited” at the end of the text) must be used in place of bibliographic footnotes in all Contributions Series and is strongly recommended in the Studies Series: “(Jones, 1910:122)” or “…Jones (1910:122).” If bibliographic footnotes are required, use the short form (author, brief title, page) with the full citation in the bibliography.

Footnotes, when few in number, whether annotative or bibliographic, should be typed on separate sheets and inserted immediately after the text pages on which the references occur. Extensive notes must be gathered together and placed at the end of the text in a notes section.

Bibliography, depending upon use, is termed “Literature Cited,” “References,” or “Bibliography.” Spell out titles of books, articles, journals, and monographic series. For book and article titles use sentence-style capitalization according to the rules of the language employed (exception: capitalize all major words in English). For journal and series titles, capitalize the initial word and all subsequent words except articles, conjunctions, and prepositions. Transliterate languages that use a non-Roman alphabet according to the Library of Congress system. Underline (for italics) titles of journals and series and titles of books that are not part of a series. Use the parentheses/colon system for volume (number): pagination: “10(2):5–9.” For alignment and arrangement of elements, follow the format of recent publications in the series for which the manuscript is intended. Guidelines for preparing bibliography may be secured from Series Section, SI Press.

Legends for illustrations must be submitted at the end of the manuscript, with as many legends typed, double-spaced, to a page as convenient.

Illustrations must be submitted as original art (not copies) accompanying, but separate from, the manuscript. Guidelines for preparing art may be secured from Series Section, SI Press. All types of illustrations (photographs, line drawings, maps, etc.) may be intermixed throughout the printed text. They should be termed Figures and should be numbered consecutively as they will appear in the monograph. If several illustrations are treated as components of a single composite figure, they should be designated by lowercase italic letters on the illustration; also, in the legend and in text references the italic letters (underlined in copy) should be used: “Figure 9b.” Illustrations that are intended to follow the printed text may be termed Plates, and any components should be similarly lettered and referenced: “Plate 9b.” Keys to any symbols within an illustration should appear on the art rather than in the legend.

Some points of style: Do not use periods after such abbreviations as “mm, ft, USNM, NNE.” Spell out numbers “one” through “nine” in expository text, but use digits in all other cases if possible. Use of the metric system of measurement is preferable; where use of the English system is unavoidable, supply metric equivalents in parentheses. Use the decimal system for precise measurements and relationships, common fractions for approximations. Use day/month/year sequence for dates: “9 April 1976.” For months in tabular listings or data sections, use three-letter abbreviations with no periods: “Jan, Mar, Jun,” etc. Omit space between initials of a personal name: “J.B. Jones.”

Arrange and paginate sequentially every sheet of manuscript in the following order: (1) title page, (2) abstract, (3) contents, (4) foreword and/or preface, (5) text, (6) appendixes, (7) notes section, (8) glossary, (9) bibliography, (10) legends, (11) tables. Index copy may be submitted at page proof stage, but plans for an index should be indicated when manuscript is submitted.